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<mods:title id="4ECEBCA87DCA1D3FB59C770DA100B650">A new subspecies ofBoletoxenus mixtus Grimm, 2014 (Coleoptera: Tenebrionidae: Tenebrioninae)</mods:title>
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<mods:title id="9B1BDA4A62B9CE5834CB872180E7F38D">A new subspecies ofBoletoxenus mixtus Grimm, 2014 (Coleoptera: Tenebrionidae: Tenebrioninae)</mods:title>
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@ -52,7 +53,7 @@
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<subSubSection id="50B7E0CC5C10E151CD295875FE23ADA8" pageId="1" pageNumber="296" type="description">
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<paragraph id="1812B3475C10E151CD295875FE23ADA8" blockId="1.[114,586,1324,1440]" pageId="1" pageNumber="296">
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(
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<figureCitation id="8096AFC25C10E151CD225875FF19AD6B" box="[121,215,1355,1379]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[624,663,1541,1565]" captionStartId-1="3.[624,663,1056,1080]" captionStartId-2="4.[189,228,898,922]" captionTargetBox-0="[630,1119,821,1518]" captionTargetBox-1="[624,1113,299,1036]" captionTargetBox-2="[194,672,340,876]" captionTargetId-0="figure-357@1.[623,1133,814,1535]" captionTargetId-1="figure-424@3.[623,1117,293,1050]" captionTargetId-2="figure-426@4.[188,674,327,890]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1.Boletoxenus mixtus dynastesssp. nov. holotype male. A – Habitus, dorsal view; B – ditto, lateral view; C – ditto, anterior view [not to scale]." captionText-1="Fig. 2.Boletoxenus mixtus dynastesssp. nov. paratype female. A – Habitus, dorsal view; B – ditto, lateral view; C – ditto, anterior view [not to scale]." captionText-2="Fig. 3.Boletoxenus mixtus dynastesssp. nov. holotype male, aedeagus. A – Dorsal view; B – Ventral view; C – Lateral view." pageId="1" pageNumber="296">Figs 1‒3</figureCitation>
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<figureCitation id="8096AFC25C10E151CD225875FF19AD6B" box="[121,215,1355,1379]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[624,663,1541,1565]" captionStartId-1="3.[624,663,1056,1080]" captionStartId-2="4.[189,228,898,922]" captionTargetBox-0="[630,1119,821,1518]" captionTargetBox-1="[624,1113,299,1036]" captionTargetBox-2="[194,672,340,876]" captionTargetId-0="figure-357@1.[623,1133,814,1535]" captionTargetId-1="figure-424@3.[623,1117,293,1050]" captionTargetId-2="figure-426@4.[188,674,327,890]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1.Boletoxenus mixtus dynastesssp. nov. holotype male. A – Habitus, dorsal view; B – ditto, lateral view; C – ditto, anterior view [not to scale]." captionText-1="Fig. 2.Boletoxenus mixtus dynastesssp. nov. paratype female. A – Habitus, dorsal view; B – ditto, lateral view; C – ditto, anterior view [not to scale]." captionText-2="Fig. 3.Boletoxenus mixtus dynastesssp. nov. holotype male, aedeagus. A – Dorsal view; B – Ventral view; C – Lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.13203421" figureDoi-1="http://doi.org/10.5281/zenodo.13203425" figureDoi-2="http://doi.org/10.5281/zenodo.13203427" httpUri-0="https://zenodo.org/record/13203421/files/figure.png" httpUri-1="https://zenodo.org/record/13203425/files/figure.png" httpUri-2="https://zenodo.org/record/13203427/files/figure.png" pageId="1" pageNumber="296">Figs 1‒3</figureCitation>
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)
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<uri id="6C3CBF455C10E151CD295854FE23ADA8" pageId="1" pageNumber="296">
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urn:lsid:zoobank.org:act:
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@ -85,7 +86,7 @@ male
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[printed], 2003 [handwritten].
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</materialsCitation>
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</paragraph>
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<caption id="4CD2E3CF5C10E151CF2B5B3BFCCCAE72" pageId="1" pageNumber="296" startId="1.[624,663,1541,1565]" targetBox="[630,1119,821,1518]" targetPageId="1" targetType="figure">
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<caption id="4CD2E3CF5C10E151CF2B5B3BFCCCAE72" ID-DOI="http://doi.org/10.5281/zenodo.13203421" ID-Zenodo-Dep="13203421" httpUri="https://zenodo.org/record/13203421/files/figure.png" pageId="1" pageNumber="296" startId="1.[624,663,1541,1565]" targetBox="[630,1119,821,1518]" targetPageId="1" targetType="figure">
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<paragraph id="1812B3475C10E151CF2B5B3BFCCCAE72" blockId="1.[624,1103,1541,1658]" pageId="1" pageNumber="296">
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Fig. 1.
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<taxonomicName id="DFADC8C45C10E151CFE35B38FC23AE16" authorityName="Telnov" authorityYear="2023" box="[696,1005,1542,1566]" class="Insecta" family="Tenebrionidae" genus="Boletoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="subSpecies" species="mixtus" status="subsp. nov." subSpecies="dynastes">
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@ -110,9 +111,9 @@ holotype male. A – Habitus, dorsal view; B – ditto, lateral view; C – ditt
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<emphasis id="2AD96F555C13E152CDE65D8FFE80A8C1" bold="true" box="[189,334,177,201]" pageId="2" pageNumber="297">Description.</emphasis>
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<typeStatus id="C7160DE55C13E152CC0C5D8CFE75A8C2" box="[343,443,178,202]" pageId="2" pageNumber="297" type="holotype">Holotype</typeStatus>
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male (
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<figureCitation id="8096AFC25C13E152CF4D5D8CFDACA8C2" box="[534,610,178,202]" captionStart="Fig" captionStartId="1.[624,663,1541,1565]" captionTargetBox="[630,1119,821,1518]" captionTargetId="figure-357@1.[623,1133,814,1535]" captionTargetPageId="1" captionText="Fig. 1.Boletoxenus mixtus dynastesssp. nov. holotype male. A – Habitus, dorsal view; B – ditto, lateral view; C – ditto, anterior view [not to scale]." pageId="2" pageNumber="297">Figs 1</figureCitation>
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||||
<figureCitation id="8096AFC25C13E152CF4D5D8CFDACA8C2" box="[534,610,178,202]" captionStart="Fig" captionStartId="1.[624,663,1541,1565]" captionTargetBox="[630,1119,821,1518]" captionTargetId="figure-357@1.[623,1133,814,1535]" captionTargetPageId="1" captionText="Fig. 1.Boletoxenus mixtus dynastesssp. nov. holotype male. A – Habitus, dorsal view; B – ditto, lateral view; C – ditto, anterior view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13203421" httpUri="https://zenodo.org/record/13203421/files/figure.png" pageId="2" pageNumber="297">Figs 1</figureCitation>
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,
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<figureCitation id="8096AFC25C13E152CF225D8CFD45A8C2" box="[633,651,178,202]" captionStart="Fig" captionStartId="4.[189,228,898,922]" captionTargetBox="[194,672,340,876]" captionTargetId="figure-426@4.[188,674,327,890]" captionTargetPageId="4" captionText="Fig. 3.Boletoxenus mixtus dynastesssp. nov. holotype male, aedeagus. A – Dorsal view; B – Ventral view; C – Lateral view." pageId="2" pageNumber="297">3</figureCitation>
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<figureCitation id="8096AFC25C13E152CF225D8CFD45A8C2" box="[633,651,178,202]" captionStart="Fig" captionStartId="4.[189,228,898,922]" captionTargetBox="[194,672,340,876]" captionTargetId="figure-426@4.[188,674,327,890]" captionTargetPageId="4" captionText="Fig. 3.Boletoxenus mixtus dynastesssp. nov. holotype male, aedeagus. A – Dorsal view; B – Ventral view; C – Lateral view." figureDoi="http://doi.org/10.5281/zenodo.13203427" httpUri="https://zenodo.org/record/13203427/files/figure.png" pageId="2" pageNumber="297">3</figureCitation>
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). Dorsum and venter entirely dark brown. Maxillary and labial palpi, antenna and legs reddish brown. Terminal setae of prothoracic processes pale rufous. Body moderately elongate-elliptical, strongly convex in dorsal aspect, opaque to subopaque. Head transverse, somewhat flattened on frons in dorsal aspect. Labrum anterior margin subtruncate. Epistoma separated from frons by weak suture. Epistoma with a long, irregularly transversely rugulose, apically slightly widened and subtruncate to shallowly emarginate process directed anterodorsad; apex of epistomal process with a row of short erect pale rufous setae. In lateral aspect, epistomal process very slightly arched. Back of epistomal process slightly concave, covered with short erect setae. Length of epistomal process
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<quantity id="DF551EA25C13E152CF4E5E29FDBEAB26" box="[533,624,791,815]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="297" unit="mm" value="1.7">1.7 mm</quantity>
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or lightly shorter than length of lateral margin of pronotum (~
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@ -122,7 +123,7 @@ or lightly shorter than length of lateral margin of pronotum (~
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(apical setation not included). Lateral and dorsal surface of a process with irregular, subacute conical tubercles, inner margin (facing another process) nearly smooth. Anterior pronotal margin broadly emarginate, anterolateral angles produced anteriad, nearly right-angled, indistinctly bifurcate, meets with canthus and encircles corresponding compound eye when head is retracted into pronotum. Basal pronotal margin slightly concave medially, shallowly emarginate laterally (bisinuous), somewhat narrowly flattened, non-punctate or tuberculate. Posterolateral angle acuteangulate, slightly produced posteriad. Lateral margin of pronotum explanate, slightly narrows anteriad, with a row of eight acute denticles (counting also denticle on each antero- and posterolateral angles). Pronotal disc except at anterior margin with irregular, variably sized subacute conical tubercles and much smaller punctures of variable size on intervening spaces between tubercles. Tubercles posterior to pronotal processes arranged into two rows from base of processes towards nearly base of pronotum; small, shallow, non-tuberculate depression each side of these rows. Near anterior margin pronotum is with numerous
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</paragraph>
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<paragraph id="1812B3475C13E153C9375B98FD9FAE63" blockId="2.[1132,1172,1702,1726]" lastBlockId="3.[114,593,178,1643]" lastPageId="3" lastPageNumber="298" pageId="2" pageNumber="297">297 subparallel longitudinally sulci. Intervening spaces microstriate. Prothoracic hypomeron rather coarsely punctate with opaque intervening spaces, non-tuberculate. Pronotal setation inconspicuous, generally as that on head. Scutellar shield subtriangular, opaque, glabrous. Elytra about 1.2–1.3× as long as combined wide, laterally slightly constricted posteriad, convex in dorsal aspect. Anterior margin of elytron with numerous short notches all along except on humeral area. Humerus near right-angled, humeral callosity small but distinctly convex. Elytral disc with series of conical tubercles of variable size and height. Four rather high, acutelyridged, three- to multifurcate (in lateral profile) tubercles each side of and some distance from suture. Remaining tubercles comparatively smaller, with single peak in lateral profile. Lateral margin of elytron densely denticulate; denticles somewhat smaller and denser than those on lateral margin of pronotum. Lateral margin of elytron slightly sinuous in lateral view. Intervening spaces between tubercles with variably large circular punctures; on each elytron punctures arranged into nine longitudinal rows strongly interrupted and disorganized by tubercular pattern of elytral dorsum. Intervening spaces between punctures microstriate. Elytral setae inconspicuous, narrowly scale-like, very sparse, suberect. Suture not or barely raised. Epipleuron wide, extending towards elytral apex, strongly narrows at inner sutural angle. Metathoracic wings fully developed. Mesoventrite V-shapely impressed to accommodate prosternal intercoxal process. Intercoxal process of first visible abdominal ventrite wide, apically widened, slightly convex in ventral aspect. Legs moderately long. Femur not clavate, densely transversely rugulose, distinctly surpassing beyond lateral margin of elytron. Each tibia dorsally with three complete carinae. Terminal tarsomere thickened, slightly arched, distinctly longer than combined length of remaining tarsomeres. Pretarsal claws with short empodium. Male last visible tergite yellowish, broadly</paragraph>
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<caption id="4CD2E3CF5C12E153CF2B591EFCF3AC9C" ID-DOI="http://doi.org/10.5281/zenodo.13203425" ID-Zenodo-Dep="13203425" httpUri="https://zenodo.org/record/13203425/files/figure.png" pageId="3" pageNumber="298" startId="3.[624,663,1056,1080]" targetBox="[624,1113,299,1036]" targetPageId="3" targetType="figure">
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||||
<paragraph id="1812B3475C12E153CF2B591EFCF3AC9C" blockId="3.[624,1103,1056,1172]" pageId="3" pageNumber="298">
|
||||
Fig. 2.
|
||||
<taxonomicName id="DFADC8C45C12E153CFE3591EFC23AC30" authorityName="Telnov" authorityYear="2023" box="[696,1005,1056,1080]" class="Insecta" family="Tenebrionidae" genus="Boletoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="298" phylum="Arthropoda" rank="subSpecies" species="mixtus" status="subsp. nov." subSpecies="dynastes">
|
||||
|
|
@ -134,13 +135,13 @@ paratype female. A – Habitus, dorsal view; B – ditto, lateral view; C – di
|
|||
</caption>
|
||||
<paragraph id="1812B3475C12E153CD295B98FC52A92D" blockId="3.[114,154,1702,1726]" lastBlockId="3.[624,1103,178,293]" pageId="3" pageNumber="298">
|
||||
298 rounded at apical margin. Male last visible ventrite broadly rounded at posterior margin. Aedeagus as in
|
||||
<figureCitation id="8096AFC25C12E153CE795DD1FCABA90F" box="[802,869,239,263]" captionStart="Fig" captionStartId="4.[189,228,898,922]" captionTargetBox="[194,672,340,876]" captionTargetId="figure-426@4.[188,674,327,890]" captionTargetPageId="4" captionText="Fig. 3.Boletoxenus mixtus dynastesssp. nov. holotype male, aedeagus. A – Dorsal view; B – Ventral view; C – Lateral view." pageId="3" pageNumber="298">Fig. 3</figureCitation>
|
||||
<figureCitation id="8096AFC25C12E153CE795DD1FCABA90F" box="[802,869,239,263]" captionStart="Fig" captionStartId="4.[189,228,898,922]" captionTargetBox="[194,672,340,876]" captionTargetId="figure-426@4.[188,674,327,890]" captionTargetPageId="4" captionText="Fig. 3.Boletoxenus mixtus dynastesssp. nov. holotype male, aedeagus. A – Dorsal view; B – Ventral view; C – Lateral view." figureDoi="http://doi.org/10.5281/zenodo.13203427" httpUri="https://zenodo.org/record/13203427/files/figure.png" pageId="3" pageNumber="298">Fig. 3</figureCitation>
|
||||
, apicale of is entire, not bifurcate in dorsal view.
|
||||
</paragraph>
|
||||
<paragraph id="1812B3475C12E154CF2B5987FE67A92D" blockId="3.[624,1103,1209,1632]" lastBlockId="4.[189,668,178,293]" lastPageId="4" lastPageNumber="299" pageId="3" pageNumber="298">
|
||||
<emphasis id="2AD96F555C12E153CF2B5987FC94ACD9" bold="true" box="[624,858,1209,1233]" pageId="3" pageNumber="298">Sexual dimorphism.</emphasis>
|
||||
Female (
|
||||
<figureCitation id="8096AFC25C12E153CEED5984FC38ACDA" box="[950,1014,1210,1234]" captionStart="Fig" captionStartId="3.[624,663,1056,1080]" captionTargetBox="[624,1113,299,1036]" captionTargetId="figure-424@3.[623,1117,293,1050]" captionTargetPageId="3" captionText="Fig. 2.Boletoxenus mixtus dynastesssp. nov. paratype female. A – Habitus, dorsal view; B – ditto, lateral view; C – ditto, anterior view [not to scale]." pageId="3" pageNumber="298">Fig. 2</figureCitation>
|
||||
<figureCitation id="8096AFC25C12E153CEED5984FC38ACDA" box="[950,1014,1210,1234]" captionStart="Fig" captionStartId="3.[624,663,1056,1080]" captionTargetBox="[624,1113,299,1036]" captionTargetId="figure-424@3.[623,1117,293,1050]" captionTargetPageId="3" captionText="Fig. 2.Boletoxenus mixtus dynastesssp. nov. paratype female. A – Habitus, dorsal view; B – ditto, lateral view; C – ditto, anterior view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.13203425" httpUri="https://zenodo.org/record/13203425/files/figure.png" pageId="3" pageNumber="298">Fig. 2</figureCitation>
|
||||
) generally smaller than male. Epistoma with transverse, medially somewhat depressed costa. Frons with two pairs of tubercles at midlength of compound eye and posterior to that; posterior pair stronger prominent dorsad and stronger transverse ridge-like. Antennomeres 5–9 with distinct subtriangular lobe. Paired pronotal process short and broad, moderately widely separated, covered with dense, acutely conical tubercles. No regular rows of tubercles posterior to base of pronotal processes. Lateral margin of elytral more regularly subparallel, comparatively less strongly constricted apicad, stronger rounded at apex. Femur slightly surpassing beyond lateral margin of elytron, its apex visible in dorsal view.
|
||||
</paragraph>
|
||||
<paragraph id="1812B3475C15E154CFE05D8CFBE0A989" blockId="4.[699,1178,178,386]" pageId="4" pageNumber="299">
|
||||
|
|
@ -195,7 +196,7 @@ is described as 1.1 (
|
|||
fig. 8).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="4CD2E3CF5C15E154CDE65EBCFDF2ABDF" pageId="4" pageNumber="299" startId="4.[189,228,898,922]" targetBox="[194,672,340,876]" targetPageId="4" targetType="figure">
|
||||
<caption id="4CD2E3CF5C15E154CDE65EBCFDF2ABDF" ID-DOI="http://doi.org/10.5281/zenodo.13203427" ID-Zenodo-Dep="13203427" httpUri="https://zenodo.org/record/13203427/files/figure.png" pageId="4" pageNumber="299" startId="4.[189,228,898,922]" targetBox="[194,672,340,876]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="1812B3475C15E154CDE65EBCFDF2ABDF" blockId="4.[189,669,898,984]" pageId="4" pageNumber="299">
|
||||
Fig. 3.
|
||||
<taxonomicName id="DFADC8C45C15E154CC5D5EBCFDF4AB92" authorityName="Telnov" authorityYear="2023" box="[262,570,898,922]" class="Insecta" family="Tenebrionidae" genus="Boletoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="299" phylum="Arthropoda" rank="subSpecies" species="mixtus" status="subsp. nov." subSpecies="dynastes">
|
||||
|
|
|
|||
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