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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1070.67779" ID-GBIF-Dataset="5b0e7362-5db2-437e-8780-ad3159c07270" ID-PMC="PMC8602214" ID-Pensoft-Pub="1313-2970-1070-87" ID-Pensoft-UUID="BF4B36D13F115A5480A28F7D0B72D8DF" ID-PubMed="34819773" ID-ZooBank="87CD8309552D43BCB8704184B09311A1" ModsDocID="1313-2970-1070-87" checkinTime="1636677281775" checkinUser="pensoft" docAuthor="Shao, Jiaqi &amp; Kang, Zehui" docDate="2021" docId="0728CE901AD65B549A1E764099AB5C0E" docLanguage="en" docName="ZooKeys 1070: 87-99" docOrigin="ZooKeys 1070" docPubDate="2021-11-11" docSource="http://dx.doi.org/10.3897/zookeys.1070.67779" docTitle="Ptychoptera longwangshana Yang &amp; Chen 1998" docType="treatment" docVersion="4" id="BF4B36D13F115A5480A28F7D0B72D8DF" lastPageNumber="87" masterDocId="BF4B36D13F115A5480A28F7D0B72D8DF" masterDocTitle="New species of the genus Ptychoptera Meigen, 1803 (Diptera, Ptychopteridae) from Zhejiang, China with an updated key to Chinese species" masterLastPageNumber="99" masterPageNumber="87" pageNumber="87" updateTime="1668151120232" updateUser="ExternalLinkService">
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<mods:title>New species of the genus Ptychoptera Meigen, 1803 (Diptera, Ptychopteridae) from Zhejiang, China with an updated key to Chinese species</mods:title>
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<mods:title id="F03ADBE0A064AC6CF63E578B17BA3DEA">New species of the genus Ptychoptera Meigen, 1803 (Diptera, Ptychopteridae) from Zhejiang, China with an updated key to Chinese species</mods:title>
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<mods:namePart>Shao, Jiaqi</mods:namePart>
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<mods:namePart>Kang, Zehui</mods:namePart>
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<treatment ID-GBIF-Taxon="190232602" LSID="urn:lsid:plazi:treatment:0728CE901AD65B549A1E764099AB5C0E" httpUri="http://treatment.plazi.org/id/0728CE901AD65B549A1E764099AB5C0E" lastPageNumber="87" pageId="0" pageNumber="87">
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<taxonomicName LSID="0728CE90-1AD6-5B54-9A1E-764099AB5C0E" authority="Yang &amp; Chen, 1998" authorityName="Yang &amp; Chen" authorityYear="1998" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera longwangshana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="longwangshana">Ptychoptera longwangshana Yang &amp; Chen, 1998</taxonomicName>
<treatment id="0728CE901AD65B549A1E764099AB5C0E" ID-GBIF-Taxon="190232602" LSID="urn:lsid:plazi:treatment:0728CE901AD65B549A1E764099AB5C0E" httpUri="http://treatment.plazi.org/id/0728CE901AD65B549A1E764099AB5C0E" lastPageNumber="87" pageId="0" pageNumber="87" scope_family="Ptychopteridae" scope_order="Diptera">
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<paragraph id="D680825050FE3060E0847F48713E90AF" pageId="0" pageNumber="87">
<taxonomicName id="E7C646A929F5D1DFF9640A353F21D788" ID-CoL="6WMTS" LSID="0728CE90-1AD6-5B54-9A1E-764099AB5C0E" authority="Yang &amp; Chen, 1998" authorityName="Yang &amp; Chen" authorityYear="1998" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera longwangshana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="longwangshana">Ptychoptera longwangshana Yang &amp; Chen, 1998</taxonomicName>
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<subSubSection pageId="0" pageNumber="87" type="description">
<paragraph pageId="0" pageNumber="87">
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Ptychoptera longwangshana Yang &amp; Chen, 1998 and Ptychoptera gutianshana Yang &amp; Chen, 1995 a wing of P. longwangshana b-f P. gutianshana (b wing c male genitalia, lateral view d epandrium, dorsal view e gonocoxite and gonostylus, dorsal view f hypandrium, ventral view). Scale bars: 1.0 mm (a, b); 0.2 mm (c-f)." figureDoi="10.3897/zookeys.1070.67779.figure5" httpUri="https://binary.pensoft.net/fig/609851" pageId="0" pageNumber="87">Figure 5a</figureCitation>
<subSubSection id="AA87387298CBB0CF215C31FD5572CFF0" pageId="0" pageNumber="87" type="description">
<paragraph id="175C7ECF0DD50EA3E4CE4ED3C087028F" pageId="0" pageNumber="87">
<figureCitation id="4918C740EBA2CFF010A4669734F34579" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Ptychoptera longwangshana Yang &amp; Chen, 1998 and Ptychoptera gutianshana Yang &amp; Chen, 1995 a wing of P. longwangshana b-f P. gutianshana (b wing c male genitalia, lateral view d epandrium, dorsal view e gonocoxite and gonostylus, dorsal view f hypandrium, ventral view). Scale bars: 1.0 mm (a, b); 0.2 mm (c-f)." figureDoi="10.3897/zookeys.1070.67779.figure5" httpUri="https://binary.pensoft.net/fig/609851" pageId="0" pageNumber="87">Figure 5a</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="87" type="reference_group">
<paragraph pageId="0" pageNumber="87">
<taxonomicName authorityName="Yang &amp; Chen" authorityYear="1998" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera longwangshana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="longwangshana">Ptychoptera longwangshana</taxonomicName>
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<paragraph id="CEB5BC759D261DF97648DF46E9300FCE" pageId="0" pageNumber="87">
<taxonomicName id="92D601D3B93C8A8A65C2CDE7DE7ABEAF" authorityName="Yang &amp; Chen" authorityYear="1998" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera longwangshana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="longwangshana">Ptychoptera longwangshana</taxonomicName>
Yang &amp; Chen, 1998: 240. Type locality: Mount Longwangshan, Anji, Zhejiang (China).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="materials_examined">
<paragraph pageId="0" pageNumber="87">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="87">
<materialsCitation collectingDate="1996-06-12" country="China" location="Mount Longwangshan" municipality="Anji District" specimenCount="1" stateProvince="Zhejiang Province" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="87">
<typeStatus>Holotype</typeStatus>
<subSubSection id="AF2D87946CC99388B3CA09353FBF3C2E" pageId="0" pageNumber="87" type="materials_examined">
<paragraph id="4B9B63EBE78F7AFA075AE0C775F5E87C" pageId="0" pageNumber="87">Specimens examined.</paragraph>
<paragraph id="B71B6A750580C8DEDBA299E0184CFA3E" pageId="0" pageNumber="87">
<materialsCitation id="8FC263843FAE0FCFB50D72289398DDF4" collectingDate="1996-06-12" country="China" location="Mount Longwangshan" municipality="Anji District" specimenCount="1" stateProvince="Zhejiang Province" typeStatus="Holotype">
<emphasis id="56C374BE9AA0CA6F09A8D9A9C5817DD1" bold="true" italics="true" pageId="0" pageNumber="87">
<typeStatus id="EA6B3E59B246DB750F5A2864D552565C">Holotype</typeStatus>
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male (CAU),
<collectingCountry name="China">China</collectingCountry>
<collectingCountry id="0C1B3FF9D45B16E6FE5509F983E90441" name="China">China</collectingCountry>
:
<collectingRegion country="China" name="Zhejiang">Zhejiang Province</collectingRegion>
<collectingRegion id="00D4DF9C6B015DBDBB0FA7E12363AFCC" country="China" name="Zhejiang">Zhejiang Province</collectingRegion>
,
<collectingMunicipality>Anji District</collectingMunicipality>
<collectingMunicipality id="D2676CC91C5E8C29ADA9B0E6A57FDE2E">Anji District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:0728CE901AD65B549A1E764099AB5C0E:44183BF133FD7D226465E94F89FBB2BB" country="China" municipality="Anji District" name="Mount Longwangshan" stateProvince="Zhejiang Province">Mount Longwangshan</location>
<location id="44183BF133FD7D226465E94F89FBB2BB" LSID="urn:lsid:plazi:treatment:0728CE901AD65B549A1E764099AB5C0E:44183BF133FD7D226465E94F89FBB2BB" country="China" municipality="Anji District" name="Mount Longwangshan" stateProvince="Zhejiang Province">Mount Longwangshan</location>
,
<collectingDate value="1996-06-12">1996.VI.12</collectingDate>
<collectingDate id="3DB0EECE7B9F5495E4B2C73993911525" value="1996-06-12">1996.VI.12</collectingDate>
, Chikun Yang.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="diagnosis">
<paragraph pageId="0" pageNumber="87">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="87">Thorax mostly black. Wing marked with two brown marks and two brown bands. Surstylus of epandrium tapering and curved downward distally, curving up at tip. Gonocoxite wide and gonostylus short and small. Apical part of hypandrium with a pair of curved up and trough-shaped projections.</paragraph>
<subSubSection id="A70F0362E6208D5F0D9E8729DE864810" pageId="0" pageNumber="87" type="diagnosis">
<paragraph id="8739F95E162293D3E0492582C6B5DE63" pageId="0" pageNumber="87">Diagnosis.</paragraph>
<paragraph id="98A1471E4AF8A7DC6DD1440826252440" pageId="0" pageNumber="87">Thorax mostly black. Wing marked with two brown marks and two brown bands. Surstylus of epandrium tapering and curved downward distally, curving up at tip. Gonocoxite wide and gonostylus short and small. Apical part of hypandrium with a pair of curved up and trough-shaped projections.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="description">
<paragraph pageId="0" pageNumber="87">Description.</paragraph>
<paragraph pageId="0" pageNumber="87">
<subSubSection id="47B9CB1BDDE114C8F308D8385E18AB9B" pageId="0" pageNumber="87" type="description">
<paragraph id="65E89E7DA582E8113B467842B0047AFD" pageId="0" pageNumber="87">Description.</paragraph>
<paragraph id="597E2C950C382E7555B2833D397A5CE4" pageId="0" pageNumber="87">
Wing length 8.0 mm. Wing (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Ptychoptera longwangshana Yang &amp; Chen, 1998 and Ptychoptera gutianshana Yang &amp; Chen, 1995 a wing of P. longwangshana b-f P. gutianshana (b wing c male genitalia, lateral view d epandrium, dorsal view e gonocoxite and gonostylus, dorsal view f hypandrium, ventral view). Scale bars: 1.0 mm (a, b); 0.2 mm (c-f)." figureDoi="10.3897/zookeys.1070.67779.figure5" httpUri="https://binary.pensoft.net/fig/609851" pageId="0" pageNumber="87">5a</figureCitation>
<figureCitation id="9630DDAAEDFEE47DC012EC1637007856" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Ptychoptera longwangshana Yang &amp; Chen, 1998 and Ptychoptera gutianshana Yang &amp; Chen, 1995 a wing of P. longwangshana b-f P. gutianshana (b wing c male genitalia, lateral view d epandrium, dorsal view e gonocoxite and gonostylus, dorsal view f hypandrium, ventral view). Scale bars: 1.0 mm (a, b); 0.2 mm (c-f)." figureDoi="10.3897/zookeys.1070.67779.figure5" httpUri="https://binary.pensoft.net/fig/609851" pageId="0" pageNumber="87">5a</figureCitation>
) 3.2 times as long as wide, subhyaline, marked with two brown marks and two brown bands as follows: an oval brown mark at base of R, a triangular brown mark at base of Rs; median band extending from anterior margin of wing, covering base of R2+3 and r-m, to middle section of CuA; subapical band extending from anterior margin of wing, covering tip of R1 and R2, to M1+2 fork, slightly separated into two marks. Veins brown; Sc ending in C at level of basal third of R2+3; Rs straight, 3 times the length of r-m.
</paragraph>
<caption doi="10.3897/zookeys.1070.67779.figure5" httpUri="https://binary.pensoft.net/fig/609851" pageId="0" pageNumber="87" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="87">
<emphasis bold="true" pageId="0" pageNumber="87">Figure 5.</emphasis>
<taxonomicName authorityName="Yang &amp; Chen" authorityYear="1998" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera longwangshana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="longwangshana">
<emphasis italics="true" pageId="0" pageNumber="87">Ptychoptera longwangshana</emphasis>
<caption id="63EB3346406817BD320024BDEF86468A" doi="10.3897/zookeys.1070.67779.figure5" httpUri="https://binary.pensoft.net/fig/609851" pageId="0" pageNumber="87" start="Figure 5" startId="F5">
<paragraph id="E726DDBF2A484C53CF48D08B9E6F0601" pageId="0" pageNumber="87">
<emphasis id="01D72CE5BF74FA89C62983216FBB17DD" bold="true" pageId="0" pageNumber="87">Figure 5.</emphasis>
<taxonomicName id="B217DA8F862D2A72FF9FF5B045FCD1E6" authorityName="Yang &amp; Chen" authorityYear="1998" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera longwangshana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="longwangshana">
<emphasis id="93C528AFCCB8C6EC9E429FD5E495A895" italics="true" pageId="0" pageNumber="87">Ptychoptera longwangshana</emphasis>
</taxonomicName>
Yang &amp; Chen, 1998 and
<taxonomicName authorityName="Yang &amp; Chen" authorityYear="1995" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera gutianshana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="gutianshana">
<emphasis italics="true" pageId="0" pageNumber="87">Ptychoptera gutianshana</emphasis>
<taxonomicName id="1BFC67B18C6C51A005F9E4406B3F4115" authorityName="Yang &amp; Chen" authorityYear="1995" class="Insecta" family="Ptychopteridae" genus="Ptychoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoptera gutianshana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="gutianshana">
<emphasis id="077907E0FAF93A4F6C99914BA18395DE" italics="true" pageId="0" pageNumber="87">Ptychoptera gutianshana</emphasis>
</taxonomicName>
Yang &amp; Chen, 1995
<emphasis bold="true" pageId="0" pageNumber="87">a</emphasis>
<emphasis id="704608FF41C4A6114BDC15CD55092CC3" bold="true" pageId="0" pageNumber="87">a</emphasis>
wing of
<taxonomicName lsidName="P. longwangshana" pageId="0" pageNumber="87" rank="species" species="longwangshana">
<emphasis italics="true" pageId="0" pageNumber="87">P. longwangshana</emphasis>
<taxonomicName id="086AC2341E4C59F03723459ECF7FF735" lsidName="P. longwangshana" pageId="0" pageNumber="87" rank="species" species="longwangshana">
<emphasis id="278B908BEC94FD53518443FCA06232AB" italics="true" pageId="0" pageNumber="87">P. longwangshana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="87">b-f</emphasis>
<taxonomicName lsidName="P. gutianshana" pageId="0" pageNumber="87" rank="species" species="gutianshana">
<emphasis italics="true" pageId="0" pageNumber="87">P. gutianshana</emphasis>
<emphasis id="5F162CAE78865BF62A28735AF8F8F2B3" bold="true" pageId="0" pageNumber="87">b-f</emphasis>
<taxonomicName id="4A6A139328BABAA761503314A6F18953" lsidName="P. gutianshana" pageId="0" pageNumber="87" rank="species" species="gutianshana">
<emphasis id="85D7B55C6445BBA72729D11C42AEB5F6" italics="true" pageId="0" pageNumber="87">P. gutianshana</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="87">b</emphasis>
<emphasis id="A112FAC4C8C41783C02DACD4A1F8631D" bold="true" pageId="0" pageNumber="87">b</emphasis>
wing
<emphasis bold="true" pageId="0" pageNumber="87">c</emphasis>
<emphasis id="8567C1BC9FF39F0829954EDD1FE73865" bold="true" pageId="0" pageNumber="87">c</emphasis>
male genitalia, lateral view
<emphasis bold="true" pageId="0" pageNumber="87">d</emphasis>
<emphasis id="E43F645B177CCBF0F16D199021A8F8F3" bold="true" pageId="0" pageNumber="87">d</emphasis>
epandrium, dorsal view
<emphasis bold="true" pageId="0" pageNumber="87">e</emphasis>
<emphasis id="6A532E7127BBE95B498BAC87CB66FF8F" bold="true" pageId="0" pageNumber="87">e</emphasis>
gonocoxite and gonostylus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="87">f</emphasis>
<emphasis id="3833888D84ACA3E6032600C4F1FA3011" bold="true" pageId="0" pageNumber="87">f</emphasis>
hypandrium, ventral view). Scale bars: 1.0 mm (
<emphasis bold="true" pageId="0" pageNumber="87">a, b</emphasis>
<emphasis id="6FECA6A813A31C697331F441ECEF927A" bold="true" pageId="0" pageNumber="87">a, b</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="87">c-f</emphasis>
<emphasis id="E7B2793445DA6F12F82D00CD68412D96" bold="true" pageId="0" pageNumber="87">c-f</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="distribution">
<paragraph pageId="0" pageNumber="87">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="87">China (Zhejiang).</paragraph>
<subSubSection id="A18B3449B316A38570A8C298A549678A" pageId="0" pageNumber="87" type="distribution">
<paragraph id="D85ACBC59A2C181AE5DC71E64897E3DF" pageId="0" pageNumber="87">Distribution.</paragraph>
<paragraph id="6ADCF6A53F1FFF51EA199AB608D56FEB" pageId="0" pageNumber="87">China (Zhejiang).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="remarks">
<paragraph pageId="0" pageNumber="87">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="87">
<subSubSection id="2D66A6E8791F7413AEF82C7350CC0FAD" pageId="0" pageNumber="87" type="remarks">
<paragraph id="337066B024AEFE63A37B72E32D35A67C" pageId="0" pageNumber="87">Remarks.</paragraph>
<paragraph id="7248B691989E1F8E7E98A8F7AD01081C" pageId="0" pageNumber="87">
Only one wing of the holotype was available during the study, while the rest of the holotype was not found in CAU. For a description and illustration of this species, also see
<bibRefCitation author="Yang, J" editor="Wu, H" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="0" pageNumber="87" pagination="240 - 241" refId="B18" refString="Yang, J, Chen, H, 1998. Diptera: Ptychopteridae. In: Wu, H, Ed., Insects of Longwangshan Nature Reserve. China Forestry Publishing House, Beijing: 240 - 241" title="Diptera: Ptychopteridae." volumeTitle="Insects of Longwangshan Nature Reserve." year="1998">Yang and Chen (1998)</bibRefCitation>
<bibRefCitation id="36845C2BAEC4A1FCF6DA9D33311A7816" author="Yang, J" editor="Wu, H" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="0" pageNumber="87" pagination="240 - 241" refId="B18" refString="Yang, J, Chen, H, 1998. Diptera: Ptychopteridae. In: Wu, H, Ed., Insects of Longwangshan Nature Reserve. China Forestry Publishing House, Beijing: 240 - 241" title="Diptera: Ptychopteridae." volumeTitle="Insects of Longwangshan Nature Reserve." year="1998">Yang and Chen (1998)</bibRefCitation>
.
</paragraph>
</subSubSection>