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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1092.75831" ID-GBIF-Dataset="c2f1268d-f466-4148-990f-2b948efcf6a7" ID-PMC="PMC9005484" ID-Pensoft-Pub="1313-2970-1092-31" ID-Pensoft-UUID="B3F12D9131AD5C878EAC9A12C068EA6D" ID-PubMed="35586293" ID-ZooBank="28BB93BFA3D54EF5A91867AECD12B808" ModsDocID="1313-2970-1092-31" checkinTime="1649136097056" checkinUser="pensoft" docAuthor="Zhao, Ming-Zhi &amp; Liu, Wei-Xin" docDate="2022" docId="DF09F378981E53868D55E4A895CD8DFD" docLanguage="en" docName="ZooKeys 1092: 31-45" docOrigin="ZooKeys 1092" docPubDate="2022-04-04" docSource="http://dx.doi.org/10.3897/zookeys.1092.75831" docTitle="Didrepanephorus mizunumai Nagai &amp; Hirasawa 1991" docType="treatment" docVersion="4" id="B3F12D9131AD5C878EAC9A12C068EA6D" lastPageNumber="31" masterDocId="B3F12D9131AD5C878EAC9A12C068EA6D" masterDocTitle="A contribution to the genus Didrepanephorus Wood-Mason, 1878 (Coleoptera, Scarabaeidae, Rutelinae)" masterLastPageNumber="45" masterPageNumber="31" pageNumber="31" updateTime="1668151609506" updateUser="ExternalLinkService">
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<mods:title id="151A844AE8EFF69CF66CC2A891D57BAF">A contribution to the genus Didrepanephorus Wood-Mason, 1878 (Coleoptera, Scarabaeidae, Rutelinae)</mods:title>
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<mods:namePart>Zhao, Ming-Zhi</mods:namePart>
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<mods:namePart>Liu, Wei-Xin</mods:namePart>
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<treatment ID-GBIF-Taxon="194444414" LSID="urn:lsid:plazi:treatment:DF09F378981E53868D55E4A895CD8DFD" httpUri="http://treatment.plazi.org/id/DF09F378981E53868D55E4A895CD8DFD" lastPageNumber="31" pageId="0" pageNumber="31">
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<taxonomicName LSID="DF09F378-981E-5386-8D55-E4A895CD8DFD" authority="Nagai &amp; Hirasawa, 1991" authorityName="Nagai &amp; Hirasawa" authorityYear="1991" class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus mizunumai" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="mizunumai">Didrepanephorus mizunumai Nagai &amp; Hirasawa, 1991</taxonomicName>
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<taxonomicName id="C783F10951394A272A3389D4578B1697" ID-CoL="6D4ZD" LSID="DF09F378-981E-5386-8D55-E4A895CD8DFD" authority="Nagai &amp; Hirasawa, 1991" authorityName="Nagai &amp; Hirasawa" authorityYear="1991" class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus mizunumai" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="mizunumai">Didrepanephorus mizunumai Nagai &amp; Hirasawa, 1991</taxonomicName>
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<figureCitation captionStart="Figures 11–18" captionStartId="F3" captionText="Figures 11 - 18. Type material of Didrepanephorus species. 11 - 15 male of D. birmanicus (Arrow, 1907) (Lectotype of Fruhstorferia birmanica Arrow, 1907) 16 - 18 female of D. mizunumai Nagai &amp; Hirasawa, 1991 (paralectotype of F. birmanica), habitus in dorsal view 11, 16 habitus in dorsal view 12, 17 habitus in lateral view 13 aedeagus showing parameres in dorsal view 14 aedeagus in dorsal view 15, 18 attached labels. All photos © Keita Matsumoto (NHMUK)." figureDoi="10.3897/zookeys.1092.75831.figures11-18" httpUri="https://binary.pensoft.net/fig/668069" pageId="0" pageNumber="31">Figs 16-18</figureCitation>
<subSubSection id="6096599E20A51EC03AC15B7BA70EB20F" pageId="0" pageNumber="31" type="description">
<paragraph id="35C5E38C23589FE75F6E2E03FD2D9EF2" pageId="0" pageNumber="31">
<figureCitation id="04F11ED24BD59B82CFD059AE2CF5DB52" captionStart="Figures 11–18" captionStartId="F3" captionText="Figures 11 - 18. Type material of Didrepanephorus species. 11 - 15 male of D. birmanicus (Arrow, 1907) (Lectotype of Fruhstorferia birmanica Arrow, 1907) 16 - 18 female of D. mizunumai Nagai &amp; Hirasawa, 1991 (paralectotype of F. birmanica), habitus in dorsal view 11, 16 habitus in dorsal view 12, 17 habitus in lateral view 13 aedeagus showing parameres in dorsal view 14 aedeagus in dorsal view 15, 18 attached labels. All photos © Keita Matsumoto (NHMUK)." figureDoi="10.3897/zookeys.1092.75831.figures11-18" httpUri="https://binary.pensoft.net/fig/668069" pageId="0" pageNumber="31">Figs 16-18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="reference_group">
<paragraph pageId="0" pageNumber="31">
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">Fruhstorferia birmanica</taxonomicName>
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<paragraph id="1BF300F2B82F60253A63980DEBDDDD83" pageId="0" pageNumber="31">
<taxonomicName id="3E8ACCF47557D6C544308273BCE43557" authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">Fruhstorferia birmanica</taxonomicName>
Arrow, 1907:
<bibRefCitation DOI="https://doi.org/10.1080/00222930709487274" author="Arrow, GJ" pageId="0" pageNumber="31" refId="B1" refString="Arrow, GJ, 1907. Some new species and genera of lamellicorn Coleoptera from the Indian Empire (part 1). The Annals and Magazine of natural History, including Zoology, Botany and Geology Series 7 19(112): 347-359. https://doi.org/10.1080/00222930709487274" title="Some new species and genera of lamellicorn Coleoptera from the Indian Empire (part 1). The Annals and Magazine of natural History, including Zoology, Botany and Geology Series 7 19 (112): 347 - 359." url="https://doi.org/10.1080/00222930709487274" year="1907">Arrow 1907</bibRefCitation>
<bibRefCitation id="3901FFD2CC2D3536721DA6DF0D6CBB58" DOI="https://doi.org/10.1080/00222930709487274" author="Arrow, GJ" pageId="0" pageNumber="31" refId="B1" refString="Arrow, GJ, 1907. Some new species and genera of lamellicorn Coleoptera from the Indian Empire (part 1). The Annals and Magazine of natural History, including Zoology, Botany and Geology Series 7 19(112): 347-359. https://doi.org/10.1080/00222930709487274" title="Some new species and genera of lamellicorn Coleoptera from the Indian Empire (part 1). The Annals and Magazine of natural History, including Zoology, Botany and Geology Series 7 19 (112): 347 - 359." url="https://doi.org/10.1080/00222930709487274" year="1907">Arrow 1907</bibRefCitation>
: 354 [partim, female];
<bibRefCitation author="Arrow, GJ" journalOrPublisher="Taylor &amp; Francis, London" pageId="0" pageNumber="31" refId="B2" refString="Arrow, GJ, 1917. The Fauna of British India, Including Ceylon and Burma. ColeopteraLamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis, London" title="The Fauna of British India, Including Ceylon and Burma. ColeopteraLamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae)." year="1917">Arrow 1917</bibRefCitation>
<bibRefCitation id="57081914B5BD4DE0FFA8E98607239114" author="Arrow, GJ" journalOrPublisher="Taylor &amp; Francis, London" pageId="0" pageNumber="31" refId="B2" refString="Arrow, GJ, 1917. The Fauna of British India, Including Ceylon and Burma. ColeopteraLamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor &amp; Francis, London" title="The Fauna of British India, Including Ceylon and Burma. ColeopteraLamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae)." year="1917">Arrow 1917</bibRefCitation>
: 49, fig. 16 [partim, female].
</paragraph>
<paragraph pageId="0" pageNumber="31">
<taxonomicName authorityName="Nagai &amp; Hirasawa" authorityYear="1991" class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus mizunumai" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="mizunumai">Didrepanephorus mizunumai</taxonomicName>
<paragraph id="1F36812A65C75F2B3691B177BC2EF5F5" pageId="0" pageNumber="31">
<taxonomicName id="F631C960DD14E0E3DC1C4E2CCD18AE4A" authorityName="Nagai &amp; Hirasawa" authorityYear="1991" class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus mizunumai" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="mizunumai">Didrepanephorus mizunumai</taxonomicName>
Nagai &amp; Hirasawa, 1991: 10, figs 2-5, 20, 32-34, 39-40 [original description];
<bibRefCitation author="Nagai, S" journalOrPublisher="Japanese Journal of Systematic Entomology" pageId="0" pageNumber="31" pagination="269 - 274" refId="B27" refString="Nagai, S, 2005. A new species and two new subspecies of the Rutelinae from southeast Asia. Japanese Journal of Systematic Entomology 11 (2): 269 - 274" title="A new species and two new subspecies of the Rutelinae from southeast Asia." volume="11" year="2005">Nagai 2005</bibRefCitation>
<bibRefCitation id="580855EC4634A1D6C2613753772E9EC4" author="Nagai, S" journalOrPublisher="Japanese Journal of Systematic Entomology" pageId="0" pageNumber="31" pagination="269 - 274" refId="B27" refString="Nagai, S, 2005. A new species and two new subspecies of the Rutelinae from southeast Asia. Japanese Journal of Systematic Entomology 11 (2): 269 - 274" title="A new species and two new subspecies of the Rutelinae from southeast Asia." volume="11" year="2005">Nagai 2005</bibRefCitation>
: 271, figs 4-6, 17;
<bibRefCitation author="Muramoto, R" journalOrPublisher="Kogane" pageId="0" pageNumber="31" pagination="57 - 59" refId="B19" refString="Muramoto, R, 2009. A new species of the genus Didrepanephorus from Vietnam. Kogane 10: 57 - 59" title="A new species of the genus Didrepanephorus from Vietnam." volume="10" year="2009">Muramoto 2009</bibRefCitation>
<bibRefCitation id="ABF5023FE2B68A77534F5E0CC589D2E9" author="Muramoto, R" journalOrPublisher="Kogane" pageId="0" pageNumber="31" pagination="57 - 59" refId="B19" refString="Muramoto, R, 2009. A new species of the genus Didrepanephorus from Vietnam. Kogane 10: 57 - 59" title="A new species of the genus Didrepanephorus from Vietnam." volume="10" year="2009">Muramoto 2009</bibRefCitation>
: 59, figs 4-6.
</paragraph>
<paragraph pageId="0" pageNumber="31">
<taxonomicName baseAuthorityName="Nagai &amp; Hirasawa" baseAuthorityYear="1991" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia mizunumai" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="mizunumai">Fruhstorferia mizunumai</taxonomicName>
<paragraph id="25C9C4B8A18D4491A9A309622E5F3CA4" pageId="0" pageNumber="31">
<taxonomicName id="8D02729CFF7B3AA5F5D3708C6E923687" baseAuthorityName="Nagai &amp; Hirasawa" baseAuthorityYear="1991" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia mizunumai" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="mizunumai">Fruhstorferia mizunumai</taxonomicName>
(Nagai &amp; Hirasawa, 1991):
<bibRefCitation author="Jameson, ML" journalOrPublisher="Bulletin of the University of Nebraska State Museum" pageId="0" pageNumber="31" pagination="1 - 184" refId="B9" refString="Jameson, ML, 1997. Phylogenetic analysis of the subtribe Rutelina and revision of the Rutela generic groups. Bulletin of the University of Nebraska State Museum 14: 1 - 184" title="Phylogenetic analysis of the subtribe Rutelina and revision of the Rutela generic groups." volume="14" year="1997">Jameson 1997</bibRefCitation>
<bibRefCitation id="B9A5088E4C8A2C6D574E27971DBE783C" author="Jameson, ML" journalOrPublisher="Bulletin of the University of Nebraska State Museum" pageId="0" pageNumber="31" pagination="1 - 184" refId="B9" refString="Jameson, ML, 1997. Phylogenetic analysis of the subtribe Rutelina and revision of the Rutela generic groups. Bulletin of the University of Nebraska State Museum 14: 1 - 184" title="Phylogenetic analysis of the subtribe Rutelina and revision of the Rutela generic groups." volume="14" year="1997">Jameson 1997</bibRefCitation>
: 170, figs 21-22 [new combination];
<bibRefCitation author="Nagai, S" journalOrPublisher="Japanese Journal of Systematic Entomology" pageId="0" pageNumber="31" pagination="145 - 157" refId="B26" refString="Nagai, S, 2004. Two new species and four new subspecies of the Rutelinae from southeast Asia. Japanese Journal of Systematic Entomology 10 (1): 145 - 157" title="Two new species and four new subspecies of the Rutelinae from southeast Asia." volume="10" year="2004">Nagai 2004</bibRefCitation>
<bibRefCitation id="8E246EBA9BBE5DE319D62DF915CFDD6B" author="Nagai, S" journalOrPublisher="Japanese Journal of Systematic Entomology" pageId="0" pageNumber="31" pagination="145 - 157" refId="B26" refString="Nagai, S, 2004. Two new species and four new subspecies of the Rutelinae from southeast Asia. Japanese Journal of Systematic Entomology 10 (1): 145 - 157" title="Two new species and four new subspecies of the Rutelinae from southeast Asia." volume="10" year="2004">Nagai 2004</bibRefCitation>
: 150, figs 13-16, 27 [recorded from Houa Phan Prov., Laos];
<bibRefCitation author="Krajcik, M" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="31" refId="B11" refString="Krajcik, M, 2007. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. x(Supplementum 4): 1-139." title="Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. x (Supplementum 4): 1 - 139." year="2007">
<normalizedToken originalValue="Krajčík">Krajcik</normalizedToken>
<bibRefCitation id="4B7ED4E998681124913F46747153E75A" author="Krajcik, M" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="31" refId="B11" refString="Krajcik, M, 2007. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. x(Supplementum 4): 1-139." title="Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. x (Supplementum 4): 1 - 139." year="2007">
<normalizedToken id="410197A54F91DE2818C222629B82F0FC" originalValue="Krajčík">Krajcik</normalizedToken>
2007
</bibRefCitation>
: 71 [catalogued];
<bibRefCitation author="Krajcik, M" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="31" refId="B12" refString="Krajcik, M, 2012. Checklist of the world Scarabaeoidea. Animma. x(Supplementum 5): 1-139." title="Checklist of the world Scarabaeoidea. Animma. x (Supplementum 5): 1 - 139." year="2012">
<normalizedToken originalValue="Krajčík">Krajcik</normalizedToken>
<bibRefCitation id="655368FAD3B54413EF42AF8920205EB4" author="Krajcik, M" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="31" refId="B12" refString="Krajcik, M, 2012. Checklist of the world Scarabaeoidea. Animma. x(Supplementum 5): 1-139." title="Checklist of the world Scarabaeoidea. Animma. x (Supplementum 5): 1 - 139." year="2012">
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2012
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: 93 [catalogued, subgeneric placement unnoted].
</paragraph>
<paragraph pageId="0" pageNumber="31">
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia yunnana" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="yunnana">Fruhstorferia yunnana</taxonomicName>
<paragraph id="D1E5B3FC4E0A5574071589CF17BAFCB5" pageId="0" pageNumber="31">
<taxonomicName id="9C0A02EF4F9E29E408B90294D8E589E2" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia yunnana" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="yunnana">Fruhstorferia yunnana</taxonomicName>
Ohaus, 1911:
<bibRefCitation author="Muramoto, R" journalOrPublisher="Gekkan-Mushi" pageId="0" pageNumber="31" pagination="7 - 10" refId="B14" refString="Muramoto, R, 1993. A new species of the genus Kibakoganea from northern Vietnam, with a key to species of Fruhstorferia (wide sense) from northern Vietnam. Gekkan-Mushi 273: 7 - 10" title="A new species of the genus Kibakoganea from northern Vietnam, with a key to species of Fruhstorferia (wide sense) from northern Vietnam." volume="273" year="1993">Muramoto 1993</bibRefCitation>
<bibRefCitation id="5AA936E48F233FB2D2D0DA5769BAF6CD" author="Muramoto, R" journalOrPublisher="Gekkan-Mushi" pageId="0" pageNumber="31" pagination="7 - 10" refId="B14" refString="Muramoto, R, 1993. A new species of the genus Kibakoganea from northern Vietnam, with a key to species of Fruhstorferia (wide sense) from northern Vietnam. Gekkan-Mushi 273: 7 - 10" title="A new species of the genus Kibakoganea from northern Vietnam, with a key to species of Fruhstorferia (wide sense) from northern Vietnam." volume="273" year="1993">Muramoto 1993</bibRefCitation>
: 7, figs 6, 10 [misidentification, recorded from Sapa, N. Vietnam].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="type locality">
<paragraph pageId="0" pageNumber="31">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="31">North Thailand, Chiang Mai Province, Fang.</paragraph>
<subSubSection id="02DFEFD398F9B2C89E9893584A46835D" pageId="0" pageNumber="31" type="type locality">
<paragraph id="2D63DB2E035C70F4C1FFAE03F9E23F42" pageId="0" pageNumber="31">Type locality.</paragraph>
<paragraph id="DE64A1D0092AC5D00BCE674CEC7CC818" pageId="0" pageNumber="31">North Thailand, Chiang Mai Province, Fang.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="materials_examined">
<paragraph pageId="0" pageNumber="31">Material examined</paragraph>
<paragraph pageId="0" pageNumber="31">
<subSubSection id="FE006F1CAEF37502525978418ABE1E49" pageId="0" pageNumber="31" type="materials_examined">
<paragraph id="B4B09EE670D3D1CD1672DFC81248F3A1" pageId="0" pageNumber="31">Material examined</paragraph>
<paragraph id="AD4A2B60E535D5C72EE86CCA5361443C" pageId="0" pageNumber="31">
(
<specimenCount type="generic">5 specimens</specimenCount>
<specimenCount id="F2A96DBD5998A2C4B7968A513DC21581" type="generic">5 specimens</specimenCount>
).
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="India" county="Fauna" location="Zhao Ming-Zhi" municipality="Arrow" specimenCount="1" specimenCount-female="1" typeStatus="PARALECTOTYPE">
<specimenCount type="female">1♀</specimenCount>
<materialsCitation id="60D5FA1D0D423CD8EBE4D1D4FDF1DB45" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="India" county="Fauna" location="Zhao Ming-Zhi" municipality="Arrow" specimenCount="1" specimenCount-female="1" typeStatus="PARALECTOTYPE">
<specimenCount id="AE474A44FC0ED0A93E05A92461BAB815" type="female">1♀</specimenCount>
(BMNH), &quot;Birmah Ruby Mes // Doherty // 64610 // Fry Coll. 1905. 100. //
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis italics="true" pageId="0" pageNumber="31">Fruhstorferia birmanica</emphasis>
<taxonomicName id="7DA2F89C13ED619696E358BEB74F5493" authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis id="9CD5EFC21069A7AD66687674BA7D78F2" italics="true" pageId="0" pageNumber="31">Fruhstorferia birmanica</emphasis>
</taxonomicName>
, Arrow type
<specimenCount>♀</specimenCount>
<specimenCount id="F93BE5099278D1C14A801BE172548F68">♀</specimenCount>
A. a M., 1907. // Figured for &quot;
<collectingCounty>Fauna</collectingCounty>
<collectingCounty id="8B7F057FC4182A5174C4C36B5A65D7F5">Fauna</collectingCounty>
of
<collectingCountry name="India">India</collectingCountry>
<collectingCountry id="196083384EFC61CEE6F28305546F1588" name="India">India</collectingCountry>
.&quot; // Type // NHMUK014379788&quot;, it will be provided with an additional red label:
<typeStatus>PARALECTOTYPE</typeStatus>
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis italics="true" pageId="0" pageNumber="31">Fruhstorferia birmanica</emphasis>
<typeStatus id="E12F42DF083EFFEF325C33C49D624054">PARALECTOTYPE</typeStatus>
<taxonomicName id="71A97EB46A1D9573DB52F96ECA643C5C" authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis id="2484FE12D4C865D21EA7C61C32B6A7DC" italics="true" pageId="0" pageNumber="31">Fruhstorferia birmanica</emphasis>
</taxonomicName>
<collectingMunicipality>Arrow</collectingMunicipality>
<collectingMunicipality id="0F85F4292E1487CDFFE633923FE0A9B8">Arrow</collectingMunicipality>
, 1907 des.
<location LSID="urn:lsid:plazi:treatment:DF09F378981E53868D55E4A895CD8DFD:C9B00C36EE32821D1EA76C4E99570418" country="India" county="Fauna" municipality="Arrow" name="Zhao Ming-Zhi">Zhao Ming-Zhi</location>
<location id="C9B00C36EE32821D1EA76C4E99570418" LSID="urn:lsid:plazi:treatment:DF09F378981E53868D55E4A895CD8DFD:C9B00C36EE32821D1EA76C4E99570418" country="India" county="Fauna" municipality="Arrow" name="Zhao Ming-Zhi">Zhao Ming-Zhi</location>
2021
</materialsCitation>
;
<materialsCitation collectingDate="2011-06-26" country="Thailand" location="Fang" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paralectotype">
<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="A3AF3715A2EB93EAD26FB8B1B46C738C" collectingDate="2011-06-26" country="Thailand" location="Fang" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paralectotype">
<specimenCount id="790EAB7905489E247D7AF1277F1A8ECA" type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="17B6322AA220778A6BC062D7E222D1F0" type="female">1♀</specimenCount>
(ZMPC),
<collectingCountry name="Thailand">Thailand</collectingCountry>
<collectingCountry id="98151CBC49B5AA039225FA074DD8E106" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
<collectingRegion id="10FC360ED1CB7C2015A2F9A9F82196AB" country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:DF09F378981E53868D55E4A895CD8DFD:8AEE6A2217B12A2504D66D7DAA679D36" country="Thailand" name="Fang" stateProvince="Chiang Mai">Fang</location>
<location id="8AEE6A2217B12A2504D66D7DAA679D36" LSID="urn:lsid:plazi:treatment:DF09F378981E53868D55E4A895CD8DFD:8AEE6A2217B12A2504D66D7DAA679D36" country="Thailand" name="Fang" stateProvince="Chiang Mai">Fang</location>
,
<collectingDate value="2011-06-26">26.VI.2011</collectingDate>
<collectingDate id="7963BB888F07CE04FBB1C9576FD605E1" value="2011-06-26">26.VI.2011</collectingDate>
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
<specimenCount id="2D5B98CDE1498B0C9A19812A7977A894" type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
<specimenCount id="D67CFFFFBFAC91D3F3D920286AE8799D" type="female">1♀</specimenCount>
(ZMPC), same as preceding but 2015.V.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="remarks">
<paragraph pageId="0" pageNumber="31">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="31">
<subSubSection id="4CB2272002EC3E7B3CC592244CEEB7F9" pageId="0" pageNumber="31" type="remarks">
<paragraph id="EC4B8E390E79EEF6699BE3AED34E4A3E" pageId="0" pageNumber="31">Remarks.</paragraph>
<paragraph id="3E7FE20916CEA95AC262C40DF6C8833B" pageId="0" pageNumber="31">
The female of
<taxonomicName lsidName="D. mizunumai" pageId="0" pageNumber="31" rank="species" species="mizunumai">
<emphasis italics="true" pageId="0" pageNumber="31">D. mizunumai</emphasis>
<taxonomicName id="90CE1624FD626C07DFD348D45ABC2F38" lsidName="D. mizunumai" pageId="0" pageNumber="31" rank="species" species="mizunumai">
<emphasis id="616E4C562C51EB15C2E1331FD3249282" italics="true" pageId="0" pageNumber="31">D. mizunumai</emphasis>
</taxonomicName>
is easily characterized by the three strongly elevated costae between humeral and apical umbone, as well as a bulge on lateral portions of elytron. The paralectotype of
<taxonomicName lsidName="F. birmanica" pageId="0" pageNumber="31" rank="species" species="birmanica">
<emphasis italics="true" pageId="0" pageNumber="31">F. birmanica</emphasis>
<taxonomicName id="28F1CF1CAAA4DB3723D9DFE86C334EB5" lsidName="F. birmanica" pageId="0" pageNumber="31" rank="species" species="birmanica">
<emphasis id="94F18A46B9A806185A4BB80E17787738" italics="true" pageId="0" pageNumber="31">F. birmanica</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figures 11–18" captionStartId="F3" captionText="Figures 11 - 18. Type material of Didrepanephorus species. 11 - 15 male of D. birmanicus (Arrow, 1907) (Lectotype of Fruhstorferia birmanica Arrow, 1907) 16 - 18 female of D. mizunumai Nagai &amp; Hirasawa, 1991 (paralectotype of F. birmanica), habitus in dorsal view 11, 16 habitus in dorsal view 12, 17 habitus in lateral view 13 aedeagus showing parameres in dorsal view 14 aedeagus in dorsal view 15, 18 attached labels. All photos © Keita Matsumoto (NHMUK)." figureDoi="10.3897/zookeys.1092.75831.figures11-18" httpUri="https://binary.pensoft.net/fig/668069" pageId="0" pageNumber="31">16-18</figureCitation>
<figureCitation id="4124415CFDA2AC529327331311B20E66" captionStart="Figures 11–18" captionStartId="F3" captionText="Figures 11 - 18. Type material of Didrepanephorus species. 11 - 15 male of D. birmanicus (Arrow, 1907) (Lectotype of Fruhstorferia birmanica Arrow, 1907) 16 - 18 female of D. mizunumai Nagai &amp; Hirasawa, 1991 (paralectotype of F. birmanica), habitus in dorsal view 11, 16 habitus in dorsal view 12, 17 habitus in lateral view 13 aedeagus showing parameres in dorsal view 14 aedeagus in dorsal view 15, 18 attached labels. All photos © Keita Matsumoto (NHMUK)." figureDoi="10.3897/zookeys.1092.75831.figures11-18" httpUri="https://binary.pensoft.net/fig/668069" pageId="0" pageNumber="31">16-18</figureCitation>
) well fits topotypical female of
<taxonomicName lsidName="D. mizunumai" pageId="0" pageNumber="31" rank="species" species="mizunumai">
<emphasis italics="true" pageId="0" pageNumber="31">D. mizunumai</emphasis>
<taxonomicName id="DBDA06C688AD1CCF31400B7E53A8CC85" lsidName="D. mizunumai" pageId="0" pageNumber="31" rank="species" species="mizunumai">
<emphasis id="E75997BF10282A701CE81AAA518B841A" italics="true" pageId="0" pageNumber="31">D. mizunumai</emphasis>
</taxonomicName>
from Fang. Therefore, the female paralectotype represents a new distributional record for Myanmar. Two similar
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">Didrepanephorus species</taxonomicName>
<taxonomicName id="5ADF41703A6871FF673C40FBB9CD3402" class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">Didrepanephorus species</taxonomicName>
, i.e., Laotian
<taxonomicName lsidName="D. lao" pageId="0" pageNumber="31" rank="species" species="lao">
<emphasis italics="true" pageId="0" pageNumber="31">D. lao</emphasis>
<taxonomicName id="B6785430F1EC5559CC4EC76CEE795D5D" lsidName="D. lao" pageId="0" pageNumber="31" rank="species" species="lao">
<emphasis id="58E79E6C20D48C6B5C1A64BE74E5032D" italics="true" pageId="0" pageNumber="31">D. lao</emphasis>
</taxonomicName>
Nagai, 2005 and Vietnamese
<taxonomicName lsidName="D. zen" pageId="0" pageNumber="31" rank="species" species="zen">
<emphasis italics="true" pageId="0" pageNumber="31">D. zen</emphasis>
<taxonomicName id="FDA79A8EBD52DD5E7CF6684DAC3295D0" lsidName="D. zen" pageId="0" pageNumber="31" rank="species" species="zen">
<emphasis id="C238594A37F4A9C92401D5925E685DC1" italics="true" pageId="0" pageNumber="31">D. zen</emphasis>
</taxonomicName>
Muramoto, 2009 have minor morphological differences, but both are more restricted in their known distribution ranges.
</paragraph>
<caption doi="10.3897/zookeys.1092.75831.figures11-18" httpUri="https://binary.pensoft.net/fig/668069" pageId="0" pageNumber="31" start="Figures 11–18" startId="F3">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figures 11-18.</emphasis>
<caption id="F5FE8E9AB05C9B9C607F2DBF986ADF14" doi="10.3897/zookeys.1092.75831.figures11-18" httpUri="https://binary.pensoft.net/fig/668069" pageId="0" pageNumber="31" start="Figures 11–18" startId="F3">
<paragraph id="4333877808197A65E3B1A37CA6B75206" pageId="0" pageNumber="31">
<emphasis id="2A582CC2C2736C8732DF67C7A5780B0C" bold="true" pageId="0" pageNumber="31">Figures 11-18.</emphasis>
Type material of
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">
<emphasis italics="true" pageId="0" pageNumber="31">Didrepanephorus species</emphasis>
<taxonomicName id="DE5A2D2EC1FFFC89C94908BEDC781D8B" class="Insecta" family="Scarabaeidae" genus="Didrepanephorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Didrepanephorus species" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species">
<emphasis id="731230C57B85A3601C871A15FF812B24" italics="true" pageId="0" pageNumber="31">Didrepanephorus species</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="31">11-15</emphasis>
<emphasis id="8C776FB02C1E58983B900688C95E5A42" bold="true" pageId="0" pageNumber="31">11-15</emphasis>
male of
<taxonomicName lsidName="D. birmanicus" pageId="0" pageNumber="31" rank="species" species="birmanicus">
<emphasis italics="true" pageId="0" pageNumber="31">D. birmanicus</emphasis>
<taxonomicName id="E6798A3093497EE35B44B1622B1AB6BF" lsidName="D. birmanicus" pageId="0" pageNumber="31" rank="species" species="birmanicus">
<emphasis id="45DBAB39C01C4D8BF351CDF8D3CD5A89" italics="true" pageId="0" pageNumber="31">D. birmanicus</emphasis>
</taxonomicName>
(Arrow, 1907) (Lectotype of
<taxonomicName authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis italics="true" pageId="0" pageNumber="31">Fruhstorferia birmanica</emphasis>
<taxonomicName id="774415E5E9B1ADBABF4A9B90616FC382" authorityName="Zhao &amp; Liu" authorityYear="2022" baseAuthorityName="Zhao &amp; Liu" baseAuthorityYear="2022" class="Insecta" family="Scarabaeidae" genus="Fruhstorferia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fruhstorferia birmanica" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis id="AEC5CAA76D6C69CAFD961EF1EFC37E08" italics="true" pageId="0" pageNumber="31">Fruhstorferia birmanica</emphasis>
</taxonomicName>
Arrow, 1907)
<emphasis bold="true" pageId="0" pageNumber="31">16-18</emphasis>
<emphasis id="C945897A593C15B10304306E68B90EFB" bold="true" pageId="0" pageNumber="31">16-18</emphasis>
female of
<taxonomicName lsidName="D. mizunumai" pageId="0" pageNumber="31" rank="species" species="mizunumai">
<emphasis italics="true" pageId="0" pageNumber="31">D. mizunumai</emphasis>
<taxonomicName id="B555D2FE089E670059C090161D7CE971" lsidName="D. mizunumai" pageId="0" pageNumber="31" rank="species" species="mizunumai">
<emphasis id="3076587E6DDF6BE720F9AD6B1C537794" italics="true" pageId="0" pageNumber="31">D. mizunumai</emphasis>
</taxonomicName>
Nagai &amp; Hirasawa, 1991 (paralectotype of
<taxonomicName lsidName="F. birmanica" pageId="0" pageNumber="31" rank="species" species="birmanica">
<emphasis italics="true" pageId="0" pageNumber="31">F. birmanica</emphasis>
<taxonomicName id="FAC4BE9812ADE01086A1C12EEF51FFB2" lsidName="F. birmanica" pageId="0" pageNumber="31" rank="species" species="birmanica">
<emphasis id="B7015EC21DD30236796B0A3DE9301CE1" italics="true" pageId="0" pageNumber="31">F. birmanica</emphasis>
</taxonomicName>
), habitus in dorsal view
<emphasis bold="true" pageId="0" pageNumber="31">11, 16</emphasis>
<emphasis id="43BFE7A22FED12AB0BCB2689E13736B9" bold="true" pageId="0" pageNumber="31">11, 16</emphasis>
habitus in dorsal view
<emphasis bold="true" pageId="0" pageNumber="31">12, 17</emphasis>
<emphasis id="273638A2A1BFAB9D8783A5A438E191E5" bold="true" pageId="0" pageNumber="31">12, 17</emphasis>
habitus in lateral view
<emphasis bold="true" pageId="0" pageNumber="31">13</emphasis>
<emphasis id="7CF5DD4CA4A419BBB7B9E69F61459444" bold="true" pageId="0" pageNumber="31">13</emphasis>
aedeagus showing parameres in dorsal view
<emphasis bold="true" pageId="0" pageNumber="31">14</emphasis>
<emphasis id="C38D6F5BD8D6756D692AF4C727D266E4" bold="true" pageId="0" pageNumber="31">14</emphasis>
aedeagus in dorsal view
<emphasis bold="true" pageId="0" pageNumber="31">15, 18</emphasis>
<emphasis id="F7E9F8307CF3BAC7477169236A0F4632" bold="true" pageId="0" pageNumber="31">15, 18</emphasis>
attached labels. All photos © Keita Matsumoto (NHMUK).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="distribution">
<paragraph pageId="0" pageNumber="31">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="31">Laos (Houaphanh); Myanmar (Mandalay) (new record); Thailand (Chiang Mai); Vietnam (Lao Cai).</paragraph>
<subSubSection id="11B889E230BF101F7B0277F8D35F11DE" pageId="0" pageNumber="31" type="distribution">
<paragraph id="6C605FB4D07D2A05176EE56377B823BD" pageId="0" pageNumber="31">Distribution.</paragraph>
<paragraph id="2C67EF52BA6C80642FA52F89EB6B5B84" pageId="0" pageNumber="31">Laos (Houaphanh); Myanmar (Mandalay) (new record); Thailand (Chiang Mai); Vietnam (Lao Cai).</paragraph>
</subSubSection>
</treatment>
</subSection>

98
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@ -1,76 +1,76 @@
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Genus
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<emphasis italics="true" pageId="0" pageNumber="115">Type species.</emphasis>
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<emphasis id="9996E17535FE464C5EEFA3C08DF122FD" italics="true" pageId="0" pageNumber="115">Type species.</emphasis>
-
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<taxonomicName class="Insecta" family="Tettigoniidae" genus="Acosmetura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acosmetura brevicerca" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="brevicerca">
<emphasis italics="true" pageId="0" pageNumber="115">Acosmetura brevicerca</emphasis>
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<taxonomicName id="96714901299E9DACF703E688E1552C32" class="Insecta" family="Tettigoniidae" genus="Acosmetura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acosmetura brevicerca" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="brevicerca">
<emphasis id="2C986814BE56546507D29144134A724A" italics="true" pageId="0" pageNumber="115">Acosmetura brevicerca</emphasis>
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Liu, 2000
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<emphasis italics="true" pageId="0" pageNumber="115">Diagnosis.</emphasis>
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<emphasis id="2175C0116E02A3F31D2E4E161A921AC5" italics="true" pageId="0" pageNumber="115">Diagnosis.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="115">The genus is characterized by the male tenth abdominal tergite with a shallow concavity on posterior margin; subgenital plate longer than wide; genitalia sclerotized, exposed, not surpassing the posterior margin of subgenital plate. Margins of ovipositor smooth or with small teeth.</paragraph>
<paragraph id="3FB811B0779792568051F69E0D62EF9D" pageId="0" pageNumber="115">The genus is characterized by the male tenth abdominal tergite with a shallow concavity on posterior margin; subgenital plate longer than wide; genitalia sclerotized, exposed, not surpassing the posterior margin of subgenital plate. Margins of ovipositor smooth or with small teeth.</paragraph>
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