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data/E8/00/61/E800617AF64870A24A47E0F8723FBE17.xml Normal file
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<document id="AFD6D00ED22B7C7E2859D18938889A70" ID-DOI="10.5281/zenodo.292249" ID-ISBN="3-7643-0556-8" ID-Zenodo-Dep="292249" checkinTime="1646762428550" checkinUser="donat" docAuthor="Hess, Hans Ernst, Landolt, Elias &amp; Hirzel, Rosmarie" docDate="1976" docId="E800617AF64870A24A47E0F8723FBE17" docLanguage="de" docName="unterfamilie_tubuliflorae.xml" docOrigin="Birkhaeuser Verlag" docSource="https://doi.org/10.5281/zenodo.292249" docTitle="Achillea collina Becker" docType="treatment" docVersion="4" lastPageNumber="556" masterDocId="9E9CE376C02228D5F297836DD94BFE82" masterDocTitle="Flora der Schweiz und angrenzender Gebiete. Band 3. Plumbaginaceae bis Compositae (2 nd edition): Unterfamilie _ tubuliflorae" pageNumber="556" updateTime="1669218408346" updateUser="tatiana">
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<mods:title id="921D39CF02278B63E7FA79A008D18FE0">Flora der Schweiz und angrenzender Gebiete. Band 3. Plumbaginaceae bis Compositae (2 nd edition): Unterfamilie _ tubuliflorae</mods:title>
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<mods:namePart id="3D65F3B442D3BDF447B36DAD6DF92BB0">Hess, Hans Ernst</mods:namePart>
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<mods:namePart id="F7E8B74BB7A7A43DE348EA2A053FE6CC">Landolt, Elias</mods:namePart>
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<mods:namePart id="F28C6BE98B874B1BA71874A3DF486870">Hirzel, Rosmarie</mods:namePart>
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<mods:identifier id="724F3A5780269CFDD2216B93ACA505B0" type="DOI">10.5281/zenodo.292249</mods:identifier>
<mods:identifier id="A4ADC6CCBB9715631209F38E17F2C395" type="ISBN">3-7643-0556-8</mods:identifier>
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<treatment id="E800617AF64870A24A47E0F8723FBE17" LSID="urn:lsid:plazi:treatment:E800617AF64870A24A47E0F8723FBE17" httpUri="http://treatment.plazi.org/id/E800617AF64870A24A47E0F8723FBE17" lastPageNumber="556" pageId="null" pageNumber="556" sec="unterfamilie_tubuliflorae.s358">
<title id="682FE17CD3A3BEDB5DCDCD26C30793B3" pageId="null" pageNumber="556">
<subSubSection id="E40BC775BC194036C4864D52D4501E5F" pageId="null" pageNumber="556" type="nomenclature">
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<taxonomicName id="1F426B80D27D216556A4B88B4F275743" authority="Becker" class="Magnoliopsida" family="Asteraceae" genus="Achillea" kingdom="Plantae" order="Asterales" pageId="null" pageNumber="556" phylum="Tracheophyta" rank="species" species="collina">
<pageBreakToken id="D3867850482D72B98B0159EB72FF204A" pageId="null" pageNumber="556" start="start">Achillea</pageBreakToken>
<normalizedToken id="F7F94475088900A3C4D1B4EE9DDA220D" originalValue="collína" pageId="null" pageNumber="556">collina</normalizedToken>
Becker
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</paragraph>
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<subSubSection id="B7FD5B67645B53F67082D94FC7936D5C" pageId="null" pageNumber="556" type="vernacular_names">
<paragraph id="FC7AA882C518FF10D5A8ACA052589BBA" pageId="null" pageNumber="556">
<normalizedToken id="324497F61808E50429E861B7BDF1EFF8" originalValue="Hügel-Schafgarbe" pageId="null" pageNumber="556">Huegel-Schafgarbe</normalizedToken>
</paragraph>
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</title>
<subSubSection id="F2B26E21C85F9E69691980F98C4610B4" pageId="null" pageNumber="556" type="description">
<paragraph id="585009E53D32E63C31281E5861B73C2D" pageId="null" pageNumber="556">
15-60 cm hoch, zerstreut behaart.
<emphasis id="5CCC78EFB59215886790EFFE6533755C" italics="true" pageId="null" pageNumber="556">
<normalizedToken id="5893FEF6AA3D18CF9D6796EBC5129255" originalValue="Grundständige" pageId="null" pageNumber="556">Grundstaendige</normalizedToken>
<normalizedToken id="9FC47371919EF65AFE4C91020F1D4447" originalValue="Blätter" pageId="null" pageNumber="556">Blaetter</normalizedToken>
0,5
</emphasis>
-
<emphasis id="43612352001EF0E715C8FA4ABB5529A3" italics="true" pageId="null" pageNumber="556">1,5 cm breit.</emphasis>
<normalizedToken id="30EBB6B1A3F68E68DDE2796CE251BB50" originalValue="Längs" pageId="null" pageNumber="556">Laengs</normalizedToken>
der obersten 12 cm des Stengels unterhalb der untersten Verzweigung 4-7
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.
<normalizedToken id="6F60A0CA6676AAD67005DA9AD791B063" originalValue="Stengelblätter" pageId="null" pageNumber="556">Stengelblaetter</normalizedToken>
mit nicht
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, nicht
<normalizedToken id="0E80DB4F7158A8F2397229F42B9F7713" originalValue="gezähnter" pageId="null" pageNumber="556">gezaehnter</normalizedToken>
, 0,6-1,2 mm breiter Mittelrippe mit aufgestellten (nicht flach ausgebreiteten), an der Basis 0,3-0,6 mm breiten, bis zum Mittelnerv gezeilten Abschnitten und mit 0,2-0,5 mm breiten, breit lanzettlichen Zipfeln;
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Endzipfel der mittleren
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1
</emphasis>
-
<emphasis id="7E5EA60F50A21BA04BFB7F2148EB058A" italics="true" pageId="null" pageNumber="556">
2mal so lang wie breit.
<normalizedToken id="BE9BDF0F6D509301756E3C2755761A8C" originalValue="Hülle" pageId="null" pageNumber="556">Huelle</normalizedToken>
3
</emphasis>
-
<emphasis id="637D5EBE1CDA9213F598C88E4D4A2E7A" italics="true" pageId="null" pageNumber="556">4 mm lang.</emphasis>
<normalizedToken id="2AABE10E7F6DC262390E48507EE3438F" originalValue="Äußere" pageId="null" pageNumber="556">Aeussere</normalizedToken>
<normalizedToken id="F96FB1595BDD6E250A4AF9C7A468FA96" originalValue="Hüllblätter" pageId="null" pageNumber="556">Huellblaetter</normalizedToken>
0,8-1,1 mm breit.
<normalizedToken id="491651152873A2FC9F687C5966B8271F" originalValue="Zungenförmige" pageId="null" pageNumber="556">Zungenfoermige</normalizedToken>
<normalizedToken id="3594159D9AF43F845109EF2C80F5A826" originalValue="Blüten" pageId="null" pageNumber="556">Blueten</normalizedToken>
<normalizedToken id="9E29048D309606FC7AC3036A06CBE168" originalValue="weiß" pageId="null" pageNumber="556">weiss</normalizedToken>
.
<normalizedToken id="52DB0049BBE64A63049D4BDF811BAA1D" originalValue="Früchte" pageId="null" pageNumber="556">Fruechte</normalizedToken>
1,5-1,8 mm lang. -
<normalizedToken id="FD01288D0BCF72C6BC61C9ECADE6C5BC" originalValue="Blüte" pageId="null" pageNumber="556">Bluete</normalizedToken>
: Sommer und
<normalizedToken id="4C402D322C4BBCBAF4C9F610B1830405" originalValue="früher" pageId="null" pageNumber="556">frueher</normalizedToken>
Herbst.
</paragraph>
<paragraph id="F4A433839660C118D7E148927092F584" pageId="null" pageNumber="556">
<emphasis id="35C3D8C0E27DFB92791994FE9B2CBA2F" italics="true" pageId="null" pageNumber="556">Zytologische Angaben. 2n</emphasis>
=
<emphasis id="F60FCEC135645BB3FAD06188AB3DCFF2" italics="true" pageId="null" pageNumber="556">36:</emphasis>
Material aus Ungarn (
<normalizedToken id="F8CA89C7AD14114F0A76C18FB919EF29" originalValue="Pólya" pageId="null" pageNumber="556">Polya</normalizedToken>
1948), von vielen Orten aus den Alpen, aus Bayern aus dem Burgenland und den
<normalizedToken id="583778EB8880318C8D7020BF92ADF378" originalValue="nördlichen" pageId="null" pageNumber="556">noerdlichen</normalizedToken>
<normalizedToken id="39014CC9A1FB85FC9866DA99F2D58773" originalValue="Balkanländern" pageId="null" pageNumber="556">Balkanlaendern</normalizedToken>
; selten mit 2 B-Chromosomen (Ehrendorfer 1959c 1961), aus der Slowakei (
<normalizedToken id="9EFDE248199CD377B1B170650FCFCA02" originalValue="Háberová" pageId="null" pageNumber="556">Haberova</normalizedToken>
1963), aus Bulgarien (Kuzmanov und
<normalizedToken id="66A7819DBA172AA4EEFCADC4F8C26C80" originalValue="Koźuharov" pageId="null" pageNumber="556">Kozuharov</normalizedToken>
in
<normalizedToken id="AA1A43FAB7B76066B560127863D2D8F5" originalValue="Löve" pageId="null" pageNumber="556">Loeve</normalizedToken>
1970a).
</paragraph>
<paragraph id="43529C75E685A0A6C28366119F45C7F3" pageId="null" pageNumber="556">
<emphasis id="B0FB331C69C92A1AAC897088519D4DF1" italics="true" pageId="null" pageNumber="556">
<pageBreakToken id="5E257B4E2C67A61135DA710F95C3D028" pageId="null" pageNumber="556">Standort</pageBreakToken>
.
</emphasis>
Kollin, montan und subalpin. Trockene, lockere
<normalizedToken id="3091B0BD26C53B64A90F1292C9948159" originalValue="Böden" pageId="null" pageNumber="556">Boeden</normalizedToken>
in warmen Lagen. Lichte
<normalizedToken id="4C7D30BF944A7006F0854BAA1672FEF6" originalValue="Wälder" pageId="null" pageNumber="556">Waelder</normalizedToken>
, Trockenwiesen,
<normalizedToken id="301A77C6BF9C28E84473CE0854B2679A" originalValue="Wegränder" pageId="null" pageNumber="556">Wegraender</normalizedToken>
,
<normalizedToken id="A411652FFA1D9C191972679E6BFBCBC2" originalValue="Schuttplätze" pageId="null" pageNumber="556">Schuttplaetze</normalizedToken>
.
</paragraph>
<paragraph id="81D4C9BEA868D55F70D8FDF6BBFA2A3F" pageId="null" pageNumber="556">
<emphasis id="FB83AF6E0ABE81B7CB6C9A9569F841CC" italics="true" pageId="null" pageNumber="556">
Verbreitung.
<normalizedToken id="E4B23B798D9F1265C62D554FA93A61BE" originalValue="Südosteuropäische" pageId="null" pageNumber="556">Suedosteuropaeische</normalizedToken>
Pflanze.
</emphasis>
Verbreitungskarte von Ehrendorfer (1962b) - Im Gebiet:
<normalizedToken id="766685B3FC15EC98D84EC7E5353B0BC3" originalValue="Elsaß" pageId="null" pageNumber="556">Elsass</normalizedToken>
, zentral- und
<normalizedToken id="2BF742A4E25846C2C09C1C5F3E1EA913" originalValue="südalpine" pageId="null" pageNumber="556">suedalpine</normalizedToken>
<normalizedToken id="A1C0A7FACCCF8927EFE2B5F0F8F3CDA1" originalValue="Täler" pageId="null" pageNumber="556">Taeler</normalizedToken>
, nicht
<normalizedToken id="D1CF8538D4B0833FA51CEA5CDEA08419" originalValue="häufig" pageId="null" pageNumber="556">haeufig</normalizedToken>
.
</paragraph>
<paragraph id="A5F224CB18A28BFB7F70402A659EE5C5" pageId="null" pageNumber="556">
<emphasis id="99D62BD55538AEFA01482C20ABC3D1A6" italics="true" pageId="null" pageNumber="556">
Bemerkungen.
<taxonomicName id="131F5966CE275D4BDF0E56C616D4EE52" class="Magnoliopsida" family="Asteraceae" genus="Achillea" kingdom="Plantae" order="Asterales" pageId="null" pageNumber="556" phylum="Tracheophyta" rank="species" species="collina">A. collina</taxonomicName>
</emphasis>
ist nach Ehrendorfer (1959e) eine tetraploide Bastardsippe, entstanden aus
<taxonomicName id="2899B7F71E7CB43FFB5A09046BA64271" authority="Vent" authorityName="Vent" class="Magnoliopsida" family="Asteraceae" genus="Achillea" kingdom="Plantae" order="Asterales" pageId="null" pageNumber="556" phylum="Tracheophyta" rank="species" species="aspleniifolia">
<emphasis id="CD028E4A534E516CA1967D003CE497DA" italics="true" pageId="null" pageNumber="556">A. aspleniifolia</emphasis>
Vent
</taxonomicName>
, (
<normalizedToken id="B5011F1AE99CD6E8E76576E1DBEE8DB8" originalValue="südosteuropäische" pageId="null" pageNumber="556">suedosteuropaeische</normalizedToken>
Pflanze feuchter, oft salzhaltiger
<normalizedToken id="DB3C112CE55EB7501C5AB7132743B63D" originalValue="Böden" pageId="null" pageNumber="556">Boeden</normalizedToken>
;
<normalizedToken id="38BF2BBA3CF89D3BD6BF8E34AB7D5CAB" originalValue="nächste" pageId="null" pageNumber="556">naechste</normalizedToken>
Fundstellen am Ostalpenrand) und
<taxonomicName id="230EED46418FC6097B8618CC8CD098DF" class="Magnoliopsida" family="Asteraceae" genus="Achillea" kingdom="Plantae" order="Asterales" pageId="null" pageNumber="556" phylum="Tracheophyta" rank="species" species="setacea">
<emphasis id="C4A0653E81F7F452169E6F6698842596" italics="true" pageId="null" pageNumber="556">A. setacea</emphasis>
</taxonomicName>
(Nr. 2a).
</paragraph>
</subSubSection>
</treatment>
</document>

58
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<document id="5399F85FECF1B01B28982954D5220A94" ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e8151" ID-PMC="PMC4867695" ID-Pensoft-Pub="1314-2828-4-8151" ID-PubMed="27226759" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828-4-e8151" ModsDocOrigin="Biodiversity Data Journal 4" ModsDocTitle="Checklist of British and Irish Hymenoptera - Braconidae" checkinTime="1461288934892" checkinUser="pensoft" docAuthor="Broad, Gavin R., Shaw, Mark R. &amp; Godfray, H. Charles J." docDate="2016" docId="E8006F4FAA98C5B84DE0DC1ED95E285D" docLanguage="en" docName="BiodivDatJour 4: e8151" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e8151" docTitle="Aphaereta tenuicornis Nixon 1939" docType="treatment" docVersion="5" lastPageNumber="8151" masterDocId="EC41FFB1FFD9782CFFAEFFF87E2EFF92" masterDocTitle="Checklist of British and Irish Hymenoptera - Braconidae" masterLastPageNumber="8151" masterPageNumber="8151" pageNumber="8151" updateTime="1668123791244" updateUser="ExternalLinkService">
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<treatment id="E8006F4FAA98C5B84DE0DC1ED95E285D" LSID="urn:lsid:plazi:treatment:E8006F4FAA98C5B84DE0DC1ED95E285D" httpUri="http://treatment.plazi.org/id/E8006F4FAA98C5B84DE0DC1ED95E285D" lastPageNumber="8151" pageId="0" pageNumber="8151">
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<taxonomicName id="A8DE69E5D9BDEC5C3D3C9E0D3B1C8DCC" ID-CoL="67KNV" authority="Nixon, 1939" authorityName="Nixon" authorityYear="1939" class="Insecta" family="Braconidae" genus="Aphaereta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphaereta tenuicornis" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="species" species="tenuicornis">Aphaereta tenuicornis Nixon, 1939</taxonomicName>
</paragraph>
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<subSubSection id="10256F4CEC51CBD07B94ECB8A9FFC9C9" pageId="0" pageNumber="8151" type="distribution">
<paragraph id="62C4530FEDD48FAA21AF62C93579D782" pageId="0" pageNumber="8151">Distribution</paragraph>
<paragraph id="43B77A8C21A40DA1156198FE259B7522" pageId="0" pageNumber="8151">England, Wales, Ireland</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/jor.31.84135" ID-GBIF-Dataset="f5afe7be-ec45-4ecd-b191-51f023aa26b4" ID-Pensoft-Pub="1937-2426-2-121" ID-Pensoft-UUID="7B47AED867BB54ED937582690C95D352" ID-ZooBank="280146F3CB024B86A05ED9DEA53D5FC6" ModsDocID="1937-2426-31-2-121" checkinTime="1661316485122" checkinUser="pensoft" docAuthor="Campos, Lucas Denadai de &amp; de Mello, Francisco de Assis Ganeo" docDate="2022" docId="E80100A238D25411B5705F51AB977FB8" docLanguage="en" docName="JourOrthoptRes 31(2): 121-141" docOrigin="Journal of Orthoptera Research 31 (2)" docPubDate="2022-08-23" docSource="http://dx.doi.org/10.3897/jor.31.84135" docTitle="Amblyrhethus alagoensis Campos &amp; de Mello 2022, sp. nov." docType="treatment" docUuid="4FDF095B-58E9-42F2-9486-55B7F8708715" docUuidSource="ZooBank" docVersion="5" id="7B47AED867BB54ED937582690C95D352" lastPageNumber="121" masterDocId="7B47AED867BB54ED937582690C95D352" masterDocTitle="Three new species of Amblyrhethus (Orthoptera, Grylloidea, Gryllidae, Paroecanthini) from Brazil" masterLastPageNumber="141" masterPageNumber="121" pageNumber="121" updateTime="1661320993016" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Three new species of Amblyrhethus (Orthoptera, Grylloidea, Gryllidae, Paroecanthini) from Brazil</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Campos, Lucas Denadai de</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2922-1571</mods:nameIdentifier>
<mods:affiliation>Universidade de Sao Paulo, Sao Paulo, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">lcdenadai@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>de Mello, Francisco de Assis Ganeo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9704-5990</mods:nameIdentifier>
<mods:affiliation>Instituto de Biociencias, UNESP, Botucatu, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Journal of Orthoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-23</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>31</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:start>121</mods:start>
<mods:end>141</mods:end>
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<mods:url>http://dx.doi.org/10.3897/jor.31.84135</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jor.31.84135</mods:identifier>
<mods:identifier type="Pensoft-Pub">1937-2426-2-121</mods:identifier>
<mods:identifier type="ZooBank">280146F3CB024B86A05ED9DEA53D5FC6</mods:identifier>
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<subSection pageId="0" pageNumber="121" type="results">
<treatment ID-GBIF-Taxon="199726745" LSID="urn:lsid:zoobank.org:act:4FDF095B-58E9-42F2-9486-55B7F8708715" httpUri="http://treatment.plazi.org/id/E80100A238D25411B5705F51AB977FB8" lastPageNumber="121" pageId="0" pageNumber="121">
<subSubSection pageId="0" pageNumber="121" type="nomenclature">
<paragraph pageId="0" pageNumber="121">
<taxonomicName LSID="https://zoobank.org/4FDF095B-58E9-42F2-9486-55B7F8708715" authority="Campos &amp; de Mello, 2022" authorityName="Campos &amp; de Mello" authorityYear="2022" class="Insecta" family="Gryllidae" genus="Amblyrhethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyrhethus alagoensis" order="Orthoptera" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="alagoensis" status="sp. nov.">Amblyrhethus alagoensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="121">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="description">
<paragraph pageId="0" pageNumber="121">
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">(Figs 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Amblyrhethus alagoensis sp. nov. Male genitalia: A. Dorsal; B. Ventral; C. Lateral; D. Posterior. Female copulatory papilla: E. Dorsal; F. Ventral; G. Lateral. Scales 1 mm. Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure6" httpUri="https://binary.pensoft.net/fig/734164" pageId="0" pageNumber="121">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Map of the known distribution of Amblyrhethus species." figureDoi="10.3897/jor.31.84135.figure7" httpUri="https://binary.pensoft.net/fig/734165" pageId="0" pageNumber="121">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="materials_examined">
<paragraph pageId="0" pageNumber="121">
<emphasis italics="true" pageId="0" pageNumber="121">Material examined.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="121">
<materialsCitation collectingDate="2002-09-08" collectingDateMax="2002-09-11" collectingDateMin="2002-09-08" country="BRAZIL" county="Bosque" location="Quebrangulo" municipality="Malaise" specimenCount="♂" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="121">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
<collectingCountry name="Brazil">BRAZIL</collectingCountry>
•
<specimenCount>♂</specimenCount>
; AL[agoas]
<location LSID="urn:lsid:plazi:treatment:E80100A238D25411B5705F51AB977FB8:B5FD1633A4799C69963BF8212FE2BB8E" country="BRAZIL" county="Bosque" municipality="Malaise" name="Quebrangulo">Quebrangulo</location>
/
<location LSID="urn:lsid:plazi:treatment:E80100A238D25411B5705F51AB977FB8:C26CE2F94918CEE8B28DE8371EB604F4" country="BRAZIL" county="Bosque" municipality="Malaise" name="Res">Res</location>
[erva]
<location LSID="urn:lsid:plazi:treatment:E80100A238D25411B5705F51AB977FB8:072162AC2C118C249739EB5FE6C4D090" country="BRAZIL" county="Bosque" municipality="Malaise" name="Biol">Biol</location>
[
<normalizedToken originalValue="ógica">ogica</normalizedToken>
]
<location LSID="urn:lsid:plazi:treatment:E80100A238D25411B5705F51AB977FB8:B1C60F75CEFB41441C8886D45AD91C04" country="BRAZIL" county="Bosque" municipality="Malaise" name="Pedra Talhada">Pedra Talhada</location>
/
<location LSID="urn:lsid:plazi:treatment:E80100A238D25411B5705F51AB977FB8:2CCC3451C3BFF20CE43D68D7D16E4335" country="BRAZIL" county="Bosque" municipality="Malaise" name="Arm">Arm</location>
[adilha];
<collectingMunicipality>Malaise</collectingMunicipality>
- 1°
<collectingCounty>Bosque</collectingCounty>
;
<collectingDate value="2002-09-08" valueMax="2002-09-11" valueMin="2002-09-08">08-11 September 2002</collectingDate>
; Refugos; Penteado-Dias &amp; eq[uipe] leg; MZSP
</materialsCitation>
.
<materialsCitation collectingDate="2012-07-25" collectorName="Dias, P. G. B. S, Costa, C. S., Alcantra, D. M. C., Nihei S. S." country="BRAZIL" location="Area de Protecao Ambiental do Catole" municipality="Satuba" specimenCount="♀" stateProvince="Alagoas" typeStatus="Allotype">
<emphasis bold="true" pageId="0" pageNumber="121">
<typeStatus>Allotype</typeStatus>
</emphasis>
:
<collectingCountry name="Brazil">BRAZIL</collectingCountry>
•
<specimenCount>♀</specimenCount>
;
<collectingMunicipality>Satuba</collectingMunicipality>
,
<collectingRegion country="Brazil" name="Alagoas">Alagoas</collectingRegion>
, Brasil /
<location LSID="urn:lsid:plazi:treatment:E80100A238D25411B5705F51AB977FB8:2A3347DE32979B153A948C0F9F72EF96" country="BRAZIL" municipality="Satuba" name="Area de Protecao Ambiental do Catole" stateProvince="Alagoas">
<normalizedToken originalValue="Área">Area</normalizedToken>
de
<normalizedToken originalValue="Proteção">Protecao</normalizedToken>
Ambiental do
<normalizedToken originalValue="Catolé">Catole</normalizedToken>
</location>
;
<collectingDate value="2012-07-25">25 July 2012</collectingDate>
;
<collectorName>Dias, P.G.B.S</collectorName>
,
<collectorName>Costa, C.S.</collectorName>
,
<collectorName>
<normalizedToken originalValue="Alcântra">Alcantra</normalizedToken>
, D.M.C.
</collectorName>
;
<collectorName>Nihei S.S.</collectorName>
leg.; LDC_014; MZSP
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="type locality">
<paragraph pageId="0" pageNumber="121">
<emphasis italics="true" pageId="0" pageNumber="121">Type locality.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="121">
Brazil, Alagoas,
<normalizedToken originalValue="Quebrângulo">Quebrangulo</normalizedToken>
and Satuba municipalities.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="etymology">
<paragraph pageId="0" pageNumber="121">
<emphasis italics="true" pageId="0" pageNumber="121">Etymology.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="121">Toponymic, referring to the State of Alagoas, Brazil.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="diagnosis">
<paragraph pageId="0" pageNumber="121">
<emphasis italics="true" pageId="0" pageNumber="121">Diagnosis.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="121">
This species is distinguished from other species of
<taxonomicName authorityName="Kirby" authorityYear="1906" class="Insecta" family="Gryllidae" genus="Amblyrhethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyrhethus" order="Orthoptera" pageId="0" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="121">Amblyrhethus</emphasis>
</taxonomicName>
by the following characters: male forewings vein A1 not connected to A2; metanotum with two clusters of bristles on the middle; first abdominal tergite with two rounded projections close to anterior margin. Male genitalia: pseudepiphallic sclerite somewhat enlarged on anterior half, especially at the middle level, rounded in dorsal and ventral views; PsP clavate, EctAp slightly curved inwards in the middle, both in dorsal and ventral views. Copulatory papilla tapered, with two anterior projections.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="description">
<paragraph pageId="0" pageNumber="121">
<emphasis italics="true" pageId="0" pageNumber="121">Description.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="121">
Head: Fastigium wide, with bristles laterally (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5E</figureCitation>
). Three ocelli present, aligned in frontal view (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5E</figureCitation>
); lateral ocelli rounded, median ocellus elliptical, smaller than lateral ones; frons smooth (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5E</figureCitation>
). Antennal scape longer than wide, narrower than fastigium in frontal view. Maxillary palpi article 3 the longest, 4 and 5 almost same-sized; article 5 straight (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5B, D</figureCitation>
). Thorax: Dorsal disk wider than long, covered by fine pubescence (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5A, C</figureCitation>
). Dorsal disk cephalic margin slightly concave, caudal margin convex (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5A, C</figureCitation>
). Lateral lobes ventro-cephalic and ventro-caudal angles rounded (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5B, D</figureCitation>
). Forewings covering abdomen, hindwings slightly longer than forewings (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5A-D</figureCitation>
). Legs: TI with tympana present on inner and outer faces; three apical spurs, two ventral, one dorsal. TII with four apical spurs, two ventral, two dorsal. TIII subapical spurs 5/4, with two spines between each spur, except proximal spurs with three spines; eight spines above subapical spurs on inner and outer sides. FIII longer than TIII (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5B, D</figureCitation>
). TIII inner apical spurs: iad&gt;iam&gt;iav; outer apical spurs: oam&gt;oav&gt;oad. Basitarsus dorsal spines 3/1; inner apical spur slightly longer than outer apical spur.
</paragraph>
<paragraph pageId="0" pageNumber="121">
<emphasis bold="true" pageId="0" pageNumber="121">Male.</emphasis>
Forewings slightly shorter than hindwings; anal area slightly bulged dorsally (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5B</figureCitation>
), A1 not connected to A2. Stridulatory vein (PCu vein) present, portion close to CuPa sinuous. Harp crossed by four veins connected to CuPa; first and second harp veins shorter than third and fourth; third and fourth harp veins connected in the proximal region. Mirror as wide as long, divided in the middle by a curved vein. Apical field longer than mirror; lateral field with 13 diagonal veins (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5F</figureCitation>
). Metanotum with two clusters of bristles on the middle, first abdominal tergite with two rounded projections close to anterior margin (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5G</figureCitation>
). Supra-anal plate posterior margin rounded (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5H</figureCitation>
); subgenital plate longer than wide, posterior margin convex (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5I</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="121">
<emphasis bold="true" italics="true" pageId="0" pageNumber="121">Male genitalia</emphasis>
: (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Amblyrhethus alagoensis sp. nov. Male genitalia: A. Dorsal; B. Ventral; C. Lateral; D. Posterior. Female copulatory papilla: E. Dorsal; F. Ventral; G. Lateral. Scales 1 mm. Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure6" httpUri="https://binary.pensoft.net/fig/734164" pageId="0" pageNumber="121">6</figureCitation>
) Pseudepiphallus: pseudepiphallic sclerite trapezoidal in dorsal and ventral views, anterior half inflated; slightly curved upwards in lateral view; anterior margin rounded on median region (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Amblyrhethus alagoensis sp. nov. Male genitalia: A. Dorsal; B. Ventral; C. Lateral; D. Posterior. Female copulatory papilla: E. Dorsal; F. Ventral; G. Lateral. Scales 1 mm. Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure6" httpUri="https://binary.pensoft.net/fig/734164" pageId="0" pageNumber="121">6A</figureCitation>
). LLophi curved upwards, posterior margin sub-straight in dorsal and ventral views; ventral face translucent (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Amblyrhethus alagoensis sp. nov. Male genitalia: A. Dorsal; B. Ventral; C. Lateral; D. Posterior. Female copulatory papilla: E. Dorsal; F. Ventral; G. Lateral. Scales 1 mm. Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure6" httpUri="https://binary.pensoft.net/fig/734164" pageId="0" pageNumber="121">6A, B</figureCitation>
). PsP clavate, longer than LLophi, not surpassing posterior margin of pseudepiphallic sclerite in dorsal and ventral views, inner margin with anterior projection finger-shaped (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Amblyrhethus alagoensis sp. nov. Male genitalia: A. Dorsal; B. Ventral; C. Lateral; D. Posterior. Female copulatory papilla: E. Dorsal; F. Ventral; G. Lateral. Scales 1 mm. Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure6" httpUri="https://binary.pensoft.net/fig/734164" pageId="0" pageNumber="121">6A, B</figureCitation>
). Rami straight in dorsal and ventral views, two times longer than pseudepiphallic sclerite (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Amblyrhethus alagoensis sp. nov. Male genitalia: A. Dorsal; B. Ventral; C. Lateral; D. Posterior. Female copulatory papilla: E. Dorsal; F. Ventral; G. Lateral. Scales 1 mm. Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure6" httpUri="https://binary.pensoft.net/fig/734164" pageId="0" pageNumber="121">6A-C</figureCitation>
). Ectophallic invagination: EctF strongly sclerotized, shorter than PsP, trapezoidal on the edge of posterior margin of pseudepiphallic sclerite (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Amblyrhethus alagoensis sp. nov. Male genitalia: A. Dorsal; B. Ventral; C. Lateral; D. Posterior. Female copulatory papilla: E. Dorsal; F. Ventral; G. Lateral. Scales 1 mm. Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure6" httpUri="https://binary.pensoft.net/fig/734164" pageId="0" pageNumber="121">6A-D</figureCitation>
). EctAp longer than pseudepiphallic sclerite, slightly curved inwards in the middle in dorsal and ventral views (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Amblyrhethus alagoensis sp. nov. Male genitalia: A. Dorsal; B. Ventral; C. Lateral; D. Posterior. Female copulatory papilla: E. Dorsal; F. Ventral; G. Lateral. Scales 1 mm. Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure6" httpUri="https://binary.pensoft.net/fig/734164" pageId="0" pageNumber="121">6A, B</figureCitation>
); arc not complete, curved posteriorly; ventral projections of ectophallic invagination shorter than arc. Endophallus: End short, u-shaped, in the middle of pseudepiphallic sclerite (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Amblyrhethus alagoensis sp. nov. Male genitalia: A. Dorsal; B. Ventral; C. Lateral; D. Posterior. Female copulatory papilla: E. Dorsal; F. Ventral; G. Lateral. Scales 1 mm. Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure6" httpUri="https://binary.pensoft.net/fig/734164" pageId="0" pageNumber="121">6B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="121">
<emphasis bold="true" pageId="0" pageNumber="121">Female.</emphasis>
Larger than male, similar coloration (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5C, D</figureCitation>
). Forewings with longitudinal veins crossed by accessory veins. Subgenital plate wider than long, posterior margin concave (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5J</figureCitation>
). Ovipositor slightly inclined upwards; apex of ovipositor lateral margins serrulate, posterior margin truncate (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5C, J</figureCitation>
). Copulatory papilla bell-shaped, with two posterior thin projections directed ventrally (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Amblyrhethus alagoensis sp. nov. Male genitalia: A. Dorsal; B. Ventral; C. Lateral; D. Posterior. Female copulatory papilla: E. Dorsal; F. Ventral; G. Lateral. Scales 1 mm. Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure6" httpUri="https://binary.pensoft.net/fig/734164" pageId="0" pageNumber="121">6E-G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="coloration.">
<paragraph pageId="0" pageNumber="121">
<emphasis italics="true" pageId="0" pageNumber="121">Coloration.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="121">
Occiput, vertex, and pronotum general coloration reddish brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5A-D</figureCitation>
). Face reddish-brown to medium brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5E</figureCitation>
); antennal scape medium brown; antennomeres yellowish with some isolated antennomeres medium brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5A-D</figureCitation>
). Forewings slightly translucent, medium brown. Metanotum light brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5G</figureCitation>
), first three abdominal tergites medium brown, the other tergites dark brown. Supra-anal plate dark brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5H</figureCitation>
). Abdominal sternites yellowish brown; subgenital plate medium brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5I</figureCitation>
). FI, FII, TI, and TII medium brown to reddish brown. FIII reddish brown, slightly striped, distal margin darker; TIII medium brown; spurs yellow with apex dark brown; tarsomeres yellowish-brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Amblyrhethus alagoensis sp. nov. A. Male habitus, dorsal; B. Male habitus, lateral; C. Female habitus, dorsal; D. Female habitus, lateral; E. Head, frontal; F. Male right forewing; G. Male metanotum and first abdominal tergite; H. Male supra-anal plate; I. Male subgenital plate; J. Female subgenital plate and ovipositor. Scale bars 5 mm (A-D); 1 mm (E-J). Abbreviations: see material and methods." figureDoi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121">5C</figureCitation>
).
</paragraph>
<caption doi="10.3897/jor.31.84135.figure5" httpUri="https://binary.pensoft.net/fig/734163" pageId="0" pageNumber="121" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="121">
<emphasis bold="true" pageId="0" pageNumber="121">Figure 5.</emphasis>
<taxonomicName authorityName="Campos &amp; de Mello" authorityYear="2022" class="Insecta" family="Gryllidae" genus="Amblyrhethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyrhethus alagoensis" order="Orthoptera" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="alagoensis">
<emphasis italics="true" pageId="0" pageNumber="121">Amblyrhethus alagoensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="121">A</emphasis>
. Male habitus, dorsal;
<emphasis bold="true" pageId="0" pageNumber="121">B</emphasis>
. Male habitus, lateral;
<emphasis bold="true" pageId="0" pageNumber="121">C</emphasis>
. Female habitus, dorsal;
<emphasis bold="true" pageId="0" pageNumber="121">D</emphasis>
. Female habitus, lateral;
<emphasis bold="true" pageId="0" pageNumber="121">E</emphasis>
. Head, frontal;
<emphasis bold="true" pageId="0" pageNumber="121">F</emphasis>
. Male right forewing;
<emphasis bold="true" pageId="0" pageNumber="121">G</emphasis>
. Male metanotum and first abdominal tergite;
<emphasis bold="true" pageId="0" pageNumber="121">H</emphasis>
. Male supra-anal plate;
<emphasis bold="true" pageId="0" pageNumber="121">I</emphasis>
. Male subgenital plate;
<emphasis bold="true" pageId="0" pageNumber="121">J</emphasis>
. Female subgenital plate and ovipositor. Scale bars 5 mm (
<emphasis bold="true" pageId="0" pageNumber="121">A-D</emphasis>
); 1 mm (
<emphasis bold="true" pageId="0" pageNumber="121">E-J</emphasis>
). Abbreviations: see material and methods.
</paragraph>
</caption>
<caption doi="10.3897/jor.31.84135.figure6" httpUri="https://binary.pensoft.net/fig/734164" pageId="0" pageNumber="121" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="121">
<emphasis bold="true" pageId="0" pageNumber="121">Figure 6.</emphasis>
<taxonomicName authorityName="Campos &amp; de Mello" authorityYear="2022" class="Insecta" family="Gryllidae" genus="Amblyrhethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyrhethus alagoensis" order="Orthoptera" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="alagoensis">
<emphasis italics="true" pageId="0" pageNumber="121">Amblyrhethus alagoensis</emphasis>
</taxonomicName>
sp. nov. Male genitalia:
<emphasis bold="true" pageId="0" pageNumber="121">A</emphasis>
. Dorsal;
<emphasis bold="true" pageId="0" pageNumber="121">B</emphasis>
. Ventral;
<emphasis bold="true" pageId="0" pageNumber="121">C</emphasis>
. Lateral;
<emphasis bold="true" pageId="0" pageNumber="121">D</emphasis>
. Posterior. Female copulatory papilla:
<emphasis bold="true" pageId="0" pageNumber="121">E</emphasis>
. Dorsal;
<emphasis bold="true" pageId="0" pageNumber="121">F</emphasis>
. Ventral;
<emphasis bold="true" pageId="0" pageNumber="121">G</emphasis>
. Lateral. Scales 1 mm. Abbreviations: see material and methods.
</paragraph>
</caption>
<caption doi="10.3897/jor.31.84135.figure7" httpUri="https://binary.pensoft.net/fig/734165" pageId="0" pageNumber="121" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="121">
<emphasis bold="true" pageId="0" pageNumber="121">Figure 7.</emphasis>
Map of the known distribution of
<taxonomicName authorityName="Kirby" authorityYear="1906" class="Insecta" family="Gryllidae" genus="Amblyrhethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyrhethus" order="Orthoptera" pageId="0" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="121">Amblyrhethus</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<caption ID-Table-UUID="117550F830A285148BF0DD6C6F81AB18" httpUri="http://table.plazi.org/id/117550F830A285148BF0DD6C6F81AB18" pageId="0" pageNumber="121" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="121">
<emphasis bold="true" pageId="0" pageNumber="121">Table 3.</emphasis>
Measurements in mm of
<taxonomicName authorityName="Campos &amp; de Mello" authorityYear="2022" class="Insecta" family="Gryllidae" genus="Amblyrhethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyrhethus alagoensis" order="Orthoptera" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="alagoensis">
<emphasis italics="true" pageId="0" pageNumber="121">Amblyrhethus alagoensis</emphasis>
</taxonomicName>
sp. nov. Abbreviations:
<emphasis bold="true" pageId="0" pageNumber="121">IOD</emphasis>
-inter ocular distance;
<emphasis bold="true" pageId="0" pageNumber="121">HW</emphasis>
-head width;
<emphasis bold="true" pageId="0" pageNumber="121">PL</emphasis>
, pronotum length;
<emphasis bold="true" pageId="0" pageNumber="121">PW</emphasis>
, pronotum width (at midline);
<emphasis bold="true" pageId="0" pageNumber="121">FWL</emphasis>
, forewing length;
<emphasis bold="true" pageId="0" pageNumber="121">LFIII</emphasis>
, length of hind femur;
<emphasis bold="true" pageId="0" pageNumber="121">LTIII</emphasis>
, length of hind tibia;
<emphasis bold="true" pageId="0" pageNumber="121">OL</emphasis>
, ovipositor length.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="121">
<table pageId="0" pageNumber="121">
<tr pageId="0" pageNumber="121">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="121" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">IOD</th>
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">HW</th>
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">PL</th>
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">PW</th>
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">FWL</th>
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">LFIII</th>
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">LTIII</th>
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">OL</th>
</tr>
<tr pageId="0" pageNumber="121">
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Female (n=1)</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5.5</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4.1</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">18.1</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">13.1</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">11</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">13</td>
</tr>
<tr pageId="0" pageNumber="121">
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Male (n=1)</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2.1</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4.2</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">3.6</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4.8</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">15.1</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">-</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>

119
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<treatment id="E8013DF3B944E096D518E18C6535D8EA" LSID="urn:lsid:plazi:treatment:E8013DF3B944E096D518E18C6535D8EA" httpUri="http://treatment.plazi.org/id/E8013DF3B944E096D518E18C6535D8EA" lastPageNumber="613" pageId="null" pageNumber="613" sec="leguminosae.s307">
<title id="2BB24C9CC092C4FD82866F916A682763" pageId="null" pageNumber="613">
<subSubSection id="3875E8E7842FCCAD3083E2A2A5704698" pageId="null" pageNumber="613" type="nomenclature">
<paragraph id="6E6C07230070BFF3D172096DE7A1CD0C" pageId="null" pageNumber="613">
<taxonomicName id="CCD0BBE7307B5E052885EE1425E0793F" authority="(Thunb.) C. K. Schneider" authorityName="C. K. Schneider" baseAuthorityName="Thunb." class="Magnoliopsida" family="Fabaceae" genus="Pueraria" kingdom="Plantae" order="Fabales" pageId="null" pageNumber="613" phylum="Tracheophyta" rank="species" species="hirsuta">
<pageBreakToken id="2D2147DF4F554FFEB908FDF6CCD286F1" pageId="null" pageNumber="613">Pueraria</pageBreakToken>
<normalizedToken id="68E190E14584E37676623DECDC1DADE0" originalValue="hirsúta" pageId="null" pageNumber="613">hirsuta</normalizedToken>
(Thunb.) C. K. Schneider
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="2BE970308A7BFC6CAF914EFC097642BB" pageId="null" pageNumber="613" type="reference_group">
<paragraph id="90E49814AE3B32708019C5C9ADAB7B6E" pageId="null" pageNumber="613">
(
<taxonomicName id="2874351A14FAD94587CC9D31A6FF2812" authority="Benth." authorityName="Benth." class="Magnoliopsida" family="Fabaceae" genus="Pueraria" kingdom="Plantae" order="Fabales" pageId="null" pageNumber="613" phylum="Tracheophyta" rank="species" species="thunbergiana">
<emphasis id="B714D9AF2BB3C962C9DE5AFBAD6B4A18" italics="true" pageId="null" pageNumber="613">P. Thunbergiana</emphasis>
Benth.
</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="C8B7B4EF81F419155B1BD371CEA1D372" pageId="null" pageNumber="613" type="vernacular_names">
<paragraph id="34B58FCAC2419B26F6465744E8F07FE4" pageId="null" pageNumber="613">Behaarte Kopoubohne</paragraph>
</subSubSection>
</title>
<subSubSection id="CF8E8DE177E9E43CA4EA7D3305612ABC" pageId="null" pageNumber="613" type="description">
<paragraph id="D38C39C5EE2FCC6F6A28A732629D3B85" pageId="null" pageNumber="613">
Ausdauernd, mit verholztem, windendem Stengel und
<emphasis id="FC50F6703A0DDEBC444CD91EDB87C495" italics="true" pageId="null" pageNumber="613">fleischiger knolliger Wurzel; bis 20 m lang.</emphasis>
Blattstiel gerillt, wie die jungen Stengel mit zahlreichen 0,1-0,3 mm langen und einzelnen bis 2 mm langen Haaren;
<normalizedToken id="ECD08A15AD04EC42E931752F1597B675" originalValue="Teilblätter" pageId="null" pageNumber="613">Teilblaetter</normalizedToken>
bis 15 cm lang, besonders unterseits mit 0,5-1 mm langen Haaren, oft 1-2mal wenig tief eingeschnitten.
<emphasis id="3A5A18C684BC059158C6F8C18C561B84" italics="true" pageId="null" pageNumber="613">
<normalizedToken id="62908F65A8617064C02DE2212092A75B" originalValue="Blütenstand" pageId="null" pageNumber="613">Bluetenstand</normalizedToken>
bis 25 cm lang, 20-50
<normalizedToken id="058B31F0AA03CC892F547E6D4B2405D3" originalValue="blütig" pageId="null" pageNumber="613">bluetig</normalizedToken>
.
</emphasis>
Kelch 0,7-1 cm lang, behaart.
<emphasis id="F883BA270F6F7EE052017CC1746B3DAC" italics="true" pageId="null" pageNumber="613">Krone 1,5-2 cm lang</emphasis>
, purpurn bis violett. Frucht flach, 4-10 cm lang und 0,7-1,2 cm breit, dicht und braun behaart, mit kurzen und bis 3 mm langen Haaren. -
<normalizedToken id="28B535FE1B36A5A5B49FCA393ABF2B37" originalValue="Blüte" pageId="null" pageNumber="613">Bluete</normalizedToken>
:
<normalizedToken id="D15C9F83EBE54D7E1C5E6887A3998F92" originalValue="Später" pageId="null" pageNumber="613">Spaeter</normalizedToken>
Sommer.
</paragraph>
<paragraph id="3B04FA04CC9F5AF42164A7214C26713A" pageId="null" pageNumber="613">
<emphasis id="15829AB0FF9CA04D89226544AF76931D" italics="true" pageId="null" pageNumber="613">Zytologische Angaben. 2n</emphasis>
=
<emphasis id="81562502203FE400D616C954819B9768" italics="true" pageId="null" pageNumber="613">22:</emphasis>
Material aus botanischen
<normalizedToken id="084AADA777C1302721772858F051FBB0" originalValue="Gärten" pageId="null" pageNumber="613">Gaerten</normalizedToken>
(Simmonds 1954).
<emphasis id="188A68B144A89FB29AAFAE9953047C45" italics="true" pageId="null" pageNumber="613">2n</emphasis>
=
<emphasis id="10974248028D023CF795B895755F94B9" italics="true" pageId="null" pageNumber="613">24:</emphasis>
Material aus Japan (Suzuka 1950, Sakai 1951).
</paragraph>
<paragraph id="8DBDC33034DF1BC2B7B8B1E530A2C400" pageId="null" pageNumber="613">
<emphasis id="79FCACD114A1F9A55C4DBDBB797BF2A5" italics="true" pageId="null" pageNumber="613">Standort.</emphasis>
Kollin, ziemlich feuchte, lockere
<normalizedToken id="4986CA96C662D2AA9D80630D611CD9F2" originalValue="Böden" pageId="null" pageNumber="613">Boeden</normalizedToken>
in milden, luftfeuchten Lagen. Gartenmauern,
<normalizedToken id="F34790CE6AE62BF33536F4A5C9BC9320" originalValue="Gebüsch" pageId="null" pageNumber="613">Gebuesch</normalizedToken>
, Schuttstellen.
</paragraph>
<paragraph id="7317436D723589160E8C739EB7DD85D7" pageId="null" pageNumber="613">
<emphasis id="0F8689581610C1CC8897390E3ADDC595" italics="true" pageId="null" pageNumber="613">Verbreitung.</emphasis>
<normalizedToken id="30648309C60833E6023AC9D9E0C98DD3" originalValue="Ursprünglich" pageId="null" pageNumber="613">Urspruenglich</normalizedToken>
<emphasis id="4E7FB5CA9CA86C811407D5FF8A76F4F1" italics="true" pageId="null" pageNumber="613">ostasiatische Pflanze;</emphasis>
in den
<normalizedToken id="682C5ECAAA748FA3B4968AD23E411E3C" originalValue="wärmeren" pageId="null" pageNumber="613">waermeren</normalizedToken>
Gebieten der ganzen Erde als Zierpflanze, Faserpflanze und Futterpflanze angebaut (die Knollen werden auch als
<normalizedToken id="71990BCC00B3B9B202B70DE549142BAD" originalValue="Wurzelgemüse" pageId="null" pageNumber="613">Wurzelgemuese</normalizedToken>
gegessen) und vielerorts
<normalizedToken id="6B02A93BEB44D28598672CB69681F01F" originalValue="eingebürgert" pageId="null" pageNumber="613">eingebuergert</normalizedToken>
. - Im Gebiet im
<normalizedToken id="223E8FCEB5EACDEF187C0B45AA18EB1C" originalValue="südlichen" pageId="null" pageNumber="613">suedlichen</normalizedToken>
Tessin oft angepflanzt und selten verwildert.
</paragraph>
</subSubSection>
</treatment>
</document>

346
data/E8/01/59/E801591D3B9E599C811E9F24FC35CAF4.xml Normal file
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@ -0,0 +1,346 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.915.46390" ID-GBIF-Dataset="87bb16b9-c7f7-4da5-96ab-082ebecd61c0" ID-PMC="PMC7052024" ID-Pensoft-Pub="1313-2970-915-107" ID-Pensoft-UUID="4ED8C22188635B28B9229C890ACEB8C1" ID-PubMed="32148425" ID-ZooBank="6245285F7A744277AD4FE81B501CDE03" ModsDocID="1313-2970-915-107" checkinTime="1582662224405" checkinUser="pensoft" docAuthor="Khormizi, Mehdi Zare &amp; Nedved, Oldrich" docDate="2020" docId="E801591D3B9E599C811E9F24FC35CAF4" docLanguage="en" docName="ZooKeys 915: 107-116" docOrigin="ZooKeys 915" docSource="http://dx.doi.org/10.3897/zookeys.915.46390" docTitle="Oenopia shirkuhensis Zare Khormizi &amp; Nedved 2020, sp. nov." docType="treatment" docUuid="69FF66FE-C039-496B-BD62-F50A564F9A51" docUuidSource="ZooBank" docVersion="4" id="4ED8C22188635B28B9229C890ACEB8C1" lastPageNumber="107" masterDocId="4ED8C22188635B28B9229C890ACEB8C1" masterDocTitle="Oenopia shirkuhensis sp. nov. (Coleoptera, Coccinellidae) from Iran mimicking Adalia bipunctata" masterLastPageNumber="116" masterPageNumber="107" pageNumber="107" updateTime="1668168511895" updateUser="ExternalLinkService">
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<mods:title>Oenopia shirkuhensis sp. nov. (Coleoptera, Coccinellidae) from Iran mimicking Adalia bipunctata</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Khormizi, Mehdi Zare</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nedved, Oldrich</mods:namePart>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>915</mods:number>
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<mods:start>107</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.915.46390</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.915.46390</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-915-107</mods:identifier>
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<treatment ID-GBIF-Taxon="162267211" LSID="urn:lsid:zoobank.org:act:69FF66FE-C039-496B-BD62-F50A564F9A51" httpUri="http://treatment.plazi.org/id/E801591D3B9E599C811E9F24FC35CAF4" lastPageNumber="107" pageId="0" pageNumber="107">
<subSubSection pageId="0" pageNumber="107" type="nomenclature">
<paragraph pageId="0" pageNumber="107">
<taxonomicName LSID="E801591D-3B9E-599C-811E-9F24FC35CAF4" authority="Zare Khormizi &amp; Nedvěd" authorityName="Zare Khormizi &amp; Nedved" authorityYear="2020" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenopia shirkuhensis" order="Coleoptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="shirkuhensis" status="sp. nov.">
Oenopia shirkuhensis Zare Khormizi &amp;
<normalizedToken originalValue="Nedvěd">Nedved</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="107">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="type material">
<paragraph pageId="0" pageNumber="107">Type material.</paragraph>
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Holotype.</emphasis>
Male, Sorgan valley, Shirkooh mountain, Seikhalishah parish Dehbala village, Taft region, Yazd province, Iran,
<geoCoordinate degrees="31" direction="north" minutes="37.50" orientation="latitude" precision="9" value="31.625">31°37.50'N</geoCoordinate>
,
<geoCoordinate degrees="54" direction="east" minutes="05.68" orientation="longitude" precision="9" value="54.094666">54°05.68'E</geoCoordinate>
, 2960 m a.s.l. 16. v. 2014. On
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Rosa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rosa canina" order="Rosales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="canina">
<emphasis italics="true" pageId="0" pageNumber="107">Rosa canina</emphasis>
</taxonomicName>
. Lgt. Mehdi Zare Khormizi, Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">2</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Paratypes.</emphasis>
5 males and 4 females in the same location and the same other data; 2 females at the beginning of the Najib valley,
<geoCoordinate degrees="31" direction="north" minutes="37.60" orientation="latitude" precision="9" value="31.626667">31°37.60'N</geoCoordinate>
,
<geoCoordinate degrees="54" direction="east" minutes="05.38" orientation="longitude" precision="9" value="54.08967">54°05.38'E</geoCoordinate>
, 3130 m a.s.l., 3 males and 2 females at the end of the Sorgan valley,
<geoCoordinate degrees="31" direction="north" minutes="37.65" orientation="latitude" precision="9" value="31.6275">31°37.65'N</geoCoordinate>
,
<geoCoordinate degrees="54" direction="east" minutes="05.35" orientation="longitude" precision="9" value="54.08917">54°05.35'E</geoCoordinate>
, 3155 m a.s.l., other data same. Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Oenopia shirkuhensis sp. nov., paratype 1 (female), dorsal view." figureDoi="10.3897/zookeys.915.46390.figure1" httpUri="https://binary.pensoft.net/fig/383100" pageId="0" pageNumber="107">1</figureCitation>
,
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">3</figureCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.915.46390.figure1" httpUri="https://binary.pensoft.net/fig/383100" pageId="0" pageNumber="107" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" pageId="0" pageNumber="107">Figure 1.</emphasis>
<taxonomicName authorityName="Zare Khormizi &amp; Nedved" authorityYear="2020" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenopia shirkuhensis" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="shirkuhensis">
<emphasis italics="true" pageId="0" pageNumber="107">Oenopia shirkuhensis</emphasis>
</taxonomicName>
sp. nov., paratype 1 (female), dorsal view.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107" start="Figures 2–10" startId="F2">
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" pageId="0" pageNumber="107">Figures 2-10.</emphasis>
<taxonomicName authorityName="Zare Khormizi &amp; Nedved" authorityYear="2020" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenopia shirkuhensis" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="shirkuhensis">
<emphasis italics="true" pageId="0" pageNumber="107">Oenopia shirkuhensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="107">2</emphasis>
holotype (male), dorsal view
<emphasis bold="true" pageId="0" pageNumber="107">3</emphasis>
paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward
<emphasis bold="true" pageId="0" pageNumber="107">4</emphasis>
individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view
<emphasis bold="true" pageId="0" pageNumber="107">5</emphasis>
prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow
<emphasis bold="true" pageId="0" pageNumber="107">6</emphasis>
paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45°) separated from the main branch and from the coxal fovea
<emphasis bold="true" pageId="0" pageNumber="107">7</emphasis>
holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep
<emphasis bold="true" pageId="0" pageNumber="107">8</emphasis>
holotype, tegmen, lateral view. The tips are inclined about 45° from the plane of penis guide main body. Parameres as long as the penis guide
<emphasis bold="true" pageId="0" pageNumber="107">9</emphasis>
holotype, penis
<emphasis bold="true" pageId="0" pageNumber="107">10</emphasis>
holotype, tip of penis. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="107">2, 3, 4, 9</emphasis>
); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="107">7, 8, 10</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="107">
The holotype and one paratype of the new species will be deposited in the National Museum, Prague, Czechia. Five paratypes will be deposited at the University of South Bohemia,
<normalizedToken originalValue="České">Ceske</normalizedToken>
<normalizedToken originalValue="Budějovice">Budejovice</normalizedToken>
, Czechia. All specimens are in alcohol, but genitalia are on microscope glass in resin. Ten paratypes will be deposited in the collection of the first author, in Yazd, Iran. All are frozen.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="materials_examined">
<paragraph pageId="0" pageNumber="107">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="107">
Iran, Kerman province, 5.5 km E of Sekonj village, Kukhbenan Mountains, at
<geoCoordinate degrees="30" direction="north" minutes="00.00" orientation="latitude" precision="9" value="30.0">30°00.00'N</geoCoordinate>
,
<geoCoordinate degrees="57" direction="east" minutes="28.53" orientation="longitude" precision="9" value="57.4755">57°28.53'E</geoCoordinate>
, 2540 m a.s.l., 28. vi. 2014, lgt. Skrylnik Yu., 1 individual, determined based on the photograph made by Alexander Slutsky, Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">4</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="etymology">
<paragraph pageId="0" pageNumber="107">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="107">Adjective referring to the place of sampling, on the slopes of Shirkooh mountain (which means Lion mountain).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="diagnosis">
<paragraph pageId="0" pageNumber="107">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="107">
This species can be separated from its congeners by the colour pattern of elytra, which is orange-red with only one pair of rounded black spots lying slightly behind the half length of the elytra, either in the middle between suture and lateral margin (Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">2</figureCitation>
) or extended to the suture (Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">3</figureCitation>
), which bears a narrow black stripe. In the closest relative, the elytra has 16 spots (Figs
<figureCitation captionStart="Figures 11–17" captionStartId="F3" captionText="Figures 11 - 17. Oenopia conglobata 11 from Yazd province, Iran, yellowish form. Dorsal view 12 from Yazd province, Iran, rose form. Dorsal view 13 from Chaharmahal and Bakhtiary province, Iran. Ventral view. Abdominal ventrites with large triangular yellow-brown margins 14 tegmen, outer view. The tips of penis guide shorter than wide, parabolic shape between the two tips more than three times wide as deep 15 tegmen, lateral view. The tips are inclined about 80 ° from the plane of penis guide main body. Parameres slightly shorter than the penis guide 16 penis 17 tip of penis. Scale bars: 1 mm (11, 13, 16); 0.5 mm (14, 15, 17)." figureDoi="10.3897/zookeys.915.46390.figures11-17" httpUri="https://binary.pensoft.net/fig/383102" pageId="0" pageNumber="107">11</figureCitation>
,
<figureCitation captionStart="Figures 11–17" captionStartId="F3" captionText="Figures 11 - 17. Oenopia conglobata 11 from Yazd province, Iran, yellowish form. Dorsal view 12 from Yazd province, Iran, rose form. Dorsal view 13 from Chaharmahal and Bakhtiary province, Iran. Ventral view. Abdominal ventrites with large triangular yellow-brown margins 14 tegmen, outer view. The tips of penis guide shorter than wide, parabolic shape between the two tips more than three times wide as deep 15 tegmen, lateral view. The tips are inclined about 80 ° from the plane of penis guide main body. Parameres slightly shorter than the penis guide 16 penis 17 tip of penis. Scale bars: 1 mm (11, 13, 16); 0.5 mm (14, 15, 17)." figureDoi="10.3897/zookeys.915.46390.figures11-17" httpUri="https://binary.pensoft.net/fig/383102" pageId="0" pageNumber="107">12</figureCitation>
). Abdominal ventrites are completely black (Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">6</figureCitation>
), while they have yellow margins in some congeners (Fig.
<figureCitation captionStart="Figures 11–17" captionStartId="F3" captionText="Figures 11 - 17. Oenopia conglobata 11 from Yazd province, Iran, yellowish form. Dorsal view 12 from Yazd province, Iran, rose form. Dorsal view 13 from Chaharmahal and Bakhtiary province, Iran. Ventral view. Abdominal ventrites with large triangular yellow-brown margins 14 tegmen, outer view. The tips of penis guide shorter than wide, parabolic shape between the two tips more than three times wide as deep 15 tegmen, lateral view. The tips are inclined about 80 ° from the plane of penis guide main body. Parameres slightly shorter than the penis guide 16 penis 17 tip of penis. Scale bars: 1 mm (11, 13, 16); 0.5 mm (14, 15, 17)." figureDoi="10.3897/zookeys.915.46390.figures11-17" httpUri="https://binary.pensoft.net/fig/383102" pageId="0" pageNumber="107">13</figureCitation>
). The two tips of penis guide are longer than wide and the parabolic shape between them, in outer view, is less than two times as wide as deep (Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">7</figureCitation>
). The tips in lateral view are inclined about 45° from the plane of main body (Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">8</figureCitation>
). Parameres are as long as the penis guide. In
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenopia conglobata" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="conglobata">
<emphasis italics="true" pageId="0" pageNumber="107">Oenopia conglobata</emphasis>
</taxonomicName>
, the tips of penis guide are shorter than wide, and the parabolic shape between the two tips are more than three times as wide as deep (Fig.
<figureCitation captionStart="Figures 11–17" captionStartId="F3" captionText="Figures 11 - 17. Oenopia conglobata 11 from Yazd province, Iran, yellowish form. Dorsal view 12 from Yazd province, Iran, rose form. Dorsal view 13 from Chaharmahal and Bakhtiary province, Iran. Ventral view. Abdominal ventrites with large triangular yellow-brown margins 14 tegmen, outer view. The tips of penis guide shorter than wide, parabolic shape between the two tips more than three times wide as deep 15 tegmen, lateral view. The tips are inclined about 80 ° from the plane of penis guide main body. Parameres slightly shorter than the penis guide 16 penis 17 tip of penis. Scale bars: 1 mm (11, 13, 16); 0.5 mm (14, 15, 17)." figureDoi="10.3897/zookeys.915.46390.figures11-17" httpUri="https://binary.pensoft.net/fig/383102" pageId="0" pageNumber="107">14</figureCitation>
), the tips are inclined about 80° from the plane of penis guide main body (Fig.
<figureCitation captionStart="Figures 11–17" captionStartId="F3" captionText="Figures 11 - 17. Oenopia conglobata 11 from Yazd province, Iran, yellowish form. Dorsal view 12 from Yazd province, Iran, rose form. Dorsal view 13 from Chaharmahal and Bakhtiary province, Iran. Ventral view. Abdominal ventrites with large triangular yellow-brown margins 14 tegmen, outer view. The tips of penis guide shorter than wide, parabolic shape between the two tips more than three times wide as deep 15 tegmen, lateral view. The tips are inclined about 80 ° from the plane of penis guide main body. Parameres slightly shorter than the penis guide 16 penis 17 tip of penis. Scale bars: 1 mm (11, 13, 16); 0.5 mm (14, 15, 17)." figureDoi="10.3897/zookeys.915.46390.figures11-17" httpUri="https://binary.pensoft.net/fig/383102" pageId="0" pageNumber="107">15</figureCitation>
), and the parameres are slightly shorter than the guide.
</paragraph>
<caption doi="10.3897/zookeys.915.46390.figures11-17" httpUri="https://binary.pensoft.net/fig/383102" pageId="0" pageNumber="107" start="Figures 11–17" startId="F3">
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" pageId="0" pageNumber="107">Figures 11-17.</emphasis>
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenopia conglobata" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="conglobata">
<emphasis italics="true" pageId="0" pageNumber="107">Oenopia conglobata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="107">11</emphasis>
from Yazd province, Iran, yellowish form. Dorsal view
<emphasis bold="true" pageId="0" pageNumber="107">12</emphasis>
from Yazd province, Iran, rose form. Dorsal view
<emphasis bold="true" pageId="0" pageNumber="107">13</emphasis>
from Chaharmahal and Bakhtiary province, Iran. Ventral view. Abdominal ventrites with large triangular yellow-brown margins
<emphasis bold="true" pageId="0" pageNumber="107">14</emphasis>
tegmen, outer view. The tips of penis guide shorter than wide, parabolic shape between the two tips more than three times wide as deep
<emphasis bold="true" pageId="0" pageNumber="107">15</emphasis>
tegmen, lateral view. The tips are inclined about 80° from the plane of penis guide main body. Parameres slightly shorter than the penis guide
<emphasis bold="true" pageId="0" pageNumber="107">16</emphasis>
penis
<emphasis bold="true" pageId="0" pageNumber="107">17</emphasis>
tip of penis. Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="107">11, 13, 16</emphasis>
); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="107">14, 15, 17</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="description">
<paragraph pageId="0" pageNumber="107">Description.</paragraph>
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Body</emphasis>
outline (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Oenopia shirkuhensis sp. nov., paratype 1 (female), dorsal view." figureDoi="10.3897/zookeys.915.46390.figure1" httpUri="https://binary.pensoft.net/fig/383100" pageId="0" pageNumber="107">1</figureCitation>
-
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">4</figureCitation>
) broad oval, 1.35
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, broadest behind middle of elytra. Dorsum convex and glabrous, except few hairs on head near eyes. Interocular distance 2.7
<normalizedToken originalValue="×">x</normalizedToken>
as wide as eye. Pronotum with fine punctures. Elytra with large shallow punctures, separated by 1-2 diameters, interspaces between punctures smooth, shiny.
</paragraph>
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Head</emphasis>
creamy white.
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Pronotum</emphasis>
creamy white with seven black spots forming triple VVV, with two pairs of additional small spots lateral to large mediolateral spots.
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Elytra</emphasis>
orange-red, shiny, with one pair of rounded black spots lying slightly behind the half length of elytra (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparative body and pattern measurements of Oenopia shirkuhensis, related O. conglobata and visually similar A. bipunctata. A: body length / width ratio; B: position of centre of the main elytral spot as relative longitudinal distance from the front margin of elytra; C: distance of the centres of the main elytral spots as transversal distance relative to the width of elytra." httpUri="http://table.plazi.org/id/C1CC4BF4C6AC36847C330108FCDCCCCE" pageId="0" pageNumber="107" tableUuid="C1CC4BF4C6AC36847C330108FCDCCCCE">1</tableCitation>
), in the middle between suture and lateral margin or eliptical, extended to the suture, which bears a narrow (quarter of width of scutellum) black stripe. Pronotum and elytra with explanate lateral margins transparent.
</paragraph>
<paragraph pageId="0" pageNumber="107">
Prosternal intercoxal process (Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">5</figureCitation>
) narrow, with a pair of carinae extending over two-thirds of the prosternum length. Ventral side of thorax black, mesepimeron white with narrow black margins. Epipleuron 8
<normalizedToken originalValue="×">x</normalizedToken>
narrower than body width. Scutellum 13
<normalizedToken originalValue="×">x</normalizedToken>
narrower than body width (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Oenopia shirkuhensis sp. nov., paratype 1 (female), dorsal view." figureDoi="10.3897/zookeys.915.46390.figure1" httpUri="https://binary.pensoft.net/fig/383100" pageId="0" pageNumber="107">1</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Abdomen</emphasis>
with ventrites completely black. Abdominal postcoxal line (Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">6</figureCitation>
) bifurcated with the main branch parallel to the posterior margin of the ventrite 1, almost reaching lateral margin, secondary branch oblique (45°) separated from the main branch and from the coxal cavity.
</paragraph>
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Legs</emphasis>
orange-brown, coxae black, fore coxae with white spot. Tarsal claws with basal tooth.
</paragraph>
<paragraph pageId="0" pageNumber="107">
Male
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">genitalia</emphasis>
(Figs
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">7-10</figureCitation>
) with penis guide of tegmen in outer view (Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">7</figureCitation>
) bifurcated; penis guide slightly constricted before tips. Tips longer than wide, parabolic shape between tips less than two-times as wide as deep. Parameres as long as penis guide, clavate, truncate at apices with dense and long hairs. Penis guide in lateral view (Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">8</figureCitation>
) gradually narrowed. Tips in lateral view inclined at about 45° from main body plane. Penis (Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">9</figureCitation>
) with well-developed basal capsule with inner and outer arms subequal; penis apex strongly narrowed, then enlarged to wide soft tip (Fig.
<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Oenopia shirkuhensis sp. nov. 2 holotype (male), dorsal view 3 paratype 3 (male), dorsal view. Notice elytral spots extended towards the suture and slightly backward 4 individual from Kerman province, collected by Yu. Skrylnik, photographed by Alexander Slutsky, dorsal view 5 prosternum and mesosternum. Prosternal carinae extending over more than half of the prosternum length. Fovea in the front margin of mesosternum is shallow 6 paratype 2, first abdominal ventrite, left side. Ventrites completely black. Abdominal postcoxal line bifurcated with the main branch parallel to the posterior margin of the ventrite 1, not reaching lateral margin, secondary branch oblique (45 °) separated from the main branch and from the coxal fovea 7 holotype, tegmen, outer view. The tips longer than wide, parabolic shape between the two tips less than two times wide as deep 8 holotype, tegmen, lateral view. The tips are inclined about 45 ° from the plane of penis guide main body. Parameres as long as the penis guide 9 holotype, penis 10 holotype, tip of penis. Scale bars: 1 mm (2, 3, 4, 9); 0.5 mm (7, 8, 10)." figureDoi="10.3897/zookeys.915.46390.figures2-10" httpUri="https://binary.pensoft.net/fig/383101" pageId="0" pageNumber="107">10</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Body measurements</emphasis>
: in mm; first value belongs to holotype, others to paratypes (smallest, largest, and most beautiful, i.e. female in Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Oenopia shirkuhensis sp. nov., paratype 1 (female), dorsal view." figureDoi="10.3897/zookeys.915.46390.figure1" httpUri="https://binary.pensoft.net/fig/383100" pageId="0" pageNumber="107">1</figureCitation>
, of the paratypes);
<emphasis bold="true" pageId="0" pageNumber="107">BL</emphasis>
= 3.87, 3.76, 4.31, 4.05;
<emphasis bold="true" pageId="0" pageNumber="107">BW</emphasis>
= 2.89, 2.87, 3.35, 2.92;
<emphasis bold="true" pageId="0" pageNumber="107">HW</emphasis>
= 1.01, 0.96, 1.12, 1.03;
<emphasis bold="true" pageId="0" pageNumber="107">PL</emphasis>
= 0.88, 0.74, 0.88, 0.80;
<emphasis bold="true" pageId="0" pageNumber="107">PW</emphasis>
= 1.90, 1.92, 2.21, 2.08;
<emphasis bold="true" pageId="0" pageNumber="107">EL</emphasis>
= 2.79, 2.87, 3.21, 3.16.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="key to the iranian oenopia">
<paragraph pageId="0" pageNumber="107">
Key to the Iranian
<taxonomicName class="Insecta" family="Lophopidae" genus="Oenopia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oenopia" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="107">Oenopia</emphasis>
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<td colspan="1" pageId="0" pageNumber="107" rowspan="1">1</td>
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<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">2</emphasis>
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<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Elytra black with light spots or at least light fore margin. Pronotum black with light anterior and lateral margins</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<taxonomicName lsidName="O. oncina" pageId="0" pageNumber="107" rank="species" species="oncina">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">O. oncina</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Elytra beige, pink, or orange to red, with rounded dark spots at least in posterior half. Mesepimeron white with narrow black margins; metaepimera and metepisterna dark. Tarsal claw with basal tooth</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Elytra beige with small dark dots in anterior half and irregular lines in posterior half. Mesepimera, metaepimera, and metepisterna pale. Tarsal claw without tooth</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<taxonomicName lsidName="O. montana" pageId="0" pageNumber="107" rank="species" species="montana">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">O. montana</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Elytra with up to 16 spots (pattern 2+2+1+3), either black or brown with lighter center. Penis guide with tip curved upwards in right angle</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<taxonomicName lsidName="O. conglobata" pageId="0" pageNumber="107" rank="species" species="conglobata">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">O. conglobata</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Elytra with one pair of black spots behind center. Penis guide with tip slightly curved upwards at 45° angle</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<taxonomicName lsidName="O. shirkuhensis" pageId="0" pageNumber="107" rank="species" species="shirkuhensis">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">O. shirkuhensis</emphasis>
</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

74
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<emphasis id="52DC1909852CFA58FF193233FEA6FA03" bold="true" box="[151,324,1394,1420]" italics="true" pageId="0" pageNumber="92">Enoplognatha</emphasis>
Pavesi, 1880
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Type species:
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(Lucas, 1846)
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<taxonomicName LSID="E8025213-947D-59D6-92C4-D9E347063647" authority="K. A. Sheph." authorityName="Shepherd &amp; Lepschi &amp; Johnson &amp; Gardner &amp; Sessa &amp; Jabaily" authorityYear="2020" class="Magnoliopsida" family="Goodeniaceae" genus="Goodenia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Goodenia capillosa" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="capillosa" status="nom. nov.">Goodenia capillosa K.A.Sheph.</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="27">nom. nov.</taxonomicNameLabel>
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<paragraph pageId="0" pageNumber="27">
≡
<taxonomicName class="Magnoliopsida" family="Goodeniaceae" genus="Velleia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Velleia hispida" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="hispida">Velleia hispida</taxonomicName>
W.Fitzg.,
<emphasis italics="true" pageId="0" pageNumber="27">W. Austral. Nat. Hist. Soc.</emphasis>
1: 25 (1904) - Lectotype (designated by Carolin, Proc. Linn. Soc. New South Wales 92(1): 42 (1967): Nannine,
<emphasis italics="true" pageId="0" pageNumber="27">W.V.Fitzgerald</emphasis>
, Sept. 1903 (NSW 75661,
<emphasis italics="true" pageId="0" pageNumber="27">n.v.</emphasis>
); isolectotype: Australia. Western Australia. Nannine, Sep. 1903, [
<emphasis italics="true" pageId="0" pageNumber="27">W.V.Fitzgerald s.n.</emphasis>
] (PERTH 01639986!)).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="notes">
<paragraph pageId="0" pageNumber="27">Note.</paragraph>
<paragraph pageId="0" pageNumber="27">
The epithet
<normalizedToken originalValue="‘hispida’">'hispida'</normalizedToken>
is unavailable in
<taxonomicName authorityName="Sm., Spec. bot. New Holland 15. 1793, nom. cons. (fide Shepherd et al. 2017; Applequist" authorityYear="2019" class="Magnoliopsida" family="Goodeniaceae" genus="Goodenia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Goodenia" order="Asterales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Goodenia</emphasis>
</taxonomicName>
as it is preoccupied by
<taxonomicName lsidName="G. hispida" pageId="0" pageNumber="27" rank="species" species="hispida">
<emphasis italics="true" pageId="0" pageNumber="27">G. hispida</emphasis>
</taxonomicName>
R.Br.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="etymology">
<paragraph pageId="0" pageNumber="27">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="27">
Named from the Latin
<emphasis italics="true" pageId="0" pageNumber="27">capillosus</emphasis>
(hairy) in reference to the indumentum present on the leaves and sepals.
</paragraph>
</subSubSection>
</treatment>
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122
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.494.8748" ID-GBIF-Dataset="e8b2a5fb-bd85-48c3-b33e-121ddc15bec9" ID-PMC="PMC4400378" ID-Pensoft-Pub="1313-2970-494-69" ID-PubMed="25901115" ID-ZBK="F73F468E4DF7479E85C4717466635BF9" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-494-69" ModsDocOrigin="ZooKeys 494" ModsDocTitle="Revision of the genus Philonome Chambers and its proposed reassignment to the family Tineidae (Lepidoptera, Tineoidea)" checkinTime="1451244473555" checkinUser="pensoft" docAuthor="Sohn, Jae-Cheon, Davis, Donald R. &amp; Lopez-Vaamonde, Carlos" docDate="2015" docId="E802732273E68385737ACA63CBFD5E3C" docLanguage="en" docName="ZooKeys 494: 69-106" docOrigin="ZooKeys 494" docSource="http://dx.doi.org/10.3897/zookeys.494.8748" docTitle="Philonome lambdagrapha Sohn, Davis &amp; Lopez-Vaamonde, sp. n." docType="treatment" docUuid="E4E1964A-D664-4342-B555-DBC16C934821" docUuidSource="ZooBank" docVersion="4" lastPageNumber="82" masterDocId="3D1D9C71FF96FFF09C07FFAC9A60FF9D" masterDocTitle="Revision of the genus Philonome Chambers and its proposed reassignment to the family Tineidae (Lepidoptera, Tineoidea)" masterLastPageNumber="106" masterPageNumber="69" pageNumber="80" updateTime="1668160171790" updateUser="ExternalLinkService">
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<paragraph pageId="11" pageNumber="80">Taxon classification Animalia Lepidoptera Lyonetiidae</paragraph>
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<paragraph pageId="11" pageNumber="80">
<taxonomicName LSID="http://zoobank.org/E4E1964A-D664-4342-B555-DBC16C934821" authority="Sohn, Davis &amp; Lopez-Vaamonde" class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome lambdagrapha" order="Lepidoptera" pageId="11" pageNumber="80" phylum="Arthropoda" rank="species" species="lambdagrapha">Philonome lambdagrapha Sohn, Davis &amp; Lopez-Vaamonde</taxonomicName>
<taxonomicNameLabel pageId="11" pageNumber="80">sp. n.</taxonomicNameLabel>
Figs 6, 37-39
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<pageBreakToken pageId="12" pageNumber="81" start="start">Diagnosis</pageBreakToken>
.
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This species is similar to
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome curvilineata" order="Lepidoptera" pageId="12" pageNumber="81" phylum="Arthropoda" rank="species" species="curvilineata">Philonome curvilineata</taxonomicName>
in external appearance but differs from the latter in having the longitudinal and costal fasciae separate (continuous in
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome curvilineata" order="Lepidoptera" pageId="12" pageNumber="81" phylum="Arthropoda" rank="species" species="curvilineata">Philonome curvilineata</taxonomicName>
and larger apical protrusion on the tegumen in the male genitalia.
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<paragraph pageId="12" pageNumber="81">Adult</paragraph>
<paragraph pageId="12" pageNumber="81">(Fig. 6). Head: Vertex orange on posterior 2/3, pale orange on anterior 1/3; scales on dorsum of occiput dark grayish brown; scales between antennal scapes lustrous pale orange; frons lustrous pale yellow. Antenna 4/5 as long as forewing; scape as long as diameter of eye, lustrous orange dorsally, lustrous pale yellowish gray laterally and ventrally, lustrous pale grayish brown apically; flagellomeres pale orange dorsally, lustrous pale yellow ventrally. Labial palpus 1/2 as long as antennal scape, lustrous pale yellowish gray.</paragraph>
<paragraph pageId="13" pageNumber="82">
<pageBreakToken pageId="13" pageNumber="82" start="start">Thorax</pageBreakToken>
: Patagium dark brown; tegula lustrous pale yellow, intermixed with orange scales basally; mesonotum silvery white with dark brown transverse band along anterior and posterior margins and at anterior 1/3, with orange transverse band at middle. Foreleg with coxa lustrous pale yellow; femur lustrous pale yellow, intermixed with pale grayish brown laterally; tibia dark brown dorsally, pale grayish yellow ventrally; tarsomeres dark brown dorsally, pale orange ventrally. Midleg with coxa and femur lustrous pale yellow; tibia and tarsomeres grayish brown dorsally, pale yellow ventrally. Hindleg consumed for DNA extraction. Forewing length 3.0 mm (n = 1), reddish brown; costa black in distal 1/3; longitudinal fascia extending to apical streak, straight, white on basal 1/2, juxtaposed with a slender, intermittent black line along lower border, sinuous, black on distal 1/2; costal fasciae slender, extending to apex; subapical streak white, juxtaposed with slender black line along lower border; apical streak white, connected with longitudinal fascia; dorsal bar white, juxtaposed with black along outer border; dorsal margin sparsely irrorated with black scales on basal 1/6 and at middle; tornal patch elongate, white, juxtaposed with black along upper border, irrorated with dark brown scales along outer border; marginal streak dark brown; fringe brown on distal 1/3 of costa, pale yellowish gray along termen. Hindwing brownish gray; fringe pale grayish brown.
</paragraph>
<paragraph pageId="13" pageNumber="82">Abdomen: Male tergum VIII and sternum VIII subquadrate; coremata piliform, as long as tergum VIII.</paragraph>
<paragraph pageId="13" pageNumber="82">Male genitalia (Figs 37-39): Tegumen nearly as long as valva, semi-elliptical on basal 3/4, rectangular on distal 1/4, with small lateral protrusion dorsoposteriorly. Valva elongate, lobate, sparsely setose on outer surface. Juxta liguiform, 1/2 as long as valva. Vinculum broad, gradually broadened anteriorly, with medial and lateral protrusions along anterior margin; Phallus slightly curved at basal 2/5, broadened posteriorly.</paragraph>
<caption pageId="13" pageNumber="82">
<paragraph pageId="13" pageNumber="82">
Figures 37-46.
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome" order="Lepidoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="genus">Philonome</taxonomicName>
, male genitalia. 37-39
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome lambdagrapha" order="Lepidoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="lambdagrapha">Philonome lambdagrapha</taxonomicName>
37 Ventral view of genital capsule and phallus 38 Lateral view 39 Phallus 40-41
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome curvilineata" order="Lepidoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="curvilineata">Philonome curvilineata</taxonomicName>
. 40 Ventral view of genital capsule and phallus 41 Lateral view 42-46
<taxonomicName class="Insecta" family="Lyonetiidae" genus="Philonome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonome euryarga" order="Lepidoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="euryarga">Philonome euryarga</taxonomicName>
42 Ventral view of genital capsule 43 Ventral view 44 abdominal tergum VIII (caudal end directed upward) 45 Lateral view of phallus 46 Anterior base of phallus, ventral view.
</paragraph>
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<paragraph pageId="13" pageNumber="82">Female unknown.</paragraph>
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<paragraph pageId="13" pageNumber="82">Type.</paragraph>
<paragraph pageId="13" pageNumber="82">
Holotype: male, &quot;ID#: CLV102310 [red letters] French Guiana: [
<normalizedToken originalValue="Régina">Regina</normalizedToken>
, Nouragues Research Station] [Lt:4.1 Ln:52/] Carlos Lopez Vaamonde 23-Jan-2010 DNA Barcode LNOUA928-10 [green letters in blue row]&quot;,
<normalizedToken originalValue="“Nou68”">&quot;Nou68&quot;</normalizedToken>
, &quot;Genitalia slide DRD ♂ USNM 34621&quot; [green label], USNM.
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<paragraph pageId="13" pageNumber="82">Distribution.</paragraph>
<paragraph pageId="13" pageNumber="82">French Guiana.</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="82" type="etymology">
<paragraph pageId="13" pageNumber="82">Etymology.</paragraph>
<paragraph pageId="13" pageNumber="82">
The species name is derived from the Greek letter
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and a suffix derived from the Greek
<normalizedToken originalValue="‘graphein’">'graphein'</normalizedToken>
meaning &quot;to write&quot;, and refers to the white fascia of the forewing resembling a lambda (λ).
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279
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<bibRefCitation id="043A4B0EB114FFF1CDB95259FF29FEAC" author="Marsh" box="[151,202,292,313]" pageId="4" pageNumber="56" refString="Marsh, O. C. (1879) Notice of new Jurassic reptiles. American Journal of Science and Arts, 18, 501 - 506." type="journal article" year="1879">1879</bibRefCitation>
<taxonomicName id="A7AB4D7CB114FFF1CDC75259FDE3FEAC" authority="Marsh" authorityName="Marsh" box="[233,512,291,314]" class="Insecta" family="Haglidae" genus="Camptonotus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="52DFEAEDB114FFF1CDC75259FE54FEAF" box="[233,439,292,314]" italics="true" pageId="4" pageNumber="56">Camptonotus dispar</emphasis>
Marsh
</taxonomicName>
, p. 501. 1885
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<emphasis id="52DFEAEDB114FFF1CDC7523EFE20FECD" box="[233,451,322,344]" italics="true" pageId="4" pageNumber="56">Camptosaurus dispar</emphasis>
Marsh
</taxonomicName>
, p. 169. 1894
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<emphasis id="52DFEAEDB114FFF1CDC7521FFE28FEE2" box="[233,459,353,375]" italics="true" pageId="4" pageNumber="56">Camptosaurus medius</emphasis>
Marsh
</taxonomicName>
, p. 85. 1894
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Marsh
</taxonomicName>
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1909
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<emphasis id="52DFEAEDB114FFF1CDC752E2FE29FE20" box="[233,458,415,437]" italics="true" pageId="4" pageNumber="56">Camptosaurus browni</emphasis>
Gilmore
</taxonomicName>
, p. 295
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.
</emphasis>
<collectionCode id="06BAAE3AB114FFF1CC6E517BFE9FFD88" LSID="urn:lsid:biocol.org:col:34880" box="[320,380,518,541]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="4" pageNumber="56">YPM</collectionCode>
1878, partial postcranium from Quarry 13.
</paragraph>
<paragraph id="601436FFB114FFF1CDEB5155FD64FD1C" blockId="4.[151,1437,481,1621]" pageId="4" pageNumber="56">
<emphasis id="52DFEAEDB114FFF1CDEB5155FE40FDD4" bold="true" box="[197,419,552,577]" pageId="4" pageNumber="56">Referred material.</emphasis>
Disarticulated cranial and postcranial material in
<collectionCode id="06BAAE3AB114FFF1CEF95154FBCBFDD4" LSID="urn:lsid:biocol.org:col:34871" box="[983,1064,553,577]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="56">USNM</collectionCode>
and
<collectionCode id="06BAAE3AB114FFF1C94E5157FB7FFDD4" LSID="urn:lsid:biocol.org:col:34880" box="[1120,1180,554,577]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="4" pageNumber="56">YPM</collectionCode>
collections, representing at least 15 additional individuals from Quarry 13 (
<bibRefCitation id="043A4B0EB114FFF1CFDC5130FC70FDF0" author="Gilmore" box="[754,915,589,613]" pageId="4" pageNumber="56" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
;
<bibRefCitation id="043A4B0EB114FFF1CEB15133FBCAFDF0" author="Foster" box="[927,1065,589,613]" pageId="4" pageNumber="56" refString="Foster, J. R. (2003) Paleoecological analysis of the vertebrate fauna of the Morrison Formation (Upper Jurassic), Rocky Mountain region, U. S. A. New Mexico Museum of Natural History and Science Bulletin, 23, 1 - 95." type="journal article" year="2003">Foster 2003</bibRefCitation>
).
<collectionCode id="06BAAE3AB114FFF1C9125133FB9BFDF0" LSID="urn:lsid:biocol.org:col:34880" box="[1084,1144,590,613]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="4" pageNumber="56">YPM</collectionCode>
PU14553, a partial skeleton from Cleveland-Lloyd Dinosaur Quarry.
</paragraph>
<paragraph id="601436FFB114FFF1CDEB51E8FEEDFD60" blockId="4.[151,1437,481,1621]" pageId="4" pageNumber="56">
<emphasis id="52DFEAEDB114FFF1CDEB51E8FC5FFD3B" bold="true" box="[197,956,661,686]" pageId="4" pageNumber="56">
Locality and horizon of
<collectionCode id="06BAAE3AB114FFF1CCCD51E8FDD5FD3B" LSID="urn:lsid:biocol.org:col:34871" box="[483,566,661,686]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="56">USNM</collectionCode>
and
<collectionCode id="06BAAE3AB114FFF1CF5E51E8FD53FD38" LSID="urn:lsid:biocol.org:col:34880" box="[624,688,661,685]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="4" pageNumber="56">YPM</collectionCode>
Quarry
<specimenCount id="76ADFD76B114FFF1CE3651E8FC54FD3B" box="[792,951,661,686]" pageId="4" pageNumber="56" type="generic">13 specimens</specimenCount>
.
</emphasis>
Quarry 13 at Como Bluff, Albany County, Wyoming; Salt Wash Member of the Morrison Formation, Kimmeridgian (
<bibRefCitation id="043A4B0EB114FFF1CECF51C4FB29FD44" author="Kowallis" box="[993,1226,697,721]" pageId="4" pageNumber="56" refString="Kowallis, B. J., Christiansen, E. H., Deino, A. L., Peterson, F., Turner, C. E., Kunk, M. J. &amp; Obradovich, J. D. (1998) The age of the Morrison Formation. Modern Geology, 22, 235 - 260." type="journal article" year="1998">
Kowallis
<emphasis id="52DFEAEDB114FFF1C96151C7FB62FD44" box="[1103,1153,697,721]" italics="true" pageId="4" pageNumber="56">et al</emphasis>
. 1998
</bibRefCitation>
;
<bibRefCitation id="043A4B0EB114FFF1C9F851C4FEE1FD60" author="Carpenter" pageId="4" pageNumber="56" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson 2008</bibRefCitation>
).
</paragraph>
<paragraph id="601436FFB114FFF1CDEB507CFB6CFCA8" blockId="4.[151,1437,481,1621]" pageId="4" pageNumber="56">
<emphasis id="52DFEAEDB114FFF1CDEB507CFD4FFC8F" bold="true" box="[197,684,769,794]" pageId="4" pageNumber="56">
Locality and horizon of
<collectionCode id="06BAAE3AB114FFF1CCC1507FFDCCFC8F" LSID="urn:lsid:biocol.org:col:34880" box="[495,559,770,794]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="4" pageNumber="56">YPM</collectionCode>
PU14553.
</emphasis>
Cleveland-Lloyd Dinosaur Quarry, Emery County, Utah; Brushy Basin Member of the Morrison Formation, lower–middle Tithonian (
<bibRefCitation id="043A4B0EB114FFF1CEB45058FB60FCA8" author="Kowallis" box="[922,1155,805,829]" pageId="4" pageNumber="56" refString="Kowallis, B. J., Christiansen, E. H., Deino, A. L., Peterson, F., Turner, C. E., Kunk, M. J. &amp; Obradovich, J. D. (1998) The age of the Morrison Formation. Modern Geology, 22, 235 - 260." type="journal article" year="1998">
Kowallis
<emphasis id="52DFEAEDB114FFF1C926505BFBDAFCA8" box="[1032,1081,805,829]" italics="true" pageId="4" pageNumber="56">et al</emphasis>
. 1998
</bibRefCitation>
).
</paragraph>
<paragraph id="601436FFB114FFF1CDEB5034FCF3FCF7" blockId="4.[151,1437,481,1621]" box="[197,784,841,866]" pageId="4" pageNumber="56">
<emphasis id="52DFEAEDB114FFF1CDEB5034FE43FCF7" bold="true" box="[197,416,841,866]" pageId="4" pageNumber="56">Generic diagnosis.</emphasis>
<collectingCountry id="18BC766FB114FFF1CC895037FE25FCF7" box="[423,454,842,866]" name="American Samoa" pageId="4" pageNumber="56">As</collectingCountry>
for the species by monotypy.
</paragraph>
<paragraph id="601436FFB114FFF1CDEB5010FC32FB80" blockId="4.[151,1437,481,1621]" pageId="4" pageNumber="56">
<emphasis id="52DFEAEDB114FFF1CDEB5010FE7CFC13" bold="true" box="[197,415,877,902]" pageId="4" pageNumber="56">Specific diagnosis.</emphasis>
Basal ankylopollexian diagnosed by the following unique combination of characters: scapula with triangular acromion process, straight dorsal margin of scapula (
<figureCitation id="F8902A7AB114FFF1CEE450ECFBEEFC3C" box="[970,1037,913,937]" captionStart="FIGURE 3" captionStartId="5.[151,250,1923,1946]" captionTargetBox="[218,1348,194,1877]" captionTargetId="figure@5.[213,1359,193,1878]" captionTargetPageId="5" captionText="FIGURE 3. (A) USNM 4282, left scapula of Camptosaurus dispar in lateral view; (B) CM 11337, cranial end of the right scapula of Uteodon aphanoecetes in lateral view; (C) CM 11337, left scapula of Uteodon aphanoecetes in lateral view; (D) OXFUM J. 3303, right scapula of Cumnoria prestwichii in lateral view. Abbreviations: cap, convex cranial margin of the acromion process; tap, triangular acromion process. Scale bar in A equals 10 cm; scale bar in B equals 5 cm." httpUri="https://zenodo.org/record/200814/files/figure.png" pageId="4" pageNumber="56">Fig. 3</figureCitation>
A), straight dorsal margin of ilium, dorsal margin of ilium thickens mediolaterally towards
<emphasis id="52DFEAEDB114FFF1CE3A50CBFCC8FC58" box="[788,811,950,973]" italics="true" pageId="4" pageNumber="56">M</emphasis>
.
<emphasis id="52DFEAEDB114FFF1CE1550C8FC5CFC58" box="[827,959,949,973]" italics="true" pageId="4" pageNumber="56">iliocaudalis</emphasis>
platform, postacetabular process of ilium tapers to a point with break in slope along dorsal margin (
<figureCitation id="F8902A7AB114FFF1CE3050A4FC80FC64" box="[798,867,985,1009]" captionStart="FIGURE 4" captionStartId="6.[151,250,720,743]" captionTargetBox="[151,1436,193,698]" captionTargetId="figure@6.[151,1436,193,699]" captionTargetPageId="6" captionText="FIGURE 4. Ilia of Camptosaurus dispar in lateral view. (A) right ilium of YPM 1877 (holotype) from Quarry 13; (B) right ilium of YPM 8622 from Quarry 13; (C) left ilium of USNM 5473 from Quarry 13; (D) left ilium of YPM PU 14553 from Cleveland-Lloyd Dinosaur Quarry. Abbreviations: mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process. Images in A, B, and D are copyright YPM. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200815/files/figure.png" pageId="4" pageNumber="56">Fig. 4</figureCitation>
), prepubic process unexpanded and concave along its dorsal margin (
<figureCitation id="F8902A7AB114FFF1CC4C5080FE46FB80" box="[354,421,1021,1045]" captionStart="FIGURE 5" captionStartId="6.[151,250,1330,1353]" captionTargetBox="[483,1104,894,1304]" captionTargetId="figure@6.[472,1115,884,1310]" captionTargetPageId="6" captionText="FIGURE 5. YPM 7334, right pubis of Camptosaurus dispar from Quarry 13 in lateral view. Abbreviations: pop, postpubic process; prp, prepubic process. Image is copyright YPM. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200816/files/figure.png" pageId="4" pageNumber="56">Fig. 5</figureCitation>
), distal end of ischium forms rounded expansion.
</paragraph>
</subSubSection>
<subSubSection id="28B16574B114FFF1CDEB575FFD0BF9C0" pageId="4" pageNumber="56" type="discussion">
<paragraph id="601436FFB114FFF1CDEB575FFD09FA84" blockId="4.[151,1437,481,1621]" pageId="4" pageNumber="56">
<emphasis id="52DFEAEDB114FFF1CDEB575FFEDAFBAF" bold="true" box="[197,313,1058,1082]" pageId="4" pageNumber="56">Remarks.</emphasis>
<taxonomicName id="A7AB4D7CB114FFF1CC11575FFDD2FBAC" box="[319,561,1057,1081]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB114FFF1CC11575FFDD2FBAC" box="[319,561,1057,1081]" italics="true" pageId="4" pageNumber="56">Camptosaurus dispar</emphasis>
</taxonomicName>
was originally named
<taxonomicName id="A7AB4D7CB114FFF1CE03575FFBF3FBAC" box="[813,1040,1057,1081]" class="Insecta" family="Haglidae" genus="Camptonotus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="52DFEAEDB114FFF1CE03575FFBF3FBAC" box="[813,1040,1057,1081]" italics="true" pageId="4" pageNumber="56">Camptonotus dispar</emphasis>
</taxonomicName>
by
<bibRefCitation id="043A4B0EB114FFF1C917575CFB32FBAC" author="Marsh" box="[1081,1233,1057,1081]" pageId="4" pageNumber="56" refString="Marsh, O. C. (1879) Notice of new Jurassic reptiles. American Journal of Science and Arts, 18, 501 - 506." type="journal article" year="1879">Marsh (1879)</bibRefCitation>
based upon material from Quarry 13, but was renamed
<taxonomicName id="A7AB4D7CB114FFF1CF74573BFC16FBC8" authority="Marsh 1885" authorityName="Marsh" authorityYear="1885" box="[602,1013,1093,1117]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB114FFF1CF74573BFCACFBC8" box="[602,847,1093,1117]" italics="true" pageId="4" pageNumber="56">Camptosaurus dispar</emphasis>
(
<bibRefCitation id="043A4B0EB114FFF1CE4E5738FC0FFBC8" author="Marsh" box="[864,1004,1093,1117]" pageId="4" pageNumber="56" refString="Marsh, O. C. (1885) Names of extinct reptiles. American Journal of Science, 29, 169." type="journal article" year="1885">Marsh 1885</bibRefCitation>
)
</taxonomicName>
because the name
<taxonomicName id="A7AB4D7CB114FFF1C9FA573BFA89FBC8" box="[1236,1386,1094,1117]" class="Insecta" family="Haglidae" genus="Camptonotus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="52DFEAEDB114FFF1C9FA573BFA89FBC8" box="[1236,1386,1094,1117]" italics="true" pageId="4" pageNumber="56">Camptonotus</emphasis>
</taxonomicName>
had already been coined by
<bibRefCitation id="043A4B0EB114FFF1CC8A5714FD84FB14" author="Uhler" box="[420,615,1129,1153]" pageId="4" pageNumber="56" refString="Uhler, P. R. (1864) Orthopterological contributions. Proceedings of the Entomological Society of Philadelphia, 2, 543 - 555." type="journal article" year="1864">Uhler (1864:548)</bibRefCitation>
for a genus of gryllacridid cricket (
<bibRefCitation id="043A4B0EB114FFF1CED75714FB05FB14" author="Capinera" box="[1017,1254,1129,1153]" pageId="4" pageNumber="56" refString="Capinera, J. L., Scott, R. D. &amp; Walker, T. J. (2004) Field Guide to Grasshoppers, Katydids, and Crickets of the United States. Comstock Publishing Associates, Ithaca, 249 pp." type="book" year="2004">
Capinera
<emphasis id="52DFEAEDB114FFF1C9465717FB79FB14" box="[1128,1178,1129,1153]" italics="true" pageId="4" pageNumber="56">et al</emphasis>
. 2004
</bibRefCitation>
). The additional species
<taxonomicName id="A7AB4D7CB114FFF1CDC157F3FDE2FB30" authority="Marsh 1894" authorityName="Marsh" authorityYear="1894" box="[239,513,1165,1189]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="medius">
<emphasis id="52DFEAEDB114FFF1CDC157F3FEE1FB30" box="[239,258,1166,1189]" italics="true" pageId="4" pageNumber="56">C</emphasis>
.
<emphasis id="52DFEAEDB114FFF1CC2157F0FEBDFB30" box="[271,350,1165,1189]" italics="true" pageId="4" pageNumber="56">medius</emphasis>
(
<bibRefCitation id="043A4B0EB114FFF1CC4357F0FE19FB30" author="Marsh" box="[365,506,1165,1189]" pageId="4" pageNumber="56" refString="Marsh, O. C. (1894) The typical Ornithopoda of the American Jurassic. American Journal of Science, 48, 85 - 90." type="journal article" year="1894">Marsh 1894</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="A7AB4D7CB114FFF1CF2357F3FCF0FB30" authority="Marsh 1894" authorityName="Marsh" authorityYear="1894" box="[525,787,1165,1189]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="nanus">
<emphasis id="52DFEAEDB114FFF1CF2357F3FDC3FB30" box="[525,544,1166,1189]" italics="true" pageId="4" pageNumber="56">C</emphasis>
.
<emphasis id="52DFEAEDB114FFF1CF0057F3FD92FB30" box="[558,625,1166,1189]" italics="true" pageId="4" pageNumber="56">nanus</emphasis>
(
<bibRefCitation id="043A4B0EB114FFF1CF5157F0FCEFFB30" author="Marsh" box="[639,780,1165,1189]" pageId="4" pageNumber="56" refString="Marsh, O. C. (1894) The typical Ornithopoda of the American Jurassic. American Journal of Science, 48, 85 - 90." type="journal article" year="1894">Marsh 1894</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="A7AB4D7CB114FFF1CE6057F3FB90FB30" authority="Gilmore 1909" authorityName="Gilmore" authorityYear="1909" box="[846,1139,1165,1189]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="browni">
<emphasis id="52DFEAEDB114FFF1CE6057F3FC82FB30" box="[846,865,1166,1189]" italics="true" pageId="4" pageNumber="56">C</emphasis>
.
<emphasis id="52DFEAEDB114FFF1CE4157F0FC5DFB30" box="[879,958,1165,1189]" italics="true" pageId="4" pageNumber="56">browni</emphasis>
(
<bibRefCitation id="043A4B0EB114FFF1CEE257F0FB88FB30" author="Gilmore" box="[972,1131,1165,1189]" pageId="4" pageNumber="56" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
)
</taxonomicName>
were subsequently named based upon material from Quarry 13, demarcated primarily by body size. However, this study follows
<bibRefCitation id="043A4B0EB114FFF1C81F57CCFED9FB78" author="Galton" pageId="4" pageNumber="56" refString="Galton, P. M. &amp; Powell, H. P. (1980) The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology, 23, 411 - 443." type="journal article" year="1980">Galton &amp; Powell (1980)</bibRefCitation>
,
<bibRefCitation id="043A4B0EB114FFF1CC6957ABFE1BFB78" author="Norman" box="[327,504,1237,1261]" pageId="4" pageNumber="56" refString="Norman, D. B. (2004) Basal Iguanodontia. In: Weishampel, D. B., Dodson, P. &amp; Osmolska, H. (eds), The Dinosauria: Second Edition. University of California Press, Berkeley, pp. 413 - 437." type="book chapter" year="2004">Norman (2004)</bibRefCitation>
, and
<bibRefCitation id="043A4B0EB114FFF1CF1857A8FC8EFB78" author="Carpenter" box="[566,877,1237,1261]" pageId="4" pageNumber="56" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson (2008)</bibRefCitation>
in regarding all the
<taxonomicName id="A7AB4D7CB114FFF1C97857ABFB1AFB78" box="[1110,1273,1238,1261]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB114FFF1C97857ABFB1AFB78" box="[1110,1273,1238,1261]" italics="true" pageId="4" pageNumber="56">Camptosaurus</emphasis>
</taxonomicName>
material from Quarry 13 as representing a single species,
<taxonomicName id="A7AB4D7CB114FFF1CF525787FD01FA84" box="[636,738,1273,1297]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB114FFF1CF525787FD6CFA84" box="[636,655,1274,1297]" italics="true" pageId="4" pageNumber="56">C</emphasis>
.
<emphasis id="52DFEAEDB114FFF1CFB25784FD01FA84" box="[668,738,1273,1297]" italics="true" pageId="4" pageNumber="56">dispar</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="601436FFB114FFF1CDE25660FD0BF9C0" blockId="4.[151,1437,481,1621]" pageId="4" pageNumber="56">
The ilium of YPM PU14553 is identical to those of
<taxonomicName id="A7AB4D7CB114FFF1CE3D5663FC98FAA0" box="[787,891,1309,1333]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB114FFF1CE3D5663FCC5FAA0" box="[787,806,1310,1333]" italics="true" pageId="4" pageNumber="56">C</emphasis>
.
<emphasis id="52DFEAEDB114FFF1CE1D5660FC98FAA0" box="[819,891,1309,1333]" italics="true" pageId="4" pageNumber="56">dispar</emphasis>
</taxonomicName>
from Quarry 13 (e.g., USNM 5473, 5818, 5959, 7631; YPM 1877, 1880, 8619, 8622), most importantly in having a straight dorsal margin, differing from the convex dorsal margins exhibited by ilia of
<taxonomicName id="A7AB4D7CB114FFF1CF615618FCADFAE8" box="[591,846,1381,1405]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB114FFF1CF615618FCADFAE8" box="[591,846,1381,1405]" italics="true" pageId="4" pageNumber="56">Uteodon aphanoecetes</emphasis>
</taxonomicName>
and
<emphasis id="52DFEAEDB114FFF1CEAB561BFB99FAE8" box="[901,1146,1381,1405]" italics="true" pageId="4" pageNumber="56">
Cumnoria
<taxonomicName id="A7AB4D7CB114FFF1CED05618FB99FAE8" box="[1022,1146,1381,1405]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
(
<figureCitation id="F8902A7AB114FFF1C9A75618FB2CFAE8" box="[1161,1231,1381,1405]" captionStart="FIGURE 4" captionStartId="6.[151,250,720,743]" captionTargetBox="[151,1436,193,698]" captionTargetId="figure@6.[151,1436,193,699]" captionTargetPageId="6" captionText="FIGURE 4. Ilia of Camptosaurus dispar in lateral view. (A) right ilium of YPM 1877 (holotype) from Quarry 13; (B) right ilium of YPM 8622 from Quarry 13; (C) left ilium of USNM 5473 from Quarry 13; (D) left ilium of YPM PU 14553 from Cleveland-Lloyd Dinosaur Quarry. Abbreviations: mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process. Images in A, B, and D are copyright YPM. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200815/files/figure.png" pageId="4" pageNumber="56">Fig. 4</figureCitation>
;
<figureCitation id="F8902A7AB114FFF1C9F55618FAC4FAE8" box="[1243,1319,1381,1405]" captionStart="FIGURE 6" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1424,927,1709]" captionTargetId="figure@7.[151,1436,913,1723]" captionTargetPageId="7" captionText="FIGURE 6. Left ilium of CM 11337 (holotype of Uteodon aphanoecetes) in (A) ventrolateral view showing the convex dorsal margin, (B) lateral view, (C), caudal view. Sacrum and ilia of CM 11337 in (D) dorsal view. Right ilium of CM 11337 in (E) lateral view. Right ilium of OXFUM J. 3303 (holotype of Cumnoria prestwichii) in (F) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process." httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="4" pageNumber="56">Figs. 6</figureCitation>
A, F). The other preserved elements (dentary, pubis, and ischium) are also quite similar to those of Quarry
<specimenCount id="76ADFD76B114FFF1C9EB56F4FAB8FA34" box="[1221,1371,1417,1441]" pageId="4" pageNumber="56" type="generic">13 specimens</specimenCount>
(e.g., YPM 1886, YPM 7334, and USNM 4697, respectively). YPM PU14553 is thus referred to
<taxonomicName id="A7AB4D7CB114FFF1C98A56D3FA76FA50" box="[1188,1429,1453,1477]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB114FFF1C98A56D3FA76FA50" box="[1188,1429,1453,1477]" italics="true" pageId="4" pageNumber="56">Camptosaurus dispar</emphasis>
</taxonomicName>
. This referral is based upon shared morphologies only; provenance is not taken into consideration. The coding of
<taxonomicName id="A7AB4D7CB114FFF1CDB9568BFE69F998" box="[151,394,1525,1549]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB114FFF1CDB9568BFE69F998" box="[151,394,1525,1549]" italics="true" pageId="4" pageNumber="56">Camptosaurus dispar</emphasis>
</taxonomicName>
for the phylogenetic analysis was based almost entirely upon the material from Quarry 13; the predentary of YPM PU14553 was used to code
<taxonomicName id="A7AB4D7CB114FFF1CFCF5567FCA9F9A4" box="[737,842,1561,1585]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB114FFF1CFCF5567FD17F9A4" box="[737,756,1562,1585]" italics="true" pageId="4" pageNumber="56">C</emphasis>
.
<emphasis id="52DFEAEDB114FFF1CE2D5564FCA9F9A4" box="[771,842,1561,1585]" italics="true" pageId="4" pageNumber="56">dispar</emphasis>
</taxonomicName>
for characters pertaining to that element because no predentaries are present in the Quarry 13 collections.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="739E4A7467FBDBB92CA48A9F2F0C3DD4">The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)</mods:title>
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<treatment id="E80287E9B114FFFDCDB95449FB43FDA9" ID-DOI="http://doi.org/10.5281/zenodo.6185213" ID-GBIF-Taxon="119391381" ID-Zenodo-Dep="6185213" LSID="urn:lsid:plazi:treatment:E80287E9B114FFFDCDB95449FB43FDA9" httpUri="http://treatment.plazi.org/id/E80287E9B114FFFDCDB95449FB43FDA9" lastPageId="8" lastPageNumber="59" pageId="4" pageNumber="56">
<subSubSection id="28B16574B114FFF1CDB95449FCD3F805" pageId="4" pageNumber="56" type="nomenclature">
<paragraph id="601436FFB114FFF1CDB95449FC6EF8D8" blockId="4.[151,909,1843,1870]" box="[151,909,1843,1870]" pageId="4" pageNumber="56">
<heading id="3B5C8193B114FFF1CDB95449FC6EF8D8" bold="true" box="[151,909,1843,1870]" fontSize="11" level="1" pageId="4" pageNumber="56" reason="1">
<emphasis id="52DFEAEDB114FFF1CDB95449FC6EF8D8" bold="true" box="[151,909,1843,1870]" pageId="4" pageNumber="56">
<taxonomicName id="A7AB4D7CB114FFF1CDB95449FD19F8D8" authority="Carpenter &amp; Wilson, 2008" authorityName="Carpenter &amp; Wilson" authorityYear="2008" box="[151,762,1843,1870]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="aphanoecetes" status="comb. nov.">
<emphasis id="52DFEAEDB114FFF1CDB95449FE4BF8DB" bold="true" box="[151,424,1844,1870]" italics="true" pageId="4" pageNumber="56">Uteodon aphanoecetes</emphasis>
<bibRefCitation id="043A4B0EB114FFF1CC81544EFD19F8D8" author="Carpenter" box="[431,762,1843,1869]" pageId="4" pageNumber="56" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson, 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="49EC5796B114FFF1CE29544EFC6EF8D8" box="[775,909,1843,1869]" pageId="4" pageNumber="56">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="601436FFB114FFF1CDB95406FCD3F805" blockId="4.[151,816,1914,1936]" box="[151,816,1914,1936]" pageId="4" pageNumber="56">
2008
<taxonomicName id="A7AB4D7CB114FFF1CDC95406FD03F805" authority="Carpenter &amp; Wilson" authorityName="Carpenter &amp; Wilson" box="[231,736,1914,1936]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="4" pageNumber="56" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB114FFF1CDC95406FDE8F805" box="[231,523,1914,1936]" italics="true" pageId="4" pageNumber="56">Camptosaurus aphanoecetes</emphasis>
Carpenter &amp; Wilson
</taxonomicName>
, p. 232.
</paragraph>
</subSubSection>
<subSubSection id="28B16574B114FFF3CDB954C0FABCF831" lastPageId="6" lastPageNumber="58" pageId="4" pageNumber="56" type="materials_examined">
<paragraph id="601436FFB114FFF1CDB954C0FD07F840" blockId="4.[151,740,1981,2005]" box="[151,740,1981,2005]" pageId="4" pageNumber="56">
<emphasis id="52DFEAEDB114FFF1CDB954C0FEE8F840" bold="true" box="[151,267,1981,2005]" pageId="4" pageNumber="56">
<typeStatus id="BF10885DB114FFF1CDB954C0FEE5F840" box="[151,262,1981,2005]" pageId="4" pageNumber="56" type="holotype">Holotype</typeStatus>
.
</emphasis>
CM 11337, nearly complete postcranium.
</paragraph>
<caption id="34D46677B115FFF0CDB954FEFB54F860" httpUri="https://zenodo.org/record/200814/files/figure.png" pageId="5" pageNumber="57" targetBox="[218,1348,194,1877]" targetPageId="5">
<paragraph id="601436FFB115FFF0CDB954FEFB54F860" blockId="5.[151,1436,1923,2037]" pageId="5" pageNumber="57">
<emphasis id="52DFEAEDB115FFF0CDB954FEFEF1F80D" bold="true" box="[151,274,1923,1946]" pageId="5" pageNumber="57">FIGURE 3.</emphasis>
(
<emphasis id="52DFEAEDB115FFF0CC0E54FEFED1F80D" bold="true" box="[288,306,1923,1944]" pageId="5" pageNumber="57">A</emphasis>
) USNM 4282, left scapula of
<taxonomicName id="A7AB4D7CB115FFF0CF4C54F9FCD8F80C" box="[610,827,1923,1945]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="57" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB115FFF0CF4C54F9FCD8F80C" box="[610,827,1923,1945]" italics="true" pageId="5" pageNumber="57">Camptosaurus dispar</emphasis>
</taxonomicName>
in lateral view; (
<emphasis id="52DFEAEDB115FFF0CEC854F9FC14F80C" bold="true" box="[998,1015,1924,1945]" pageId="5" pageNumber="57">B</emphasis>
) CM 11337, cranial end of the right scapula of
<taxonomicName id="A7AB4D7CB115FFF0CDCF54DFFE2EF82D" box="[225,461,1954,1976]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="57" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB115FFF0CDCF54DFFE2EF82D" box="[225,461,1954,1976]" italics="true" pageId="5" pageNumber="57">Uteodon aphanoecetes</emphasis>
</taxonomicName>
in lateral view; (
<emphasis id="52DFEAEDB115FFF0CFA254DCFD7DF82D" bold="true" box="[652,670,1953,1976]" pageId="5" pageNumber="57">C</emphasis>
) CM 11337, left scapula of
<taxonomicName id="A7AB4D7CB115FFF0CEF854DFFB21F82D" box="[982,1218,1954,1976]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="57" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB115FFF0CEF854DFFB21F82D" box="[982,1218,1954,1976]" italics="true" pageId="5" pageNumber="57">Uteodon aphanoecetes</emphasis>
</taxonomicName>
in lateral view; (
<emphasis id="52DFEAEDB115FFF0C8AF54DFFA70F822" bold="true" box="[1409,1427,1954,1975]" pageId="5" pageNumber="57">D</emphasis>
) OXFUM J.3303, right scapula of
<emphasis id="52DFEAEDB115FFF0CCC054BFFD2FF842" box="[494,716,1985,2007]" italics="true" pageId="5" pageNumber="57">
Cumnoria
<taxonomicName id="A7AB4D7CB115FFF0CF7254BCFD2FF842" box="[604,716,1985,2007]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="5" pageNumber="57" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
in lateral view. Abbreviations: cap, convex cranial margin of the acromion process; tap, triangular acromion process. Scale bar in
<emphasis id="52DFEAEDB115FFF0CFD5549DFCEEF860" bold="true" box="[763,781,2016,2037]" pageId="5" pageNumber="57">A</emphasis>
equals 10 cm; scale bar in
<emphasis id="52DFEAEDB115FFF0C90E549DFBD2F860" bold="true" box="[1056,1073,2016,2037]" pageId="5" pageNumber="57">B</emphasis>
equals 5 cm.
</paragraph>
</caption>
<caption id="34D46677B116FFF3CDB951ADFBD1FCD7" httpUri="https://zenodo.org/record/200815/files/figure.png" pageId="6" pageNumber="58" targetBox="[151,1436,193,698]" targetPageId="6">
<paragraph id="601436FFB116FFF3CDB951ADFBD1FCD7" blockId="6.[151,1436,720,834]" pageId="6" pageNumber="58">
<emphasis id="52DFEAEDB116FFF3CDB951ADFEF7FD70" bold="true" box="[151,276,720,743]" pageId="6" pageNumber="58">FIGURE 4.</emphasis>
Ilia of
<taxonomicName id="A7AB4D7CB116FFF3CC4D51ACFDA3FD73" box="[355,576,720,742]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB116FFF3CC4D51ACFDA3FD73" box="[355,576,720,742]" italics="true" pageId="6" pageNumber="58">Camptosaurus dispar</emphasis>
</taxonomicName>
in lateral view. (
<emphasis id="52DFEAEDB116FFF3CFDF51ADFCE0FD70" bold="true" box="[753,771,720,741]" pageId="6" pageNumber="58">A</emphasis>
) right ilium of YPM 1877 (holotype) from Quarry 13; (
<emphasis id="52DFEAEDB116FFF3C86551ACFABFFD73" bold="true" box="[1355,1372,721,742]" pageId="6" pageNumber="58">B</emphasis>
) right ilium of YPM 8622 from Quarry 13; (
<emphasis id="52DFEAEDB116FFF3CF065193FDD9FC90" bold="true" box="[552,570,750,773]" pageId="6" pageNumber="58">C</emphasis>
) left ilium of USNM 5473 from Quarry 13; (
<emphasis id="52DFEAEDB116FFF3C9345192FBCFFC91" bold="true" box="[1050,1068,751,772]" pageId="6" pageNumber="58">D</emphasis>
) left ilium of YPM PU14553 from Cleveland-Lloyd Dinosaur Quarry. Abbreviations: mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process. Images in
<emphasis id="52DFEAEDB116FFF3CCDF5051FDE0FCD4" bold="true" box="[497,515,812,833]" pageId="6" pageNumber="58">A</emphasis>
,
<emphasis id="52DFEAEDB116FFF3CF3E5051FDC2FCD4" bold="true" box="[528,545,812,833]" pageId="6" pageNumber="58">B</emphasis>
, and
<emphasis id="52DFEAEDB116FFF3CF775051FD88FCD4" bold="true" box="[601,619,812,833]" pageId="6" pageNumber="58">D</emphasis>
are copyright YPM. Scale bar equals 10 cm.
</paragraph>
</caption>
<caption id="34D46677B116FFF3CDB9564FFC2DFAFD" httpUri="https://zenodo.org/record/200816/files/figure.png" pageId="6" pageNumber="58" targetBox="[483,1104,894,1304]" targetPageId="6">
<paragraph id="601436FFB116FFF3CDB9564FFC2DFAFD" blockId="6.[151,1436,1330,1384]" pageId="6" pageNumber="58">
<emphasis id="52DFEAEDB116FFF3CDB9564FFEF6FADD" bold="true" box="[151,277,1330,1353]" pageId="6" pageNumber="58">FIGURE 5.</emphasis>
YPM 7334, right pubis of
<taxonomicName id="A7AB4D7CB116FFF3CF1D5649FCECFADC" box="[563,783,1331,1353]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB116FFF3CF1D5649FCECFADC" box="[563,783,1331,1353]" italics="true" pageId="6" pageNumber="58">Camptosaurus dispar</emphasis>
</taxonomicName>
from Quarry 13 in lateral view. Abbreviations: pop, postpubic process; prp, prepubic process. Image is copyright YPM. Scale bar equals 10 cm.
</paragraph>
</caption>
<paragraph id="601436FFB116FFF3CDEB56E8FC67FA39" blockId="6.[151,1436,1428,2029]" box="[197,900,1428,1453]" pageId="6" pageNumber="58">
<emphasis id="52DFEAEDB116FFF3CDEB56E8FEA7FA38" bold="true" box="[197,324,1429,1453]" pageId="6" pageNumber="58">
<typeStatus id="BF10885DB116FFF3CDEB56E8FEDCFA38" box="[197,319,1429,1453]" pageId="6" pageNumber="58" type="paratype">Paratypes</typeStatus>
.
</emphasis>
CM 15780, 41689, 79050;
<collectionCode id="06BAAE3AB116FFF3CF5456E9FD23FA39" box="[634,704,1428,1452]" httpUri="http://grbio.org/cool/h4n0-drs5" name="Dinosaur National Monument" pageId="6" pageNumber="58">DINO</collectionCode>
556, 1030, 1032.
</paragraph>
<paragraph id="601436FFB116FFF3CDEB56C5FA95FA45" blockId="6.[151,1436,1428,2029]" box="[197,1398,1464,1489]" pageId="6" pageNumber="58">
<emphasis id="52DFEAEDB116FFF3CDEB56C5FE40FA44" bold="true" box="[197,419,1464,1489]" pageId="6" pageNumber="58">Referred material.</emphasis>
Disarticulated elements held in CM and
<collectionCode id="06BAAE3AB116FFF3CE4056C5FC57FA45" box="[878,948,1464,1488]" httpUri="http://grbio.org/cool/h4n0-drs5" name="Dinosaur National Monument" pageId="6" pageNumber="58">DINO</collectionCode>
collections (
<bibRefCitation id="043A4B0EB116FFF3C96D56C5FA88FA45" author="Carpenter" box="[1091,1387,1464,1488]" pageId="6" pageNumber="58" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson 2008</bibRefCitation>
).
</paragraph>
<paragraph id="601436FFB116FFF3CDEB56A1FF00F9F5" blockId="6.[151,1436,1428,2029]" pageId="6" pageNumber="58">
<emphasis id="52DFEAEDB116FFF3CDEB56A1FE48FA60" bold="true" box="[197,427,1500,1525]" pageId="6" pageNumber="58">Generic etymology.</emphasis>
<taxonomicName id="A7AB4D7CB116FFF3CC9F56A3FE39FA60" box="[433,474,1502,1525]" class="Calcarea" family="Grantiidae" genus="Ute" kingdom="Animalia" order="Leucosolenida" pageId="6" pageNumber="58" phylum="Porifera" rank="genus">
<emphasis id="52DFEAEDB116FFF3CC9F56A3FE39FA60" box="[433,474,1502,1525]" italics="true" pageId="6" pageNumber="58">Ute</emphasis>
</taxonomicName>
(pronounced “yewt”) is the name of a Native American people who inhabit northeastern Utah, in the vicinity of Dinosaur National Monument.
<emphasis id="52DFEAEDB116FFF3CE68557DFC66F98D" box="[838,901,1536,1560]" italics="true" pageId="6" pageNumber="58">Odon</emphasis>
comes from the transliterated Greek meaning “tooth”, and is a common root in the generic names of basal iguanodonts. The gender of the generic name is masculine.
</paragraph>
<paragraph id="601436FFB116FFF3CDEB5511FBAAF93D" blockId="6.[151,1436,1428,2029]" pageId="6" pageNumber="58">
<emphasis id="52DFEAEDB116FFF3CDEB5511FE25F910" bold="true" box="[197,454,1644,1669]" pageId="6" pageNumber="58">Locality and horizon.</emphasis>
Carnegie Quarry at Dinosaur National Monument, Uintah County, Utah; Brushy Basin Member of the Morrison Formation, lower–middle Tithonian (
<bibRefCitation id="043A4B0EB116FFF3CE7D55EDFBDFF93D" author="Kowallis" box="[851,1084,1680,1704]" pageId="6" pageNumber="58" refString="Kowallis, B. J., Christiansen, E. H., Deino, A. L., Peterson, F., Turner, C. E., Kunk, M. J. &amp; Obradovich, J. D. (1998) The age of the Morrison Formation. Modern Geology, 22, 235 - 260." type="journal article" year="1998">
Kowallis
<emphasis id="52DFEAEDB116FFF3CEEF55ECFC12F93D" box="[961,1009,1680,1704]" italics="true" pageId="6" pageNumber="58">et al</emphasis>
. 1998
</bibRefCitation>
).
</paragraph>
<paragraph id="601436FFB116FFF3CDEB55C9FCF3F958" blockId="6.[151,1436,1428,2029]" box="[197,784,1716,1741]" pageId="6" pageNumber="58">
<emphasis id="52DFEAEDB116FFF3CDEB55C9FE43F958" bold="true" box="[197,416,1716,1741]" pageId="6" pageNumber="58">Generic diagnosis.</emphasis>
<collectingCountry id="18BC766FB116FFF3CC8955C8FE25F958" box="[423,454,1717,1741]" name="American Samoa" pageId="6" pageNumber="58">As</collectingCountry>
for the species by monotypy.
</paragraph>
<paragraph id="601436FFB116FFF3CDEB55A5FABCF831" blockId="6.[151,1436,1428,2029]" pageId="6" pageNumber="58">
<emphasis id="52DFEAEDB116FFF3CDEB55A5FE42F964" bold="true" box="[197,417,1752,1777]" pageId="6" pageNumber="58">Specific diagnosis.</emphasis>
Basal styracosternan distinguished by a single autapomorphy: occipital condyle projects farther ventrally than the basal tubera (
<figureCitation id="F8902A7AB116FFF3CF605581FD70F881" box="[590,659,1788,1812]" captionStart="FIGURE 7" captionStartId="8.[151,250,1622,1645]" captionTargetBox="[472,1114,625,1600]" captionTargetId="figure@8.[472,1115,625,1601]" captionTargetPageId="8" captionText="FIGURE 7. Partial braincases of Camptosaurus dispar, Uteodon aphanoecetes, and Cumnoria prestwichii. (A) YPM 1880 in left lateral view (image copyright YPM); (B) CM 15780 in right lateral view (image courtesy of M. Lamanna); (C) OXFUM J. 3303 in right lateral view. Abbreviations: bt, basal tubera; oc, occipital condyle. Scale bar equals 5 cm." httpUri="https://zenodo.org/record/200818/files/figure.png" pageId="6" pageNumber="58">Fig. 7</figureCitation>
). Also distinguished from all other iguanodontians except
<emphasis id="52DFEAEDB116FFF3C8045580FEF0F8AD" italics="true" pageId="6" pageNumber="58">
Cumnoria
<taxonomicName id="A7AB4D7CB116FFF3CDB9545DFEF0F8AD" box="[151,275,1824,1848]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
by the following unique combination of characters: scapula with acromion process that is convex along its cranial margin (
<figureCitation id="F8902A7AB116FFF3CC475439FE4FF8C9" box="[361,428,1860,1884]" captionStart="FIGURE 3" captionStartId="5.[151,250,1923,1946]" captionTargetBox="[218,1348,194,1877]" captionTargetId="figure@5.[213,1359,193,1878]" captionTargetPageId="5" captionText="FIGURE 3. (A) USNM 4282, left scapula of Camptosaurus dispar in lateral view; (B) CM 11337, cranial end of the right scapula of Uteodon aphanoecetes in lateral view; (C) CM 11337, left scapula of Uteodon aphanoecetes in lateral view; (D) OXFUM J. 3303, right scapula of Cumnoria prestwichii in lateral view. Abbreviations: cap, convex cranial margin of the acromion process; tap, triangular acromion process. Scale bar in A equals 10 cm; scale bar in B equals 5 cm." httpUri="https://zenodo.org/record/200814/files/figure.png" pageId="6" pageNumber="58">Fig. 3</figureCitation>
B), convex dorsal margin of scapula (
<figureCitation id="F8902A7AB116FFF3CE7E5439FC71F8C9" box="[848,914,1860,1884]" captionStart="FIGURE 3" captionStartId="5.[151,250,1923,1946]" captionTargetBox="[218,1348,194,1877]" captionTargetId="figure@5.[213,1359,193,1878]" captionTargetPageId="5" captionText="FIGURE 3. (A) USNM 4282, left scapula of Camptosaurus dispar in lateral view; (B) CM 11337, cranial end of the right scapula of Uteodon aphanoecetes in lateral view; (C) CM 11337, left scapula of Uteodon aphanoecetes in lateral view; (D) OXFUM J. 3303, right scapula of Cumnoria prestwichii in lateral view. Abbreviations: cap, convex cranial margin of the acromion process; tap, triangular acromion process. Scale bar in A equals 10 cm; scale bar in B equals 5 cm." httpUri="https://zenodo.org/record/200814/files/figure.png" pageId="6" pageNumber="58">Fig. 3</figureCitation>
C), convex dorsal margin of ilium, dorsal margin of ilium thickens mediolaterally towards
<emphasis id="52DFEAEDB116FFF3CFB65414FD4CF815" box="[664,687,1897,1920]" italics="true" pageId="6" pageNumber="58">M</emphasis>
.
<emphasis id="52DFEAEDB116FFF3CF915415FCA0F815" box="[703,835,1896,1920]" italics="true" pageId="6" pageNumber="58">iliocaudalis</emphasis>
platform, postacetabular process of ilium tapers to a point with break in slope along dorsal margin (
<figureCitation id="F8902A7AB116FFF3CF8D54F1FD06F831" box="[675,741,1932,1956]" captionStart="FIGURE 6" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1424,927,1709]" captionTargetId="figure@7.[151,1436,913,1723]" captionTargetPageId="7" captionText="FIGURE 6. Left ilium of CM 11337 (holotype of Uteodon aphanoecetes) in (A) ventrolateral view showing the convex dorsal margin, (B) lateral view, (C), caudal view. Sacrum and ilia of CM 11337 in (D) dorsal view. Right ilium of CM 11337 in (E) lateral view. Right ilium of OXFUM J. 3303 (holotype of Cumnoria prestwichii) in (F) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process." httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="6" pageNumber="58">Fig. 6</figureCitation>
A), distal end of ischium forms cranially expanded boot.
</paragraph>
</subSubSection>
<subSubSection id="28B16574B116FFFDCDEB54CCFB43FDA9" lastPageId="8" lastPageNumber="60" pageId="6" pageNumber="58" type="discussion">
<paragraph id="601436FFB116FFF2CDEB54CCFD29FE5A" blockId="6.[151,1436,1428,2029]" lastBlockId="7.[151,1437,151,860]" lastPageId="7" lastPageNumber="59" pageId="6" pageNumber="58">
<emphasis id="52DFEAEDB116FFF3CDEB54CCFEDAF85C" bold="true" box="[197,313,1969,1993]" pageId="6" pageNumber="58">Remarks.</emphasis>
The
<typeStatus id="BF10885DB116FFF3CC5054CDFE03F85D" box="[382,480,1968,1992]" pageId="6" pageNumber="58" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A7AB4D7CB116FFF3CF3C54CDFCF4F85D" box="[530,791,1968,1992]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB116FFF3CF3C54CDFCF4F85D" box="[530,791,1968,1992]" italics="true" pageId="6" pageNumber="58">Uteodon aphanoecetes</emphasis>
</taxonomicName>
, CM 11337, was originally referred to
<taxonomicName id="A7AB4D7CB116FFF3C9D754CCFA7FF85D" box="[1273,1436,1969,1992]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB116FFF3C9D754CCFA7FF85D" box="[1273,1436,1969,1992]" italics="true" pageId="6" pageNumber="58">Camptosaurus</emphasis>
</taxonomicName>
“
<taxonomicName id="A7AB4D7CB116FFF3CD8D54A8FF11F878" box="[163,242,2005,2029]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="medius">
<emphasis id="52DFEAEDB116FFF3CD8D54A8FF11F878" box="[163,242,2005,2029]" italics="true" pageId="6" pageNumber="58">medius</emphasis>
</taxonomicName>
” (=
<taxonomicName id="A7AB4D7CB116FFF3CC0654A8FE72F878" box="[296,401,2005,2029]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="6" pageNumber="58" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB116FFF3CC0654A8FED8F879" box="[296,315,2005,2028]" italics="true" pageId="6" pageNumber="58">C</emphasis>
.
<emphasis id="52DFEAEDB116FFF3CC6454A8FE72F878" box="[330,401,2005,2029]" italics="true" pageId="6" pageNumber="58">dispar</emphasis>
</taxonomicName>
) by
<bibRefCitation id="043A4B0EB116FFF3CCE654A9FD9FF879" author="Gilmore" box="[456,636,2004,2028]" pageId="6" pageNumber="58" refString="Gilmore, C. W. (1925) Osteology of ornithopodous dinosaurs from the Dinosaur National Monument, Utah. Memoirs of the Carnegie Museum, 10, 385 - 411." type="journal article" year="1925">Gilmore (1925)</bibRefCitation>
.
<bibRefCitation id="043A4B0EB116FFF3CFA954A9FC5DF879" author="Carpenter" box="[647,958,2004,2028]" pageId="6" pageNumber="58" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson (2008)</bibRefCitation>
reassessed the specimen and found that it differed sufficiently from specimens of
<taxonomicName id="A7AB4D7CB117FFF2CF7953E5FD23FF25" box="[599,704,152,176]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB117FFF2CF7953E5FD89FF3A" box="[599,618,152,175]" italics="true" pageId="7" pageNumber="59">C</emphasis>
.
<emphasis id="52DFEAEDB117FFF2CF5753E5FD23FF25" box="[633,704,152,176]" italics="true" pageId="7" pageNumber="59">dispar</emphasis>
</taxonomicName>
to warrant the creation of the new species
<taxonomicName id="A7AB4D7CB117FFF2C98653E5FA82FF25" box="[1192,1377,152,176]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB117FFF2C98653E5FB58FF3A" box="[1192,1211,152,175]" italics="true" pageId="7" pageNumber="59">C</emphasis>
.
<emphasis id="52DFEAEDB117FFF2C9E753E5FA82FF25" box="[1225,1377,152,176]" italics="true" pageId="7" pageNumber="59">aphanoecetes</emphasis>
</taxonomicName>
. The new phylogenetic analysis supports the distinction of
<taxonomicName id="A7AB4D7CB117FFF2CFC053C1FC0EFF41" box="[750,1005,188,212]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB117FFF2CFC053C1FC0EFF41" box="[750,1005,188,212]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
</taxonomicName>
from
<taxonomicName id="A7AB4D7CB117FFF2C91F53C0FB7BFF41" box="[1073,1176,188,212]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB117FFF2C91F53C0FBA7FF41" box="[1073,1092,189,212]" italics="true" pageId="7" pageNumber="59">C</emphasis>
.
<emphasis id="52DFEAEDB117FFF2C97C53C1FB7BFF41" box="[1106,1176,188,212]" italics="true" pageId="7" pageNumber="59">dispar</emphasis>
</taxonomicName>
, but does not recover a monophyletic
<taxonomicName id="A7AB4D7CB117FFF2CC12539DFE3CFF62" box="[316,479,224,247]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB117FFF2CC12539DFE3CFF62" box="[316,479,224,247]" italics="true" pageId="7" pageNumber="59">Camptosaurus</emphasis>
</taxonomicName>
as conceived by
<bibRefCitation id="043A4B0EB117FFF2CF9E53A2FC10FF62" author="Carpenter" box="[688,1011,223,247]" pageId="7" pageNumber="59" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson (2008)</bibRefCitation>
. Instead,
<taxonomicName id="A7AB4D7CB117FFF2C94A539DFA8BFF6D" box="[1124,1384,224,248]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB117FFF2C94A539DFA8BFF6D" box="[1124,1384,224,248]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
</taxonomicName>
and
<emphasis id="52DFEAEDB117FFF2CDB95278FE68FE89" box="[151,395,260,284]" italics="true" pageId="7" pageNumber="59">
Cumnoria
<taxonomicName id="A7AB4D7CB117FFF2CC3E5279FE68FE89" box="[272,395,260,284]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
(see below) are more derived than
<taxonomicName id="A7AB4D7CB117FFF2CE355278FC60FE89" box="[795,899,260,284]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB117FFF2CE355278FCCDFE89" box="[795,814,261,284]" italics="true" pageId="7" pageNumber="59">C</emphasis>
.
<emphasis id="52DFEAEDB117FFF2CE125279FC60FE89" box="[828,899,260,284]" italics="true" pageId="7" pageNumber="59">dispar</emphasis>
</taxonomicName>
in the reduced consensus tree and are therefore the most basal members of Styracosterna, which is defined as those species more closely related to
<taxonomicName id="A7AB4D7CB117FFF2C8285255FEC4FEF1" class="Reptilia" family="Hadrosauridae" genus="Parasaurolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="walkeri">
<emphasis id="52DFEAEDB117FFF2C8285255FEC4FEF1" italics="true" pageId="7" pageNumber="59">Parasaurolophus walkeri</emphasis>
</taxonomicName>
than to
<taxonomicName id="A7AB4D7CB117FFF2CCAA5230FCFDFEF1" authority="Sereno 2005" authorityName="Sereno" authorityYear="2005" box="[388,798,332,356]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB117FFF2CCAA5230FD94FEF1" box="[388,631,332,356]" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
(
<bibRefCitation id="043A4B0EB117FFF2CFA85231FCF5FEF1" author="Sereno" box="[646,790,332,356]" pageId="7" pageNumber="59" refString="Sereno, P. C. (2005) Stem Archosauria version 1. 0, TaxonSearch. Available from: http: // www. taxonsearch. org / Archive / stemarchosauria- 1.0. php (21 April 2009)." type="journal article" year="2005">Sereno 2005</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="F8902A7AB117FFF2CE005231FC93FEF1" box="[814,880,332,356]" captionStart="FIGURE 1" captionStartId="2.[151,250,1854,1877]" captionTargetBox="[207,1262,450,1790]" captionTargetId="figure@2.[191,1274,445,1796]" captionTargetPageId="2" captionText="FIGURE 1. Reduced consensus tree of 11,850 MPTs of 358 steps each, following ordering of 22 multistate characters and safe taxonomic reduction. Camptosaurus dispar is highlighted in yellow and Uteodon aphanoecetes and Cumnoria prestwichii in blue. Numbers below and to the left of some nodes correspond to the following clade names: 1, Ornithopoda; 2, Iguanodontia; 3, Rhabdodontidae; 4, Dryomorpha; 5, Dryosauridae; 6, Ankylopollexia; 7, Styracosterna; 8, Hadrosauriformes; 9, Hadrosauroidea." httpUri="https://zenodo.org/record/200812/files/figure.png" pageId="7" pageNumber="59">Fig. 1</figureCitation>
). Furthermore,
<taxonomicName id="A7AB4D7CB117FFF2C90D5231FB60FEF1" box="[1059,1155,332,356]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="66" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="52DFEAEDB117FFF2C90D5231FB60FEF1" box="[1059,1155,332,356]" italics="true" pageId="7" pageNumber="59">Uteodon</emphasis>
</taxonomicName>
is also found to be more derived than
<taxonomicName id="A7AB4D7CB117FFF2CC04520DFE70FE1D" box="[298,403,368,392]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB117FFF2CC04520DFEDEFE12" box="[298,317,368,391]" italics="true" pageId="7" pageNumber="59">C</emphasis>
.
<emphasis id="52DFEAEDB117FFF2CC65520DFE70FE1D" box="[331,403,368,392]" italics="true" pageId="7" pageNumber="59">dispar</emphasis>
</taxonomicName>
in the agreement subtree (
<figureCitation id="F8902A7AB117FFF2CF935212FCE1FE12" box="[701,770,367,391]" captionStart="FIGURE 2" captionStartId="3.[151,250,1430,1453]" captionTargetBox="[296,1231,202,1382]" captionTargetId="figure@3.[275,1239,193,1396]" captionTargetPageId="3" captionText="FIGURE 2. Agreement subtree of 11,850 MPTs of 358 steps each, following ordering of 22 multistate characters and safe taxonomic reduction. Camptosaurus dispar is highlighted in yellow and Uteodon aphanoecetes in blue. Numbers below and to the left of some nodes correspond to the following clade names: 1, Iguanodontia; 2, Rhabdodontidae; 3, Dryomorpha; 4, Dryosauridae; 5, Ankylopollexia; 6, Styracosterna; 7, Hadrosauriformes; 8, Hadrosauroidea; 9, Hadrosauridae." httpUri="https://zenodo.org/record/200813/files/figure.png" pageId="7" pageNumber="59">Fig. 2</figureCitation>
). For the analysis,
<taxonomicName id="A7AB4D7CB117FFF2CEF9520DFB34FE1D" box="[983,1239,368,392]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB117FFF2CEF9520DFB34FE1D" box="[983,1239,368,392]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
</taxonomicName>
was coded based upon the
<typeStatus id="BF10885DB117FFF2CC2C52E9FE84FE39" box="[258,359,404,428]" pageId="7" pageNumber="59" type="holotype">holotype</typeStatus>
,
<typeStatus id="BF10885DB117FFF2CC5D52E8FE02FE38" box="[371,481,405,429]" pageId="7" pageNumber="59" type="paratype">paratypes</typeStatus>
, and referred specimens designated by
<bibRefCitation id="043A4B0EB117FFF2CE8E52E9FB38FE39" author="Carpenter" box="[928,1243,404,428]" pageId="7" pageNumber="59" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson (2008)</bibRefCitation>
. Only specimens in the CM collection could be examined firsthand.
</paragraph>
<paragraph id="601436FFB117FFF2CDE952A1FECAFCC9" blockId="7.[151,1437,151,860]" pageId="7" pageNumber="59">
<taxonomicName id="A7AB4D7CB117FFF2CDE952A1FE2BFE61" box="[199,456,476,500]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB117FFF2CDE952A1FE2BFE61" box="[199,456,476,500]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
</taxonomicName>
and
<emphasis id="52DFEAEDB117FFF2CF2D52A0FD1AFE61" box="[515,761,476,500]" italics="true" pageId="7" pageNumber="59">
Cumnoria
<taxonomicName id="A7AB4D7CB117FFF2CF5352A1FD1AFE61" box="[637,761,476,500]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
are quite similar to each other and different from
<taxonomicName id="A7AB4D7CB117FFF2C81452A0FED2FD8D" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB117FFF2C81452A0FED2FD8D" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
</taxonomicName>
. The
<typeStatus id="BF10885DB117FFF2CC585282FE3BFD82" box="[374,472,511,535]" pageId="7" pageNumber="59" type="holotype">holotype</typeStatus>
skeletons of
<taxonomicName id="A7AB4D7CB117FFF2CF5D517DFD37FD8D" box="[627,724,512,536]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="66" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="52DFEAEDB117FFF2CF5D517DFD37FD8D" box="[627,724,512,536]" italics="true" pageId="7" pageNumber="59">Uteodon</emphasis>
</taxonomicName>
and
<emphasis id="52DFEAEDB117FFF2CE20517DFC9CFD82" box="[782,895,512,535]" italics="true" pageId="7" pageNumber="59">Cumnoria</emphasis>
both exhibit scapulae with convex dorsal margins, whereas scapulae of
<taxonomicName id="A7AB4D7CB117FFF2CC935158FD4CFDA9" box="[445,687,548,572]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB117FFF2CC935158FD4CFDA9" box="[445,687,548,572]" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
</taxonomicName>
are straight or somewhat concave along their dorsal margins (e.g., USNM 2210, 4282, 5473, 5954, 5955; YPM 8611) (
<figureCitation id="F8902A7AB117FFF2CFCE513AFCC7FDCA" box="[736,804,583,607]" captionStart="FIGURE 3" captionStartId="5.[151,250,1923,1946]" captionTargetBox="[218,1348,194,1877]" captionTargetId="figure@5.[213,1359,193,1878]" captionTargetPageId="5" captionText="FIGURE 3. (A) USNM 4282, left scapula of Camptosaurus dispar in lateral view; (B) CM 11337, cranial end of the right scapula of Uteodon aphanoecetes in lateral view; (C) CM 11337, left scapula of Uteodon aphanoecetes in lateral view; (D) OXFUM J. 3303, right scapula of Cumnoria prestwichii in lateral view. Abbreviations: cap, convex cranial margin of the acromion process; tap, triangular acromion process. Scale bar in A equals 10 cm; scale bar in B equals 5 cm." httpUri="https://zenodo.org/record/200814/files/figure.png" pageId="7" pageNumber="59">Fig. 3</figureCitation>
). Furthermore, as noted above, the ilia of
<emphasis id="52DFEAEDB117FFF2C9D55135FA8EFDCA" box="[1275,1389,584,607]" italics="true" pageId="7" pageNumber="59">Cumnoria</emphasis>
and
<taxonomicName id="A7AB4D7CB117FFF2CDB95111FF1BFD11" box="[151,248,620,644]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="66" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="52DFEAEDB117FFF2CDB95111FF1BFD11" box="[151,248,620,644]" italics="true" pageId="7" pageNumber="59">Uteodon</emphasis>
</taxonomicName>
exhibit convex dorsal margins (
<figureCitation id="F8902A7AB117FFF2CF5E5111FD54FD11" box="[624,695,620,644]" captionStart="FIGURE 6" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1424,927,1709]" captionTargetId="figure@7.[151,1436,913,1723]" captionTargetPageId="7" captionText="FIGURE 6. Left ilium of CM 11337 (holotype of Uteodon aphanoecetes) in (A) ventrolateral view showing the convex dorsal margin, (B) lateral view, (C), caudal view. Sacrum and ilia of CM 11337 in (D) dorsal view. Right ilium of CM 11337 in (E) lateral view. Right ilium of OXFUM J. 3303 (holotype of Cumnoria prestwichii) in (F) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process." httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="7" pageNumber="59">Fig. 6</figureCitation>
), similar to some more derived iguanodonts such as
<taxonomicName id="A7AB4D7CB117FFF2C80D5111FEA2FD3D" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="52DFEAEDB117FFF2C80D5111FEA2FD3D" italics="true" pageId="7" pageNumber="59">Iguanodon bernissartensis</emphasis>
</taxonomicName>
(IRSNB 1534) and
<emphasis id="52DFEAEDB117FFF2CF0F51EDFD1BFD3D" box="[545,760,656,680]" italics="true" pageId="7" pageNumber="59">Equijubus normani</emphasis>
(
<bibRefCitation id="043A4B0EB117FFF2CE2851F2FC54FD32" author="You" box="[774,951,655,680]" pageId="7" pageNumber="59" refString="You, H., Luo, Z., Shubin, N. H., Witmer, L. M., Tang, Z. &amp; Tang, F. (2003) The earliest-known duck-billed dinosaur from deposits of late Early Cretaceous age in northwest China and hadrosaur evolution. Cretaceous Research, 24, 347 - 355." type="journal article" year="2003">
You
<emphasis id="52DFEAEDB117FFF2CE1251ECFC8EFD3D" box="[828,877,656,680]" italics="true" pageId="7" pageNumber="59">et al</emphasis>
. 2003
</bibRefCitation>
), but different from the straight dorsal margin of the ilium of
<taxonomicName id="A7AB4D7CB117FFF2CC4351C8FD83FD59" box="[365,608,692,716]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB117FFF2CC4351C8FD83FD59" box="[365,608,692,716]" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
</taxonomicName>
(
<figureCitation id="F8902A7AB117FFF2CF5E51C9FD55FD59" box="[624,694,692,716]" captionStart="FIGURE 4" captionStartId="6.[151,250,720,743]" captionTargetBox="[151,1436,193,698]" captionTargetId="figure@6.[151,1436,193,699]" captionTargetPageId="6" captionText="FIGURE 4. Ilia of Camptosaurus dispar in lateral view. (A) right ilium of YPM 1877 (holotype) from Quarry 13; (B) right ilium of YPM 8622 from Quarry 13; (C) left ilium of USNM 5473 from Quarry 13; (D) left ilium of YPM PU 14553 from Cleveland-Lloyd Dinosaur Quarry. Abbreviations: mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process. Images in A, B, and D are copyright YPM. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200815/files/figure.png" pageId="7" pageNumber="59">Fig. 4</figureCitation>
). Furthermore,
<taxonomicName id="A7AB4D7CB117FFF2CE4851C9FC25FD59" box="[870,966,692,716]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="66" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="52DFEAEDB117FFF2CE4851C9FC25FD59" box="[870,966,692,716]" italics="true" pageId="7" pageNumber="59">Uteodon</emphasis>
</taxonomicName>
possesses an autapomorphy in the partial braincase of
<typeStatus id="BF10885DB117FFF2CC0951A5FE64FD65" box="[295,391,728,752]" pageId="7" pageNumber="59" type="paratype">paratype</typeStatus>
specimen CM 15780: the occipital condyle projects far ventrally relative to the basal tubera. This condition differs from the basioccipitals of
<taxonomicName id="A7AB4D7CB117FFF2CFE85180FC59FC81" box="[710,954,764,788]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB117FFF2CFE85180FC59FC81" box="[710,954,764,788]" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
</taxonomicName>
(USNM 5473, 5996, 5997; YPM 1856A, 1880) and
<emphasis id="52DFEAEDB117FFF2CC21505DFE62FCA2" box="[271,385,800,823]" italics="true" pageId="7" pageNumber="59">Cumnoria</emphasis>
(OXFUM J.3303), in which the basal tubera protrude farther ventrally than the occipital condyle (
<figureCitation id="F8902A7AB117FFF2CDF85039FEF8FCC9" box="[214,283,836,860]" captionStart="FIGURE 7" captionStartId="8.[151,250,1622,1645]" captionTargetBox="[472,1114,625,1600]" captionTargetId="figure@8.[472,1115,625,1601]" captionTargetPageId="8" captionText="FIGURE 7. Partial braincases of Camptosaurus dispar, Uteodon aphanoecetes, and Cumnoria prestwichii. (A) YPM 1880 in left lateral view (image copyright YPM); (B) CM 15780 in right lateral view (image courtesy of M. Lamanna); (C) OXFUM J. 3303 in right lateral view. Abbreviations: bt, basal tubera; oc, occipital condyle. Scale bar equals 5 cm." httpUri="https://zenodo.org/record/200818/files/figure.png" pageId="7" pageNumber="59">Fig. 7</figureCitation>
).
</paragraph>
<caption id="34D46677B117FFF2CDB955ADFAEAF8D7" httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="7" pageNumber="59" targetBox="[151,1424,927,1709]" targetPageId="7">
<paragraph id="601436FFB117FFF2CDB955ADFAEAF8D7" blockId="7.[151,1436,1744,1858]" pageId="7" pageNumber="59">
<emphasis id="52DFEAEDB117FFF2CDB955ADFEF1F973" bold="true" box="[151,274,1744,1767]" pageId="7" pageNumber="59">FIGURE 6.</emphasis>
Left ilium of CM 11337 (holotype of
<taxonomicName id="A7AB4D7CB117FFF2CFB855ACFC9EF972" box="[662,893,1745,1767]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB117FFF2CFB855ACFC9EF972" box="[662,893,1745,1767]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
</taxonomicName>
) in (
<emphasis id="52DFEAEDB117FFF2CE8055ACFC23F973" bold="true" box="[942,960,1745,1766]" pageId="7" pageNumber="59">A</emphasis>
) ventrolateral view showing the convex dorsal margin, (
<emphasis id="52DFEAEDB117FFF2CDDA558DFEE6F890" bold="true" box="[244,261,1776,1797]" pageId="7" pageNumber="59">B</emphasis>
) lateral view, (
<emphasis id="52DFEAEDB117FFF2CC8F5592FE50F893" bold="true" box="[417,435,1775,1798]" pageId="7" pageNumber="59">C</emphasis>
), caudal view. Sacrum and ilia of CM 11337 in (
<emphasis id="52DFEAEDB117FFF2CE83558DFC5CF890" bold="true" box="[941,959,1776,1797]" pageId="7" pageNumber="59">D</emphasis>
) dorsal view. Right ilium of CM 11337 in (
<emphasis id="52DFEAEDB117FFF2C8AD558DFA77F890" bold="true" box="[1411,1428,1776,1797]" pageId="7" pageNumber="59">E</emphasis>
) lateral view. Right ilium of OXFUM J.3303 (holotype of
<emphasis id="52DFEAEDB117FFF2CFF05472FC58F8B1" box="[734,955,1806,1828]" italics="true" pageId="7" pageNumber="59">
Cumnoria
<taxonomicName id="A7AB4D7CB117FFF2CE655473FC58F8B1" box="[843,955,1806,1828]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
) in (
<emphasis id="52DFEAEDB117FFF2CEC25473FC18F8B6" bold="true" box="[1004,1019,1806,1827]" pageId="7" pageNumber="59">F</emphasis>
) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process.
</paragraph>
</caption>
<paragraph id="601436FFB117FFFDCDE95412FB43FDA9" blockId="7.[151,1436,1903,2036]" lastBlockId="8.[151,1436,151,573]" lastPageId="8" lastPageNumber="60" pageId="7" pageNumber="59">
The results of the phylogenetic analysis and the morphological differences explicated above strongly suggest that
<taxonomicName id="A7AB4D7CB117FFF2CDE554E9FE2DF839" box="[203,462,1940,1964]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB117FFF2CDE554E9FE2DF839" box="[203,462,1940,1964]" italics="true" pageId="7" pageNumber="59">Uteodon aphanoecetes</emphasis>
</taxonomicName>
and
<emphasis id="52DFEAEDB117FFF2CF2554E9FCE1F839" box="[523,770,1940,1964]" italics="true" pageId="7" pageNumber="59">
Cumnoria
<taxonomicName id="A7AB4D7CB117FFF2CFA954E9FCE1F839" box="[647,770,1940,1964]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
should not be considered congeneric with
<taxonomicName id="A7AB4D7CB117FFF2C9D754E9FF3CF85A" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB117FFF2C9D754E9FF3CF85A" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
</taxonomicName>
;
<taxonomicName id="A7AB4D7CB117FFF2CDC054CAFEADF85A" box="[238,334,1975,1999]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="66" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="52DFEAEDB117FFF2CDC054CAFEADF85A" box="[238,334,1975,1999]" italics="true" pageId="7" pageNumber="59">Uteodon</emphasis>
</taxonomicName>
and
<emphasis id="52DFEAEDB117FFF2CCA854C5FE14F85A" box="[390,503,1976,1999]" italics="true" pageId="7" pageNumber="59">Cumnoria</emphasis>
are recovered as basal styracosternans more derived than
<taxonomicName id="A7AB4D7CB117FFF2C9A554C5FA9FF85A" box="[1163,1404,1975,1999]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB117FFF2C9A554C5FA9FF85A" box="[1163,1404,1975,1999]" italics="true" pageId="7" pageNumber="59">Camptosaurus dispar</emphasis>
</taxonomicName>
. It is thus a straightforward matter to reinstate the existing genus
<emphasis id="52DFEAEDB117FFF2CE7D54A1FC27F866" box="[851,964,2012,2035]" italics="true" pageId="7" pageNumber="59">Cumnoria</emphasis>
(
<bibRefCitation id="043A4B0EB117FFF2CEFD54A6FB83F866" author="Seeley" box="[979,1120,2011,2035]" pageId="7" pageNumber="59" refString="Seeley, H. G. (1888) On Cumnoria, an iguanodont genus founded upon the Iguanodon prestwichi, Hulke. Report of the British Association of the Advancement of Science, 57, 698." type="journal article" year="1888">Seeley 1888</bibRefCitation>
) for “
<taxonomicName id="A7AB4D7CB117FFF2C98A54A1FAA4F866" box="[1188,1351,2012,2035]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB117FFF2C98A54A1FAA4F866" box="[1188,1351,2012,2035]" italics="true" pageId="7" pageNumber="59">Camptosaurus</emphasis>
</taxonomicName>
”
<taxonomicName id="A7AB4D7CB117FFFDC87254A0FF3FFF25" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="8" lastPageNumber="60" order="Ornithischia" pageId="7" pageNumber="59" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="52DFEAEDB117FFFDC87254A0FF3FFF25" italics="true" lastPageId="8" lastPageNumber="60" pageId="7" pageNumber="59">prestwichii</emphasis>
</taxonomicName>
(see below). However, a more difficult dilemma is the status of “
<taxonomicName id="A7AB4D7CB118FFFDCE9053E5FB82FF3A" box="[958,1121,152,175]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB118FFFDCE9053E5FB82FF3A" box="[958,1121,152,175]" italics="true" pageId="8" pageNumber="60">Camptosaurus</emphasis>
</taxonomicName>
”
<taxonomicName id="A7AB4D7CB118FFFDC95B53E5FAEEFF25" box="[1141,1293,152,176]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB118FFFDC95B53E5FAEEFF25" box="[1141,1293,152,176]" italics="true" pageId="8" pageNumber="60">aphanoecetes</emphasis>
</taxonomicName>
, which does not have a pre-existing genus into which it can be conveniently placed.
<collectingCountry id="18BC766FB118FFFDCEFD53C1FC11FF41" box="[979,1010,188,212]" name="American Samoa" pageId="8" pageNumber="60">As</collectingCountry>
described above, this species is very similar to
<emphasis id="52DFEAEDB118FFFDCC24539DFE1DFF6D" box="[266,510,224,248]" italics="true" pageId="8" pageNumber="60">
Cumnoria
<taxonomicName id="A7AB4D7CB118FFFDCCAD539DFE1DFF6D" box="[387,510,224,248]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
, distinguished only by a single autapomorphy of the braincase. Thus, the possible taxonomic revision that immediately arises is to create the new combination
<emphasis id="52DFEAEDB118FFFDC9225278FAFCFE89" box="[1036,1311,260,284]" italics="true" pageId="8" pageNumber="60">
Cumnoria
<taxonomicName id="A7AB4D7CB118FFFDC9A95279FAFCFE89" box="[1159,1311,260,284]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">aphanoecetes</taxonomicName>
</emphasis>
. However, such a move presupposes that future phylogenetic analyses and discoveries will find a sister-taxa relationship of
<emphasis id="52DFEAEDB118FFFDCDB95230FE73FEF1" box="[151,400,332,356]" italics="true" pageId="8" pageNumber="60">
Cumnoria
<taxonomicName id="A7AB4D7CB118FFFDCC3A5231FE73FEF1" box="[276,400,332,356]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
and “
<taxonomicName id="A7AB4D7CB118FFFDCCF25230FD9CFEF1" box="[476,639,333,356]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB118FFFDCCF25230FD9CFEF1" box="[476,639,333,356]" italics="true" pageId="8" pageNumber="60">Camptosaurus</emphasis>
</taxonomicName>
”
<taxonomicName id="A7AB4D7CB118FFFDCFB65231FCD2FEF1" box="[664,817,332,356]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB118FFFDCFB65231FCD2FEF1" box="[664,817,332,356]" italics="true" pageId="8" pageNumber="60">aphanoecetes</emphasis>
</taxonomicName>
. The topologies presented herein could change with additional new taxa and new material, most especially additional cranial elements of “
<taxonomicName id="A7AB4D7CB118FFFDC9B0520DFAA2FE12" box="[1182,1345,368,391]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB118FFFDC9B0520DFAA2FE12" box="[1182,1345,368,391]" italics="true" pageId="8" pageNumber="60">Camptosaurus</emphasis>
</taxonomicName>
”
<taxonomicName id="A7AB4D7CB118FFFDC875520DFF1BFE39" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB118FFFDC875520DFF1BFE39" italics="true" pageId="8" pageNumber="60">aphanoecetes</emphasis>
</taxonomicName>
(in addition to the known right dentary DINO 556 and partial braincase of CM 15780). It is this labile nature of the phylogeny that prompts the creation of the new genus and combination
<taxonomicName id="A7AB4D7CB118FFFDC97552C5FAB8FE45" box="[1115,1371,440,464]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB118FFFDC97552C5FAB8FE45" box="[1115,1371,440,464]" italics="true" pageId="8" pageNumber="60">Uteodon aphanoecetes</emphasis>
</taxonomicName>
. This species can thus occupy any position relative to
<emphasis id="52DFEAEDB118FFFDCF9D52A0FC45FE61" box="[691,934,476,500]" italics="true" pageId="8" pageNumber="60">
Cumnoria
<taxonomicName id="A7AB4D7CB118FFFDCE0552A1FC45FE61" box="[811,934,476,500]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
and
<taxonomicName id="A7AB4D7CB118FFFDCEF352A0FB2CFE61" box="[989,1231,476,500]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB118FFFDCEF352A0FB2CFE61" box="[989,1231,476,500]" italics="true" pageId="8" pageNumber="60">Camptosaurus dispar</emphasis>
</taxonomicName>
in future analyses without necessitating still another taxonomic reassignment. From the standpoint of taxonomic stability, giving
<taxonomicName id="A7AB4D7CB118FFFDC844517CFE8DFDA9" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB118FFFDC844517CFE8DFDA9" italics="true" pageId="8" pageNumber="60">Uteodon aphanoecetes</emphasis>
</taxonomicName>
its own new genus is preferable to committing it to the genus
<emphasis id="52DFEAEDB118FFFDC9065158FB79FDA9" box="[1064,1178,549,572]" italics="true" pageId="8" pageNumber="60">Cumnoria</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="7147B3712306857C76ACBC2AE1A7084B">The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)</mods:title>
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<treatment id="E80287E9B118FFFCCDB95442FE0EFC55" ID-DOI="http://doi.org/10.5281/zenodo.6185215" ID-GBIF-Taxon="119391385" ID-Zenodo-Dep="6185215" LSID="urn:lsid:plazi:treatment:E80287E9B118FFFCCDB95442FE0EFC55" httpUri="http://treatment.plazi.org/id/E80287E9B118FFFCCDB95442FE0EFC55" lastPageId="9" lastPageNumber="61" pageId="8" pageNumber="60">
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<paragraph id="601436FFB118FFFDCDB95442FDD9F8CC" blockId="8.[151,570,1855,1881]" box="[151,570,1855,1881]" pageId="8" pageNumber="60">
<heading id="3B5C8193B118FFFDCDB95442FDD9F8CC" bold="true" box="[151,570,1855,1881]" fontSize="11" level="1" pageId="8" pageNumber="60" reason="1">
<emphasis id="52DFEAEDB118FFFDCDB95442FDD9F8CC" bold="true" box="[151,570,1855,1881]" pageId="8" pageNumber="60">
<emphasis id="52DFEAEDB118FFFDCDB95442FEF0F8CC" bold="true" box="[151,275,1855,1881]" italics="true" pageId="8" pageNumber="60">Cumnoria</emphasis>
<taxonomicName id="A7AB4D7CB118FFFDCC345442FDD9F8CC" authority="Hulke, 1880" authorityName="Hulke" authorityYear="1880" box="[282,570,1855,1881]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="52DFEAEDB118FFFDCC345442FE7DF8CC" bold="true" box="[282,414,1855,1881]" italics="true" pageId="8" pageNumber="60">prestwichii</emphasis>
<bibRefCitation id="043A4B0EB118FFFDCC8B5442FDD9F8CC" author="Hulke" box="[421,570,1855,1881]" pageId="8" pageNumber="60" refString="Hulke, J. W. (1880) Iguanodon prestwichii, a new species from the Kimmeridge Clay, founded on numerous fossil remains lately discovered at Cumnor, near Oxford. Quarterly Journal of the Geological Society of London, 36, 433 - 456." type="journal article" year="1880">Hulke, 1880</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="601436FFB118FFFDCDB954FAFDBCF809" blockId="8.[151,710,1926,2010]" box="[151,607,1926,1949]" pageId="8" pageNumber="60">
<bibRefCitation id="043A4B0EB118FFFDCDB954FAFF29F809" author="Hulke" box="[151,202,1927,1948]" pageId="8" pageNumber="60" refString="Hulke, J. W. (1880) Iguanodon prestwichii, a new species from the Kimmeridge Clay, founded on numerous fossil remains lately discovered at Cumnor, near Oxford. Quarterly Journal of the Geological Society of London, 36, 433 - 456." type="journal article" year="1880">1880</bibRefCitation>
<taxonomicName id="A7AB4D7CB118FFFDCDCB54FAFDECF809" authority="Hulke" authorityName="Hulke" box="[229,527,1926,1949]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="52DFEAEDB118FFFDCDCB54FAFE2AF808" box="[229,457,1927,1949]" italics="true" pageId="8" pageNumber="60">Iguanodon prestwichii</emphasis>
Hulke
</taxonomicName>
, p. 454.
</paragraph>
<paragraph id="601436FFB118FFFDCDB954DBFD25F84C" blockId="8.[151,710,1926,2010]" pageId="8" pageNumber="60">
1888
<emphasis id="52DFEAEDB118FFFDCDCB54DBFE59F82E" box="[229,442,1957,1979]" italics="true" pageId="8" pageNumber="60">
Cumnoria
<taxonomicName id="A7AB4D7CB118FFFDCC7C54D8FE59F82E" box="[338,442,1957,1979]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="prestwichi">prestwichi</taxonomicName>
</emphasis>
(
<emphasis id="52DFEAEDB118FFFDCCE754DBFE06F82E" box="[457,485,1958,1979]" italics="true" pageId="8" pageNumber="60">sic</emphasis>
) Seeley, p. 698. 1889
<taxonomicName id="A7AB4D7CB118FFFDCDCB54B9FE0BF84F" box="[229,488,1988,2010]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="8" pageNumber="60" phylum="Chordata" rank="species" species="prestwichi">
<emphasis id="52DFEAEDB118FFFDCDCB54B9FE0BF84F" box="[229,488,1988,2010]" italics="true" pageId="8" pageNumber="60">Camptosaurus prestwichi</emphasis>
</taxonomicName>
(
<emphasis id="52DFEAEDB118FFFDCCD854B8FDF1F84F" box="[502,530,1989,2010]" italics="true" pageId="8" pageNumber="60">sic</emphasis>
) Lydekker, p. 48.
</paragraph>
<paragraph id="601436FFB119FFFCCDB953E5FD7FFF3B" blockId="9.[151,755,152,205]" box="[151,668,152,174]" pageId="9" pageNumber="61">
1909
<taxonomicName id="A7AB4D7CB119FFFCCDCB53E4FDA8FF3B" authority="Gilmore" authorityName="Gilmore" box="[229,587,152,174]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="52DFEAEDB119FFFCCDCB53E4FE0CFF3B" box="[229,495,152,174]" italics="true" pageId="9" pageNumber="61">Camptosaurus prestwichii</emphasis>
Gilmore
</taxonomicName>
, p. 285.
</paragraph>
<paragraph id="601436FFB119FFFCCDB953CAFD10FF59" blockId="9.[151,755,152,205]" box="[151,755,182,205]" pageId="9" pageNumber="61">
1980
<taxonomicName id="A7AB4D7CB119FFFCCDCB53CAFD40FF59" authority="Galton &amp; Powell" authorityName="Galton &amp; Powell" box="[229,675,182,205]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="52DFEAEDB119FFFCCDCB53CAFE0CFF58" box="[229,495,183,205]" italics="true" pageId="9" pageNumber="61">Camptosaurus prestwichii</emphasis>
Galton &amp; Powell
</taxonomicName>
, p. 412.
</paragraph>
</subSubSection>
<subSubSection id="28B16574B119FFFCCDB95384FC26FD98" pageId="9" pageNumber="61" type="materials_examined">
<paragraph id="601436FFB119FFFCCDB95384FCC0FE84" blockId="9.[151,1436,249,960]" box="[151,803,249,273]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDB95384FEE8FE84" bold="true" box="[151,267,249,273]" pageId="9" pageNumber="61">
<typeStatus id="BF10885DB119FFFCCDB95384FEE5FE84" box="[151,262,249,273]" pageId="9" pageNumber="61" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="06BAAE3AB119FFFCCC3C5384FE94FE84" box="[274,375,249,273]" pageId="9" pageNumber="61">OXFUM</collectionCode>
J.3303, partial skull and postcranium.
</paragraph>
<paragraph id="601436FFB119FFFCCDEB5260FD80FECC" blockId="9.[151,1436,249,960]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB5260FE2EFEA3" bold="true" box="[197,461,285,310]" pageId="9" pageNumber="61">Locality and horizon.</emphasis>
Chawley Brick Pit, near Cumnor, Oxfordshire,
<collectingCountry id="18BC766FB119FFFCC92E5263FBCAFEA0" box="[1024,1065,286,309]" name="United Kingdom" pageId="9" pageNumber="61">UK</collectingCountry>
(
<bibRefCitation id="043A4B0EB119FFFCC9155260FAAAFEA0" author="Galton" box="[1083,1353,285,309]" pageId="9" pageNumber="61" refString="Galton, P. M. &amp; Powell, H. P. (1980) The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology, 23, 411 - 443." type="journal article" year="1980">Galton &amp; Powell 1980</bibRefCitation>
); Kimmeridge Clay Formation, Kimmeridgian.
</paragraph>
<paragraph id="601436FFB119FFFCCDEB5218FCF3FEEB" blockId="9.[151,1436,249,960]" box="[197,784,357,382]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB5218FE43FEEB" bold="true" box="[197,416,357,382]" pageId="9" pageNumber="61">Generic diagnosis.</emphasis>
<collectingCountry id="18BC766FB119FFFCCC89521BFE25FEEB" box="[423,454,358,382]" name="American Samoa" pageId="9" pageNumber="61">As</collectingCountry>
for the species by monotypy.
</paragraph>
<paragraph id="601436FFB119FFFCCDEB52F5FC26FD98" blockId="9.[151,1436,249,960]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB52F5FE41FE34" bold="true" box="[197,418,392,417]" pageId="9" pageNumber="61">Specific diagnosis.</emphasis>
Basal styracosternan distinguished from all other iguanodontians except
<taxonomicName id="A7AB4D7CB119FFFCC9DE52F4FF1BFE50" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB119FFFCC9DE52F4FF1BFE50" italics="true" pageId="9" pageNumber="61">Uteodon aphanoecetes</emphasis>
</taxonomicName>
by a unique combination of characters; the same unique combination of characters distinguishes
<taxonomicName id="A7AB4D7CB119FFFCC81252D0FED3FE7C" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB119FFFCC81252D0FED3FE7C" italics="true" pageId="9" pageNumber="61">Uteodon aphanoecetes</emphasis>
</taxonomicName>
(see ‘Specific diagnosis’ above). Distinguished from
<taxonomicName id="A7AB4D7CB119FFFCCEA052ACFB6DFE7C" box="[910,1166,465,489]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB119FFFCCEA052ACFB6DFE7C" box="[910,1166,465,489]" italics="true" pageId="9" pageNumber="61">Uteodon aphanoecetes</emphasis>
</taxonomicName>
by an occipital condyle that does not project farther ventrally than the basal tubera (
<figureCitation id="F8902A7AB119FFFCCE1A5288FC61FD98" box="[820,898,501,525]" captionStart="FIGURE 7" captionStartId="8.[151,250,1622,1645]" captionTargetBox="[472,1114,625,1600]" captionTargetId="figure@8.[472,1115,625,1601]" captionTargetPageId="8" captionText="FIGURE 7. Partial braincases of Camptosaurus dispar, Uteodon aphanoecetes, and Cumnoria prestwichii. (A) YPM 1880 in left lateral view (image copyright YPM); (B) CM 15780 in right lateral view (image courtesy of M. Lamanna); (C) OXFUM J. 3303 in right lateral view. Abbreviations: bt, basal tubera; oc, occipital condyle. Scale bar equals 5 cm." httpUri="https://zenodo.org/record/200818/files/figure.png" pageId="9" pageNumber="61">Figs. 7</figureCitation>
B, C).
</paragraph>
</subSubSection>
<subSubSection id="28B16574B119FFFCCDEB5164FE0EFC55" pageId="9" pageNumber="61" type="discussion">
<paragraph id="601436FFB119FFFCCDEB5164FE03FC99" blockId="9.[151,1436,249,960]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB5164FEDAFDA4" bold="true" box="[197,313,537,561]" pageId="9" pageNumber="61">Remarks.</emphasis>
<emphasis id="52DFEAEDB119FFFCCC6F5167FDD5FDA4" box="[321,566,537,561]" italics="true" pageId="9" pageNumber="61">
Cumnoria
<taxonomicName id="A7AB4D7CB119FFFCCC945164FDD5FDA4" box="[442,566,537,561]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
was originally named as a new species of
<taxonomicName id="A7AB4D7CB119FFFCC9305164FADAFDA4" authority="Hulke 1880" authorityName="Hulke" authorityYear="1880" box="[1054,1337,537,561]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB119FFFCC9305164FB74FDA4" box="[1054,1175,537,561]" italics="true" pageId="9" pageNumber="61">Iguanodon</emphasis>
(
<bibRefCitation id="043A4B0EB119FFFCC9895164FAD2FDA4" author="Hulke" box="[1191,1329,537,561]" pageId="9" pageNumber="61" refString="Hulke, J. W. (1880) Iguanodon prestwichii, a new species from the Kimmeridge Clay, founded on numerous fossil remains lately discovered at Cumnor, near Oxford. Quarterly Journal of the Geological Society of London, 36, 433 - 456." type="journal article" year="1880">Hulke 1880</bibRefCitation>
)
</taxonomicName>
, as were numerous different basal iguanodonts in the 19th Century.
<bibRefCitation id="043A4B0EB119FFFCCE0F513DFC5EFDCD" author="Seeley" box="[801,957,576,600]" pageId="9" pageNumber="61" refString="Seeley, H. G. (1888) On Cumnoria, an iguanodont genus founded upon the Iguanodon prestwichi, Hulke. Report of the British Association of the Advancement of Science, 57, 698." type="journal article" year="1888">Seeley (1888)</bibRefCitation>
subsequently erected the new genus
<emphasis id="52DFEAEDB119FFFCC870513CFF37FDE9" italics="true" pageId="9" pageNumber="61">Cumnoria</emphasis>
for “
<emphasis id="52DFEAEDB119FFFCCC3E5118FEFAFDE9" box="[272,281,613,636]" italics="true" pageId="9" pageNumber="61">I</emphasis>
.”
<taxonomicName id="A7AB4D7CB119FFFCCC1B5119FE53FDE9" box="[309,432,612,636]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="52DFEAEDB119FFFCCC1B5119FE53FDE9" box="[309,432,612,636]" italics="true" pageId="9" pageNumber="61">prestwichii</emphasis>
</taxonomicName>
. This designation was almost immediately challenged by
<bibRefCitation id="043A4B0EB119FFFCC96E5119FB1DFDE9" author="Lydekker" box="[1088,1278,612,636]" pageId="9" pageNumber="61" refString="Lydekker, R. (1889) On the remains and affinities of give genera of Mesozoic reptiles. Quarterly Journal of the Geological Society of London, 45, 41 - 59." type="journal article" year="1889">Lydekker (1889)</bibRefCitation>
, who referred the species to
<taxonomicName id="A7AB4D7CB119FFFCCC1A51F4FE3BFD35" box="[308,472,649,672]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB119FFFCCC1A51F4FE3BFD35" box="[308,472,649,672]" italics="true" pageId="9" pageNumber="61">Camptosaurus</emphasis>
</taxonomicName>
. Later authors have largely followed Lydekker’s decision (e.g.,
<bibRefCitation id="043A4B0EB119FFFCC98A51F5FAA7FD35" author="Gilmore" box="[1188,1348,648,672]" pageId="9" pageNumber="61" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
;
<bibRefCitation id="043A4B0EB119FFFCC86151F5FEA8FD51" author="Galton" pageId="9" pageNumber="61" refString="Galton, P. M. &amp; Powell, H. P. (1980) The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England. Palaeontology, 23, 411 - 443." type="journal article" year="1980">Galton &amp; Powell 1980</bibRefCitation>
;
<bibRefCitation id="043A4B0EB119FFFCCC7451D0FE1FFD51" author="Norman" box="[346,508,684,708]" pageId="9" pageNumber="61" refString="Norman, D. B. (2004) Basal Iguanodontia. In: Weishampel, D. B., Dodson, P. &amp; Osmolska, H. (eds), The Dinosauria: Second Edition. University of California Press, Berkeley, pp. 413 - 437." type="book chapter" year="2004">Norman 2004</bibRefCitation>
), although in a recent historical overview of British dinosaurs,
<bibRefCitation id="043A4B0EB119FFFCC9CC51D1FF02FD7D" author="Naish" pageId="9" pageNumber="61" refString="Naish, D. &amp; Martill, D. M. (2008) Dinosaurs of Great Britain and the role of the Geological Society of London in their discovery: Ornithischia. Journal of the Geological Society, London, 165, 613 - 623." type="journal article" year="2008">Naish &amp; Martill (2008)</bibRefCitation>
suggested that using
<emphasis id="52DFEAEDB119FFFCCCFA51ACFDA5FD7D" box="[468,582,721,744]" italics="true" pageId="9" pageNumber="61">Cumnoria</emphasis>
might be preferable in light of new information on the skull of
<taxonomicName id="A7AB4D7CB119FFFCC83D51ACFE3FFC99" authority="Brill &amp; Carpenter 2007" authorityName="Brill &amp; Carpenter" authorityYear="2007" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB119FFFCC83D51ACFF58FC99" italics="true" pageId="9" pageNumber="61">Camptosaurus</emphasis>
(
<bibRefCitation id="043A4B0EB119FFFCCDE75189FE36FC99" author="Brill" box="[201,469,756,780]" pageId="9" pageNumber="61" refString="Brill, K. &amp; Carpenter, K. (2007) A description of a new ornithopod from the Lytle Member of the Purgatoire Formation (Lower Cretaceous) and a reassessment of the skull of Camptosaurus. In: Carpenter, K. (ed), Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Indiana University Press, Bloomington, pp. 49 - 67." type="book chapter" year="2007">Brill &amp; Carpenter 2007</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="601436FFB119FFFCCDEB5065FE0EFC55" blockId="9.[151,1436,249,960]" pageId="9" pageNumber="61">
The phylogenetic analysis supports Naish &amp; Martill’s tentative proposal, recovering “
<taxonomicName id="A7AB4D7CB119FFFCC98C5064FAA6FCA5" box="[1186,1349,793,816]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB119FFFCC98C5064FAA6FCA5" box="[1186,1349,793,816]" italics="true" pageId="9" pageNumber="61">Camptosaurus</emphasis>
</taxonomicName>
”
<taxonomicName id="A7AB4D7CB119FFFCC8725064FF3FFCC1" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="52DFEAEDB119FFFCC8725064FF3FFCC1" italics="true" pageId="9" pageNumber="61">prestwichii</emphasis>
</taxonomicName>
in a polytomy at the base of Styracosterna with
<taxonomicName id="A7AB4D7CB119FFFCCFD25041FC1FFCC1" box="[764,1020,828,852]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB119FFFCCFD25041FC1FFCC1" box="[764,1020,828,852]" italics="true" pageId="9" pageNumber="61">Uteodon aphanoecetes</emphasis>
</taxonomicName>
in the reduced consensus tree. Thus, the generic name
<emphasis id="52DFEAEDB119FFFCCC4A501CFE35FCED" box="[356,470,865,888]" italics="true" pageId="9" pageNumber="61">Cumnoria</emphasis>
is reinstated for the species
<taxonomicName id="A7AB4D7CB119FFFCCE31501CFC2FFCED" box="[799,972,864,888]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="prestwichii">
<emphasis id="52DFEAEDB119FFFCCE31501CFCDCFCED" box="[799,831,865,888]" italics="true" pageId="9" pageNumber="61">Cu</emphasis>
.
<emphasis id="52DFEAEDB119FFFCCE7E501DFC2FFCED" box="[848,972,864,888]" italics="true" pageId="9" pageNumber="61">prestwichii</emphasis>
</taxonomicName>
.
<emphasis id="52DFEAEDB119FFFCCEF2501CFB31FCED" box="[988,1234,864,888]" italics="true" pageId="9" pageNumber="61">
Cumnoria
<taxonomicName id="A7AB4D7CB119FFFCC978501DFB31FCED" box="[1110,1234,864,888]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="prestwichii">prestwichii</taxonomicName>
</emphasis>
was coded based upon the
<typeStatus id="BF10885DB119FFFCCDD050F9FE83FC09" box="[254,352,900,924]" pageId="9" pageNumber="61" type="holotype">holotype</typeStatus>
and only known specimen, OXFUM J.3303. See ‘Remarks’ on
<taxonomicName id="A7AB4D7CB119FFFCC90750F9FB69FC09" box="[1065,1162,900,924]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="66" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="52DFEAEDB119FFFCC90750F9FB69FC09" box="[1065,1162,900,924]" italics="true" pageId="9" pageNumber="61">Uteodon</emphasis>
</taxonomicName>
for further discussion of the relationships of
<emphasis id="52DFEAEDB119FFFCCC5A50D4FE06FC55" box="[372,485,937,960]" italics="true" pageId="9" pageNumber="61">Cumnoria</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="418EB8730E7548BBF6212C5AA486C2CE">The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)</mods:title>
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<subSubSection id="28B16574B119FFFCCDB95727FD35FB63" pageId="9" pageNumber="61" type="nomenclature">
<paragraph id="601436FFB119FFFCCDB95727FD1AFBE6" blockId="9.[151,761,1114,1139]" box="[151,761,1114,1139]" pageId="9" pageNumber="61">
<heading id="3B5C8193B119FFFCCDB95727FD1AFBE6" bold="true" box="[151,761,1114,1139]" fontSize="10" level="2" pageId="9" pageNumber="61" reason="2">
<emphasis id="52DFEAEDB119FFFCCDB95727FD1AFBE6" bold="true" box="[151,761,1114,1139]" pageId="9" pageNumber="61">
<taxonomicName id="A7AB4D7CB119FFFCCDB95727FD88FBE6" authority="Gilmore, 1909" authorityName="Gilmore" authorityYear="1909" box="[151,619,1114,1139]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressus" status="comb. nov.">
<emphasis id="52DFEAEDB119FFFCCDB95727FE59FBE6" bold="true" box="[151,442,1114,1139]" italics="true" pageId="9" pageNumber="61">Osmakasaurus depressus</emphasis>
<bibRefCitation id="043A4B0EB119FFFCCCEF5727FD88FBE6" author="Gilmore" box="[449,619,1114,1139]" pageId="9" pageNumber="61" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 197 - 332." type="journal article" year="1909">Gilmore, 1909</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="49EC5796B119FFFCCF565726FD1AFBE6" box="[632,761,1115,1139]" pageId="9" pageNumber="61">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="601436FFB119FFFCCDB957DEFD75FB2D" blockId="9.[151,726,1186,1270]" box="[151,662,1186,1209]" pageId="9" pageNumber="61">
1909
<taxonomicName id="A7AB4D7CB119FFFCCDC557DEFDA6FB2D" authority="Gilmore" authorityName="Gilmore" box="[235,581,1186,1209]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB119FFFCCDC557DEFE0BFB2C" box="[235,488,1187,1209]" italics="true" pageId="9" pageNumber="61">Camptosaurus depressus</emphasis>
Gilmore
</taxonomicName>
, p. 292.
</paragraph>
<paragraph id="601436FFB119FFFCCDB957BFFD7CFB42" blockId="9.[151,726,1186,1270]" box="[151,671,1217,1239]" pageId="9" pageNumber="61">
2004?
<taxonomicName id="A7AB4D7CB119FFFCCDD857BFFDACFB42" authority="Norman" authorityName="Norman" box="[246,591,1217,1239]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB119FFFCCDD857BFFE10FB42" box="[246,499,1217,1239]" italics="true" pageId="9" pageNumber="61">Camptosaurus depressus</emphasis>
Norman
</taxonomicName>
, p. 415.
</paragraph>
<paragraph id="601436FFB119FFFCCDB9579DFD35FB63" blockId="9.[151,726,1186,1270]" box="[151,726,1248,1270]" pageId="9" pageNumber="61">
2008
<taxonomicName id="A7AB4D7CB119FFFCCDC5579DFD65FB63" authority="Carpenter &amp; Wilson" authorityName="Carpenter &amp; Wilson" box="[235,646,1248,1270]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressa">
<emphasis id="52DFEAEDB119FFFCCDC5579DFE52FB63" box="[235,433,1248,1270]" italics="true" pageId="9" pageNumber="61">Planicoxa depressa</emphasis>
Carpenter &amp; Wilson
</taxonomicName>
, p. 257.
</paragraph>
</subSubSection>
<subSubSection id="28B16574B119FFFCCDB9565EFF3AF952" pageId="9" pageNumber="61" type="materials_examined">
<paragraph id="601436FFB119FFFCCDB9565EFD32FAAF" blockId="9.[151,1437,1314,1986]" box="[151,721,1314,1339]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDB9565EFEE8FAAE" bold="true" box="[151,267,1315,1339]" pageId="9" pageNumber="61">
<typeStatus id="BF10885DB119FFFCCDB9565EFEE5FAAE" box="[151,262,1315,1339]" pageId="9" pageNumber="61" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="06BAAE3AB119FFFCCC3C565FFE80FAAF" LSID="urn:lsid:biocol.org:col:34871" box="[274,355,1314,1338]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="61">USNM</collectionCode>
4753, fragmentary postcranium.
</paragraph>
<paragraph id="601436FFB119FFFCCDEB563BFF2BFA5F" blockId="9.[151,1437,1314,1986]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB563BFE4FFACA" bold="true" box="[197,428,1350,1375]" pageId="9" pageNumber="61">Generic etymology.</emphasis>
The generic name is derived from
<emphasis id="52DFEAEDB119FFFCCE19563AFC6FFACA" box="[823,908,1351,1375]" italics="true" pageId="9" pageNumber="61">ósmaka</emphasis>
, which means “canyon” in the language of the Lakota People, in reference to the name of the
<typeStatus id="BF10885DB119FFFCCF905616FD0DFA16" box="[702,750,1387,1411]" pageId="9" pageNumber="61">type</typeStatus>
locality in western South Dakota, an area inhabited by the Lakota.
<taxonomicName id="A7AB4D7CB119FFFCCDDA56F2FEA1FA33" box="[244,322,1423,1446]" family="Synodontidae" genus="Saurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Aulopiformes" pageId="9" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB119FFFCCDDA56F2FEA1FA33" box="[244,322,1423,1446]" italics="true" pageId="9" pageNumber="61">Saurus</emphasis>
</taxonomicName>
comes from the transliterated Greek meaning “lizard”. The gender of the generic name is masculine.
</paragraph>
<paragraph id="601436FFB119FFFCCDEB56ABFCDFF987" blockId="9.[151,1437,1314,1986]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB56ABFE33FA7A" bold="true" box="[197,464,1494,1519]" pageId="9" pageNumber="61">Locality and horizon.</emphasis>
Calico Canyon, near Buffalo Gap, Custer County, South Dakota (
<bibRefCitation id="043A4B0EB119FFFCC9C956AAFA6CFA7A" author="Gilmore" box="[1255,1423,1495,1519]" pageId="9" pageNumber="61" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 197 - 332." type="journal article" year="1909">Gilmore 1909</bibRefCitation>
); Lakota Formation, Barremian–Aptian (
<bibRefCitation id="043A4B0EB119FFFCCF635687FCD3F987" author="Tschudy" box="[589,816,1530,1555]" pageId="9" pageNumber="61" refString="Tschudy, R. H., Tschudy, B. D. &amp; Craig, L. C. (1984) Palynological evaluation of Cedar Mountain and Burro Canyon formations, Colorado Plateau. United States Geological Survey Professional Paper, 1281, 1 - 24." type="journal article" year="1984">
Tschudy
<emphasis id="52DFEAEDB119FFFCCF9A5681FD07F986" box="[692,740,1531,1555]" italics="true" pageId="9" pageNumber="61">et al</emphasis>
. 1984
</bibRefCitation>
).
</paragraph>
<paragraph id="601436FFB119FFFCCDEB5563FCF3F9AD" blockId="9.[151,1437,1314,1986]" box="[197,784,1566,1592]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB5563FE43F9A2" bold="true" box="[197,416,1566,1591]" pageId="9" pageNumber="61">Generic diagnosis.</emphasis>
<collectingCountry id="18BC766FB119FFFCCC895562FE25F9A2" box="[423,454,1567,1591]" name="American Samoa" pageId="9" pageNumber="61">As</collectingCountry>
for the species by monotypy.
</paragraph>
<paragraph id="601436FFB119FFFCCDEB553FFF3AF952" blockId="9.[151,1437,1314,1986]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB553FFE43F9CE" bold="true" box="[197,416,1602,1627]" pageId="9" pageNumber="61">Specific diagnosis.</emphasis>
Basal styracosternan distinguished from all other iguanodontians by the following unique combination of characters: cranial end of preacetabular process of ilium modified into horizontal boot, straight dorsal margin of ilium, dorsal margin of ilium thickens mediolaterally towards the
<emphasis id="52DFEAEDB119FFFCC93D55F1FBC9F936" box="[1043,1066,1676,1699]" italics="true" pageId="9" pageNumber="61">M</emphasis>
.
<emphasis id="52DFEAEDB119FFFCC91655F6FB5FF936" box="[1080,1212,1675,1699]" italics="true" pageId="9" pageNumber="61">iliocaudalis</emphasis>
platform (
<figureCitation id="F8902A7AB119FFFCC81D55F7FA63F937" box="[1331,1408,1674,1698]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="9" pageNumber="61">Figs. 8</figureCitation>
A, B, E).
</paragraph>
</subSubSection>
<subSubSection id="28B16574B119FFF9CDEB55AEFC10F979" lastPageId="12" lastPageNumber="64" pageId="9" pageNumber="61" type="discussion">
<paragraph id="601436FFB119FFFCCDEB55AEFC11F857" blockId="9.[151,1437,1314,1986]" pageId="9" pageNumber="61">
<emphasis id="52DFEAEDB119FFFCCDEB55AEFEDAF97E" bold="true" box="[197,313,1747,1771]" pageId="9" pageNumber="61">Remarks.</emphasis>
<taxonomicName id="A7AB4D7CB119FFFCCC6F55AEFDB9F97E" box="[321,602,1747,1771]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB119FFFCCC6F55AEFDB9F97E" box="[321,602,1747,1771]" italics="true" pageId="9" pageNumber="61">Camptosaurus depressus</emphasis>
</taxonomicName>
was originally distinguished from other species of
<taxonomicName id="A7AB4D7CB119FFFCC98D55AEFAA5F97F" box="[1187,1350,1747,1770]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB119FFFCC98D55AEFAA5F97F" box="[1187,1350,1747,1770]" italics="true" pageId="9" pageNumber="61">Camptosaurus</emphasis>
</taxonomicName>
chiefly by the “narrowness or depressed nature of the ilia” (
<bibRefCitation id="043A4B0EB119FFFCCFF5558AFC4CF89A" author="Gilmore" box="[731,943,1783,1807]" pageId="9" pageNumber="61" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 197 - 332." type="journal article" year="1909">Gilmore 1909:293</bibRefCitation>
). This condition is exhibited by only the left ilium of USNM 4753, on which the dorsal margin of the ilium above the acetabulum faces dorsolaterally and the postacetabular process is nearly horizontal (
<figureCitation id="F8902A7AB119FFFCCFAF5442FD27F8C2" box="[641,708,1855,1879]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="9" pageNumber="61">Fig. 8</figureCitation>
A).
<bibRefCitation id="043A4B0EB119FFFCCFC15442FBC0F8C2" author="Carpenter" box="[751,1059,1855,1879]" pageId="9" pageNumber="61" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson (2008)</bibRefCitation>
considered the horizontal postacetabular process of USNM 4753 similar to that of DMNH 42504, the
<typeStatus id="BF10885DB119FFFCCECC541FFBA7F8EF" box="[994,1092,1890,1914]" pageId="9" pageNumber="61" type="holotype">holotype</typeStatus>
ilium of
<taxonomicName id="A7AB4D7CB119FFFCC995541EFE38F80A" authority="DiCroce &amp; Carpenter 2001" authorityName="DiCroce &amp; Carpenter" authorityYear="2001" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB119FFFCC995541EFA7FF8EE" box="[1211,1436,1891,1915]" italics="true" pageId="9" pageNumber="61">Planicoxa venenica</emphasis>
(
<bibRefCitation id="043A4B0EB119FFFCCDB054FAFE31F80A" author="DiCroce" box="[158,466,1927,1951]" pageId="9" pageNumber="61" refString="DiCroce, T. &amp; Carpenter, K. (2001) New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah. In: Tanke, D. &amp; Carpenter, K. (eds), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, pp. 183 - 196." type="book chapter" year="2001">DiCroce &amp; Carpenter 2001</bibRefCitation>
)
</taxonomicName>
; thus, they referred
<taxonomicName id="A7AB4D7CB119FFFCCF9954F5FCA7F80A" box="[695,836,1927,1951]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB119FFFCCF9954F5FD29F80A" box="[695,714,1928,1951]" italics="true" pageId="9" pageNumber="61">C</emphasis>
.
<emphasis id="52DFEAEDB119FFFCCFF954FAFCA7F80A" box="[727,836,1927,1951]" italics="true" pageId="9" pageNumber="61">depressus</emphasis>
</taxonomicName>
to
<taxonomicName id="A7AB4D7CB119FFFCCE4954FAFC3AF80A" box="[871,985,1927,1951]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB119FFFCCE4954FAFC3AF80A" box="[871,985,1927,1951]" italics="true" pageId="9" pageNumber="61">Planicoxa</emphasis>
</taxonomicName>
, as
<taxonomicName id="A7AB4D7CB119FFFCC92A54F5FB62F80A" box="[1028,1153,1927,1951]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="61" phylum="Chordata" rank="species" species="depressa">
<emphasis id="52DFEAEDB119FFFCC92A54F5FBF1F80A" box="[1028,1042,1928,1951]" italics="true" pageId="9" pageNumber="61">P</emphasis>
.
<emphasis id="52DFEAEDB119FFFCC93154FAFB62F80A" box="[1055,1153,1927,1951]" italics="true" pageId="9" pageNumber="61">depressa</emphasis>
</taxonomicName>
. However, the horizontal postacetabular process of USNM 4753 is more likely a product of distortion.
</paragraph>
<paragraph id="601436FFB11AFFFFCDE953EAFB9CFE61" blockId="10.[151,1436,151,500]" pageId="10" pageNumber="62">
The blade of the left ilium of USNM 4753 is heavily fractured by corresponding cracks on the lateral and medial surfaces (
<figureCitation id="F8902A7AB11AFFFFCC7953C1FE47FF41" box="[343,420,188,212]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="10" pageNumber="62">Figs. 8</figureCitation>
A, B). Four roughly vertical cracks divide the blade of the ilium into four segments; these four cracks all merge with a roughly horizontal elongate crack extending craniocaudally dorsal to the acetabulum and peduncles. These cracks mean that the three more cranial segments are suspended in filling substance and that their orientations relative to the rest of the ilium are not certain. Furthermore, there is a point of offset on the lateral surface of the ilium at which one of the blade fragments overhangs the acetabular portion (
<figureCitation id="F8902A7AB11AFFFFC9BB5231FB3BFEF1" box="[1173,1240,332,356]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="10" pageNumber="62">Fig. 8</figureCitation>
C); this overhang is due to filler between the two pieces and does not reflect the actual morphology of the ilium. Notwithstanding uncertainty regarding the reconstruction of the iliac blade fragments, the postacetabular process does project far dorsolaterally relative to the ischial peduncle (
<figureCitation id="F8902A7AB11AFFFFCF9E52CAFD15FE5A" box="[688,758,439,463]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="10" pageNumber="62">Fig. 8</figureCitation>
D). However, comparing the case of USNM 4753 to other basal iguanodonts, it becomes clear that this is probably a product of plastic deformation.
</paragraph>
<caption id="34D46677B11AFFFFCDB95619FD67FA22" httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="10" pageNumber="62" targetBox="[151,1436,537,1358]" targetPageId="10">
<paragraph id="601436FFB11AFFFFCDB95619FD67FA22" blockId="10.[151,1436,1380,1463]" pageId="10" pageNumber="62">
<emphasis id="52DFEAEDB11AFFFFCDB95619FEF1FAEC" bold="true" box="[151,274,1380,1403]" pageId="10" pageNumber="62">FIGURE 8.</emphasis>
Left ilium of USNM 4753 (holotype of
<taxonomicName id="A7AB4D7CB11AFFFFCF835619FC4EFAEF" box="[685,941,1380,1402]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="62" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11AFFFFCF835619FC4EFAEF" box="[685,941,1380,1402]" italics="true" pageId="10" pageNumber="62">Osmakasaurus depressus</emphasis>
</taxonomicName>
) in (
<emphasis id="52DFEAEDB11AFFFFCEF05619FC13FAEC" bold="true" box="[990,1008,1380,1401]" pageId="10" pageNumber="62">A</emphasis>
) lateral, (
<emphasis id="52DFEAEDB11AFFFFC97A5618FB86FAEF" bold="true" box="[1108,1125,1381,1402]" pageId="10" pageNumber="62">B</emphasis>
) medial, (
<emphasis id="52DFEAEDB11AFFFFC9E05619FB03FAEE" bold="true" box="[1230,1248,1380,1403]" pageId="10" pageNumber="62">C</emphasis>
) caudolateral, and (
<emphasis id="52DFEAEDB11AFFFFCDB156FEFF52FA0D" bold="true" box="[159,177,1411,1432]" pageId="10" pageNumber="62">D</emphasis>
) caudal views. Right ilium of USNM 4753 in (
<emphasis id="52DFEAEDB11AFFFFCF8756FEFD59FA0D" bold="true" box="[681,698,1411,1432]" pageId="10" pageNumber="62">E</emphasis>
) lateral and (
<emphasis id="52DFEAEDB11AFFFFCE6756FEFCBBFA0D" bold="true" box="[841,856,1411,1432]" pageId="10" pageNumber="62">F</emphasis>
) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm.
</paragraph>
</caption>
<paragraph id="601436FFB11AFFFFCDE95698FD64F889" blockId="10.[151,1436,1508,2036]" pageId="10" pageNumber="62">
Among the skeletons of
<taxonomicName id="A7AB4D7CB11AFFFFCCF05699FCEAFA69" box="[478,777,1508,1532]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="62" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="52DFEAEDB11AFFFFCCF05699FCEAFA69" box="[478,777,1508,1532]" italics="true" pageId="10" pageNumber="62">Iguanodon bernissartensis</emphasis>
</taxonomicName>
excavated from the Bernissart Mine in
<collectingCountry id="18BC766FB11AFFFFC9E05699FACDFA69" box="[1230,1326,1508,1532]" name="Belgium" pageId="10" pageNumber="62">Belgium</collectingCountry>
(
<bibRefCitation id="043A4B0EB11AFFFFC8105699FF30F9B5" author="Norman" pageId="10" pageNumber="62" refString="Norman, D. B. (1980) On the ornithischian dinosaur Iguanodon bernissartensis from Belgium. Memoires Institut Royal des Sciences Naturelles de Belgique, 178, 1 - 103." type="journal article" year="1980">Norman 1980</bibRefCitation>
), there is variation in the forms of the ilia that can be attributed to deformation. The left ilium of the
<typeStatus id="BF10885DB11AFFFFC8005575FA74F9B5" box="[1326,1431,1544,1568]" pageId="10" pageNumber="62" type="lectotype">lectotype</typeStatus>
, IRSNB 1534, is well preserved with a nearly intact, gently convex dorsal margin along which there extends a thickened, laterally everted rim (
<figureCitation id="F8902A7AB11AFFFFCCE1552DFDF0F9FD" box="[463,531,1616,1640]" captionStart="FIGURE 9" captionStartId="11.[151,250,1766,1789]" captionTargetBox="[152,1390,343,1723]" captionTargetId="figure@11.[151,1436,307,1745]" captionTargetPageId="11" captionText="FIGURE 9. Left ilia of Iguanodon bernissartensis in lateral view. (A) IRSNB 1534 (lectotype), dorsal margin outlined in red; (B) IRSNB 1536, dorsal margin outlined in red; (C) IRSNB 1561." httpUri="https://zenodo.org/record/200820/files/figure.png" pageId="10" pageNumber="62">Fig. 9</figureCitation>
A). The postacetabular process tapers towards its caudal end. This is taken to be the undistorted form of the ilium. In contrast, the left ilia of IRSNB
<date id="1415103FB11AFFFFCEBF5509FBDBF919" box="[913,1080,1652,1676]" pageId="10" pageNumber="62" value="1536" valueMax="1561">1536 and 1561</date>
resemble those of USNM
<quantity id="A7539B1AB11AFFFFC84A5509FF4EF925" metricMagnitude="2" metricUnit="m" metricValue="1.207262" pageId="10" pageNumber="62" unit="in" value="4753.0">4753 in</quantity>
their dorsolaterally projecting dorsal margins dorsal to the ischial peduncle (
<figureCitation id="F8902A7AB11AFFFFC93F55E5FB83F925" box="[1041,1120,1688,1712]" captionStart="FIGURE 9" captionStartId="11.[151,250,1766,1789]" captionTargetBox="[152,1390,343,1723]" captionTargetId="figure@11.[151,1436,307,1745]" captionTargetPageId="11" captionText="FIGURE 9. Left ilia of Iguanodon bernissartensis in lateral view. (A) IRSNB 1534 (lectotype), dorsal margin outlined in red; (B) IRSNB 1536, dorsal margin outlined in red; (C) IRSNB 1561." httpUri="https://zenodo.org/record/200820/files/figure.png" pageId="10" pageNumber="62">Figs. 9</figureCitation>
B, C), giving the ilia a sinuous appearance in lateral view. That this morphology is likely a result of deformation is evinced in IRSNB 1536 by the sacral neural spines being inclined noticeably to the left side of the animal (
<figureCitation id="F8902A7AB11AFFFFC922559DFBBCF96D" box="[1036,1119,1760,1784]" captionStart="FIGURE 10" captionStartId="12.[151,250,1212,1235]" captionTargetBox="[501,1087,298,1189]" captionTargetId="figure@12.[496,1091,294,1191]" captionTargetPageId="12" captionText="FIGURE 10. Sacrum and ilia of IRSNB 1536 (Iguanodon bernissartensis) in dorsal view." httpUri="https://zenodo.org/record/200821/files/figure.png" pageId="10" pageNumber="62">Fig. 10</figureCitation>
), an obvious result of deformation that could have also affected the ilia.
</paragraph>
<paragraph id="601436FFB11AFFFECDEB5455FBB6FE89" blockId="10.[151,1436,1508,2036]" lastBlockId="11.[151,1436,151,284]" lastPageId="11" lastPageNumber="63" pageId="10" pageNumber="62">
Furthermore, the
<typeStatus id="BF10885DB11AFFFFCCA75455FE08F8D5" box="[393,491,1832,1856]" pageId="10" pageNumber="62" type="holotype">holotype</typeStatus>
skeleton of
<taxonomicName id="A7AB4D7CB11AFFFFCF585455FC94F8D5" box="[630,887,1832,1856]" class="Reptilia" family="Hadrosauridae" genus="Uteodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="10" pageNumber="62" phylum="Chordata" rank="species" species="aphanoecetes">
<emphasis id="52DFEAEDB11AFFFFCF585455FC94F8D5" box="[630,887,1832,1856]" italics="true" pageId="10" pageNumber="62">Uteodon aphanoecetes</emphasis>
</taxonomicName>
, CM 11337, exhibits a left ilium strikingly similar to that of USNM
<quantity id="A7539B1AB11AFFFFCCA95431FE3EF8F1" box="[391,477,1868,1892]" metricMagnitude="2" metricUnit="m" metricValue="1.207262" pageId="10" pageNumber="62" unit="in" value="4753.0">4753 in</quantity>
lateral and caudal views, with the dorsal margin of the ilium facing almost laterally and the blade of the ilium projecting dorsolaterally far past the ischial peduncle (
<figureCitation id="F8902A7AB11AFFFFC90C540DFB92F81D" box="[1058,1137,1904,1928]" captionStart="FIGURE 6" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1424,927,1709]" captionTargetId="figure@7.[151,1436,913,1723]" captionTargetPageId="7" captionText="FIGURE 6. Left ilium of CM 11337 (holotype of Uteodon aphanoecetes) in (A) ventrolateral view showing the convex dorsal margin, (B) lateral view, (C), caudal view. Sacrum and ilia of CM 11337 in (D) dorsal view. Right ilium of CM 11337 in (E) lateral view. Right ilium of OXFUM J. 3303 (holotype of Cumnoria prestwichii) in (F) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process." httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="10" pageNumber="62">Figs. 6</figureCitation>
B, C). That this is a consequence of plastic deformation in CM 11337 is indicated by the condition of the sacral neural spines, which incline sharply towards the left side of the skeleton, as documented above in IRSNB 1536 (
<figureCitation id="F8902A7AB11AFFFFC96754C5FB6DF845" box="[1097,1166,1976,2000]" captionStart="FIGURE 6" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1424,927,1709]" captionTargetId="figure@7.[151,1436,913,1723]" captionTargetPageId="7" captionText="FIGURE 6. Left ilium of CM 11337 (holotype of Uteodon aphanoecetes) in (A) ventrolateral view showing the convex dorsal margin, (B) lateral view, (C), caudal view. Sacrum and ilia of CM 11337 in (D) dorsal view. Right ilium of CM 11337 in (E) lateral view. Right ilium of OXFUM J. 3303 (holotype of Cumnoria prestwichii) in (F) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process." httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="10" pageNumber="62">Fig. 6</figureCitation>
D; see also
<bibRefCitation id="043A4B0EB11AFFFFC83E54C5FEC8F861" author="Carpenter" pageId="10" pageNumber="62" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson 2008</bibRefCitation>
, fig. 13E); the blade of the left ilium likely was subject to the same force that distorted the neural spines. Furthermore, while the blade of the right ilium of CM 11337 is incomplete, enough is preserved to indicate that the iliac blade was nearly vertical and projected dorsally relative to the ischial peduncle, as in ilia of
<taxonomicName id="A7AB4D7CB11BFFFEC81453C0FED2FF6D" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="63" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB11BFFFEC81453C0FED2FF6D" italics="true" pageId="11" pageNumber="63">Camptosaurus dispar</emphasis>
</taxonomicName>
(e.g., YPM 1877, YPM 8622, USNM 5473;
<figureCitation id="F8902A7AB11BFFFECE1F53A2FC9CFF62" box="[817,895,223,247]" captionStart="FIGURE 4" captionStartId="6.[151,250,720,743]" captionTargetBox="[151,1436,193,698]" captionTargetId="figure@6.[151,1436,193,699]" captionTargetPageId="6" captionText="FIGURE 4. Ilia of Camptosaurus dispar in lateral view. (A) right ilium of YPM 1877 (holotype) from Quarry 13; (B) right ilium of YPM 8622 from Quarry 13; (C) left ilium of USNM 5473 from Quarry 13; (D) left ilium of YPM PU 14553 from Cleveland-Lloyd Dinosaur Quarry. Abbreviations: mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process. Images in A, B, and D are copyright YPM. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200815/files/figure.png" pageId="11" pageNumber="63">Figs. 4</figureCitation>
A–C;
<figureCitation id="F8902A7AB11BFFFECEE253A2FBECFF62" box="[972,1039,223,247]" captionStart="FIGURE 6" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1424,927,1709]" captionTargetId="figure@7.[151,1436,913,1723]" captionTargetPageId="7" captionText="FIGURE 6. Left ilium of CM 11337 (holotype of Uteodon aphanoecetes) in (A) ventrolateral view showing the convex dorsal margin, (B) lateral view, (C), caudal view. Sacrum and ilia of CM 11337 in (D) dorsal view. Right ilium of CM 11337 in (E) lateral view. Right ilium of OXFUM J. 3303 (holotype of Cumnoria prestwichii) in (F) lateral view. Abbreviations: dmi, dorsal margin of ilium; mlt, mediolateral thickening; pas, break in slope along dorsal margin of postacetabular process." httpUri="https://zenodo.org/record/200817/files/figure.png" pageId="11" pageNumber="63">Fig. 6</figureCitation>
E). The situation in USNM 4753 is akin to that of CM 11337, i.e. one ilium is plastically deformed while the other is not.
</paragraph>
<caption id="34D46677B11BFFFECDB9559BFCD6F88E" httpUri="https://zenodo.org/record/200820/files/figure.png" pageId="11" pageNumber="63" targetBox="[152,1390,343,1723]" targetPageId="11">
<paragraph id="601436FFB11BFFFECDB9559BFCD6F88E" blockId="11.[151,1436,1766,1820]" pageId="11" pageNumber="63">
<emphasis id="52DFEAEDB11BFFFECDB9559BFEF0F969" bold="true" box="[151,275,1766,1789]" pageId="11" pageNumber="63">FIGURE 9.</emphasis>
Left ilia of
<taxonomicName id="A7AB4D7CB11BFFFECCA3559AFD78F968" box="[397,667,1767,1789]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="63" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="52DFEAEDB11BFFFECCA3559AFD78F968" box="[397,667,1767,1789]" italics="true" pageId="11" pageNumber="63">Iguanodon bernissartensis</emphasis>
</taxonomicName>
in lateral view. (
<emphasis id="52DFEAEDB11BFFFECE69559AFCBAF969" bold="true" box="[839,857,1767,1788]" pageId="11" pageNumber="63">A</emphasis>
) IRSNB 1534 (lectotype), dorsal margin outlined in red; (
<emphasis id="52DFEAEDB11BFFFECDB1547BFF53F88E" bold="true" box="[159,176,1798,1819]" pageId="11" pageNumber="63">B</emphasis>
) IRSNB 1536, dorsal margin outlined in red; (
<emphasis id="52DFEAEDB11BFFFECFA45478FD7FF889" bold="true" box="[650,668,1797,1820]" pageId="11" pageNumber="63">C</emphasis>
) IRSNB 1561.
</paragraph>
</caption>
<paragraph id="601436FFB11BFFF9CDEB5435FCAFFF62" blockId="11.[151,1436,1864,2032]" lastBlockId="12.[151,1436,151,248]" lastPageId="12" lastPageNumber="64" pageId="11" pageNumber="63">
The preserved morphology of USNM 4753 is itself evidence against the supposed horizontal postacetabular process. The preserved part of the right ilium of USNM 4753 does not exhibit any hint of a folded, horizontally projecting iliac blade (
<emphasis id="52DFEAEDB11BFFFECCB654ECFE02F83D" box="[408,481,1937,1960]" italics="true" pageId="11" pageNumber="63">contra</emphasis>
<bibRefCitation id="043A4B0EB11BFFFECCC454EDFCF1F83D" author="Carpenter" box="[490,786,1936,1960]" pageId="11" pageNumber="63" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson 2008</bibRefCitation>
) (
<figureCitation id="F8902A7AB11BFFFECE0454EDFC8DF83D" box="[810,878,1936,1960]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="11" pageNumber="63">Fig. 8</figureCitation>
E). Indeed, the cross-section of the right ilium in caudal view indicates that the iliac blade was nearly vertical, as in
<taxonomicName id="A7AB4D7CB11BFFFECE5054C8FB8CF859" box="[894,1135,1972,1996]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="11" pageNumber="63" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB11BFFFECE5054C8FB8CF859" box="[894,1135,1972,1996]" italics="true" pageId="11" pageNumber="63">Camptosaurus dispar</emphasis>
</taxonomicName>
(
<figureCitation id="F8902A7AB11BFFFEC95054C9FB22F859" box="[1150,1217,1972,1996]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="11" pageNumber="63">Fig. 8</figureCitation>
F). Moreover, comparing the left ilium with the preserved portion of the right reveals that if the morphology of the left ilium were genuine, the right ilium should exhibit the cranial region of the folded iliac blade (
<figureCitation id="F8902A7AB11CFFF9C91453EAFB6EFF3A" box="[1082,1165,151,175]" captionStart="FIGURE 11" captionStartId="13.[151,250,1103,1126]" captionTargetBox="[151,1436,327,1082]" captionTargetId="figure@13.[151,1436,327,1082]" captionTargetPageId="13" captionText="FIGURE 11. Left (A) and right (B, reversed) ilia of USNM 4753 (holotype of Osmakasaurus depressus) in lateral view, set to the same scale and with the notch between the preacetabular process and pubic peduncle lined up. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200822/files/figure.png" pageId="12" pageNumber="64">Fig. 11</figureCitation>
). Thus, the morphology used by
<bibRefCitation id="043A4B0EB11CFFF9CDDD53C1FE42FF41" author="Gilmore" box="[243,417,188,212]" pageId="12" pageNumber="64" refString="Gilmore, C. W. (1909) Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum, 36, 197 - 332." type="journal article" year="1909">Gilmore (1909)</bibRefCitation>
to demarcate
<taxonomicName id="A7AB4D7CB11CFFF9CF1353C0FD29FF41" box="[573,714,188,212]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CF1353C0FDB3FF41" box="[573,592,189,212]" italics="true" pageId="12" pageNumber="64">C</emphasis>
.
<emphasis id="52DFEAEDB11CFFF9CF7353C1FD29FF41" box="[605,714,188,212]" italics="true" pageId="12" pageNumber="64">depressus</emphasis>
</taxonomicName>
and by
<bibRefCitation id="043A4B0EB11CFFF9CE0C53C1FBB0FF41" author="Carpenter" box="[802,1107,188,212]" pageId="12" pageNumber="64" refString="Carpenter, K. &amp; Wilson, Y. (2008) A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb. Annals of the Carnegie Museum, 76, 227 - 263." type="journal article" year="2008">Carpenter &amp; Wilson (2008)</bibRefCitation>
to assign the species to
<taxonomicName id="A7AB4D7CB11CFFF9C87053C1FF37FF6D" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB11CFFF9C87053C1FF37FF6D" italics="true" pageId="12" pageNumber="64">Planicoxa</emphasis>
</taxonomicName>
is the product of plastic deformation and fragmentation.
</paragraph>
<caption id="34D46677B11CFFF9CDB957C1FBCAFB46" box="[151,1065,1212,1235]" httpUri="https://zenodo.org/record/200821/files/figure.png" pageId="12" pageNumber="64" targetBox="[501,1087,298,1189]" targetPageId="12">
<paragraph id="601436FFB11CFFF9CDB957C1FBCAFB46" blockId="12.[151,1065,1212,1235]" box="[151,1065,1212,1235]" pageId="12" pageNumber="64">
<emphasis id="52DFEAEDB11CFFF9CDB957C1FEFCFB47" bold="true" box="[151,287,1212,1235]" pageId="12" pageNumber="64">FIGURE 10.</emphasis>
Sacrum and ilia of IRSNB 1536 (
<taxonomicName id="A7AB4D7CB11CFFF9CF5657C0FC66FB46" box="[632,901,1213,1235]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="52DFEAEDB11CFFF9CF5657C0FC66FB46" box="[632,901,1213,1235]" italics="true" pageId="12" pageNumber="64">Iguanodon bernissartensis</emphasis>
</taxonomicName>
) in dorsal view.
</paragraph>
</caption>
<paragraph id="601436FFB11CFFF9CDEB5782FC10F979" blockId="12.[151,1436,1279,1773]" pageId="12" pageNumber="64">
Although crushed and broken, a unique combination of characters can be discerned in the ilia of USNM 4753 (see “Specific diagnosis” above). The dorsal margin of the left ilium faces dorsolaterally due to distortion of the iliac blade (
<figureCitation id="F8902A7AB11CFFF9CC34563AFE85FACA" box="[282,358,1351,1375]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="12" pageNumber="64">Figs. 8</figureCitation>
A, C), making the ilium appear dorsally convex; however, the dorsal margin itself lacks any curvature and is straight. The dorsal margin of the ilium thickens mediolaterally towards the postacetabular process (
<figureCitation id="F8902A7AB11CFFF9CDB156F2FF08FA32" box="[159,235,1423,1447]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="12" pageNumber="64">Figs. 8</figureCitation>
A, C).
<taxonomicName id="A7AB4D7CB11CFFF9CC1656EDFE25FA3D" box="[312,454,1424,1448]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CC1656EDFEA8FA32" box="[312,331,1424,1447]" italics="true" pageId="12" pageNumber="64">C</emphasis>
.
<emphasis id="52DFEAEDB11CFFF9CC7756EDFE25FA3D" box="[345,454,1424,1448]" italics="true" pageId="12" pageNumber="64">depressus</emphasis>
</taxonomicName>
is more derived than
<taxonomicName id="A7AB4D7CB11CFFF9CF9756EDFC49FA3D" box="[697,938,1424,1448]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="dispar">
<emphasis id="52DFEAEDB11CFFF9CF9756EDFC49FA3D" box="[697,938,1424,1448]" italics="true" pageId="12" pageNumber="64">Camptosaurus dispar</emphasis>
</taxonomicName>
in exhibiting a horizontal boot at the cranial end of the preacetabular process (
<figureCitation id="F8902A7AB11CFFF9CF3056C9FD80FA59" box="[542,611,1460,1484]" captionStart="FIGURE 8" captionStartId="10.[151,250,1380,1403]" captionTargetBox="[151,1436,537,1358]" captionTargetId="figure@10.[151,1436,537,1359]" captionTargetPageId="10" captionText="FIGURE 8. Left ilium of USNM 4753 (holotype of Osmakasaurus depressus) in (A) lateral, (B) medial, (C) caudolateral, and (D) caudal views. Right ilium of USNM 4753 in (E) lateral and (F) caudal views. Abbreviations: hb, horizontal boot; mlt, mediolateral thickening. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200819/files/figure.png" pageId="12" pageNumber="64">Fig. 8</figureCitation>
E), a feature shared with other more derived basal iguanodonts such as
<taxonomicName id="A7AB4D7CB11CFFF9CDB956A5FE6CFA65" box="[151,399,1496,1520]" class="Reptilia" genus="Cedrorestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="crichtoni">
<emphasis id="52DFEAEDB11CFFF9CDB956A5FE6CFA65" box="[151,399,1496,1520]" italics="true" pageId="12" pageNumber="64">Cedrorestes crichtoni</emphasis>
</taxonomicName>
(DMNH 47994),
<taxonomicName id="A7AB4D7CB11CFFF9CF4256A5FC7FFA65" box="[620,924,1496,1520]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="bernissartensis">
<emphasis id="52DFEAEDB11CFFF9CF4256A5FC7FFA65" box="[620,924,1496,1520]" italics="true" pageId="12" pageNumber="64">Iguanodon bernissartensis</emphasis>
</taxonomicName>
(IRSNB 1534), and
<emphasis id="52DFEAEDB11CFFF9C9B056A5FA7FFA65" box="[1182,1436,1496,1520]" italics="true" pageId="12" pageNumber="64">Eolambia caroljonesa</emphasis>
(CEUM 52090). It might seem acceptable to refer
<taxonomicName id="A7AB4D7CB11CFFF9CFFA5680FC81F981" box="[724,866,1532,1556]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CFFA5680FD04F981" box="[724,743,1533,1556]" italics="true" pageId="12" pageNumber="64">C</emphasis>
.
<emphasis id="52DFEAEDB11CFFF9CFDB5681FC81F981" box="[757,866,1532,1556]" italics="true" pageId="12" pageNumber="64">depressus</emphasis>
</taxonomicName>
to
<taxonomicName id="A7AB4D7CB11CFFF9CEA65681FBE6F981" box="[904,1029,1532,1556]" class="Reptilia" family="Iguanodontidae" genus="Dakotadon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB11CFFF9CEA65681FBE6F981" box="[904,1029,1532,1556]" italics="true" pageId="12" pageNumber="64">Dakotadon</emphasis>
</taxonomicName>
because USNM 4753 and the
<typeStatus id="BF10885DB11CFFF9C84F5681FF24F9AC" pageId="12" pageNumber="64" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A7AB4D7CB11CFFF9CDDE555CFE71F9AD" box="[240,402,1568,1592]" class="Reptilia" family="Iguanodontidae" genus="Dakotadon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="lakotaensis">
<emphasis id="52DFEAEDB11CFFF9CDDE555CFEE7F9AD" box="[240,260,1569,1592]" italics="true" pageId="12" pageNumber="64">D</emphasis>
.
<emphasis id="52DFEAEDB11CFFF9CC3A555DFE71F9AD" box="[276,402,1568,1592]" italics="true" pageId="12" pageNumber="64">lakotaensis</emphasis>
</taxonomicName>
, SDSM 8656, both came from the Lakota Formation of South Dakota. However, the only overlapping elements between the two specimens are poorly preserved caudal vertebrae that display no features that would positively link “
<emphasis id="52DFEAEDB11CFFF9CCE95515FE39F9EA" box="[455,474,1640,1663]" italics="true" pageId="12" pageNumber="64">C</emphasis>
.”
<taxonomicName id="A7AB4D7CB11CFFF9CCDA5515FD81F915" box="[500,610,1640,1664]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CCDA5515FD81F915" box="[500,610,1640,1664]" italics="true" pageId="12" pageNumber="64">depressus</emphasis>
</taxonomicName>
with
<taxonomicName id="A7AB4D7CB11CFFF9CFB15514FCDCF915" box="[671,831,1640,1664]" class="Reptilia" family="Iguanodontidae" genus="Dakotadon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="lakotaensis">
<emphasis id="52DFEAEDB11CFFF9CFB15514FD50F915" box="[671,691,1641,1664]" italics="true" pageId="12" pageNumber="64">D</emphasis>
.
<emphasis id="52DFEAEDB11CFFF9CFEF5515FCDCF915" box="[705,831,1640,1664]" italics="true" pageId="12" pageNumber="64">lakotaensis</emphasis>
</taxonomicName>
.
<collectingCountry id="18BC766FB11CFFF9CE635515FC8FF915" box="[845,876,1640,1664]" name="American Samoa" pageId="12" pageNumber="64">As</collectingCountry>
the specimen clearly does not pertain to the genus
<taxonomicName id="A7AB4D7CB11CFFF9CDB955F0FED9F931" box="[151,314,1677,1700]" class="Reptilia" family="Camptosauridae" genus="Camptosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="52DFEAEDB11CFFF9CDB955F0FED9F931" box="[151,314,1677,1700]" italics="true" pageId="12" pageNumber="64">Camptosaurus</emphasis>
</taxonomicName>
, cannot be confidently referred to another existing genus, and exhibits a unique combination of characters, a new genus should be created to accommodate “
<emphasis id="52DFEAEDB11CFFF9CE6F55CDFCB7F952" box="[833,852,1712,1735]" italics="true" pageId="12" pageNumber="64">C</emphasis>
.”
<taxonomicName id="A7AB4D7CB11CFFF9CE4155CDFC3FF95D" box="[879,988,1712,1736]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CE4155CDFC3FF95D" box="[879,988,1712,1736]" italics="true" pageId="12" pageNumber="64">depressus</emphasis>
</taxonomicName>
and achieve taxonomic stability. Hence the novel genus and combination proposed herein,
<taxonomicName id="A7AB4D7CB11CFFF9CFFC55A9FC0FF979" box="[722,1004,1748,1772]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11CFFF9CFFC55A9FC0FF979" box="[722,1004,1748,1772]" italics="true" pageId="12" pageNumber="64">Osmakasaurus depressus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

159
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<subSubSection id="28B16574B11CFFF9CDB954FBFD01F877" pageId="12" pageNumber="64" type="nomenclature">
<paragraph id="601436FFB11CFFF9CDB954FBFD01F835" blockId="12.[151,738,1926,1952]" box="[151,738,1926,1952]" pageId="12" pageNumber="64">
<heading id="3B5C8193B11CFFF9CDB954FBFD01F835" bold="true" box="[151,738,1926,1952]" fontSize="11" level="1" pageId="12" pageNumber="64" reason="1">
<taxonomicName id="A7AB4D7CB11CFFF9CDB954FBFD01F835" authority="DiCroce &amp; Carpenter, 2001" authorityName="DiCroce &amp; Carpenter" authorityYear="2001" box="[151,738,1926,1952]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11CFFF9CDB954FBFD01F835" bold="true" box="[151,738,1926,1952]" pageId="12" pageNumber="64">
<emphasis id="52DFEAEDB11CFFF9CDB954FBFE60F835" bold="true" box="[151,387,1926,1952]" italics="true" pageId="12" pageNumber="64">Planicoxa venenica</emphasis>
<bibRefCitation id="043A4B0EB11CFFF9CCA554FBFD01F835" author="DiCroce" box="[395,738,1926,1952]" pageId="12" pageNumber="64" refString="DiCroce, T. &amp; Carpenter, K. (2001) New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah. In: Tanke, D. &amp; Carpenter, K. (eds), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, pp. 183 - 196." type="book chapter" year="2001">DiCroce &amp; Carpenter, 2001</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="601436FFB11CFFF9CDB954B0FD01F877" blockId="12.[151,738,1997,2019]" box="[151,738,1997,2019]" pageId="12" pageNumber="64">
<bibRefCitation id="043A4B0EB11CFFF9CDB954B0FF29F877" author="Carpenter" box="[151,202,1997,2018]" pageId="12" pageNumber="64" refString="DiCroce, T. &amp; Carpenter, K. (2001) New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah. In: Tanke, D. &amp; Carpenter, K. (eds), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, pp. 183 - 196." type="book chapter" year="2001">2001</bibRefCitation>
<taxonomicName id="A7AB4D7CB11CFFF9CDC754B0FD72F877" authority="DiCroce &amp; Carpenter" authorityName="DiCroce &amp; Carpenter" box="[233,657,1997,2019]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="12" pageNumber="64" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11CFFF9CDC754B0FE53F876" box="[233,432,1997,2019]" italics="true" pageId="12" pageNumber="64">Planicoxa venenica</emphasis>
DiCroce &amp; Carpenter
</taxonomicName>
, p. 190.
</paragraph>
</subSubSection>
<subSubSection id="28B16574B11DFFFBCDB953E5FAE4FEAA" lastPageId="14" lastPageNumber="66" pageId="13" pageNumber="65" type="materials_examined">
<paragraph id="601436FFB11DFFF8CDB953E5FDD2FF3A" blockId="13.[151,1436,151,284]" box="[151,561,151,176]" pageId="13" pageNumber="65">
<emphasis id="52DFEAEDB11DFFF8CDB953E5FEE8FF25" bold="true" box="[151,267,152,176]" pageId="13" pageNumber="65">
<typeStatus id="BF10885DB11DFFF8CDB953E5FEE5FF25" box="[151,262,152,176]" pageId="13" pageNumber="65" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="06BAAE3AB11DFFF8CC3C53E5FE8BFF3A" LSID="urn:lsid:biocol.org:col:34940" box="[274,360,152,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34940" name="Delaware Museum of Natural History" pageId="13" pageNumber="65">DMNH</collectionCode>
42504, left ilium.
</paragraph>
<paragraph id="601436FFB11DFFF8CDEB53C1FE91FE89" blockId="13.[151,1436,151,284]" pageId="13" pageNumber="65">
<emphasis id="52DFEAEDB11DFFF8CDEB53C1FEA7FF41" bold="true" box="[197,324,188,212]" pageId="13" pageNumber="65">
<typeStatus id="BF10885DB11DFFF8CDEB53C1FEDCFF41" box="[197,319,188,212]" pageId="13" pageNumber="65" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="06BAAE3AB11DFFF8CC6253C0FE41FF41" LSID="urn:lsid:biocol.org:col:34940" box="[332,418,189,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34940" name="Delaware Museum of Natural History" pageId="13" pageNumber="65">DMNH</collectionCode>
42511, 42516, 42518, 42519, 42520, 42521, 42522, 42524, 42513, 42515, 42525, 42523, 42526, 42527, 42510, 42514, 42517, 42508, 42507, 42505, 40917, 40914, 40918, 42506, 42509, 42512 (
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).
</paragraph>
<caption id="34D46677B11DFFF8CDB95732FA8EFB11" httpUri="https://zenodo.org/record/200822/files/figure.png" pageId="13" pageNumber="65" targetBox="[151,1436,327,1082]" targetPageId="13">
<paragraph id="601436FFB11DFFF8CDB95732FA8EFB11" blockId="13.[151,1436,1103,1156]" pageId="13" pageNumber="65">
<emphasis id="52DFEAEDB11DFFF8CDB95732FEFDFBF0" bold="true" box="[151,286,1103,1126]" pageId="13" pageNumber="65">FIGURE 11.</emphasis>
Left (
<emphasis id="52DFEAEDB11DFFF8CC70572DFE93FBF0" bold="true" box="[350,368,1104,1125]" pageId="13" pageNumber="65">A</emphasis>
) and right (
<emphasis id="52DFEAEDB11DFFF8CCC7572DFE19FBF0" bold="true" box="[489,506,1104,1125]" pageId="13" pageNumber="65">B</emphasis>
, reversed) ilia of USNM 4753 (holotype of
<taxonomicName id="A7AB4D7CB11DFFF8CE945732FB59FBF0" box="[954,1210,1103,1125]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="65" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11DFFF8CE945732FB59FBF0" box="[954,1210,1103,1125]" italics="true" pageId="13" pageNumber="65">Osmakasaurus depressus</emphasis>
</taxonomicName>
) in lateral view, set to the same scale and with the notch between the preacetabular process and pubic peduncle lined up. Scale bar equals 10 cm.
</paragraph>
</caption>
<caption id="34D46677B11DFFF8CDB954C3FD1DF866" httpUri="https://zenodo.org/record/200823/files/figure.png" pageId="13" pageNumber="65" targetBox="[151,1436,1206,1961]" targetPageId="13">
<paragraph id="601436FFB11DFFF8CDB954C3FD1DF866" blockId="13.[151,1437,1982,2035]" pageId="13" pageNumber="65">
<emphasis id="52DFEAEDB11DFFF8CDB954C3FEC3F841" bold="true" box="[151,288,1982,2005]" pageId="13" pageNumber="65">FIGURE 12.</emphasis>
Left ilium of
<taxonomicName id="A7AB4D7CB11DFFF8CC9D54C2FD98F840" box="[435,635,1983,2005]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="13" pageNumber="65" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11DFFF8CC9D54C2FD98F840" box="[435,635,1983,2005]" italics="true" pageId="13" pageNumber="65">Planicoxa venenica</emphasis>
</taxonomicName>
(DMNH 42504, holotype) in (
<emphasis id="52DFEAEDB11DFFF8CE9754C2FC28F841" bold="true" box="[953,971,1983,2004]" pageId="13" pageNumber="65">A</emphasis>
) dorsal and (
<emphasis id="52DFEAEDB11DFFF8C97B54C2FB85F841" bold="true" box="[1109,1126,1983,2004]" pageId="13" pageNumber="65">B</emphasis>
) lateral views. Abbreivations: cdm, convex dorsal margin of ilium. Scale bar equals 10 cm.
</paragraph>
</caption>
<paragraph id="601436FFB11EFFFBCDEB53EAFC47FF41" blockId="14.[151,1436,151,716]" pageId="14" pageNumber="66">
<emphasis id="52DFEAEDB11EFFFBCDEB53EAFE24FF25" bold="true" box="[197,455,151,176]" pageId="14" pageNumber="66">Locality and horizon.</emphasis>
Tony’s Bone Bed, near Moab, Grand County, Utah (
<bibRefCitation id="043A4B0EB11EFFFBC93153EAFABBFF3A" author="DiCroce" box="[1055,1368,151,175]" pageId="14" pageNumber="66" refString="DiCroce, T. &amp; Carpenter, K. (2001) New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah. In: Tanke, D. &amp; Carpenter, K. (eds), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, pp. 183 - 196." type="book chapter" year="2001">DiCroce &amp; Carpenter 2001</bibRefCitation>
); Poison Strip Member, Cedar Mountain Formation, lower–middle Aptian.
</paragraph>
<paragraph id="601436FFB11EFFFBCDEB53A2FD04FF6D" blockId="14.[151,1436,151,716]" box="[197,743,223,248]" pageId="14" pageNumber="66">
<emphasis id="52DFEAEDB11EFFFBCDEB53A2FE43FF6D" bold="true" box="[197,416,223,248]" pageId="14" pageNumber="66">Generic diagnosis.</emphasis>
<collectingCountry id="18BC766FB11EFFFBCC89539DFE25FF6D" box="[423,454,224,248]" name="American Samoa" pageId="14" pageNumber="66">As</collectingCountry>
for species by monotypy.
</paragraph>
<paragraph id="601436FFB11EFFFBCDEB527EFAE4FEAA" blockId="14.[151,1436,151,716]" pageId="14" pageNumber="66">
<emphasis id="52DFEAEDB11EFFFBCDEB527EFE7DFE89" bold="true" box="[197,414,259,284]" pageId="14" pageNumber="66">Specific diagnosis.</emphasis>
Provisionally diagnostic basal styracosternan distinguished from
<taxonomicName id="A7AB4D7CB11EFFFBC9555279FA77FE89" box="[1147,1428,260,284]" class="Reptilia" family="Hadrosauridae" genus="Osmakasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="66" phylum="Chordata" rank="species" species="depressus">
<emphasis id="52DFEAEDB11EFFFBC9555279FA77FE89" box="[1147,1428,260,284]" italics="true" pageId="14" pageNumber="66">Osmakasaurus depressus</emphasis>
</taxonomicName>
,
<taxonomicName id="A7AB4D7CB11EFFFBCDB95255FE6BFED5" box="[151,392,296,320]" class="Reptilia" genus="Cedrorestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="66" phylum="Chordata" rank="species" species="crichtoni">
<emphasis id="52DFEAEDB11EFFFBCDB95255FE6BFED5" box="[151,392,296,320]" italics="true" pageId="14" pageNumber="66">Cedrorestes crichtoni</emphasis>
</taxonomicName>
, and
<emphasis id="52DFEAEDB11EFFFBCCEB5255FD55FED5" box="[453,694,296,320]" italics="true" pageId="14" pageNumber="66">Iguanacolossus fortis</emphasis>
by the convex dorsal margin of the ilium (
<figureCitation id="F8902A7AB11EFFFBC9BA525AFB05FEAA" box="[1172,1254,295,319]" captionStart="FIGURE 12" captionStartId="13.[151,250,1982,2005]" captionTargetBox="[151,1436,1206,1961]" captionTargetId="figure@13.[151,1436,1206,1962]" captionTargetPageId="13" captionText="FIGURE 12. Left ilium of Planicoxa venenica (DMNH 42504, holotype) in (A) dorsal and (B) lateral views. Abbreivations: cdm, convex dorsal margin of ilium. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200823/files/figure.png" pageId="14" pageNumber="66">Fig. 12</figureCitation>
B).
</paragraph>
</subSubSection>
<subSubSection id="28B16574B11EFFFBCDEB5231FDF5FD59" pageId="14" pageNumber="66" type="discussion">
<paragraph id="601436FFB11EFFFBCDEB5231FDF5FD59" blockId="14.[151,1436,151,716]" pageId="14" pageNumber="66">
<emphasis id="52DFEAEDB11EFFFBCDEB5231FEDAFEF1" bold="true" box="[197,313,332,356]" pageId="14" pageNumber="66">Remarks.</emphasis>
The reassessment of USNM 4753 given above casts doubt upon the supposed horizontal postacetabular process of DMNH 42504, which was the primary feature used by
<bibRefCitation id="043A4B0EB11EFFFBCE995212FB1FFE12" author="DiCroce" box="[951,1276,367,391]" pageId="14" pageNumber="66" refString="DiCroce, T. &amp; Carpenter, K. (2001) New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah. In: Tanke, D. &amp; Carpenter, K. (eds), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, pp. 183 - 196." type="book chapter" year="2001">DiCroce &amp; Carpenter (2001)</bibRefCitation>
in diagnosing
<taxonomicName id="A7AB4D7CB11EFFFBCDB952E9FE96FE39" box="[151,373,404,428]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="66" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11EFFFBCDB952E9FE96FE39" box="[151,373,404,428]" italics="true" pageId="14" pageNumber="66">Planicoxa venenica</emphasis>
</taxonomicName>
. In dorsal view, the postacetabular process of DMNH 42504 is heavily fractured (
<figureCitation id="F8902A7AB11EFFFBC80B52E9FA9BFE39" box="[1317,1400,404,428]" captionStart="FIGURE 12" captionStartId="13.[151,250,1982,2005]" captionTargetBox="[151,1436,1206,1961]" captionTargetId="figure@13.[151,1436,1206,1962]" captionTargetPageId="13" captionText="FIGURE 12. Left ilium of Planicoxa venenica (DMNH 42504, holotype) in (A) dorsal and (B) lateral views. Abbreivations: cdm, convex dorsal margin of ilium. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200823/files/figure.png" pageId="14" pageNumber="66">Fig. 12</figureCitation>
A). There are additional smaller cracks along the dorsal margin of the ilium visible in lateral view (
<figureCitation id="F8902A7AB11EFFFBC9C152CAFAA0FE5A" box="[1263,1347,439,463]" captionStart="FIGURE 12" captionStartId="13.[151,250,1982,2005]" captionTargetBox="[151,1436,1206,1961]" captionTargetId="figure@13.[151,1436,1206,1962]" captionTargetPageId="13" captionText="FIGURE 12. Left ilium of Planicoxa venenica (DMNH 42504, holotype) in (A) dorsal and (B) lateral views. Abbreivations: cdm, convex dorsal margin of ilium. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200823/files/figure.png" pageId="14" pageNumber="66">Fig. 12</figureCitation>
B). It is likely that the fractures on the postacetabular process and dorsal margin are the results of deformation of the ilium, akin to what is proposed above for the left ilium of USNM 4753. If so, then the diagnosis of
<taxonomicName id="A7AB4D7CB11EFFFBC990517DFA7FFD8D" box="[1214,1436,512,536]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="66" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11EFFFBC990517DFA7FFD8D" box="[1214,1436,512,536]" italics="true" pageId="14" pageNumber="66">Planicoxa venenica</emphasis>
</taxonomicName>
loses its key feature. Nevertheless, pending recovery of additional iguanodont specimens from the Poison Strip Member,
<taxonomicName id="A7AB4D7CB11EFFFBCC2A5135FE01FDF5" box="[260,482,584,608]" class="Reptilia" family="Dryosauridae" genus="Planicoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="14" pageNumber="66" phylum="Chordata" rank="species" species="venenica">
<emphasis id="52DFEAEDB11EFFFBCC2A5135FE01FDF5" box="[260,482,584,608]" italics="true" pageId="14" pageNumber="66">Planicoxa venenica</emphasis>
</taxonomicName>
is herein retained as a provisionally viable taxon, as its ilium can still be differentiated from all other basal styracosternan ilia known from the Early Cretaceous of North
<collectingCountry id="18BC766FB11EFFFBC9445111FB29FD11" box="[1130,1226,620,644]" name="United States of America" pageId="14" pageNumber="66">America</collectingCountry>
by its convex dorsal margin. The dorsal margin of the ilium is convex cranial to the distorted part of the bone, indicating that this is probably a real feature (
<figureCitation id="F8902A7AB11EFFFBCC8D51C9FE16FD59" box="[419,501,692,716]" captionStart="FIGURE 12" captionStartId="13.[151,250,1982,2005]" captionTargetBox="[151,1436,1206,1961]" captionTargetId="figure@13.[151,1436,1206,1962]" captionTargetPageId="13" captionText="FIGURE 12. Left ilium of Planicoxa venenica (DMNH 42504, holotype) in (A) dorsal and (B) lateral views. Abbreivations: cdm, convex dorsal margin of ilium. Scale bar equals 10 cm." httpUri="https://zenodo.org/record/200823/files/figure.png" pageId="14" pageNumber="66">Fig. 12</figureCitation>
B).
</paragraph>
</subSubSection>
</treatment>
</document>

114
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<mods:role>
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</mods:role>
<mods:namePart>Knoff, Marcelo</mods:namePart>
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<mods:name type="personal">
<mods:role>
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<mods:namePart>Gomes, Delir Correa</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<treatment ID-GBIF-Taxon="127892287" LSID="urn:lsid:plazi:treatment:E802AB34B97CFC2481A90C5B716C882A" httpUri="http://treatment.plazi.org/id/E802AB34B97CFC2481A90C5B716C882A" lastPageNumber="23" pageId="22" pageNumber="23">
<subSubSection pageId="22" pageNumber="23" type="multiple">
<paragraph pageId="22" pageNumber="23">Taxon classification Animalia Dactylogyridea Dactylogyridae</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="nomenclature">
<paragraph pageId="22" pageNumber="23">
*
<taxonomicName authority="Kritsky &amp; Boeger, 1995" authorityName="Kritsky &amp; Boeger" authorityYear="1995" class="Monogenea" family="Dactylogyridae" genus="Annulotrematoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Annulotrematoides amazonicus" order="Dactylogyridea" pageId="22" pageNumber="23" phylum="Platyhelminthes" rank="species" species="amazonicus">Annulotrematoides amazonicus Kritsky &amp; Boeger, 1995</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="type host">
<paragraph pageId="22" pageNumber="23">Type host.</paragraph>
<paragraph pageId="22" pageNumber="23">
<taxonomicName class="Actinopterygii" family="Curimatidae" genus="Psectrogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psectrogaster rutiloides" order="Characiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="rutiloides">Psectrogaster rutiloides</taxonomicName>
(Kner, 1858) (
<taxonomicName lsidName="Osteichthyes" pageId="22" pageNumber="23" rank="superclass" superclass="Osteichthyes">Osteichthyes</taxonomicName>
:
<taxonomicName family="Curimatidae" lsidName="" pageId="22" pageNumber="23" rank="family">Curimatidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="infection site">
<paragraph pageId="22" pageNumber="23">Infection site.</paragraph>
<paragraph pageId="22" pageNumber="23">Gills.</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="type locality">
<paragraph pageId="22" pageNumber="23">Type locality.</paragraph>
<paragraph pageId="22" pageNumber="23">
Brazil, Amazonas State, Manaus, Furo do
<normalizedToken originalValue="Catalão">Catalao</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="holotype">
<paragraph pageId="22" pageNumber="23">Holotype.</paragraph>
<paragraph pageId="22" pageNumber="23">CHIOC 33364.</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="remarks">
<paragraph pageId="22" pageNumber="23">Remarks.</paragraph>
<paragraph pageId="22" pageNumber="23">
Collection number referred to as CHIOC
<normalizedToken originalValue="“33664”">&quot;33664&quot;</normalizedToken>
in the original description due to a mistake. Paratypes deposited in HWML and USNPC.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="reference">
<paragraph pageId="22" pageNumber="23">Reference.</paragraph>
<paragraph pageId="22" pageNumber="23">
<bibRefCitation author="Kritsky, DC" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="68" pageNumber="69" pagination="528 - 532" title="Neotropical Monogenoidea. 26. Annulotrematoidesamazonicus, a new genus and species (Dactylogyridae: Ancyrocephalinae), from the gills of Psectrogasterrutiloides (Kner) (Teleostei: Characiformes: Curimatidae) from the Brazilian Amazon." volume="103" year="1995">Kritsky and Boeger (1995)</bibRefCitation>
.
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<mods:namePart id="34427485A536A341B1F93C34383F520A">Bhushan, S</mods:namePart>
<mods:nameIdentifier id="1FF8171A078C1CFCB7197AD4A8648A2D" type="email">shashi@cife.edu.in</mods:nameIdentifier>
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<mods:namePart id="9AC9D97FB3935039FDFB98CCBE25A008">Jaiswar, A K</mods:namePart>
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<mods:namePart id="CDF07CCE79BEBAB5F70B7BDF31A1B69A">Ramteke, K</mods:namePart>
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<mods:namePart id="76FE01DEFA8973D6EED61B77DC96BB36">Pathak, V</mods:namePart>
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<mods:titleInfo id="D112E5797F0742925C2DFDD8C7E7804A">
<mods:title id="04DDC92D7949BC96F32FD7BDFDFE67CD">Indian Journal of Geo Marine Sciences</mods:title>
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<mods:part id="706BC6C3C50C970755C747B0D51D4435">
<mods:date id="6C71227F94B7B5C0C9B4349E02E0E1EC">2022</mods:date>
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<mods:number id="C39FE745094DC301AA1F6A9ED7578409">2022-09-30</mods:number>
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<mods:number id="BF92EAADB6578FA944C8B54ECC5F277D">51</mods:number>
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<mods:number id="5950F62BF401AAAD89D525D612E0BB68">9</mods:number>
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<mods:url id="5DB7CD96623D427D7367A06DC4984152">https://www.mendeley.com/catalogue/60c3db70-3ffa-37a7-be4e-1cd88eac0ab8/</mods:url>
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<mods:classification id="1E27194EF170499975B6362AF23724A0">journal article</mods:classification>
<mods:identifier id="C9D871F1540C1F84BF9F4D7EB6AF97E6" type="DOI">10.56042/ijms.v51i09.2332</mods:identifier>
<mods:identifier id="87893D4801818DA5A3BBE29249DAB443" type="ISSN">2582-6727</mods:identifier>
<mods:identifier id="D0019215B06EA0E4068EE6F5AC8394E4" type="Zenodo-Dep">11622251</mods:identifier>
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<heading id="3B5D5771FFFCFFD3FCA6FC700CB45484" bold="true" box="[848,939,971,997]" fontSize="11" level="1" pageId="2" pageNumber="762" reason="1">
<taxonomicName id="A7AA9B9EFFFCFFD3FCA6FC700CB45484" ID-CoL="7DM6V" authority="Forsskal, 1775" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[848,939,971,997]" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="762" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="52DE3C0FFFFCFFD3FCA6FC700CB45484" bold="true" box="[848,939,971,997]" pageId="2" pageNumber="762">Results</emphasis>
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<subSubSection id="28B0B396FFFCFFD3FC86FC490CF350DC" pageId="2" pageNumber="762" type="description">
<paragraph id="6015E01DFFFCFFD3FC86FC490CF350DC" blockId="2.[848,1489,971,1981]" pageId="2" pageNumber="762">
In this study, a total of 237 samples were collected randomly from three landing centers. The results of the descriptive statistics are given in
<tableCitation id="2D28D5A6FFFCFFD3FA96FB8E0AD5532E" box="[1376,1482,1077,1103]" captionStart="Table 1" captionStartId="2.[365,420,287,309]" captionTargetBox="[160,1488,361,738]" captionTargetPageId="2" captionText="Table 1 — Fish length and otolith dimensions measurement of U. vittatus from the Indian coast Sampling Station Sampling location N Measurement Min Max Mean S. D SE" pageId="2" pageNumber="762">Table 1</tableCitation>
. The findings revealed that the left and right otoliths of
<taxonomicName id="A7AA9B9EFFFCFFD3FC80FBC20CE853F3" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[886,1015,1145,1170]" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="762" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="52DE3C0FFFFCFFD3FC80FBC20CE853F3" box="[886,1015,1145,1170]" italics="true" pageId="2" pageNumber="762">U. vittatus</emphasis>
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are not significantly different in size from one another. For the
<collectingCountry id="18BDA08DFFFCFFD3FB51FB210A1553D5" box="[1191,1290,1178,1204]" name="India" pageId="2" pageNumber="762">Mumbai</collectingCountry>
population, the total length, otolith length, otolith width and otolith weight ranged between
<quantity id="A7524DF8FFFCFFD3FB8AFB650A405399" box="[1148,1375,1246,1273]" metricMagnitude="-1" metricUnit="m" metricValue="1.5945" metricValueMax="2.144" metricValueMin="1.045" pageId="2" pageNumber="762" unit="cm" value="15.945" valueMax="21.44" valueMin="10.45">10.45 – 21.44 cm</quantity>
,
<quantity id="A7524DF8FFFCFFD3FA84FB650CD2527B" metricMagnitude="-3" metricUnit="m" metricValue="4.550999999999999" metricValueMax="5.962" metricValueMin="3.14" pageId="2" pageNumber="762" unit="cm" value="0.45509999999999995" valueMax="0.5962" valueMin="0.314">
<geoCoordinate id="059E86DAFFFCFFD3FA84FB650AAB5399" box="[1394,1460,1246,1272]" degrees="0.314" direction="north" orientation="latitude" pageId="2" pageNumber="762" precision="55" value="0.314">0.314</geoCoordinate>
<geoCoordinate id="059E86DAFFFCFFD3FA34FB650C81527A" degrees="0.5962" direction="west" orientation="longitude" pageId="2" pageNumber="762" precision="5" value="-0.5962">– 0.5962</geoCoordinate>
cm
</quantity>
,
<quantity id="A7524DF8FFFCFFD3FC2FFABB0BBA527B" box="[985,1189,1280,1306]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="3.98" metricValueMin="1.12" pageId="2" pageNumber="762" unit="cm" value="0.255" valueMax="0.398" valueMin="0.112">
<geoCoordinate id="059E86DAFFFCFFD3FC2FFABB0B06527B" box="[985,1049,1280,1306]" degrees="0.112" direction="north" orientation="latitude" pageId="2" pageNumber="762" precision="55" value="0.112">0.112</geoCoordinate>
<geoCoordinate id="059E86DAFFFCFFD3FBD4FABB0B66527B" box="[1058,1145,1280,1306]" degrees="0.398" direction="west" orientation="longitude" pageId="2" pageNumber="762" precision="55" value="-0.398">– 0.398</geoCoordinate>
cm
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and
<geoCoordinate id="059E86DAFFFCFFD3FB16FABB0A31527B" box="[1248,1326,1280,1306]" degrees="0.0029" direction="north" orientation="latitude" pageId="2" pageNumber="762" precision="5" value="0.0029">0.0029</geoCoordinate>
<geoCoordinate id="059E86DAFFFCFFD3FAC0FABB0A83527B" box="[1334,1436,1280,1306]" degrees="0.0089" direction="west" orientation="longitude" pageId="2" pageNumber="762" precision="5" value="-0.0089">– 0.0089</geoCoordinate>
gm, respectively (
<tableCitation id="2D28D5A6FFFCFFD3FC07FA9A0B50525A" box="[1009,1103,1313,1339]" captionStart="Table 1" captionStartId="2.[365,420,287,309]" captionTargetBox="[160,1488,361,738]" captionTargetPageId="2" captionText="Table 1 — Fish length and otolith dimensions measurement of U. vittatus from the Indian coast Sampling Station Sampling location N Measurement Min Max Mean S. D SE" pageId="2" pageNumber="762">Table 1</tableCitation>
). For the Kakinada population, the total length, otolith length, otolith width, and otolith weight were ranged between
<quantity id="A7524DF8FFFCFFD3FB0AFADE0AD3521E" box="[1276,1484,1381,1408]" metricMagnitude="-1" metricUnit="m" metricValue="1.4594999999999998" metricValueMax="1.752" metricValueMin="1.167" pageId="2" pageNumber="762" unit="cm" value="14.594999999999999" valueMax="17.52" valueMin="11.67">11.67 – 17.52 cm</quantity>
,
<quantity id="A7524DF8FFFCFFD3FCA6FA3C0B3252C0" box="[848,1069,1415,1441]" metricMagnitude="-3" metricUnit="m" metricValue="4.1049999999999995" metricValueMax="4.87" metricValueMin="3.34" pageId="2" pageNumber="762" unit="cm" value="0.4105" valueMax="0.487" valueMin="0.334">
<geoCoordinate id="059E86DAFFFCFFD3FCA6FA3C0C8D52C0" box="[848,914,1415,1441]" degrees="0.334" direction="north" orientation="latitude" pageId="2" pageNumber="762" precision="55" value="0.334">0.334</geoCoordinate>
<geoCoordinate id="059E86DAFFFCFFD3FC68FA3C0CE452C0" box="[926,1019,1415,1441]" degrees="0.487" direction="west" orientation="longitude" pageId="2" pageNumber="762" precision="55" value="-0.487">– 0.487</geoCoordinate>
cm
</quantity>
,
<quantity id="A7524DF8FFFCFFD3FBC9FA3C0A0352C0" box="[1087,1308,1415,1441]" metricMagnitude="-3" metricUnit="m" metricValue="2.58" metricValueMax="3.04" metricValueMin="2.12" pageId="2" pageNumber="762" unit="cm" value="0.258" valueMax="0.304" valueMin="0.212">
<geoCoordinate id="059E86DAFFFCFFD3FBC9FA3C0B9E52C0" box="[1087,1153,1415,1441]" degrees="0.212" direction="north" orientation="latitude" pageId="2" pageNumber="762" precision="55" value="0.212">0.212</geoCoordinate>
<geoCoordinate id="059E86DAFFFCFFD3FB7BFA3C0BF552C0" box="[1165,1258,1415,1441]" degrees="0.304" direction="west" orientation="longitude" pageId="2" pageNumber="762" precision="55" value="-0.304">– 0.304</geoCoordinate>
cm
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, and
<geoCoordinate id="059E86DAFFFCFFD3FA93FA3C0AAA52C0" box="[1381,1461,1415,1441]" degrees="0.0031" direction="north" orientation="latitude" pageId="2" pageNumber="762" precision="5" value="0.0031">0.0031</geoCoordinate>
<geoCoordinate id="059E86DAFFFCFFD3FA37FA3C0CBE52A2" degrees="0.0075" direction="west" orientation="longitude" pageId="2" pageNumber="762" precision="5" value="-0.0075">– 0.0075</geoCoordinate>
gm, respectively (
<tableCitation id="2D28D5A6FFFCFFD3FB89FA130BC452A3" box="[1151,1243,1448,1474]" captionStart="Table 1" captionStartId="2.[365,420,287,309]" captionTargetBox="[160,1488,361,738]" captionTargetPageId="2" captionText="Table 1 — Fish length and otolith dimensions measurement of U. vittatus from the Indian coast Sampling Station Sampling location N Measurement Min Max Mean S. D SE" pageId="2" pageNumber="762">Table 1</tableCitation>
). Similarly, for Puri population the total length, otolith length, otolith width and otolith weight were ranged between
<quantity id="A7524DF8FFFCFFD3FA8EFA570CDB5149" metricMagnitude="-1" metricUnit="m" metricValue="1.423" metricValueMax="1.659" metricValueMin="1.1869999999999998" pageId="2" pageNumber="762" unit="cm" value="14.23" valueMax="16.59" valueMin="11.87">11.87 – 16.59 cm</quantity>
,
<quantity id="A7524DF8FFFCFFD3FC22F9B50BAF5149" box="[980,1200,1550,1576]" metricMagnitude="-3" metricUnit="m" metricValue="3.6799999999999997" metricValueMax="4.09" metricValueMin="3.27" pageId="2" pageNumber="762" unit="cm" value="0.368" valueMax="0.409" valueMin="0.327">
<geoCoordinate id="059E86DAFFFCFFD3FC22F9B50B095149" box="[980,1046,1550,1576]" degrees="0.327" direction="north" orientation="latitude" pageId="2" pageNumber="762" precision="55" value="0.327">0.327</geoCoordinate>
<geoCoordinate id="059E86DAFFFCFFD3FBD4F9B50B615149" box="[1058,1150,1550,1576]" degrees="0.409" direction="west" orientation="longitude" pageId="2" pageNumber="762" precision="55" value="-0.409">– 0.409</geoCoordinate>
cm
</quantity>
,
<quantity id="A7524DF8FFFCFFD3FB37F9B50A855149" box="[1217,1434,1550,1576]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.72" metricValueMin="2.08" pageId="2" pageNumber="762" unit="cm" value="0.24" valueMax="0.272" valueMin="0.208">
<geoCoordinate id="059E86DAFFFCFFD3FB37F9B50A1C5149" box="[1217,1283,1550,1576]" degrees="0.208" direction="north" orientation="latitude" pageId="2" pageNumber="762" precision="55" value="0.208">0.208</geoCoordinate>
<geoCoordinate id="059E86DAFFFCFFD3FAF8F9B50A755149" box="[1294,1386,1550,1576]" degrees="0.272" direction="west" orientation="longitude" pageId="2" pageNumber="762" precision="55" value="-0.272">– 0.272</geoCoordinate>
cm
</quantity>
and
<geoCoordinate id="059E86DAFFFCFFD3FCA6F9940C8D5128" box="[848,914,1583,1609]" degrees="0.002" direction="north" orientation="latitude" pageId="2" pageNumber="762" precision="55" value="0.002">0.002</geoCoordinate>
<geoCoordinate id="059E86DAFFFCFFD3FC53F98B0B075128" box="[933,1048,1583,1610]" degrees="0.0099" direction="west" orientation="longitude" pageId="2" pageNumber="762" precision="5" value="-0.0099">– 0.0099</geoCoordinate>
gm, respectively (
<tableCitation id="2D28D5A6FFFCFFD3FAE2F9940A655128" box="[1300,1402,1583,1609]" captionStart="Table 1" captionStartId="2.[365,420,287,309]" captionTargetBox="[160,1488,361,738]" captionTargetPageId="2" captionText="Table 1 — Fish length and otolith dimensions measurement of U. vittatus from the Indian coast Sampling Station Sampling location N Measurement Min Max Mean S. D SE" pageId="2" pageNumber="762">Table 1</tableCitation>
).
<collectingCounty id="89749891FFFCFFD3FA69F9940ACF5128" box="[1439,1488,1583,1609]" pageId="2" pageNumber="762">Puri</collectingCounty>
population had the highest correlation between fish length and otolith weight (
<emphasis id="52DE3C0FFFFCFFD3FB65F9CF0BB1511F" box="[1171,1198,1645,1677]" italics="true" pageId="2" pageNumber="762">
R
<superScript id="97DF4D55FFFCFFD3FB53F9D60BB1511F" attach="left" box="[1189,1198,1645,1662]" fontSize="7" pageId="2" pageNumber="762">2</superScript>
</emphasis>
= 0.9101), followed by
<collectingMunicipality id="80717A67FFFCFFD3FCA6F92F0CDF51CF" box="[848,960,1684,1710]" pageId="2" pageNumber="762">Kakinada</collectingMunicipality>
and
<collectingCountry id="18BDA08DFFFCFFD3FBFFF92F0B7351CF" box="[1033,1132,1684,1710]" name="India" pageId="2" pageNumber="762">Mumbai</collectingCountry>
populations (
<figureCitation id="F891FC98FFFCFFD3FAEDF92F0ADD51CE" box="[1307,1474,1684,1711]" captionStart="Fig" captionStartId="3.[160,195,1588,1610]" captionTargetBox="[182,1472,288,1540]" captionTargetId="figure-227@3.[170,1482,275,1549]" captionTargetPageId="3" captionText="Fig. 4 — Relationship between (a – c: Mumbai): a) total length and otolith weight, b) total length and otolith width and c) total length and otolith length; (d – f: Kakinada): d) total length and otolith weight, e) total length and otolith width and f) total length and otolith length; (g – i: Odisha): g) total length and otolith weight, h) total length and otolith width and i) total length and otolith length; of U. vittatus" figureDoi="http://doi.org/10.5281/zenodo.11622261" httpUri="https://zenodo.org/record/11622261/files/figure.png" pageId="2" pageNumber="762">Fig. 4a, d, g</figureCitation>
).
<location id="6575B6C6FFFCFFD3FCA6F90D0C6151B1" LSID="urn:lsid:plazi:treatment:E803510BFFFCFFD3FCA6FC700CF350DC:6575B6C6FFFCFFD3FCA6F90D0C6151B1" box="[848,894,1718,1744]" country="India" county="Puri" latitude="0.002" longLatPrecision="55" longitude="-0.0099" municipality="Kakinada" name="The" pageId="2" pageNumber="762" stateProvince="Odisha">The</location>
maximum correlation between fish length and otolith width was observed in the
<location id="6575B6C6FFFCFFD3FA96F9630ACF5193" LSID="urn:lsid:plazi:treatment:E803510BFFFCFFD3FCA6FC700CF350DC:6575B6C6FFFCFFD3FA96F9630ACF5193" box="[1376,1488,1752,1778]" country="India" county="Puri" latitude="0.002" longLatPrecision="55" longitude="-0.0099" municipality="Kakinada" name="Kakinada" pageId="2" pageNumber="762" stateProvince="Odisha">Kakinada</location>
population (
<emphasis id="52DE3C0FFFFCFFD3FC10F9410B1E5064" box="[998,1025,1780,1811]" italics="true" pageId="2" pageNumber="762">
R
<superScript id="97DF4D55FFFCFFD3FC0EF94F0B1E5064" attach="left" box="[1016,1025,1780,1797]" fontSize="7" pageId="2" pageNumber="762">2</superScript>
</emphasis>
= 0.8957), followed by
<collectingRegion id="A26E2EFFFFFCFFD3FAB2F9410A885075" box="[1348,1431,1786,1812]" country="India" name="Odisha" pageId="2" pageNumber="762">Odisha</collectingRegion>
and
<collectingCountry id="18BDA08DFFFCFFD3FCA6F8A00CAC5054" box="[848,947,1819,1845]" name="India" pageId="2" pageNumber="762">Mumbai</collectingCountry>
populations (
<figureCitation id="F891FC98FFFCFFD3FB96F8A00A1C5057" box="[1120,1283,1819,1846]" captionStart="Fig" captionStartId="3.[160,195,1588,1610]" captionTargetBox="[182,1472,288,1540]" captionTargetId="figure-227@3.[170,1482,275,1549]" captionTargetPageId="3" captionText="Fig. 4 — Relationship between (a – c: Mumbai): a) total length and otolith weight, b) total length and otolith width and c) total length and otolith length; (d – f: Kakinada): d) total length and otolith weight, e) total length and otolith width and f) total length and otolith length; (g – i: Odisha): g) total length and otolith weight, h) total length and otolith width and i) total length and otolith length; of U. vittatus" figureDoi="http://doi.org/10.5281/zenodo.11622261" httpUri="https://zenodo.org/record/11622261/files/figure.png" pageId="2" pageNumber="762">Fig. 4b, e, h</figureCitation>
).
<location id="6575B6C6FFFCFFD3FAE8F8A00A545054" LSID="urn:lsid:plazi:treatment:E803510BFFFCFFD3FCA6FC700CF350DC:6575B6C6FFFCFFD3FAE8F8A00A545054" box="[1310,1355,1819,1845]" country="India" county="Puri" latitude="0.002" longLatPrecision="55" longitude="-0.0099" municipality="Kakinada" name="The" pageId="2" pageNumber="762" stateProvince="Odisha">The</location>
maximum correlation between fish length and otolith length was observed in the
<collectingCountry id="18BDA08DFFFCFFD3FBE9F8E40B9D5018" box="[1055,1154,1887,1913]" name="India" pageId="2" pageNumber="762">Mumbai</collectingCountry>
population (
<emphasis id="52DE3C0FFFFCFFD3FAD0F8DB0A5E500B" box="[1318,1345,1881,1913]" italics="true" pageId="2" pageNumber="762">
R
<superScript id="97DF4D55FFFCFFD3FACEF8E20A5E500B" attach="left" box="[1336,1345,1881,1898]" fontSize="7" pageId="2" pageNumber="762">2</superScript>
</emphasis>
= 0.9306), followed by
<collectingRegion id="A26E2EFFFFFCFFD3FBFCF83B0B4250FB" box="[1034,1117,1920,1946]" country="India" name="Odisha" pageId="2" pageNumber="762">Odisha</collectingRegion>
and
<location id="6575B6C6FFFCFFD3FB4BF83B0A3250FB" LSID="urn:lsid:plazi:treatment:E803510BFFFCFFD3FCA6FC700CF350DC:6575B6C6FFFCFFD3FB4BF83B0A3250FB" box="[1213,1325,1920,1946]" country="India" county="Puri" latitude="0.002" longLatPrecision="55" longitude="-0.0099" municipality="Kakinada" name="Kakinada" pageId="2" pageNumber="762" stateProvince="Odisha">Kakinada</location>
populations (
<figureCitation id="F891FC98FFFCFFD3FCAFF8190CC250DC" box="[857,989,1954,1981]" captionStart="Fig" captionStartId="3.[160,195,1588,1610]" captionTargetBox="[182,1472,288,1540]" captionTargetId="figure-227@3.[170,1482,275,1549]" captionTargetPageId="3" captionText="Fig. 4 — Relationship between (a – c: Mumbai): a) total length and otolith weight, b) total length and otolith width and c) total length and otolith length; (d – f: Kakinada): d) total length and otolith weight, e) total length and otolith width and f) total length and otolith length; (g – i: Odisha): g) total length and otolith weight, h) total length and otolith width and i) total length and otolith length; of U. vittatus" figureDoi="http://doi.org/10.5281/zenodo.11622261" httpUri="https://zenodo.org/record/11622261/files/figure.png" pageId="2" pageNumber="762">Fig. 4c, f, i</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

100
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Genus
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<paragraph id="985F54F4DDA3FE2D8BC55E1B406906BE" pageId="1" pageNumber="84">Type species.</paragraph>
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<taxonomicName id="22FA3FE56F5E664F854F3FD44CA41144" class="Arachnida" family="Telemidae" genus="Pinelema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinelema bailongensis" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="bailongensis">Pinelema bailongensis</taxonomicName>
Wang &amp; Li, 2012 from Guangxi.
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<paragraph id="DF36423F9FBC9B1DC4C9381915141A22" pageId="1" pageNumber="84">Diagnosis and description.</paragraph>
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See Wang and Li (2012,
<bibRefCitation id="6BD8F78E78F666CC178594E127D7FFF6" author="Wang, C" journalOrPublisher="Ecologica Montenegrina" pageId="3" pageNumber="86" pagination="546 - 566" title="Four new species of the spider genus Pinelema (Araneae, Telemidae) from caves in south China." volume="7" year="2016">2016</bibRefCitation>
).
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<paragraph id="4205883F379C29E001DD0218F225446A" pageId="1" pageNumber="84">Composition.</paragraph>
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<taxonomicName id="583527936D9A416048E343C60A7DBADF" class="Arachnida" family="Telemidae" genus="Pinelema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinelema" order="Araneae" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">Pinelema</taxonomicName>
currently comprises eight species, including three new species described here. Five of them occur in the Guangxi Zhuang Autonomous Region:
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,
<taxonomicName id="82E33A993DC3762BD3D444F28565B799" lsidName="P. cunfengensis" pageId="1" pageNumber="84" rank="species" species="cunfengensis">P. cunfengensis</taxonomicName>
sp. n.,
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sp. n.,
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sp. n.,
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; the others are known from Yunnan Province in southern China:
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,
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,
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<normalizedToken id="F6EAF9D78CD5D7614B7672786920D279" originalValue="Varietät">Varietaet</normalizedToken>
ISFS: Checklist: 1008251
<taxonomicName id="F28DE13E780566249C8E4EFDE51AFCBB" class="Magnoliopsida" family="Brassicaceae" kingdom="Plantae" order="Brassicales" phylum="Tracheophyta" rank="family">Brassicaceae</taxonomicName>
<taxonomicName id="9A06CC200206D5D38AD8A39B4201B9D7" class="Magnoliopsida" family="Brassicaceae" genus="Camelina" kingdom="Plantae" order="Brassicales" phylum="Tracheophyta" rank="genus">Camelina</taxonomicName>
<taxonomicName id="9828E7DFD65E8AD9F743B63B349FD636" authority="(L.) Crantz" authorityName="Crantz" baseAuthorityName="L." class="Magnoliopsida" family="Brassicaceae" genus="Camelina" kingdom="Plantae" order="Brassicales" phylum="Tracheophyta" rank="species" species="sativa">Camelina sativa (L.) Crantz</taxonomicName>
<taxonomicName id="B05FA4806FEC070AE01D77C75876FF05" authority="(L.) Crantz" authorityName="Crantz" baseAuthorityName="L." class="Magnoliopsida" family="Brassicaceae" genus="Camelina" kingdom="Plantae" order="Brassicales" phylum="Tracheophyta" rank="species" species="sativa">Camelina sativa (L.) Crantz</taxonomicName>
<taxonomicName id="42AAB5FC823D983EBD5316911DA27947" class="Magnoliopsida" family="Brassicaceae" genus="Camelina" kingdom="Plantae" order="Brassicales" phylum="Tracheophyta" rank="variety" species="sativa" variety="sativa">var. sativa</taxonomicName>
</paragraph>
<paragraph id="93C2C5D155B22B34DAF5D9DD4E2D220D">
<heading id="B9D704A5EC8F611AE005439717823841">Zusammenfassung</heading>
KEINE ANGABE
</paragraph>
<paragraph id="E3B01847BE2BCC2898FD4593AFE55AA7">
<heading id="ED1C767574DC861BEEA0EFE5032116F3">
<normalizedToken id="D8F219146262298EAAE25E214CD06115" originalValue="Ökologie">Oekologie</normalizedToken>
</heading>
</paragraph>
<paragraph id="D6159CDC4DD495734F1ED19029E54C1C">
<heading id="EA434797D566C831C7D6D6BD766E6861">Lebensraum Lebensraum</heading>
nach
<bibRefCitation id="3A9A2772DD8B8E213BAA0CDA3FAA3CBB" author="Delarze, R. &amp; Gonseth, Y. &amp; Eggenberger, S. &amp; Vust, M." location="Bad Hersfeld, Germany" publisher="Ott Verlag" refString="Delarze, R., Gonseth, Y., Eggenberger, S., Vust, M. 2015. Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition). Ott Verlag, Bad Hersfeld, Germany, 456 pp." title="Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition)" year="2015">Delarze &amp; al. 2015</bibRefCitation>
</paragraph>
<paragraph id="2F1279106E3DB5758C5BA5D32DE36B07">
<table id="5B6CC648EE74F7CE61F3FE310CB43244" class="table" inLine="true">
<tr id="2D4BBBD2E51ACFECC166BC0383B22AFA">
<td id="DC6C883F4603005E4158BC75A4844CC2">
<tbody id="FF45B300D8B7A612E8CBB7E4AEF98615">KEINE ANGABE</tbody>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="7371BE64760E501BA4E3EC754EAB71CD" type="biology_ecology">
<paragraph id="FEB993C7B0F4BC2E0C094F8CA11EFFE7">
<normalizedToken id="2AA4AD54FDDA8588D61ABEBA86933977" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="61CE2D24939A4A580A6468DA1AE7A1EC" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="437F95970FE189D665A53096B7351A09">
<table id="773DA324605DC39EE352198188179326" class="table" inLine="true">
<tbody id="0754DC0F914A2F00A6C7E97297C653B5">
<tr id="6E88C6B4E90FB752BA3E07E1266C0816">
<th id="12725B612815C8853AA54D644E5C5D25" colspan="2">
<emphasis id="D93F32E8DB02DD7890A2E5983D216C24" bold="true">Bodenfaktoren</emphasis>
</th>
<th id="E6D027636CFCAFD148719E07BD7FD1E8" colspan="2">
<emphasis id="526498D10EA95F5EF36DFCE169DBE55C" bold="true">Klimafaktoren</emphasis>
</th>
<th id="D35F4B919607E53C3F574297A1B323D0" colspan="2">
<emphasis id="F8D7E2D2325FAD4E18C609AD480B939F" bold="true">Salztoleranz</emphasis>
</th>
</tr>
<tr id="7CE04D862ED53EB73F47905163FA9028">
<td id="A9DF1878057072702B1BD78CF7C72C28">Feuchtezahl F</td>
<td id="95527C76B395D51D8595AC1BC63D19D9" style="text-align:left;width:100px">--</td>
<td id="28C62BD7984977C7BE8D5D4E4917D285">Lichtzahl L</td>
<td id="8EFD56AD23BBA11E7A675E68CEBB8965" style="text-align:left;width:60px">--</td>
<td id="A080197046E98756ADA44F7190AE4FE7">Salzzeichen</td>
<td id="58038E11E16278DC3BFBF269975BB3D4" style="text-align:left;width:60px">--</td>
</tr>
<tr id="2C7403926279A067DA6662506B2609DC">
<td id="6B8F74B17264DF23C4BE7979E5F9897F">Reaktionszahl R</td>
<td id="554D3244BF220DF990B0B511F3A0596C" style="text-align:left">--</td>
<td id="C593DA46F8255542093B0BD5B2B0ADBB">Temperaturzahl T</td>
<td id="AEE720B69B13304E86F71E6132E2239C" style="text-align:left">--</td>
</tr>
<tr id="861FA3CC731A2097AADCDF9401039870">
<td id="044A012E93B90A9FBA4B53EBCBE8D451">
<normalizedToken id="5145A60F26F2B4AC1C7E4EE1B111828D" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="BEBE60809E33E5F7A2B9259A2AE51C98" style="text-align:left">--</td>
<td id="B8988C47B48A330555D726802DAD9E68">
<normalizedToken id="74F0D20937EB9A2C7CFF4596BBBC1557" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="3B3AD3AEBE82B18E72B8D58F1C6E5CF6" style="text-align:left">--</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="CF2CCD2A44764E00A690560FECE8A75B" type="reference_group">
<paragraph id="A97E25282D8F38ADDB1A578F74BAC0F3">
<heading id="2DA9254A4D98BF6648F235C903E059CE">Nomenklatur</heading>
</paragraph>
<paragraph id="F92695DF6980A9F6D105F1B3A2B900EA">
<treatmentCitation id="B6118CAD65EB885C2589DE4C175E83D6">
<heading id="BDEEC55C1A8564E7D167C46507E7E80E">
<normalizedToken id="D72797ED1BF6008981491C6D9B46F91B" originalValue="Gültiger">Gueltiger</normalizedToken>
Name (
<bibRefCitation id="870CB18DE6B10C903119250E54467FE8" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</heading>
:
<taxonomicName id="EC4697E9D351F934F1F9977748553FEF" ID-CoL="Q9DM" authority="(L.) Crantz" authorityName="Crantz" baseAuthorityName="L." class="Magnoliopsida" family="Brassicaceae" genus="Camelina" kingdom="Plantae" order="Brassicales" phylum="Tracheophyta" rank="species" species="sativa">
<emphasis id="8410A40851CF9C0DF406324505D12E4E" italics="true">Camelina sativa</emphasis>
(L.) Crantz
</taxonomicName>
</treatmentCitation>
<taxonomicName id="5E4AC49D259B41CF28C9595E4352C958" class="Magnoliopsida" family="Brassicaceae" genus="Camelina" kingdom="Plantae" order="Brassicales" phylum="Tracheophyta" rank="variety" species="sativa" variety="sativa">
var.
<emphasis id="C7F8E534104C87537A13BCF2E1306286" italics="true">sativa</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E37FAC591F6221E40ABBCADBF2021846" type="vernacular_names">
<paragraph id="0B0A6F70B2DBEB9A743279B4486EFC95">Volksname</paragraph>
</subSubSection>
<subSubSection id="09064CC9B5164A3FCE79A071BF2F6A3D" type="multiple">
<paragraph id="DF9DCA74A6A8217F91136DF73C83C39C">
Deutscher Name: -- Nom
<normalizedToken id="3D67F47A27D9176AD4C2B48BAD5B900D" originalValue="français">francais</normalizedToken>
: -- Nome italiano: --
</paragraph>
</subSubSection>
<subSubSection id="8AE7D96A58D91084DF39A5359809AF64" type="conservation">
<paragraph id="B165707DC0F96EE92B12475A714C2508">
<heading id="594168AE6C76CE454FA5712391E0CEA7">Status Indigenat</heading>
: -
</paragraph>
<paragraph id="62EC1C869794ACA3459B458A860D3494">
<emphasis id="CAFCEA8E8D19AE51865B451973C88433" bold="true">
<heading id="FF1A7E90557CF1666D99CEAA7F62AF25">
Liste der
<normalizedToken id="E6CE2C1A733E339E15BF85C469EC9BF7" originalValue="gefährdeten">gefaehrdeten</normalizedToken>
Pflanzen IUCN
</heading>
(nach
<bibRefCitation id="884C570830476396A0EE745E1FAC4F9C" author="Walter, K.S. &amp; Gillett, H.J." location="Gland, Switzerland and Cambridge, UK" publicationUrl="https://doi.org/10.5962/bhl.title.44833" publisher="IUCN" refString="Walter, K.S. and Gillett, H.J. (editors). 1998. 1997 IUCN Red List of Threatened Plants. IUCN, Gland, Switzerland and Cambridge, UK. DOI: https://doi.org/10.5962/bhl.title.44833" title="IUCN Red List of Threatened Plants" year="1997">Walter &amp; Gillett 1997</bibRefCitation>
):
</emphasis>
Nein
</paragraph>
</subSubSection>
</treatment>
</document>

111
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<mods:namePart>Wei, Na-sen</mods:namePart>
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6.
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Cleptes mandsuricus
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<taxonomicName authorityName="Moczar" authorityYear="1968" class="Insecta" family="Chrysididae" genus="Cleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cleptes (Holcocleptes) mandsuricus" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mandsuricus" subGenus="Holcocleptes">Cleptes (Holcocleptes) mandsuricus</taxonomicName>
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, 1968: 171. Holotype ♂, China: Manchuria: Erzendjanzsy (171 (descr.), 172 (figs 5-7), depository: NHMW)*.
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<paragraph pageId="3" pageNumber="4">
<taxonomicName authorityName="Moczar" authorityYear="1968" class="Insecta" family="Chrysididae" genus="Cleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cleptes (Holcocleptes) mandsuricus" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mandsuricus" subGenus="Holcocleptes">Cleptes (Holcocleptes) mandsuricus</taxonomicName>
:
<bibRefCitation author="Moczar, L" journalOrPublisher="Acta Zoologica" pageId="62" pageNumber="63" pagination="323 - 343" refId="B100" refString="Moczar, L, 1998. Revision of the Cleptes (Holocleptes) species of the World. Acta Zoologica 43 (4): 323 - 343" title="Revision of the Cleptes (Holocleptes) species of the World." volume="43" year="1998">
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1998
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: 325 (cat., aerosus group), 329 (key), 332 (figs 12-13), 337 (China: Manchuria: Erzendjanzsy, tax., descr.).
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<paragraph pageId="3" pageNumber="4">
<taxonomicName authorityName="Moczar" authorityYear="1968" class="Insecta" family="Chrysididae" genus="Cleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cleptes mandsuricus" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mandsuricus">Cleptes mandsuricus</taxonomicName>
:
<bibRefCitation author="Kimsey, LS" journalOrPublisher="Oxford University Press, New York" pageId="60" pageNumber="61" refId="B56" refString="Kimsey, LS, Bohart, RM, 1991. The Chrysidid Wasps of the World. Oxford University Press, New York" title="The Chrysidid Wasps of the World." year="1991">Kimsey and Bohart 1991</bibRefCitation>
: 61 (China: Manchuria: Erzendjanzsy, cat.,
<emphasis italics="true" pageId="3" pageNumber="4">aerosus</emphasis>
group);
<bibRefCitation author="Kurzenko, NV" editor="Lelej, AS" journalOrPublisher="Dalnauka, Vladivostok" pageId="60" pageNumber="61" pagination="998 - 1006" refId="B60" refString="Kurzenko, NV, Lelej, AS, 2007. [Fam. Chrysididae -Chrysidid wasps]. In: Lelej, AS, Ed., [Key to the Insect of Russian Far East, Vol. 4, Part 5]. Dalnauka, Vladivostok: 998 - 1006" title="[Fam. Chrysididae - Chrysidid wasps]." volumeTitle="[Key to the Insect of Russian Far East, Vol. 4, Part 5]." year="2007">Kurzenko and Lelej 2007</bibRefCitation>
: 1002 (Northeast China, cat.);
<bibRefCitation author="Wei, NS" journalOrPublisher="ZooKey" pageId="66" pageNumber="67" pagination="55 - 96" publicationUrl="10.3897/zookeys.362.6175" refId="B168" refString="Wei, NS, Rosa, P, Xu, ZF, 2013. Revision of the Chinese Cleptes (Hymenoptera, Chrysididae) with description of new species. ZooKey 362: 55 - 96, DOI: 10.3897/zookeys.362.6175" title="Revision of the Chinese Cleptes (Hymenoptera, Chrysididae) with description of new species." url="10.3897/zookeys.362.6175" volume="362" year="2013">Wei et al. 2013</bibRefCitation>
: 56 (tab.), 60 (key), 69 (Northeast China, tax.).
</paragraph>
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<paragraph pageId="3" pageNumber="4">Distribution.</paragraph>
<paragraph pageId="3" pageNumber="4">China.</paragraph>
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<subSubSection pageId="3" pageNumber="4" type="remarks">
<paragraph pageId="3" pageNumber="4">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="4">
<bibRefCitation author="Moczar, L" journalOrPublisher="Acta Zoologica" pageId="62" pageNumber="63" pagination="323 - 343" refId="B100" refString="Moczar, L, 1998. Revision of the Cleptes (Holocleptes) species of the World. Acta Zoologica 43 (4): 323 - 343" title="Revision of the Cleptes (Holocleptes) species of the World." volume="43" year="1998">
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(1998)
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added some morphological characteristics to the original description.
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482
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<mods:titleInfo>
<mods:title>Clastoderma confusum (Myxomycetes: Amoebozoa), a remarkable new species of slime mould from Western Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Knight, Karina J.</mods:namePart>
<mods:affiliation>Western Australian Herbarium, Biodiversity and Conservation Science, Department of Biodiversity, Conservation and Attractions, Locked Bag 104, Bentley Delivery Centre, Western Australia 6983</mods:affiliation>
<mods:nameIdentifier type="email">Karina.Knight62@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lado, Carlos</mods:namePart>
<mods:affiliation>Real Jardín Botánico (CSIC), Plaza de Murillo 2, 28014 Madrid, Spain</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Nuytsia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-02-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>31</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>35</mods:start>
<mods:end>40</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4064843</mods:identifier>
<mods:identifier type="GBIF-Dataset">d4a84d4a-1ff2-4786-9229-f8790e7091ab</mods:identifier>
<mods:identifier type="Zenodo-Dep">4064843</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4323648" ID-GBIF-Taxon="168219663" ID-Zenodo-Dep="4323648" LSID="urn:lsid:plazi:treatment:E804878EB72CFF9FFF2DA441FB51FAE1" httpUri="http://treatment.plazi.org/id/E804878EB72CFF9FFF2DA441FB51FAE1" lastPageId="4" lastPageNumber="39" pageId="1" pageNumber="36">
<subSubSection box="[136,707,308,332]" pageId="1" pageNumber="36" type="nomenclature">
<paragraph blockId="1.[136,867,308,332]" box="[136,707,308,332]" pageId="1" pageNumber="36">
<taxonomicName authority="K.J.Knight &amp; Lado" authorityName="K. J. Knight &amp; Lado" authorityYear="2020" box="[136,615,308,332]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="1" pageNumber="36" phylum="Mycetozoa" rank="species" species="confusum" status="sp. nov.">
<emphasis bold="true" box="[136,400,308,332]" pageId="1" pageNumber="36">Clastoderma confusum</emphasis>
K.J.Knight &amp; Lado
</taxonomicName>
,
<emphasis box="[626,707,309,332]" italics="true" pageId="1" pageNumber="36">
<taxonomicNameLabel box="[626,707,309,332]" pageId="1" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="36" type="materials_examined">
<paragraph blockId="1.[136,867,308,332]" box="[713,867,308,332]" pageId="1" pageNumber="36">
(
<accessionNumber box="[721,855,308,332]" pageId="1" pageNumber="36">
<collectionCode box="[721,764,308,332]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="1" pageNumber="36">MB</collectionCode>
832527
</accessionNumber>
).
</paragraph>
<paragraph blockId="1.[136,1202,378,466]" pageId="1" pageNumber="36">
<materialsCitation ID-GBIF-Occurrence="2867605303" collectingDate="2018-07-24" collectingDateMax="2019-01" collectingDateMin="2018-07-24" collectionCode="K, J, CANB" county="Eagle Highway" location="Little Sandy Desert" municipality="Wiluna" pageId="1" pageNumber="36" specimenCode="MC 154, PERTH 09078509" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis box="[136,186,379,402]" italics="true" pageId="1" pageNumber="36">
<typeStatus box="[136,186,379,402]" pageId="1" pageNumber="36">Type</typeStatus>
</emphasis>
:
<collectingCounty box="[200,367,378,402]" pageId="1" pageNumber="36">Eagle Highway</collectingCounty>
,
<quantity box="[378,459,378,402]" metricMagnitude="5" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="36" unit="km" value="450.0">450 km</quantity>
north-east of
<collectingMunicipality box="[608,688,378,402]" pageId="1" pageNumber="36">Wiluna</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E804878EB72CFF9FFF2DA441FB51FAE1:65726043B72CFF9AFD1EA40FFC74FE28" box="[699,909,378,402]" county="Eagle Highway" municipality="Wiluna" name="Little Sandy Desert" pageId="1" pageNumber="36" stateProvince="Western Australia">Little Sandy Desert</location>
,
<collectingRegion box="[920,1113,378,402]" country="Australia" name="Western Australia" pageId="1" pageNumber="36">Western Australia</collectingRegion>
,
<date pageId="1" pageNumber="36" value="2018-07-24">
<collectingDate pageId="1" pageNumber="36" value="2018-07-24">24 July 2018</collectingDate>
</date>
,
<emphasis box="[201,322,410,434]" italics="true" pageId="1" pageNumber="36">
<collectionCode box="[201,222,411,434]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="36" type="Herbarium">K</collectionCode>
.
<collectionCode box="[226,241,411,434]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="1" pageNumber="36" type="Herbarium">J</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:E804878EB72CFF9FFF2DA441FB51FAE1:65726043B72CFF9AFF5CA4EFFEBBFE0B" box="[249,322,410,434]" county="Eagle Highway" municipality="Wiluna" name="Knight" pageId="1" pageNumber="36" stateProvince="Western Australia">Knight</location>
</emphasis>
<specimenCode box="[327,413,410,434]" collectionCode="MC" country="0" name="Museo de Cipolleti" pageId="1" pageNumber="36">MC 154</specimenCode>
[from moist chamber culture of bark of prone dead
<taxonomicName box="[953,1108,410,434]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="36" phylum="Tracheophyta" rank="species" species="aneura">
<emphasis box="[953,1108,410,434]" italics="true" pageId="1" pageNumber="36">Acacia aneura</emphasis>
</taxonomicName>
,
<date pageId="1" pageNumber="36" value="2019-01">
<collectingDate pageId="1" pageNumber="36" value="2019-01">January 2019</collectingDate>
</date>
] (
<emphasis box="[214,261,442,466]" italics="true" pageId="1" pageNumber="36">holo</emphasis>
:
<specimenCode box="[275,475,442,466]" collectionCode="PERTH" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15701" lsid="urn:lsid:biocol.org:col:15701" name="Western Australian Herbarium" pageId="1" pageNumber="36" type="Herbarium">PERTH 09078509</specimenCode>
;
<emphasis box="[486,517,442,465]" italics="true" pageId="1" pageNumber="36">iso</emphasis>
:
<collectionCode box="[531,608,442,466]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15596" lsid="urn:lsid:biocol.org:col:15596" name="Australian National Botanic Gardens" pageId="1" pageNumber="36" type="Herbarium">CANB</collectionCode>
,
<collectionCode box="[618,640,442,465]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="36" type="Herbarium">K</collectionCode>
, MA-Fungi 90498)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="36" type="reference_group">
<paragraph blockId="1.[136,1202,511,568]" pageId="1" pageNumber="36">
<taxonomicName box="[136,274,512,536]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="1" pageNumber="36" phylum="Mycetozoa" rank="genus">
<emphasis box="[136,274,512,536]" italics="true" pageId="1" pageNumber="36">Clastoderma</emphasis>
</taxonomicName>
sp. Mungilli (
<collectionCode box="[419,442,512,536]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="36" type="Herbarium">K</collectionCode>
.
<collectionCode box="[446,458,512,536]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="1" pageNumber="36" type="Herbarium">J</collectionCode>
. Knight
<specimenCode box="[544,633,511,536]" collectionCode="MC" country="0" name="Museo de Cipolleti" pageId="1" pageNumber="36">MC 154</specimenCode>
),
<location LSID="urn:lsid:plazi:treatment:E804878EB72CFF9FFF2DA441FB51FAE1:65726043B72CFF9AFD2CA775FC32FDA1" box="[649,971,512,536]" municipality="FloraBase" name="Western Australian Herbarium" pageId="1" pageNumber="36">Western Australian Herbarium</location>
, in
<emphasis box="[1004,1117,512,536]" italics="true" pageId="1" pageNumber="36">
<collectingMunicipality box="[1004,1117,512,536]" pageId="1" pageNumber="36">FloraBase</collectingMunicipality>
</emphasis>
, https:// florabase.dpaw.wa.gov.au/ [accessed
<date box="[532,704,544,568]" pageId="1" pageNumber="36" value="2019-10-03">
<collectingDate box="[532,704,544,568]" pageId="1" pageNumber="36" value="2019-10-03">3 October 2019</collectingDate>
</date>
].
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="36" type="description">
<paragraph blockId="1.[136,1202,613,1470]" pageId="1" pageNumber="36">
<emphasis box="[136,258,614,637]" italics="true" pageId="1" pageNumber="36">Sporocarps</emphasis>
gregarious, stalked,
<quantity box="[473,625,613,637]" metricMagnitude="-4" metricUnit="m" metricValue="3.55" metricValueMax="4.6" metricValueMin="2.5" pageId="1" pageNumber="36" unit="mm" value="0.355" valueMax="0.46" valueMin="0.25">0.25–0.46 mm</quantity>
high.
<emphasis box="[688,818,613,637]" italics="true" pageId="1" pageNumber="36">Hypothallus</emphasis>
bulbous and/or disc-like, sometimes insignificant, black, white or light brownish.
<emphasis box="[633,725,645,669]" italics="true" pageId="1" pageNumber="36">Stalks c.</emphasis>
75% total height of the sporocarp, curved, frequently nodding, cylindrical, gently tapering from base to near the apex, with a slightly flared apex subtended by a small, straight section
<emphasis box="[600,619,710,733]" italics="true" pageId="1" pageNumber="36">c.</emphasis>
10–40
<emphasis bold="true" box="[699,712,710,734]" pageId="1" pageNumber="36">×</emphasis>
<quantity box="[719,815,709,734]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="36" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 µm</quantity>
; surface longitudinally rugose, dull, black or dark brown, sometimes with a pale, almost colourless section below the apex; walls pale ochraceous to colourless by transmitted light, grading to jet black
<emphasis box="[843,861,774,797]" italics="true" pageId="1" pageNumber="36">c.</emphasis>
<quantity box="[868,938,773,798]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="36" unit="mm" value="25.0">25 µm</quantity>
below the apex, densely filled with dark, granular refuse basally which becomes lighter and less dense distally.
<emphasis box="[1081,1201,805,829]" italics="true" pageId="1" pageNumber="36">Sporotheca</emphasis>
globose,
<quantity box="[235,360,838,862]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="1" pageNumber="36" unit="mm" value="105.0" valueMax="130.0" valueMin="80.0">80–130 µm</quantity>
diam., dark brown or black.
<emphasis box="[685,784,837,861]" italics="true" pageId="1" pageNumber="36">Peridium</emphasis>
membranous, shiny, fugacious except for a collar and numerous minute and irregular fragments persisting either directly on the external surface of the capillitial mesh or on minute capillitial stubs of variable length along the capillitium, fragments forming an interrupted patterning of a shiny mesh with copper-coloured reflections; brown or violaceous-brown by transmitted light; collar relatively large,
<emphasis box="[825,843,966,989]" italics="true" pageId="1" pageNumber="36">c.</emphasis>
<quantity box="[850,919,966,990]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="36" unit="mm" value="30.0">30 µm</quantity>
diam., irregularly stellatewebbed between the capillitial threads, attached to the capillitium at the base and for a short distance up the threads, minutely striate.
<emphasis box="[442,553,1029,1053]" italics="true" pageId="1" pageNumber="36">Columella</emphasis>
absent.
<emphasis box="[640,762,1029,1053]" italics="true" pageId="1" pageNumber="36">Capillitium</emphasis>
with 4–6 threads radiating at almost right angles from the stalk apex; threads branching and anastomosing, forming a globose, rigid, complete, many-meshed net (the net sometimes collapsed in at the nodes but quickly expanding to globose in Hoyer’s medium); meshes mostly angular, (4–)5–7-sided, nodes not differentiated; threads straight, purple-brown or brown by transmitted light, hollow, thick-walled, flat,
<emphasis box="[920,938,1158,1181]" italics="true" pageId="1" pageNumber="36">c.</emphasis>
<quantity box="[948,1007,1158,1182]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="36" unit="mm" value="2.0">2 µm</quantity>
wide throughout, without free ends, rarely with short, blunt branches within.
<emphasis box="[781,853,1190,1213]" italics="true" pageId="1" pageNumber="36">Spores</emphasis>
globose, (12–)
<quantity box="[1016,1127,1189,1214]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="1" pageNumber="36" unit="mm" value="14.5" valueMax="16.0" valueMin="13.0">13–16 µm</quantity>
diam., dull brown-black or black in mass, mid-brown, brown or pink-brown by transmitted light; surface verruculose, with minute, variably sized and unevenly distributed warts, sometimes arranged in lax, short lines at ×100 with patchy areas of smooth surface, inconspicuous darker patches of warts seen at ×40; by SEM the spore surface is densely minutely ornamented with laxly sinuous ridges, the ridges decorated with struts from apex down to the spore wall and projections on the ridges forming undulating crests, ridges and struts are perforated in patches, warts rounded and occurring on the side of the ridges or between ridges in denser patches, with some thin projections emerging from the base of the wart forming fenestrations. (
<figureCitation box="[508,609,1445,1470]" captionStart="Figure 1" captionStartId="2.[174,230,1045,1064]" captionTargetBox="[172,1241,172,1167]" captionTargetId="figure-18@2.[174,1240,174,1032]" captionTargetPageId="2" captionText="Figure 1. Clastoderma confusum. A – a group of sporocarps in situ showing the gregarious habit; B – sporocarp showing the globose sporotheca with wide-meshed angular capillitium appearing almost completely shiny; C – sporocarp showing the highly reflective large collar; D – sporocarp by transmitted light showing the dark, narrow straight section of stalk just prior to the flared apex; E – spores by transmitted light showing patches of denser warts on surface. Images from K.J. Knight MC 154. Photographs by C. de Mier (A–C) and J.M. Huisman (D, E). Scale bars = 0.1 mm (A–C); 100 µm (D); 50 µm (E)." figureDoi="http://doi.org/10.5281/zenodo.4064845" httpUri="https://zenodo.org/record/4064845/files/figure.png" pageId="1" pageNumber="36">Figures 1</figureCitation>
,
<figureCitation box="[623,637,1446,1470]" captionStart="Figure 2" captionStartId="3.[136,192,1490,1509]" captionTargetBox="[135,1203,172,1638]" captionTargetId="figure-50@3.[136,1202,174,1476]" captionTargetPageId="3" captionText="Figure 2. Clastoderma confusum by Scanning Electron Microscopy.A – sporocarp showing rugose striations on the stalk, which narrows toward the apex; B – collar showing striations, the web-stellate shape and attachment to the capillitium for a short distance up the threads; C – capillitium, with threads radiating at almost right angles from top of the stalk; D – capillitium, showing numerous peridial fragments on capillitial stubs or directly on capillitial net; E, F – different spores; G – spore surface, showing detail of ornamentation. Images from K.J. Knight MC 154. Photographs by K.A. Shepherd (A–D) and C. Lado and Y. Ruiz (E–G). Scale bars: = 50 µm (A); 20 µm (B, D); 10 µm (C, E, F); 3 µm (G)." figureDoi="http://doi.org/10.5281/zenodo.4064847" httpUri="https://zenodo.org/record/4064847/files/figure.png" pageId="1" pageNumber="36">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="36" type="diagnosis">
<paragraph blockId="1.[136,1202,1515,1699]" pageId="1" pageNumber="36">
<emphasis box="[136,346,1515,1539]" italics="true" pageId="1" pageNumber="36">Diagnostic features</emphasis>
. This species is readily distinguished from all other species of
<taxonomicName box="[1030,1168,1515,1539]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="1" pageNumber="36" phylum="Mycetozoa" rank="genus">
<emphasis box="[1030,1168,1515,1539]" italics="true" pageId="1" pageNumber="36">Clastoderma</emphasis>
</taxonomicName>
by the following combination of features: a distinctive, rigid, capillitial net that arises at almost right angles from the top of the stalk; a mostly angular mesh that is ornamented with abundant, highly reflective, peridial fragments that resemble a shiny, copper-coloured net; a stellate-webbed peridial collar situated in close proximity to the capillitium; the absence of a columella; and patchy, verruculose spore ornamentation.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4064845" ID-Zenodo-Dep="4064845" httpUri="https://zenodo.org/record/4064845/files/figure.png" pageId="2" pageNumber="37" startId="2.[174,230,1045,1064]" targetBox="[172,1241,172,1167]" targetPageId="2">
<paragraph blockId="2.[174,1241,1045,1167]" pageId="2" pageNumber="37">
Figure 1.
<taxonomicName authorityName="K. J. Knight &amp; Lado" authorityYear="2020" box="[257,455,1045,1064]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="2" pageNumber="37" phylum="Mycetozoa" rank="species" species="confusum">
<emphasis box="[257,455,1045,1064]" italics="true" pageId="2" pageNumber="37">Clastoderma confusum</emphasis>
</taxonomicName>
. A – a group of sporocarps
<emphasis box="[692,745,1045,1063]" italics="true" pageId="2" pageNumber="37">in situ</emphasis>
showing the gregarious habit; B – sporocarp showing the globose sporotheca with wide-meshed angular capillitium appearing almost completely shiny; C – sporocarp showing the highly reflective large collar; D – sporocarp by transmitted light showing the dark, narrow straight section of stalk just prior to the flared apex; E – spores by transmitted light showing patches of denser warts on surface. Images from
<emphasis box="[1062,1160,1122,1141]" italics="true" pageId="2" pageNumber="37">K.J. Knight</emphasis>
MC 154. Photographs by C. de Mier (A–C) and J.M. Huisman (D, E). Scale bars = 0.1 mm (A–C); 100 µm (D); 50 µm (E).
</paragraph>
</caption>
<subSubSection pageId="2" pageNumber="37" type="materials_examined">
<paragraph blockId="2.[174,1240,1222,1310]" pageId="2" pageNumber="37">
<emphasis box="[174,461,1222,1246]" italics="true" pageId="2" pageNumber="37">Other specimens examined</emphasis>
.
<materialsCitation ID-GBIF-Occurrence="2867605304" box="[475,1119,1222,1246]" collectingDate="2003-04-19" collectionCode="M, H" location="Mt Caroline" pageId="2" pageNumber="37" specimenCount="1" stateProvince="Western Australia">
<collectingRegion box="[475,761,1222,1246]" country="Australia" name="Western Australia" pageId="2" pageNumber="37">WESTERN AUSTRALIA</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:E804878EB72CFF9FFF2DA441FB51FAE1:65726043B72FFF99FCA6A1B3FC71FB67" box="[771,904,1222,1246]" name="Mt Caroline" pageId="2" pageNumber="37" stateProvince="Western Australia">Mt Caroline</location>
,
<date box="[915,1056,1222,1246]" pageId="2" pageNumber="37" value="2003-04-19">
<collectingDate box="[915,1056,1222,1246]" pageId="2" pageNumber="37" value="2003-04-19">19 Apr. 2003</collectingDate>
</date>
,
<emphasis box="[1067,1193,1223,1246]" italics="true" pageId="2" pageNumber="37">
<collectionCode box="[1067,1093,1223,1246]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="2" pageNumber="37" type="Herbarium">M</collectionCode>
.
<collectionCode box="[1096,1119,1223,1246]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="2" pageNumber="37" type="Herbarium">H</collectionCode>
.
<materialsCitation ID-GBIF-Occurrence="2867605302" collectingDate="2019-04-01" collectionCode="PERTH, NE, K, J" county="Brims" location="Laverton on Great Central Road" municipality="Giles Breakaway" pageId="2" pageNumber="37" specimenCount="1" stateProvince="WESTERN AUSTRALIA">
<collectingCounty box="[1129,1193,1223,1246]" pageId="2" pageNumber="37">Brims</collectingCounty>
608 (
<collectionCode box="[181,270,1254,1278]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15701" lsid="urn:lsid:biocol.org:col:15701" name="Western Australian Herbarium" pageId="2" pageNumber="37" type="Herbarium">PERTH</collectionCode>
);
<collectingMunicipality box="[288,472,1254,1278]" pageId="2" pageNumber="37">Giles Breakaway</collectingMunicipality>
,
<quantity box="[482,549,1254,1278]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="37" unit="km" value="50.0">50 km</quantity>
<collectionCode box="[555,591,1254,1277]" country="Australia" lsid="urn:lsid:biocol.org:col:14575" name="University of New England" pageId="2" pageNumber="37" type="Herbarium">NE</collectionCode>
of
<location LSID="urn:lsid:plazi:treatment:E804878EB72CFF9FFF2DA441FB51FAE1:65726043B72FFF99FDD7A193FC33FB47" box="[626,970,1254,1278]" county="Brims" municipality="Giles Breakaway" name="Laverton on Great Central Road" pageId="2" pageNumber="37" stateProvince="WESTERN AUSTRALIA">Laverton on Great Central Road</location>
,
<date box="[981,1108,1254,1278]" pageId="2" pageNumber="37" value="2019-04-01">
<collectingDate box="[981,1108,1254,1278]" pageId="2" pageNumber="37" value="2019-04-01">1 Apr. 2019</collectingDate>
</date>
,
<emphasis box="[1118,1240,1254,1278]" italics="true" pageId="2" pageNumber="37">
<collectionCode box="[1118,1139,1255,1278]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="2" pageNumber="37" type="Herbarium">K</collectionCode>
.
<collectionCode box="[1143,1158,1255,1278]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="2" pageNumber="37" type="Herbarium">J</collectionCode>
.
<materialsCitation ID-GBIF-Occurrence="2867605301" collectionCode="PERTH" location="Knight" pageId="2" pageNumber="37" specimenCode="MC 169" specimenCount="1" stateProvince="WESTERN AUSTRALIA">
<location LSID="urn:lsid:plazi:treatment:E804878EB72CFF9FFF2DA441FB51FAE1:65726043B72FFF99FB2AA193FB21FB47" box="[1167,1240,1254,1278]" name="Knight" pageId="2" pageNumber="37" stateProvince="WESTERN AUSTRALIA">Knight</location>
<specimenCode box="[174,262,1286,1310]" collectionCode="MC" country="0" name="Museo de Cipolleti" pageId="2" pageNumber="37">MC 169</specimenCode>
(
<collectionCode box="[276,366,1286,1310]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15701" lsid="urn:lsid:biocol.org:col:15701" name="Western Australian Herbarium" pageId="2" pageNumber="37" type="Herbarium">PERTH</collectionCode>
)
</materialsCitation>
</emphasis>
</materialsCitation>
</emphasis>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="37" type="biology_ecology">
<paragraph blockId="2.[174,1240,1356,1572]" pageId="2" pageNumber="37">
<emphasis box="[174,528,1356,1380]" italics="true" pageId="2" pageNumber="37">Ecology, distribution and habitat</emphasis>
. Associated with acidic bark (bark of living
<taxonomicName box="[1004,1233,1356,1380]" class="Pinopsida" family="Cupressaceae" genus="Callitris" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="columellaris">
<emphasis box="[1004,1233,1356,1380]" italics="true" pageId="2" pageNumber="37">Callitris columellaris</emphasis>
</taxonomicName>
, pH 4.4 –
<emphasis box="[271,393,1388,1412]" italics="true" pageId="2" pageNumber="37">
<collectionCode box="[271,292,1388,1411]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="2" pageNumber="37" type="Herbarium">K</collectionCode>
.
<collectionCode box="[296,311,1388,1411]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="2" pageNumber="37" type="Herbarium">J</collectionCode>
. Knight
</emphasis>
<collectionCode box="[399,440,1388,1412]" country="0" name="Museo de Cipolleti" pageId="2" pageNumber="37">MC</collectionCode>
169, bark of prone dead
<taxonomicName box="[705,861,1388,1411]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="37" phylum="Tracheophyta" rank="species" species="aneura">
<emphasis box="[705,861,1388,1411]" italics="true" pageId="2" pageNumber="37">Acacia aneura</emphasis>
</taxonomicName>
, pH 5.3 –
<emphasis box="[971,1093,1388,1412]" italics="true" pageId="2" pageNumber="37">
<collectionCode box="[971,992,1388,1411]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="2" pageNumber="37" type="Herbarium">K</collectionCode>
.
<collectionCode box="[996,1011,1388,1411]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="2" pageNumber="37" type="Herbarium">J</collectionCode>
. Knight
</emphasis>
<collectionCode box="[1099,1140,1388,1412]" country="0" name="Museo de Cipolleti" pageId="2" pageNumber="37">MC</collectionCode>
154) and wood-based insect casings.The three known records are disjunct in WesternAustralia (WesternAustralian
<bibRefCitation accessDate="2019-10-03" author="Western Australian Herbarium" box="[174,349,1452,1476]" journalOrPublisher="Department of Biodiversity, Conservation and Attractions" pageId="2" pageNumber="37" publicationUrl="https://florabase.dpaw.wa.gov.au/" refId="ref3015" refString="Western Australian Herbarium (1998 -). FloraBase-the Western Australian Flora. Department of Biodiversity, Conservation and Attractions. https: // florabase. dpaw. wa. gov. au / [accessed 3 October 2019]." title="FloraBase-the Western Australian Flora" type="url" year="1998">Herbarium 1998</bibRefCitation>
–), occurring in semi-arid to arid areas in the Little Sandy Desert, Murchison and Avon Wheatbelt bioregions (
<bibRefCitation accessDate="2019-06-23" author="Department of the Environment" box="[420,833,1484,1508]" journalOrPublisher="Commonwealth of Australia" pageId="2" pageNumber="37" publicationUrl="http://www.environment.gov.au/land/nrs/science/ibraibra" refId="ref2653" refString="Department of the Environment (2013). Australia's bioregions (IBRA), IBRA 7, Commonwealth of Australia. http: // www. environment. gov. au / land / nrs / science / ibra ibra [accessed 23 June 2019]." title="Australia's bioregions (IBRA), IBRA 7" type="url" year="2013">Department of the Environment 2013</bibRefCitation>
). The
<typeStatus box="[901,947,1484,1508]" pageId="2" pageNumber="37">type</typeStatus>
collection and
<emphasis box="[1116,1240,1484,1508]" italics="true" pageId="2" pageNumber="37">
<collectionCode box="[1116,1137,1484,1507]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="2" pageNumber="37" type="Herbarium">K</collectionCode>
.
<collectionCode box="[1141,1156,1484,1507]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="2" pageNumber="37" type="Herbarium">J</collectionCode>
. Knight
</emphasis>
<collectionCode box="[174,215,1516,1540]" country="0" name="Museo de Cipolleti" pageId="2" pageNumber="37">MC</collectionCode>
169 are from sparse mulga over spinifex rangeland with summer rainfall, while
<emphasis box="[1069,1194,1516,1539]" italics="true" pageId="2" pageNumber="37">
<collectionCode box="[1069,1095,1516,1539]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="2" pageNumber="37" type="Herbarium">M</collectionCode>
.
<collectionCode box="[1098,1121,1516,1539]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="2" pageNumber="37" type="Herbarium">H</collectionCode>
. Brims
</emphasis>
608 is from a region characterised by eucalypt woodland or proteaceous scrub with winter rainfall.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="37" type="etymology">
<paragraph blockId="2.[174,1240,1617,1673]" pageId="2" pageNumber="37">
<emphasis box="[174,288,1617,1641]" italics="true" pageId="2" pageNumber="37">Etymology</emphasis>
. From the Latin
<emphasis box="[476,569,1618,1642]" italics="true" pageId="2" pageNumber="37">confusus</emphasis>
(confused), in reference to its previous misidentification as a species of
<taxonomicName box="[286,390,1649,1673]" class="Myxomycetes" family="Cribrariaceae" genus="Cribraria" kingdom="Animalia" order="Liceales" pageId="2" pageNumber="37" phylum="Mycetozoa" rank="genus">
<emphasis box="[286,390,1649,1673]" italics="true" pageId="2" pageNumber="37">Cribraria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="38" pageId="2" pageNumber="37" type="vernacular_names">
<paragraph blockId="2.[174,671,1719,1743]" lastBlockId="3.[136,1202,1673,1761]" lastPageId="3" lastPageNumber="38" pageId="2" pageNumber="37">
<emphasis box="[174,364,1719,1743]" italics="true" pageId="2" pageNumber="37">Vernacular name.</emphasis>
Copper-netted
<taxonomicName box="[529,667,1719,1743]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="2" pageNumber="37" phylum="Mycetozoa" rank="genus">Clastoderma</taxonomicName>
.
<emphasis box="[136,350,1673,1697]" italics="true" pageId="3" pageNumber="38">Conservation status</emphasis>
. Although
<taxonomicName authorityName="K. J. Knight &amp; Lado" authorityYear="2020" box="[472,606,1673,1697]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="3" pageNumber="38" phylum="Mycetozoa" rank="species" species="confusum">
<emphasis box="[472,606,1673,1697]" italics="true" pageId="3" pageNumber="38">C. confusum</emphasis>
</taxonomicName>
is known from only three disjunct collections, it is not considered to be under conservation threat as its occurrence is likely to be more common and widespread.
<taxonomicName box="[136,287,1737,1761]" class="Myxomycetes" kingdom="Protozoa" pageId="3" pageNumber="38" phylum="Mycetozoa" rank="class">Myxomycetes</taxonomicName>
in
<collectingRegion box="[321,511,1737,1761]" country="Australia" name="Western Australia" pageId="3" pageNumber="38">Western Australia</collectingRegion>
are poorly known and rarely collected.
</paragraph>
</subSubSection>
<paragraph pageId="3" pageNumber="38">
<table box="[149,886,610,1469]" gridcols="3" gridrows="3" pageId="3" pageNumber="38">
<tr box="[149,886,610,640]" gridrow="0" pageId="3" pageNumber="38" rowspan-2="1">
<th box="[149,175,610,640]" gridcol="0" gridrow="0" pageId="3" pageNumber="38">A</th>
<th box="[507,530,610,640]" gridcol="1" gridrow="0" pageId="3" pageNumber="38">B</th>
</tr>
<tr box="[149,886,1083,1115]" gridrow="1" pageId="3" pageNumber="38" rowspan-2="1">
<th box="[149,175,1083,1115]" gridcol="0" gridrow="1" pageId="3" pageNumber="38">C</th>
<td box="[507,530,1083,1115]" gridcol="1" gridrow="1" pageId="3" pageNumber="38">D</td>
</tr>
<tr box="[149,886,1438,1469]" gridrow="2" pageId="3" pageNumber="38">
<th box="[149,175,1438,1469]" gridcol="0" gridrow="2" pageId="3" pageNumber="38">E</th>
<td box="[507,530,1438,1469]" gridcol="1" gridrow="2" pageId="3" pageNumber="38">F</td>
<td box="[861,886,1438,1469]" gridcol="2" gridrow="2" pageId="3" pageNumber="38">G</td>
</tr>
</table>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4064847" ID-Zenodo-Dep="4064847" httpUri="https://zenodo.org/record/4064847/files/figure.png" pageId="3" pageNumber="38" startId="3.[136,192,1490,1509]" targetBox="[135,1203,172,1638]" targetPageId="3">
<paragraph blockId="3.[136,1203,1490,1638]" pageId="3" pageNumber="38">
Figure 2.
<taxonomicName authorityName="K. J. Knight &amp; Lado" authorityYear="2020" box="[215,411,1490,1509]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="3" pageNumber="38" phylum="Mycetozoa" rank="species" species="confusum">
<emphasis box="[215,411,1490,1509]" italics="true" pageId="3" pageNumber="38">Clastoderma confusum</emphasis>
</taxonomicName>
by Scanning Electron Microscopy.A – sporocarp showing rugose striations on the stalk, which narrows toward the apex; B – collar showing striations, the web-stellate shape and attachment to the capillitium for a short distance up the threads; C – capillitium, with threads radiating at almost right angles from top of the stalk; D – capillitium, showing numerous peridial fragments on capillitial stubs or directly on capillitial net; E, F – different spores; G – spore surface, showing detail of ornamentation. Images from
<emphasis box="[539,638,1592,1611]" italics="true" pageId="3" pageNumber="38">K.J. Knight</emphasis>
MC 154. Photographs by K.A. Shepherd (A–D) and C. Lado and Y. Ruiz (E–G). Scale bars: = 50 µm (A); 20 µm (B, D); 10 µm (C, E, F); 3 µm (G).
</paragraph>
</caption>
<subSubSection pageId="4" pageNumber="39" type="discussion">
<paragraph blockId="4.[174,1240,174,422]" pageId="4" pageNumber="39">
<emphasis box="[174,270,174,198]" italics="true" pageId="4" pageNumber="39">Affinities</emphasis>
.
<taxonomicName authorityName="K. J. Knight &amp; Lado" authorityYear="2020" box="[283,529,174,198]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="4" pageNumber="39" phylum="Mycetozoa" rank="species" species="confusum">
<emphasis box="[283,529,174,198]" italics="true" pageId="4" pageNumber="39">Clastoderma confusum</emphasis>
</taxonomicName>
appears to be most closely allied to
<taxonomicName authorityName="Kowalski" authorityYear="1975" baseAuthorityName="Meyl." box="[913,1086,175,198]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="4" pageNumber="39" phylum="Mycetozoa" rank="species" species="microcarpum">
<emphasis box="[913,1086,175,198]" italics="true" pageId="4" pageNumber="39">C. microcarpum</emphasis>
</taxonomicName>
, a species that also has a complete, wide-meshed net and large spores,
<quantity box="[778,908,206,231]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.5" metricValueMin="1.35" pageId="4" pageNumber="39" unit="mm" value="14.25" valueMax="15.0" valueMin="13.5">13.5–15 µm</quantity>
diam. (
<bibRefCitation author="Kowalski, D. T." box="[991,1159,206,230]" journalOrPublisher="Mycologia" pageId="4" pageNumber="39" pagination="448 - 494" part="67" refId="ref2825" refString="Kowalski, D. T. (1975). The Myxomycete taxa described by Charles Meylan. Mycologia 67 (3): 448 - 494." title="The Myxomycete taxa described by Charles Meylan" type="journal article" year="1975">Kowalski 1975</bibRefCitation>
). It can readily be distinguished from
<taxonomicName authorityName="Kowalski" authorityYear="1975" baseAuthorityName="Meyl." box="[488,660,239,262]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="4" pageNumber="39" phylum="Mycetozoa" rank="species" species="microcarpum">
<emphasis box="[488,660,239,262]" italics="true" pageId="4" pageNumber="39">C. microcarpum</emphasis>
</taxonomicName>
by its dark brown or black sporocarps (vs ferruginous), black stalk apex (vs red-brown), patchy, verruculose spore ornamentation (vs evenly ornamented with numerous, scattered, minute papillae) and the absence of a columella. It also has a morphologically distinct capillitium with 4–6 threads that emerge at almost right angles from the stalk apex (vs branching from the columella apex into 2–4 main threads), and numerous persistent peridial fragments (vs peridial fragments entirely lacking or occasionally persisting as minute membranous expansions).
</paragraph>
<paragraph blockId="4.[174,1240,468,812]" pageId="4" pageNumber="39">
<taxonomicName authorityName="K. J. Knight &amp; Lado" authorityYear="2020" box="[174,423,468,492]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="4" pageNumber="39" phylum="Mycetozoa" rank="species" species="confusum">
<emphasis box="[174,423,468,492]" italics="true" pageId="4" pageNumber="39">Clastoderma confusum</emphasis>
</taxonomicName>
does not adhere to the most recent diagnosis of the genus and new order
<taxonomicName authority="Leontyev, Schnittler, S. L. Stephenson, Novozhilov &amp; Shchepin" authorityName="Leontyev, Schnittler, S. L. Stephenson, Novozhilov &amp; Shchepin" box="[174,1007,500,525]" order="Clastodermatales" pageId="4" pageNumber="39" rank="order">Clastodermatales Leontyev, Schnittler, S.L. Stephenson, Novozhilov &amp; Shchepin</taxonomicName>
proposed by
<bibRefCitation author="Leontyev, D. V. &amp; Schnittler, M. &amp; Stephenson, S. L. &amp; Novozhilov, Y. K. &amp; Shchepin, O. N." journalOrPublisher="Phytotaxa" pageId="4" pageNumber="39" pagination="209 - 238" part="399" refId="ref2898" refString="Leontyev, D. V., Schnittler, M., Stephenson, S. L., Novozhilov, Y. K. &amp; Shchepin, O. N. (2019). Towards a phylogenetic classification of the Myxomycetes. Phytotaxa 399 (3): 209 - 238." title="Towards a phylogenetic classification of the Myxomycetes" type="journal article" year="2019">
Leontyev
<emphasis box="[174,230,532,556]" italics="true" pageId="4" pageNumber="39">et al.</emphasis>
(2019)
</bibRefCitation>
. These authors describe the columella as always ‘present, gradually turning into the capillitium’ and the capillitial threads as ‘branched and anastomosed, merging at the periphery to form plate-like swellings’. We note, however, that previous studies have observed that the columella may be lacking in
<taxonomicName authorityName="A. Blytt" authorityYear="1880" box="[322,488,628,652]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="4" pageNumber="39" phylum="Mycetozoa" rank="species" species="debaryanum">
<emphasis box="[322,488,628,652]" italics="true" pageId="4" pageNumber="39">C. debaryanum</emphasis>
</taxonomicName>
(Martin &amp; Alexopolous 1969;
<bibRefCitation author="Eliasson, U. H. &amp; Keller, H. W." box="[825,1084,628,652]" journalOrPublisher="Mycological Research" pageId="4" pageNumber="39" pagination="601 - 609" part="100" refId="ref2701" refString="Eliasson, U. H. &amp; Keller, H. W. (1996). The genus Clastoderma: taxonomic evaluation and infraspecific variation of C. debaryanum. Mycological Research 100 (5): 601 - 609." title="The genus Clastoderma: taxonomic evaluation and infraspecific variation of C. debaryanum" type="journal article" year="1996">Eliasson &amp; Keller 1996</bibRefCitation>
;
<bibRefCitation author="Poulain, M. &amp; Meyer, M. &amp; Bozonnet, J." journalOrPublisher="Federation Mycologique et Botanique Dauphine-Savoie: Delemont" pageId="4" pageNumber="39" refId="ref2982" refString="Poulain, M., Meyer, M. &amp; Bozonnet, J. (2011). Les Myxomyc e tes. (Federation Mycologique et Botanique Dauphine-Savoie: Delemont.)" title="Les Myxomyc e tes" type="book" year="2011">
Poulain
<emphasis box="[1185,1240,628,652]" italics="true" pageId="4" pageNumber="39">et al.</emphasis>
2011
</bibRefCitation>
). Furthermore, the peridial fragments in
<taxonomicName authorityName="K. J. Knight &amp; Lado" authorityYear="2020" box="[673,807,660,684]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="4" pageNumber="39" phylum="Mycetozoa" rank="species" species="confusum">
<emphasis box="[673,807,660,684]" italics="true" pageId="4" pageNumber="39">C. confusum</emphasis>
</taxonomicName>
are shiny like the collar and irregularly shaped and thus appear to be of peridial rather than capillitial origin. As such, this contradicts what is indicated by
<bibRefCitation author="Frederick, L. &amp; Roth, I. L. &amp; Pendergrass, L." box="[312,554,724,748]" journalOrPublisher="Mycologia" pageId="4" pageNumber="39" pagination="263 - 268" part="78" refId="ref2743" refString="Frederick, L., Roth, I. L. &amp; Pendergrass, L. (1986). Observations on the ' Peridial Platelets' of Clastoderma debaryanum. Mycologia 78 (2): 263 - 268." title="Observations on the ' Peridial Platelets' of Clastoderma debaryanum" type="journal article" year="1986">
Frederick
<emphasis box="[422,476,724,748]" italics="true" pageId="4" pageNumber="39">et al.</emphasis>
(1986)
</bibRefCitation>
for
<taxonomicName authorityName="A. Blytt" authorityYear="1880" box="[599,763,724,748]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="4" pageNumber="39" phylum="Mycetozoa" rank="species" species="debaryanum">
<emphasis box="[599,763,724,748]" italics="true" pageId="4" pageNumber="39">C. debaryanum</emphasis>
</taxonomicName>
. The new species, therefore, seems to fit the broader concept of the genus
<taxonomicName box="[490,628,756,780]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="4" pageNumber="39" phylum="Mycetozoa" rank="genus">
<emphasis box="[490,628,756,780]" italics="true" pageId="4" pageNumber="39">Clastoderma</emphasis>
</taxonomicName>
, but an exhaustive review of their species is necessary to clarify its distinctive features.
</paragraph>
<paragraph blockId="4.[174,1241,858,1074]" pageId="4" pageNumber="39">
<emphasis box="[174,235,859,882]" italics="true" pageId="4" pageNumber="39">Notes</emphasis>
. The collection
<emphasis box="[409,535,859,882]" italics="true" pageId="4" pageNumber="39">
<collectionCode box="[409,435,859,882]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="4" pageNumber="39" type="Herbarium">M</collectionCode>
.
<collectionCode box="[438,461,859,882]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="4" pageNumber="39" type="Herbarium">H</collectionCode>
. Brims
</emphasis>
608 is in poor condition due to an attack by a filamentous fungus and depauperate since it was split to confirm identification in 2004. It is somewhat atypical in having sporotheca that have mostly collapsed inwards at the capillitial nodes (as some have in the other two collections), slightly darker and smaller spores (brown and
<emphasis box="[820,838,955,978]" italics="true" pageId="4" pageNumber="39">c.</emphasis>
<quantity box="[847,918,954,978]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="39" unit="mm" value="12.0">12 µm</quantity>
diam. vs mid-brown or pinkbrown and
<quantity box="[295,406,986,1010]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="4" pageNumber="39" unit="mm" value="14.5" valueMax="16.0" valueMin="13.0">13–16 µm</quantity>
diam.), and a stalk with a distinctly paler section distally rather than black or dark-brown as in the other collections where the paler section can only be seen by transmitted light. These differences are minor and not considered to be taxonomically significant.
</paragraph>
<paragraph blockId="4.[174,1240,1120,1368]" pageId="4" pageNumber="39">
The
<typeStatus box="[221,267,1120,1144]" pageId="4" pageNumber="39">type</typeStatus>
substrate was not originally collected for use in a moist chamber but rather it was forwarded to the Western Australian Herbarium with the fungi
<taxonomicName box="[753,1020,1152,1176]" class="Agaricomycetes" family="Gloeophyllaceae" genus="Gloeophyllum" kingdom="Fungi" order="Gloeophyllales" pageId="4" pageNumber="39" phylum="Basidiomycota" rank="species" species="abietinum">
<emphasis box="[753,1020,1152,1176]" italics="true" pageId="4" pageNumber="39">Gloeophyllum abietinum</emphasis>
</taxonomicName>
growing on it. It is of interest to note that the substrate for the
<typeStatus box="[648,694,1184,1208]" pageId="4" pageNumber="39">type</typeStatus>
species of the genus,
<taxonomicName authorityName="A. Blytt" authorityYear="1880" box="[937,1103,1184,1208]" class="Myxomycetes" family="Clastodermataceae" genus="Clastoderma" kingdom="Protozoa" order="Echinosteliales" pageId="4" pageNumber="39" phylum="Mycetozoa" rank="species" species="debaryanum">
<emphasis box="[937,1103,1184,1208]" italics="true" pageId="4" pageNumber="39">C. debaryanum</emphasis>
</taxonomicName>
, is probably
<taxonomicName authority="(Eliasson &amp; Keller 1996)" baseAuthorityName="Eliasson &amp; Keller" baseAuthorityYear="1996" box="[174,575,1216,1240]" class="Agaricomycetes" family="Gloeophyllaceae" genus="Gloeophyllum" kingdom="Fungi" order="Gloeophyllales" pageId="4" pageNumber="39" phylum="Basidiomycota" rank="species" species="odoratus">
<emphasis box="[174,301,1216,1240]" italics="true" pageId="4" pageNumber="39">G. odoratus</emphasis>
(
<bibRefCitation author="Eliasson, U. H. &amp; Keller, H. W." box="[313,567,1216,1240]" journalOrPublisher="Mycological Research" pageId="4" pageNumber="39" pagination="601 - 609" part="100" refId="ref2701" refString="Eliasson, U. H. &amp; Keller, H. W. (1996). The genus Clastoderma: taxonomic evaluation and infraspecific variation of C. debaryanum. Mycological Research 100 (5): 601 - 609." title="The genus Clastoderma: taxonomic evaluation and infraspecific variation of C. debaryanum" type="journal article" year="1996">Eliasson &amp; Keller 1996</bibRefCitation>
)
</taxonomicName>
, and although this new species was not growing directly on the hymenium of the fungi as per
<taxonomicName baseAuthorityName="Eliasson &amp; Keller" baseAuthorityYear="1996" box="[500,628,1248,1272]" class="Agaricomycetes" family="Gloeophyllaceae" genus="Gloeophyllum" kingdom="Fungi" order="Gloeophyllales" pageId="4" pageNumber="39" phylum="Basidiomycota" rank="species" species="odoratus">
<emphasis box="[500,628,1248,1272]" italics="true" pageId="4" pageNumber="39">G. odoratus</emphasis>
</taxonomicName>
, there is a possibility of a relationship with this genus of fungus. For Herbarium quarantine purposes, the material was frozen at -18°
<collectionCode box="[985,1003,1280,1304]" country="Denmark" name="University of Copenhagen" pageId="4" pageNumber="39" type="Herbarium">C</collectionCode>
for seven consecutive days and accordingly the viability of spores was expected to be low, yet five species of slime mould were harvested, confirming that this procedure is a safe practice prior to moist chamber culture.
</paragraph>
</subSubSection>
</treatment>
</document>

168
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@ -0,0 +1,168 @@
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<paragraph id="601236E6DE01385EFF790E769FFDC327" blockId="1.[151,502,311,337]" box="[151,502,311,337]" pageId="1" pageNumber="48">
<heading id="3B5A818ADE01385EFF790E769FFDC327" bold="true" box="[151,502,311,337]" fontSize="11" level="1" pageId="1" pageNumber="48" reason="1">
<emphasis id="52D9EAF4DE01385EFF790E769FFDC327" bold="true" box="[151,502,311,337]" pageId="1" pageNumber="48">
<taxonomicName id="A7AD4D65DE01385EFF790E769F9DC327" ID-CoL="5MGM" authority="Wheeler" authorityName="Wheeler" box="[151,406,311,337]" class="Insecta" family="Chloropidae" genus="Medeventor" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE01385EFF790E769F2DC327" bold="true" box="[151,294,311,337]" italics="true" pageId="1" pageNumber="48">Medeventor</emphasis>
Wheeler
</taxonomicName>
, gen. n.
</emphasis>
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</paragraph>
</subSubSection>
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<paragraph id="601236E6DE01385EFF790EC39D5DC3EF" blockId="1.[151,854,386,410]" box="[151,854,386,410]" pageId="1" pageNumber="48">
<typeStatus id="BF168844DE01385EFF790EC39EC7C3EF" box="[151,204,386,409]" pageId="1" pageNumber="48">Type</typeStatus>
species:
<taxonomicName id="A7AD4D65DE01385EFEC30EC39C00C3EF" box="[301,523,386,409]" class="Insecta" family="Chloropidae" genus="Medeventor" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="nubosus" status="sp. nov.">
<emphasis id="52D9EAF4DE01385EFEC30EC39C00C3EF" box="[301,523,386,409]" italics="true" pageId="1" pageNumber="48">Medeventor nubosus</emphasis>
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</emphasis>
(by present designation)
</paragraph>
<paragraph id="601236E6DE01385EFF790E8B9D2FC07A" blockId="1.[151,1437,458,2016]" pageId="1" pageNumber="48">
Diagnosis: Very large (
<quantity id="A7559B03DE01385EFE4A0E8B9C02C392" box="[420,521,458,484]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="48" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
)
<taxonomicName id="A7AD4D65DE01385EFDF60E8B9CA1C392" box="[536,682,458,484]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="subFamily" subFamily="Oscinellinae">Oscinellinae</taxonomicName>
with extensively setulose body and reduced setae, especially on the head; weak, broad facial carina; broad, rugose gena.
</paragraph>
<paragraph id="601236E6DE01385EFF280D5F9C15C126" blockId="1.[151,1437,458,2016]" pageId="1" pageNumber="48">
Description:
<taxonomicName id="A7AD4D65DE01385EFE8E0D5F9FFFC04E" box="[352,500,542,568]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
,
<taxonomicName id="A7AD4D65DE01385EFDEC0D5F9C92C04E" box="[514,665,542,568]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="subFamily" subFamily="Oscinellinae">Oscinellinae</taxonomicName>
. Head about as high as deep in lateral view; frontal triangle distinct, bare except for row of setulae along margin; frons densely setulose, most cephalic setae short and weak, fronto-orbital, vertical, ocellar and postocellar setae barely differentiated from surrounding setulae, ocellar and postocellar setae convergent; eye bare, rounded; gena broad, swollen, divided into densely pruinose dorsal region and broader, shining, rugose ventral region, postgena swollen, setulose (
<figureCitation id="F8962A63DE01385EFB8A0DFF9AA2C0AE" box="[1124,1193,702,728]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="1" pageNumber="48">Fig. 1</figureCitation>
); vibrissal angle not projecting, vibrissa short; face with broad but weak carina between antennal bases, less pronounced ventrally (
<figureCitation id="F8962A63DE01385EFF710C4F9EE2C15E" box="[159,233,782,808]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="1" pageNumber="48">Fig. 2</figureCitation>
); antenna with scape and pedicel short, first flagellomere rounded, arista pubescent; proboscis short; palpus elongate, swollen (
<figureCitation id="F8962A63DE01385EFE260C779C07C126" box="[456,524,822,848]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="1" pageNumber="48">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="601236E6DE01385EFF280C1F9B29C6E6" blockId="1.[151,1437,458,2016]" pageId="1" pageNumber="48">
Scutum convex, densely setulose, thoracic setae weak, 1 postpronotal, 1 anterior and 1–2 posterior notopleural, 1 postsutural supra-alar, 1 postsutural intra-alar, 1 dorsocentral seta; scutellum short, strongly convex, with long, dense lateral and dorsal setae, apical and subapical scutellar setae longer and stronger than surrounding setae; thoracic pleurites bare except for dorsal patch of setulae on katepisternum. Wing hyaline but heavily microtrichose, alula and anal angle well-developed, cell c long and broad, especially in male, second costal sector longer than third; crossvein r-m beyond midpoint of cell dm; crossvein dm-cu perpendicular or slightly oblique; pronounced ventral bulge at wing base (
<figureCitation id="F8962A63DE01385EFCB30B0F9DADC61E" box="[861,934,1102,1128]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="1" pageNumber="48">Fig. 3</figureCitation>
). Legs without outstanding setae or spurs; femoral organ not differentiated from surrounding setulae; tibial organ elongate oval, dark, velvety.
</paragraph>
<paragraph id="601236E6DE01385EFF280BDF9CAFC696" blockId="1.[151,1437,458,2016]" pageId="1" pageNumber="48">
Abdominal tergites with long, dense setae; sternites narrow but distinct, with long setae; male spiracles
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membrane near lateral margin of tergite.
</paragraph>
<paragraph id="601236E6DE01385EFF280BAF9F58C406" blockId="1.[151,1437,458,2016]" pageId="1" pageNumber="48">
Male postabdomen: remnant of sternite 6 median, symmetrical, represented by a pair of sensory setulae and a patch of microtrichia distinct from surrounding membrane; sensory setulae of sternite 7 present posterior to this (
<figureCitation id="F8962A63DE01385EFEC90A7F9F7AC72E" box="[295,369,1342,1368]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="1" pageNumber="48">Fig. 4</figureCitation>
); dorsal pregenital sclerite large, symmetrical, divided by a transverse suture into narrow anterior and broad posterior portions, posterior portion densely setose; spiracles 6 and
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membrane ventral to lateral margin of dorsal sclerite; epandrium simple, convex, with short setae; surstylus simple, elongate, freely articulated with epandrium; hypandrium open posteriorly; pregonite fused with postgonite, sclerotized; postgonite with ventral setulae and campaniform sensillae; basiphallus elongate, sclerotized, distiphallus short, membranous; phallapodeme simple, phallic guide well sclerotized, fused anteriorly with hypandrium (
<figureCitation id="F8962A63DE01385EFF71096F9EE1C43E" box="[159,234,1582,1608]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="1" pageNumber="48">Fig. 7</figureCitation>
); cerci broad, flat, setulose; subepandrial sclerite well-developed, with setulose lobe dorsomedial to surstyli (
<figureCitation id="F8962A63DE01385EFF1309179F4EC406" box="[253,325,1622,1648]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="1" pageNumber="48">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="601236E6DE01385EFF28093F9AD8C4EE" blockId="1.[151,1437,458,2016]" box="[198,1235,1662,1688]" pageId="1" pageNumber="48">Female terminalia not modified, cerci separate, dorsoventrally flattened, broad, setulose.</paragraph>
<paragraph id="601236E6DE01385EFF2809EB9F87C49A" blockId="1.[151,1437,458,2016]" pageId="1" pageNumber="48">
Etymology: The generic name is an anagram of Monteverde, the
<typeStatus id="BF168844DE01385EFC3209EB9A04C4B2" box="[988,1039,1706,1732]" pageId="1" pageNumber="48">type</typeStatus>
locality of the
<typeStatus id="BF168844DE01385EFB2409EB9AF7C4B2" box="[1226,1276,1706,1732]" pageId="1" pageNumber="48">type</typeStatus>
species. It is treated as masculine.
</paragraph>
<paragraph id="601236E6DE01385EFF2809BF9AC0C5CE" blockId="1.[151,1437,458,2016]" pageId="1" pageNumber="48">
Remarks:
<taxonomicName id="A7AD4D65DE01385EFEAF09BF9FC6C561" box="[321,461,1790,1815]" class="Insecta" family="Chloropidae" genus="Medeventor" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE01385EFEAF09BF9FC6C561" box="[321,461,1790,1815]" italics="true" pageId="1" pageNumber="48">Medeventor</emphasis>
</taxonomicName>
appears to be most closely related to the Old World genera
<taxonomicName id="A7AD4D65DE01385EFB5D09BF9B6DC56E" authority="Meigen" authorityName="Meigen" box="[1203,1382,1790,1816]" class="Insecta" family="Chloropidae" genus="Lipara" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE01385EFB5D09BF9B08C561" box="[1203,1283,1790,1815]" italics="true" pageId="1" pageNumber="48">Lipara</emphasis>
Meigen
</taxonomicName>
and
<taxonomicName id="A7AD4D65DE01385EFF7908679F20C549" box="[151,299,1830,1855]" class="Insecta" family="Chloropidae" genus="Pseudeurina" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE01385EFF7908679F20C549" box="[151,299,1830,1855]" italics="true" pageId="1" pageNumber="48">Pseudeurina</emphasis>
</taxonomicName>
de Meijere (
<bibRefCitation id="043C4B17DE01385EFE2D08679C8CC536" author="Andersson" box="[451,647,1830,1856]" pageId="1" pageNumber="53" refString="Andersson, H. (1977) Taxonomic and phylogenetic studies on Chloropidae (Diptera) with special reference to Old World genera. Entomologica scandinavica Supplement, 8, 1 - 200." type="journal article" year="1977">Andersson 1977</bibRefCitation>
,
<bibRefCitation id="043C4B17DE01385EFD7B08679D43C536" author="Kanmiya" box="[661,840,1830,1856]" pageId="1" pageNumber="53" refString="Kanmiya, K. (1983) A systematic study of the Japanese Chloropidae (Diptera). Memoirs of the Entomological Society of Washington, 11, 1 - 370." type="journal article" year="1983">Kanmiya 1983</bibRefCitation>
). These three genera share an extensively setulose body, large size, a broad rugose gena and a broad facial carina (less developed in
<taxonomicName id="A7AD4D65DE01385EFB6D080F9B04C511" box="[1155,1295,1870,1895]" class="Insecta" family="Chloropidae" genus="Medeventor" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE01385EFB6D080F9B04C511" box="[1155,1295,1870,1895]" italics="true" pageId="1" pageNumber="48">Medeventor</emphasis>
</taxonomicName>
). However,
<taxonomicName id="A7AD4D65DE01385EFF7908379F28C5F9" box="[151,291,1910,1935]" class="Insecta" family="Chloropidae" genus="Medeventor" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE01385EFF7908379F28C5F9" box="[151,291,1910,1935]" italics="true" pageId="1" pageNumber="48">Medeventor</emphasis>
</taxonomicName>
cannot be accommodated in either of those genera without significantly modifying their current limits, and the autapomorphic characters of the
<typeStatus id="BF168844DE01385EFD2B08DF9CFCC5CE" box="[709,759,1950,1976]" pageId="1" pageNumber="48">type</typeStatus>
species justify erection of a new genus.
</paragraph>
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The male postabdomen of
<taxonomicName id="A7AD4D65DE01385EFDEE08879C87C5A9" box="[512,652,1990,2015]" class="Insecta" family="Chloropidae" genus="Medeventor" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE01385EFDEE08879C87C5A9" box="[512,652,1990,2015]" italics="true" pageId="1" pageNumber="48">Medeventor</emphasis>
</taxonomicName>
is atypical in that ventral, symmetrical remnants of sternites 6 and 7 are present. In the subfamily
<taxonomicName id="A7AD4D65DE02385DFE110FD99CC7C2C4" box="[511,716,152,178]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="subFamily" subFamily="Siphonellopsinae">Siphonellopsinae</taxonomicName>
, sternites 6 and 7 are often retained, but are usually asymmetrical, as is the case with many other families of acalyptrate
<taxonomicName id="A7AD4D65DE02385DFCBE0F819A8DC2AC" authority="Andersson 1977" authorityName="Andersson" authorityYear="1977" box="[848,1158,192,218]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="order">
Diptera (
<bibRefCitation id="043C4B17DE02385DFC590F819A76C2AC" author="Andersson" box="[951,1149,192,218]" pageId="2" pageNumber="53" refString="Andersson, H. (1977) Taxonomic and phylogenetic studies on Chloropidae (Diptera) with special reference to Old World genera. Entomologica scandinavica Supplement, 8, 1 - 200." type="journal article" year="1977">Andersson 1977</bibRefCitation>
)
</taxonomicName>
. The reduction and loss of sternites 6 and 7, and the secondary symmetry of the postabdomen are apparently apomorphic for
<taxonomicName id="A7AD4D65DE02385DFAD70FA99EDBC35C" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="subFamily" subFamily="Oscinellinae">Oscinellinae</taxonomicName>
plus
<taxonomicName id="A7AD4D65DE02385DFEFE0E519FAAC35C" box="[272,417,272,298]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="subFamily" subFamily="Chloropinae">Chloropinae</taxonomicName>
(e.g.,
<bibRefCitation id="043C4B17DE02385DFE020E519CBAC35C" author="Andersson" box="[492,689,272,298]" pageId="2" pageNumber="53" refString="Andersson, H. (1977) Taxonomic and phylogenetic studies on Chloropidae (Diptera) with special reference to Old World genera. Entomologica scandinavica Supplement, 8, 1 - 200." type="journal article" year="1977">Andersson 1977</bibRefCitation>
). Although the postabdomen is symmetrical in
<taxonomicName id="A7AD4D65DE02385DFB300E519B62C35F" box="[1246,1385,272,297]" class="Insecta" family="Chloropidae" genus="Medeventor" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE02385DFB300E519B62C35F" box="[1246,1385,272,297]" italics="true" pageId="2" pageNumber="49">Medeventor</emphasis>
</taxonomicName>
, the distinct remnant of sternite 6 is unusual among
<taxonomicName id="A7AD4D65DE02385DFD2D0E799D52C324" box="[707,857,312,338]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="subFamily" subFamily="Oscinellinae">Oscinellinae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="56AD903E2E79046E7842FCFB10104712">Two new genera of oscinelline Chloropidae (Diptera) from Costa Rica</mods:title>
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<subSubSection id="28B7656DDE02385DFF7908219F1CC5EB" pageId="2" pageNumber="49" type="nomenclature">
<paragraph id="601236E6DE02385DFF7908219C5AC50C" blockId="2.[151,593,1888,1949]" box="[151,593,1888,1914]" pageId="2" pageNumber="49">
<heading id="3B5A818ADE02385DFF7908219C5AC50C" bold="true" box="[151,593,1888,1914]" fontSize="11" level="1" pageId="2" pageNumber="49" reason="1">
<emphasis id="52D9EAF4DE02385DFF7908219C5AC50C" bold="true" box="[151,593,1888,1914]" pageId="2" pageNumber="49">
<taxonomicName id="A7AD4D65DE02385DFF7908219C08C50C" ID-CoL="6R8C8" authority="Wheeler" authorityName="Wheeler" box="[151,515,1888,1914]" class="Insecta" family="Chloropidae" genus="Medeventor" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="nubosus" status="sp. nov.">
<emphasis id="52D9EAF4DE02385DFF7908219F99C50C" bold="true" box="[151,402,1888,1914]" italics="true" pageId="2" pageNumber="49">Medeventor nubosus</emphasis>
Wheeler
</taxonomicName>
,
<taxonomicNameLabel id="49EA578FDE02385DFDE108219C5AC50C" box="[527,593,1888,1914]" pageId="2" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="601236E6DE02385DFF7908C29F1CC5EB" blockId="2.[151,593,1888,1949]" box="[151,279,1923,1949]" pageId="2" pageNumber="49">
(
<figureCitation id="F8962A63DE02385DFF7108C29F05C5EB" box="[159,270,1923,1949]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="2" pageNumber="49">Figs. 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="28B7656DDE02385CFF7908929F21C4F3" lastPageId="3" lastPageNumber="50" pageId="2" pageNumber="49" type="description">
<paragraph id="601236E6DE02385CFF7908929B7BC30C" blockId="2.[151,1436,2003,2029]" lastBlockId="3.[151,1437,152,1669]" lastPageId="3" lastPageNumber="50" pageId="2" pageNumber="49">
Description: Total length
<quantity id="A7559B03DE02385DFE2408929C23C59B" box="[458,552,2003,2029]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="49" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5mm</quantity>
. Overall colour medium brown; head paler, with frontal triangle pruinose but distinct, not reaching anterior margin of frons; most cephalic setae reduced, outer vertical seta strongest, inner vertical, ocellars and postocellars only slightly differentiated from interfrontal setulae (
<figureCitation id="F8962A63DE03385CFB200F819B19C2AC" box="[1230,1298,192,218]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="3" pageNumber="50">Fig. 1</figureCitation>
); interfrontal setulae and fronto-orbital setulae dense, fine, pale; gena dark yellow, genal height 0.5–0.6 times eye height (
<figureCitation id="F8962A63DE03385CFF710E519EEFC35C" box="[159,228,272,298]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="3" pageNumber="50">Fig. 1</figureCitation>
), face with small, relatively broad carina between antennal bases, extending ventrally halfway to ventral margin of face (
<figureCitation id="F8962A63DE03385CFE6C0E799FC0C324" box="[386,459,312,338]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="3" pageNumber="50">Fig. 2</figureCitation>
); antenna with patch of short dorsal setae on pedicel, first flagellomere round, yellow, arista black, pubescent; palpus yellow, swollen; proboscis black, short but sclerotized and geniculate.
</paragraph>
<paragraph id="601236E6DE03385CFF280EC99A5AC034" blockId="3.[151,1437,152,1669]" pageId="3" pageNumber="50">
Scutum apparently shining (although
<typeStatus id="BF168844DE03385CFD640EC99CFBC3D4" box="[650,752,392,418]" pageId="3" pageNumber="50" type="holotype">holotype</typeStatus>
is covered with dull residue), evenly covered with long, pale setulae,
<typeStatus id="BF168844DE03385CFEDF0EF19F9CC3BC" box="[305,407,432,458]" pageId="3" pageNumber="50" type="holotype">holotype</typeStatus>
with 1 anterior, 1 posterior notopleural seta (
<typeStatus id="BF168844DE03385CFC5F0EF19A28C3BC" box="[945,1059,432,458]" pageId="3" pageNumber="50" type="paratype">paratypes</typeStatus>
with 1+2 notopleural setae, see Remarks); scutellum with apical and subapical setae only slightly longer and darker than surrounding setae. Legs long, fore femur inflated in
<typeStatus id="BF168844DE03385CFDCD0D419C87C06C" box="[547,652,512,538]" pageId="3" pageNumber="50" type="holotype">holotype</typeStatus>
, at least 3 times diameter of fore tibia (not as broad in
<typeStatus id="BF168844DE03385CFAF40D419B9BC06C" box="[1306,1424,512,538]" pageId="3" pageNumber="50" type="paratype">paratypes</typeStatus>
). Wing pale gray, veins dark, ratio of costal sectors C1: C2: C3: C4 – 1: 1: 0.5: 0.3.
</paragraph>
<paragraph id="601236E6DE03385CFF280D119C65C01C" blockId="3.[151,1437,152,1669]" box="[198,622,592,618]" pageId="3" pageNumber="50">Abdomen slightly paler than thorax.</paragraph>
<paragraph id="601236E6DE03385CFF280D399FEDC1F4" blockId="3.[151,1437,152,1669]" pageId="3" pageNumber="50">
Male genitalia (
<figureCitation id="F8962A63DE03385CFE6A0D399FF2C0E4" box="[388,505,632,658]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,269,1696,1720]" captionTargetBox="[200,1346,339,1695]" captionTargetId="figure@2.[198,1389,323,1770]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Medeventor nubosus. 1. head, lateral. 2. head, anterior. 3. wing. 4. male abdomen, posterior portion, ventral. 5. male genitalia, lateral. 6. male genitalia, posterior. 7. male genitalia, ventral. Abbreviations: st — sternite; ses — subepandrial sclerite. Scale bars = 1.0 mm (Fig. 3), 0.2 mm (Figs. 6 – 8)." httpUri="https://zenodo.org/record/175671/files/figure.png" pageId="3" pageNumber="50">Figs. 5–7</figureCitation>
): epandrium approximately as long as high in lateral view, broad in posterior view, with dense setae on posterior half; surstylus pointed, evenly tapering, recurved in lateral view, with dense patch of fine setae anteromedially near base, and scattered fine setae distally; hypandrium robust anteriorly, pregonites, postgonites and phallic guide all well-sclerotized and fused with hypandrium anteriorly; basiphallus long and well-sclerotized, distiphallus shorter than basiphallus, membranous; cerci broad, only slightly rounded ventrally in posterior view, with fine setae; subepandrial sclerite with setulose lobe between base of surstylus and cercus.
</paragraph>
<paragraph id="601236E6DE03385CFF280CD29DE7C603" blockId="3.[151,1437,152,1669]" pageId="3" pageNumber="50">
<typeStatus id="BF168844DE03385CFF280CD29F0BC1DB" box="[198,256,915,941]" pageId="3" pageNumber="50">Type</typeStatus>
material:
<typeStatus id="BF168844DE03385CFE900CD29FE0C1DB" box="[382,491,915,941]" pageId="3" pageNumber="50" type="holotype">Holotype</typeStatus>
ɗ:
<collectingCountry id="18BA7676DE03385CFDF30CD29CDAC1DB" box="[541,721,915,941]" name="Costa Rica" pageId="3" pageNumber="50">COSTA RICA</collectingCountry>
: Prov. Puntarenas, Monteverde, 1995, B.J. Sinclair (CNC).
<typeStatus id="BF168844DE03385CFF790CFA9F05C1A3" box="[151,270,955,981]" pageId="3" pageNumber="50" type="paratype">Paratypes</typeStatus>
: 1Ψ:
<collectingCountry id="18BA7676DE03385CFEA10CFA9C09C1A3" box="[335,514,955,981]" name="Costa Rica" pageId="3" pageNumber="50">COSTA RICA</collectingCountry>
: Prov. Guanacaste,
<collectingCountry id="18BA7676DE03385CFD000CFA9D4DC1A3" box="[750,838,955,981]" name="Liberia" pageId="3" pageNumber="50">Liberia</collectingCountry>
, Parque Nacional Santa Rosa, Sector Santa Rosa, Area Adm.
<quantity id="A7559B03DE03385CFED70CA29F74C18B" box="[313,383,995,1021]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="50" unit="m" value="300.0">300m</quantity>
,
<date id="14131026DE03385CFE760CA29FF0C18B" box="[408,507,995,1021]" pageId="3" pageNumber="50" value="1999-07">vii.1999</date>
, J. Sullivan and A.A. Pérez, Malaise, L N 313000 359800, #56027, INB0003115663 (INBio); 1? (sex unknown, abdomen damaged):
<collectingCountry id="18BA7676DE03385CFC420B4A9A54C653" box="[940,1119,1035,1061]" name="Costa Rica" pageId="3" pageNumber="50">COSTA RICA</collectingCountry>
: Prov. Guanacaste, Parque Nacional Rincón de la Vieja, Est. Las Pailas,
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,
<date id="14131026DE03385CFCC80B729DD9C63B" box="[806,978,1075,1101]" pageId="3" pageNumber="50" value="1993-11-10" valueMax="1993-11-21" valueMin="1993-11-10">10–21.xi.1993</date>
, M.A. Zumbado, D.G. García and G. Rodríguez, L N 306300_388600, #2559, INBIOCRI001827899 (INBio).
</paragraph>
<paragraph id="601236E6DE03385CFF280BC99FAEC6BC" blockId="3.[151,1437,152,1669]" pageId="3" pageNumber="50">
Etymology: The species name is from the Spanish
<emphasis id="52D9EAF4DE03385CFCF70BC99D65C6D7" box="[793,878,1160,1185]" italics="true" pageId="3" pageNumber="50">nuboso</emphasis>
(cloudy), referring to the cloud forests that surround the
<typeStatus id="BF168844DE03385CFEE30BF19F4BC6BC" box="[269,320,1200,1226]" pageId="3" pageNumber="50">type</typeStatus>
locality.
</paragraph>
<paragraph id="601236E6DE03385CFF280B9A9AD4C7CB" blockId="3.[151,1437,152,1669]" pageId="3" pageNumber="50">
Remarks: Differences in the texture of the frontal triangle and scutum; number of notopleural setae and diameter of the fore femur between the
<typeStatus id="BF168844DE03385CFD990A429CD5C76B" box="[631,734,1283,1309]" pageId="3" pageNumber="50" type="holotype">holotype</typeStatus>
and
<typeStatus id="BF168844DE03385CFCF50A429D80C76B" box="[795,907,1283,1309]" pageId="3" pageNumber="50" type="paratype">paratypes</typeStatus>
suggest that multiple species may be represented or that sexual dimorphism may exist in the species. Unfortunately, all
<typeStatus id="BF168844DE03385CFBF20A6A9A44C733" box="[1052,1103,1323,1349]" pageId="3" pageNumber="50">type</typeStatus>
specimens are dirty or damaged to some degree so it is difficult to speculate on which of the two possible explanations is more likely. Additional specimens in good condition will be required to resolve the question. For the present, the species description is based on the
<typeStatus id="BF168844DE03385CFE3B0AE29C30C7CB" box="[469,571,1443,1469]" pageId="3" pageNumber="50" type="holotype">holotype</typeStatus>
and differences in the
<typeStatus id="BF168844DE03385CFCA90AE29DBCC7CB" box="[839,951,1443,1469]" pageId="3" pageNumber="50" type="paratype">paratypes</typeStatus>
are noted where relevant
</paragraph>
<paragraph id="601236E6DE03385CFF280A8A9F21C4F3" blockId="3.[151,1437,152,1669]" pageId="3" pageNumber="50">
Although known from only three specimens, this species is widespread in a range of habitats in
<collectingCountry id="18BA7676DE03385CFAB70A8A9EC6C47B" name="Costa Rica" pageId="3" pageNumber="50">Costa Rica</collectingCountry>
from high elevation cloud forest at Monteverde to lowland dry forest in Guanacaste. Because its size and appearance are atypical for
<taxonomicName id="A7AD4D65DE03385CFE09095A9C70C443" box="[487,635,1563,1589]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
, additional specimens may be present but unidentified in museum collections.
<taxonomicName id="A7AD4D65DE03385CFECB09059C13C42B" box="[293,536,1604,1629]" class="Insecta" family="Chloropidae" genus="Medeventor" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="nubosus">
<emphasis id="52D9EAF4DE03385CFECB09059C13C42B" box="[293,536,1604,1629]" italics="true" pageId="3" pageNumber="50">Medeventor nubosus</emphasis>
</taxonomicName>
is the largest known chloropid in
<collectingCountry id="18BA7676DE03385CFC4409029A25C42B" box="[938,1070,1603,1629]" name="Costa Rica" pageId="3" pageNumber="50">Costa Rica</collectingCountry>
; the species described below is the smallest.
</paragraph>
</subSubSection>
</treatment>
</document>

136
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<mods:title id="5D99136FFD67FD996652C9DD3A15B169">Two new genera of oscinelline Chloropidae (Diptera) from Costa Rica</mods:title>
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<subSubSection id="28B7656DDE03385CFF7909929FEBC49B" box="[151,480,1747,1773]" pageId="3" pageNumber="50" type="nomenclature">
<paragraph id="601236E6DE03385CFF7909929FEBC49B" blockId="3.[151,480,1747,1773]" box="[151,480,1747,1773]" pageId="3" pageNumber="50">
<heading id="3B5A818ADE03385CFF7909929FEBC49B" bold="true" box="[151,480,1747,1773]" fontSize="11" level="1" pageId="3" pageNumber="50" reason="1">
<emphasis id="52D9EAF4DE03385CFF7909929FEBC49B" bold="true" box="[151,480,1747,1773]" pageId="3" pageNumber="50">
<taxonomicName id="A7AD4D65DE03385CFF7909929F8AC49B" ID-CoL="63GMX" authority="Wheeler" authorityName="Wheeler" box="[151,385,1747,1773]" class="Insecta" family="Chloropidae" genus="Oscinicita" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE03385CFF7909929F1AC49B" bold="true" box="[151,273,1747,1773]" italics="true" pageId="3" pageNumber="50">Oscinicita</emphasis>
Wheeler
</taxonomicName>
, gen. n.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="28B7656DDE03385BFF7908579ECCC489" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50" type="materials_examined">
<paragraph id="601236E6DE03385CFF7908579CF0C55D" blockId="3.[151,763,1814,1835]" box="[151,763,1814,1835]" pageId="3" pageNumber="50">
<typeStatus id="BF168844DE03385CFF7908579ECCC55D" box="[151,199,1814,1835]" pageId="3" pageNumber="50">Type</typeStatus>
species:
<taxonomicName id="A7AD4D65DE03385CFEF008579FD8C55D" box="[286,467,1814,1835]" class="Insecta" family="Chloropidae" genus="Oscinicita" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="hansoni" status="sp. nov.">
<emphasis id="52D9EAF4DE03385CFEF008579FD8C55D" box="[286,467,1814,1835]" italics="true" pageId="3" pageNumber="50">Oscinicita hansoni</emphasis>
</taxonomicName>
<emphasis id="52D9EAF4DE03385CFE3708579C04C55D" bold="true" box="[473,527,1814,1835]" pageId="3" pageNumber="50">
<taxonomicNameLabel id="49EA578FDE03385CFE3708579C04C55D" box="[473,527,1814,1835]" pageId="3" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(by present designation)
</paragraph>
<paragraph id="601236E6DE03385CFF7908189C81C5B0" blockId="3.[151,1436,1881,2034]" pageId="3" pageNumber="50">
Diagnosis: Minute (
<quantity id="A7559B03DE03385CFE6608189C17C505" box="[392,540,1881,1907]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="3" pageNumber="50" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
)
<taxonomicName id="A7AD4D65DE03385CFDDE08189CC9C505" box="[560,706,1881,1907]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Oscinellinae">Oscinellinae</taxonomicName>
with antennae in deep foveae, incurved vibrissal angle and gena, deep postgena, strongly convex scutum, and reduced wing venation (dm-cu absent, CuA1 straight and incomplete, other longitudinal veins short)
</paragraph>
<paragraph id="601236E6DE03385BFF2808999A50C01C" blockId="3.[151,1436,1881,2034]" lastBlockId="4.[151,1437,152,1791]" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50">
Description:
<taxonomicName id="A7AD4D65DE03385CFEB008999FF9C584" box="[350,498,2008,2034]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
,
<taxonomicName id="A7AD4D65DE03385CFE1008999C9EC584" box="[510,661,2008,2034]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Oscinellinae">Oscinellinae</taxonomicName>
. Head higher than long in lateral view, with frons strongly sloping toward lunule (
<figureCitation id="F8962A63DE04385BFEA70FD99F84C2C4" box="[329,399,152,178]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 – 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 8</figureCitation>
), wider than long in dorsal view, with occipital margin concave; frontal triangle distinct, bare except for row of setulae along margin, ocellar tubercle large relative to size of frons; frons with dense interfrontal setulae, fronto-orbital setae in long row barely distinguished from interfrontal setulae, vertical, ocellar and postocellar setae short and weak, ocellar setae reclinate, convergent, postocellar setae convergent; eye rounded, with short ommatrichia; gena incurved ventrally, sparsely setulose, with bare shining strip anteroventral to eye, continuous with shining parafacial; postgena deep and swollen posteroventral to eye, densely tomentose (
<figureCitation id="F8962A63DE04385BFE910EC99FCEC3D4" box="[383,453,392,418]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 – 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 8</figureCitation>
); vibrissal angle broadly rounded, strongly incurved toward midline of face (
<figureCitation id="F8962A63DE04385BFAA80EC99B86C3D4" box="[1350,1421,392,418]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 – 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 9</figureCitation>
), barely visible in lateral view, vibrissa short but distinct; face short, scarcely extending beyond ventral margin of first flagellomere, oral margin high and U-shaped (
<figureCitation id="F8962A63DE04385BFCE00E999D5EC384" box="[782,853,472,498]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 – 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 9</figureCitation>
), face narrow because of incurved vibrissal angle and parafacial, most of face forming deep antennal foveae, facial carina small and narrow in ventral part of antennal foveae; antenna small, only slightly projecting in lateral view, scape and pedicel small, first flagellomere small, rounded, arista short, pubescent; proboscis short; palpus short.
</paragraph>
<paragraph id="601236E6DE04385BFF280D399D0AC1A1" blockId="4.[151,1437,152,1791]" pageId="4" pageNumber="51">
Scutum strongly convex, postpronotum small; scutal setulae evenly distributed, 1–2 postpronotal, 1 anterior and 2 posterior notopleural, 1 postsutural supra-alar, 1 postsutural intra-alar, 1 dorsocentral seta apparently present (setae missing or damaged to some degree in all
<typeStatus id="BF168844DE04385BFC400D899DFAC094" box="[942,1009,712,738]" pageId="4" pageNumber="51">types</typeStatus>
); scutellum short, convex, densely tomentose, with short apical, subapical and dorsal setae; subscutellum swollen; thoracic pleurites bare. Wing hyaline, alula small, anal angle well-developed, R2+3 and R4+5 upcurved distally toward C, second costal sector longer than third; M1+2 ending near apex of wing, CuA1 straight, without trace of kink near midpoint, ending well before margin of wing, crossvein r-m present, crossvein dm-cu absent (
<figureCitation id="F8962A63DE04385BFBF30C2C9A7DC1F1" box="[1053,1142,877,903]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 – 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 10</figureCitation>
). Legs without outstanding setae or spurs; femoral organ not evident in
<typeStatus id="BF168844DE04385BFD0A0CD49D1CC1D9" box="[740,791,917,943]" pageId="4" pageNumber="51">type</typeStatus>
specimens; tibial organ may be present as indistinct, small oval patch (not obvious in all
<typeStatus id="BF168844DE04385BFDD20CFC9C64C1A1" box="[572,623,957,983]" pageId="4" pageNumber="51">type</typeStatus>
specimens).
</paragraph>
<paragraph id="601236E6DE04385BFF280CA49FE6C639" blockId="4.[151,1437,152,1791]" pageId="4" pageNumber="51">
Abdominal tergites with sparse setae laterally and posteriorly, evenly covered with tiny setulae; sternites relatively broad, with sparse setae, abdominal membrane densely spinulose; male spiracles
<quantity id="A7559B03DE04385BFB230B4C9B12C651" box="[1229,1305,1037,1063]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="4" pageNumber="51" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">3–5 in</quantity>
membrane near lateral margin of tergite.
</paragraph>
<paragraph id="601236E6DE04385BFF280B1F9B3DC71E" blockId="4.[151,1437,152,1791]" pageId="4" pageNumber="51">
Male postabdomen: dorsal pregenital sclerite narrow; spiracles 6 and
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membrane ventral to lateral margin of dorsal sclerite; epandrium simple, convex, with sparse setae; surstylus simple, broadly rounded, freely articulated with epandrium; hypandrium open posteriorly; pregonite fused with postgonite, weakly sclerotized; postgonite with ventral seta and campaniform sensillae; basiphallus elongate, sclerotized, distiphallus short, membranous (
<figureCitation id="F8962A63DE04385BFE100BBF9C9AC76E" box="[510,657,1278,1304]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 – 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Figs. 11–12</figureCitation>
); phallapodeme simple, phallic guide sclerotized, rounded; cerci small, with triangular ventral points broadly separated across midline, setulose (
<figureCitation id="F8962A63DE04385BFBD30A679A9FC736" box="[1085,1172,1318,1344]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 – 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 12</figureCitation>
); subepandrial sclerite well-developed, with sclerotized projection extending dorsally above base of surstylus (
<figureCitation id="F8962A63DE04385BFB750A0F9B22C71E" box="[1179,1321,1358,1384]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 – 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Figs. 11–12</figureCitation>
).
</paragraph>
<paragraph id="601236E6DE04385BFF280A379F92C7E6" blockId="4.[151,1437,152,1791]" box="[198,409,1398,1424]" pageId="4" pageNumber="51">Female unknown.</paragraph>
<paragraph id="601236E6DE04385BFF280AE39F92C47A" blockId="4.[151,1437,152,1791]" pageId="4" pageNumber="51">
Etymology: The generic name is derived from
<emphasis id="52D9EAF4DE04385BFD070AE39D2CC7CD" box="[745,807,1442,1467]" italics="true" pageId="4" pageNumber="51">oscin</emphasis>
- a common generic root in the
<taxonomicName id="A7AD4D65DE04385BFB780AE39B2CC7CA" box="[1174,1319,1442,1468]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
and
<emphasis id="52D9EAF4DE04385BFAB10AE39B9EC7CD" box="[1375,1429,1442,1467]" italics="true" pageId="4" pageNumber="51">-cita</emphasis>
, a Spanish term widely used as a diminutive in
<collectingCountry id="18BA7676DE04385BFD5B0A8B9D3CC792" box="[693,823,1482,1508]" name="Costa Rica" pageId="4" pageNumber="51">Costa Rica</collectingCountry>
, referring to the minute size of these flies. The name is treated as feminine.
</paragraph>
<paragraph id="601236E6DE04385BFF28095C9ECCC489" blockId="4.[151,1437,152,1791]" pageId="4" pageNumber="51">
Remarks: Although the autapomorphic character states of
<taxonomicName id="A7AD4D65DE04385BFC9E095F9DE2C441" box="[880,1001,1566,1591]" class="Insecta" family="Chloropidae" genus="Oscinicita" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE04385BFC9E095F9DE2C441" box="[880,1001,1566,1591]" italics="true" pageId="4" pageNumber="51">Oscinicita</emphasis>
</taxonomicName>
make these flies easily recognizable, they do not resolve the phylogenetic relationships of the genus. The head characters (genal structure, incurved vibrissal angle, short face with distinct antennal foveae) are not informative about relationships. The reduction in wing venation, especially loss of dm-cu, is a homoplasious state, related to small size, in
<taxonomicName id="A7AD4D65DE04385BFB0C09D49B7DC4D9" box="[1250,1398,1685,1711]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
, as well as other acalyptrates, although it occurs more frequently in the subfamily
<taxonomicName id="A7AD4D65DE04385BFBD909FC9AC3C4A1" box="[1079,1224,1725,1751]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="subFamily" subFamily="Chloropinae">Chloropinae</taxonomicName>
than in
<taxonomicName id="A7AD4D65DE04385BFAC709FC9EC8C489" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="subFamily" subFamily="Oscinellinae">Oscinellinae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="FAD1F043B4EF50C8332C136FA44BD4CA" ID-DOI="10.5281/zenodo.175670" ID-GBIF-Dataset="7f25f9fd-d335-42e5-b89a-6ea67e8ed713" ID-ISSN="1175-5326" ID-Zenodo-Dep="175670" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459816256441" checkinUser="plazi" docAuthor="Wheeler, Terry A." docDate="2007" docId="E80487F0DE043859FF79081C9D8BC113" docLanguage="en" docName="zt01413p053.pdf" docOrigin="Zootaxa 1413" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Oscinicita hansoni Wheeler, sp. n." docType="treatment" docVersion="9" lastPageNumber="53" masterDocId="143DFF88DE00385FFFEE0F419E0BC276" masterDocTitle="Two new genera of oscinelline Chloropidae (Diptera) from Costa Rica" masterLastPageNumber="53" masterPageNumber="47" pageNumber="51" updateTime="1698221377221" updateUser="plazi">
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<mods:title id="D6A6CBF12C7671606737D29989BBAA14">Two new genera of oscinelline Chloropidae (Diptera) from Costa Rica</mods:title>
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<heading id="3B5A818ADE04385BFF79081C9C32C501" bold="true" box="[151,569,1885,1911]" fontSize="11" level="1" pageId="4" pageNumber="51" reason="1">
<emphasis id="52D9EAF4DE04385BFF79081C9C32C501" bold="true" box="[151,569,1885,1911]" pageId="4" pageNumber="51">
<taxonomicName id="A7AD4D65DE04385BFF79081C9FE1C501" ID-CoL="74YWP" authority="Wheeler" authorityName="Wheeler" box="[151,490,1885,1911]" class="Insecta" family="Chloropidae" genus="Oscinicita" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="hansoni" status="sp. nov.">
<emphasis id="52D9EAF4DE04385BFF79081C9F71C501" bold="true" box="[151,378,1885,1911]" italics="true" pageId="4" pageNumber="51">Oscinicita hansoni</emphasis>
Wheeler
</taxonomicName>
,
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<paragraph id="601236E6DE04385BFF7908C19F2DC5EC" blockId="4.[151,569,1885,1946]" box="[151,294,1920,1946]" pageId="4" pageNumber="51">
(
<figureCitation id="F8962A63DE04385BFF7108C19F15C5EC" box="[159,286,1920,1946]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 – 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Figs. 8–13</figureCitation>
)
</paragraph>
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<subSubSection id="28B7656DDE043859FF7908919D8BC113" lastPageId="6" lastPageNumber="53" pageId="4" pageNumber="51" type="description">
<paragraph id="601236E6DE04385AFF7908919B23C324" blockId="4.[151,1436,2000,2026]" lastBlockId="5.[151,1437,152,498]" lastPageId="5" lastPageNumber="52" pageId="4" pageNumber="51">
Description: Total length
<quantity id="A7559B03DE04385BFE2208919C5DC59C" box="[460,598,2000,2026]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="4" pageNumber="51" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2mm</quantity>
. Overall colour medium brown; frontal triangle shining, not reaching anterior margin of frons; frons microscopically striate, appearing dull; most setae and setulae on head pale, weak; genal height 0.2 times eye height directly below ventral midpoint of eye, much broader posterior to this (
<figureCitation id="F8962A63DE05385AFF710FA99EE3C374" box="[159,232,232,258]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 – 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="5" pageNumber="52">Fig. 8</figureCitation>
); postgena slightly paler than gena and parafacial; face shining, especially in antennal foveae, darker than rest of head, carina narrow, most pronounced ventrally, spinulose membrane below ventral margin of face continuous with oral cavity; antenna yellow-brown, arista brown; palpus black; proboscis pale.
</paragraph>
<paragraph id="601236E6DE053859FF280E219FF0C2C4" blockId="5.[151,1437,152,498]" lastBlockId="6.[151,1437,152,869]" lastPageId="6" lastPageNumber="53" pageId="5" pageNumber="52">Scutum shining, evenly covered with sparse, pale setulae; notopleural setae stronger and darker than other thoracic setae; scutellum with dense microtrichia giving it a dull appearance, sparse, short, dark apical, subapical and dorsal setae. Legs short, coxae and femora medium brown, tibiae and tarsi yellow. Wing hyaline but with dense microtrichia, veins pale, ratio of costal sectors C1: C2: C3: C4 – 0.9: 1: 0.6: 0.5. Halter large, Abdomen medium brown.</paragraph>
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<paragraph id="601236E6DE05385AFF79082B9A96C5BD" blockId="5.[151,1436,1898,1995]" pageId="5" pageNumber="52">
<emphasis id="52D9EAF4DE05385AFF79082B9F45C5F4" bold="true" box="[151,334,1898,1922]" pageId="5" pageNumber="52">FIGURES 8–13.</emphasis>
<taxonomicName id="A7AD4D65DE05385AFEBA082B9C17C5F7" box="[340,540,1898,1921]" class="Insecta" family="Chloropidae" genus="Oscinicita" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="hansoni">
<emphasis id="52D9EAF4DE05385AFEBA082B9C17C5F7" box="[340,540,1898,1921]" italics="true" pageId="5" pageNumber="52">Oscinicita hansoni</emphasis>
</taxonomicName>
. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11–13).
</paragraph>
</caption>
<paragraph id="601236E6DE063859FF280F819F2DC384" blockId="6.[151,1437,152,869]" pageId="6" pageNumber="53">
Male genitalia (
<figureCitation id="F8962A63DE063859FE680F819C12C2AC" box="[390,537,192,218]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 – 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="6" pageNumber="53">Figs. 11–13</figureCitation>
): epandrium approximately as long as high, broad in posterior view, with sparse setae on posterior half; surstylus broadly rounded, projecting ventrally in lateral view, ventral margin dentate, with sparse, fine setae distally; hypandrium relatively narrow; pregonite weakly sclerotized, postgonite clavate, sclerotized marginally, with single ventral seta and 2–3 campaniform sensillae; phallic guide sclerotized, rounded, separated from hypandrium anteriorly; basiphallus long and well-sclerotized, distiphallus shorter than basiphallus, membranous; cerci small ventral projections separated by broad, U-shaped ventral margin, with fine setae; subepandrial sclerite simple, well-sclerotized, strongly projecting dorsally in lateral view.
</paragraph>
<paragraph id="601236E6DE063859FF280D419F92C06C" blockId="6.[151,1437,152,869]" box="[198,409,512,538]" pageId="6" pageNumber="53">Female unknown.</paragraph>
<paragraph id="601236E6DE063859FF280D6A9DE3C01B" blockId="6.[151,1437,152,869]" pageId="6" pageNumber="53">
<typeStatus id="BF168844DE063859FF280D6A9F0BC033" box="[198,256,555,581]" pageId="6" pageNumber="53">Type</typeStatus>
material:
<typeStatus id="BF168844DE063859FE940D6A9FECC033" box="[378,487,555,581]" pageId="6" pageNumber="53" type="holotype">Holotype</typeStatus>
ɗ:
<collectingCountry id="18BA7676DE063859FDFB0D6A9CCCC033" box="[533,711,555,581]" name="Costa Rica" pageId="6" pageNumber="53">COSTA RICA</collectingCountry>
: Alajuela:
<collectingCountry id="18BA7676DE063859FCA90D6A9DCAC033" box="[839,961,555,581]" name="Saint Pierre and Miquelon" pageId="6" pageNumber="53">San Pedro</collectingCountry>
de la Tigra, cacao,
<quantity id="A7559B03DE063859FB400D6A9AFFC033" box="[1198,1268,555,581]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="53" unit="m" value="200.0">200m</quantity>
, Malaise trap,
<date id="14131026DE063859FF790D129EEFC01B" box="[151,228,595,621]" pageId="6" pageNumber="53" value="1990-01">i.1990</date>
, P. Hanson (LEM).
<typeStatus id="BF168844DE063859FE3D0D129C41C01B" box="[467,586,595,621]" pageId="6" pageNumber="53" type="paratype">Paratypes</typeStatus>
: 2ɗ, same data as
<typeStatus id="BF168844DE063859FCCF0D129D8CC01B" box="[801,903,595,621]" pageId="6" pageNumber="53" type="holotype">holotype</typeStatus>
(LEM).
</paragraph>
<paragraph id="601236E6DE063859FF280D3E9A45C0B7" blockId="6.[151,1437,152,869]" pageId="6" pageNumber="53">
Etymology: The species name is a genitive patronym in honour of Dr. Paul Hanson who collected the
<typeStatus id="BF168844DE063859FA870D3E9B90C0EF" box="[1385,1435,639,665]" pageId="6" pageNumber="53">type</typeStatus>
series of this species, along with vast numbers of other Costa Rican
<taxonomicName id="A7AD4D65DE063859FC5B0DE69A41C0B7" box="[949,1098,679,705]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="53" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
.
</paragraph>
<paragraph id="601236E6DE063859FF280D929D8BC113" blockId="6.[151,1437,152,869]" pageId="6" pageNumber="53">
Remarks: The
<typeStatus id="BF168844DE063859FE9D0D929FAEC09B" box="[371,421,723,749]" pageId="6" pageNumber="53">type</typeStatus>
locality is a cacao plantation, which suggests that this species may be associated with, or at least tolerant of, disturbed habitats. The fact that more specimens have not been encountered despite extensive collecting in other disturbed habitats throughout
<collectingCountry id="18BA7676DE063859FCA30C629DC6C14B" box="[845,973,803,829]" name="Costa Rica" pageId="6" pageNumber="53">Costa Rica</collectingCountry>
also raises the possibility that this species may be associated with cacao. Further research is required.
</paragraph>
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<taxonomicName LSID="E8056006-30A0-5ABC-B8EE-49E72475A4D8" authority="Chiu, 1954" authorityName="Chiu" authorityYear="1954" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus multidens" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="multidens" status="stat. rev.">Enicospilus multidens Chiu, 1954</taxonomicName>
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<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Enicospilus multidens Chiu, 1954, stat. rev. ♂ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure29" httpUri="https://binary.pensoft.net/fig/473795" pageId="0" pageNumber="1">Figure 29</figureCitation>
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<taxonomicName authorityName="Chiu" authorityYear="1954" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus multidens" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="multidens">Enicospilus multidens</taxonomicName>
Chiu, 1954: 75; HT ♀ from Japan, TARI, examined;
<emphasis bold="true" pageId="0" pageNumber="1">stat. rev.</emphasis>
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</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">Total of 30 specimens (14♀♀16♂♂): Japan (14♀♀16♂♂).</paragraph>
<paragraph pageId="0" pageNumber="1">
Type series: HT ♀ of
<taxonomicName authorityName="Chiu" authorityYear="1954" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus multidens" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="multidens">
<emphasis italics="true" pageId="0" pageNumber="1">Enicospilus multidens</emphasis>
</taxonomicName>
Chiu, 1954, Minoh,
<normalizedToken originalValue="Ōsaka">Ōsaka</normalizedToken>
, Kinki, JAPAN, 20.VII.1918, N. Tosawa leg. (TARI).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Eastern Palaearctic region (
<bibRefCitation author="Chiu, SC" journalOrPublisher="Bulletin of the Taiwan Agricultural Research Institute" pageId="0" pageNumber="1" pagination="1 - 79" refId="B18" refString="Chiu, SC, 1954. On some Enicospilus -species from the Orient (Hymenoptera: Ichneumonidae). Bulletin of the Taiwan Agricultural Research Institute 13: 1 - 79" title="On some Enicospilus - species from the Orient (Hymenoptera: Ichneumonidae)." volume="13" year="1954">Chiu 1954</bibRefCitation>
; present study); new to the Oriental region; this is a predominantly Eastern Palaearctic species.
</paragraph>
<paragraph pageId="0" pageNumber="1">
JAPAN: [
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]*; [
<normalizedToken originalValue="Tôhoku">Tohoku</normalizedToken>
] Aomori*; [Hokuriku] Niigata*; [
<normalizedToken originalValue="Kantô-Kôshin">Kanto-Koshin</normalizedToken>
] Gunma*, Nagano*, Yamanashi*, and Saitama*; [
<normalizedToken originalValue="Tôkai">Tokai</normalizedToken>
] Shizuoka* and Mie*; [Kinki]
<normalizedToken originalValue="Ōsaka">Ōsaka</normalizedToken>
(
<bibRefCitation author="Chiu, SC" journalOrPublisher="Bulletin of the Taiwan Agricultural Research Institute" pageId="0" pageNumber="1" pagination="1 - 79" refId="B18" refString="Chiu, SC, 1954. On some Enicospilus -species from the Orient (Hymenoptera: Ichneumonidae). Bulletin of the Taiwan Agricultural Research Institute 13: 1 - 79" title="On some Enicospilus - species from the Orient (Hymenoptera: Ichneumonidae)." volume="13" year="1954">Chiu 1954</bibRefCitation>
; present study) and
<normalizedToken originalValue="Hyôgo">Hyogo</normalizedToken>
*; [
<normalizedToken originalValue="Chûgoku">Chugoku</normalizedToken>
] Tottori (
<bibRefCitation author="Chiu, SC" journalOrPublisher="Bulletin of the Taiwan Agricultural Research Institute" pageId="0" pageNumber="1" pagination="1 - 79" refId="B18" refString="Chiu, SC, 1954. On some Enicospilus -species from the Orient (Hymenoptera: Ichneumonidae). Bulletin of the Taiwan Agricultural Research Institute 13: 1 - 79" title="On some Enicospilus - species from the Orient (Hymenoptera: Ichneumonidae)." volume="13" year="1954">Chiu 1954</bibRefCitation>
), Shimane* and Hiroshima*; [Shikoku] Ehime* and
<normalizedToken originalValue="Kôchi">Kochi</normalizedToken>
*; [
<normalizedToken originalValue="Kyûshû">Kyushu</normalizedToken>
] Kumamoto*; [
<normalizedToken originalValue="Ryûkyûs">Ryukyus</normalizedToken>
] Kagoshima*. *New records.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="bionomics">
<paragraph pageId="0" pageNumber="1">Bionomics.</paragraph>
<paragraph pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
In some ichneumonid collections, this species has been confused with
<taxonomicName lsidName="E. shikokuensis" pageId="0" pageNumber="1" rank="species" species="shikokuensis">
<emphasis italics="true" pageId="0" pageNumber="1">E. shikokuensis</emphasis>
</taxonomicName>
. However,
<taxonomicName lsidName="E. multidens" pageId="0" pageNumber="1" rank="species" species="multidens">
<emphasis italics="true" pageId="0" pageNumber="1">E. multidens</emphasis>
</taxonomicName>
stat. rev. can be distinguished from
<taxonomicName lsidName="E. shikokuensis" pageId="0" pageNumber="1" rank="species" species="shikokuensis">
<emphasis italics="true" pageId="0" pageNumber="1">E. shikokuensis</emphasis>
</taxonomicName>
by shallow concavity of proximal outer mandibular surface (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Enicospilus multidens Chiu, 1954, stat. rev. ♂ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure29" httpUri="https://binary.pensoft.net/fig/473795" pageId="0" pageNumber="1">29B, D</figureCitation>
) (proximal outer mandibular surface with very wide subtriangular concavity in
<taxonomicName lsidName="E. shikokuensis" pageId="0" pageNumber="1" rank="species" species="shikokuensis">
<emphasis italics="true" pageId="0" pageNumber="1">E. shikokuensis</emphasis>
</taxonomicName>
, as in Fig.
<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Enicospilus shikokuensis (Uchida, 1928) ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure44" httpUri="https://binary.pensoft.net/fig/473810" pageId="0" pageNumber="1">44B, D</figureCitation>
); separated proximal and distal sclerites (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Enicospilus multidens Chiu, 1954, stat. rev. ♂ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure29" httpUri="https://binary.pensoft.net/fig/473795" pageId="0" pageNumber="1">29F</figureCitation>
) (proximal and distal sclerites usually confluent, as in Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Intraspecific variation of the fore wing sclerite development in E. shikokuensis (Uchida, 1928) A the proximal and distal sclerites separated and the central sclerite weak (SEN 42) B the proximal and distal sclerites confluent and the central sclerite strong (SEN 41)." figureDoi="10.3897/zookeys.990.55542.figure7" httpUri="https://binary.pensoft.net/fig/473773" pageId="0" pageNumber="1">7B</figureCitation>
,
<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Enicospilus shikokuensis (Uchida, 1928) ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure44" httpUri="https://binary.pensoft.net/fig/473810" pageId="0" pageNumber="1">44F</figureCitation>
, or rarely separated, as in Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Intraspecific variation of the fore wing sclerite development in E. shikokuensis (Uchida, 1928) A the proximal and distal sclerites separated and the central sclerite weak (SEN 42) B the proximal and distal sclerites confluent and the central sclerite strong (SEN 41)." figureDoi="10.3897/zookeys.990.55542.figure7" httpUri="https://binary.pensoft.net/fig/473773" pageId="0" pageNumber="1">7A</figureCitation>
, in
<taxonomicName lsidName="E. shikokuensis" pageId="0" pageNumber="1" rank="species" species="shikokuensis">
<emphasis italics="true" pageId="0" pageNumber="1">E. shikokuensis</emphasis>
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); stouter, shorter, and evenly narrowed mandible (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Enicospilus multidens Chiu, 1954, stat. rev. ♂ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure29" httpUri="https://binary.pensoft.net/fig/473795" pageId="0" pageNumber="1">29B, D</figureCitation>
) (mandible much longer, slender, proximally strongly narrowed, and apical 0.7 parallel-sided in
<taxonomicName lsidName="E. shikokuensis" pageId="0" pageNumber="1" rank="species" species="shikokuensis">
<emphasis italics="true" pageId="0" pageNumber="1">E. shikokuensis</emphasis>
</taxonomicName>
, as in Fig.
<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Enicospilus shikokuensis (Uchida, 1928) ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure44" httpUri="https://binary.pensoft.net/fig/473810" pageId="0" pageNumber="1">44B, D</figureCitation>
); narrower lower face (Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Enicospilus multidens Chiu, 1954, stat. rev. ♂ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure29" httpUri="https://binary.pensoft.net/fig/473795" pageId="0" pageNumber="1">29B</figureCitation>
) (lower face wider in
<taxonomicName lsidName="E. shikokuensis" pageId="0" pageNumber="1" rank="species" species="shikokuensis">
<emphasis italics="true" pageId="0" pageNumber="1">E. shikokuensis</emphasis>
</taxonomicName>
, as in Fig.
<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Enicospilus shikokuensis (Uchida, 1928) ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure44" httpUri="https://binary.pensoft.net/fig/473810" pageId="0" pageNumber="1">44B</figureCitation>
), etc.
</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is also similar to
<taxonomicName lsidName="E. combustus" pageId="0" pageNumber="1" rank="species" species="combustus">
<emphasis italics="true" pageId="0" pageNumber="1">E. combustus</emphasis>
</taxonomicName>
, and
<bibRefCitation author="Uchida, T" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido University" pageId="0" pageNumber="1" pagination="95 - 133" refId="B183" refString="Uchida, T, 1955. Die von Dr. K. Tsuneki in Korea gesammelten Ichneumoniden. Journal of the Faculty of Agriculture, Hokkaido University 50: 95 - 133" title="Die von Dr. K. Tsuneki in Korea gesammelten Ichneumoniden." volume="50" year="1955">Uchida (1955)</bibRefCitation>
synonymised
<taxonomicName lsidName="E. multidens" pageId="0" pageNumber="1" rank="species" species="multidens">
<emphasis italics="true" pageId="0" pageNumber="1">E. multidens</emphasis>
</taxonomicName>
stat. rev. under
<taxonomicName lsidName="E. combustus" pageId="0" pageNumber="1" rank="species" species="combustus">
<emphasis italics="true" pageId="0" pageNumber="1">E. combustus</emphasis>
</taxonomicName>
. However, they are morphologically easily distinguished by body colour pattern (i.e., body entirely testaceous in
<taxonomicName lsidName="E. multidens" pageId="0" pageNumber="1" rank="species" species="multidens">
<emphasis italics="true" pageId="0" pageNumber="1">E. multidens</emphasis>
</taxonomicName>
stat. rev., as in Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Enicospilus multidens Chiu, 1954, stat. rev. ♂ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure29" httpUri="https://binary.pensoft.net/fig/473795" pageId="0" pageNumber="1">29A</figureCitation>
, but body with black marks in
<taxonomicName lsidName="E. combustus" pageId="0" pageNumber="1" rank="species" species="combustus">
<emphasis italics="true" pageId="0" pageNumber="1">E. combustus</emphasis>
</taxonomicName>
, as in Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Enicospilus combustus (Gravenhorst, 1829) ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure13" httpUri="https://binary.pensoft.net/fig/473779" pageId="0" pageNumber="1">13A</figureCitation>
), and DNA barcodes also separate them. Hence, we revise the status of
<taxonomicName lsidName="E. multidens" pageId="0" pageNumber="1" rank="species" species="multidens">
<emphasis italics="true" pageId="0" pageNumber="1">E. multidens</emphasis>
</taxonomicName>
stat. rev. as a valid species.
</paragraph>
<caption doi="10.3897/zookeys.990.55542.figure29" httpUri="https://binary.pensoft.net/fig/473795" pageId="0" pageNumber="1" start="Figure 29" startId="F29">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 29.</emphasis>
<taxonomicName authorityName="Chiu" authorityYear="1954" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus multidens" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="multidens">
<emphasis italics="true" pageId="0" pageNumber="1">Enicospilus multidens</emphasis>
</taxonomicName>
Chiu, 1954, stat. rev. ♂ from Japan
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
head, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
head, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
head, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
mesosoma, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
central part of fore wing.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1013</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>665</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1013.55600</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1013.55600</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1013-1</mods:identifier>
<mods:identifier type="ZooBank">CFDCEFBB523040339D46E302F66E9886</mods:identifier>
<mods:identifier type="Pensoft-UUID">E4329863A39E5EEBA395938413BDD579</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="177205932" LSID="urn:lsid:zoobank.org:act:BEB26BEC-9AE7-41BD-A266-B27AF8A33C35" httpUri="http://treatment.plazi.org/id/E80566B8F2608AC5C4793EEA6D01F2AD" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/BEB26BEC-9AE7-41BD-A266-B27AF8A33C35" authority="Sharkey &amp; van Achterberg" authorityName="Sharkey &amp; van Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Hymenochaonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymenochaonia keithlangdoni" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlangdoni" status="sp. nov.">Hymenochaonia keithlangdoni Sharkey &amp; van Achterberg</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 291" captionStartId="F291" captionText="Figure 291. Hymenochaonia keithlangdoni, holotype." figureDoi="10.3897/zookeys.1013.55600.figure291" httpUri="https://binary.pensoft.net/fig/503746" pageId="0" pageNumber="1">Figure 291</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnostics">
<paragraph pageId="0" pageNumber="1">Diagnostics.</paragraph>
<paragraph pageId="0" pageNumber="1">
BOLD:AAC3252. Consensus barcode. TATATTATATTTTTTATTTGGAATATGATCGGGAATAATTGGTTTATCAATAAGTATAATTATTCGTTTAGAATTAGGTCAATCAGGAACTTTAATTAATAATGATCAAATTTATAATAGATTTGTAACTTCTCATGCATTTATTATAATTTTTTTTATAGTTATACCAATTATAATTGGAGGGTTTGGAAATTGATTAATTCCATTAATATTAGGAAGAGTTGATATGGCTTTCCCTCGAATAAATAATATAAGATTTTGATTGTTAATTCCTTCATTAATAATATTAATTTTAAGAGGATTTATAAATATTGGAGTTGGTACAGGATGAACAGTTTATCCACCTTTATCTTTAAATGTAAGTCATATAGGAATTTCAGTTGATATAGCTATTTTTTCTTTACATTTAGCTGGAATTTCATCTATTATGGGAGCTATTAATTTTATTGTTACTATTTTAAATATACGAAATTATGGGGTGTTAATAGATAAAATTAGTTTATTATGTTGATCAATTTTAATTACAGCTATTTTATTATTATTATCATTACCAGTTTTAGCTGGAGCTATTACTATATTATTAACTGATCGAAATTTAAATACATCTTTTTTTGATCCTTCAGGGGGGGGGGATCCAATTTTGTATCAACATTTATTT. Both
<taxonomicName lsidName="H. keithlangdoni" pageId="0" pageNumber="1" rank="species" species="keithlangdoni">
<emphasis italics="true" pageId="0" pageNumber="1">H. keithlangdoni</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="H. kazumifukunagae" pageId="0" pageNumber="1" rank="species" species="kazumifukunagae">
<emphasis italics="true" pageId="0" pageNumber="1">H. kazumifukunagae</emphasis>
</taxonomicName>
have the ovipositor sheath and antenna of ♀ medially white or ivory combined with a strongly widened vein 1Cua of forewing which is shorter than curved vein cu-a. Differs from
<taxonomicName lsidName="H. kazumifukunagae" pageId="0" pageNumber="1" rank="species" species="kazumifukunagae">
<emphasis italics="true" pageId="0" pageNumber="1">H. kazumifukunagae</emphasis>
</taxonomicName>
by the shorter ovipositor sheath (approx. as long as body) and largely ivory head, mesopleuron and metapleuron.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="holotype ♀">
<paragraph pageId="0" pageNumber="1">Holotype ♀.</paragraph>
<paragraph pageId="0" pageNumber="1">
Guanacaste, Sector Cacao, Sendero Segundo,
<geoCoordinate degrees="10.92679" direction="north" orientation="latitude" precision="1" value="10.92679">10.92679</geoCoordinate>
,
<geoCoordinate degrees="85.45332" direction="west" orientation="longitude" precision="1" value="-85.45332">-85.45332</geoCoordinate>
, 1180 meters, caterpillar collection date: 30/vii/2007, wasp eclosion date: 1/ix/2007. Depository: CNC.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Host data</emphasis>
.
</emphasis>
gelJanzen01 Janzen356 (
<taxonomicName authorityName="Meyrick" authorityYear="1883" class="Insecta" family="Depressariidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Depressariidae</taxonomicName>
) feeding on
<taxonomicName authorityName="Standl" authorityYear="1927" baseAuthorityName="J. D. Smith" class="Magnoliopsida" family="Malvaceae" genus="Hampea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hampea appendiculata" order="Malvales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="appendiculata">
<emphasis italics="true" pageId="0" pageNumber="1">Hampea appendiculata</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="D.F.L.Schlechtendal" authorityYear="1837" class="Magnoliopsida" family="Malvaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Malvales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="family">Malvaceae</taxonomicName>
)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Caterpillar and holotype voucher codes</emphasis>
.
</emphasis>
07-SRNP-36311, DHJPAR0021647.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="paratypes">
<paragraph pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="1">
Hosts =
<taxonomicName authorityName="H.Druce" authorityYear="1895" class="Insecta" family="Crambidae" genus="Conchylodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Conchylodes salamisalis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="salamisalis">
<emphasis italics="true" pageId="0" pageNumber="1">Conchylodes salamisalis</emphasis>
</taxonomicName>
(
<taxonomicName class="Insecta" family="Crambidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Crambidae</taxonomicName>
)
<taxonomicName authorityName="Hubner" authorityYear="1818" class="Insecta" family="Gelechiidae" genus="Dichomeris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Dichomeris" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Dichomeris</emphasis>
</taxonomicName>
Janzen169 (
<taxonomicName authorityName="Stainton" authorityYear="1854" class="Insecta" family="Gelechiidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Gelechiidae</taxonomicName>
). DHJPAR0040592, DHJPAR0040597. Depository: CNC.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sharkey &amp; van Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Hymenochaonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymenochaonia keithlangdoni" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlangdoni">
<emphasis italics="true" pageId="0" pageNumber="1">Hymenochaonia keithlangdoni</emphasis>
</taxonomicName>
is named to honor Mr. Keith Langdon of the US National Park Service for his initiation of the ongoing biodiversity inventory of Great Smoky Mountains National Park.
</paragraph>
<caption doi="10.3897/zookeys.1013.55600.figure291" httpUri="https://binary.pensoft.net/fig/503746" pageId="0" pageNumber="1" start="Figure 291" startId="F291">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 291.</emphasis>
<taxonomicName authorityName="Sharkey &amp; van Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Hymenochaonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymenochaonia keithlangdoni" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="keithlangdoni">
<emphasis italics="true" pageId="0" pageNumber="1">Hymenochaonia keithlangdoni</emphasis>
</taxonomicName>
, holotype.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

149
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•
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. •
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<taxonomicName id="A7ACA2DCDE72FFE1DF96FDE2B285FA48" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[984,1220,521,544]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="1147" phylum="Arthropoda" rank="species" species="atriceps">
<emphasis id="52D8054DDE72FFE1DF96FDE2B285FA48" box="[984,1220,521,544]" italics="true" pageId="2" pageNumber="1147">Camponotus atriceps</emphasis>
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identified through a combination of characters described by
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and
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. Body thickly covered by hairs, from head to gaster; in mesosoma, indistinctly marked, without defined lateral and dorsal regions; head sub-square, with occipital region presenting higher density of hairs; clypeus with a well-developed longitudinal median carina; antennal scapes fully covered with hairs; gaster usually with more hairs in dorsal view. There is a great variation in coloration, including bicolored with head dark brown and body yellow, or body entirely dark brown or entirely yellow.
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128
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. Body covered by golden-yellow or reddish hairs which vary in length; anterior border of clypeus strongly concave, with angled sides and longitudinal median carina well developed; scapes dilated and flattened at apex, with hairs in their whole extension, from the base to apex; mesosoma with well-defined promesonotal suture and poorly defined mesometanotal suture; legs usually reddish-brown.
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•
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coll.,
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(
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,
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<emphasis id="52D8054DDE73FFE0DF63FDCFB58FFA54" bold="true" box="[813,974,548,572]" pageId="3" pageNumber="1148">Identification.</emphasis>
<taxonomicName id="A7ACA2DCDE73FFE0DF92FDCFB2F9FA54" baseAuthorityName="Klug" baseAuthorityYear="1824" box="[988,1208,548,572]" class="Insecta" family="Formicidae" genus="Cephalotes" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="1148" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="52D8054DDE73FFE0DF92FDCFB2F9FA54" box="[988,1208,548,572]" italics="true" pageId="3" pageNumber="1148">Cephalotes pusillus</emphasis>
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(
<figureCitation id="F897C5DADE73FFE0D881FDCFB30AFA54" box="[1231,1355,548,572]" captionStart="Figure 2" captionStartId="3.[168,231,1333,1353]" captionTargetBox="[168,775,156,1309]" captionTargetId="figure-602@3.[168,775,157,1310]" captionTargetPageId="3" captionText="Figure 2. Species of ants found in the stomach contents of eight specimens of Tamandua tetradactyla roadkilled on the BR-040 (Rio de Janeiro–Juiz de Fora) between 2009 and 2014. A–F. Species more frequent in the samples: (A, B) Camponotus atriceps; (C, D) C. rufipes; (E, F) Cephalotes pusillus. G–J. Ant species avoided by anteaters due to their aggressive defenses: (G, H) Neoponera crenata; (I–J) N. villosa." pageId="3" pageNumber="1148">Fig. 2E, F</figureCitation>
) was
</emphasis>
identified with the combination of characters described by
<bibRefCitation id="043DA4AEDE73FFE0DF01FD83B5A3FAE8" author="Kempf WW" box="[847,994,616,640]" pageId="3" pageNumber="1148" pagination="1 - 244" refId="ref5256" refString="Kempf WW (1951) A taxonomic study on the ant tribe Cephalotini (Hymenoptera: Formicidae). Revista de Entomologia 22 (1 - 3): 1 - 244" type="journal article" year="1951">Kempf (1951)</bibRefCitation>
and
<bibRefCitation id="043DA4AEDE73FFE0D857FD83B309FAE8" author="De Andrade ML &amp; Baroni Urbani C" box="[1049,1352,616,640]" pageId="3" pageNumber="1148" pagination="1 - 889" refId="ref4561" refString="De Andrade ML, Baroni Urbani C (1999) Diversity and adaptation in the ant genus Cephalotes, past and present. Stuttgarter Beitrage zur Naturkunde. Serie B 271: 1 - 889." type="journal article" year="1999">Andrade and Urbani (1999)</bibRefCitation>
. Head sub-opaque, shorter than broad, with its upper border convex; occipital region with defined projections; thorax opaque, with anterior edge arched and humerus angulate. Mesosoma with pronotum laterally expanded backward into a narrow, denticulate crest, and with two short teeth on each side, posterior tooth markedly shorter than anterior. Posterior corner of pronotum rectangular and protruding; promesonotal suture well defined. Mesonotum on each side with a very small and sub-rectangular tooth. Basal surface of metanotum in same plane as mesonotum, with a short lateral tooth, larger than second pronotal tooth, and a very long subcylindrical posterior spine originating from posterior corner and diverging at about 45 degrees from longitudinal axis. Propodeum moder- ately convex. Entire thorax finely, quite densely, and strongly reticulate-punctuate, containing a brilliant silver scale. Petiole opaque, finely reticulated, and with an inconspicuous and slightly curved lateral denticle. Postpetiole opaque, as broad as petiole, apically truncated, with a wide basal projection, also finely reticulated with sparse and scaly hairs, and subdivided on each side at the apex, curving slightly forward. Gaster opaque, elliptical, and emarginate anterior medially; first gastral tergite with very narrow lateral border, not distinctly defined and without crest; tergites and sternites finely and rather reticulate-punctuate.
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120
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<emphasis id="52D8054DDE73FFE0DF63F9F3B242FE47" bold="true" box="[813,1027,1560,1583]" italics="true" pageId="3" pageNumber="1148">Neoponera crenata</emphasis>
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<emphasis id="52D8054DDE73FFE0DF63F9A2B255FE09" bold="true" box="[813,1044,1609,1633]" pageId="3" pageNumber="1148">Material examined.</emphasis>
<materialsCitation id="D0C4D302DE73FFE0D86EF9A1B5A8FEEB" country="Brazil" location="Rio de Janeiro" pageId="3" pageNumber="1148" specimenCount="1" stateProvince="Rio de Janeiro">
<collectingCountry id="18BB99CFDE73FFE0D86EF9A1B2C7FE0A" box="[1056,1158,1610,1634]" name="Brazil" pageId="3" pageNumber="1148">BRAZIL</collectingCountry>
:
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: along highway BR-040
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•
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8 (
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),
<collectorName id="CD59BC89DE73FFE0DF93F966B20FFECD" box="[989,1102,1677,1701]" pageId="3" pageNumber="1148">C. Bueno</collectorName>
coll.,
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<collectingDate id="04560677DE73FFE0D8E6F966B360FECD" box="[1192,1313,1677,1701]" pageId="3" pageNumber="1148" value="2014-11-02">02-XI-2014</collectingDate>
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(
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to
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<emphasis id="52D8054DDE73FFE0DF92F909B2F5FE91" box="[988,1204,1762,1785]" italics="true" pageId="3" pageNumber="1148">Neoponera crenata</emphasis>
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(
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identified following the description of
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. Anterior medial region of clypeus with a strong protuberance; eyes relatively large; carina malar not defined; mesosoma with pronotum carina present but poorly defined; propodeal spiracle oval; petiole, in lateral view, as long as high, with anterior and posterior faces almost parallel and dorsal margin convex; head and back of mesosoma punctuate; side of pronotum weakly punctuate, smooth, and shiny; gaster with poorly developed stridulatory organ on second tergite; the whole body usually reddish-brown.
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•
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(
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following descriptions by
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and
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works where they highlighted several conspicuous features. Body large (about
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), black, covered by long golden-yellow hairs. Legs reddish-brown. Petiole thick in profile, with anterior face slightly concave and posterior face rounded and convex. Anterior clypeal margin convex but concave medially with a median notch; malar carina well defined; stridulatory organ is well developed in second tergite on gaster.
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<paragraph pageNumber="676">&quot;Habitat in Zeylona.&quot; RCN: 7652.</paragraph>
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in Plukenet, Phytographia: t. 211, f. 5. 1692; Almag. Bot.: 294. 1696.
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Current name:
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(L. f.) H.S. Irwin &amp; Barneby (
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24.
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<paragraph id="077CBF83D379CCD2965D34F8ED7FEB4E" pageNumber="148">[[ worker ]] maj. 5 — 5.5 Millim. [[ worker ]] min. 4.5 Millim. lang, jener dunkelroth, dieser hell gelbroth, beide mit schwarzem Hinterleib, ohne Glanz, reichlich mit abstehenden weisslich gelben Haaren besetzt, besonders am Rande des Metanotum « und auf der Schuppe; sehr feine, kurze weissliche Haerchen findet man an Kopf und Thorax sehr sparsam, am Abdomen reichlicher, so dass dieses bei reinen Exemplaren gelblich schimmert. Die Beine sind bald heller, bald dunkler rotii, bei einem Stuecke mit Ausnahme der Vorderschienen und Tarsen fast schwarz. Fuehler hellroth.</paragraph>
<paragraph id="87027DF7A0150850658DB9A8DD9A1D2D" pageNumber="148">Die Mandibeln, ganz vorn fast glatt, sonst aeusserst fein laengs gerunzelt und mit zerstreuten, grossen Punkten besetzt, haben 6 — 7 schwarze Zaehne. Clipeus in der Mitte lappig vorgezogen kaum gekielt. Der Kopf ist dicht fingerhutartig punktirt und hat ausserdem noch auf dem Clipeus und den Wangen zahlreiche, groessere Punkte, die aber beim kleinen [[ worker ]] fast fehlen.</paragraph>
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Der Thorax ist oben breit, fast flach, an den Ecken des Pronotums am breitesten, nach hinten allmaelig verschmaelert, ganz aehnlich wie bei
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. Die drei Bruststuecke sind durch Querlinien deutlich getrennt. Das Metanotum ¡ st hinten abgestutzt und beim groesseren [[ worker ]] etwas ausgehoehlt. Der Thorax ist fingerhutartig dicht punktirt, seitlich etwas laengs gerunzelt. Die Schuppe ist bei dem grossen [[ worker ]], platt, oben gerundet, beim kleinen [[ worker ]] dicker, vorn ziemlich gewoelbt, oben ebenfalls bogenfoermig. Das Abdomen ist aeusserst dicht
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punktirt, die Segmente sind gelblich gesaeumt. Schienen mit kurzen, kaum absiebenden Haerchen.
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Diese Art hat grosse Aehnlichkeit mit
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Mayr, um so mehr, als von diesem Stuecke vorkommen, welche einen- ganz rothen Kopf haben; die neue Art ist aber entschieden kleiner und hat eine andere Skulptur des Hinterleibs, indem dieser einfach dicht fingerhutartig punktirt ist, bei
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aber ausser der Punktirung noch eine Menge grosser, eingestochener Punkte bat, die die Skulptur groeber, fast schuppenartig erscheinen lassen.
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<paragraph id="4C7C9F7E0EFCF401F4B2926E470C5274" pageNumber="149">[[ queen ]] 8 Millim. lang, hellroth mit schwarzem Hinterleib, Hinterbeine dunkler roth. Behaarung und Skulptur ganz wie beim [[ worker ]]. Der Kopf ist laenglich, nur so breit ak der Thorax. Die Fluegel sind leicht gelblich.</paragraph>
<paragraph id="EC2514D27BAF288CF29CFCE926E106BB" pageNumber="149">[[ male ]] etwas ueber 5 Millim. lang. schwach glaenzend, an Kopf und Thorax kuerzer and spaerlicher, am Hinterleib laenger und reichlicher abstehend behaart. Der Kopf ist dicht fingerhutartig punktirt, Mandibeln braeunlich, schneidend mit zahnfoermiger Spitze. Fuehlerschaft lang, schwarz, Radicula gelb, Geissel braeunlich. Pronotum sehr schmal, vom Mesonotum ganz ueberragt; dieses ist vorn lederartig gerunzelt mit grossem, flachen Punkten und hat vom Pronotum an bis zu seiner hoechsten Woelbung eine eingedrueckte Laengslinie; seine Scheibe ist roth durchscheinend, fingerhutartig punktirt; das Scutellum ebenso, aber etwas weitlaeufiger und feiner punktirt. Schuppe sehr niedrig, quer, dick. Cuba.</paragraph>
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<taxonomicName LSID="http://species-id.net/wiki/Onthophagus_kokodanus_hagenaltus" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus kokodanus" order="Coleoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="kokodanus" subSpecies="hagenaltus">Onthophagus kokodanus hagenaltus</taxonomicName>
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Figs 7-816, 2039-44
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<paragraph pageId="12" pageNumber="61">Type material.</paragraph>
<paragraph pageId="12" pageNumber="61">5 males, 2 collection records. Holotype male (CMNC) from Papua New Guinea: 25 mi NE Mount Hagen, 6000 ft, 5.vii.1974, H.F. Howden.</paragraph>
<paragraph pageId="12" pageNumber="61">Paratypes: Papua New Guinea: 25 mi NE Mount Hagen, 05/vii/1974, Howden, 6000ft, 3 spm., incl. holotype, in CMNC. Western Highlands: Mt. Hagen, 05-08/vii/1974, Peck, 6000ft, oak forest, 2 spm., in CMNC</paragraph>
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<paragraph pageId="12" pageNumber="61">Diagnosis.</paragraph>
<paragraph pageId="12" pageNumber="61">The vertexal horns of this subspecies are more upright, even in major males, with angular tubercle on top of the interconnecting basal ridge. Dorsum black-brown, largely shining (particularly the elytra), with slight greenish metallic lustre. Elytral strial punctures all equally small, distinct, scatteredly crenulating finely punctate interstriae. Lateral border of pronotum distinctly rounded at about 0.4 of length behind anterolateral angle (not angular, in dorsal view, Fig. 40). Pronotum usually with blunt bisinuate rim behind depressed, medially more or less concave anterior declivity. Clypeofrontal ridge low, very slightly arcuate, ends on either side angularly connected to clypeogenal ridge. Most of dorsal side finely, sparsely punctate. Clypeal tip distinctly, broadly reflexed, apical border bisinuate. Dorsal eye parts broadly elliptic, separated by about three eye widths. Pygidium abundantly, finely, distinctly punctate. Body length usually 10-11 mm. Female unknown.</paragraph>
<paragraph pageId="12" pageNumber="61">Apparently a close highland relative of the preceding subspecies.</paragraph>
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<paragraph pageId="12" pageNumber="61">Description</paragraph>
<paragraph pageId="12" pageNumber="61">(holotype, male). Body length ca 11 mm. Habitus generally convex, robust. Colour of dorsal side black, generally shiny, with slight metallic lustre; ventral parts largely black, partly matt (microreticulate); legs black-brown, shiny. Dorsal side and pygidium virtually glabrous (apart from any inconspicuous micro-stubbles); ventral side and legs with numerous long, light-brown setae.</paragraph>
<paragraph pageId="12" pageNumber="61">Clypeal border broadly, distinctly marginate, sides virtually straight from genae to bisinuate, reflexed, shortly lobiform apex; clypeal surface shiny, with sparse to abundant, minute punctation; clypeogenal transition at border obtusely angular, slightly elevated, on either side with straight ridge curving to end of virtually straight, distinct clypeofrontal ridge. Genal and frontal surface sparsely, minutely punctate; genal border evenly, widely rounded. Vertex between posterior end of eyes with pair of long, basally very broad horns, connected at base by angular ridge, with distinct anteromedian angle; horns slightly curved, directed upright behind eyes (not beyond, in axial view), evenly tapering to blunt tip, inner edge of horns evenly concave (in axial view); horn surface with convex anterior surface, minutely, sparsely punctate. Frontal disc limited by fine frontogenal ridge, on either side extending along eye. Dorsal eye parts widely elliptic, with ca 28 facet rows across widest point. Ratio of interocular distance to maximum (transverse) eye width ca 3.0.</paragraph>
<paragraph lastPageId="13" lastPageNumber="62" pageId="12" pageNumber="61">
Pronotum generally strongly convex, surface shining, with greenish lustre; disc posteriorly slightly convex (midline impression effaced), anterior surface broadly depressed, medially shallowly concave, up to posterior, strongly bisinuate, transverse protrusion coming from disc forward; anterior and lateral borders of pron
<pageBreakToken pageId="13" pageNumber="62" start="start">otum</pageBreakToken>
marginate; anterior section of lateral border slightly concave; anterolateral angle rectangular, shortly rounded; posterior section of lateral border slightly sinuate, posterolateral angle rounded; base finely marginate, with very obtuse basomedian angle. Most of pronotal surface finely, sparsely punctate, anterior depression minutely, sparsely punctate, this punctation interspersed with vague micropunctation; lateral declivities more matt (microreticulate).
</paragraph>
<paragraph pageId="13" pageNumber="62">Elytra generally shiny, with greenish metallic lustre; scutellum indistinct in dorsal view, shape of base and apex unmodified; anterior half of epipleuron distinctly punctate-setose. Elytra broad, with 8 straight to slightly curved, fine, distinct striae; stria 7 slightly sinuate in front, extending onto shining humeral umbone; strial punctures distinct, small, widely separated (mostly 5-8 puncture diameters), distinctly crenulating interstrial edges. Interstrial surfaces virtually flat to very slightly convex (from disc to lateral declivity), with sparse to abundant, fine micropunctation.</paragraph>
<paragraph pageId="13" pageNumber="62">Antennal club medium-brown, scapus unmodified. Mentum shallowly emarginate in front. Prothoracic sides with coxal-marginal ridge distinct, curving forward onto lateral border, much of surface annulate-punctate-setose, including large, seta-bearing hemi-punctures. Anterior lobe of metasternum slightly convex, abundantly punctate-setose, disc micropunctate; metasternal sides and adjacent parts matt, abundantly to densely annulate-punctate-setose. Abdominal ventrites matt, each laterally with row of seta-bearing annulate punctures. Pygidium black, weakly shiny (sericeous), with abundant, shallow, fine punctation, without macro-setae; surface slightly convex, base with transverse ridge, apex marginate.</paragraph>
<paragraph pageId="13" pageNumber="62">Legs robust. Protibia with 3+1 larger external denticles (distal 3 longer, sub-acuminate), separated by some serration; proximal serration consisting of 7 smaller denticles; apico-external denticle oblique to tibial axis; apical edge slightly rounded, with down-curved spur (tip worn away); protibial underside with low longitudinal crest; protarsus unmodified. Profemur robust, underside finely punctate, and with numerous larger seta-bearing hemi-punctures. Meso- and metafemoral undersides also finely punctate, with fewer large seta-bearing hemi-punctures. Meso- and metatibiae strongly dilated distad to transversely subelliptic crest, which is fringed with fine fossorial spines; external side with ca 4 sets of distinct, spine-bearing fossorial protrusions. Tarsi generally slender, with well-developed sickle-shaped claws; meso- and metatarsomeres 1 straight, unmodified. Spurs on meso- and metatibiae elongate-acuminate. Relative length proportionsof metatibial spur to metatarsomeres 1-5: ca 12, 13, 6, 4, 3, 6.</paragraph>
<paragraph pageId="13" pageNumber="62">Parameres narrow (dorsal view), tapering, curved downward (lateral view), Fig. 20.</paragraph>
<paragraph pageId="13" pageNumber="62">Measurements in dorsal view: maximum width of head 3.2 mm, median length of pronotum 3.5 mm, maximum width 5.5 mm, sutural length of elytra 4.6 mm, maximum width of combined elytra 5.8 mm.</paragraph>
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<paragraph pageId="13" pageNumber="62">Variability and sexual dimorphism.</paragraph>
<paragraph pageId="13" pageNumber="62">Female unknown; females probably have a transverse vertexal ridge, not the variably long horns of the males. Strongly varying: there is a minor male looking like a female, having a transverse vertexal ridge (Fig. 44), its aedeagus being similar to that of the holotype. Body length 9.5-11 mm.</paragraph>
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<paragraph pageId="13" pageNumber="62">Etymology.</paragraph>
<paragraph pageId="13" pageNumber="62">Name refers to the highland nature of the type locality.</paragraph>
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63
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<mods:title>A new genus from Madagascar with strongly enlarged labium (Ephemeroptera, Baetidae)</mods:title>
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<mods:namePart>Kaltenbach, Thomas</mods:namePart>
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<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
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<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<taxonomicName LSID="http://zoobank.org/871DA221-4FDB-4CF6-B81B-88E6C35527AB" authority="Kaltenbach &amp; Gattolliat, 2021" authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Megalabiops" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Megalabiops" order="Diptera" pageId="0" pageNumber="465" rank="genus" status="gen. nov.">Megalabiops</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="465">gen. nov.</taxonomicNameLabel>
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<paragraph pageId="0" pageNumber="465">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megalabiops madagasikara gen. et sp. nov., habitus, nymph a dorsal view b ventral view c, d lateral view. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure1" httpUri="https://binary.pensoft.net/fig/610392" pageId="0" pageNumber="465">Figures 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megalabiops madagasikara gen. et sp. nov., nymph morphology a frontal section of head b head, ventral view c labium, lateral view d head, ventrolateral view. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure2" httpUri="https://binary.pensoft.net/fig/610393" pageId="0" pageNumber="465">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megalabiops madagasikara gen. et sp. nov., nymph morphology a base of antenna b labrum (left: ventral view, right: dorsal view) c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h left incisor and kinetodontium. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure3" httpUri="https://binary.pensoft.net/fig/610394" pageId="0" pageNumber="465">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megalabiops madagasikara gen. et sp. nov., nymph morphology a hypopharynx and superlinguae b maxilla (ds: denti-seta) c seta at tip of maxilla d labium (left: ventral view, right: dorsal view) e apex of paraglossa (ventral view). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure4" httpUri="https://binary.pensoft.net/fig/610395" pageId="0" pageNumber="465">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megalabiops madagasikara gen. et sp. nov., nymph morphology a foreleg b fore claw c hind leg d head. Scale bars: 0.2 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure5" httpUri="https://binary.pensoft.net/fig/610396" pageId="0" pageNumber="465">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megalabiops madagasikara gen. et sp. nov., nymph morphology a tergite IV (posterior margin) b paraproct c gill IV (with enlarged margins). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure6" httpUri="https://binary.pensoft.net/fig/610397" pageId="0" pageNumber="465">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megalabiops madagasikara gen. et sp. nov., distribution in Madagascar." figureDoi="10.3897/afrinvertebr.62.73911.figure7" httpUri="https://binary.pensoft.net/fig/610398" pageId="0" pageNumber="465">, 7</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="465" type="type species">
<paragraph pageId="0" pageNumber="465">Type species.</paragraph>
<paragraph pageId="0" pageNumber="465">
<taxonomicName authorityName="gen. et" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Megalabiops" kingdom="Animalia" lsidName="Megalabiops madagasikara" order="Ephemeroptera" pageId="0" pageNumber="465" rank="species" species="madagasikara">
<emphasis italics="true" pageId="0" pageNumber="465">Megalabiops madagasikara</emphasis>
</taxonomicName>
gen. et sp. nov., by present designation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="465" type="diagnosis">
<paragraph pageId="0" pageNumber="465">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Nymph.</emphasis>
This new genus is distinguished by the combination of the following characters: A) body rather short and stocky (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megalabiops madagasikara gen. et sp. nov., habitus, nymph a dorsal view b ventral view c, d lateral view. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure1" httpUri="https://binary.pensoft.net/fig/610392" pageId="0" pageNumber="465">1</figureCitation>
); B) base of antennae close to each other, with rather broad carina between them (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megalabiops madagasikara gen. et sp. nov., nymph morphology a frontal section of head b head, ventral view c labium, lateral view d head, ventrolateral view. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure2" httpUri="https://binary.pensoft.net/fig/610393" pageId="0" pageNumber="465">2a</figureCitation>
); C) pedicellus with many long, stout, pointed setae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megalabiops madagasikara gen. et sp. nov., nymph morphology a base of antenna b labrum (left: ventral view, right: dorsal view) c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h left incisor and kinetodontium. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure3" httpUri="https://binary.pensoft.net/fig/610394" pageId="0" pageNumber="465">3a</figureCitation>
); D) labrum rather small, lateral margins straight, distal margin largely V-shaped; dorsal surface with one long, submedian seta and submarginal arc of setae restricted to a few simple setae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megalabiops madagasikara gen. et sp. nov., nymph morphology a base of antenna b labrum (left: ventral view, right: dorsal view) c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h left incisor and kinetodontium. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure3" httpUri="https://binary.pensoft.net/fig/610394" pageId="0" pageNumber="465">3b</figureCitation>
); E) both mandibles with brush of medium, fine setae between prostheca and mola (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megalabiops madagasikara gen. et sp. nov., nymph morphology a base of antenna b labrum (left: ventral view, right: dorsal view) c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h left incisor and kinetodontium. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure3" httpUri="https://binary.pensoft.net/fig/610394" pageId="0" pageNumber="465">3c, f</figureCitation>
); F) hypopharynx with large, medial tuft of stout setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megalabiops madagasikara gen. et sp. nov., nymph morphology a hypopharynx and superlinguae b maxilla (ds: denti-seta) c seta at tip of maxilla d labium (left: ventral view, right: dorsal view) e apex of paraglossa (ventral view). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure4" httpUri="https://binary.pensoft.net/fig/610395" pageId="0" pageNumber="465">4a</figureCitation>
); G) maxillary palp 2-segmented (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megalabiops madagasikara gen. et sp. nov., nymph morphology a hypopharynx and superlinguae b maxilla (ds: denti-seta) c seta at tip of maxilla d labium (left: ventral view, right: dorsal view) e apex of paraglossa (ventral view). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure4" httpUri="https://binary.pensoft.net/fig/610395" pageId="0" pageNumber="465">4b</figureCitation>
); H) labium with strongly enlarged mentum (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megalabiops madagasikara gen. et sp. nov., nymph morphology a frontal section of head b head, ventral view c labium, lateral view d head, ventrolateral view. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure2" httpUri="https://binary.pensoft.net/fig/610393" pageId="0" pageNumber="465">2b-d</figureCitation>
); glossae basally broad, narrowing toward apex, shorter than paraglossae; inner ventrolateral margin of glossae with a row of long, simple setae, close to ventrolateral margin with a row of very long, simple setae, apically with some very robust, long, curved setae; stout paraglossae, slightly curved inward, apex with three rows of stout, pectinate setae; labial palps segment II without distolateral protuberance, segment III slightly pentagonal (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megalabiops madagasikara gen. et sp. nov., nymph morphology a hypopharynx and superlinguae b maxilla (ds: denti-seta) c seta at tip of maxilla d labium (left: ventral view, right: dorsal view) e apex of paraglossa (ventral view). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure4" httpUri="https://binary.pensoft.net/fig/610395" pageId="0" pageNumber="465">4d, e</figureCitation>
); I) forelegs with a row of long, spine-like setae on dorsal margin of femur; femoral patch absent on all legs; patellotibial suture absent on foreleg and present on middle and hind legs; claw strongly elongated, pointed, with two rows of numerous, minute denticles (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megalabiops madagasikara gen. et sp. nov., nymph morphology a foreleg b fore claw c hind leg d head. Scale bars: 0.2 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure5" httpUri="https://binary.pensoft.net/fig/610396" pageId="0" pageNumber="465">5a, b</figureCitation>
); J) posterior margin of abdominal tergites with triangular spines, lateral margins without spines (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megalabiops madagasikara gen. et sp. nov., nymph morphology a tergite IV (posterior margin) b paraproct c gill IV (with enlarged margins). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure6" httpUri="https://binary.pensoft.net/fig/610397" pageId="0" pageNumber="465">6a</figureCitation>
).
</paragraph>
<caption doi="10.3897/afrinvertebr.62.73911.figure1" httpUri="https://binary.pensoft.net/fig/610392" pageId="0" pageNumber="465" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Figure 1.</emphasis>
<taxonomicName authorityName="gen. et" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Megalabiops" kingdom="Animalia" lsidName="Megalabiops madagasikara" order="Ephemeroptera" pageId="0" pageNumber="465" rank="species" species="madagasikara">
<emphasis italics="true" pageId="0" pageNumber="465">Megalabiops madagasikara</emphasis>
</taxonomicName>
gen. et sp. nov., habitus, nymph
<emphasis bold="true" pageId="0" pageNumber="465">a</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="465">b</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="465">c,d</emphasis>
lateral view. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="465" type="imagines">
<paragraph pageId="0" pageNumber="465">Imagines.</paragraph>
<paragraph pageId="0" pageNumber="465">Unknown.</paragraph>
<caption doi="10.3897/afrinvertebr.62.73911.figure2" httpUri="https://binary.pensoft.net/fig/610393" pageId="0" pageNumber="465" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Figure 2.</emphasis>
<taxonomicName authorityName="gen. et" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Megalabiops" kingdom="Animalia" lsidName="Megalabiops madagasikara" order="Ephemeroptera" pageId="0" pageNumber="465" rank="species" species="madagasikara">
<emphasis italics="true" pageId="0" pageNumber="465">Megalabiops madagasikara</emphasis>
</taxonomicName>
gen. et sp. nov., nymph morphology
<emphasis bold="true" pageId="0" pageNumber="465">a</emphasis>
frontal section of head
<emphasis bold="true" pageId="0" pageNumber="465">b</emphasis>
head, ventral view
<emphasis bold="true" pageId="0" pageNumber="465">c</emphasis>
labium, lateral view
<emphasis bold="true" pageId="0" pageNumber="465">d</emphasis>
head, ventrolateral view. Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="465" type="etymology">
<paragraph pageId="0" pageNumber="465">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="465">
<taxonomicName authorityName="Kaltenbach &amp; Gattolliat" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Megalabiops" kingdom="Animalia" lsidName="Megalabiops" order="Ephemeroptera" pageId="0" pageNumber="465" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="465">Megalabiops</emphasis>
</taxonomicName>
is an arbitrary combination of letters with allusion to the Greek words
<emphasis italics="true" pageId="0" pageNumber="465">mega</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="465">iops</emphasis>
, and the Latin word
<emphasis italics="true" pageId="0" pageNumber="465">labium</emphasis>
.
<emphasis italics="true" pageId="0" pageNumber="465">Megalab</emphasis>
- is with reference to the strongly enlarged
<emphasis italics="true" pageId="0" pageNumber="465">labium</emphasis>
and -
<emphasis italics="true" pageId="0" pageNumber="465">iops</emphasis>
with reference to the
<taxonomicName authorityName="Leach" authorityYear="1815" class="Insecta" family="Baetidae" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="0" pageNumber="465" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
which look and move like small fishes. The gender is feminine.
</paragraph>
<caption doi="10.3897/afrinvertebr.62.73911.figure3" httpUri="https://binary.pensoft.net/fig/610394" pageId="0" pageNumber="465" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Figure 3.</emphasis>
<taxonomicName authorityName="gen. et" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Megalabiops" kingdom="Animalia" lsidName="Megalabiops madagasikara" order="Ephemeroptera" pageId="0" pageNumber="465" rank="species" species="madagasikara">
<emphasis italics="true" pageId="0" pageNumber="465">Megalabiops madagasikara</emphasis>
</taxonomicName>
gen. et sp. nov., nymph morphology
<emphasis bold="true" pageId="0" pageNumber="465">a</emphasis>
base of antenna
<emphasis bold="true" pageId="0" pageNumber="465">b</emphasis>
labrum (left: ventral view, right: dorsal view)
<emphasis bold="true" pageId="0" pageNumber="465">c</emphasis>
right mandible
<emphasis bold="true" pageId="0" pageNumber="465">d</emphasis>
right prostheca
<emphasis bold="true" pageId="0" pageNumber="465">e</emphasis>
right incisor and kinetodontium
<emphasis bold="true" pageId="0" pageNumber="465">f</emphasis>
left mandible
<emphasis bold="true" pageId="0" pageNumber="465">g</emphasis>
left prostheca
<emphasis bold="true" pageId="0" pageNumber="465">h</emphasis>
left incisor and kinetodontium. Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="465" type="description">
<paragraph pageId="0" pageNumber="465">Description.</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Nymph.</emphasis>
(Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megalabiops madagasikara gen. et sp. nov., habitus, nymph a dorsal view b ventral view c, d lateral view. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure1" httpUri="https://binary.pensoft.net/fig/610392" pageId="0" pageNumber="465">1</figureCitation>
-
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megalabiops madagasikara gen. et sp. nov., distribution in Madagascar." figureDoi="10.3897/afrinvertebr.62.73911.figure7" httpUri="https://binary.pensoft.net/fig/610398" pageId="0" pageNumber="465">7</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Body.</emphasis>
Short and stocky (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megalabiops madagasikara gen. et sp. nov., habitus, nymph a dorsal view b ventral view c, d lateral view. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure1" httpUri="https://binary.pensoft.net/fig/610392" pageId="0" pageNumber="465">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megalabiops madagasikara gen. et sp. nov., nymph morphology a frontal section of head b head, ventral view c labium, lateral view d head, ventrolateral view. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure2" httpUri="https://binary.pensoft.net/fig/610393" pageId="0" pageNumber="465">2</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Head</emphasis>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Antenna</emphasis>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megalabiops madagasikara gen. et sp. nov., nymph morphology a frontal section of head b head, ventral view c labium, lateral view d head, ventrolateral view. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure2" httpUri="https://binary.pensoft.net/fig/610393" pageId="0" pageNumber="465">2a</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megalabiops madagasikara gen. et sp. nov., nymph morphology a base of antenna b labrum (left: ventral view, right: dorsal view) c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h left incisor and kinetodontium. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure3" httpUri="https://binary.pensoft.net/fig/610394" pageId="0" pageNumber="465">3a</figureCitation>
). Antennae bases close to each other, with broad carina between them. Pedicellus with many long, stout, pointed setae; scapus bare.
</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Labrum</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megalabiops madagasikara gen. et sp. nov., nymph morphology a base of antenna b labrum (left: ventral view, right: dorsal view) c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h left incisor and kinetodontium. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure3" httpUri="https://binary.pensoft.net/fig/610394" pageId="0" pageNumber="465">3b</figureCitation>
). Rather small, lateral margins straight and distal margin largely V-shaped, wider than long; on dorsal surface with one submedian, long, simple seta and submarginal arc of setae restricted to few simple setae; on ventral margin anterolaterally and medially with a row of long, fine setae.
</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Right mandible</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megalabiops madagasikara gen. et sp. nov., nymph morphology a base of antenna b labrum (left: ventral view, right: dorsal view) c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h left incisor and kinetodontium. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure3" httpUri="https://binary.pensoft.net/fig/610394" pageId="0" pageNumber="465">3c, d, e</figureCitation>
). Incisor and kinetodontium cleft, kinetodontium with distinct denticles; inner margin of innermost denticle without a row of thin setae; prostheca stick-like, apicolaterally denticulate; margin between prostheca and mola with brush of abundant, medium, fine setae.
</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Left mandible</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Megalabiops madagasikara gen. et sp. nov., nymph morphology a base of antenna b labrum (left: ventral view, right: dorsal view) c right mandible d right prostheca e right incisor and kinetodontium f left mandible g left prostheca h left incisor and kinetodontium. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure3" httpUri="https://binary.pensoft.net/fig/610394" pageId="0" pageNumber="465">3f, g, h</figureCitation>
). Incisor and kinetodontium partially cleft, kinetodontium with distinct denticles; prostheca robust, apically with denticles and comb-shaped structure; margin between prostheca and mola with brush of abundant, medium, fine setae; mola slightly reduced in size.
</paragraph>
<paragraph pageId="0" pageNumber="465">
<taxonomicName genus="Hypopharynx" lsidName="Hypopharynx" pageId="0" pageNumber="465" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Hypopharynx</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megalabiops madagasikara gen. et sp. nov., nymph morphology a hypopharynx and superlinguae b maxilla (ds: denti-seta) c seta at tip of maxilla d labium (left: ventral view, right: dorsal view) e apex of paraglossa (ventral view). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure4" httpUri="https://binary.pensoft.net/fig/610395" pageId="0" pageNumber="465">4a</figureCitation>
). Lingua rather broad and expanded, with large dorsomedial tuft of stout setae.
</paragraph>
<caption doi="10.3897/afrinvertebr.62.73911.figure4" httpUri="https://binary.pensoft.net/fig/610395" pageId="0" pageNumber="465" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Figure 4.</emphasis>
<taxonomicName authorityName="gen. et" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Megalabiops" kingdom="Animalia" lsidName="Megalabiops madagasikara" order="Ephemeroptera" pageId="0" pageNumber="465" rank="species" species="madagasikara">
<emphasis italics="true" pageId="0" pageNumber="465">Megalabiops madagasikara</emphasis>
</taxonomicName>
gen. et sp. nov., nymph morphology
<emphasis bold="true" pageId="0" pageNumber="465">a</emphasis>
hypopharynx and superlinguae
<emphasis bold="true" pageId="0" pageNumber="465">b</emphasis>
maxilla (ds: denti-seta)
<emphasis bold="true" pageId="0" pageNumber="465">c</emphasis>
seta at tip of maxilla
<emphasis bold="true" pageId="0" pageNumber="465">d</emphasis>
labium (left: ventral view, right: dorsal view)
<emphasis bold="true" pageId="0" pageNumber="465">e</emphasis>
apex of paraglossa (ventral view). Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Maxilla</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megalabiops madagasikara gen. et sp. nov., nymph morphology a hypopharynx and superlinguae b maxilla (ds: denti-seta) c seta at tip of maxilla d labium (left: ventral view, right: dorsal view) e apex of paraglossa (ventral view). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure4" httpUri="https://binary.pensoft.net/fig/610395" pageId="0" pageNumber="465">4b, c</figureCitation>
). Apically with three canines and three denti-setae; distal denti-seta tooth-like, closely attached to canines, following same direction as canines, other denti-setae slender, bifid and pectinate; maxillary palp 2-segmented, apically pointed and with stout seta at the tip.
</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Labium</emphasis>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Megalabiops madagasikara gen. et sp. nov., nymph morphology a frontal section of head b head, ventral view c labium, lateral view d head, ventrolateral view. Scale bars: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure2" httpUri="https://binary.pensoft.net/fig/610393" pageId="0" pageNumber="465">2b-d</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Megalabiops madagasikara gen. et sp. nov., nymph morphology a hypopharynx and superlinguae b maxilla (ds: denti-seta) c seta at tip of maxilla d labium (left: ventral view, right: dorsal view) e apex of paraglossa (ventral view). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure4" httpUri="https://binary.pensoft.net/fig/610395" pageId="0" pageNumber="465">4d, e</figureCitation>
). Mentum strongly enlarged (long, very thick). Glossae basally broad, narrowing toward apex, shorter than paraglossae; inner ventrolateral margin with row of long, simple setae and close to inner margin with row of very long, simple setae; apically with some very robust, long, curved setae. Paraglossae broad, slightly curved inward, with convex outer margin; apically with three rows of long, stout, pectinate setae. Labial palps long, segment II without distolateral protuberance, segment III almost pentagonal, distal margin straight, lateral margin curved.
</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">
Thorax.
<emphasis italics="true" pageId="0" pageNumber="465">Hind protoptera</emphasis>
</emphasis>
well developed.
</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Foreleg</emphasis>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megalabiops madagasikara gen. et sp. nov., nymph morphology a foreleg b fore claw c hind leg d head. Scale bars: 0.2 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure5" httpUri="https://binary.pensoft.net/fig/610396" pageId="0" pageNumber="465">5a, b</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Femur</emphasis>
stocky with a row of long, spine-like setae on dorsal margin and many long, spine-like setae along ventral margin; femoral patch absent.
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Tibia</emphasis>
stocky, slightly shorter than tarsus. Patellotibial suture absent, connection between tibia and tarsus tight.
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Claw</emphasis>
strongly elongated, pointed; with two long rows of numerous, minute denticles; subapical setae absent.
</paragraph>
<caption doi="10.3897/afrinvertebr.62.73911.figure5" httpUri="https://binary.pensoft.net/fig/610396" pageId="0" pageNumber="465" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Figure 5.</emphasis>
<taxonomicName authorityName="gen. et" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Megalabiops" kingdom="Animalia" lsidName="Megalabiops madagasikara" order="Ephemeroptera" pageId="0" pageNumber="465" rank="species" species="madagasikara">
<emphasis italics="true" pageId="0" pageNumber="465">Megalabiops madagasikara</emphasis>
</taxonomicName>
gen. et sp. nov., nymph morphology
<emphasis bold="true" pageId="0" pageNumber="465">a</emphasis>
foreleg
<emphasis bold="true" pageId="0" pageNumber="465">b</emphasis>
fore claw
<emphasis bold="true" pageId="0" pageNumber="465">c</emphasis>
hind leg
<emphasis bold="true" pageId="0" pageNumber="465">d</emphasis>
head. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Middle and hind legs</emphasis>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megalabiops madagasikara gen. et sp. nov., nymph morphology a foreleg b fore claw c hind leg d head. Scale bars: 0.2 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure5" httpUri="https://binary.pensoft.net/fig/610396" pageId="0" pageNumber="465">5c</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Femur</emphasis>
slenderer than at foreleg; femoral patch absent.
<emphasis bold="true" pageId="0" pageNumber="465">
<emphasis italics="true" pageId="0" pageNumber="465">Tibia</emphasis>
.
</emphasis>
Patellotibial suture present.
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Claw</emphasis>
as in foreleg.
</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">
Abdomen.
<emphasis italics="true" pageId="0" pageNumber="465">Tergites</emphasis>
</emphasis>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megalabiops madagasikara gen. et sp. nov., nymph morphology a tergite IV (posterior margin) b paraproct c gill IV (with enlarged margins). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure6" httpUri="https://binary.pensoft.net/fig/610397" pageId="0" pageNumber="465">6a</figureCitation>
). Posterior margin with triangular spines on all segments; lateral margins without spines.
</paragraph>
<caption doi="10.3897/afrinvertebr.62.73911.figure6" httpUri="https://binary.pensoft.net/fig/610397" pageId="0" pageNumber="465" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Figure 6.</emphasis>
<taxonomicName authorityName="gen. et" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Megalabiops" kingdom="Animalia" lsidName="Megalabiops madagasikara" order="Ephemeroptera" pageId="0" pageNumber="465" rank="species" species="madagasikara">
<emphasis italics="true" pageId="0" pageNumber="465">Megalabiops madagasikara</emphasis>
</taxonomicName>
gen. et sp. nov., nymph morphology
<emphasis bold="true" pageId="0" pageNumber="465">a</emphasis>
tergite IV (posterior margin)
<emphasis bold="true" pageId="0" pageNumber="465">b</emphasis>
paraproct
<emphasis bold="true" pageId="0" pageNumber="465">c</emphasis>
gill IV (with enlarged margins). Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Gills</emphasis>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megalabiops madagasikara gen. et sp. nov., nymph morphology a tergite IV (posterior margin) b paraproct c gill IV (with enlarged margins). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure6" httpUri="https://binary.pensoft.net/fig/610397" pageId="0" pageNumber="465">6c</figureCitation>
). Seven pairs of gills on segments I-VII, dorsally oriented.
</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Paraproct</emphasis>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Megalabiops madagasikara gen. et sp. nov., nymph morphology a tergite IV (posterior margin) b paraproct c gill IV (with enlarged margins). Scale bar: 0.1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure6" httpUri="https://binary.pensoft.net/fig/610397" pageId="0" pageNumber="465">6b</figureCitation>
). With slight, rounded expansion at posterior margin; with numerous, stout, marginal spines.
</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" italics="true" pageId="0" pageNumber="465">Caudal filaments</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Megalabiops madagasikara gen. et sp. nov., habitus, nymph a dorsal view b ventral view c, d lateral view. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.62.73911.figure1" httpUri="https://binary.pensoft.net/fig/610392" pageId="0" pageNumber="465">1</figureCitation>
). Inner margin of cerci with 3 to more than 15 long, thin primary swimming setae per segment, increasing in number distally; paracercus bilaterally with 4 to more than 13 long, thin primary swimming setae per segment, increasing in number distally.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="465" type="description">
<paragraph pageId="0" pageNumber="465">Description.</paragraph>
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Imagines.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="465" type="distribution">
<paragraph pageId="0" pageNumber="465">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="465">
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Megalabiops madagasikara gen. et sp. nov., distribution in Madagascar." figureDoi="10.3897/afrinvertebr.62.73911.figure7" httpUri="https://binary.pensoft.net/fig/610398" pageId="0" pageNumber="465">7</figureCitation>
). Madagascar: Eastern coast.
</paragraph>
<caption doi="10.3897/afrinvertebr.62.73911.figure7" httpUri="https://binary.pensoft.net/fig/610398" pageId="0" pageNumber="465" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="465">
<emphasis bold="true" pageId="0" pageNumber="465">Figure 7.</emphasis>
<taxonomicName authorityName="gen. et" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Megalabiops" kingdom="Animalia" lsidName="Megalabiops madagasikara" order="Ephemeroptera" pageId="0" pageNumber="465" rank="species" species="madagasikara">
<emphasis italics="true" pageId="0" pageNumber="465">Megalabiops madagasikara</emphasis>
</taxonomicName>
gen. et sp. nov., distribution in Madagascar.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="F1C3FC930A5ED6231BB60CEC8FCA4529" ID-CoL="4HTQQ" authority="(Gray 1868)" baseAuthorityName="Gray" baseAuthorityYear="1868" class="Mammalia" family="Cercopithecidae" genus="Piliocolobus" kingdom="Animalia" order="Primates" pageNumber="169" phylum="Chordata" rank="species" species="kirkii">
<emphasis id="A8E780F9ABF844B5F3CDB0D38B906DC6" italics="true" pageNumber="169">Piliocolobus kirkii</emphasis>
(Gray 1868)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="30D690B00CBB9CCADE567EE893BE0BC9" pageNumber="169" type="reference_group">
<paragraph id="0E260787A80C6CCB8608C87935A68069" pageNumber="169">
<treatmentCitation id="AC0F92E1290FF92FC9A9FC530017D575" author="Gray" authority="Gray 1868" authorityName="Gray" authorityYear="1868" class="Mammalia" family="Cercopithecidae" genus="Colobus" journal="Proc. Zool. Soc. Lond." kingdom="Animalia" order="Primates" page="180" pageNumber="169" phylum="Chordata" rank="species" species="kirkii" title="Colobus kirkii" volume="1868" year="1868">
<taxonomicName id="2D4B1C4DE7700090A83C2BD351575B9D" authority="Gray 1868" authorityName="Gray" authorityYear="1868" class="Mammalia" family="Cercopithecidae" genus="Colobus" kingdom="Animalia" order="Primates" pageNumber="169" phylum="Chordata" rank="species" species="kirkii">
<emphasis id="9806D5427C0E43C9696E8D79AAEA45BD" italics="true" pageNumber="169">[Colobus] kirkii</emphasis>
Gray 1868
</taxonomicName>
,
<bibCitation id="448E201C64FA02D0A1DE87D456E30B81" author="Gray" journal="Proc. Zool. Soc. Lond." pageNumber="169" pagination="180" title="Colobus kirkii" volume="1868" year="1868">Proc. Zool. Soc. Lond., 1868: 180</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8114A1D8EDA59504AA40E465DE456A64" pageNumber="169" type="materials_examined">
<paragraph id="45A07CAF4C5BBDF5B1FC7BB6D32C3CF2" pageNumber="169">
<emphasis id="FB0F48F286DED0C7E445FAD6DA2B7039" inLineHeading="true" pageNumber="169">Type Locality:</emphasis>
<materialsCitation id="F885A3E31AEE83C809DDB5C03B587D18" ID-GBIF-Occurrence="4867331574" country="Tanzania" pageNumber="169">
<collectingCountry id="D7948A7B0D4E4E478C7E763EFC710FC2" name="Tanzania">Tanzania</collectingCountry>
,
<collectingCountry id="D897CA9E2180DB08609E4E66F49B1D77" name="Tanzania">Zanzibar</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9C0DA68AD076FEDAB75E4E1E67C70A07" pageNumber="169" type="vernacular_names">
<paragraph id="F62774F58E81243D6AFBBB537CB2F26E" pageNumber="169">
<emphasis id="6869F9944E5DC863A8EF183BF3D5B9DC" inLineHeading="true" pageNumber="169">Vernacular Names:</emphasis>
<vernacularName id="C77EEB8F927DAAD91A0C1A0A5EC022C8" language="eng" pageNumber="169">
<collectingCountry id="FDA7335D589DADF8706239B41F384DC0" name="Tanzania">Zanzibar</collectingCountry>
Red Colobus
</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="890B7246191A8770EC183B24412039A4" pageNumber="169" type="distribution">
<paragraph id="465F136BDC96842937E973FFDC3AA672" pageNumber="169">
<emphasis id="64F21C50D614959BFD7ED341042D8019" inLineHeading="true" pageNumber="169">Distribution:</emphasis>
<collectingCountry id="0447D4C5547E0EF0CD0B0A02AE3D8056" name="Tanzania">Zanzibar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0607626660E830CE5A7465C3E5A22E2F" pageNumber="169" type="conservation">
<paragraph id="77090A088FB8C6C0FB98A622F9071F03" pageNumber="169">
<emphasis id="3C6FFE043E74131B6E831B0A3E833901" inLineHeading="true" pageNumber="169">Conservation:</emphasis>
<collectionCode id="FE8BE4A13A9C8D937C107B3A93E3A6EC">CITES</collectionCode>
– Appendix I and
<collectingCountry id="5C09811EB65ED4932823DB444B982907" name="United States of America">U.S.</collectingCountry>
<collectionCode id="FE122EF63E5801F013BDBD771B160A9C" country="Brazil" lsid="urn:lsid:biocol.org:col:14319" name="Universidade de São Paulo" type="Herbarium">ESA</collectionCode>
– Endangered as
<taxonomicName id="496A387A29A83400E6D4A74510ACFE6F" class="Mammalia" family="Cercopithecidae" genus="Procolobus" kingdom="Animalia" order="Primates" phylum="Chordata" rank="subSpecies" species="pennantii" subSpecies="kirki">
<emphasis id="7939E58C2E8C3CC4E475E263442EF7D2" italics="true" pageNumber="169">Procolobus pennantii kirki</emphasis>
</taxonomicName>
;
<collectionCode id="83001E4C88E92FBCF8A005A2F7C9F71D">IUCN</collectionCode>
– Endangered.
</paragraph>
</subSubSection>
<subSubSection id="E561B17004FF244C2BD3A70C665116CB" pageNumber="169" type="discussion">
<paragraph id="BDC24CDA942301762214E5CA2FAA8388" pageNumber="169">
<emphasis id="185598E775589E4CA3F67599BDF514C9" inLineHeading="true" pageNumber="169">Discussion:</emphasis>
Considered a separate species by
<bibRefCitation id="434A16D93C4EA07B8F08C87FFD20883F" author="Dandelot, P." refId="ref73122" refString="Dandelot, P. 1974. Order Primates. Part 3. Pp. 1 - 45, in The mammals of Africa: An identification manual (J. Meester and H. W. Setzer, eds.) [issued 10 Sep 1974]. Smithsonian Institution Press, Washington, D. C., not continuously paginated." year="1974">Dandelot (1974)</bibRefCitation>
and
<bibRefCitation id="8A069680B932B5299BC3679AAA030503" author="Groves, C. P." refId="ref132585" refString="Groves, C. P. 2001 c. Primate taxonomy. Smithsonian Institution Press, Washington, DC, 350 pp." year="2001">
Groves (2001
<emphasis id="CFD59DD598EF13810E37FE8E95462AF6" italics="true" pageNumber="169">c</emphasis>
)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

52
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Taymyr and Severnaya Zemlya, Russia (
<bibRefCitation id="1D281BD371183EBD0319DC8F3ED01682" author="Gagarin, V. G." journalOrPublisher="Zoologicheskii Zhurnal" pageId="0" pageNumber="1165" pagination="323 - 334" title="Free-living nematodes from some water bodies of Taimyr peninsula" volume="75" year="1996">Gagarin 1996</bibRefCitation>
,
<bibRefCitation id="2B4D85F344E337B70B57B6C3382DC5A1" author="Gagarin, V. G." journalOrPublisher="Biologia Vnutrennikh Vod" pageId="0" pageNumber="1165" pagination="32 - 37" title="Review of free-living nematode fauna from fresh waters of the Russian Arctic and Subarctic" volume="2" year="2001">Gagarin 2001b</bibRefCitation>
).
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784
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<taxonomicName LSID="http://species-id.net/wiki/Allotrichoma_simplex" authority="Loew" class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma (Allotrichoma) simplex" order="Diptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="simplex" subGenus="Allotrichoma">Allotrichoma (Allotrichoma) simplex (Loew)</taxonomicName>
Figs 6368
</paragraph>
</subSubSection>
<subSubSection lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="reference_group">
<paragraph pageId="52" pageNumber="53">
<taxonomicName class="Insecta" family="Ephydridae" genus="Discocerina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discocerina simplex" order="Diptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="simplex">Discocerina simplex</taxonomicName>
<bibRefCitation author="Loew, H" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="77" pageNumber="78" pagination="307 - 359" title="Diptera Americae septentrionalis indigena. Centuria prima." volume="5" year="1861">Loew 1861</bibRefCitation>
: 354.
<bibRefCitation pageId="52" pageNumber="53">Osten Sacken 1878</bibRefCitation>
: 201 [Nearctic catalog].
<bibRefCitation author="Aldrich, JM" journalOrPublisher="Smithsonian Institution, Smithsonian Miscellaneous Collections" pageId="76" pageNumber="77" pagination="1 - 680" title="A catalogue of North American Diptera (or two-winged flies)." volume="46" year="1905">Aldrich 1905</bibRefCitation>
: 626 [Nearctic catalog].
<bibRefCitation author="Jones, BJ" journalOrPublisher="University of California Publications in Entomology" pageId="77" pageNumber="78" pagination="153 - 198" title="Catalogue of the Ephydridae, with bibliography and description of new species." volume="1" year="1906">Jones 1906</bibRefCitation>
: 178 [key, list].
</paragraph>
<paragraph lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma simplex" order="Diptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="simplex">Allotrichoma simplex</taxonomicName>
.
<bibRefCitation author="Johnson, CW" journalOrPublisher="Occasional Papers of the Boston Society of Natural History" pageId="77" pageNumber="78" pagination="1 - 326" title="List of the Diptera or two-winged flies. In: Fauna of New England, 15." volume="7" year="1925">Johnson 1925</bibRefCitation>
: 271 [generic combination; list, Massachusetts].
<bibRefCitation author="Cresson, ET Jr" journalOrPublisher="Transactions of the American Entomological Society" pageId="77" pageNumber="78" pagination="101 - 128" title="Synopses of North American Ephydridae (Diptera) I. The subfamily Psilopinae, with descriptions of new species." volume="68" year="1942">Cresson 1942</bibRefCitation>
: 108 [list, general distribution in North America];
<bibRefCitation author="Cresson, ET Jr" journalOrPublisher="Transactions of the American Entomological Society" pageId="77" pageNumber="78" pagination="225 - 260" title="A systematic annotated arrangement of the genera and species of the North American Ephydridae (Diptera). IV. The subfamily Napaeinae." volume="74" year="1949">1949</bibRefCitation>
: 260 [syn
<pageBreakToken pageId="53" pageNumber="54" start="start">onymy</pageBreakToken>
].
<bibRefCitation pageId="53" pageNumber="54">Wirth and Stone 1956</bibRefCitation>
: 467 [key, California].
<bibRefCitation author="Deonier, D. L" journalOrPublisher="Iowa State Journal of Science" pageId="77" pageNumber="78" pagination="103 - 126" title="Keys to the shore flies of Iowa (Diptera, Ephydridae)." volume="39" year="1964">Deonier 1964</bibRefCitation>
: 120 [key, Iowa];
<bibRefCitation author="Deonier, DL" journalOrPublisher="Iowa Academy of Science" pageId="77" pageNumber="78" pagination="496 - 510" title="Ecological observations on Iowa shore flies (Diptera, Ephydridae)." volume="71" year="1965">1965</bibRefCitation>
: 501 [natural history].
<bibRefCitation pageId="53" pageNumber="54">Wirth 1965</bibRefCitation>
: 736 [Nearctic catalog].
<bibRefCitation pageId="53" pageNumber="54">Cole 1969</bibRefCitation>
: 397 [discussion, western United States].
<bibRefCitation pageId="53" pageNumber="54">Runyan and Deonier 1979</bibRefCitation>
: 123-137 [discussion, Figs of head and immature stages].
<bibRefCitation pageId="53" pageNumber="54">Mathis and Zatwarnicki 1995</bibRefCitation>
: 153 [world catalog].
<bibRefCitation pageId="53" pageNumber="54">Mathis and Mathis 2008</bibRefCitation>
: 181-182 [shore flies of Plummers Island].
</paragraph>
<paragraph pageId="53" pageNumber="54">
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma filiforme" order="Diptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="filiforme">Allotrichoma filiforme</taxonomicName>
<bibRefCitation author="Becker, Th" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="76" pageNumber="77" pagination="91 - 276" title="Dipterologische Studien IV. Ephydridae." volume="41" year="1896">Becker 1896</bibRefCitation>
: 123 [Russia.
<normalizedToken originalValue="“Sarepta”">&quot;Sarepta&quot;</normalizedToken>
(= Volgograd-Krasnoarmejsk); LT ♂ (designated by
<bibRefCitation author="Papp, L" journalOrPublisher="Folia Entomologica Hungarica series nova," pageId="78" pageNumber="79" pagination="97 - 104" title="A contribution to the revision of the Palaearctic Ephydridae (Diptera)." volume="32" year="1979">Papp 1979</bibRefCitation>
: 98), ZMHU]; 1905: 189 [Palearctic catalog].
<bibRefCitation author="Krivosheina, MG" journalOrPublisher="Polskie Pismo Entomologiczne" pageId="77" pageNumber="78" pagination="291 - 310" title="Some notes on the Old World Allotrichoma (Diptera: Ephydridae) with description of three new species." volume="66" year="1997">Krivosheina and Zatwarnicki 1997</bibRefCitation>
: 301-303 [revision]. New Synonym
</paragraph>
<paragraph pageId="53" pageNumber="54">
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma trispinum" order="Diptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="trispinum">Allotrichoma trispinum</taxonomicName>
<bibRefCitation author="Becker, Th" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="76" pageNumber="77" pagination="91 - 276" title="Dipterologische Studien IV. Ephydridae." volume="41" year="1896">Becker 1896</bibRefCitation>
: 124 [Poland. &quot;Oderwalde dei Maltsch, Schlesien&quot; [= Malczyce, Lower Silesia]; LT ♂ (designated by
<bibRefCitation author="Papp, L" journalOrPublisher="Folia Entomologica Hungarica series nova," pageId="78" pageNumber="79" pagination="97 - 104" title="A contribution to the revision of the Palaearctic Ephydridae (Diptera)." volume="32" year="1979">Papp 1979</bibRefCitation>
: 99), ZMHU]; 1905: 189 [Palearctic catalog].
<bibRefCitation author="Krivosheina, MG" journalOrPublisher="Polskie Pismo Entomologiczne" pageId="77" pageNumber="78" pagination="291 - 310" title="Some notes on the Old World Allotrichoma (Diptera: Ephydridae) with description of three new species." volume="66" year="1997">Krivosheina and Zatwarnicki 1997</bibRefCitation>
: 301 [synonymy with
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma filiforme" order="Diptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="filiforme">Allotrichoma filiforme</taxonomicName>
Becker]. New Synonym
</paragraph>
<paragraph pageId="53" pageNumber="54">
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma dahli" order="Diptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="dahli">Allotrichoma dahli</taxonomicName>
<bibRefCitation author="Beschovski, VL" journalOrPublisher="Comptes rendus de l'Academie bulgare des Sciences" pageId="77" pageNumber="78" pagination="937 - 939" title="Allotrichoma dahli n. sp. Eine neue Art von Diptera (Ephydridae) aus Bulgarien." volume="19" year="1966 b">Beschovski 1966b</bibRefCitation>
: 937 [Bulgaria. Kap Galata; HT ♂, ZMS].
<bibRefCitation author="Papp, L" journalOrPublisher="Folia Entomologica Hungarica series nova," pageId="78" pageNumber="79" pagination="97 - 104" title="A contribution to the revision of the Palaearctic Ephydridae (Diptera)." volume="32" year="1979">Papp 1979</bibRefCitation>
: 98 [synonymy with
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma filiforme" order="Diptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="filiforme">Allotrichoma filiforme</taxonomicName>
Becker]. New Synonym
</paragraph>
<paragraph pageId="53" pageNumber="54">
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma trispinum" order="Diptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="trispinum">Allotrichoma trispinum</taxonomicName>
of authors, not Becker, 1896 [misidentification].
<bibRefCitation author="Johnson, CW" journalOrPublisher="Occasional Papers of the Boston Society of Natural History" pageId="77" pageNumber="78" pagination="1 - 326" title="List of the Diptera or two-winged flies. In: Fauna of New England, 15." volume="7" year="1925">Johnson 1925</bibRefCitation>
: 271 [list, Connecticut].
<bibRefCitation pageId="53" pageNumber="54">Wirth 1965</bibRefCitation>
: 736 [Nearctic catalog].
<bibRefCitation pageId="53" pageNumber="54">Wirth and Stone 1956</bibRefCitation>
: 467 [key].
<bibRefCitation pageId="53" pageNumber="54">Cole 1969</bibRefCitation>
: 397 [list].
<bibRefCitation pageId="53" pageNumber="54">Mathis and Zatwarnicki 1995</bibRefCitation>
: 154 [world catalog].
</paragraph>
</subSubSection>
<subSubSection lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="description">
<paragraph pageId="53" pageNumber="54">Description.</paragraph>
<paragraph pageId="53" pageNumber="54">This species is distinguished from congeners by the following combination of characters: Small to moderately small shore flies, body length 1.75-2.15 mm. Head: Medial facial carina above facial prominence shallow; labella broad, fleshy, shorter than mediproboscis; clypeus microtomentose, usually gray.</paragraph>
<paragraph pageId="53" pageNumber="54">Thorax: Presutural supra-alar seta present. Pleural area lacking stripes. Wing with costal vein ratio 0.25-0.31; M vein ratio 0.42-0.50. Legs blackish brown to black; midtibia bearing numerous, erect setae along dorsal surface.</paragraph>
<paragraph lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
Abdomen: Male 5th sternal flap in lateral view (Fig. 67) broadly developed to it apex, bearing approximately 15 apical setulae; 5th medial process in lateral view (Fig. 67) narrow, digitiform. Male terminalia (Figs 63-66): Epandrium in posterior view (Fig. 63) rounded dorsally, constricted laterally at base of surstyli; epandrium in lateral view (Fig. 64) somewhat rectangular dorsally with anterior and posterior margins parallel sided, ventral portion with an anterior bulge and a narrow, sharp incision at about level or base of surstylus; cerci in posterior view (Fig. 63) approximate dorsally then extended laterally to form a lyre-like structure that curves laterally than ventrally and medially, apex of cercus bearing 4 short setulae; cercus in lateral view (Fig. 64) shallowly sinuous, nearly parallel sided, apex slightly expanded, bluntly rounded; surstylus (ventral extensions of epandrium) in posterior view (Fig. 63) as relatively wide at base third, thereafter ventrally abruptly narrowed, especially along medial margin, to form an elongate process that is slightly spatulate apically, apical portion bearing several short setulae; surstylus in lateral view (Fig. 64) moderately narrow, slightly expanding with apex obliquely pointed and bearing several short setulae; aedeagus in ventral view (Fig. 65) elongate, narrowly ovate, very slightly tapered apically, in lateral view
<pageBreakToken pageId="54" pageNumber="55" start="start">(</pageBreakToken>
Fig. 66) elongate, nearly parallel sided, apex bluntly rounded; phallapodeme in lateral view (Fig. 66) triangular, keel abruptly tapered to narrow, digitiform apex; gonite in ventral view (Fig. 65) bar-like, slightly wider basally, apex with short, lateral process, apex bearing setula, in lateral view (Fig. 66) wide basally, gently but distinctly arched, tapered to apical point.
</paragraph>
</subSubSection>
<subSubSection pageId="54" pageNumber="55" type="type material">
<paragraph pageId="54" pageNumber="55">Type material.</paragraph>
<paragraph pageId="54" pageNumber="55">
The lectotype female of
<taxonomicName class="Insecta" family="Ephydridae" genus="Discocerina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discocerina simplex" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="simplex">Discocerina simplex</taxonomicName>
Loew, here designated to preserve stability and make more universal the use of this name, is labeled &quot;[United States.] M[arylan]d./Loew Coll./simplex m[ihi]. [handwritten]/Type 11150 [red; number handwritten]/Prop. of Harvard Univ. [yellow]/LECTOTYPE ♀
<taxonomicName class="Insecta" family="Ephydridae" genus="Discocerina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discocerina simplex" order="Diptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="simplex">Discocerina simplex</taxonomicName>
Loew by Mathis &amp; Zatwarnicki 2011 [red].&quot; The lectotype is double mounted (minuten in a block of cork), is in good condition (abdomen removed, dissected, stored in glycerin in an attached microvial), and is deposited in the MCZ (11150).
</paragraph>
</subSubSection>
<subSubSection pageId="54" pageNumber="55" type="type locality">
<paragraph pageId="54" pageNumber="55">Type locality.</paragraph>
<paragraph pageId="54" pageNumber="55">
United States.
<normalizedToken originalValue="“Maryland.”">&quot;Maryland.&quot;</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection lastPageId="57" lastPageNumber="58" pageId="54" pageNumber="55" type="other specimens examined from the new world">
<paragraph pageId="54" pageNumber="55">Other specimens examined from the New World.</paragraph>
<paragraph pageId="54" pageNumber="55">
CANADA. ONTARIO. Fort Erie (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.89">42°53.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-78.925">78°55.5'W</geoCoordinate>
), 18 Apr 1909, M. C. Van Duzee (1♂; CAS).
</paragraph>
<paragraph pageId="54" pageNumber="55">
UNITED STATES. CONNECTICUT. Fairfield: Putnam Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.338333">41°20.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-73.38">73°22.8'W</geoCoordinate>
), 18 Jul 1939, A. L. Melander (1♂; USNM).
</paragraph>
<paragraph pageId="54" pageNumber="55">
DELAWARE. Kent: Woodland Beach (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.331665">39°19.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-75.471664">75°28.3'W</geoCoordinate>
; beach), 18 May 2006, D. and W. N. Mathis (5♂, 3♀; USNM). Sussex: Rehoboth (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.72">38°43.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-75.07667">75°04.6'W</geoCoordinate>
), 25 Jun 1939, A. L. Melander (1♂; ANSP).
</paragraph>
<paragraph pageId="54" pageNumber="55">
DISTRICT OF COLUMBIA. Rock Creek, Milkhouse Ford (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.965">38°57.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.04833">77°02.9'W</geoCoordinate>
), 18 May 2007, D. and W. N. Mathis (1♀; USU).
</paragraph>
<paragraph pageId="54" pageNumber="55">
ILLINOIS. McHenry: Algonquin (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.165">42°09.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-88.295">88°17.7'W</geoCoordinate>
), 1 Jun 1896 (2♂; USNM). Peoria: Peoria (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="40.75">40°45'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-89.615">89°36.9'W</geoCoordinate>
), 28 Aug 1917, J. M. Aldrich (2♂; USNM). Washington: Ashley (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.33">38°19.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-89.191666">89°11.5'W</geoCoordinate>
), 7 Aug 1917 (1♂; USNM).
</paragraph>
<paragraph pageId="54" pageNumber="55">
INDIANA. Porter: Dunes (near Porter;
<geoCoordinate direction="north" orientation="latitude" precision="925" value="41.616665">41°37'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-87.066666">87°04'W</geoCoordinate>
), 19 Jul 1933, A. L. Melander (1♂, 1♀; USNM).
</paragraph>
<paragraph pageId="54" pageNumber="55">
IOWA. Allamakee: Waterville (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="43.208332">43°12.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-91.29667">91°17.8'W</geoCoordinate>
), 2 Aug 1960, D. L. Deonier (1♀; USNM). Boone: Fraser Dam (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.125">42°07.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-93.973335">93°58.4'W</geoCoordinate>
), 4 Sep 1960, D. L. Deonier (2♂; USNM); Ledges State Park (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="41.983334">41°59'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-93.886665">93°53.2'W</geoCoordinate>
), 19
<normalizedToken originalValue="May–">May-</normalizedToken>
9 Oct 1949, 1955, 1958, 1960, D. L. Deonier, J. L. Laffoon (4♂, 4♀; USNM). Dickenson: Gull Point State Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.388332">42°23.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-95.165">95°09.9'W</geoCoordinate>
), 27 Jun 1960, D. L. Deonier (1♀; USNM). Hamilton: Little Wall Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.27">42°16.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-93.636665">93°38.2'W</geoCoordinate>
), 13-14 Jul 1960, D. L. Deonier (3♂; USNM). Louisa: Oakville (6.5 km N,
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.163334">41°09.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-91.13333">91°08'W</geoCoordinate>
), 10 Aug 1980, D. L. Deonier (14♂, 4♀; USNM); Wapello (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.231667">41°13.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-91.11667">91°07'W</geoCoordinate>
), 9 Aug 1960, D. L. Deonier (3♂, 6♀; USNM). Monona: Lewis and Clark State Park (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="42.033333">42°02'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-96.166664">96°10'W</geoCoordinate>
), 6 Jun 1960, D. L. Deonier (1♂, 2♀; USNM). Story: Ames (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.035">42°02.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-93.62">93°37.2'W</geoCoordinate>
), 12 May-3 Oct 1959, D. L. Deonier (5♂, 5♀; USNM); Gilbert (6.5 km E;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.108334">42°06.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-93.61667">93°37'W</geoCoordinate>
), 25
<normalizedToken originalValue="Jun–">Jun-</normalizedToken>
22 Jul 1960, 1961, D. L. Deonier (7♂, 4♀; USNM). Warren: Banner Mine (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.44">41°26.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-93.56333">93°33.8'W</geoCoordinate>
), 7 Aug 1960, D. L. Deonier (1♀; USNM).
</paragraph>
<paragraph lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
MARYLAND. Anne Arundel: Bristol (3 km WSW;
<geoCoordinate direction="north" orientation="latitude" precision="925" value="38.783333">38°47'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-76.666664">76°40'W</geoCoordinate>
; at Jug Bay), 19 Sep 1988, S. F. Larcher, W. E. Steiner (2♂; USNM). Baltimore: Hart-Miller Island (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.251667">39°15.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-76.37334">76°22.4'W</geoCoordinate>
), 31 May 1987, W. E. Steiner, J. M. Swearingen (1♂, 1♀; USNM). Calvert: Calvert Beach (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.465">38°27.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-76.47667">76°28.6'W</geoCoordinate>
), 17 Sep 1989, J. M. Hill, W. E. Steiner, J. M. Swearingen (1♂; USNM); Chesapeake Beach (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.68667">38°41.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-76.535">76°32.1'W</geoCoordinate>
), 18 Aug 1922, J. R. Malloch (1♀; ANSP). Carroll: Eldersburg (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.40333">39°24.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-76.95">76°57'W</geoCoordinate>
), 2 Jun 1985, J. E. Lowry, W. E. Steiner (1♂; USNM). Garrett: Broadford Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.411667">39°24.7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-79.37334">79°22.4'W</geoCoordinate>
; 750 m), 24 Aug 2006, D. and W. N. Mathis (23♂,
<pageBreakToken pageId="55" pageNumber="56" start="start">2</pageBreakToken>
♀; USNM). Montgomery: Clarksburg, Little Bennett Regional Park (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="39.266666">39°16'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.28">77°16.8'W</geoCoordinate>
), 21 Sep 1990, M. J. and R. Molineaux, W. E. Steiner (1♂; USNM); Kensington (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="39.016666">39°01'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.06834">77°04.1'W</geoCoordinate>
), 13 Apr 2006, D. and W. N. Mathis (1♂; USNM); Little Bennett Regional Park (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="39.266666">39°16'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.28">77°16.8'W</geoCoordinate>
), 20 Apr 2005, D. and W. N. Mathis (1♂; USNM); Plummers Island (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.97">38°58.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.17667">77°10.6'W</geoCoordinate>
), 12 Apr 1914, W. L. McAtee (1♂, 1♀; ANSP). Prince George's: (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="38.833332">38°50'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-76.85">76°51'W</geoCoordinate>
), 4 Jul 1954, C. W. Sabrosky (1♂; USNM). Talbot: Wittman (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.793335">38°47.6'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-76.293335">76°17.6'W</geoCoordinate>
), 19 May 1986, W. E. Steiner (1♀; USNM).
</paragraph>
<paragraph pageId="55" pageNumber="56">
MASSACHUSETTS. Barnstable: Woods Hole (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.525">41°31.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-70.67167">70°40.3'W</geoCoordinate>
), 24 Aug 1922, A. H. Sturtevant (1♂; USNM).
</paragraph>
<paragraph pageId="55" pageNumber="56">
MICHIGAN. Monroe: Monroe (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="41.916668">41°55'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.39833">83°23.9'W</geoCoordinate>
), 2 Jul 1939, G. C. Steyskal (1♂, 4♀; USNM). Newaygo: (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="43.42">43°25.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-85.8">85°48'W</geoCoordinate>
), 27 Jun 1953, R. R. Dreisbach (1♂; USNM). Wayne: Detroit (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="42.383335">42°23'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.10167">83°06.1'W</geoCoordinate>
), 17 Sep 1939, G. C. Steyskal (1♂; USNM); Grosse Ile (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.128334">42°07.7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.145">83°08.7'W</geoCoordinate>
), 8 Aug 1954, G. C. Steyskal (2♂; USNM).
</paragraph>
<paragraph pageId="55" pageNumber="56">
MINNESOTA. Becker: Detroit Lake (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="46.816666">46°49'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-95.845">95°50.7'W</geoCoordinate>
), 9 Aug 1935, A. L. Melander (2♂; USNM). Crow Wing: Pequot Lakes (8 km SE;
<geoCoordinate direction="north" orientation="latitude" precision="925" value="46.6">46°36'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-84.28333">84°17'W</geoCoordinate>
), 6 Jul 1957, J. L. Laffoon (1♂; USNM). Goodhue: Lake Pepin (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="44.433334">44°26'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-92.175">92°10.5'W</geoCoordinate>
), 31 May 1951, R. Namba (2♂; USNM). Lake: Basswood Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="48.075">48°04.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-91.575">91°34.5'W</geoCoordinate>
), 17 Aug 1950, R. Namba (1♂, 2♀; USNM). Mille Lacs: Mille Lacs Lake (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="46.233334">46°14'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-93.65">93°39'W</geoCoordinate>
), 2 Jun 1937, D. G. Denning (1♂; USNM).
</paragraph>
<paragraph pageId="55" pageNumber="56">
NEW JERSEY. Burlington: Crowley's Landing (39°
<geoCoordinate direction="north" orientation="latitude" precision="5555" value="37.6">37.6N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-74.62">74°37.2'W</geoCoordinate>
), 17 Jun 2004, D. and W. N. Mathis (3♂, 1♀; USNM). Ocean: Tuckerton (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.603333">39°36.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-74.346664">74°20.8'W</geoCoordinate>
), 17 Jun 2004, D. and W. N. Mathis (19♂, 6♀; USNM).
</paragraph>
<paragraph pageId="55" pageNumber="56">
NEW YORK. Niagara: Niagara Falls (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="43.095">43°05.7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-79.05666">79°03.4'W</geoCoordinate>
), 26 Mar 1911, M. C. Van Duzee (2♂; CAS). Tompkins: Inlet Valley (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.41">42°24.6'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-76.53833">76°32.3'W</geoCoordinate>
), 15 May 1956, B. A. Foote (1♂; USNM). Wyoming: Portageville (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.57">42°34.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-78.04">78°02.4'W</geoCoordinate>
), 13 Jun 1963, W. W. Wirth (1♂; USNM).
</paragraph>
<paragraph lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
OHIO. Ashtabula: Geneva State Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.855">41°51.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-80.971664">80°58.3'W</geoCoordinate>
), 16 Apr 1976, D. L. Deonier (1♂; USNM). Auglaize: St. Marys Lake (= Grand Lake;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="40.521667">40°31.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-84.42167">84°25.3'W</geoCoordinate>
; sedge meadow), 26 May 1977, J. T. Runyan, B. A. Steinly (10♂, 4♀; USNM). Brown: Grant Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.788334">38°47.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.915">83°54.9'W</geoCoordinate>
), 7 Jul 1975, J. Regensburg (1♂, 1♀; USNM). Butler: Hueston Woods State Park (Acton Lake;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.571667">39°34.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-84.74167">84°44.5'W</geoCoordinate>
), 16 Jul 1974, J. Regensburg (12♂, 15♀; USNM); Oxford, Peffer Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.495">39°29.7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-84.73167">84°43.9'W</geoCoordinate>
), 1 Jul 1975, J. Regensburg (1♂; USNM). Champaign: Kiser Lake State Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="40.191666">40°11.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-83.96667">83°58'W</geoCoordinate>
), 5 Aug 1976, B. A. Steinly (4♂, 7♀; USNM). Clermont: Stonelick State Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.226665">39°13.6'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-84.058334">84°03.5'W</geoCoordinate>
), 1 Aug 1974, J. Regensburg (6♂, 5♀; USNM). Delaware: Delaware Wildlife Refuge (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="40.408333">40°24.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.045">83°02.7'W</geoCoordinate>
), 28 Jun 1977, B. A. Steinly (3♂, 7♀; USNM). Fulton: Harrison Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.643333">41°38.6'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-84.371666">84°22.3'W</geoCoordinate>
), 21 Aug 1977, B. A. Steinly (2♂, 5♀; USNM). Hamilton: Whitewater River (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.235">39°14.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-84.785">84°47.1'W</geoCoordinate>
), 5 Aug 1974, J. Regensburg (2♀; USNM). Hocking: Hocking Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.508335">39°30.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-82.45333">82°27.2'W</geoCoordinate>
), 8 Nov 1974, J. Regensburg (3♂, 3♀; USNM). Lake: Mentor marsh (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="41.666668">41°40'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.33833">81°20.3'W</geoCoordinate>
), 16 Jun 1976, D. L. Deonier (1♂; USNM). Lawrence:
<pageBreakToken pageId="56" pageNumber="57" start="start">Vesuvius</pageBreakToken>
Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.58">38°34.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-82.625">82°37.5'W</geoCoordinate>
), 23 Aug 1974, J. Regensburg (1♀; USNM). Lorain: Amherst, Beaver Creek (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="40.488335">40°29.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-84.54833">84°32.9'W</geoCoordinate>
), 22-24 Aug 1977, B. A. Steinly (10♂, 5♀; USNM); Vermillion River, Mill Hollow (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.381668">41°22.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-82.316666">82°19'W</geoCoordinate>
), 22 Sep 1976, B. A. Steinly (1♂; USNM). Lucas: Presque Isle (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.695">41°41.7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.456665">83°27.4'W</geoCoordinate>
), 6 May 1977, B. A. Steinly (2♂, 3♀; USNM). Madison: Madison Lake State Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.515">39°30.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.355">83°21.3'W</geoCoordinate>
), 22 Jul 1974, J. Regensburg (1♂, 1♀; USNM). Meigs: Belleville Dam (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.095">39°05.7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.74834">81°44.9'W</geoCoordinate>
), 20 Aug 1974, J. Regensburg (9♂, 14♀; USNM); Fork Run State Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.065">39°03.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.77167">81°46.3'W</geoCoordinate>
), 27 Aug 1974, J. Regensburg (4♂, 2♀; USNM). Mercer: Montezuma (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="40.488335">40°29.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-84.54833">84°32.9'W</geoCoordinate>
), 26 May 1977, B. A. Steinly (10♂, 7♀; USNM); Montezuma Creek (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="40.493332">40°29.6'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-84.55">84°33'W</geoCoordinate>
), 11 Oct 1976, B. A. Steinly (2♂, 4♀; USNM); Windy Point (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="40.516666">40°31'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-84.53667">84°32.2'W</geoCoordinate>
), 26 May 1977, B. A. Steinly (14♂, 9♀; USNM). Morgan: Burr Oak State Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.555">39°33.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-82.04667">82°02.8'W</geoCoordinate>
), 26 Aug 1974, J. Regensburg (5♂, 2♀; USNM); Muskingum River near Big Bottom State Memorial (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.518333">39°31.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.791664">81°47.5'W</geoCoordinate>
), 26 Aug 1974, J. Regensburg (1♂, 2♀; USNM). Ottawa: South Bass Island (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.643333">41°38.6'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-82.83833">82°50.3'W</geoCoordinate>
), 22 Jul-22 Aug 1977, B. A. Steinly (6♂, 5♀; USNM). Portage: Kent (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.15333">41°09.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.35833">81°21.5'W</geoCoordinate>
), 25 Aug 1969, J. Scheiring (1♂; USNM); Kent (1.6 km E;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.15333">41°09.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-81.35">81°21'W</geoCoordinate>
; dead snail), 2-11 Sep 1972, B. A. Foote (5♂, 1♀; USNM); Kent (5 km E;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.15333">41°09.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-81.333336">81°20'W</geoCoordinate>
; muskrat dung), 10 Sep 1975, B. A. Foote (1♂, 3♀; USNM); Ravenna (5 km E;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="41.158333">41°09.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-81.21667">81°13'W</geoCoordinate>
), 3 May 1969, J. Scheiring (3♂, 5♀; USNM). Preble: Four-Mile Creek, Route 725 (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="39.583332">39°35'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-84.763336">84°45.8'W</geoCoordinate>
), 2 Jun-26 Sep 1978, 1979, B. A. Steinly (14♂, 13♀; USNM). Stark: Berlin Reservoir (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="40.981667">40°58.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-81.1">81°06'W</geoCoordinate>
), 7 Jul 1976, B. A. Steinly (1♂, 3♀; USNM).
</paragraph>
<paragraph pageId="56" pageNumber="57">
TEXAS. Brewster: Big Bend National Park, Boquillas Canyon (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="29.211666">29°12.7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-102.886665">102°53.2'W</geoCoordinate>
), 20 Jun 1953, W. W. Wirth (1♂, 1♀; USNM).
</paragraph>
<paragraph pageId="56" pageNumber="57">
VERMONT. Chittenden: Burlington (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="44.475">44°28.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-73.21167">73°12.7'W</geoCoordinate>
), 1 Sep 1957, A. L. Melander (6♂, 10♀; USNM).
</paragraph>
<paragraph lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
VIRGINIA. Accomack: Assateague Island, near Refuge headquarters (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.908333">37°54.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-75.36">75°21.6'W</geoCoordinate>
), 3 Oct 2005, D. and W. N. Mathis (2♂, 1♀; USNM). Arlington: 4-Mile Run (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.84">38°50.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.045">77°02.7'W</geoCoordinate>
), 28 May 1977, W. N. Mathis (2♂, 3♀; USNM). Culpeper: Lake Pelham (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.463333">38°27.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-78.045">78°02.7'W</geoCoordinate>
), 28 Apr 2006, D. and W. N. Mathis (4♂, 2♀; USNM). Fairfax: Annandale (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.83">38°49.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.19666">77°11.8'W</geoCoordinate>
), 28 Jul 1968, J. W. Adams (1♂; USNM); Great Falls (Clay Pond;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.001667">39°00.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.25667">77°15.4'W</geoCoordinate>
), 13 Jun-17 Aug 2006, W. N. Mathis, T. Zatwarnicki (11♂, 2♀; USNM); Great Falls (Patowmack Canal;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.001667">39°00.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.253334">77°15.2'W</geoCoordinate>
), 5 Jul 2007, D. and W. N. Mathis (1♂; USNM); Great Falls (Potomac River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.003334">39°0.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.253334">77°15.2'W</geoCoordinate>
), 4 Oct 2007, D. and W. N. Mathis (1♂; USNM); Great Falls (quarry;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.985">38°59.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.246666">77°14.8'W</geoCoordinate>
; 50 m), 13 Jun 2007, D. and W. N. Mathis (5♂; USNM); Great Falls (river trail; Potomac River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.985">38°59.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.24333">77°14.6'W</geoCoordinate>
; 60 m), 3 Apr 2007, D. and W. N. Mathis (1♂, 1♀; USNM); Scott Run (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.968334">38°58.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.20333">77°12.2'W</geoCoordinate>
), 7 Jun, 1955, C. W. Sabrosky (2♀; USNM); Turkey Run (mouth;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.965">38°57.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.15667">77°09.4'W</geoCoordinate>
), 22 May-7 Sep 2006, 2007, W. N. Mathis, T. Zatwarnicki (65♂, 9♀; USNM). Giles: Mountain Lake Bio Station (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.375">37°22.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-80.52167">80°31.3'W</geoCoordinate>
; 1176 m), 23 Sep 2007, D. and W. N. Mathis (1♂; USNM). Grayson: Galax (15 km S; Chestnut Creek;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="36.581665">36°34.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-80.89167">80°53.5'W</geoCoordinate>
), 28 Jun 1986, W. E. Steiner (1♂; USNM).
<pageBreakToken pageId="57" pageNumber="58" start="start">Henry</pageBreakToken>
: Martinsville Reservoir (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="36.745">36°44.7'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-79.87">79°52.2'W</geoCoordinate>
), 17 May 2005, D. and W. N. Mathis (13♂, 1♀; USNM). Loudoun: Lucketts (6 km N,
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.271667">39°16.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.54833">77°32.9'W</geoCoordinate>
), 15 Apr 2005, D. and W. N. Mathis (1♂; USNM). Louisa: Lake Anna (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.09667">38°05.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.89333">77°53.6'W</geoCoordinate>
; Christopher Run Campground), 4 May 2007, D. and W. N. Mathis (1♀; USNM). Patrick: Woolwine (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="36.79">36°47.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-80.278336">80°16.7'W</geoCoordinate>
; 300 m), 17 May 2005, D. and W. N. Mathis (4♂, 1♀; USNM). Rappahannock: Hazel River (NW Culpeper;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.56333">38°33.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-78.19334">78°11.6'W</geoCoordinate>
, 171 m), 28 Jun 2008, W. N. Mathis and T. Zatwarnicki (1♂; USNM). Roanoke: Salem (Roanoke River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.268333">37°16.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-80.03667">80°02.2'W</geoCoordinate>
; 300 m), 23 Sep 2007, D. and W. N. Mathis (5♂, 1♀; USNM). Smyth: Saltville (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="36.871666">36°52.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.77333">81°46.4'W</geoCoordinate>
, 1710 ft), 22 Jun 2007, D. and W. N. Mathis (1♀; USNM). Spotsylvania: Rappahannock River (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.31333">38°18.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.541664">77°32.5'W</geoCoordinate>
), 15 Apr-10 Oct 2006, 2007, D. and W. N. Mathis (21♂, 7♀; USNM). Stafford: Aquia Harbour (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.465">38°27.9'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.388336">77°23.3'W</geoCoordinate>
), 15 May 2000, D. and W. N. Mathis (2♂; USNM); Aquia Harbour, Aquia Creek (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.463333">38°27.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.385">77°23.1'W</geoCoordinate>
), 12 Apr-2 Sep 2006, D. and W. N. Mathis (2♂, 1♀; USNM); Aquia Harbour, Lions Park (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="38.45">38°27'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.388336">77°23.3'W</geoCoordinate>
), 10 Mar-18 Dec 2004, 2005, 2006, 2007, D. and W. N. Mathis (27♂, 14♀; USNM); Aquia Landing (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.386665">38°23.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-77.316666">77°19'W</geoCoordinate>
), 7 May-8 Sep 2003, 2005, D. and W. N. Mathis (12♂, 4♀; USNM); Curtis Park Lake (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="38.433334">38°26'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.555">77°33.3'W</geoCoordinate>
), 30 Mar-2 Jul 2005, 2007, D. Mathis (16♂, 4♀; USNM); Falmouth (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.32">38°19.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.46833">77°28.1'W</geoCoordinate>
; Rappahannock River; 9 m), 11 Apr-30 Jun 2007, 2008, D. and W. N. Mathis (23♂, 2♀; USNM). York: Maury Lake (ca. James River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.041668">37°02.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-76.486664">76°29.2'W</geoCoordinate>
), 22 Mar-19 Aug 2006, 2007, D. and W. N. Mathis (1♂, 1♀; USNM). Westmoreland: Potomac Beach (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.273335">38°16.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-76.99333">76°59.6'W</geoCoordinate>
; Potomac River), 8 Jun 2007, D. and W. N. Mathis (1♀; USNM). Independent City: Fredericksburg (Rappahannock River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.305">38°18.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.458336">77°27.5'W</geoCoordinate>
), 14 Apr 2006, D. and W. N. Mathis (1♀; USNM).
</paragraph>
<paragraph pageId="57" pageNumber="58">
WEST VIRGINIA. Cavell: Culloden (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.42">38°25.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-82.055">82°03.3'W</geoCoordinate>
), Jul 1954, M. R. Wheeler (1♂; USNM). Hardy: Baker (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.041668">39°02.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-78.74834">78°44.9'W</geoCoordinate>
; 405 m), 20 Jun-13 Jul 2007, D. and W. N. Mathis (13♂; USNM); Lost City (1.6 km N,
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.941666">38°56.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-78.815">78°48.9'W</geoCoordinate>
; 460 m), 12 Jul 2007, D. and W. N. Mathis (1♀; USNM); Mathias (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.876667">38°52.6'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-78.86667">78°52'W</geoCoordinate>
; 465 m), 13 Jul 2007, D. and W. N. Mathis (1♂; USNM); Trout Pond (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.956665">38°57.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-78.736664">78°44.2'W</geoCoordinate>
; 595 m), 13 Jul 2007, D. and W. N. Mathis (1♂; USNM). Mercer: Ceres (Kee Reservoir;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.306667">37°18.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.17333">81°10.4'W</geoCoordinate>
; 757 m), 24 Sep 2007, D. and W. N. Mathis (5♂; USNM). Morgan: Great Cacapon (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.62">39°37.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-78.293335">78°17.6'W</geoCoordinate>
), 3 Jul 1977, W. W. Wirth (1♂; USNM). Perkins: Silver Lake (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.255">39°15.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-79.496666">79°29.8'W</geoCoordinate>
; 762 m), 21 Jun 2007, D. and W. N. Mathis (3♂; USNM). Pocahontas: Marlinton (Greenbrier River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.225">38°13.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-80.095">80°05.7'W</geoCoordinate>
), 21 Jun 2007, D. and W. N. Mathis (3♂; USNM); Marlinton, Knapp Creek (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="38.216667">38°13'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-80.1">80°06'W</geoCoordinate>
), 29 Jun 1982, W. N. Mathis (2♀; USNM). Richie: North Bend State Park (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.22333">39°13.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.11">81°06.6'W</geoCoordinate>
), 23 Jun 1970, G. C. Steyskal (1♂♀; USNM). Roanoke: Salem (Roanoke River;
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.268333">37°16.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-80.03667">80°02.2'W</geoCoordinate>
; 300 m), 23 Sep 2007, D. and W. N. Mathis (4♂, 1♀; USNM). Summers: Hinton (
<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.696667">37°41.8'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-80.88333">80°53'W</geoCoordinate>
; New River; 427 m), 26 Sep 2007, D. and W. N. Mathis (1♂, 2♀; USNM).
</paragraph>
</subSubSection>
<subSubSection lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="distribution">
<paragraph pageId="57" pageNumber="58">Distribution.</paragraph>
<paragraph lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
(Fig. 68) Nearctic: Canada (Ontario), United States (Connecticut, Delaware, District of Columbia, Illinois, Indiana, Iowa, Maryland, Massachusetts,
<pageBreakToken pageId="58" pageNumber="59" start="start">Michigan</pageBreakToken>
, Minnesota, New Jersey, New York, Ohio, Texas, Vermont, Virginia, West Virginia). Palearctic: Austria, Bulgaria, China (Tibet), Czech Republic, France, Hungary, Israel, Italy, Morocco, Poland, Spain, Russia (European Territory), Switzerland, Yugoslavia.
</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="natural history">
<paragraph pageId="58" pageNumber="59">Natural history.</paragraph>
<paragraph pageId="58" pageNumber="59">
<bibRefCitation author="Deonier, DL" journalOrPublisher="Iowa Academy of Science" pageId="77" pageNumber="78" pagination="496 - 510" title="Ecological observations on Iowa shore flies (Diptera, Ephydridae)." volume="71" year="1965">Deonier (1965</bibRefCitation>
: 501) found this species to be
<normalizedToken originalValue="“occasional”">&quot;occasional&quot;</normalizedToken>
in mud shore and sand shore habitats but rare in sedge meadow and
<taxonomicName class="Liliopsida" family="Poaceae" genus="Eragrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eragrostis" order="Poales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="genus">Eragrostis</taxonomicName>
mat habitats.
<bibRefCitation pageId="58" pageNumber="59">Runyan and Deonier (1979)</bibRefCitation>
produced the only illustrations of immature stages, including an egg, the cephalopharyngeal skeleton of the third instar larva, and the puparium.
</paragraph>
</subSubSection>
<subSubSection pageId="58" pageNumber="59" type="remarks">
<paragraph pageId="58" pageNumber="59">Remarks.</paragraph>
<paragraph pageId="58" pageNumber="59">
This species has a Holarctic distribution, and Palearctic specimens have been named at least three different times, with
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma simplex" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="simplex">Allotrichoma simplex</taxonomicName>
being the senior synonym. The junior synonyms in chronological order are:
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<bibRefCitation author="Becker, Th" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="76" pageNumber="77" pagination="91 - 276" title="Dipterologische Studien IV. Ephydridae." volume="41" year="1896">Becker (1896)</bibRefCitation>
,
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma trispinum" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="trispinum">Allotrichoma trispinum</taxonomicName>
<bibRefCitation author="Becker, Th" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="76" pageNumber="77" pagination="91 - 276" title="Dipterologische Studien IV. Ephydridae." volume="41" year="1896">Becker (1896)</bibRefCitation>
, and
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma dahli" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="dahli">Allotrichoma dahli</taxonomicName>
<bibRefCitation author="Beschovski, VL" journalOrPublisher="Comptes rendus de l'Academie bulgare des Sciences" pageId="77" pageNumber="78" pagination="851 - 853" title="Allotrichoma valkanovi n. sp. Eine neue Art von (Diptera Ephydridae) aus Bulgarien." volume="19" year="1966 a">Beschovski (1966a)</bibRefCitation>
. In Europe, there is also a very similar species,
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma strandi" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="strandi">Allotrichoma strandi</taxonomicName>
Duda, that differs only slightly but apparently consistently from this species. In males of
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma simplex" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="simplex">Allotrichoma simplex</taxonomicName>
, the cercus has four apical setulae, compared to three in
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma strandi" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="strandi">Allotrichoma strandi</taxonomicName>
, and the mesonotal stripes are more evident in
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma strandi" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="strandi">Allotrichoma strandi</taxonomicName>
than in
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma simplex" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="simplex">Allotrichoma simplex</taxonomicName>
.
</paragraph>
<paragraph pageId="58" pageNumber="59">In the Nearctic Region, this is a common species that may have the most widespread distribution.</paragraph>
<caption pageId="58" pageNumber="59">
<paragraph pageId="58" pageNumber="59">
Figures 63-67. Illustration of
<taxonomicName class="Insecta" family="Ephydridae" genus="Allotrichoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allotrichoma simplex" order="Diptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="simplex">Allotrichoma simplex</taxonomicName>
(Loew) (male) 63 epandrium, cerci, surstylus, posterior aspect 64 same, lateral aspect 65 aedeagus, phallapodeme, gonite, hypandrium, ventral aspect 66 same, lateral aspect 67 male 5th sternal flap and medial process, lateral aspect. Scale bar = 0.1 mm.
</paragraph>
</caption>
<caption pageId="58" pageNumber="59">
<paragraph pageId="58" pageNumber="59">
Figure 68. Distribution of
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(Loew).
</paragraph>
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Yalanzou, une ouvriere. Seul type arboricole du genre. La
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Ashmead, 1893
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Ashmead, 1893
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Ashmead, 1896
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(Nagamori, 1925,
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.327.5908" ID-GBIF-Dataset="a14e4da5-3053-4f01-b3a9-9a1e0092a3a3" ID-PMC="PMC3804766" ID-Pensoft-Pub="1313-2970-327-65" ID-PubMed="24167422" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-327-65" ModsDocOrigin="ZooKeys 327" ModsDocTitle="Review of Canadian species of the genus Dinaraea Thomson, with descriptions of six new species (Coleoptera, Staphylinidae, Aleocharinae, Athetini)" checkinTime="1451246991397" checkinUser="pensoft" docAuthor="Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Caroline Bourdon, &amp; Jacobs, Jenna" docDate="2013" docId="E808ADA36F6924BB8898D0E26161A1E0" docLanguage="en" docName="ZooKeys 327: 65-101" docOrigin="ZooKeys 327" docSource="http://dx.doi.org/10.3897/zookeys.327.5908" docTitle="Dinaraea borealis Lohse" docType="treatment" docVersion="3" lastPageNumber="87" masterDocId="B741FF8DFFA79653E35CFFD3FF852475" masterDocTitle="Review of Canadian species of the genus Dinaraea Thomson, with descriptions of six new species (Coleoptera, Staphylinidae, Aleocharinae, Athetini)" masterLastPageNumber="101" masterPageNumber="65" pageNumber="86" updateTime="1668156376624" updateUser="ExternalLinkService">
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<mods:title>Review of Canadian species of the genus Dinaraea Thomson, with descriptions of six new species (Coleoptera, Staphylinidae, Aleocharinae, Athetini)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Klimaszewski, Jan</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Webster, Reginald P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Langor, David W.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Caroline Bourdon,</mods:namePart>
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<mods:namePart>Jacobs, Jenna</mods:namePart>
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<mods:date>2013</mods:date>
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<mods:number>327</mods:number>
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<treatment ID-GBIF-Taxon="152047406" LSID="urn:lsid:plazi:treatment:E808ADA36F6924BB8898D0E26161A1E0" httpUri="http://treatment.plazi.org/id/E808ADA36F6924BB8898D0E26161A1E0" lastPageId="23" lastPageNumber="87" pageId="21" pageNumber="86">
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<paragraph pageId="21" pageNumber="86">
8.
<taxonomicName LSID="http://species-id.net/wiki/Dinaraea_borealis" authority="Lohse" class="Insecta" family="Staphylinidae" genus="Dinaraea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dinaraea borealis" order="Coleoptera" pageId="21" pageNumber="86" phylum="Arthropoda" rank="species" species="borealis">Dinaraea borealis Lohse</taxonomicName>
Fig. 8
<normalizedToken originalValue="a–g">a-g</normalizedToken>
, 12, Map 8
</paragraph>
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<subSubSection pageId="21" pageNumber="86" type="reference_group">
<paragraph pageId="21" pageNumber="86">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dinaraea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dinaraea borealis" order="Coleoptera" pageId="21" pageNumber="86" phylum="Arthropoda" rank="species" species="borealis">Dinaraea borealis</taxonomicName>
Lohse, in
<bibRefCitation author="Lohse, GA" journalOrPublisher="The Coleopterists Bulletin" pageId="29" pageNumber="94" pagination="121 - 202" title="Revision of arctic Aleocharinae of North America (Coleoptera: Staphylinidae)." volume="44" year="1990">Lohse et al. 1990</bibRefCitation>
: 198 (based on a single female description).
</paragraph>
<paragraph pageId="21" pageNumber="86">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dinaraea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dinaraea borealis" order="Coleoptera" pageId="21" pageNumber="86" phylum="Arthropoda" rank="species" species="borealis">Dinaraea borealis</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="86" type="holotype">
<paragraph pageId="21" pageNumber="86">HOLOTYPE</paragraph>
<paragraph pageId="21" pageNumber="86">(female): CANADA, QUEBEC, Gt. Whale River, 30.VI.1949, J.R. Vockeroth, No. 20341 (CNC). Holotype examined.</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="87" pageId="21" pageNumber="86" type="diagnosis">
<paragraph pageId="21" pageNumber="86">Diagnosis.</paragraph>
<paragraph pageId="22" pageNumber="87">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dinaraea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dinaraea borealis" order="Coleoptera" pageId="22" pageNumber="87" phylum="Arthropoda" rank="species" species="borealis">
<pageBreakToken pageId="22" pageNumber="87" start="start">Dinaraea</pageBreakToken>
borealis
</taxonomicName>
(habitus Fig. 8a) may be distinguished from congeners by the following combination of characters: body length 2.8-3.0 mm; head, pronotum and elytra slightly glossy with dense microsculpture; pronotum broadest in apical third and narrowest at base; elytra at suture as long as pronotum, with dense punctation similar to that on pronotum; antennal articles 7-10 moderately transverse; male tergite VIII without apical teeth (Fig. 8c); median lobe of aedeagus with straight, long tubus pointed at apex and slightly pointed ventrally (Fig. 8b); spermatheca elongate with pear-shaped capsule, and long apical invagination, stem long and looped posteriorly, with strongly swollen apical part (Fig. 8e).
</paragraph>
<caption pageId="22" pageNumber="87">
<paragraph pageId="22" pageNumber="87">
Figure 8.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dinaraea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dinaraea borealis" order="Coleoptera" pageId="22" pageNumber="87" phylum="Arthropoda" rank="species" species="borealis">Dinaraea borealis</taxonomicName>
Lohse: a habitus b median lobe of aedeagus in lateral view c male tergite VIII d male sternite VIII e spermatheca in lateral view f female tergite VIII g female sternite VIII. Habitus scale bar = 1.0 mm, other scale bars = 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="22" pageNumber="87" type="description">
<paragraph pageId="22" pageNumber="87">Description.</paragraph>
<paragraph pageId="22" pageNumber="87">Body length 2.8-3.0 mm; body variable in colour, either entirely black with brown or reddish appendages and part of elytra, or dark brown with legs, antennae (at least basally), and labial palpi appearing reddish-brown; head, pronotum and elytra slightly glossy, the latter more so, with dense microsculpture; abdominal microsculpture less dense and integument more glossy than that of pronotum and elytra; head moderately large, as broad as pronotum, genae slightly longer than eyes in dorsal view; pronotum broadest in apical third, slightly transverse, usually as long as elytra at suture; elytra transverse, truncate posteriorly; abdomen arcuate laterally. MALE (new description): tergite VIII truncate apically and without apical teeth (Fig. 8c); sternite VIII rounded apically (Fig. 8d); median lobe of aedeagus with long and straight tubus with apex pointed ventrally (Fig. 8b). FEMALE: tergite VIII concave basally and truncate apically (Fig. 8f); sternite VIII rounded apically, antecostal suture strongly sinuate (Fig. 8g); spermatheca elongate with pear-shaped capsule, and long apical invagination, stem long and looped posteriorly, with strongly swollen apical part (Figs 8e, 12).</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="87" type="distribution">
<paragraph pageId="22" pageNumber="87">Distribution.</paragraph>
<paragraph pageId="22" pageNumber="87">
This species was previously recorded only from Gt. Whale River in Quebec (
<bibRefCitation author="Lohse, GA" journalOrPublisher="The Coleopterists Bulletin" pageId="29" pageNumber="94" pagination="121 - 202" title="Revision of arctic Aleocharinae of North America (Coleoptera: Staphylinidae)." volume="44" year="1990">Lohse et al. 1990</bibRefCitation>
). Here, we provide new distribution data for New Brunswick, Quebec, and Ontario.
</paragraph>
<caption pageId="22" pageNumber="87">
<paragraph pageId="22" pageNumber="87">
Map 7-10. Distribution map of: 7
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dinaraea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dinaraea piceana" order="Coleoptera" pageId="22" pageNumber="87" phylum="Arthropoda" rank="species" species="piceana">Dinaraea piceana</taxonomicName>
sp. n. 8
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dinaraea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dinaraea borealis" order="Coleoptera" pageId="22" pageNumber="87" phylum="Arthropoda" rank="species" species="borealis">Dinaraea borealis</taxonomicName>
9
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dinaraea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dinaraea curtipenis" order="Coleoptera" pageId="22" pageNumber="87" phylum="Arthropoda" rank="species" species="curtipenis">Dinaraea curtipenis</taxonomicName>
sp. n. 10
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dinaraea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dinaraea longipenis" order="Coleoptera" pageId="22" pageNumber="87" phylum="Arthropoda" rank="species" species="longipenis">Dinaraea longipenis</taxonomicName>
sp. n.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="22" pageNumber="87" type="collection and habitat data">
<paragraph pageId="22" pageNumber="87">Collection and habitat data.</paragraph>
<paragraph pageId="22" pageNumber="87">Adults were collected from March to August from: under bark of large fallen spruce in old-growth eastern white cedar forests; leaf litter under alders near a small stream in a mixed forest; and moss in a black spruce forest. Specimens were also captured in Lindgren funnel traps in an old-growth eastern white cedar forest and an old-growth white spruce and balsam fir forest.</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="88" pageId="22" pageNumber="87" type="material examined">
<paragraph pageId="22" pageNumber="87">Material examined.</paragraph>
<paragraph lastPageId="23" lastPageNumber="88" pageId="22" pageNumber="87">
CANADA, NEW BRUNSWICK: Charlotte Co., 10 km NW of New River Beach,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.211">45.2110°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.617">66.6170°W</geoCoordinate>
, 15-29.VI.2010, R. Webster &amp; C. MacKay // Old-growth eastern white cedar forest, Lindgren funnel trap (LFC) 1 female; Restigouche Co., Little Tobique River near Red Brook,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="47.44616">47.44616°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-67.06888">67.06888°W</geoCoordinate>
, 24.V.2007, R.P. Webster // Old-growth eastern white cedar forest, under bark of large fallen spruce (LFC, RWC) 2 males, 2 females; Restigouche Co., MacFarlane Brook Protected Area,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="47.6018">47.6018°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-67.6263">67.6263°W</geoCoordinate>
, 25.V.2007, R.P. Webster // Old-growth eastern white cedar swamp, under bark of large fallen spruce (RWC) 1 male, 2 females; Restigouche Co., Dionne Brook P.N.A.,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="47.9064">47.9064°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-68.3441">68.3441°W</geoCoordinate>
, 31.
<normalizedToken originalValue="V–">V-</normalizedToken>
15.VI.2011, M. Roy &amp; V. Webster // Old-growth white spruce and balsam fir forest, Lindgren funnel trap (RWC) 3 males, 2 females; York Co. New Maryland, Charters Settlement,
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.8395">45.8395°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.7391">66.7391°W</geoCoordinate>
, 29.III.2006, R.P. Webster // Mixed forest, under alders near small stream, in leaf litter (RWC) 1 female. QUEBEC: MRC Manic,
<normalizedToken originalValue="Réservoir">Reservoir</normalizedToken>
Outardes 4,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="50.6">50.60°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-69.37">69.37°W</geoCoordinate>
, 30.
<normalizedToken originalValue="VII–">VII-</normalizedToken>
07.VIII.2007, Chaire
<normalizedToken originalValue="Côte-Nord">Cote-Nord</normalizedToken>
, J.P.
<normalizedToken originalValue="Légaré">Legare</normalizedToken>
,
<pageBreakToken pageId="23" pageNumber="88" start="start">Block</pageBreakToken>
Abitibi Nord,
<normalizedToken originalValue="Témoin">Temoin</normalizedToken>
1-10 m,
<normalizedToken originalValue="Piège">Piege</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
impact, 2007-3-1306 (LFC) 1 male; MRC Manic, Lac
<normalizedToken originalValue="Lacoursière">Lacoursiere</normalizedToken>
,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="51.27">51.27°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-67.99">67.99°W</geoCoordinate>
, 18.
<normalizedToken originalValue="VI–">VI-</normalizedToken>
26.VI.2007, Chaire
<normalizedToken originalValue="Côte-Nord">Cote-Nord</normalizedToken>
, J.P.
<normalizedToken originalValue="Légaré">Legare</normalizedToken>
, Bloc Arbec,
<normalizedToken originalValue="Témoin">Temoin</normalizedToken>
3-10 m,
<normalizedToken originalValue="Piège">Piege</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
impact, 2007-3-0168, CPRS 4-15 m, 2007-3-01153 (LFC) 2 females; same data except: 30.VII-07.VIII.2007, CPRS 3-10 m, 2007-3-1352 (LFC) 1 female; Lac-St-Jean, Compagnie
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
Arbec,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="50.381664">50°22'54&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-70.55806">70°33'29&quot;W</geoCoordinate>
, 3.
<normalizedToken originalValue="VI–">VI-</normalizedToken>
16.VI.2009, Annelage 2009, C.
<normalizedToken originalValue="Hébert">Hebert</normalizedToken>
,
<normalizedToken originalValue="pessière">pessiere</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
mousses (LFC) 1 female; Lac-St-Jean, Compagnie
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
Arbec,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="50.35611">50°21'22&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-70.52139">70°31'17&quot;W</geoCoordinate>
, 4.
<normalizedToken originalValue="VI–">VI-</normalizedToken>
18.VI.2009, Annelage 2009, C.
<normalizedToken originalValue="Hébert">Hebert</normalizedToken>
,
<normalizedToken originalValue="pessière">pessiere</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
mousses (LFC) 2 males; Lac-St-Jean, Compagnie
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
Arbec,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="50.381664">50°22'54&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-70.55806">70°33'29&quot;W</geoCoordinate>
, 16.
<normalizedToken originalValue="VI–">VI-</normalizedToken>
01.VII.2009, Annelage 2009, C.
<normalizedToken originalValue="Hébert">Hebert</normalizedToken>
,
<normalizedToken originalValue="pessière">pessiere</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
mousses (LFC) 1 male, 2 females; Compagnie
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
Arbec,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="50.37694">50°22'37&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-70.55222">70°33'08&quot;W</geoCoordinate>
, 17.
<normalizedToken originalValue="VI–">VI-</normalizedToken>
01.VII.2009, Annelage 2009, C.
<normalizedToken originalValue="Hébert">Hebert</normalizedToken>
,
<normalizedToken originalValue="pessière">pessiere</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
mousses (LFC) 3 males, 2 females; Lac-St-Jean, Compagnie
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
Arbec,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="50.35611">50°21'22&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-70.52139">70°31'17&quot;W</geoCoordinate>
, 18.
<normalizedToken originalValue="VI–">VI-</normalizedToken>
02.VII.2009, Annelage 2009, C.
<normalizedToken originalValue="Hébert">Hebert</normalizedToken>
,
<normalizedToken originalValue="pessière">pessiere</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
mousses (LFC) 1 female. ONTARIO: Moosonee, 2.VII.1973, Parry &amp; Campbell (CNC) 1 female.
</paragraph>
</subSubSection>
</treatment>
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102
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<taxonomicName class="Arachnida" lsidName="" pageId="0" pageNumber="977" rank="class">Arachnida</taxonomicName>
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<taxonomicName kingdom="Animalia" lsidName="" pageId="0" pageNumber="977" rank="kingdom">Animalia</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="977" type="nomenclature">
<paragraph pageId="0" pageNumber="977">
<taxonomicName authority="Panzer, 1801" authorityName="Panzer" authorityYear="1801" class="Arachnida" family="Theridiidae" genus="Asagena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Asagena phalerata" order="Araneae" pageId="0" pageNumber="977" phylum="Arthropoda" rank="species" species="phalerata">Asagena phalerata (Panzer, 1801)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="977" type="distribution">
<paragraph pageId="0" pageNumber="977">Distribution</paragraph>
<paragraph pageId="0" pageNumber="977">Palearctic.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="977" type="notes">
<paragraph pageId="0" pageNumber="977">Notes</paragraph>
<paragraph pageId="0" pageNumber="977">
Previously recorded from unspecified locality between Resen and Ohrid (
<bibRefCitation author="Drensky, P." journalOrPublisher="Spisanie na Bulgarska Akademiya na Naukite" pageId="0" pageNumber="977" pagination="1 - 76" title="Spinnen (Araneae) aus Mittel und Sued-West Mazedonien" volume="39" year="1929">Drensky 1929</bibRefCitation>
,
<bibRefCitation author="Drensky, P" journalOrPublisher="Spisanie na Bulgarska Akademiya na Naukite" pageId="0" pageNumber="977" pagination="1 - 223" title="Katalog der echten Spinnen (Araneae) der Balkanhalbinsel" volume="32" year="1936">Drensky 1936</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

151
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<treatment id="E80987C6FFA2F131FF32FF2BFEBFFB70" ID-DOI="http://doi.org/10.5281/zenodo.6134309" ID-GBIF-Taxon="119420379" ID-Zenodo-Dep="6134309" LSID="urn:lsid:plazi:treatment:E80987C6FFA2F131FF32FF2BFEBFFB70" httpUri="http://treatment.plazi.org/id/E80987C6FFA2F131FF32FF2BFEBFFB70" lastPageNumber="172" pageId="1" pageNumber="172">
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<paragraph id="601F36D0FFA2F131FF32FF2BFE23FF15" blockId="1.[151,707,220,280]" box="[151,450,220,247]" pageId="1" pageNumber="172">
<heading id="3B5781BCFFA2F131FF32FF2BFE23FF15" bold="true" box="[151,450,220,247]" fontSize="11" level="1" pageId="1" pageNumber="172" reason="1">
<taxonomicName id="A7A04D53FFA2F131FF32FF2BFE23FF15" ID-CoL="75ca6120-d519-4b4c-a8c4-7fe4ab501c56" authority="Bruce, 1971" authorityName="Bruce" authorityYear="1971" box="[151,450,220,247]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="52D4EAC2FFA2F131FF32FF2BFE23FF15" bold="true" box="[151,450,220,247]" pageId="1" pageNumber="172">
<emphasis id="52D4EAC2FFA2F131FF32FF2BFEC7FF14" bold="true" box="[151,294,220,246]" italics="true" pageId="1" pageNumber="172">Eupontonia</emphasis>
<bibRefCitation id="04314B21FFA2F131FE88FF2AFE23FF15" author="Bruce" box="[301,450,221,247]" pageId="1" pageNumber="172" refString="Bruce, A. J. (1971) Notes on some Indo-Pacific Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp. nov., a new pontoniinid shrimp from Mahe, the Seychelle Islands. Crustaceana, 20, 225 - 236. http: // dx. doi. org / 10.1163 / 156854071 x 00012" type="journal article" year="1971">Bruce, 1971</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="601F36D0FFA2F131FF32FEF7FD22FEFA" blockId="1.[151,707,220,280]" box="[151,707,255,280]" pageId="1" pageNumber="172">[New Japanese name: Awamori-kakure-ebi-zoku]</paragraph>
</subSubSection>
<subSubSection id="28BA655BFFA2F131FF32FEBEFA7DFC74" pageId="1" pageNumber="172" type="reference_group">
<paragraph id="601F36D0FFA2F131FF32FEBEFE5EFE9F" blockId="1.[151,1435,328,381]" pageId="1" pageNumber="172">
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<taxonomicName id="A7A04D53FFA2F131FF32FEBEFE58FEBC" ID-CoL="75ca6120-d519-4b4c-a8c4-7fe4ab501c56" authority="Bruce, 1971: 15" authorityName="Bruce" authorityPageNumber="15" authorityYear="1971" box="[151,441,328,350]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="52D4EAC2FFA2F131FF32FEBEFEECFEBC" box="[151,269,329,350]" italics="true" pageId="1" pageNumber="172">Eupontonia</emphasis>
<treatmentCitation id="E10110C1FFA2F131FEB1FEBFFE58FEBC" author="Bruce" box="[276,441,328,350]" page="15" pageId="1" pageNumber="172" year="1971">
<bibRefCitation id="04314B21FFA2F131FEB1FEBFFE72FEBF" author="Bruce" box="[276,403,328,349]" pageId="1" pageNumber="172" refString="Bruce, A. J. (1971) Notes on some Indo-Pacific Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp. nov., a new pontoniinid shrimp from Mahe, the Seychelle Islands. Crustaceana, 20, 225 - 236. http: // dx. doi. org / 10.1163 / 156854071 x 00012" type="journal article" year="1971">Bruce, 1971</bibRefCitation>
: 15
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; 1995; 2010: 413 (key).—
<treatmentCitation id="E10110C1FFA2F131FD61FEBFFC6BFEBF" author="Holthuis" box="[708,906,328,350]" page="118" pageId="1" pageNumber="172" year="1993">
<bibRefCitation id="04314B21FFA2F131FD61FEBFFCBBFEBF" author="Holthuis" box="[708,858,328,350]" pageId="1" pageNumber="172" refString="Holthuis, L. B. (1993) The Recent Genera of the Caridean and Stenopodidean Shrimps (Crustacea, Decapoda) with an Appendix on the Order Amphionidacea. Nationaal Natuurhistorisch Museum, Leiden, 328 pp." type="book" year="1993">Holthuis 1993</bibRefCitation>
: 118
</treatmentCitation>
(key), 142.—
<treatmentCitation id="E10110C1FFA2F131FBB2FEBFFB73FEBF" author="Li" box="[1047,1170,328,350]" page="48" pageId="1" pageNumber="172" year="2000">
<bibRefCitation id="04314B21FFA2F131FBB2FEBFFB8CFEBF" author="Li" box="[1047,1133,328,350]" pageId="1" pageNumber="172" refString="Li, X. (2000) Catalog of the Genera and Species of Pontoniinae Kingsley, 1878 (Decapoda, Palaemonidae). Xueyuan Press, Beijing, VIII + 319 pp." type="book" year="2000">Li 2000</bibRefCitation>
: 48
</treatmentCitation>
;
<treatmentCitation id="E10110C1FFA2F131FB38FEBFFAB8FEBF" author="Li" box="[1181,1369,328,350]" page="171" pageId="1" pageNumber="172" year="2007">
<bibRefCitation id="04314B21FFA2F131FB38FEBFFAC8FEBF" author="Li" box="[1181,1321,328,350]" pageId="1" pageNumber="172" refString="Li, X., Liu, R., Lian, X. &amp; Chen, G. (2007) Crustacea Decapoda Palaemonoidea. Fauna Sinica Invertebrata Vol. 44. Science Press, Beijing, ix + 369 pp." type="book" year="2007">Li et al. 2007</bibRefCitation>
: 171
</treatmentCitation>
(key), 198;
<treatmentCitation id="E10110C1FFA2F131FF5FFE90FE9CFE9E" author="Li" box="[250,381,359,381]" page="887" pageId="1" pageNumber="172" year="2009">
<bibRefCitation id="04314B21FFA2F131FF5FFE90FEACFE9E" author="Li" box="[250,333,359,381]" pageId="1" pageNumber="172" refString="Li, X. (2009) Sandimenes nov. gen., for Periclimenes hirsutus Bruce, 1971 (Decapoda, Caridea, Pontoniinae). Crustaceana, 82, 881 - 896. http: // dx. doi. org / 10.1163 / 156854009 x 427414" type="journal article" year="2009">Li 2009</bibRefCitation>
: 887
</treatmentCitation>
(key).
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</paragraph>
<paragraph id="601F36D0FFA2F131FF32FE5EFDE1FC90" blockId="1.[151,1437,425,1170]" pageId="1" pageNumber="172">
<emphasis id="52D4EAC2FFA2F131FF32FE5EFE6EFE20" bold="true" box="[151,399,425,450]" pageId="1" pageNumber="172">Emended diagnosis.</emphasis>
Body moderately robust, subcylindrical or slightly compressed laterally; integument glabrous. Rostrum slender, straight, dentate dorsally and ventrally, compressed laterally, lateral carinae distinct. Carapace with distinct postorbital ridge; suborbital angle not delimited; pterygostomial angle rounded; antennal tooth present; supraorbital and hepatic teeth absent. Fourth thoracic sternite with spiniform median process. Pleon with first to fourth pleura rounded; third pleomere with posterodorsal margin rounded, not produced. Telson with 2 pairs of long dorsolateral spines, posterior margin with 3 pairs of spines. Antennule normal. Antennal scale well developed. Eye with cornea narrower than eyestalk. Epistome unarmed. Mandible robust, with simple or 2- articulated palp; incisor and molar processes well developed. Maxillule with weakly divided palp, endites broad. Maxilla moderately broad scaphognathite. First and second maxillipeds normal. Third maxilliped with ischiomerus (antepenultimate segment) partially fused with basis; exopod well developed; rudimentary arthrobranch present. First pereopod slender. Second pereopods similar, subequal, fingers without conspicuous teeth. Third to fifth pereopods with simple dactyli; propodi each with spinules on flexor margin. Uropod with exopod armed with 1 or 2 movable posterolateral spines.
</paragraph>
<paragraph id="601F36D0FFA2F131FF62FC8AFA7DFC74" blockId="1.[151,1437,425,1170]" box="[199,1436,893,918]" pageId="1" pageNumber="172">
<emphasis id="52D4EAC2FFA2F131FF62FC8AFE87FC74" bold="true" box="[199,358,893,918]" pageId="1" pageNumber="172">Composition.</emphasis>
<taxonomicName id="A7A04D53FFA2F131FEC8FC89FD1CFC74" ID-CoL="5b20d688-6ef1-4d70-997c-0315fda2ff37" authority="Bruce, 1971" authorityName="Bruce" authorityYear="1971" box="[365,765,893,918]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="172" phylum="Arthropoda" rank="species" species="noctalbata">
<emphasis id="52D4EAC2FFA2F131FEC8FC89FD8FFC77" box="[365,622,893,917]" italics="true" pageId="1" pageNumber="172">Eupontonia noctalbata</emphasis>
<bibRefCitation id="04314B21FFA2F131FDD0FC89FD1CFC74" author="Bruce" box="[629,765,894,918]" pageId="1" pageNumber="172" refString="Bruce, A. J. (1971) Notes on some Indo-Pacific Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp. nov., a new pontoniinid shrimp from Mahe, the Seychelle Islands. Crustaceana, 20, 225 - 236. http: // dx. doi. org / 10.1163 / 156854071 x 00012" type="journal article" year="1971">Bruce, 1971</bibRefCitation>
</taxonomicName>
(
<typeStatus id="BF1B8872FFA2F131FCA9FC89FCDFFC74" box="[780,830,894,918]" pageId="1" pageNumber="172">type</typeStatus>
species),
<taxonomicName id="A7A04D53FFA2F131FC09FC89FB73FC74" ID-CoL="ace12d29-5de3-4457-8990-1cf3b1491455" authority="Bruce, 2010" authorityName="Bruce" authorityYear="2010" box="[940,1170,893,918]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="172" phylum="Arthropoda" rank="species" species="oahu">
<emphasis id="52D4EAC2FFA2F131FC09FC89FBE2FC77" box="[940,1027,893,917]" italics="true" pageId="1" pageNumber="172">E. oahu</emphasis>
<bibRefCitation id="04314B21FFA2F131FBACFC89FB73FC74" author="Bruce" box="[1033,1170,894,918]" pageId="1" pageNumber="172" refString="Bruce, A. J. (2010) Eupontonia oahu sp. nov., a second species of the genus Eupontonia Bruce, 1971, (Crustacea: Decapoda: Pontoniinae) from Oahu, Hawai'ian Islands. In: De Grave, S. &amp; Fransen, C. H. J. M. (Eds.), Contributions to Shrimp Taxonomy. Zootaxa, 2372, 405 - 414." type="journal article" year="2010">Bruce, 2010</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="A7A04D53FFA2F131FB6DFC89FAB7FC77" ID-CoL="336495c5-0031-4df9-a4c4-ee5125a7d4ae" box="[1224,1366,893,917]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="172" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA2F131FB6DFC89FAB7FC77" box="[1224,1366,893,917]" italics="true" pageId="1" pageNumber="172">E. gracilipes</emphasis>
</taxonomicName>
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<emphasis id="52D4EAC2FFA2F131FAF8FC8AFA7DFC74" bold="true" box="[1373,1436,893,918]" pageId="1" pageNumber="172">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="28BA655BFFA2F131FF62FC56FEBFFB70" pageId="1" pageNumber="172" type="discussion">
<paragraph id="601F36D0FFA2F131FF62FC56FEBFFB70" blockId="1.[151,1437,425,1170]" pageId="1" pageNumber="172">
<emphasis id="52D4EAC2FFA2F131FF62FC56FEDAFC58" bold="true" box="[199,315,929,954]" pageId="1" pageNumber="172">Remarks.</emphasis>
The present new species generally agrees with the generic diagnosis of
<taxonomicName id="A7A04D53FFA2F131FBC9FC55FB0EFC5B" box="[1132,1263,930,953]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="52D4EAC2FFA2F131FBC9FC55FB0EFC5B" box="[1132,1263,930,953]" italics="true" pageId="1" pageNumber="172">Eupontonia</emphasis>
</taxonomicName>
, particularly in the presence of a postorbital ridge on the carapace, the absence of a hepatic tooth on the carapace, the reduced cornea and stout eyestalk, and the spinulate propodi of the third and fourth pereopods (
<bibRefCitation id="04314B21FFA2F131FB10FC1DFAA3FBE0" author="Bruce" box="[1205,1346,1002,1026]" pageId="1" pageNumber="172" refString="Bruce, A. J. (1971) Notes on some Indo-Pacific Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp. nov., a new pontoniinid shrimp from Mahe, the Seychelle Islands. Crustaceana, 20, 225 - 236. http: // dx. doi. org / 10.1163 / 156854071 x 00012" type="journal article" year="1971">Bruce 1971</bibRefCitation>
;
<bibRefCitation id="04314B21FFA2F131FAF6FC1DFA71FBE0" author="Bruce" box="[1363,1424,1002,1026]" pageId="1" pageNumber="172" refString="Bruce, A. J. (2010) Eupontonia oahu sp. nov., a second species of the genus Eupontonia Bruce, 1971, (Crustacea: Decapoda: Pontoniinae) from Oahu, Hawai'ian Islands. In: De Grave, S. &amp; Fransen, C. H. J. M. (Eds.), Contributions to Shrimp Taxonomy. Zootaxa, 2372, 405 - 414." type="journal article" year="2010">2010</bibRefCitation>
). However, the presence of a postrostral tooth on the carapace in the new species requires a minor emendation to the generic diagnosis once emended by
<bibRefCitation id="04314B21FFA2F131FDF0FBC5FD15FBA8" author="Bruce" box="[597,756,1074,1098]" pageId="1" pageNumber="172" refString="Bruce, A. J. (2010) Eupontonia oahu sp. nov., a second species of the genus Eupontonia Bruce, 1971, (Crustacea: Decapoda: Pontoniinae) from Oahu, Hawai'ian Islands. In: De Grave, S. &amp; Fransen, C. H. J. M. (Eds.), Contributions to Shrimp Taxonomy. Zootaxa, 2372, 405 - 414." type="journal article" year="2010">Bruce (2010)</bibRefCitation>
, in differentiating
<taxonomicName id="A7A04D53FFA2F131FC7EFBC5FBBFFBAB" box="[987,1118,1074,1097]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="52D4EAC2FFA2F131FC7EFBC5FBBFFBAB" box="[987,1118,1074,1097]" italics="true" pageId="1" pageNumber="172">Eupontonia</emphasis>
</taxonomicName>
from
<taxonomicName id="A7A04D53FFA2F131FB17FBC5FB33FBAB" box="[1202,1234,1074,1097]" class="Malacostraca" family="Palaemonidae" genus="Vir" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="52D4EAC2FFA2F131FB17FBC5FB33FBAB" box="[1202,1234,1074,1097]" italics="true" pageId="1" pageNumber="172">Vir</emphasis>
</taxonomicName>
. In spite of the elimination of the character on the postrostral tooth,
<taxonomicName id="A7A04D53FFA2F131FD4CFBA1FC8DFB8F" box="[745,876,1110,1133]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="52D4EAC2FFA2F131FD4CFBA1FC8DFB8F" box="[745,876,1110,1133]" italics="true" pageId="1" pageNumber="172">Eupontonia</emphasis>
</taxonomicName>
is easily differentiated from
<taxonomicName id="A7A04D53FFA2F131FB16FBA1FB32FB8F" box="[1203,1235,1110,1133]" class="Malacostraca" family="Palaemonidae" genus="Vir" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="52D4EAC2FFA2F131FB16FBA1FB32FB8F" box="[1203,1235,1110,1133]" italics="true" pageId="1" pageNumber="172">Vir</emphasis>
</taxonomicName>
by the characters mentioned above.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="FCC7B766CE0BE116E42047EEF7C4ACE1">A new species of the pontoniine shrimp genus Eupontonia (Crustacea: Decapoda: Caridea: Palaemonidae) from the Ryukyu Islands, Japan</mods:title>
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<mods:namePart id="2EB772991BEFFBB980B5DFD0961405E1">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="2F2C3AFE440A8B74E64176083F4308CE">Minemizu, Ryo</mods:namePart>
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<treatment id="E80987C6FFA2F137FF32FB13FC5EFEFF" ID-DOI="http://doi.org/10.5281/zenodo.6134311" ID-GBIF-Taxon="119420380" ID-Zenodo-Dep="6134311" LSID="urn:lsid:plazi:treatment:E80987C6FFA2F137FF32FB13FC5EFEFF" httpUri="http://treatment.plazi.org/id/E80987C6FFA2F137FF32FB13FC5EFEFF" lastPageId="7" lastPageNumber="178" pageId="1" pageNumber="172">
<subSubSection id="28BA655BFFA2F131FF32FB13FEE1FAA6" pageId="1" pageNumber="172" type="nomenclature">
<paragraph id="601F36D0FFA2F131FF32FB13FE0DFB1D" blockId="1.[151,643,1252,1349]" box="[151,492,1252,1279]" pageId="1" pageNumber="172">
<heading id="3B5781BCFFA2F131FF32FB13FE0DFB1D" bold="true" box="[151,492,1252,1279]" fontSize="11" level="1" pageId="1" pageNumber="172" reason="1">
<emphasis id="52D4EAC2FFA2F131FF32FB13FE0DFB1D" bold="true" box="[151,492,1252,1279]" pageId="1" pageNumber="172">
<taxonomicName id="A7A04D53FFA2F131FF32FB13FE43FB1C" ID-CoL="336495c5-0031-4df9-a4c4-ee5125a7d4ae" box="[151,418,1252,1278]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="172" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA2F131FF32FB13FE43FB1C" bold="true" box="[151,418,1252,1278]" italics="true" pageId="1" pageNumber="172">Eupontonia gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E757B9FFA2F131FE0FFB12FE0DFB1D" box="[426,492,1253,1279]" pageId="1" pageNumber="172" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="601F36D0FFA2F131FF32FAFFFEE1FAA6" blockId="1.[151,643,1252,1349]" pageId="1" pageNumber="172">
[New Japanese name: Awamori-kakure-ebi]
<figureCitation id="F89B2A55FFA2F131FF32FADBFEE1FAA6" box="[151,256,1324,1349]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1845,1867]" captionStartId-1="4.[151,250,1360,1382]" captionStartId-2="5.[151,250,1890,1912]" captionStartId-3="6.[151,250,1071,1093]" captionTargetBox-0="[231,1373,216,1827]" captionTargetBox-1="[185,1417,232,1312]" captionTargetBox-2="[217,1400,193,1832]" captionTargetBox-3="[151,1436,193,1049]" captionTargetId-0="figure@3.[195,1390,193,1840]" captionTargetId-1="figure@4.[151,1436,193,1339]" captionTargetId-2="figure@5.[194,1400,193,1859]" captionTargetId-3="figure@6.[151,1436,193,1050]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." captionText-1="FIGURE 2. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. Left appendages. A, mandible, inner view; B, maxillule, outer view; C, same, close up of distal part of palp, inner view; D, first maxilliped, outer view; E, second maxilliped, outer view; F, third maxilliped, lateral (outer) view; G, same, coxa, dorsal view; H, first pleopod, dorsal view (setae omitted); second pleopod, dorsal view (setae omitted). Scale bars: 0.5 mm." captionText-2="FIGURE 3. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, left first pereopod, lateral view; B, same, chela, extensor view; C, same, mesial view; D, left second pereopod, lateral view; E, same, chela, extensor view; F, left third pereopod, lateral view; G, same, dactylus, lateral view; H, left fourth pereopod, lateral view; I, same, dactylus, lateral view; J, right fifth pereopod, lateral view; K, same, dactylus and distal part of propodus, mesial view. Scale bars: 1 mm for A, D, F, H, J; 0.5 mm for B, C, G, I, K." captionText-3="FIGURE 4. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300, habitus in situ, photo taken by R. Minemizu." httpUri-0="https://zenodo.org/record/224485/files/figure.png" httpUri-1="https://zenodo.org/record/224486/files/figure.png" httpUri-2="https://zenodo.org/record/224487/files/figure.png" httpUri-3="https://zenodo.org/record/224488/files/figure.png" pageId="1" pageNumber="172">Figs. 1–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="28BA655BFFA2F131FF32FA83FB1AFA52" pageId="1" pageNumber="172" type="materials_examined">
<paragraph id="601F36D0FFA2F131FF32FA83FB1AFA52" blockId="1.[151,1436,1396,2033]" pageId="1" pageNumber="172">
<emphasis id="52D4EAC2FFA2F131FF32FA83FE65FA6F" bold="true" box="[151,388,1396,1421]" pageId="1" pageNumber="172">Material examined.</emphasis>
<typeStatus id="BF1B8872FFA2F131FE2AFA83FE1DFA6F" box="[399,508,1396,1421]" pageId="1" pageNumber="172" type="holotype">Holotype</typeStatus>
: off Osaki, Ishigaki Island, Yaeyama Islands, Ryukyu Islands,
<quantity id="A7589B35FFA2F131FB7CFA83FAEBFA6E" box="[1241,1290,1396,1421]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="172" unit="m" value="5.0">5 m</quantity>
, under coral rubble,
<date id="141E1010FFA2F131FF49FA6FFE59FA52" box="[236,440,1431,1456]" pageId="1" pageNumber="172" value="2011-11-04">4 November 2011</date>
, coll. R. Minemizu,
<collectionCode id="06B1AE15FFA2F131FD38FA6FFD1BFA52" box="[669,762,1432,1456]" pageId="1" pageNumber="172">SCUBA</collectionCode>
diving, female (cl
<quantity id="A7589B35FFA2F131FC6AFA6FFBC8FA52" box="[975,1065,1432,1456]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="172" unit="mm" value="3.2">3.2 mm</quantity>
), CBM-ZC 12300.
</paragraph>
</subSubSection>
<subSubSection id="28BA655BFFA2F134FF62FA4BFEE3F834" lastPageId="4" lastPageNumber="175" pageId="1" pageNumber="172" type="description">
<paragraph id="601F36D0FFA2F131FF62FA4BFBF6FA37" blockId="1.[151,1436,1396,2033]" box="[199,1047,1468,1493]" pageId="1" pageNumber="172">
<emphasis id="52D4EAC2FFA2F131FF62FA4BFEB8FA37" bold="true" box="[199,345,1468,1493]" pageId="1" pageNumber="172">Description.</emphasis>
Body (
<figureCitation id="F89B2A55FFA2F131FE0EFA4BFE08FA36" box="[427,489,1468,1493]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="1" pageNumber="172">Fig. 1</figureCitation>
A, D) subcylindrical; tegmental surface glabrous.
</paragraph>
<paragraph id="601F36D0FFA2F131FF62FA17FCC4F932" blockId="1.[151,1436,1396,2033]" pageId="1" pageNumber="172">
Rostrum (
<figureCitation id="F89B2A55FFA2F131FE9EFA28FE9FFA1A" box="[315,382,1503,1528]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="1" pageNumber="172">Fig. 1</figureCitation>
A, B) moderately slender, straight, about 0.4 of carapace length, directed forward, nearly reaching distal margin of second segment of antennular peduncle; dorsal margin armed with 6 teeth, including 1 posterior to rostral base located at 0.1 of carapace length; ventral margin slightly convex in lateral view, with 2 small teeth in distal 0.4; lateral surface with sharp carina in proximal half, merging into orbital margin. Carapace (
<figureCitation id="F89B2A55FFA2F131FF3AF998FF03F96A" box="[159,226,1647,1672]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="1" pageNumber="172">Fig. 1</figureCitation>
A, B) with low but distinct postrostral median ridge extending to midlength; postorbital ridge distinct, delimiting deep orbital concavity; suborbital angle not produced; antennal tooth moderately small, directed forward; pterygostomial angle slightly produced, rounded.
</paragraph>
<paragraph id="601F36D0FFA2F131FF62F92BFD6EF882" blockId="1.[151,1436,1396,2033]" pageId="1" pageNumber="172">
Epistome with pair of low protuberances on either side of midline. Labrum large, well produced, seen in lateral view (
<figureCitation id="F89B2A55FFA2F131FF78F908FEFCF8FA" box="[221,285,1791,1816]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="1" pageNumber="172">Fig. 1</figureCitation>
A). Fourth thoracic sternite (
<figureCitation id="F89B2A55FFA2F131FDC7F908FD40F8FA" box="[610,673,1791,1816]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="1" pageNumber="172">Fig. 1</figureCitation>
C) with spine-like median process directed anteroventrally and pair of small teeth laterally on anterior margin. Fifth sternite (
<figureCitation id="F89B2A55FFA2F131FCBEF8D3FCBBF8DE" box="[795,858,1828,1853]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="1" pageNumber="172">Fig. 1</figureCitation>
C) with pair of acute submedian teeth separated by U-shaped notch. Posterior sternites unarmed.
</paragraph>
<paragraph id="601F36D0FFA2F132FF62F89BFF29FEFF" blockId="1.[151,1436,1396,2033]" lastBlockId="2.[151,1436,151,2013]" lastPageId="2" lastPageNumber="173" pageId="1" pageNumber="172">
Pleon (
<figureCitation id="F89B2A55FFA2F131FEB9F89BFEBEF866" box="[284,351,1900,1925]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="1" pageNumber="172">Fig. 1</figureCitation>
D) dorsally rounded. First to fourth pleura rounded, fifth and sixth pleura each with small posteroventral tooth. Sixth pleomere 1.6 times as long as fifth pleomere and 1.4 times longer than high; posterolateral process terminating in sharp tooth. Telson (
<figureCitation id="F89B2A55FFA2F131FCBFF843FCB8F82E" box="[794,857,1972,1997]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="1" pageNumber="172">Fig. 1</figureCitation>
D, E) about 1.3 times as long as sixth pleomere and 2.4 times as long as wide, armed with 1 left and 2 right long dorsolateral spines (anterior spine located at 0.3 of telson length, posterior pair at about midlength); lateral margin parallel in anterior half, tapering to convex posterior margin in posterior half; posterior margin armed with 3 pairs of spines, lateral pair moderately short, intermediate pair elongate, about 0.3 times as long as telson, mesial pair slender, about 0.6 length of intermediate pair.
</paragraph>
<paragraph id="601F36D0FFA1F132FF62FEDFFA6AFE86" blockId="2.[151,1436,151,2013]" pageId="2" pageNumber="173">
Eye (
<figureCitation id="F89B2A55FFA1F132FEA2FED0FEAAFEA2" box="[263,331,295,320]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="2" pageNumber="173">Fig. 1</figureCitation>
A, B) with cornea fairly reduced in size, semispherical, darkly pigmented, much shorter and narrower than eyestalk; no ocellar spot evident. Eyestalk somewhat inflated with lateral and mesial faces convex.
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<paragraph id="601F36D0FFA1F132FF62FE98FB81FDDF" blockId="2.[151,1436,151,2013]" pageId="2" pageNumber="173">
Antennular peduncle (
<figureCitation id="F89B2A55FFA1F132FE63FE98FDE6FE6A" box="[454,519,367,392]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="2" pageNumber="173">Fig. 1</figureCitation>
A, B) reaching distal 0.3 of antennal scale. First segment broad, lateral margin slightly convex, distolateral angle produced, terminating in small sharp tooth; dorsal surface concave; ventromesial margin with small acute tooth at about midlength; stylocerite short, acute, located proximal to midlength of first peduncular segment. Second segment less than 0.3 length of first segment. Third segment slightly longer and narrower than second segment. Lateral flagellum biramous, 10 proximal articles fused; shorter ramus (
<figureCitation id="F89B2A55FFA1F132FA98FE08FA9EFDFA" box="[1341,1407,511,536]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="2" pageNumber="173">Fig. 1</figureCitation>
F) very short, consisting of 5 articles, distal 4 article each bearing 1 or 2 long aesthetascs.
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<paragraph id="601F36D0FFA1F132FF62FDBFFB91FD2F" blockId="2.[151,1436,151,2013]" pageId="2" pageNumber="173">
Antenna (
<figureCitation id="F89B2A55FFA1F132FE90FDB0FE93FD82" box="[309,370,583,608]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="2" pageNumber="173">Fig. 1</figureCitation>
A, B, G) with stout basicerite bearing small acute tooth on distolateral margin. Fifth segment of antennal peduncle (carpocerite) cylindrical, far falling short of midlength of antennal scale. Antennal scale about 0.4 of carapace length, 2.8 times longer than wide, lateral margin straight; distolateral tooth moderately strong, slightly overreaching rounded distal lamella. Flagellum exceeding twice length of body.
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<paragraph id="601F36D0FFA1F132FF62FD20FDB7FA9A" blockId="2.[151,1436,151,2013]" pageId="2" pageNumber="173">
Mandible (left) (
<figureCitation id="F89B2A55FFA1F132FE24FD20FE25FD12" box="[385,452,727,752]" captionStart="FIGURE 2" captionStartId="4.[151,250,1360,1382]" captionTargetBox="[185,1417,232,1312]" captionTargetId="figure@4.[151,1436,193,1339]" captionTargetPageId="4" captionText="FIGURE 2. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. Left appendages. A, mandible, inner view; B, maxillule, outer view; C, same, close up of distal part of palp, inner view; D, first maxilliped, outer view; E, second maxilliped, outer view; F, third maxilliped, lateral (outer) view; G, same, coxa, dorsal view; H, first pleopod, dorsal view (setae omitted); second pleopod, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/224486/files/figure.png" pageId="2" pageNumber="173">Fig. 2</figureCitation>
A) with small 2-articulated palp, distal article subequal to proximal article, with 1 short terminal seta; molar process robust, truncate distally, with 3 principal peripheral teeth; incisor process large, armed with 3 unequal teeth distally. Maxillule (
<figureCitation id="F89B2A55FFA1F132FDC5FCE8FD45FCDA" box="[608,676,799,824]" captionStart="FIGURE 2" captionStartId="4.[151,250,1360,1382]" captionTargetBox="[185,1417,232,1312]" captionTargetId="figure@4.[151,1436,193,1339]" captionTargetPageId="4" captionText="FIGURE 2. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. Left appendages. A, mandible, inner view; B, maxillule, outer view; C, same, close up of distal part of palp, inner view; D, first maxilliped, outer view; E, second maxilliped, outer view; F, third maxilliped, lateral (outer) view; G, same, coxa, dorsal view; H, first pleopod, dorsal view (setae omitted); second pleopod, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/224486/files/figure.png" pageId="2" pageNumber="173">Fig. 2</figureCitation>
B) with palp deeply bilobed, outer lobe tapering distally; inner lobe short, strongly curved inward, with minute apical seta (
<figureCitation id="F89B2A55FFA1F132FCADFCB3FCADFCBE" box="[776,844,836,861]" captionStart="FIGURE 2" captionStartId="4.[151,250,1360,1382]" captionTargetBox="[185,1417,232,1312]" captionTargetId="figure@4.[151,1436,193,1339]" captionTargetPageId="4" captionText="FIGURE 2. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. Left appendages. A, mandible, inner view; B, maxillule, outer view; C, same, close up of distal part of palp, inner view; D, first maxilliped, outer view; E, second maxilliped, outer view; F, third maxilliped, lateral (outer) view; G, same, coxa, dorsal view; H, first pleopod, dorsal view (setae omitted); second pleopod, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/224486/files/figure.png" pageId="2" pageNumber="173">Fig. 2</figureCitation>
C); proximal endite extending as far as distal endite, slightly tapering to roundly truncate tip, bearing stiff setae distally; distal endite subrectangular, with double row of spines and stiff setae on truncate distal margin. Maxilla damaged during dissection (not figured). First maxilliped (
<figureCitation id="F89B2A55FFA1F132FF3AFC58FF03FC2A" box="[159,226,943,968]" captionStart="FIGURE 2" captionStartId="4.[151,250,1360,1382]" captionTargetBox="[185,1417,232,1312]" captionTargetId="figure@4.[151,1436,193,1339]" captionTargetPageId="4" captionText="FIGURE 2. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. Left appendages. A, mandible, inner view; B, maxillule, outer view; C, same, close up of distal part of palp, inner view; D, first maxilliped, outer view; E, second maxilliped, outer view; F, third maxilliped, lateral (outer) view; G, same, coxa, dorsal view; H, first pleopod, dorsal view (setae omitted); second pleopod, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/224486/files/figure.png" pageId="2" pageNumber="173">Fig. 2</figureCitation>
D) with proximal endite somewhat thickened, mesial margin doubled; basial endite suboval, separated from proximal endite by shallow notch followed by transverse suture; endopod relatively wide, tapering distally, far falling short of distal margin of distal endite; exopod well developed, with long flagellum, caridean lobe moderately broad; epipod slightly bilobed. Second maxilliped (
<figureCitation id="F89B2A55FFA1F132FC2CFBEBFC33FBD6" box="[905,978,1052,1077]" captionStart="FIGURE 2" captionStartId="4.[151,250,1360,1382]" captionTargetBox="[185,1417,232,1312]" captionTargetId="figure@4.[151,1436,193,1339]" captionTargetPageId="4" captionText="FIGURE 2. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. Left appendages. A, mandible, inner view; B, maxillule, outer view; C, same, close up of distal part of palp, inner view; D, first maxilliped, outer view; E, second maxilliped, outer view; F, third maxilliped, lateral (outer) view; G, same, coxa, dorsal view; H, first pleopod, dorsal view (setae omitted); second pleopod, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/224486/files/figure.png" pageId="2" pageNumber="173">Fig. 2</figureCitation>
E) with dactylus and propodus clearly separated; mesial margin of dactylus with numerous stiff setae; propodus with row of moderately long stiff setae on mesial margin; ischium and basis fused; coxa with convex mesial margin; exopod moderately long; epipod moderately large, subrectangular, without podobranch. Third maxilliped (
<figureCitation id="F89B2A55FFA1F132FC5AFB70FBA9FB42" box="[1023,1096,1159,1184]" captionStart="FIGURE 2" captionStartId="4.[151,250,1360,1382]" captionTargetBox="[185,1417,232,1312]" captionTargetId="figure@4.[151,1436,193,1339]" captionTargetPageId="4" captionText="FIGURE 2. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. Left appendages. A, mandible, inner view; B, maxillule, outer view; C, same, close up of distal part of palp, inner view; D, first maxilliped, outer view; E, second maxilliped, outer view; F, third maxilliped, lateral (outer) view; G, same, coxa, dorsal view; H, first pleopod, dorsal view (setae omitted); second pleopod, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/224486/files/figure.png" pageId="2" pageNumber="173">Fig. 2</figureCitation>
F) with endopod moderately slender, not reaching distal margin of antennal scale; coxa (
<figureCitation id="F89B2A55FFA1F132FC94FB5BFC70FB26" box="[817,913,1196,1221]" captionStart="FIGURE 2" captionStartId="4.[151,250,1360,1382]" captionTargetBox="[185,1417,232,1312]" captionTargetId="figure@4.[151,1436,193,1339]" captionTargetPageId="4" captionText="FIGURE 2. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. Left appendages. A, mandible, inner view; B, maxillule, outer view; C, same, close up of distal part of palp, inner view; D, first maxilliped, outer view; E, second maxilliped, outer view; F, third maxilliped, lateral (outer) view; G, same, coxa, dorsal view; H, first pleopod, dorsal view (setae omitted); second pleopod, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/224486/files/figure.png" pageId="2" pageNumber="173">Figure 2</figureCitation>
G) with oval lateral plate (= epipod) and small setose tubercle on mesial margin; basis short, partially fused to ischiomerus; ischiomerus somewhat bowed in dorsal view, slightly tapering distally; carpus about 0.7 length of ischiomerus, about 4.9 times longer than high; ultimate segment 0.7 length of carpus, 4.9 times longer than high, tapering distally, with long terminal unguis and numerous stiff setae on mesial face; exopod well developed, reaching distal end of ischiomerus; single arthrobranch poorly lamellate, bud-like.
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<paragraph id="601F36D0FFA1F132FF62FA73FE1BF9B2" blockId="2.[151,1436,151,2013]" pageId="2" pageNumber="173">
First pereopod (
<figureCitation id="F89B2A55FFA1F132FED8FA73FE21FA7E" box="[381,448,1412,1437]" captionStart="FIGURE 3" captionStartId="5.[151,250,1890,1912]" captionTargetBox="[217,1400,193,1832]" captionTargetId="figure@5.[194,1400,193,1859]" captionTargetPageId="5" captionText="FIGURE 3. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, left first pereopod, lateral view; B, same, chela, extensor view; C, same, mesial view; D, left second pereopod, lateral view; E, same, chela, extensor view; F, left third pereopod, lateral view; G, same, dactylus, lateral view; H, left fourth pereopod, lateral view; I, same, dactylus, lateral view; J, right fifth pereopod, lateral view; K, same, dactylus and distal part of propodus, mesial view. Scale bars: 1 mm for A, D, F, H, J; 0.5 mm for B, C, G, I, K." httpUri="https://zenodo.org/record/224487/files/figure.png" pageId="2" pageNumber="173">Fig. 3</figureCitation>
A) slender, reaching distal margin of antennal scale by mero-carpal articulation. Chela (
<figureCitation id="F89B2A55FFA1F132FF3AFA50FF00FA22" box="[159,225,1447,1472]" captionStart="FIGURE 3" captionStartId="5.[151,250,1890,1912]" captionTargetBox="[217,1400,193,1832]" captionTargetId="figure@5.[194,1400,193,1859]" captionTargetPageId="5" captionText="FIGURE 3. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, left first pereopod, lateral view; B, same, chela, extensor view; C, same, mesial view; D, left second pereopod, lateral view; E, same, chela, extensor view; F, left third pereopod, lateral view; G, same, dactylus, lateral view; H, left fourth pereopod, lateral view; I, same, dactylus, lateral view; J, right fifth pereopod, lateral view; K, same, dactylus and distal part of propodus, mesial view. Scale bars: 1 mm for A, D, F, H, J; 0.5 mm for B, C, G, I, K." httpUri="https://zenodo.org/record/224487/files/figure.png" pageId="2" pageNumber="173">Fig. 3</figureCitation>
B, C) 4.4 times longer than broad; dactylus about 1.2 of palm length; cutting edges of fingers entire, forming thin edge; palm subcylindrical. Carpus 1.2 length of chela, 7.3 times longer than wide. Merus subequal in length to carpus. Ischium 0.7 length of merus. Grooming apparatus consisting of some short transverse rows of setae on proximomesial portion of palm and transverse row of short setae plus tuft of long setae located near ventromesial distal angle of carpus (
<figureCitation id="F89B2A55FFA1F132FE30F9C0FE36F9B2" box="[405,471,1591,1616]" captionStart="FIGURE 3" captionStartId="5.[151,250,1890,1912]" captionTargetBox="[217,1400,193,1832]" captionTargetId="figure@5.[194,1400,193,1859]" captionTargetPageId="5" captionText="FIGURE 3. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, left first pereopod, lateral view; B, same, chela, extensor view; C, same, mesial view; D, left second pereopod, lateral view; E, same, chela, extensor view; F, left third pereopod, lateral view; G, same, dactylus, lateral view; H, left fourth pereopod, lateral view; I, same, dactylus, lateral view; J, right fifth pereopod, lateral view; K, same, dactylus and distal part of propodus, mesial view. Scale bars: 1 mm for A, D, F, H, J; 0.5 mm for B, C, G, I, K." httpUri="https://zenodo.org/record/224487/files/figure.png" pageId="2" pageNumber="173">Fig. 3</figureCitation>
B).
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<paragraph id="601F36D0FFA1F132FF62F9ABFB91F8AE" blockId="2.[151,1436,151,2013]" pageId="2" pageNumber="173">
Second pereopods (
<figureCitation id="F89B2A55FFA1F132FE0EF9ABFE10F996" box="[427,497,1628,1653]" captionStart="FIGURE 3" captionStartId="5.[151,250,1890,1912]" captionTargetBox="[217,1400,193,1832]" captionTargetId="figure@5.[194,1400,193,1859]" captionTargetPageId="5" captionText="FIGURE 3. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, left first pereopod, lateral view; B, same, chela, extensor view; C, same, mesial view; D, left second pereopod, lateral view; E, same, chela, extensor view; F, left third pereopod, lateral view; G, same, dactylus, lateral view; H, left fourth pereopod, lateral view; I, same, dactylus, lateral view; J, right fifth pereopod, lateral view; K, same, dactylus and distal part of propodus, mesial view. Scale bars: 1 mm for A, D, F, H, J; 0.5 mm for B, C, G, I, K." httpUri="https://zenodo.org/record/224487/files/figure.png" pageId="2" pageNumber="173">Fig. 3</figureCitation>
D) subequal, similar, moderately slender, overreaching distal margin of antennal scale by 0.2 length of merus. Chela (
<figureCitation id="F89B2A55FFA1F132FDE5F988FD62F97A" box="[576,643,1663,1688]" captionStart="FIGURE 3" captionStartId="5.[151,250,1890,1912]" captionTargetBox="[217,1400,193,1832]" captionTargetId="figure@5.[194,1400,193,1859]" captionTargetPageId="5" captionText="FIGURE 3. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, left first pereopod, lateral view; B, same, chela, extensor view; C, same, mesial view; D, left second pereopod, lateral view; E, same, chela, extensor view; F, left third pereopod, lateral view; G, same, dactylus, lateral view; H, left fourth pereopod, lateral view; I, same, dactylus, lateral view; J, right fifth pereopod, lateral view; K, same, dactylus and distal part of propodus, mesial view. Scale bars: 1 mm for A, D, F, H, J; 0.5 mm for B, C, G, I, K." httpUri="https://zenodo.org/record/224487/files/figure.png" pageId="2" pageNumber="173">Fig. 3</figureCitation>
E) 1.4 of carapace length, 5.9 times longer than wide in dorsal view; surfaces smooth and glabrous. Dactylus slender, 1.2 length of palm, terminating in curved, acute tip; fixed finger similarly slender, slightly deflexed, terminating in curved, acute tip crossing tip of dactylus; cutting edges of both fingers forming thin plate, without conspicuous teeth. Palm subcylindrical, slightly swollen, 2.8 times longer than wide. Carpus slightly widened distally, 0.70 of chela length, 1.7 length of merus, 6.5 times longer than high; surfaces smooth. Merus 6.8 times longer than high. Ischium slightly shorter than merus.
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<paragraph id="601F36D0FFA1F134FF62F8A0FA99F98C" blockId="2.[151,1436,151,2013]" lastBlockId="4.[151,1436,1513,2006]" lastPageId="4" lastPageNumber="175" pageId="2" pageNumber="173">
Ambulatory legs (third to fifth pereopods) slender compared with two other congeneric species. Third pereopod (
<figureCitation id="F89B2A55FFA1F132FEB7F88BFEB7F876" box="[274,342,1916,1941]" captionStart="FIGURE 3" captionStartId="5.[151,250,1890,1912]" captionTargetBox="[217,1400,193,1832]" captionTargetId="figure@5.[194,1400,193,1859]" captionTargetPageId="5" captionText="FIGURE 3. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, left first pereopod, lateral view; B, same, chela, extensor view; C, same, mesial view; D, left second pereopod, lateral view; E, same, chela, extensor view; F, left third pereopod, lateral view; G, same, dactylus, lateral view; H, left fourth pereopod, lateral view; I, same, dactylus, lateral view; J, right fifth pereopod, lateral view; K, same, dactylus and distal part of propodus, mesial view. Scale bars: 1 mm for A, D, F, H, J; 0.5 mm for B, C, G, I, K." httpUri="https://zenodo.org/record/224487/files/figure.png" pageId="2" pageNumber="173">Fig. 3</figureCitation>
F) overreaching distal margin of antennal scale by full length of propodus; dactylus (
<figureCitation id="F89B2A55FFA1F132FA9CF88BFA9CF876" box="[1337,1405,1916,1941]" captionStart="FIGURE 3" captionStartId="5.[151,250,1890,1912]" captionTargetBox="[217,1400,193,1832]" captionTargetId="figure@5.[194,1400,193,1859]" captionTargetPageId="5" captionText="FIGURE 3. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, left first pereopod, lateral view; B, same, chela, extensor view; C, same, mesial view; D, left second pereopod, lateral view; E, same, chela, extensor view; F, left third pereopod, lateral view; G, same, dactylus, lateral view; H, left fourth pereopod, lateral view; I, same, dactylus, lateral view; J, right fifth pereopod, lateral view; K, same, dactylus and distal part of propodus, mesial view. Scale bars: 1 mm for A, D, F, H, J; 0.5 mm for B, C, G, I, K." httpUri="https://zenodo.org/record/224487/files/figure.png" pageId="2" pageNumber="173">Fig. 3</figureCitation>
G) simple, compressed laterally, feebly curved, 0.25 of propodal length, 7.5 times longer than high, unguis not clearly demarcated, with tuft of short setae on extensor margin slightly distal to midlength; propodus with 4 widely spaced spinules on flexor margin, including 1 pair on distal margin; carpus 0.7 of propodal length; merus about 16 times longer than high; ischium 0.6 of meral length. Fourth pereopod (
<figureCitation id="F89B2A55FFA7F134FCC9F9FAFC4FF9C4" box="[876,942,1549,1574]" captionStart="FIGURE 3" captionStartId="5.[151,250,1890,1912]" captionTargetBox="[217,1400,193,1832]" captionTargetId="figure@5.[194,1400,193,1859]" captionTargetPageId="5" captionText="FIGURE 3. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, left first pereopod, lateral view; B, same, chela, extensor view; C, same, mesial view; D, left second pereopod, lateral view; E, same, chela, extensor view; F, left third pereopod, lateral view; G, same, dactylus, lateral view; H, left fourth pereopod, lateral view; I, same, dactylus, lateral view; J, right fifth pereopod, lateral view; K, same, dactylus and distal part of propodus, mesial view. Scale bars: 1 mm for A, D, F, H, J; 0.5 mm for B, C, G, I, K." httpUri="https://zenodo.org/record/224487/files/figure.png" pageId="4" pageNumber="175">Fig. 3</figureCitation>
H, I) overreaching distal margin of antennal scale by 0.7 length of propodus, similar to third pereopod. Fifth pereopod (
<figureCitation id="F89B2A55FFA7F134FC7CF9C6FBFAF9A8" box="[985,1051,1585,1610]" captionStart="FIGURE 3" captionStartId="5.[151,250,1890,1912]" captionTargetBox="[217,1400,193,1832]" captionTargetId="figure@5.[194,1400,193,1859]" captionTargetPageId="5" captionText="FIGURE 3. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, left first pereopod, lateral view; B, same, chela, extensor view; C, same, mesial view; D, left second pereopod, lateral view; E, same, chela, extensor view; F, left third pereopod, lateral view; G, same, dactylus, lateral view; H, left fourth pereopod, lateral view; I, same, dactylus, lateral view; J, right fifth pereopod, lateral view; K, same, dactylus and distal part of propodus, mesial view. Scale bars: 1 mm for A, D, F, H, J; 0.5 mm for B, C, G, I, K." httpUri="https://zenodo.org/record/224487/files/figure.png" pageId="4" pageNumber="175">Fig. 3</figureCitation>
J, K) generally similar to third and fourth pereopods, but longer than them, overreaching distal margin of antennal scale by 0.7 length of propodus.
</paragraph>
<caption id="34DF6658FFA0F133FF32F8C2FE9EF806" httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="3" pageNumber="174" targetBox="[231,1373,216,1827]" targetPageId="3">
<paragraph id="601F36D0FFA0F133FF32F8C2FE9EF806" blockId="3.[151,1436,1845,2021]" pageId="3" pageNumber="174">
<emphasis id="52D4EAC2FFA0F133FF32F8C2FEFBF8A8" bold="true" box="[151,282,1845,1867]" pageId="3" pageNumber="174">FIGURE 1.</emphasis>
<taxonomicName id="A7A04D53FFA0F133FE82F8C1FDEEF8A9" box="[295,527,1845,1867]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA0F133FE82F8C1FDEEF8A9" box="[295,527,1845,1867]" italics="true" pageId="3" pageNumber="174">Eupontonia gracilipes</emphasis>
</taxonomicName>
<emphasis id="52D4EAC2FFA0F133FDB8F8C1FDBCF8A9" bold="true" box="[541,605,1846,1867]" pageId="3" pageNumber="174">
<taxonomicNameLabel id="49E757B9FFA0F133FDB8F8C1FDBCF8A9" box="[541,605,1846,1867]" pageId="3" pageNumber="174" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E–H.
</paragraph>
</caption>
<caption id="34DF6658FFA7F134FF32FAA7FC6CFA20" httpUri="https://zenodo.org/record/224486/files/figure.png" pageId="4" pageNumber="175" targetBox="[185,1417,232,1312]" targetPageId="4">
<paragraph id="601F36D0FFA7F134FF32FAA7FC6CFA20" blockId="4.[151,1436,1360,1474]" pageId="4" pageNumber="175">
<emphasis id="52D4EAC2FFA7F134FF32FAA7FEF3FA87" bold="true" box="[151,274,1360,1382]" pageId="4" pageNumber="175">FIGURE 2.</emphasis>
<taxonomicName id="A7A04D53FFA7F134FEBDFAA6FE18FA84" box="[280,505,1360,1382]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA7F134FEBDFAA6FE18FA84" box="[280,505,1360,1382]" italics="true" pageId="4" pageNumber="175">Eupontonia gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E757B9FFA7F134FDA5FAA6FDD8FA84" box="[512,569,1361,1382]" pageId="4" pageNumber="175" rank="species">
<emphasis id="52D4EAC2FFA7F134FDA5FAA6FDD8FA84" bold="true" box="[512,569,1361,1382]" pageId="4" pageNumber="175">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, female (cl 3.2 mm), CBM-ZC 12300. Left appendages. A, mandible, inner view; B, maxillule, outer view; C, same, close up of distal part of palp, inner view; D, first maxilliped, outer view; E, second maxilliped, outer view; F, third maxilliped, lateral (outer) view; G, same, coxa, dorsal view; H, first pleopod, dorsal view (setae omitted); second pleopod, dorsal view (setae omitted). Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="601F36D0FFA7F134FF62F98EFA9AF954" blockId="4.[151,1436,1513,2006]" pageId="4" pageNumber="175">
Protopods of pleopods slender. First pleopod (
<figureCitation id="F89B2A55FFA7F134FD69F98EFCEEF970" box="[716,783,1657,1682]" captionStart="FIGURE 2" captionStartId="4.[151,250,1360,1382]" captionTargetBox="[185,1417,232,1312]" captionTargetId="figure@4.[151,1436,193,1339]" captionTargetPageId="4" captionText="FIGURE 2. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. Left appendages. A, mandible, inner view; B, maxillule, outer view; C, same, close up of distal part of palp, inner view; D, first maxilliped, outer view; E, second maxilliped, outer view; F, third maxilliped, lateral (outer) view; G, same, coxa, dorsal view; H, first pleopod, dorsal view (setae omitted); second pleopod, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/224486/files/figure.png" pageId="4" pageNumber="175">Fig. 2</figureCitation>
H) with endopod about 0.40 length of exopod, moderately narrow. Second to fifth pleopods each with appendix interna arising at about midlength of endopod (cf.
<figureCitation id="F89B2A55FFA7F134FA85F96AFA82F954" box="[1312,1379,1693,1718]" captionStart="FIGURE 2" captionStartId="4.[151,250,1360,1382]" captionTargetBox="[185,1417,232,1312]" captionTargetId="figure@4.[151,1436,193,1339]" captionTargetPageId="4" captionText="FIGURE 2. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. Left appendages. A, mandible, inner view; B, maxillule, outer view; C, same, close up of distal part of palp, inner view; D, first maxilliped, outer view; E, second maxilliped, outer view; F, third maxilliped, lateral (outer) view; G, same, coxa, dorsal view; H, first pleopod, dorsal view (setae omitted); second pleopod, dorsal view (setae omitted). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/224486/files/figure.png" pageId="4" pageNumber="175">
Fig.
<date id="141E1010FFA7F134FAF6F969FA82F954" box="[1363,1379,1694,1718]" pageId="4" pageNumber="175">2</date>
</figureCitation>
I).
</paragraph>
<paragraph id="601F36D0FFA7F134FF62F936FDA5F8A4" blockId="4.[151,1436,1513,2006]" pageId="4" pageNumber="175">
Uropod (
<figureCitation id="F89B2A55FFA7F134FE8EF936FE88F938" box="[299,361,1729,1754]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="4" pageNumber="175">Fig. 1</figureCitation>
D) with protopod bearing small acute posterolateral tooth. Exopod distinctly overreaching tip of telson (except for terminal spines), with submarginal row of setae along lateral margin; lateral margin nearly straight, terminating in small acute tooth, with 2 small movable spines just mesial to posterolateral tooth (
<figureCitation id="F89B2A55FFA7F134FA90F8FEFA95F8C0" box="[1333,1396,1801,1826]" captionStart="FIGURE 1" captionStartId="3.[151,250,1845,1867]" captionTargetBox="[231,1373,216,1827]" captionTargetId="figure@3.[195,1390,193,1840]" captionTargetPageId="3" captionText="FIGURE 1. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300. A, carapace and cephalic appendages, lateral view (antennular flagella partially omitted; labrum shown); B, anterior part of carapace and cephalic appendages, dorsal view; C, fourth and fifth thoracic sternites, ventral view; D, pleon, lateral view; E, telson, dorsal view; F, bifurcated part of outer flagellum of left antennule; G, left antenna, ventral view (flagellum and marginal setae on antennal scale omitted); H, posterolateral tooth and spines of exopod of left uropod, dorsal view (setae omitted). Scale bars: 1 mm for A, D; 0.5 mm for B, C, E – H." httpUri="https://zenodo.org/record/224485/files/figure.png" pageId="4" pageNumber="175">Fig. 1</figureCitation>
H). Endopod slightly shorter than exopod.
</paragraph>
<paragraph id="601F36D0FFA7F134FF62F8A6FEE3F834" blockId="4.[151,1436,1513,2006]" pageId="4" pageNumber="175">
<emphasis id="52D4EAC2FFA7F134FF62F8A6FE7BF888" bold="true" box="[199,410,1873,1898]" pageId="4" pageNumber="175">Coloration in life.</emphasis>
Body and appendages entirely translucent. Corneas darkly pigmented. See
<figureCitation id="F89B2A55FFA7F134FB42F8A6FACDF888" box="[1255,1324,1873,1898]" captionStart="FIGURE 4" captionStartId="6.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@6.[151,1436,193,1050]" captionTargetPageId="6" captionText="FIGURE 4. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300, habitus in situ, photo taken by R. Minemizu." httpUri="https://zenodo.org/record/224488/files/figure.png" pageId="4" pageNumber="175">Fig. 4</figureCitation>
.
<emphasis id="52D4EAC2FFA7F134FF62F882FE81F86C" bold="true" box="[199,352,1909,1934]" pageId="4" pageNumber="175">Distribution.</emphasis>
Known only from Ishigaki Island, Yaeyama Islands, southern Ryukyu Islands, at a depth of
<specimenCount id="76A6FD59FFA7F134FACFF882FA7DF86C" box="[1386,1436,1909,1934]" pageId="4" pageNumber="175" type="generic">
<quantity id="A7589B35FFA7F134FACFF882FA79F86C" box="[1386,1432,1909,1934]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="175" unit="m" value="5.0">5 m</quantity>
.
</specimenCount>
<emphasis id="52D4EAC2FFA7F134FF62F86EFECBF850" bold="true" box="[199,298,1945,1970]" pageId="4" pageNumber="175">Habitat.</emphasis>
The unique
<typeStatus id="BF1B8872FFA7F134FE10F86EFDF6F850" box="[437,535,1945,1970]" pageId="4" pageNumber="175" type="holotype">holotype</typeStatus>
was found under coral rubble. No symbiotic association with other invertebrates was seen.
</paragraph>
</subSubSection>
<caption id="34DF6658FFA6F135FF32F895FDE0F811" httpUri="https://zenodo.org/record/224487/files/figure.png" pageId="5" pageNumber="176" targetBox="[217,1400,193,1832]" targetPageId="5">
<paragraph id="601F36D0FFA6F135FF32F895FDE0F811" blockId="5.[151,1436,1890,2035]" pageId="5" pageNumber="176">
<emphasis id="52D4EAC2FFA6F135FF32F895FEF5F895" bold="true" box="[151,276,1890,1912]" pageId="5" pageNumber="176">FIGURE 3.</emphasis>
<taxonomicName id="A7A04D53FFA6F135FEBEF894FE1CF89A" box="[283,509,1890,1912]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA6F135FEBEF894FE1CF89A" box="[283,509,1890,1912]" italics="true" pageId="5" pageNumber="176">Eupontonia gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E757B9FFA6F135FDA1F894FDDFF89A" box="[516,574,1891,1912]" pageId="5" pageNumber="176" rank="species">
<emphasis id="52D4EAC2FFA6F135FDA1F894FDDFF89A" bold="true" box="[516,574,1891,1912]" pageId="5" pageNumber="176">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, female (cl 3.2 mm), CBM-ZC 12300. A, left first pereopod, lateral view; B, same, chela, extensor view; C, same, mesial view; D, left second pereopod, lateral view; E, same, chela, extensor view; F, left third pereopod, lateral view; G, same, dactylus, lateral view; H, left fourth pereopod, lateral view; I, same, dactylus, lateral view; J, right fifth pereopod, lateral view; K, same, dactylus and distal part of propodus, mesial view. Scale bars: 1 mm for A, D, F, H, J; 0.5 mm for B, C, G, I, K.
</paragraph>
</caption>
<caption id="34DF6658FFA5F136FF32FBD8FEE7FB86" httpUri="https://zenodo.org/record/224488/files/figure.png" pageId="6" pageNumber="177" targetBox="[151,1436,193,1049]" targetPageId="6">
<paragraph id="601F36D0FFA5F136FF32FBD8FEE7FB86" blockId="6.[151,1436,1071,1124]" pageId="6" pageNumber="177">
<emphasis id="52D4EAC2FFA5F136FF32FBD8FEF5FBA6" bold="true" box="[151,276,1071,1093]" pageId="6" pageNumber="177">FIGURE 4.</emphasis>
<taxonomicName id="A7A04D53FFA5F136FEB9FBC6FE1EFBA7" box="[284,511,1071,1094]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA5F136FEB9FBC6FE1EFBA7" box="[284,511,1071,1094]" italics="true" pageId="6" pageNumber="177">Eupontonia gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E757B9FFA5F136FDA2FBC7FDA3FBA7" box="[519,578,1072,1093]" pageId="6" pageNumber="177" rank="species">
<emphasis id="52D4EAC2FFA5F136FDA2FBC7FDA3FBA7" bold="true" box="[519,578,1072,1093]" pageId="6" pageNumber="177">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, female (cl 3.2 mm), CBM-ZC 12300, habitus in situ, photo taken by R. Minemizu.
</paragraph>
</caption>
<subSubSection id="28BA655BFFA5F137FF62FB67FD40FF36" lastPageId="7" lastPageNumber="178" pageId="6" pageNumber="177" type="discussion">
<paragraph id="601F36D0FFA5F136FF62FB67FBD6FB13" blockId="6.[151,1436,1168,2022]" pageId="6" pageNumber="177">
<emphasis id="52D4EAC2FFA5F136FF62FB67FEDAFB4B" bold="true" box="[199,315,1168,1193]" pageId="6" pageNumber="177">Remarks.</emphasis>
The
<typeStatus id="BF1B8872FFA5F136FED3FB67FE39FB4B" box="[374,472,1168,1193]" pageId="6" pageNumber="177" type="holotype">holotype</typeStatus>
was externally infested by an unidentified isopod parasite, which was attached to the right side of the carapace (
<figureCitation id="F89B2A55FFA5F136FE6AFB43FDF7FB2F" box="[463,534,1204,1229]" captionStart="FIGURE 4" captionStartId="6.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@6.[151,1436,193,1050]" captionTargetPageId="6" captionText="FIGURE 4. Eupontonia gracilipes n. sp., holotype, female (cl 3.2 mm), CBM-ZC 12300, habitus in situ, photo taken by R. Minemizu." httpUri="https://zenodo.org/record/224488/files/figure.png" pageId="6" pageNumber="177">Fig. 4</figureCitation>
), although the parasite specimen was not preserved. The right first pleuron is modified to form an expanded fold to receive the anterior part of the parasite body.
</paragraph>
<paragraph id="601F36D0FFA5F136FF62FB0AFDF6F943" blockId="6.[151,1436,1168,2022]" pageId="6" pageNumber="177">
The present new species is readily distinguished from the other two congeneric species,
<taxonomicName id="A7A04D53FFA5F136FB16FB09FAAAFAF7" box="[1203,1355,1277,1301]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="noctalbata">
<emphasis id="52D4EAC2FFA5F136FB16FB09FAAAFAF7" box="[1203,1355,1277,1301]" italics="true" pageId="6" pageNumber="177">E. noctalbata</emphasis>
</taxonomicName>
and
<taxonomicName id="A7A04D53FFA5F136FA21FB09FF2EFADB" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="oahu">
<emphasis id="52D4EAC2FFA5F136FA21FB09FF2EFADB" italics="true" pageId="6" pageNumber="177">E. oahu</emphasis>
</taxonomicName>
, by the presence of a distinct postrostral median ridge on the carapace, extending to the midlength of the carapace, and of a postrostral median tooth located at about anterior 0.1 of the carapace. In the latter two species, there is no postrostral median ridge and no postrostral tooth on the carapace (
<bibRefCitation id="04314B21FFA5F136FC5AFA9EFB66FA63" author="Bruce" box="[1023,1159,1385,1409]" pageId="6" pageNumber="177" refString="Bruce, A. J. (1971) Notes on some Indo-Pacific Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp. nov., a new pontoniinid shrimp from Mahe, the Seychelle Islands. Crustaceana, 20, 225 - 236. http: // dx. doi. org / 10.1163 / 156854071 x 00012" type="journal article" year="1971">Bruce 1971</bibRefCitation>
: figs. 2, 3a, b; 2010: fig. 1A, C). Furthermore, ambulatory legs are comparatively slenderer in
<taxonomicName id="A7A04D53FFA5F136FC1BFA79FBB1FA47" box="[958,1104,1421,1445]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA5F136FC1BFA79FBB1FA47" box="[958,1104,1421,1445]" italics="true" pageId="6" pageNumber="177">E. gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E757B9FFA5F136FBFEFA7AFB7DFA44" box="[1115,1180,1421,1446]" pageId="6" pageNumber="177" rank="species">
<emphasis id="52D4EAC2FFA5F136FBFEFA7AFB7DFA44" bold="true" box="[1115,1180,1421,1446]" pageId="6" pageNumber="177">n. sp.</emphasis>
</taxonomicNameLabel>
than in the latter two species (cf.
<bibRefCitation id="04314B21FFA5F136FEBEFA46FE40FA2B" author="Bruce" box="[283,417,1457,1481]" pageId="6" pageNumber="177" refString="Bruce, A. J. (1971) Notes on some Indo-Pacific Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp. nov., a new pontoniinid shrimp from Mahe, the Seychelle Islands. Crustaceana, 20, 225 - 236. http: // dx. doi. org / 10.1163 / 156854071 x 00012" type="journal article" year="1971">Bruce 1971</bibRefCitation>
: fig. 5h; 2010: fig. 3D, E). The lack of a supraorbital tooth on the carapace and the rounded posterior margin of the telson also distinguish
<taxonomicName id="A7A04D53FFA5F136FD66FA21FCB6FA0F" box="[707,855,1493,1517]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA5F136FD66FA21FCB6FA0F" box="[707,855,1493,1517]" italics="true" pageId="6" pageNumber="177">E. gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E757B9FFA5F136FCC1FA22FC49FA0C" box="[868,936,1493,1518]" pageId="6" pageNumber="177" rank="species">
<emphasis id="52D4EAC2FFA5F136FCC1FA22FC49FA0C" bold="true" box="[868,936,1493,1518]" pageId="6" pageNumber="177">n. sp.</emphasis>
</taxonomicNameLabel>
from
<taxonomicName id="A7A04D53FFA5F136FC52FA21FB75FA0F" box="[1015,1172,1493,1517]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="noctalbata">
<emphasis id="52D4EAC2FFA5F136FC52FA21FB75FA0F" box="[1015,1172,1493,1517]" italics="true" pageId="6" pageNumber="177">E. noctalbata</emphasis>
</taxonomicName>
. In
<taxonomicName id="A7A04D53FFA5F136FB6FFA21FA86FA0F" box="[1226,1383,1493,1517]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="noctalbata">
<emphasis id="52D4EAC2FFA5F136FB6FFA21FA86FA0F" box="[1226,1383,1493,1517]" italics="true" pageId="6" pageNumber="177">E. noctalbata</emphasis>
</taxonomicName>
, the carapace is provided with a pair of supraorbital teeth (
<bibRefCitation id="04314B21FFA5F136FD5CFA0EFC60F9F3" author="Bruce" box="[761,897,1529,1553]" pageId="6" pageNumber="177" refString="Bruce, A. J. (1971) Notes on some Indo-Pacific Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp. nov., a new pontoniinid shrimp from Mahe, the Seychelle Islands. Crustaceana, 20, 225 - 236. http: // dx. doi. org / 10.1163 / 156854071 x 00012" type="journal article" year="1971">Bruce 1971</bibRefCitation>
: fig. 3a, b); the posterior margin of the telson is acutely pointed at the apex (
<bibRefCitation id="04314B21FFA5F136FE78F9EAFD86F9D7" author="Bruce" box="[477,615,1565,1589]" pageId="6" pageNumber="177" refString="Bruce, A. J. (1971) Notes on some Indo-Pacific Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp. nov., a new pontoniinid shrimp from Mahe, the Seychelle Islands. Crustaceana, 20, 225 - 236. http: // dx. doi. org / 10.1163 / 156854071 x 00012" type="journal article" year="1971">Bruce 1971</bibRefCitation>
: fig. 3c;
<bibRefCitation id="04314B21FFA5F136FD6AF9EAFCCFF9D7" author="Li" box="[719,814,1565,1590]" pageId="6" pageNumber="177" refString="Li, X. (1997) Report on Gnathophyllidae and Pontoniinae (Decapoda, Palaemonoidea) shrimps from the Xisha Islands and adjacent waters, South China Sea. Studia Marina Sinica, 38, 223 - 250." type="journal article" year="1997">Li 1997</bibRefCitation>
: fig.
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). The elongate, unarmed carpus of the second pereopods, which is distinctly longer than the palm of the chela, further separates
<taxonomicName id="A7A04D53FFA5F136FB83F9B6FB54F9BB" box="[1062,1205,1601,1625]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA5F136FB83F9B6FB54F9BB" box="[1062,1205,1601,1625]" italics="true" pageId="6" pageNumber="177">E. gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E757B9FFA5F136FB19F9B7FB1AF9BB" box="[1212,1275,1600,1625]" pageId="6" pageNumber="177" rank="species">
<emphasis id="52D4EAC2FFA5F136FB19F9B7FB1AF9BB" bold="true" box="[1212,1275,1600,1625]" pageId="6" pageNumber="177">n. sp.</emphasis>
</taxonomicNameLabel>
from
<taxonomicName id="A7A04D53FFA5F136FA9BF9B6FA74F9BB" box="[1342,1429,1601,1625]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="oahu">
<emphasis id="52D4EAC2FFA5F136FA9BF9B6FA74F9BB" box="[1342,1429,1601,1625]" italics="true" pageId="6" pageNumber="177">E. oahu</emphasis>
</taxonomicName>
. In
<taxonomicName id="A7A04D53FFA5F136FF10F991FEEDF99F" box="[181,268,1637,1661]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="oahu">
<emphasis id="52D4EAC2FFA5F136FF10F991FEEDF99F" box="[181,268,1637,1661]" italics="true" pageId="6" pageNumber="177">E. oahu</emphasis>
</taxonomicName>
, the carpus of the second pereopod is much shorter than the palm and is armed with a small distomesial tooth (cf.
<bibRefCitation id="04314B21FFA5F136FEA1F97EFE6BF943" author="Bruce" box="[260,394,1673,1697]" pageId="6" pageNumber="177" refString="Bruce, A. J. (2010) Eupontonia oahu sp. nov., a second species of the genus Eupontonia Bruce, 1971, (Crustacea: Decapoda: Pontoniinae) from Oahu, Hawai'ian Islands. In: De Grave, S. &amp; Fransen, C. H. J. M. (Eds.), Contributions to Shrimp Taxonomy. Zootaxa, 2372, 405 - 414." type="journal article" year="2010">Bruce 2010</bibRefCitation>
: fig. 4A, D).
</paragraph>
<paragraph id="601F36D0FFA5F136FF62F95AFF01F89B" blockId="6.[151,1436,1168,2022]" pageId="6" pageNumber="177">
As mentioned above,
<taxonomicName id="A7A04D53FFA5F136FE19F959FDF2F927" box="[444,531,1709,1733]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="oahu">
<emphasis id="52D4EAC2FFA5F136FE19F959FDF2F927" box="[444,531,1709,1733]" italics="true" pageId="6" pageNumber="177">E. oahu</emphasis>
</taxonomicName>
is associated with a scleractinian coral
<taxonomicName id="A7A04D53FFA5F136FC6AF95AFB34F927" box="[975,1237,1709,1733]" class="Anthozoa" family="Pocilloporidae" genus="Pocillopora" kingdom="Animalia" order="Scleractinia" pageId="6" pageNumber="177" phylum="Cnidaria" rank="species" species="meandrina">
<emphasis id="52D4EAC2FFA5F136FC6AF95AFB34F927" box="[975,1237,1709,1733]" italics="true" pageId="6" pageNumber="177">Pocillopora meandrina</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="04314B21FFA5F136FAACF959FA6CF927" author="Bruce" box="[1289,1421,1709,1733]" pageId="6" pageNumber="177" refString="Bruce, A. J. (2010) Eupontonia oahu sp. nov., a second species of the genus Eupontonia Bruce, 1971, (Crustacea: Decapoda: Pontoniinae) from Oahu, Hawai'ian Islands. In: De Grave, S. &amp; Fransen, C. H. J. M. (Eds.), Contributions to Shrimp Taxonomy. Zootaxa, 2372, 405 - 414." type="journal article" year="2010">Bruce 2010</bibRefCitation>
). In contrast, there is no evidence of a symbiotic association for
<taxonomicName id="A7A04D53FFA5F136FCEBF926FC05F90B" box="[846,996,1745,1769]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="noctalbata">
<emphasis id="52D4EAC2FFA5F136FCEBF926FC05F90B" box="[846,996,1745,1769]" italics="true" pageId="6" pageNumber="177">E. noctalbata</emphasis>
</taxonomicName>
and
<taxonomicName id="A7A04D53FFA5F136FBBCF926FB49F90B" box="[1049,1192,1745,1769]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA5F136FBBCF926FB49F90B" box="[1049,1192,1745,1769]" italics="true" pageId="6" pageNumber="177">E. gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E757B9FFA5F136FB0BF926FAE6F90A" box="[1198,1287,1745,1768]" pageId="6" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
, although the
<typeStatus id="BF1B8872FFA5F136FF32F902FF18F8EC" box="[151,249,1781,1806]" pageId="6" pageNumber="177" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A7A04D53FFA5F136FE84F901FE5BF8EF" box="[289,442,1781,1805]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="noctalbata">
<emphasis id="52D4EAC2FFA5F136FE84F901FE5BF8EF" box="[289,442,1781,1805]" italics="true" pageId="6" pageNumber="177">E. noctalbata</emphasis>
</taxonomicName>
was discovered when a colony of the coral
<taxonomicName id="A7A04D53FFA5F136FC11F902FA89F8EC" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" box="[948,1384,1781,1806]" class="Anthozoa" family="Oculinidae" genus="Galaxea" kingdom="Animalia" order="Scleractinia" pageId="6" pageNumber="177" phylum="Cnidaria" rank="species" species="fascicularis">
<emphasis id="52D4EAC2FFA5F136FC11F902FB7FF8EF" box="[948,1182,1781,1805]" italics="true" pageId="6" pageNumber="177">Galaxea fascicularis</emphasis>
(Linnaeus, 1767)
</taxonomicName>
was removed (
<bibRefCitation id="04314B21FFA5F136FEA2F8EEFE6EF8D3" author="Bruce" box="[263,399,1817,1841]" pageId="6" pageNumber="177" refString="Bruce, A. J. (1971) Notes on some Indo-Pacific Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp. nov., a new pontoniinid shrimp from Mahe, the Seychelle Islands. Crustaceana, 20, 225 - 236. http: // dx. doi. org / 10.1163 / 156854071 x 00012" type="journal article" year="1971">Bruce 1971</bibRefCitation>
).
<bibRefCitation id="04314B21FFA5F136FE07F8EEFDD7F8D3" author="Bruce" box="[418,566,1817,1841]" pageId="6" pageNumber="177" refString="Bruce, A. J. (1971) Notes on some Indo-Pacific Pontoniinae. XVII. Eupontonia noctalbata gen. nov., sp. nov., a new pontoniinid shrimp from Mahe, the Seychelle Islands. Crustaceana, 20, 225 - 236. http: // dx. doi. org / 10.1163 / 156854071 x 00012" type="journal article" year="1971">Bruce (1971)</bibRefCitation>
considered that
<taxonomicName id="A7A04D53FFA5F136FD4AF8EEFC64F8D3" box="[751,901,1817,1841]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="noctalbata">
<emphasis id="52D4EAC2FFA5F136FD4AF8EEFC64F8D3" box="[751,901,1817,1841]" italics="true" pageId="6" pageNumber="177">E. noctalbata</emphasis>
</taxonomicName>
was a free-living species. As mentioned above, the
<typeStatus id="BF1B8872FFA5F136FF66F8CAFEC4F8B4" box="[195,293,1853,1878]" pageId="6" pageNumber="177" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A7A04D53FFA5F136FEF5F8C9FE02F8B7" box="[336,483,1853,1877]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA5F136FEF5F8C9FE02F8B7" box="[336,483,1853,1877]" italics="true" pageId="6" pageNumber="177">E. gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E757B9FFA5F136FE49F8CAFDCFF8B4" box="[492,558,1853,1878]" pageId="6" pageNumber="177" rank="species">
<emphasis id="52D4EAC2FFA5F136FE49F8CAFDCFF8B4" bold="true" box="[492,558,1853,1878]" pageId="6" pageNumber="177">n. sp.</emphasis>
</taxonomicNameLabel>
was found under coral rubble, suggesting that the new species is also freeliving.
</paragraph>
<paragraph id="601F36D0FFA5F137FF62F872FD40FF36" blockId="6.[151,1436,1168,2022]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="178" pageId="6" pageNumber="177">
The dorsolateral spines on the telson in the
<typeStatus id="BF1B8872FFA5F136FD0AF872FCF0F87C" box="[687,785,1925,1950]" pageId="6" pageNumber="177" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A7A04D53FFA5F136FC93F871FC27F87F" box="[822,966,1925,1949]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="gracilipes" status="sp. nov.">
<emphasis id="52D4EAC2FFA5F136FC93F871FC27F87F" box="[822,966,1925,1949]" italics="true" pageId="6" pageNumber="177">E. gracilipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E757B9FFA5F136FC68F872FBEDF87C" box="[973,1036,1925,1950]" pageId="6" pageNumber="177" rank="species">
<emphasis id="52D4EAC2FFA5F136FC68F872FBEDF87C" bold="true" box="[973,1036,1925,1950]" pageId="6" pageNumber="177">n. sp.</emphasis>
</taxonomicNameLabel>
are markedly asymmetrical in their arrangement (the left has one spine and the right has two spines) and are considered to be abnormal. It is probable that two pairs of dorsolateral spines are normally present, as in vast majority of pontoniine species. In fact, the nontype specimen of
<taxonomicName id="A7A04D53FFA4F137FEC5FF6EFE19FF52" box="[352,504,152,176]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="noctalbata">
<emphasis id="52D4EAC2FFA4F137FEC5FF6EFE19FF52" box="[352,504,152,176]" italics="true" pageId="7" pageNumber="178">E. noctalbata</emphasis>
</taxonomicName>
reported by
<bibRefCitation id="04314B21FFA4F137FD2FFF60FD14FF52" author="Li" box="[650,757,151,176]" pageId="7" pageNumber="178" refString="Li, X. (1997) Report on Gnathophyllidae and Pontoniinae (Decapoda, Palaemonoidea) shrimps from the Xisha Islands and adjacent waters, South China Sea. Studia Marina Sinica, 38, 223 - 250." type="journal article" year="1997">Li (1997)</bibRefCitation>
and specimens of the congeneric
<taxonomicName id="A7A04D53FFA4F137FBD2FF6EFB2EFF52" box="[1143,1231,152,176]" class="Malacostraca" family="Palaemonidae" genus="Eupontonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="oahu">
<emphasis id="52D4EAC2FFA4F137FBD2FF6EFB2EFF52" box="[1143,1231,152,176]" italics="true" pageId="7" pageNumber="178">E. oahu</emphasis>
</taxonomicName>
have two pairs of dorsolateral spines on the telson (
<bibRefCitation id="04314B21FFA4F137FDA8FF4BFD74FF36" author="Bruce" box="[525,661,188,212]" pageId="7" pageNumber="178" refString="Bruce, A. J. (2010) Eupontonia oahu sp. nov., a second species of the genus Eupontonia Bruce, 1971, (Crustacea: Decapoda: Pontoniinae) from Oahu, Hawai'ian Islands. In: De Grave, S. &amp; Fransen, C. H. J. M. (Eds.), Contributions to Shrimp Taxonomy. Zootaxa, 2372, 405 - 414." type="journal article" year="2010">Bruce 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="28BA655BFFA4F137FF62FF28FC5EFEFF" pageId="7" pageNumber="178" type="etymology">
<paragraph id="601F36D0FFA4F137FF62FF28FC5EFEFF" blockId="7.[151,1436,151,285]" pageId="7" pageNumber="178">
<emphasis id="52D4EAC2FFA4F137FF62FF28FEAFFF1A" bold="true" box="[199,334,223,248]" pageId="7" pageNumber="178">Etymology.</emphasis>
From the combination of the Latin,
<emphasis id="52D4EAC2FFA4F137FD40FF17FCDAFF1A" box="[741,827,224,248]" italics="true" pageId="7" pageNumber="178">gracilis</emphasis>
(= slender) and
<emphasis id="52D4EAC2FFA4F137FC56FF16FBF9FF1A" box="[1011,1048,225,248]" italics="true" pageId="7" pageNumber="178">pes</emphasis>
(= leg), in reference to the slender ambulatory legs (third to fifth pereopods). Used as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

95
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<paragraph id="601F36D0FFA4F137FF32FE87FE1DFE6B" blockId="7.[151,508,367,394]" box="[151,508,367,394]" pageId="7" pageNumber="178">
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Key to species of
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<emphasis id="52D4EAC2FFA4F137FEC8FE98FE1DFE6B" bold="true" box="[365,508,367,393]" italics="true" pageId="7" pageNumber="178">Eupontonia</emphasis>
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1. Carapace with supraorbital tooth................................................................
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<emphasis id="52D4EAC2FFA4F137FABEFE41FA7DFE29" box="[1307,1436,437,459]" italics="true" pageId="7" pageNumber="178">E. noctalbata</emphasis>
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<keyLead id="DB5195E5FFA4F137FF32FE24FA7AFE0B" box="[151,1435,467,489]" pageId="7" pageNumber="178">- Carapace without supraorbital tooth....................................................................... 2</keyLead>
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2. Posteriormost tooth of dorsal rostral series distinctly postrostral; second pereopod with carpus distinctly longer than palm, unarmed...............................................................................
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- Posteriormost tooth of dorsal rostral series not postrostral; second pereopod with carpus distinctly shorter than carpus, armed with small distomesial tooth.........................................................................
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<emphasis id="52D4EAC2FFA4F137FAE8FDBEFA7DFDBC" box="[1357,1436,584,606]" italics="true" pageId="7" pageNumber="178">E. oahu</emphasis>
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54. -
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(fig. 4 b.).
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[[ worker ]]. Long. 1,7 - 1,8 mm. D'un jaune roussatre. Le gastre un peu rembruni. Reticulee et mate; les mandibules et le gastre lisses et luisants avec des stries espacees a la base de celui-ci. Pilosite comme chez
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Sant.
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Tete comme chez
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Sants., un peu plus petite et le bord posterieur un peu moins echancre que chez
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. L'epistome triangulaire plus court que large, a le bord anterieur droit. Mandibules un peu plus courtes que chez
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, leur bord denticule plus long d'un quart que l'epistome. Premier article du funicule tres legerement plus long que le quatrieme, ce dernier environ le double plus long qu'epais. Les deuxieme et troisieme un peu plus epais que longs. Le dernier de l'antenne aussi epais au milieu que la longueur des articles trois et quatre reunis, et aussi long que le reste du funicule. Thorax comme chez
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, la face basale de l'epinotum aussi longue que large, non bordee, sauf entre les epines ou elle est concave. Epines assez fines, longues comme la moitie de leur intervalle. La face declive bordee d'une membrane spongieuse plus distincte que chez
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et
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. Petiole a pedicule aussi long que chez
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, mais avec une bande membraneuse etendue sur toute sa face inferieure; le n oe ud plus arrondi au sommet. Postpetiole le double plus large que long, les cotes arrondis, un peu plus finement reticules que le thorax, avec passablement de tissus spongieux autour et surtout dessous.
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Tres voisine de
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Sants. mais cette derniere a l'epistome convexe devant, la couleur plus pale. Peut etre une simple race de
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.
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<paragraph id="66C4CA3063769D8B26B9711F297CF849" pageNumber="290">Congo belge: Haut Uele, Watsa (L. Burgeon, nov. 1919) (Mus. Tervueren).</paragraph>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Living snail. A Hemiplecta humphreysiana from Singapore (width about 45 mm) B, C Hemiplecta distincta B from Saraburi, Thailand (width about 65 mm) and C mating pairs D, E Hemiplecta funerea from Nan, Thailand (width about 50 mm) D yellow shell form and E dark shell form (width about 50 mm) F Hemiplecta esculenta from Chiang Mai, Thailand (width about 30 mm)." figureDoi="10.3897/zookeys.1047.65735.figure1" httpUri="https://binary.pensoft.net/fig/560488" pageId="0" pageNumber="101">Figures 1B, C</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A, B Hemiplecta humphreysiana A specimen CUMZ 4573 from Singapore and B specimen CUMZ 4648 from Narathivat, Thailand C Hemiplecta distincta, specimen CUMZ 4531 from Chacheongsao, Thailand." figureDoi="10.3897/zookeys.1047.65735.figure2" httpUri="https://binary.pensoft.net/fig/560489" pageId="0" pageNumber="101">, 2C</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Genitalia, mantle edge structure, and caudal region of Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand A whole genital organ B internal wall sculpture of penis, vagina, and dart chamber C dorsal view of mantle edge D ventral view of mantle edge and E right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure4" httpUri="https://binary.pensoft.net/fig/560491" pageId="0" pageNumber="101">, 4</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Representative SEM images of radula A-C Hemiplecta humphreysiana, specimen CUMZ 4573 from Singapore A central and lateral teeth B transition from lateral teeth to marginal teeth and C outermost marginal teeth D, E Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand D central and lateral teeth and E outermost marginal teeth F, G Hemiplecta funerea, specimen CUMZ 4575 from Nan, Thailand F central and lateral teeth and G outermost marginal teeth H, I Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand H central and lateral teeth I transition from lateral teeth to marginal teeth J Hemiplecta nemorosa sp. nov., paratype CUMZ 5253 from Maehongsorn, Thailand. Central tooth indicated by ' C'. Numbers indicate the tooth order from lateral to marginal end." figureDoi="10.3897/zookeys.1047.65735.figure10" httpUri="https://binary.pensoft.net/fig/560497" pageId="0" pageNumber="101">, 10D, E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="reference_group">
<paragraph pageId="0" pageNumber="101">
<taxonomicName authorityName="L. Pfeiffer" authorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta">Helix distincta</taxonomicName>
Pfeiffer, 1850: 69, 70. Type locality: Insulis Moluccis [possibly error or mislabeling].
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Zeitschrift fuer Malakozoologie" pageId="0" pageNumber="101" refId="B89" refString="Pfeiffer, L, 1853. Die Schnirkelschnecken nebst den zunaechst verwandten Gattungen. Dritter Theil. Systematisches Conchylien-Cabinet von Martini und Chemnitz. 1 (12) (3): 291-524. [pp. 291-362, pls 145-156 (1853)] [Published in parts, dates follow Welter-Schultes (1999)]" title="Die Schnirkelschnecken nebst den zunaechst verwandten Gattungen. Dritter Theil. Systematisches Conchylien-Cabinet von Martini und Chemnitz. 1 (12) (3): 291 - 524. [pp. 291 - 362, pls 145 - 156 (1853)] [Published in parts, dates follow Welter-Schultes (1999)]" year="1853">Pfeiffer 1853</bibRefCitation>
: 346, pl. 134, figs 1, 2.
<bibRefCitation author="Reeve, LA" journalOrPublisher="Records of the Zoological Survey of India, Occasional Paper No," pageId="0" pageNumber="101" refId="B104" refString="Reeve, LA, 1854. Conchologia Iconica: Illustrations of the shells of molluscous animals (Vol. 7), Helix. Lovell Reeve &amp; Co., London." title="Conchologia Iconica: Illustrations of the shells of molluscous animals (Vol. 7), Helix. Lovell Reeve &amp; Co., London." year="1854">Reeve 1854</bibRefCitation>
:
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="genus">Helix</taxonomicName>
pl. 86, species 465.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName authorityName="L.Pfeiffer" authorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta">Nanina distincta</taxonomicName>
: Martens, 1860: 7.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName authorityName="L.Pfeiffer" authorityYear="1861" class="Gastropoda" family="Ariophantidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix neptunus" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="neptunus">Helix neptunus</taxonomicName>
Pfeiffer, 1861a: 190. Type locality: Siam [Thailand].
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="101" refId="B95" refString="Pfeiffer, L, 1861b. Novitates Conchologicae. Series prima. Mollusca extramarina. Descriptions et figures de coquilles, extramarines nouvelles, ou peu connues. Beschreibung und Abbildung neuer oder kritischer Land- und Suesswasser Mollusken. Tome 2, Lief. 15, 16: 161-178. [Published in parts, dates follow Johnson (1969)]" title="Novitates Conchologicae. Series prima. Mollusca extramarina. Descriptions et figures de coquilles, extramarines nouvelles, ou peu connues. Beschreibung und Abbildung neuer oder kritischer Land- und Suesswasser Mollusken. Tome 2, Lief. 15, 16: 161 - 178. [Published in parts, dates follow Johnson (1969)]" year="1861 b">Pfeiffer 1861b</bibRefCitation>
: 176, 177. pl. 48, figs 1, 2. New synonym
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName authorityName="L.Pfeiffer" authorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina (Rhyssota) distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta" subGenus="Rhyssota">Nanina (Rhyssota) distincta</taxonomicName>
: Martens, 1867: 69, 70, pl. 6, fig. 8.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName authorityName="L.Pfeiffer" authorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina (Hemiplecta) distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta" subGenus="Hemiplecta">Nanina (Hemiplecta) distincta</taxonomicName>
: Tryon, 1886: 30, pl. 8, fig. 26.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina (Hemiplecta) neptunus" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="neptunus" subGenus="Hemiplecta">Nanina (Hemiplecta) neptunus</taxonomicName>
: Tryon, 1886: 34, pl. 8, fig. 27.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName authorityName="Godwin-Austen" authorityYear="1888" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hemiplecta zimmayensis" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="zimmayensis">Hemiplecta zimmayensis</taxonomicName>
Godwin-Austen, 1888c: 241, 242. Type locality: Zimme, Siam territory [Chiang Mai Province, Thailand]. New synonym
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Ariophanta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ariophanta (Hemiplecta) distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta" subGenus="Hemiplecta">Ariophanta (Hemiplecta) distincta</taxonomicName>
: Morelet, 1891: 231.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hemiplecta distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta">Hemiplecta distincta</taxonomicName>
: Morelet, 1889: 124.
<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a065982" author="Blanford, WT" journalOrPublisher="Proceeding of the Malacological Society of London" pageId="0" pageNumber="101" pagination="274 - 284" refId="B15" refString="Blanford, WT, 1903. Note on Mr. WM Daly's collections of land and freshwater Mollusca from Siam. Proceeding of the Malacological Society of London 5: 274 - 284, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a065982" title="Note on Mr. WM Daly's collections of land and freshwater Mollusca from Siam." url="https://doi.org/10.1093/oxfordjournals.mollus.a065982" volume="5" year="1903">Blanford 1903</bibRefCitation>
: 277, 278.
<bibRefCitation author="Panha, S" journalOrPublisher="Venus" pageId="0" pageNumber="101" pagination="108 - 115" refId="B82" refString="Panha, S, 1987a. Histochemical and ultrastructural studies on the amatorial organ of Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae). Venus 46: 108 - 115" title="Histochemical and ultrastructural studies on the amatorial organ of Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae)." volume="46" year="1987 a">Panha 1987a</bibRefCitation>
: 108-115, figs 1-3.
<bibRefCitation author="Panha, S" journalOrPublisher="Venus" pageId="0" pageNumber="101" pagination="25 - 34" refId="B83" refString="Panha, S, 1987b. The breeding data of Thai edible land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae). Venus 46: 25 - 34" title="The breeding data of Thai edible land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae)." volume="46" year="1987 b">Panha 1987b</bibRefCitation>
: 25-34, fig. 9.
<bibRefCitation author="Panha, S" journalOrPublisher="Venus" pageId="0" pageNumber="101" pagination="197 - 206" refId="B84" refString="Panha, S, 1988a. Effect of temperature on the growth and behavior of a tropical land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae). Venus 47: 197 - 206" title="Effect of temperature on the growth and behavior of a tropical land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae)." volume="47" year="1988 a">Panha 1988a</bibRefCitation>
: 197-206, figs 6, 7.
<bibRefCitation author="Panha, S" journalOrPublisher="Journal of Science Society of Thailand" pageId="0" pageNumber="101" pagination="233 - 239" refId="B85" refString="Panha, S, 1988b. Natural infection of the rat lungworm Angiostrongylus cantonensis in Thai edible land snail, Hemiplecta distincta. Journal of Science Society of Thailand 14: 233 - 239" title="Natural infection of the rat lungworm Angiostrongylus cantonensis in Thai edible land snail, Hemiplecta distincta." volume="14" year="1988 b">Panha 1988b</bibRefCitation>
233-239.
<bibRefCitation author="Panha, S" journalOrPublisher="The Papustyla" pageId="0" pageNumber="101" pagination="4 - 15" refId="B86" refString="Panha, S, 1994. Biology of a Thai edible land snail Hemiplecta distincta (Pfeiffer, 1850) (Gastropoda: Ariophantidae). The Papustyla 8: 4 - 15" title="Biology of a Thai edible land snail Hemiplecta distincta (Pfeiffer, 1850) (Gastropoda: Ariophantidae)." volume="8" year="1994">Panha 1994</bibRefCitation>
: 4-15.
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="101" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
: 76, figs 35b, c, 56c.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName authorityName="L.Pfeiffer" authorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina (Rhysota) distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta" subGenus="Rhysota">Nanina (Rhysota) distincta</taxonomicName>
:
<bibRefCitation author="Fischer, H" journalOrPublisher="In: Mission Pavie, Etudes diverses" pageId="0" pageNumber="101" pagination="390 - 450" refId="B27" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Mission Pavie, Etudes diverses 3: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volume="3" year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
: 393.
<bibRefCitation author="Dautzenberg, P" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="101" pagination="343 - 471" refId="B23" refString="Dautzenberg, P, Fischer, H, 1906. Liste des mollusques recoltes par M. H. Mansuy en Indo-Chine et au Yunnan et description d'especes nouvelles. Journal de Conchyliologie 54: 343 - 471" title="Liste des mollusques recoltes par M. H. Mansuy en Indo-Chine et au Yunnan et description d'especes nouvelles." volume="54" year="1906">Dautzenberg and Fischer 1906</bibRefCitation>
: 346, 347.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina (Rhysota) distincta subsp. var. var. neptunus" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="variety" species="distincta" subGenus="Rhysota" subSpecies="var." variety="neptunus">Nanina (Rhysota) distincta var. neptunus</taxonomicName>
:
<bibRefCitation author="Dautzenberg, P" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="101" pagination="169 - 217" refId="B24" refString="Dautzenberg, P, Fischer, H, 1908. Liste des mollusques recoltes par M. H. Mansuy en Indo-Chine et description d'especes nouvelles. Journal de Conchyliologie 56: 169 - 217" title="Liste des mollusques recoltes par M. H. Mansuy en Indo-Chine et description d'especes nouvelles." volume="56" year="1908">Dautzenberg and Fischer 1908</bibRefCitation>
: 171.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Koratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Koratia distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta">Koratia distincta</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.25931" author="Godwin-Austen, HH" journalOrPublisher="Records of the Indian Museum" pageId="0" pageNumber="101" pagination="199 - 202" refId="B34" refString="Godwin-Austen, HH, 1919. On the position of Helix distincta Pfr. of Siam. Records of the Indian Museum 16: 199 - 202, DOI: https://doi.org/10.5962/bhl.part.25931" title="On the position of Helix distincta Pfr. of Siam." url="https://doi.org/10.5962/bhl.part.25931" volume="16" year="1919">Godwin-Austen 1919</bibRefCitation>
: 199-202, figs 1, 2.
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Materialien zur Naturgeschichte der Insel Celebes" pageId="0" pageNumber="101" refId="B106" refString="Schileyko, AA, 2002. Treatise on Recent terrestrial pulmonate molluscs: Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica, Supplement 2, Part 9: 1167-1307." title="Treatise on Recent terrestrial pulmonate molluscs: Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica, Supplement 2, Part 9: 1167 - 1307." year="2002">Schileyko 2002</bibRefCitation>
: 1282, 1283, fig. 1685.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Nanina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanina (Rhysota) distincta subsp. neptunus" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="subSpecies" species="distincta" subGenus="Rhysota" subSpecies="neptunus">Nanina (Rhysota) distincta neptunus</taxonomicName>
: Dautzenberg and Fischer, 1906: 347, 348.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hemiplecta (Koratia) distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta" subGenus="Koratia">Hemiplecta (Koratia) distincta</taxonomicName>
: Solem, 1966: 27.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hemiplecta (Hemiplecta) distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta" subGenus="Hemiplecta">Hemiplecta (Hemiplecta) distincta</taxonomicName>
: Hemmen and Hemmen, 2001: 44, fig. 12.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hemiplecta (Hemiplecta) neptunus" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="neptunus" subGenus="Hemiplecta">Hemiplecta (Hemiplecta) neptunus</taxonomicName>
: Hemmen and Hemmen, 2001: 44.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName authorityName="Godwin-Austen" authorityYear="1888" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hemiplecta (Hemiplecta) zimmayensis" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="zimmayensis" subGenus="Hemiplecta">Hemiplecta (Hemiplecta) zimmayensis</taxonomicName>
: Hemmen and Hemmen, 2001: 44.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="type specimen">
<paragraph pageId="0" pageNumber="101">Type specimen.</paragraph>
<paragraph pageId="0" pageNumber="101">
See the species list of Indochinese species (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Hemiplecta auriettae, specimen NHMUK 1912.4.16.497 B Hemiplecta cymatium, specimen UMZC I. 104350 ex. Benson collection C Hemiplecta distincta, possible syntype NHMUK 20200199 D Hemiplecta floweri, syntype NHMUK 1899.3.16.1 - 2." figureDoi="10.3897/zookeys.1047.65735.figure11" httpUri="https://binary.pensoft.net/fig/560498" pageId="0" pageNumber="101">11C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="materials_examined">
<paragraph pageId="0" pageNumber="101">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="101">
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Thailand" county="Klong Lann National Park" location="Sam Larn National Park" municipality="Huay Kla Klang National Park" specimenCount="6" stateProvince="Chiang Mai">
<emphasis bold="true" pageId="0" pageNumber="101">
<collectingCountry name="Thailand">Thailand</collectingCountry>
</emphasis>
: Tam Chiang Dao,
<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
: CUMZ 4550 (1 shell), CUMZ 4558 (5 shells). Wang Chao Waterfall, Kampangphet: CUMZ 4641 (4 shells).
<collectingCounty>Klong Lann National Park</collectingCounty>
, Kampangphet: CUMZ 4579 (
<specimenCount type="generic">2 specimens</specimenCount>
in ethanol). Kaeng Jed Kwae, Watbot, Phitsanuloke: CUMZ 4638 (7 shells). Tam Wang Daeng, Nern Maprang, Phitsanuloke: CUMZ 4632 (1 shell). Khao Nang Rum,
<collectingMunicipality>Huay Kla Klang National Park</collectingMunicipality>
, Uthaithani: CUMZ 4502 (6 shells), CUMZ 4510 (3 shells), CUMZ 4538 (3 shells), CUMZ 4541 (6 shells), CUMZ 4607 (6 shells), CUMZ 4610 (9 shells), CUMZ 4611 (3 shells). Jed Sow Noi Waterfall, Muaklek,
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
: CUMZ 4548 (3 shells). Pu Kare Botanic Garden,
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
: CUMZ 4505 (4 shells), CUMZ 4506 (1 shell), CUMZ 4534 (5 shells). Tam Dao Khao Kaew, Muaklek,
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
: CUMZ 4624 (1 shell). Wat Tharahat,
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
: CUMZ 4508 (1 shell), 4530 (1 shell).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:7AEFDE7CD04DE6372F4971BB24338920" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Sam Larn National Park" stateProvince="Chiang Mai">Sam Larn National Park</location>
,
<collectingRegion country="Thailand" name="Sara Buri">Saraburi</collectingRegion>
: CUMZ 4578 (
<specimenCount type="generic">2 specimens</specimenCount>
in ethanol). Bang Srithong, Bang Kruay,
<collectingRegion country="Thailand" name="Nonthaburi">Nonthaburi</collectingRegion>
: CUMZ 4555 (5 shells). Khao Look Chang, Pakchong, Nakhonratchasima: CUMZ 4501 (8 shells), CUMZ 4606 (9 shells), CUMZ 4612 (9 shells), CUMZ 4535 (3 shells).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:EEDE1074BBAE2FCB1FDEF1B6D7C1EFE6" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Tub Lann National Park" stateProvince="Chiang Mai">Tub Lann National Park</location>
, Nakhonratchasima: CUMZ 4617 (1 shell). Nawang, Nongbualumphu: CUMZ 4529 (1 shell). Tam Suwankuha, Nongbualumphu: CUMZ 4633 (3 shells), 4637 (3 shells). Thung Kra-Mang, Phu Kiew Wildlife Sanctuary,
<collectingRegion country="Thailand" name="Chaiyaphum">Chaiyaphum</collectingRegion>
: CUMZ 4608 (5 shells). Pang Khone, Sakonnakhon: CUMZ 4619 (4 shells).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:46B012506C3511F10A80551F7F941B31" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Phuphan Mountains" stateProvince="Chiang Mai">Phuphan Mountains</location>
, Sakonnakhon: CUMZ 4504 (6 shells), CUMZ 4507 (5 shells). Phu Kum Khao, Sahatsakhan,
<collectingRegion country="Thailand" name="Kalasin">Kalasin</collectingRegion>
: CUMZ 4557 (8 shells). Phu Sri Tharn Wildlife Sanctuary,
<collectingRegion country="Thailand" name="Kalasin">Kalasin</collectingRegion>
: CUMZ 4621 (1 shell). Huay Lao Waterfall, Phuluang Wildlife Sanctuary,
<collectingRegion country="Thailand" name="Loei">Loei</collectingRegion>
: CUMZ 4634 (1 shell). Tam Pha Bend, Chiang Karn,
<collectingRegion country="Thailand" name="Loei">Loei</collectingRegion>
: CUMZ 4532 (1 shell). Tam Pha Bing, Wangsapung,
<collectingRegion country="Thailand" name="Loei">Loei</collectingRegion>
: CUMZ 4636 (1 shell). Tam Piya,
<collectingRegion country="Thailand" name="Loei">Loei</collectingRegion>
: CUMZ 4639 (3 shells). Tam Mahasombat, Lomsak,
<collectingRegion country="Thailand" name="Phetchabun">Phetchabun</collectingRegion>
: CUMZ 4567 (1 shell). Tam Yai Namnao,
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:3795E57E801F6788E5B5207B1180F794" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Namnao National Park" stateProvince="Chiang Mai">Namnao National Park</location>
,
<collectingRegion country="Thailand" name="Phetchabun">Phetchabun</collectingRegion>
: CUMZ 4566 (1 shell), CUMZ 4622 (1 shell). Tam Phraya Nakarat,
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:9E5899F5317748DFA263C83DF5BC5B7D" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Phuphaman National Park" stateProvince="Chiang Mai">Phuphaman National Park</location>
, Khonkaen: CUMZ 4635 (1 shell).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:73A8ABB53BE92FF95F97A08EF161FFDF" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Pha Tam National Park" stateProvince="Chiang Mai">Pha Tam National Park</location>
, Ubonratchathani: CUMZ 4604 (3 shells), CUMZ 4616 (3 shells).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:B051BE83F2302BB2020CA797C19A5BB9" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Yod Dome National Park" stateProvince="Chiang Mai">Yod Dome National Park</location>
,
<collectingRegion country="Thailand" name="Buriram">Buriram</collectingRegion>
: CUMZ 4629 (2 shells). Wang Ta Krai Waterfall, Nakhonnayok: CUMZ 4540 (2 shells), CUMZ 4549 (2 shells), CUMZ 4605 (1 shell), CUMZ 4640 (5 shells), CUMZ 4577 (
<specimenCount type="generic">1 specimen</specimenCount>
in ethanol). Khao Ang Rue Nai Wildlife Sanctuary,
<collectingRegion country="Thailand" name="Chachoengsao">Chachoengsao</collectingRegion>
: CUMZ 4531 (1 shell; Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A, B Hemiplecta humphreysiana A specimen CUMZ 4573 from Singapore and B specimen CUMZ 4648 from Narathivat, Thailand C Hemiplecta distincta, specimen CUMZ 4531 from Chacheongsao, Thailand." figureDoi="10.3897/zookeys.1047.65735.figure2" httpUri="https://binary.pensoft.net/fig/560489" pageId="0" pageNumber="101">2B</figureCitation>
), CUMZ 4546 (7 shells), CUMZ 4609 (3 shells), CUMZ 4613 (4 shells), CUMZ 4620 (2 shells), CUMZ 4627 (1 shell), CUMZ 4630 (1 shell).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:854D7E01D1DE901A3AB474591CA0A78F" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Pang Srida National Park" stateProvince="Chiang Mai">Pang Srida National Park</location>
,
<collectingRegion country="Thailand" name="Prachin Buri">Prachinburi</collectingRegion>
: CUMZ 4631 (4 shells). Ra-Ru, Taphraya, Srakeow: CUMZ 4628 (1 shell). Tam Leum, Klonghad, Srakeow: CUMZ 4625 (1 shell). Khao Cha Ang-Oan, Borthong,
<collectingRegion country="Thailand" name="Chon Buri">Chonburi</collectingRegion>
: CUMZ 4542 (4 shells), CUMZ 4618 (1 shell), CUMZ 4626 (4 shells). Khao Cha Mao Waterfall,
<collectingRegion country="Thailand" name="Rayong">Rayong</collectingRegion>
: CUMZ 4543 (1 shell). Tam Suwanphupha, Khao Chamao,
<collectingRegion country="Thailand" name="Rayong">Rayong</collectingRegion>
: CUMZ 4545 (1 shell). Wat Ma-deau (Tam Khao Loi), Khao Chamao,
<collectingRegion country="Thailand" name="Rayong">Rayong</collectingRegion>
: CUMZ 4544 (4 shells).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:BEB5FA84A9890E7F6EAF0A3FA6F04E5D" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Plieu National Park" stateProvince="Chiang Mai">Plieu National Park</location>
,
<collectingRegion country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
: CUMZ 4509 (1 shell), CUMZ 4536 (1 shell), CUMZ 4539 (9 shells), CUMZ 4560 (
<specimenCount type="generic">1 specimen</specimenCount>
in ethanol; Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Genitalia, mantle edge structure, and caudal region of Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand A whole genital organ B internal wall sculpture of penis, vagina, and dart chamber C dorsal view of mantle edge D ventral view of mantle edge and E right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure4" httpUri="https://binary.pensoft.net/fig/560491" pageId="0" pageNumber="101">4</figureCitation>
), CUMZ 4601 (1 shell), CUMZ 4615 (4 shells).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:73F16538C299540BB2713FF85C5D499C" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Sibha Shan Waterfall" stateProvince="Chiang Mai">Sibha Shan Waterfall</location>
,
<collectingRegion country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
: CUMZ 4547 (6 shells).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:4AB89085755439F420FFCE48F5E75651" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Tha Mai District" stateProvince="Chiang Mai">Tha Mai District</location>
,
<collectingRegion country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
: CUMZ 4603 (1 shell).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:8DBB13BF1ACA36974789F9DCF67A9958" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Koh Kud" stateProvince="Chiang Mai">Koh Kud</location>
,
<collectingRegion country="Thailand" name="Trat">Trat</collectingRegion>
: CUMZ 4559 (9 shells), CUMZ 4614 (7 shells).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:2B08638875CB1B0524C7215541FC2950" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Kaeng Kracharn National Park" stateProvince="Chiang Mai">Kaeng Kracharn National Park</location>
,
<collectingRegion country="Thailand" name="Phetchaburi">Phetchaburi</collectingRegion>
: CUMZ 4527 (1 shell).
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:59879A8A69163CCE5ADF470B73510CD9" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Tam Nam Pud" stateProvince="Chiang Mai">Tam Nam Pud</location>
,
<location LSID="urn:lsid:plazi:treatment:E80A2E8DC02D5D248AF307DD2235C9DF:281E10542652F826A9B8F44A8F398C19" country="Thailand" county="Klong Lann National Park" municipality="Huay Kla Klang National Park" name="Pangnga" stateProvince="Chiang Mai">Pangnga</location>
: CUMZ 4623 (1 shell)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1047.65735.figure4" httpUri="https://binary.pensoft.net/fig/560491" pageId="0" pageNumber="101" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" pageId="0" pageNumber="101">Figure 4.</emphasis>
Genitalia, mantle edge structure, and caudal region of
<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" lsidName="Hemiplecta distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta">
<emphasis italics="true" pageId="0" pageNumber="101">Hemiplecta distincta</emphasis>
</taxonomicName>
, specimen CUMZ 4560 from
<collectingRegion country="Thailand" name="Chanthaburi">Chanthaburi</collectingRegion>
,
<collectingCountry name="Thailand">Thailand</collectingCountry>
<emphasis bold="true" pageId="0" pageNumber="101">A</emphasis>
whole genital organ
<emphasis bold="true" pageId="0" pageNumber="101">B</emphasis>
internal wall sculpture of penis, vagina, and dart chamber
<emphasis bold="true" pageId="0" pageNumber="101">C</emphasis>
dorsal view of mantle edge
<emphasis bold="true" pageId="0" pageNumber="101">D</emphasis>
ventral view of mantle edge and
<emphasis bold="true" pageId="0" pageNumber="101">E</emphasis>
right view of caudal region.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" italics="true" pageId="0" pageNumber="101">Shell.</emphasis>
Shell large (height up to
<quantity metricMagnitude="-2" metricUnit="m" metricValue="5.5" unit="mm" value="55.0">55 mm</quantity>
, width up to
<quantity metricMagnitude="-2" metricUnit="m" metricValue="7.5" unit="mm" value="75.0">75 mm</quantity>
), yellowish with white narrow peripheral band, and paler color below on lower shell surface. Upper shell surface with thin growth lines interrupted with spiral wrinkles. Last whorl large and rounded; aperture large ovate; lip simple but slightly thickened in adult snails (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A, B Hemiplecta humphreysiana A specimen CUMZ 4573 from Singapore and B specimen CUMZ 4648 from Narathivat, Thailand C Hemiplecta distincta, specimen CUMZ 4531 from Chacheongsao, Thailand." figureDoi="10.3897/zookeys.1047.65735.figure2" httpUri="https://binary.pensoft.net/fig/560489" pageId="0" pageNumber="101">2C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" italics="true" pageId="0" pageNumber="101">Genitalia.</emphasis>
The external genital organs were described in
<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a065776" author="Godwin-Austen, HH" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="101" pagination="31 - 36" refId="B33" refString="Godwin-Austen, HH, 1900. On the anatomy of Hemiplecta floweri E.A. Smith, from Perak, Malay Peninsula; with notes on some other eastern genera. Proceedings of the Malacological Society of London 4: 31 - 36, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a065776" title="On the anatomy of Hemiplecta floweri E. A. Smith, from Perak, Malay Peninsula; with notes on some other eastern genera." url="https://doi.org/10.1093/oxfordjournals.mollus.a065776" volume="4" year="1900">Godwin-Austen (1900</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.25931" author="Godwin-Austen, HH" journalOrPublisher="Records of the Indian Museum" pageId="0" pageNumber="101" pagination="199 - 202" refId="B34" refString="Godwin-Austen, HH, 1919. On the position of Helix distincta Pfr. of Siam. Records of the Indian Museum 16: 199 - 202, DOI: https://doi.org/10.5962/bhl.part.25931" title="On the position of Helix distincta Pfr. of Siam." url="https://doi.org/10.5962/bhl.part.25931" volume="16" year="1919">1919</bibRefCitation>
). Gametolytic sac (gs) bulbous with undifferentiated duct. Internal wall of penis exhibits closely packed papilla knobs that abruptly cease near atrium; penial verge absent. Internal sculpture of vagina with thin and smooth longitudinal vaginal pilasters (vp). Internal surface of dart apparatus smooth; dart papilla (dp) conic, and with a smooth surface (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Genitalia, mantle edge structure, and caudal region of Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand A whole genital organ B internal wall sculpture of penis, vagina, and dart chamber C dorsal view of mantle edge D ventral view of mantle edge and E right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure4" httpUri="https://binary.pensoft.net/fig/560491" pageId="0" pageNumber="101">4A, B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" italics="true" pageId="0" pageNumber="101">Radula.</emphasis>
Each row with about 543 teeth (276-(15-20)-1-(15-20)-276). Central tooth unicuspid triangular with rounded head (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Representative SEM images of radula A-C Hemiplecta humphreysiana, specimen CUMZ 4573 from Singapore A central and lateral teeth B transition from lateral teeth to marginal teeth and C outermost marginal teeth D, E Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand D central and lateral teeth and E outermost marginal teeth F, G Hemiplecta funerea, specimen CUMZ 4575 from Nan, Thailand F central and lateral teeth and G outermost marginal teeth H, I Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand H central and lateral teeth I transition from lateral teeth to marginal teeth J Hemiplecta nemorosa sp. nov., paratype CUMZ 5253 from Maehongsorn, Thailand. Central tooth indicated by ' C'. Numbers indicate the tooth order from lateral to marginal end." figureDoi="10.3897/zookeys.1047.65735.figure10" httpUri="https://binary.pensoft.net/fig/560497" pageId="0" pageNumber="101">10D</figureCitation>
). Lateral teeth unicuspid, oblique and triangular. Outer lateral teeth unicuspid, sickle-shaped, with transition to curved and narrow sickle form; latero-marginal transition starts from tooth numbers 15 to 20. Marginal teeth narrowly curved, bicuspid; endocone and ectocone small and pointed (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Representative SEM images of radula A-C Hemiplecta humphreysiana, specimen CUMZ 4573 from Singapore A central and lateral teeth B transition from lateral teeth to marginal teeth and C outermost marginal teeth D, E Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand D central and lateral teeth and E outermost marginal teeth F, G Hemiplecta funerea, specimen CUMZ 4575 from Nan, Thailand F central and lateral teeth and G outermost marginal teeth H, I Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand H central and lateral teeth I transition from lateral teeth to marginal teeth J Hemiplecta nemorosa sp. nov., paratype CUMZ 5253 from Maehongsorn, Thailand. Central tooth indicated by ' C'. Numbers indicate the tooth order from lateral to marginal end." figureDoi="10.3897/zookeys.1047.65735.figure10" httpUri="https://binary.pensoft.net/fig/560497" pageId="0" pageNumber="101">10E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" italics="true" pageId="0" pageNumber="101">External features.</emphasis>
Living snails have a similar soft body morphology and pulmonary cavity to that of
<taxonomicName higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. humphreysiana" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="humphreysiana">
<emphasis italics="true" pageId="0" pageNumber="101">H. humphreysiana</emphasis>
</taxonomicName>
. The distinct characters are pale brown to brownish body (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Living snail. A Hemiplecta humphreysiana from Singapore (width about 45 mm) B, C Hemiplecta distincta B from Saraburi, Thailand (width about 65 mm) and C mating pairs D, E Hemiplecta funerea from Nan, Thailand (width about 50 mm) D yellow shell form and E dark shell form (width about 50 mm) F Hemiplecta esculenta from Chiang Mai, Thailand (width about 30 mm)." figureDoi="10.3897/zookeys.1047.65735.figure1" httpUri="https://binary.pensoft.net/fig/560488" pageId="0" pageNumber="101">1B, C</figureCitation>
). Sole of foot brownish; caudal horn not overhanging; caudal foss a long vertical slit arranged on tail above sole margin. Pedal groove typical aulacopod and well defined (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Genitalia, mantle edge structure, and caudal region of Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand A whole genital organ B internal wall sculpture of penis, vagina, and dart chamber C dorsal view of mantle edge D ventral view of mantle edge and E right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure4" httpUri="https://binary.pensoft.net/fig/560491" pageId="0" pageNumber="101">4E</figureCitation>
). Mantle edge narrow with large dorsal lobes. Right dorsal lobe (rdl) to right of pneumostome, large and thick; left dorsal lobe to left of pneumostome, composed of anterior left dorsal lobe (aldl) and posterior left dorsal lobe (pldl); shell lobe absent (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Genitalia, mantle edge structure, and caudal region of Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand A whole genital organ B internal wall sculpture of penis, vagina, and dart chamber C dorsal view of mantle edge D ventral view of mantle edge and E right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure4" httpUri="https://binary.pensoft.net/fig/560491" pageId="0" pageNumber="101">4C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="distribution">
<paragraph pageId="0" pageNumber="101">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="101">
Ranges from Cambodia to Laos, Thailand and southern Vietnam (
<bibRefCitation DOI="https://doi.org/10.1080/00222939608680338" author="Smith, EA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="101" refId="B111" refString="Smith, EA, 1896. Notes on some land shells from Vanbu, Tonkin, with description of two new species. Annals and Magazine of Natural History, Series 6, 17: 128-130. https://doi.org/10.1080/00222939608680338" title="Notes on some land shells from Vanbu, Tonkin, with description of two new species. Annals and Magazine of Natural History, Series 6, 17: 128 - 130." url="https://doi.org/10.1080/00222939608680338" year="1896">Smith 1896</bibRefCitation>
;
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Proceeding of the Malacological Society of London" pageId="0" pageNumber="101" pagination="80 - 94" refId="B54" refString="Laidlaw, FF, 1932a. Notes on Ariophantidae from the Malay Peninsula, with descriptions of new genera. Proceeding of the Malacological Society of London 20: 80 - 94" title="Notes on Ariophantidae from the Malay Peninsula, with descriptions of new genera." volume="20" year="1932 a">Laidlaw 1932a</bibRefCitation>
;
<bibRefCitation author="Panha, S" journalOrPublisher="Venus" pageId="0" pageNumber="101" pagination="197 - 206" refId="B84" refString="Panha, S, 1988a. Effect of temperature on the growth and behavior of a tropical land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae). Venus 47: 197 - 206" title="Effect of temperature on the growth and behavior of a tropical land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae)." volume="47" year="1988 a">Panha 1988a</bibRefCitation>
,
<bibRefCitation author="Panha, S" journalOrPublisher="The Papustyla" pageId="0" pageNumber="101" pagination="4 - 15" refId="B86" refString="Panha, S, 1994. Biology of a Thai edible land snail Hemiplecta distincta (Pfeiffer, 1850) (Gastropoda: Ariophantidae). The Papustyla 8: 4 - 15" title="Biology of a Thai edible land snail Hemiplecta distincta (Pfeiffer, 1850) (Gastropoda: Ariophantidae)." volume="8" year="1994">1994</bibRefCitation>
;
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="101" pagination="1 - 68" refId="B108" refString="Schileyko, AA, 2011. Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica 21: 1 - 68" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="21" year="2011">Schileyko 2011</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="101" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
). In Thailand,
<taxonomicName higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. distincta" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta">
<emphasis italics="true" pageId="0" pageNumber="101">H. distincta</emphasis>
</taxonomicName>
is fairly abundant and occurs throughout the country, except for southern Thailand (
<bibRefCitation author="Panha, S" journalOrPublisher="Journal of Science Society of Thailand" pageId="0" pageNumber="101" pagination="233 - 239" refId="B85" refString="Panha, S, 1988b. Natural infection of the rat lungworm Angiostrongylus cantonensis in Thai edible land snail, Hemiplecta distincta. Journal of Science Society of Thailand 14: 233 - 239" title="Natural infection of the rat lungworm Angiostrongylus cantonensis in Thai edible land snail, Hemiplecta distincta." volume="14" year="1988 b">Panha 1988b</bibRefCitation>
,
<bibRefCitation author="Panha, S" journalOrPublisher="The Papustyla" pageId="0" pageNumber="101" pagination="4 - 15" refId="B86" refString="Panha, S, 1994. Biology of a Thai edible land snail Hemiplecta distincta (Pfeiffer, 1850) (Gastropoda: Ariophantidae). The Papustyla 8: 4 - 15" title="Biology of a Thai edible land snail Hemiplecta distincta (Pfeiffer, 1850) (Gastropoda: Ariophantidae)." volume="8" year="1994">1994</bibRefCitation>
). The southern limit of the species appears to be near the Isthmus of Kra (10°N). We have a single and old shell from Pangnga Province, southern Thailand that needs to be confirmed.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="remarks">
<paragraph pageId="0" pageNumber="101">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="101">
The type specimens of
<taxonomicName authorityName="L.Pfeiffer" authorityYear="1861" class="Gastropoda" family="Ariophantidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix neptunus" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="neptunus">
<emphasis italics="true" pageId="0" pageNumber="101">Helix neptunus</emphasis>
</taxonomicName>
Pfeiffer, 1861 and
<taxonomicName authorityName="Godwin-Austen" authorityYear="1888" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" lsidName="Hemiplecta zimmayensis" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="zimmayensis">
<emphasis italics="true" pageId="0" pageNumber="101">Hemiplecta zimmayensis</emphasis>
</taxonomicName>
Godwin-Austen, 1888 exhibit a shell morphology and color patterns identical to that of
<taxonomicName higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. distincta" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta">
<emphasis italics="true" pageId="0" pageNumber="101">H. distincta</emphasis>
</taxonomicName>
. The absence of a whitish peripheral band in
<taxonomicName authorityName="L.Pfeiffer" authorityYear="1861" class="Gastropoda" family="Ariophantidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix neptunus" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="neptunus">
<emphasis italics="true" pageId="0" pageNumber="101">Helix neptunus</emphasis>
</taxonomicName>
Pfeiffer, 1861 and the strong growth lines of
<taxonomicName authorityName="Godwin-Austen" authorityYear="1888" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" lsidName="Hemiplecta zimmayensis" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="zimmayensis">
<emphasis italics="true" pageId="0" pageNumber="101">Hemiplecta zimmayensis</emphasis>
</taxonomicName>
are the only observed differences from
<taxonomicName higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. distincta" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta">
<emphasis italics="true" pageId="0" pageNumber="101">H. distincta</emphasis>
</taxonomicName>
. Therefore, we recognize these two nominal species as junior subjective synonyms of
<taxonomicName higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. distincta" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta">
<emphasis italics="true" pageId="0" pageNumber="101">H. distincta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="101">
<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" lsidName="Hemiplecta distincta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="distincta">
<emphasis italics="true" pageId="0" pageNumber="101">Hemiplecta distincta</emphasis>
</taxonomicName>
has long been considered a food item for local people in northeastern Thailand (
<bibRefCitation author="Panha, S" journalOrPublisher="Venus" pageId="0" pageNumber="101" pagination="25 - 34" refId="B83" refString="Panha, S, 1987b. The breeding data of Thai edible land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae). Venus 46: 25 - 34" title="The breeding data of Thai edible land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae)." volume="46" year="1987 b">Panha 1987b</bibRefCitation>
), as well as in Cambodia and Laos (personal observation). It has also been found to be an intermediate host of the rat lungworm, a human pathogen (
<bibRefCitation author="Panha, S" journalOrPublisher="Venus" pageId="0" pageNumber="101" pagination="25 - 34" refId="B83" refString="Panha, S, 1987b. The breeding data of Thai edible land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae). Venus 46: 25 - 34" title="The breeding data of Thai edible land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae)." volume="46" year="1987 b">Panha 1987b</bibRefCitation>
,
<bibRefCitation author="Panha, S" journalOrPublisher="Journal of Science Society of Thailand" pageId="0" pageNumber="101" pagination="233 - 239" refId="B85" refString="Panha, S, 1988b. Natural infection of the rat lungworm Angiostrongylus cantonensis in Thai edible land snail, Hemiplecta distincta. Journal of Science Society of Thailand 14: 233 - 239" title="Natural infection of the rat lungworm Angiostrongylus cantonensis in Thai edible land snail, Hemiplecta distincta." volume="14" year="1988 b">1988b</bibRefCitation>
). The life cycle and breeding biology of this species have been extensively studied (
<bibRefCitation author="Panha, S" journalOrPublisher="Venus" pageId="0" pageNumber="101" pagination="108 - 115" refId="B82" refString="Panha, S, 1987a. Histochemical and ultrastructural studies on the amatorial organ of Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae). Venus 46: 108 - 115" title="Histochemical and ultrastructural studies on the amatorial organ of Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae)." volume="46" year="1987 a">Panha 1987a</bibRefCitation>
,
<bibRefCitation author="Panha, S" journalOrPublisher="Venus" pageId="0" pageNumber="101" pagination="25 - 34" refId="B83" refString="Panha, S, 1987b. The breeding data of Thai edible land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae). Venus 46: 25 - 34" title="The breeding data of Thai edible land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae)." volume="46" year="1987 b">b</bibRefCitation>
,
<bibRefCitation author="Panha, S" journalOrPublisher="Venus" pageId="0" pageNumber="101" pagination="197 - 206" refId="B84" refString="Panha, S, 1988a. Effect of temperature on the growth and behavior of a tropical land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae). Venus 47: 197 - 206" title="Effect of temperature on the growth and behavior of a tropical land snail Hemiplecta distincta (Pfeiffer) (Pulmonata: Ariophantidae)." volume="47" year="1988 a">1988a</bibRefCitation>
,
<bibRefCitation author="Panha, S" journalOrPublisher="Journal of Science Society of Thailand" pageId="0" pageNumber="101" pagination="233 - 239" refId="B85" refString="Panha, S, 1988b. Natural infection of the rat lungworm Angiostrongylus cantonensis in Thai edible land snail, Hemiplecta distincta. Journal of Science Society of Thailand 14: 233 - 239" title="Natural infection of the rat lungworm Angiostrongylus cantonensis in Thai edible land snail, Hemiplecta distincta." volume="14" year="1988 b">b</bibRefCitation>
,
<bibRefCitation author="Panha, S" journalOrPublisher="The Papustyla" pageId="0" pageNumber="101" pagination="4 - 15" refId="B86" refString="Panha, S, 1994. Biology of a Thai edible land snail Hemiplecta distincta (Pfeiffer, 1850) (Gastropoda: Ariophantidae). The Papustyla 8: 4 - 15" title="Biology of a Thai edible land snail Hemiplecta distincta (Pfeiffer, 1850) (Gastropoda: Ariophantidae)." volume="8" year="1994">1994</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

337
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<treatment id="E80A6E4C17DDDE8BC49C391737F630A7" ID-GBIF-Taxon="231546687" LSID="urn:lsid:plazi:treatment:E80A6E4C17DDDE8BC49C391737F630A7" httpUri="http://treatment.plazi.org/id/E80A6E4C17DDDE8BC49C391737F630A7" lastPageNumber="1551" pageNumber="1551" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=13400135">
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<paragraph id="02B97075928D3CED0ED8E1F6D92916B9" pageNumber="1551">
<taxonomicName id="E030DC7EB0A7A4B92493DD2AEDDCBD63" ID-CoL="63832" authority="Meyen 1833" authorityName="Meyen" authorityYear="1833" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="genus">
<emphasis id="B76CB633F28BB20146584F3F0DD77D16" italics="true" pageNumber="1551">Lagidium</emphasis>
Meyen 1833
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D8C214F369C37FFF94B3E3087FE36A9C" pageNumber="1551" type="reference_group">
<paragraph id="0855EC4E7BADA10F3ED05655D368090B" pageNumber="1551">
<treatmentCitation id="325515E85CA122A7F4C9541F8E02BFDB" author="Meyen" authority="Meyen 1833" authorityName="Meyen" authorityYear="1833" class="Mammalia" family="Chinchillidae" genus="Lagidium" issue="2" journal="Nova. Acta Acad. Caes. Leop.-Carol." kingdom="Animalia" order="Rodentia" page="576" pageNumber="1551" phylum="Chordata" rank="genus" title="Lagidium" volume="16" year="1833">
<taxonomicName id="637BFBE420FE29923ADD3870E5B59F3A" ID-CoL="63832" authority="Meyen 1833" authorityName="Meyen" authorityYear="1833" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="genus">
<emphasis id="3F2CE9B7AEB34A917B37AC65A693C41E" italics="true" pageNumber="1551">Lagidium</emphasis>
Meyen 1833
</taxonomicName>
,
<bibCitation id="B8675701B22080F6831AB926AE27BBA8" author="Meyen" issue="2" journal="Nova. Acta Acad. Caes. Leop.-Carol." pageNumber="1551" pagination="576" title="Lagidium" volume="16" year="1833">Nova. Acta Acad. Caes. Leop.-Carol., 16 (2): 576</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6289F405087D3B70625C247D44085A35" pageNumber="1551" type="type_taxon">
<paragraph id="C1C5F5488F232410DD6446B97DF8CE39" pageNumber="1551">
<emphasis id="87BC1B438BBC22EDCEF2A0FD1B971631" inLineHeading="true" pageNumber="1551">Type Species:</emphasis>
<taxonomicName id="E56DCB6DBF1800118457C7DE504980E0" authority="Meyen 1833" authorityName="Meyen" authorityYear="1833" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="species" species="peruanum">
<emphasis id="C0445113DF92EA8967825BB6E8E37F2D" italics="true" pageNumber="1551">Lagidium peruanum</emphasis>
Meyen 1833
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="7232D2A4EB4E25908F560B48E1185B34" pageNumber="1551" type="synonym_list">
<paragraph id="E6AE3A21D9157809F8E5AFADB84994E6" pageNumber="1551">
<emphasis id="C2A21E0F7D98BE4CC128EBE856DA2F45" inLineHeading="true" pageNumber="1551">Synonyms:</emphasis>
<taxonomicName id="08EEDC1F5A6A29D321261FBE11202B73" authority="Bennett 1833" authorityName="Bennett" authorityYear="1833" class="Mammalia" family="Chinchillidae" genus="Lagotis" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="genus">
<emphasis id="D30E80F1E6ECF5CB84C5DBBF7A2AF850" italics="true" pageNumber="1551">Lagotis</emphasis>
Bennett 1833
</taxonomicName>
;
<taxonomicName id="7FD3390D459DB2285D9F2D9DD8D6B697" authority="Oken 1816" authorityName="Oken" authorityYear="1816" class="Mammalia" family="Chinchillidae" genus="Viscaccia" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="genus">
<emphasis id="D727021F667C6C1E29C05EC89B48904E" italics="true" pageNumber="1551">Viscaccia</emphasis>
Oken 1816
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="64BC3E4B7E31FAF32F817EE38011A6D0" pageNumber="1551" type="child_taxa">
<paragraph id="2BBFAE4B74DEF525439D101EC17E65DA" pageNumber="1551">
<emphasis id="8553D77D0D4F92BDEC66BD3912B578C9" inLineHeading="true" pageNumber="1551">Species and subspecies:</emphasis>
3 species with 20 subspecies:
</paragraph>
<paragraph id="3C7B5F4508B9DF6EAC38DA09F1ACB5E4" pageNumber="1551">
Species
<taxonomicName id="42731F639256907FEC88BC3C00432E45" authority="Meyen 1833" authorityName="Meyen" authorityYear="1833" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="species" species="peruanum">
<emphasis id="7FD0162CA55B093E3CBB7C3378FFE0C1" italics="true" pageNumber="1551">Lagidium peruanum</emphasis>
Meyen 1833
</taxonomicName>
</paragraph>
<paragraph id="1E5ABB89827966D37276A1213A40FD7C" pageNumber="1551">
Subspecies
<taxonomicName id="6AB6A23A290AF912676CC7DC3DCCE6B5" authority="Meyen 1833" authorityName="Meyen" authorityYear="1833" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="peruanum" subSpecies="peruanum">
<emphasis id="9A42CACF01AB9C7FB113C23F4C040B15" italics="true" pageNumber="1551">Lagidium peruanum</emphasis>
subsp.
<emphasis id="39C4E8D0EA132B6C11DDA3ABD079DEB6" italics="true" pageNumber="1551">peruanum</emphasis>
Meyen 1833
</taxonomicName>
</paragraph>
<paragraph id="F5939058F84937176B2A52CAC920838C" pageNumber="1551">
Subspecies
<taxonomicName id="E0EE9F15F097A3D4967C2C61C09A4BF7" authority="Thomas 1907" authorityName="Thomas" authorityYear="1907" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="peruanum" subSpecies="arequipe">
<emphasis id="878FADB9552CF6D83A97AD1142F26BAA" italics="true" pageNumber="1551">Lagidium peruanum</emphasis>
subsp.
<emphasis id="A6E72FA31350BC34BC9ABB17D2D7DF41" italics="true" pageNumber="1551">arequipe</emphasis>
Thomas 1907
</taxonomicName>
</paragraph>
<paragraph id="8F81D3E6801DCA6D942F852093DBB78C" pageNumber="1551">
Subspecies
<taxonomicName id="C1135B3E4B7AD6D2FDBB1E1E33B2541B" authority="Thomas 1907" authorityName="Thomas" authorityYear="1907" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="peruanum" subSpecies="inca">
<emphasis id="BE058F04BCAFA9BDCD6E9C7CBC9B3458" italics="true" pageNumber="1551">Lagidium peruanum</emphasis>
subsp.
<emphasis id="C41DAEA886191BC5164DF5F99C68D1DE" italics="true" pageNumber="1551">inca</emphasis>
Thomas 1907
</taxonomicName>
</paragraph>
<paragraph id="EF20EDAE357DFB471B1F14880C4E91CE" pageNumber="1551">
Subspecies
<taxonomicName id="DF313D674ABDAE7E714C6D52A93AFBF9" authority="Bennett 1835" authorityName="Bennett" authorityYear="1835" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="peruanum" subSpecies="pallipes">
<emphasis id="CE13C8B590CA511CD20C74AE840E28A4" italics="true" pageNumber="1551">Lagidium peruanum</emphasis>
subsp.
<emphasis id="DBBCC2E82B550773410CA1CDDA219116" italics="true" pageNumber="1551">pallipes</emphasis>
Bennett 1835
</taxonomicName>
</paragraph>
<paragraph id="5BF6B04D2F838A6272B3B92CA7395AC0" pageNumber="1551">
Subspecies
<taxonomicName id="1A0F5A34190E33B2FF36CBA074685FFC" authority="Thomas 1907" authorityName="Thomas" authorityYear="1907" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="peruanum" subSpecies="punensis">
<emphasis id="9219C3AF77B0A67048FAFF9F2959AA3C" italics="true" pageNumber="1551">Lagidium peruanum</emphasis>
subsp.
<emphasis id="7455BDD697E83C2F71988E1111DBF299" italics="true" pageNumber="1551">punensis</emphasis>
Thomas 1907
</taxonomicName>
</paragraph>
<paragraph id="9924F832CB39D680A8F0A6422AB6E446" pageNumber="1551">
Subspecies
<taxonomicName id="3B7EF5A24A984C4A50EE2A0986DC1388" authority="Thomas 1907" authorityName="Thomas" authorityYear="1907" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="peruanum" subSpecies="saturata">
<emphasis id="4DFD94D50C1A9AF11189F2DBF487FCD0" italics="true" pageNumber="1551">Lagidium peruanum</emphasis>
subsp.
<emphasis id="108B3DBCEDDDDE019A4E41543399EC59" italics="true" pageNumber="1551">saturata</emphasis>
Thomas 1907
</taxonomicName>
</paragraph>
<paragraph id="0D77E742CEFBF920BE61965D65C57813" pageNumber="1551">
Subspecies
<taxonomicName id="FDB7AE6CF08078A6C373CD3BFC6652CE" authority="Thomas 1907" authorityName="Thomas" authorityYear="1907" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="peruanum" subSpecies="subrosea">
<emphasis id="FFBC773B9D2316D37187A24981A97EED" italics="true" pageNumber="1551">Lagidium peruanum</emphasis>
subsp.
<emphasis id="3FE60325F6812484771C7B3BF3CB4EA0" italics="true" pageNumber="1551">subrosea</emphasis>
Thomas 1907
</taxonomicName>
</paragraph>
<paragraph id="5C7F3D88E6DBC83B7E43C4AFD0C7FB06" pageNumber="1551">
Species
<taxonomicName id="6DC8C3DD892B9D7BD8DFE5321E878C31" authority="(Molina 1782)" baseAuthorityName="Molina" baseAuthorityYear="1782" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="species" species="viscacia">
<emphasis id="60F9A34C872BDD825DECDFF985D257D7" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
(
<bibRefCitation id="731C2493E0D3CAE934E5D3C3CB6ED049" author="Molina, G. I." refId="ref244945" refString="Molina, G. I. 1782. Saggio sulla storia naturale del Chili. Stamperia di S. Tommaso d'Aquino, Bologna." year="1782">Molina 1782</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="80A0C347F588CC5E7A71838D6E8C9331" pageNumber="1551">
Subspecies
<taxonomicName id="4B01C9B828A9CEC452A84C7E4F178793" authority="Molina 1782" authorityName="Molina" authorityYear="1782" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="viscacia">
<emphasis id="D6411B5179D20E4F1D9957487EF8A83C" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="199302F5A5BD16D6049288EE3D1C1A61" italics="true" pageNumber="1551">viscacia</emphasis>
<bibRefCitation id="C8DB976136B056190A224BD3013E4C56" author="Molina, G. I." refId="ref244945" refString="Molina, G. I. 1782. Saggio sulla storia naturale del Chili. Stamperia di S. Tommaso d'Aquino, Bologna." year="1782">Molina 1782</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="727C776EB52ECDBC1B2EC63E93E5415D" pageNumber="1551">
Subspecies
<taxonomicName id="33BB29227EF72DAC380092BAF7E8231F" authority="Thomas 1921" authorityName="Thomas" authorityYear="1921" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="boxi">
<emphasis id="23C56E1521E2A25A47127EEF4FCDBE99" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="A33742D05E6CFD5116E13EE301ABDCCC" italics="true" pageNumber="1551">boxi</emphasis>
Thomas 1921
</taxonomicName>
</paragraph>
<paragraph id="DA95A9614B5DD7CA039B3C76DD11671B" pageNumber="1551">
Subspecies
<taxonomicName id="681B00B2B3DE70D547F99CF4D35E3674" authority="Thomas 1907" authorityName="Thomas" authorityYear="1907" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="cuscus">
<emphasis id="07EE324A137F853F7256CA07A6A0C71C" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="5DEDC04DC4B8315B5585CF1C8BAA4599" italics="true" pageNumber="1551">cuscus</emphasis>
Thomas 1907
</taxonomicName>
</paragraph>
<paragraph id="F8323CD417C7112EA15622A0CABFA729" pageNumber="1551">
Subspecies
<taxonomicName id="AB8551166AB49B48DB7EF618A041AE40" authority="Bennett 1833" authorityName="Bennett" authorityYear="1833" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="cuvieri">
<emphasis id="C3359A2EF75E4E1EDDFA21EF190199BC" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="55B1D4DC56EF3177DD1FC62A5A540D6B" italics="true" pageNumber="1551">cuvieri</emphasis>
Bennett 1833
</taxonomicName>
</paragraph>
<paragraph id="AA9735CA7B0F9D8196467B6EAC525C21" pageNumber="1551">
Subspecies
<taxonomicName id="55F3E6F7A23CBBF5073160A799425BAE" authority="Thomas 1920" authorityName="Thomas" authorityYear="1920" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="famatinae">
<emphasis id="FF5C4266E018C5A971B4ECB2723687BB" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="3901FE17D40F777EB3F7B0B8C5EF75DC" italics="true" pageNumber="1551">famatinae</emphasis>
Thomas 1920
</taxonomicName>
</paragraph>
<paragraph id="E7D5FC06C6EA67CC8A8EDFE830225093" pageNumber="1551">
Subspecies
<taxonomicName id="F6641E058F694C95E1B65A40D709443D" authority="Thomas 1919" authorityName="Thomas" authorityYear="1919" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="lockwoodi">
<emphasis id="9EBC23E986C4431DF0CFF394D7FC4E3F" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="006C62EDAAA8BADE18AC89553AD63559" italics="true" pageNumber="1551">lockwoodi</emphasis>
<bibRefCitation id="923C172C537C89572B3C53DF17F1938F" author="Thomas, O." refId="ref354472" refString="Thomas, O. 1919. On small mammals from &quot; Otro Cerro &quot; north-eastern Rioja, collected by Sr. E. Budin. Annals and Magazine of Natural History, ser. 9, 3: 489 - 500." year="1919">Thomas 1919</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="04A19B6FE73CB1FDDF7F1C733CC28C20" pageNumber="1551">
Subspecies
<taxonomicName id="00096C5203CFEC492F1508CC19B677D7" authority="Thomas 1897" authorityName="Thomas" authorityYear="1897" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="moreni">
<emphasis id="9C96C2EC809F0E3A4D9E569E784D5DED" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="C72C35CA80BFD7CD11E9D33D1D7057FC" italics="true" pageNumber="1551">moreni</emphasis>
Thomas 1897
</taxonomicName>
</paragraph>
<paragraph id="4D21817D4442E965C3881ADB9341B596" pageNumber="1551">
Subspecies
<taxonomicName id="DBFAA06D6AEABF565624785055289449" authority="Thomas 1907" authorityName="Thomas" authorityYear="1907" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="perlutea">
<emphasis id="7E6B1AC6EDAB46409DEA0D24401BF780" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="54B173C354AEDB88BF16E223D73F9565" italics="true" pageNumber="1551">perlutea</emphasis>
Thomas 1907
</taxonomicName>
</paragraph>
<paragraph id="6EC037A55DAB4033272A9960BA688A07" pageNumber="1551">
Subspecies
<taxonomicName id="6183DFDFEBA4B2CB712E27B0C1892672" authority="Thomas and St. Leger 1926" authorityName="Thomas and St. Leger" authorityYear="1926" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="sarae">
<emphasis id="D646C6E02050CDE8306773531B189D66" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="D73680BFB708B58CEA0ACEFE8CD79B95" italics="true" pageNumber="1551">sarae</emphasis>
Thomas and St. Leger 1926
</taxonomicName>
</paragraph>
<paragraph id="33019ABF9EC38BE812E92EBDB300E8E3" pageNumber="1551">
Subspecies
<taxonomicName id="2763BEFAC5B53FFD7F4DA1F41FD80AC0" authority="Thomas 1921" authorityName="Thomas" authorityYear="1921" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="tontalis">
<emphasis id="5615E89DBE03BC5D432564B63C755A5A" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="4F5220A29D7A0DCD59BC99CCFCE5CEC7" italics="true" pageNumber="1551">tontalis</emphasis>
Thomas 1921
</taxonomicName>
</paragraph>
<paragraph id="0B510F71EB8DE8D8DE3D1B33F514F514" pageNumber="1551">
Subspecies
<taxonomicName id="58873E2094BB1122381B48CE587EC7DD" authority="Thomas 1907" authorityName="Thomas" authorityYear="1907" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="tucumana">
<emphasis id="F3F74A43D8E1D339DB0463C6F0A65CFB" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="2C8D4D754EADBFF465F5F8650832FE4B" italics="true" pageNumber="1551">tucumana</emphasis>
Thomas 1907
</taxonomicName>
</paragraph>
<paragraph id="EDD4339C66ECA29FC78866923630CB18" pageNumber="1551">
Subspecies
<taxonomicName id="B71213F2BD3C1572C7BA8477982C9F9C" authority="Thomas 1921" authorityName="Thomas" authorityYear="1921" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="viatorum">
<emphasis id="9F8E3EE5825879793C224E6F7822686C" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="9AED6A91E7277B984C05523D62A38E5F" italics="true" pageNumber="1551">viatorum</emphasis>
Thomas 1921
</taxonomicName>
</paragraph>
<paragraph id="033F8CC0F1BFE19D6D03CB4578A84C19" pageNumber="1551">
Subspecies
<taxonomicName id="40D56F07E9DF289BC563441F9580BB8D" authority="Thomas 1919" authorityName="Thomas" authorityYear="1919" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="subSpecies" species="viscacia" subSpecies="vulcani">
<emphasis id="0853BF4F56C2BB7F83501F9C6B2699E5" italics="true" pageNumber="1551">Lagidium viscacia</emphasis>
subsp.
<emphasis id="018A54CBF7300975C4716934ED3A970B" italics="true" pageNumber="1551">vulcani</emphasis>
<bibRefCitation id="DAC5C15CDC5716C6FC0E27CE450CC78A" author="Thomas, O." refId="ref354472" refString="Thomas, O. 1919. On small mammals from &quot; Otro Cerro &quot; north-eastern Rioja, collected by Sr. E. Budin. Annals and Magazine of Natural History, ser. 9, 3: 489 - 500." year="1919">Thomas 1919</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A5A5B36FE3069D72A6B4EE5CE3D27E08" pageNumber="1551">
Species
<taxonomicName id="786A451D298D6E730AE7198BB83A6BBF" authority="Thomas 1907" authorityName="Thomas" authorityYear="1907" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" pageNumber="1551" phylum="Chordata" rank="species" species="wolffsohni">
<emphasis id="F4E743ED820FD005E0833DC326760356" italics="true" pageNumber="1551">Lagidium wolffsohni</emphasis>
Thomas 1907
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="3223CF9BB9B1B5AA59A72F5C391B77F8" pageNumber="1551" type="discussion">
<paragraph id="30497BAC79055471415B6493E55200B7" pageNumber="1551">
<emphasis id="BA5515895FC66229CEE6EFC943189C59" inLineHeading="true" pageNumber="1551">Discussion:</emphasis>
Revised by
<bibRefCitation id="BD97F71DE72D4C12F117CEA8B42567E2" author="Osgood, W. H." refId="ref269062" refString="Osgood, W. H. 1943. The mammals of Chile. Field Museum of Natural History, Zoological Series, 30: 1 - 268." year="1943">Osgood (1943)</bibRefCitation>
.
<taxonomicName id="CFCEBA81561C67780C269F1DED8D7336" authority="Oken, 1816" authorityName="Oken" authorityYear="1816" class="Mammalia" family="Chinchillidae" genus="Viscaccia" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="AB65DBB9BD5EA21AF248D66BC1239AFA" italics="true" pageNumber="1551">Viscaccia</emphasis>
Oken, 1816
</taxonomicName>
has precedence over
<taxonomicName id="0460A9AC66676A5039A8E85F3EA4E9D7" authorityName="Meyen" authorityYear="1833" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="C9ABCF4764195EF1FFB149C4B38B614C" italics="true" pageNumber="1551">Lagidium</emphasis>
</taxonomicName>
, but
<taxonomicName id="43F9C674A39A37B30A1D6D28482824D7" authorityName="Meyen" authorityYear="1833" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="20729A5F836408251A6C045E1B5317F4" italics="true" pageNumber="1551">Lagidium</emphasis>
</taxonomicName>
was adopted in preference to
<taxonomicName id="F66DC0C04A9CC64A25240131A6AD8563" authorityName="Oken" authorityYear="1816" class="Mammalia" family="Chinchillidae" genus="Viscaccia" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="5F09B288A051587430372EDDCE144A2D" italics="true" pageNumber="1551">Viscaccia</emphasis>
</taxonomicName>
by suspension of the rules (Opinion 110,
<bibRefCitation id="EA34FE2BF52C2433E70E9830DD171C24" author="International Commission on Zoological Nomenclature" refId="ref166065" refString="International Commission on Zoological Nomenclature. 1929 a. Opinion 110. Suspension of Rules for Lagidium 1833. Smithsonian Miscellaneous Collections, 73 (6): 415." year="1929">
International Commission on Zoological Nomenclature, 1929
<emphasis id="EAB40021C5A8E01EC232C6B70B46FFF8" italics="true" pageNumber="1551">a</emphasis>
</bibRefCitation>
). However, names from Oken’s 1816 &quot;Lehrbuch der Naturgeschichte&quot; are non-Linnaean and not available and neither an opinion nor suspension of the Rules was required for the recognition of
<taxonomicName id="F370874473F428E4EB6B8AFE56F3DC71" authority="(Hershkovitz, 1949 b)" baseAuthorityName="Hershkovitz" baseAuthorityYear="1949" class="Mammalia" family="Chinchillidae" genus="Lagidium" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="41E1200D0F39AADA6CE4C044FAF45184" italics="true" pageNumber="1551">Lagidium</emphasis>
(
<bibRefCitation id="ECE59BF41CCAE703824654A203505956" author="Hershkovitz, P." refId="ref148751" refString="Hershkovitz, P. 1949 b. Status of names credited to Oken, 1816. Journal of Mammalogy, 30: 289 - 301." year="1949">
Hershkovitz, 1949
<emphasis id="073B47B8C651FEBF1CF904F487CFCAA2" italics="true" pageNumber="1551">b</emphasis>
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart>Zhi, Yan</mods:namePart>
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<mods:namePart>Chen, Xiang-Sheng</mods:namePart>
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<treatment ID-GBIF-Taxon="146002429" LSID="urn:lsid:zoobank.org:act:AD015429-772E-4B7F-809D-5CD8CBE35EF9" httpUri="http://treatment.plazi.org/id/E80AE6D0F689CE9CE0E0DC836D21BAC4" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/AD015429-772E-4B7F-809D-5CD8CBE35EF9" authority="Zhi &amp; Chen" class="Insecta" family="Cixiidae" genus="Oecleopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecleopsis productus" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="productus">Oecleopsis productus Zhi &amp; Chen</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
Figs 3-4, 21-34, 57-60
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Holotype: ♂, China: Yunnan Province, Baoshan, Baihua ridge (
<geoCoordinate direction="north" orientation="latitude" precision="925" value="25.283333">25°17'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="98.8">98°48'E</geoCoordinate>
), 15 June 2011, Jian-Kun Long; paratypes: 1♂, 14♀♀, same data as holotype, 13-15 June 2011; 3♂♂, 1♀, collection area same as holotype, 6 May 2010, Bin Zhang, Yan-Li Zheng; 2♂♂, 1♀, Fugong County, Yunnan Province, 17 May 2011, Bin Zhang, Yan-Li Zheng, Bin Yan.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Body length: male 6.7-6.9mm (N = 7), female 6.7-7.8mm (N = 16).</paragraph>
<paragraph pageId="0" pageNumber="1">Coloration. General color blackish brown (Figs 3-4, 21-24). Eyes mid brown, ocelli light yellow. Vertex blackish brown. Face mid to blackish brown, carinae lighter; rostrum mid to blackish brown. Pronotum mid to blackish brown with carinae lighter; mesonotum mid to blackish brown, area between lateral carinae lighter. Forewing semi-translucent, light brown, stigma mid brown. Hind tibiae and abdominal sternites blackish brown.</paragraph>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figures 21-34.
<taxonomicName class="Insecta" family="Cixiidae" genus="Oecleopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecleopsis productus" order="Hemiptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="productus">Oecleopsis productus</taxonomicName>
sp. n., male 21 Head and thorax, dorsal view 22 Face, ventral view 23-24 Head 25 Forewing 26 Genitalia, lateral view 27 Pygofer and genital styles, ventral view 28-29 Anal segment 28 dorsal view 29 right lateral view 30 Genital styles, inner lateral view 31 Aedeagus, right side 32 Aedeagus, left side 33 Aedeagus, dorsal view 34 Aedeagus, ventral view. Scale bars: 0.5 mm (21-24, 26-34); 1.0 mm (25).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">Head and thorax. Vertex (Fig. 21) narrow, 3.5 times longer than wide; frons (Fig. 22) 1.7 times as long as wide. Forewing (Fig. 25) 3.0 times longer than wide, with 12 apical and 6 subapical cells; fork Sc+RP distal to fork CuA1+CuA2; RP 4 branches, MA 3 branches, MP 2 branches. Hind tibia with 4 lateral spines; chaetotaxy of hind tarsi: 7/5.</paragraph>
<paragraph pageId="0" pageNumber="1">Male genitalia. Pygofer (Figs 26-27), dorsal margin concave and U-shaped ventrally, widened towards apex; in lateral view, lateral lobes triangularly extended caudally. Medioventral process narrowly triangular ventrally. Anal segment (Figs 26, 28-29) tubular, asymmetrical, widened towards apex in left side view; in right side view, left ventral margin convex and right ventral margin excavated near apex; 1.8 times longer than wide in dorsal view; anal style fingerlike, beyond anal segment. Genital styles illustrated in Fig. 30. Aedeagus (Figs 31-34) in total with five processes. Spinose process on right side near apex of periandrium long, directed right-dorsocephalically, more than 1/2 of periandrium in length; left side near apex of periandrium with a short reversed spinose process, curving slightly, directed dorsocaudally. Flagellum tapering, apex curved in a semi-circle; a long spinose process arising from dorsal margin, reaching to basal margin of periandrium, curving right side, directed ventrocephalically; left side with a short triangular process, directed ventrocephalically.</paragraph>
<paragraph pageId="0" pageNumber="1">Female genitalia. Genitalia as shown in Fig. 57 ventrally. Anal tube (Fig. 58) 1.9 times longer than wide in dorsal view. Posterior vagina (Figs 59-60) elongate, with a long longitudinal sclerite dorsally, large. In ventral view, left side with a long longitudinal sclerite, semi-sclerotized, right side with a large median constricted sclerite, which with a process, the process directed left-ventrally and longer than shown in the figure.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">China (Yunnan).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific name is derived from the Latin
<taxonomicName lsidName="productus" pageId="0" pageNumber="1" rank="species" species="productus">productus</taxonomicName>
, referring to the dorsal margin of flagellum with a long process.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Male genitalia of
<taxonomicName lsidName="O. productus" pageId="0" pageNumber="1" rank="species" species="productus">O. productus</taxonomicName>
sp. n. is similar to
<taxonomicName lsidName="O. articara" pageId="0" pageNumber="1" rank="species" species="articara">O. articara</taxonomicName>
, but differs in: (1) left side near apex of periandrium with a short process (in
<taxonomicName lsidName="O. articara" pageId="0" pageNumber="1" rank="species" species="articara">O. articara</taxonomicName>
, left side without process); (2) a very long spinose process arising from dorsal margin of flagellum (dorsal margin of flagellum without process in
<taxonomicName lsidName="O. articara" pageId="0" pageNumber="1" rank="species" species="articara">O. articara</taxonomicName>
); (3) left side near apex of flagellum with a very short process (in
<taxonomicName lsidName="O. articara" pageId="0" pageNumber="1" rank="species" species="articara">O. articara</taxonomicName>
, left side near middle of flagellum with a much longer process); 4) forewing with RP 4 branches (in
<taxonomicName lsidName="O. articara" pageId="0" pageNumber="1" rank="species" species="articara">O. articara</taxonomicName>
, 3 branches).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Female genitalia of
<taxonomicName lsidName="O. productus" pageId="0" pageNumber="1" rank="species" species="productus">O. productus</taxonomicName>
sp. n. is similar to
<taxonomicName lsidName="O. yoshikawai" pageId="0" pageNumber="1" rank="species" species="yoshikawai">O. yoshikawai</taxonomicName>
, but differs in: (1) right side of posterior vagina with one sclerite, which constricted medially in ventral view in
<taxonomicName lsidName="O. productus" pageId="0" pageNumber="1" rank="species" species="productus">O. productus</taxonomicName>
(with two sclerites, terminal irregular sclerite forming three projecting oblong structures in ventral view in
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); (2) in ventral view, distal part of the sclerite on left side of posterior vagina not extended and curved to right in
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(extended and curved to right in
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).
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<mods:title id="B026D71709B76E826BD9CB714F7A85A6">A new wobbegong shark, Orectolobus hutchinsi n. sp. (Orectolobiformes: Orectolobidae) from southwestern Australia.</mods:title>
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<mods:namePart id="FEE26A5AC8BAA8FBBC9A0276CD7F6FA8">Peter R. Last</mods:namePart>
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[[ Family
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Gill ]]
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The wobbegongs (F.
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) are represented by three genera and seven species in the Indo -Pacific (Compagno et al., 2005). The group has not been subjected to a formal review and several taxonomic problems exist. At least six species are thought to occur in Australian seas (Last and Stevens, 1994):
<taxonomicName id="A27B18853528EA4B39A7571C4EBF8C4E" class="Elasmobranchii" family="Orectolobidae" genus="Eucrossorhinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Orectolobiformes" pageNumber="35" phylum="Chordata" rank="species" species="dasypogon">Eucrossorhinus dasypogon (Bleeker, 1867)</taxonomicName>
,
<taxonomicName id="9470E8CA2545496C46A02FF00FE4142B" class="Elasmobranchii" family="Orectolobidae" genus="Orectolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Orectolobiformes" pageNumber="35" phylum="Chordata" rank="species" species="maculatus">O. maculatus (Bonnaterre, 1788)</taxonomicName>
,
<taxonomicName id="88B44236035771848EDA074A145F2408" class="Elasmobranchii" family="Orectolobidae" genus="Orectolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Orectolobiformes" pageNumber="35" phylum="Chordata" rank="species" species="ornatus">O. ornatus (de Vis, 1883)</taxonomicName>
,
<taxonomicName id="209AFEF5963D1C7FEBF5F57B09052539" LSID-ZBK="urn:lsid:zoobank.org:act:D4FB912C-72AD-4D37-837C-E5B8C34B6C8E" class="Elasmobranchii" family="Orectolobidae" genus="Orectolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Orectolobus wardi Whitley 1939:264" order="Orectolobiformes" pageNumber="35" phylum="Chordata" rank="species" species="wardi">O. wardi Whitley, 1939</taxonomicName>
,
<taxonomicName id="F2D40B8AF3DA7F629E041D40776A7404" class="Elasmobranchii" family="Orectolobidae" genus="Sutorectus" higherTaxonomySource="CoL" kingdom="Animalia" order="Orectolobiformes" pageNumber="35" phylum="Chordata" rank="species" species="tentaculatus">Sutorectus tentaculatus (Peters, 1864)</taxonomicName>
and an un-named species,
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. All of theses species, apart from
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and
<taxonomicName id="4A0E61236FEC31164B9B83F8BB1196B6" class="Elasmobranchii" family="Orectolobidae" genus="Eucrossorhinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Orectolobiformes" pageNumber="35" phylum="Chordata" rank="species" species="dasypogon">E. dasypogon</taxonomicName>
, were thought to occur off southwestern Australia. More recent research suggests that other undescribed species occur off Western Australia (Last and Chidlow, unpublished data). The new species was informally described but not named by Last and Stevens (1994) and illustrated as a composite of multiple specimens from a variety of colour prints and slides. More recently, Chidlow (2003) completed a study of the biology of this species as part of a higher university degree. The new wobbegong is formally described below and aspects of its biology are discussed.
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<taxonomicName LSID="urn:lsid:zoobank.org:act:730024CA-3646-4660-9FBD-C307773D0E94" authority="Saldaitis, Ivinskis &amp; Yakovlev" class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia hackeri" order="Lepidoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="hackeri">Meharia hackeri Saldaitis, Ivinskis &amp; Yakovlev</taxonomicName>
<taxonomicNameLabel pageId="3" pageNumber="48">sp. n.</taxonomicNameLabel>
Figs 9, 10, 13, 14
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<paragraph pageId="3" pageNumber="48">Type material.</paragraph>
<paragraph pageId="3" pageNumber="48">Holotype ♀ (Fig. 9), NE Socotra Island, Wadi Difarroha, North side, 19 January 2010. leg. A. Saldaitis (deposited in MWM/ZSM); (slide No PI 2011/1) Paratypes: 3 ♀ (Fig. 10), S Socotra Island, Wadi Difarroha, South side, 15 January 2010. leg. A. Saldaitis; (coll. ASV; MWM/ZSM); (slide No BJ 1523).</paragraph>
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<subSubSection pageId="3" pageNumber="48" type="diagnosis">
<paragraph pageId="3" pageNumber="48">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="48">
The new species differs from the related species
<taxonomicName class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia acuta" order="Lepidoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="acuta">Meharia acuta</taxonomicName>
Wiltshire, 1982 (Figs 8, 12) by forewing pattern, DNA and distribution. In
<taxonomicName class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia acuta" order="Lepidoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="acuta">Meharia acuta</taxonomicName>
, the basal spot at the costal wing margin is missing.
<taxonomicName class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia hackeri" order="Lepidoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="hackeri">Meharia hackeri</taxonomicName>
has a straight basal fascia at the costal wing margin for
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the length of forewing and a narrow white fascia, with a wide interruption antemedially and a narrow interruption tornally, running along the entire inner margin.
<taxonomicName class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia acuta" order="Lepidoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="acuta">Meharia acuta</taxonomicName>
has no such fascia, but has a wide subterminal band. DNA barcodes clearly separate
<taxonomicName class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia hackeri" order="Lepidoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="hackeri">Meharia hackeri</taxonomicName>
from
<taxonomicName class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia acuta" order="Lepidoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="acuta">Meharia acuta</taxonomicName>
. Three identical sequences of
<taxonomicName class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia hackeri" order="Lepidoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="hackeri">Meharia hackeri</taxonomicName>
were compared to those of a single
<taxonomicName class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia acuta" order="Lepidoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="acuta">Meharia acuta</taxonomicName>
specimen resulting in a significant 7.48% variation.
</paragraph>
<paragraph pageId="3" pageNumber="48">
<taxonomicName class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia acuta" order="Lepidoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="acuta">Meharia acuta</taxonomicName>
is distributed in the Arabic peninsula,
<taxonomicName class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia hackeri" order="Lepidoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="hackeri">Meharia hackeri</taxonomicName>
is endemic to Socotra Island.
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<paragraph pageId="3" pageNumber="48">Description.</paragraph>
<paragraph lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
Female: Forewing costal margin length of holotype 10 mm, wingspan 21 mm; forewing length of paratypes 11 mm, wingspan 22 mm; antennae slightly longer than half the length of forewing; bipectinate, color white, black at base; head and tegular yellowish-white; labial palpi yellowish brown, white at base; ground colour of forewings yellowish-brown with white longitudinal fascia forming wing pattern, basal fascia in the costal area straight, running to
<normalizedToken originalValue="¼">1/4</normalizedToken>
the length of forewing, curved fascia extending medially from inner margin to ⅔ the length of forewing, its extension ends at terminal wing margin, medially the fascia and its interrupted portion in terminal area bordered by dark brown scales with black inserts; narrow white fascia, widely i
<pageBreakToken pageId="4" pageNumber="49" start="start">nterrupted</pageBreakToken>
antemedially and narrowly interrupted tornally, runs along the entire inner margin, cilia yellowish-white, ventral forewing brown; hindwing greyish-yellow, cilia light brown, ventral hindwing brown.
</paragraph>
<paragraph pageId="4" pageNumber="49">Male genitalia: unknown.</paragraph>
<paragraph pageId="4" pageNumber="49">Female genitalia (Figs 13, 14): Papilla analis triangular, covered with short, thin, very long setae; apophysis posterioris about the same length as papilla analis; apophysis anterioris very short, broad, with V-shaped sclerotisation apically; ostium concave; antrum weakly sclerotised basally with a loop forming very narrow ductus bursae; corpus bursae not sclerotised, shaped like a small sac.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="49" type="distribution">
<paragraph pageId="4" pageNumber="49">Bionomics and distribution.</paragraph>
<paragraph pageId="4" pageNumber="49">
Known only from the central part of Socotra Island.
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is likely endemic to Socotra Island. All specimens were collected in mid-January;
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females were attracted to light and appear to have a very local distribution as the species was discovered only in Difarroha Valley (Fig. 31). The new species was collected in the central part of the country in an oasis-type valley dominated by various tree and shrub species such as:
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,
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,
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,
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<taxonomicName class="Magnoliopsida" family="Moraceae" genus="Ficus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ficus vasta" order="Rosales" pageId="4" pageNumber="49" phylum="Tracheophyta" rank="species" species="vasta">Ficus vasta</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Euphorbia socotrana" order="Malpighiales" pageId="4" pageNumber="49" phylum="Tracheophyta" rank="species" species="socotrana">Euphorbia socotrana</taxonomicName>
,
<taxonomicName genus="Jathropha" lsidName="Jathropha unicostata" pageId="4" pageNumber="49" rank="species" species="unicostata">Jathropha unicostata</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Mitolepis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Mitolepis intricata" order="Gentianales" pageId="4" pageNumber="49" phylum="Tracheophyta" rank="species" species="intricata">Mitolepis intricata</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Asphodelaceae" genus="Aloe" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aloe perryi" order="Asparagales" pageId="4" pageNumber="49" phylum="Tracheophyta" rank="species" species="perryi">Aloe perryi</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Apocynaceae" genus="Adenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Adenium obesum" order="Gentianales" pageId="4" pageNumber="49" phylum="Tracheophyta" rank="species" species="obesum">Adenium obesum</taxonomicName>
(
<bibRefCitation author="Miller, AG" journalOrPublisher="Edinburgh" pageId="14" pageNumber="59" pagination="1 - 556" title="Flora of the Arabian Peninsula and Socotra 1" year="1996">Miller and Cope 1996</bibRefCitation>
). It flies with several other Socotra Archipelago endemic moths such as
<taxonomicName class="Insecta" family="Cossidae" genus="Meharia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meharia yakovlevi" order="Lepidoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="yakovlevi">Meharia yakovlevi</taxonomicName>
Saldaitis &amp; Ivinskis, 2010, (
<taxonomicName family="Cossidae" lsidName="" pageId="4" pageNumber="49" rank="family">Cossidae</taxonomicName>
),
<taxonomicName class="Insecta" family="Arctiidae" genus="Pelosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelosia sokotrensis" order="Lepidoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="sokotrensis">Pelosia sokotrensis</taxonomicName>
(Hampson, 1900), (
<taxonomicName family="Arctiidae" lsidName="" pageId="4" pageNumber="49" rank="family">Arctiidae</taxonomicName>
),
<taxonomicName class="Insecta" family="Noctuidae" genus="Cerocala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cerocala socotrensis" order="Lepidoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="socotrensis">Cerocala socotrensis</taxonomicName>
Hampson, 1899,
<taxonomicName class="Insecta" family="Noctuidae" genus="Agrotis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agrotis brachypecten" order="Lepidoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="brachypecten">Agrotis brachypecten</taxonomicName>
Hampson, 1899,
<taxonomicName class="Insecta" family="Noctuidae" genus="Plecoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plecoptera butkevicii" order="Lepidoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="butkevicii">Plecoptera butkevicii</taxonomicName>
Hacker &amp; Saldaitis, 2010,
<taxonomicName class="Insecta" family="Noctuidae" genus="Acantholipes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholipes canofusca" order="Lepidoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="canofusca">Acantholipes canofusca</taxonomicName>
Hacker &amp; Saldaitis, 2010,
<taxonomicName class="Insecta" family="Noctuidae" genus="Stenosticta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenosticta wiltshirei" order="Lepidoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="wiltshirei">Stenosticta wiltshirei</taxonomicName>
Hacker, Saldaitis &amp; Ivinskis, 2010 (
<taxonomicName family="Noctuidae" lsidName="" pageId="4" pageNumber="49" rank="family">Noctuidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="49" type="etymology">
<paragraph pageId="4" pageNumber="49">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="49">The new species name is dedicated to Hermann Hacker, a prominent German lepidopterist, who has contributed much to the investigation of macro-moths of the Arabian peninsula and Africa.</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.99.26489" ID-GBIF-Dataset="cd80a3c5-60b9-4bda-9780-c03269f98d11" ID-PMC="PMC5990585" ID-Pensoft-Pub="1314-2003-99-107" ID-Pensoft-UUID="036DC676945FFFA8FFE6D1180807FF89" ID-PubMed="29881324" ID-Zenodo-Dep="1284663" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-2003-99-107" ModsDocOrigin="PhytoKeys 99" ModsDocTitle="Anemonastrumtenuicaule and A.antucense (Ranunculaceae), new combinations for a New Zealand endemic species and its South American relative" checkinTime="1527698462165" checkinUser="pensoft" docAuthor="Mosyakin, Sergei L. &amp; Lange, Peter J. de" docDate="2018" docId="E80D394FDA0933BF2168E5A0185A6B17" docLanguage="en" docName="PhytoKeys 99: 107-124" docOrigin="PhytoKeys 99" docSource="http://dx.doi.org/10.3897/phytokeys.99.26489" docTitle="Anemonastrum tenuicaule de Lange &amp; Mosyakin 2018, comb. nov." docType="treatment" docVersion="6" id="036DC676945FFFA8FFE6D1180807FF89" lastPageNumber="107" masterDocId="036DC676945FFFA8FFE6D1180807FF89" masterDocTitle="Anemonastrum tenuicaule and A. antucense (Ranunculaceae), new combinations for a New Zealand endemic species and its South American relative" masterLastPageNumber="124" masterPageNumber="107" pageNumber="107" updateTime="1668142300223" updateUser="ExternalLinkService">
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<mods:title>Anemonastrum tenuicaule and A. antucense (Ranunculaceae), new combinations for a New Zealand endemic species and its South American relative</mods:title>
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<mods:namePart>Mosyakin, Sergei L.</mods:namePart>
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<mods:namePart>Lange, Peter J. de</mods:namePart>
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<treatment ID-GBIF-Taxon="144192116" LSID="urn:lsid:plazi:treatment:E80D394FDA0933BF2168E5A0185A6B17" httpUri="http://treatment.plazi.org/id/E80D394FDA0933BF2168E5A0185A6B17" lastPageNumber="107" pageId="0" pageNumber="107">
<subSubSection pageId="0" pageNumber="107" type="nomenclature">
<paragraph pageId="0" pageNumber="107">
<taxonomicName LSID="C715301A-D70D-5326-B1AF-30A13A93FBA1" authority="(Cheeseman) de Lange &amp; Mosyakin" authorityName="de Lange &amp; Mosyakin" authorityYear="2018" baseAuthorityName="de Lange &amp; Mosyakin" baseAuthorityYear="2018" class="Magnoliopsida" family="Ranunculaceae" genus="Anemonastrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anemonastrum tenuicaule" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaule" status="comb. nov.">Anemonastrum tenuicaule (Cheeseman) de Lange &amp; Mosyakin</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="107">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="reference_group">
<paragraph pageId="0" pageNumber="107">
≡
<taxonomicName authorityName="Christenh &amp; Byng" authorityYear="2018" baseAuthorityName="Cheeseman" class="Magnoliopsida" family="Ranunculaceae" genus="Anemonidium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anemonidium tenuicaule" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaule">Anemonidium tenuicaule</taxonomicName>
(Cheeseman) Christenh. &amp; Byng in Christenhusz et al. (Eds)
<emphasis italics="true" pageId="0" pageNumber="107">The Global Flora</emphasis>
4: 73. 2018.
</paragraph>
<paragraph pageId="0" pageNumber="107">
≡
<taxonomicName authorityName="Parkin &amp; Sledge" authorityYear="1935" baseAuthorityName="Cheesem." class="Magnoliopsida" family="Ranunculaceae" genus="Anemone" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anemone tenuicaulis" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaulis">Anemone tenuicaulis</taxonomicName>
(Cheeseman) Parkin &amp; Sledge,
<emphasis italics="true" pageId="0" pageNumber="107">J. Linn. Soc., Bot.</emphasis>
49: 647. 1935.
</paragraph>
<paragraph pageId="0" pageNumber="107">
≡
<taxonomicName authorityName="Cheesem" authorityYear="1885" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus tenuicaulis" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaulis">Ranunculus tenuicaulis</taxonomicName>
Cheeseman,
<emphasis italics="true" pageId="0" pageNumber="107">Trans. &amp; Proc. New Zealand Inst.</emphasis>
17: 235. 1885.
<emphasis bold="true" pageId="0" pageNumber="107">Lectotype</emphasis>
(designated by
<bibRefCitation author="Allan, HH" pageId="0" pageNumber="107" refId="B1" refString="Allan, HH, 1961. Flora of New Zealand, Vol. I. Government Printer, Wellington, liv + 1085 pp." title="Flora of New Zealand, Vol. I. Government Printer, Wellington, liv + 1085 pp." year="1961">Allan 1961</bibRefCitation>
: 164; accepted by
<bibRefCitation author="Burrows, CJ" journalOrPublisher="New Zealand Journal of Botany" pageId="0" pageNumber="107" pagination="9 - 68" publicationUrl="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" refId="B3" refString="Burrows, CJ, 1986. Botany of Arthur's Pass National Park, South Island, New Zealand. I. History of botanical studies and checklist of the vascular flora. New Zealand Journal of Botany 24: 9 - 68, https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" title="Botany of Arthur's Pass National Park, South Island, New Zealand. I. History of botanical studies and checklist of the vascular flora." url="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" volume="24" year="1986">Burrows 1986</bibRefCitation>
: 15, and confirmed and specified here). NEW ZEALAND. South Island, Mountains above
<normalizedToken originalValue="Arthur’s">Arthur's</normalizedToken>
Pass, Canterbury Alps. Printed and handwritten label: &quot;Herb. T.F. Cheeseman |
<taxonomicName class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">Ranunculus</taxonomicName>
sp. [
<normalizedToken originalValue="“sp.”">&quot;sp.&quot;</normalizedToken>
crossed out-S.M. &amp; P.dL.] tenuicaulis n. sp. [new identification added in pencil-S.M. &amp; P.dL.] | Locality:-South Island, N.Z. | Mts above
<normalizedToken originalValue="Arthur’s">Arthur's</normalizedToken>
Pass, Canterbury Alps, alt. | 4,500 ft. | Jany [January-S.M. &amp; P.dL.] 1883 | Collector-T.F.C.&quot; Small slip attached in the upper part of the sheet: &quot;TYPE SELECTED. Dec. 1941. [signature of Lucy Cranwell]&quot; (AK4232!;
<emphasis bold="true" pageId="0" pageNumber="107">isolectotypes</emphasis>
: &quot;Herb. T.F. Cheeseman.
<emphasis bold="true" pageId="0" pageNumber="107">Com.</emphasis>
[communicated?]
<emphasis bold="true" pageId="0" pageNumber="107">9/83</emphasis>
[text in bold added in black ink, handwritten-S.M. &amp; P.dL.] |
<taxonomicName class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">Ranunculus</taxonomicName>
n. sp.? | LOCALITY: -South Island, N.Z. | mountains above
<normalizedToken originalValue="Arthur’s">Arthur's</normalizedToken>
Pass, Canterbury, | alt. 4,500 ft. January 1883 | Collector-T.F.C. [T.F. Cheeseman-S.M. &amp; P.dL.]&quot;. Identification added directly on the sheet under the label: &quot;
<taxonomicName authorityName="Cheesem" authorityYear="1885" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus tenuicaulis" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaulis">Ranunculus tenuicaulis</taxonomicName>
, Cheeseman&quot; K000692121!, reported by Ziman
<emphasis italics="true" pageId="0" pageNumber="107">et al</emphasis>
. 2006: 217 as
<normalizedToken originalValue="“lectotype”">&quot;lectotype&quot;</normalizedToken>
, E
<emphasis italics="true" pageId="0" pageNumber="107">s.n.</emphasis>
reported by Ziman
<emphasis italics="true" pageId="0" pageNumber="107">et al</emphasis>
. 2006: 217,
<emphasis italics="true" pageId="0" pageNumber="107">non vidi</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="notes">
<paragraph pageId="0" pageNumber="107">Notes.</paragraph>
<paragraph pageId="0" pageNumber="107">
Cheeseman (
<bibRefCitation author="Cheeseman, TF" journalOrPublisher="Transactions and Proceedings of the New Zealand Institute" pageId="0" pageNumber="107" pagination="235 - 236" publicationUrl="http://rsnz.natlib.govt.nz/volume/rsnz_17/rsnz_17_00_003820.html" refId="B4" refString="Cheeseman, TF, 1885 [1884]. New species of plants. Transactions and Proceedings of the New Zealand Institute 17: 235 - 236, http://rsnz.natlib.govt.nz/volume/rsnz_17/rsnz_17_00_003820.html" title="New species of plants." url="http://rsnz.natlib.govt.nz/volume/rsnz_17/rsnz_17_00_003820.html" volume="17" year="1885 [1884]">1885</bibRefCitation>
) reported his new species (as
<taxonomicName authorityName="Cheesem" authorityYear="1885" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus tenuicaulis" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaulis">
<emphasis italics="true" pageId="0" pageNumber="107">Ranunculus tenuicaulis</emphasis>
</taxonomicName>
) from &quot;Canterbury mountains above
<normalizedToken originalValue="Arthur’s">Arthur's</normalizedToken>
Pass, altitude 4,000-5,000 feet.
<emphasis italics="true" pageId="0" pageNumber="107">T.F.C.</emphasis>
&quot; and all his collections from that locality should be considered syntypes. Ziman et al. (
<bibRefCitation author="Ziman, SN" journalOrPublisher="Journal of Japanese Botany" pageId="0" pageNumber="107" pagination="1 - 19" publicationUrl="http://www.jjbotany.com/pdf/JJB_081_1_19.pdf" refId="B99" refString="Ziman, SN, Ehrendorfer, F, Kadota, Y, Keener, CS, Tsarenko, ON, Bulakh, EV, Dutton, BE, 2006a. A taxonomic revision of Anemone L. section Omalocarpus DC. sensu lato (Ranunculaceae). II. Journal of Japanese Botany 81: 1 - 19, http://www.jjbotany.com/pdf/JJB_081_1_19.pdf" title="A taxonomic revision of Anemone L. section Omalocarpus DC. sensu lato (Ranunculaceae). II." url="http://www.jjbotany.com/pdf/JJB_081_1_19.pdf" volume="81" year="2006 a">2006</bibRefCitation>
: 217) provided for
<taxonomicName authorityName="Parkin &amp; Sledge" authorityYear="1935" baseAuthorityName="Cheesem." class="Magnoliopsida" family="Ranunculaceae" genus="Anemone" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anemone tenuicaulis" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaulis">
<emphasis italics="true" pageId="0" pageNumber="107">Anemone tenuicaulis</emphasis>
</taxonomicName>
the following type information: &quot;Type: NEW ZEALAND. South Island, Auckland, South Alps, Mountains above
<normalizedToken originalValue="Arthur’s">Arthur's</normalizedToken>
Pass, Canterbury, 4000-5000 ft. 1.1883. Lannary (lectotype-K!; isolectotype-E!)&quot;. They, however, cited
<normalizedToken originalValue="“Auckland”">&quot;Auckland&quot;</normalizedToken>
(printed on the label, indicating the location of
<normalizedToken originalValue="Cheeseman’s">Cheeseman's</normalizedToken>
herbarium) as part of the type locality information, misunderstood the handwritten word
<normalizedToken originalValue="“January”">&quot;January&quot;</normalizedToken>
for a collector name (
<normalizedToken originalValue="“Lannary”">&quot;Lannary&quot;</normalizedToken>
), and erroneously listed the combination
<taxonomicName authorityName="Parkin &amp; Sledge" authorityYear="1935" baseAuthorityName="Cheesem." class="Magnoliopsida" family="Ranunculaceae" genus="Anemone" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anemone tenuicaulis" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaulis">
<emphasis italics="true" pageId="0" pageNumber="107">Anemone tenuicaulis</emphasis>
</taxonomicName>
as validated in &quot;Nat. 1 (1932)&quot;, the incomplete citation evidently corresponding to the article in
<emphasis italics="true" pageId="0" pageNumber="107">Nature</emphasis>
(
<bibRefCitation DOI="https://doi.org/10.1038/130023b0" author="Parkin, J" journalOrPublisher="Nature" pageId="0" pageNumber="107" pagination="23 - 24" refId="B58" refString="Parkin, J, Sledge, WA, 1932. A genus of Ranunculaceae hitherto unrecorded for New Zealand. Nature 130 (3270): 23 - 24, DOI: https://doi.org/10.1038/130023b0" title="A genus of Ranunculaceae hitherto unrecorded for New Zealand." url="https://doi.org/10.1038/130023b0" volume="130" year="1932">Parkin and Sledge 1932</bibRefCitation>
) in which only preliminary information on the new generic placement of
<taxonomicName authorityName="Cheesem" authorityYear="1885" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus tenuicaulis" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaulis">
<emphasis italics="true" pageId="0" pageNumber="107">Ranunculus tenuicaulis</emphasis>
</taxonomicName>
was reported, but no new combination has been validated. When listing and designating types of
<taxonomicName class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="107">Ranunculus</emphasis>
</taxonomicName>
names from New Zealand,
<bibRefCitation author="Garnock-Jones, PJ" journalOrPublisher="New Zealand Journal of Botany" pageId="0" pageNumber="107" pagination="115 - 123" publicationUrl="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1990.10412351" refId="B27" refString="Garnock-Jones, PJ, 1990. Typification of Ranunculus names in New Zealand (Ranunculaceae). New Zealand Journal of Botany 28: 115 - 123, https://www.tandfonline.com/doi/abs/10.1080/0028825X.1990.10412351" title="Typification of Ranunculus names in New Zealand (Ranunculaceae)." url="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1990.10412351" volume="28" year="1990">Garnock-Jones (1990)</bibRefCitation>
only mentioned
<taxonomicName authorityName="Cheesem" authorityYear="1885" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus tenuicaulis" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaulis">
<emphasis italics="true" pageId="0" pageNumber="107">Ranunculus tenuicaulis</emphasis>
</taxonomicName>
among the taxa that are excluded from that genus but gave no type information.
</paragraph>
<paragraph pageId="0" pageNumber="107">
The following type information was provided by
<bibRefCitation author="Allan, HH" pageId="0" pageNumber="107" refId="B1" refString="Allan, HH, 1961. Flora of New Zealand, Vol. I. Government Printer, Wellington, liv + 1085 pp." title="Flora of New Zealand, Vol. I. Government Printer, Wellington, liv + 1085 pp." year="1961">Allan (1961</bibRefCitation>
: 164): &quot;Type locality: &quot;Mountains above
<normalizedToken originalValue="Arthur’s">Arthur's</normalizedToken>
Pass, alt. 4000-5000 feet.&quot; Type: A, T. F. Cheeseman&quot;, which constitutes effective lectotypification (Art. 7.10 of the ICN:
<bibRefCitation author="McNeill, J" journalOrPublisher="Regnum Vegetabile" pageId="0" pageNumber="107" pagination="1 - 274" publicationUrl="http://www.iapt-taxon.org/historic/2012.htm" refId="B52" refString="McNeill, J, Barrie, FR, Buck, WR, Demoulin, V, Greuter, W, Hawksworth, DL, Herendeen, PS, Knapp, S, Marhold, K, Prado, J, Prud'homme, van Reine WF, Smith, GF, Wiersema, JH, Turland, NJ, 2012. International Code of Nomenclature for algae, fungi, and plants (Melbourne Code): Adopted by the eighteenth International Botanical Congress Melbourne, Australia, July 2011. Regnum Vegetabile 154: 1 - 274, http://www.iapt-taxon.org/historic/2012.htm" title="International Code of Nomenclature for algae, fungi, and plants (Melbourne Code): Adopted by the eighteenth International Botanical Congress Melbourne, Australia, July 2011." url="http://www.iapt-taxon.org/historic/2012.htm" volume="154" year="2012">McNeill et al. 2012</bibRefCitation>
). In this citation, the letter
<normalizedToken originalValue="“A”">&quot;A&quot;</normalizedToken>
indicates the Herbarium of Auckland Institute and Museum (AK).
<bibRefCitation author="Burrows, CJ" journalOrPublisher="New Zealand Journal of Botany" pageId="0" pageNumber="107" pagination="9 - 68" publicationUrl="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" refId="B3" refString="Burrows, CJ, 1986. Botany of Arthur's Pass National Park, South Island, New Zealand. I. History of botanical studies and checklist of the vascular flora. New Zealand Journal of Botany 24: 9 - 68, https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" title="Botany of Arthur's Pass National Park, South Island, New Zealand. I. History of botanical studies and checklist of the vascular flora." url="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" volume="24" year="1986">Burrows (1986)</bibRefCitation>
in his article also provided a table entitled &quot;List of vascular plant taxa described originally from
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Pass National Park&quot; and listed
<taxonomicName authorityName="Parkin &amp; Sledge" authorityYear="1935" baseAuthorityName="Cheesem." class="Magnoliopsida" family="Ranunculaceae" genus="Anemone" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anemone tenuicaulis" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaulis">
<emphasis italics="true" pageId="0" pageNumber="107">Anemone tenuicaulis</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="Cheesem" authorityYear="1885" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus tenuicaulis" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaulis">
<emphasis italics="true" pageId="0" pageNumber="107">Ranunculus tenuicaulis</emphasis>
</taxonomicName>
), with proper references to the authors of the basionym and combination and their original publications. He reported (
<bibRefCitation author="Burrows, CJ" journalOrPublisher="New Zealand Journal of Botany" pageId="0" pageNumber="107" pagination="9 - 68" publicationUrl="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" refId="B3" refString="Burrows, CJ, 1986. Botany of Arthur's Pass National Park, South Island, New Zealand. I. History of botanical studies and checklist of the vascular flora. New Zealand Journal of Botany 24: 9 - 68, https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" title="Botany of Arthur's Pass National Park, South Island, New Zealand. I. History of botanical studies and checklist of the vascular flora." url="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" volume="24" year="1986">Burrows 1986</bibRefCitation>
: 15) the date and place of the original collection of Cheeseman (&quot;Jan 1883 Mts above
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Pass&quot;) and the location of the type specimen (
<normalizedToken originalValue="“AUCK”">&quot;AUCK&quot;</normalizedToken>
, meaning &quot;Auckland Institute &amp; Museum [AK]&quot;; see explanation in
<bibRefCitation author="Burrows, CJ" journalOrPublisher="New Zealand Journal of Botany" pageId="0" pageNumber="107" pagination="9 - 68" publicationUrl="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" refId="B3" refString="Burrows, CJ, 1986. Botany of Arthur's Pass National Park, South Island, New Zealand. I. History of botanical studies and checklist of the vascular flora. New Zealand Journal of Botany 24: 9 - 68, https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" title="Botany of Arthur's Pass National Park, South Island, New Zealand. I. History of botanical studies and checklist of the vascular flora." url="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" volume="24" year="1986">Burrows 1986</bibRefCitation>
: 17). Considering the lectotypification information provided above, the Kew specimen is not the lectotype of
<taxonomicName authorityName="Cheesem" authorityYear="1885" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ranunculus tenuicaulis" order="Ranunculales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tenuicaulis">
<emphasis italics="true" pageId="0" pageNumber="107">Ranunculus tenuicaulis</emphasis>
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, but an isolectotype.
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<paragraph pageId="0" pageNumber="107">
There are several specimens of the species at AK collected by Cheeseman, e.g., AK4233, AK4234 (data and images available from the Auckland War Memorial Museum: http://www.aucklandmuseum.com), but only one collected in January 1883 near
<normalizedToken originalValue="Arthur’s">Arthur's</normalizedToken>
Pass and matching other data provided by
<bibRefCitation author="Allan, HH" pageId="0" pageNumber="107" refId="B1" refString="Allan, HH, 1961. Flora of New Zealand, Vol. I. Government Printer, Wellington, liv + 1085 pp." title="Flora of New Zealand, Vol. I. Government Printer, Wellington, liv + 1085 pp." year="1961">Allan (1961)</bibRefCitation>
and
<bibRefCitation author="Burrows, CJ" journalOrPublisher="New Zealand Journal of Botany" pageId="0" pageNumber="107" pagination="9 - 68" publicationUrl="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" refId="B3" refString="Burrows, CJ, 1986. Botany of Arthur's Pass National Park, South Island, New Zealand. I. History of botanical studies and checklist of the vascular flora. New Zealand Journal of Botany 24: 9 - 68, https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" title="Botany of Arthur's Pass National Park, South Island, New Zealand. I. History of botanical studies and checklist of the vascular flora." url="https://www.tandfonline.com/doi/abs/10.1080/0028825X.1986.10409720" volume="24" year="1986">Burrows (1986)</bibRefCitation>
. Lucy M. Cranwell, who incorporated the Cheeseman collections (ca. 10 000 specimens) into AK, in December 1941 annotated the specimen AK4232 as the type (see above), but her type designation was not formally published. It is documented (
<bibRefCitation author="Goulding, JH" journalOrPublisher="Records of the Auckland Institute and Museum" pageId="0" pageNumber="107" pagination="105 - 117" refId="B28" refString="Goulding, JH, 1974. Notes on the Cheeseman Herbarium: Part 1. Exchange with U.S.A. herbaria before 1900. Records of the Auckland Institute and Museum 11: 105 - 117" title="Notes on the Cheeseman Herbarium: Part 1. Exchange with U. S. A. herbaria before 1900." volume="11" year="1974">Goulding 1974</bibRefCitation>
,
<bibRefCitation author="Goulding, JH" journalOrPublisher="Records of the Auckland Institute and Museum" pageId="0" pageNumber="107" pagination="95 - 120" refId="B29" refString="Goulding, JH, 1975. Notes on the Cheeseman Herbarium: Part 2. Exchange with European herbaria before 1900. Records of the Auckland Institute and Museum 12: 95 - 120" title="Notes on the Cheeseman Herbarium: Part 2. Exchange with European herbaria before 1900." volume="12" year="1975">1975</bibRefCitation>
,
<bibRefCitation author="Goulding, JH" journalOrPublisher="Records of the Auckland Institute and Museum" pageId="0" pageNumber="107" pagination="101 - 108" refId="B30" refString="Goulding, JH, 1976. Notes on the Cheeseman Herbarium: Part 3. Exchange with Australian herbaria from before 1900 to 1923. Records of the Auckland Institute and Museum 13: 101 - 108" title="Notes on the Cheeseman Herbarium: Part 3. Exchange with Australian herbaria from before 1900 to 1923." volume="13" year="1976">1976</bibRefCitation>
) that Cheeseman exchanged herbarium specimens with several European, American, and Australian herbaria and individual botanists; thus, additional isolectotypes or syntypes could be found in some other collections, in addition to the specimens known to be at K and E.
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.996.59075</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-996-93</mods:identifier>
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<mods:identifier type="Pensoft-UUID">2C7EB5892FBE5F74B4DFB88CE33B2FC1</mods:identifier>
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<treatment ID-GBIF-Taxon="170515935" LSID="urn:lsid:zoobank.org:act:FC39B76A-3AC3-41D7-9420-7148410F6D2C" httpUri="http://treatment.plazi.org/id/E80E167AE9EA56D1B74F036C4920D938" lastPageNumber="93" pageId="0" pageNumber="93">
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<paragraph pageId="0" pageNumber="93">
<taxonomicName LSID="E80E167A-E9EA-56D1-B74F-036C4920D938" authority="Kang" authorityName="Kang" authorityYear="2020" class="Insecta" family="Braconidae" genus="Bohayella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bohayella geraldinae" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="geraldinae" status="sp. nov.">Bohayella geraldinae Kang</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="93">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">Figure 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="materials_examined">
<paragraph pageId="0" pageNumber="93">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Holotype</emphasis>
Costa Rica • ♀; female, Heredia, 3 km S. Puerto Viejo OTS, La Selva; 100 m; x.1992; P. Hanson; huertos Malaise trap set by G. Wright.
<emphasis bold="true" italics="true" pageId="0" pageNumber="93">Paratypes</emphasis>
Costa Rica • 1 ♀; same data as for holotype; xi.1992 • 1 ♂; male; same collecting data as for preceding;
<geoCoordinate degrees="10" direction="north" minutes="26" orientation="latitude" precision="925" value="10.433333">10°26'N</geoCoordinate>
,
<geoCoordinate degrees="84" direction="west" minutes="01" orientation="longitude" precision="925" value="-84.01667">84°01'W</geoCoordinate>
; 4. iv. 1987; H. A. Hespenheide.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="diagnosis">
<paragraph pageId="0" pageNumber="93">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Kang" authorityYear="2020" class="Insecta" family="Braconidae" genus="Bohayella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bohayella geraldinae" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="geraldinae">
<emphasis italics="true" pageId="0" pageNumber="93">Bohayella geraldinae</emphasis>
</taxonomicName>
sp. nov. can be recognized by the following combination of characters: apical maxillary palpomere as long as fifth maxillary palpomere; median crenula of notauli as long as median crenula of scutellar sulcus; scutellar sulcus with one median crenula; hind basitarsus antero-posteriorly slightly expanded; dorsal metasoma mostly pale.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="description">
<paragraph pageId="0" pageNumber="93">Description.</paragraph>
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Female.</emphasis>
Body 4.6-4.8 mm. Forewing length: ~ 4.2 mm Antenna length: ~ 4.8 mm.
<emphasis bold="true" pageId="0" pageNumber="93">
<emphasis italics="true" pageId="0" pageNumber="93">Head</emphasis>
.
</emphasis>
Antenna 34-segmented. Interantennal space with well-developed median carina. POL ~ 1.38
<normalizedToken originalValue="×">x</normalizedToken>
longer than diameter of anterior ocellus (11:8) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3F</figureCitation>
). Eye sparsely setose with short setae; median width of eye 0.75
<normalizedToken originalValue="×">x</normalizedToken>
longer than median width of gena in lateral view (36:48). Width of clypeus ~ 2.07
<normalizedToken originalValue="×">x</normalizedToken>
longer than height (60:29) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3B</figureCitation>
). Malar space ~ 2.62
<normalizedToken originalValue="×">x</normalizedToken>
longer than basal width of mandible (34:13) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3B</figureCitation>
). Mandible bidentate. Maxillary palpus 6-segmented; apical maxillary palpomere as long as fifth maxillary palpomere.
<emphasis bold="true" pageId="0" pageNumber="93">
<emphasis italics="true" pageId="0" pageNumber="93">Mesosoma</emphasis>
.
</emphasis>
Mesoscutum with sharp margin (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3C, F</figureCitation>
). Notauli broadly converging at base, with 11 crenulae; median crenula of notauli as long as median crenula of scutellar sulcus (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3C, F</figureCitation>
). Scutellar sulcus with one median crenula (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3C, F</figureCitation>
). Postscutellar depression present (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3C, F</figureCitation>
). Propodeum rugulose with well-defined median areola; median transverse carina on propodeum reaching lateral margin (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3D, F</figureCitation>
). Pronotum dorso-posteriorly crenulate and antero-ventrally smooth. Mesopleuron dorsally and posteriorly with crenulate margin (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3A</figureCitation>
). Mesosternal sulcus broad and crenulate. Metapleuron rugulose.
<emphasis bold="true" pageId="0" pageNumber="93">
<emphasis italics="true" pageId="0" pageNumber="93">Legs</emphasis>
.
</emphasis>
Basal spur on fore tibia ~ 0.86
<normalizedToken originalValue="×">x</normalizedToken>
longer than basitarsus (30:35). Width of hind femur ~ 0.34
<normalizedToken originalValue="×">x</normalizedToken>
longer than its length (42:124). Basal spur on hind tibia ~ 0.76
<normalizedToken originalValue="×">x</normalizedToken>
longer than basitarsus (58:76). Hind tarsal claw pectinate, with four sharp teeth (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="93">
<emphasis italics="true" pageId="0" pageNumber="93">Wings</emphasis>
.
</emphasis>
Forewing second submarginal cell trapezoidal, ~ 0.35
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum width (30:85); 3r absent (Note: one specimen has basally present 3r vein in particular angle); Rs sharply angled at basal third; stigma ~ 2.67
<normalizedToken originalValue="×">x</normalizedToken>
longer than medial width (80:30). 1CUa short, 0.23
<normalizedToken originalValue="×">x</normalizedToken>
longer than 1CUb (12:52). Hind wing 2-1A absent.
<emphasis bold="true" pageId="0" pageNumber="93">
<emphasis italics="true" pageId="0" pageNumber="93">Metasoma</emphasis>
.
</emphasis>
T1 with a pair of lateral sutures posteriorly reduced, median length of T1 ~ 5.07
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width (71:14) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3D, F</figureCitation>
). T2 with a ball-like projection, medially 0.21
<normalizedToken originalValue="×">x</normalizedToken>
longer than T1 (15:71) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3D, F</figureCitation>
). T3 ~ 2.13
<normalizedToken originalValue="×">x</normalizedToken>
longer than T2 medially (32:15) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3D, F</figureCitation>
). Protruded ovipositor sheath ~ 0.13
<normalizedToken originalValue="×">x</normalizedToken>
longer than hind tibia and apically setose (20:154) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. B. geraldinae sp. nov., holotype A lateral habitus B anterior head C dorsal mesonotum D dorsal propodeum and T 1 - T 3 E hind tarsal claw F dorsal habitus." figureDoi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93">3A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Male.</emphasis>
Body ~ 5.0 mm. Same as female except for the following characters: antenna 32-segmented, melanistic color does not reach the dorsal margin of foramen magnum.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Color.</emphasis>
Body mostly pale; the following areas are melanistic: antenna, vertex, frons, dorsal occiput, maxillary palpus, labial palpus, lateral mesonotal lobe (pale basally), lateral scutellum, margin of metanotum, apical fore femur, fore tibia, apical fore tarsus, apical mid femur, mid tibia, apical mid tarsus, apical hind femur, basal and apical hind tibia, apical hind tarsus, posterior T5 and T6 (weakly), entire T7 and T8, ovipositor sheath. Wings entirely infuscate, stigma entirely melanistic.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="host">
<paragraph pageId="0" pageNumber="93">Host.</paragraph>
<paragraph pageId="0" pageNumber="93">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="distribution">
<paragraph pageId="0" pageNumber="93">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Kang" authorityYear="2020" class="Insecta" family="Braconidae" genus="Bohayella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bohayella geraldinae" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="geraldinae">
<emphasis italics="true" pageId="0" pageNumber="93">Bohayella geraldinae</emphasis>
</taxonomicName>
sp. nov. is known only from the La Selva Biological Station owned and managed by Organization for Tropical Studies (OTS) in Heredia, Costa Rica at an elevation of 100 m (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the species of Bohayella in Costa Rica. Map data 2020 Google." figureDoi="10.3897/zookeys.996.59075.figure1" httpUri="https://binary.pensoft.net/fig/478783" pageId="0" pageNumber="93">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution map of B. geraldinae sp. nov. in La Selva Biological Station in Costa Rica. Map data 2020 Google." figureDoi="10.3897/zookeys.996.59075.figure2" httpUri="https://binary.pensoft.net/fig/478784" pageId="0" pageNumber="93">2</figureCitation>
). The station is located in the Caribbean lowlands, at a confluence of the
<normalizedToken originalValue="Sarapiquí">Sarapiqui</normalizedToken>
river and Puerto Viejo (
<bibRefCitation author="McDade, LA" editor="McDade, LA" journalOrPublisher="University of Chicago Press, Chicago" pageId="0" pageNumber="93" pagination="6 - 14" refId="B18" refString="McDade, LA, Hartshorn, GS, 1994. La Selva Biological Station. In: McDade, LA, Bawa, KS, Hespenheide, HA, Hartshorn, GS, Eds., La Selva: ecology and natural history of a neotropical rain forest. University of Chicago Press, Chicago: 6 - 14" title="La Selva Biological Station." volumeTitle="La Selva: ecology and natural history of a neotropical rain forest." year="1994">McDade and Hartshorn 1994</bibRefCitation>
). According to
<normalizedToken originalValue="Holdridge’s">Holdridge's</normalizedToken>
life zone system (
<bibRefCitation author="Holdridge, LR" journalOrPublisher="San Jose, Costa Rica" pageId="0" pageNumber="93" refId="B14" refString="Holdridge, LR, 1967. Life zone ecology. Tropical science center. San Jose, Costa Rica" title="Life zone ecology. Tropical science center." year="1967">Holdridge 1967</bibRefCitation>
), the station is in the tropical wet forest region (
<bibRefCitation author="Hartshorn, GS" editor="Almeda, F" journalOrPublisher="California Academy of Science, San Francisco" pageId="0" pageNumber="93" pagination="281 - 295" refId="B12" refString="Hartshorn, GS, Peralta, R, 1987. Preliminary description of primary forests along the La Selva-Volcan Barva altitudinal transect, Costa Rica. In: Almeda, F, Pringle, CM, Eds., Tropical Rainforests: Diversity and Conservation. California Academy of Science, San Francisco: 281 - 295" title="Preliminary description of primary forests along the La Selva-Volcan Barva altitudinal transect, Costa Rica." volumeTitle="Tropical Rainforests: Diversity and Conservation." year="1987">Hartshorn and Peralta 1987</bibRefCitation>
), and the average annual precipitation in the area is ~ 4,000 mm (
<bibRefCitation author="Sanford Jr, RL" editor="McDade, LA" journalOrPublisher="University of Chicago Press, Chicago" pageId="0" pageNumber="93" pagination="19 - 33" refId="B22" refString="Sanford Jr, RL, Paaby, P, Luvall, JC, Phillips, E, 1994. Climate, Geomorphology, and Aquatic systems. In: McDade, LA, Bawa, KS, Hespenheide, HA, Hartshorn, GS, Eds., La Selva: ecology and natural history of a neotropical rain forest. University of Chicago Press, Chicago: 19 - 33" title="Climate, Geomorphology, and Aquatic systems." volumeTitle="La Selva: ecology and natural history of a neotropical rain forest." year="1994">Sanford et al. 1994</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.996.59075.figure1" httpUri="https://binary.pensoft.net/fig/478783" pageId="0" pageNumber="93" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Figure 1.</emphasis>
Distribution map of the species of
<taxonomicName authorityName="Belokobylskij" authorityYear="1987" class="Insecta" family="Braconidae" genus="Bohayella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bohayella" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="93">Bohayella</emphasis>
</taxonomicName>
in Costa Rica. Map data 2020 Google.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.996.59075.figure2" httpUri="https://binary.pensoft.net/fig/478784" pageId="0" pageNumber="93" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Figure 2.</emphasis>
Distribution map of
<taxonomicName lsidName="B. geraldinae" pageId="0" pageNumber="93" rank="species" species="geraldinae">
<emphasis italics="true" pageId="0" pageNumber="93">B. geraldinae</emphasis>
</taxonomicName>
sp. nov. in La Selva Biological Station in Costa Rica. Map data 2020 Google.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.996.59075.figure3" httpUri="https://binary.pensoft.net/fig/478782" pageId="0" pageNumber="93" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Figure 3.</emphasis>
<taxonomicName lsidName="B. geraldinae" pageId="0" pageNumber="93" rank="species" species="geraldinae">
<emphasis italics="true" pageId="0" pageNumber="93">B. geraldinae</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="93">A</emphasis>
lateral habitus
<emphasis bold="true" pageId="0" pageNumber="93">B</emphasis>
anterior head
<emphasis bold="true" pageId="0" pageNumber="93">C</emphasis>
dorsal mesonotum
<emphasis bold="true" pageId="0" pageNumber="93">D</emphasis>
dorsal propodeum and T1-T3
<emphasis bold="true" pageId="0" pageNumber="93">E</emphasis>
hind tarsal claw
<emphasis bold="true" pageId="0" pageNumber="93">F</emphasis>
dorsal habitus.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="etymology">
<paragraph pageId="0" pageNumber="93">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="93">This species is named in honor of Dr Geraldine Wright, a former student of the second author (SRS), Rhodes Scholar, professor in the Department of Zoology in the University of Oxford (United Kingdom), and the person who set the trap that collected the specimens.</paragraph>
</subSubSection>
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<paragraph id="167383807B66A5F5730BAB8D38B2C118">
<taxonomicName id="E00DFE24C39080FC7555B8171BF8C1C7" ID-CoL="69HBG" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">
<emphasis id="27B97E7AF3F57C804A9C41F3485F372D" italics="true">Cephalanthera longifolia</emphasis>
(L.) Fritsch
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A3A3F12AB42CFE7B1C49E7E8B6C68D5F" type="vernacular_names">
<paragraph id="5050FB33541F7FB66828ACADF81B6C77">
<vernacularName id="6375F479C064DEFEFE43A79385F6C743">
<normalizedToken id="034D748C9D40DB633B6FE1161695DAF7" originalValue="Langblättriges">Langblaettriges</normalizedToken>
<normalizedToken id="55EF72D4C325D3AE7B48051CBB14A7AE" originalValue="Waldvögelein">Waldvoegelein</normalizedToken>
</vernacularName>
</paragraph>
</subSubSection>
<subSubSection id="9C87368311E4B693A08143E6AD43CADB" type="multiple">
<paragraph id="F66CB60051E34455BEC112115888EDFB">
Art ISFS: 101900 Checklist: 1011200
<taxonomicName id="188190B5A4F95FD46582771757962D32" class="Liliopsida" family="Orchidaceae" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="family">Orchidaceae</taxonomicName>
<taxonomicName id="BD5E8164DAE43246C7994132D6FCF29B" class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="genus">Cephalanthera</taxonomicName>
<taxonomicName id="D3160269D761520DFF69C4940ACEC470" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">Cephalanthera longifolia (L.) Fritsch</taxonomicName>
</paragraph>
<paragraph id="73BDE0AA1A61F263FBE2C3542BE36D64">
<heading id="85E4A4FB64EB5EE936CABA8ABF77B7C4">Zusammenfassung</heading>
</paragraph>
</subSubSection>
<subSubSection id="7AB70F476794AB2F3101186DC224AD38" type="description">
<paragraph id="FF7F59C561B48BD9D5A5CE76AD83EE49">
Artbeschreibung
<emphasis id="A0A29DFE9AA8C3F9515C49DE35EB3DF7" italics="true">
(nach
<bibRefCitation id="4C6B807CA8E36EA99F2C1BE90C11CF06" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
:
<normalizedToken id="5C373B568C89B21D088A82EEE49C39CE" originalValue="Ähnlich">Aehnlich</normalizedToken>
wie
<taxonomicName id="4AD13AFFBDD416A41AA293F885CF42BF" class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="damasonium">
<emphasis id="870FC403719BD1511F05FC76A1EEC2D3" italics="true">C. damasonium</emphasis>
</taxonomicName>
, aber
<emphasis id="E20683998099CF50FA293263D30514A4" bold="true">
<normalizedToken id="F0638040905A340D6E6D70C4232361F6" originalValue="Blätter">Blaetter</normalizedToken>
lineal-lanzettlich, gefaltet und 2zeilig angeordnet
</emphasis>
,
<normalizedToken id="56C29ABA56F149C226A40D792A440EC8" originalValue="Tragblätter">Tragblaetter</normalizedToken>
meist viel
<normalizedToken id="610DB32814D59907973C8CB4F16C484F" originalValue="kürzer">kuerzer</normalizedToken>
als der Fruchtknoten,
<normalizedToken id="75329121AA8FF97EB993559F78B96F75" originalValue="Blütenstand">Bluetenstand</normalizedToken>
bis
<normalizedToken id="E9D6E0F2020FF2BBE10D12F5F46AAFB1" originalValue="über">ueber</normalizedToken>
30
<normalizedToken id="89BBF379DDD6B1F06DF9999C93209451" originalValue="blütig">bluetig</normalizedToken>
.
<emphasis id="95017132824C2E14EE5FA68CE5D8D79C" bold="true">
<normalizedToken id="618CDC7417F0D1795140E48BD061F854" originalValue="Perigonblätter">Perigonblaetter</normalizedToken>
reinweiss, spitz
</emphasis>
, weniger zusammenneigend.
</paragraph>
</subSubSection>
<subSubSection id="D41A92011738251B2CE171853F555B81" type="biology_ecology">
<paragraph id="CBEEF24A894C870CBD2A8A8DBECCE199">
<normalizedToken id="430A81226AF7EAC82B3C1CD77356DF58" originalValue="Blütezeit">Bluetezeit</normalizedToken>
<emphasis id="555E14B9097AFF64E3BAB925E4FB5BFA" italics="true">
(nach
<bibRefCitation id="9973A015C316DCD83D69CE83823B4F89" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
: 5-6
</paragraph>
<paragraph id="3566C0712F7886CD762E204EBF5C5D89">
Standort und Verbreitung in der Schweiz
<emphasis id="E349F822566828C4188422DC3C83312E" italics="true">
(nach
<bibRefCitation id="A8396CFB4557EC7A2FF249436B5531D8" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
: Bergwiesen, Waldlichtungen,
<normalizedToken id="59B5135306BAB0C74F245611C32CBFA1" originalValue="Föhrenwälder">Foehrenwaelder</normalizedToken>
/ kollin-montan / CH
</paragraph>
</subSubSection>
<subSubSection id="2D196A10B845B47D4E9D6FFDB1178913" type="distribution">
<paragraph id="75384F6C3F767DF5D73B12DB1732D469">
Verbreitung global
<emphasis id="627E015EE605E5453FF1F45F5D0DC7F4" italics="true">
(nach
<bibRefCitation id="12150FC7FA272C6902799B70C930D671" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
: Eurasiatisch
</paragraph>
</subSubSection>
<subSubSection id="78FCE1D268BE7DB80EDFB82119F37585" type="biology_ecology">
<paragraph id="BFFC733729DBFE8A922D41991A2F2D97">
<normalizedToken id="75F719414236E0DE588DFE623C92E3BD" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte
<emphasis id="95FC72CD0E75845DCFB263091E8AF78F" italics="true">
(nach
<bibRefCitation id="BF1221978772C420D261A1F6321A6FE9" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. 2010</bibRefCitation>
)
</emphasis>
2 + w + 42-33 + 4.g.2n=32
</paragraph>
</subSubSection>
<subSubSection id="23B68E9F30D603F3D25C5EE4B404FB1A" type="multiple">
<paragraph id="A767C66D6DB36D1FAA15B76A8228F7ED">Status</paragraph>
</subSubSection>
<subSubSection id="58D66D7B08F40FE15BE939765BFA1692" type="conservation">
<paragraph id="F780E81CD9DB4C1848C6EFB2956B615C">
<heading id="7BD214918ECFF8D8F477E3244AB9A6C8">Status IUCN</heading>
: Nicht
<normalizedToken id="A91A613CD33A3FE4420B5097B1EC4F5C" originalValue="gefährdet">gefaehrdet</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="88E79C0E374B07472EFC578D3CFEA335" type="multiple">
<paragraph id="4FA093A37AB91082781617D1DDFD6467">Anatomie</paragraph>
<paragraph id="926085B21DF4AA143F4709E2E4946C83">
<heading id="E67961C66767ED42BDCFDDD69ACFACDB">Zusammenfassung der Stammanatomie</heading>
</paragraph>
<paragraph id="FB1D1A505C9C07704A7E536CFD168A68">
Umriss rund oder oval.
<normalizedToken id="FE0E99A18977FB767FF74D61A7A10F5D" originalValue="Leitbündel">Leitbuendel</normalizedToken>
diffus verteilt. Kleine Interzellularen, oft dreieckig. Epidermiszellen nicht verholzt.
</paragraph>
<paragraph id="A460734CD1E49CB01B24D95059DDD5D3">
<heading id="809C1606F49142C23E5304A83E9FE013">Beschreibung (Englisch)</heading>
</paragraph>
<paragraph id="67DF1505D7A6DA4A21A71E7E7E05CC70">
Culm-diameter
<quantity id="87C21B66329183FBDDD7754B7C61F3B1" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5-10 mm</quantity>
, center full, radius of culm in relation to wall thickness 1:1. Outline circular, with rips. Culm-center full, containing unlignified cells. Culm without endodermis but with a sheath of lignified cells around the center. Epidermis smooth. Large vascular bundles distributed in the whole culm. Chlorenchyma present, continuous peripheral belt with unlignified round cells (like a large cortex). Sclerenchyma belt absent but sometimes with a belt of unlignified cells. Sheath around vascular bundles absent or not lignified. Vessel arrangement horseshoe-like. Largest vessel in the bundle 20-50
<normalizedToken id="77173F36AA47E8DAEA68B6D88FA20A7F" originalValue="μm">μm</normalizedToken>
. Cavities (intercellulars) between parenchyma-cells small, often triangular.
</paragraph>
<paragraph id="FBDBB50AFFAAC7991CD2D7EFE307CF5A">
<heading id="78E3585081E60443724DD08AB71227FF">
<normalizedToken id="F61917C790EDB0C2B22588992A07D253" originalValue="Ökologie">Oekologie</normalizedToken>
</heading>
</paragraph>
<paragraph id="7E5E18BF9E7719E102DB089F3B75762A">Lebensform Geophyt</paragraph>
<paragraph id="1E614F1AC62AE1E3E07960B4CB8F768D">
<heading id="27FD5B27C7BB96CB7ECDBFBAED4B5A4D">Lebensraum Lebensraum</heading>
nach
<bibRefCitation id="7DEFB9C8E845FFE2867DE3C8009B0506" author="Delarze, R. &amp; Gonseth, Y. &amp; Eggenberger, S. &amp; Vust, M." location="Bad Hersfeld, Germany" publisher="Ott Verlag" refString="Delarze, R., Gonseth, Y., Eggenberger, S., Vust, M. 2015. Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition). Ott Verlag, Bad Hersfeld, Germany, 456 pp." title="Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition)" year="2015">Delarze &amp; al. 2015</bibRefCitation>
</paragraph>
<paragraph id="93F8B69702209330482E6416B03142EA">
<table id="7FBD1AE2D4E520DFF52153F51CA05AEF" class="table" inLine="true">
<tr id="EEF247B17985C7C5290265A734C87435">
<td id="17AC4447B887B39014DE030B79E79CBE">
<tbody id="BFE24DBCE2E0FA46E765C97F33BA28E2">
6.2.1 - Orchideen-Buchenwald (
<emphasis id="49749F9AB7F82C27A2FB756D6784F3FD" italics="true">Cephalanthero-Fagenion</emphasis>
)
</tbody>
</td>
</tr>
</table>
</paragraph>
<paragraph id="484A80B8D5C9CE0B6E39915FBAEAC44B">
<tableNote id="88571AA42A3303D88FDB2B97C9F7B4FB">
<emphasis id="91A4455DDEEB25B1B34F6C7C5DC012E0" bold="true">fett</emphasis>
<emphasis id="A8963C3543A3D7E8C74827605F6C894B" italics="true">
Dominante Art, welche das Aussehen des Lebensraumes
<normalizedToken id="012224114C5482DEC41D667759A1D5F9" originalValue="mitprägt">mitpraegt</normalizedToken>
</emphasis>
<emphasis id="5BAB666E53C4B559EBCD81EFD89F7C1C" italics="true">Charakterart</emphasis>
<emphasis id="8AA427CFF4AE1109A1F28991FF8217F1" italics="true">Weniger strikt an den Lebensraum gebundene Art</emphasis>
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="704A8B87F62FDE22CE06A762376D055F" type="biology_ecology">
<paragraph id="DBFF0A067A5A4867D6AEA90BF3AA0599">
<normalizedToken id="4F23DA09D80A9500A9030D2B23F970C2" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="197A506BCB6E55E5EB02C8B9148D49FF" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="6D662571F2D1BCDC6CA2782129853D87">
<table id="049DC1BEDF9AB5312731D6D019D4298C" class="table" inLine="true">
<tbody id="AE12105B5D9458BE9EF727440BE75422">
<tr id="A63CD681F4EDA9DE4753ACE43C82D96C">
<th id="EAFE3D091588395F9DB07833A69E2EB7" colspan="2">
<emphasis id="D4C60AAACC81849D0C228D1A7941D058" bold="true">Bodenfaktoren</emphasis>
</th>
<th id="1EB209765F0B65454880DA5E06985164" colspan="2">
<emphasis id="2FD9353FA809E181D145908C3DD40DA7" bold="true">Klimafaktoren</emphasis>
</th>
<th id="E7F69BC7A00108765D9425DEAF8DAF63" colspan="2">
<emphasis id="6A1D4AEE09484C629226BBE692A893BF" bold="true">Salztoleranz</emphasis>
</th>
</tr>
<tr id="60354440D499F8C21101AB103E09D9FD">
<td id="FA6130977462F84B063FAF36118D92CC">Feuchtezahl F</td>
<td id="7D9579AAFF7A1B9FA83E8FDDD7151655" style="text-align:left;width:100px">
frisch; Feuchtigkeit stark wechselnd (mehr als
<normalizedToken id="8CF882C9B61C7704BDFE25E83A9500DA" originalValue="±">+/-</normalizedToken>
2 Stufen)
</td>
<td id="79DCF521364A3A96229D4FB31684B705">Lichtzahl L</td>
<td id="CE6177316EF79DA66919BC02F80F9C39" style="text-align:left;width:60px">halbschattig</td>
<td id="F716B2682BA3134A37D173346CE9568C">Salzzeichen</td>
<td id="682C3D03E439034FAF78AD8D62B09535" style="text-align:left;width:60px">--</td>
</tr>
<tr id="DB02D9A5576AB00F3569D4A31E28CF95">
<td id="492C814F57CDC6E04593853316D55C2A">Reaktionszahl R</td>
<td id="1A3CD7951E815615845F05F151B11011" style="text-align:left">neutral bis basisch (pH 5.5-8.5)</td>
<td id="91E1FC52681388D22561EF10ABF850E4">Temperaturzahl T</td>
<td id="EC3329319AEC9626DF9FAD93E5632420" style="text-align:left">unter-montan und ober-kollin</td>
</tr>
<tr id="BE332CB7388DC62414C324B91F375055">
<td id="20DF9FCCE54E4CFF8496EEFBF40715D5">
<normalizedToken id="E565A5CA64730156E277A778420D5DA2" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="913C22A200CCA034E6CB33D0407B7ECB" style="text-align:left">
<normalizedToken id="295495A2FC28667A028D3DB97E2AE381" originalValue="nährstoffarm">naehrstoffarm</normalizedToken>
</td>
<td id="E38632DBCA5B44ABAF72EAD8F67BE88B">
<normalizedToken id="ADFA68410E9AD75B735D68AC7F6C4AEA" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="1C3FB7018C6B9BAF1206118F720E58CA" style="text-align:left">subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter)</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="B22FACA17F7A8CB8140DC71C9502E420" type="reference_group">
<paragraph id="B4F2D2EC17F67392BEEF9D7DFE99A67D">
<heading id="A2B0A0BD2B64B990D5891A78DAB15F0C">Nomenklatur</heading>
</paragraph>
<paragraph id="31C1D12483F978E5FD1F1A35BE9274B9">
<treatmentCitation id="62F67F917A086CEFC4A432518FC90BD9">
<heading id="407309889AE7DD856F11E3137B384FEC">
<normalizedToken id="6EB362B0F448A542B3BD11122CE2F06E" originalValue="Gültiger">Gueltiger</normalizedToken>
Name (
<bibRefCitation id="E2087ADBB22EC9C59AD8392217709F1C" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</heading>
:
<taxonomicName id="332F3BDCC56CB34F7F3AC8979CDF7D51" ID-CoL="69HBG" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">
<emphasis id="1DC2BA5F08D038B16CD20FA0E8D9ADF3" italics="true">Cephalanthera longifolia</emphasis>
(L.) Fritsch
</taxonomicName>
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="A1357B5AA506E84C428443A7C926F47D" type="vernacular_names">
<paragraph id="2EB84886D6370A9599F261BFBDD8436B">
Volksname Deutscher Name:
<vernacularName id="A58B143F9C280A1548673DBEE09A210A" language="deu">
<normalizedToken id="19C71C2A88DC3F3BA3F8D47417927CF2" originalValue="Langblättriges">Langblaettriges</normalizedToken>
<normalizedToken id="BD41D530EE395739CFE79C3DEBCD64E7" originalValue="Waldvögelein">Waldvoegelein</normalizedToken>
</vernacularName>
,
<vernacularName id="45E52EA91349AEEE8EB7167653F947AD" language="deu">
<normalizedToken id="8519CDF4CC105FFDB4B2EBA85A70EDB1" originalValue="Langblättrige">Langblaettrige</normalizedToken>
Kopforche
</vernacularName>
Nom
<normalizedToken id="03F41ECD88F98BAE39BB8B64A3CBB0A3" originalValue="français">francais</normalizedToken>
:
<vernacularName id="60687827B555A43615BBA0248195C452" language="fra">
<normalizedToken id="D7A22059068A126F8702A6BC6AC5946E" originalValue="Céphalanthère">Cephalanthere</normalizedToken>
<normalizedToken id="E23FAEDF2FC19B391E5CA01C6A155402" originalValue="à">a</normalizedToken>
longues feuilles
</vernacularName>
Nome italiano:
<vernacularName id="071A2509485554F4F0C6237A5075DCC3" language="ita">Cefalantera maggiore</vernacularName>
</paragraph>
</subSubSection>
<subSubSection id="11D71A461545D2DD9AFB0E204012E592" type="reference_group">
<paragraph id="AA6D8402506469385741AD7372BC68C6">
<normalizedToken id="E344F426FA19128E423486F9F4AACC13" originalValue="Übereinstimmung">Uebereinstimmung</normalizedToken>
mit anderen Referenzwerken
</paragraph>
<paragraph id="B3C8A4C0BCD48B700F79D1626AE3FD39">
<table id="7D674A6B6BA6AF7BDB8848EEE2C23FC6" class="table table-condensed" inLine="true">
<tbody id="A96B726AAF8DFF9D7C0F48A61664D981">
<tr id="26272498095E7ECDAC8BF94087FC246E">
<th id="0A609CA56871BFB5B917F5B7544DCE71" width="80px">Relation</th>
<th id="EDEF3132B86F1303D59EFCD6A2D137EC">Nom</th>
<th id="96FEEAD7F2DF436023E82650284BF3FE">Referenzwerke</th>
<th id="736CE01B7360314C8725992046C8662F" width="90px">No</th>
</tr>
<tr id="F62826DF87A2EE45C7FDF048F4555AB2">
<td id="7054643E8441F1F0F9D5BE27BA035605" class="taxon-relation">=</td>
<td id="41F0AC10CA980F61482A3AFB1DFACE19">
<taxonomicName id="B958A8B918B5B047A8D2606694823C7C" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">Cephalanthera longifolia (L.) Fritsch</taxonomicName>
</td>
<td id="7922F6566D94CCCBB2C52CE861B04815">
<treatmentCitation id="54011BB265837CA5F275AB52D086525C">
<bibRefCitation id="01F83B690C83EB9B31253868DC2DBAA4" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
</treatmentCitation>
</td>
<td id="8BBC3CC49C4CD2DCC21B1B74C2F25AB3">101900</td>
</tr>
<tr id="CA351F463C375142B6B7522BA4CB15E0">
<td id="2D677E20B0462BCF521B81A08773FC07" class="taxon-relation">=</td>
<td id="73D5BBD7A7B10F8114C2E279FAC66F5F">
<taxonomicName id="A11809116E5449D8987DD8C6E911BCDC" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">Cephalanthera longifolia (L.) Fritsch</taxonomicName>
</td>
<td id="CD2A57505C4B17237148848F9574AB8F">
<treatmentCitation id="2F702983033340DD74C982A48ADCE873">
<bibRefCitation id="5FA1935A65DB6651479F0B722E900A52">Flora Helvetica 2001</bibRefCitation>
</treatmentCitation>
</td>
<td id="3D108BA24E0D7D81FF23F34BAA800C78">2936</td>
</tr>
<tr id="F78A6CC319399F7F1ED295C78FA79CB1">
<td id="B8B8405EE422C9772DF5BA68794D577D" class="taxon-relation">=</td>
<td id="11E307A9BBD708F61271898930651C6B">
<taxonomicName id="F84F7CCE0075A93B2379AD61B0CFCC42" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">Cephalanthera longifolia (L.) Fritsch</taxonomicName>
</td>
<td id="B022203922261A057A256CF00A42047E">
<treatmentCitation id="0FC99FA9864D89028508B13355BAA7A1">
<bibRefCitation id="64678AF3AA746562626980B8A887382A">Flora Helvetica 2012</bibRefCitation>
</treatmentCitation>
</td>
<td id="1D35C1FE4ADD265A45FE233A863555CE">2520</td>
</tr>
<tr id="E07A5817055FB95AE975A7BFB740596A">
<td id="A3A0F887A17BB839540426102330B984" class="taxon-relation">=</td>
<td id="7434BDAE9D61EC066C2BB7D7AC6B327D">
<taxonomicName id="918367A2C4722C2A7E43BF4531A54959" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">Cephalanthera longifolia (L.) Fritsch</taxonomicName>
</td>
<td id="8EC308E5F6F1FDBABEBD736467D9A4E6">
<treatmentCitation id="2B8E254B7CA47E7F8D514A04F7767B71" httpUri="http://treatment.plazi.org/id/673A83FC5C0BA9AF8EC5BBF3CF6EFCD3">
<bibRefCitation id="0721AA5CD51EDF629540BC5EAC4D6C15">Flora Helvetica 2018</bibRefCitation>
</treatmentCitation>
</td>
<td id="5126262DF24070A6085F3C56DE857BB2">2520</td>
</tr>
<tr id="9450D9388BB55BB79C041BDE8D23CDEB">
<td id="5B03D6B0C75AEDBCAEB5419DC200A3A4" class="taxon-relation">=</td>
<td id="1F3703B93210EEB605F5F430103CEB23">
<taxonomicName id="0D53B7152A16F8828CB57B4838D53005" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">Cephalanthera longifolia (L.) Fritsch</taxonomicName>
</td>
<td id="B6A7022310F21A1657162AAF99DB71ED">
<treatmentCitation id="1697426E3EF487DB453D92607FAB4BAC">
<bibRefCitation id="C34F1EE5DC2BE6DA32CFEE9DA582AC09">Index synonymique 1996</bibRefCitation>
</treatmentCitation>
</td>
<td id="D7506CCE35452CFED0300F7133D16115">101900</td>
</tr>
<tr id="218F1C8E17D5473E8E5EC657B254EEED">
<td id="898C42AF5E6ADF3C9280B2CE8C29730E" class="taxon-relation">=</td>
<td id="2DE7B488C351AC074B2B15BEC9472F7E">
<taxonomicName id="4CB607C077DEC38738451348FB3594A4" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">Cephalanthera longifolia (L.) Fritsch</taxonomicName>
</td>
<td id="D08BC2FE492965BD60194C4778050718">
<treatmentCitation id="03BF52F8897CEC9419AAA3797A13B3DB">
<bibRefCitation id="1B57B03C9DAFC18A32EE5D08796B2744">Landolt 1977</bibRefCitation>
</treatmentCitation>
</td>
<td id="11D0160ECAC5189B2DCD9E6036AAFA18">729</td>
</tr>
<tr id="283248357230A79D61209503F78DBF41">
<td id="B9EF715FA53738CB442AF15741DD4BDD" class="taxon-relation">=</td>
<td id="FC125D02527983383CBBD54D694CAB80">
<taxonomicName id="233D0A90673BD9210646F76A3573DF4B" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">Cephalanthera longifolia (L.) Fritsch</taxonomicName>
</td>
<td id="9A8995FDFF91255E19D61990AE741992">
<treatmentCitation id="06527823A3F55E8A4C7CFDC1C2FBE5A1">
<bibRefCitation id="5AC9F5B70BC13FA4D8DFABC63737C18B">Landolt 1991</bibRefCitation>
</treatmentCitation>
</td>
<td id="90B40AE4A3A4DC91E6484D33400DFF4E">636</td>
</tr>
<tr id="ABEE813A7C45BAC9B3E2A8A29D7F2B13">
<td id="392639DC5C73D48EBF053025FD544454" class="taxon-relation">=</td>
<td id="1094797E2D083703BF699E853E97212D">
<taxonomicName id="494F2A0600CD631EEE258F5B8C51470C" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">Cephalanthera longifolia (L.) Fritsch</taxonomicName>
</td>
<td id="E25C66603F5FDAF645BFFD1EE5E31B63">
<treatmentCitation id="CC3D0D62B9F0F5FCD56CE2F6AE6712E6">
<bibRefCitation id="989ECB8B98B95311999366D80837DE6F">SISF/ISFS 2</bibRefCitation>
</treatmentCitation>
</td>
<td id="1C387A519F0E4D1361A6537D09A6E0F9">101900</td>
</tr>
<tr id="266D30E02CE9B8381B78AE1CD5102327">
<td id="38B72AC9532DF201BB2F869C430AD01F" class="taxon-relation">=</td>
<td id="479DF2FBC0EACCE22A0DA12F2C44FBD8">
<taxonomicName id="8E6B6F75B8ABF0726FAD338C2CEA8D2E" authority="(L.) Fritsch" authorityName="Fritsch" baseAuthorityName="L." class="Liliopsida" family="Orchidaceae" genus="Cephalanthera" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="species" species="longifolia">Cephalanthera longifolia (L.) Fritsch</taxonomicName>
</td>
<td id="BFA00D632342750FC4EA6BF32BEA1EB6">
<treatmentCitation id="8A0965C144966D204CD98BE472D1FCDA">
<bibRefCitation id="F9750814E373E6B241E4CB5560246C8A">Welten &amp; Sutter 1982</bibRefCitation>
</treatmentCitation>
</td>
<td id="FF0BF6A37EBBE8B059226D8EA610FA50">2522</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="D99489DDCC81F829583377BF7F9CF81B">
<tableNote id="D3A6B80AA8E93BD6AF18DB22EFD34683">
= Taxon stimmt mit akzeptiertem Taxon
<normalizedToken id="30EEDF89331BAD36F3D159D652066B95" originalValue="überein">ueberein</normalizedToken>
(
<bibRefCitation id="BFF89C76DDAF5ACFD4CFFE1D4532D130" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) &lt;Taxon ist im akzeptierten Taxon (
<bibRefCitation id="5FD3FA62E93D87F6800D6422059AB8EB" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) enthalten&gt; Taxon
<normalizedToken id="3CE8DC63B0B2AC4C13D32275C27D2367" originalValue="enthält">enthaelt</normalizedToken>
(neben anderen) auch das akzeptierte Taxon (
<bibRefCitation id="B544E9F956A5ACE49B8D5F6C094A4D67" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="4B6BA844CA993C74E946667AACEDF874" type="conservation">
<paragraph id="0E11DF8079A3BE01765206C19B40F916">
<heading id="31D77110A38AF5985CDCD0AC6D5F30BE">Status Indigenat</heading>
: Indigen
</paragraph>
<paragraph id="908AD2B21B2E9795C569CDB6B91404B9">
<emphasis id="61C04D9871445ADFCC21D6832F37F5A5" bold="true">
<heading id="BF8B5B42C4C5CBF1B71AF1AE89B18FF1">
Liste der
<normalizedToken id="831E95838CD3B613AD6205E6B41186F5" originalValue="gefährdeten">gefaehrdeten</normalizedToken>
Pflanzen IUCN
</heading>
(nach
<bibRefCitation id="F6056FCD27E1FDF2627A0E1E96B80F92" author="Walter, K.S. &amp; Gillett, H.J." location="Gland, Switzerland and Cambridge, UK" publicationUrl="https://doi.org/10.5962/bhl.title.44833" publisher="IUCN" refString="Walter, K.S. and Gillett, H.J. (editors). 1998. 1997 IUCN Red List of Threatened Plants. IUCN, Gland, Switzerland and Cambridge, UK. DOI: https://doi.org/10.5962/bhl.title.44833" title="IUCN Red List of Threatened Plants" year="1997">Walter &amp; Gillett 1997</bibRefCitation>
):
</emphasis>
Nein
</paragraph>
<paragraph id="14BFCB2DD65236222E324DE72AE1905A">
<heading id="27CA79A8A8E70B305517FB2A43FCA6B2">Status Rote Liste national 2016</heading>
</paragraph>
<paragraph id="4874F0DED59FE24DD9ED71E1C92D5177">
<heading id="FF03BF23D7F2AACC0CDD4D83EFE8CB0A">Status IUCN</heading>
: Nicht
<normalizedToken id="8805D16DBB7CEFE1314AB34566E93DD6" originalValue="gefährdet">gefaehrdet</normalizedToken>
</paragraph>
<paragraph id="76DF3BD1F286AD64FF89AF64471507B0">
<heading id="5BE50162240D47937BE4C7140DFCB1A7">
<normalizedToken id="CC914BBB6F1F4B079898C7CFDBFD1B1F" originalValue="Zusätzliche">Zusaetzliche</normalizedToken>
Informationen
</heading>
Kriterien IUCN: --
</paragraph>
<paragraph id="ACB09D7EBBC047605CD16C1AAE619854">
<heading id="DF89CE8AD3A92C4ED6BCA4FF53D0ED96">Status Rote Liste regional 2019</heading>
</paragraph>
<paragraph id="98660BC327930E0FF33A36D367FA21EA">
<table id="59F5A7A357DE1B7D71D6DF05B9721CCD" class="table table-condensed" inLine="true">
<tbody id="547043ECD1C6DD3E8FF568091AF3E458">
<tr id="3A87C03B14808AC1B274E7A2C79C19F9">
<th id="D56864537BE45F074925E3212FDD057D">Biogeografische Regionen</th>
<th id="987CF058C0805C7863D53C503B201729">Status</th>
<th id="359E9856B740DC34F4DE9925C8614B10">Kriterien IUCN</th>
</tr>
<tr id="BAEA73CF3D049A9B89686F7F7E4E40F4">
<td id="136C9528A4CBB01153CDFD75C9972D55" class="regional_status">Jura (JU)</td>
<td id="DC4FD02E71530A2CC0DD3E093E85017C">
nicht
<normalizedToken id="604732169D5086C874E2593B710A49AC" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="3367509F379FC7C479AC25DABCDD49C4">
<td id="5994E148A31533559B0012866554D1F2" class="regional_status">Mittelland (MP)</td>
<td id="1A437CBD5BC25A306B2D32C5EB021E5B">
potenziell
<normalizedToken id="A4440DEABDE32A0D2DB23C1F7369D5B6" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Near Threatened)
</td>
<td id="6CA92F5CECCAE823F228F698A527E45C">A3c</td>
</tr>
<tr id="C432A4D5A26CECBD7DE10313697E8240">
<td id="76AF3249627B7DFB3EF2E15C9A222BC2" class="regional_status">Alpennordflanke (NA)</td>
<td id="68474CA1461AF5576791CEF0C099B66F">
nicht
<normalizedToken id="0CA483F857AC909A7D086EC9B2B0F409" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="445E1234403FEC38199D29B105C2876B">
<td id="A537AC2DDA8C2A585D33DDF308D6A82F" class="regional_status">
<normalizedToken id="71F28EC119A936FC8C81C46733D89326" originalValue="Alpensüdflanke">Alpensuedflanke</normalizedToken>
(SA)
</td>
<td id="8EE4C49193C66F00AB2C682BC0694F01">
nicht
<normalizedToken id="9087CC39EB9F2CEF711B2D906F0D3DBC" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="99A41A44458B63E8D7C82C0A51BBACBB">
<td id="1EE56D35EF397E09A07B0E21D3043696" class="regional_status">
<normalizedToken id="CDE38D5B6388291BA0A141BD176B5304" originalValue="Östliche">Oestliche</normalizedToken>
Zentralalpen (EA)
</td>
<td id="C604DD50DE24998155AF02494492E274">
nicht
<normalizedToken id="09B51356C4B210953E4BAD7E305ADC33" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
<tr id="8ED28EF545C820FBBA9B62E47824873B">
<td id="9D580189B0004099C209254306C0CEF4" class="regional_status">Westliche Zentralalpen (WA)</td>
<td id="3BA78BA62DCEB53C90CC94A59DC956A2">
nicht
<normalizedToken id="3FEF047CC504CE118C07DE9D0D6CE735" originalValue="gefährdet">gefaehrdet</normalizedToken>
(Least Concern)
</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="3E8FDBAFB310A97EFF57B1F063E3B4AF">
<heading id="F58AE93981707C0C0316D9D38421C170">
Status nationale
<normalizedToken id="054D8A5577E86A56CEB6A46A41745704" originalValue="Priorität">Prioritaet</normalizedToken>
/Verantwortung
</heading>
</paragraph>
<paragraph id="BF6E9D9264708D4BF6EF5F29541B2546">
<table id="94274B740AE5A5259BAD034239181A06" class="table" inLine="true">
<tr id="2FA20C5BF0F854E255F7A35E55B86C6C">
<td id="A873078FA4E77D2ACA324B24859EA8D7">
Keine nationale
<normalizedToken id="38BDBA990FEF72EE1F8BC8629043C2F0" originalValue="Priorität">Prioritaet</normalizedToken>
oder internationale Verantwortung
</td>
</tr>
</table>
</paragraph>
<paragraph id="DBECD35EDF985B21415844E5E1445522">
<heading id="D2FE4A9E9338EA60BBCAAF3944E43DCC">Schutzstatus</heading>
</paragraph>
<paragraph id="81C617C345502471393DAFEB589EBBBB">
<table id="4FA24A26DAAC7CD921ED5015D5B604D8" class="table table-condensed" inLine="true">
<tbody id="368AEF3EDB59E6FDA4008537DA161314">
<tr id="7381F058DBECF99C76138DA8D9FA0CFF">
<td id="5BB6FDCC07984679FAB86834F31C4087" colspan="2">
<emphasis id="B1F1B0CB0E1F6C9707A3B8A2F07A1FDB" bold="true">International (Berner Konvention)</emphasis>
</td>
<td id="F81A4843729A385C776D1A336BED1171" width="90px">Nein</td>
</tr>
<tr id="20E6C872FA925A77D925386921052D96">
<td id="B606D01AC706DB63051D0597B30D3523">
<emphasis id="FF602982733776DC1AA70E05DE132628" bold="true">AG</emphasis>
</td>
<td id="F82FF8A76F6F77AC6EB935BE4957D4AC" width="150px">
<normalizedToken id="E7136B4EB00F81973F9E263CF78DD6E3" originalValue="Vollständig">Vollstaendig</normalizedToken>
<normalizedToken id="3E61FAD9B393F45B963250979FBBE567" originalValue="geschützt">geschuetzt</normalizedToken>
</td>
<td id="0259A313D47437FF9953AFE50FBB8958">(01.01.2010)</td>
</tr>
<tr id="52EFEC56D5F24FE3BC8166EF17989374">
<td id="DF684AB79D49875F9ED1EB6F6352608D">
<emphasis id="F9EB1F1E12D2E7D223B9B07116C360EF" bold="true">BE</emphasis>
</td>
<td id="B9C99A82390E6D704B542899F5ABB189" width="150px">
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<td id="9FD1CF1219CC22B88D243FE22219E5F7">(01.01.2016)</td>
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<emphasis id="6E6937CA2982901203FF7F24225853F3" bold="true">FR</emphasis>
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<td id="A4DEA7195FE41EB80E430EF4F709C138" width="150px">
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<td id="0527293FCC1F7B4A003D64DCD51AD622">(12.03.1973)</td>
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<td id="CE38424E67B5E1B0419693589CA2FC44">
<emphasis id="2B061F18FAF0227716C1FC172133E540" bold="true">GL</emphasis>
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<td id="E7D261A699FBC6B871A16C73C8A2B5C9" width="150px">
<normalizedToken id="700C9613D30848A50BF83C52C4B067C6" originalValue="Vollständig">Vollstaendig</normalizedToken>
<normalizedToken id="76ED64716DDB58A7DD03630DA3B3CC9E" originalValue="geschützt">geschuetzt</normalizedToken>
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<td id="B9CB960BDEDBB33306E3BA0482A59078">(07.05.2006)</td>
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<tr id="4A071B7CBA9A552208C86D0B1C3B9851">
<td id="E9F211768290578EE4564B44B5CA5991">
<emphasis id="6041D575D5D0217480F6B217810142EC" bold="true">GR</emphasis>
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<td id="C5CD76B4CAE96A827BF27B7527A8AFBB" width="150px">
<normalizedToken id="B9C8331EFA4FBFE6C0B0AF93DCF82814" originalValue="Vollständig">Vollstaendig</normalizedToken>
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</td>
<td id="4A7BAA7064DD0F319CD986CB031B8B8C">(01.12.2012)</td>
</tr>
<tr id="7D147F7D8812B96D7E9D43FBBA9D7B23">
<td id="27D45DB667E200FADA326A4A9B7823FC">
<emphasis id="05A3130D158E02A22D3963FC3858B0C7" bold="true">JU</emphasis>
</td>
<td id="4B222B99F6F0B46BD80A5F76C25C3718" width="150px">
<normalizedToken id="3EC959D4C05F1556BCA9D2A9E0ABABD1" originalValue="Vollständig">Vollstaendig</normalizedToken>
<normalizedToken id="80F41D226B31A1DF7869C94D0AA44419" originalValue="geschützt">geschuetzt</normalizedToken>
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<td id="4242DB86EC72E53EC992577A8078608E">(06.12.1978)</td>
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<tr id="5ED696B181BD449BAEC57168F42D7980">
<td id="D5477FD38452CEB1121FF85E733DC44D">
<emphasis id="584FF0E2D4D0EFA25E3104202772733E" bold="true">OW</emphasis>
</td>
<td id="CA95BD2A1C2F65E62F584F61139C8285" width="150px">
<normalizedToken id="4C1C5E6E50E3BA9D701C34FB0743FE21" originalValue="Vollständig">Vollstaendig</normalizedToken>
<normalizedToken id="212A3EEE2948CCBF9380DB0EA2878DD9" originalValue="geschützt">geschuetzt</normalizedToken>
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<td id="22BC43FD66AFB3D24C8C003869AADD9C">(01.04.2013)</td>
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</tbody>
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</paragraph>
<paragraph id="98D1C8252EC15C35166E5E7C961929E1">
<table id="573266E7FB0AC4311CE3BEA188E8D0E3" class="table table-condensed" inLine="true">
<tbody id="B567579600D5AE4798F7A61281044AD3">
<tr id="831597F055B7F279822000857DA45458">
<td id="69EDD25B7F71558C885445FF03B167FA">
<emphasis id="46E1BEF79007F26DA7A964EBD334DC4F" bold="true">Schweiz</emphasis>
</td>
<td id="E459B9EB692ECF4F4A8B276877FC1E06" width="150px">
<normalizedToken id="27886C9544712D1CDB9EE1FB070E8626" originalValue="Vollständig">Vollstaendig</normalizedToken>
<normalizedToken id="332DB29027B67D47DCE2F94948943E28" originalValue="geschützt">geschuetzt</normalizedToken>
</td>
</tr>
<tr id="3A4F56B17FFEAF3E15A4339404597CAE">
<td id="BDD2DF5FB685B3DA2D50DA062505270B">
<emphasis id="B7E8650C36A4AAAB153E651EECAA8CE8" bold="true">SO</emphasis>
</td>
<td id="C8B85893051629FCF7F7CEB7835D849B">
Teilweise
<normalizedToken id="CFD232C0BB8277DC00A5AB8ABEDCA81D" originalValue="geschützt">geschuetzt</normalizedToken>
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<td id="7E344167756D579E319BDD202082D3A1">(23.02.1972)</td>
</tr>
<tr id="550029ABFDFA4486AD28048AAD135040">
<td id="352C4264580B8FB04829DD66701A4636">
<emphasis id="1CC09CD98CBADC49FA0B4FFB32DD2BAC" bold="true">VD</emphasis>
</td>
<td id="EE7C6D1E863EE2BDC23258712D29E606">
<normalizedToken id="28E786DE9F9F3870E6EED8803C7D3D30" originalValue="Vollständig">Vollstaendig</normalizedToken>
<normalizedToken id="422ACE667BE11714067A08047C491169" originalValue="geschützt">geschuetzt</normalizedToken>
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<td id="F642D120A982C3C84EB5805BF0557D75">(02.03.2005)</td>
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<tr id="355633F391744386547FCD885085B050">
<td id="884D491C32E77F8A6FF8076CEB79D2B2">
<emphasis id="B1EFEA7F379DD9BE3709349203DEBE08" bold="true">ZG</emphasis>
</td>
<td id="0B27A895FE8D7657ABF21F71025DA3B8">
<normalizedToken id="9931795355102D26F9C1D610547204CB" originalValue="Vollständig">Vollstaendig</normalizedToken>
<normalizedToken id="24D255C7B8EA2A4E24F84371D6B58352" originalValue="geschützt">geschuetzt</normalizedToken>
</td>
<td id="A9E0A04C27D8C364EAC331363744282B">(01.10.2013)</td>
</tr>
<tr id="4EB3C07CB384EB2C2574875FD36AB0F5">
<td id="B34F1D524B01FB6A713FCF1931A95B50">
<emphasis id="D056811F6D2E418A785ED4F35FBF6BC8" bold="true">ZH</emphasis>
</td>
<td id="CE20FE43C1AAD8164215C6002A5D6003">
<normalizedToken id="34769EC6E82EDC2A1C434AAF5A925502" originalValue="Vollständig">Vollstaendig</normalizedToken>
<normalizedToken id="93D78453287CFB713B7BF166E337AB9D" originalValue="geschützt">geschuetzt</normalizedToken>
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<td id="514BA1250F41AC58F9C7B2232EC75E93">(03.12.1964)</td>
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<tr id="8E20DDE16E831F76B0A173B68C85CEB6">
<td id="3D185380AA8B1E6186B0425A11D1032A">
<emphasis id="56E83BB8B66899702E6156B323087CB6" bold="true">TI</emphasis>
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<td id="C8055D89A5619B197E7123843D62C48F">
<normalizedToken id="223006D34114B4F301EB5F9D8996D037" originalValue="Vollständig">Vollstaendig</normalizedToken>
<normalizedToken id="54311737A5FEBF81194168D135432D46" originalValue="geschützt">geschuetzt</normalizedToken>
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<td id="AAAA3B8682B6F3600A7C6C008F63A938">(23.01.2013)</td>
</tr>
<tr id="25113C91381BF76F6AAC757A6A49D296">
<td id="BA162F23C32C7D049F11EC4353E84224">
<emphasis id="7F1466660B67685CD0AC020BCB6F13D5" bold="true">AI</emphasis>
</td>
<td id="C6624511EACD50C4912750DDB97158B4">
<normalizedToken id="30836D0606EC59379C4B36D2161D9414" originalValue="Vollständig">Vollstaendig</normalizedToken>
<normalizedToken id="5C98585A5C9DFF6395777F4BD752879C" originalValue="geschützt">geschuetzt</normalizedToken>
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<td id="5CAA19DF324B84585CC1908DF684AB16">(13.03.1989)</td>
</tr>
<tr id="5282DEA7FEE3FF599BBEA3B19CBBAE0F">
<td id="A247CD1EB5E70081761A8449492B4AA3">
<emphasis id="F81809B4D31FE642DE7EF4341530821A" bold="true">BL</emphasis>
</td>
<td id="F63AFC3472F1ADF0EC6117730E9A1525">
<normalizedToken id="C1B96C38BE6714D7B3D03B06FFACAAD3" originalValue="Vollständig">Vollstaendig</normalizedToken>
<normalizedToken id="CB78AE61F454548E8C4BE11DE22AA2A5" originalValue="geschützt">geschuetzt</normalizedToken>
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<td id="241BE742632CA28B68313C25A95FFF52">(01.01.2012)</td>
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<mods:detail id="9ECF872338CFF273F78562FE15E72BEA" type="issue">
<mods:number id="E02520C4360AA490F2CF1681E7717AB4">4</mods:number>
</mods:detail>
<mods:extent id="F41B35A081D41881B488EC9AA31649E7" unit="page">
<mods:start id="A220A79B09EC4CE9898926D9580CE69B">572</mods:start>
<mods:end id="07217063BC252A5B65D2A39B0CC1E162">580</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8BA0B65AE326C80BD8633B75E7A8F333">journal article</mods:classification>
<mods:identifier id="D34CD7FF9321148B54DD5C7AF2425717" type="DOI">10.11646/zootaxa.3640.4.5</mods:identifier>
<mods:identifier id="BDDF71BC72C7A7A64E581265DF554DF5" type="GBIF-Dataset">9f4f85be-dc51-447e-b1bf-ad5b9c3f086f</mods:identifier>
<mods:identifier id="F2784A10903944D0D425D6CA94AC27E6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B7B2C0837CD75FB6FE87F8C437E15CE0" type="Zenodo-Dep">215844</mods:identifier>
<mods:identifier id="0C22D3E39E18C6E37E8F27DA24FBF4BB" type="ZooBank">A584752D-CA26-498C-B468-02F3A49EB5E7</mods:identifier>
</mods:mods>
<treatment id="E80E87B1FF96FF811C8FF8FCFCD2FEB9" ID-DOI="http://doi.org/10.5281/zenodo.6159400" ID-GBIF-Taxon="119566392" ID-Zenodo-Dep="6159400" LSID="urn:lsid:plazi:treatment:E80E87B1FF96FF811C8FF8FCFCD2FEB9" httpUri="http://treatment.plazi.org/id/E80E87B1FF96FF811C8FF8FCFCD2FEB9" lastPageId="7" lastPageNumber="578" pageId="5" pageNumber="577">
<subSubSection id="28BD652CFF96FF811C8FF8FCFCD2FEB9" lastPageId="7" lastPageNumber="579" pageId="5" pageNumber="577" type="nomenclature">
<paragraph id="601836A7FF96FF831C8FF8FCFD4BF8FB" blockId="5.[151,688,1860,1919]" box="[151,688,1860,1886]" pageId="5" pageNumber="577">
<heading id="3B5081CBFF96FF831C8FF8FCFD4BF8FB" bold="true" box="[151,688,1860,1886]" fontSize="11" level="1" pageId="5" pageNumber="577" reason="1">
<emphasis id="52D3EAB5FF96FF831C8FF8FCFD4BF8FB" bold="true" box="[151,688,1860,1886]" pageId="5" pageNumber="577">
<taxonomicName id="A7A74D24FF96FF831C8FF8FCFD51F8FB" ID-CoL="TGLM" authority="Waterhouse 1882:1" authorityName="Waterhouse" authorityPageNumber="1" authorityYear="1882" box="[151,682,1860,1886]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF96FF831C8FF8FCFE4AF8FB" bold="true" box="[151,433,1860,1886]" italics="true" pageId="5" pageNumber="577">Chalcophora mexicana</emphasis>
Waterhouse 1882:1
</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
<paragraph id="601836A7FF96FF831C8FF8DFFEA2F8DA" blockId="5.[151,688,1860,1919]" box="[151,345,1895,1919]" pageId="5" pageNumber="577">(Figures 5–8, 14)</paragraph>
<paragraph id="601836A7FF96FF801C8FF817FC83FAF1" blockId="5.[151,1436,1967,2028]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="578" pageId="5" pageNumber="577">
This species too was described from a single female specimen (BMNH), and is labeled “
<typeStatus id="BF1C8805FF96FF83189BF817FB3BF862" box="[1155,1216,1967,1991]" pageId="5" pageNumber="577">Type</typeStatus>
/S.
<collectingCountry id="18B07637FF96FF8318FBF817FAC2F862" box="[1251,1337,1967,1991]" name="Mexico" pageId="5" pageNumber="577">Mexico</collectingCountry>
Oaxaca/
<taxonomicName id="A7A74D24FF96FF831C8FF86BFE5AF84E" box="[151,417,2003,2027]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF96FF831C8FF86BFE5AF84E" box="[151,417,2003,2027]" italics="true" pageId="5" pageNumber="577">Chalcophora mexicana</emphasis>
</taxonomicName>
,
<typeStatus id="BF1C8805FF96FF831DACF86CFE10F849" box="[436,491,2004,2028]" pageId="5" pageNumber="577">Type</typeStatus>
C. Waterh.” It measures
<quantity id="A75F9B42FF96FF831F03F86BFC7FF84E" box="[795,900,2003,2027]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" pageId="5" pageNumber="577" unit="mm" value="26.5">26.5 mm</quantity>
long. One of us (RLW) examined nine other specimens from BMNH labeled ”Zapotlan, Colima, Höge/B.C.A., III.
<taxonomicName id="A7A74D24FF95FF801FB4FF20FB33FF15" box="[940,1224,152,176]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="virginiensis">
<emphasis id="52D3EAB5FF95FF801FB4FF20FB33FF15" box="[940,1224,152,176]" italics="true" pageId="6" pageNumber="578">Chalcophora virginiensis</emphasis>
</taxonomicName>
.” They are instead from Jalisco, Ciudad Guzmán (Selander and Vaurie 1962). This fact must have been recognized by Waterhouse (1889:167), who discussed those specimens and recorded the locality to be in that state. Ciudad Guzmán is in the municipality of Zapotlán el Grande, which name, though it is old, currently sometimes is used for the city. It is within about 25 kms of the Colima border, so it is possible that the actual collection was made in that state. Regardless,
<taxonomicName id="A7A74D24FF95FF801D39FEF5FE55FEC1" box="[289,430,333,356]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF95FF801D39FEF5FE55FEC1" box="[289,430,333,356]" italics="true" pageId="6" pageNumber="578">C. mexicana</emphasis>
</taxonomicName>
must occur there. Another specimen sent from BMNH is from Juquila [Oaxaca, 16°14’, 97°18’, 4900’, fide Selander and Vaurie 1962]. All
<specimenCount id="76A1FD2EFF95FF801EEDFED7FC6AFE22" box="[757,913,367,391]" pageId="6" pageNumber="578" type="generic">10 specimens</specimenCount>
bear Waterhouse’s determination label as
<taxonomicName id="A7A74D24FF95FF80199AFEC8FEE3FE09" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="virginiensis">
<emphasis id="52D3EAB5FF95FF80199AFEC8FEE3FE09" italics="true" pageId="6" pageNumber="578">C. virginiensis</emphasis>
</taxonomicName>
and all are male. They are part of the
<emphasis id="52D3EAB5FF95FF801EFDFE2CFBC9FE09" box="[741,1074,404,428]" italics="true" pageId="6" pageNumber="578">Biologia Centrali-Americana</emphasis>
Collection, and RLW placed a determination label on each. In a series of
<specimenCount id="76A1FD2EFF95FF801EB9FE0FFCC5FE6A" box="[673,830,439,463]" pageId="6" pageNumber="578" type="generic">48 specimens</specimenCount>
(RLWE) of this species examined, 15 are female. Additional records from Jalisco, based on
<specimenCount id="76A1FD2EFF95FF801E9FFE64FCDAFE51" box="[647,801,476,500]" pageId="6" pageNumber="578" type="generic">43 specimens</specimenCount>
we examined, are: [near] Mazamitla, 6800’,
<date id="14191067FF95FF80192DFE64FA63FE51" box="[1333,1432,476,500]" pageId="6" pageNumber="578" value="1950-10-16">16-X-50</date>
; Tecalitlán [the town is at
<quantity id="A75F9B42FF95FF801DDBFE47FDE6FDB2" box="[451,541,511,535]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="6" pageNumber="578" unit="m" value="1140.0">1140 m</quantity>
, but likely the specimens were collected at about
<quantity id="A75F9B42FF95FF801870FE47FB3AFDB2" box="[1128,1217,511,535]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="578" unit="m" value="1300.0">1300 m</quantity>
around the nearby microwave station where there are pines],
<date id="14191067FF95FF801E76FD9CFD18FD99" box="[622,739,548,572]" pageId="6" pageNumber="578" value="1963-07-31">31-VII-63</date>
;
<quantity id="A75F9B42FF95FF801EF6FD9CFCDDFD99" box="[750,806,548,572]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="578" unit="km" value="5.0">5 km</quantity>
N,
<quantity id="A75F9B42FF95FF801F56FD9CFC4BFD99" box="[846,944,548,572]" metricMagnitude="4" metricUnit="m" metricValue="1.0" metricValueMax="1.7" metricValueMin="0.3" pageId="6" pageNumber="578" unit="km" value="10.0" valueMax="17.0" valueMin="3.0">3–17 km</quantity>
E El Tuito,
<quantity id="A75F9B42FF95FF80182EFD9CFB38FD99" box="[1078,1219,548,572]" metricMagnitude="2" metricUnit="m" metricValue="9.925" metricValueMax="11.3" metricValueMin="8.55" pageId="6" pageNumber="578" unit="m" value="992.5" valueMax="1130.0" valueMin="855.0">855–1130 m</quantity>
, on pine logs, 23 &amp;
<date id="14191067FF95FF801C8FFDFFFEF0FDFA" box="[151,267,583,607]" pageId="6" pageNumber="578" value="1990-07-27">27-VII-90</date>
,
<date id="14191067FF95FF801D07FDFFFE89FDFA" box="[287,370,583,607]" pageId="6" pageNumber="578" value="1991-10-02">2-X-91</date>
,
<date id="14191067FF95FF801D9EFDFFFE2CFDFA" box="[390,471,583,607]" pageId="6" pageNumber="578" value="1999-10-04">4-X-99</date>
(FSCA, RLWE, WFBM); Tapalpa, km 2 La Frontera-Tapalpa, BPE, ex
<taxonomicName id="A7A74D24FF95FF801944FDF1FEF7FD21" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="578" phylum="Tracheophyta" rank="species" species="devoniana">
<emphasis id="52D3EAB5FF95FF801944FDF1FEF7FD21" italics="true" pageId="6" pageNumber="578">Pinus devoniana</emphasis>
</taxonomicName>
,
<date id="14191067FF95FF801D02FDD4FE7AFD21" box="[282,385,620,644]" pageId="6" pageNumber="578" value="1994-11-13">13-XI-94</date>
(CZUG); Sierra Tapalpa,
<quantity id="A75F9B42FF95FF801EBEFDD4FD24FD21" box="[678,735,620,644]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="578" unit="km" value="9.0">9 km</quantity>
SE Chiquilistlán,
<geoCoordinate id="05935060FF95FF801FB5FDD4FBEBFD21" box="[941,1040,620,644]" direction="north" orientation="latitude" pageId="6" pageNumber="578" precision="925" value="20.033333">20°02’N</geoCoordinate>
, 103°50W,
<quantity id="A75F9B42FF95FF80188BFDD4FB12FD21" box="[1171,1257,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="578" unit="m" value="1800.0">1800 m</quantity>
,
<date id="14191067FF95FF8018EDFDD4FAACFD21" box="[1269,1367,620,644]" pageId="6" pageNumber="578" value="1996-10-21">21-X-96</date>
, pineoak forest, on standing dead trunk or log of
<taxonomicName id="A7A74D24FF95FF801E91FD29FCBAFD0D" box="[649,833,656,680]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="578" phylum="Tracheophyta" rank="species" species="lumholtzii">
<emphasis id="52D3EAB5FF95FF801E91FD29FCBAFD0D" box="[649,833,656,680]" italics="true" pageId="6" pageNumber="578">Pinus lumholtzii</emphasis>
</taxonomicName>
(RLWE). Apparently, other than as mentioned above and listed in catalogs and checklists, there has been nothing published about this species since Dugès (1891), who recorded a specimen from Moroleon, Guanajuato. Forty-six specimens from the following localities represent new state records:
<emphasis id="52D3EAB5FF95FF801D2AFD43FE38FCB1" bold="true" box="[306,451,763,788]" pageId="6" pageNumber="578">DURANGO</emphasis>
,
<quantity id="A75F9B42FF95FF801DC8FD44FDEDFCB1" box="[464,534,764,788]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="578" unit="km" value="30.0">30 km</quantity>
W Durango,
<date id="14191067FF95FF801EB5FD44FCF4FCB1" box="[685,783,764,788]" pageId="6" pageNumber="578" value="1982-10-13">13-X-82</date>
,
<taxonomicName id="A7A74D24FF95FF801F02FD45FC1BFCB1" box="[794,992,764,788]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="578" phylum="Tracheophyta" rank="species" species="cembroides">
<emphasis id="52D3EAB5FF95FF801F02FD45FC1BFCB1" box="[794,992,764,788]" italics="true" pageId="6" pageNumber="578">Pinus cembroides</emphasis>
</taxonomicName>
(UANL).
<emphasis id="52D3EAB5FF95FF80184DFD43FAF0FCB1" bold="true" box="[1109,1291,763,788]" pageId="6" pageNumber="578">MICHOACAN</emphasis>
,
<quantity id="A75F9B42FF95FF801900FD44FAA4FCB1" box="[1304,1375,764,788]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="578" unit="km" value="16.0">16 km</quantity>
ENE Uruapan,
<quantity id="A75F9B42FF95FF801D1EFCA7FEA5FC9D" box="[262,350,799,824]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="578" unit="m" value="1900.0">1900 m</quantity>
,
<date id="14191067FF95FF801D73FCA7FE2EFC92" box="[363,469,799,823]" pageId="6" pageNumber="578" value="1976-11-12">12-XI-76</date>
, on pine log (RLWE); and Km 98 Carr. Ario de Rosales – La Huacana, 19°04.401’, - 101°46.689’,
<quantity id="A75F9B42FF95FF801D21FCFCFE62FCF9" box="[313,409,836,860]" metricMagnitude="3" metricUnit="m" metricValue="1.285" pageId="6" pageNumber="578" unit="m" value="1285.0">1285 m</quantity>
,
<date id="14191067FF95FF801DB5FCFCFDBCFCF9" box="[429,583,836,860]" pageId="6" pageNumber="578" value="2002-08-25">25-VIII-2002</date>
, on fallen trunk of
<taxonomicName id="A7A74D24FF95FF801F50FCFDFC73FCF9" box="[840,904,837,860]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="578" phylum="Tracheophyta" rank="genus">
<emphasis id="52D3EAB5FF95FF801F50FCFDFC73FCF9" box="[840,904,837,860]" italics="true" pageId="6" pageNumber="578">Pinus</emphasis>
</taxonomicName>
sp. (CEAM, CLBC, RLWE).
<emphasis id="52D3EAB5FF95FF80191EFCFBFA6EFCF9" bold="true" box="[1286,1429,835,860]" pageId="6" pageNumber="578">MORELOS</emphasis>
, Cuernavaca,
<date id="14191067FF95FF801D32FCDFFE76FCDA" box="[298,397,871,895]" pageId="6" pageNumber="578" value="1981-07-02">2-VII-81</date>
(RLWE) &amp;
<date id="14191067FF95FF801E03FCDFFD7AFCDA" box="[539,641,871,895]" pageId="6" pageNumber="578" value="1981-04-30">30-IV-81</date>
(UAEM); Cuernavaca, Chamilpa,
<quantity id="A75F9B42FF95FF801814FCDFFB9FFC25" box="[1036,1124,871,896]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="578" unit="m" value="1900.0">1900 m</quantity>
, on
<taxonomicName id="A7A74D24FF95FF80188CFCD1FA93FC25" box="[1172,1384,873,896]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="578" phylum="Tracheophyta" rank="species" species="montezumae">
<emphasis id="52D3EAB5FF95FF80188CFCD1FA93FC25" box="[1172,1384,873,896]" italics="true" pageId="6" pageNumber="578">Pinus montezumae</emphasis>
</taxonomicName>
,
<date id="14191067FF95FF80196EFCDFFF21FC01" pageId="6" pageNumber="578" value="1981-09-23">23- IX-81</date>
(UAEM).
<emphasis id="52D3EAB5FF95FF801D4CFC33FE2BFC01" bold="true" box="[340,464,907,932]" pageId="6" pageNumber="578">NAYARIT</emphasis>
,
<quantity id="A75F9B42FF95FF801DC6FC34FDDBFC01" box="[478,544,908,932]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="6" pageNumber="578" unit="mi" value="20.0">20 mi</quantity>
E Tepic,
<date id="14191067FF95FF801E94FC34FD1EFC01" box="[652,741,908,932]" pageId="6" pageNumber="578" value="1965-09-05">5-IX-65</date>
(WFBM).
<emphasis id="52D3EAB5FF95FF801F7CFC33FC1FFC01" bold="true" box="[868,996,907,932]" pageId="6" pageNumber="578">SINALOA</emphasis>
, El Palmetto,
<quantity id="A75F9B42FF95FF80189CFC34FB20FC01" box="[1156,1243,908,932]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="578" unit="m" value="2000.0">2000 m</quantity>
,
<date id="14191067FF95FF8018FFFC34FABFFC01" box="[1255,1348,908,932]" pageId="6" pageNumber="578" value="1992-09-06">6-IX-92</date>
, and
<quantity id="A75F9B42FF95FF801998FC34FF4EFC62" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="6" pageNumber="578" unit="mi" value="20.0">20 mi</quantity>
W El Palmetto,
<date id="14191067FF95FF801D9CFC17FE0EFC62" box="[388,501,943,967]" pageId="6" pageNumber="578" value="1984-07-20">20-VII-84</date>
(CLBC);
<quantity id="A75F9B42FF95FF801E6AFC17FD4CFC62" box="[626,695,943,967]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="6" pageNumber="578" unit="mi" value="30.0">30 mi</quantity>
[N]E Villa Union,
<date id="14191067FF95FF801FBCFC17FBD9FC62" box="[932,1058,943,967]" pageId="6" pageNumber="578" value="1965-08-30">30-VIII-65</date>
, pine logs (FSCA); Hwy 40 @ Aserradero El Batel cutoff, 1900 M,
<date id="14191067FF95FF801E21FC6CFD6CFC49" box="[569,663,980,1004]" pageId="6" pageNumber="578" value="1992-09-04">4-IX-92</date>
; La Capilla del Taxte, 3800’,
<date id="14191067FF95FF801FF2FC6CFBA9FC49" box="[1002,1106,980,1004]" pageId="6" pageNumber="578" value="1991-06-22">22-VI-91</date>
(RLWE); Hwy. 40, El Batel,
<quantity id="A75F9B42FF95FF801C8FFC4FFF14FBB5" box="[151,239,1015,1040]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="578" unit="m" value="1700.0">1700 m</quantity>
, on pine slash,
<date id="14191067FF95FF801DB8FC4FFDF6FBAA" box="[416,525,1015,1039]" pageId="6" pageNumber="578" value="1990-08-06">6-VIII-90</date>
(WFBM), and Hwy. 40 @ El Batel cutoff,
<quantity id="A75F9B42FF95FF801FE5FC4FFBAEFBB5" box="[1021,1109,1015,1040]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="578" unit="m" value="1900.0">1900 m</quantity>
,
<date id="14191067FF95FF80187AFC4FFB2DFBAA" box="[1122,1238,1015,1039]" pageId="6" pageNumber="578" value="1992-09-04" valueMax="1992-09-07" valueMin="1992-09-04">4-7-IX-92</date>
, and
<quantity id="A75F9B42FF95FF80190CFC4FFAB5FBAA" box="[1300,1358,1015,1039]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="578" unit="km" value="2.0">2 km</quantity>
NE El Batel,
<quantity id="A75F9B42FF95FF801CF9FBA4FEC1FB91" box="[225,314,1052,1076]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="6" pageNumber="578" unit="m" value="1970.0">1970 m</quantity>
,
<date id="14191067FF95FF801D5FFBA4FE5AFB91" box="[327,417,1052,1076]" pageId="6" pageNumber="578" value="1992-09-04">4-IX-92</date>
(CLBC); Hwy 40, Loberas,
<date id="14191067FF95FF801EF2FBA4FCBFFB91" box="[746,836,1052,1076]" pageId="6" pageNumber="578" value="1992-09-04">4-IX-92</date>
(WFBM).
<emphasis id="52D3EAB5FF95FF801FDCFBA3FBC7FB91" bold="true" box="[964,1084,1051,1076]" pageId="6" pageNumber="578">SONORA</emphasis>
,
<quantity id="A75F9B42FF95FF801854FBA4FB76FB91" box="[1100,1165,1052,1076]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="6" pageNumber="578" unit="mi" value="20.0">20 mi</quantity>
SE Alamos, 4700’,
<date id="14191067FF95FF80196EFBA4FF21FBF2" pageId="6" pageNumber="578" value="1970-11-28">28- XI-70</date>
(FSCA) [Assuming the elevation is correct, this location would have to be in a more easterly direction and the miles by air]; Yécora (airport),
<geoCoordinate id="05935060FF95FF801E3FFBDCFD60FBD9" box="[551,667,1124,1148]" direction="north" orientation="latitude" pageId="6" pageNumber="578" precision="92" value="28.498333">28°29.9'N</geoCoordinate>
,
<geoCoordinate id="05935060FF95FF801EB0FBDCFCD5FBD9" box="[680,814,1124,1148]" direction="west" orientation="longitude" pageId="6" pageNumber="578" precision="92" value="-108.92667">108°55.6'W</geoCoordinate>
,
<quantity id="A75F9B42FF95FF801F22FBDCFC62FBD9" box="[826,921,1124,1148]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="6" pageNumber="578" unit="m" value="1550.0">1,550 m</quantity>
,
<date id="14191067FF95FF801FBEFBDCFBC8FBD9" box="[934,1075,1124,1148]" pageId="6" pageNumber="578" value="2005-07-18">18-VII-2005</date>
(TCMC). Johnson et al. (2012) mentioned “
<taxonomicName id="A7A74D24FF95FF801D3AFB30FE4DFB05" box="[290,438,1160,1184]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="52D3EAB5FF95FF801D3AFB30FE4DFB05" box="[290,438,1160,1184]" italics="true" pageId="6" pageNumber="578">Chalcophora</emphasis>
</taxonomicName>
sp.” on “standing dead pine” found near Yécora, but did not say if any were collected. According to Carl Olson (pers. comm.) three specimens, identified as
<taxonomicName id="A7A74D24FF95FF801FC8FB15FB9BFB61" box="[976,1120,1197,1220]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF95FF801FC8FB15FB9BFB61" box="[976,1120,1197,1220]" italics="true" pageId="6" pageNumber="578">C. mexicana</emphasis>
</taxonomicName>
by one of us (RLW) from images, were collected: Municipio de Yécora,
<geoCoordinate id="05935060FF95FF801EBBFB68FCE5FB4D" box="[675,798,1232,1256]" direction="north" orientation="latitude" pageId="6" pageNumber="578" precision="5" value="28.3825">28.3825°N</geoCoordinate>
<geoCoordinate id="05935060FF95FF801F3DFB68FC2BFB4D" box="[805,976,1232,1256]" direction="west" orientation="longitude" pageId="6" pageNumber="578" precision="1" value="-108.88556">108.885556°W</geoCoordinate>
,
<quantity id="A75F9B42FF95FF801FC3FB68FBCAFB4D" box="[987,1073,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.655" pageId="6" pageNumber="578" unit="m" value="1655.0">1655 m</quantity>
elev., pine-oak forest,
<date id="14191067FF95FF801929FB68FA6CFB4D" box="[1329,1431,1232,1256]" pageId="6" pageNumber="578" value="1993-07-03">3-VII-93</date>
, on dead standing pine in association with
<taxonomicName id="A7A74D24FF95FF801E6AFB4CFBDCFAA9" authority="Erichson" authorityName="Erichson" box="[626,1063,1268,1292]" class="Insecta" family="Elateridae" genus="Chalcolepidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="approximatus">
<emphasis id="52D3EAB5FF95FF801E6AFB4CFC47FAA9" box="[626,956,1268,1292]" italics="true" pageId="6" pageNumber="578">Chalcolepidius approximatus</emphasis>
Erichson
</taxonomicName>
(UAIC). The known range of
<taxonomicName id="A7A74D24FF95FF80199AFB4DFEF9FA95" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF95FF80199AFB4DFEF9FA95" italics="true" pageId="6" pageNumber="578">C. mexicana</emphasis>
</taxonomicName>
, then, is from the Sierra Madre Occidental in southern Sonora, through the western portion of the Transverse Neovolcanic Belt, south into the Sierra Madre del Sur.
</paragraph>
<paragraph id="601836A7FF95FF811CD1FAD8FCD2FEB9" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1436,151,284]" lastPageId="7" lastPageNumber="579" pageId="6" pageNumber="578">
As mentioned above, Waterhouse (1882, 1889) considered all the specimens he studied to be
<taxonomicName id="A7A74D24FF95FF8018E0FAD8FA60FADD" box="[1272,1435,1376,1400]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="virginiensis">
<emphasis id="52D3EAB5FF95FF8018E0FAD8FA60FADD" box="[1272,1435,1376,1400]" italics="true" pageId="6" pageNumber="578">C. virginiensis</emphasis>
</taxonomicName>
except for one, which is the
<typeStatus id="BF1C8805FF95FF801DC5FA3DFDF6FA38" box="[477,525,1413,1437]" pageId="6" pageNumber="578">type</typeStatus>
of
<taxonomicName id="A7A74D24FF95FF801E2DFA3DFD38FA39" box="[565,707,1413,1436]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF95FF801E2DFA3DFD38FA39" box="[565,707,1413,1436]" italics="true" pageId="6" pageNumber="578">C. mexicana</emphasis>
</taxonomicName>
. However, he did mention a form represented by the specimens from Zapotlán that at first he suggested could be a distinct species, then considered it best to call it a variety of the former. Later, after seeing more specimens and discussing more variation, he ended on the same note, i.e. lumping everything but his
<typeStatus id="BF1C8805FF95FF801D7FFA49FE6CF9AC" box="[359,407,1521,1545]" pageId="6" pageNumber="578">type</typeStatus>
of
<taxonomicName id="A7A74D24FF95FF801DA4FA48FDBCF9AD" box="[444,583,1520,1544]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF95FF801DA4FA48FDBCF9AD" box="[444,583,1520,1544]" italics="true" pageId="6" pageNumber="578">C. mexicana</emphasis>
</taxonomicName>
under
<taxonomicName id="A7A74D24FF95FF801E8CFA48FCCDF9AD" box="[660,822,1520,1544]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="virginiensis">
<emphasis id="52D3EAB5FF95FF801E8CFA48FCCDF9AD" box="[660,822,1520,1544]" italics="true" pageId="6" pageNumber="578">C. virginiensis</emphasis>
</taxonomicName>
. The
<typeStatus id="BF1C8805FF95FF801F6EFA49FC5DF9AC" box="[886,934,1521,1545]" pageId="6" pageNumber="578">type</typeStatus>
(Figs. 5, 6; showing how the same specimen can differ in appearance depending upon lighting) does exhibit very distinct coppery reflections in punctured depressed areas of the surface, and that is not a common variant. However, many specimens do appear somewhat coppery in those areas, but on closer inspection one can see the color comes more from a
<typeStatus id="BF1C8805FF95FF8018A0F9E5FB13F9D0" box="[1208,1256,1629,1653]" pageId="6" pageNumber="578">type</typeStatus>
of flocculence. Sometimes that flocculence is more grayish or whitish (Fig. 14), as can also be seen on some specimens of
<taxonomicName id="A7A74D24FF95FF80199AF938FED8F919" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="hondurasica">
<emphasis id="52D3EAB5FF95FF80199AF938FED8F919" italics="true" pageId="6" pageNumber="578">C. hondurasica</emphasis>
</taxonomicName>
. In summary,
<taxonomicName id="A7A74D24FF95FF801DD4F91DFDA1F919" box="[460,602,1701,1724]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF95FF801DD4F91DFDA1F919" box="[460,602,1701,1724]" italics="true" pageId="6" pageNumber="578">C. mexicana</emphasis>
</taxonomicName>
is a variable species, not only in its general appearance, but also in the shape of the sides of the pronotum—that too was noted by Waterhouse (1882, 1889)—and in females the apex of the last abdominal ventrite ranges from narrowly, truncately rounded to variably emarginate or notched. Based on the aforementioned
<specimenCount id="76A1FD2EFF95FF801D63F8A8FDE3F88D" box="[379,536,1808,1832]" pageId="6" pageNumber="578" type="generic">48 specimens</specimenCount>
, the size ranges from
<quantity id="A75F9B42FF95FF801F00F8A8FC6DF88D" box="[792,918,1808,1832]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.2" metricValueMin="2.3" pageId="6" pageNumber="578" unit="mm" value="27.5" valueMax="32.0" valueMin="23.0">23–32 mm</quantity>
. Although it is usually distinguished from
<taxonomicName id="A7A74D24FF95FF80199AF8A8FED8F8E9" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="hondurasica">
<emphasis id="52D3EAB5FF95FF80199AF8A8FED8F8E9" italics="true" pageId="6" pageNumber="578">C. hondurasica</emphasis>
</taxonomicName>
by general appearance,
<taxonomicName id="A7A74D24FF95FF801E2CF88DFD3AF8E9" box="[564,705,1845,1868]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF95FF801E2CF88DFD3AF8E9" box="[564,705,1845,1868]" italics="true" pageId="6" pageNumber="578">C. mexicana</emphasis>
</taxonomicName>
can be readily separated by the distinctly wider smooth elevated median line of the pronotum. The male genitalia (Fig. 8), which are uniform across the known populations, also are distinct, much longer than in either that species or
<emphasis id="52D3EAB5FF95FF801EF2F8C5FC6CF831" box="[746,919,1917,1940]" italics="true" pageId="6" pageNumber="578">
<taxonomicName id="A7A74D24FF95FF801EF2F8C5FC68F831" box="[746,915,1917,1940]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="virginiensis">C. virginiensis</taxonomicName>
.
</emphasis>
The latter species is unknown in
<collectingCountry id="18B07637FF95FF80193DF8C4FA81F831" box="[1317,1402,1916,1940]" name="Mexico" pageId="6" pageNumber="578">Mexico</collectingCountry>
or Central
<collectingCountry id="18B07637FF95FF801CE9F818FEAEF81D" box="[241,341,1952,1976]" name="United States of America" pageId="6" pageNumber="578">America</collectingCountry>
. It might be expected to occur in the Sierra Madre Oriental, but we are unaware of any specimen of this genus having been collected there or, excluding the two aforementioned specimens of
<taxonomicName id="A7A74D24FF95FF8018FCF87DFA6EF879" box="[1252,1429,1988,2012]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="hondurasica">
<emphasis id="52D3EAB5FF95FF8018FCF87DFA6EF879" box="[1252,1429,1988,2012]" italics="true" pageId="6" pageNumber="578">C. hondurasica</emphasis>
</taxonomicName>
, anywhere along the eastern side of the country. The nearest occurrence of
<taxonomicName id="A7A74D24FF94FF811FCEFF20FB82FF15" box="[982,1145,152,176]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="virginiensis">
<emphasis id="52D3EAB5FF94FF811FCEFF20FB82FF15" box="[982,1145,152,176]" italics="true" pageId="7" pageNumber="579">C. virginiensis</emphasis>
</taxonomicName>
to
<collectingCountry id="18B07637FF94FF811884FF2FFB09FF0A" box="[1180,1266,151,175]" name="Mexico" pageId="7" pageNumber="579">Mexico</collectingCountry>
known to us is Bastrop, Texas (S. G. Wellso, pers. comm.), which is about
<quantity id="A75F9B42FF94FF811F24FF04FC50FF71" box="[828,939,188,212]" metricMagnitude="5" metricUnit="m" metricValue="4.828032" pageId="7" pageNumber="579" unit="mi" value="300.0">300 miles</quantity>
across arid lands to any apparently suitable habitat (pines) in
<collectingCountry id="18B07637FF94FF811D43FF67FE4AFF52" box="[347,433,223,247]" name="Mexico" pageId="7" pageNumber="579">Mexico</collectingCountry>
for the genus. Thus, any earlier reference to
<taxonomicName id="A7A74D24FF94FF811FBCFF58FBBDFF5D" box="[932,1094,224,248]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="virginiensis">
<emphasis id="52D3EAB5FF94FF811FBCFF58FBBDFF5D" box="[932,1094,224,248]" italics="true" pageId="7" pageNumber="579">C. virginiensis</emphasis>
</taxonomicName>
in
<collectingCountry id="18B07637FF94FF811870FF67FB46FF52" box="[1128,1213,223,247]" name="Mexico" pageId="7" pageNumber="579">Mexico</collectingCountry>
or Central
<collectingCountry id="18B07637FF94FF811924FF67FA67FF52" box="[1340,1436,223,247]" name="United States of America" pageId="7" pageNumber="579">America</collectingCountry>
must be regarded as either
<taxonomicName id="A7A74D24FF94FF811DDCFEBDFD8BFEB9" box="[452,624,260,284]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="hondurasica">
<emphasis id="52D3EAB5FF94FF811DDCFEBDFD8BFEB9" box="[452,624,260,284]" italics="true" pageId="7" pageNumber="579">C. hondurasica</emphasis>
</taxonomicName>
or
<taxonomicName id="A7A74D24FF94FF811E8DFEBDFCDAFEB9" box="[661,801,261,284]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF94FF811E8DFEBDFCDAFEB9" box="[661,801,261,284]" italics="true" pageId="7" pageNumber="579">C. mexicana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

201
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<mods:title id="7E877C9C9C9ADF8DDC98FD75609E6A8F">Mexican Buprestidae: two new species of Acmaeodera Eschscholtz and a review of the genus Chalcophora Dejean</mods:title>
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<heading id="3B5081CBFF97FF821C8FFA37FD6CFA0C" bold="true" box="[151,663,1423,1449]" fontSize="11" level="1" pageId="4" pageNumber="576" reason="1">
<emphasis id="52D3EAB5FF97FF821C8FFA37FD6CFA0C" bold="true" box="[151,663,1423,1449]" pageId="4" pageNumber="576">
<taxonomicName id="A7A74D24FF97FF821C8FFA37FD6BFA0C" ID-CoL="TGL4" authority="Casey 1909:79" authorityName="Casey" authorityPageNumber="79" authorityYear="1909" box="[151,656,1423,1449]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="hondurasica">
<emphasis id="52D3EAB5FF97FF821C8FFA37FE28FA0C" bold="true" box="[151,467,1423,1449]" italics="true" pageId="4" pageNumber="576">Chalcophora hondurasica</emphasis>
Casey 1909:79
</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
<paragraph id="601836A7FF97FF821C8FFA0AFEC6FA6F" blockId="4.[151,663,1423,1482]" box="[151,317,1458,1482]" pageId="4" pageNumber="576">(Figures 9, 10)</paragraph>
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This species was described from a single female from
<collectingCountry id="18B07637FF97FF821EE1FA42FC92F9B7" box="[761,873,1530,1554]" name="Honduras" pageId="4" pageNumber="576">Honduras</collectingCountry>
, with no other data. One of us (RLW) studied five specimens of
<taxonomicName id="A7A74D24FF97FF821D21F9A7FE12F993" box="[313,489,1566,1590]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="hondurasica">
<emphasis id="52D3EAB5FF97FF821D21F9A7FE12F993" box="[313,489,1566,1590]" italics="true" pageId="4" pageNumber="576">C. hondurasica</emphasis>
</taxonomicName>
from Guacamaya, San Geronimo, 1800’,
<collectingCountry id="18B07637FF97FF821FC0F9A6FBAEF993" box="[984,1109,1566,1590]" name="Guatemala" pageId="4" pageNumber="576">Guatemala</collectingCountry>
; and Bonaca Island, Bay of
<collectingCountry id="18B07637FF97FF821C8FF9FAFEFCF9FF" box="[151,263,1602,1626]" name="Honduras" pageId="4" pageNumber="576">Honduras</collectingCountry>
,
<collectingCountry id="18B07637FF97FF821D0BF9FAFE7BF9FF" box="[275,384,1602,1626]" name="Honduras" pageId="4" pageNumber="576">Honduras</collectingCountry>
in the
<emphasis id="52D3EAB5FF97FF821DCAF9FAFCE6F9FF" box="[466,797,1602,1626]" italics="true" pageId="4" pageNumber="576">Biologia Centrali-Americana</emphasis>
Collection (BMNH) that had been determined by C. O. Waterhouse as
<taxonomicName id="A7A74D24FF97FF821D59F9DFFE1EF9DB" box="[321,485,1639,1662]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="virginiensis">
<emphasis id="52D3EAB5FF97FF821D59F9DFFE1EF9DB" box="[321,485,1639,1662]" italics="true" pageId="4" pageNumber="576">C. virginiensis</emphasis>
</taxonomicName>
and the data published by him (1882, 1889). A new determination label was placed on each specimen. Waterhouse (1882) also mentioned that there are four specimens from
<collectingCountry id="18B07637FF97FF8218ADF932FAF1F907" box="[1205,1290,1674,1698]" name="Mexico" pageId="4" pageNumber="576">Mexico</collectingCountry>
(no specific locality) in BMNH, but we have not seen them. One could be the specimen of
<taxonomicName id="A7A74D24FF97FF821825F917FAA0F963" authority="Waterhouse" authorityName="Waterhouse" box="[1085,1371,1710,1734]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF97FF821825F917FB36F963" box="[1085,1229,1711,1734]" italics="true" pageId="4" pageNumber="576">C. mexicana</emphasis>
Waterhouse
</taxonomicName>
from Juquila (q.v.). Our concept of
<taxonomicName id="A7A74D24FF97FF821DE8F96BFD64F94F" box="[496,671,1746,1770]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="hondurasica">
<emphasis id="52D3EAB5FF97FF821DE8F96BFD64F94F" box="[496,671,1746,1770]" italics="true" pageId="4" pageNumber="576">C. hondurasica</emphasis>
</taxonomicName>
is largely based on a female compared with the
<typeStatus id="BF1C8805FF97FF8218C8F96AFAC9F94F" box="[1232,1330,1746,1770]" pageId="4" pageNumber="576" type="holotype">holotype</typeStatus>
by G. H. Nelson, labeled from
<collectingCountry id="18B07637FF97FF821D88F94EFDF7F8AB" box="[400,524,1782,1806]" name="Guatemala" pageId="4" pageNumber="576">Guatemala</collectingCountry>
, Alta Verapaz, Mpio.
<collectingCountry id="18B07637FF97FF821F12F94EFC58F8AB" box="[778,931,1782,1806]" name="Saint Kitts and Nevis" pageId="4" pageNumber="576">San Cristobal</collectingCountry>
Verapaz, Baleu, +
<quantity id="A75F9B42FF97FF821860F94EFB2AF8AB" box="[1144,1233,1782,1806]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="4" pageNumber="576" unit="m" value="1350.0">1350 m</quantity>
,
<date id="14191067FF97FF8218F8F94EFAAFF8AB" box="[1248,1364,1782,1806]" pageId="4" pageNumber="576" value="1966-07-16">16-VII-66</date>
, G. C. Walters (FSCA). Two other Guatemalan specimens are from Baja Verapaz, Los Limas,
<quantity id="A75F9B42FF97FF821867F8A2FB2DF897" box="[1151,1238,1818,1842]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="4" pageNumber="576" unit="m" value="1125.0">1125 m</quantity>
,
<date id="14191067FF97FF8218FCF8A3FACAF897" box="[1252,1329,1819,1842]" pageId="4" pageNumber="576" value="1977-07">VII-77</date>
(CLBC). We examined 15 additional specimens from the following localities in
<collectingCountry id="18B07637FF97FF821FF3F886FBA7F8F3" box="[1003,1116,1854,1878]" name="Honduras" pageId="4" pageNumber="576">Honduras</collectingCountry>
: Siguatepeque &amp;
<quantity id="A75F9B42FF97FF82192CF886FA81F8F3" box="[1332,1402,1854,1878]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="4" pageNumber="576" unit="mi" value="10.0">10 mi</quantity>
N Siguatepeque, 10-20-VIII-78,
<date id="14191067FF97FF821DF6F8DAFD92F8DF" box="[494,617,1890,1914]" pageId="4" pageNumber="576" value="1979-08-17">17-VIII-79</date>
,
<date id="14191067FF97FF821E62F8DAFD2FF8DF" box="[634,724,1890,1914]" pageId="4" pageNumber="576" value="1987-06-04">4-VI-87</date>
(FSCA, WFBM); Zamorano, F[rancisco] M[orazán],
<date id="14191067FF97FF821958F8DAFA67F8DF" box="[1344,1436,1890,1914]" pageId="4" pageNumber="576" value="1976-05-22">22-V-76</date>
(FSCA); Siguatepeque,
<date id="14191067FF97FF821DBDF83EFDFFF83B" box="[421,516,1926,1950]" pageId="4" pageNumber="576" value="1975-05-29">29-V-75</date>
, “under loose bark – dead pine 14 caught together”, and
<date id="14191067FF97FF821894F83EFB08F83B" box="[1164,1267,1926,1950]" pageId="4" pageNumber="576" value="1974-06-11">11-VI-74</date>
(RLWE). This species has heretofore been recorded only from
<collectingCountry id="18B07637FF97FF821EDDF812FCC5F867" box="[709,830,1962,1986]" name="Guatemala" pageId="4" pageNumber="576">Guatemala</collectingCountry>
and
<collectingCountry id="18B07637FF97FF821F63F812FC13F867" box="[891,1000,1962,1986]" name="Honduras" pageId="4" pageNumber="576">Honduras</collectingCountry>
(see above),
<collectingCountry id="18B07637FF97FF82189EF812FB02F867" box="[1158,1273,1962,1986]" name="Nicaragua" pageId="4" pageNumber="576">Nicaragua</collectingCountry>
(Maes,
<emphasis id="52D3EAB5FF97FF821942F813FA75F867" box="[1370,1422,1962,1986]" italics="true" pageId="4" pageNumber="576">et al</emphasis>
., 1993), and
<collectingCountry id="18B07637FF97FF821D00F876FE9BF843" box="[280,352,1998,2022]" name="Belize" pageId="4" pageNumber="576">Belize</collectingCountry>
(Westcott, 2008; as
<taxonomicName id="A7A74D24FF97FF821E51F877FD16F843" box="[585,749,1999,2022]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="virginiensis">
<emphasis id="52D3EAB5FF97FF821E51F877FD16F843" box="[585,749,1999,2022]" italics="true" pageId="4" pageNumber="576">C. virginiensis</emphasis>
</taxonomicName>
). We have examined several other specimens of this species from
<collectingCountry id="18B07637FF96FF831CCCFF2FFEE5FF0A" box="[212,286,151,175]" name="Belize" pageId="5" pageNumber="577">Belize</collectingCountry>
, some of which were labeled as taken on pine or pine slash. The following specimens represent the first authenticated records for
<collectingCountry id="18B07637FF96FF831DA5FF04FDCBFF71" box="[445,560,188,212]" name="Mexico" pageId="5" pageNumber="577">MEXICO</collectingCountry>
: Chiapas, [Cerro?] La Sepultura, Arriaga,
<quantity id="A75F9B42FF96FF831805FF04FB92FF71" box="[1053,1129,188,212]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="5" pageNumber="577" unit="m" value="850.0">850 m</quantity>
,
<date id="14191067FF96FF831860FF04FB18FF71" box="[1144,1251,188,212]" pageId="5" pageNumber="577" value="1975-06-27">27-VI-75</date>
, “Sierra Madre: Matorral orillas Selva Baja Decidua” (RLWE); San Quintín,
<date id="14191067FF96FF831F5AFF67FC48FF52" box="[834,947,223,247]" pageId="5" pageNumber="577" value="1977-07-27">27-VII-77</date>
(UNAM). The only place we could find by the latter name in Chiapas is at 16.406°, -91.347°,
<quantity id="A75F9B42FF96FF831EFAFEBCFCD6FEB9" box="[738,813,260,284]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="577" unit="m" value="220.0">220 m</quantity>
, and it seems an unlikely locality for pines. However, according to the collector, Peter Hubbell (pers. com.), this is the correct locality. One can check a map and see there are nearby areas of significantly higher elevation where pines should occur. A total of about
<specimenCount id="76A1FD2EFF96FF8318A7FEF4FAA2FEC1" box="[1215,1369,332,356]" pageId="5" pageNumber="577" type="generic">30 specimens</specimenCount>
of
<taxonomicName id="A7A74D24FF96FF83199AFEF5FED8FE2D" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="hondurasica">
<emphasis id="52D3EAB5FF96FF83199AFEF5FED8FE2D" italics="true" pageId="5" pageNumber="577">C. hondurasica</emphasis>
</taxonomicName>
were examined, ranging in size from
<quantity id="A75F9B42FF96FF831ED3FED7FCBCFE22" box="[715,839,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="5" pageNumber="577" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">25–30 mm</quantity>
.
</paragraph>
<paragraph id="601836A7FF96FF831CD1FE2CFAC4FD99" blockId="5.[151,1437,151,572]" pageId="5" pageNumber="577">
Obviously
<taxonomicName id="A7A74D24FF96FF831D5EFE2DFE0FFE09" box="[326,500,404,428]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="hondurasica">
<emphasis id="52D3EAB5FF96FF831D5EFE2DFE0FFE09" box="[326,500,404,428]" italics="true" pageId="5" pageNumber="577">C. hondurasica</emphasis>
</taxonomicName>
very closely resembles
<taxonomicName id="A7A74D24FF96FF831F10FE2DFC57FE09" box="[776,940,405,428]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="virginiensis">
<emphasis id="52D3EAB5FF96FF831F10FE2DFC57FE09" box="[776,940,405,428]" italics="true" pageId="5" pageNumber="577">C. virginiensis</emphasis>
</taxonomicName>
(Fig. 11), as it has been confused with that species. The former differs by having the front of the head more widely, somewhat more deeply depressed and the surface more punctate; the median smooth line of the pronotum is narrower, and the pronotal punctation on either side is less dense. See Fig. 10 for the male genitalia of
<taxonomicName id="A7A74D24FF96FF831EE4FDB8FC53FDBD" box="[764,936,512,536]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="hondurasica">
<emphasis id="52D3EAB5FF96FF831EE4FDB8FC53FDBD" box="[764,936,512,536]" italics="true" pageId="5" pageNumber="577">C. hondurasica</emphasis>
</taxonomicName>
; the male parameres of
<taxonomicName id="A7A74D24FF96FF8318ADFDB8FAA3FDBD" box="[1205,1368,512,536]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="virginiensis">
<emphasis id="52D3EAB5FF96FF8318ADFDB8FAA3FDBD" box="[1205,1368,512,536]" italics="true" pageId="5" pageNumber="577">C. virginiensis</emphasis>
</taxonomicName>
differ by being slightly swollen apically (Fig. 12). See discussion of the latter species in the following treatment.
</paragraph>
</subSubSection>
<subSubSection id="28BD652CFF96FF831C8FF907FAC6F956" pageId="5" pageNumber="577" type="description">
<paragraph id="601836A7FF96FF831C8FF907FAC6F956" blockId="5.[151,1437,1727,1780]" pageId="5" pageNumber="577">
<emphasis id="52D3EAB5FF96FF831C8FF907FEFEF971" bold="true" box="[151,261,1727,1748]" pageId="5" pageNumber="577">PLATE 3.</emphasis>
Live specimens
<emphasis id="52D3EAB5FF96FF831DA0F978FE02F970" box="[440,505,1728,1749]" italics="true" pageId="5" pageNumber="577">in situ</emphasis>
: Figure 13,
<taxonomicName id="A7A74D24FF96FF831E65F907FCA5F970" box="[637,862,1727,1749]" class="Insecta" family="Buprestidae" genus="Acmaeodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="inusitata" status="sp. nov.">
<emphasis id="52D3EAB5FF96FF831E65F907FCA5F970" box="[637,862,1727,1749]" italics="true" pageId="5" pageNumber="577">Acmaeodera inusitata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E057CEFF96FF831F70F907FC58F971" box="[872,931,1727,1748]" pageId="5" pageNumber="577" rank="species">
<emphasis id="52D3EAB5FF96FF831F70F907FC58F971" bold="true" box="[872,931,1727,1748]" pageId="5" pageNumber="577">n. sp.</emphasis>
</taxonomicNameLabel>
on
<taxonomicName id="A7A74D24FF96FF831FD6F978FBEFF970" box="[974,1044,1728,1749]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="577" phylum="Tracheophyta" rank="genus">
<emphasis id="52D3EAB5FF96FF831FD6F978FBEFF970" box="[974,1044,1728,1749]" italics="true" pageId="5" pageNumber="577">Acacia</emphasis>
</taxonomicName>
sp. at
<typeStatus id="BF1C8805FF96FF831847F978FB70F970" box="[1119,1163,1728,1749]" pageId="5" pageNumber="577">type</typeStatus>
locality,
<typeStatus id="BF1C8805FF96FF8318F5F978FABCF970" box="[1261,1351,1728,1749]" pageId="5" pageNumber="577" type="paratype">paratype</typeStatus>
, CLBC. Figure 14,
<taxonomicName id="A7A74D24FF96FF831D1DF966FD94F956" authority="Waterhouse" authorityName="Waterhouse" box="[261,623,1757,1780]" class="Insecta" family="Buprestidae" genus="Chalcophora" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="52D3EAB5FF96FF831D1DF966FE09F951" box="[261,498,1758,1780]" italics="true" pageId="5" pageNumber="577">Chalcophora mexicana</emphasis>
Waterhouse
</taxonomicName>
on trunk of fallen pine, Michoacán, 19°04.401’, -101°46.689’, CLBC.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.419.7481" ID-GBIF-Dataset="10a6ae8a-2677-4356-aea0-008675da7c32" ID-PMC="PMC4109454" ID-Pensoft-Pub="1313-2970-419-103" ID-PubMed="25061363" ID-ZBK="B6F178208C8447F09D3F96627F11B8F0" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-419-103" ModsDocOrigin="ZooKeys 419" ModsDocTitle="Two new species of the genus Abrus Dai &amp; Zhang, 2002 (Hemiptera, Cicadellidae, Deltocephalinae) from China" checkinTime="1451245705013" checkinUser="pensoft" docAuthor="Xing, Jichun &amp; Li, Zizhong" docDate="2014" docId="E80EFDD9D791584909E3EEA04C72855B" docLanguage="en" docName="ZooKeys 419: 103-109" docOrigin="ZooKeys 419" docSource="http://dx.doi.org/10.3897/zookeys.419.7481" docTitle="Abrus expansivus Xing &amp; Li, sp. n." docType="treatment" docUuid="9C25F88C-D706-481F-B6B1-089C0EE2D4D8" docUuidSource="ZooBank" docVersion="4" lastPageNumber="106" masterDocId="A174FFE5634FFFBEFFAB2702FFB7FFAD" masterDocTitle="Two new species of the genus Abrus Dai &amp; Zhang, 2002 (Hemiptera, Cicadellidae, Deltocephalinae) from China" masterLastPageNumber="109" masterPageNumber="103" pageNumber="105" updateTime="1668158716817" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species of the genus Abrus Dai &amp; Zhang, 2002 (Hemiptera, Cicadellidae, Deltocephalinae) from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xing, Jichun</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Zizhong</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>419</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>103</mods:start>
<mods:end>109</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.419.7481</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.419.7481</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-419-103</mods:identifier>
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<mods:identifier type="ZooBank">B6F178208C8447F09D3F96627F11B8F0</mods:identifier>
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<treatment ID-GBIF-Taxon="152053748" LSID="urn:lsid:zoobank.org:act:9C25F88C-D706-481F-B6B1-089C0EE2D4D8" httpUri="http://treatment.plazi.org/id/E80EFDD9D791584909E3EEA04C72855B" lastPageId="3" lastPageNumber="106" pageId="2" pageNumber="105">
<subSubSection pageId="2" pageNumber="105" type="multiple">
<paragraph pageId="2" pageNumber="105">Taxon classification Animalia Hemiptera Cicadellidae</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="105" type="nomenclature">
<paragraph pageId="2" pageNumber="105">
<taxonomicName LSID="http://zoobank.org/9C25F88C-D706-481F-B6B1-089C0EE2D4D8" authority="Xing &amp; Li" class="Insecta" family="Cicadellidae" genus="Abrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abrus expansivus" order="Hemiptera" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="expansivus">Abrus expansivus Xing &amp; Li</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="105">sp. n.</taxonomicNameLabel>
Figs 12-22
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="106" pageId="2" pageNumber="105" type="description">
<paragraph pageId="2" pageNumber="105">Description.</paragraph>
<paragraph pageId="2" pageNumber="105">
External features resemble as
<taxonomicName class="Insecta" family="Cicadellidae" genus="Abrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abrus damingshanensis" order="Hemiptera" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="damingshanensis">Abrus damingshanensis</taxonomicName>
sp. n., but color light yellow and body slightly small.
</paragraph>
<paragraph pageId="3" pageNumber="106">
<pageBreakToken pageId="3" pageNumber="106" start="start">Male</pageBreakToken>
genitalia. Male pygofer side elongate with many macrosetae posteriorly and a few at midventral margin; posterior margin lobe elongate and with a long membranous process at inner apex (Fig. 16). Valve subtriangular with anterior margin concaved and posterior margin strongly produced medially (Fig. 17). Subgenital plate broad and short, outer margin rounded, with many macrosetae on lateral margin (Fig. 18). Aedeagus with broad and flat basal projection from dorsal margin, deeply concave at dorsal margin, the basal projection without processes; aedeagal shaft slightly shorter than basal projection, with a pair of slender apical processes and its length equal to aedeagal shaft, gonopore apical (Figs 19, 20). Connective
<normalizedToken originalValue="Y–shaped">Y-shaped</normalizedToken>
, stem robust, arms well developed, articulated with the aedeagus (Fig. 21). Style long, broad at base, narrow at middle, apex slightly widening (Fig. 22).
</paragraph>
<caption pageId="3" pageNumber="106">
<paragraph pageId="3" pageNumber="106">
Figures 12-22.
<taxonomicName class="Insecta" family="Cicadellidae" genus="Abrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abrus expansivus" order="Hemiptera" pageId="3" pageNumber="106" phylum="Arthropoda" rank="species" species="expansivus">Abrus expansivus</taxonomicName>
sp. n., 12 ♂, dorsal view 13 ♂, lateral view 14 ♂, head and thorax, dorsal view 15 ♂, face 16 Male pygofer side, lateral view 17 Valve, ventral view 18 Subgenital plate, ventral view 19 Aedeagus, lateral view 20 Aedeagus, caudal view 21 Connective, ventral view 22 Style, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="106" type="measurement">
<paragraph pageId="3" pageNumber="106">Measurement.</paragraph>
<paragraph pageId="3" pageNumber="106">Length (including tegmen): ♂, 8.1 mm.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="106" type="type material">
<paragraph pageId="3" pageNumber="106">Type material.</paragraph>
<paragraph pageId="3" pageNumber="106">Holotype ♂, China: Guizhou Province, Dushan County, 16 July 2012, coll. Qiongzhang Song (GUGC).</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="106" type="diagnosis">
<paragraph pageId="3" pageNumber="106">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="106">
This new species is very similar to
<taxonomicName class="Insecta" family="Cicadellidae" genus="Abrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abrus brevis" order="Hemiptera" pageId="3" pageNumber="106" phylum="Arthropoda" rank="species" species="brevis">Abrus brevis</taxonomicName>
Dai &amp; Zhang, 2002 in aedeagal shaft with a pair of long apical appendages, but can be distinguished from the latter by the aedeagal shaft longer than half length of basal projection; aedeagal shaft with apical processes located medially in lateral view and its length equal to aedeagal shaft; the apical process of style wide and flat; and the subgenital plate narrow apically.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="106" type="etymology">
<paragraph pageId="3" pageNumber="106">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="106">
The new species name is derived from the Latin word
<normalizedToken originalValue="“expansivus”">&quot;expansivus&quot;</normalizedToken>
, referring to the apical process of style wide and expand.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="D9357400B64DA5EC035EBDB0609669C2">A New Goblin Spider Genus of the Zyngoonops Group from Costa Rica, With Notes on Coxapopha (Araneae, Oonopidae)</mods:title>
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<figureCitation id="F89DAFC62300FFD1E959E45CAB6AFCF3" box="[620,772,833,859]" captionStart-0="FIGURES 1–15" captionStart-1="FIGURES 16–30" captionStart-2="FIGURES 31–45" captionStart-3="FIGURES 46–60" captionStartId-0="7.[144,242,1647,1668]" captionStartId-1="8.[144,242,1618,1639]" captionStartId-2="9.[144,242,1649,1670]" captionStartId-3="10.[144,242,1676,1697]" captionTargetBox-0="[144,1248,248,1622]" captionTargetBox-1="[161,1237,248,1592]" captionTargetBox-2="[144,1248,248,1622]" captionTargetBox-3="[145,1239,248,1650]" captionTargetId-0="figure-20@7.[144,880,798,1074]" captionTargetId-1="figure-23@8.[519,877,785,1592]" captionTargetId-2="figure-106@9.[513,881,797,1073]" captionTargetId-3="figure-88@10.[404,975,794,1156]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURES 1–15. Hexapopha reimoseri (Fage), male. 1. Habitus, dorsal view. 2. Same, lateral view. 3. Carapace, dorsal view. 4. Same, lateral view. 5. Same, anterior view. 6. Sternum, ventral view. 7. Chelicerae, anterior view. 8. Same, posterior view. 9. Mouthparts, ventral view. 10. Endites, dorsal view. 11. Same, lateral view. 12. Same, anterior view. 13. Tip of endites, dorsal view. 14. Same, ventral view. 15. Labium, ventral view." captionText-1="FIGURES 16–30. Hexapopha reimoseri (Fage), male. 16. Left palp, prolateral view. 17. Same, ventral view. 18. Same, retrolateral view. 19. Same, dorsal view. 20. Palpal tibia, dorsal view. 21. Left embolus, prolateral view. 22. Same, ventral view. 23. Same, retrolateral view. 24. Same, dorsal view. 25. Epigastric region, ventral view. 26. Spinnerets, ventral view. 27. Same, distal view. 28. Anterior lateral spinneret, same. 29. Posterior median spinneret, same. 30. Posterior lateral spinneret, same." captionText-2="FIGURES 31–45. Hexapopha reimoseri (Fage), male. 31. Claws of leg I, distal view. 32. Same, leg II. 33. Same, leg III. 34. Same, leg IV. 35. Claws of leg I, lateral view. 36. Same, leg II. 37. Same, leg III. 38. Same, leg IV. 39. Tarsal organ of leg I, dorsal view. 40. Same, leg II. 41. Same, leg III. 42. Same, leg IV. 43. Same, palp. 44. Trichobothrial base, metatarsus II, dorsal view. 45. Labrum, dorsal view." captionText-3="FIGURES 46–60. Hexapopha reimoseri (Fage), male (46–53) and female (54–60). 46, 54. Habitus, dorsal view. 47. Carapace, dorsal view. 48, 55. Sternum, ventral view. 49. Endites, ventral view. 50, 57. Habitus, lateral view. 51. Left palp, prolateral view. 52. Same, ventral view. 53. Same, retrolateral view. 56. Abdomen, ventral view. 58, 59. Genitalia, ventral view. 60. Same, dorsal view." pageId="6">Figures 1–60</figureCitation>
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<emphasis id="52D26F512300FFD1EBA5E474A939FC28" box="[144,343,873,896]" italics="true" pageId="6">Xestaspis reimoseri</emphasis>
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<bibRefCitation id="0437CEB22300FFD1EA6BE474AA6BFC29" author="Fage, L." box="[350,517,873,897]" pageId="6" pagination="369 - 376" refId="ref7242" refString="Fage, L. 1938. Quelques arachnides provenant de fourmilieres ou de termitieres du Costa Rica. Bulletin du Museum National d'Histoire Naturelle Paris (2) 10: 369 - 376." type="journal article" year="1938">Fage, 1938: 371</bibRefCitation>
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, figs. 1A–D (male
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from
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,
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,
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, should be in the
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, lost).
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<paragraph id="6019B3432300FFD1EBF5E4DCABD2FB8E" blockId="6.[144,1248,961,1734]" pageId="6">DIAGNOSIS: Males can be recognized by the small squared projection on the squared tip of the endites (fig. 9), females by the relatively long and narrow posterior genitalic plate (fig. 59) and the large, straight arms of the anterior genitalic process (fig. 60).</paragraph>
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<subSubSection id="28BCE0C82300FFD1EBF5E32CA910FAF9" pageId="6" type="description">
<paragraph id="6019B3432300FFD1EBF5E32CAB0CFB3D" blockId="6.[144,1248,961,1734]" pageId="6">MALE (PBI_OON 49619, figs. 1–53): Total length 1.54. Labium with squared, narrow tip situated on slightly wider base; endites with squared tip followed posteriorly by obliquely set elevated projections. Conductor with gradually narrowed tip.</paragraph>
<paragraph id="6019B3432300FFD1EBF5E3BCA910FAF9" blockId="6.[144,1248,961,1734]" pageId="6">FEMALE (PBI_OON 10125, figs. 54–60): Total length 1.83. ALE separated by their radius to diameter. Dorsal scutum covering more than 3/4 of abdomen length, more than 1/2 to most of abdomen width. Postepigastric scutum covering about 3/4 of abdomen length. Anterior genitalic process with long, straight arms; posterior genitalic plate relatively long, narrow.</paragraph>
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<collectionCode id="06B72B862300FFD1EBF5E241A9A0FADE" box="[192,462,1372,1398]" pageId="6">MATERIAL EXAMINED</collectionCode>
:
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:
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:
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<collectingCounty id="8978CBCF2300FFD1E824E241AC08FADE" box="[785,1126,1372,1398]" pageId="6">Estación Biológica La Selva</collectingCounty>
,
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,
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,
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<collectingDate id="045C6C6B2300FFD1EA31E29CA9F5FA33" box="[260,411,1409,1436]" pageId="6" value="1973-03-05">Mar. 5, 1973</collectingDate>
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,
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, tree hole above floor in coarse wood chips with large ants (
<collectorName id="CD53D6952300FFD1EFFAE29CA882FA69" pageId="6">J. Wagner</collectorName>
,
<collectorName id="CD53D6952300FFD1EBCDE2BAA904FA68" box="[248,362,1446,1473]" pageId="6">J. Kethley</collectorName>
,
<specimenCode id="30001B382300FFD1EA43E2BAAA73FA69" box="[374,541,1446,1473]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="6" type="Museum">FMNH 33527</specimenCode>
,
<collectionCode id="06B72B862300FFD1E91FE2BBAA39FA68" box="[554,599,1446,1472]" pageId="6">PBI</collectionCode>
_OON 10034),
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,
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<collectingDate id="045C6C6B2300FFD1E877E2BAABB6FA68" box="[834,984,1446,1473]" pageId="6" value="1973-03-07">Mar. 7, 1973</collectingDate>
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,
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, concentrated epiphytic root fibers (
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,
<collectorName id="CD53D6952300FFD1E924E2D1AAECFA4E" box="[529,642,1484,1510]" pageId="6">J. Kethley</collectorName>
,
<collectionCode id="06B72B862300FFD1E9BAE2D1AA8BFA4E" box="[655,741,1484,1510]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="6" type="Museum">FMNH</collectionCode>
<collectionCode id="06B72B862300FFD1E9D8E2D1AB74FA4E" box="[749,794,1484,1510]" pageId="6">PBI</collectionCode>
_OON 10542),
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;
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,
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,
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,
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, fogging
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<emphasis id="52D26F512300FFD1E9CAE2ECABDAF9A3" box="[767,948,1521,1547]" italics="true" pageId="6">Virola koschnyi</emphasis>
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, elev.
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, Camino Experimental Sur,
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marker,
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project,
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_ OON 36994),
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<collectingMunicipality id="807D29392300FFD1EAF8E17CAAE9F9D4" box="[461,647,1633,1660]" pageId="6">Hamburg Farm</collectingMunicipality>
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,
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<collectingDate id="045C6C6B2300FFD1E81FE17CABAAF9D3" box="[810,964,1633,1659]" pageId="6" value="1937-02-19">Feb. 19, 1937</collectingDate>
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(
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<collectionCode id="06B72B862300FFD1EBD0E19BA97CF908" box="[229,274,1670,1696]" pageId="6">PBI</collectionCode>
_OON 10125),
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.
</paragraph>
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<materialsCitation id="D0CEB91E2300FFD1EBF5E1B1ACFCF96E" box="[192,1170,1708,1734]" country="Costa Rica" location="Heredia" pageId="6" specimenCount="1" stateProvince="Heredia">
DISTRIBUTION: Known only from two localities in
<collectingRegion id="A2627DA12300FFD1E82CE1B1AB19F96E" box="[793,887,1708,1734]" country="Costa Rica" name="Heredia" pageId="6">Heredia</collectingRegion>
and
<collectingRegion id="A2627DA12300FFD1E887E1B1AC6DF96E" box="[946,1027,1708,1734]" country="Costa Rica" name="Limon" pageId="6">Limón</collectingRegion>
,
<collectingCountry id="18B1F3D32300FFD1EF3AE1B1ACFCF96E" box="[1039,1170,1708,1734]" name="Costa Rica" pageId="6">Costa Rica</collectingCountry>
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.
</paragraph>
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110
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<mods:title id="AFA2BF70470D5DFBB943917634D85E2E">A New Goblin Spider Genus of the Zyngoonops Group from Costa Rica, With Notes on Coxapopha (Araneae, Oonopidae)</mods:title>
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<emphasis id="52D26F512305FFD4E918E269AAD6FA27" bold="true" box="[557,696,1396,1423]" italics="true" pageId="3">Hexapopha</emphasis>
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,
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<taxonomicName id="A7A6C8C02305FFD4EA5BE2B0AAB0FA6E" authority="Fage (1938)" authorityName="Fage" authorityYear="1938" box="[366,734,1452,1479]" class="Arachnida" family="Oonopidae" genus="Xestaspis" kingdom="Animalia" order="Araneae" pageId="3" phylum="Arthropoda" rank="species" species="reimoseri">
<emphasis id="52D26F512305FFD4EA5BE2B0AA27FA6E" box="[366,585,1453,1478]" italics="true" pageId="3">Xestaspis reimoseri</emphasis>
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.
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ETYMOLOGY: The generic name refers to the presence of six eyes and the similarities to
<taxonomicName id="A7A6C8C02305FFD4EBA5E2EAA97DF9B9" authorityName="Platnick" authorityYear="2000" box="[144,275,1527,1553]" class="Arachnida" family="Oonopidae" genus="Coxapopha" kingdom="Animalia" order="Araneae" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="52D26F512305FFD4EBA5E2EAA97DF9B9" box="[144,275,1527,1553]" italics="true" pageId="3">Coxapopha</emphasis>
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, and is feminine in gender.
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DIAGNOSIS: These spiders are easily recognized as members of the
<taxonomicName id="A7A6C8C02305FFD4E8D0E100AC1EF99E" authorityName="Benoit" authorityYear="1977" box="[997,1136,1565,1590]" class="Arachnida" family="Oonopidae" genus="Zyngoonops" kingdom="Animalia" order="Araneae" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="52D26F512305FFD4E8D0E100AC1EF99E" box="[997,1136,1565,1590]" italics="true" pageId="3">Zyngoonops</emphasis>
</taxonomicName>
group of genera by the heavily sclerotized endites of males (figs. 49, 79, 114). The combined presence of six eyes, a palpal conductor in males, and a female epigastric region without an external orifice or scape distinguishes these species from those of the other described genera of the group. Additional differences from
<taxonomicName id="A7A6C8C02305FFD4E90FE1AFAAABF963" authorityName="Benoit" authorityYear="1977" box="[570,709,1714,1739]" class="Arachnida" family="Oonopidae" genus="Zyngoonops" kingdom="Animalia" order="Araneae" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="52D26F512305FFD4E90FE1AFAAABF963" box="[570,709,1714,1739]" italics="true" pageId="3">Zyngoonops</emphasis>
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,
<taxonomicName id="A7A6C8C02305FFD4E9E1E1AFAB3FF964" authorityName="Fannes and Jocque" authorityYear="2008" box="[724,849,1714,1740]" class="Arachnida" family="Oonopidae" genus="Antoonops" kingdom="Animalia" order="Araneae" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="52D26F512305FFD4E9E1E1AFAB3FF964" box="[724,849,1714,1740]" italics="true" pageId="3">Antoonops</emphasis>
</taxonomicName>
, and
<taxonomicName id="A7A6C8C02305FFD4E8A1E1AFAC77F964" authorityName="Platnick" authorityYear="2000" box="[916,1049,1714,1740]" class="Arachnida" family="Oonopidae" genus="Coxapopha" kingdom="Animalia" order="Araneae" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="52D26F512305FFD4E8A1E1AFAC77F964" box="[916,1049,1714,1740]" italics="true" pageId="3">Coxapopha</emphasis>
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are discussed in the Introduction.
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115
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<paragraph id="A85E28821F9D7B50A2F7C29E7496C955" pageNumber="351">
<treatmentCitation id="D8AA97A3B4B3C6A7280533E9CEB9B91C" author="Linnaeus" pageNumber="831" year="1753">
<bibCitation id="DB49FED70B3A5B7EA777B272EFD0914E" author="Linnaeus" pagination="831" title="Bidens tripartita" volumeTitle="Species Plantarum 2" year="1753">
<taxonomicName id="EF467E59A55523EF6D3873531F26C6B8" ID-CoL="LSLN" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Asteraceae" genus="Bidens" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="tripartita">
<emphasis id="06DB11A5D522FFFC235E1DB088C7168E" bold="true" pageNumber="351">Bidens tripartita</emphasis>
Linnaeus
</taxonomicName>
,
<volumeTitle id="564ABDE916FC774F59EEDD249E5F6E29">
<emphasis id="6833697B3790D7292E22393C93E0EE66" italics="true" pageNumber="351">Species Plantarum</emphasis>
2
</volumeTitle>
: 831. 1753
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C95BA47DB4C5E2EB52B75FE6D9C565" pageNumber="351" type="distribution">
<paragraph id="A7ADACC45EED84A73A4D4129D3515B99" pageNumber="351">&quot;Habitat in Europae inundatis.&quot; RCN: 6017.</paragraph>
</subSubSection>
<subSubSection id="C561D66E792519496AB3691867AFCFAF" pageNumber="351" type="materials_examined">
<paragraph id="7CA169206714FDE52D87E8A14B98AFD6" pageNumber="351">
<materialsCitation id="6B06EFB424A68E2A3BB79031BC673E22" ID-GBIF-Occurrence="2612079455" collectionCode="BM" specimenCode="BM-000646986" typeStatus="lectotype">
<typeStatus id="BF14D79BC9285ADFCC9A5C61B824F0B6">Lectotype</typeStatus>
(Sherff in
<emphasis id="44A42EEDAF90BB9A0A1391C86FB83E9E" italics="true" pageNumber="351">Publ. Field Mus. Nat. Hist., Bot. Ser.</emphasis>
16: 271. 1937): Herb. Clifford: 399,
<taxonomicName id="9B1DEF90C1C22907A91E8F7260425D9B" class="Magnoliopsida" family="Asteraceae" genus="Bidens" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="genus">
<emphasis id="607B908B4CEB18CFC268894DFDE8EF4E" italics="true" pageNumber="351">Bidens</emphasis>
</taxonomicName>
1 α (BM-000646986)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="37976DE5A9A05C2AFB90C110B926526E" pageNumber="351" type="materials_examined">
<paragraph id="6793F26A95DA6E8B2DAF358EC6F0487D" pageNumber="351">
<typeStatus id="5358281811A8C5F65F6D5CEDA75A4737">Generitype</typeStatus>
of
<taxonomicName id="37661937DA5ECCBE987B461934ECC00A" authority="Linnaeus" authorityName="Linnaeus" class="Magnoliopsida" family="Asteraceae" genus="Bidens" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="genus">
<emphasis id="46BEECA7E782D7A958C024ACADF2560F" italics="true" pageNumber="351">Bidens</emphasis>
Linnaeus
</taxonomicName>
(vide Green,
<emphasis id="4B16D7FC2CF1AA2E3AAF4F3A302661C6" italics="true" pageNumber="351">Prop. Brit. Bot</emphasis>
.: 179. 1929).
</paragraph>
</subSubSection>
<subSubSection id="CDA3167118777564CBE059C988BEF9EB" pageNumber="351" type="nomenclature">
<paragraph id="EE67CFF311323BD297828CD480897C99" pageNumber="351">
Current name:
<taxonomicName id="E3DA12F73E56E5770E280BF85F1B220F" ID-CoL="LSLN" authority="L." authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Asteraceae" genus="Bidens" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="tripartita">
<emphasis id="F1598B89A4143F631D3C8753BDA722B8" bold="true" italics="true" pageNumber="351">Bidens tripartita</emphasis>
L.
</taxonomicName>
(
<taxonomicName id="B4D47117F5485F55132504CED4C1B6C1" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="A80E7B08361CD2523DECE6386BED5965" pageNumber="351" type="discussion">
<paragraph id="DE18853FB68813298645818D0E7B160F" pageNumber="351">
<emphasis id="FA93485E3E054244A8CA1C9DD98C4916" italics="true" pageNumber="351">Note:</emphasis>
Harriman (in
<emphasis id="2A1D5CAB7DCD5E336092AA2BEF99CD88" italics="true" pageNumber="351">Taxon</emphasis>
47: 485. 1998) proposed the conservation of the generic name with feminine gender, recommended by the Committee for Spermatophyta (in
<emphasis id="08F7D180931BF14632AD110439B0F835" italics="true" pageNumber="351">Taxon</emphasis>
49: 272. 2000). However, the General Committee (in
<emphasis id="B0A52C4D43267E521225B9466C3365A0" italics="true" pageNumber="351">Taxon</emphasis>
55: 796-797. 2006) referred this back to the Committee for
<volumeTitle id="73BF7DB4837C670D18BC6120ABC26548">Vascular Plants</volumeTitle>
for reconsideration.
</paragraph>
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<emphasis id="52D2EA873F74FFFC30D5B82CA0577F3B" bold="true" box="[178,332,547,569]" italics="true" pageId="1" pageNumber="1862">Corynis similis</emphasis>
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M a t e r i a l:3 Lambousa, Nordküste,
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,
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, in Blüten von
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, 1 leg., in coll. et det. W. Sch 08.
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W i r t s p f l a n z e n: Die Larven fressen Blütenblätter, später die Blütenzentren von
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und
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(
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).
</paragraph>
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V e r b r e i t u n g:JonischeInseln,Peleponnes, Attika, Kreta, Thira, Chios, S-Zypern,
<collectingCountry id="18B176053F74FFFC30D5B8E2A1EC7E01" box="[178,247,749,771]" name="Syria" pageId="1" pageNumber="1862">Syrien</collectingCountry>
,
<collectingCountry id="18B176053F74FFFC3163B8E2A0247E01" box="[260,319,749,771]" name="Israel" pageId="1" pageNumber="1862">Israel</collectingCountry>
,
<collectingCountry id="18B176053F74FFFC312BB8E2A0B57E01" box="[332,430,749,771]" name="Jordan" pageId="1" pageNumber="1862">Jordanien</collectingCountry>
(
<bibRefCitation id="04374B643F74FFFC31A7B8E2A3487E01" author="BENSON R" box="[448,595,749,771]" pageId="1" pageNumber="1862" pagination="109 - 207" refId="ref2280" refString="BENSON R. B. (1968): Hymenoptera from Turkey. Symphyta. - Bull. Brit. Mus. (Nat. Hist.) Ent. 22: 109 - 207." type="journal article" year="1968">BENSON 1968</bibRefCitation>
;
<bibRefCitation id="04374B643F74FFFC3207B8E2A2077E01" author="GREATHEAD D" box="[608,796,749,771]" pageId="1" pageNumber="1862" pagination="107 - 111" refId="ref2340" refString="GREATHEAD D. G. (1978): The larvae and notes on the life history of Corynis similis (MOCSARY) (Hymenoptera: Cimbicidae). - J. nat. Hist., London 12: 107 - 111." type="journal article" year="1978">GREATHEAD 1978</bibRefCitation>
;
<bibRefCitation id="04374B643F74FFFC334DB8E2A2A27E01" author="SCHEDL W" box="[810,953,749,771]" pageId="1" pageNumber="1862" pagination="137 - 140" refId="ref2621" refString="SCHEDL W. (2005): Eine neue minierende Blattwespe des Genus Heterarthrus von Zypern (Hymenoptera: Symphyta: Tenthredinidae). - Ber. nat. - med. Ver. Innsbruck 92: 137 - 140." type="journal article" year="2005">SCHEDL 2005</bibRefCitation>
,
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). Neu für N-Zypern.
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<emphasis id="52D2EA873F74FFFC30D5BF02A08C7820" bold="true" box="[178,407,1293,1315]" italics="true" pageId="1" pageNumber="1862">Heterarthrus cypricus</emphasis>
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2005
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M a t e r i a l: 5 Platzminen mit Kot und Larvenhäuten an
<emphasis id="52D2EA873F74FFFC32BDBF33A26C7852" box="[730,887,1339,1361]" italics="true" pageId="1" pageNumber="1862">Acer obtusifolium</emphasis>
(Sibth. et Smith.), St. Hilarion, 1. Parkplatz Umgebung, ca.
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, Pentadaktylos-Gebirge.
<date id="141810553F74FFFC335CBF5AA28F786B" box="[827,916,1365,1385]" pageId="1" pageNumber="1862" value="2008-10-04">10.4.2008</date>
, leg., in coll. et det. W. Sch. 08.
</paragraph>
</subSubSection>
<subSubSection id="28BC651E3F74FFFC30D5BF9FA3B07B20" pageId="1" pageNumber="1862" type="description">
<paragraph id="601936953F74FFFC30D5BF9FA21678E2" blockId="1.[178,1085,1423,1504]" pageId="1" pageNumber="1862">
W i r t s p f l a n z e (n): bisher nur an Blättern von
<emphasis id="52D2EA873F74FFFC336DBF9FA2DF78A7" box="[778,964,1424,1445]" italics="true" pageId="1" pageNumber="1862">Acer obtusifolium</emphasis>
(SIBTH. &amp; SMITH.) nachgewiesen. Auf der gesamten
<collectingCountry id="18B176053F74FFFC3201BFA2A3F378C0" box="[614,744,1453,1475]" name="Insel Zypern" pageId="1" pageNumber="1862">Insel Zypern</collectingCountry>
soll nur diese
<emphasis id="52D2EA873F74FFFC33E1BFA2A2AD78C0" box="[902,950,1453,1474]" italics="true" pageId="1" pageNumber="1862">Acer</emphasis>
Art vorkommen. Andere Artnamen auf
<collectingCountry id="18B176053F74FFFC31ABBFC4A30E78E2" box="[460,533,1483,1504]" name="Cyprus" pageId="1" pageNumber="1862">Zypern</collectingCountry>
bezogen stimmen nicht.
</paragraph>
<paragraph id="601936953F74FFFC30D5BFFFA3B07B20" blockId="1.[178,1084,1519,1571]" pageId="1" pageNumber="1862">
V e r b r e i t u n g:BishernurinS-Zyperngefunden., leg. A.C. &amp; W.N. Ellis (
<bibRefCitation id="04374B643F74FFFC338BBFE0A1FC7B21" author="SCHEDL W" pageId="1" pageNumber="1862" pagination="137 - 140" refId="ref2621" refString="SCHEDL W. (2005): Eine neue minierende Blattwespe des Genus Heterarthrus von Zypern (Hymenoptera: Symphyta: Tenthredinidae). - Ber. nat. - med. Ver. Innsbruck 92: 137 - 140." type="journal article" year="2005">SCHEDL 2005</bibRefCitation>
, LISTON &amp; SPÄTH 2008). Neu für N-Zypern.
</paragraph>
</subSubSection>
</treatment>
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126
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<paragraph id="601936953F77FFFF30D5BF05A3587822" blockId="2.[178,579,1290,1312]" box="[178,579,1290,1312]" pageId="2" pageNumber="1863">
<heading id="3B5181F93F77FFFF30D5BF05A3587822" box="[178,579,1290,1312]" fontSize="8" level="2" pageId="2" pageNumber="1863" reason="2">
<taxonomicName id="A7A64D163F77FFFF30D5BF05A3587822" authority="(STEIN 1876)" baseAuthorityName="STEIN" baseAuthorityYear="1876" box="[178,579,1290,1312]" class="Insecta" family="Cephidae" genus="Pachycephus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="1863" phylum="Arthropoda" rank="species" species="smyrnensis">
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<emphasis id="52D2EA873F77FFFF30D5BF05A0AD7822" bold="true" box="[178,438,1290,1312]" italics="true" pageId="2" pageNumber="1863">Pachycephus smyrnensis</emphasis>
(STEIN 1876)
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<materialsCitation id="D0CE3CC83F77FFFF30D5BF35A50D784C" box="[178,1046,1338,1358]" collectionCode="BMNH" collectorName="Mavr." location="Kyrenia" pageId="2" pageNumber="1863" specimenCode="BENSON 1954" specimenCount="1" stateProvince="Kyrenia">
M a t e r i a l:? Ex.
<collectingRegion id="A262F8773F77FFFF311ABF35A0D8784C" box="[381,451,1338,1358]" country="Cyprus" name="Kyrenia" pageId="2" pageNumber="1863">Kyrenia</collectingRegion>
(=
<collectingRegion id="A262F8773F77FFFF31BCBF35A30A784C" box="[475,529,1338,1358]" country="Cyprus" name="Kyrenia" pageId="2" pageNumber="1863">Girne</collectingRegion>
),
<date id="141810553F77FFFF3278BF35A36E784C" box="[543,629,1338,1358]" pageId="2" pageNumber="1863" value="1931-04-03">3.-4-1931</date>
-35, leg.
<collectorName id="CD5353433F77FFFF32A6BF35A3E2784C" box="[705,761,1338,1358]" pageId="2" pageNumber="1863">Mavr.</collectorName>
, in coll.
<collectionCode id="06B7AE503F77FFFF3325BF35A29E784C" box="[834,901,1338,1358]" pageId="2" pageNumber="1863">BMNH</collectionCode>
(
<bibRefCitation id="04374B643F77FFFF33F6BF35A50B784C" author="BENSON R" box="[913,1040,1338,1358]" pageId="2" pageNumber="1863" pagination="267 - 2296" refId="ref2234" refString="BENSON R. B. (1954): Some sawflies of the European Alps and the Mediterranean Region (Hymenoptera: Symphyta). - Bull. brit. Mus. (Nat. Hist.) Ent. 3: 267 - 2296." type="journal article" year="1954">
<specimenCode id="30009EEE3F77FFFF33F6BF35A50B784C" box="[913,1040,1338,1358]" collectionCode="BENSON" pageId="2" pageNumber="1863">BENSON 1954</specimenCode>
</bibRefCitation>
)
</materialsCitation>
; 1 Lambousa, N-Küste, ca.
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,
<date id="141810553F77FFFF3197BF5DA3257864" box="[496,574,1362,1382]" pageId="2" pageNumber="1863" value="2008-04-06">6.4.2008</date>
, in Blüte von
<emphasis id="52D2EA873F77FFFF32DEBF5EA2587867" box="[697,835,1360,1382]" italics="true" pageId="2" pageNumber="1863">Papaver rhoeas</emphasis>
, leg., in coll. et det. W. Sch. 08.
</paragraph>
</subSubSection>
<subSubSection id="28BC651E3F77FFFE30D5BF82A2E17C4A" lastPageId="3" lastPageNumber="1864" pageId="2" pageNumber="1863" type="description">
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W i r t s p f l a n z e n: Larven in Stengeln von
<emphasis id="52D2EA873F77FFFF32CCBF82A25778A0" box="[683,844,1421,1442]" italics="true" pageId="2" pageNumber="1863">Papaver rhoeas</emphasis>
,
<emphasis id="52D2EA873F77FFFF333FBF82A2A278A0" box="[856,953,1421,1442]" italics="true" pageId="2" pageNumber="1863">P. dubius</emphasis>
,
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<emphasis id="52D2EA873F77FFFF33A2BF82A52778A0" box="[965,1084,1421,1442]" italics="true" pageId="2" pageNumber="1863">P. orientale</emphasis>
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und
<emphasis id="52D2EA873F77FFFF30B9BFA4A06A78BD" box="[222,369,1450,1472]" italics="true" pageId="2" pageNumber="1863">P. somniferum</emphasis>
(
<bibRefCitation id="04374B643F77FFFF31E7BFA5A37D78C2" author="SCHEIBELREITER G" box="[384,614,1450,1472]" pageId="2" pageNumber="1863" pagination="19 - 25" refId="ref2745" refString="SCHEIBELREITER G. (1978): The poppy-cephid, Pachycephus smyrnensis STEIN (Hymenoptera: Cephidae). - Ztsch. ang. Ent. 86: 19 - 25." type="journal article" year="1978">SCHEIBELREITER 1978</bibRefCitation>
).
</paragraph>
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V e r b r e i t u n g Balkan,
<collectingCountry id="18B176053F77FFFF318CBFC0A33178E7" box="[491,554,1487,1509]" name="Turkey" pageId="2" pageNumber="1863">Türkei</collectingCountry>
, Transkaukasien,
<collectingCountry id="18B176053F77FFFF328BBFC0A23278E7" box="[748,809,1487,1509]" name="Syria" pageId="2" pageNumber="1863">Syrien</collectingCountry>
, Isreal,
<collectingCountry id="18B176053F77FFFF33E2BFC0A2F978E7" box="[901,994,1487,1509]" name="Jordan" pageId="2" pageNumber="1863">Jordanien</collectingCountry>
, S- und N-Zypern (
<bibRefCitation id="04374B643F77FFFF3143BFE3A0A87B00" author="BENSON R" box="[292,435,1516,1538]" pageId="2" pageNumber="1863" pagination="109 - 207" refId="ref2280" refString="BENSON R. B. (1968): Hymenoptera from Turkey. Symphyta. - Bull. Brit. Mus. (Nat. Hist.) Ent. 22: 109 - 207." type="journal article" year="1968">BENSON 1968</bibRefCitation>
; SCHEDL &amp; KRAUS 1988;
<bibRefCitation id="04374B643F77FFFF32DDBFE3A25C7B00" author="SCHEDL W" box="[698,839,1516,1538]" pageId="2" pageNumber="1863" pagination="535 - 541" refId="ref2662" refString="SCHEDL W. (2007): Ein Beitrag zur Kenntnis der Pflanzenwespenfauna von Jordanien (Insecta: Hymenoptera; Symphyta). - Linzer biol. Beitr. 39 (1): 535 - 541." type="journal article" year="2007">SCHEDL 2007</bibRefCitation>
).
</paragraph>
<paragraph id="601936953F76FFFE30D5BAD4A3B47DF2" blockId="3.[178,1085,219,328]" box="[178,687,219,241]" pageId="3" pageNumber="1864">
Nachtrag zur Symphyten-Fauna von ganz
<collectingCountry id="18B176053F76FFFE3239BAD4A3B07DF2" box="[606,683,219,240]" name="Cyprus" pageId="3" pageNumber="1864">Zypern</collectingCountry>
:
</paragraph>
<paragraph id="601936953F76FFFE30D5BAF7A2E17C4A" blockId="3.[178,1085,219,328]" pageId="3" pageNumber="1864">
<emphasis id="52D2EA873F76FFFE30D5BAF7A08B7C0F" box="[178,400,248,269]" italics="true" pageId="3" pageNumber="1864">Calameuta filiformis</emphasis>
(EVERSMANN 1847): 1
<collectingCountry id="18B176053F76FFFE32A0BAF6A20B7C0C" box="[711,784,249,270]" name="Cyprus" pageId="3" pageNumber="1864">Zypern</collectingCountry>
NW [S-Zypern], Akamas Halbinsel, Umg. Palis-Latchi,
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,
<date id="141810553F76FFFE3255BB1AA3CB7C29" box="[562,720,277,299]" pageId="3" pageNumber="1864" value="2007-04-24" valueMax="2007-04-30" valueMin="2007-04-24">24.-30.IV.2007</date>
, E. Heiss (leg.), in coll. et det. W. Sch. 2008 – Zweitnachweis für
<collectingCountry id="18B176053F76FFFE3193BB3CA3267C4A" box="[500,573,307,328]" name="Cyprus" pageId="3" pageNumber="1864">Zypern</collectingCountry>
als Ganzes (siehe SCHEDL &amp; KRAUS 1988).
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174
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<mods:namePart id="80C2315D4F7EA75514255E03C6ED3D33">Dai, Li</mods:namePart>
<mods:affiliation id="9108786E7FAEC3EF5F27322D5A2908AA">Shanghai Entomological Museum, Chinese Academy of Sciences, Shanghai, 200032, China</mods:affiliation>
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<mods:namePart id="C98969EED92E43026DBB8C14E09D74B4">Yin, Zhan</mods:namePart>
<mods:affiliation id="72C7BEEC0FAC9951315638B044588EBE">College of Life Sciences, Hebei University, Baoding, Hebei, 071002, China &amp; College of Forestry Engineering, Shandong Agriculture and Engineering University, Jinan, Shandong, 250100, China</mods:affiliation>
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<mods:date id="3E0481940D5344A680D1A1B57BE6654D">2021</mods:date>
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<treatment id="E80F87D6DC67FFB0FF22FB78342C5A9C" ID-DOI="http://doi.org/10.5281/zenodo.5706803" ID-GBIF-Taxon="190302587" ID-Zenodo-Dep="5706803" LSID="urn:lsid:plazi:treatment:E80F87D6DC67FFB0FF22FB78342C5A9C" httpUri="http://treatment.plazi.org/id/E80F87D6DC67FFB0FF22FB78342C5A9C" lastPageId="1" lastPageNumber="449" pageId="0" pageNumber="448">
<subSubSection id="28BC654BDC67FFB1FF22FB78313E5B4B" pageId="0" pageNumber="448" type="nomenclature">
<paragraph id="601936C0DC67FFB1FF22FB78332A59C4" blockId="0.[151,1437,1215,2043]" box="[151,773,1215,1242]" pageId="0" pageNumber="448">
<heading id="3B5181ACDC67FFB1FF22FB78332A59C4" bold="true" box="[151,773,1215,1242]" fontSize="11" level="1" pageId="0" pageNumber="448" reason="1">
<emphasis id="52D2EAD2DC67FFB1FF22FB78332A59C4" bold="true" box="[151,773,1215,1242]" pageId="0" pageNumber="448">
<taxonomicName id="A7A64D43DC67FFB1FF22FB78323559C7" ID-CoL="9GZQS" authority="Zhang &amp; Wang &amp; Dai &amp; Yin, 2021" authorityName="Zhang &amp; Wang &amp; Dai &amp; Yin" authorityYear="2021" box="[151,538,1215,1242]" class="Insecta" family="Acrididae" genus="Mongolotettix" kingdom="Animalia" order="Orthoptera" pageId="0" pageNumber="448" phylum="Arthropoda" rank="species" species="baohuashanensis" status="sp. nov.">
<emphasis id="52D2EAD2DC67FFB1FF22FB78323559C7" bold="true" box="[151,538,1215,1242]" italics="true" pageId="0" pageNumber="448">Mongolotettix baohuashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E157A9DC67FFB1FD97FB78325059C4" box="[546,639,1215,1241]" pageId="0" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="F89D2A45DC67FFB1FD3AFB7832D559C4" box="[655,762,1215,1241]" captionStart="FIGURES 1-9" captionStartId="1.[151,264,1693,1717]" captionTargetBox="[213,1378,175,1667]" captionTargetId="figure-26@1.[202,1385,161,1680]" captionTargetPageId="1" captionText="FIGURES 1-9. Mongolotettix baohuashanensis sp. nov. 1. Body in lateral view ♂; 2. Epiphallus, ♂; 3. Head lateral view ♂; 4. Stridulatory pegs ♂; 5. Tegmen ♂; 6. End of abdomen in dorsal view ♂; 7. Body in lateral view ♀; 8. Tegmen ♀; 9. End of abdomen in lateral view ♀. FIGURES 10-14 M. fujianensis Chen et al, 2018 10. Head lateral view ♂; 11. Tegmen ♂; 12. Epiphallus; 13. Tegmen ♀; 14. End of abdomen in lateral view ♀. (figs. 10-14 after Chen et al, 2018)." figureDoi="http://doi.org/10.5281/zenodo.5706801" httpUri="https://zenodo.org/record/5706801/files/figure.png" pageId="0" pageNumber="448">Figs. 1-9</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="601936C0DC67FFB1FF22FB23313E5B4B" blockId="0.[151,1437,1215,2043]" pageId="0" pageNumber="448">
Male: Body small in size. Head shorter than pronotum. Face oblique in profile. widened downward gradually. Antennae ensiform. Eyes oval, vertical diameter 1.5 times horizontal diameter and 1.2 times length of subocular furrow. Pronotum median keel visible, distinctly cut by hind transverse sulcus, lateral keels almost parallel, prozona 1.6 times of metazona in length. Tegmina longer, reaching the hind margin of 7
<superScript id="97D39B88DC67FFB1FD4FFA8D33295845" attach="left" box="[762,774,1354,1368]" fontSize="6" pageId="0" pageNumber="448">th</superScript>
abdominal tergite, apically concave in the middle, maximum width of cubital area 1.6 times maximum width of medial area. Interspace of mesosternum distinctly narrowed in the middle
<emphasis id="52D2EAD2DC67FFB1FF54FA5630D358BA" bold="true" box="[225,252,1425,1447]" pageId="0" pageNumber="448">•</emphasis>
length 2.3 times minimum width. Lateral lobes of metasternum separated. Upper keel of hind femur smooth, length of hind femur 5.6 times its maximum width, end of lower knee lobes angular, inner side with 88 stridulatory pegs. Hind tibia with 13 spines on inner and 12 spines on outer sides, external apical spine absent. Second joint of hind tarsus shorter than the first one. Tympanum large and round. Furculae distinct, apex rounded. Cercus long conical, almost reaching tip of epiproct. Subgenital plate long conical. Width of epiphallus larger than high, with indistinct projection on both inner sides.
</paragraph>
</subSubSection>
<subSubSection id="28BC654BDC67FFB0FF72F9A6342C5A9C" lastPageId="1" lastPageNumber="449" pageId="0" pageNumber="448" type="description">
<paragraph id="601936C0DC67FFB1FF72F9A631665A19" blockId="0.[151,1437,1215,2043]" pageId="0" pageNumber="448">Female: Body more robust than male. Vertical diameter of eyes 1.3 times horizontal diameter and 1.1 times the length of subocular furrow. Prozona 1.7 times metazoan in length. Tegmina short, extending over the middle of 2nd abdominal tergite slightly, length 2.8 times its maximum width. Length of hind femur 5.4 times its maximum width. Cercus shortconical, not reaching end of epiproct. Upper ovipositor valve longer, length 3.3 times maximum width, with small teeth on outer margin.</paragraph>
<paragraph id="601936C0DC67FFB1FF72F8C934D05A71" blockId="0.[151,1437,1215,2043]" pageId="0" pageNumber="448">Coloration: Male body brown. Antennae yellow. Pronotum yellowish-brown. Tegmina of male without a white stripe on the fore margin at base. Tegmina of female with a black longitudinal stripe in the middle. Subgenital plate brown. Abdomen of female yellowish-brown, with a brown longitudinal stripe on both sides, other as same male.</paragraph>
<paragraph id="601936C0DC67FFB1FF72F8B0326F5AAC" blockId="0.[151,1437,1215,2043]" pageId="0" pageNumber="448">
Measurement (in mm): Length of body:
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20.1-20.7,
<specimenCount id="76A0FD49DC67FFB1FD4BF8B0333D5A93" box="[766,786,1911,1934]" pageId="0" pageNumber="448" type="female">♀</specimenCount>
27.9-28.6. Length of tegmina:
<specimenCount id="76A0FD49DC67FFB1FBEFF8B034415A93" box="[1114,1134,1911,1934]" pageId="0" pageNumber="448" type="male">♂</specimenCount>
11.1-11.3,
<specimenCount id="76A0FD49DC67FFB1FB50F8B134DC5A93" box="[1253,1267,1910,1934]" pageId="0" pageNumber="448" type="female">♀</specimenCount>
4.5-5.5. Length of hind femur:
<specimenCount id="76A0FD49DC67FFB1FE8DF85E31645AAC" box="[312,331,1945,1969]" pageId="0" pageNumber="448" type="male">♂</specimenCount>
11.5-13.2,
<specimenCount id="76A0FD49DC67FFB1FE0BF85E31E35AAC" box="[446,460,1945,1969]" pageId="0" pageNumber="448" type="female">♀</specimenCount>
15.8-16.4.
</paragraph>
<paragraph id="601936C0DC67FFB1FF72F87B34DF5AC9" blockId="0.[151,1437,1215,2043]" box="[199,1264,1980,2004]" pageId="0" pageNumber="448">
<materialsCitation id="D0CE3C9DDC67FFB1FF72F87B31635ACE" ID-GBIF-Occurrence="3407191302" box="[199,332,1980,2004]" collectingDate="1984-07-23" latitude="32.166668" location="Baohuashan" longLatPrecision="1212" longitude="119.26667" pageId="0" pageNumber="448" specimenCount="4" specimenCount-male="1" stateProvince="Jiangsu" typeStatus="holotype">
<typeStatus id="BF1D8862DC67FFB1FF72F87B31055AC9" box="[199,298,1980,2004]" pageId="0" pageNumber="448" type="holotype">Holotype</typeStatus>
<specimenCount id="76A0FD49DC67FFB1FE84F87B31685ACE" box="[305,327,1980,2003]" pageId="0" pageNumber="448" type="male">♂</specimenCount>
,
</materialsCitation>
<materialsCitation id="D0CE3C9DDC67FFB1FEE7F87B34DF5AC9" ID-GBIF-Occurrence="3407191301" box="[338,1264,1980,2004]" collectingDate="1984-07-23" latitude="32.166668" location="Baohuashan" longLatPrecision="1212" longitude="119.26667" pageId="0" pageNumber="448" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Jiangsu" typeStatus="paratype">
<typeStatus id="BF1D8862DC67FFB1FEE7F87B31975AC9" box="[338,440,1980,2004]" pageId="0" pageNumber="448" type="paratype">paratypes</typeStatus>
<specimenCount id="76A0FD49DC67FFB1FE0AF87B31CD5AC9" box="[447,482,1980,2004]" pageId="0" pageNumber="448" type="male">1♂</specimenCount>
,
<specimenCount id="76A0FD49DC67FFB1FE5BF87B323F5AC9" box="[494,528,1980,2004]" pageId="0" pageNumber="448" type="female">2♀</specimenCount>
,
<collectingRegion id="A262F822DC67FFB1FDA8F87B325F5AC9" box="[541,624,1980,2004]" country="China" name="Jiangsu" pageId="0" pageNumber="448">Jiangsu</collectingRegion>
,
<location id="6579601BDC67FFB1FDCFF87B32D35AC9" LSID="urn:lsid:plazi:treatment:E80F87D6DC67FFB0FF22FB78342C5A9C:6579601BDC67FFB1FDCFF87B32D35AC9" box="[634,764,1980,2004]" latitude="32.166668" longLatPrecision="1212" longitude="119.26667" name="Baohuashan" pageId="0" pageNumber="448" stateProvince="Jiangsu">Baohuashan</location>
<geoCoordinate id="05925007DC67FFB1FCB6F87B33705AC9" box="[771,863,1980,2004]" degrees="32" direction="north" minutes="10" orientation="latitude" pageId="0" pageNumber="448" precision="925" value="32.166668">32º10´N</geoCoordinate>
,
<geoCoordinate id="05925007DC67FFB1FCDFF87B33FF5AC9" box="[874,976,1980,2004]" degrees="119" direction="east" minutes="16" orientation="longitude" pageId="0" pageNumber="448" precision="925" value="119.26667">119º16´E</geoCoordinate>
,
<date id="14181000DC67FFB1FC6FF87B347E5AC9" box="[986,1105,1980,2004]" pageId="0" pageNumber="448" value="1984-07-23">
<collectingDate id="045CE9E8DC67FFB1FC6FF87B347E5AC9" box="[986,1105,1980,2004]" pageId="0" pageNumber="448" value="1984-07-23">1984-7-23.</collectingDate>
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leg. unknown.
</materialsCitation>
</paragraph>
<paragraph id="601936C0DC67FFB1FF22F824308B5AE6" blockId="0.[151,1437,1215,2043]" box="[151,164,2019,2043]" pageId="0" pageNumber="448">1</paragraph>
<caption id="34D96648DC66FFB0FF22F95A348F5A04" ID-DOI="http://doi.org/10.5281/zenodo.5706801" ID-Zenodo-Dep="5706801" httpUri="https://zenodo.org/record/5706801/files/figure.png" pageId="1" pageNumber="449" startId="1.[151,264,1693,1717]" targetBox="[213,1378,175,1667]" targetPageId="1">
<paragraph id="601936C0DC66FFB0FF22F95A348F5A04" blockId="1.[151,1437,1692,1817]" pageId="1" pageNumber="449">
<emphasis id="52D2EAD2DC66FFB0FF22F95A32CC5BA8" bold="true" box="[151,739,1692,1717]" pageId="1" pageNumber="449">
FIGURES 1-9.
<taxonomicName id="A7A64D43DC66FFB0FE88F95B32A35BA9" authority="Zhang &amp; Wang &amp; Dai &amp; Yin, 2021" authorityName="Zhang &amp; Wang &amp; Dai &amp; Yin" authorityYear="2021" box="[317,652,1692,1716]" class="Insecta" family="Acrididae" genus="Mongolotettix" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="baohuashanensis" status="sp. nov.">
<emphasis id="52D2EAD2DC66FFB0FE88F95B32A35BA9" bold="true" box="[317,652,1692,1716]" italics="true" pageId="1" pageNumber="449">Mongolotettix baohuashanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49E157A9DC66FFB0FD26F95A32CC5BA8" box="[659,739,1693,1717]" pageId="1" pageNumber="449" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1. Body in lateral view ♂; 2. Epiphallus, ♂; 3. Head lateral view ♂; 4. Stridulatory pegs ♂; 5. Tegmen ♂; 6. End of abdomen in dorsal view ♂; 7. Body in lateral view ♀; 8. Tegmen ♀; 9. End of abdomen in lateral view ♀.
<emphasis id="52D2EAD2DC66FFB0FE0FF91A33E55BE8" bold="true" box="[442,970,1756,1781]" pageId="1" pageNumber="449">
FIGURES 10-14
<taxonomicName id="A7A64D43DC66FFB0FDC2F91A33E55BE8" authority="Chen et al, 2018" authorityName="Chen et al" authorityYear="2018" box="[631,970,1756,1781]" class="Insecta" family="Acrididae" genus="Mongolotettix" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="fujianensis">
<emphasis id="52D2EAD2DC66FFB0FDC2F91A33205BE9" bold="true" box="[631,783,1756,1780]" italics="true" pageId="1" pageNumber="449">M. fujianensis</emphasis>
<bibRefCitation id="04374B31DC66FFB0FCA2F91A33E55BE8" author="Chen, S. - Z. &amp; Shi, J. P. &amp; Dai, Li" box="[791,970,1757,1781]" pageId="1" pageNumber="449" pagination="387 - 389" refId="ref1354" refString="Chen, S. - Z., Shi, J. P. &amp; Dai, Li (2018) Description of a new species of the genus Mongolotettix Rehn, 1928 (Orthoptera: Acrididae) from Fujian, China. Zootaxa, 4410 (2), 387 - 389. https: // doi. org / 10.11646 / zootaxa. 4410.2.8" type="journal article" year="2018">Chen et al, 2018</bibRefCitation>
</taxonomicName>
</emphasis>
10. Head lateral view ♂; 11. Tegmen ♂; 12. Epiphallus; 13. Tegmen ♀; 14. End of abdomen in lateral view ♀. (figs. 10-14 after
<bibRefCitation id="04374B31DC66FFB0FC5FF8C634BA5A04" author="Chen, S. - Z. &amp; Shi, J. P. &amp; Dai, Li" box="[1002,1173,1793,1817]" pageId="1" pageNumber="449" pagination="387 - 389" refId="ref1354" refString="Chen, S. - Z., Shi, J. P. &amp; Dai, Li (2018) Description of a new species of the genus Mongolotettix Rehn, 1928 (Orthoptera: Acrididae) from Fujian, China. Zootaxa, 4410 (2), 387 - 389. https: // doi. org / 10.11646 / zootaxa. 4410.2.8" type="journal article" year="2018">
Chen
<emphasis id="52D2EAD2DC66FFB0FB93F8C534175A04" box="[1062,1080,1794,1817]" italics="true" pageId="1" pageNumber="449">et</emphasis>
al, 2018
</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="601936C0DC66FFB0FF72F88135B85A43" blockId="1.[199,1431,1862,1921]" box="[199,1431,1862,1886]" pageId="1" pageNumber="449">
Diagnosis. The new species is similar to
<taxonomicName id="A7A64D43DC66FFB0FDCDF88033E95A43" authority="Chen et al, 2018" authorityName="Chen et al" authorityYear="2018" box="[632,966,1862,1886]" class="Insecta" family="Acrididae" genus="Mongolotettix" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="fujianensis">
<emphasis id="52D2EAD2DC66FFB0FDCDF88033215A43" box="[632,782,1862,1886]" italics="true" pageId="1" pageNumber="449">M. fujianensis</emphasis>
<bibRefCitation id="04374B31DC66FFB0FCA0F88133E95A43" author="Chen, S. - Z. &amp; Shi, J. P. &amp; Dai, Li" box="[789,966,1862,1886]" pageId="1" pageNumber="449" pagination="387 - 389" refId="ref1354" refString="Chen, S. - Z., Shi, J. P. &amp; Dai, Li (2018) Description of a new species of the genus Mongolotettix Rehn, 1928 (Orthoptera: Acrididae) from Fujian, China. Zootaxa, 4410 (2), 387 - 389. https: // doi. org / 10.11646 / zootaxa. 4410.2.8" type="journal article" year="2018">Chen et al, 2018</bibRefCitation>
</taxonomicName>
. The major differences are listed in
<tableCitation id="2D24037BDC66FFB0FAF7F88135BE5A43" box="[1346,1425,1862,1886]" captionStart="TABLE 1" captionStartId="2.[151,244,151,177]" captionTargetPageId="2" captionText="TABLE 1. Comparison of M. baohuashanensis sp. nov. and M. fujianensis Chen et al, 2018" httpUri="http://table.plazi.org/id/34D96648DC65FFB3FF22FF50348F5DAC" pageId="1" pageNumber="449" tableUuid="34D96648DC65FFB3FF22FF50348F5DAC">Table 1</tableCitation>
.
</paragraph>
<paragraph id="601936C0DC66FFB0FF72F8AE342C5A9C" blockId="1.[199,1431,1862,1921]" box="[199,1027,1897,1921]" pageId="1" pageNumber="449">
<emphasis id="52D2EAD2DC66FFB0FF72F8AE31645A9C" bold="true" box="[199,331,1897,1921]" pageId="1" pageNumber="449">Etymology:</emphasis>
The specific epithet is named after Baohuashan the
<typeStatus id="BF1D8862DC66FFB0FCCCF8AE33885A9C" box="[889,935,1897,1921]" pageId="1" pageNumber="449">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

71
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<mods:title>Cryptocarya kaengkrachanensis, a new species of Lauraceae from Kaeng Krachan National Park, southwest Thailand</mods:title>
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<mods:namePart>Yahara, Tetsukazu</mods:namePart>
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<mods:namePart>Tagane, Shuichiro</mods:namePart>
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<mods:namePart>Suddee, Somran</mods:namePart>
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<mods:namePart>Moritsuka, Etsuko</mods:namePart>
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<taxonomicName LSID="E8102981-1BD3-5137-800B-04A6D0B0B0C9" authority="M. Z. Zhang, Yahara &amp; Tagane" authorityName="M. Z. Zhang, Yahara &amp; Tagane" authorityYear="2020" class="Magnoliopsida" family="Lauraceae" genus="Cryptocarya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptocarya kaengkrachanensis" order="Laurales" pageId="0" pageNumber="139" phylum="Tracheophyta" rank="species" species="kaengkrachanensis" status="sp. nov.">Cryptocarya kaengkrachanensis M.Z.Zhang, Yahara &amp; Tagane</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="139">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cryptocarya kaengkrachanensis M. Z. Zhang, Yahara &amp; Tagane A branch with immature fruit B lower leaf surface C holotype: Tagane et al. T 2069 (KYO) D young branchlet E part of an infructescence with immature fruits F longitudinal sections of an immature fruit." figureDoi="10.3897/phytokeys.140.34574.figure2" httpUri="https://binary.pensoft.net/fig/385178" pageId="0" pageNumber="139">Fig. 2</figureCitation>
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<paragraph pageId="0" pageNumber="139">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="139">
<taxonomicName authorityName="M. Z. Zhang, Yahara &amp; Tagane" authorityYear="2020" class="Magnoliopsida" family="Lauraceae" genus="Cryptocarya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptocarya kaengkrachanensis" order="Laurales" pageId="0" pageNumber="139" phylum="Tracheophyta" rank="species" species="kaengkrachanensis">
<emphasis italics="true" pageId="0" pageNumber="139">Cryptocarya kaengkrachanensis</emphasis>
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resembles
<taxonomicName lsidName="C. amygdalina" pageId="0" pageNumber="139" rank="species" species="amygdalina">
<emphasis italics="true" pageId="0" pageNumber="139">C. amygdalina</emphasis>
</taxonomicName>
in having pinnately veined, leathery leaves apparently glabrous (microscopically hairy) below, young twigs with yellowish brown hairs and fruits 1.36-1.85 times longer than width. However,
<taxonomicName lsidName="C. kaengkrachanensis" pageId="0" pageNumber="139" rank="species" species="kaengkrachanensis">
<emphasis italics="true" pageId="0" pageNumber="139">C. kaengkrachanensis</emphasis>
</taxonomicName>
differs from
<taxonomicName lsidName="C. amygdalina" pageId="0" pageNumber="139" rank="species" species="amygdalina">
<emphasis italics="true" pageId="0" pageNumber="139">C. amygdalina</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cryptocarya amygdalina A leafy branchlet B lower leaf surface C specimen Tagane et al. T 3090 (KYO) D lower leaf surface (dry) E fresh immature fruit F dried fruit." figureDoi="10.3897/phytokeys.140.34574.figure3" httpUri="https://binary.pensoft.net/fig/385179" pageId="0" pageNumber="139">3</figureCitation>
) in having the leaves ovate and elliptic to narrowly elliptic (vs. oblong-lanceolate) with leaf aspect ratio from 1.38 to 2.28 (vs. 2.46-3.43) (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scatter plot of leaf aspect ratio (horizontal axis) and circularity (vertical axis). Solid circles: Cryptocarya amygdalina, solid triangles: C. kaengkrachanensis." figureDoi="10.3897/phytokeys.140.34574.figure4" httpUri="https://binary.pensoft.net/fig/385180" pageId="0" pageNumber="139">4</figureCitation>
), and fruits ovoid (vs. ellipsoid) with the stalk distinctly swollen (vs. not or only slightly swollen). While
<taxonomicName lsidName="C. kaengkrachanensis" pageId="0" pageNumber="139" rank="species" species="kaengkrachanensis">
<emphasis italics="true" pageId="0" pageNumber="139">C. kaengkrachanensis</emphasis>
</taxonomicName>
was sister to
<taxonomicName lsidName="C. albiramea" pageId="0" pageNumber="139" rank="species" species="albiramea">
<emphasis italics="true" pageId="0" pageNumber="139">C. albiramea</emphasis>
</taxonomicName>
in MIG-seq tree,
<taxonomicName lsidName="C. kaengkrachanensis" pageId="0" pageNumber="139" rank="species" species="kaengkrachanensis">
<emphasis italics="true" pageId="0" pageNumber="139">C. kaengkrachanensis</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName lsidName="C. albiramea" pageId="0" pageNumber="139" rank="species" species="albiramea">
<emphasis italics="true" pageId="0" pageNumber="139">C. albiramea</emphasis>
</taxonomicName>
by elliptic leaves with leaf aspect ratio less than 2.5 (vs. oblong-lanceolate leaves with leaf aspect ratio more than 2.5).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="materials_examined">
<paragraph pageId="0" pageNumber="139">Type.</paragraph>
<paragraph pageId="0" pageNumber="139">
THAILAND. Phetchaburi Province: Kaeng Krachan National Park, 960 m elev.,
<geoCoordinate degrees="12" direction="north" minutes="49" orientation="latitude" precision="1" seconds="19.7" value="12.822139">12°49'19.7&quot;N</geoCoordinate>
,
<geoCoordinate degrees="99" direction="east" minutes="21" orientation="longitude" precision="1" seconds="57.7" value="99.36603">99°21'57.7&quot;E</geoCoordinate>
, 23 Oct. 2013,
<emphasis italics="true" pageId="0" pageNumber="139">Tagane S.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Nagamasu H.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Naiki A.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Rueangruea S.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Suddee S.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Fuse K.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Keiwbang W.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Pansamrong P. T2069</emphasis>
[fr.] (holotype KYO!, isotypes BKF!, FU!, KAG!).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="description">
<paragraph pageId="0" pageNumber="139">Description.</paragraph>
<paragraph pageId="0" pageNumber="139">
Trees up to 12 m tall. Young twigs densely covered with appressed short yellowish to brown hairs, old twigs lenticellate, terete and slightly hairy. Leaves alternate; blade leathery, ovate, elliptic to narrowly elliptic, the range (and average
<normalizedToken originalValue="±">+/-</normalizedToken>
SD) of leaf length (cm) is 2.6-10.3 long (7.2
<normalizedToken originalValue="±">+/-</normalizedToken>
2.5, n=17), the range (and average
<normalizedToken originalValue="±">+/-</normalizedToken>
SD) of leaf width (cm) is 1.5-6.4 (4.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6, n=17) wide leaf aspect ratio from 1.38 to 2.28, obtuse or retuse at apex in adult trees, acuminate in young trees, broadly cuneate at base, green and not lustrous above and slightly glaucous below when fresh, brown above and grey brown below when dry, apparently glabrous but microscopically sparsely hairy below; pinnately veined, midrib sunken above, raised below, secondary veins 6 or 7 pairs, slightly sunken above, raised below, tertiary veins scalariform-reticulate, faintly visible above, raised below; the range (and average
<normalizedToken originalValue="±">+/-</normalizedToken>
SD) of petiole length (cm) is 0.7-1.5 long (1.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.22, n=10), flat above, rounded below, dark brown when dry, covered with short yellowish hairs. Inflorescences and flowers not seen. Infructescence axillary, 4-17 cm long (8.4
<normalizedToken originalValue="±">+/-</normalizedToken>
3.4, n=16) (the range, average
<normalizedToken originalValue="±">+/-</normalizedToken>
SD), rachis hairy, lenticellate; bracteoles not seen. Immature fruits ovoid, 9.9-13.8 mm long (11.49
<normalizedToken originalValue="±">+/-</normalizedToken>
1.28, n=14), 5.8-9.7 mm wide (7.53
<normalizedToken originalValue="±">+/-</normalizedToken>
2.36, n=14) with aspect ratio 1.36-1.85 (1.54
<normalizedToken originalValue="±">+/-</normalizedToken>
0.14, n=14). yellow green when fresh, dark brown when dry, shortly hairy. Fruiting stalk slightly swollen, rough, light brown when fresh, dark brown when dry. Mature fruits not seen.
</paragraph>
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<subSubSection pageId="0" pageNumber="139" type="other specimens of c. kaengkrachanensis examined">
<paragraph pageId="0" pageNumber="139">
Other specimens of
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<emphasis italics="true" pageId="0" pageNumber="139">C. kaengkrachanensis</emphasis>
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examined.
</paragraph>
<paragraph pageId="0" pageNumber="139">
THAILAND. Phrae Province: between Ban Nam Krai and Pha Tuem, 16 Apr 1970,
<emphasis italics="true" pageId="0" pageNumber="139">Smitinand T.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Cheke A.S. 10817</emphasis>
[BKF 46511!]. Phetchaburi Province: Kaeng Krachan National Park, 960 m elev.,
<geoCoordinate degrees="12" direction="north" minutes="49" orientation="latitude" precision="1" seconds="19.7" value="12.822139">12°49'19.7&quot;N</geoCoordinate>
,
<geoCoordinate degrees="99" direction="east" minutes="21" orientation="longitude" precision="1" seconds="57.7" value="99.36603">99°21'57.7&quot;E</geoCoordinate>
, 23 Oct. 2013,
<emphasis italics="true" pageId="0" pageNumber="139">Tagane S.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Nagamasu H.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Naiki A.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Rueangruea S.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Suddee S.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Fuse K.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Keiwbang W.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Pansamrong P. T1883</emphasis>
(BKF!, FU!). Kanchanaburi Province: Thong Pha Phum District, Pilok, at the Thai-Burmanese border. C. 900 m. 14 41.0'N, 98 21.8'E, tree 12m, 25 January 2009 [fr.],
<emphasis italics="true" pageId="0" pageNumber="139">Middleton D.J.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Karaket P.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Lindsay S.</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="139">Suddee S. 4785</emphasis>
[BKF 182421!].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="distribution">
<paragraph pageId="0" pageNumber="139">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="139">Endemic to Thailand. The new species is currently only known in a few protected areas of Phrae, Phetchaburi and Kanchanaburi Provinces including Kaeng Krachan National Park.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="etymology">
<paragraph pageId="0" pageNumber="139">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="139">
The specific epithet
<emphasis italics="true" pageId="0" pageNumber="139">kaengkrachanensis</emphasis>
is derived from the name of the national park from which the species has first been recorded.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="conservation status">
<paragraph pageId="0" pageNumber="139">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="139">
Least Concern (
<bibRefCitation author="IUCN" journalOrPublisher="IUCN, Gland and Cambridge" pageId="0" pageNumber="139" refId="B10" refString="IUCN, 2012. IUCN Red List Categories and Criteria, Version 3.1. (2nd edn). IUCN, Gland and Cambridge" title="IUCN Red List Categories and Criteria, Version 3.1. (2 nd edn)." year="2012">IUCN 2012</bibRefCitation>
,
<bibRefCitation author="IUCN" journalOrPublisher="IUCN, Gland and Cambridge" pageId="0" pageNumber="139" refId="B11" refString="IUCN, 2017. Guidelines for Using the IUCN Red List Categories and Criteria. Version 13. Prepared by the Standards and Petitions Subcommittee." title="Guidelines for Using the IUCN Red List Categories and Criteria. Version 13. Prepared by the Standards and Petitions Subcommittee." year="2017">2017</bibRefCitation>
). This species occurs in hill evergreen forests of some protected areas including Kaeng Krachan National Park and there is no sign of declining trends.
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName LSID="urn:lsid:ipni.org:names:77145586-1" authority="RD Gore, SP Gaikwad &amp; SD Randive, 2015" authorityName="RD Gore, SP Gaikwad &amp; SD Randive" authorityYear="2015" class="Magnoliopsida" family="Fabaceae" genus="Vigna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vigna pandeyana" order="Fabales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="species" species="pandeyana">Vigna pandeyana RD Gore, SP Gaikwad &amp; SD Randive, 2015</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="4606">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4606" type="materials_examined">
<paragraph pageId="0" pageNumber="4606">Materials</paragraph>
<paragraph pageId="0" pageNumber="4606">
<materialsCitation collectingDate="2012-09-27" country="India" latitude="17.596085" location="near Chalkewadi" longLatPrecision="1" longitude="73.84425" pageId="0" pageNumber="4606" specimenCount="1" typeStatus="Holotype">
Type status:
<typeStatus pageId="0" pageNumber="4606">Holotype</typeStatus>
. Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="4606">India</collectingCountry>
; countryCode: IND; stateProvince: Maharashtra; county: Sahyadri Ranges; municipality: Satara district; locality:
<location LSID="urn:lsid:plazi:treatment:E810301B37B5573346755260FD7CD26E:DE5B228296738124167B6A79AC234056" country="India" latitude="17.596085" longLatPrecision="1" longitude="73.84425" name="near Chalkewadi" pageId="0" pageNumber="4606">near Chalkewadi</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="4606">1050 m</elevation>
; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.596085">17°35'45.9&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="east" orientation="longitude" precision="1" value="73.84425">73°50'39.3&quot;E</geoCoordinate>
; verbatimCoordinateSystem: degrees minutes seconds; Event: eventDate:
<collectingDate pageId="0" pageNumber="4606" value="2012-09-27">27 September 2012</collectingDate>
; habitat: It grows in lateritic gravelly soil on hill slopes in or on grasses and herbs; fieldNumber: RD Gore 1007; fieldNotes: Twining annual herbs; stems terete; flowers are of two kinds; the chasmogamous flowers present on leafy aerial shoots while cleistogamous flowers present on leafless shoots, which are close to soil surface; pods hairy; Record Level: language: English; institutionID: CAL; basisOfRecord: Herbarium specimen
</materialsCitation>
<materialsCitation collectingDate="2012-09-27" country="India" latitude="17.596085" location="near Chalkewadi" longLatPrecision="1" longitude="73.84425" pageId="0" pageNumber="4606" specimenCount="1" typeStatus="Isotype">
Type status:
<typeStatus pageId="0" pageNumber="4606">Isotype</typeStatus>
. Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="4606">India</collectingCountry>
; countryCode: IND; stateProvince: Maharashtra; county: Sahyadri Ranges; municipality: Satara district; locality:
<location LSID="urn:lsid:plazi:treatment:E810301B37B5573346755260FD7CD26E:7F4962D6BE77C91730573D832EB8C1D3" country="India" latitude="17.596085" longLatPrecision="1" longitude="73.84425" name="near Chalkewadi" pageId="0" pageNumber="4606">near Chalkewadi</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="4606">1050 m</elevation>
; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.596085">17°35'45.9&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="east" orientation="longitude" precision="1" value="73.84425">73°50'39.3&quot;E</geoCoordinate>
; verbatimCoordinateSystem: degrees minutes seconds; Event: eventDate:
<collectingDate pageId="0" pageNumber="4606" value="2012-09-27">27 September 2012</collectingDate>
; habitat: It grows in lateritic gravelly soil on hill slopes in or on grasses and herbs; fieldNumber: RD Gore 1007; fieldNotes: Twining annual herbs; stems terete; flowers are of two kinds; the chasmogamous flowers present on leafy aerial shoots while cleistogamous flowers present on leafless shoots, which are close to soil surface; pods hairy; Record Level: language: English; institutionID: BSI, K, SUK; basisOfRecord: Herbarium specimen
</materialsCitation>
<materialsCitation collectingDate="2012-11-10" country="India" latitude="19.683916" location="Kasara-Ghat near Igatpuri" longLatPrecision="1" longitude="73.49953" pageId="0" pageNumber="4606" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="4606">Other material</typeStatus>
. Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="4606">India</collectingCountry>
; countryCode: IND; stateProvince: Maharashtra; municipality: Nasik District; locality:
<location LSID="urn:lsid:plazi:treatment:E810301B37B5573346755260FD7CD26E:36AA10FE4A8DF7681769BD8DB7910221" country="India" latitude="19.683916" longLatPrecision="1" longitude="73.49953" name="Kasara-Ghat near Igatpuri" pageId="0" pageNumber="4606">Kasara-Ghat near Igatpuri</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="4606">365 m</elevation>
; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="19.683916">19°41'02.1&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="east" orientation="longitude" precision="1" value="73.49953">73°29'58.3&quot;E</geoCoordinate>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="4606" value="2012-11-10">10-11-2012</collectingDate>
; fieldNumber: RD Gore 1042; fieldNotes: Twining herbs; leaves stipulate; stipules submedifixed; chasmogamous flowers yellow &amp; Cleistogamous flowers white; pods falcate to straight; seeds well developed; Record Level: language: English; institutionID: Botanical Survey of India, Calcutta (CAL).
</materialsCitation>
<materialsCitation collectingDate="2012-11-10" country="India" latitude="19.683916" location="Kasara-Ghat near Igatpuri" longLatPrecision="1" longitude="73.49953" pageId="0" pageNumber="4606" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="4606">Other material</typeStatus>
. Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="4606">India</collectingCountry>
; countryCode: IND; stateProvince: Maharashtra; municipality: Nasik District; locality:
<location LSID="urn:lsid:plazi:treatment:E810301B37B5573346755260FD7CD26E:3DDD3CEAE728FE8226AFE88689527A5C" country="India" latitude="19.683916" longLatPrecision="1" longitude="73.49953" name="Kasara-Ghat near Igatpuri" pageId="0" pageNumber="4606">Kasara-Ghat near Igatpuri</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="4606">365 m</elevation>
; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="19.683916">19°41'02.1&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="east" orientation="longitude" precision="1" value="73.49953">73°29'58.3&quot;E</geoCoordinate>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="4606" value="2012-11-10">10-11-2012</collectingDate>
; fieldNumber: RD Gore 1042a; fieldNotes: Twining herbs; leaves stipulate; stipules submedifixed; chasmogamous flowers yellow &amp; Cleistogamous flowers white; pods falcate to straight; seeds well developed; Record Level: language: English; institutionID: Botanical Survey of India, Pune (BSI).
</materialsCitation>
<materialsCitation collectingDate="2012-11-10" country="India" location="Kasara-Ghat near Igatpuri" pageId="0" pageNumber="4606" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="4606">Other material</typeStatus>
. Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="4606">India</collectingCountry>
; countryCode: IND; stateProvince: Maharashtra; municipality: Nasik District; locality:
<location LSID="urn:lsid:plazi:treatment:E810301B37B5573346755260FD7CD26E:98D8D53DA08E66C4C23C3D91ECF38417" country="India" name="Kasara-Ghat near Igatpuri" pageId="0" pageNumber="4606">Kasara-Ghat near Igatpuri</location>
; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="4606">SP Gaikwad &amp; RD Gore</determinerName>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="4606" value="2012-11-10">10-11-2012</collectingDate>
; fieldNumber: SD Randive 322; fieldNotes: Twining herbs; flowers yellow; pods slightly hairy; Record Level: institutionID: Herbarium of Walchand College of Arts &amp; Science, Solapur (WCAS).
</materialsCitation>
<materialsCitation collectingDate="2012-11-09" country="India" location="Saptshrungi hills (Kalvan tehsil)" pageId="0" pageNumber="4606" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="4606">Other material</typeStatus>
. Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="4606">India</collectingCountry>
; countryCode: IND; stateProvince: Maharashtra; municipality: Nasik District; locality:
<location LSID="urn:lsid:plazi:treatment:E810301B37B5573346755260FD7CD26E:82A05452C061A961BDFE9EA28D798A4D" country="India" name="Saptshrungi hills (Kalvan tehsil)" pageId="0" pageNumber="4606">Saptshrungi hills (Kalvan tehsil)</location>
; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="4606">SP Gaikwad &amp; RD Gore</determinerName>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="4606" value="2012-11-09">9-11-2012</collectingDate>
; fieldNumber: RD Gore 1040; fieldNotes: Twining herbs; flowers both chasmogamous (yellow) and cleistogamous (white/albino); Record Level: institutionID: Herbarium of Walchand College of Arts &amp; Science, Solapur (WCAS).
</materialsCitation>
<materialsCitation collectingDate="2011-10-21" collectionCode="K 000978011" country="India" location="Pasarnighat" pageId="0" pageNumber="4606" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="4606">Other material</typeStatus>
. Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="4606">India</collectingCountry>
; countryCode: IND; stateProvince: Maharashtra; municipality: Satara District; locality:
<location LSID="urn:lsid:plazi:treatment:E810301B37B5573346755260FD7CD26E:101E88DB465AB0F0235A42E29D211316" country="India" name="Pasarnighat" pageId="0" pageNumber="4606">Pasarnighat</location>
; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="4606">MM Aitawade</determinerName>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="4606" value="2011-10-21">21-10-2011</collectingDate>
; fieldNumber: SP Sutar 156; fieldNotes: Closely resembles with Vignasilvestris but differs in pod &amp; seed number; seeds rectangular; Record Level: institutionID: Kew herbarium (K); collectionCode:
<collectionCode pageId="0" pageNumber="4606">K000978011</collectionCode>
; source: http://apps.kew.org/herbcat/getImage.do?imageBarcode=K000978011
</materialsCitation>
<materialsCitation collectingDate="2011-10-21" country="India" location="Pasarnighat" pageId="0" pageNumber="4606" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="4606">Other material</typeStatus>
. Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="4606">India</collectingCountry>
; countryCode: IND; stateProvince: Maharashtra; municipality: Satara District; locality:
<location LSID="urn:lsid:plazi:treatment:E810301B37B5573346755260FD7CD26E:69E76905AF7245C614B14227B7762928" country="India" name="Pasarnighat" pageId="0" pageNumber="4606">Pasarnighat</location>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="4606" value="2011-10-21">21-10-2011</collectingDate>
; fieldNumber: SP Sutar 156; fieldNotes: Closely resembles with Vignasilvestris but differs in pod &amp; seed number; seeds rectangular; Record Level: institutionID: Herbarium of Shivaji University, Kolhapur (SUK).
</materialsCitation>
<materialsCitation collectingDate="1989-09-28" country="India" location="Dandoba hills" pageId="0" pageNumber="4606" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="4606">Other material</typeStatus>
. Location: continent: Asia; country:
<collectingCountry pageId="0" pageNumber="4606">India</collectingCountry>
; countryCode: IND; stateProvince: Maharashtra; municipality: Sangli; locality:
<location LSID="urn:lsid:plazi:treatment:E810301B37B5573346755260FD7CD26E:2EB24F9E99AF4B7AD76A8A96C3ADF8CC" country="India" name="Dandoba hills" pageId="0" pageNumber="4606">Dandoba hills</location>
; Event: eventDate:
<collectingDate pageId="0" pageNumber="4606" value="1989-09-28">28-09-1989</collectingDate>
; fieldNumber: AN Londhe 170037; Record Level: institutionID: Botanical Survey of India, Pune (BSI).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4606" type="description">
<paragraph pageId="0" pageNumber="4606">Description</paragraph>
<paragraph pageId="0" pageNumber="4606">
Twining annual herbs. Stems terete, up to 2 m long, covered with 1-3 mm long, retrorse or spreading bulbous based hairs. Leaves 3-foliolate; petioles 3-5(-7) cm long, densely covered with retrorse or spreading bulbous based hairs; stipules elliptically lanceolate, medifixed, 7-9 mm long, 5-7 nerved, rounded at base, acute at apex, hairy on dorsal surface; stipels 2, linear-lanceolate, 2-3 mm long, acute or acuminate at apex, glabrous. Leaflets membranous, entire; lateral ones obliquely ovate or rhomboid to lanceolate, 2.2-5.5 x 1.5-2.3 cm, obliquely rounded at base, acute or shortly acuminate at apex, sparsely hairy; terminal leaflet ovate or rhomboid-lanceolate, 3-6.4 x 1.2-4 cm, base rounded (not oblique as in lateral leaflets), apex acute or shortly acuminate, sparsely hairy, margins entire or sometimes wavy; rachis 2-10 mm long, densely covered with whitish hairs. Flowers are of two kinds; the chasmogamous flowers present on leafy aerial shoots while cleistogamous flowers present on leafless subterranean shoots, which are close to soil surface. Chasmogamous flowers: yellow, 2-4 in axillary and terminal pseudo-racemes; peduncle 5-7.5 cm long, densely hairy with whitish-brown, 1-1.5 mm long, retrosely spreading hairs; pedicels 2-2.5 mm long, densely hairy as peduncle; bracts lanceolate, 2.5-3 x 0.8-1 mm, herbaceous, acute at apex, densely hairy; bracteoles 2, linear, 4-4.5 mm long, densely hairy. Calyx campanulate, ca 2 mm long; teeth triangular, 0.2-1.2 x 0.7-1 mm, sparsely hairy along margins. Standard petal yellow, asymmetrical, broadly ovate, 6-6.5 x 7.5-8.5 mm, emarginate at apex, central protuberance present inside (up to 0.9 mm long); claw ca 1 mm long. Right wing concealing the upper portion of the keel petals; claws 0.7-1 mm long; lamina obliquely obovate, 3.5-6.5 x 2.7-3.5 mm, notched at apex. Left wing claws 1-1.2 mm long; lamina obliquely obovate, 4-6.2 x 1.2-4 mm. Keel petals spirally incurved to left, 6-6.5 x 2.5-3.5 mm; horn-like pocket present (1.5-1.7 mm long) on the left side of keel petal. Stamens 9+1, included; staminal tube 4.5-5 mm long; free filament 8-9 mm long; anthers dorsifixed, 0.2-0.3 mm long. Pistil 1.5-1.8 cm long; ovary linear, 2.2-2.4 mm long, densely covered with long whitish hairs; style filiform, 1.2-1.3 cm long,
<normalizedToken originalValue="‘S’">'S'</normalizedToken>
shaped before stigma, beaked beyond stigma; beak 0.8-1 mm long, upwardly curved at apex. Pods ascending linear, cylindrical, 2-5 x 0.3-0.5 cm, straight or slightly curved, densely covered with brownish, 2-3 mm long spreading hairs, acute at apex. Seeds 4-10 per pod, broadly rectangular, 3-3.1 x 2-2.1 mm, brown, mottled with faint black patches, rounded or rectangular at both ends; seed coat porous with mesh-like reticulation; hilum well developed, rim-aril protruded, elliptic, 1.8-2 x 0.7-0.9 mm, whitish-brown. Germination epigeal; ecophylls petioles, simple and minutely pulvinous. Cleistogamous flowers: 2-5, whitish-yellow on leafless sub-aerial branches which produce on lower nodes of stem; pedicels 1-2 mm long; bracts lanceolate, 5-6 mm long, acute at apex, densely hairy; bracteoles 2, linear, 7-9 mm long, densely covered with 1-3 mm long bulbous based hairs. Calyx as in chasmogamous flowers. Standard petal symmetrical, broadly ovate, 4.5-5 x 5.5-6 mm, emarginate at apex, inside central protuberance absent; claws 0.3-0.5 mm long. Right wing petal slightly concealing the upper portion of the keel petal; claws 0.3-0.5 mm long; lamina obliquely ovate, 4.5-5 x 2-3 mm; auricle 0.1-0.2 mm long. Left wing petal spreading; claws 0.5-0.7 mm long; lamina ovate-elliptic, 4.9-5 x 2-2.7 mm, base oblique; auricle 0.3-0.5 mm long. Keel petals slightly curved, 4.8-5 x 1.5-2.5 mm. Stamens 9+1, dimorphic, out of 9 stamens in a bundle 4 short and 5 long; short stamens sterile with ca 1 mm long filaments, whereas long stamens fertile with ca 2 mm long filaments; staminal tube ca 1.5 mm long; free stamen sterile with ca 2 mm long filament; anthers dorsifixed 0.2-0.6 mm long. Pistil filiform, up to 6.5 mm long; ovary linear, 1.7-2 mm long, glabrescent or sparsely hairy; style filiform, 3-3.5 mm long, curved (not
<normalizedToken originalValue="‘S’">'S'</normalizedToken>
shaped), densely hairy before stigma with 0.4-0.6 mm long hairs, beaked beyond stigma; beak short, 0.3-0.5 mm long, straight, pointed at apex. Pods linear, cylindrical, straight, 2.5-4 x 0.3-0.4 cm, narrowed at both ends, sparsely hairy with short hairs (0.5 mm long), white hairs. Seeds 8-9 per pod, obliquely rounded, 3.5-4 x 3-3.5 mm, dark brown; seed coat foveolatewith mesh-like reticulation; hilum well developed, protruded, broadly elliptic, 2-2.2 x 0.4-0.7 mm, whitish. Germination epigeal; ecophylls petioles, simple and minutely pulvinous. (Figs 1, 2, 7).
</paragraph>
<subSection pageId="0" pageNumber="4606" type="flowering and fruiting">
<paragraph pageId="0" pageNumber="4606">Flowering and fruiting</paragraph>
<paragraph pageId="0" pageNumber="4606">August-October.</paragraph>
</subSection>
</subSubSection>
<subSubSection pageId="0" pageNumber="4606" type="diagnosis">
<paragraph pageId="0" pageNumber="4606">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="4606">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Vigna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vigna yadavii" order="Fabales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="species" species="yadavii">Vigna yadavii</taxonomicName>
similis, ramis dimorphis, floribus cleistogamis vexillo sine processo carina sine marsupio corniformi, styli rostro applanato, seminum testa foveolata, hilo bene evoluto differt.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4606" type="etymology">
<paragraph pageId="0" pageNumber="4606">Etymology</paragraph>
<paragraph pageId="0" pageNumber="4606">The specific epithet honors Prof AK Pandey, Department of Botany, Delhi University, New Delhi (India), in recognition of his valuable contribution to the taxonomy of flowering plants of India.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4606" type="distribution">
<paragraph pageId="0" pageNumber="4606">Distribution</paragraph>
<paragraph pageId="0" pageNumber="4606">India, Maharashtra, Satara district, near Chalkewadi in Patan tahsil.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4606" type="ecology">
<paragraph pageId="0" pageNumber="4606">Ecology</paragraph>
<paragraph pageId="0" pageNumber="4606">
It is a twining annual herb, grows on lateritic gravelly soil on hill slopes amonggrasses and herbs at about 1200 m altitude above mean sea level in Satara district of Maharashtra, India. The species has adapted to the monsoon seasonality. It thrives in humid climate with heavy rainfall during growth season. The seed germination takes place with onset of monsoon rain in the first week of June and the plant completes its life cycle with formation of seeds when rains ceasein mid October. The common associates of the species are
<taxonomicName class="Magnoliopsida" family="Acanthaceae" genus="Carvia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Carvia callosa" order="Lamiales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="species" species="callosa">Carvia callosa</taxonomicName>
(Nees) Bremek.,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Crotalaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Crotalaria nana" order="Fabales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="species" species="nana">Crotalaria nana</taxonomicName>
Burm. f.,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Crotalaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Crotalaria vestita" order="Fabales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="species" species="vestita">Crotalaria vestita</taxonomicName>
Baker,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Cajanus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cajanus lineatus" order="Fabales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="species" species="lineatus">Cajanus lineatus</taxonomicName>
(Wight &amp; Arn.) Maesen,
<taxonomicName class="Liliopsida" family="Poaceae" genus="Eragrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eragrostis" order="Poales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="genus">Eragrostis</taxonomicName>
spp.,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pseudarthria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudarthria" order="Fabales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="genus">Pseudarthria</taxonomicName>
spp.,
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Nogra" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nogra dalzellii" order="Fabales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="species" species="dalzellii">Nogra dalzellii</taxonomicName>
(Baker) Merr. and
<taxonomicName class="Liliopsida" family="Poaceae" genus="Themeda" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Themeda" order="Poales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="genus">Themeda</taxonomicName>
spp.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="4606" type="notes">
<paragraph pageId="0" pageNumber="4606">Notes</paragraph>
<paragraph pageId="0" pageNumber="4606">
Interestingly, two types of shoots are observed in the species, one a normal aerial leafy shoot and the other subterranean (close to soil surface) leafless shoot producedatthe lower nodes of the stem. The later shoots produce cleistogamous flowers, which remain closed. They show differences in the structure of their floral parts as compared to chasmogamous flowers such as standard petal without central protuberance inside, keel petals without horn-like pocket, curved style and short style beak (0.3-0.5 mm long). (Figs 3, 4, 5, 6). The floral parts of chasmogamous flowers roughly correspond to the floral parts of the species of
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Vigna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vigna" order="Fabales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="genus">Vigna</taxonomicName>
sect.
<taxonomicName genus="Ceratotropis" lsidName="Ceratotropis" pageId="0" pageNumber="4606" rank="section" section="Ceratotropis">Ceratotropis</taxonomicName>
while cleistogamous floral parts correspond to the species of
<taxonomicName genus="V." lsidName="V." pageId="0" pageNumber="4606" rank="genus">V.</taxonomicName>
sect.
<taxonomicName genus="Aconitifoliae" lsidName="Aconitifoliae" pageId="0" pageNumber="4606" rank="section" section="Aconitifoliae">Aconitifoliae</taxonomicName>
of the same subgenus. Thus, it shows a combination of characters of species of both sections. The new species shows morphological similarities with
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Vigna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vigna yadavii" order="Fabales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="species" species="yadavii">Vigna yadavii</taxonomicName>
Gaikwad et al. and
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Vigna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vigna dalzelliana" order="Fabales" pageId="0" pageNumber="4606" phylum="Tracheophyta" rank="species" species="dalzelliana">Vigna dalzelliana</taxonomicName>
(Kuntze) Verdc. but differs in its dimorphic branches, foveolate seed coat and absence of standard protuberance and horn-like keel pocket in cleistogamous flowers. A comparative account between above mentioned three species is given in Table 1.
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95
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<bibCitation author="Linnaeus" pagination="435" title="Oxalis stricta" volumeTitle="Species Plantarum 1" year="1753">
<taxonomicName ID-CoL="7NJ5M" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis bold="true" pageNumber="717">Oxalis stricta</emphasis>
Linnaeus
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1
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<paragraph pageNumber="717">&quot;Habitat in Virginia.&quot; RCN: 3381.</paragraph>
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<typeStatus>
<typeStatus>Lectotype</typeStatus>
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(Eiten in
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4: 103. 1955): [icon]
<emphasis italics="true" pageNumber="717">&quot;Oxys s. trifolium luteum corniculatum, virginianum rectum majus&quot;</emphasis>
in Morison, Pl. Hist. Univ. 2: 184, s. 2, t. 17, f. 3. 1680.
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<paragraph pageNumber="717">
Current name:
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<emphasis italics="true" pageNumber="717">
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L.
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(
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<emphasis italics="true" pageNumber="717">Note:</emphasis>
Watson (in
<emphasis italics="true" pageNumber="717">Bot. J. Linn. Soc.</emphasis>
101: 358-361. 1989) gives a detailed review, and reproduces the type illustration (as f. 3).
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<emphasis id="52CDCC71FA70282FFF6EAD32FE9EFDBF" bold="true" box="[151,340,583,609]" italics="true" pageId="7" pageNumber="268">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA70282FFE9CAD32FDE8FDBF" bold="true" box="[357,546,583,609]" italics="true" pageId="7" pageNumber="268">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA70282FFDCAAD32FD66FDBF" bold="true" box="[563,684,583,609]" italics="true" pageId="7" pageNumber="268">inaxeyeus</emphasis>
Sasa, Katami &amp; Suzuki
</emphasis>
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(
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<emphasis id="52CDCC71FA70282FFF6EADC7FEFCFD16" box="[151,310,690,712]" italics="true" pageId="7" pageNumber="268">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA70282FFEBDADC7FE29FD16" box="[324,483,690,712]" italics="true" pageId="7" pageNumber="268">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA70282FFE0BADC6FD9CFD16" box="[498,598,691,712]" italics="true" pageId="7" pageNumber="268">inaxeyeus</emphasis>
Sasa, Katami &amp;
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<emphasis id="52CDCC71FA70282FFF6EAD80FE4BFCD0" bold="true" box="[151,385,757,782]" pageId="7" pageNumber="268">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA70282FFE70AD80FE3EFCD0" box="[393,500,757,782]" pageId="7" pageNumber="268" type="holotype">Holotype</typeStatus>
, male, on the shore of Lake Inawashiro, The Dr.Noguchi Memorial Hall,
<collectingCountry id="18AE50F3FA70282FFAC1AD83FAB1FCD3" box="[1336,1403,758,781]" name="Japan" pageId="7" pageNumber="268">Japan</collectingCountry>
.
<date id="140736A3FA70282FFA7EAD80FEDAFCEC" pageId="7" pageNumber="268" value="2000-08-05">5. VIII. 2000</date>
, light trap (No.401: 79).
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<emphasis id="52CDCC71FA70282FFF3EAC48FE1BFC88" bold="true" box="[199,465,829,854]" pageId="7" pageNumber="268">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by superior volsella developed and with rounded margin, inferior volsella a little hooked and like nose-shaped, with many setae; anal point with 2 long lateral setae on each side, transverse sternapodeme without oral projections.
</paragraph>
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<emphasis id="52CDCC71FA70282FFF6EA8F6FE84F846" bold="true" box="[151,334,1923,1945]" pageId="7" pageNumber="268">FIGURE 7 A–B.</emphasis>
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(
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)
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Sasa, Kitami &amp; Suzuki
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, Male imago.
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Hypopygium, dorsal view.
<emphasis id="52CDCC71FA70282FFF2AA8D7FF20F869" bold="true" box="[211,234,1954,1975]" pageId="7" pageNumber="268">B.</emphasis>
Hypopygium, ventral view.
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<emphasis id="52CDCC71FA7F2820FF3EAFE2FD55FF6E" bold="true" box="[199,671,151,176]" pageId="8" pageNumber="269">Additional description and corrections.</emphasis>
AR should be 0.50 (AR
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original description). Based on the analysis of
<typeStatus id="BF02AEC1FA7F2820FEE7AFC9FE4AFF0B" box="[286,384,188,213]" pageId="8" pageNumber="269" type="holotype">holotype</typeStatus>
material, the examined characters were consistent with the original description, but some additional characters should be added: Total length / wing length 2.2. Wing length / length of profemur 2.6. Tentorium 135 µm long, 23 µm wide. Dorsocentrals 11, scutellars 6. Squama with 6 setae. Palpomeres length (in µm): 35, 38, 78, 98, 148. Palpomere 2 ellipsoid, 3 and 4 rectangular, 5 long and slender, Palpomeres 5/3: 1.9. Laterosternite IX with 8 long setae. Anal point 25 µm long, basal wide 20 µm, peak wide 5 µm, with 2 long lateral setae on each side. Transverse sternapodeme 60 µm long, without oral projections. Gonocoxite 153 µm long. Gonostylus 65 µm long, with triangularly pointed costa dorsalis. Megaseta 13 µm long. HR 2.3, HV 3.8. The hypopygium of
<typeStatus id="BF02AEC1FA7F2820FEB3AEC2FE66FE0E" box="[330,428,439,464]" pageId="8" pageNumber="269" type="holotype">holotype</typeStatus>
are re-drawn as in
<figureCitation id="F8820CE6FA7F2820FD7AAEC2FCCDFE0E" box="[643,775,439,464]" captionStart="FIGURE 7 A – B" captionStartId="7.[151,249,1923,1945]" captionTargetBox="[237,1358,987,1901]" captionTargetId="figure@7.[226,1360,978,1901]" captionTargetPageId="7" captionText="FIGURE 7 A – B. Rhecricotopus (Rheocricotopus) inaxeyeus Sasa, Kitami &amp; Suzuki, Male imago. A. Hypopygium, dorsal view. B. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255143/files/figure.png" pageId="8" pageNumber="269">Figs 7 A–B</figureCitation>
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<emphasis id="52CDCC71FA7F2820FF3EAEA9FEAAFE2B" bold="true" box="[199,352,476,501]" pageId="8" pageNumber="269">Distribution.</emphasis>
The species occurs in
<collectingCountry id="18AE50F3FA7F2820FDA7AEA8FD54FE2A" box="[606,670,477,500]" name="Japan" pageId="8" pageNumber="269">Japan</collectingCountry>
(Palaearctic Region).
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165
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<mods:title id="A399194EBD186E456035415B7E387ED3">Redescription of 13 holotypes of Rheocricotopus Brundin, 1956 (Diptera: Chironomidae) from the Sino – Indian Region</mods:title>
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<emphasis id="52CDCC71FA71282EFF6EAD78FE9EFDF9" bold="true" box="[151,340,525,551]" italics="true" pageId="6" pageNumber="267">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA71282EFE9CAD78FDE8FDF9" bold="true" box="[357,546,525,551]" italics="true" pageId="6" pageNumber="267">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA71282EFDCAAD78FD70FDF9" bold="true" box="[563,698,525,551]" italics="true" pageId="6" pageNumber="267">inaquereus</emphasis>
Sasa, Katami &amp; Suzuki
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<paragraph id="60061063FA71282EFF6EAD5AFEE1FD96" blockId="6.[151,996,525,584]" box="[151,299,559,584]" pageId="6" pageNumber="267">
(
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)
</paragraph>
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<emphasis id="52CDCC71FA71282EFF6EAD07FEFCFD56" box="[151,310,626,648]" italics="true" pageId="6" pageNumber="267">Rheocricotopus</emphasis>
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(R
<emphasis id="52CDCC71FA71282EFEACAD07FE2EFD56" box="[341,484,626,648]" italics="true" pageId="6" pageNumber="267">heocricotopus</emphasis>
)
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<emphasis id="52CDCC71FA71282EFE0DAD06FDAEFD56" box="[500,612,627,648]" italics="true" pageId="6" pageNumber="267">inaquereus</emphasis>
Sasa, Katami &amp;
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<bibRefCitation id="04286D92FA71282EFCF6AD07FC52FD56" author="Suzuki" box="[783,920,626,648]" pageId="6" pageNumber="267" refString="Sasa, M., Kitami, K. &amp; Suzuki, H. (2001) Additional studies on the chironomid midges collected on the shore of Lake Inawashiro. Research Report of the Dr. Noguchi Memorial Hall, 38 pp." type="book" year="2001">Suzuki, 2001</bibRefCitation>
: 19
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;
<treatmentCitation id="E1183672FA71282EFC33AD07FAC8FD56" author="Ashe" box="[970,1282,626,648]" page="571" pageId="6" pageNumber="267" year="2012">
<bibRefCitation id="04286D92FA71282EFC33AD07FB07FD56" author="Ashe" box="[970,1229,626,648]" pageId="6" pageNumber="267" refString="Ashe, P. &amp; O'Connor, J. P. (2012) A world catalogue of Chironomidae (Diptera). Part 2 B. Orthocladiinae. Irish Biogeographical Society &amp; National Museum of Ireland, Dublin, pp. 558 - 575." type="book chapter" year="2012">Ashe &amp; O’Connor 2012</bibRefCitation>
: 571
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.
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<paragraph id="60061063FA71282EFF6EADDAFDD2FD35" blockId="6.[151,1437,687,851]" pageId="6" pageNumber="267">
<emphasis id="52CDCC71FA71282EFF6EADDAFE4BFD16" bold="true" box="[151,385,687,712]" pageId="6" pageNumber="267">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA71282EFE70ADDAFE3EFD16" box="[393,500,687,712]" pageId="6" pageNumber="267" type="holotype">Holotype</typeStatus>
, male, on the shore of Lake Inawashiro, The Dr.Noguchi Memorial Hall,
<collectingCountry id="18AE50F3FA71282EFAC1ADC5FAB1FD19" box="[1336,1403,688,711]" name="Japan" pageId="6" pageNumber="267">Japan</collectingCountry>
.
<date id="140736A3FA71282EFA7EADC5FF37FD35" pageId="6" pageNumber="267" value="1999-09-04">4. IX. 1999</date>
, sweeping (No. 397: 10).
</paragraph>
<paragraph id="60061063FA71282EFF3EAD80FDF1FC8D" blockId="6.[151,1437,687,851]" pageId="6" pageNumber="267">
<emphasis id="52CDCC71FA71282EFF3EAD80FE1BFCD0" bold="true" box="[199,465,757,782]" pageId="6" pageNumber="267">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by superior volsella developed and with almost right-angle corner, inferior volsella extended and double-layered, Anal point with 3 long lateral setae on each side.
</paragraph>
<caption id="34C640EBFA71282EFF6EA8C3FD86F831" httpUri="https://zenodo.org/record/255142/files/figure.png" pageId="6" pageNumber="267" targetBox="[236,1352,896,1958]" targetPageId="6">
<paragraph id="60061063FA71282EFF6EA8C3FD86F831" blockId="6.[151,1436,1974,2031]" pageId="6" pageNumber="267">
<emphasis id="52CDCC71FA71282EFF6EA8C3FE80F812" bold="true" box="[151,330,1974,1996]" pageId="6" pageNumber="267">FIGURE 6 A–C.</emphasis>
<taxonomicName id="A7B96BE0FA71282EFEA9A8C3FC35F812" authority="Sasa, Kitami &amp; Suzuki" authorityName="Sasa, Kitami &amp; Suzuki" box="[336,1023,1974,1997]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="267" phylum="Arthropoda" rank="species" species="inaquereus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA71282EFEA9A8C3FE28F812" box="[336,482,1974,1996]" italics="true" pageId="6" pageNumber="267">Rhecricotopus</emphasis>
(
<emphasis id="52CDCC71FA71282EFE08A8C3FD5AF812" box="[497,656,1974,1996]" italics="true" pageId="6" pageNumber="267">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA71282EFD66A8CDFCC5F813" box="[671,783,1976,1997]" italics="true" pageId="6" pageNumber="267">inaquereus</emphasis>
Sasa, Kitami &amp; Suzuki
</taxonomicName>
, Male imago.
<emphasis id="52CDCC71FA71282EFB76A8C3FB62F815" bold="true" box="[1167,1192,1974,1995]" pageId="6" pageNumber="267">A.</emphasis>
Wing.
<emphasis id="52CDCC71FA71282EFB0BA8C2FAC3F812" bold="true" box="[1266,1289,1975,1996]" pageId="6" pageNumber="267">B.</emphasis>
Hypopygium, dorsal view.
<emphasis id="52CDCC71FA71282EFEEEA8ACFEFAF831" bold="true" box="[279,304,2009,2031]" pageId="6" pageNumber="267">C.</emphasis>
Hypopygium, ventral view.
</paragraph>
</caption>
<paragraph id="60061063FA70282FFF3EAFE2FF19FE0E" blockId="7.[151,1437,151,501]" pageId="7" pageNumber="268">
<emphasis id="52CDCC71FA70282FFF3EAFE2FD51FF6E" bold="true" box="[199,667,151,176]" pageId="7" pageNumber="268">Additional description and corrections.</emphasis>
Based on the analysis of
<typeStatus id="BF02AEC1FA70282FFC43AFE2FBD6FF6E" box="[954,1052,151,176]" pageId="7" pageNumber="268" type="holotype">holotype</typeStatus>
material, the examined characters were consistent with the original description, but some additional characters should be added: Total length / wing length 2.0. Wing length / length of profemur 2.5. Tentorium 140 µm long, 28 µm wide. Dorsocentrals 11, scutellars 6. Squama with 10 setae. Palpomeres length (in µm): 35, 50, 85, 110, 175. Palpomere 2 ellipsoid, 3 and 4 rectangular, 5 long and slender, Palpomeres 5/3: 2.1. Laterosternite IX with 7 long setae. Anal point 38 µm long, basal wide 18 µm, peak wide 5 µm, with 3 long lateral setae on each side. Transverse sternapodeme 75 µm long, with weak oral projections. Gonocoxite 175 µm long. Gonostylus 80 µm long, with small triangularly costa dorsalis. Megaseta 13 µm long. HR 2.2, HV 3.3. The wing and hypopygium of
<typeStatus id="BF02AEC1FA70282FFBE3AEE1FBB6FE73" box="[1050,1148,404,429]" pageId="7" pageNumber="268" type="holotype">holotype</typeStatus>
are re-drawn as in
<figureCitation id="F8820CE6FA70282FFAAFAEE1FF05FE0E" captionStart="FIGURE 6 A – C" captionStartId="6.[151,249,1974,1996]" captionTargetBox="[236,1352,896,1958]" captionTargetId="figure@6.[226,1360,886,1959]" captionTargetPageId="6" captionText="FIGURE 6 A – C. Rhecricotopus (Psilocricotopus) inaquereus Sasa, Kitami &amp; Suzuki, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255142/files/figure.png" pageId="7" pageNumber="268">Figs 6 A–C</figureCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="60061063FA70282FFF3EAEA9FC58FE2B" blockId="7.[151,1437,151,501]" box="[199,914,476,501]" pageId="7" pageNumber="268">
<emphasis id="52CDCC71FA70282FFF3EAEA9FEAAFE2B" bold="true" box="[199,352,476,501]" pageId="7" pageNumber="268">Distribution.</emphasis>
The species occurs in
<collectingCountry id="18AE50F3FA70282FFDA7AEA8FD54FE2A" box="[606,670,477,500]" name="Japan" pageId="7" pageNumber="268">Japan</collectingCountry>
(Palaearctic Region).
</paragraph>
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</treatment>
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164
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<mods:title id="0D81A24EE02AE211A1D2A08DC53E9BA9">Redescription of 13 holotypes of Rheocricotopus Brundin, 1956 (Diptera: Chironomidae) from the Sino – Indian Region</mods:title>
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<emphasis id="52CDCC71FA72282DFF6EAEC2FE9EFE0F" bold="true" box="[151,340,439,465]" italics="true" pageId="5" pageNumber="266">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA72282DFE9CAEC2FDD4FE0F" bold="true" box="[357,542,439,465]" italics="true" pageId="5" pageNumber="266">Psilocricotopus</emphasis>
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Sasa &amp; Suzuki
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(
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)
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<emphasis id="52CDCC71FA72282DFF6EAD57FEFCFDE6" box="[151,310,546,568]" italics="true" pageId="5" pageNumber="266">Rheocricotopus</emphasis>
(
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<emphasis id="52CDCC71FA72282DFE0BAD57FDBEFDE6" box="[498,628,546,568]" italics="true" pageId="5" pageNumber="266">tokarakeleus</emphasis>
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: 270
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;
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<emphasis id="52CDCC71FA72282DFF6EA9B0FE80F905" bold="true" box="[151,330,1733,1755]" pageId="5" pageNumber="266">FIGURE 5 A–C.</emphasis>
<taxonomicName id="A7B96BE0FA72282DFEB6A9B0FC76F905" authority="Sasa &amp; Suzuki" authorityName="Sasa &amp; Suzuki" box="[335,956,1733,1755]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="266" phylum="Arthropoda" rank="species" species="tokarakeleus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA72282DFEB6A9B0FE2BF905" box="[335,481,1733,1755]" italics="true" pageId="5" pageNumber="266">Rhecricotopus</emphasis>
(
<emphasis id="52CDCC71FA72282DFE09A9B0FD45F905" box="[496,655,1733,1755]" italics="true" pageId="5" pageNumber="266">Psilocricotopus</emphasis>
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Sasa &amp; Suzuki
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, Male imago.
<emphasis id="52CDCC71FA72282DFBB5A9B0FBAFF904" bold="true" box="[1100,1125,1733,1754]" pageId="5" pageNumber="266">A.</emphasis>
Wing.
<emphasis id="52CDCC71FA72282DFB57A9B0FB0FF904" bold="true" box="[1198,1221,1733,1754]" pageId="5" pageNumber="266">B.</emphasis>
Hypopygium, dorsal view.
<emphasis id="52CDCC71FA72282DFF2AA991FF21F924" bold="true" box="[211,235,1764,1786]" pageId="5" pageNumber="266">C.</emphasis>
Hypopygium, ventral view.
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<emphasis id="52CDCC71FA72282DFF6EA84AFE4AF886" bold="true" box="[151,384,1855,1880]" pageId="5" pageNumber="266">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA72282DFE7FA84AFE3BF886" box="[390,497,1855,1880]" pageId="5" pageNumber="266" type="holotype">Holotype</typeStatus>
, male, Kuchinoshima, Kokara, Kagoshima Prefecture,
<collectingCountry id="18AE50F3FA72282DFBAEA835FB50F889" box="[1111,1178,1856,1879]" name="Japan" pageId="5" pageNumber="266">Japan</collectingCountry>
.
<date id="140736A3FA72282DFB5DA835FAE2F886" box="[1188,1320,1856,1880]" pageId="5" pageNumber="266" value="1994-05-19">19. V. 1994</date>
. (No. 287: 37).
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<paragraph id="60061063FA72282DFF3EA8F2FB54F836" blockId="5.[151,1437,1855,2024]" pageId="5" pageNumber="266">
<emphasis id="52CDCC71FA72282DFF3EA8F2FE1DF87E" bold="true" box="[199,471,1927,1952]" pageId="5" pageNumber="266">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by laterosternite IX with 7 long setae, anal point with 5 long lateral setae on each side, superior volsella with rounded corner, inferior volsella slightly hooked, gonostylus with a triangular pointed crista dorsalis.
</paragraph>
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<subSubSection id="28A343E8FA71282EFF3EAFE2FAA4FE7D" pageId="6" pageNumber="267" type="description">
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<emphasis id="52CDCC71FA71282EFF3EAFE2FD67FF6E" bold="true" box="[199,685,151,176]" pageId="6" pageNumber="267">Additional description and corrections.</emphasis>
Based on the examination of
<typeStatus id="BF02AEC1FA71282EFBE6AFE2FB4BFF6E" box="[1055,1153,151,176]" pageId="6" pageNumber="267" type="holotype">holotype</typeStatus>
material, we found the character consistent with the original description. But some additional characters should be added: Total length / wing length 1.9. Wing length / length of profemur 2.4. Tentorium 133 µm long, 30 µm wide. Brachilous 1. Dorsocentrals 15, scutellars 10. Palpomeres length (in µm): 35, 50, 120, 158, 210. Palpomere 2 ellipsoid, 3 rectangular, 4 and 5 long and slender, Palpomeres 5/3: 1.8. Laterosternite IX with 7 long setae. Anal point 40 µm long, basal wide 15 µm, peak wide 3 µm, with 5 long lateral setae on each side. Transverse sternapodeme 100 µm long, with oral projections. Gonocoxite 220 µm long. Gonostylus 100 µm long, with triangular crista dorsalis. Megaseta 13 µm long. HR 2.2, HV 3.7. The wing and hypopygium of
<typeStatus id="BF02AEC1FA71282EFC55AEFFFBC4FE7D" box="[940,1038,394,419]" pageId="6" pageNumber="267" type="holotype">holotype</typeStatus>
are re-drawn as in
<figureCitation id="F8820CE6FA71282EFB1CAEFFFAA0FE7D" box="[1253,1386,394,419]" captionStart="FIGURE 5 A – C" captionStartId="5.[151,249,1733,1755]" captionTargetBox="[235,1360,630,1711]" captionTargetId="figure@5.[226,1360,621,1712]" captionTargetPageId="5" captionText="FIGURE 5 A – C. Rhecricotopus (Psilocricotopus) tokarakeleus Sasa &amp; Suzuki, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255141/files/figure.png" pageId="6" pageNumber="267">Figs 5 A–C</figureCitation>
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<emphasis id="52CDCC71FA71282EFF3EAED8FEAAFE18" bold="true" box="[199,352,429,454]" pageId="6" pageNumber="267">Distribution.</emphasis>
The species occurs in
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(Palaearctic Region).
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<emphasis id="52CDCC71FA73282CFF6EAD89FE9EFCC8" bold="true" box="[151,340,764,790]" italics="true" pageId="4" pageNumber="265">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA73282CFE9CAD89FDD4FCC8" bold="true" box="[357,542,764,790]" italics="true" pageId="4" pageNumber="265">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA73282CFDD6AD89FD33FCC8" bold="true" box="[559,761,764,790]" italics="true" pageId="4" pageNumber="265">tobatervicesimus</emphasis>
Kikuchi &amp; Sasa
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<paragraph id="60061063FA73282CFF6EAC6BFEE1FCE9" blockId="4.[151,959,764,823]" box="[151,299,798,823]" pageId="4" pageNumber="265">
(
<figureCitation id="F8820CE6FA73282CFF66AC6BFEE9FCE9" box="[159,291,798,823]" captionStart="FIGURE 4 A – B" captionStartId="4.[151,249,1680,1702]" captionTargetBox="[264,1323,952,1651]" captionTargetId="figure@4.[256,1331,945,1659]" captionTargetPageId="4" captionText="FIGURE 4 A – B. Rhecricotopus (Psilocricotopus) tobatervicesimus Kikuchi &amp; Sasa, Male imago. A. Hypopygium, dorsal view. B. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255140/files/figure.png" pageId="4" pageNumber="265">Figs 4 A–B</figureCitation>
)
</paragraph>
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<emphasis id="52CDCC71FA73282CFF6EAC13FEFCFCA2" box="[151,310,870,892]" italics="true" pageId="4" pageNumber="265">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA73282CFEBDAC13FE29FCA2" box="[324,483,870,892]" italics="true" pageId="4" pageNumber="265">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA73282CFE0BAC13FD54FCA2" box="[498,670,870,892]" italics="true" pageId="4" pageNumber="265">tobatervicesimus</emphasis>
<treatmentCitation id="E1183672FA73282CFD5DAC13FC70FCA2" author="Kikuchi" box="[676,954,870,892]" page="326" pageId="4" pageNumber="265" year="1990">
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: 326
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;
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: 567
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="34C640EBFA73282CFF6EA9E5FDCCF91B" httpUri="https://zenodo.org/record/255140/files/figure.png" pageId="4" pageNumber="265" targetBox="[264,1323,952,1651]" targetPageId="4">
<paragraph id="60061063FA73282CFF6EA9E5FDCCF91B" blockId="4.[151,1436,1680,1733]" pageId="4" pageNumber="265">
<emphasis id="52CDCC71FA73282CFF6EA9E5FE85F97B" bold="true" box="[151,335,1680,1702]" pageId="4" pageNumber="265">FIGURE 4 A–B.</emphasis>
<taxonomicName id="A7B96BE0FA73282CFEA1A9E5FBC6F978" authority="Kikuchi &amp; Sasa" authorityName="Kikuchi &amp; Sasa" box="[344,1036,1680,1702]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="tobatervicesimus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA73282CFEA1A9E5FE20F978" box="[344,490,1680,1702]" italics="true" pageId="4" pageNumber="265">Rhecricotopus</emphasis>
(
<emphasis id="52CDCC71FA73282CFE05A9E5FD51F978" box="[508,667,1680,1702]" italics="true" pageId="4" pageNumber="265">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA73282CFD54A9E5FC93F978" box="[685,857,1680,1702]" italics="true" pageId="4" pageNumber="265">tobatervicesimus</emphasis>
Kikuchi &amp; Sasa
</taxonomicName>
, Male imago.
<emphasis id="52CDCC71FA73282CFB5FA9E5FB75F97B" bold="true" box="[1190,1215,1680,1701]" pageId="4" pageNumber="265">A.</emphasis>
Hypopygium, dorsal view.
<emphasis id="52CDCC71FA73282CFF2AA9DAFF20F91A" bold="true" box="[211,234,1711,1732]" pageId="4" pageNumber="265">B.</emphasis>
Hypopygium, ventral view.
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<subSubSection id="28A343E8FA73282DFF6EA999FCC6FE9E" lastPageId="5" lastPageNumber="266" pageId="4" pageNumber="265" type="materials_examined">
<paragraph id="60061063FA73282CFF6EA999FEFCF8F6" blockId="4.[151,1437,1772,2013]" pageId="4" pageNumber="265">
<emphasis id="52CDCC71FA73282CFF6EA999FE4BF8DB" bold="true" box="[151,385,1772,1797]" pageId="4" pageNumber="265">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA73282CFE70A999FE3EF8DB" box="[393,500,1772,1797]" pageId="4" pageNumber="265" type="holotype">Holotype</typeStatus>
, male, At the side of Lake Toba, Sumatra,
<collectingCountry id="18AE50F3FA73282CFC24A999FB87F8DB" box="[989,1101,1772,1797]" name="Indonesia" pageId="4" pageNumber="265">Indonesia</collectingCountry>
,
<date id="140736A3FA73282CFBA0A998FB00F8DB" box="[1113,1226,1773,1797]" pageId="4" pageNumber="265" value="1988-01-01">1. I. 1988</date>
, Kikuchi and Sasa (No. 200: 22).
</paragraph>
<paragraph id="60061063FA73282CFF3EA841FDEFF84B" blockId="4.[151,1437,1772,2013]" pageId="4" pageNumber="265">
<emphasis id="52CDCC71FA73282CFF3EA841FE1BF893" bold="true" box="[199,465,1844,1869]" pageId="4" pageNumber="265">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by superior volsella reduced, inferior volsella developed and rounded margins, with many microtrichia, gonostylus with a triangularly rounded crista dorsalis.
</paragraph>
<paragraph id="60061063FA73282CFF3EA8D5FA57F802" blockId="4.[151,1437,1772,2013]" pageId="4" pageNumber="265">
<emphasis id="52CDCC71FA73282CFF3EA8D5FD51F867" bold="true" box="[199,667,1952,1977]" pageId="4" pageNumber="265">Additional description and corrections.</emphasis>
Based on the analysis of
<typeStatus id="BF02AEC1FA73282CFC43A8D5FBD6F867" box="[954,1052,1952,1977]" pageId="4" pageNumber="265" type="holotype">holotype</typeStatus>
material, the examined characters were consistent with the original description, but some additional characters should be added: Clypeus with 9 setae.
</paragraph>
<paragraph id="60061063FA72282DFF6EAFE2FCC6FE9E" blockId="5.[151,1437,151,357]" pageId="5" pageNumber="266">
Tentorium 128 µm long, 30 µm wide. Dorsocentrals 8. Wing of
<typeStatus id="BF02AEC1FA72282DFCA7AFE2FC0AFF6E" box="[862,960,151,176]" pageId="5" pageNumber="266" type="holotype">holotype</typeStatus>
destoryed. Palpomeres length (in µm): 33, 48, 75, 92, 125. Palpomere 2 ellipsoid, 3 and 4 rectangular, 5 long and slender, Palpomeres 5/3: 1.7. Anal point 43 µm long, basal wide 20 µm, peak wide 2 µm. Transverse sternapodeme 70 µm long, with weak oral projections. Gonocoxite 152 µm long. Gonostylus 75 µm long, with crista dorsalis. Megaseta 10 µm long. HR 2.0, HV 2.2. The hypopygium of
<typeStatus id="BF02AEC1FA72282DFEB3AE52FE66FE9E" box="[330,428,295,320]" pageId="5" pageNumber="266" type="holotype">holotype</typeStatus>
are re-drawn as in
<figureCitation id="F8820CE6FA72282DFD7AAE52FCCDFE9E" box="[643,775,295,320]" captionStart="FIGURE 4 A – B" captionStartId="4.[151,249,1680,1702]" captionTargetBox="[264,1323,952,1651]" captionTargetId="figure@4.[256,1331,945,1659]" captionTargetPageId="4" captionText="FIGURE 4 A – B. Rhecricotopus (Psilocricotopus) tobatervicesimus Kikuchi &amp; Sasa, Male imago. A. Hypopygium, dorsal view. B. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255140/files/figure.png" pageId="5" pageNumber="266">Figs 4 A–B</figureCitation>
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<emphasis id="52CDCC71FA72282DFF3EAE39FEAAFEBB" bold="true" box="[199,352,332,357]" pageId="5" pageNumber="266">Distribution.</emphasis>
The species occurs in
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(Oriental Region).
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164
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<emphasis id="52CDCC71FA74282BFF6EADD3FE9EFD1E" bold="true" box="[151,340,678,704]" italics="true" pageId="3" pageNumber="264">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA74282BFE9CADD3FDD4FD1E" bold="true" box="[357,542,678,704]" italics="true" pageId="3" pageNumber="264">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA74282BFDD6ADD3FD7CFD1E" bold="true" box="[559,694,678,704]" italics="true" pageId="3" pageNumber="264">kurocedeus</emphasis>
Sasa
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="60061063FA74282BFF6EADBCFEE0FD3C" blockId="3.[151,756,678,738]" box="[151,298,713,738]" pageId="3" pageNumber="264">
(
<figureCitation id="F8820CE6FA74282BFF66ADBCFEE8FD3C" box="[159,290,713,738]" captionStart="FIGURE 3 A – C" captionStartId="3.[151,249,1968,1990]" captionTargetBox="[241,1345,849,1946]" captionTargetId="figure@3.[231,1355,841,1947]" captionTargetPageId="3" captionText="FIGURE 3 A – C. Rhecricotopus (Psilocricotopus) kurocedeus Sasa, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255139/files/figure.png" pageId="3" pageNumber="264">Figs 3 A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA74282BFF6EAC7BFBF1FCFA" box="[151,1083,782,804]" pageId="3" pageNumber="264" type="reference_group">
<paragraph id="60061063FA74282BFF6EAC7BFBF1FCFA" blockId="3.[151,1083,782,804]" box="[151,1083,782,804]" pageId="3" pageNumber="264">
<treatmentCitationGroup id="40A9374DFA74282BFF6EAC7BFBF1FCFA" box="[151,1083,782,804]" pageId="3" pageNumber="264">
<taxonomicName id="A7B96BE0FA74282BFF6EAC7BFD3EFCFA" ID-CoL="9HS9Q" authority="Sasa, 1996: 19" authorityName="Sasa" authorityPageNumber="19" authorityYear="1996" box="[151,756,782,804]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="kurocedeus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA74282BFF6EAC7BFEFCFCFA" box="[151,310,782,804]" italics="true" pageId="3" pageNumber="264">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA74282BFEBDAC7BFE14FCFA" box="[324,478,782,804]" italics="true" pageId="3" pageNumber="264">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA74282BFE15AC7BFD91FCFA" box="[492,603,782,804]" italics="true" pageId="3" pageNumber="264">kurocedeus</emphasis>
<treatmentCitation id="E1183672FA74282BFD98AC7BFD3EFCFA" author="Sasa" box="[609,756,782,804]" page="19" pageId="3" pageNumber="264" year="1996">
<bibRefCitation id="04286D92FA74282BFD98AC7BFD05FCFA" author="Sasa" box="[609,719,782,804]" pageId="3" pageNumber="264" refString="Sasa, M. (1996) Studies on the chironomids in Japan, 1996. Research Report from Toyama Prefectural Environment Pollution Research Center, 12 - 102." type="journal article" year="1996">Sasa, 1996</bibRefCitation>
: 19
</treatmentCitation>
</taxonomicName>
;
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: 565
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="34C640EBFA74282BFF6EA8C5FE66F83B" httpUri="https://zenodo.org/record/255139/files/figure.png" pageId="3" pageNumber="264" targetBox="[241,1345,849,1946]" targetPageId="3">
<paragraph id="60061063FA74282BFF6EA8C5FE66F83B" blockId="3.[151,1436,1968,2021]" pageId="3" pageNumber="264">
<emphasis id="52CDCC71FA74282BFF6EA8C5FE87F818" bold="true" box="[151,333,1968,1990]" pageId="3" pageNumber="264">FIGURE 3 A–C.</emphasis>
<emphasis id="52CDCC71FA74282BFEADA8C5FE2CF818" box="[340,486,1968,1990]" italics="true" pageId="3" pageNumber="264">Rhecricotopus</emphasis>
<taxonomicName id="A7B96BE0FA74282BFE16A8C5FC9BF818" authority="Sasa" authorityName="Sasa" box="[495,849,1968,1990]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="kurocedeus" subGenus="Psilocricotopus">
(
<emphasis id="52CDCC71FA74282BFE0EA8C5FD5CF818" box="[503,662,1968,1990]" italics="true" pageId="3" pageNumber="264">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA74282BFD5FA8C5FCD3F818" box="[678,793,1968,1990]" italics="true" pageId="3" pageNumber="264">kurocedeus</emphasis>
Sasa
</taxonomicName>
, Male imago.
<emphasis id="52CDCC71FA74282BFC1EA8C5FC35F81B" bold="true" box="[999,1023,1968,1989]" pageId="3" pageNumber="264">A.</emphasis>
Wing.
<emphasis id="52CDCC71FA74282BFBB4A8C4FBAEF818" bold="true" box="[1101,1124,1969,1990]" pageId="3" pageNumber="264">B.</emphasis>
Hypopygium, dorsal view.
<emphasis id="52CDCC71FA74282BFA7AA8C5FA56F818" bold="true" box="[1411,1436,1968,1990]" pageId="3" pageNumber="264">C.</emphasis>
Hypopygium, ventral view.
</paragraph>
</caption>
<subSubSection id="28A343E8FA73282CFF6EAFE2FF19FD5A" pageId="4" pageNumber="265" type="materials_examined">
<paragraph id="60061063FA73282CFF6EAFE2FEFFFF0A" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="265">
<emphasis id="52CDCC71FA73282CFF6EAFE2FE48FF6E" bold="true" box="[151,386,151,176]" pageId="4" pageNumber="265">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA73282CFE72AFE2FE3CFF6E" box="[395,502,151,176]" pageId="4" pageNumber="265" type="holotype">Holotype</typeStatus>
, male, Kuroyon Lake, Toyama Prefecture, Honshu,
<collectingCountry id="18AE50F3FA73282CFBB5AFEDFB45FF71" box="[1100,1167,152,175]" name="Japan" pageId="4" pageNumber="265">Japan</collectingCountry>
.
<date id="140736A3FA73282CFB65AFEDFAEDFF6E" box="[1180,1319,152,176]" pageId="4" pageNumber="265" value="1994-10-14">14. X. 1994</date>
, light trap (No. 284: 31).
</paragraph>
<paragraph id="60061063FA73282CFF3EAFAAFC53FE9E" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="265">
<emphasis id="52CDCC71FA73282CFF3EAFAAFE1BFF26" bold="true" box="[199,465,223,248]" pageId="4" pageNumber="265">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by inferior volsella rounded and double-layered, anal point with 2 long lateral setae on each side, gonostylus almost straight and median expanded, with small triangularly pointed costa dorsalis.
</paragraph>
<paragraph id="60061063FA73282CFF3EAE39FF19FD5A" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="265">
<emphasis id="52CDCC71FA73282CFF3EAE39FD51FEBB" bold="true" box="[199,667,332,357]" pageId="4" pageNumber="265">Additional description and corrections.</emphasis>
Based on the analysis of
<typeStatus id="BF02AEC1FA73282CFC43AE39FBD6FEBB" box="[954,1052,332,357]" pageId="4" pageNumber="265" type="holotype">holotype</typeStatus>
material, the examined characters were consistent with the original description, b
<emphasis id="52CDCC71FA73282CFD53AE1AFD73FE56" bold="true" box="[682,697,367,392]" pageId="4" pageNumber="265">u</emphasis>
t some additional characters should be added: Total length / wing length 1.5. Wing length / length of profemur 2.8. Outer verticals 4. Tentorium 150 µm long, 45 µm wide. Palpomeres length (in µm): 33, 53, 110, 120, 213. Palpomere 2 ellipsoid, 3 and 4 rectangular, 5 long and slender, Palpomeres 5/3: 1.9. Laterosternite IX with 7 long setae. Anal point 48 µm long, basal wide 15 µm, peak wide 3 µm, with 2 long lateral setae on each side. Transverse sternapodeme 115 µm long, without oral projections. Superior volsella reduced. Gonocoxite 188 µm long. Gonostylus 70 µm long, with small triangularly pointed costa dorsalis. Megaseta 15 µm long. HR 2.7, HV 3.9. The wing and hypopygium of
<typeStatus id="BF02AEC1FA73282CFBE3AD32FBB6FDBE" box="[1050,1148,583,608]" pageId="4" pageNumber="265" type="holotype">holotype</typeStatus>
are re-drawn as in
<figureCitation id="F8820CE6FA73282CFAAFAD32FF05FD5A" captionStart="FIGURE 3 A – C" captionStartId="3.[151,249,1968,1990]" captionTargetBox="[241,1345,849,1946]" captionTargetId="figure@3.[231,1355,841,1947]" captionTargetPageId="3" captionText="FIGURE 3 A – C. Rhecricotopus (Psilocricotopus) kurocedeus Sasa, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255139/files/figure.png" pageId="4" pageNumber="265">Figs 3 A–C</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA73282CFF3EADFAFC58FD76" box="[199,914,655,680]" pageId="4" pageNumber="265" type="distribution">
<paragraph id="60061063FA73282CFF3EADFAFC58FD76" blockId="4.[151,1437,151,680]" box="[199,914,655,680]" pageId="4" pageNumber="265">
<emphasis id="52CDCC71FA73282CFF3EADFAFEAAFD76" bold="true" box="[199,352,655,680]" pageId="4" pageNumber="265">Distribution.</emphasis>
The species occurs in
<collectingCountry id="18AE50F3FA73282CFDA7ADE5FD54FD79" box="[606,670,656,679]" name="Japan" pageId="4" pageNumber="265">Japan</collectingCountry>
(Palaearctic Region).
</paragraph>
</subSubSection>
</treatment>
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159
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<heading id="3B4EA70FFA75282AFF6EAD51FCB8FDE0" bold="true" box="[151,882,548,574]" fontSize="11" level="1" pageId="2" pageNumber="263" reason="1">
<taxonomicName id="A7B96BE0FA75282AFF6EAD51FCB8FDE0" ID-CoL="4S5Z8" authority="Sasa &amp; Suzuki" authorityName="Sasa &amp; Suzuki" box="[151,882,548,574]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="isigadeeus" subGenus="Psilocricotopus">
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<emphasis id="52CDCC71FA75282AFF6EAD51FE9EFDE0" bold="true" box="[151,340,548,574]" italics="true" pageId="2" pageNumber="263">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA75282AFE9CAD51FDD4FDE0" bold="true" box="[357,542,548,574]" italics="true" pageId="2" pageNumber="263">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA75282AFDD6AD51FD66FDE0" bold="true" box="[559,684,548,574]" italics="true" pageId="2" pageNumber="263">isigadeeus</emphasis>
Sasa &amp; Suzuki
</emphasis>
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(
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)
</paragraph>
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<emphasis id="52CDCC71FA75282AFF6EADFBFEFCFD7A" box="[151,310,654,676]" italics="true" pageId="2" pageNumber="263">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA75282AFEBDADFBFE29FD7A" box="[324,483,654,676]" italics="true" pageId="2" pageNumber="263">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA75282AFE0BADFBFD97FD7A" box="[498,605,654,676]" italics="true" pageId="2" pageNumber="263">isigadeeus</emphasis>
Sasa &amp; Suzuki, 2000:7
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;
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</subSubSection>
<caption id="34C640EBFA75282AFF6EA844FDC2F8B8" httpUri="https://zenodo.org/record/255138/files/figure.png" pageId="2" pageNumber="263" targetBox="[226,1356,735,1819]" targetPageId="2">
<paragraph id="60061063FA75282AFF6EA844FDC2F8B8" blockId="2.[151,1436,1841,1894]" pageId="2" pageNumber="263">
<emphasis id="52CDCC71FA75282AFF6EA844FE86F899" bold="true" box="[151,332,1841,1863]" pageId="2" pageNumber="263">FIGURE 2 A–C.</emphasis>
<taxonomicName id="A7B96BE0FA75282AFEADA844FC78F899" authority="Sasa &amp; Suzuki" authorityName="Sasa &amp; Suzuki" box="[340,946,1841,1863]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="isigadeeus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA75282AFEADA844FE2CF899" box="[340,486,1841,1863]" italics="true" pageId="2" pageNumber="263">Rhecricotopus</emphasis>
(
<emphasis id="52CDCC71FA75282AFE0FA844FD5FF899" box="[502,661,1841,1863]" italics="true" pageId="2" pageNumber="263">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA75282AFD5CA844FCDAF899" box="[677,784,1841,1863]" italics="true" pageId="2" pageNumber="263">isigadeeus</emphasis>
Sasa &amp; Suzuki
</taxonomicName>
, Male imago.
<emphasis id="52CDCC71FA75282AFBBFA844FB95F898" bold="true" box="[1094,1119,1841,1862]" pageId="2" pageNumber="263">A.</emphasis>
Wing.
<emphasis id="52CDCC71FA75282AFB55A844FB09F898" bold="true" box="[1196,1219,1841,1862]" pageId="2" pageNumber="263">B.</emphasis>
Hypopygium, dorsal view.
<emphasis id="52CDCC71FA75282AFF2AA825FF21F8B8" bold="true" box="[211,235,1872,1894]" pageId="2" pageNumber="263">C.</emphasis>
Hypopygium, ventral view.
</paragraph>
</caption>
<subSubSection id="28A343E8FA75282BFF6EA8F8FD87FF26" lastPageId="3" lastPageNumber="264" pageId="2" pageNumber="263" type="materials_examined">
<paragraph id="60061063FA75282AFF6EA8F8FE6DF817" blockId="2.[151,1436,1933,1994]" pageId="2" pageNumber="263">
<emphasis id="52CDCC71FA75282AFF6EA8F8FE4AF878" bold="true" box="[151,384,1933,1958]" pageId="2" pageNumber="263">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA75282AFE7FA8F8FE3BF878" box="[390,497,1933,1958]" pageId="2" pageNumber="263" type="holotype">Holotype</typeStatus>
, male, Nakuragawa River, Ishigaki Island, Okinawa Prefecture, Ryukyus,
<collectingCountry id="18AE50F3FA75282AFAD5A8FBFAA5F87B" box="[1324,1391,1934,1957]" name="Japan" pageId="2" pageNumber="263">Japan</collectingCountry>
.
<date id="140736A3FA75282AFA80A8FBFF37F817" pageId="2" pageNumber="263" value="1999-06-30">30. VI. 1999</date>
, (No. 385: 08).
</paragraph>
<paragraph id="60061063FA74282BFF3EAFE2FD87FF26" blockId="3.[151,1437,151,608]" pageId="3" pageNumber="264">
<emphasis id="52CDCC71FA74282BFF3EAFE2FE1BFF6E" bold="true" box="[199,465,151,176]" pageId="3" pageNumber="264">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by inferior volsella extended and hooked posteriorly, Anal point with 5 long lateral setae on each side. Transverse sternapodeme without oral projections.
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA74282BFF3EAE71FD93FDE2" pageId="3" pageNumber="264" type="description">
<paragraph id="60061063FA74282BFF3EAE71FD93FDE2" blockId="3.[151,1437,151,608]" pageId="3" pageNumber="264">
<emphasis id="52CDCC71FA74282BFF3EAE71FD56FEC3" bold="true" box="[199,668,260,285]" pageId="3" pageNumber="264">Additional description and corrections.</emphasis>
Anal point with 5 long lateral setae on each side (Anal point with 4 pairs of marginal setae in original description). But some additional characters should be added: Total length / wing length 2.4. Wing length / length of profemur 2.4. Inner verticals 2, outer verticals 3. Clypeus with 19 setae. Tentorium 125 µm long, 20 µm wide. Dorsocentrals 13. Squama with 7 setae. Palpomeres length (in µm): 28, 45, 70, 90, 150. Palpomere 2 ellipsoid, 3 and 4 rectangular, 5 long and slender, Palpomeres 5/3: 2.1. Laterosternite IX with 6 long setae. Anal point 30 µm long, basal wide 20 µm, peak wide 3 µm, with 5 long lateral setae on each side. Transverse sternapodeme 58 µm long, without oral projections. Gonocoxite 165 µm long. Gonostylus 75 µm long, with triangularly pointed costa dorsalis. Megaseta 13 µm long. HR 2.2, HV 3.4. The wing and hypopygium of
<typeStatus id="BF02AEC1FA74282BFF6EAD51FF33FDE3" box="[151,249,548,573]" pageId="3" pageNumber="264" type="holotype">holotype</typeStatus>
are re-drawn as in
<figureCitation id="F8820CE6FA74282BFE29AD51FD9FFDE2" box="[464,597,548,573]" captionStart="FIGURE 2 A – C" captionStartId="2.[151,249,1841,1863]" captionTargetBox="[226,1356,735,1819]" captionTargetId="figure@2.[226,1360,729,1820]" captionTargetPageId="2" captionText="FIGURE 2 A – C. Rhecricotopus (Psilocricotopus) isigadeeus Sasa &amp; Suzuki, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255138/files/figure.png" pageId="3" pageNumber="264">Figs 2 A–C</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA74282BFF3EAD32FC58FDBE" box="[199,914,583,608]" pageId="3" pageNumber="264" type="distribution">
<paragraph id="60061063FA74282BFF3EAD32FC58FDBE" blockId="3.[151,1437,151,608]" box="[199,914,583,608]" pageId="3" pageNumber="264">
<emphasis id="52CDCC71FA74282BFF3EAD32FEAAFDBE" bold="true" box="[199,352,583,608]" pageId="3" pageNumber="264">Distribution.</emphasis>
The species occurs in
<collectingCountry id="18AE50F3FA74282BFDA7AD3DFD54FD81" box="[606,670,584,607]" name="Japan" pageId="3" pageNumber="264">Japan</collectingCountry>
(Palaearctic Region).
</paragraph>
</subSubSection>
</treatment>
</document>

155
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@ -0,0 +1,155 @@
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<mods:title id="7AF47386CB74256A5C95AB35427B95F1">Redescription of 13 holotypes of Rheocricotopus Brundin, 1956 (Diptera: Chironomidae) from the Sino – Indian Region</mods:title>
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<subSubSection id="28A343E8FA762829FF6EAE99FEE1FDF9" pageId="1" pageNumber="262" type="nomenclature">
<paragraph id="60061063FA762829FF6EAE99FC47FDD8" blockId="1.[151,909,492,551]" box="[151,909,492,518]" pageId="1" pageNumber="262">
<heading id="3B4EA70FFA762829FF6EAE99FC47FDD8" bold="true" box="[151,909,492,518]" fontSize="11" level="1" pageId="1" pageNumber="262" reason="1">
<taxonomicName id="A7B96BE0FA762829FF6EAE99FC47FDD8" ID-CoL="4S5Z4" authority="Sasa &amp; Suzuki" authorityName="Sasa &amp; Suzuki" box="[151,909,492,518]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="hidakadeeus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA762829FF6EAE99FC47FDD8" bold="true" box="[151,909,492,518]" pageId="1" pageNumber="262">
<emphasis id="52CDCC71FA762829FF6EAE99FE9EFDD8" bold="true" box="[151,340,492,518]" italics="true" pageId="1" pageNumber="262">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA762829FE9CAE99FDD4FDD8" bold="true" box="[357,542,492,518]" italics="true" pageId="1" pageNumber="262">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA762829FDD6AE99FD0CFDD8" bold="true" box="[559,710,492,518]" italics="true" pageId="1" pageNumber="262">hidakadeeus</emphasis>
Sasa &amp; Suzuki
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="60061063FA762829FF6EAD7BFEE1FDF9" blockId="1.[151,909,492,551]" box="[151,299,526,551]" pageId="1" pageNumber="262">
(
<figureCitation id="F8820CE6FA762829FF66AD7BFEE8FDF9" box="[159,290,526,551]" captionStart="FIGURE 1 A – C" captionStartId="1.[151,249,1765,1787]" captionTargetBox="[273,1320,667,1740]" captionTargetId="figure@1.[258,1328,661,1752]" captionTargetPageId="1" captionText="FIGURE 1 A – C. Rhecricotopus (Psilocricotopus) hidakadeeus Sasa &amp; Suzuki, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255137/files/figure.png" pageId="1" pageNumber="262">Figs 1 A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA762829FF6EAD23FBEAFDB2" box="[151,1056,598,620]" pageId="1" pageNumber="262" type="reference_group">
<paragraph id="60061063FA762829FF6EAD23FBEAFDB2" blockId="1.[151,1056,598,620]" box="[151,1056,598,620]" pageId="1" pageNumber="262">
<treatmentCitationGroup id="40A9374DFA762829FF6EAD23FBEAFDB2" box="[151,1056,598,620]" pageId="1" pageNumber="262">
<taxonomicName id="A7B96BE0FA762829FF6EAD23FD13FDB2" ID-CoL="4S5Z4" authority="Sasa &amp; Suzuki, 2000: 184" authorityName="Sasa &amp; Suzuki" authorityPageNumber="184" authorityYear="2000" box="[151,729,598,620]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="hidakadeeus">
<emphasis id="52CDCC71FA762829FF6EAD23FE71FDB2" box="[151,443,598,620]" italics="true" pageId="1" pageNumber="262">Rheocricotopus hidakadeeus</emphasis>
Sasa &amp; Suzuki, 2000: 184
</taxonomicName>
;
<treatmentCitation id="E1183672FA762829FD1DAD23FBD6FDB2" author="Ashe" box="[740,1052,598,620]" page="564" pageId="1" pageNumber="262" year="2012">
<bibRefCitation id="04286D92FA762829FD1DAD23FC2DFDB2" author="Ashe" box="[740,999,598,620]" pageId="1" pageNumber="262" refString="Ashe, P. &amp; O'Connor, J. P. (2012) A world catalogue of Chironomidae (Diptera). Part 2 B. Orthocladiinae. Irish Biogeographical Society &amp; National Museum of Ireland, Dublin, pp. 558 - 575." type="book chapter" year="2012">Ashe &amp; O’Connor 2012</bibRefCitation>
: 564
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="34C640EBFA762829FF6EA990FDC2F8C4" httpUri="https://zenodo.org/record/255137/files/figure.png" pageId="1" pageNumber="262" targetBox="[273,1320,667,1740]" targetPageId="1">
<paragraph id="60061063FA762829FF6EA990FDC2F8C4" blockId="1.[151,1436,1765,1818]" pageId="1" pageNumber="262">
<emphasis id="52CDCC71FA762829FF6EA990FE80F925" bold="true" box="[151,330,1765,1787]" pageId="1" pageNumber="262">FIGURE 1 A–C.</emphasis>
<taxonomicName id="A7B96BE0FA762829FEA9A990FC71F925" authority="Sasa &amp; Suzuki" authorityName="Sasa &amp; Suzuki" box="[336,955,1765,1787]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="hidakadeeus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA762829FEA9A990FE28F925" box="[336,482,1765,1787]" italics="true" pageId="1" pageNumber="262">Rhecricotopus</emphasis>
(
<emphasis id="52CDCC71FA762829FE08A990FD5AF925" box="[497,656,1765,1787]" italics="true" pageId="1" pageNumber="262">Psilocricotopus</emphasis>
)
<emphasis id="52CDCC71FA762829FD66A990FCD7F925" box="[671,797,1765,1787]" italics="true" pageId="1" pageNumber="262">hidakadeeus</emphasis>
Sasa &amp; Suzuki
</taxonomicName>
, Male imago.
<emphasis id="52CDCC71FA762829FBB5A993FBAEF925" bold="true" box="[1100,1124,1766,1787]" pageId="1" pageNumber="262">A.</emphasis>
Wing.
<emphasis id="52CDCC71FA762829FB57A993FB0FF925" bold="true" box="[1198,1221,1766,1787]" pageId="1" pageNumber="262">B.</emphasis>
Hypopygium, dorsal view.
<emphasis id="52CDCC71FA762829FF2AA871FF21F8C4" bold="true" box="[211,235,1796,1818]" pageId="1" pageNumber="262">C.</emphasis>
Hypopygium, ventral view.
</paragraph>
</caption>
<subSubSection id="28A343E8FA76282AFF6EA834FCD7FE72" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262" type="materials_examined">
<paragraph id="60061063FA762829FF6EA834FE1BF8A0" blockId="1.[151,1437,1857,2026]" pageId="1" pageNumber="262">
<emphasis id="52CDCC71FA762829FF6EA834FE49F884" bold="true" box="[151,387,1857,1882]" pageId="1" pageNumber="262">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA762829FE74A834FE32F884" box="[397,504,1857,1882]" pageId="1" pageNumber="262" type="holotype">Holotype</typeStatus>
, male, Hidaka Mountain areas, Nakasatsunai Village, Tokachi, Hokkaido,
<collectingCountry id="18AE50F3FA762829FAACA837FA52F887" box="[1365,1432,1858,1881]" name="Japan" pageId="1" pageNumber="262">Japan</collectingCountry>
.
<date id="140736A3FA762829FF6EA810FEEDF8A0" box="[151,295,1893,1918]" pageId="1" pageNumber="262" value="1998-09-26">26. IX. 1998</date>
, (No. 401: 51).
</paragraph>
<paragraph id="60061063FA762829FF3EA8FCFEA2F834" blockId="1.[151,1437,1857,2026]" pageId="1" pageNumber="262">
<emphasis id="52CDCC71FA762829FF3EA8FCFE1DF87C" bold="true" box="[199,471,1929,1954]" pageId="1" pageNumber="262">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by developed superior volsella with rounded corner, inferior volsella double-layered, anal point with 2 long lateral setae on each side.
</paragraph>
<paragraph id="60061063FA75282AFF3EAFE2FCD7FE72" blockId="2.[151,1437,151,464]" pageId="2" pageNumber="263">
<emphasis id="52CDCC71FA75282AFF3EAFE2FD51FF6E" bold="true" box="[199,667,151,176]" pageId="2" pageNumber="263">Additional description and corrections.</emphasis>
Based on the analysis of
<typeStatus id="BF02AEC1FA75282AFC43AFE2FBD6FF6E" box="[954,1052,151,176]" pageId="2" pageNumber="263" type="holotype">holotype</typeStatus>
material, the examined characters were consistent with the original description. But some additional characters should be added: Total length/ wing length 1.8. Wing length/ profemur length 2.5. Tentorium 188 µm long, 43 µm wide. Palpomeres length (in µm): –, 70, 120, 175, 275, palpomere 2 ellipsoid, 3 and 4 rectangular, 5 long and slender, palpomeres 5/ 3: 2.3. Laterosternite IX with 3 long setae. Anal point 58 µm long, basal wide 20 µm, peak wide 3 µm, with 2 long lateral setae on each side. Transverse sternapodeme 108 µm long, with weak oral projections. Gonocoxite 238 µm long. Gonostylus 108 µm long, with small triangularly costa dorsalis. Megaseta 12 µm long. HR 2.2, HV 3.3. The wing and hypopygium of
<typeStatus id="BF02AEC1FA75282AFE80AEE1FE11FE73" box="[377,475,404,429]" pageId="2" pageNumber="263" type="holotype">holotype</typeStatus>
are re-drawn in
<figureCitation id="F8820CE6FA75282AFD6DAEE1FCD3FE72" box="[660,793,404,429]" captionStart="FIGURE 1 A – C" captionStartId="1.[151,249,1765,1787]" captionTargetBox="[273,1320,667,1740]" captionTargetId="figure@1.[258,1328,661,1752]" captionTargetPageId="1" captionText="FIGURE 1 A – C. Rhecricotopus (Psilocricotopus) hidakadeeus Sasa &amp; Suzuki, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255137/files/figure.png" pageId="2" pageNumber="263">Figs 1 A–C</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA75282AFF3EAEC2FC53FE0E" box="[199,921,439,464]" pageId="2" pageNumber="263" type="distribution">
<paragraph id="60061063FA75282AFF3EAEC2FC53FE0E" blockId="2.[151,1437,151,464]" box="[199,921,439,464]" pageId="2" pageNumber="263">
<emphasis id="52CDCC71FA75282AFF3EAEC2FEAAFE0E" bold="true" box="[199,352,439,464]" pageId="2" pageNumber="263">Distribution.</emphasis>
The species occurs in
<collectingCountry id="18AE50F3FA75282AFDA7AECDFD54FE11" box="[606,670,440,463]" name="Japan" pageId="2" pageNumber="263">Japan</collectingCountry>
(Palaearctic Region).
</paragraph>
</subSubSection>
</treatment>
</document>

309
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<mods:title id="CE4BCD9846664BE6E5F6BCA56BA23ECD">Redescription of 13 holotypes of Rheocricotopus Brundin, 1956 (Diptera: Chironomidae) from the Sino – Indian Region</mods:title>
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<subSubSection id="28A343E8FA792826FF6EAFEDFB8EFF6C" box="[151,1092,151,178]" pageId="14" pageNumber="275" type="nomenclature">
<paragraph id="60061063FA792826FF6EAFEDFB8EFF6C" blockId="14.[151,1092,151,178]" box="[151,1092,151,178]" pageId="14" pageNumber="275">
<heading id="3B4EA70FFA792826FF6EAFEDFB8EFF6C" bold="true" box="[151,1092,151,178]" fontSize="11" level="1" pageId="14" pageNumber="275" reason="1">
<emphasis id="52CDCC71FA792826FF6EAFEDFB8EFF6C" bold="true" box="[151,1092,151,178]" pageId="14" pageNumber="275">
Key to of
<specimenCount id="76BFDBEAFA792826FEE9AFEDFEB6FF6C" box="[272,380,152,178]" pageId="14" pageNumber="275" type="male">13 males</specimenCount>
of
<taxonomicName id="A7B96BE0FA792826FE5AAFE2FD1BFF6C" ID-CoL="78HQ" ID-ENA="611384" authority="Brundin" authorityName="Brundin" box="[419,721,151,178]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="52CDCC71FA792826FE5AAFE2FDAAFF6F" bold="true" box="[419,608,151,177]" italics="true" pageId="14" pageNumber="275">Rheocricotopus</emphasis>
Brundin
</taxonomicName>
from the Sino–Indian Region
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA792826FF6EAFABFA56FC30" pageId="14" pageNumber="275" type="key">
<keyStep id="DB4D08C6FA792826FF6EAFABFA51FECE" pageId="14" pageNumber="275">
<paragraph id="60061063FA792826FF6EAFABFA56FF2D" blockId="14.[151,1436,222,1006]" box="[151,1436,222,243]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAFABFA56FF2D" box="[151,1436,222,243]" pageId="14" pageNumber="275">1. Tergite IX reduced, the posterior margin of tergite IX almost coincide with the base of anal point...................... 2</keyLead>
</paragraph>
<paragraph id="60061063FA792826FF6EAF8EFA51FECE" blockId="14.[151,1436,222,1006]" box="[151,1435,251,272]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAF8EFA51FECE" box="[151,1435,251,272]" pageId="14" pageNumber="275">- Tergite IX developed, the posterior margin of tergite IX exceed the apex of anal point............................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4D08C6FA792826FF6EAE6CFA51FE95" pageId="14" pageNumber="275">
<paragraph id="60061063FA792826FF6EAE6CFA51FEF0" blockId="14.[151,1436,222,1006]" box="[151,1435,281,302]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAE6CFA51FEF0" box="[151,1435,281,302]" pageId="14" pageNumber="275">2. Superior volsella triangular or rounded or absent, relatively small............................................... 3</keyLead>
</paragraph>
<paragraph id="60061063FA792826FF6EAE43FA51FE95" blockId="14.[151,1436,222,1006]" box="[151,1435,310,331]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAE43FA51FE95" box="[151,1435,310,331]" pageId="14" pageNumber="275">– Superior volsella longitypical or tooth–like, relatively large.................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4D08C6FA792826FF6EAE26FA51FE58" pageId="14" pageNumber="275">
<paragraph id="60061063FA792826FF6EAE26FA56FEB6" blockId="14.[151,1436,222,1006]" box="[151,1436,339,360]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAE26FA56FEB6" box="[151,1436,339,360]" pageId="14" pageNumber="275">
3. Inferior volsella triangular and hooked...........................................
<taxonomicName id="A7B96BE0FA792826FBA4AE26FA56FEB6" authority="Sasa &amp; Suzuki" authorityName="Sasa &amp; Suzuki" box="[1117,1436,339,360]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="isigadeeus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA792826FBA4AE26FBBFFEB6" box="[1117,1141,339,360]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FB7DAE26FB5EFEB6" box="[1156,1172,339,360]" italics="true" pageId="14" pageNumber="275">P.</emphasis>
)
<emphasis id="52CDCC71FA792826FB5AAE26FAC2FEB6" box="[1187,1288,339,360]" italics="true" pageId="14" pageNumber="275">isigadeeus</emphasis>
Sasa &amp; Suzuki
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="60061063FA792826FF6EAE04FA51FE58" blockId="14.[151,1436,222,1006]" box="[151,1435,369,390]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAE04FA51FE58" box="[151,1435,369,390]" pageId="14" pageNumber="275">- Inferior volsella rounded................................................................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4D08C6FA792826FF6EAEFBFA56FE25" pageId="14" pageNumber="275">
<paragraph id="60061063FA792826FF6EAEFBFA56FE1E" blockId="14.[151,1436,222,1006]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAEFBFA56FE1E" pageId="14" pageNumber="275">
4. Anal point with 2 lateral setae on each side, inferior volsella double–layered..........................................................................................................................
<taxonomicName id="A7B96BE0FA792826FB42AEDEFA56FE1E" ID-CoL="9HS9Q" authority="Sasa" authorityName="Sasa" box="[1211,1436,427,448]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="kurocedeus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA792826FB42AEDEFB1AFE1E" box="[1211,1232,427,448]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FB27AEDEFB24FE1E" box="[1246,1262,427,448]" italics="true" pageId="14" pageNumber="275">P.</emphasis>
)
<emphasis id="52CDCC71FA792826FB04AEDEFAA3FE1E" box="[1277,1385,427,448]" italics="true" pageId="14" pageNumber="275">kurocedeus</emphasis>
Sasa
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="60061063FA792826FF6EAEBCFA56FE25" blockId="14.[151,1436,222,1006]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAEBCFA56FE25" pageId="14" pageNumber="275">
- Anal point with without lateral setae on each side, inferior volsella single–layered........................................................................................................
<taxonomicName id="A7B96BE0FA792826FBEFAE93FA56FE25" authority="Kikuchi &amp; Sasa" authorityName="Kikuchi &amp; Sasa" box="[1046,1436,486,507]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="tobatervicesimus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA792826FBEFAE93FBE4FE25" box="[1046,1070,486,507]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FBC5AE93FB86FE25" box="[1084,1100,486,507]" italics="true" pageId="14" pageNumber="275">P.</emphasis>
)
<emphasis id="52CDCC71FA792826FBA5AE93FB34FE25" box="[1116,1278,486,507]" italics="true" pageId="14" pageNumber="275">tobatervicesimus</emphasis>
Kikuchi &amp; Sasa
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4D08C6FA792826FF6EAD76FA56FDE8" pageId="14" pageNumber="275">
<paragraph id="60061063FA792826FF6EAD76FA56FDC6" blockId="14.[151,1436,222,1006]" box="[151,1436,515,536]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAD76FA56FDC6" box="[151,1436,515,536]" pageId="14" pageNumber="275">
5. Superior volsella longitypical, inferior volsella pointed triangular...........................
<taxonomicName id="A7B96BE0FA792826FB6DAD76FA56FDC6" authority="Sasa" authorityName="Sasa" box="[1172,1436,515,536]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="tamahumeralis" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA792826FB6DAD76FB60FDC6" box="[1172,1194,515,536]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FB4EAD76FB07FDC6" box="[1207,1229,515,536]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
)
<emphasis id="52CDCC71FA792826FB23AD76FAA0FDC6" box="[1242,1386,515,536]" italics="true" pageId="14" pageNumber="275">tamahumeralis</emphasis>
Sasa
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="60061063FA792826FF6EAD54FA56FDE8" blockId="14.[151,1436,222,1006]" box="[151,1436,545,566]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAD54FA56FDE8" box="[151,1436,545,566]" pageId="14" pageNumber="275">
- Superior volsella tooth–like, inferior volsella rounded....................................
<taxonomicName id="A7B96BE0FA792826FB6AAD54FA56FDE8" ID-CoL="9J2G2" authority="Sasa" authorityName="Sasa" box="[1171,1436,545,566]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="tedorisecundus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA792826FB6AAD54FB62FDE8" box="[1171,1192,545,566]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FB4FAD54FB01FDE8" box="[1206,1227,545,566]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
)
<emphasis id="52CDCC71FA792826FB20AD54FAA0FDE8" box="[1241,1386,545,566]" italics="true" pageId="14" pageNumber="275">tedorisecundus</emphasis>
Sasa
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4D08C6FA792826FF6EAD4BFA51FDAF" pageId="14" pageNumber="275">
<paragraph id="60061063FA792826FF6EAD4BFA51FD8D" blockId="14.[151,1436,222,1006]" box="[151,1435,574,595]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAD4BFA51FD8D" box="[151,1435,574,595]" pageId="14" pageNumber="275">6. Gonostylus with apical pointed triangular crista dorsalis....................................................... 7</keyLead>
</paragraph>
<paragraph id="60061063FA792826FF6EAD29FA51FDAF" blockId="14.[151,1436,222,1006]" box="[151,1435,604,625]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAD29FA51FDAF" box="[151,1435,604,625]" pageId="14" pageNumber="275">- Gonostylus with preapical or middle rounded triangular crista dorsalis or without................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4D08C6FA792826FF6EAD0CFA56FD17" pageId="14" pageNumber="275">
<paragraph id="60061063FA792826FF6EAD0CFA56FD50" blockId="14.[151,1436,222,1006]" box="[151,1436,633,654]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAD0CFA56FD50" box="[151,1436,633,654]" pageId="14" pageNumber="275">
7. Anal point with 2 lateral setae on each side, inferior volsella rounded, double–layered....
<taxonomicName id="A7B96BE0FA792826FBB7AD0CFA56FD50" authority="Sasa &amp; Suzuki" authorityName="Sasa &amp; Suzuki" box="[1102,1436,633,654]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="hidakadeeus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA792826FBB7AD0CFBAEFD50" box="[1102,1124,633,654]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FB88AD0CFB4BFD50" box="[1137,1153,633,654]" italics="true" pageId="14" pageNumber="275">P.</emphasis>
)
<emphasis id="52CDCC71FA792826FB69AD0CFAC2FD50" box="[1168,1288,633,654]" italics="true" pageId="14" pageNumber="275">hidakadeeus</emphasis>
Sasa &amp; Suzuki
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="60061063FA792826FF6EADE3FA56FD17" blockId="14.[151,1436,222,1006]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EADE3FA56FD17" pageId="14" pageNumber="275">
- Anal point with with 5 lateral setae on each side, inferior volsella slightly hooked, single–layered................................................................................................
<taxonomicName id="A7B96BE0FA792826FBB5ADC1FA56FD17" authority="Sasa &amp; Suzuki" authorityName="Sasa &amp; Suzuki" box="[1100,1436,692,713]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="tokarakeleus" subGenus="Psilocricotopus">
<emphasis id="52CDCC71FA792826FBB5ADC1FBABFD17" box="[1100,1121,692,713]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FB97ADC1FBB5FD17" box="[1134,1151,692,713]" italics="true" pageId="14" pageNumber="275">P.</emphasis>
)
<emphasis id="52CDCC71FA792826FB77ADC1FAC3FD17" box="[1166,1289,692,713]" italics="true" pageId="14" pageNumber="275">tokarakeleus</emphasis>
Sasa &amp; Suzuki
</taxonomicName>
</keyLead>
</paragraph>
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<keyLead id="DB48B356FA792826FF6EADA4FA56FD38" box="[151,1436,721,742]" pageId="14" pageNumber="275">
8. Gonostylus without crista dorsalis..........................................
<taxonomicName id="A7B96BE0FA792826FBDCADA4FA56FD38" ID-CoL="9HT6F" authority="Sasa &amp; Okazawa" authorityName="Sasa &amp; Okazawa" box="[1061,1436,721,742]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="togapeniculus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA792826FBDCADA4FBF1FD38" box="[1061,1083,721,742]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FBB0ADA4FB94FD38" box="[1097,1118,721,742]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
)
<emphasis id="52CDCC71FA792826FB95ADA4FB38FD38" box="[1132,1266,721,742]" italics="true" pageId="14" pageNumber="275">togapeniculus</emphasis>
Sasa &amp; Okazawa
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<paragraph id="60061063FA792826FF6EAD9BFA51FCDD" blockId="14.[151,1436,222,1006]" box="[151,1435,750,771]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAD9BFA51FCDD" box="[151,1435,750,771]" pageId="14" pageNumber="275">- Gonostylus with crista dorsalis........................................................................... 9</keyLead>
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<paragraph id="60061063FA792826FF64AC79FA51FCFF" blockId="14.[151,1436,222,1006]" box="[157,1435,780,801]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF64AC79FA51FCFF" box="[157,1435,780,801]" pageId="14" pageNumber="275">9. Superior volsella with almost right–angle corner............................................................ 10</keyLead>
</paragraph>
<paragraph id="60061063FA792826FF6EAC5CFA51FCE0" blockId="14.[151,1436,222,1006]" box="[151,1435,809,830]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAC5CFA51FCE0" box="[151,1435,809,830]" pageId="14" pageNumber="275">- Superior volsella with rounded corner or tooth–like or absent.................................................. 11</keyLead>
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<keyStep id="DB4D08C6FA792826FF6EAC33FA56FCA7" pageId="14" pageNumber="275">
<paragraph id="60061063FA792826FF6EAC33FA56FC85" blockId="14.[151,1436,222,1006]" box="[151,1436,838,859]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAC33FA56FC85" box="[151,1436,838,859]" pageId="14" pageNumber="275">
10. Transverse sternapodeme without oral projections, inferior vosella single–layered..
<taxonomicName id="A7B96BE0FA792826FBEFAC33FA56FC85" ID-CoL="9HVHH" authority="Sasa, Kitami &amp; Suzuki" authorityName="Sasa, Kitami &amp; Suzuki" box="[1046,1436,838,859]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="inaxeyeus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA792826FBEFAC33FBE1FC85" box="[1046,1067,838,859]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FBC0AC33FB84FC85" box="[1081,1102,838,859]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
)
<emphasis id="52CDCC71FA792826FBA5AC33FB70FC85" box="[1116,1210,838,859]" italics="true" pageId="14" pageNumber="275">inaxeyeus</emphasis>
Sasa, Kitami &amp; Suzuki
</taxonomicName>
</keyLead>
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<paragraph id="60061063FA792826FF6EAC11FA56FCA7" blockId="14.[151,1436,222,1006]" box="[151,1436,868,889]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EAC11FA56FCA7" box="[151,1436,868,889]" pageId="14" pageNumber="275">
- Transverse sternapodeme with oral projections, inferior vosella double–layered...
<taxonomicName id="A7B96BE0FA792826FBF2AC11FA56FCA7" ID-CoL="9HRXG" authority="Sasa, Kitami &amp; Suzuki" authorityName="Sasa, Kitami &amp; Suzuki" box="[1035,1436,868,889]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="inaquereus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA792826FBF2AC11FBEAFCA7" box="[1035,1056,868,889]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FBD7AC11FB89FCA7" box="[1070,1091,868,889]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
)
<emphasis id="52CDCC71FA792826FBA9AC11FB70FCA7" box="[1104,1210,868,889]" italics="true" pageId="14" pageNumber="275">inaquereus</emphasis>
Sasa, Kitami &amp; Suzuki
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<keyLead id="DB48B356FA792826FF6EACF4FA56FC48" box="[151,1436,897,918]" pageId="14" pageNumber="275">
11. Superior volsella tooth–shaped.........................................................
<taxonomicName id="A7B96BE0FA792826FB40ACF4FA56FC48" ID-CoL="9HV5Q" authority="Sasa" authorityName="Sasa" box="[1209,1436,897,918]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="tatequintus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA792826FB40ACF4FB04FC48" box="[1209,1230,897,918]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FB25ACF4FB38FC48" box="[1244,1266,897,918]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
)
<emphasis id="52CDCC71FA792826FB06ACF4FAA3FC48" box="[1279,1385,897,918]" italics="true" pageId="14" pageNumber="275">tatequintus</emphasis>
Sasa
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="60061063FA792826FF6EACEBFA51FC6D" blockId="14.[151,1436,222,1006]" box="[151,1435,926,947]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EACEBFA51FC6D" box="[151,1435,926,947]" pageId="14" pageNumber="275">- Superior volsella rounded or absent...................................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4D08C6FA792826FF6EACC9FA56FC30" pageId="14" pageNumber="275">
<paragraph id="60061063FA792826FF6EACC9FA56FC0F" blockId="14.[151,1436,222,1006]" box="[151,1436,956,977]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EACC9FA56FC0F" box="[151,1436,956,977]" pageId="14" pageNumber="275">
12. Inferior volsella foot–shaped, gonostylus with preapical crista dorsalis...................
<taxonomicName id="A7B96BE0FA792826FB8FACC9FA56FC0F" authority="Sasa" authorityName="Sasa" box="[1142,1436,956,977]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="shoufukusecundus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA792826FB8FACC9FB46FC0F" box="[1142,1164,956,977]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FB60ACC9FB64FC0F" box="[1177,1198,956,977]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
)
<emphasis id="52CDCC71FA792826FB42ACC9FAA3FC0F" box="[1211,1385,956,977]" italics="true" pageId="14" pageNumber="275">shoufukusecundus</emphasis>
Sasa
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="60061063FA792826FF6EACACFA56FC30" blockId="14.[151,1436,222,1006]" box="[151,1436,985,1006]" pageId="14" pageNumber="275">
<keyLead id="DB48B356FA792826FF6EACACFA56FC30" box="[151,1436,985,1006]" pageId="14" pageNumber="275">
- Inferior volsella rounded with a small lobe, gonostylus with middle crista dorsalis.......
<taxonomicName id="A7B96BE0FA792826FBAAACACFA56FC30" authority="Sasa &amp; Suzuki" authorityName="Sasa &amp; Suzuki" box="[1107,1436,985,1006]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="275" phylum="Arthropoda" rank="species" species="yakulemeus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA792826FBAAACACFBA3FC30" box="[1107,1129,985,1006]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
(
<emphasis id="52CDCC71FA792826FB8FACACFB41FC30" box="[1142,1163,985,1006]" italics="true" pageId="14" pageNumber="275">R.</emphasis>
)
<emphasis id="52CDCC71FA792826FB61ACACFAC2FC30" box="[1176,1288,985,1006]" italics="true" pageId="14" pageNumber="275">yakulemeus</emphasis>
Sasa &amp; Suzuki
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
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<heading id="3B4EA70FFA7A2825FF6EAFE2FCB5FF6C" bold="true" box="[151,895,151,178]" fontSize="11" level="1" pageId="13" pageNumber="274" reason="1">
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<emphasis id="52CDCC71FA7A2825FF6EAFE2FE9EFF6F" bold="true" box="[151,340,151,177]" italics="true" pageId="13" pageNumber="274">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA7A2825FE9CAFE2FDE8FF6F" bold="true" box="[357,546,151,177]" italics="true" pageId="13" pageNumber="274">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7A2825FDCAAFE2FD08FF6F" bold="true" box="[563,706,151,177]" italics="true" pageId="13" pageNumber="274">yakulemeus</emphasis>
Sasa &amp; Suzuki
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<paragraph id="60061063FA7A2825FF6EAFCFFEF3FF0D" blockId="13.[151,895,151,211]" box="[151,313,186,211]" pageId="13" pageNumber="274">
(
<figureCitation id="F8820CE6FA7A2825FF66AFCFFEFAFF0D" box="[159,304,186,211]" captionStart="FIGURE 13 A – C" captionStartId="13.[151,249,1982,2004]" captionTargetBox="[236,1355,882,1960]" captionTargetId="figure@13.[226,1360,873,1961]" captionTargetPageId="13" captionText="FIGURE 13 A – C. Rhecricotopus (Rheocricotopus) yakulemeus Sasa &amp; Okazawa, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255149/files/figure.png" pageId="13" pageNumber="274">Figs 13 A–C</figureCitation>
)
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<emphasis id="52CDCC71FA7A2825FF6EAE77FEFCFEC6" box="[151,310,258,280]" italics="true" pageId="13" pageNumber="274">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA7A2825FEBDAE77FE29FEC6" box="[324,483,258,280]" italics="true" pageId="13" pageNumber="274">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7A2825FE0BAE77FDA2FEC6" box="[498,616,258,280]" italics="true" pageId="13" pageNumber="274">yakulemeus</emphasis>
Sasa &amp; Suzuki, 2000: 69
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;
<treatmentCitation id="E1183672FA7A2825FC81AE77FB65FEC6" author="Ashe" box="[888,1199,258,280]" page="573" pageId="13" pageNumber="274" year="2012">
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: 573
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.
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<paragraph id="60061063FA7A2825FF6EAE30FF36FE5F" blockId="13.[151,1437,325,818]" pageId="13" pageNumber="274">
<emphasis id="52CDCC71FA7A2825FF6EAE30FE4BFE80" bold="true" box="[151,385,325,350]" pageId="13" pageNumber="274">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA7A2825FE71AE30FE39FE80" box="[392,499,325,350]" pageId="13" pageNumber="274" type="holotype">Holotype</typeStatus>
, male, Nagata, Yakushima Island, Kagoshima Prefecture,
<collectingCountry id="18AE50F3FA7A2825FB79AE33FB09FE83" box="[1152,1219,326,349]" name="Japan" pageId="13" pageNumber="274">Japan</collectingCountry>
.
<date id="140736A3FA7A2825FB37AE33FA94FE80" box="[1230,1374,326,350]" pageId="13" pageNumber="274" value="1999-03-23">23. III. 1999</date>
. (No. 381: 44).
</paragraph>
<paragraph id="60061063FA7A2825FF3EAEF8FDDEFE30" blockId="13.[151,1437,325,818]" pageId="13" pageNumber="274">
<emphasis id="52CDCC71FA7A2825FF3EAEF8FE1DFE78" bold="true" box="[199,471,397,422]" pageId="13" pageNumber="274">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by laterosternite IX with 1 long seta, anal point with 5 long lateral setae on each side, inferior volsella developed and double-layered, with a small lobe.
</paragraph>
<paragraph id="60061063FA7A2825FF3EAE8CFBD1FCD0" blockId="13.[151,1437,325,818]" pageId="13" pageNumber="274">
<emphasis id="52CDCC71FA7A2825FF3EAE8CFD51FDCC" bold="true" box="[199,667,505,530]" pageId="13" pageNumber="274">Additional description and corrections.</emphasis>
Based on the analysis of
<typeStatus id="BF02AEC1FA7A2825FC43AE8CFBD6FDCC" box="[954,1052,505,530]" pageId="13" pageNumber="274" type="holotype">holotype</typeStatus>
material, the examined characters were consistent with the original description, but some additional characters should be added: Total length / wing length 2.0. Wing length / length of profemur 2.5. Tentorium 155 µm long, 38 µm wide. Dorsocentrals 20. Palpomeres length (in µm): 45, 63, 100, 125, 198. Palpomere 2 ellipsoid, 3 and 4 rectangular, 5 long and slender, Palpomeres 5/3: 2.0. Laterosternite IX with 2 long setae. Anal point 45 µm long, basal wide 32 µm, peak wide 10 µm, with 5 long lateral setae on each side. Transverse sternapodeme 65 µm long, with weak oral projections. Gonocoxite 195 µm long. Gonostylus 83 µm long, with triangular crista dorsalis. Megaseta 15 µm long. HR 2.4, HV 3.6. The wing and hypopygium of
<typeStatus id="BF02AEC1FA7A2825FDB2AD80FD67FCD0" box="[587,685,757,782]" pageId="13" pageNumber="274" type="holotype">holotype</typeStatus>
are re-drawn as in
<figureCitation id="F8820CE6FA7A2825FC7CAD80FBDDFCD0" box="[901,1047,757,782]" captionStart="FIGURE 13 A – C" captionStartId="13.[151,249,1982,2004]" captionTargetBox="[236,1355,882,1960]" captionTargetId="figure@13.[226,1360,873,1961]" captionTargetPageId="13" captionText="FIGURE 13 A – C. Rhecricotopus (Rheocricotopus) yakulemeus Sasa &amp; Okazawa, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255149/files/figure.png" pageId="13" pageNumber="274">Figs 13 A–C</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA7A2825FF3EAC6CFC58FCEC" box="[199,914,793,818]" pageId="13" pageNumber="274" type="distribution">
<paragraph id="60061063FA7A2825FF3EAC6CFC58FCEC" blockId="13.[151,1437,325,818]" box="[199,914,793,818]" pageId="13" pageNumber="274">
<emphasis id="52CDCC71FA7A2825FF3EAC6CFEAAFCEC" bold="true" box="[199,352,793,818]" pageId="13" pageNumber="274">Distribution.</emphasis>
The species occurs in
<collectingCountry id="18AE50F3FA7A2825FDA7AC6FFD54FCEF" box="[606,670,794,817]" name="Japan" pageId="13" pageNumber="274">Japan</collectingCountry>
(Palaearctic Region).
</paragraph>
</subSubSection>
</treatment>
</document>

154
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<mods:title id="3F65676E58E1558A4389278EB244A4AE">Redescription of 13 holotypes of Rheocricotopus Brundin, 1956 (Diptera: Chironomidae) from the Sino – Indian Region</mods:title>
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<paragraph id="60061063FA7B2824FF6EAFE2FC72FF6C" blockId="12.[151,952,151,211]" box="[151,952,151,178]" pageId="12" pageNumber="273">
<heading id="3B4EA70FFA7B2824FF6EAFE2FC72FF6C" bold="true" box="[151,952,151,178]" fontSize="11" level="1" pageId="12" pageNumber="273" reason="1">
<taxonomicName id="A7B96BE0FA7B2824FF6EAFE2FC72FF6C" ID-CoL="9HT6F" authority="Sasa &amp; Okazawa" authorityName="Sasa &amp; Okazawa" box="[151,952,151,178]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="togapeniculus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA7B2824FF6EAFE2FC72FF6C" bold="true" box="[151,952,151,178]" pageId="12" pageNumber="273">
<emphasis id="52CDCC71FA7B2824FF6EAFE2FD16FF6F" bold="true" box="[151,732,151,177]" italics="true" pageId="12" pageNumber="273">Rheocricotopus (Rheocricotopus) togapeniculus</emphasis>
Sasa &amp; Okazawa
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="60061063FA7B2824FF6EAFCFFEF3FF0D" blockId="12.[151,952,151,211]" box="[151,313,186,211]" pageId="12" pageNumber="273">
(
<figureCitation id="F8820CE6FA7B2824FF66AFCFFEFAFF0D" box="[159,304,186,211]" captionStart="FIGURE 12 A – C" captionStartId="12.[151,249,1447,1469]" captionTargetBox="[227,1354,344,1426]" captionTargetId="figure@12.[226,1360,333,1426]" captionTargetPageId="12" captionText="FIGURE 12 A – C. Rhecricotopus (Rheocricotopus) togapeniculus Sasa &amp; Okazawa, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255148/files/figure.png" pageId="12" pageNumber="273">Figs 12 A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA7B2824FF6EAE77FB3AFEC6" box="[151,1264,258,280]" pageId="12" pageNumber="273" type="reference_group">
<paragraph id="60061063FA7B2824FF6EAE77FB3AFEC6" blockId="12.[151,1264,258,280]" box="[151,1264,258,280]" pageId="12" pageNumber="273">
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<taxonomicName id="A7B96BE0FA7B2824FF6EAE77FC63FEC6" ID-CoL="9HT6F" authority="Sasa &amp; Okazawa, 1992: 104" authorityName="Sasa &amp; Okazawa" authorityPageNumber="104" authorityYear="1992" box="[151,937,258,280]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="togapeniculus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA7B2824FF6EAE77FD4AFEC6" box="[151,640,258,280]" italics="true" pageId="12" pageNumber="273">Rheocricotopus (Rheocricotopus) togapeniculus</emphasis>
<treatmentCitation id="E1183672FA7B2824FD7FAE77FC63FEC6" author="Sasa" box="[646,937,258,280]" page="104" pageId="12" pageNumber="273" year="1992">
<bibRefCitation id="04286D92FA7B2824FD7FAE77FCBCFEC6" author="Sasa" box="[646,886,258,280]" pageId="12" pageNumber="273" refString="Sasa, M. &amp; Okazawa, T. (1992) Studies on the chironomid midges (yusurika) of Toga Mura, Toyama. Part 2. The subfamily Orthocladiinae. Research Report from Toyama Prefectural Environment Pollution Research Center, 92 - 204." type="book chapter" year="1992">Sasa &amp; Okazawa, 1992</bibRefCitation>
: 104
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="E1183672FA7B2824FC4DAE77FB26FEC6" author="Ashe" box="[948,1260,258,280]" page="572" pageId="12" pageNumber="273" year="2012">
<bibRefCitation id="04286D92FA7B2824FC4DAE77FB7DFEC6" author="Ashe" box="[948,1207,258,280]" pageId="12" pageNumber="273" refString="Ashe, P. &amp; O'Connor, J. P. (2012) A world catalogue of Chironomidae (Diptera). Part 2 B. Orthocladiinae. Irish Biogeographical Society &amp; National Museum of Ireland, Dublin, pp. 558 - 575." type="book chapter" year="2012">Ashe &amp; O’Connor 2012</bibRefCitation>
: 572
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="34C640EBFA7B2824FF6EAAD2FD86FA02" httpUri="https://zenodo.org/record/255148/files/figure.png" pageId="12" pageNumber="273" targetBox="[227,1354,344,1426]" targetPageId="12">
<paragraph id="60061063FA7B2824FF6EAAD2FD86FA02" blockId="12.[151,1436,1447,1500]" pageId="12" pageNumber="273">
<emphasis id="52CDCC71FA7B2824FF6EAAD2FE93FA63" bold="true" box="[151,345,1447,1469]" pageId="12" pageNumber="273">FIGURE 12 A–C.</emphasis>
<taxonomicName id="A7B96BE0FA7B2824FE99AAD2FC33FA63" authority="Sasa &amp; Okazawa" authorityName="Sasa &amp; Okazawa" box="[352,1017,1447,1469]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="togapeniculus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA7B2824FE99AAD2FE38FA63" box="[352,498,1447,1469]" italics="true" pageId="12" pageNumber="273">Rhecricotopus</emphasis>
(
<emphasis id="52CDCC71FA7B2824FDFBAAD2FD6BFA63" box="[514,673,1447,1469]" italics="true" pageId="12" pageNumber="273">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7B2824FD48AAD2FCF5FA63" box="[689,831,1447,1469]" italics="true" pageId="12" pageNumber="273">togapeniculus</emphasis>
Sasa &amp; Okazawa
</taxonomicName>
, Male imago.
<emphasis id="52CDCC71FA7B2824FB75AAD2FB6FFA62" bold="true" box="[1164,1189,1447,1468]" pageId="12" pageNumber="273">A.</emphasis>
Wing.
<emphasis id="52CDCC71FA7B2824FB08AADDFAC3FA63" bold="true" box="[1265,1289,1448,1469]" pageId="12" pageNumber="273">B.</emphasis>
Hypopygium, dorsal view.
<emphasis id="52CDCC71FA7B2824FEEEAAB3FEFAFA02" bold="true" box="[279,304,1478,1500]" pageId="12" pageNumber="273">C.</emphasis>
Hypopygium, ventral view.
</paragraph>
</caption>
<subSubSection id="28A343E8FA7B2824FF6EA97CFC71F873" pageId="12" pageNumber="273" type="materials_examined">
<paragraph id="60061063FA7B2824FF6EA97CFA5FF9FF" blockId="12.[151,1437,1545,2002]" box="[151,1429,1545,1570]" pageId="12" pageNumber="273">
<emphasis id="52CDCC71FA7B2824FF6EA97CFE4AF9FC" bold="true" box="[151,384,1545,1570]" pageId="12" pageNumber="273">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA7B2824FE7FA97CFE3BF9FC" box="[390,497,1545,1570]" pageId="12" pageNumber="273" type="holotype">Holotype</typeStatus>
, male, Toga-Mura, Toyama Prefecture, Honshu,
<collectingCountry id="18AE50F3FA7B2824FBEBA97FFB9FF9FF" box="[1042,1109,1546,1569]" name="Japan" pageId="12" pageNumber="273">Japan</collectingCountry>
.
<date id="140736A3FA7B2824FB99A97CFB27F9FF" box="[1120,1261,1545,1569]" pageId="12" pageNumber="273" value="1990-04-27">27. IV. 1990</date>
. (No. 181: 29).
</paragraph>
<paragraph id="60061063FA7B2824FF3EA959FBD6F950" blockId="12.[151,1437,1545,2002]" pageId="12" pageNumber="273">
<emphasis id="52CDCC71FA7B2824FF3EA959FE1DF99B" bold="true" box="[199,471,1580,1605]" pageId="12" pageNumber="273">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by laterosternite IX with 6 long setae, anal point with 2 long lateral setae on each side and many basal long microtrichia, inferior volsella double-layered, gonostylus without crista dorsalis.
</paragraph>
<paragraph id="60061063FA7B2824FF3EA9ECFC71F873" blockId="12.[151,1437,1545,2002]" pageId="12" pageNumber="273">
<emphasis id="52CDCC71FA7B2824FF3EA9ECFD51F96C" bold="true" box="[199,667,1689,1714]" pageId="12" pageNumber="273">Additional description and corrections.</emphasis>
Based on the analysis of
<typeStatus id="BF02AEC1FA7B2824FC43A9ECFBD6F96C" box="[954,1052,1689,1714]" pageId="12" pageNumber="273" type="holotype">holotype</typeStatus>
material, the examined characters were consistent with the original description, but some additional characters should be added: Total length / wing length 1.6. Wing length / length of profemur 2.6. Tentorium 130 µm long, 30 µm wide. Palpomeres length (in µm): 30, 65, 133, 138, 240. Palpomere 2 ellipsoid, 3 rectangular, 4 and 5 long and slender, Palpomeres 5/3: 1.8. Laterosternite IX with 6 long setae. Anal point 40 µm long, basal wide 20 µm, peak wide 3 µm, with 2 long lateral setae on each side and many basal long microtrichia. Transverse sternapodeme 120 µm long, with oral projections. Gonocoxite 188 µm long. Gonostylus 70 µm long, without crista dorsalis. Megaseta 15 µm long. HR 2.7, HV 4.0. The hypopygium and wing of
<typeStatus id="BF02AEC1FA7B2824FE12A8E0FD87F870" box="[491,589,1941,1966]" pageId="12" pageNumber="273" type="holotype">holotype</typeStatus>
are re-drawn as in
<figureCitation id="F8820CE6FA7B2824FCDDA8E0FC7DF873" box="[804,951,1941,1966]" captionStart="FIGURE 12 A – C" captionStartId="12.[151,249,1447,1469]" captionTargetBox="[227,1354,344,1426]" captionTargetId="figure@12.[226,1360,333,1426]" captionTargetPageId="12" captionText="FIGURE 12 A – C. Rhecricotopus (Rheocricotopus) togapeniculus Sasa &amp; Okazawa, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255148/files/figure.png" pageId="12" pageNumber="273">Figs 12 A–C</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA7B2824FF3EA8CCFC58F80C" box="[199,914,1977,2002]" pageId="12" pageNumber="273" type="distribution">
<paragraph id="60061063FA7B2824FF3EA8CCFC58F80C" blockId="12.[151,1437,1545,2002]" box="[199,914,1977,2002]" pageId="12" pageNumber="273">
<emphasis id="52CDCC71FA7B2824FF3EA8CCFEAAF80C" bold="true" box="[199,352,1977,2002]" pageId="12" pageNumber="273">Distribution.</emphasis>
The species occurs in
<collectingCountry id="18AE50F3FA7B2824FDA7A8CFFD54F80F" box="[606,670,1978,2001]" name="Japan" pageId="12" pageNumber="273">Japan</collectingCountry>
(Palaearctic Region).
</paragraph>
</subSubSection>
</treatment>
</document>

137
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Sasa
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(
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Sasa, 1994: 51
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<emphasis id="52CDCC71FA7C2823FF6EADC9FE48FD0B" bold="true" box="[151,386,700,725]" pageId="11" pageNumber="272">Material examined.</emphasis>
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, male, Tedori Dam, Ishikawa Prefecture., Honshu,
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, (No. 267: 24).
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<emphasis id="52CDCC71FA7C2823FF3EAC71FE1BFCC3" bold="true" box="[199,465,772,797]" pageId="11" pageNumber="272">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by superior volsella developed and long, like a stick, inferior volsella rounded margins, gonostylus with a small triangularly crista dorsalis.
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<emphasis id="52CDCC71FA7C2823FF3EAC1AFD51FC56" bold="true" box="[199,667,879,904]" pageId="11" pageNumber="272">Additional description and corrections.</emphasis>
Based on the analysis of
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material, the examined characters were consistent with the original description, but some additional characters should be added: Clypeus with 12 setae. Tentorium 165 µm long, 30 µm wide. Dorsocentrals 8. Wing of
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destoryed. Palpomeres length (in µm): 40, 55, 80, 105, 185. Palpomere 2 ellipsoid, 3 and 4 rectangular, 5 long and slender, Palpomeres 5/3: 2.3. Anal point 40 µm long, basal wide 22 µm, peak wide 2.5 µm. Transverse sternapodeme 75 µm long, with weak oral projections. Gonocoxite 178 µm long. Gonostylus 80 µm long, with small crista dorsalis. Megaseta 12 µm long. HR 2.2, HV 2.8. The hypopygium of
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are re-drawn as in
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<emphasis id="52CDCC71FA7C2823FF3EAB19FEAAFB5B" bold="true" box="[199,352,1132,1157]" pageId="11" pageNumber="272">Distribution.</emphasis>
The species occurs in
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<paragraph id="60061063FA7D2822FF6EAEE1FD30FE70" blockId="10.[151,762,404,463]" box="[151,762,404,430]" pageId="10" pageNumber="271">
<heading id="3B4EA70FFA7D2822FF6EAEE1FD30FE70" bold="true" box="[151,762,404,430]" fontSize="11" level="1" pageId="10" pageNumber="271" reason="1">
<taxonomicName id="A7B96BE0FA7D2822FF6EAEE1FD30FE70" ID-CoL="9HV5Q" authority="Sasa" authorityName="Sasa" box="[151,762,404,430]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="tatequintus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA7D2822FF6EAEE1FD30FE70" bold="true" box="[151,762,404,430]" pageId="10" pageNumber="271">
<emphasis id="52CDCC71FA7D2822FF6EAEE1FE9EFE70" bold="true" box="[151,340,404,430]" italics="true" pageId="10" pageNumber="271">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA7D2822FE9CAEE1FDE8FE70" bold="true" box="[357,546,404,430]" italics="true" pageId="10" pageNumber="271">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7D2822FDCAAEE1FD70FE70" bold="true" box="[563,698,404,430]" italics="true" pageId="10" pageNumber="271">tatequintus</emphasis>
Sasa
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="60061063FA7D2822FF6EAEC3FEF3FE11" blockId="10.[151,762,404,463]" box="[151,313,438,463]" pageId="10" pageNumber="271">
(
<figureCitation id="F8820CE6FA7D2822FF66AEC3FEFAFE11" box="[159,304,438,463]" captionStart="FIGURE 10 A – C" captionStartId="10.[151,249,1741,1763]" captionTargetBox="[230,1354,605,1718]" captionTargetId="figure@10.[226,1360,585,1719]" captionTargetPageId="10" captionText="FIGURE 10 A – C. Rhecricotopus (Rheocricotopus) tatequintus Sasa, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255146/files/figure.png" pageId="10" pageNumber="271">Figs 10 A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA7D2822FF6EAE8BFB8DFDCA" box="[151,1095,510,533]" pageId="10" pageNumber="271" type="reference_group">
<paragraph id="60061063FA7D2822FF6EAE8BFB8DFDCA" blockId="10.[151,1095,510,533]" box="[151,1095,510,533]" pageId="10" pageNumber="271">
<treatmentCitationGroup id="40A9374DFA7D2822FF6EAE8BFB8DFDCA" box="[151,1095,510,533]" pageId="10" pageNumber="271">
<taxonomicName id="A7B96BE0FA7D2822FF6EAE8BFCCBFDCA" ID-CoL="9HV5Q" authority="Sasa, 1996: 32" authorityName="Sasa" authorityPageNumber="32" authorityYear="1996" box="[151,769,510,533]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="tatequintus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA7D2822FF6EAE8BFEFCFDCA" box="[151,310,510,532]" italics="true" pageId="10" pageNumber="271">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA7D2822FEBDAE8BFE29FDCA" box="[324,483,510,532]" italics="true" pageId="10" pageNumber="271">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7D2822FE0BAD75FDA9FDCB" box="[498,611,512,533]" italics="true" pageId="10" pageNumber="271">tatequintus</emphasis>
<treatmentCitation id="E1183672FA7D2822FD90AE8BFCCBFDCA" author="Sasa" box="[617,769,510,532]" page="32" pageId="10" pageNumber="271" year="1996">
<bibRefCitation id="04286D92FA7D2822FD90AE8BFD10FDCA" author="Sasa" box="[617,730,510,532]" pageId="10" pageNumber="271" refString="Sasa, M. (1996) Studies on the chironomids in Japan, 1996. Research Report from Toyama Prefectural Environment Pollution Research Center, 12 - 102." type="journal article" year="1996">Sasa, 1996</bibRefCitation>
: 32
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="E1183672FA7D2822FCF5AE8BFB8EFDCA" author="Ashe" box="[780,1092,510,532]" page="572" pageId="10" pageNumber="271" year="2012">
<bibRefCitation id="04286D92FA7D2822FCF5AE8BFBC4FDCA" author="Ashe" box="[780,1038,510,532]" pageId="10" pageNumber="271" refString="Ashe, P. &amp; O'Connor, J. P. (2012) A world catalogue of Chironomidae (Diptera). Part 2 B. Orthocladiinae. Irish Biogeographical Society &amp; National Museum of Ireland, Dublin, pp. 558 - 575." type="book chapter" year="2012">Ashe &amp; O’Connor 2012</bibRefCitation>
: 572
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="34C640EBFA7D2822FF6EA9B8FE66F8DF" httpUri="https://zenodo.org/record/255146/files/figure.png" pageId="10" pageNumber="271" targetBox="[230,1354,605,1718]" targetPageId="10">
<paragraph id="60061063FA7D2822FF6EA9B8FE66F8DF" blockId="10.[151,1436,1740,1793]" pageId="10" pageNumber="271">
<emphasis id="52CDCC71FA7D2822FF6EA9B8FE92F93D" bold="true" box="[151,344,1741,1763]" pageId="10" pageNumber="271">FIGURE 10 A–C.</emphasis>
<taxonomicName id="A7B96BE0FA7D2822FEA6A9B9FC92F93C" authority="Sasa" authorityName="Sasa" box="[351,856,1740,1763]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="271" phylum="Arthropoda" rank="species" species="tatequintus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA7D2822FEA6A9B9FE3BF93C" box="[351,497,1740,1762]" italics="true" pageId="10" pageNumber="271">Rhecricotopus</emphasis>
(
<emphasis id="52CDCC71FA7D2822FDF8A9B9FD6AF93C" box="[513,672,1740,1762]" italics="true" pageId="10" pageNumber="271">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7D2822FD56A9BBFCEAF93D" box="[687,800,1742,1763]" italics="true" pageId="10" pageNumber="271">tatequintus</emphasis>
Sasa
</taxonomicName>
, Male imago.
<emphasis id="52CDCC71FA7D2822FC12A9B8FBCEF93C" bold="true" box="[1003,1028,1741,1762]" pageId="10" pageNumber="271">A.</emphasis>
Wing.
<emphasis id="52CDCC71FA7D2822FBA9A9B8FBADF93C" bold="true" box="[1104,1127,1741,1762]" pageId="10" pageNumber="271">B.</emphasis>
Hypopygium, dorsal view.
<emphasis id="52CDCC71FA7D2822FA7AA9B8FA56F93D" bold="true" box="[1411,1436,1741,1763]" pageId="10" pageNumber="271">C.</emphasis>
Hypopygium, ventral view.
</paragraph>
</caption>
<subSubSection id="28A343E8FA7D2823FF6EA85DFEE7FE72" lastPageId="11" lastPageNumber="272" pageId="10" pageNumber="271" type="materials_examined">
<paragraph id="60061063FA7D2822FF6EA85DFAA6F89F" blockId="10.[151,1436,1832,2037]" box="[151,1388,1832,1857]" pageId="10" pageNumber="271">
<emphasis id="52CDCC71FA7D2822FF6EA85DFE4AF89F" bold="true" box="[151,384,1832,1857]" pageId="10" pageNumber="271">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA7D2822FE7EA85DFE38F89F" box="[391,498,1832,1857]" pageId="10" pageNumber="271" type="holotype">Holotype</typeStatus>
, male, Toyama Prefecture, Honshu,
<collectingCountry id="18AE50F3FA7D2822FC70A85CFC06F89E" box="[905,972,1833,1856]" name="Japan" pageId="10" pageNumber="271">Japan</collectingCountry>
.
<date id="140736A3FA7D2822FC2EA85CFB9FF89F" box="[983,1109,1832,1857]" pageId="10" pageNumber="271" value="1986-04-09">9. IV. 1986</date>
, light trap (No. 284: 58).
</paragraph>
<paragraph id="60061063FA7D2822FF3EA839FE5EF872" blockId="10.[151,1436,1832,2037]" pageId="10" pageNumber="271">
<emphasis id="52CDCC71FA7D2822FF3EA839FE1AF8BB" bold="true" box="[199,464,1868,1893]" pageId="10" pageNumber="271">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by AR 2.55, superior volsella developed and horn-like, inferior volsella extended and acutely angular, anal point sharp pointed and with 4 basal setae.
</paragraph>
<paragraph id="60061063FA7D2823FF3EA8CDFEE7FE72" blockId="10.[151,1436,1832,2037]" lastBlockId="11.[151,1436,151,464]" lastPageId="11" lastPageNumber="272" pageId="10" pageNumber="271">
<emphasis id="52CDCC71FA7D2822FF3EA8CDFD56F80F" bold="true" box="[199,668,1976,2001]" pageId="10" pageNumber="271">Additional description and corrections.</emphasis>
AR should be 2.55. Clypeus with 13 setae. (AR 1.13–1.34, Clypeus with 8, 10 setae in the original description). Based on the analysis of
<typeStatus id="BF02AEC1FA7D2822FC4DA8A9FBDCF82B" box="[948,1046,2012,2037]" pageId="10" pageNumber="271" type="holotype">holotype</typeStatus>
material, the examined characters were consistent with the original description, but some additional characters should be added: Total length / wing length 1.6. Wing length / length of profemur 2.7. Costa extension 50 µm long. Tentorium 185 µm long, 40 µm wide. Palpomeres length (in µm): –, –, 115, 145, 225. Palpomere 2 ellipsoid, 3 and 4 rectangular, 5 long and slender, Palpomeres 5/3: 2.0. Laterosternite IX with 5 long setae. Anal point sharp pointed and 55 µm long, basal wide 25 µm, peak wide 3 µm, with 4 long lateral setae on each side and 4 basal setae. Transverse sternapodeme 110 µm long, with oral projections. Gonocoxite 213 µm long. Gonostylus 105 µm long, with large triangularly rounded costa dorsalis. Megaseta 18 µm long. HR 2.0, HV 3.2. The wing and hypopygium of
<typeStatus id="BF02AEC1FA7C2823FB9CAE1AFB0DFE56" box="[1125,1223,367,392]" pageId="11" pageNumber="272" type="holotype">holotype</typeStatus>
are re-drawn as in
<figureCitation id="F8820CE6FA7C2823FF6EAEE1FEE3FE72" box="[151,297,404,429]" captionStart="FIGURE 10 A – C" captionStartId="10.[151,249,1741,1763]" captionTargetBox="[230,1354,605,1718]" captionTargetId="figure@10.[226,1360,585,1719]" captionTargetPageId="10" captionText="FIGURE 10 A – C. Rhecricotopus (Rheocricotopus) tatequintus Sasa, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255146/files/figure.png" pageId="11" pageNumber="272">Figs 10 A–C</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA7C2823FF3EAEC2FC58FE0E" box="[199,914,439,464]" pageId="11" pageNumber="272" type="distribution">
<paragraph id="60061063FA7C2823FF3EAEC2FC58FE0E" blockId="11.[151,1436,151,464]" box="[199,914,439,464]" pageId="11" pageNumber="272">
<emphasis id="52CDCC71FA7C2823FF3EAEC2FEAAFE0E" bold="true" box="[199,352,439,464]" pageId="11" pageNumber="272">Distribution.</emphasis>
The species occurs in
<collectingCountry id="18AE50F3FA7C2823FDA7AECDFD54FE11" box="[606,670,440,463]" name="Japan" pageId="11" pageNumber="272">Japan</collectingCountry>
(Palaearctic Region).
</paragraph>
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</treatment>
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160
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<paragraph id="60061063FA7E2821FF6EADC1FCE3FD10" blockId="9.[151,809,692,751]" box="[151,809,692,718]" pageId="9" pageNumber="270">
<heading id="3B4EA70FFA7E2821FF6EADC1FCE3FD10" bold="true" box="[151,809,692,718]" fontSize="11" level="1" pageId="9" pageNumber="270" reason="1">
<taxonomicName id="A7B96BE0FA7E2821FF6EADC1FCE3FD10" ID-CoL="4S5ZW" authority="Sasa" authorityName="Sasa" box="[151,809,692,718]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="tamahumeralis" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA7E2821FF6EADC1FCE3FD10" bold="true" box="[151,809,692,718]" pageId="9" pageNumber="270">
<emphasis id="52CDCC71FA7E2821FF6EADC1FE9EFD10" bold="true" box="[151,340,692,718]" italics="true" pageId="9" pageNumber="270">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA7E2821FE9CADC1FDE8FD10" bold="true" box="[357,546,692,718]" italics="true" pageId="9" pageNumber="270">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7E2821FDCAADC1FD23FD10" bold="true" box="[563,745,692,718]" italics="true" pageId="9" pageNumber="270">tamahumeralis</emphasis>
Sasa
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="60061063FA7E2821FF6EADA3FEE1FD31" blockId="9.[151,809,692,751]" box="[151,299,726,751]" pageId="9" pageNumber="270">
(
<figureCitation id="F8820CE6FA7E2821FF66ADA3FEE9FD31" box="[159,291,726,751]" captionStart="FIGURE 9 A – B" captionStartId="9.[151,249,1639,1661]" captionTargetBox="[226,1356,883,1610]" captionTargetId="figure@9.[226,1360,873,1617]" captionTargetPageId="9" captionText="FIGURE 9 A – B. Rhecricotopus (Rheocricotopus) tamahumeralis Sasa, Male imago. A. Hypopygium, dorsal view. B. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255145/files/figure.png" pageId="9" pageNumber="270">Figs 9 A–B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA7E2821FF6EAC6BFBA5FCEA" box="[151,1135,798,820]" pageId="9" pageNumber="270" type="reference_group">
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<treatmentCitationGroup id="40A9374DFA7E2821FF6EAC6BFBA5FCEA" box="[151,1135,798,820]" pageId="9" pageNumber="270">
<taxonomicName id="A7B96BE0FA7E2821FF6EAC6BFCE2FCEA" ID-CoL="4S5ZW" authority="Sasa, 1981: 92" authorityName="Sasa" authorityPageNumber="92" authorityYear="1981" box="[151,808,798,820]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="tamahumeralis" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA7E2821FF6EAC6BFEFCFCEA" box="[151,310,798,820]" italics="true" pageId="9" pageNumber="270">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA7E2821FEBDAC6BFE29FCEA" box="[324,483,798,820]" italics="true" pageId="9" pageNumber="270">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7E2821FE0BAC6BFD40FCEA" box="[498,650,798,820]" italics="true" pageId="9" pageNumber="270">tamahumeralis</emphasis>
<treatmentCitation id="E1183672FA7E2821FD69AC6BFCE2FCEA" author="Sasa" box="[656,808,798,820]" page="92" pageId="9" pageNumber="270" year="1981">
<bibRefCitation id="04286D92FA7E2821FD69AC6BFCCBFCEA" author="Sasa" box="[656,769,798,820]" pageId="9" pageNumber="270" refString="Sasa, M. (1981) Studies on chironomid midges of the Tama River. Part 4. Chironomidae recorded at a winter survey. Research Report from the National Institute for Environmental Studies, 29, 79 - 148." type="journal article" year="1981">Sasa, 1981</bibRefCitation>
: 92
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="E1183672FA7E2821FCCAAC6BFBA1FCEA" author="Ashe" box="[819,1131,798,820]" page="572" pageId="9" pageNumber="270" year="2012">
<bibRefCitation id="04286D92FA7E2821FCCAAC6BFBFCFCEA" author="Ashe" box="[819,1078,798,820]" pageId="9" pageNumber="270" refString="Ashe, P. &amp; O'Connor, J. P. (2012) A world catalogue of Chironomidae (Diptera). Part 2 B. Orthocladiinae. Irish Biogeographical Society &amp; National Museum of Ireland, Dublin, pp. 558 - 575." type="book chapter" year="2012">Ashe &amp; O’Connor 2012</bibRefCitation>
: 572
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="34C640EBFA7E2821FF6EA912FE66F945" httpUri="https://zenodo.org/record/255145/files/figure.png" pageId="9" pageNumber="270" targetBox="[226,1356,883,1610]" targetPageId="9">
<paragraph id="60061063FA7E2821FF6EA912FE66F945" blockId="9.[151,1435,1639,1691]" pageId="9" pageNumber="270">
<emphasis id="52CDCC71FA7E2821FF6EA912FE9CF9A2" bold="true" box="[151,342,1639,1661]" pageId="9" pageNumber="270">FIGURE 9 A–B.</emphasis>
<taxonomicName id="A7B96BE0FA7E2821FE9AA912FC5EF9A3" authority="Sasa" authorityName="Sasa" box="[355,916,1639,1661]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="tamahumeralis" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA7E2821FE9AA912FE3FF9A3" box="[355,501,1639,1661]" italics="true" pageId="9" pageNumber="270">Rhecricotopus</emphasis>
(
<emphasis id="52CDCC71FA7E2821FDF3A912FD63F9A3" box="[522,681,1639,1661]" italics="true" pageId="9" pageNumber="270">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7E2821FD47A912FC9CF9A3" box="[702,854,1639,1661]" italics="true" pageId="9" pageNumber="270">tamahumeralis</emphasis>
Sasa
</taxonomicName>
, Male imago.
<emphasis id="52CDCC71FA7E2821FBC1A912FB9BF9A2" bold="true" box="[1080,1105,1639,1660]" pageId="9" pageNumber="270">A.</emphasis>
Hypopygium, dorsal view.
<emphasis id="52CDCC71FA7E2821FA7DA912FA51F9A2" bold="true" box="[1412,1435,1639,1660]" pageId="9" pageNumber="270">B.</emphasis>
Hypopygium, ventral view.
</paragraph>
</caption>
<subSubSection id="28A343E8FA7E2822FF6EA9BDFBE4FEC2" lastPageId="10" lastPageNumber="271" pageId="9" pageNumber="270" type="materials_examined">
<paragraph id="60061063FA7E2821FF6EA9BDFF24F8DB" blockId="9.[151,1437,1736,2013]" pageId="9" pageNumber="270">
<emphasis id="52CDCC71FA7E2821FF6EA9BDFE49F93F" bold="true" box="[151,387,1736,1761]" pageId="9" pageNumber="270">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA7E2821FE75A9BDFE3DF93F" box="[396,503,1736,1761]" pageId="9" pageNumber="270" type="holotype">Holotype</typeStatus>
, male, Kobotoke, Minamiasagawa River, Tokyo, Honshu,
<collectingCountry id="18AE50F3FA7E2821FB61A9BCFB11F93E" box="[1176,1243,1737,1760]" name="Japan" pageId="9" pageNumber="270">Japan</collectingCountry>
.
<date id="140736A3FA7E2821FB10A9BDFA96F93E" box="[1257,1372,1736,1760]" pageId="9" pageNumber="270" value="1980-01-02">2. I. 1980</date>
, (No. 56: 01).
</paragraph>
<paragraph id="60061063FA7E2821FF3EA865FEF1F8AF" blockId="9.[151,1437,1736,2013]" pageId="9" pageNumber="270">
<emphasis id="52CDCC71FA7E2821FF3EA865FE1AF8F7" bold="true" box="[199,464,1808,1833]" pageId="9" pageNumber="270">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by AR 2.55, inferior volsella extended and acutely angular, anal point and with 8 basal setae, gonostylus median expanded and apical hooked.
</paragraph>
<paragraph id="60061063FA7E2822FF3EA809FBE4FEC2" blockId="9.[151,1437,1736,2013]" lastBlockId="10.[151,1437,151,320]" lastPageId="10" lastPageNumber="271" pageId="9" pageNumber="270">
<emphasis id="52CDCC71FA7E2821FF3EA809FD51F84B" bold="true" box="[199,667,1916,1941]" pageId="9" pageNumber="270">Additional description and corrections.</emphasis>
Based on the analysis of
<typeStatus id="BF02AEC1FA7E2821FC43A809FBD6F84B" box="[954,1052,1916,1941]" pageId="9" pageNumber="270" type="holotype">holotype</typeStatus>
material, the examined characters were consistent with the original description, but some additional characters should be added: Tentorium 130 µm long, 40 µm wide. Palpomeres length (in µm): 38, 50, 90, 108, 140. Palpomere 2 ellipsoid, 3 and 4 rectangular, 5 long and slender, Palpomeres 5/3: 1.5. Laterosternite IX with 4 long setae. Anal point 65 µm long, basal wide 25 µm, peak wide 3 µm, with 8 long lateral setae on each side. Transverse sternapodeme 90 µm long, with oral projections. Gonocoxite 195 µm long. Gonostylus 75 µm long, apical hooked. Megaseta 10 µm long. HR 2.6, HV 3.8. Wing destoryed. The hypopygium of
<typeStatus id="BF02AEC1FA7D2822FD95AE71FD04FEC3" box="[620,718,260,285]" pageId="10" pageNumber="271" type="holotype">holotype</typeStatus>
are re-drawn as in
<figureCitation id="F8820CE6FA7D2822FC5FAE71FBC2FEC2" box="[934,1032,260,285]" captionStart="FIGURE 9 A – B" captionStartId="9.[151,249,1639,1661]" captionTargetBox="[226,1356,883,1610]" captionTargetId="figure@9.[226,1360,873,1617]" captionTargetPageId="9" captionText="FIGURE 9 A – B. Rhecricotopus (Rheocricotopus) tamahumeralis Sasa, Male imago. A. Hypopygium, dorsal view. B. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255145/files/figure.png" pageId="10" pageNumber="271">Figs 9 A</figureCitation>
–C.
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA7D2822FF3EAE52FC58FE9E" box="[199,914,295,320]" pageId="10" pageNumber="271" type="distribution">
<paragraph id="60061063FA7D2822FF3EAE52FC58FE9E" blockId="10.[151,1437,151,320]" box="[199,914,295,320]" pageId="10" pageNumber="271">
<emphasis id="52CDCC71FA7D2822FF3EAE52FEAAFE9E" bold="true" box="[199,352,295,320]" pageId="10" pageNumber="271">Distribution.</emphasis>
The species occurs in
<collectingCountry id="18AE50F3FA7D2822FDA7AE5DFD54FEE1" box="[606,670,296,319]" name="Japan" pageId="10" pageNumber="271">Japan</collectingCountry>
(Palaearctic Region).
</paragraph>
</subSubSection>
</treatment>
</document>

162
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<paragraph id="60061063FA7F2820FF6EAD32FC9DFDBC" blockId="8.[151,855,583,643]" box="[151,855,583,610]" pageId="8" pageNumber="269">
<heading id="3B4EA70FFA7F2820FF6EAD32FC9DFDBC" bold="true" box="[151,855,583,610]" fontSize="11" level="1" pageId="8" pageNumber="269" reason="1">
<taxonomicName id="A7B96BE0FA7F2820FF6EAD32FC9DFDBC" authority="Sasa" authorityName="Sasa" box="[151,855,583,610]" class="Insecta" family="Chironomidae" genus="Rheocricotopus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="shoufukusecundus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA7F2820FF6EAD32FC9DFDBC" bold="true" box="[151,855,583,610]" pageId="8" pageNumber="269">
<emphasis id="52CDCC71FA7F2820FF6EAD32FE9EFDBF" bold="true" box="[151,340,583,609]" italics="true" pageId="8" pageNumber="269">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA7F2820FE9CAD32FDE8FDBF" bold="true" box="[357,546,583,609]" italics="true" pageId="8" pageNumber="269">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7F2820FDCAAD32FCDDFDBF" bold="true" box="[563,791,583,609]" italics="true" pageId="8" pageNumber="269">shoufukusecundus</emphasis>
Sasa
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="60061063FA7F2820FF6EAD1FFEE1FD5D" blockId="8.[151,855,583,643]" box="[151,299,618,643]" pageId="8" pageNumber="269">
(
<figureCitation id="F8820CE6FA7F2820FF66AD1FFEE8FD5D" box="[159,290,618,643]" captionStart="FIGURE 8 A – C" captionStartId="8.[151,249,1972,1994]" captionTargetBox="[226,1355,840,1950]" captionTargetId="figure@8.[226,1360,829,1950]" captionTargetPageId="8" captionText="FIGURE 8 A – C. Rhecricotopus (Rheocricotopus) shoufukusecundus Sasa, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255144/files/figure.png" pageId="8" pageNumber="269">Figs 8 A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA7F2820FF6EADC7FB45FD16" box="[151,1167,690,712]" pageId="8" pageNumber="269" type="reference_group">
<paragraph id="60061063FA7F2820FF6EADC7FB45FD16" blockId="8.[151,1167,690,712]" box="[151,1167,690,712]" pageId="8" pageNumber="269">
<treatmentCitationGroup id="40A9374DFA7F2820FF6EADC7FB45FD16" box="[151,1167,690,712]" pageId="8" pageNumber="269">
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<emphasis id="52CDCC71FA7F2820FF6EADC7FEFCFD16" box="[151,310,690,712]" italics="true" pageId="8" pageNumber="269">Rheocricotopus</emphasis>
(
<emphasis id="52CDCC71FA7F2820FEBDADC7FE29FD16" box="[324,483,690,712]" italics="true" pageId="8" pageNumber="269">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7F2820FE0BADC7FD60FD16" box="[498,682,690,712]" italics="true" pageId="8" pageNumber="269">shoufukusecundus</emphasis>
<treatmentCitation id="E1183672FA7F2820FD48ADC7FC82FD16" author="Sasa" box="[689,840,690,712]" page="30" pageId="8" pageNumber="269" year="1998">
<bibRefCitation id="04286D92FA7F2820FD48ADC7FCEBFD16" author="Sasa" box="[689,801,690,712]" pageId="8" pageNumber="269" refString="Sasa, M. (1998) Chironomidae of Japan. - List of species recorded, and supplemental keys for identification. Research Report Institute of Environmental and Welfare Studies, Sunaba, Kurobe-shi, Japan, 1998, 156 pp." type="book" year="1998">Sasa, 1998</bibRefCitation>
: 30
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="E1183672FA7F2820FCAAADC7FB41FD16" author="Ashe" box="[851,1163,690,712]" page="572" pageId="8" pageNumber="269" year="2012">
<bibRefCitation id="04286D92FA7F2820FCAAADC7FB9CFD16" author="Ashe" box="[851,1110,690,712]" pageId="8" pageNumber="269" refString="Ashe, P. &amp; O'Connor, J. P. (2012) A world catalogue of Chironomidae (Diptera). Part 2 B. Orthocladiinae. Irish Biogeographical Society &amp; National Museum of Ireland, Dublin, pp. 558 - 575." type="book chapter" year="2012">Ashe &amp; O’Connor 2012</bibRefCitation>
: 572
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA7F2821FF6EAD9AFB36FDE2" lastPageId="9" lastPageNumber="270" pageId="8" pageNumber="269" type="materials_examined">
<paragraph id="60061063FA7F2820FF6EAD9AFABBFCD6" blockId="8.[151,1393,751,776]" box="[151,1393,751,776]" pageId="8" pageNumber="269">
<emphasis id="52CDCC71FA7F2820FF6EAD9AFE4AFCD6" bold="true" box="[151,384,751,776]" pageId="8" pageNumber="269">Material examined.</emphasis>
<typeStatus id="BF02AEC1FA7F2820FE7EAD9AFE38FCD6" box="[391,498,751,776]" pageId="8" pageNumber="269" type="holotype">Holotype</typeStatus>
, male, Kurobe, Toyama Prefecture, Honshu,
<collectingCountry id="18AE50F3FA7F2820FC12AD85FBE4FCD9" box="[1003,1070,752,775]" name="Japan" pageId="8" pageNumber="269">Japan</collectingCountry>
.
<date id="140736A3FA7F2820FBC0AD85FB0DFCD6" box="[1081,1223,752,776]" pageId="8" pageNumber="269" value="1997-03-13">13. III. 1997</date>
, (No. 329: 28).
</paragraph>
<caption id="34C640EBFA7F2820FF6EA8C1FDC2F836" httpUri="https://zenodo.org/record/255144/files/figure.png" pageId="8" pageNumber="269" targetBox="[226,1355,840,1950]" targetPageId="8">
<paragraph id="60061063FA7F2820FF6EA8C1FDC2F836" blockId="8.[151,1436,1972,2024]" pageId="8" pageNumber="269">
<emphasis id="52CDCC71FA7F2820FF6EA8C1FE9BF814" bold="true" box="[151,337,1972,1994]" pageId="8" pageNumber="269">FIGURE 8 A–C.</emphasis>
<taxonomicName id="A7B96BE0FA7F2820FEA3A8C1FC69F814" authority="Sasa" authorityName="Sasa" box="[346,931,1972,1994]" class="Insecta" family="Chironomidae" genus="Rhecricotopus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="shoufukusecundus" subGenus="Rheocricotopus">
<emphasis id="52CDCC71FA7F2820FEA3A8C1FE26F814" box="[346,492,1972,1994]" italics="true" pageId="8" pageNumber="269">Rhecricotopus</emphasis>
(
<emphasis id="52CDCC71FA7F2820FE06A8C1FD54F814" box="[511,670,1972,1994]" italics="true" pageId="8" pageNumber="269">Rheocricotopus</emphasis>
)
<emphasis id="52CDCC71FA7F2820FD49A8C1FCA2F814" box="[688,872,1972,1994]" italics="true" pageId="8" pageNumber="269">shoufukusecundus</emphasis>
Sasa
</taxonomicName>
, Male imago.
<emphasis id="52CDCC71FA7F2820FBC4A8C1FB9CF817" bold="true" box="[1085,1110,1972,1993]" pageId="8" pageNumber="269">A.</emphasis>
Wing.
<emphasis id="52CDCC71FA7F2820FB5EA8C1FB75F817" bold="true" box="[1191,1215,1972,1993]" pageId="8" pageNumber="269">B.</emphasis>
Hypopygium, dorsal view.
<emphasis id="52CDCC71FA7F2820FF2AA8A7FF21F836" bold="true" box="[211,235,2002,2024]" pageId="8" pageNumber="269">C.</emphasis>
Hypopygium, ventral view.
</paragraph>
</caption>
<paragraph id="60061063FA7E2821FF3EAFE2FD0AFF29" blockId="9.[151,1437,151,608]" pageId="9" pageNumber="270">
<emphasis id="52CDCC71FA7E2821FF3EAFE2FE1BFF6E" bold="true" box="[199,465,151,176]" pageId="9" pageNumber="270">Diagnostic characters.</emphasis>
The species can be separated from other East Asian members of the genus by superior volsella absent, inferior volsella expanded apically and somewhat foot-shaped. Anal point sharp pointed and with 10 basal setae. Gonocoxite with many long setae.
</paragraph>
<paragraph id="60061063FA7E2821FF3EAE71FB36FDE2" blockId="9.[151,1437,151,608]" pageId="9" pageNumber="270">
<emphasis id="52CDCC71FA7E2821FF3EAE71FD51FEC3" bold="true" box="[199,667,260,285]" pageId="9" pageNumber="270">Additional description and corrections.</emphasis>
Based on the analysis of
<typeStatus id="BF02AEC1FA7E2821FC43AE71FBD6FEC3" box="[954,1052,260,285]" pageId="9" pageNumber="270" type="holotype">holotype</typeStatus>
material, the examined characters were consistent with the original description, but some additional characters should be added: Total length / wing length 1.6. Wing length / length of profemur 3.0. Outer verticals 2, postorbitals 4. Clypeus with 12 setae. Tentorium 185 µm long, 40 µm wide. Dorsocentrals 35, scutellars 13. Squama with 33 setae. Palpomeres length (in µm): –, 55, 110, 133, 158. Palpomere 2 ellipsoid, 3 and 4 rectangular, 5 long and slender, Palpomeres 5/3: 1.4. Laterosternite IX with 10 long setae. Anal point sharp pointed and 45 µm long, basal wide 13 µm, peak wide 3 µm, with 2 long lateral setae on each side and 9 basal setae. Transverse sternapodeme 188 µm long, with obvious oral projections. Gonocoxite 275 µm long. Gonostylus 58 µm long, with triangularly rounded costa dorsalis. Megaseta 13 µm long. HR 4.8, HV 7.4. The wing and hypopygium of
<typeStatus id="BF02AEC1FA7E2821FCC3AD51FC56FDE3" box="[826,924,548,573]" pageId="9" pageNumber="270" type="holotype">holotype</typeStatus>
are re-drawn as in
<figureCitation id="F8820CE6FA7E2821FB8AAD51FB32FDE2" box="[1139,1272,548,573]" captionStart="FIGURE 8 A – C" captionStartId="8.[151,249,1972,1994]" captionTargetBox="[226,1355,840,1950]" captionTargetId="figure@8.[226,1360,829,1950]" captionTargetPageId="8" captionText="FIGURE 8 A – C. Rhecricotopus (Rheocricotopus) shoufukusecundus Sasa, Male imago. A. Wing. B. Hypopygium, dorsal view. C. Hypopygium, ventral view." httpUri="https://zenodo.org/record/255144/files/figure.png" pageId="9" pageNumber="270">Figs 8 A–C</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="28A343E8FA7E2821FF3EAD32FC58FDBE" box="[199,914,583,608]" pageId="9" pageNumber="270" type="distribution">
<paragraph id="60061063FA7E2821FF3EAD32FC58FDBE" blockId="9.[151,1437,151,608]" box="[199,914,583,608]" pageId="9" pageNumber="270">
<emphasis id="52CDCC71FA7E2821FF3EAD32FEAAFDBE" bold="true" box="[199,352,583,608]" pageId="9" pageNumber="270">Distribution.</emphasis>
The species occurs in
<collectingCountry id="18AE50F3FA7E2821FDA7AD3DFD54FD81" box="[606,670,584,607]" name="Japan" pageId="9" pageNumber="270">Japan</collectingCountry>
(Palaearctic Region).
</paragraph>
</subSubSection>
</treatment>
</document>

431
data/E8/10/B9/E810B924E00E5676A21F5D5ED71C4EEC.xml Normal file
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@ -0,0 +1,431 @@
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<mods:title>Molecular phylogeny of the riffle beetle genus Hexanchorus revealed a presence of a new genus (Coleoptera: Elmidae)</mods:title>
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<mods:namePart>Linsky, Marek</mods:namePart>
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<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
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<mods:namePart>Ciamporova-Zatovicova, Zuzana</mods:namePart>
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<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
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<mods:namePart>Lassova, Kristina</mods:namePart>
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<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia</mods:affiliation>
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<mods:namePart>Ciampor Jr, Fedor</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6269-3592</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
<mods:nameIdentifier type="email">f.ciampor@savba.sk</mods:nameIdentifier>
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<paragraph pageId="0" pageNumber="575">
3.2.6.
<taxonomicName LSID="https://zoobank.org/4BA8996F-8FA8-4A73-A7C0-3A20FAF7CE92" authority="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr, 2022" authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rumilara paterna" order="Coleoptera" pageId="0" pageNumber="575" rank="species" species="paterna" status="sp. nov.">Rumilara paterna</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="575">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="description">
<paragraph pageId="0" pageNumber="575">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575">Figs 2d</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">, 3d</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Rumilara gen. nov. and Hexanchorus: a R. obscura sp. nov.; b R. suppressa sp. nov.; c R. riberai sp. nov.; d R. paterna sp. nov.; e R. leleupi comb. nov.; f H. mcdiarmidi; g H. virilis; h H. shannoni; i H. angeli; j H. rostratus; k H. cordillierae; l H. onorei; m H. tarsalis; n H. usitatus; o H. gracilipes; p H. caraibus; q H. crinitus; r H. dimorphus; s H. emarginatus; t H. browni. (a, k based on dissected antennae; b-j, l-r based on photographs, not to scale, setae omitted)." figureDoi="10.3897/asp.80.e84013.figure5" httpUri="https://binary.pensoft.net/fig/770428" pageId="0" pageNumber="575">, 5d</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronota of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575">, 6d</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ventral habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male paratype; b R. suppressa sp. nov. male paratype; c R. riberai sp. nov. female paratype; d R. paterna sp. nov. paratype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure7" httpUri="https://binary.pensoft.net/fig/770430" pageId="0" pageNumber="575">, 7d</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Abdomens of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure8" httpUri="https://binary.pensoft.net/fig/770431" pageId="0" pageNumber="575">, 8d</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larvae of Rumilara gen. nov. species: a R. obscura sp. nov., dorsal; b R. suppressa sp. nov., dorsal; c R. riberai sp. nov., dorsal; d R. paterna sp. nov., dorsal; e R. obscura sp. nov., ventral; f R. suppressa sp. nov., ventral. Scale: 1 mm." figureDoi="10.3897/asp.80.e84013.figure9" httpUri="https://binary.pensoft.net/fig/770432" pageId="0" pageNumber="575">, 9d</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Rumilara paterna sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d male sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure14" httpUri="https://binary.pensoft.net/fig/770437" pageId="0" pageNumber="575">, 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="materials_examined">
<paragraph pageId="0" pageNumber="575">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="575">
<emphasis bold="true" pageId="0" pageNumber="575">
<emphasis italics="true" pageId="0" pageNumber="575">Type material</emphasis>
:
<materialsCitation collectingDate="2013-08-24" country="Ecuador" elevation="1299" latitude="-3.1439445" location="Indanza" longLatPrecision="1" longitude="-78.53625" specimenCount="♂" stateProvince="Morona-Santiago prov." typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
<specimenCount>♂</specimenCount>
(PUCE) &quot;
<collectingCountry name="Ecuador">Ecuador</collectingCountry>
,
<collectingRegion country="Ecuador" name="Morona-Santiago">Morona-Santiago prov.</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:E810B924E00E5676A21F5D5ED71C4EEC:C3A3C831D193EA4F8B8149306660EE69" country="Ecuador" latitude="-3.1439445" longLatPrecision="1" longitude="-78.53625" name="Indanza" stateProvince="Morona-Santiago prov.">Indanza</location>
env.,,
<geoCoordinate degrees="03" direction="south" minutes="08" orientation="latitude" precision="1" seconds="38.2" value="-3.1439445">03°08′38.2″ S</geoCoordinate>
,
<geoCoordinate degrees="78" direction="west" minutes="32" orientation="longitude" precision="1" seconds="10.5" value="-78.53625">78°32′10.5″ W</geoCoordinate>
,
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.299" unit="m" value="1299.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.299" unit="m" value="1299.0">1299m</quantity>
a.s.l.
</elevation>
,
<collectingDate value="2013-08-24">24.8.2013</collectingDate>
, stream ca
<quantity metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">
<elevation metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">5m</elevation>
</quantity>
wide, with gravel, boulders, leaf debris, submerged wood,
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp;
<normalizedToken originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
</emphasis>
;
<materialsCitation collectingDate="2013-08-17" collectingDateMax="2013-08-25" collectingDateMin="2013-08-17" collectorName="Ciampor Jr, Ciamporova-Zatovicova" country="Ecuador" elevation="698" latitude="-3.0874166" location="Cascada la Encanada" longLatPrecision="1" longitude="-78.41" municipality="Santa Rosa de Mamanguy" specimenCount="2" specimenCount-larva="1" stateProvince="Morona-Santiago prov." typeStatus="Paratype">
<emphasis bold="true" pageId="0" pageNumber="575">
<typeStatus>Paratype</typeStatus>
</emphasis>
<specimenCount type="generic">1 ex</specimenCount>
(CCB) &quot;
<collectingCountry name="Ecuador">Ecuador</collectingCountry>
,
<collectingRegion country="Ecuador" name="Morona-Santiago">Morona-Santiago prov.</collectingRegion>
,
<collectingMunicipality>Santa Rosa de Mamanguy</collectingMunicipality>
env.,
<location LSID="urn:lsid:plazi:treatment:E810B924E00E5676A21F5D5ED71C4EEC:DAC632C7B24553185168171B44B2C698" country="Ecuador" latitude="-3.0874166" longLatPrecision="1" longitude="-78.41" municipality="Santa Rosa de Mamanguy" name="Cascada la Encanada" stateProvince="Morona-Santiago prov.">
Cascada la
<normalizedToken originalValue="Encañada">Encanada</normalizedToken>
</location>
,
<geoCoordinate degrees="03" direction="south" minutes="05" orientation="latitude" precision="1" seconds="14.7" value="-3.0874166">03°05′14.7″ S</geoCoordinate>
,
<geoCoordinate degrees="78" direction="west" minutes="24" orientation="longitude" precision="1" seconds="36.0" value="-78.41">78°24′36.0″ W</geoCoordinate>
,
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" unit="m" value="698.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" unit="m" value="698.0">698m</quantity>
a.s.l.
</elevation>
,
<collectingDate value="2013-08-25">25.8.2013</collectingDate>
, stream ca
<quantity metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">
<elevation metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2-3m</elevation>
</quantity>
wide under waterfall, with clay, gravel, stones, submerged wood,
<collectorName>
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
Jr
</collectorName>
&amp;
<normalizedToken originalValue="Čiamporová-Zaťovičová">
<collectorName>Ciamporova-Zatovicova</collectorName>
</normalizedToken>
lgt.&quot;
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">
<typeStatus isOtherTypeStatus="true">Other material</typeStatus>
</emphasis>
:
<specimenCount type="larva">1 larva</specimenCount>
&quot;
<collectingCountry name="Ecuador">Ecuador</collectingCountry>
,
<collectingRegion country="Ecuador" name="Napo">Napo prov.</collectingRegion>
, road to Coca,
<geoCoordinate degrees="00" direction="south" minutes="43" orientation="latitude" precision="1" seconds="39.6" value="-0.7276666">00°43′39.6″ S</geoCoordinate>
,
<geoCoordinate degrees="77" direction="west" minutes="45" orientation="longitude" precision="1" seconds="56.1" value="-77.76559">77°45′56.1″ W</geoCoordinate>
,
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.129" unit="m" value="1129.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.129" unit="m" value="1129.0">1129m</quantity>
a.s.l.
</elevation>
,
<collectingDate value="2013-08-17">17.8.2013</collectingDate>
, small waterfall,
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp;
<normalizedToken originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
;
<materialsCitation collectingDate="2013-08-24" country="Ecuador" elevation="979" latitude="-3.0491667" location="Rio Crusado" longLatPrecision="1" longitude="-78.50089" specimenCount="2" specimenCount-larva="2" stateProvince="Morona-Santiago prov." typeStatus="paratype">
<specimenCount type="larva">2 larvae</specimenCount>
&quot;
<collectingCountry name="Ecuador">Ecuador</collectingCountry>
,
<collectingRegion country="Ecuador" name="Morona-Santiago">Morona-Santiago prov.</collectingRegion>
, tributary of
<location LSID="urn:lsid:plazi:treatment:E810B924E00E5676A21F5D5ED71C4EEC:95B3477B895E4CE199A4B4EDFDA86623" country="Ecuador" latitude="-3.0491667" longLatPrecision="1" longitude="-78.50089" name="Rio Crusado" stateProvince="Morona-Santiago prov.">
<normalizedToken originalValue="Río">Rio</normalizedToken>
Crusado
</location>
,
<geoCoordinate degrees="03" direction="south" minutes="02" orientation="latitude" precision="1" seconds="57.0" value="-3.0491667">03°02′57.0″ S</geoCoordinate>
,
<geoCoordinate degrees="78" direction="west" minutes="30" orientation="longitude" precision="1" seconds="03.2" value="-78.50089">78°30′03.2″ W</geoCoordinate>
,
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.79" unit="m" value="979.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.79" unit="m" value="979.0">979m</quantity>
a.s.l.
</elevation>
,
<collectingDate value="2013-08-24">24.8.2013</collectingDate>
, stream ca
<quantity metricMagnitude="0" metricUnit="m" metricValue="2.0" unit="m" value="2.0">
<elevation metricMagnitude="0" metricUnit="m" metricValue="2.0" unit="m" value="2.0">2m</elevation>
</quantity>
wide, in forest, with gravel, stones, leaf debris, submerged wood with moss,
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp;
<normalizedToken originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
;
<materialsCitation collectingDate="2013-08-25" country="Ecuador" elevation="698" latitude="-3.0874166" location="Cascada la Encanada" longLatPrecision="1" longitude="-78.41" municipality="Santa Rosa de Mamanguy" specimenCount="1" specimenCount-larva="1" stateProvince="Morona-Santiago prov." typeStatus="paratype">
<specimenCount type="larva">1 larva</specimenCount>
&quot;
<collectingCountry name="Ecuador">Ecuador</collectingCountry>
,
<collectingRegion country="Ecuador" name="Morona-Santiago">Morona-Santiago prov.</collectingRegion>
,
<collectingMunicipality>Santa Rosa de Mamanguy</collectingMunicipality>
env.,
<location LSID="urn:lsid:plazi:treatment:E810B924E00E5676A21F5D5ED71C4EEC:FA47464094779A3B31C10291A9829D60" country="Ecuador" latitude="-3.0874166" longLatPrecision="1" longitude="-78.41" municipality="Santa Rosa de Mamanguy" name="Cascada la Encanada" stateProvince="Morona-Santiago prov.">
Cascada la
<normalizedToken originalValue="Encañada">Encanada</normalizedToken>
</location>
,
<geoCoordinate degrees="03" direction="south" minutes="05" orientation="latitude" precision="1" seconds="14.7" value="-3.0874166">03°05′14.7″ S</geoCoordinate>
,
<geoCoordinate degrees="78" direction="west" minutes="24" orientation="longitude" precision="1" seconds="36.0" value="-78.41">78°24′36.0″ W</geoCoordinate>
,
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" unit="m" value="698.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" unit="m" value="698.0">698m</quantity>
a.s.l.
</elevation>
,
<collectingDate value="2013-08-25">25.8.2013</collectingDate>
, stream ca
<quantity metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">
<elevation metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2-3m</elevation>
</quantity>
wide under waterfall, with clay, gravel, stones, submerged wood,
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp;
<normalizedToken originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="adult diagnosis">
<paragraph pageId="0" pageNumber="575">Adult diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="575">The species can be distinguished by a combination of following characters: 1) head and clypeus fully clothed by pubescence; 2) femora fully pubescent; 3) elytra with dense pubescence, not arranged in stripes; 4) aedeagus with rod-like sclerite; 5) parameres moderately asymmetrical.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="description">
<paragraph pageId="0" pageNumber="575">Adult diagnostic description.</paragraph>
<paragraph pageId="0" pageNumber="575">
Male.
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Colour</emphasis>
: Body (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575">2d</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">3d</figureCitation>
) dark brown; coxae, trochanters, femora, tibiae, and tarsi brown; tarsal claws pale brown with reddish tincture.
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Pubescence</emphasis>
: Head and clypeus fully clothed with short, prone pubescence, most densely set behind eyes, and with numerous slightly longer, curved, semi-erect, hair-like setae. Pronotum (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronota of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575">6d</figureCitation>
) covered by a short, moderately dense, prone pubescence, and with moderately long, curved, semi-erect, hair-like setae. Elytra with densely arranged, short, recumbent pubescence, and numerous moderately long, curved, semi-erect, hair-like setae. Femora, pro- and metatibiae densely covered with moderately long, recumbent, hair-like setae; mesotibiae with pubescence only at extreme base. Metaventrite densely clothed with short, prone pubescence.
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Surface</emphasis>
: Head and clypeus sparsely microreticulate, concealed by pubescence. Pronotum densely punctate, partly concealed by pubescence. Elytra with rows of punctures shallowly impressed; intervals nearly flat. Prosternal process (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ventral habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male paratype; b R. suppressa sp. nov. male paratype; c R. riberai sp. nov. female paratype; d R. paterna sp. nov. paratype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure7" httpUri="https://binary.pensoft.net/fig/770430" pageId="0" pageNumber="575">7d</figureCitation>
) subparallel between procoxae, then subtriangular with rounded apex.
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Aedeagus</emphasis>
(Figs
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Rumilara paterna sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d male sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure14" httpUri="https://binary.pensoft.net/fig/770437" pageId="0" pageNumber="575">14a-c</figureCitation>
) with long, dorsal, rod-like sclerite on penis; parameres asymmetrical, left one slightly longer. - Female unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="description">
<paragraph pageId="0" pageNumber="575">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="575">♂ - CL: 2.67 mm; PL: 0.61 mm; PW: 0.75 mm; EL: 2.07 mm; EW: 0.83 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="variation">
<paragraph pageId="0" pageNumber="575">Variation.</paragraph>
<paragraph pageId="0" pageNumber="575">No significant differences were observed between the two known specimens.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="etymology">
<paragraph pageId="0" pageNumber="575">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="575">
From Latin
<emphasis bold="true" pageId="0" pageNumber="575">paternus</emphasis>
(of or connected with
<normalizedToken originalValue="one’s">one's</normalizedToken>
forefathers, ancestral) due to its similarity with the genus
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="larva">
<paragraph pageId="0" pageNumber="575">Larva</paragraph>
<paragraph pageId="0" pageNumber="575">
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larvae of Rumilara gen. nov. species: a R. obscura sp. nov., dorsal; b R. suppressa sp. nov., dorsal; c R. riberai sp. nov., dorsal; d R. paterna sp. nov., dorsal; e R. obscura sp. nov., ventral; f R. suppressa sp. nov., ventral. Scale: 1 mm." figureDoi="10.3897/asp.80.e84013.figure9" httpUri="https://binary.pensoft.net/fig/770432" pageId="0" pageNumber="575">9d</figureCitation>
) with length 3.48-3.55 mm, greatest width 0.99-1.10 mm. Pleurites on abdominal segments 1-5. Abdominal segments 1-7 explanate. Surface around the frontal pair of pronotal gibbosities paler than the rest of pronotum. Tergum of abdominal segments 1-7 with a pair of distinct, longitudinally arranged small gibbosities.
</paragraph>
<caption doi="10.3897/asp.80.e84013.figure16" httpUri="https://binary.pensoft.net/fig/770439" pageId="0" pageNumber="575" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="575">
<emphasis bold="true" pageId="0" pageNumber="575">Figure 16.</emphasis>
Sampling sites of
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
in Ecuador:
<emphasis bold="true" pageId="0" pageNumber="575">a</emphasis>
Cotopaxi prov., Otonga env.,
<normalizedToken originalValue="Río">Rio</normalizedToken>
Esmeralda;
<emphasis bold="true" pageId="0" pageNumber="575">b</emphasis>
Cotopaxi prov., Artos env., stream above confluence with
<normalizedToken originalValue="Río">Rio</normalizedToken>
Esmeralda;
<emphasis bold="true" pageId="0" pageNumber="575">c</emphasis>
Santo Domingo de los Tsachilas prov., Otongachi env., tributary of
<normalizedToken originalValue="Río">Rio</normalizedToken>
Toachi;
<emphasis bold="true" pageId="0" pageNumber="575">d</emphasis>
Santo Domingo de los Tsachilas prov., thin water layer on rock, Otongachi env.;
<emphasis bold="true" pageId="0" pageNumber="575">e</emphasis>
Morona-Santiago prov., small stream near Indanza;
<emphasis bold="true" pageId="0" pageNumber="575">f</emphasis>
Morona-Santiago prov., Santa Rosa de Mamanguy env., Cascada la
<normalizedToken originalValue="Encañada">Encanada</normalizedToken>
;
<emphasis bold="true" pageId="0" pageNumber="575">g</emphasis>
fast-flowing stream in Napo prov.;
<emphasis bold="true" pageId="0" pageNumber="575">h</emphasis>
Morona-Santiago prov., tributary of
<normalizedToken originalValue="Río">Rio</normalizedToken>
Tinajillas;
<emphasis bold="true" pageId="0" pageNumber="575">i</emphasis>
Morona-Santiago prov., tributary of
<normalizedToken originalValue="Río">Rio</normalizedToken>
Tinajillas; (localities of
<taxonomicName lsidName="R. obscura" pageId="0" pageNumber="575" rank="species" species="obscura">
<emphasis italics="true" pageId="0" pageNumber="575">R. obscura</emphasis>
</taxonomicName>
-
<emphasis bold="true" pageId="0" pageNumber="575">a</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="575">b</emphasis>
;
<taxonomicName lsidName="R. riberai" pageId="0" pageNumber="575" rank="species" species="riberai">
<emphasis italics="true" pageId="0" pageNumber="575">R. riberai</emphasis>
</taxonomicName>
-
<emphasis bold="true" pageId="0" pageNumber="575">c</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="575">d</emphasis>
;
<taxonomicName lsidName="R. paterna" pageId="0" pageNumber="575" rank="species" species="paterna">
<emphasis italics="true" pageId="0" pageNumber="575">R. paterna</emphasis>
</taxonomicName>
-
<emphasis bold="true" pageId="0" pageNumber="575">e</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="575">f</emphasis>
;
<taxonomicName lsidName="R. suppressa" pageId="0" pageNumber="575" rank="species" species="suppressa">
<emphasis italics="true" pageId="0" pageNumber="575">R. suppressa</emphasis>
</taxonomicName>
-
<emphasis bold="true" pageId="0" pageNumber="575">g</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="575">i</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="distribution">
<paragraph pageId="0" pageNumber="575">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="575">
Known from one locality in Napo Province and three localities in Morona-Santiago Province (Figs
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sampling sites of Rumilara gen. nov. in Ecuador: a Cotopaxi prov., Otonga env., Rio Esmeralda; b Cotopaxi prov., Artos env., stream above confluence with Rio Esmeralda; c Santo Domingo de los Tsachilas prov., Otongachi env., tributary of Rio Toachi; d Santo Domingo de los Tsachilas prov., thin water layer on rock, Otongachi env.; e Morona-Santiago prov., small stream near Indanza; f Morona-Santiago prov., Santa Rosa de Mamanguy env., Cascada la Encanada; g fast-flowing stream in Napo prov.; h Morona-Santiago prov., tributary of Rio Tinajillas; i Morona-Santiago prov., tributary of Rio Tinajillas; (localities of R. obscura - a, b; R. riberai - c, d; R. paterna - e, f; R. suppressa - g - i)." figureDoi="10.3897/asp.80.e84013.figure16" httpUri="https://binary.pensoft.net/fig/770439" pageId="0" pageNumber="575">16e-f</figureCitation>
).
</paragraph>
<caption doi="10.3897/asp.80.e84013.figure17" httpUri="https://binary.pensoft.net/fig/770440" pageId="0" pageNumber="575" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="575">
<emphasis bold="true" pageId="0" pageNumber="575">Figure 17.</emphasis>
Altitudinal distribution of
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
in Ecuador (with one record from Colombia); circles -
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
, triangles -
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
Hollow symbols represent material not examined in this study (Modified from Google Earth).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="remarks">
<paragraph pageId="0" pageNumber="575">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="575">Only two specimens are known, one of them of unidentified sex, with abdomen lacking last two segments.</paragraph>
</subSubSection>
</treatment>
</document>

222
data/E8/11/10/E811100CF6396EDA884FA73A3C68B1EF.xml Normal file
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<treatment ID-GBIF-Taxon="152034794" LSID="urn:lsid:plazi:treatment:E811100CF6396EDA884FA73A3C68B1EF" httpUri="http://treatment.plazi.org/id/E811100CF6396EDA884FA73A3C68B1EF" lastPageId="9" lastPageNumber="128" pageId="7" pageNumber="126">
<subSubSection pageId="7" pageNumber="126" type="nomenclature">
<paragraph pageId="7" pageNumber="126">
<taxonomicName LSID="http://species-id.net/wiki/Dexiogyia_angustiventris" authority="Casey, 1894" authorityName="Casey" authorityYear="1894" class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia angustiventris" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="angustiventris">Dexiogyia angustiventris (Casey, 1894)</taxonomicName>
Figs 1383-89Map 13
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="127" pageId="7" pageNumber="126" type="reference_group">
<paragraph pageId="7" pageNumber="126">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Thiasophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thiasophila angustiventris" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="angustiventris">Thiasophila angustiventris</taxonomicName>
Casey, 1894: 303; 1911: 16. Lectotype (male). Iowa; angustiventris-3, paratype NMNH 39754; Casey bequest 1925; male; our lectotype designation label, present designation (NMNH). Paralectotypes: Iowa; Type NMNH 39754; Casey bequest 192 (NMNH) 1 female [dissected, missing spermatheca]; Iowa; angustiventris-2, paratype NMNH 39754; Casey bequest 1925 (NMNH) 1 female [undissected].
</paragraph>
<paragraph pageId="7" pageNumber="126">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Thiasophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thiasophila asperata" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="asperata">Thiasophila asperata</taxonomicName>
Casey, 1894: 303 syn. n.. Lectotype (female). California;
<taxonomicName class="Insecta" family="Staphylinidae" genus="Thiasophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thiasophila asperata" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="asperata">Thiasophila asperata</taxonomicName>
Casey; Type NMNH 39757; Casey bequest 1925; our lectotype designation label, present designation (NMNH) [dissected].
</paragraph>
<paragraph lastPageId="8" lastPageNumber="127" pageId="7" pageNumber="126">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnoglossa abscissa" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="abscissa">Ischnoglossa abscissa</taxonomicName>
Casey, 1911: 16 syn. n.
<pageBreakToken pageId="8" pageNumber="127" start="start">.</pageBreakToken>
Holotype (male). Rhode Island (Boston Neck in orig. descr.); male;
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnoglossa abscissa" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="abscissa">Ischnoglossa abscissa</taxonomicName>
Casey; Type NMNH 39753, Casey bequest 1925 (NMNH) [dissected].
</paragraph>
<paragraph pageId="8" pageNumber="127">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnoglossa tenuicauda" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuicauda">Ischnoglossa tenuicauda</taxonomicName>
Casey, 1911: 17 syn. n.. Holotype (male). Florida; male;
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnoglossa tenuicauda" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuicauda">Ischnoglossa tenuicauda</taxonomicName>
Casey; Type NMNH 39755, Casey bequest 1925 (NMNH) [dissected].
</paragraph>
<paragraph pageId="8" pageNumber="127">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnoglossa intenta" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="intenta">Ischnoglossa intenta</taxonomicName>
Casey, 1911: 17 syn. n.. Lectotype (male). Iowa, Iowa City, Wickham; intenta Casey; Type NMNH 39756, Casey bequest 1925; our lectotype designation label, present designation (NMNH), [dissected].
</paragraph>
<paragraph pageId="8" pageNumber="127">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnoglossa alticola" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="alticola">Ischnoglossa alticola</taxonomicName>
Casey, 1911: 18 syn. n.. Holotype (female). California (Truckee in orig. descr.);alticola Casey;Type NMNH 39758; Casey bequest 1925 (NMNH) [dissected, missing spermatheca].
</paragraph>
<paragraph pageId="8" pageNumber="127">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia angustiventris" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="angustiventris">Dexiogyia angustiventris</taxonomicName>
(Casey);
<bibRefCitation pageId="8" pageNumber="127">Seevers 1978</bibRefCitation>
: 68 (as
<normalizedToken originalValue="‘anguliventris’">'anguliventris'</normalizedToken>
). New Canadian Record
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="127" type="material examined">
<paragraph pageId="8" pageNumber="127">Material examined.</paragraph>
<paragraph pageId="8" pageNumber="127">
(Type material - see above). CANADA: ON:Hald.-Norfolk Reg., Cronmiller prop., 6 km W St Williams, site 2,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="42.67167">42°40'18&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-80.49">80°29'24&quot;W</geoCoordinate>
, forest, malaise pans, 17.v to 31.v.2011, A. Brunke &amp; S.M. Paiero, 1 (DEBU) same data except: 31.v. to 15.vi; Cronmiller prop., 6 km W St Williams,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="42.6725">42°40'21&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-80.490555">80°29'26&quot;W</geoCoordinate>
, forest, 5.vii.2011, A. Brunke, 1 (DEBU); Lambton Co.,Pinery Prov. Pk., under white pine bark, 17.iv.2010, A. Brunke, 2 (DEBU).
</paragraph>
<caption pageId="8" pageNumber="127">
<paragraph pageId="8" pageNumber="127">
Figures 13-18. Dorsal habitus of: 13
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia angustiventris" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="angustiventris">Dexiogyia angustiventris</taxonomicName>
(Casey) 14
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ilyobates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyobates bennetti" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="bennetti">Ilyobates bennetti</taxonomicName>
Donisthorpe 15
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ocyusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ocyusa canadensis" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="canadensis">Ocyusa canadensis</taxonomicName>
Lohse 16
<taxonomicName class="Insecta" family="Staphylinidae" genus="Oxypoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxypoda rubescans" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="rubescans">Oxypoda rubescans</taxonomicName>
Casey 17
<taxonomicName class="Insecta" family="Staphylinidae" genus="Parocyusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parocyusa americana" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="americana">Parocyusa americana</taxonomicName>
(Casey) 18
<taxonomicName class="Insecta" family="Staphylinidae" genus="Parocyusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parocyusa fuliginosa" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="fuliginosa">Parocyusa fuliginosa</taxonomicName>
(Casey). Scale 1mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="8" pageNumber="127" type="distribution">
<paragraph pageId="8" pageNumber="127">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="127">Canada: ON; USA: CA, FL, IA, RI. Native.</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="128" pageId="8" pageNumber="127" type="comments">
<paragraph pageId="8" pageNumber="127">Comments.</paragraph>
<paragraph pageId="8" pageNumber="127">
All North American species of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="genus">Dexiogyia</taxonomicName>
were described by
<bibRefCitation pageId="8" pageNumber="127">Casey (1894</bibRefCitation>
, 1911) (as
<taxonomicName class="Insecta" family="Staphylinidae" genus="Thiasophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thiasophila" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="genus">Thiasophila</taxonomicName>
and
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnoglossa" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="genus">Ischnoglossa</taxonomicName>
) and differentiated based on slight differences in body proportions, punctation, pubescence and color. An examination of the relevant types revealed no differences between them in their aedeagi or spermathecae and slight differences in external morphology, which were attributed to intraspecific variation. Therefore,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia angustiventris" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="angustiventris">Dexiogyia angustiventris</taxonomicName>
was selected as the valid name for this species based on its appearance before
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia asperata" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="asperata">Dexiogyia asperata</taxonomicName>
(Casey) in
<bibRefCitation pageId="8" pageNumber="127">Casey (1894)</bibRefCitation>
, and
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia asperata" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="asperata">Dexiogyia asperata</taxonomicName>
(Casey) syn. n.,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia abscissa" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="abscissa">Dexiogyia abscissa</taxonomicName>
(Casey) syn. n.,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia tenuicauda" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="tenuicauda">Dexiogyia tenuicauda</taxonomicName>
(Casey) syn. n.,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia intenta" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="intenta">Dexiogyia intenta</taxonomicName>
(Casey) syn. n.and
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia alticola" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="alticola">Dexiogyia alticola</taxonomicName>
(Casey) syn. n.are here placed in synonymy with
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia angustiventris" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="angustiventris">Dexiogyia angustiventris</taxonomicName>
(Casey). To provide nomenclatural stability we have selected and designated lectotypes for
<taxonomicName class="Insecta" family="Staphylinidae" genus="Thiasophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thiasophila angustiventris" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="angustiventris">Thiasophila angustiventris</taxonomicName>
Casey,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Thiasophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thiasophila asperata" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="asperata">Thiasophila asperata</taxonomicName>
Casey and
<taxonomicName class="Insecta" family="Staphylinidae" genus="Ischnoglossa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ischnoglossa intenta" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="intenta">Ischnoglossa intenta</taxonomicName>
Casey. Additionally, one non-type specimen (Iowa, male) of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia angustiventris" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="angustiventris">Dexiogyia angustiventris</taxonomicName>
and five non-type specimens in
<normalizedToken originalValue="Casey’s">Casey's</normalizedToken>
collection (NMNH) of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia alticola" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="alticola">Dexiogyia alticola</taxonomicName>
(California, Siskiou Co., 3 females, 1 male, 1 sex?) were examined.
</paragraph>
<paragraph pageId="8" pageNumber="127">
<bibRefCitation pageId="8" pageNumber="127">Seevers (1978)</bibRefCitation>
noted that the European species
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia corticina" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="corticina">Dexiogyia corticina</taxonomicName>
(Erichson) was probably distinct from
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia angustiventris" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="angustiventris">Dexiogyia angustiventris</taxonomicName>
based on the longer and shaper teeth on male tergite 8 in the latter species. After examination of dissected specimens of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia corticina" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="corticina">Dexiogyia corticina</taxonomicName>
from Leipzig, Saxonia, Germany (ZMB), we consider both as valid but extremely similar species.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia corticina" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="corticina">Dexiogyia corticina</taxonomicName>
may be distinguished from the Nearctic
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia angustiventris" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="angustiventris">Dexiogyia angustiventris</taxonomicName>
based on the tubus of the median lobe with a ventral swelling in lateral view (straight in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia angustiventris" order="Coleoptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="species" species="angustiventris">Dexiogyia angustiventris</taxonomicName>
(Fig. 84) and the shorter, obtuse teeth of male tergite 8 (Fig. 85).
</paragraph>
<paragraph pageId="9" pageNumber="128">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Dexiogyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dexiogyia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="genus">
<pageBreakToken pageId="9" pageNumber="128" start="start">Dexiogyia</pageBreakToken>
</taxonomicName>
has been associated with subcortical microhabitats, especially those of pine and in the 'burrows of wood-boring
<normalizedToken originalValue="beetles’">beetles'</normalizedToken>
(
<bibRefCitation pageId="9" pageNumber="128">Seevers 1978</bibRefCitation>
). This is the first record of the genus from Canada and due to its association with pine (
<bibRefCitation pageId="9" pageNumber="128">Seevers 1978</bibRefCitation>
), we suspect this species to be transcontinental in Canada.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicNameLabel pageId="0" pageNumber="11223">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11223" type="materials_examined">
<paragraph pageId="0" pageNumber="11223">Materials</paragraph>
<paragraph pageId="0" pageNumber="11223">
<materialsCitation collectingDate="2003-09-28" collectingMethod="Reared from the larvae of the erebid moth, Dysschema jansonis" collectionCode="CNC, Insects" collectorName="D. H. Janzen, W. Hallwachs &amp; Asitente de Diana" country="Costa Rica" location="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223" specimenCode="DHJPAR 0011711" specimenCount="1" typeStatus="Holotype">
Type status:
<typeStatus pageId="0" pageNumber="11223">Holotype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="11223">DHJPAR0011711</specimenCode>
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<specimenType pageId="0" pageNumber="11223">M</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="11223">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: 03-SRNP-21135, BOLD:ABY7242, ASTAS437-06; Taxon: scientificName: Calolydellafredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming &amp; Wood, 2016; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="11223">Costa Rica</collectingCountry>
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<location LSID="urn:lsid:plazi:treatment:E811A9FF142D779B2FCD19787B3FC304:345835BC792CC0E2807107BD8DA4BC4C" country="Costa Rica" name="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223">Area de Conservacion Guanacaste</location>
; verbatimLocality: Estacion Pitilla; verbatimElevation:
<elevation pageId="0" pageNumber="11223">675</elevation>
; verbatimLatitude: 10.9893; verbatimLongitude: -85.4258; verbatimCoordinateSystem: Decimal; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="11223">AJ Fleming</determinerName>
; dateIdentified: 2016; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="11223">Reared from the larvae of the erebid moth, Dysschema jansonis</collectingMethod>
; verbatimEventDate:
<collectingDate pageId="0" pageNumber="11223" value="2003-09-28">28-Sep-2003</collectingDate>
; Record Level: language: en; institutionCode:
<collectionCode pageId="0" pageNumber="11223">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="11223">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
<materialsCitation collectingDate="2013-11-21" collectingMethod="Reared from the larvae of the nymphalid butterfly, Danaus plexippus" collectionCode="CNC, Insects" collectorName="D. H. Janzen, W. Hallwachs &amp; Cirilo Umana" country="Costa Rica" location="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223" specimenCode="DHJPAR 0054013" specimenCount="1" typeStatus="Paratype">
Type status:
<typeStatus pageId="0" pageNumber="11223">Paratype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="11223">DHJPAR0054013</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="11223">D.H. Janzen, W. Hallwachs &amp; Cirilo Umana</collectorName>
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; sex:
<specimenType pageId="0" pageNumber="11223">F</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="11223">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: 13-SRNP-77901, BOLD:ABY7242, ASHYD3181-14; Taxon: scientificName: Calolydellafredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming &amp; Wood, 2016; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="11223">Costa Rica</collectingCountry>
; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality:
<location LSID="urn:lsid:plazi:treatment:E811A9FF142D779B2FCD19787B3FC304:6E4FF5191C2CBADBB946623A037DC7D0" country="Costa Rica" name="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223">Area de Conservacion Guanacaste</location>
; verbatimLocality: Finca Esmeralda; verbatimElevation:
<elevation pageId="0" pageNumber="11223">123</elevation>
; verbatimLatitude: 10.9355; verbatimLongitude: -85.2531; verbatimCoordinateSystem: Decimal; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="11223">AJ Fleming</determinerName>
; dateIdentified: 2016; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="11223">Reared from the larvae of the nymphalid butterfly, Danaus plexippus</collectingMethod>
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<collectingDate pageId="0" pageNumber="11223" value="2013-11-21">21-Nov-2013</collectingDate>
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<collectionCode pageId="0" pageNumber="11223">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="11223">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
<materialsCitation collectingDate="1999-06-17" collectingMethod="Reared from the larvae of the nymphalid butterfly, Anaea aidea" collectionCode="CNC, Insects" collectorName="D. H. Janzen, W. Hallwachs &amp; gusaneros" country="Costa Rica" location="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223" specimenCode="DHJPAR 0017784" specimenCount="1" typeStatus="Paratype">
Type status:
<typeStatus pageId="0" pageNumber="11223">Paratype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="11223">DHJPAR0017784</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="11223">D.H. Janzen, W. Hallwachs &amp; gusaneros</collectorName>
; individualID: DHJPAR0017784; individualCount:
<specimenCount pageId="0" pageNumber="11223">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="11223">F</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="11223">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: 99-SRNP-8296, BOLD:ABY7242, ASTAR495-07; Taxon: scientificName: Calolydellafredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming &amp; Wood, 2016; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="11223">Costa Rica</collectingCountry>
; countryCode: CR; stateProvince: Guanacaste; county: Sector Horizontes; locality:
<location LSID="urn:lsid:plazi:treatment:E811A9FF142D779B2FCD19787B3FC304:B8ACC401639D2A7F6E607799A99F9055" country="Costa Rica" name="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223">Area de Conservacion Guanacaste</location>
; verbatimLocality: Sitio La Dama; verbatimElevation:
<elevation pageId="0" pageNumber="11223">105</elevation>
; verbatimLatitude: 10.7863; verbatimLongitude: -85.5583; verbatimCoordinateSystem: Decimal; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="11223">AJ Fleming</determinerName>
; dateIdentified: 2016; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="11223">Reared from the larvae of the nymphalid butterfly, Anaea aidea</collectingMethod>
; verbatimEventDate:
<collectingDate pageId="0" pageNumber="11223" value="1999-06-17">17-Jun-1999</collectingDate>
; Record Level: language: en; institutionCode:
<collectionCode pageId="0" pageNumber="11223">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="11223">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
<materialsCitation collectingDate="2012-12-16" collectingMethod="Reared from the larvae of the erebid moth, Melese Espinoza 01" collectionCode="CNC, Insects" collectorName="D. H. Janzen, W. Hallwachs &amp; Cirilo Umana" country="Costa Rica" location="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223" specimenCode="DHJPAR 0050643" specimenCount="1" typeStatus="Paratype">
Type status:
<typeStatus pageId="0" pageNumber="11223">Paratype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="11223">DHJPAR0050643</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="11223">D.H. Janzen, W. Hallwachs &amp; Cirilo Umana</collectorName>
; individualID: DHJPAR0050643; individualCount:
<specimenCount pageId="0" pageNumber="11223">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="11223">M</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="11223">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: 12-SRNP-77815, BOLD:ABY7242, ACGBA3235-13; Taxon: scientificName: Calolydellafredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming &amp; Wood, 2016; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="11223">Costa Rica</collectingCountry>
; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality:
<location LSID="urn:lsid:plazi:treatment:E811A9FF142D779B2FCD19787B3FC304:C38DACDA6AFA292EB2412DD89813C85E" country="Costa Rica" name="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223">Area de Conservacion Guanacaste</location>
; verbatimLocality: Finca Esmeralda; verbatimElevation:
<elevation pageId="0" pageNumber="11223">123</elevation>
; verbatimLatitude: 10.9355; verbatimLongitude: -85.2531; verbatimCoordinateSystem: Decimal; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="11223">AJ Fleming</determinerName>
; dateIdentified: 2016; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="11223">Reared from the larvae of the erebid moth, Melese Espinoza01</collectingMethod>
; verbatimEventDate:
<collectingDate pageId="0" pageNumber="11223" value="2012-12-16">16-Dec-2012</collectingDate>
; Record Level: language: en; institutionCode:
<collectionCode pageId="0" pageNumber="11223">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="11223">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
<materialsCitation collectingDate="2009-08-09" collectingMethod="Reared from the larvae of the nymphalid butterfly, Chlosyne rosita" collectionCode="CNC, Insects" collectorName="D. H. Janzen, W. Hallwachs &amp; Mariano Pereira" country="Costa Rica" location="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223" specimenCode="DHJPAR 0035673" specimenCount="1" typeStatus="Paratype">
Type status:
<typeStatus pageId="0" pageNumber="11223">Paratype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="11223">DHJPAR0035673</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="11223">D.H. Janzen, W. Hallwachs &amp; Mariano Pereira</collectorName>
; individualID: DHJPAR0035673; individualCount:
<specimenCount pageId="0" pageNumber="11223">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="11223">M</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="11223">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: 09-SRNP-56992, BOLD:ABY7242, ASHYD1054-09; Taxon: scientificName: Calolydellafredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming &amp; Wood, 2016; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="11223">Costa Rica</collectingCountry>
; countryCode: CR; stateProvince: Guanacaste; county: Sector Mundo Nuevo; locality:
<location LSID="urn:lsid:plazi:treatment:E811A9FF142D779B2FCD19787B3FC304:CA72215AF325A376C066A3137CDD6E8F" country="Costa Rica" name="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223">Area de Conservacion Guanacaste</location>
; verbatimLocality: Camino Pozo Tres; verbatimElevation:
<elevation pageId="0" pageNumber="11223">733</elevation>
; verbatimLatitude: 10.7708; verbatimLongitude: -85.3742; verbatimCoordinateSystem: Decimal; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="11223">AJ Fleming</determinerName>
; dateIdentified: 2016; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="11223">Reared from the larvae of the nymphalid butterfly, Chlosyne rosita</collectingMethod>
; verbatimEventDate:
<collectingDate pageId="0" pageNumber="11223" value="2009-08-09">09-Aug-2009</collectingDate>
; Record Level: language: en; institutionCode:
<collectionCode pageId="0" pageNumber="11223">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="11223">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
<materialsCitation collectingDate="2008-10-02" collectingMethod="Reared from the larvae of the crambid moth, Cliniodes opalalis" collectionCode="CNC, Insects" collectorName="D. H. Janzen, W. Hallwachs &amp; Dunia Garcia" country="Costa Rica" location="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223" specimenCode="DHJPAR 0029600" specimenCount="1" typeStatus="Paratype">
Type status:
<typeStatus pageId="0" pageNumber="11223">Paratype</typeStatus>
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
<specimenCode pageId="0" pageNumber="11223">DHJPAR0029600</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="11223">D.H. Janzen, W. Hallwachs &amp; Dunia Garcia</collectorName>
; individualID: DHJPAR0029600; individualCount:
<specimenCount pageId="0" pageNumber="11223">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="11223">F</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="11223">adult</specimenType>
; preparations: pinned; otherCatalogNumbers: 08-SRNP-36696, BOLD:ABY7242, ASHYM1021-09; Taxon: scientificName: Calolydellafredriksjobergi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: fredriksjobergi; scientificNameAuthorship: Fleming &amp; Wood, 2016; Location: continent: Central America; country:
<collectingCountry pageId="0" pageNumber="11223">Costa Rica</collectingCountry>
; countryCode: CR; stateProvince: Guanacaste; county: Sector Cacao; locality:
<location LSID="urn:lsid:plazi:treatment:E811A9FF142D779B2FCD19787B3FC304:DFFA757F9A322764B4F40F80D450388F" country="Costa Rica" name="Area de Conservacion Guanacaste" pageId="0" pageNumber="11223">Area de Conservacion Guanacaste</location>
; verbatimLocality: Sendero Ponderosa; verbatimElevation:
<elevation pageId="0" pageNumber="11223">1060</elevation>
; verbatimLatitude: 10.9146; verbatimLongitude: -85.4626; verbatimCoordinateSystem: Decimal; Identification: identifiedBy:
<determinerName pageId="0" pageNumber="11223">AJ Fleming</determinerName>
; dateIdentified: 2016; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="11223">Reared from the larvae of the crambid moth, Cliniodes opalalis</collectingMethod>
; verbatimEventDate:
<collectingDate pageId="0" pageNumber="11223" value="2008-10-02">02-Oct-2008</collectingDate>
; Record Level: language: en; institutionCode:
<collectionCode pageId="0" pageNumber="11223">CNC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="11223">Insects</collectionCode>
; basisOfRecord: Pinned Specimen
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11223" type="description">
<paragraph pageId="0" pageNumber="11223">Description</paragraph>
<paragraph pageId="0" pageNumber="11223">Male (Fig. 19a, b, c). Length: 7-9mm. Head (Fig. 19b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, with a single row of fine setulae outside of frontal setae; parafacial silver throughout. Thorax (Fig. 19a, c): gold on both dorsal and lateral surfaces; outermost two thoracic vittae, twice as wide as innermost two; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite with three or more hair-like setae, often in a small tuft; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with 6-7 setulae dorsally, reaching from base almost to crossvein R-M. Abdomen (Fig. 19a): ground color dark brown-orange, with uninterrupted transverse marginal pollinose bands; abdominal pollinosity gold dorsally, silver ventrally, with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae and two pairs of discal setae; T4 with a row of marginal setae and one pair of discal setae. Terminalia (Fig. 20): sternite 5 (Fig. 20c) with two small lobes and a wide U-shaped median cleft, 0.47X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; entire lobe of sternite with short setae, of varying lengths. Cerci (Fig. 20b), in dorsal view, separated by a narrow gap not widened at apex; each cercus very slightly tapered from its already narrow base; cercus long, slender and with an upward bend when viewed laterally; setose along its entire length. Surstylus (Fig. 20a) slender and digitiform with a strong divot at tip, giving it an overall bent look; appearing 0.2X shorter than cercus when viewed laterally; with short setae along entire length; tip of surstylus slightly curved inwards when viewed dorsally.</paragraph>
<paragraph pageId="0" pageNumber="11223">Female (Fig. 19d, e, f). Length: 5-6mm. Fronto-orbital plate 2.2X wider than in male; thoracic vittae 0.5X as wide as in male.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11223" type="diagnosis">
<paragraph pageId="0" pageNumber="11223">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="11223">
<taxonomicName class="Insecta" family="Tachinidae" genus="Calolydella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calolydella fredriksjobergi" order="Diptera" pageId="0" pageNumber="11223" phylum="Arthropoda" rank="species" species="fredriksjobergi">Calolydella fredriksjobergi</taxonomicName>
can be distinguished from all other species of
<taxonomicName class="Insecta" family="Tachinidae" genus="Calolydella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calolydella" order="Diptera" pageId="0" pageNumber="11223" phylum="Arthropoda" rank="genus">Calolydella</taxonomicName>
by the following combination of characters: parafacial mostly gold, anatergite with three or more hair-like setae arranged in a small tuft, two pairs of discal setae on T3 and one pair of discal setae on T4.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11223" type="etymology">
<paragraph pageId="0" pageNumber="11223">Etymology</paragraph>
<paragraph pageId="0" pageNumber="11223">
The specific epithet is in honor of Fredrik Sjoberg of Runmaro Island, Sweden, in recognition of his wonderful book &quot;The Fly Trap&quot; (
<bibRefCitation author="Sjoeberg, Fredrik" journalOrPublisher="Pantheon" pageId="0" pageNumber="11223" title="The Fly Trap" year="2014">
<normalizedToken originalValue="Sjöberg">Sjoeberg</normalizedToken>
2014
</bibRefCitation>
, translation by Thomas Teal); a book that explains much about Malaise traps, which have captured so many ACG
<taxonomicName family="Tachinidae" lsidName="" pageId="0" pageNumber="11223" rank="family">Tachinidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11223" type="distribution">
<paragraph pageId="0" pageNumber="11223">Distribution</paragraph>
<paragraph pageId="0" pageNumber="11223">Costa Rica, ACG, Alajuela and Guanacaste provinces, 105-1060m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11223" type="ecology">
<paragraph pageId="0" pageNumber="11223">Ecology</paragraph>
<paragraph pageId="0" pageNumber="11223">
<taxonomicName class="Insecta" family="Tachinidae" genus="Calolydella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calolydella fredriksjobergi" order="Diptera" pageId="0" pageNumber="11223" phylum="Arthropoda" rank="species" species="fredriksjobergi">Calolydella fredriksjobergi</taxonomicName>
has been reared six times from six different species belonging to various families of
<taxonomicName lsidName="Bernales" order="Lepidoptera" pageId="0" pageNumber="11223" rank="order">Lepidoptera</taxonomicName>
(
<taxonomicName family="Nymphalidae" lsidName="" pageId="0" pageNumber="11223" rank="family">Nymphalidae</taxonomicName>
;
<taxonomicName family="Erebidae" lsidName="" pageId="0" pageNumber="11223" rank="family">Erebidae</taxonomicName>
; and
<taxonomicName family="Crambidae" lsidName="" pageId="0" pageNumber="11223" rank="family">Crambidae</taxonomicName>
), in rain forest, cloud forest, and dry forest ecosystems.
</paragraph>
</subSubSection>
</treatment>
</document>

203
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</mods:role>
<mods:namePart>Yakovlev, R. V.</mods:namePart>
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<mods:date>2018</mods:date>
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<mods:number>2018-04-10</mods:number>
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<heading box="[227,691,1394,1417]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="19" reason="2">
<taxonomicName authority="Felder et Rogenhofer, 1874" authorityName="Felder et Rogenhofer" authorityYear="1874" box="[227,691,1394,1417]" class="Insecta" family="Noctuidae" genus="Amphipyra" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="19" phylum="Arthropoda" rank="species" species="surnia">
<emphasis bold="true" box="[227,691,1394,1417]" pageId="1" pageNumber="19">
<emphasis bold="true" box="[227,408,1394,1416]" italics="true" pageId="1" pageNumber="19">Amphipyra surnia</emphasis>
Felder et Rogenhofer, 1874
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[227,283,1422,1444]" pageId="1" pageNumber="19" type="description">
<paragraph blockId="1.[227,691,1394,1444]" box="[227,283,1422,1444]" pageId="1" pageNumber="19">
<figureCitation box="[227,283,1422,1444]" captionStart="Figs" captionStartId="2.[265,309,1371,1393]" captionTargetBox="[229,1121,270,1332]" captionTargetId="figure-1@2.[227,1123,257,1338]" captionTargetPageId="2" captionText="Figs. 1–7. Imago of Lepidoptera from the Russian Far East (coll. S. Rybalkin): 1 ‒ Gas-" figureDoi="http://doi.org/10.5281/zenodo.7164423" httpUri="https://zenodo.org/record/7164423/files/figure.png" pageId="1" pageNumber="19">Fig. 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="19" type="materials_examined">
<paragraph blockId="1.[227,1127,1477,1609]" box="[265,1127,1477,1499]" pageId="1" pageNumber="19">
MATERIAL.
<emphasis bold="true" box="[403,471,1477,1499]" pageId="1" pageNumber="19">
<collectingCountry box="[403,471,1477,1499]" name="Russia" pageId="1" pageNumber="19">Russia</collectingCountry>
</emphasis>
: Kunashir Island, 4.5 km NW Mendeleevo airport, Tretyakovo vill.,
</paragraph>
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<date box="[227,346,1505,1527]" pageId="1" pageNumber="19" value="2015-07-16">16.VII 2015</date>
,
<specimenCount box="[356,394,1505,1526]" pageId="1" pageNumber="19" type="male">1 ♂</specimenCount>
, leg. S. Rybalkin.
</paragraph>
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<subSubSection lastPageId="2" lastPageNumber="20" pageId="1" pageNumber="19" type="distribution">
<paragraph blockId="1.[227,1127,1477,1609]" box="[265,1127,1532,1554]" pageId="1" pageNumber="19">
DISTRIBUTION.
<collectingCountry box="[450,511,1533,1554]" name="South Korea" pageId="1" pageNumber="19">Korea</collectingCountry>
,
<collectingCountry box="[526,586,1532,1554]" name="China" pageId="1" pageNumber="19">China</collectingCountry>
,
<collectingCountry box="[600,674,1532,1554]" name="Taiwan" pageId="1" pageNumber="19">Taiwan</collectingCountry>
,
<collectingCountry box="[688,745,1533,1554]" name="Japan" pageId="1" pageNumber="19">Japan</collectingCountry>
,
<collectingCountry box="[760,824,1532,1554]" name="Russia" pageId="1" pageNumber="19">Russia</collectingCountry>
(Khabarovskii krai,
<collectingRegion box="[1037,1123,1532,1554]" country="Russia" name="Sakhalin" pageId="1" pageNumber="19">Sakhalin</collectingRegion>
,
</paragraph>
<paragraph blockId="1.[227,1127,1477,1609]" box="[227,1126,1560,1582]" pageId="1" pageNumber="19">
Shikotan Island) (Anikin
<emphasis box="[481,527,1560,1582]" italics="true" pageId="1" pageNumber="19">et al</emphasis>
., 2016). Here this species is firstly recorded from Kunashir
</paragraph>
<paragraph blockId="1.[227,1127,1477,1609]" box="[227,293,1587,1609]" pageId="1" pageNumber="19">Island.</paragraph>
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Figs. 1–7. Imago of
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from the Russian Far East (coll. S. Rybalkin): 1 ‒ Gas-
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1887),
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(
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(
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surnia
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Felder et Rogenhofer, 1874,
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Sugi, 1982
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(
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‒
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<specimenCount box="[690,708,1482,1503]" pageId="2" pageNumber="20" type="male">♂</specimenCount>
(
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niphonica
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Kishida et Yoshimoto, 1979,
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(Kunashir); 8 ‒
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<emphasis box="[704,919,1509,1531]" italics="true" pageId="2" pageNumber="20">Helicoverpa armigera</emphasis>
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<paragraph blockId="2.[227,1127,1371,1559]" box="[227,338,1537,1559]" pageId="2" pageNumber="20">(Kunashir).</paragraph>
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MATERIAL.
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:
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,
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NW Lesnoe
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, leg.
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DISTRIBUTION.
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, NE
<collectingCountry id="18AC75E5FFD0050BFDB3FC57FDEAFC11" box="[551,611,925,947]" name="China" pageId="1" pageNumber="19">China</collectingCountry>
, Russian Far East (Khabarovskii and Primorskii krai,
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Kunashir and
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Islands) (Anikin
<emphasis id="52CFE967FFD0050BFDF9FC73FD11FC6C" box="[621,664,953,974]" italics="true" pageId="1" pageNumber="19">et al</emphasis>
., 2016). This species was firstly recorded from
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<materialsCitation id="D0D33F28FFD0050BFF70FC1EFBEEFC48" box="[228,1127,980,1002]" location="Sakhalin" pageId="1" pageNumber="19" specimenCount="1" stateProvince="Sakhalin">
<collectingRegion id="A27FFB97FFD0050BFF70FC1EFEB1FC48" box="[228,312,980,1002]" country="Russia" name="Sakhalin" pageId="1" pageNumber="19">Sakhalin</collectingRegion>
based on
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from the Yuzhno-Sakhalinsk and Nevelsky (Vertyankin,
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2015). Our finding indicates its distribution in the south part of
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.
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Subfamily
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: Kunashir Island, 4.5 km NW Mendeleevo airport, Tretyakovo vill.,
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<date id="140513B5FFD0050BFF77FD98FED3FDCA" box="[227,346,594,616]" pageId="1" pageNumber="19" value="2015-07-23">23.VII 2015</date>
,
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DISTRIBUTION. Transpalaearctic species. Eastern part of area includes
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,
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, E
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MATERIAL.
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DISTRIBUTION. Transpalaearctic species (Anikin
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<emphasis id="52CFE967FFD20509FF77FE2EFE49FE5B" bold="true" box="[227,448,484,506]" italics="true" pageId="3" pageNumber="21">Sineugraphe oceanica</emphasis>
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<materialsCitation id="D0D33F28FFD20509FE9DFDFDFD24FDCA" collectingDate="2015-08-05" collectingDateMax="2015-08-06" collectingDateMin="2015-08-05" collectorName="S. Rybalkin" country="Russia" county="Island" location="Dolinka river" municipality="Korsakovo distr." pageId="3" pageNumber="21" specimenCount="8" stateProvince="Sakhalin">
MATERIAL.
<emphasis id="52CFE967FFD20509FE01FDFCFE50FDEE" bold="true" box="[405,473,566,588]" pageId="3" pageNumber="21">
<collectingCountry id="18AC75E5FFD20509FE01FDFCFE50FDEE" box="[405,473,566,588]" name="Russia" pageId="3" pageNumber="21">Russia</collectingCountry>
</emphasis>
:
<collectingRegion id="A27FFB97FFD20509FE7EFDFCFDB7FDEE" box="[490,574,566,588]" country="Russia" name="Sakhalin" pageId="3" pageNumber="21">Sakhalin</collectingRegion>
<collectingCounty id="89654DF9FFD20509FDDCFDFCFD0CFDEE" box="[584,645,566,588]" pageId="3" pageNumber="21">Island</collectingCounty>
,
<collectingMunicipality id="8060AF0FFFD20509FD07FDFCFCB3FDEE" box="[659,826,566,588]" pageId="3" pageNumber="21">Korsakovo distr.</collectingMunicipality>
,
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NW Lesnoe
</locationDeviation>
,
<location id="656463AEFFD20509FB8DFDFCFE9AFDCA" LSID="urn:lsid:plazi:treatment:E8128463FFD20509FF77FE2EFCFFFD74:656463AEFFD20509FB8DFDFCFE9AFDCA" country="Russia" county="Island" municipality="Korsakovo distr." name="Dolinka river" pageId="3" pageNumber="21" stateProvince="Sakhalin">Dolinka river</location>
,
<date id="140513B5FFD20509FEB5FD98FE3BFDCA" box="[289,434,594,616]" pageId="3" pageNumber="21" value="2015-08-05" valueMax="2015-08-06" valueMin="2015-08-05">
<collectingDate id="0441EA5DFFD20509FEB5FD98FE3BFDCA" box="[289,434,594,616]" pageId="3" pageNumber="21" value="2015-08-05" valueMax="2015-08-06" valueMin="2015-08-05">5‒6.VIII 2015</collectingDate>
</date>
,
<specimenCount id="76BDFEFCFFD20509FE55FD98FE7EFDCA" box="[449,503,594,616]" pageId="3" pageNumber="21" type="generic">8 ex.</specimenCount>
, leg.
<collectorName id="CD4E50A3FFD20509FDA0FD98FD24FDCA" box="[564,685,594,616]" pageId="3" pageNumber="21">S. Rybalkin</collectorName>
</materialsCitation>
;
<materialsCitation id="D0D33F28FFD20509FD28FD98FCBFFD26" collectingDate="2015-07-18" collectorName="NW Mendeleevo &amp; S. Rybalkin" country="Russia" location="Tretyakovo" municipality="Kunashir Island" pageId="3" pageNumber="21" specimenCount="4" stateProvince="Sakhalin">
<collectingMunicipality id="8060AF0FFFD20509FD28FD98FCD5FDCA" box="[700,860,594,616]" pageId="3" pageNumber="21">Kunashir Island</collectingMunicipality>
, 4.5 km
<collectorName id="CD4E50A3FFD20509FC29FD99FBEBFDCA" box="[957,1122,594,616]" pageId="3" pageNumber="21">NW Mendeleevo</collectorName>
airport,
<location id="656463AEFFD20509FEA4FDA4FE16FD26" LSID="urn:lsid:plazi:treatment:E8128463FFD20509FF77FE2EFCFFFD74:656463AEFFD20509FEA4FDA4FE16FD26" box="[304,415,622,644]" country="Russia" municipality="Kunashir Island" name="Tretyakovo" pageId="3" pageNumber="21" stateProvince="Sakhalin">Tretyakovo</location>
vill.,
<date id="140513B5FFD20509FE4DFDA4FDD8FD21" box="[473,593,621,643]" pageId="3" pageNumber="21" value="2015-07-18">
<collectingDate id="0441EA5DFFD20509FE4DFDA4FDD8FD21" box="[473,593,621,643]" pageId="3" pageNumber="21" value="2015-07-18">18.VII 2015</collectingDate>
</date>
,
<specimenCount id="76BDFEFCFFD20509FDC9FDA4FD06FD21" box="[605,655,622,643]" pageId="3" pageNumber="21" type="generic">4 ex.</specimenCount>
, leg.
<collectorName id="CD4E50A3FFD20509FD50FDA4FCBFFD26" box="[708,822,622,644]" pageId="3" pageNumber="21">S. Rybalkin</collectorName>
</materialsCitation>
;
<materialsCitation id="D0D33F28FFD20509FCD0FDA4FBEFFD21" box="[836,1126,621,644]" collectingDate="2015-07-18" collectorName="NW Mendeleevo &amp; S. Rybalkin" country="Russia" location="Tretyakovo" municipality="Kunashir Island" pageId="3" pageNumber="21" specimenCount="1" stateProvince="Sakhalin">same locality, 26.VII ‒ 5.VIII</materialsCitation>
</paragraph>
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2017,
<specimenCount id="76BDFEFCFFD20509FE8BFD43FED5FD3D" box="[287,348,649,671]" pageId="3" pageNumber="21" type="generic">19 ex.</specimenCount>
, leg. S. Rybalkin.
</paragraph>
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<paragraph id="60043575FFD20509FE9DFD6FFBE1FD19" blockId="3.[227,1128,566,726]" box="[265,1128,677,699]" pageId="3" pageNumber="21">
DISTRIBUTION.
<collectingCountry id="18AC75E5FFD20509FE55FD6FFE73FD18" box="[449,506,677,698]" name="Japan" pageId="3" pageNumber="21">Japan</collectingCountry>
,
<collectingCountry id="18AC75E5FFD20509FD93FD6FFDCDFD18" box="[519,580,677,698]" name="South Korea" pageId="3" pageNumber="21">Korea</collectingCountry>
,
<collectingCountry id="18AC75E5FFD20509FDC5FD6FFD04FD19" box="[593,653,677,699]" name="China" pageId="3" pageNumber="21">China</collectingCountry>
,
<collectingCountry id="18AC75E5FFD20509FD0EFD6FFD53FD19" box="[666,730,677,699]" name="Russia" pageId="3" pageNumber="21">Russia</collectingCountry>
(Primorskii krai) (Anikin
<emphasis id="52CFE967FFD20509FC71FD6FFB9AFD18" box="[997,1043,677,698]" italics="true" pageId="3" pageNumber="21">et al</emphasis>
., 2016).
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<paragraph id="60043575FFD20509FF77FD0BFCFFFD74" blockId="3.[227,1128,566,726]" box="[227,886,704,726]" pageId="3" pageNumber="21">
Here this species is firstly recorded from
<collectingRegion id="A27FFB97FFD20509FDFBFD0AFD4AFD74" box="[623,707,704,726]" country="Russia" name="Sakhalin" pageId="3" pageNumber="21">Sakhalin</collectingRegion>
and Kuril Islands.
</paragraph>
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121
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<emphasis id="52CFE967FFD20509FF77FECDFE20FEBF" bold="true" box="[227,425,263,285]" italics="true" pageId="3" pageNumber="21">Melanchra postalba</emphasis>
Sugi, 1982
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<materialsCitation id="D0D33F28FFD20509FE9DFE90FD0AFE29" collectingDate="2015-08-05" collectingDateMax="2015-08-06" collectingDateMin="2015-08-05" collectorName="S. Rybalkin." country="Russia" county="Island" location="Dolinka river" municipality="Korsakovo distr." pageId="3" pageNumber="21" specimenCount="2" specimenCount-male="2" stateProvince="Sakhalin">
MATERIAL.
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<collectingCountry id="18AC75E5FFD20509FE01FE93FE50FECD" box="[405,473,345,367]" name="Russia" pageId="3" pageNumber="21">Russia</collectingCountry>
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:
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<collectingCounty id="89654DF9FFD20509FDDCFE90FD0CFED2" box="[584,645,346,368]" pageId="3" pageNumber="21">Island</collectingCounty>
,
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NW Lesnoe
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,
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DISTRIBUTION.
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,
<collectingCountry id="18AC75E5FFD20509FD93FE5BFDCDFE04" box="[519,580,401,422]" name="South Korea" pageId="3" pageNumber="21">Korea</collectingCountry>
,
<collectingCountry id="18AC75E5FFD20509FDC5FE5BFD04FE05" box="[593,653,401,423]" name="China" pageId="3" pageNumber="21">China</collectingCountry>
,
<collectingCountry id="18AC75E5FFD20509FD0EFE5BFD53FE05" box="[666,730,401,423]" name="Russia" pageId="3" pageNumber="21">Russia</collectingCountry>
(Primorskii krai) (Anikin
<emphasis id="52CFE967FFD20509FC71FE58FB9AFE04" box="[997,1043,401,423]" italics="true" pageId="3" pageNumber="21">et al</emphasis>
., 2016).
</paragraph>
<paragraph id="60043575FFD20509FF77FE67FC9DFE60" blockId="3.[227,1128,345,450]" box="[227,788,428,450]" pageId="3" pageNumber="21">
Here this species is firstly recorded from
<collectingRegion id="A27FFB97FFD20509FDFBFE66FD4AFE60" box="[623,707,428,450]" country="Russia" name="Sakhalin" pageId="3" pageNumber="21">Sakhalin</collectingRegion>
Islands.
</paragraph>
</subSubSection>
</treatment>
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308
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@ -0,0 +1,308 @@
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<mods:title id="66364D118F31C302CF947FB6B869402B">A new species of Polyophthalmus (Annelida, Opheliidae) from the Arabian Gulf with an insight on internal anatomy and diversity of the genus</mods:title>
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<paragraph id="6004368E2C4EF763FF518AC0FB5BFBB8" blockId="21.[151,1164,1014,1041]" box="[151,1164,1014,1041]" pageId="21" pageNumber="522">
<heading id="3B4C81E22C4EF763FF518AC0FB5BFBB8" bold="true" box="[151,1164,1014,1041]" fontSize="11" level="1" pageId="21" pageNumber="522" reason="1">
<emphasis id="52CFEA9C2C4EF763FF518AC0FB5BFBB8" bold="true" box="[151,1164,1014,1041]" pageId="21" pageNumber="522">
Key to species of
<taxonomicName id="A7BB4D0D2C4EF763FEA88AC1FCCAFBB8" ID-CoL="84SK5" authority="Quatrefages, 1850" authorityName="Quatrefages" authorityYear="1850" box="[366,797,1014,1041]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="genus">
<emphasis id="52CFEA9C2C4EF763FEA88AC1FDE4FBB9" bold="true" box="[366,563,1014,1041]" italics="true" pageId="21" pageNumber="522">Polyophthalmus</emphasis>
<bibRefCitation id="042A4B7F2C4EF763FDFD8AC1FCCAFBB8" author="Quatrefages, A. de" box="[571,797,1014,1040]" pageId="21" pageNumber="522" pagination="5 - 46" refId="ref14480" refString="Quatrefages, A. de (1850) Etudes sur les types inferieurs de l'embranchement des Anneles. Memoire sur la famille des Polyphthalmiens (Polyphthalmea Nob.). Annales des Sciences Naturelles, Paris, Series 3, 13, 5 - 46." type="journal article" year="1850">Quatrefages, 1850</bibRefCitation>
</taxonomicName>
of Indian and Pacific Oceans
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="28A165052C4EF763FF518D08FD96F9A1" pageId="21" pageNumber="522" type="materials_examined">
<paragraph id="6004368E2C4EF763FF518D08FC7DFA99" blockId="21.[151,1437,1087,1545]" pageId="21" pageNumber="522">
The following tentative key comprises the seven
<taxonomicName id="A7BB4D0D2C4EF763FD0A8D08FC0BFBF1" box="[716,988,1087,1113]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="52CFEA9C2C4EF763FD0A8D08FC55FBF1" box="[716,898,1087,1113]" italics="true" pageId="21" pageNumber="522">Polyophthalmus</emphasis>
species
</taxonomicName>
previously described in the Indian and Pacific oceans by
<bibRefCitation id="042A4B7F2C4EF763FEA78D54FE2DFBD6" author="Grube, A. E." box="[353,506,1123,1150]" pageId="21" pageNumber="522" pagination="59 - 68" refId="ref12791" refString="Grube, A. E. (1869) Familie der Opheliaceen. Schlesische gesellschaft fur vaterlandische kultur Breslau Jahresbericht, 46, 59 - 68." type="journal article" year="1869">Grube (1869)</bibRefCitation>
,
<bibRefCitation id="042A4B7F2C4EF763FDC08D54FD19FBD5" author="Kukenthal, W." box="[518,718,1123,1149]" pageId="21" pageNumber="522" pagination="361 - 373" refId="ref13441" refString="Kukenthal, W. (1887) Die Opheliaceen der expedition der Vettore Pisani. Jenaische Zeitschrift fur Naturwissenschaft, 21, 361 - 373." type="journal article" year="1887">Kükenthal (1887)</bibRefCitation>
,
<bibRefCitation id="042A4B7F2C4EF763FD1F8D54FC79FBD5" author="Michaelsen, W." box="[729,942,1123,1149]" pageId="21" pageNumber="522" pagination="1 - 23" refId="ref13643" refString="Michaelsen, W. (1892) Polychaeten von Ceylon. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 9 (2), 1 - 23." type="journal article" year="1892">Michaelsen (1892)</bibRefCitation>
,
<bibRefCitation id="042A4B7F2C4EF763FC7F8D54FB78FBD5" author="Purschke, G. &amp; Ding, Z. &amp; Muller, M. C." box="[953,1199,1122,1149]" pageId="21" pageNumber="522" pagination="229 - 241" refId="ref14413" refString="Purschke, G., Ding, Z. &amp; Muller, M. C. (1995) Ultrastructural differences as a taxonomic marker: the segmental ocelli of Polyophthalmus pictus and Polyophthalmus qingdaoensis sp. n. (Polychaeta, Opheliidae). Zoomorphology, 115 (4), 229 - 241. https: // doi. org / 10.1007 / BF 00393803" type="journal article" year="1995">
Purschke
<emphasis id="52CFEA9C2C4EF763FBE08D54FB89FBD5" box="[1062,1118,1123,1149]" italics="true" pageId="21" pageNumber="522">et al.</emphasis>
(1995)
</bibRefCitation>
and
<bibRefCitation id="042A4B7F2C4EF763FB238D54FF33FB09" author="Magalhaes, W. F. &amp; Rizzo, A. E. &amp; Bailey-Brock, J. H." pageId="21" pageNumber="522" pagination="209 - 235" refId="ref13534" refString="Magalhaes, W. F., Rizzo, A. E. &amp; Bailey-Brock, J. H. (2019) Opheliidae (Annelida: Polychaeta) from the western Pacific islands, including five new species. Zootaxa, 4555 (2), 209 - 235. https: // doi. org / 10.11646 / zootaxa. 4555.2.3" type="journal article" year="2019">
Magalhães
<emphasis id="52CFEA9C2C4EF763FAA38D54FA4AFBD5" box="[1381,1437,1123,1149]" italics="true" pageId="21" pageNumber="522">et al.</emphasis>
(2019)
</bibRefCitation>
, as well as the new species described here,
<taxonomicName id="A7BB4D0D2C4EF763FD078DB0FC91FB09" authority="Parapar &amp; Al-Kandari &amp; Candás &amp; Moreira, 2021" authorityName="Parapar &amp; Al-Kandari &amp; Candás &amp; Moreira" authorityYear="2021" box="[705,838,1159,1185]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="zhadanae" status="sp. nov.">
<emphasis id="52CFEA9C2C4EF763FD078DB0FC91FB09" box="[705,838,1159,1185]" italics="true" pageId="21" pageNumber="522">P. zhadanae</emphasis>
</taxonomicName>
<emphasis id="52CFEA9C2C4EF763FC8B8DB0FC72FB09" bold="true" box="[845,933,1159,1185]" pageId="21" pageNumber="522">
<taxonomicNameLabel id="49FC57E72C4EF763FC8B8DB0FC72FB09" box="[845,933,1159,1185]" pageId="21" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Some of these species had been synonimized with the NE Atlantic
<taxonomicName id="A7BB4D0D2C4EF763FE558D9CFE22FB6C" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[403,501,1195,1220]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="pictus">
<emphasis id="52CFEA9C2C4EF763FE558D9CFE22FB6C" box="[403,501,1195,1220]" italics="true" pageId="21" pageNumber="522">P. pictus</emphasis>
</taxonomicName>
and therefore the latter is also included in the key for comparisons. Material of previously described species has not been examined here; original descriptions and
<bibRefCitation id="042A4B7F2C4EF763FB8D8DF8FA84FB41" author="Magalhaes, W. F. &amp; Rizzo, A. E. &amp; Bailey-Brock, J. H." box="[1099,1363,1231,1257]" pageId="21" pageNumber="522" pagination="209 - 235" refId="ref13534" refString="Magalhaes, W. F., Rizzo, A. E. &amp; Bailey-Brock, J. H. (2019) Opheliidae (Annelida: Polychaeta) from the western Pacific islands, including five new species. Zootaxa, 4555 (2), 209 - 235. https: // doi. org / 10.11646 / zootaxa. 4555.2.3" type="journal article" year="2019">
Magalhães
<emphasis id="52CFEA9C2C4EF763FB0B8DF8FB28FB41" box="[1229,1279,1231,1257]" italics="true" pageId="21" pageNumber="522">et al</emphasis>
. (2019
</bibRefCitation>
;
<tableCitation id="2D3903352C4EF763FA988DF8FF74FAA5" captionStart="TABLE 1" captionStartId="4.[151,239,471,497]" captionTargetPageId="4" captionText="TABLE 1. Locality and collection data, museum registration numbers for type material of new species (except for KISR) and reference to figures, of examined specimens of Polyophthalmus zhadanae sp. nov. Abbreviations: H—high intertidal; L—low intertidal; M—middle intertidal; n.d. = no data; ST—subtidal. Specimens deposited at the KISR do not have collection numbers. (*) SEM stub. Geographical coordinates in decimal degrees (degree symbols omitted); latitude is North (N) and longitude is East (E) unless noted (West: W)." httpUri="http://table.plazi.org/id/34C466062C5FF772FF5188E0FEDCFDF5" pageId="21" pageNumber="522" tableUuid="34C466062C5FF772FF5188E0FEDCFDF5">Table 1</tableCitation>
) were consulted to build the key. Some characters not explicitly mentioned in original descriptions were extracted from original illustrations.
<typeStatus id="BF00882C2C4EF763FE028C20FE2BFA99" box="[452,508,1303,1329]" pageId="21" pageNumber="522">Type</typeStatus>
locality of each species is in brackets.
</paragraph>
<paragraph id="6004368E2C4EF763FF018C0CFD96F9A1" blockId="21.[151,1437,1087,1545]" pageId="21" pageNumber="522">
Characters used in
<taxonomicName id="A7BB4D0D2C4EF763FE558C0CFD9EFAFD" authorityName="Quatrefages" authorityYear="1850" box="[403,585,1339,1365]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="genus">
<emphasis id="52CFEA9C2C4EF763FE558C0CFD9EFAFD" box="[403,585,1339,1365]" italics="true" pageId="21" pageNumber="522">Polyophthalmus</emphasis>
</taxonomicName>
keys are traditionally focused on body pigmentation, distribution of segmental eyes and chaetal length. We avoided diagnostic characters such as presence of pygidial cirri and parapodial cirri because they can only be visualized properly with SEM and therefore have been only described in recent works. In the same way, the presence and distribution of interparapodial organs is unknown for most of species. Study of
<typeStatus id="BF00882C2C4EF763FAAA8C90FA4BFA69" box="[1388,1436,1447,1473]" pageId="21" pageNumber="522">type</typeStatus>
material and newly collected specimens in the Indo-Pacific will help to better characterize these taxa regarding the aforementioned diagnostic characters.
</paragraph>
</subSubSection>
<subSubSection id="28A165052C4EF760FF518F03FA4BFE8B" lastPageId="22" lastPageNumber="523" pageId="21" pageNumber="522" type="key">
<keyStep id="DB4F2E2B2C4EF763FF518F03FA4BF9CE" pageId="21" pageNumber="522">
<paragraph id="6004368E2C4EF763FF518F03FA4BF9E2" blockId="21.[151,1437,1588,2020]" box="[151,1436,1588,1610]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518F03FA4BF9E2" box="[151,1436,1588,1610]" pageId="21" pageNumber="522">
1. Segmental eyes in almost all body chaetigers....................................
<taxonomicName id="A7BB4D0D2C4EF763FB8B8F03FA4BF9E2" ID-CoL="4LCTF" authority="(Dujardin, 1839) (France)" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[1101,1436,1588,1610]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="pictus">
<emphasis id="52CFEA9C2C4EF763FB8B8F03FB4AF9E1" box="[1101,1181,1588,1609]" italics="true" pageId="21" pageNumber="522">P. pictus</emphasis>
(
<bibRefCitation id="042A4B7F2C4EF763FB6C8F03FAE9F9E2" author="Dujardin, F." box="[1194,1342,1588,1610]" pageId="21" pageNumber="522" pagination="287 - 294" refId="ref12092" refString="Dujardin, F. (1839) Observations sur quelques Annelides marines. Annales des Sciences Naturelles, Paris, 11, 287 - 294." type="journal article" year="1839">Dujardin, 1839</bibRefCitation>
) (
<collectingCountry id="18AC761E2C4EF763FA978F03FA42F9E2" box="[1361,1429,1588,1610]" name="France" pageId="21" pageNumber="522">France</collectingCountry>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="6004368E2C4EF763FF518F66FA4BF9CE" blockId="21.[151,1437,1588,2020]" box="[151,1436,1617,1639]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518F66FA4BF9CE" box="[151,1436,1617,1639]" pageId="21" pageNumber="522">- Segmental eyes absent in anterior most and posterior most chaetigers............................................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4F2E2B2C4EF763FF518F59FA4BF909" pageId="21" pageNumber="522">
<paragraph id="6004368E2C4EF763FF518F59FA4BF92B" blockId="21.[151,1437,1588,2020]" box="[151,1436,1646,1668]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518F59FA4BF92B" box="[151,1436,1646,1668]" pageId="21" pageNumber="522">2. Chaetae of last 5–7 chaetigers longer than previous ones and surpassing pygidium................................. 3</keyLead>
</paragraph>
<paragraph id="6004368E2C4EF763FF518FBBFA4BF909" blockId="21.[151,1437,1588,2020]" box="[151,1436,1675,1698]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518FBBFA4BF909" box="[151,1436,1675,1698]" pageId="21" pageNumber="522">- Chaetae of last 5–7 chaetigers same length as previous ones................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4F2E2B2C4EF763FF518F9EFA4AF974" pageId="21" pageNumber="522">
<paragraph id="6004368E2C4EF763FF518F9EFA4BF917" blockId="21.[151,1437,1588,2020]" box="[151,1436,1705,1727]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518F9EFA4BF917" box="[151,1436,1705,1727]" pageId="21" pageNumber="522">
3. Anal marginal papillae elongate......................................
<taxonomicName id="A7BB4D0D2C4EF763FC2F8F9DFAFFF916" ID-CoL="8RDG8" authority="Michaelsen, 1892" authorityName="Michaelsen" authorityYear="1892" box="[1001,1320,1705,1727]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="longisetosus">
<emphasis id="52CFEA9C2C4EF763FC2F8F9DFBAFF916" box="[1001,1144,1705,1727]" italics="true" pageId="21" pageNumber="522">P. longisetosus</emphasis>
<bibRefCitation id="042A4B7F2C4EF763FBB88F9EFAFFF916" author="Michaelsen, W." box="[1150,1320,1705,1727]" pageId="21" pageNumber="522" pagination="1 - 23" refId="ref13643" refString="Michaelsen, W. (1892) Polychaeten von Ceylon. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 9 (2), 1 - 23." type="journal article" year="1892">Michaelsen, 1892</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="18AC761E2C4EF763FAF38F9EFA41F917" box="[1333,1430,1705,1727]" name="Sri Lanka" pageId="21" pageNumber="522">Sri Lanka</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="6004368E2C4EF763FF518FF0FA4AF974" blockId="21.[151,1437,1588,2020]" box="[151,1437,1734,1756]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518FF0FA4AF974" box="[151,1437,1734,1756]" pageId="21" pageNumber="522">
- Anal marginal papillae digitate, short...................
<taxonomicName id="A7BB4D0D2C4EF763FCF08FF0FA4AF974" ID-CoL="4LCTB" authority="Magalhaes, Rizzo and Bailey-Brock, 2019" authorityName="Magalhaes, Rizzo and Bailey-Brock" authorityYear="2019" box="[822,1437,1734,1756]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="mauliola">
<emphasis id="52CFEA9C2C4EF763FCF08FF0FC74F973" box="[822,931,1734,1756]" italics="true" pageId="21" pageNumber="522">P. mauliola</emphasis>
<bibRefCitation id="042A4B7F2C4EF763FC6F8FF1FA96F973" author="Magalhaes, W. F. &amp; Rizzo, A. E. &amp; Bailey-Brock, J. H." box="[937,1345,1734,1756]" pageId="21" pageNumber="522" pagination="209 - 235" refId="ref13534" refString="Magalhaes, W. F., Rizzo, A. E. &amp; Bailey-Brock, J. H. (2019) Opheliidae (Annelida: Polychaeta) from the western Pacific islands, including five new species. Zootaxa, 4555 (2), 209 - 235. https: // doi. org / 10.11646 / zootaxa. 4555.2.3" type="journal article" year="2019">Magalhães, Rizzo and Bailey-Brock, 2019</bibRefCitation>
(Hawaii)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4F2E2B2C4EF763FF518FD3FA4BF8BF" pageId="21" pageNumber="522">
<paragraph id="6004368E2C4EF763FF518FD3FA4BF951" blockId="21.[151,1437,1588,2020]" box="[151,1436,1763,1786]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518FD3FA4BF951" box="[151,1436,1763,1786]" pageId="21" pageNumber="522">4. Prostomium with distinctive pigmentation.................................................................. 5</keyLead>
</paragraph>
<paragraph id="6004368E2C4EF763FF518E36FA4BF8BF" blockId="21.[151,1437,1588,2020]" box="[151,1436,1793,1815]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518E36FA4BF8BF" box="[151,1436,1793,1815]" pageId="21" pageNumber="522">- Prostomium lacking distinctive pigmentation............................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4F2E2B2C4EF763FF518E29FA4AF8C7" pageId="21" pageNumber="522">
<paragraph id="6004368E2C4EF763FF518E29FA4BF89C" blockId="21.[151,1437,1588,2020]" box="[151,1436,1822,1844]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518E29FA4BF89C" box="[151,1436,1822,1844]" pageId="21" pageNumber="522">
5. Pigmentation present in dorsal region of prostomium and peristomium...........
<taxonomicName id="A7BB4D0D2C4EF763FBD18E28FAFFF89B" ID-CoL="8RDG7" authority="Michaelsen, 1892" authorityName="Michaelsen" authorityYear="1892" box="[1047,1320,1822,1844]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="collaris">
<emphasis id="52CFEA9C2C4EF763FBD18E28FBA0F89B" box="[1047,1143,1822,1844]" italics="true" pageId="21" pageNumber="522">P. collaris</emphasis>
<bibRefCitation id="042A4B7F2C4EF763FBB88E29FAFFF89B" author="Michaelsen, W." box="[1150,1320,1822,1844]" pageId="21" pageNumber="522" pagination="1 - 23" refId="ref13643" refString="Michaelsen, W. (1892) Polychaeten von Ceylon. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 9 (2), 1 - 23." type="journal article" year="1892">Michaelsen, 1892</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="18AC761E2C4EF763FAF38E29FA41F89C" box="[1333,1430,1822,1844]" name="Sri Lanka" pageId="21" pageNumber="522">Sri Lanka</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="6004368E2C4EF763FF518E0BFA4AF8C7" blockId="21.[151,1437,1588,2020]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518E0BFA4AF8C7" pageId="21" pageNumber="522">
- About 10 dark pigmented structures inside brain including ocelli; peristomium not pigmented.........................................................................................
<taxonomicName id="A7BB4D0D2C4EF763FC218E6DFA4AF8C7" ID-CoL="4LCTG" authority="Purschke et al., 1995" authorityName="Purschke" authorityYear="1995" box="[999,1437,1881,1903]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="qingdaoensis">
<emphasis id="52CFEA9C2C4EF763FC218E6DFBAAF8C6" box="[999,1149,1881,1903]" italics="true" pageId="21" pageNumber="522">P. qingdaoensis</emphasis>
<bibRefCitation id="042A4B7F2C4EF763FB458E6EFA9AF8C7" author="Purschke, G. &amp; Ding, Z. &amp; Muller, M. C." box="[1155,1357,1881,1903]" pageId="21" pageNumber="522" pagination="229 - 241" refId="ref14413" refString="Purschke, G., Ding, Z. &amp; Muller, M. C. (1995) Ultrastructural differences as a taxonomic marker: the segmental ocelli of Polyophthalmus pictus and Polyophthalmus qingdaoensis sp. n. (Polychaeta, Opheliidae). Zoomorphology, 115 (4), 229 - 241. https: // doi. org / 10.1007 / BF 00393803" type="journal article" year="1995">
Purschke
<emphasis id="52CFEA9C2C4EF763FB278E6DFADCF8C6" box="[1249,1291,1881,1903]" italics="true" pageId="21" pageNumber="522">et al</emphasis>
., 1995
</bibRefCitation>
(
<collectingCountry id="18AC761E2C4EF763FA9C8E6EFA41F8C7" box="[1370,1430,1881,1903]" name="China" pageId="21" pageNumber="522">China</collectingCountry>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4F2E2B2C4EF763FF518E41FA4BF801" pageId="21" pageNumber="522">
<paragraph id="6004368E2C4EF763FF518E41FA4BF824" blockId="21.[151,1437,1588,2020]" box="[151,1436,1910,1932]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518E41FA4BF824" box="[151,1436,1910,1932]" pageId="21" pageNumber="522">6. Transverse bands of brown pigmentation similar in almost all segments.......................................... 7</keyLead>
</paragraph>
<paragraph id="6004368E2C4EF763FF518EA3FA4BF801" blockId="21.[151,1437,1588,2020]" box="[151,1436,1940,1962]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518EA3FA4BF801" box="[151,1436,1940,1962]" pageId="21" pageNumber="522">- Transverse bands of brown pigmentation in two different patterns............................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4F2E2B2C4EF763FF518E86FA4AF84C" pageId="21" pageNumber="522">
<paragraph id="6004368E2C4EF763FF518E86FA4BF86F" blockId="21.[151,1437,1588,2020]" box="[151,1436,1969,1991]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518E86FA4BF86F" box="[151,1436,1969,1991]" pageId="21" pageNumber="522">
7. Segmental eyes from
<collectingCountry id="18AC761E2C4EF763FE748E86FE03F86E" box="[434,468,1969,1990]" name="Switzerland" pageId="21" pageNumber="522">CH</collectingCountry>
7 to
<collectingCountry id="18AC761E2C4EF763FE388E86FDF6F86E" box="[510,545,1969,1990]" name="Switzerland" pageId="21" pageNumber="522">CH</collectingCountry>
17........................................
<taxonomicName id="A7BB4D0D2C4EF763FBE68E85FADDF86F" ID-CoL="8RDG5" authority="Grube, 1869" authorityName="Grube" authorityYear="1869" box="[1056,1290,1969,1991]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="australis">
<emphasis id="52CFEA9C2C4EF763FBE68E85FB5BF86E" box="[1056,1164,1969,1991]" italics="true" pageId="21" pageNumber="522">P. australis</emphasis>
<bibRefCitation id="042A4B7F2C4EF763FB548E86FADDF86F" author="Grube, A. E." box="[1170,1290,1969,1991]" pageId="21" pageNumber="522" pagination="59 - 68" refId="ref12791" refString="Grube, A. E. (1869) Familie der Opheliaceen. Schlesische gesellschaft fur vaterlandische kultur Breslau Jahresbericht, 46, 59 - 68." type="journal article" year="1869">Grube, 1869</bibRefCitation>
</taxonomicName>
(NE
<collectingCountry id="18AC761E2C4EF763FAFA8E86FA41F86F" box="[1340,1430,1969,1991]" name="Australia" pageId="21" pageNumber="522">Australia</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="6004368E2C4EF763FF518EF8FA4AF84C" blockId="21.[151,1437,1588,2020]" box="[151,1437,1998,2020]" pageId="21" pageNumber="522">
<keyLead id="DB4A95BB2C4EF763FF518EF8FA4AF84C" box="[151,1437,1998,2020]" pageId="21" pageNumber="522">
- Segmental eyes from
<collectingCountry id="18AC761E2C4EF763FE748EF9FE02F84C" box="[434,469,1998,2020]" name="Switzerland" pageId="21" pageNumber="522">CH</collectingCountry>
5 to
<collectingCountry id="18AC761E2C4EF763FE388EF9FDF6F84C" box="[510,545,1998,2020]" name="Switzerland" pageId="21" pageNumber="522">CH</collectingCountry>
16.......................................
<taxonomicName id="A7BB4D0D2C4EF763FBD68EF8FAC1F84B" ID-CoL="8RDG9" authority="Kukenthal, 1887" authorityName="Kukenthal" authorityYear="1887" box="[1040,1302,1998,2020]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="striatus">
<emphasis id="52CFEA9C2C4EF763FBD68EF8FBA7F84C" box="[1040,1136,1999,2020]" italics="true" pageId="21" pageNumber="522">P. striatus</emphasis>
<bibRefCitation id="042A4B7F2C4EF763FBB08EF9FAC1F84B" author="Kukenthal, W." box="[1142,1302,1998,2020]" pageId="21" pageNumber="522" pagination="361 - 373" refId="ref13441" refString="Kukenthal, W. (1887) Die Opheliaceen der expedition der Vettore Pisani. Jenaische Zeitschrift fur Naturwissenschaft, 21, 361 - 373." type="journal article" year="1887">Kükenthal, 1887</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="18AC761E2C4EF763FAE48EF9FA41F84C" box="[1314,1430,1998,2020]" name="Hong Kong" pageId="21" pageNumber="522">Hong Kong</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB4F2E2B2C4DF760FF5189AFFA4BFE8B" pageId="22" pageNumber="523">
<paragraph id="6004368E2C4DF760FF5189AFFA4BFF63" blockId="22.[151,1437,151,291]" pageId="22" pageNumber="523">
<keyLead id="DB4A95BB2C4DF760FF5189AFFA4BFF63" pageId="22" pageNumber="523">
8. Nine pairs of lateral eyespots (
<collectingCountry id="18AC761E2C4DF760FDCC89A0FDF8FF05" box="[522,559,151,173]" name="Switzerland" pageId="22" pageNumber="523">CH</collectingCountry>
7–15), all same size. Anterior body region (
<collectingCountry id="18AC761E2C4DF760FC1489A0FC20FF05" box="[978,1015,151,173]" name="Switzerland" pageId="22" pageNumber="523">CH</collectingCountry>
1–5) with two light brown bands; following segments with dorsal dark brown patches................................
<taxonomicName id="A7BB4D0D2C4DF760FC3A8982FAFFFF62" ID-CoL="8RDG6" authority="Kukenthal, 1887" authorityName="Kukenthal" authorityYear="1887" box="[1020,1320,181,203]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="22" pageNumber="523" phylum="Annelida" rank="species" species="ceylonensis">
<emphasis id="52CFEA9C2C4DF760FC3A8982FB54FF62" box="[1020,1155,181,202]" italics="true" pageId="22" pageNumber="523">P. ceylonensis</emphasis>
<bibRefCitation id="042A4B7F2C4DF760FB4E8982FAFFFF62" author="Kukenthal, W." box="[1160,1320,181,203]" pageId="22" pageNumber="523" pagination="361 - 373" refId="ref13441" refString="Kukenthal, W. (1887) Die Opheliaceen der expedition der Vettore Pisani. Jenaische Zeitschrift fur Naturwissenschaft, 21, 361 - 373." type="journal article" year="1887">Kükenthal, 1887</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="18AC761E2C4DF760FAF38982FA41FF63" box="[1333,1430,181,203]" name="Sri Lanka" pageId="22" pageNumber="523">Sri Lanka</collectingCountry>
)
</keyLead>
</paragraph>
<paragraph id="6004368E2C4DF760FF5189E4FA4BFE8B" blockId="22.[151,1437,151,291]" pageId="22" pageNumber="523">
<keyLead id="DB4A95BB2C4DF760FF5189E4FA4BFE8B" pageId="22" pageNumber="523">
- Twelve pairs of lateral eyespots (
<collectingCountry id="18AC761E2C4DF760FDE589E5FD9FFF40" box="[547,584,210,232]" name="Switzerland" pageId="22" pageNumber="523">CH</collectingCountry>
7–18); anterior and posterior ones smaller. Anterior body region (
<collectingCountry id="18AC761E2C4DF760FB0789E5FB31FF40" box="[1217,1254,210,232]" name="Switzerland" pageId="22" pageNumber="523">CH</collectingCountry>
1–10) with a dense aggregation of dark brown spots and stripes (see Description), following segments with a combination of one anterior dorsal transverse band and two posterior dark brown spots........................
<emphasis id="52CFEA9C2C4DF760FC35883AFB6BFE8A" bold="true" box="[1011,1212,268,290]" pageId="22" pageNumber="523">
<taxonomicName id="A7BB4D0D2C4DF760FC35883AFBBDFE8A" ID-CoL="8RW9V" authority="Parapar &amp; Al-Kandari &amp; Candás &amp; Moreira, 2021" authorityName="Parapar &amp; Al-Kandari &amp; Candás &amp; Moreira" authorityYear="2021" box="[1011,1130,268,290]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="22" pageNumber="523" phylum="Annelida" rank="species" species="zhadanae" status="sp. nov.">
<emphasis id="52CFEA9C2C4DF760FC35883AFBBDFE8A" bold="true" box="[1011,1130,268,290]" italics="true" pageId="22" pageNumber="523">P. zhadanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="49FC57E72C4DF760FBB6883AFB6BFE8A" box="[1136,1212,269,290]" pageId="22" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="18AC761E2C4DF760FB0F883AFAC6FE8B" box="[1225,1297,269,291]" name="Kuwait" pageId="22" pageNumber="523">Kuwait</collectingCountry>
/
<collectingCountry id="18AC761E2C4DF760FADE883AFA41FE8B" box="[1304,1430,269,291]" name="Saudi Arabia" pageId="22" pageNumber="523">Saudi Arabia</collectingCountry>
)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

2792
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