diff --git a/data/03/8E/87/038E87DAFFC0E42FFCFFF936FD46FD63.xml b/data/03/8E/87/038E87DAFFC0E42FFCFFF936FD46FD63.xml new file mode 100644 index 00000000000..a01213e8244 --- /dev/null +++ b/data/03/8E/87/038E87DAFFC0E42FFCFFF936FD46FD63.xml @@ -0,0 +1,387 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + +Crassimurex +( +Pliocrassimurex +) +hirtus + +n. subgen., n. sp. + + + + +( +Figs 21 +A-F; 28C-E) + + + +urn:lsid:zoobank.org:act: +D443EA04-13CB-4AC7-8082-87A2DD4CF2D2 + + + + + +TYPE +MATERIAL +. — + + +Holotype +. +Spain + +• +Llobregat Basin +, +Molins del Rei + +; + +Yellow +and grey clays of +Molins del Rei +(quarry +Anna +); Pliocene (Zanclean); +MNHN +.F.A77751 ( +MNHN +coll.),H: +28 mm +( +Figs 21A,B +; +28E +). + + + +Paratypes +. +Spain + +• 2 spm; same as for the holotype; +MNHN +.F.A77752 ( +MNHN +coll.) ( +Figs 21C, D +; +28D +, +MNHN +.F.A77753 ( +MNHN +coll.) ( +Figs 21E, F +; +28C +). + + + + + +FIG +. 21. — Spiral cords and internal denticles of the outer lip in + +Crassimurex +( +Pliocrassimurex +) +hirtus + +n. subgen., n. sp. +( +A -F +) and + +Crassimurex + +( +s.s. +) + +calcitrapa +( +Lamarck, 1803 +) + +( +G +, +H +) for comparison: +A +, +B +, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei, Llobregat Basin, Spain, Early Pliocene (Zanclean); +C +, +D +, paratype, MNHN.F.A77752, Molins del Rei; +E +, +F +, paratype (juvenile), MNHN.F.A77753 (MNHN coll.), Molins del Rei; +G +, +H +, MNHN.F.A32445 (Morlet coll.), Beynes (Ferme de l’Orme), Paris Basin, France, middle Eocene (Lutetian). Scale bars: A, B, G, H, 10 mm; C-F, 5 mm. Credits: A-F, P. Loubry (MNHN/CNRS); G, H, C. Lemzaouda (MNHN/CNRS). + + + + +ETYMOLOGY +. — From the Latin adjective + +hirtus + +meaning bristly, because of the cord spines of the species; gender masculine. + + + + + +TYPE + +HORIZON +. — Yellow and grey clays of Molins del Rei (quarry Anna). + + + +TYPE + +LOCALITY +. — +Spain +, +Llobregat Basin +, +Pliocene +(Zanclean; see +Salvany & Aguirre 2020 +), +Molins del Rei +(quarry Anna). + + + + +DISTRIBUTION +. — Only known from the +type +locality. + + + + +DESCRIPTION + + +Protoconch not preserved. Teleoconch up to +28 mm +in height ( +holotype +), up to +7.6 mm +in width, biconic in profile, composed of six whorls. Moderately low spire with carinate whorls. Last whorl up to 76% of total length. Apical angle 68°. Spiral sculpture with marked primary cords. First whorl: not preserved; from second to fourth whorl: presence of P1 and P2, P2 close to abapical suture; fifth whorl: appearance of IP; sixth whorl: appearance of abis infrasutural ramp. P1 to P5 and s1 to s3 on convex part of whorl; P6, ADP, MP and ABP on siphonal canal. Axial sculpture with high, sublamellose varices. On second whorl: 9-10 varices; on third whorl: ten varices; from fourth whorl to fifth whorl: 8-10 varices; on sixth whorl: eight varices. On spire, short cord spines on P1-P2; P1 delimiting periphery. On last whorl, short cord spines on IP, P1-P6, ADP, MP, ABP, but more developed on P1, P2 and P3; P2 delimiting periphery. Ovate aperture up to 31% of diameter and up to 74% of length of last whorl (including siphonal canal). Columellar lip, smooth, adherent. Parietal lip smooth, adherent. Outer lip lacking internal denticles. Pseudoumbilicus moderately large. Siphonal canal open, up to 41% of aperture length, slightly dorsally recurved. Aragonitic microstructure. + + + + +COMMENTS +By its shape and its sculpture, + +C. +( +Pliocrassimurex +) +hirtus + +n. subgen., n. sp. +resembles members of + +Crassimurex +Merle, 1990 + +. However, until now, + +Crassimurex + +was not known in the Pliocene and seemed extinct after the Middle Miocene. + +Crassimurex + +is currently represented by two subgenera, + +Crassimurex + +( +s.s. +): early Eocene to middle Eocene, +France +, and + +Crassimurex +( +Eopaziella +) + +: late Eocene to Middle Miocene (Europe). Therefore, the discovery of a species of + +Crassimurex + +in the Early Pliocene of Spain is surprising and extends the stratigraphic range of the genus. + +Crassimurex +( +Pliocrassimurex +) + +differs from + +Crassimurex + +( +s.s. +) and +C. +( +Eopaziella +) by having a more obtuse shoulder angle. In +C. +( + +Pliocrassimurex + +), P2 delimits the periphery on the last whorl, whereas in + +Crassimurex + +( +s.s. +) and +C. +( +Eopaziella +), P1 delimits the periphery. + +Crassimurex + +( +s.s. +) 1990 can share cord spines with +C. +( + +Pliocrassimurex + +), as seen in the +type +species +C. +( +s.s. +) + +calcitrapa +( +Lamarck, 1803 +) + +( +Fig. 21G, H +), but it differs by having varices which are not axially aligned. + +Crassimurex +( +Eopaziella +) + +resembles +C. +( + +Pliocrassimurex + +) by having varices axially aligned but differs in lacking cord spines (see + +Merle +et al. +2011: 178 + +, fig. 62, pl. 144). + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFC3E423FE99FDCEFC48F93E.xml b/data/03/8E/87/038E87DAFFC3E423FE99FDCEFC48F93E.xml new file mode 100644 index 00000000000..4efeefeb9bc --- /dev/null +++ b/data/03/8E/87/038E87DAFFC3E423FE99FDCEFC48F93E.xml @@ -0,0 +1,400 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + + +Flexopteron liancurtense + +n. sp. + + + + + +( +Figs 20 +E-H; 28A, B) + + + +urn:lsid:zoobank.org:act: + +6CA77DFC-A09D-4A63-A2B4-55305DD23 +BEC + + + + + + + +Murex +( +Poirieria +) +foliaceus + +– +Cossmann & Pissarro 1911 +: pl. 36, fig. 169-17 [ +non +Melleville, 1843 +]. + + + + +Poirieria +( +Flexopteron +) +foliacea + +– + +Le Renard & Pacaud 1995: 114 + +, Ref. GA169-17 [ +non +Melleville, 1843 +]. + + + + + +Vauxyrytis liancurtensis + +– + +Merle 1999: 459 + +, pl. 68 (unpublished thesis) ( +nomen nudum +). + + + + + + +TYPE +MATERIAL +. — + +Holotype +. +France + +• +Paris Basin +, +Oise +, Liancourt-Saint-Pierre; +Sables d’Hérouval +(biozone NP12); early +Eocene +( +Ypresian +); +MNHN +.F.J02578 ( +Cossmann +coll.), figured specimen in +Cossmann & Pissarro (1911 +: pl. 39, fig. 169-17), H: +7.7 mm +( +Figs 20E, F +; +28A +). + + + + + +Paratypes +. +France + +• 1 spm; same as for the holotype; +MNHN +.F.A77758 ( +Merle +coll.) ( +Figs 20G, H +; +28B +) + +• + +17 spm; idem; +MNHN +.F.A77759 ( +Merle +coll.) + +• + +9 spm; idem; +MNHN +.F.J17633 ( +Cossmann +coll.); + +• + +1 spm; +Paris Basin +, Oise, +Hérouval +; +Sables d’Hérouval +(biozone NP12); early +Eocene +( +Ypresian +); +MNHN +.F.B63662 ( +Morlet +coll.) + +• + +29 spm; idem; +MNHN +.F.A90913 ( +Pacaud +coll.). + + + + + +ETYMOLOGY +. — From the Latin adjective + +liancurtense + +in reference to the +type +locality, Liancourt-Saint-Pierre (Oise). + + + + +TYPE +HORIZON +. — Sables d’Hérouval (biozone NP12), see +Gély & Lorenz (1991 +: pl. 1), early Eocene (Ypresian). + + +TYPE +LOCALITY +. — +France +, Paris Basin, Oise, Liancourt-Saint-Pierre, see +Fritel (1910: 190) +. + + + + +DISTRIBUTION +. — Paris Basin ( +France +), early Eocene (Ypresian, Hérouval Formation only). + + + + +DESCRIPTION + + +Smooth, conical multispiral protoconch of around three whorls. Sinusigeral scar well marked. Biconic teleoconch up to +7 mm +in height, up to +4 mm +in width (including cord spines). Moderately high spire, up to four subcarinate whorls. Last whorl (fourth whorl) up to 68% of total length. Apical angle 54° including P1 spine. Spiral sculpture with marked primary cords. First whorl: no cord; second whorl: appearance of P2 weak; third whorl: development of P1 spine and appearance of P3 above suture line. Fourth and last whorl: no primary cord on sutural ramp, P1 to P5, s1 and s2 moderately marked, P6 and ADP weak on siphonal canal. Axial sculpture with 11 lamellose varices on first whorl, 13 on second, and 14 on third and fourth. Varices axially aligned in a zig-zag fashion. Ovate aperture up to 36% of diameter and up to 77% of length of the last whorl (including siphonal canal). Columellar lip slightly erect anteriorly, adherent posteriorly, usually smooth or occasionnally with one tubercle. No anal sulcus, no parietal callus. Outer lip with small denticles. D1 to D5 simple, D1 more developed than other denticles. Outer lip not crenulate. Shoulder sinus of P1 spine widely open, shallow. Pseudoumbilicus narrow. Siphonal canal open, up to 36% of the apertural length and slightly dorsally recurved. + + +COMPARISONS + + +Melleville (1843) +described + +Murex foliaceous + +for a small Ypresian + +Flexopteron + +displaying axially zizgaging lamellae and a short P1 spine. Later, +Cossmann (1889) +described + +Murex dyscritus + +which bears less marked primary cords than + +Murex foliaceous + +. However, numerous specimens housed in the MNHN show many transitional forms suggesting these differences represent intraspecific variation. Thus, the older binome + +Murex foliaceous + +takes nomenclatural priority. Moreover, +Cossmann & Pissarro (1911) +illustrated an atypic specimen from the Ypresian of Liancourt-Saint-Pierre (Hérouval Formation), squatter than usual for + +F. foliaceum + +, with a bent siphonal canal, poorly developed P1 spines, more marked primary cords and occasionally a tubercle on the columellar lip. This specimen from Liancourt is not a pathological specimen and all specimens from this locality and Hérouval display similar morphology. They differ from + +F. foliaceum + +(also found at Liancourt-Saint-Pierre) by having spiral cords P1 and P2 appearing earlier in ontogeny. + +Merle +(1999 + +, unpublished thesis) illustrated several other specimens and the ontogenetic changes in their spiral sculpture. He considered this population as a different species and named it + +Vauxyrytis +liancurtensis + +. However, + +Merle +(1999) + +is an unpublished thesis and the name is a +nomen nudum +made available herein. Placement of this species in the genus + +Vauxyrytis +Merle, 1999 + +( +nomen nudum +), an unpublished genus corresponding to + +Beyregrex +Merle + +, +n. gen. +(see below), was based on the shape of the varices, which are not spiny, and the occasional presence of a columellar tubercle. Nevertheless, we found some specimens of + +F. liancurtense + +n. sp. +with P1 spines and several species of + +Flexopteron + +can have a columellar tubercle [e.g., + +F. ogormani +( +Cossmann, 1923 +) + +]. In addition, + +F. liancurtense + +n. sp. +clearly resembles a Tortonian species from NW +France +: + +F. gallicum +( +Landau, Merle, Ceulemans & Van Dingenen, 2019 +) + +, see + +Landau +et al. +(2019 + +: pl. 11). Thus, we attribute this species to + +Flexopteron + +. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFC4E424FEAAFE20FD5DFBDD.xml b/data/03/8E/87/038E87DAFFC4E424FEAAFE20FD5DFBDD.xml new file mode 100644 index 00000000000..26c73e705d4 --- /dev/null +++ b/data/03/8E/87/038E87DAFFC4E424FEAAFE20FD5DFBDD.xml @@ -0,0 +1,166 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + +Genus + +Textiliomurex +Merle, 2011 + + + + + + + +TYPE + +SPECIES +. — + +Murex textiliosus +Lamarck, 1803 + +by original designation. Middle Eocene: Paris basin, +France +and +Hampshire +Basin, +England +. + + + + +COMMENTS +ON +THE +GENUS + + + +Textiliomurex + +shares similarities with + +Pterynotus + +( +s.s. +) and + +Pterynotus +( +Pterymarchia +) + +such as a scabrous surface, columellar denticles and a series of internal denticles including ID, D1 to D6. + +Textiliomurex + +does not display trivaricate axial sculpture and lacks intervarices, usually found in + +Pterynotus + +( +s.s. +). The number of varices on the last whorls of + +Textiliomurex + +is higher (9-11) than in + +Pterymarchia + +(3-5, see +Houart 1995 +). As with + +Pterynotus + +, we cannot give a subfamilial placement to this genus. + +Merle +et al. +(2011: 127) + +included six species in the genus. Recently, +Lozouet (2023) +added (with some doubt) one species from the Chattian of the +Aquitaine +Basin ( + +Textiliomurex +? +problematicus +Lozouet, 2023 + +), and we add a new species from the Lutetian of the Paris Basin herein. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFC4E424FEB5FB63FAC1FB7D.xml b/data/03/8E/87/038E87DAFFC4E424FEB5FB63FAC1FB7D.xml new file mode 100644 index 00000000000..f0ba66cb02a --- /dev/null +++ b/data/03/8E/87/038E87DAFFC4E424FEB5FB63FAC1FB7D.xml @@ -0,0 +1,249 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + + +Textiliomurex chodmonensis + +n. sp. + + + + + +( +Figs 19 +A-D; 27I, J) + + + +urn:lsid:zoobank.org:act: +C498F221-B504-40D3-A082-2DBCAC1E7D78 + + + + + + +TYPE +MATERIAL +. — + +Holotype +. +France + +• +Paris Basin +, +Oise +, +Chaumont-en-Vexin +( +Carrière Darcy +); +Calcaire Grossier +inférieur (biozone NP14); middle +Eocene +( +lower Lutetian +); +MNHN +.F.A81144 ( +Pacaud +coll.), H: +28 mm +( +Figs 19A, B +; +27I +). + + + + + +Paratype +. +France + +• 1 spm; same as for the holotype; +MNHN +.F.A81145 ( +Pons +coll.) ( +Figs 19C, D +; +27J +) + +. + + + + +ETYMOLOGY +. — Name from the +type +locality Chaumont-en-Vexin. This place name comes from the Celtic words Chod “wood” and +Mon +“montain”. + + + + +TYPE +HORIZON +. — Calcaire Grossier inférieur (biozone NP14), see +Gély & Lorenz (1991 +: pl. 1), middle Eocene (lower Lutetian). + + +TYPE +LOCALITY +. — +France +, Paris Basin, Oise, Chaumont-en-Vexin (Carrière Darcy), see + +Ott d’Estevou +et al. +(2014) + +. + + + + +DISTRIBUTION +. — Only known from the +type +locality. + + + + +DESCRIPTION + + +Protoconch not preserved. Fusiform teleoconch up to +28 mm +in height, up to +21 mm +in width. High spire, up to seven rounded whorls. Last whorl (seventh whorl) up to 65% of total length. Apical angle 55°. Spiral sculpture with fine, narrow, well marked primary and secondary cords. First to third whorl not preserved. Fourth whorl: IP, s1 and P2. Fifth whorl: appearance of the secondary cord adis (sutural ramp). Sixth whorl: addition of tertiary cords on sutural ramp and convex part of whorl. Seventh and last whorl: IP, adis and abis (sutural ramp), P1 to P6 (convex part of the whorl); P6 placed on base on convex part of whorl; ADP, MP, ABP (siphonal canal), MP, ABP poorly distinct from secondary cords; primary and secondary cords of same thickness. Surface of shell with numerous threads. Axial sculpture with nine varices on fourth whorl, seven on fifth whorls, six on sixth whorls and five on seventh and last whorl. Nodulose varices. No intervarices. No varical spine. Numerous, fine microlamellae covering entire surface forming scabrous microsculpture. Ovate aperture up to 36% of diameter and up to 70% of length of last whorl (including siphonal canal). Columellar lip adherent, smooth. Parietal lip adherent. Short anal sulcus. No shoulder channel. Outer lip with subobsolete internal denticles. Outer lip finely crenulated. Pseudoumbilicus wide. Siphonal canal open, up to 38% of apertural length, slightly dorsally recurved. + + +COMPARISONS + + + +Textiliomurex chodmonensis + +n. sp. +can be compared to two other species from the middle Lutetian: + +T +. +textiliosus +( +Lamarck, 1803 +) + +and + +T. bernayi +( +Deshayes, 1865 +) + +. + +Textiliomurex textiliosus + +differs in having a shorter spire, a more inflated last whorl, a wider aperture, and a longer siphonal canal bearing a narrower pseudoumbilicus. + +Textiliomurex bernayi + +has angular teleoconch whorls with a marked P1 shoulder, bears well developed internal denticles within the outer lip and cannot be confused with + +T. chodmonensis + +n. sp. +(see + +Merle +et al. +2011: 126 + +, text. fig. 47 and pl. 94 about + +Textiliomurex + +). + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFC7E424FF69FA4EFDECFE02.xml b/data/03/8E/87/038E87DAFFC7E424FF69FA4EFDECFE02.xml new file mode 100644 index 00000000000..ccadacc82e9 --- /dev/null +++ b/data/03/8E/87/038E87DAFFC7E424FF69FA4EFDECFE02.xml @@ -0,0 +1,425 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + +Pterynotus +( +Pterymarchia +) +pelouatensis + +n. sp. + + + + +( +Figs 18 +E-H; 27G, H) + + + +urn:lsid:zoobank.org:act: +4852FE61-DAEE-4316-B3D0-A7A7162A231E + + + + + + + +Pterynotus +( +Pterymarchia +) + +sp. – + + +Merle +et al. +2011: 424 + + +, pl. 90, fig.7a, b. + + + + + + +TYPE +MATERIAL +. — + +Holotype +. +France + +• +Aquitaine +Basin +, +Gironde +, +Saucats +( +Peloua +); +Faluns de Saucats +; Early Miocene ( +lower Burdigalian +); +MNHN +.F.J09945 ( +Neuville +coll.), figured specimen in + +Merle +et al. +(2011: 424 + +, pl. 90, fig. 7), H: +21 mm +( +Figs 18E, F +; +27G +). + + + +Paratype +. +France + +• 1 spm; same as for the holotype; +MNHN +.F.J17632 ( +Neuville +coll.) ( +Figs 18G, H +; +27H +). + + + + + +ETYMOLOGY +. — From the +type +locality, Peloua, Gironde, +France +. + + + + + +TYPE + +HORIZON +. — Faluns de Saucats, Early Miocene (lower Burdigalian), see + +Parize +et al. +(2008: 396) + +. + + + +TYPE + +LOCALITY +. — +France +, +Aquitaine +Basin, Gironde, Saucats (Peloua), seeParize +et al. +(2008: fig. 1). + + + + +DISTRIBUTION +. — Only known from the +type +locality. + + + + +DESCRIPTION + + +Two smooth protoconch whorls preserved delimited by sinusigera scar (protoconch tip not preserved) (see + +Merle +et al. +2011: 122 + +, text-fig. 46E). Teleoconch up to +21 mm +in height, up to +12.5 mm +in width, biconic in profile, composed of five whorls. Moderately high spire with subcarinate whorls. Last whorl up to 73% of total length. Apical angle 64°. Spiral sculpture with marked primary cords. First whorl: appearance of P1 and P2; second whorl: appearance of IP; third whorl: appearance of abis, adis and s2. Fourth whorl: no change or appearance of threads. Fifth whorl: IP, adis and abis, P1 to P6, s1 to s6; s3 and s4 more developed than other secondary cords; P6 slightly atrophied, on base of convex part of whorl; ADP, MP, ads and ms; ABP missing. Surface of shell with numerous threads appearing on fourth whorl. Axial sculpture with thick varices. On first whorl: 10-11 lamellose varices; on second whorl: 10-12 varices; on third whorl: 11-12 varices; on fourth whorl: 10-12 varices; on fifth whorl 8-10 varices. From second whorl to fifth whorl: thick varices. No evidence of intervarices on last whorl. From third to last whorl numerous, fine microlamellae forming scabrous microsculpture covering entire surface. Ovate aperture up to 31% of diameter and up to 72% of length of last whorl (including siphonal canal). Columellar lip adherent posteriorly, becoming progressively more erect anteriorly. One or two weak columellar tubercles. Parietal lip adherent. Outer lip with strong internal denticles: ID, D2 to D6 present, D1 missing; D4 split, D6 poorly developed. Pseudoumbilicus moderately large. Siphonal canal open, up to 43% of aperture length, slightly dorsally recurved. + + +COMPARISONS + + +The European Neogene contains two species of + +Pterynotus +( +Pterymarchia +) + +: + +P. +( +P. +) +pelouatensis + +n. sp. +and + +P. +( +P. +) +sandbergeri +(Hörnes, 1856) + +n. comb. +from the Langhian of Paratethys ( +Austria +, +Hungary +, +Romania +, +Poland +) and the Serravallian of +Turkey +. The type material of + +P. +( +P. +) +sandbergeri + +is housed in Naturhistorisches Museum of +Wien +and contains the figured +syntype +of the species (Hörnes 1856: pl. 51, fig. 5a, b, NHMW 1855/0045/0435). Here, we designate this specimen as the +lectotype +( +Fig. 18I, J +). + +Pterynotus +( +P. +) +sandbergeri + +was often attributed to + +Murexsul +Iredale, 1915 + +( + +Landau +et al. +2013 + +; + +Kovács +et al. +2018 + +; Kovács 2019). Although they share several similarities (biconic shells, internal denticles, columellar denticles), members of + +Murexsul + +have a more rounded aperture, their internal denticles are close together, ID is weaker and their siphonal canal is narrower. The spires of + +P. +( +P. +) +pelouatensis + +n. sp. +and the +lectotype +of + +P. +( +P. +) +sandbergeri + +are very similar, but their last whorls are very different. The +lectotype +of + +P. +( +P. +) +sandbergeri + +has a larger aperture, a surface covered by numerous threads and atrophy of P2, P5 and P6; P4 is more developed and D6 is missing. The differences between the primary cords are great, but in other specimens attributed from + +P. +( +P. +) +sandbergeri + +( +Fig. 18K, L +), the atrophy of P2, P5 and P6 is less strong than in the +lectotype +from +Austria +(see + +Landau +et al. +2013 + +; + +Kovács +et al. +2018 + +; Kovács 2019). Compared to extant +P. +( + +Pterymarchia + +), + +P. +( +P. +) +pelouatensis + +n. sp. +and + +P. +( +P. +) +sandbergeri + +share greatest morphological similarities with + +P. +( +P. +) +martinetana +(Röding, 1798) + +and particularly with specimens from the Red Sea ( + +Merle +et al. +2011 + +: pl. 92, figs 11, 12), but + +P. +( +P. +) +sandbergeri + +differs by having develop cord spines on P1, P2, P4, P5, ADP and MP. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFC9E434FC4BF983FE5FFD81.xml b/data/03/8E/87/038E87DAFFC9E434FC4BF983FE5FFD81.xml new file mode 100644 index 00000000000..aa491bdba70 --- /dev/null +++ b/data/03/8E/87/038E87DAFFC9E434FC4BF983FE5FFD81.xml @@ -0,0 +1,663 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + + +Pronucellopsis pacaudi + + +Merle, Ledon & Goret, +n. gen., n sp. + + +( +Figs 24 +; +28H, I +) + + + + + +urn:lsid:zoobank.org:act: +A1CD0282-7026-43EB-9992-5C106808B8CA + + + + + + + +Murex +( +Muricopsis +) +multistriatus + +– + +Cossmann 1889: 129 + +[ +non +Deshayes, 1865 +]. + + + + + +Jania +( +Muricopsis +) +multistriatus + +– + +Cossmann 1892: 69 + +[ +non +Deshayes, 1865 +]. + + + + +Muricopsis multistriatus + +– +Cossmann & Pissarro 1911 +: pl. 36, fig. 169bis-5 [ +non +Deshayes, 1865 +]. + + + +FIG +. 28. — New species of + +Flexopteron +Shuto, 1969 + +, + +Crassimurex +( +Pliocrassimurex +) + +n. subgen. +, + +Nucellopsis +Merle, 2005 + +and + +Pronucellopsis +Merle + +, +n. gen. +: +A +, +B +, + +Flexopteron liancurtense + +n. sp. +: +A +, holotype, MNHN.F.J02578 (Cossmann coll.), Liancourt-Saint-Pierre, France, Ypresian; +B +, paratype: MNHN.F.A77758 ( +Merle +coll.), Liancourt-Saint-Pierre; +C -E +, + +Crassimurex +( +Pliocrassimurex +) +hirtus + +n. subgen., n. sp. +: +C +, paratype (juvenile), MNHN.F.A77753, Molins del Rei, Spain, Zanclean; +D +, paratype, MNHN.F.A77752, Molins del Rei; +E +, holotype, MNHN.F.A77751 (MNHN coll.), Molins del Rei; +F +, +G +, + +Nucellopsis parisiensis + +n. sp. +: +F +, holotype, MNHN.F.R64955 ( +Merle +coll.), Saint-Witz (Le Guépelle), France, Bartonian; +G +, paratype, MNHN.F.A90963 (Faullummel coll.), Saint-Witz (Le Guépelle); +H +, +I +, + +Pronucellopsis pacaudi +Merle, Ledon & Goret + +, +n. gen., n. sp. +: +H +, holotype, MNHN.F.J02587 (Cossmann coll.), Châlons-sur-Vesle, France, Thanetian; +I +, paratype, MNHN.F.A90829 (Pacaud coll.), Jonchery-sur-Vesle, France, Thanetian. Scale bars: 5 mm. Scale bar: A, B, 3 mm; C, D, F-I, 5 mm; E, 10 mm. Credits: P. Loubry (MNHN/CNRS). + + + + +“ + +Murex +” +multistriatus + +– + +Glibert 1963: 30 + +, no. 8 [ +non +Deshayes, 1865 +]. + + + + + +Trophonopsis multistriata + +– + +Le Renard & Pacaud 1995: 114 + +, Ref.GA-169bis-5 [ +non +Deshayes, 1865 +]. + + + + + +TYPE +MATERIAL +. — + + +Holotype +. +France + +• +Paris Basin +, +Marne +, +Châlons-sur-Vesle +; + +Sables +de Châlons-sur-Vesle + +, biozone NP9; +Paleocene +( +Thanetian +); +MNHN +.F.J02587 ( +Cossmann +coll.), figured specimen in +Cossmann & Pissarro (1911 +: pl. 36, fig. 169bis-5), H: +22 mm +( +Figs 24A, B +; +28H +). + + + + + +Paratypes +. +France + +• 3 spm; same as for the holotype; +MNHN +.F.A90831 ( +Pacaud +coll.) + +• + +2 spm; idem; +MNHN +.F.A90832 ( +MNHN +coll.) • 7 spm; + + +Paris Basin +, Marne, +Jonchery-sur-Vesle + +; + +Paleocene +(Thanetian); +MNHN +.F.A90833 ( +Staadt +coll.) + +• + +1 spm; idem; +MNHN +.F.A90829 ( +Pacaud +coll.) ( +Figs 24C, D +; +28I +) + +• + +8 spm; idem; +MNHN +.F.A90830 ( +Pacaud +coll.) + +• + +6 spm; +Paris Basin +, +Oise +, +Abbecourt +(Bois-des-Godins), +Paleocene +( +Thanetian +); +MNHN +.F.A90834 ( +Pacaud +coll.). + + + + + +ADDITIONAL +MATERIAL +. — + +1 spm; Paris Basin, Marne, Jonchery-sur-Vesle; Paleocene (Thanetian); UPMC (Munier-Chlamas coll.), figured specimen of + +Merle +(1999 + +: pl. 66, b-d) ( +Fig. 24E, F +) + +• + +1 spm; idem; MNHN-IM (Staadt coll.), figured specimen of + +Merle +(1999 + +: pl. 66, c-e) ( +Fig. 24G, H +) + +• +5 spm; idem; MNHN.F.A91007 (Staadt coll.). + + + + +ETYMOLOGY +. — Dedicated to Jean-Michel Pacaud (MNHN) who has for a long time observed that + +Murex multistriatus +Deshayes, 1865 + +is a junior synonym of + +Fusus angusticostatus +Melleville, 1843 + +(now + +Lyrofusus angusticostatus + +). + + + + +TYPE +HORIZON +. — Sables de Châlons-sur-Vesle, biozone NP9, see + +Huyghe +et al. +(2015 + +: fig. 2), Paleocene (Thanetian). + + +TYPE +LOCALITY +. — +France +, Paris Basin, Marne, Châlons-sur-Vesles, see +Fritel (1910: 77) +. + + + + +DISTRIBUTION +. — Paris Basin ( +France +), Paleocene (Thanetian), Sables de Bracheux and Sables de Châlons-sur-Vesle. + + + + +FIG +. 29. — Histogram showing the evolution of the species richness during the Paleogene in Western and Eastern Atlantic, Tethys and Austral Pacific (original histogram of + +Merle +et al. +(2011: 38 + +, fig. 11B) updated from this paper, +Pacaud 2018 +; + +Pacaud +et al. +2017 + +; + +Merle +& Pacaud 2019 + +; +Kovács & Vicián 2019 +; + +Merle +& Landau 2020 + +; + +Ledon +et al. +2023 + +; +Lozouet 2023 +). It illustrates the Switch phase with the increase of the proportion of the Tethysian and Pacific species. + + + + +DESCRIPTION + + +Unknown protoconch. Teleoconch, up to +22 mm +in height, up to +12.5 mm +in width, subfusiform in profile, composed of seven whorls. High spire with rounded whorls. Last whorl up to 67% of total length. Apical angle 45°. Spiral sculpture with weak primary cords. First and second whorls: no spiral scultpure; third and fourth whorls, P1 weak. Fifth whorl: appearance of P2. Sixth whorl: appearance of s1. Last whorl and seventh whorl: P1 to P6, s1, s2, s3 and s4 (convex part of whorls), ADP, MP (siphonal canal). Axial sculpture on first to second whorl: not observed; third whorl: nine varices; fourth whorl: 14 varices; fifth whorl: 14-16 varices; sixth whorl: 11-19 varices; seventh whorl: 12-23. Varices finely lamellose. No spine on P1. Wide, ovate aperture up to 45% of diameter and up to 84% of length of last whorl (including siphonal canal). Posterior sinus deep, no shoulder sinus. Columella slightly angular. Columellar lip smooth, narrow and adherent. Outer lip with occasional denticles. D1 to D6 small, poorly developed, ID missing. Siphonal canal, short, open, up to 23% of aperture length, slightly dorsally recurved. Pseudoumbilicus narrow. + + +COMPARISONS +Deshayes (1865) +described + +Murex multistriatus + +and illustrated a specimen bearing a buccinid shape (pl. 88, fig. 810). In fact, this species is a junior synonym of + +Fusus angusticostatus +Melleville, 1843 + +(now + +Lyrofusus angusticostatus + +). However, +Cossmann (1889) +andCossmann & Pissarro (1911) confused this buccinoid species with a Thanetian +Muricidae +, which they mistakenly called + +Murex multistriatus + +. We described this species as + +Pronucellopsis pacaudi +Merle, Ledon & Goret + +, +n. gen., n. sp. +It can be compared to + +Nucellopsis ponsi +Merle, + + + +2005c or + +N. plicatilis +( +Deshayes, 1835 +) + +from the late Paleocene-early Eocene of the Paris Basin. Juveniles of + +P. pacaudi +Merle, Ledon & Goret + +, +n. gen., n. sp. +share with these + +Nucellopsis + +a buccinid shape, but at the end of its growth + +P. pacaudi +Merle, Ledon & Goret + +, +n. gen., n. sp. +displays a subfusiform shape with a high spire. The sculptures (both axial and spiral) of + +P. pacaudi +Merle, Ledon & Goret + +, +n. gen., n. sp. +and the internal denticles of outer lip are also less developed. Because of its high spire, + +P. pacaudi +Merle, Ledon & Goret + +, +n. gen., n. sp. +resembles some elongated + +Beyregrex sarroniensis +( +Carez, 1879 +) + +, +n. comb. +but in + +B. sarroniensis + +n. comb. +the spiral sculpture and the internal denticles are always more strongly developed. In addition, P1 spine is present in + +B. sarroniensis + +n. comb. +, whereas it is missing in + +P. pacaudi +Merle, Ledon & Goret + +, +n. gen., n. sp. + +CONCLUSION + +Based on new field material and a re-examination of collection material, this paper presents 24 new species, four new genera, one new subgenus and one new subfamily and therefore it deeply increases the knowledge of muricid paleobiodiversity. At species level, this paper highlights the strong diversity of the Eocene + +Timbellus + +which was underestimated. In addition, with 19 new species, this paper highlights the species richness of European Paleogene muricids and allows updating the diagram showing its evolution given by + +Merle +et al. +(2011: 36 + +, fig. 21b). This update ( +Fig. 29 +) illustrates the high muricid species richness in the Lutetian and confirms the Switch phase (Oligocene) which displays an increase of the proportion of Tethysian and Pacific species after the Eocene. + + +At supraspecific level, this paper improves the understanding of European Eocene-Oligocene basal muricids ( + +Pronucellopsis + +n. gen, + +Beyregrex +Merle + +, n. gen, + +Pseudotrophonopsis +Merle + +, +n. gen. +, +Nucellopsinae Merle +, +n. subfam. +), that have, until now, been shoehorned uncomfortably into extant genera and subfamilies. Supraspecific taxonomy today relies heavily on molecular data, but this paper illustrates the value of using precise homologies basing data gathered on the careful observation of shell characters with which one can construct fossil phylogenies of groups that have no living descendants (see also +Merle +2002, 2005b, c). + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFCAE429FE96F8C5FAEBFDC3.xml b/data/03/8E/87/038E87DAFFCAE429FE96F8C5FAEBFDC3.xml new file mode 100644 index 00000000000..252fa89617f --- /dev/null +++ b/data/03/8E/87/038E87DAFFCAE429FE96F8C5FAEBFDC3.xml @@ -0,0 +1,636 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + + +Nucellopsis parisiensis + +n. sp. + + + + + +( +Figs 23 +A-F; 28F, G) + + + +urn:lsid:zoobank.org:act: +36606EEC-3983-4334-8A8C-E7FAF7894FEE + + + + + + +Muricopsis depauperatus + +– +Cossmann & Pissarro 1911 +: fig. 169bis-2 [ +non +Deshayes, 1865 +]. + +Muricopsis +sp. + + +non +depauperata + +– Merle +in +Le Renard & Pacaud 1995: 114 +, réf.: GA 169bis-2. + + + +Nucellopsis dudariensis + +– +Merle 2005c +[ +non +Strausz, 1966 +]. + + + + +TYPE +MATERIAL +. — + + +Holotype +. +France + +• Paris Basin, Val d’Oise, Saint-Witz (Le Guépelle); Sable du Guépelle (biozone NP16); middle Eocene (early Bartonian, Auversian); MNHN.F.R64955 ( +Merle +coll.), figured specimen in + +Merle +(2005c + +: fig. 2H), H: +21.5 mm +( +Figs 23E, F +; +28F +). + + + +Paratypes +. +France + +• 1 spm; same as for the holotype; MNHN.F.A71797 (Pons coll.), figured specimen in + +Merle +(2005c + +: fig. 1G) + +• + +1 spm; idem; MNHN.F.A71798 (Pons coll.), figured specimen in + +Merle +(2005c + +: fig. 1H, I) + +• + +1 spm; idem; MNHN.F.J02585 (Cossmann coll.); figured in +Cossmann & Pissarro (1911 +: pl. 36 fig. 169bis-2) + +• + +1 spm; idem; MNHN.F.A90963 (Faullummel coll.) ( +Figs 23C, D +; +28G +) + +• + +1 spm; idem; MNHN.F.A90964 (Faullummel coll.) ( +Fig. 23A, B +) + +• +6 spm; idem; MNHN.F.A90962 (Faullummel coll.). +ADDITIONAL +MATERIAL +. — + + +France + +• 1 spm; Paris Basin, Val d’Oise, Saint-Witz (Le Guépelle); Sable du Guépelle (biozone NP16); early Bartonian; MNHN.F.A27771 (Pacaud coll.) + +• +1 spm; Paris Basin, Val d’Oise, Auvers-sur-Oise (Bois-le-Roi); idem; MNHN.F.A27770 (Pacaud coll.) +• +1 spm; Paris Basin, Aisne, Nogent-l’Artaud; idem; MNHN.F.A90565 (Margerie coll.) +• +1 spm; Paris Basin, Aisne, Bézu-le-Guéry; idem;MNHN.F.A90825 (Faullummel coll.) +• +1 spm; Paris Basin, Aisne, Mont-Saint-Martin; idem; MNHN.F.J07787 (UMPC coll.) +• +1 spm; Paris Basin, Seine-et-Marne, Sainte-Aulde (Caumont); idem; MNHN.F.A90826 – (Faullummel coll.) +• +1 spm; Paris Basin, Seine-et-Marne, Luzancy; idem; MNHN.F.A90827 (Faullummel coll.) +• +Paris Basin, Oise, Loisy; idem; MNHN.F.A90828 (Faullummel coll.). + + + + +FIG +. 26. — New species of + +Timbellus +De Gregorio, 1885 + +: +A +, +B +, + +Timbellus occidentalis + +n. sp. +: +A +, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian; +B +, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian; +C +, +D +, + +T.calciacus + +n. sp. +: +C +, holotype,MNHN.F.A90536 ( +Merle +coll.), Chaussy (Les Garennes), France, Lutetian; +D +, paratype, MNHN.F.A90537 ( +Merle +coll.), Chaussy (Les Garennes); +E +, +F +, + +T. longicanalis + +n. sp. +: +E +, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; +F +, paratype, MNHN.F.A24100, Fresville; +G -I +, + +T. magnei + +n. name: +G +, MNHN.F.A32508 ( +Merle +coll.), juvenile specimen, Gan (Tuilerie), France; +H +, MNHN.F.A90545 ( +Merle +coll.), juvenile specimen with protoconch, Gan (Tuilerie); +I +, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); +J -L +, + +T. curvispina + +n. sp. +: +J +, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; +K +, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; +L +, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; +M +, + +T. sixi + +n. sp. +, holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; +N -P +, + +T. radulfiensis + +n. sp. +: +N +, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; +O +, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; +P +, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm. + + + + +FIG +. 27. — New species of + +Timbellus +De Gregorio, 1885 + +, + +Pterynotus +Swainson, 1833 + +( + +Pterymarchia +Houart, 1995 + +), + +Textiliomurex +Merle, 2011 + +, + +Ponderia +Houart, 1986 + +and + +Flexopteron +Shuto, 1969 + +: +A -C +, + +Timbellus submicropterus + +n. sp. +: +A +, holotype, MNHN.F.A24111 (MNHN coll.), Fresville, France, Lutetian; +B +, paratype, MNHN.F.A24126 (Munier-Chalmas coll.), Gourbesville, France, Lutetian; +C +, paratype, UCBL-EM 33438 (Bourdot coll.), Fresville; +D +, + +T. +cf. +submicropterus + +n. sp. +, MNHN.F.A90557 (Brébion coll.), Saint-Aignan-Grand-Lieu (Pierre-Aiguë), France, Bartonian; +E +, +F +, + +Pterynotus +( +Pterymarchia +) +gaasensis + +n. sp. +: +E +, holotype, MNHN.F.A32492 ( +Merle +coll.), Gaas (Espibos), France, Rupelian; +F +, paratype, MNHN.F.A90554 ( +Merle +coll.), Gaas (Espibos); +G +, +H +, + +P. +( +P. +) +pelouatensis + +n. sp. +: +G +, holotype, MNHN.F.J09945, Saucats (Peloua), France, Burdigalian; +H +, paratype, MNHN.F.J17632, Saucats (Peloua); +I +, +J +, + +Textiliomurex chodmonensis + +n. sp. +: +I +, holotype, MNHN.F.A81144 (Pacaud coll.), Chaumont-en-Vexin (Carrière Darcy), France, Lutetian; +J +, paratype, MNHN.F.A81145 (Pons coll.), Chaumont-enVexin (Carrière Darcy); +K +, +L +, + +Ponderia remyi + +n. sp. +: +K +, holotype, MNHN.F.A90548 (Remy coll.), Fresville, Lutetian; +L +, paratype, MNHN.F.A90555, Fresville (Remy coll.); +M +, +N +, + +Flexopteron constantinense + +n. sp. +: +M +, paratype, MNHN.F.A77761 (MNHN coll.), Carentan-les-Marais (Cauvin), France, Priabonian; +N +, holotype, MNHN.F.A77760 (MNHN coll.), Carentan-les-Marais (Cauvin). Scale bars: A-D, I-L, 10 mm; E-H, N, 5 mm; M, 3 mm. Credits: A, B, D-F, N, P. Loubry (MNHN/ CNRS); C, E. Robert (UCBL). + + + + +ETYMOLOGY. — From the Latin adjective + +parisiensis + +, the geographic origin of this species being the Paris Basin. + + + + + +TYPE + +HORIZON +. — Sable du Guépelle (biozone NP16, see +Gély & Lorenz (1991 +: pl. 1), middle Eocene (early Bartonian, Auversian). + + + +TYPE + +LOCALITY +. — +France +, Paris Basin, Val d’Oise, Saint-Witz (Le Guépelle), +INPN (2024) +. + + + + +DISTRIBUTION +. — Paris Basin ( +France +), middle Eocene (early Bartonian, Auversian). + + + + +DESCRIPTION + + +Smooth, multispiral protoconch of up to 2.5 whorls (see + +Merle +1999 + +: pl. 23, fig. 5, +holotype +). Biconic teleoconch, up to +21.5 mm +in height, up to +14 mm +in width, biconic in profile and composed of six whorls. Moderately high spire with rounded or subcarinate whorls. Last whorl up to 73% of total length. Apical angle 64°. Spiral sculpture with strongly marked primary cords. First whorl: appearance of P1, s1, P2; P1 less marked than s1-P2; from second to third whorl, no addition of cord, P1 of equal strength to s1-P2; on fourth whorl, appearance of IP poorly marked. Fifth whorl: no addition of cord. Last whorl: IP (weak), P1 to P6, P6 not atrophied (on convex part of whorl), ADP, MP and ABP; s1 to s3 dominant, equal in strength to primary cords. First to second whorl: twelve varices. Third to fourth whorl: eleven varices. Fifth whorl: ten varices. Sixth whorl: nine varices. Varices more or less sublamellose. No true spine except angular point on P1. Suboval aperture up to 31% of diameter and up to 66% of length of last whorl (including siphonal canal). Columella slightly angular. Columellar lip narrow, adherent, smooth or with one tubercle. Outer lip with five or six denticles including D1 to D6. D1 appearing late in ontogeny. D6 occasional. ID missing. Siphonal canal, short, open, up to 31% of aperture length, slightly dorsally recurved. Pseudo-umbilicus deep, rather wide. + + +COMPARISONS + + + +Nucellopsis ponsi +Merle, 2005c + +differs from + +N. parisiensis + +n. sp. +, + +N. plicatilis +( +Deshayes, 1835 +) + +and + +N. dudariensis +(Strauzs, 1966) + +by having a deeper posterior sinus and weaker spiral cords. Its columellar lip is occasionally ornamented by a weak denticle mid-columella, whereas it is usually smooth in the other three species. + +Nucellopsis parisiensis + +n. sp. +, + +N. plicatilis + +and + +N. dudariensis + +bear more numerous cords on the siphonal canal (MP and ABP) and more secondary cords than + +N. ponsi + +. + +Nucellopsis parisiensis + +n. sp. +differs from + +N. plicatilis + +by the appearance of P1 and s1 on the first teleoconch whorls, by having more angular teleoconch whorls, and by usually lacking D6. Both species share a multispiral protoconch. + +Nucellopsis parisiensis + +n. sp. +was erroneously identified as + +Jsowerbya depauperata +( +Deshayes, 1865 +) + +by +Cossmann & Pissarro (1911) +in spite of numerous differences, such as a narrower shell shape, IP on the sutural ramp and a deeper anal sulcus ( + +Merle +2005b + +). + +Merle +(1999 + +, +2005c +) identified + +N. parisiensis + +n. sp. +as + +N. dudariensis + +due to several similarities with the +holotype +of + +Cantharus +( +Pollia +) +dudariensis + +( +Fig. 23G, H +) from the upper Lutetian of +Hungary +[apical angle near 64°, subangular whorls, thick varices, poorly developed shoulder sinus, five internal denticles (D1 to D5, D6 missing), angular columella with one small tubercle]. These similarities suggest that + +N. parisiensis + +n. sp. +and + +N. dudariensis + +are closely related. However, close examinaton of the +holotype +of + +N. dudariensis + +( +Fig. 23G, H +) reveals several differences with the Parisian species, such as the late appearance of s1 on the last whorl, s1 less coarse than the primary cords P1 and P2, fewer secondary cords (only s1 instead s1 to s3), and the stronger development of IP. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFCFE42FFF11FCC3FE1BF858.xml b/data/03/8E/87/038E87DAFFCFE42FFF11FCC3FE1BF858.xml new file mode 100644 index 00000000000..b2c4c028d57 --- /dev/null +++ b/data/03/8E/87/038E87DAFFCFE42FFF11FCC3FE1BF858.xml @@ -0,0 +1,189 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + +Informal group + +Jsowerbya + +, + +Beyregrex +Merle + +, n. gen, + +Pseudotrophonopsis +Merle + +, +n. gen. + + + + + +DIAGNOSIS +. — Shell medium sized for family, biconic. P6 not atrophied. IP, ADP, MP and ABP present. Poorly developed axial sculpture formed by slightly lamellose varices. Posterior sinus of sutural ramp shallow. Shoulder sinus of P1 reduced or missing. ID missing, D1 to D6 present. Columellar lip occasionally ornamented by 1-3 tubercles. Microstructure of three aragonite layers. + + +INCLUDED +GENERA +. — + +Jsowerbya +Merle, 2005b + +, + +Beyregrex +Merle + +, +n. gen. +and + +Pseudotrophonopsis +Merle + +, +n. gen. + + + + +DISCUSSION + + +This informal group includes basal taxa often erroneously attributed to the +Muricopsinae +(synonym of +Aspellinae +) because of their biconic shape and the presence of 1-3 columellar tubercles or attributed to the +Trophoninae +, because of the more or less lamellose varices. In the phylogenetic analysis, they represent basal taxa (Clades G, H, I, J, L of Winclada [ +Fig. 1 +; Appendix 3] and Clades 44, 43, 42, 41, 40 of PAUP [Appendix 5]). They essentially differ from members from the other subfamilies by having a combination of characters including a posterior sinus, a D6 denticle, occasional columellar tubercles, and by lacking the ID denticle. The primary cords appear late in ontogeny and IP is missing in the most basal species of this group (e.g., + +Beyregrex sarroniensis +( +Carez, 1879 +) + +, +n. comb. +) from the Thanetian, whereas in the derived members (e.g., members of + +Jsowerbya +Merle, 2005b + +) they appear earlier, and the primary cord IP is often present. From an evolutionary point of view, members of this group are diversified from the late Paleocene to the middle Eocene with + +Beyregrex +Merle + +, +n. gen. +and + +Jsowerbya + +. The decline of the group begins at the Priabonian in which only + +Beyregrex +Merle + +, +n. gen. +and + +Pseudotrophonopsis +Merle + +, +n. gen. +are recorded. + +Beyregrex +Merle + +, +n. gen. +survives after the Eocene/ Oligocene climatic crisis and is known in the Early Miocene (Burdigalian) with + +Beyregrex cedillatus +( +Cossmann & Peyrot, 1924 +) + +, +n. comb +.. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFEAE40BFC49F9AEFD08F85F.xml b/data/03/8E/87/038E87DAFFEAE40BFC49F9AEFD08F85F.xml new file mode 100644 index 00000000000..887681c67ac --- /dev/null +++ b/data/03/8E/87/038E87DAFFEAE40BFC49F9AEFD08F85F.xml @@ -0,0 +1,313 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + + +Favartia jansseni + +n. sp. + + + + + +( +Figs 4 +A-D; 25E, F) + + + +urn:lsid:zoobank.org:act: +D34B3691-CAE8-4FB4-9492-05EFDF71FCD3 + + + + + + + +Favartia collega + +– + +Janssen 1984: 214 + +, pl. 59, fig.7 [ +non +Boettger,1906 +]. + + + + +Favartia +sp. 2 + +– Merle +et al. +2022: pl. 58, figs 3, 4. + + + + + +TYPE +MATERIAL +. — + +Holotype +. +The Netherlands + +• +North Sea Basin +, +Miste +, +Winterswijk +; +Miste +bed; +Middle Miocene +(middle Langhian); +RGM +.225979 (figured specimen in +Janssen [1984 +: pl. 59, fig. 7]), H: +12.5 mm +( +Figs 4A, B +; +25E +). + + + + + +Paratype +. +The Netherlands + +• 1 spm; same as for the holotype; +MNHN +.F.A77748 ( +Ledon +coll., +Figs 4C, D +; +25F +) + +. + + + + +ETYMOLOGY +. — Dedicated to Arie W. Janssen for his paleontogical and malacological work. + + + + +TYPE +HORIZON +. — Miste bed, Middle Miocene (middle Langhian), see +Janssen (1984) +and +Schneider & Heissig (2005) +. + + +TYPE +LOCALITY +. — +The Netherlands +, North Sea Basin, Miste, Winterswijk (see +Janssen 1984 +). + + + + +DISTRIBUTION +. — Only known from the +type +locality. + + + + +DESCRIPTION + + +Protoconch not preserved. Delicate teleoconch up to +12.5 mm +in height, up to +7.5 mm +in width, composed of five whorls. Moderately high spire with subangular whorls. Last whorl up to 72% of total length. Apical angle 72° (including spines) and 48° (excluding spines). Spiral sculpture with strong primary cords. From first to fourth whorl: P1 and P2; on fourth whorl, appearance of IP. Fifth whorl: IP and abis, P1 to P5, P6 atrophied (on siphonal canal), ADP and MP present, ABP missing. Surface of P1, P2 and P3 bearing deep spiral grooves. Second whorl: 8-9 varices; from third to fourth whorl: 8 varices; fifth whorl: 6-7 varices. Adapertural face of varices with some imbricated growth lamellae. Axial ornamentation forming short spines on P1 to P5, ADP and MP. Axial microsculpture with small erect lamellae. Suboval aperture up to 29% of diameter and up to 71% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, erect anteriorly. Outer lip lacking denticles. Siphonal canal open, up to 42% of apertural length and slightly dorsally recurved. Pseudo-umbilicus rather wide. + + +COMPARISONS + + +This rare species from the Langhian of Miste (North Sea) was identified as + +Favartia collega +( +Boettger, 1906 +) + +by +Janssen (1984) +. The +holotype +of + +F. collega + +(SMF +XII2258 +a, figured in +Merle +et al. +2022: pl. 58, fig. 2) comes from the Langhian of Costeiu de Sus ( +Romania +, Paratethys Sea). It differs from + +F. jansseni + +n. sp. +by having a slenderer shell, a rounded shoulder sinus in the aperture, and weak internal denticles within the outer lip (D1 to D5). Secondary cords are poorly developed, but the +holotype +of + +F. collega + +is a juvenile specimen of only +7.5 mm +height. + +Favartia collega + +and + +F. czjzeki +( +Hoërnes & Auinger, 1885 +) + +from the Langhian of +Austria +(Paratethys Sea) and +Hungary +( + +Kovács +et al. +2018 + +) share a slender shell, a rounded shoulder sinus, and a multispiral protoconch ( + +Kovács +et al. +2018 + +). The shells of + +F. collega + +and + +F. czjzeki + +are very similar, and therefore, + +F. collega + +is considered to be a junior synonym of + +F. czjzeki + +herein. The +syntype +of + +F. czjzeki + +(NHMW 1855/0045/008 figured in +Merle +et al. +2022: pl. 58, fig. 1) and other adult specimens of this species display the secondary cords (s1, s2, s3 and s5) and weak internal denticles missing in + +F. jansseni + +n. sp. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFEBE40BFC13FDC0FC64FD2F.xml b/data/03/8E/87/038E87DAFFEBE40BFC13FDC0FC64FD2F.xml new file mode 100644 index 00000000000..a714567d1fa --- /dev/null +++ b/data/03/8E/87/038E87DAFFEBE40BFC13FDC0FC64FD2F.xml @@ -0,0 +1,97 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + +Subgenus + +Gracilimurex +Thiele, 1929 + + + + + + +TYPE +SPECIES +. — + +Murex bicolor +Thiele, 1929 + +(synonym of + +Aspella bakeri +Hertlein & Strong, 1951 + +) by monotypy. Recent: Southern Gulf of +California +. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFECE40DFEA7F8A7FD4FF85F.xml b/data/03/8E/87/038E87DAFFECE40DFEA7F8A7FD4FF85F.xml new file mode 100644 index 00000000000..9ddcc6f076b --- /dev/null +++ b/data/03/8E/87/038E87DAFFECE40DFEA7F8A7FD4FF85F.xml @@ -0,0 +1,361 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + + +Muricopsis pontileviensis + +n. sp. + + + + + +( +Figs 3 +A-D; 25A, B) + + + +urn:lsid:zoobank.org:act: +0D061422-80CC-47A5-897C-47EA1FBA313F + + + + + + +Muricopsis dujardini + +– +Merle 1999 +: pl. 12, fig. D, pl. 13, fig. D, pl. IX, figs 5, 6 [ +non + +Murex dujardini +Peyrot, 1938 + +]. + + + +Muricopsis +sp. 2 + +– Merle +et al. +2022: 63, 70, fig. 29D; pl. 3, figs 4-6. + + + + + +TYPE +MATERIAL +. — + +Holotype +. +France + +• +Loire +Basin +, +Loir-et-Cher +, +Pontlevoy +; + +Faluns du Blésois + +(mammal biozone MN5b); +Middle Miocene +(base of the +Langhian +); +MNHN +.F.A71120 ( +Dollfus +coll.), figured specimen in +Merle +et al. +(2022: pl. 3, fig. 4), H: +14 mm +( +Figs 3A, B +; +25A +). + + + + + +Paratypes +. +France + +• 1 spm + +; +same data as for the holotype +; +MNHN +.F.A71122, figured specimen inMerle +et al. +(2022: pl. 3, fig. 5) • 11 spm; +idem +; +MNHN +.F.A77742 ( +Dollfus +coll.) • 1 spm; +idem +; +MNHN +.F.A77743 ( +Dollfus +coll., +Figs 3C, D +; +25B +) • spm; +idem +; +MNHN +.F.B26846 ( +MNHN +coll.) • 5 spm; +idem +; +MNHN +.F.A77744 ( +Lhomme +coll.); • 2 spm; + +Loire +Basin +, Indre-et-Loire, +Paulmy +( +Pauvrelay +) + +; +idem +; +MNHN +.F.A77745 ( +Faullummel +coll.). + + + + +ETYMOLOGY. — The name + +pontileviensis + +refers to the +type +locality: Pontlevoy. + + + + +TYPE +HORIZON +. — Faluns du Blésois (mammal biozone MN5b), see +Ginsburg (2000) +, Middle Miocene (base of the Langhian). + + +TYPE +LOCALITY +. — +France +, +Loire +Basin, Loir-et-Cher, Pontlevoy (see + +Macaire +et al. +2020 + +). + + + + +DISTRIBUTION +. — +Loire +Basin ( +France +), Faluns du Blésois, Middle Miocene (Langhian). + + + + +DESCRIPTION + + +Paucispiral and rounded protoconch of 1.5 whorls.Teleoconch up to +14 mm +in height, up to +6.5 mm +in width, biconic in profile, composed of six whorls. Moderately high spire with subcarinate whorls. Last whorl up to 67.7% of total length. Apical angle 42°. Spiral sculpture with marked primary cords. First whorl: appearance of P1 and P2 (P2 more developed than P1) at beginning of whorl and appearance of IP on second half of whorl; second to fifth-sixth whorl: no change, but sometimes s3 placed slightly above suture. Sixth whorl: IP (sutural ramp), P1 to P5 (convex part of whorl, P6 atrophied (on siphonal canal), ADP and MP well developed, ABP poorly developed. Secondary cords s1, s4, s5 and s6 appear between end of fourth whorl and fifth whorl; s2 and s3 appear earlier. On first whorl: 14 varices; from second to fourth whorl: 10-12 varices; from fifth to sixth whorl: 10-11 varices. On spire axial ornamentation forming small nodules placed on IP, P1 and P2. P1 and P2 nodules more developed than IP. On convex part of last whorl, P1 to P4 nodules more or less equally developed; P5 slightly more developed; on siphonal canal, P6 not atrophied, ADP and MP more developed than ABP when present. Ovate aperture up to 42% of diameter and up to 70% of length of last whorl (including siphonal canal). Columellar lip narrow, entirely adherent, bearing two weak tubercles anteriorly. No anal sulcus, no parietal callus. Outer lip with strong internal denticles. Series of denticles including: ID, D2 to D5. D1 missing. D2 stronger than other denticles. Outer lip not crenulate. Siphonal canal open, up to 33% of apertural length and slightly dorsally recurved. Pseudoumbilicus narrow. + + +COMPARISONS + + +This species is comparable to + +Muricopsis crassicostata +(Benoist, 1880) + +from the late Oligocene/Early Miocene of +Aquitaine +Basin ( +France +) and + +M. dujardini +Peyrot, 1938 + +from the Middle Miocene of +Loire +Basin ( +France +). + +Muricopsis crassicostata + +clearly differs from + +M. pontileviensis + +n. sp. +in being larger ( +19-23 mm +for 6-7 whorls vs 12-14 for 5.5-6 whorls) and having a multispiral protoconch of three convex whorls (for more comparative details, see + +Merle +1999: 348 + +, fig. 115). + +Muricopsis dujardini + +and + +M. pontileviensis + +n. sp. +both occur at Pontlevoy. Specimens of + +M. dujardini + +are larger ( +19-24 mm +for 6-6.5 whorls), but they are mainly distinguishing from + +M. pontileviensis + +n. sp. +by having short spines on P1 to P +5 in +which P3 is atrophied and by having stronger denticles within the outer lip. + +Muricopsis pontileviensis + +n. sp. +never bears spines. + +Merle +(1999) + +wrongly attributed studied specimens of + +M. pontileviensis + +n. sp. +to + +M. dujardini + +, however, he demonstrated that they display a paedomorphic shape (particularly because of the post-displacement of the secondary cords s1, s4 and s6) compared to larger shells as + +M. crassicostata + +. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFEFE40FFF10F8C2FB2CF85F.xml b/data/03/8E/87/038E87DAFFEFE40FFF10F8C2FB2CF85F.xml new file mode 100644 index 00000000000..fbad6d43290 --- /dev/null +++ b/data/03/8E/87/038E87DAFFEFE40FFF10F8C2FB2CF85F.xml @@ -0,0 +1,324 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + +Paleochicoreus bicostatus +( +Deshayes, 1835 +) + +, +n. comb. +( +Fig. 2D, E +) + + + + + + + +Murex bicostatus +Deshayes,1835: 602 + + +, 603, no. 17, pl. 81, figs 28-30. + + + + + +Pterynotus bicostatus + +– + +Vokes 1971: 23 + +. + + + + +? + +Chicoreus bicostatus + +– + + +Merle +et al. +2011: 107 + + +, text-fig. 41a, b. + + + + + + +TYPE +MATERIAL +. — + +Holotype +(by monotypy). +France + +• +Paris Basin +, +Val d’Oise +, +Valmondois +; +Sables d’Auvers +(biozone NP16); middle +Eocene +( +lower Bartonian +); +UCBL-EM 33437 +( +Deshayes +coll.); H: +21.5 mm +(apex incomplete) ( +Fig. 2D, E +). + + + + +TYPE + +HORIZON +. — Sables d’Auvers (biozone NP16), see +Gély & Lorenz (1991 +: pl. 1), middle Eocene (lower Bartonian). + + + +TYPE + +LOCALITY +. — +France +, Paris Basin, Val d’Oise, Valmondois (see +Fritel 1910: 315 +). + + + + +DISTRIBUTION +. — Only known from the +type +locality. + + + + +DESCRIPTION + + +Teleoconch of +21.5 mm +in height (apex incomplete), of +13.2 mm +in width, biconic in profile, composed of 3.5 preserved whorls. Moderately high spire with convex whorls. Last whorl up to 79% of total length of teleoconch. Apical angle 63°. Spiral sculpture with well-marked primary cords. Spire with P1 and P2 cords. Last whorl: IP, P1 to P6 (P6 placed on top of siphonal canal), ADP, MP and ABP; secondary cords (adis, abis, s3-s6, ads and ms). Axial sculpture not known on early teleoconch whorls. On spire and last whorl, three varices and between them two intervarices.Intervarices strongly developed. Intersection between axial and spiral sculpture displaying small nodules on IP, P1 to P5, ADP and MP. Intervarical nodules, present and more developed on P1 and P2. Ovate aperture of 24% of diameter, up to 64% of length of last whorl. Parietal lip smooth, adherent. Columellar lip smooth, slightly erect. Anal notch deep. Parietal callus well developed. Outer lip showing internal denticles: ID stronger than D1, D2 and D3, D4, D5 and D6 stronger than D1 to D3, D6 split. Siphonal canal of 50% of apertural length. Pseudoumbilicus narrow. + + +COMPARISONS + + +No new specimens of this species have been found since its original description byDeshayes (1835). +Cossmann & Pissarro (1911 +: pl. 36, fig. 170bis-1) attributed an eroded specimen (MNHN.F.J02588 refigured inMerle +et al. +2022:pl. 147, fig.4a, b) to this species under the name + +Ocenebra +( +Ocinebrina +) +bicostata +( +Deshayes,1835 +) + +.However,re-examination of this specimen ( + +Merle +1999 + +) reveals that it represents + +Eofavartia subrudis +(d’Orbigny, 1850) + +. + +Paleochicoreus bicostatus +( +Deshayes,1835 +) + +, +n. comb. +shares with + +P. newtoni +( +Eames, 1957 +) + +and + +P. initialis +( +Vokes, 1990 +) + +, +n. comb. +an aperture with developed internal denticles, a strong parietal callus, and a deep anal notch. + +P.newtoni + +differs in having a slenderer shape and lower, less coarse intervarices. In + +P. initialis + +n. comb. +, the intervarices are also lower than in + +P. bicostatus +( +Deshayes, 1835 +) + +, +n. comb. + +Paleochicoreus packardi +( +Dickerson, 1915 +) + +is poorly preserved (see + +Merle +et al. +2011 + +), but it bears slightly winged varices as do + +P. newtoni + +and + +P. initialis + +n. comb. + +Paleochicoreus diderrichi +( +Vincent,1913 +) + +displays well developed denticulation within the outer lip(see + +Merle +et al. +2011 + +: fig.41.3). + +Paleochicoreus bicostatus +( +Deshayes, 1835 +) + +, +n. comb. +differs from these species in lacking winged varices. However, the +holotype +is eroded and in the early Bartonian (Auversian) from the Paris Basin, many species tend to lose their delicate ornaments (wings, spines) and thicken their shell, as is the case in another muricid, + +Eofavartia subrudis +( +Deshayes, 1835 +) + +. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFF1E41FFE8EFA21FEE9FBDC.xml b/data/03/8E/87/038E87DAFFF1E41FFE8EFA21FEE9FBDC.xml new file mode 100644 index 00000000000..eb4299b3479 --- /dev/null +++ b/data/03/8E/87/038E87DAFFF1E41FFE8EFA21FEE9FBDC.xml @@ -0,0 +1,569 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + + +Timbellus magnei + +n. name + + + + + +( +Figs 12 +A-F; 26G-I) + + + + + + + +Murex trigonus +Rouault, 1850: 204 + + +, pl. XVII, fig. 17a + +[ +non + +Murex trigonus +Gmelin, 1791 + +]. + + + + +Pterynotus trigonus + +– + +Glibert 1963: 11 + + +[ +non + +Murex trigonus +Gmelin, 1791 + +]. + + + + +Pterynotus tricarinatus trigonus + +– + + +Merle 1990: 158 + +, pl. 2, figs 13, 14 only [ +non +Murex trigonus +Gmelin, 1791 +]. + + + + +Pterynotus tripteroides + +– + +Merle 1990: 156 + +, pl. 3, fig. 1 only [ +non +Lamarck, 1822 +]. + + + + + +Timbellus tripteroides + +– + + +Merle +et al. +2011: 44 + + +, pl. 100 figs 3, 4 only [ +non +Lamarck, 1822 +]. + + + + + +Timbellus crenulatus palensis + +– + + +Merle +et al. +2011: 440 + + +, pl. 98, fig. 1 only [ +non +Magne, 1941 +]. + + + + + + +TYPE +MATERIAL +. — + +Neotype +. +France + +• +Aquitaine +Basin, Pyrénées-Atlantiques, Bos d’Arros; uppermost part of the + +Marnes +de Gan + +only (top biozone N12/base biozone NP13); Early Eocene (Ypresian); BMNH 69802 ( +Seaman +coll.) designated by + +Merle +(1990: 158 + +, pl. 2, fig. 14) ( +Fig. 12A +). + + + +ADDITIONAL +MATERIAL +. — + + +France + +• 1 spm +Aquitaine +Basin, Pyrénées-Atlantiques, Gan (Tuilerie); uppermost part of the Marnes de Gan only (top biozone N12/base biozone NP13); Early Eocene (Ypresian, uppermost part of the Marnes de Gan); +MNHN +.F.A32506 ( +Merle +coll.), figured specimen of + +Merle +et al. +(2011 + +: pl. 100, fig. 3) + +• + +1 spm; idem; +MNHN +.F.A32508 ( +Merle +coll.), figured specimen of + +Merle +et al. +(2011 + +: pl. 100, fig. 1) ( +Figs 12C, D +; +26G +) + +• + +1 spm; idem; +MNHN +.F.R61437, +Merle +coll., figured specimen of + +Merle +(1990 + +: pl. 3, fig. 1) + +• + +1 spm; idem; +MNHN +.F.A24115 ( +Merle +coll.) + +• + +2 spm; idem; +MNHN +.F.A72443 (Varone coll.) ( +Figs 12E, F +; +26I +) + +• + +35 spm; idem; +MNHN +.F.A24114 ( +Merle +coll.) + +• + +1 spm (juvenile with protoconch); idem; +MNHN +.F.A90545 ( +Merle +coll.) ( +Figs 12B +; +26H +). + + + + +TYPE + +HORIZON +. — Uppermost part of the Marnes de Gan only (top biozone N12/base biozone NP13), see + +Nolf +et al. +(2002: 173 + +, fig. 4), Early Eocene (Ypresian). + + + +TYPE + +LOCALITY +. — +France +, +Aquitaine +Basin, Pyrénées-Atlantiques, Bos d’Arros ( + +Nolf +et al. +2002: 185 + +). + + + + +DISTRIBUTION +. — Known from the +type +locality and Gan (tuilerie), +Aquitaine +Basin ( +France +), early Eocene (Ypresian). + + + + +DESCRIPTION + + +Smooth, multispiral protoconch of 3.5 whorls ( +Figs 12B +; +26H +). Sinusigeral scar well marked.Teleoconch up to +46 mm +in height, up to +39 mm +in width, subfusiform in profile, composed of eight whorls. Moderately high spire. Last whorl up to 70% of total length. Apical angle 53° excluding wings, 70° including wings. Spiral sculpture with marked primary cords on varices and intevarices. First whorl: no cord. Second whorl: appearance of weak P1, P2 and s1. Third whorl: P1, P2, s1 well marked; appearance of s2 weak. Fourth whorl: s2 well marked. From fifth to seventh whorl: no change or appearance of threads. Eighth whorl: P1 to P6, s1, s2, s3 (well developed), P6; ADP, MP weak or missing on siphonal canal. First whorl:around ten protovarices.Second whorl: appearance of varices, three varices and two intervarices between. From third to last whorl, three to five low intervarices. Higher relief of intervarices on P2. Varices lamellose and strongly winged. Appearance of short P1 spine on second whorl.P1 spine short, almost included in wing. No other cord spines. Ovate aperture up to 22% of diameter and up to 71% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, erect anteriorly, forming curved inductura at base. Parietal lip slightly adherent. Outer lip with well developed internal denticles including ID, D1 to D6. D1 and D2 bifid. Outer lip not crenulate. Siphonal canal narrow, open, straight, up to 57% of apertural length. Pseudoumbilicus narrow. + + +COMPARISONS + + +Rouault (1850) +described from Bos d’Arros (a locality very close to Gan), + +Murex trigonus + +based on juvenile specimens. Unfortunately, + +Murex trigonus +Rouault, 1850 + +is a junior homonym of + +Murex trigonus +Gmelin, 1791 + +and +Magne (1941) +, a homonymy as yet uncorrected. Therefore, we propose the new name + +Timbellus magnei + +n. name herein. The juvenile specimens illustrated by +Rouault (1850) +are characterized by low intervarices varying from three to five (compare +Fig. 12C, D and G +). This character is present on the early whorls of the adults of + +T. tripteroides sensu +Merle (1990 + +: pl. 2, figs 13, 14 only). Thus, one can accept that these large + +T. tripteroides sensu +Merle (1990) + +correspond to adult specimens of + +T. magnei + +n. name. + +Timbellus magnei + +n. name is easily separated from typical + +T. tripteroides + +by having a multispiral protoconch ( +Fig. 12B +), whilst typical + +tripteroides + +has a paucipiral protoconch (see +Fig. 8D, E +) and by having more and lower intervarices. + + + +FIG +. 12. — Spiral cords and internal denticles of the outer lip in + +Timbellus magnei + +n. name ( +A -F +) and + +T. palensis +( +Magne, 1941 +) + +( +G -I +): +A +, neotype, BMNH 69802 (Seaman coll.), Bos d’Arros,Pyrénées-Atlantiques,Aquitaine Basin, France,early Eocene (Ypresian); +B +, MNHN.F.A90545 ( +Merle +coll.), juvenile specimen and protoconch,Gan (Tuilerie),Pyrénées-Atlantiques,early Eocene (Ypresian); +C +, +D +, MNHN.F.A32508 ( +Merle +coll.), juvenile specimen,Gan (Tuilerie); +E +, +F +, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); +G +, MNHN.F.A90553 ( +Merle +coll.), juvenile specimen, Gan (Tuilerie); +H +, +I +, MNHN.F.A32509 ( +Merle +coll.), Gan (Tuilerie). Scale bars: A, G, 5 mm; B, 2 mm; C-F, H, I, 10 mm. Credits: A, D. Serrette (MNHN); B, L. Cazes; E-I, P. Loubry (MNHN/CNRS). + + + +Cossmann & O’Gorman (1923) +misinterpreted + +Murex trigonus sensu +Rouault + +and illustrated a specimen typical of the genus + +Ponderia + +: + +P. daguini +( +Magne, 1941 +) + +. In addition, +Cossmann & O’Gorman (1923) +described and figured an adult specimen of + +Timbellus + +that Cosmann named + +Murex +cf. +tricarinatus +Lamarck, 1803 + +. +Magne (1941) +gave the name + +Murex +( +Pteropurpura +) +palensis + +to + +Murex +cf. +tricarinatus +Lamarck, 1803 + +( +sensu +Cossmann 1923 +) and demonstrated that the juvenile specimens illustrated by +Rouault (1850) +as + +Murex trigonus + +were not juveniles of + +Murex +( +Pteropurpura +) +palensis + +. We discovered a young specimen from Gan differing from the juveniles of + +T. magnei + +n. name. It displays one high intervarix between each varix ( +Fig. 12G +), a character also present on the early teleoconch whorls of well preserved adults of + +T. palensis + +( +Fig. 12H, I +). +Douvillé & O’Gorman (1929 +: pl. 29, fig. 1) figured a specimen from the early Eocene of Biron (near Orthez) and called it + +Murex tripteroides + +. That specimen bears one high intervarix between each varix and possibly represents + +T. palensis + +. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFFAE41BFC7AFC40FC86F858.xml b/data/03/8E/87/038E87DAFFFAE41BFC7AFC40FC86F858.xml new file mode 100644 index 00000000000..33bf9bb1e6c --- /dev/null +++ b/data/03/8E/87/038E87DAFFFAE41BFC7AFC40FC86F858.xml @@ -0,0 +1,425 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + + +Timbellus submicropterus + +n. sp. + + + + + +( +Figs 16 +A-H, K; 27A-D) + + + +urn:lsid:zoobank.org:act: +77B79E49-DDB1-413B-983A-A19753441F55 + + + + + + + +Murex +( +Pterymurex +) +tricarinatus + +– + +Cossmann & Pissarro 1901: 128 + +, pl. 14, fig. 1 [ +non +Lamarck, 1803 +]. + + + + + +Timbellus micropterus + +– + + +Merle +et al. +2011: 448 + + +, pl. 102, fig. 4a, b only [ +non +Deshayes, 1865 +]. + + + + + + +TYPE +MATERIAL +. — + +Holotype +. +France + +• +Normandy +, Manche, Fresville; + +Calcaire +de Fresville + +; middle Eocene (uppermost Lutetian); +MNHN +.F.A24111 ( +MNHN +coll.), H: +23.6 mm +( +Figs 16A, B +; +27A +). + + + + + +Paratypes +. +France + +• 1 spm; same as for the holotype; +UCBL-EM 33438 +( +Bourdot +coll., figured specimen of +Cossmann & Pissarro 1901 +)( +Figs 16E, F +; +27C +) + + +• 1 spm; idem; +MNHN +.F.A90556 ( +MNHN +coll.) ( +Fig. 16K +) + +• + +1 spm; idem; +MNHN +.F.B63700 ( +de Morgan +coll.) + +• + +1 spm; +Normandy +, +Manche +, +Gourbesville +; +Calcaire de Fresville +; +upper Lutetian +; +MNHN +.F.A24126 ( +Munier-Chalmas +coll.), figured specimen of + +Merle +et al. +(2011: 135 + +, pl. 102, fig. 4a, b) ( +Figs 16C, D +; +27B + +). + + + + +ADDITIONAL +MATERIAL +(material referring to + +Timbellus +cf. +submicropterus + +n. sp. +). — + + +France + +• 3 spm; Loire-Atlantique, +Loire +Basin, Saffré (Bois Gouët); middle Eocene (lower Bartonian); MNHN.F.A29660 (Pacaud coll.) + +• + +2 spm; Saint-Aignan-GrandLieu (Pierre-Aiguë), Loire-Atlantique; idem; MNHN.F.A29662, (Brébion coll.),MNHN.F.A90557 (Brébion coll. ( +Figs 16G, H +; +27D +). + + + + + +ETYMOLOGY +. — Name formed by the prefix +sub- +and + +micropterus + +for this species older than Bartonian + +Timbellus micropterus +( +Deshayes, 1835 +) + +. + + + + +TYPE +HORIZON +. — Calcaire de Fresville, middle Eocene (uppermost Lutetian), see + +Bignot +et al. +(1968) + +. + + +TYPE +LOCALITY +. — +France +, +Normandy +, Manche, Fresville, see +APGN (2024) +. + + + + +DISTRIBUTION +. — +Normandy +(Calcaire de Fresville) middle Eocene (uppermost Lutetian) and? +Loire +Basin (Sables du Bois-Gouët), lowermost Bartonian, see +Mathelin & Bignot (1989) +. + + + + +DESCRIPTION + + +Bulbous protoconch of one whorl ( +Fig. 16K +). Teleoconch up to +23.6 mm +in height, up to +11.6 mm +in width, biconic in profile, composed of six whorls. Moderately high spire. Last whorl up to 70% of total length. Apical angle 59°. Spiral sculpture with coarse primary cords. First to third whorl: no cord. Fourth whorl: appearance of weak P1, s1 and P2. Fifth whorl: P1, s2 and P2 coarser. Sixth whorl: P1 to P6, s1; P6 not atrophied on convex part of whorl; ADP, MP and ABP on siphonal canal. From first whorl to second whorl: 6-7 protovarices. On third whorl: appearance of varices; three varices and one intervarix intercalated. From third to sixth whorl: no change. Higher relief of intervarices on P2. Varices rather thick. Appearance of short P1 spine on third whorl. Ovate aperture up to 27% of diameter and up to 71% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, poorly erect anteriorly, forming curved inductura at base. Parietal lip slightly adherent. Outer lip with developed denticles including ID, D1 to D6. D +6 in +some specimens. Outer lip not crenulate. Siphonal canal open, straight, up to 40% of apertural length. Pseudoumbilicus narrow. + + +COMPARISONS + + +A specimen of this species ( +Figs 16E, F +; +27C +) was identified as + +Murex +( +Pterymurex +) +tricarinatus +Lamarck, 1803 + +(synonym of + +Timbellus crenulatus tricarinatus + +) by +Cossmann & Pissarro (1901) +. + +Timbellus crenulatus tricarinatus + +differs greatly by apertural (with weak internal denticles within the outer lip), sculptural and shape characters, and belongs to another group species (see above +Fig. 14 +). In fact, this specimen illustrated by +Cossmann & Pissarro (1901) +more closely resembles + +T. micropterus + +from the Bartonian Paris Basin. + +Merle +et al. +(2011) + +attributed another Norman specimen (pl. 102, fig. 4a, b) to + +T. micropterus + +( +Figs 16C, D +; +27B +). It is true that these +two specimens +are superficially similar to + +T. micropterus + +, but further comparison reveals some differences, such as only one intervarice (instead two in typical + +T. micropterus + +) and finer secondary cords on the last whorl. The bivariate plot of total length against width ( +Fig. 17 +) does not suggest strong differences +Normandy +and Paris Basin populations. Finally, it is tempting to conclude that the observed teleoconch differences in + +T. submicropterus + +n. sp. +fall within the intraspecific variation of + +T. micropterus + +. However, + +T. submicropterus + +n. sp. +has a paucispiral protoconch, whereas + +T. micropterus + +has a multispiral ( +Fig. 16K, L +), suggesting two separate species with different modes of larval development. In the Bartonian from Loire-Atlantique, there are specimens showing coarse spiral cords like those of typical + +T. submicropterus + +n. sp. +, but they bear a more pronounced shoulder ( +Fig. 16G, H +). Their protoconch is not preserved. We therefore attribute these specimens to + +T. submicropterus + +n. sp. +with hesitation. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFFBE41BFC1EFBC2FAA5F89F.xml b/data/03/8E/87/038E87DAFFFBE41BFC1EFBC2FAA5F89F.xml new file mode 100644 index 00000000000..02bea689973 --- /dev/null +++ b/data/03/8E/87/038E87DAFFFBE41BFC1EFBC2FAA5F89F.xml @@ -0,0 +1,168 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + +Subgenus + +Pterymarchia +Houart, 1995 + + + + + + + +TYPE + +SPECIES +. — + +Murex tripterus +Born, 1778 + +, by original designation. Recent: Indo-Pacific. + + + + +COMMENTS +ON +THE +SUBGENUS + + +According toHouart (1995), + +Pterymarchia + +can be distinguished from + +Pterynotus + +( +s.s. +) by having more numerous axial varices on the early teleoconch whorls (9-11 vs 6-8), later appearance in ontogeny of the trivaricate sculpture, when present, sometimes more varices on the last whorls (3-5 vs 3), and more strongly developed denticles. +Houart (1995 +, +2018 +) proposed full generic status for + +Pterymarchia + +. However, we prefer to maintain it at subgeneric rank because: 1) based on molecular data, + +Pterynotus + +and + +Pterymarchia + +are very close ( + +Barco +et al. +2010 + +); and 2) based on shell characters, + +Pterymarchia + +shares with + +Pterynotus + +a scabrous surface appearing early in ontogeny, generally fine to moderately fine primary cords (from IP to ABP), and a series of internal denticles from ID to D6. Furthermore, several other characters are occasionally shared by both genera, such as the trivaricate sculpture, columellar tubercles [e.g., +P. +( +s.s. +) +patagiatus +(Hedley, 1912), +P. +( +s.s. +) +praepatagiatus +Vokes, 1992 +and +P. +( +s.s. +) +granuliferus +(Grateloup, 1833)], and even developed denticles in some +P. +( +s.s. +) +albobruneus +Bertsch & D’Attilio, 1980. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFFBE427FC33F8A4FE65FAFC.xml b/data/03/8E/87/038E87DAFFFBE427FC33F8A4FE65FAFC.xml new file mode 100644 index 00000000000..97c0701e9fa --- /dev/null +++ b/data/03/8E/87/038E87DAFFFBE427FC33F8A4FE65FAFC.xml @@ -0,0 +1,649 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + +Pterynotus +( +Pterymarchia +) +gaasensis + +n. sp. + + + + +( +Figs 18 +A-D; 27E, F) + + + +urn:lsid:zoobank.org:act: +B1E15CF2-7B4F-4D02-8F34-1D5848D596F5 + + + + + +FIG +. 16. — Spiral cords and internal denticles of the outer lip in + +Timbellus submicropterus + +n. sp. +( +A -H +, +K +) and + +T. micropterus +( +Deshayes, 1835 +) + +( +I +, +J +, +L +): +A +, +B +, holotype, MNHN.F.A24111 (MNHN coll.), Fresville, Manche, Normandy, France, middle Eocene (Lutetian); +C +, +D +, paratype, MNHN.F.A24126 (Munier-Chalmas coll.), Gourbesville, Manche, middle Eocene (Lutetian); +E +, +F +, paratype, UCBL-EM 33438 (Bourdot coll.), Fresville; +G +, +H +, MNHN.F.A90557 (Brébion coll.), Saint-AignanGrand-Lieu (Pierre-Aiguë), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); +I +, +J +, MNHN.F.A32522 (Faullummel coll.), Le Limon, Seine-et-Marne, Paris Basin, France,middle Eocene (Bartonian); +K +, paratype (protoconch), MNHN.F.A90556 (MNHN coll.), Fresville; +L +, MNHN.F.A90558 (Faullummel coll., protoconch), Chavençon (Carrière Tuyttens), Oise, Paris Basin, France, middle Eocene (Bartonian). Scale bars: A-J, 10 mm; K, L, 0.5 mm. Credits: A-D, I, J, P. Loubry (MNHN/CNRS); E, F, E. Robert (UCBL); G, H, K, L, L. Cazes (MNHN/CNRS). + + + + +FIG +. 17. — Bivariate plot of total length against width comparing + +T. micropterus +( +Deshayes, 1835 +) + +from the lower Bartonian of the Paris Basin (50 complete specimens) and + +T. submicropterus + +n. sp. +from the uppper Lutetian of Normandy (five complete specimens). + + + + + +Murex quadrifrons + +– +Grateloup 1845 +: pl. 3 [pl. 31], fig. 10 [ +non + +Murex quadrifrons +Lamarck, 1822 + +]. + + + +Pterynotus +( +Pterymarchia +) cf. +subquadrifrons + +– + +Merle +et al. +2011 + +( +partim +): pl. 90, figs 3-74 only. + + + + +Pterynotus +cf. +subquadrifrons + +– + +Lozouet 2023: 21 + +, pl. 17, figs 1-9, text-fig. 10E. + + + + + +TYPE +MATERIAL +. — + + +Holotype +. +France + +• +Aquitaine +Basin, Landes, Gaas (Espibos); Gaas Formation (biozone NP23); early Oligocene (Rupelian); +MNHN +.F.A32492 ( +Merle +coll.), figured specimen in + +Merle +et al. +(2011 + +: pl. 90, fig. 3), H: +27 mm +( +Figs 18A, B +; +27E +). + + + +Paratypes +. +France + +• 6 spm; same as for the holotype; +MNHN +.F.A77749 ( +Merle +coll.) • + + +1 spm; idem; +MNHN +.F.A90554 ( +Merle +coll.) ( +Figs 18C, D +; +27F +) • + + +1 spm; idem; +MNHN +.F.A86913 (Lozouet coll.), figured in +Lozouet (2023 +: text-fig. 10a, b) + +. + + + +ADDITIONAL +MATERIAL + +. + +— + +France + +• 1 spm; +Aquitaine +Basin +, Landes, +Gaas +( +La Gouarde +); same as for the +holotype +; Aucoin coll., figured +in + +Merle +et al. +(2011 + +: pl. 90, fig. 4a, b). + + + + + +ETYMOLOGY +. — From the +type +locality, Gaas (Landes), +France +. + + + + + +TYPE + +HORIZON +. — Gaas Formation (biozone NP23), see +Sztrákos & Steurbaut (2017: 752) +, early Oligocene (Rupelian). + + + +TYPE + +LOCALITY +. — +France +, +Aquitaine +Basin, Landes, Gaas (Espibos), see +Sztrákos & Steurbaut (2017: 752) +. + + + + +DISTRIBUTION +. — +Aquitaine +Basin ( +France +), early Oligocene (Rupelian). + + + + +DESCRIPTION + + +Paucispiral protoconch ( +fide +Lozouet 2023 +: text-fig. 10e). Teleoconch up to +27 mm +in height, up to +12 mm +in width, biconic in profile, composed of six whorls. Moderately high spire, first two whorls subcarinate; next whorls rounded. Last whorl up to 75% of total length. Apical angle 56°. Spiral sculpture with marked primary cords. First whorl: appearance of primary cords P1 and P2; second whorl: P1 and P2; third whorl: appearance of s1. Fourth whorl: appearance of abis, adis and s2. Fifth whorl: no change or appearance of threads. Sixth whorl: IP, adis and abis, P1 to P6, s1 to s6; s2 and s3 well developed; P6 on base on convex part of whorl; ADP, MP and ABP slightly more developed than ads and ms. Surface of shell with numerous threads appearing on fourth whorl. Axial sculpture with thick varices. On first whorl: nine lamellose varices. From the second to fifth whorl: 9-10 thick varices. On sixth: appearance of three or four varices and one intervarices intercalated. Varices and intervarices thick. From the third to last whorl, numerous, fine microlamellae covering entire surface forming scabrous microsculpture. Ovate aperture up to 25% of diameter and up to 66% of length of last whorl (including siphonal canal). Columellar lip, adherent posteriorly, becoming progressively more erect anteriorly. One or two small columellar tubercles. Parietal lip adherent. Outer lip bearing strong internal denticles: ID, D2 to D6 present, D1 missing. Pseudoumbilicus moderately large. Siphonal canal open, up to 48% of apertural length, slightly dorsally recurved. + + +COMPARISONS + + + +Merle +et al. +(2011 + +: pl. 90, figs 3-4) attributed with hesitation +two specimens +from Gaas to + +Murex subquadrifrons +d’Orbigny, 1852 + +, originally described as + +Murex quadrifrons + +by +Grateloup (1845) +from this same Rupelian locality. The Grateloup’s collection is housed in the University of Talence, but the +type +material of + +M. quadrifrons + +was not found and is likely lost (personal observation of the first author). Therefore, only the drawing of +Grateloup (1845 +: pl. 3 [pl. 31], fig. 10, refigured in + +Merle +et al. +2011 + +: pl. 90, fig. 2) can be used to identify the species. According to + +Merle +et al. +(2011) + +, the drawing shows a shell with a textiliose surface and a biconic shape, as in the Gaas material at hand. Moreover, to our knowledge, no other muricid from Gaas except +P. +( + +Pterymarchia + +) is comparable to the specimen illustrated by +Grateloup (1845) +. However, the shell illustrated by +Grateloup (1845) +lacks internal denticles within the outer lip, lacks columellar denticles, shows more numerous varices (six on the ventral face vs four in the specimens at hand), has a narrower siphonal canal, and is more likely to represent a specimen of + +Favartia + +. These numerous differences between +Grateloup’s (1845) +figure and the studied material suggest that it obviously corresponds to a species other than + +M. subquadrifrons + +. + +Murex subquadrifrons + +is therefore not recognizable using the figure of Grateloup, lacks +type +material and it should be considered a +nomen dubium +. + + + +FIG +. 18. — Spiral cords and internal denticles of the outer lip in + +Pterynotus +( +Pterymarchia +) +gaasensis + +n. sp. +( +A -D +), + +P. +( +P. +) +pelouatensis + +n. sp. +( +E -H +) and +P. +( +P +) + +sandbergeri +(HÖrnes,1856) + +( +I -L +): +A +, +B +, holotype,MNHN.F.A32492 ( +Merle +coll.), Gaas (Espibos),Landes,Aquitaine Basin,France,early Oligocene (Rupelian); +C +, +D +, paratype, MNHN.F.A90554 ( +Merle +coll.), Gaas (Espibos); +E +, +F +, holotype, MNHN.F.J09945, Saucats (Peloua), Gironde, Aquitaine Basin, Early Miocene (Burdigalian); +G +, +H +, paratype, MNHN.F.J17632, Saucats (Peloua); +I +, +J +, lectotype designated herein,NHMW1855/0045/0435,Gainfarn,Baden, Austria,Middle Miocene (Langhian); +K +, +L +, Landau coll., Bad VÖslau, Baden. Scale bars: A-H, 5 mm; I-L, 10 mm. Credits:A-H, P. Loubry (MNHN/CNRS); I, J, M. Harzhauser (NHMW); K, L, B. Landau. + + + +Eocene + +Pterynotus +( +Pterymarchia +) + +, such as + +P. +( +P. +) +antiquus +( +Merle 1990 +) + +, + +P. +( +P. +) +denudatus +( +Deshayes, 1835 +) + +and + +P. +( +P. +) +defensus +(Fuchs, 1870) + +strongly differ from the + +P. +( +P. +) +gaasensis + +n. sp. +in having trivaricate sculpture. + +Pterynotus +( +P. +) +fascistria +( +von Koenen, 1889 +) + +from the Priabonian of North +Germany +is more similar shape, as it lacks trivaricate sculpture and because their primary cords are poorly distinguishable from the secondary cords on the last whorls. + +Pterynotus +( +P. +) +fascistria + +is distinguished from + +P. +( +P. +) +gaasensis + +n. sp. +in having a lower spire and by lacking intervarices, D1 and ID denticles. + +Pterynotus +( +P. +) +gaasensis + +n. sp. +can be compared to two other Neogene species: + +P. +( +P. +) +pelouatensis + +n. sp. +from the Burdigalian of Aquitaine Basin ( +Figs 18 +EH; 27G, H) and + +P. +( +P. +) +sandbergeri +(Hörnes, 1856) + +from the Langhian of the Paratethys. Both species differs from + +P. +( +P. +) +gaasensis + +n. sp. +in having more angular whorls with a shoulder delimited by a subspiny P1 and by other primary cords more distinguishable from the secondary cords on the last whorl. +Lozouet (2023) +followed + +Merle +et al. +(2011) + +and adopted the name + +Pterynotus +cf. +subquadrifrons + +for the Rupelian material from Gaas, but he did not include Chattian and Aquitanian specimens figured in + +Merle +et al. +(2011 + +: pl. 90, figs 5, 6), because Rupelian shells have a paucispiral protoconch, whereas the Chattian/Aquitanian species has a multispiral protoconch, and is therefore a separate species. Thus, we consider for instance that + +P. +( +P. +) +gaasensis + +n. sp. +to be present only in the Rupelian. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFFCE41DFC63FA22FB82FAA3.xml b/data/03/8E/87/038E87DAFFFCE41DFC63FA22FB82FAA3.xml new file mode 100644 index 00000000000..4f5650221a7 --- /dev/null +++ b/data/03/8E/87/038E87DAFFFCE41DFC63FA22FB82FAA3.xml @@ -0,0 +1,497 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + + +Timbellus curvispina + +n. sp. + + + + + +( +Figs 14 +D-H; 26J-L) + + + +urn:lsid:zoobank.org:act: +00EFA23B-DFE2-4272-AC88-BF1C6253EBA7 + + + + + + + +Murex tricarinatus + +– + +Deshayes 1835: 597 + +, pl. 82, figs 9, 10 [ +non +Lamarck, 1803 +]. + + + + + +Timbellus crenulatus tricarinatus + +– + + +Merle +et al. +2011: 440 + + +, pl. 98, figs 6, 7 only [ +non +Lamarck, 1803 +]. + + + + + + +TYPE +MATERIAL +. — + +Holotype +. +France + +• Paris Basin, Oise, Parnes (Les Bôves); + +Falun +de Parnes + +(biozone NP15); middle Eocene (middle Lutetian); +MNHN +.F.A90541 (ex B63594), H: +51.8 mm +( +Figs 14E, F +; +26J +). + + + + + +Paratypes +. +France + +• 3 spm +Paris Basin +; +Eure +; +Fontenay-en-Vexin +(Bois du but); +Falun de Parnes +(biozone NP15); middle +Eocene +( +Lutetian +); +MNHN +.F.A73848 ( +Figs 14D +; +26L +), +MNHN +.F.A90542, +MNHN +.F.A90543 ( +Pons +coll.) ( +Figs 14G, H +, +26K +) + +• + +1 spm; idem; +MNHN +.F.A32514 ( +Pacaud +coll.), figured +in + +Merle +et al. +(2011: 440 + +, pl. 98, fig. 7) + +• + +1 spm; +Paris Basin +, Yvelines, Villiers-Saint-Frédéric ( +Butte Saint-Léonard +); +Falun de Parnes +(biozone NP15); +Lutetian +; +MNHN +.F.A32516 ( +Schtrock +coll.), figured +in + +Merle +et al. +(2011: 440 + +pl. 98, fig. 6) + +• + +1 spm; +Paris Basin +, +Yvelines +, +Parnes +; +Falun de Parnes +(biozone NP15); +Lutetian +; +MNHN +.F.A90544 (ex +Galerie de Zoologie +coll.) + +. + + + + +ETYMOLOGY +. — Name derived from Latin +curv +- (meaning curved) and +spina +(meaning spine) and refers to the P1 spine dorsally curved; gender female. + + + + +TYPE +HORIZON +. — Falun de Parnes (biozone NP15), see +Gély & Lorenz (1991 +: pl. 1), middle Eocene (middle Lutetian). + + +TYPE +LOCALITY +. — +France +, Paris Basin, Oise, Parnes (Les Bôves), see +Fritel (1910: 221) +. + + + + +DISTRIBUTION +. — Calcaire grossier moyen, biozone NP15, Paris Basin ( +France +), middle Eocene (middle Lutetian). + + + + +DESCRIPTION + + +Protoconch not preserved. Teleoconch up to +51.8 mm +in height, up to +30 mm +in width, subfusiform in profile, composed of six whorls. Moderately high spire. Last whorl up to 85% of total length. Apical angle 63°. Spiral sculpture with marked primary cords on varices, but weaker on intervarices. First and second whorls eroded. From third: appearance of weak P1 and P2. Third word: no change. Fourth whorl: appearance of s1. Fifth whorl: occasional appearance of s2. Sixth whorl: P1 to P6, s1, s2, P6 not atrophied, s1 well developed; ADP developed, MP, ABP weak or missing on siphonal canal. Axial sculpture eroded on two early whorls. On second whorl, eight protovarices. Third whorl: appearance of varices; three varices and one intervarix between. From fourth to last whorl, no change. Higher relief of intervarices on P2. Varices sublamellose. Appearance of short straight P1 spine on fourth whorl. On last whorl, axial ornamentation forming very short spinelets, dorsally recurved on P1 to P6. P1 spine more strongly developed than other cord spines. Rounded aperture up to 34% of diameter and up to 74% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, slightly erect anteriorly, forming curved inductura at base. Parietal lip slightly adherent. Outer lip with very weak denticles including D1 to D6. Small crenulations at edge of outer lip. Siphonal canal narrow, open, bent, slightly dorsally recurved, up to 81% of apertural length. Pseudoumbilicus narrow + + +COMPARISONS + + +This species can be included in the group of + +T. crenulatus + +defined by +Harasewych & Jensen (1979) +, +Vokes (1992) +, and updated by + +Merle +et al. +(2011: 15) + +. In this group the last whorl is generally wider than in species of the + +T. tripteroides + +group, the shoulder spine (P1) is more delineated and the internal denticles within the outer lip are weaker. The spiral sculpture corresponds to fine primary cords, as in the + +T.tripteroides + +group. + +Timbellus curvispina + +n. sp. +was mistaken for the Lutetian species + +T. crenulatus tricarinatus +( +Lamarck, 1803 +) + +but it differs by the construction of the P1 spine. This spine is straight, long and oriented abaperturally in + +T. crenulatus tricarinatus + +, whereas it is shorter, curved and dorsally oriented in + +T. curvispina + +n. sp. +(see +Fig. 14 +). + +Timbellus curvispina + +n. sp. +also bears cord spines that are always more developed than in + +T. crenulatus tricarinatus + +. Moreover, the spiral cords of + +T. curvispina + +n. sp. +are weak in the intervarical spaces, whereas those of + +T. crenulatus tricarinatus + +are well marked. + +Pacaud +et al. +(2017) + +recently designated the +lectotype +of + +Murex tricarinoides +Deshayes, 1835 + +demonstrating that authors confused this species with another: + +T. capitaneus +Pacaud, Ledon & Goret, 2017 + +. In addition, they synonymised + +T. tricarinoides + +with + +T. crenulatus tricarinatus + +. However, + +T. tricarinoides + +differs from + +T. crenulatus tricarinatus + +in lacking true P1 spines and in displaying a rounded outer lip. + +T. crenulatus crenulatus +(Röding, 1798) + +from the Bartonian is easily distinguishable from + +T. curvispina + +n. sp. +by its acute and long P1 spine pointing abaperturally. + + +SPECIES +GROUP +OF +T. + + +PHYLLOPTERUS + +( + +LAMARCK +, 1822 + +) + +Members of this group are usually large sized and share a strong development of the internal denticles within the outer lip and strong primary cords; secondary cords are poorly developed. All species lack columellar denticles, except in the +holotype +of + +T +. +xenos +(Harasewych, 1982) + +, which is presumably a pathological specimen ( +Vokes 1992: 18 +). In the middle Eocene from Europe, + +Timbellus submicropterus + +n. sp. +(upper Lutetian) and the Bartonian + +T. micropterus +( +Deshayes, 1835 +) + +represent the oldest members of this species group. It becomes more diversified in the late Eocene/early Oligocene with + +T. lamarckii +( +Grateloup, 1845 +) + +, + +T. detritus +(von Koenen, 1890) + +, + +T. arenarius +( +Steuer, 1912 +) + +, + +T. danapris +Pacaud, 2018 + +and + +T. radulfiensis + +n. sp. +It contains three Miocene species: + +T. perlongus +(Bellardi, 1872) + +, + +T. rosavendae +(Bellardi, 1872) + +and + +T. pseudolamarcki +( +Magne, 1941 +) + +, and becomes extinct before the Pliocene. + + + + \ No newline at end of file diff --git a/data/03/8E/87/038E87DAFFFFE41CFEE1FB63FF77F858.xml b/data/03/8E/87/038E87DAFFFFE41CFEE1FB63FF77F858.xml new file mode 100644 index 00000000000..e11a8016770 --- /dev/null +++ b/data/03/8E/87/038E87DAFFFFE41CFEE1FB63FF77F858.xml @@ -0,0 +1,379 @@ + + + +New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe + + + +Author + +Merle, Didier + + + +Author + +Pacaud, Jean-Michel + + + +Author + +Ledon, Daniel +Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr +didier.merle@mnhn.fr + + + +Author + +Goret, Bernard +Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com +bernardgoret@hotmail.com + +text + + +Geodiversitas + + +2024 + +2024-10-24 + + +46 + + +15 + + +495 +551 + + + + +https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf + +journal article +10.5252/geodiversitas2024v46a15 +1638-9395 +13985760 +urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713 + + + + + + +Timbellus sixi + +n. sp. + + + + + +( +Figs 13 +; +26M +) + + + +urn:lsid:zoobank.org:act: +98875389-52BE-40D1-920C-5D8E4BB7B580 + + + + + + +TYPE +MATERIAL +. — + +Holotype +. +France + +• +Paris Basin +, +Oise +, +Trosly-Breuil +( +Fond Couturier +); + +Sables +de Cuise + +, (biozone NP12); early +Eocene +( +Ypresian +, +lower Cuisian +); +MNHN +.F.A90544 ( +Six +coll.), H: +14 mm +( +Figs 13 +; +26M +). + + + + + +ETYMOLOGY +. — Dedicated to Didier Six who collected this rare species. + + + + + +TYPE + +HORIZON +. — Sables de Cuise, (biozone NP12), see +Gély & Lorenz (1991 +: pl. 1), early Eocene (Ypresian, lower Cuisian). + + + +TYPE + +LOCALITY +. — +France +, Paris Basin, Oise, Trosly-Breuil (Fond Couturier). + + + + +DISTRIBUTION +. — Only known from the +type +locality. + + + + +DESCRIPTION + + +Unknow protoconch. Teleoconch up to +14 mm +in height, up to +7.6 mm +in width, biconic in profile, composed of five whorls. Moderately high spire. Last whorl 74% of total length. Apical angle 74° including spines, 66° excluding spines. Spiral sculpture with fine, narrow, well developed primary cords. First whorl, no spiral sculpture. Second whorl, appearance of weak P1 and P2. Third whorl, P1 and P2 well marked, appearance of s1. Fourth whorl, appearance of threads on infrasutural ramp. Fifth whorl: P1 to P6, s1 to s3; primary and secondary cords almost same relief; ADP to ABP on siphonal canal. On first whorl, nine protovarices. On second whorl, appearance of varices with two intervarices intercalated. On third whorl: three varices and three intervarices intercalated. From fourth to fifth whorl, three varices and two or three intervarices intercalated. Intevarices low. Higher relief of the intervarices on P2. Varices sublamellose on early whorls, rather thick on last whorls. Appearance of short P1 spine on second whorl. On last whorl, short P1 spine oriented adapically; no other cord spines. Ovate aperture up to 34% of diameter and up to 68% of length of last whorl (including the siphonal canal). Columellar lip smooth, narrow, slightly erect anteriorly, forming flat curved inductura at base. Parietal lip adherent. Outer lip with well developed denticles including ID, D1 to D3 (D4-D5 probably not formed); all denticles simple. Siphonal canal open, slightly dorsally recurved, up to 43% of apertural length. Pseudoumbilicus narrow. + + +COMPARISONS + + +This shell represents the oldest known + +Timbellus + +from the Paris Basin, as no + +Timbellus + +has been recorded from the Early Cuisian (biozone NP12, Ypresian) despite intensive fieldwork. It is easily distinguishable from the middle Eocene Paris Basin + +Timbellus +species + +: + +T. tripteroides +( +Lamarck, 1822 +) + +, + +T. crenulatus tricarinatus +( +Lamarck, 1803 +) + +, + +T. micropterus +( +Deshayes, 1835 +) + +and + +T. capitaneus +Pacaud, Ledon & Goret, 2017 + +. Juveniles of + +T. tripteroides + +and + +T. capitaneus + +have weak spiral sculpture; + +Timbellus crenulatus tricarinatus + +lacks internal denticles, and juveniles of + +T. micropterus + +have coarser spiral sculpture. + +Timbellus sixi + +n. sp. +resembles juveniles of + +T. magnei + +n. name from Bos d’Arros and Gan ( +Fig. 12C, D +) in sharing a similar shape of its cords (fine, narrow, but well marked) and by having flat intervarices between the varices. However, + +T. sixi + +n. sp. +differs from + +T. magnei + +n. name in having less foliated varices, having less numerous intervarices (usually 2-3 instead +4-5 in + +T. magnei + +n. name), and by having MP and ABP on the siphonal canal. + + +SPECIES +GROUP +OF + +T. + +CRENULATUS + +( +RÖDING +, 1798) + + + +In this species group, the last whorl is generally broader than in the species group of + +T +. +tripteroides + +and the shoulder spine (P1) is more strongly developed. The spiral sculpture corresponds to fine primary cords, as in the + +T +. +tripteroides + +species group.Two species have an acute open shoulder spine, similar to that seen in the genus + +Pterochelus +Jousseaume, 1880 + +: the Eocene species + +T +. +crenulatus +(Röding, 1798) + +and the Miocene species + +T +. +valdintortus +( +Cossmann & Peyrot, 1924 +) + +. In the + +T. crenulatus + +species group the rounded aperture bears poorly developed denticles or lirae within the outer lip, except in the oldest known member ( + +T +. +palensis + +[ +Magne, 1941 +]) from the early Eocene of +Aquitaine +Basin of Gan ( +Fig. 12 +G-I). However, according +Cossmann & O’Gorman (1923) +, the short shape and the rounded aperture of shells of this species resemble more closely + +T. crenulatus tricarinatus +( +Lamarck, 1803 +) + +than + +T. tripteroides + +. Probably, the presence of internal denticles in + +T. palensis + +represents a plesiomorphic character state and its loss in younger species of this group (e.g., + +T. crenulatus tricarinatus + +and + +T. crenulatus crenulatus + +) represents a derived state. For this group, we describe a new species: + +T. curvispina + +n. sp. +from the Lutetian of the Paris Basin. As for + +T. tripteroides + +, the type material of + +Murex tricarinatus +Lamarck, 1803 + +is lost, as it is also missing in the collections of Geneva and Paris. In order to avoid future taxonomic mistakes, we designate a +neotype +herein (MNHN.F.A90547, +Fig. 14B, C +) from the original type locality of Grignon (precisely Falunière of Grignon; Calcaire à Orbitolites complanatus Formation, biozone NP15, see +Gély & Lorenz [1991 +: pl. 1], middle Lutetian). + + + + \ No newline at end of file