Add updates up until 2024-09-20 17:26:40
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data/41/D4/55/41D455A5216C50D0931DAAAD1607ABA8.xml
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<document id="F744254E354DF9396B60297DE34A52FF" ID-DOI="10.3897/vz.74.e130147" ID-ZooBank="9C1F5BF9-B733-463C-B085-792EE730EEFB" ID-publisher-id="130147" URI-arpha="5112DC7F-8C35-5534-A208-D8A08397FA88" URI-zoobank="http://zoobank.org/9C1F5BF9-B733-463C-B085-792EE730EEFB" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="research-article" checkinTime="1726844247928" checkinUser="pensoft" docAuthor="Torres-Carvajal, Omar, Sandoval, Camila & Paucar, Diego A." docDate="2024" docId="41D455A5216C50D0931DAAAD1607ABA8" docLanguage="en" docName="Vertebrate Zoology 74: 551-564" docOrigin="Vertebrate Zoology 74" docSource="https://vertebrate-zoology.arphahub.com/article/130147/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Mabuya yasuniensis Torres-Carvajal, Sandoval & Paucar, 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="564" masterDocId="5112DC7F8C355534A208D8A08397FA88" masterDocTitle="The skinks (Squamata: Scincidae) of Ecuador, with description of a new Amazonian species" masterLastPageNumber="564" masterPageNumber="551" pageNumber="551" updateTime="1726853198118" updateUser="admin">
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<mods:mods id="53C7D120D1D5E96C3F3C568054DC1CB6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A4F905AE143A21F8AF61DFCC9E56921C">
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<mods:title id="D44BB085F3C97B390D2BAEBCFB3BCF37">The skinks (Squamata: Scincidae) of Ecuador, with description of a new Amazonian species</mods:title>
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</mods:titleInfo>
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<mods:name id="A0FD28006C1D712EBB0B6F9FB9CDB0D1" type="personal">
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</mods:role>
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<mods:namePart id="278A3471799DA558A11D196F51E09D85">Torres-Carvajal, Omar</mods:namePart>
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<mods:nameIdentifier id="4CAA1781ED7109F0C7107FC80CB94584" type="ORCID">0000-0003-0041-9250</mods:nameIdentifier>
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<mods:affiliation id="D905C4CE2441B3B22FEC38B55CADCA94">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Quito, Ecuador</mods:affiliation>
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</mods:name>
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<mods:name id="02F3707C828C45438170316E60486963" type="personal">
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<mods:roleTerm id="7313FD74D6403ECB07FC247A6BC5BE05">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BDEB922F0A0758BC34B0B8530C1D5D10">Sandoval, Camila</mods:namePart>
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<mods:nameIdentifier id="3665F8161DE7F3650B72F70A98220F37" type="ORCID">https://orcid.org/0009-0004-1909-5211</mods:nameIdentifier>
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<mods:affiliation id="1A3A9881E92D3D713DB76C99410A8981">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Quito, Ecuador</mods:affiliation>
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<mods:name id="6E8EAB84D46BDEEBD3A4C7A84228298B" type="personal">
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<mods:roleTerm id="B2BF872ED8462A0F4FAF7B5044021AF0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BF835A3DD292254AE57F333FF32D72E0">Paucar, Diego A.</mods:namePart>
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||||
<mods:affiliation id="1E06D85E3C184006D368E17C37AD045C">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Quito, Ecuador</mods:affiliation>
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<mods:title id="2E2B35FE013246B46E87E93F005A2BC7">Vertebrate Zoology</mods:title>
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</mods:titleInfo>
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<mods:part id="C5C1B5F84815569025652531510961FA">
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<mods:date id="5F473A103B174F18FC17C9810ECD9CE2">2024</mods:date>
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<mods:detail id="66FF0B58ABFB9DD9FBEF4AF4C046CCCF" type="pubDate">
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<mods:number id="E3A774F399E9A3CDC56F0347A50D571D">2024-09-20</mods:number>
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<mods:detail id="9E7110F8C2E55CED7202F5FB7C06B4B4" type="volume">
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<mods:number id="B993F10EC0CCE1BC36330CCE8DDCC73D">74</mods:number>
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<mods:start id="E64AB4AF4A39C6C0BE10EBD8C51D95AE">551</mods:start>
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<mods:classification id="67645685D713CA240711775AE47D5BB5">journal article</mods:classification>
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<mods:identifier id="34454C853DB8AB9CFBFB4B7C16EFD219" type="DOI">10.3897/vz.74.e130147</mods:identifier>
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<mods:identifier id="8D43E0CD586A04ECA1309F19510351BE" type="ZooBank">9C1F5BF9-B733-463C-B085-792EE730EEFB</mods:identifier>
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</mods:mods>
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<treatment id="41D455A5216C50D0931DAAAD1607ABA8" ID-arpha="41D455A5-216C-50D0-931D-AAAD1607ABA8" ID-zoobank="https://zoobank.org/41A31B4B-EC6F-42DC-9D40-8DDE5D22B00E" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8" LSID-ZBK="urn:lsid:zoobank.org:act:41A31B4B-EC6F-42DC-9D40-8DDE5D22B00E" httpUri="http://treatment.plazi.org/id/41D455A5216C50D0931DAAAD1607ABA8">
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<subSubSection id="03DF7504BAE7CF3C11462C17EA3EBAD7" type="nomenclature">
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<paragraph id="6E7D58E327C149FB1377706C57FA82F9">
|
||||
<taxonomicName id="142DAAD61DDAAD7B497069853CE5380E" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis" status="sp. nov.">
|
||||
<emphasis id="6BC7FC7E89357812A15BE8CC5450E135" italics="true">Mabuya yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="6B6F9A46CCFB3B5CB2FD2C524953C02C" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EXABG" type="Holotype">
|
||||
<paragraph id="3F5FA41B0428ACF63FF65757D716E489">
|
||||
<heading id="1EF79B71DE89F706DDE1C8A92316C22A" reason="title">
|
||||
<typeStatus id="19C97C196EB03281F697807EAFDF3252" type="holotype">Holotype</typeStatus>
|
||||
.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="1E3361AC660182DE772EEE8417828CAE">
|
||||
<materialsCitation id="B3DB312A7E25BFC40B9CBCBF86F9AC6F" collectingDate="2016-07-29" collectionCode="QCAZ" collectorName="Ruben Jarrin" country="Ecuador" county="Yasuni River" elevation="190" latitude="-0.967" location="lagoon close to Yasuni River" longLatPrecision="78" longitude="-75.875" municipality="Yasuni National Park" specimenCode="QCAZ 14814" specimenCount="1" specimenCount-male="1" stateProvince="Orellana Province" typeStatus="holotype">
|
||||
An
|
||||
<specimenCount id="C93A981937ED261C5180BDEF840F0298" count="1" type="male">adult male</specimenCount>
|
||||
,
|
||||
<specimenCode id="2BC696101657ED6A2EE59DD8599B1085" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">
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<named-content id="442F061A6E8EBF39F6FBB95E9D94BFEB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
14814
|
||||
</specimenCode>
|
||||
(Figs
|
||||
<figureCitation id="AE9DDDDF390C410C37AA4DE8F91E8E2C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Four species of skinks Mabuya in life, from top to bottom: M. yasuniensis sp. nov. (QCAZ 14814, holotype, SVL = 72.98 mm), M. altamazonica (QCAZ 13087, SVL = 92.11 mm), M. nigropunctata (QCAZ 8107, SVL = 104.58 mm), and M. bistriata. Photographs from Bioweb. bio, except M. bistriata (Laurie J. Vitt)." figureDoi="10.3897/vz.74.e130147.figure2" httpUri="https://binary.pensoft.net/fig/1137914">2</figureCitation>
|
||||
,
|
||||
<figureCitation id="1B2FAD7A744359E8D48A39B0ADCD1111" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male holotype (QCAZ 14814; SVL = 72.98 mm) of Mabuya yasuniensis sp. nov. in dorsal (top), lateral (middle) and ventral (bottom) views. Photographs by D. A. Paucar. Scale bar: 10 mm." figureDoi="10.3897/vz.74.e130147.figure4" httpUri="https://binary.pensoft.net/fig/1137916">4</figureCitation>
|
||||
,
|
||||
<figureCitation id="0EBFB1C317CF458C39768E906B67956A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Head of the holotype (QCAZ 14814) of Mabuya yasuniensis sp. nov. in dorsal (top), ventral (middle), and lateral (bottom) views. Photographs by D. A. Paucar. Scale bar: 5 mm." figureDoi="10.3897/vz.74.e130147.figure5" httpUri="https://binary.pensoft.net/fig/1137917">5</figureCitation>
|
||||
), field number SC 55292, collected on
|
||||
<date id="8B643E3B9EE4D826B20CF5C2B13E56BB" value="2016-07-29">
|
||||
<collectingDate id="A22F51C6C47C4A32AD27456BEEF940FD" value="2016-07-29">29 July 2016</collectingDate>
|
||||
</date>
|
||||
at 1630 h by
|
||||
<collectorName id="395D1DE12059F71D622B2791E9D23A47">Rubén Jarrín</collectorName>
|
||||
at
|
||||
<collectingMunicipality id="237EF0A018307C7F80CE8BAE088BACD0">Yasuní National Park</collectingMunicipality>
|
||||
in a
|
||||
<location id="36247848109C5BB9B70453264B277133" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:36247848109C5BB9B70453264B277133" country="Ecuador" county="Yasuni River" latitude="-0.967" longLatPrecision="78" longitude="-75.875" municipality="Yasuni National Park" name="lagoon close to Yasuni River" stateProvince="Orellana Province">lagoon close to Yasuní River</location>
|
||||
,
|
||||
<locationDeviation id="134CEC031AACC87F7879C463ED6542B4">
|
||||
<quantity id="B32EEF0DBBEB6F82FFAFB23014140BFC" metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
|
||||
S Rumiyacu River mouth
|
||||
</locationDeviation>
|
||||
(
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQBBG" specific-use="{"type":"Point","coordinates":[-75.875000,-0.967000]}">
|
||||
<geoCoordinate id="9596CEFCC47DC19EFC241660209B09CC" degrees="0.967" direction="south" orientation="latitude" precision="55" value="-0.967">0.967 ° S</geoCoordinate>
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,
|
||||
<geoCoordinate id="42607784916381F3FA555BD7DB44C03C" degrees="75.875" direction="west" orientation="longitude" precision="55" value="-75.875">75.875 ° W</geoCoordinate>
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</named-content>
|
||||
,
|
||||
<quantity id="86FD0901A8CCAD4E2110D1C2783736BD" metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">
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||||
<elevation id="FF49FA81C9FB18B33D65AF9C78F8F1BA" metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">190 m</elevation>
|
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</quantity>
|
||||
),
|
||||
<collectingRegion id="6257E75F6C23D6AC764DA5A41F1B5212" country="Ecuador" name="Orellana">Orellana Province</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="97F1095A9923985DF917750AEAE4A543" name="Ecuador">Ecuador</collectingCountry>
|
||||
.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
<caption id="F5386EEBD1EE311BE34B74FA3CF64788" ID-DOI="10.3897/vz.74.e130147.figure4" ID-arpha="C734701E-5BFC-5F57-8008-A8EBC28A7E26" httpUri="https://binary.pensoft.net/fig/1137916" startId="F4">
|
||||
<paragraph id="CFA29BB905C2404D8E99736940BC560E">
|
||||
<label id="FCE7022050A1A542EDFDA3AC76E27774">Figure 4.</label>
|
||||
</paragraph>
|
||||
<paragraph id="D4496F62B87B4A4DBC973F69EEC7841A">
|
||||
Male holotype (
|
||||
<specimenCode id="8A7BCD291AF9FC7F85F306AF18810BFC">
|
||||
<named-content id="A247BBDC50B641A070023197EA97B5EA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
14814
|
||||
</specimenCode>
|
||||
;
|
||||
<abbrev id="ABBRID0EBCBG" xlink_title="snout-vent length">SVL</abbrev>
|
||||
= 72.98 mm) of
|
||||
<taxonomicName id="4CE2A31F774C158E2296ED31E61622E1" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis" status="sp. nov.">
|
||||
<emphasis id="0BA07E3FECC0F8F141678A18FF948912" italics="true">Mabuya yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="41F55501EC0A7EAF818C58BEEE7A6A7D" bold="true">
|
||||
<taxonomicNameLabel id="583D7B574DB8493C95F4505DB7AC9AC3" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
in dorsal (top), lateral (middle) and ventral (bottom) views. Photographs by D. A. Paucar. Scale bar: 10 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EXCBG" type="Paratypes (13)">
|
||||
<paragraph id="307BEB99F3E34546E46835A8CD77C66E">
|
||||
<heading id="2D5E1260B1D7D049FA614850CFB892BA" reason="title">
|
||||
<typeStatus id="56DE18D28F87D7D2A5FAF5A497779F84" type="paratype">Paratypes</typeStatus>
|
||||
(13).
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="E39D3F196284626AB3AFD34422121D1D">
|
||||
<materialsCitation id="26E82B7E0BEEAF10F0425D4EE42E8435" collectionCode="QCAZ" country="Ecuador" elevation="220" latitude="-0.46" location="500 m from Napo River" longLatPrecision="78" longitude="-76.135" municipality="Oasis community" specimenCode="QCAZ 10740" specimenCount="1" stateProvince="Orellana" typeStatus="paratype">
|
||||
<collectingCountry id="756768B4D91F9089DA023C0F55108AF7" name="Ecuador">ECUADOR</collectingCountry>
|
||||
:
|
||||
<underline id="123DB207FBE26631A1B54E8B6B157BD0">
|
||||
<collectingRegion id="3029CA578ED8415CEEBA3B8B67B78842" country="Ecuador" name="Orellana">Orellana</collectingRegion>
|
||||
</underline>
|
||||
:
|
||||
<collectingMunicipality id="9DD76BC9A6AB02487C4F76EECBCBDF05">Oasis community</collectingMunicipality>
|
||||
,
|
||||
<location id="25C7F22EC4F73126903C391BD3E95213" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:25C7F22EC4F73126903C391BD3E95213" country="Ecuador" latitude="-0.46" longLatPrecision="78" longitude="-76.135" municipality="Oasis community" name="500 m from Napo River" stateProvince="Orellana">
|
||||
<quantity id="9F55AB9C5DB6D563499E25557716B1AA" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</quantity>
|
||||
from Napo River
|
||||
</location>
|
||||
,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECDBG" specific-use="{"type":"Point","coordinates":[-76.135000,-0.460000]}">
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<geoCoordinate id="16EEAFD9E5C0C06A8555B6CCE96DEE05" degrees="0.460" direction="south" orientation="latitude" precision="55" value="-0.46">0.460 ° S</geoCoordinate>
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,
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<geoCoordinate id="B4A7CA0D7F0CB6E83F8DF5F33C9AC63D" degrees="76.135" direction="west" orientation="longitude" precision="55" value="-76.135">76.135 ° W</geoCoordinate>
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</named-content>
|
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,
|
||||
<quantity id="C223A741F0F3EC371DC90E33EA234319" metricMagnitude="2" metricUnit="m" metricValue="2.2" unit="m" value="220.0">
|
||||
<elevation id="14CA5E0DA801463F7AA3DBE529E80DDE" metricMagnitude="2" metricUnit="m" metricValue="2.2" unit="m" value="220.0">220 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<specimenCode id="FFA0E0C90229B14DB999E9BDA76BDB1F">
|
||||
<named-content id="41AACBCF53949A8F43332A957FA9DE76" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
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10740
|
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</specimenCode>
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="99C9F92B0E191359C4083A35D5F29E16" collectionCode="QCAZ" elevation="230" latitude="-0.557" location="Puerto Yuturi" longLatPrecision="78" longitude="-76.008" specimenCode="QCAZ 3609" specimenCount="1" typeStatus="paratype">
|
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<location id="0C1151858FF18F162D5A213F98958310" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:0C1151858FF18F162D5A213F98958310" latitude="-0.557" longLatPrecision="78" longitude="-76.008" name="Puerto Yuturi">Puerto Yuturi</location>
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,
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EODBG" specific-use="{"type":"Point","coordinates":[-76.008000,-0.557000]}">
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<geoCoordinate id="7F664C4F3E64C598BC34CCC48B315D82" degrees="0.557" direction="south" orientation="latitude" precision="55" value="-0.557">0.557 ° S</geoCoordinate>
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,
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<geoCoordinate id="15960EAE78BF090C4810ACE83BF3B194" degrees="76.008" direction="west" orientation="longitude" precision="55" value="-76.008">76.008 ° W</geoCoordinate>
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</named-content>
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,
|
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<quantity id="1BECA61CFA760FEC0418B749E89A287F" metricMagnitude="2" metricUnit="m" metricValue="2.3" unit="m" value="230.0">
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<elevation id="0040E5A6FAD1E1C65613D87C79BCA5F1" metricMagnitude="2" metricUnit="m" metricValue="2.3" unit="m" value="230.0">230 m</elevation>
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</quantity>
|
||||
,
|
||||
<specimenCode id="839A6EC70DA49545B956DA8BC2438C8C">
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<named-content id="EFB356123818BF549EE52A5C1C9347A0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
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3609
|
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</specimenCode>
|
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</materialsCitation>
|
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;
|
||||
<materialsCitation id="7FEF85C46C8CD8AD33F9DEB26832A914" collectionCode="QCAZ" elevation="273" latitude="-0.677" location="near PUCE (Pontificia Universidad Catolica del Ecuador) scientific station" longLatPrecision="78" longitude="-76.401" municipality="Yasuni National Park" specimenCode="QCAZ 5901" specimenCount="1" typeStatus="paratype">
|
||||
<collectingMunicipality id="1D98E4239339B51187695D6365CBCE14">Yasuní National Park</collectingMunicipality>
|
||||
,
|
||||
<location id="354A9ED189421BD9AA8559ADE2C46D5D" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:354A9ED189421BD9AA8559ADE2C46D5D" latitude="-0.677" longLatPrecision="78" longitude="-76.401" municipality="Yasuni National Park" name="near PUCE (Pontificia Universidad Catolica del Ecuador) scientific station">near PUCE (Pontificia Universidad Católica del Ecuador) scientific station</location>
|
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,
|
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1DBG" specific-use="{"type":"Point","coordinates":[-76.401000,-0.677000]}">
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<geoCoordinate id="2F14B2216935D109FBC39EEA6066FBFB" degrees="0.677" direction="south" orientation="latitude" precision="55" value="-0.677">0.677 ° S</geoCoordinate>
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,
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<geoCoordinate id="FD7F873701961D5655A5120941E2341B" degrees="76.401" direction="west" orientation="longitude" precision="55" value="-76.401">76.401 ° W</geoCoordinate>
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</named-content>
|
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,
|
||||
<quantity id="CCAFF8305054F0B0B986D057AB663980" metricMagnitude="2" metricUnit="m" metricValue="2.73" unit="m" value="273.0">
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||||
<elevation id="FFA6F6A0240F08C0194B9BECDD8EDCE3" metricMagnitude="2" metricUnit="m" metricValue="2.73" unit="m" value="273.0">273 m</elevation>
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||||
</quantity>
|
||||
,
|
||||
<specimenCode id="8925719D0C7AFFA4E639EE0FE2A5D519">
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<named-content id="F35992575FC667CF069A4E9D05332419" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
5901
|
||||
</specimenCode>
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="3CFA49A7045CA6619C2E6ECE599C9ED8" collectionCode="QCAZ" elevation="264" latitude="-0.674" location="near PUCE scientific station" longLatPrecision="78" longitude="-76.398" municipality="Yasuni National Park" specimenCode="QCAZ 17519" specimenCount="1" typeStatus="paratype">
|
||||
<collectingMunicipality id="BE4037AF2253994E10EED37C38FBE8C1">Yasuní National Park</collectingMunicipality>
|
||||
,
|
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<location id="807D3567EF57E6A1605BB27F4352E3A8" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:807D3567EF57E6A1605BB27F4352E3A8" latitude="-0.674" longLatPrecision="78" longitude="-76.398" municipality="Yasuni National Park" name="near PUCE scientific station">near PUCE scientific station</location>
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,
|
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGEBG" specific-use="{"type":"Point","coordinates":[-76.398000,-0.674000]}">
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<geoCoordinate id="CDB1536FF1C86B33EB0031C9F599E56E" degrees="0.674" direction="south" orientation="latitude" precision="55" value="-0.674">0.674 ° S</geoCoordinate>
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,
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<geoCoordinate id="AEE3D9F2E575731C782C6F5BD96174AE" degrees="76.398" direction="west" orientation="longitude" precision="55" value="-76.398">76.398 ° W</geoCoordinate>
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</named-content>
|
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,
|
||||
<quantity id="ED954282EE761F8627CB4E603359027A" metricMagnitude="2" metricUnit="m" metricValue="2.6399999999999997" unit="m" value="264.0">
|
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<elevation id="F321336C9F01C1DFCFA7EE39F98EDE49" metricMagnitude="2" metricUnit="m" metricValue="2.6399999999999997" unit="m" value="264.0">264 m</elevation>
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</quantity>
|
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,
|
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<specimenCode id="9BA4DC1191E368BFF8D771D25A898593">
|
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<named-content id="932C17248EEF4914DC053A22D3EC23BB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
17519
|
||||
</specimenCode>
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="D89508D0D64281A2ADDBADC34D375470" collectionCode="QCAZ" elevation="285" latitude="-0.681" location="near PUCE scientific station, north trail of 50 ha dynamic forest plot" longLatPrecision="78" longitude="-76.403" municipality="Yasuni National Park" specimenCode="QCAZ 10229" specimenCount="1" typeStatus="paratype">
|
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<collectingMunicipality id="0916A2CF86041D9D908FA684B5861329">Yasuní National Park</collectingMunicipality>
|
||||
,
|
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<location id="940C3120B941F57B9860BADFEDCD7E80" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:940C3120B941F57B9860BADFEDCD7E80" latitude="-0.681" longLatPrecision="78" longitude="-76.403" municipality="Yasuni National Park" name="near PUCE scientific station, north trail of 50 ha dynamic forest plot">near PUCE scientific station, north trail of 50 ha dynamic forest plot</location>
|
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,
|
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESEBG" specific-use="{"type":"Point","coordinates":[-76.403000,-0.681000]}">
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<geoCoordinate id="703BE26D565015A324FFC83647DEE098" degrees="0.681" direction="south" orientation="latitude" precision="55" value="-0.681">0.681 ° S</geoCoordinate>
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,
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<geoCoordinate id="661C92489A37E525C97E8701D02D9AB9" degrees="76.403" direction="west" orientation="longitude" precision="55" value="-76.403">76.403 ° W</geoCoordinate>
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</named-content>
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,
|
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<quantity id="B90F9A8B95084761ADB14F56FAB73D42" metricMagnitude="2" metricUnit="m" metricValue="2.85" unit="m" value="285.0">
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<elevation id="EBBF2DF1BC5AC06552F8B8C0C63F07F1" metricMagnitude="2" metricUnit="m" metricValue="2.85" unit="m" value="285.0">285 m</elevation>
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</quantity>
|
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,
|
||||
<specimenCode id="A575DB03CC7C7117C84B8D4E67670D17">
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<named-content id="D90B9F6428C5987B28FE79A842737C6B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
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10229
|
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</specimenCode>
|
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</materialsCitation>
|
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;
|
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<materialsCitation id="BE7B0AA7704A67450DE3F395DC44D553" collectionCode="QCAZ" latitude="-0.467" location="El Coca" longLatPrecision="78" longitude="-76.983" municipality="Puerto Francisco de Orellana" specimenCode="QCAZ 185" specimenCount="1" typeStatus="paratype">
|
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<collectingMunicipality id="3848E2CF1AF8B54602D23A248814887B">Puerto Francisco de Orellana</collectingMunicipality>
|
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(
|
||||
<location id="AC1E07A31F1B6F79A902D74F8E4F1C40" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:AC1E07A31F1B6F79A902D74F8E4F1C40" latitude="-0.467" longLatPrecision="78" longitude="-76.983" municipality="Puerto Francisco de Orellana" name="El Coca">El Coca</location>
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),
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5EBG" specific-use="{"type":"Point","coordinates":[-76.983000,-0.467000]}">
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<geoCoordinate id="0DC93D0B94A407E32DBE2DCDC3730369" degrees="0.467" direction="south" orientation="latitude" precision="55" value="-0.467">0.467 ° S</geoCoordinate>
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,
|
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<geoCoordinate id="07EB012653CB89190572485B50D422A2" degrees="76.983" direction="west" orientation="longitude" precision="55" value="-76.983">76.983 ° W</geoCoordinate>
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</named-content>
|
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,
|
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<specimenCode id="7021CBBB6DE45491EE9972125B089183">
|
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<named-content id="CE59CF449DDA620CC295B1A68ECAE0EE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
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185
|
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</specimenCode>
|
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</materialsCitation>
|
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;
|
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<materialsCitation id="AD3CA014CC35FF0B6801B43C60982EB5" collectionCode="QCAZ" elevation="955" latitude="-0.537" location="Rio Bigal Biological Reserve" longLatPrecision="78" longitude="-77.425" specimenCode="QCAZ 17646" specimenCount="1" typeStatus="paratype">
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<location id="D38C1850CD1C7DEC8DA900F47C8015FC" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:D38C1850CD1C7DEC8DA900F47C8015FC" latitude="-0.537" longLatPrecision="78" longitude="-77.425" name="Rio Bigal Biological Reserve">Río Bigal Biological Reserve</location>
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,
|
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKFBG" specific-use="{"type":"Point","coordinates":[-77.425000,-0.537000]}">
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<geoCoordinate id="26970AC5AB60A1B82CA4D6E5FD0FB631" degrees="0.537" direction="south" orientation="latitude" precision="55" value="-0.537">0.537 ° S</geoCoordinate>
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,
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<geoCoordinate id="D51C944F1CBD2A24A4F51A7444588A86" degrees="77.425" direction="west" orientation="longitude" precision="55" value="-77.425">77.425 ° W</geoCoordinate>
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</named-content>
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,
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<quantity id="4FD1FEF8A17CEC3BB670CC2C90BF2DDC" metricMagnitude="2" metricUnit="m" metricValue="9.55" unit="m" value="955.0">
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<elevation id="CBB368A1E2026D498590BBAED010FEA8" metricMagnitude="2" metricUnit="m" metricValue="9.55" unit="m" value="955.0">955 m</elevation>
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</quantity>
|
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,
|
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<specimenCode id="C987CDA935F47A32EC82778CF28C20AB">
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<named-content id="59FA00C2D6DC1A2ABBF15110B3AD4ABB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
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17646
|
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</specimenCode>
|
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</materialsCitation>
|
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;
|
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<materialsCitation id="D7A2C227F81C1F82038C05C011DC5D9D" collectionCode="QCAZ" elevation="251" latitude="-0.49" location="Taracoa" longLatPrecision="78" longitude="-76.773" specimenCode="QCAZ 1118" specimenCount="1" typeStatus="paratype">
|
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<location id="B4E733ECDD9A660AA92F2CB7D516FD43" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:B4E733ECDD9A660AA92F2CB7D516FD43" latitude="-0.49" longLatPrecision="78" longitude="-76.773" name="Taracoa">Taracoa</location>
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,
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWFBG" specific-use="{"type":"Point","coordinates":[-76.773000,-0.490000]}">
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<geoCoordinate id="780A7C097164C80469E56FE4CB5D5C11" degrees="0.490" direction="south" orientation="latitude" precision="55" value="-0.49">0.490 ° S</geoCoordinate>
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,
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<geoCoordinate id="7E88C1FE11E4DB81BE54446413B240F1" degrees="76.773" direction="west" orientation="longitude" precision="55" value="-76.773">76.773 ° W</geoCoordinate>
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</named-content>
|
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,
|
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<quantity id="FF914E7DA2E0F972662A4F86F672BD98" metricMagnitude="2" metricUnit="m" metricValue="2.51" unit="m" value="251.0">
|
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<elevation id="486EA4122142F9E480E8D9B43C2F50F3" metricMagnitude="2" metricUnit="m" metricValue="2.51" unit="m" value="251.0">251 m</elevation>
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</quantity>
|
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,
|
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<specimenCode id="F402939F49E05B94D3EBDFECCC6582B0">
|
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<named-content id="120F87F7849D48492B4360E223A64075" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
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1118
|
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</specimenCode>
|
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</materialsCitation>
|
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.
|
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<materialsCitation id="673FCB46400A80624EEC521E2FD62FB6" collectionCode="QCAZ" elevation="286" latitude="-2.084" location="Montalvo" longLatPrecision="78" longitude="-76.921" municipality="Jatun Yaku community" specimenCode="QCAZ 4839" specimenCount="1" stateProvince="Pastaza" typeStatus="paratype">
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<underline id="ED21B930FAD19C2B2BF278B990CB3D2F">
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<collectingRegion id="95EFE3682FC257643471F25973D4FF72" country="Ecuador" name="Pastaza">Pastaza</collectingRegion>
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</underline>
|
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:
|
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<collectingMunicipality id="CEAD2845ED6D7BBAC714F1C32BB2CEB7">Jatun Yaku community</collectingMunicipality>
|
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,
|
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<locationDeviation id="A097A85D2506A537C1DC9BA3989ACB0B" bearing="135.0" distance="5000.0">
|
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<quantity id="CFBD9ADA0B23ED32C2EE6E498D2E140F" metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
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SE
|
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</locationDeviation>
|
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<location id="192F24F220239E4087E9A1ECFB0834F8" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:192F24F220239E4087E9A1ECFB0834F8" latitude="-2.084" longLatPrecision="78" longitude="-76.921" municipality="Jatun Yaku community" name="Montalvo" stateProvince="Pastaza">Montalvo</location>
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,
|
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EEGBG" specific-use="{"type":"Point","coordinates":[-76.921000,-2.084000]}">
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<geoCoordinate id="0931D3CEABBAF431982FEA56C65F305B" degrees="2.084" direction="south" orientation="latitude" precision="55" value="-2.084">2.084 ° S</geoCoordinate>
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,
|
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<geoCoordinate id="C6BE3FBF2F45BBB5B858290D04A8D795" degrees="76.921" direction="west" orientation="longitude" precision="55" value="-76.921">76.921 ° W</geoCoordinate>
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</named-content>
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,
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<quantity id="FF4AC236BB4E24F77C92877D17D4D7D9" metricMagnitude="2" metricUnit="m" metricValue="2.86" unit="m" value="286.0">
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<elevation id="67976665C463DF7341DB61F255C0AB3E" metricMagnitude="2" metricUnit="m" metricValue="2.86" unit="m" value="286.0">286 m</elevation>
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</quantity>
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,
|
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<specimenCode id="A2F71C7E9DDD098536BB5D98BC3A7B23">
|
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<named-content id="2B1A68F72DE628CCCE00DBD76273E0B9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
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4839
|
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</specimenCode>
|
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</materialsCitation>
|
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;
|
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<materialsCitation id="7244DD0D78B14AD01F1E8B465BF8A578" collectionCode="QCAZ" elevation="468" latitude="-1.475" location="Bloque 10 - Agip Oil" longLatPrecision="78" longitude="-77.534" municipality="Villano camp" specimenCode="QCAZ 11832" specimenCount="1" typeStatus="paratype">
|
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K 10 camp,
|
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<collectingMunicipality id="06B44D2982C50D6FCCA3F5B179A94DFF">Villano camp</collectingMunicipality>
|
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,
|
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<location id="0698C7A6AD51C86F72D49576FFE15701" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:0698C7A6AD51C86F72D49576FFE15701" latitude="-1.475" longLatPrecision="78" longitude="-77.534" municipality="Villano camp" name="Bloque 10 - Agip Oil">Bloque 10 - Agip Oil</location>
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,
|
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQGBG" specific-use="{"type":"Point","coordinates":[-77.534000,-1.475000]}">
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<geoCoordinate id="A865CB29710192C0F82A8866D7097E28" degrees="1.475" direction="south" orientation="latitude" precision="55" value="-1.475">1.475 ° S</geoCoordinate>
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,
|
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<geoCoordinate id="479C9F21DF8997DAE7C5809B3A794639" degrees="77.534" direction="west" orientation="longitude" precision="55" value="-77.534">77.534 ° W</geoCoordinate>
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</named-content>
|
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,
|
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<quantity id="50E7FBF6D387C3A9E6F51E3EFDD97747" metricMagnitude="2" metricUnit="m" metricValue="4.68" unit="m" value="468.0">
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<elevation id="463D1F99506BC5D25295DB14E345D340" metricMagnitude="2" metricUnit="m" metricValue="4.68" unit="m" value="468.0">468 m</elevation>
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</quantity>
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,
|
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<specimenCode id="C6B47F07874EA1CD58C40DD842E327B0">
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<named-content id="364510286ED3AC0F3132E659326C3C80" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
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11832
|
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</specimenCode>
|
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</materialsCitation>
|
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.
|
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<materialsCitation id="ED00CCB6D44924669B8F53B3AB6FDF8F" collectionCode="QCAZ" elevation="229" latitude="-0.498" location="La Selva Amazon Ecolodge" longLatPrecision="78" longitude="-76.374" specimenCode="QCAZ 2926" specimenCount="1" stateProvince="Sucumbios" typeStatus="paratype">
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<underline id="E39351A7C6D6A6DEB230158231D5C844">
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<collectingRegion id="F27F99FD90E21678DE2B67A0F4C36937" country="Ecuador" name="Sucumbios">Sucumbíos</collectingRegion>
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</underline>
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:
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<location id="371E0DEE2938D09C846060342D71910F" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:371E0DEE2938D09C846060342D71910F" latitude="-0.498" longLatPrecision="78" longitude="-76.374" name="La Selva Amazon Ecolodge" stateProvince="Sucumbios">La Selva Amazon Ecolodge</location>
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,
|
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5GBG" specific-use="{"type":"Point","coordinates":[-76.374000,-0.498000]}">
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<geoCoordinate id="2F4C714EB8724096FE52E9B3910A2545" degrees="0.498" direction="south" orientation="latitude" precision="55" value="-0.498">0.498 ° S</geoCoordinate>
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,
|
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<geoCoordinate id="EF9BCA48163092D8BB5E084D27E58F2A" degrees="76.374" direction="west" orientation="longitude" precision="55" value="-76.374">76.374 ° W</geoCoordinate>
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</named-content>
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,
|
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<quantity id="2DDDD794D32F1E40729C76E4AEB06DA3" metricMagnitude="2" metricUnit="m" metricValue="2.29" unit="m" value="229.0">
|
||||
<elevation id="7366F052D0A07328E37F5486F0F186EA" metricMagnitude="2" metricUnit="m" metricValue="2.29" unit="m" value="229.0">229 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<specimenCode id="BAA1232ECE0C9E0D21A0458935160E68">
|
||||
<named-content id="B0E20A6395810C4EACE6EA030D8AD6A1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
2926
|
||||
</specimenCode>
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="C30242CF463DF4F3CBF30084C37CC8E2" collectionCode="QCAZ" latitude="-0.002" location="PUCE scientific station" longLatPrecision="78" longitude="-76.176" municipality="Cuyabeno Wildlife Reserve" specimenCode="QCAZ 2623" specimenCount="1" typeStatus="paratype">
|
||||
<collectingMunicipality id="81E4BEACE37F315E9124DD79B604E35D">Cuyabeno Wildlife Reserve</collectingMunicipality>
|
||||
,
|
||||
<location id="B1A9FB0832C465D4737602677F2124F8" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:B1A9FB0832C465D4737602677F2124F8" latitude="-0.002" longLatPrecision="78" longitude="-76.176" municipality="Cuyabeno Wildlife Reserve" name="PUCE scientific station">PUCE scientific station</location>
|
||||
,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKHBG" specific-use="{"type":"Point","coordinates":[-76.176000,-0.002000]}">
|
||||
<geoCoordinate id="03E97359E0DAE8EB52B9B982100315DF" degrees="0.002" direction="south" orientation="latitude" precision="55" value="-0.002">0.002 ° S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="D50B283027AE5E786C804E27718E5FD1" degrees="76.176" direction="west" orientation="longitude" precision="55" value="-76.176">76.176 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
,
|
||||
<specimenCode id="601A3AA0AA5E89D6693E6D838EDF4080">
|
||||
<named-content id="7CA848336D39A4BA4FAD778C465FAED9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
2623
|
||||
</specimenCode>
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="51C2F4B232A4AF5E10120792068882BB" collectionCode="QCAZ" latitude="-0.031" location="bridge on Cuyabeno River" longLatPrecision="78" longitude="-76.321" municipality="Cuyabeno Wildlife Reserve" specimenCode="QCAZ 7138" specimenCount="1" typeStatus="paratype">
|
||||
<collectingMunicipality id="BFF9534B8C3907D2BBC5390E62C9DB5D">Cuyabeno Wildlife Reserve</collectingMunicipality>
|
||||
,
|
||||
<location id="5BAE17E52539CBF11597D692C713F95A" LSID="urn:lsid:plazi:treatment:41D455A5216C50D0931DAAAD1607ABA8:5BAE17E52539CBF11597D692C713F95A" latitude="-0.031" longLatPrecision="78" longitude="-76.321" municipality="Cuyabeno Wildlife Reserve" name="bridge on Cuyabeno River">bridge on Cuyabeno River</location>
|
||||
,
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWHBG" specific-use="{"type":"Point","coordinates":[-76.321000,-0.031000]}">
|
||||
<geoCoordinate id="EE070E569F177031E7D5B4986B04BA03" degrees="0.031" direction="south" orientation="latitude" precision="55" value="-0.031">0.031 ° S</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="B853FB38377F9EFCB0200EA510E0F003" degrees="76.321" direction="west" orientation="longitude" precision="55" value="-76.321">76.321 ° W</geoCoordinate>
|
||||
</named-content>
|
||||
,
|
||||
<specimenCode id="0FDE23E67C2F983C825AFC2BA2F0F396">
|
||||
<named-content id="B29564CDFD818F8B84235508D361D6EE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
7138
|
||||
</specimenCode>
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="974F088D7C880EA2EBAF788A94DFCEF7" ID-DOI="10.3897/vz.74.e130147.figure5" ID-arpha="32942626-8A09-59F8-86A6-1B76A9B57B76" httpUri="https://binary.pensoft.net/fig/1137917" startId="F5">
|
||||
<paragraph id="B01E8CBC6E1764A03E1209E6F7534359">
|
||||
<label id="02FB3CAE6066145B00E091C7E586A35E">Figure 5.</label>
|
||||
</paragraph>
|
||||
<paragraph id="9FD64668D9FF3F0C659655CC2667C7ED">
|
||||
Head of the holotype (
|
||||
<specimenCode id="C0BC4D957D87522ACBD275284E414792">
|
||||
<named-content id="20FC04616249168893E2CA44EB2436B3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
14814
|
||||
</specimenCode>
|
||||
) of
|
||||
<taxonomicName id="38E5095C5337F975F537967AC75AA884" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis" status="sp. nov.">
|
||||
<emphasis id="9E5B46EAE0C44AC6505C75E6519A79FE" italics="true">Mabuya yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="FC1FBF89449AD09D052A72D4F843E64F" bold="true">
|
||||
<taxonomicNameLabel id="801BE61E52F3464510A611AF645FC2E8" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
in dorsal (top), ventral (middle), and lateral (bottom) views. Photographs by D. A. Paucar. Scale bar: 5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E4IBG" type="diagnosis">
|
||||
<paragraph id="FBF5CED50704D87D6C0E955BE6D8DD7F">
|
||||
<heading id="A6B2C10066DAE969FC481CF056F92B9F" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="55B8C3458DF2157AA41C8C6836ACBB26">
|
||||
<taxonomicName id="E64F2AA4ED5CC0528F2F89FB854648E7" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis" status="sp. nov.">
|
||||
<emphasis id="65B91DDC051D2066F91132879F85BA74" italics="true">Mabuya yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="28836C11820DA6CE8127776702993ECF" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
can be distinguished from other congeners by the combination of the following character states: (1) prefrontals paired; (2) frontoparietals paired; (3) supraciliaries four, second longest; (4) supraoculars four, second largest; (5) supralabials seven, fifth longest and below eye; (6) nuchals in one pair; (7) dorsals smooth; (8) scales around midbody 26–30; (9) dorsals 47–52; (10) ventrals 32–40; (11) caudals smaller and similar in shape to dorsals; (12) lamellae under fourth finger 12–16; (13) lamellae under fourth toe 16–19; (14) palms and soles dark; (15) dark dorsolateral stripes absent; (16) pale dorsolateral stripes absent.
|
||||
</paragraph>
|
||||
<paragraph id="2CF73526D612182B4E7B55E39C6C1A15">
|
||||
Among species of
|
||||
<taxonomicName id="DE9D6978807FB17EB6F01EE989843DA6" authorityName="Fitzinger" authorityYear="1826" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="genus">
|
||||
<emphasis id="33595C4F9068570D8731C3EFFB151C58" italics="true">Mabuya</emphasis>
|
||||
</taxonomicName>
|
||||
known to occur in
|
||||
<collectingCountry id="FB23CC186672EFF297AEDAEC79692E2A" name="Ecuador">Ecuador</collectingCountry>
|
||||
—
|
||||
<taxonomicName id="E9B1DBF48FC76C36323F03C70A4E75B8" authorityName="Miralles, Barrio-Amoros, Rivas & Chaparro-Auza" authorityYear="2006" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="altamazonica">
|
||||
<emphasis id="D3899B74181BA7064BAB4DAA6813904B" italics="true">M. altamazonica</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="E103EAEE80C0CF4F43342246425C32BB" authorityName="Spix" authorityYear="1825" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="nigropunctata">
|
||||
<emphasis id="7AC2F13B852686D1455152168AEF3778" italics="true">M nigropunctata</emphasis>
|
||||
</taxonomicName>
|
||||
—,
|
||||
<taxonomicName id="D9D0430A447944A18601D384348E4B3B" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis">
|
||||
<emphasis id="F533C0D9F3F5395AC8E51BC42DAF11CE" italics="true">M. yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
is most similar to
|
||||
<taxonomicName id="4B0633B22C69B153B9E19002E1DC5082" authorityName="Miralles, Barrio-Amoros, Rivas & Chaparro-Auza" authorityYear="2006" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="altamazonica">
|
||||
<emphasis id="768DDB1012F0B3B11EE5B712A21F5F37" italics="true">M. altamazonica</emphasis>
|
||||
</taxonomicName>
|
||||
in having (1) seven supralabials, with the fifth being the largest and located under the eye (eight supralabials, with the sixth being the largest and located under the eye in
|
||||
<taxonomicName id="EC5BAB6C1159D08D1A751B51E97159E9" authorityName="Spix" authorityYear="1825" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="nigropunctata">
|
||||
<emphasis id="68565950A03C0D9ECB51A756F4882987" italics="true">M. nigropunctata</emphasis>
|
||||
</taxonomicName>
|
||||
), and (2) parietals in contact posterior to interparietal (no contact in
|
||||
<taxonomicName id="75C5B6BAD013F8674448638B8041E847" authorityName="Spix" authorityYear="1825" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="nigropunctata">
|
||||
<emphasis id="1FB7277B40DAA981A21E68E543B71D7D" italics="true">M. nigropunctata</emphasis>
|
||||
</taxonomicName>
|
||||
). However,
|
||||
<taxonomicName id="C2CC08AF48C8A73FD757DD40C2C18AF4" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis">
|
||||
<emphasis id="E8D239DA53E5307476C1CDAF966C2F84" italics="true">M. yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
can be readily distinguished by having four supraciliaries, of which the second is longer than the others (usually five subequal supraciliaries in both
|
||||
<taxonomicName id="905A1B9184739DB66B4DA3E6ECD91C91" authorityName="Miralles, Barrio-Amoros, Rivas & Chaparro-Auza" authorityYear="2006" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="altamazonica">
|
||||
<emphasis id="29A8BD3AEA208F20E9F2F33F76B60C13" italics="true">M. altamazonica</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="C999299AE181BB11565FCC3E856E2EEC" authorityName="Spix" authorityYear="1825" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="nigropunctata">
|
||||
<emphasis id="D1CAFDBDBB61C2414BEE31828A0099C0" italics="true">M. nigropunctata</emphasis>
|
||||
</taxonomicName>
|
||||
). The new species is closely related to
|
||||
<taxonomicName id="B97F6FD7A474E7DAA35A050B785BD7D3" authority="Spix, 1825" authorityName="Spix" authorityYear="1825" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="bistriata">
|
||||
<emphasis id="60200DF433B231D66E8237AF55255184" italics="true">M. bistriata</emphasis>
|
||||
Spix, 1825
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="1DB13C9D336FB07FF91071474EB7F92B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogeny of Mabuya, with a close-up of clades containing M. nigropunctata (blue) and both M. altamazonica and M. yasuniensis sp. nov. (red). All samples of M. yasuniensis correspond to paratypes. This maximum likelihood tree was obtained from an analysis of five genes (two mitochondrial, three nuclear) and 256 terminals. Numbers above branches are ultrafast bootstrap support values, not shown in short branches for clarity. Specimens sampled in this study are shown in bold. Taxon name (M.: Mabuya) followed by voucher number and country of origin are provided. BR: Brazil, CO: Colombia, EC: Ecuador, FG: French Guiana, LA: Lesser Antilles, PE: Peru, TT: Trinidad and Tobago, VE: Venezuela." figureDoi="10.3897/vz.74.e130147.figure1" httpUri="https://binary.pensoft.net/fig/1137913">1</figureCitation>
|
||||
), which also has four supraciliaries, the second the longest. It differs from
|
||||
<taxonomicName id="EE5EFE1411529D9FC2C62CD2BDC42C81" authorityName="Spix" authorityYear="1825" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="bistriata">
|
||||
<emphasis id="19A34025425EE61E0A58F73F0B3555AC" italics="true">M. bistriata</emphasis>
|
||||
</taxonomicName>
|
||||
in (1) lacking a pair of thin, pale dorsolateral stripes from neck to midbody (Fig.
|
||||
<figureCitation id="55DE7FA4248615FCAF9FBE9BBE7141A5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Four species of skinks Mabuya in life, from top to bottom: M. yasuniensis sp. nov. (QCAZ 14814, holotype, SVL = 72.98 mm), M. altamazonica (QCAZ 13087, SVL = 92.11 mm), M. nigropunctata (QCAZ 8107, SVL = 104.58 mm), and M. bistriata. Photographs from Bioweb. bio, except M. bistriata (Laurie J. Vitt)." figureDoi="10.3897/vz.74.e130147.figure2" httpUri="https://binary.pensoft.net/fig/1137914">2</figureCitation>
|
||||
), and having (2) dark palms and soles (Fig.
|
||||
<figureCitation id="22FC13C851E2505760D438F6C67153F4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male holotype (QCAZ 14814; SVL = 72.98 mm) of Mabuya yasuniensis sp. nov. in dorsal (top), lateral (middle) and ventral (bottom) views. Photographs by D. A. Paucar. Scale bar: 10 mm." figureDoi="10.3897/vz.74.e130147.figure4" httpUri="https://binary.pensoft.net/fig/1137916">4</figureCitation>
|
||||
), darker than venter (palms and soles cream or tan, similar in color to venter in
|
||||
<taxonomicName id="370B8D043ED696F316AB51B1F80F1A63" authorityName="Spix" authorityYear="1825" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="bistriata">
|
||||
<emphasis id="5A13C8867FD340C7D6DB0DC12E41B0E2" italics="true">M. bistriata</emphasis>
|
||||
</taxonomicName>
|
||||
), and (3) 47–52 transverse rows of dorsal scales (
|
||||
<quantity id="CD8B1DE8C791087730D79C604CBE2FC5" metricMagnitude="0" metricUnit="m" metricValue="1.4224" metricValueMax="1.4986" metricValueMin="1.3462" unit="in" value="56.0" valueMax="59.0" valueMin="53.0">53–59 in</quantity>
|
||||
<taxonomicName id="5BE5914C589FDF41F9DA11FD9ECFD65D" authorityName="Spix" authorityYear="1825" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="bistriata">
|
||||
<emphasis id="20ED4C394FD9DDAB91E43C42975A6B6C" italics="true">M. bistriata</emphasis>
|
||||
</taxonomicName>
|
||||
; Table
|
||||
<tableCitation id="BF7BAE564EA8E377239631EAF477D697" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Summary of lepidosis and measurements of Mabuya altamazonica, M. bistriata, M. nigropunctata, and M. yasuniensis sp. nov. Range followed by mean ± standard deviation is presented. Sample size is given in parentheses if different from heading." httpUri="http://table.plazi.org/id/33535B323D0E53EF2C22504299BAC971" tableUuid="33535B323D0E53EF2C22504299BAC971">2</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EWOBG" type="description">
|
||||
<paragraph id="204168D94BABA11F26D10CFC26D3B1D2">
|
||||
<heading id="C88E22E1FA1E87BB93DF7584F5F1D627" reason="title">
|
||||
Description of the
|
||||
<typeStatus id="EC52D6C3EA5B1D5BAF150640A6A09C73" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="4203CF2B09CE5D389BB2B09321FBB58C">
|
||||
Adult male (
|
||||
<specimenCode id="D53BEC496E0AEE5FDBD7A7B3E71B67DC">
|
||||
<named-content id="3B4D0F63E844298E8541D4C0BBE9A36E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">
|
||||
<collectionCode id="EEAEE097EA089E3ADA6FD6E25CD076BF" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" type="Museum">QCAZ</collectionCode>
|
||||
</named-content>
|
||||
14814
|
||||
</specimenCode>
|
||||
) (Figs
|
||||
<figureCitation id="77F88363F23A0C9B281E87A2B419D2A7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Four species of skinks Mabuya in life, from top to bottom: M. yasuniensis sp. nov. (QCAZ 14814, holotype, SVL = 72.98 mm), M. altamazonica (QCAZ 13087, SVL = 92.11 mm), M. nigropunctata (QCAZ 8107, SVL = 104.58 mm), and M. bistriata. Photographs from Bioweb. bio, except M. bistriata (Laurie J. Vitt)." figureDoi="10.3897/vz.74.e130147.figure2" httpUri="https://binary.pensoft.net/fig/1137914">2</figureCitation>
|
||||
,
|
||||
<figureCitation id="0411D45ACB53ABDE5C551CC14C728E9E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male holotype (QCAZ 14814; SVL = 72.98 mm) of Mabuya yasuniensis sp. nov. in dorsal (top), lateral (middle) and ventral (bottom) views. Photographs by D. A. Paucar. Scale bar: 10 mm." figureDoi="10.3897/vz.74.e130147.figure4" httpUri="https://binary.pensoft.net/fig/1137916">4</figureCitation>
|
||||
,
|
||||
<figureCitation id="E1C7219E6AA17D6F4E1721F84824A385" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Head of the holotype (QCAZ 14814) of Mabuya yasuniensis sp. nov. in dorsal (top), ventral (middle), and lateral (bottom) views. Photographs by D. A. Paucar. Scale bar: 5 mm." figureDoi="10.3897/vz.74.e130147.figure5" httpUri="https://binary.pensoft.net/fig/1137917">5</figureCitation>
|
||||
);
|
||||
<abbrev id="ABBRID0EMPBG" xlink_title="snout-vent length">SVL</abbrev>
|
||||
<quantity id="7CF6D88482BE8BFA8EFABCC15FA268B1" metricMagnitude="-2" metricUnit="m" metricValue="7.298" unit="mm" value="72.98">72.98 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0EQPBG" xlink_title="tail length">TL</abbrev>
|
||||
<quantity id="4889506CB6BC84076B20DA9C28250C96" metricMagnitude="-1" metricUnit="m" metricValue="1.2672" unit="mm" value="126.72">126.72 mm</quantity>
|
||||
; head length
|
||||
<quantity id="D16D2ECF4271FB546EB77592745ACB6C" metricMagnitude="-2" metricUnit="m" metricValue="1.439" unit="mm" value="14.39">14.39 mm</quantity>
|
||||
, head width
|
||||
<quantity id="ABD09772E32964CD162B0D537FBBC4F6" metricMagnitude="-2" metricUnit="m" metricValue="1.129" unit="mm" value="11.29">11.29 mm</quantity>
|
||||
, head depth
|
||||
<quantity id="2BF81EDD8BE5D9D704D6DE4422D33E72" metricMagnitude="-3" metricUnit="m" metricValue="8.45" unit="mm" value="8.45">8.45 mm</quantity>
|
||||
; rostral wider than high (2.86 ×
|
||||
<quantity id="3705EBD3788818D626F54E86B59C1AFD" metricMagnitude="-3" metricUnit="m" metricValue="1.31" unit="mm" value="1.31">1.31 mm</quantity>
|
||||
), in contact with first supralabials, nasals and supranasals; paired, posterolaterally oriented supranasals in median contact, with posterolateral end contacting anteriormost loreal; frontonasal with rounded anterior border and pointed posterior border, wider than long (3.76 ×
|
||||
<quantity id="EC16DAEC7FFEBCC006251D13503A61FF" metricMagnitude="-3" metricUnit="m" metricValue="2.5" unit="mm" value="2.5">2.50 mm</quantity>
|
||||
), in contact with anterior loreal laterally; paired prefrontals roughly rectangular, anterolaterally oriented, in contact medially, and contacting frontonasal anteriorly, both anterior and posterior loreals laterally, first supraciliaries and first and second supraoculars posterolaterally, and frontal posteromedially; frontal rhomboid, longer than wide (3.93 ×
|
||||
<quantity id="9DB1B7F5CBD418900DAADC2542C86604" metricMagnitude="-3" metricUnit="m" metricValue="2.59" unit="mm" value="2.59">2.59 mm</quantity>
|
||||
), wider anteriorly, in contact with prefrontals anteriorly, second supraoculars laterally, and frontoparietals posteriorly; supraoculars four, the first the smallest, the second the largest; first supraocular in contact with prefrontal anteriorly, first and second supraciliaries laterally, and second supraocular medially; second supraocular in contact with prefrontal anteromedially, frontal medially, frontoparietal and third supraocular posteriorly, and second supraciliary laterally; third supraocular in contact with frontoparietal laterally, fourth supraocular posteriorly, second and third supraciliaries laterally and second supraocular anteriorly; fourth supraocular in contact with frontoparietal medially, parietal and one pretemporal posteriorly, and third and fourth supraciliaries laterally; supraciliaries four, the second more than twice the length of others, which are similar in length; frontoparietals paired, longer than wide, in broad contact at midline, in contact with frontal anteriorly, all supraoculars except the first laterally, parietal and interparietal posteriorly; interparietal rhomboid, longer than wide (3.03 ×
|
||||
<quantity id="7D04B45E51DB8ED5CD0349A74A035C7F" metricMagnitude="-3" metricUnit="m" metricValue="2.54" unit="mm" value="2.54">2.54 mm</quantity>
|
||||
); parietal eye distinct on posterior end of interparietal; parietals more than twice as long as interparietal, wider than long, in contact with each other posterior to interparietal and overlapping the upper anterior temporal scales; each parietal in contact with frontoparietal and fourth supraocular anteriorly, two pretemporals anterolaterally, two temporals (overlapped by parietal) posterolaterally, nuchal posteromedially, and interparietal medially; nasal subrectangular; nostril approximately half the size of nasal, located posteriorly; postnasal small, in contact with supranasal dorsally, anterior loreal and first and second supralabials ventrally, and nasal anteriorly; two subrectangular loreals posterior to nasal, subequal in size, the posteriormost slightly higher; anterior loreal in contact with second supralabial, posterior loreal in contact with second and third supralabials; presubocular single, in contact with fourth and fifth supralabials; preocular single, between presubocular and posterior loreal, in contact with third and fourth supralabials; lower eyelid with large transparent disk, one elongated scale across its dorsal edge (an additional small, anterior scale on right side); supralabials seven, the fifth ventral to eye and more than twice as long as the others; infralabials seven on right side and six on left side; temporals imbricate, smooth, cycloid, not distinctly delimited from scales on the nape or sides of the neck; pretemporals two; primary temporal single, two secondary temporals in contact with three tertiary temporals; ear opening relatively small (maximum diameter =
|
||||
<quantity id="5DFCED7DABEE7244ABBE2A45701F4604" metricMagnitude="-3" metricUnit="m" metricValue="1.04" unit="mm" value="1.04">1.04 mm</quantity>
|
||||
), round, with two scales overlapping anterior margin; auricular lobules absent; mental about three times as wide as long (3.21 ×
|
||||
<quantity id="335E34871AA8EDF6AC68794C64CE9D5B" metricMagnitude="-3" metricUnit="m" metricValue="1.05" unit="mm" value="1.05">1.05 mm</quantity>
|
||||
), with straight posterior margin; postmental wider than long, in contact with first and second infralabials laterally and one pair of chin shields posteriorly; three pairs of chin shields, anterior one in contact medially, posterior ones separated by three longitudinally arranged gular scales; anterior pair of chin shields in contact with second infralabial and first sublabial laterally, posterior pairs separated from infralabials by sublabials; gulars similar in size and shape to ventrals, separated from supralabials by sublabials; palms and soles covered with small tubercles, subequal in size, both regions delimited by a row of larger and flatter scales; subdigital lamellae smooth, single, 14 under fourth finger and 18 under fourth toe on both sides; finger and toes clawed; relative length of the toes in the following order: I <
|
||||
<collectionCode id="916E48E336E778F3F2D5E31EE5724680">II</collectionCode>
|
||||
<
|
||||
<collectionCode id="15C8965850C150305965FA607EF3EBBF" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
<
|
||||
<collectionCode id="ED73C4BEC418204F271AC3862D5832E3">III</collectionCode>
|
||||
<
|
||||
<collectionCode id="BE0C6EBB23675B6F6A896F40ECA12942">IV</collectionCode>
|
||||
; conspicuous posthumeral mite pocket with slit-like opening; all scales, except head shields and scales on sole and digits, cycloid, smooth and imbricate; longitudinal scale rows around midbody 26, transverse rows of dorsal scales 49, transverse rows of ventral scales 36; preanals four, larger than adjacent ventral scales; median subcaudal series of scales twice as wide as long on posterior half of tail.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EUPBG" type="Coloration of the holotype (Figs 2, 4, 5)">
|
||||
<paragraph id="DAF9F392259BB7B9E1CE0D3305D38BC7">
|
||||
<heading id="50737BD6332082B01184E8FF2506A3BB" reason="title">
|
||||
Coloration of the
|
||||
<typeStatus id="52B2271F44A80975248FF4D7F0701A3E" type="holotype">holotype</typeStatus>
|
||||
(Figs
|
||||
<figureCitation id="6281592A161CBA04B4A8A6DFFD37C035" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Four species of skinks Mabuya in life, from top to bottom: M. yasuniensis sp. nov. (QCAZ 14814, holotype, SVL = 72.98 mm), M. altamazonica (QCAZ 13087, SVL = 92.11 mm), M. nigropunctata (QCAZ 8107, SVL = 104.58 mm), and M. bistriata. Photographs from Bioweb. bio, except M. bistriata (Laurie J. Vitt)." figureDoi="10.3897/vz.74.e130147.figure2" httpUri="https://binary.pensoft.net/fig/1137914">2</figureCitation>
|
||||
,
|
||||
<figureCitation id="5C45BE6E1571B6D33102D38C53C3E39C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male holotype (QCAZ 14814; SVL = 72.98 mm) of Mabuya yasuniensis sp. nov. in dorsal (top), lateral (middle) and ventral (bottom) views. Photographs by D. A. Paucar. Scale bar: 10 mm." figureDoi="10.3897/vz.74.e130147.figure4" httpUri="https://binary.pensoft.net/fig/1137916">4</figureCitation>
|
||||
,
|
||||
<figureCitation id="092902766E696FB9D7BDEA4E4D21514C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Head of the holotype (QCAZ 14814) of Mabuya yasuniensis sp. nov. in dorsal (top), ventral (middle), and lateral (bottom) views. Photographs by D. A. Paucar. Scale bar: 5 mm." figureDoi="10.3897/vz.74.e130147.figure5" httpUri="https://binary.pensoft.net/fig/1137917">5</figureCitation>
|
||||
).
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="96CB966D2648F0D749E687E3F3DD1E05">Dorsal background olive-bronze; ventral side of head, body, and limbs immaculate bluish-grey; ventral aspect of tail with dense brown pigmentation; dorsum with series of small (~ 1 / 3 of a dorsal scale) dark brown spots more or less longitudinally arranged from scapular region onto tail; lateral and dorsal sides of limbs spotted with small, fused dark dots; palms and soles dark brown; preanals pale cream; dark lateral and ventrolateral stripes present; dark lateral stripe wider, about three scales wide at midbody, extending from nostrils through loreals, upper half of supralabials, around eyes, temporals, upper half of ear openings, neck, above arms, and along flanks up to insertion of hindlimbs; dark ventrolateral stripe diffuse, extending from corner of mouth, along ventral aspect of ear opening, above forelimbs, up to insertion of hindlimbs; dark lateral and ventrolateral stripes separated from each other by ill-defined pale lateral stripe, same color as venter, about one scale wide at midbody, with brown pigmentation (denser on posterior half).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EGQBG" type="Hemipenis">
|
||||
<paragraph id="0A653B9B5733B45122230F06A75B32D2">
|
||||
<heading id="F2DF2616CEC48F475320FA8BBCFED0CA" reason="title">Hemipenis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="6D781B56EEF93A56F0FE50C891BDEE8E">
|
||||
The following description is based on two partially everted hemipenes of
|
||||
<specimenCount id="22E03229D675B43490037DF116ED9D82" count="2" type="male">two male</specimenCount>
|
||||
<typeStatus id="FABAD35F02DBDC40F8B80F5BC82DC66D" type="paratype">paratypes</typeStatus>
|
||||
of
|
||||
<taxonomicName id="18FEC62C2F836A3BFF4EB6B247F13895" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis">
|
||||
<emphasis id="7FED0CD5466DFC705DE261A412EFCD7D" italics="true">Mabuya yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="432DE0E54387CB62CBC21C13B3397F45" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Hemipenes of two male paratypes of Mabuya yasuniensis sp. nov. (top: QCAZ 4839, SVL = 77.46 mm; bottom: QCAZ 11832, SVL = 76.64 mm) in asulcate, sulcate and lateral views from left to right. Number labels correspond to characters described in the text based on Sánchez-Martínez et al. (2020). Photographs by D. A. Paucar. Scale bar: 2 mm." figureDoi="10.3897/vz.74.e130147.figure6" httpUri="https://binary.pensoft.net/fig/1137918">6</figureCitation>
|
||||
); character numbers in parentheses correspond to those listed by
|
||||
<bibRefCitation id="530A3E1E2FA9E499D9BC0A866B0C84C0" DOI="10.1002/ar.24329" author="Sánchez-Martínez PM & Ramírez-Pinilla MP & Meneses-Pelayo E & Nunes PMS" etAl="et al." firstAuthor="Sanchez-Martinez" journalOrPublisher="Anatomical Record" pagination="2917-2930" refId="B28" refString="Sánchez-Martínez PM, Ramírez-Pinilla MP, Meneses-Pelayo E, Nunes PMS (2020) Hemipenial morphology of nine South American species of Mabuya (Scincidae: Lygosominae) with comments on the morphology of the family. Anatomical Record 303: 2917–2930. https://doi.org/10.1002/ar.24329" title="Hemipenial morphology of nine South American species of Mabuya (Scincidae: Lygosominae) with comments on the morphology of the family." volume="303" year="2020">Sánchez-Martínez et al. (2020)</bibRefCitation>
|
||||
. Organ tubular, bilobed, not capitate, with both external and internal membranous layers;
|
||||
<emphasis id="DE927A7C9D65D630FFA638545789A4FD" italics="true">sulcus spermaticus</emphasis>
|
||||
diagonally oriented, bifurcating at base of lobes, with crotch before branches deep and exposed; proximal half of
|
||||
<emphasis id="CAD4541E00AF433DE1E415D17428C0E3" italics="true">sulcus spermaticus</emphasis>
|
||||
overlapped by large thick fold (1); two small horizontal ridges below the base of each lobe (2) on sulcate side; obtuse projection (lower fold, 3) proximal to each pair of small horizontal ridges; broad, medial bell-shaped fold (4) below lobes on asulcate side, its upper tip (5) extending on surface between lobes; two large protuberances (6) proximal to bell-shaped fold; lateral fold (7) connecting lower fold (3) of sulcate side with large protuberance (6) of asulcate side.
|
||||
</paragraph>
|
||||
<caption id="55EF4CF1B3534597033E47D63F87F743" ID-DOI="10.3897/vz.74.e130147.figure6" ID-arpha="52424FF1-8417-56FA-A292-C2FD816C395F" httpUri="https://binary.pensoft.net/fig/1137918" startId="F6">
|
||||
<paragraph id="A27E7B2DEE7118D98021A3216914318B">
|
||||
<label id="6D8478C5BC3A3218A2844DB881AFE53C">Figure 6.</label>
|
||||
</paragraph>
|
||||
<paragraph id="77934B4558BB34CA033771BF86A9884F">
|
||||
Hemipenes of two male paratypes of
|
||||
<taxonomicName id="571A40BB7E56AF605A5EE781304379FC" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis" status="sp. nov.">
|
||||
<emphasis id="6DFFFAE5429373468F3A494CF7A92F4B" italics="true">Mabuya yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="C28B8A422A30F266F8F4CCE8A2433D50" bold="true">
|
||||
<taxonomicNameLabel id="4ABEA7EAB07E5431CB856741098458CE" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(top:
|
||||
<specimenCode id="602702C5EBCF0BECF0379A462C4D5AE4">
|
||||
<named-content id="6234DD0C644AC212081F7E1B55142EFB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
4839
|
||||
</specimenCode>
|
||||
,
|
||||
<abbrev id="ABBRID0E3RBG" xlink_title="snout-vent length">SVL</abbrev>
|
||||
= 77.46 mm; bottom:
|
||||
<specimenCode id="4DD023365D0AFCF1892AC83EB935A1E9">
|
||||
<named-content id="78D13B4E46E717F43506BD1B62B70265" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
11832
|
||||
</specimenCode>
|
||||
,
|
||||
<abbrev id="ABBRID0EESBG" xlink_title="snout-vent length">SVL</abbrev>
|
||||
= 76.64 mm) in asulcate, sulcate and lateral views from left to right. Number labels correspond to characters described in the text based on
|
||||
<bibRefCitation id="9626EFCF85C6A38B22B66D8A7C15C4B5" DOI="10.1002/ar.24329" author="Sánchez-Martínez PM & Ramírez-Pinilla MP & Meneses-Pelayo E & Nunes PMS" etAl="et al." firstAuthor="Sanchez-Martinez" journalOrPublisher="Anatomical Record" pagination="2917-2930" refId="B28" refString="Sánchez-Martínez PM, Ramírez-Pinilla MP, Meneses-Pelayo E, Nunes PMS (2020) Hemipenial morphology of nine South American species of Mabuya (Scincidae: Lygosominae) with comments on the morphology of the family. Anatomical Record 303: 2917–2930. https://doi.org/10.1002/ar.24329" title="Hemipenial morphology of nine South American species of Mabuya (Scincidae: Lygosominae) with comments on the morphology of the family." volume="303" year="2020">Sánchez-Martínez et al. (2020)</bibRefCitation>
|
||||
. Photographs by D. A. Paucar. Scale bar: 2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ERSBG" type="variation">
|
||||
<paragraph id="B60E3F5BF07EFDBA3D4B3A10FB078F32">
|
||||
<heading id="4D4DF3FED6194F1BE6A78D89BEE1B7D1" reason="title">Variation.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="BD5397E55B3CBDAC90F28EE5EA3EC203">
|
||||
Intraspecific morphological variation in
|
||||
<taxonomicName id="D4B815412564B58F02DF4B101F285F8C" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis">
|
||||
<emphasis id="9741ABB461AC99E876F89BB822BE3C0F" italics="true">M. yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
is presented in Table
|
||||
<tableCitation id="BB4DDA5E5067EC2228DA967635A0B20E" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Summary of lepidosis and measurements of Mabuya altamazonica, M. bistriata, M. nigropunctata, and M. yasuniensis sp. nov. Range followed by mean ± standard deviation is presented. Sample size is given in parentheses if different from heading." httpUri="http://table.plazi.org/id/33535B323D0E53EF2C22504299BAC971" tableUuid="33535B323D0E53EF2C22504299BAC971">2</tableCitation>
|
||||
. One of the most distinctive characters of
|
||||
<taxonomicName id="D6A9C99A38F63F04FE1582FC6A8151F4" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis">
|
||||
<emphasis id="D8F3D35118848FEF0D981928C8C34C9E" italics="true">M. yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
is having four supraciliaries, of which the second is elongated; however, of 14 examined specimens, one had only three supraciliaries, possibly as a result of scale fusion. Contact of parietals behind interparietal, a diagnostic character for
|
||||
<taxonomicName id="B1740F6764E05F8EE1C8114B4C011162" authorityName="Miralles, Barrio-Amoros, Rivas & Chaparro-Auza" authorityYear="2006" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="altamazonica">
|
||||
<emphasis id="42BC94D7EDB7EDBD47739B2D8CA5E3AA" italics="true">M. altamazonica</emphasis>
|
||||
</taxonomicName>
|
||||
, was observed in 79 % of the specimens. The dorsal color pattern of
|
||||
<taxonomicName id="5FB6ED12B126ACE16F5B2D4A61617110" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis">
|
||||
<emphasis id="E0732D91750353FA4856DB64C5E1FFA8" italics="true">M. yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
varies from having a few longitudinally aligned dark brown spots (usually restricted to scale tips) to a densely spotted back, whereas the dark ventrolateral stripe can be ill-defined as a few longitudinally arranged dark spots on flank scales (Fig.
|
||||
<figureCitation id="D3433B07425981AE5B240C24EEC38FF7" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Paratypes (top: QCAZ 4839, SVL = 77.46 mm; bottom QCAZ 17646, SVL = 54.81 mm) of Mabuya yasuniensis sp. nov. in dorsal and lateral views. Photographs by D. A. Paucar. Scale bar: 10 mm." figureDoi="10.3897/vz.74.e130147.figure7" httpUri="https://binary.pensoft.net/fig/1137919">7</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="F6A40FBDCCD4404586B49CF258AC5994" ID-DOI="10.3897/vz.74.e130147.figure7" ID-arpha="05878AE7-466D-5107-B678-BB881BAA33F9" httpUri="https://binary.pensoft.net/fig/1137919" startId="F7">
|
||||
<paragraph id="497ED2C894C76193452304555213E613">
|
||||
<label id="BF4440973E109042676E09E3E0414B5B">Figure 7.</label>
|
||||
</paragraph>
|
||||
<paragraph id="33362905E96CE9423C99DEBE3D693D05">
|
||||
Paratypes (top:
|
||||
<specimenCode id="A129003277FFE055E3E3971A6E45D242">
|
||||
<named-content id="5CDAC1A75B2727549D1C633DD03ECB12" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
4839
|
||||
</specimenCode>
|
||||
,
|
||||
<abbrev id="ABBRID0EXUBG" xlink_title="snout-vent length">SVL</abbrev>
|
||||
= 77.46 mm; bottom
|
||||
<specimenCode id="0F7D6479E4CA2851270B3F8D2236D947">
|
||||
<named-content id="1D25505B33EB7F3889A6B5CCD30CB68C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-de-zoologia-pontificia-universidad-catolica-del-ecuador" xlink_title="Pontificia Universidad Católica del Ecuador, Quito">QCAZ</named-content>
|
||||
17646
|
||||
</specimenCode>
|
||||
,
|
||||
<abbrev id="ABBRID0E6UBG" xlink_title="snout-vent length">SVL</abbrev>
|
||||
= 54.81 mm) of
|
||||
<taxonomicName id="3494F5EDCBBDE64CE6CDF92957945813" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis" status="sp. nov.">
|
||||
<emphasis id="43CD23249710413678E5C1A89C90A407" italics="true">Mabuya yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="A1C037E6D4ED076945D2BF0CD03FB09A" bold="true">
|
||||
<taxonomicNameLabel id="CB5614C4B55DCCDE046F5CE28C68E30D" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
in dorsal and lateral views. Photographs by D. A. Paucar. Scale bar: 10 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EVVBG" type="etymology">
|
||||
<paragraph id="174A65EEE052B7C7D7DD677F2CA22DCD">
|
||||
<heading id="3054D3DFF424B03525CA8EEAAF60FB01" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="7E4BFC91871D21E433194BC39C1D3E1F">
|
||||
The specific epithet
|
||||
<taxonomicName id="575B83B3CE721814291B815C7BF8AFBE" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis">
|
||||
<emphasis id="6272B2FE7F0B0EC9B964FFE7DFAF140A" italics="true">yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
derives from the words “ Yasuní ” (Spanish) and “ - ensis ” (Latin adjectival suffix denoting place or locality). It refers to Yasuní National Park, where
|
||||
<taxonomicName id="6743E5305299772A5BEE1437E95E53A5" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis">
|
||||
<emphasis id="9CB48AE36A4942E1F39AACE6F6D713A0" italics="true">M. yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
was discovered. With
|
||||
<quantity id="19A571B156616936443D402F55BF0AE3" metricMagnitude="6" metricUnit="m" metricValue="9.82" unit="km" value="9820.0">9,820 km</quantity>
|
||||
<superScript id="C0BBD255C55E3617FE4471AA2C0C893C">2</superScript>
|
||||
, Yasuní is the largest protected area in continental
|
||||
<collectingCountry id="758A541EADE1CF8B8E9FC6D0513841A8" name="Ecuador">Ecuador</collectingCountry>
|
||||
and one of the most biodiverse places on Earth (
|
||||
<bibRefCitation id="82D020F204E65B99447794A27D1FCE82" DOI="10.1371/journal.pone.0008767" author="Bass" etAl="et al." firstAuthor="Bass" refId="B2" refString="Bass MS, Finer M, Jenkins CN, Kreft H, Cisneros-Heredia DF, McCracken SF, Pitman NCA, English PH, Swing K, Villa G, Di Fiore A, Voigt CC, Kunz TH (2010) Global conservation significance of Ecuador’s Yasuní National Park. PLoS ONE 5: e 8767. https://doi.org/10.1371/journal.pone.0008767" year="2010">Bass et al. 2010</bibRefCitation>
|
||||
). In 1989 Yasuní was declared a UNESCO Biosphere Reserve.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EWWBG" type="distribution">
|
||||
<paragraph id="68DC23E902722D36BF272B4A80EE9C64">
|
||||
<heading id="168F98064E7F6F66B9D2687642A30126" reason="title">Distribution and natural history.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="53D87CF1F2E19D0D38A7EC0167A03E66">
|
||||
<taxonomicName id="5CB030DE50BDD2C8C7DCEC5F24298071" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis">
|
||||
<emphasis id="01CFCED9F7A7C753D97E877070753C2D" italics="true">Mabuya yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
is known from the Amazonian lowlands of
|
||||
<collectingCountry id="55864D4E9FE021D972CDB457571815C3" name="Ecuador">Ecuador</collectingCountry>
|
||||
—
|
||||
<collectingRegion id="D578D3447D542E1B5048EB9A9DF0BB23" country="Ecuador" name="Orellana">Orellana</collectingRegion>
|
||||
,
|
||||
<collectingRegion id="6B24EC1DA171FE091F6A9E7433C6B139" country="Ecuador" name="Pastaza">Pastaza</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="9103CDA09E0F1F8135113A3718A83620" country="Ecuador" name="Sucumbios">Sucumbíos</collectingRegion>
|
||||
provinces, at elevations between
|
||||
<quantity id="B6C274503FBAD990E102ABD3A35F10EE" metricMagnitude="2" metricUnit="m" metricValue="5.775" metricValueMax="9.55" metricValueMin="2.0" unit="m" value="577.5" valueMax="955.0" valueMin="200.0">200–955 m</quantity>
|
||||
(Fig.
|
||||
<figureCitation id="A180AD37D396B208A3D7C7C86F9A4496" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Known distribution of Mabuya in Ecuador: M. altamazonica (squares), M. nigropunctata (circles), M. yasuniensis sp. nov. (triangles). Red-filled symbols correspond to localities of sequenced specimens. Red star indicates type locality of M. yasuniensis sp. nov." figureDoi="10.3897/vz.74.e130147.figure3" httpUri="https://binary.pensoft.net/fig/1137915">3</figureCitation>
|
||||
). Some localities lie within protected areas, including Yasuní National Park, Cuyabeno Wildlife Reserve and Río Bigal Biological Reserve, suggesting that some populations of
|
||||
<taxonomicName id="531AD73E7E4C43C1E14386472120D42E" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis">
|
||||
<emphasis id="24442D4DE57377911D6DC37C5DE4E9FD" italics="true">M. yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
are well protected. Specimens were found active mostly on the ground between 1030 h and 1630 h in primary and secondary forests. The
|
||||
<typeStatus id="1DBE0A5A260A0293065EF2206E4E0FEF" type="holotype">holotype</typeStatus>
|
||||
was found within moss
|
||||
<quantity id="5D6332C87BD5EABBD0870ED1F0B7E344" metricMagnitude="0" metricUnit="m" metricValue="1.0" unit="m" value="1.0">1 m</quantity>
|
||||
above water on a tree standing within a lagoon.
|
||||
<taxonomicName id="B4814A2E4DB06F9D01B1EBF2CB6623E5" authorityName="Torres-Carvajal & Sandoval & Paucar" authorityYear="2024" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="yasuniensis">
|
||||
<emphasis id="C57072BC9F153B777BC680F5B33EEB3E" italics="true">Mabuya yasuniensis</emphasis>
|
||||
</taxonomicName>
|
||||
occurs in sympatry with both
|
||||
<taxonomicName id="D4D4184F1364984A39DA7A4A634D26E0" authorityName="Miralles, Barrio-Amoros, Rivas & Chaparro-Auza" authorityYear="2006" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="altamazonica">
|
||||
<emphasis id="4CA2C79B3B7B1993227689AA1B660F1B" italics="true">M. altamazonica</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4F954EFE130214D9EB396774CD0AD2C5" authorityName="Spix" authorityYear="1825" class="Squamata" family="Scincidae" genus="Mabuya" kingdom="Animalia" phylum="Chordata" rank="species" species="nigropunctata">
|
||||
<emphasis id="70E78FE163D6007621BFFDA7D24A8B47" italics="true">M. nigropunctata</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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