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<document id="662B94F955468E5D4E47BA126901A497" ID-CLB-Dataset="28466" ID-DOI="http://dx.doi.org/10.3897/dez.65.27424" ID-GBIF-Dataset="290e099d-cb77-4ad7-975e-32ce4a51926f" ID-Pensoft-Pub="1860-1324-2-177" ID-ZooBank="A047B48DD161424FB8800428DCC5888A" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1860-1324-2-177" ModsDocOrigin="Deutsche Entomologische Zeitschrift 65 (2)" ModsDocTitle="Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M.lophoides (Chaudoir) species complex" checkinTime="1552154842744" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="19EB255A7A94DC7D12BC30E2E85BCFF1" docLanguage="en" docName="DeutEntomolZeit 65(2): 177-224" docOrigin="Deutsche Entomologische Zeitschrift 65 (2)" docSource="http://dx.doi.org/10.3897/dez.65.27424" docTitle="Mecyclothorax moorei Baehr" docType="treatment" docVersion="7" lastPageNumber="177" masterDocId="FFB33A56FFD86772FFAAFFA9FF882245" masterDocTitle="Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M. lophoides (Chaudoir) species complex" masterLastPageNumber="224" masterPageNumber="177" pageNumber="177" updateTime="1701378747773" updateUser="ExternalLinkService">
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<taxonomicName id="CFCF418001C1461241F11FE314C6072F" ID-CoL="8M6ND" authority="Baehr" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax moorei" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="moorei">Mecyclothorax moorei Baehr</taxonomicName>
Figures 2A, 3A, 7
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, 8A, 9A, 10A, 11A
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<paragraph id="3B16E937B871A7B82546B79007A1E1B8" pageId="0" pageNumber="177">
<taxonomicName id="7F0E01C5B7EE7BAA07B7BD3038A69D83" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax moorei" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="moorei">Mecyclothorax moorei</taxonomicName>
Baehr, 2009: 90.
</paragraph>
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<subSubSection id="18E5308F041ECD5C26AF7ABF025CA179" pageId="0" pageNumber="177" type="diagnosis">
<paragraph id="E8ED88784F9DFF8253BB4041610ED517" pageId="0" pageNumber="177">Diagnosis</paragraph>
<paragraph id="D9900996CEA130568EE3287016CD3976" pageId="0" pageNumber="177">
(n = 2). This species is diagnosed by the broadly punctate pronotal disc (Fig. 2A), punctate head with a transverse line of about 5 large punctures between the posterior supraorbital setae, and convex, smooth elytra with only striae 1-4 in evidence, striae 2-4 only a series of small isolated punctures. The prosternum and proepisternum are also broadly punctate, the punctures large and separated from each other by a distances equal to the punctural diameters (
<bibRefCitation id="7E160274C637B6B60857CCF6CF8E226A" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="177" pagination="1 - 63" title="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel." url="https://doi.org/10.3897/dez.65.21000" volume="65" year="2018">Liebherr 2018</bibRefCitation>
, fig. 2F). The punctate pronotal disc, head, and prosternum + proepisternum are shared with
<taxonomicName id="69341E336D16CA319420FF73DA87859E" lsidName="M. punctatus" pageId="0" pageNumber="177" rank="species" species="punctatus">M. punctatus</taxonomicName>
, but the pronotal punctures are more numerous and less separated in
<taxonomicName id="D57657D8E89E43390D260500983F9D0E" lsidName="M. punctatus" pageId="0" pageNumber="177" rank="species" species="punctatus">M. punctatus</taxonomicName>
(Fig. 2
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), with about 30 punctures each side in
<taxonomicName id="90F1BB0BF1ACCED277C190B475FB15BA" lsidName="M. moorei" pageId="0" pageNumber="177" rank="species" species="moorei">M. moorei</taxonomicName>
, and about 60 each side in
<taxonomicName id="1FB01F1A9BD628592A38BB441479FD8C" lsidName="M. punctatus" pageId="0" pageNumber="177" rank="species" species="punctatus">M. punctatus</taxonomicName>
. In addition, the elytral striae are more distinct in
<taxonomicName id="E79676B2AB6B2C6CC1B787B1D86A94AF" lsidName="M. punctatus" pageId="0" pageNumber="177" rank="species" species="punctatus">M. punctatus</taxonomicName>
, with striae 1-5 present in the basal half of the elytra, though the punctures of stria 5 are smaller and shallower. The elytra of
<taxonomicName id="8EA21BEA1E7D768F5B8CED7C40569E3C" lsidName="M. moorei" pageId="0" pageNumber="177" rank="species" species="moorei">M. moorei</taxonomicName>
are more ovoid, with the lateral margins more narrowly rounded behind the subangulate humeri (Fig. 3
<normalizedToken id="CCCA0AB3BBD6FF293FF502E314564477" originalValue="A–B">A-B</normalizedToken>
). Whereas the parascutellar seta is present or absent in
<taxonomicName id="D569159E415DB9651D2132B74079AA0A" lsidName="M. punctatus" pageId="0" pageNumber="177" rank="species" species="punctatus">M. punctatus</taxonomicName>
, and if present appearing short and narrow, this seta is well developed in
<taxonomicName id="0CC10D205D8FF88709A42CDE93935E8A" lsidName="M. moorei" pageId="0" pageNumber="177" rank="species" species="moorei">M. moorei</taxonomicName>
, with the setal articulatory socket set within a depression coincident with the base of the parascutellar striole, the seta as long as the breadth of 2-3 elytral intervals. The eyes of
<taxonomicName id="859DEB4052330AE164623F8B2FB109B7" lsidName="M. moorei" pageId="0" pageNumber="177" rank="species" species="moorei">M. moorei</taxonomicName>
are convex but they cover only the anterior 4/5 of the ocular lobe: EyL/OLL = 0.83-0.86. Finally, males of this species,
<taxonomicName id="9A26539FA4DB0CAD1EDB30024DDF0021" lsidName="M. punctatus" pageId="0" pageNumber="177" rank="species" species="punctatus">M. punctatus</taxonomicName>
, and
<taxonomicName id="2EFB7563D9DCA80011882485B3D79101" lsidName="M. curtus" pageId="0" pageNumber="177" rank="species" species="curtus">M. curtus</taxonomicName>
all exhibit 2 setae each side of the apical abdominal ventrite, an unusual character state within
<taxonomicName id="CBB5E832230E5AE8FE5266872EA3966C" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
, and evidence of their monophyletic relationship within subgenus
<taxonomicName id="370D3CAD8FA48397337EDBA6AB148792" class="Insecta" family="Carabidae" genus="Moriomorphina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Moriomorphina (Eucyclothorax)" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="subGenus" subGenus="Eucyclothorax">Eucyclothorax</taxonomicName>
(Fig. 1B, character 83). Standardized body length 3.2-3.3 mm. Setal formula ++/++/+2++.
</paragraph>
<paragraph id="6C29AF94B8CCC48139D600432C1E38C8" pageId="0" pageNumber="177">Male genitalia (n = 2). Aedeagal median lobe apex narrowly rounded and only slightly projected beyond apical ostial margin (Fig. 7A); aedeagal internal sac elongate, membranous, with apical dorsal plate, flagellum, and flagellar sheath (Fig. 7B); right paramere narrow, elongate, with 6 setae along the ventral margin, 1-2 apical setae, and the dorsal margin glabrous except for a small apical seta (Fig. 8A); left paramere broadly quadrate basally, the apical 1/3 of length very narrow in contrast to broad base.</paragraph>
<paragraph id="17031AD10E677427DE78ACCD90DBD6CC" pageId="0" pageNumber="177">
Female reproductive tract (n = 1). Bursa copulatrix elongate, columnar (Fig. 9A); helminthoid sclerite elongate; spermathecal duct basally expanded and sclerotized, the more apical portion of duct membranous and of lesser diameter; basal gonocoxite with 2 larger setae laterally along apical margin (Fig. 10A), a very small seta medially near margin; apical gonocoxite broad basally, breadth more than half length; lateral ensiform setae elongate, ~ 0.60
<normalizedToken id="C431504AE30535D99A8B1D7CD96C540D" originalValue="×">x</normalizedToken>
length of apical gonocoxite; apical nematiform setae in subbasal sensory furrow.
</paragraph>
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<paragraph id="D1350FD21480E4828F1EAEF3F0695274" pageId="0" pageNumber="177">Type information.</paragraph>
<paragraph id="7FEB945C9DCA1942EA90CC28BC37C6E7" pageId="0" pageNumber="177">
Holotype male (AMS): site 32AR NSW Banda Rd about 4.5 km E Hastings Forest Hwy
<geoCoordinate id="8D728B7C93298F051D5CEA6D416B670A" direction="south" orientation="latitude" precision="925" value="-31.15">31°09'S</geoCoordinate>
<geoCoordinate id="C1401A72B888188DE4AC1647187408EB" direction="east" orientation="longitude" precision="925" value="152.41667">152°25'E</geoCoordinate>
Mount Boss State Forest 17 1100m (NPWS Survey) 4
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9 Apr 1993 M. Gray, G. Cassis (AMS K241125) (
<bibRefCitation id="FA081665E7658FAE105EC269F9824035" author="Baehr, M" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="177" pagination="89 - 92" title="A new species of the genus Mecyclothorax Sharp from New South Wales (Insecta: Carabidae: Psydrinae)." url="https://doi.org/10.3853/j.0067-1975.61.2009.1519" volume="61" year="2009">Baehr 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3E18F7CAB548773C550FD74ECAA3C3F6" pageId="0" pageNumber="177" type="distribution">
<paragraph id="2D096C7E908FFA1208E671EA19D28775" pageId="0" pageNumber="177">Distribution and habitat.</paragraph>
<paragraph id="0C70F63CC6BC123CDCD49AD7B59EB841" pageId="0" pageNumber="177">
<taxonomicName id="EF34CFC830F668B859DF44615F8997AC" lsidName="M. moorei" pageId="0" pageNumber="177" rank="species" species="moorei">M. moorei</taxonomicName>
is restricted to northeastern New South Wales (Fig. 11A), with populations allopatrically distributed relative to eastern populations of the more southerly distributed
<taxonomicName id="04365C168585E26492F9DF990FA48E9F" lsidName="M. punctatus" pageId="0" pageNumber="177" rank="species" species="punctatus">M. punctatus</taxonomicName>
. Recorded elevations of collecting localities range from 110 m near Ramornie and 1100 m in Mt. Boss State Forest (
<bibRefCitation id="FE915AB12EB2B84B0598CFA5E14D3DAA" author="Baehr, M" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="177" pagination="89 - 92" title="A new species of the genus Mecyclothorax Sharp from New South Wales (Insecta: Carabidae: Psydrinae)." url="https://doi.org/10.3853/j.0067-1975.61.2009.1519" volume="61" year="2009">Baehr 2009</bibRefCitation>
). Philip Darlington collected this species in Bellangry Forest NW of Wauchope (MCZ). All specimens are vestigially winged.
</paragraph>
</subSubSection>
</treatment>
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<treatmentCitation id="5B3B7B602CDF29A3B7EB37263DCF5995" author="Linnaeus" pageNumber="794" year="1753">
<bibCitation id="41C728A9CFBCF16AF738F54A46579B2E" author="Linnaeus" pagination="794" title="Sonchus oleraceus Linnaeus var. asper" volumeTitle="Species Plantarum 2" year="1753">
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Linnaeus var.
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Linnaeus
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,
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2
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: 794. 1753
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.
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<paragraph id="0C86AD3E2DB588F23F993FE5D863D985" pageNumber="864">RCN: 5808.</paragraph>
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(Boulos in Jarvis &amp; Turland in
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47: 368. 1998): Herb. Burser VI: 14 (
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="F40E1450685C0E8E6523543F8582C20C" pageNumber="865" type="nomenclature">
<paragraph id="61420596FE0711169B0EA2492B00514F" pageNumber="865">
Current name:
<taxonomicName id="D0D7CBEE7F4A4E49ADBEA35C19287B1D" ID-CoL="4Y96B" authority="(L.) Hill subsp. asper" baseAuthorityName="Linnaeus" baseAuthorityYear="1753" class="Magnoliopsida" family="Asteraceae" genus="Sonchus" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="subSpecies" species="asper" subSpecies="asper">
<emphasis id="528847274D4AFF05EA740D20C5EC948D" italics="true" pageNumber="865">
<emphasis id="B398ABCA49623F5F9201FB0F09727287" bold="true" italics="true" pageNumber="865">Sonchus asper</emphasis>
</emphasis>
(L.) Hill subsp.
<emphasis id="4EEDBBEDC28963F0B2628E3A5F2904BB" italics="true" pageNumber="865">
<emphasis id="FE1CAA1492D8F58FB8B292CCA174A414" bold="true" italics="true" pageNumber="865">asper</emphasis>
</emphasis>
</taxonomicName>
(
<taxonomicName id="F5C53C61EB3CF08FC700F744A7601941" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="47C5B5821D6A07430B08A2FC01A50AF9" pageNumber="865" type="discussion">
<paragraph id="2C2C293E5142709D596C09EAA174D8B9" pageNumber="865">
<emphasis id="1614E52A4FB180E02DD06172F9BB8787" italics="true" pageNumber="865">Note:</emphasis>
Boulos (in
<emphasis id="6169C5D64C4AD5DF55DA8266C28E1395" italics="true" pageNumber="865">Bot. Not.</emphasis>
126: 164. 1973) and a number of later authors all treated
<specimenReference id="39D66FEA83D81295C15C3BD70DB15659" collectionCode="LINN" specimenCode="949.8">
Herb. Linn. No. 949.8 (
<collectionCode id="B37FFE8000AB6A42C6BD2600C91B36D4">LINN</collectionCode>
)
</specimenReference>
as the type, but this sheet is unannotated as to species by Linnaeus and is not original material for the name. Boulos therefore superseded this choice with a new typification in 1998.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.137.46544" ID-PMC="PMC6962254" ID-Pensoft-Pub="1314-2003-137-1" ID-Pensoft-UUID="FB2C14EFA257564AA84CD47711BBC273" ID-PubMed="31969792" ModsDocID="1314-2003-137-1" checkinTime="1578614985992" checkinUser="pensoft" docAuthor="Zaika, Maxim A., Kilian, Norbert, Jones, Katy, Krinitsina, Anastasiya A., Nilova, Maya V., Speranskaya, Anna S. &amp; Sukhorukov, Alexander P." docDate="2020" docId="19EBB35BCA7558339C97C8192AFB5695" docLanguage="en" docName="PhytoKeys 137: 1-85" docOrigin="PhytoKeys 137" docSource="http://dx.doi.org/10.3897/phytokeys.137.46544" docTitle="Koelpinia Pall., Reise Russ. Reich. 3: 755. 1776" docType="treatment" docVersion="5" id="FB2C14EFA257564AA84CD47711BBC273" lastPageNumber="1" masterDocId="FB2C14EFA257564AA84CD47711BBC273" masterDocTitle="Scorzonera sensu lato (Asteraceae, Cichorieae) - taxonomic reassessment in the light of new molecular phylogenetic and carpological analyses" masterLastPageNumber="85" masterPageNumber="1" pageNumber="1" updateTime="1668139594603" updateUser="ExternalLinkService">
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<mods:title>Scorzonera sensu lato (Asteraceae, Cichorieae) - taxonomic reassessment in the light of new molecular phylogenetic and carpological analyses</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zaika, Maxim A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kilian, Norbert</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jones, Katy</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Krinitsina, Anastasiya A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nilova, Maya V.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Speranskaya, Anna S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sukhorukov, Alexander P.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>137</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>85</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.137.46544</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.137.46544</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-137-1</mods:identifier>
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<treatment ID-GBIF-Taxon="161671731" LSID="urn:lsid:plazi:treatment:19EBB35BCA7558339C97C8192AFB5695" httpUri="http://treatment.plazi.org/id/19EBB35BCA7558339C97C8192AFB5695" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="19EBB35B-CA75-5833-9C97-C8192AFB5695" authority="Pall., Reise Russ. Reich. 3: 755. 1776." class="Magnoliopsida" family="Asteraceae" genus="Koelpinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Koelpinia" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Koelpinia Pall., Reise Russ. Reich. 3: 755. 1776.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pall" authorityYear="1776" class="Magnoliopsida" family="Asteraceae" genus="Koelpinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Koelpinia linearis" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="linearis">
<emphasis italics="true" pageId="0" pageNumber="1">Koelpinia linearis</emphasis>
</taxonomicName>
Pall.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnostic features">
<paragraph pageId="0" pageNumber="1">Diagnostic features.</paragraph>
<paragraph pageId="0" pageNumber="1">
Annual herbs; pollen with 15 lacunae; at least outer achenes columnar-scorpioid, surface with hooked emergences and elongated papillae looking like glandular hairs, (sometimes achenes smooth, without emergences and papillae:
<taxonomicName lsidName="K. tenuissima" pageId="0" pageNumber="1" rank="species" species="tenuissima">
<emphasis italics="true" pageId="0" pageNumber="1">K. tenuissima</emphasis>
</taxonomicName>
), without pappus, but apically with hooked, retrorse spines.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Habit, life form, subterranean parts:</emphasis>
annual herbs with a taproot.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Leaves:</emphasis>
subsessile, filiform to narrowly oblong, entire.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Stem, synflorescence:</emphasis>
stem solitary or mostly branched, glabrous or with scattered simple hairs; synflorescence mostly branched, with several capitula.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Capitula:</emphasis>
involucre glabrous to hairy, phyllaries in several series, narrowly oblong to lanceolate, outer phyllaries much shorter than inner; receptacle naked or with scattered setae; capitula with 5-12 yellow florets, much exceeding the involucre.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pollen:</emphasis>
echinolophate, tricolporate and each colpus divided into 2 lacunae, with 15 (6 abporal, 3 equatorial, 6 interporal) lacunae, often pollen tetracolporate in polyploids (
<bibRefCitation DOI="https://doi.org/10.1111/j.1095-8339.1981.tb00947.x" author="Blackmore, S" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="1 - 13" refId="B8" refString="Blackmore, S, 1981. Palynology and intergeneric relationships in subtribe Hyoseridinae (Compositae: Lactuceae). Botanical Journal of the Linnean Society 82 (1): 1 - 13, DOI: https://doi.org/10.1111/j.1095-8339.1981.tb00947.x" title="Palynology and intergeneric relationships in subtribe Hyoseridinae (Compositae: Lactuceae)." url="https://doi.org/10.1111/j.1095-8339.1981.tb00947.x" volume="82" year="1981">Blackmore 1981</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Achenes:</emphasis>
6-25 mm, usually columnar-scorpioid, (sub)annular (
<taxonomicName lsidName="K. macrantha" pageId="0" pageNumber="1" rank="species" species="macrantha">
<emphasis italics="true" pageId="0" pageNumber="1">K. macrantha</emphasis>
</taxonomicName>
) or almost straight (
<taxonomicName lsidName="K. tenuissima" pageId="0" pageNumber="1" rank="species" species="tenuissima">
<emphasis italics="true" pageId="0" pageNumber="1">K. tenuissima</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="K. turanica" pageId="0" pageNumber="1" rank="species" species="turanica">
<emphasis italics="true" pageId="0" pageNumber="1">K. turanica</emphasis>
</taxonomicName>
), with 5 or 10 more or less prominent ribs, surface often covered with emergences resembling hooked spines and glandular-like (elongated) papillae or surface almost glabrous in
<taxonomicName lsidName="K. tenuissima" pageId="0" pageNumber="1" rank="species" species="tenuissima">
<emphasis italics="true" pageId="0" pageNumber="1">K. tenuissima</emphasis>
</taxonomicName>
, without carpopodium; achene wall with or without air cavities, parenchyma continuous above sclerenchyma represented by thick-walled cells, sclerenchyma continuous, its fibres orientated parallel to the fruit axis (perpendicular in emergences), tannins absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pappus:</emphasis>
bristles absent, achenes with several retrorse hooked spines (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Images of living plants. A, B - Tourneuxia variifolia (Morocco, near Mrhimina). Photographs by A. Garcin (www. teline. fr) C - Gelasia villosa (cult. in BG Berlin, 13 Jun 2010). Photograph by N. Kilian D, E, F - Gelasia rigida (Jordan, Nakeb, 1250 m alt., 8 May 2019). Photograph by O. Fragman-Sapir. D general view E capitula in flowering F capitula in fruiting, a part of achenes had fallen off G Geropogon hybridus (Greece, Lefkas, 15 May 2016). Photograph by E. Willing H Koelpinia linearis (Qatar, Khashem Al Nekhsh area, 18 Mar 2016). Photograph by A. Sergeev." figureDoi="10.3897/phytokeys.137.46544.figure15" httpUri="https://binary.pensoft.net/fig/370363" pageId="0" pageNumber="1">15H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="chromosome number">
<paragraph pageId="0" pageNumber="1">Chromosome number.</paragraph>
<paragraph pageId="0" pageNumber="1">
x = 6 for
<taxonomicName lsidName="K. macrantha" pageId="0" pageNumber="1" rank="species" species="macrantha">
<emphasis italics="true" pageId="0" pageNumber="1">K. macrantha</emphasis>
</taxonomicName>
(
<bibRefCitation author="Podlech, D" journalOrPublisher="Mitteilungen aus der Botanischen Staatssammlung Muenchen" pageId="0" pageNumber="1" pagination="457 - 488" refId="B81" refString="Podlech, D, Bader, O, 1974. Chromosomenstudien an afghanischen Pflanzen. 2. Mitteilungen aus der Botanischen Staatssammlung Muenchen 11: 457 - 488" title="Chromosomenstudien an afghanischen Pflanzen. 2." volume="11" year="1974">Podlech and Bader 1974</bibRefCitation>
;
<bibRefCitation author="Nazarova, EA" journalOrPublisher="Botanicheskii Zhurnal" pageId="0" pageNumber="1" pagination="1755 - 1758" refId="B71" refString="Nazarova, EA, 1981. Tsitosistematicheskoe izuchenie roda Koelpinia (Asteraceae). Botanicheskii Zhurnal 66: 1755 - 1758" title="Tsitosistematicheskoe izuchenie roda Koelpinia (Asteraceae)." volume="66" year="1981">Nazarova 1981</bibRefCitation>
) or 7 (all others species) (
<bibRefCitation author="Stebbins, GL" journalOrPublisher="University of California Publications in Botany" pageId="0" pageNumber="1" pagination="401 - 430" refId="B93" refString="Stebbins, GL, Jenkins, JA, Walters, MS, 1953. Chromosomes and phylogeny in the Compositae, tribe Chichorieae. University of California Publications in Botany 26: 401 - 430" title="Chromosomes and phylogeny in the Compositae, tribe Chichorieae." volume="26" year="1953">Stebbins et al. 1953</bibRefCitation>
;
<bibRefCitation author="Podlech, D" journalOrPublisher="Mitteilungen aus der Botanischen Staatssammlung Muenchen" pageId="0" pageNumber="1" pagination="457 - 488" refId="B81" refString="Podlech, D, Bader, O, 1974. Chromosomenstudien an afghanischen Pflanzen. 2. Mitteilungen aus der Botanischen Staatssammlung Muenchen 11: 457 - 488" title="Chromosomenstudien an afghanischen Pflanzen. 2." volume="11" year="1974">Podlech and Bader 1974</bibRefCitation>
;
<bibRefCitation author="Nazarova, EA" journalOrPublisher="Botanicheskii Zhurnal" pageId="0" pageNumber="1" pagination="1755 - 1758" refId="B71" refString="Nazarova, EA, 1981. Tsitosistematicheskoe izuchenie roda Koelpinia (Asteraceae). Botanicheskii Zhurnal 66: 1755 - 1758" title="Tsitosistematicheskoe izuchenie roda Koelpinia (Asteraceae)." volume="66" year="1981">Nazarova 1981</bibRefCitation>
), diploids to octaploids.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="species">
<paragraph pageId="0" pageNumber="1">Species.</paragraph>
<paragraph pageId="0" pageNumber="1">
(1)
<taxonomicName lsidName="K. chrysoglochis" pageId="0" pageNumber="1" rank="species" species="chrysoglochis">
<emphasis italics="true" pageId="0" pageNumber="1">K. chrysoglochis</emphasis>
</taxonomicName>
Rech.f.
</paragraph>
<paragraph pageId="0" pageNumber="1">
(2)
<taxonomicName lsidName="K. linearis" pageId="0" pageNumber="1" rank="species" species="linearis">
<emphasis italics="true" pageId="0" pageNumber="1">K. linearis</emphasis>
</taxonomicName>
Pall.
</paragraph>
<paragraph pageId="0" pageNumber="1">
(3)
<taxonomicName lsidName="K. macrantha" pageId="0" pageNumber="1" rank="species" species="macrantha">
<emphasis italics="true" pageId="0" pageNumber="1">K. macrantha</emphasis>
</taxonomicName>
C.Winkl.
</paragraph>
<paragraph pageId="0" pageNumber="1">
(4)
<taxonomicName lsidName="K. tenuissima" pageId="0" pageNumber="1" rank="species" species="tenuissima">
<emphasis italics="true" pageId="0" pageNumber="1">K. tenuissima</emphasis>
</taxonomicName>
Pavlov &amp; Lipsch.
</paragraph>
<paragraph pageId="0" pageNumber="1">
(5)
<taxonomicName lsidName="K. turanica" pageId="0" pageNumber="1" rank="species" species="turanica">
<emphasis italics="true" pageId="0" pageNumber="1">K. turanica</emphasis>
</taxonomicName>
Vassilcz.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution area.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">N Africa</emphasis>
: DZ; EG; LY; TN.
<emphasis bold="true" pageId="0" pageNumber="1">Asia-Temperate</emphasis>
: AF; AM; AZ; BH; CN; CY; GE; IL; IQ; IR; KG; KW; KZ; LB; QA; SA; SY; TJ; TM; TR; UZ.
<emphasis bold="true" pageId="0" pageNumber="1">Asia-Tropical</emphasis>
: IN; PK.
<emphasis bold="true" pageId="0" pageNumber="1">Europe</emphasis>
: ES; RU; UA.
</paragraph>
</subSubSection>
</treatment>
</document>