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<document id="450CB250B5C6E8EE3EE4A35A60B71EB7" ID-DOI="10.1080/00222930802598409" ID-ISSN="1464-5262" ID-Zenodo-Dep="4651952" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617211462487" checkinUser="carolina" docAuthor="Marusik, Y. M. &amp; Fritzén, N. R." docDate="2009" docId="6111322DDB076F2CFE2810C8FE0CE1CB" docLanguage="en" docName="JNATHIST.43.7-8.411-422.pdf" docOrigin="Journal of Natural History 43 (7 - 8)" docSource="http://dx.doi.org/10.1080/00222930802598409" docStyle="DocumentStyle:FCA295E821436EF9307F50C7AC261B53.5:JNatHist.2008-2012.journal_article.1cover" docStyleId="FCA295E821436EF9307F50C7AC261B53" docStyleName="JNatHist.2008-2012.journal_article.1cover" docStyleVersion="5" docTitle="Pardosa pseudomixta Marusik &amp; Fritzén 2009, n.sp." docType="treatment" docVersion="3" lastPageNumber="417" masterDocId="9D284A55DB056F24FFF11330FFF0E473" masterDocTitle="A new wolf spider species in the Pardosa monticola-group (Araneae: Lycosidae) from Xinjiang, China" masterLastPageNumber="422" masterPageNumber="411" pageNumber="412" updateTime="1698897642781" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="29F28B6AC9CEC08A0110C8968A73DBB5">A new wolf spider species in the Pardosa monticola-group (Araneae: Lycosidae) from Xinjiang, China</mods:title>
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<mods:namePart id="55BC8A5780DE2174301E1349618EEFE3">Marusik, Y. M.</mods:namePart>
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<mods:namePart id="F5FEE76CA9CD0B793D245A807D8EE580">Fritzén, N. R.</mods:namePart>
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<mods:title id="6ABBA25FD641B1951F56540F4D4DF41B">Journal of Natural History</mods:title>
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<mods:date id="935914AA94B9D29F701DBB52077BEED9">2009</mods:date>
<mods:detail id="4A951B71E2A697DC1D93E46895E12B90" type="pubDate">
<mods:number id="6636656B6DABD893DD5AFA9EAFF46749">2009-02-28</mods:number>
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<mods:detail id="69A57080A1110CDCD67C1A6AA00BECBD" type="volume">
<mods:number id="547FC65BFA2C2A242E3412FB0D918873">43</mods:number>
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<mods:detail id="1672ABA1EA6FD71BE15A6B0C47FAC852" type="issue">
<mods:number id="A3419ECE559E61591E3A7D5088F43C1B">7 - 8</mods:number>
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<mods:start id="F417B334BDDAE7D0A646C67CB470407F">411</mods:start>
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<mods:classification id="54FECFFEF925D23F0C11A0A8F55C30AC">journal article</mods:classification>
<mods:identifier id="4B469614591E6936603D84994B521276" type="DOI">10.1080/00222930802598409</mods:identifier>
<mods:identifier id="38C279BC093E5D5BF70946BADC0D0DFB" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="C11A425FE0F8D1CC9F35A3B511AA9BE9" type="Zenodo-Dep">4651952</mods:identifier>
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<treatment id="6111322DDB076F2CFE2810C8FE0CE1CB" LSID="urn:lsid:plazi:treatment:6111322DDB076F2CFE2810C8FE0CE1CB" httpUri="http://treatment.plazi.org/id/6111322DDB076F2CFE2810C8FE0CE1CB" lastPageId="8" lastPageNumber="417" pageId="2" pageNumber="412">
<subSubSection id="A1A2D0B0DB076F26FE2810C8FCF8E063" box="[473,776,1016,1040]" pageId="2" pageNumber="412" type="nomenclature">
<paragraph id="E907833BDB076F26FE2810C8FCF8E063" blockId="2.[306,943,1016,1072]" box="[473,776,1016,1040]" pageId="2" pageNumber="412">
<heading id="B24F3457DB076F26FE2810C8FCF8E063" box="[473,776,1016,1040]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="412" reason="3">
<taxonomicName id="2EB8F8B8DB076F26FE2810C8FD39E063" ID-CoL="4DJZF" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[473,713,1016,1040]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="412" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB076F26FE2810C8FD39E063" bold="true" box="[473,713,1016,1040]" italics="true" pageId="2" pageNumber="412">Pardosa pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB076F26FD2310C8FCF8E063" box="[722,776,1016,1040]" pageId="2" pageNumber="412" rank="species">n.sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1A2D0B0DB076F26FEC31728FC5FE043" box="[306,943,1048,1072]" pageId="2" pageNumber="412" type="description">
<paragraph id="E907833BDB076F26FEC31728FC5FE043" blockId="2.[306,943,1016,1072]" box="[306,943,1048,1072]" pageId="2" pageNumber="412">
(
<figureCitation id="71839FBEDB076F26FEC81728FE6EE043" box="[313,414,1048,1072]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="2" pageNumber="412">Figures 1</figureCitation>
A–C,E,F; 2A,B,F–J; 3B–D,G; and 4B,E–G,J)
</paragraph>
</subSubSection>
<subSubSection id="A1A2D0B0DB076F26FF63177BFC4EE0F0" pageId="2" pageNumber="412" type="reference_group">
<paragraph id="E907833BDB076F26FF63177BFC4EE0F0" blockId="2.[146,1072,1098,1155]" pageId="2" pageNumber="412">
<taxonomicName id="2EB8F8B8DB076F26FF63177BFD7DE011" ID-CoL="4DJVZ" authority=": Chen and Song 2002: p. 342" authorityName="Chen and Song" authorityPageNumber="342" authorityYear="2002" box="[146,653,1098,1123]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="412" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB076F26FF63177BFEC7E010" box="[146,311,1099,1123]" italics="true" pageId="2" pageNumber="412">Pardosa mixta</emphasis>
:
<bibRefCitation id="8D29FECADB076F26FEB6177AFDCBE011" author="Chen J &amp; Song DX" box="[327,571,1098,1122]" pageId="2" pageNumber="412" pagination="341 - 344" refId="ref4077" refString="Chen J, Song DX. 2002. A new species and a new record of monticola group of the genus Pardosa from China (Araneae, Lycosidae). J Shanxi Univ (Nat Sci). 25: 341 - 344. Chinese." type="journal article" year="2002">Chen and Song 2002</bibRefCitation>
: p. 342
</taxonomicName>
, figures 4–11 (
<emphasis id="DBCC5F29DB076F26FCC0177BFCB3E011" box="[817,835,1099,1122]" italics="true" pageId="2" pageNumber="412">„</emphasis>
ǫ, misidentification).
<taxonomicName id="2EB8F8B8DB076F26FF63175BFD87E0F0" ID-CoL="4DJVZ" authority="sensu Chen and Song 2002" authorityName="sensu Chen and Song" authorityYear="2002" box="[146,631,1131,1155]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="412" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB076F26FF63175BFE8BE0F0" box="[146,379,1131,1155]" italics="true" pageId="2" pageNumber="412">Pardosa mixta sensu</emphasis>
<bibRefCitation id="8D29FECADB076F26FE75175BFD87E0F0" author="Chen J &amp; Song DX" box="[388,631,1131,1155]" pageId="2" pageNumber="412" pagination="341 - 344" refId="ref4077" refString="Chen J, Song DX. 2002. A new species and a new record of monticola group of the genus Pardosa from China (Araneae, Lycosidae). J Shanxi Univ (Nat Sci). 25: 341 - 344. Chinese." type="journal article" year="2002">Chen and Song 2002</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="8D29FECADB076F26FD75175BFC97E0F0" author="Marusik YM &amp; Fritzen NR &amp; Song DX" box="[644,871,1131,1155]" pageId="2" pageNumber="412" pagination="259 - 276" refId="ref4327" refString="Marusik YM, Fritzen NR, Song DX. 2007. On spiders (Aranei) collected in central Xinjiang, China. Arthropod Select. 15: 259 - 276." type="journal article" year="2007">Marusik et al. 2007</bibRefCitation>
, p. 268.
</paragraph>
</subSubSection>
<subSubSection id="A1A2D0B0DB076F23FF63178BFEC8E212" lastPageId="7" lastPageNumber="417" pageId="2" pageNumber="412" type="materials_examined">
<paragraph id="E907833BDB076F26FF63178BFEDFE0A0" blockId="2.[146,303,1211,1235]" box="[146,303,1211,1235]" pageId="2" pageNumber="412">
<emphasis id="DBCC5F29DB076F26FF63178BFEDFE0A0" box="[146,303,1211,1235]" italics="true" pageId="2" pageNumber="412">Type material</emphasis>
</paragraph>
<paragraph id="E907833BDB076F26FF6317D3FC3DE149" blockId="2.[146,1103,1250,1338]" pageId="2" pageNumber="412">
<materialsCitation id="59D08966DB076F26FF6317D3FC3DE149" collectingDate="2004-05-03" collectingDateMax="2004-06-28" collectingDateMin="2004-05-03" collectionCode="ZIB, ZMUT, ZMMU, R" country="China" elevation="1950" latitude="43.438" location="Nantaizi" longLatPrecision="68" longitude="87.262" municipality="Urumqi" pageId="2" pageNumber="412" specimenCount="2" stateProvince="Xinjiang Uygur" typeStatus="holotype">
<typeStatus id="36033D99DB076F26FF6317D3FF0DE088" box="[146,253,1251,1275]" pageId="2" pageNumber="412" type="holotype">Holotype</typeStatus>
<emphasis id="DBCC5F29DB076F26FEF717D3FEE8E089" box="[262,280,1251,1274]" italics="true" pageId="2" pageNumber="412">„</emphasis>
(
<collectionCode id="8FA91BFEDB076F26FED817D3FEA9E088" box="[297,345,1251,1275]" pageId="2" pageNumber="412">ZIB</collectionCode>
) and
<typeStatus id="36033D99DB076F26FE6C17D3FDFBE088" box="[413,523,1251,1275]" pageId="2" pageNumber="412" type="paratype">paratypes</typeStatus>
19
<emphasis id="DBCC5F29DB076F26FDC817D3FDACE089" box="[569,604,1251,1274]" italics="true" pageId="2" pageNumber="412">„„</emphasis>
and 21 ǫǫ (
<collectionCode id="8FA91BFEDB076F26FD1B17D3FCEBE088" box="[746,795,1251,1275]" pageId="2" pageNumber="412">ZIB</collectionCode>
,
<collectionCode id="8FA91BFEDB076F26FCD817D3FC70E088" box="[809,896,1251,1275]" country="Japan" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="2" pageNumber="412">ZMUT</collectionCode>
,
<collectionCode id="8FA91BFEDB076F26FC7C17D3FC1DE088" box="[909,1005,1251,1275]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="412" type="Museum">ZMMU</collectionCode>
),
<collectingCountry id="91AFC3ABDB076F26FBF317D3FBBAE088" box="[1026,1098,1251,1275]" name="China" pageId="2" pageNumber="412">China</collectingCountry>
,
<collectingRegion id="2B7C4DD9DB076F26FF631633FF07E168" box="[146,247,1283,1307]" country="China" name="Xinjiang Uygur" pageId="2" pageNumber="412">Xinjiang</collectingRegion>
,
<quantity id="2E402EDEDB076F26FEF51633FEB8E168" box="[260,328,1283,1307]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="412" unit="km" value="70.0">70 km</quantity>
southwest of
<collectingMunicipality id="09631941DB076F26FE181633FDB7E168" box="[489,583,1283,1307]" pageId="2" pageNumber="412">Urumqi</collectingMunicipality>
,
<location id="EC67D5E0DB076F26FDA21633FD48E168" LSID="urn:lsid:plazi:treatment:6111322DDB076F2CFE2810C8FE0CE1CB:EC67D5E0DB076F26FDA21633FD48E168" box="[595,696,1283,1307]" country="China" latitude="43.438" longLatPrecision="68" longitude="87.262" municipality="Urumqi" name="Nantaizi" pageId="2" pageNumber="412" stateProvince="Xinjiang Uygur">Nantaizi</location>
,
<geoCoordinate id="8C8CE5FCDB076F26FD341633FCC7E168" box="[709,823,1283,1307]" degrees="43.399" direction="north" orientation="latitude" pageId="2" pageNumber="412" precision="55" value="43.399">
43.399
<emphasis id="DBCC5F29DB076F26FCFF1633FCE8E168" bold="true" box="[782,792,1283,1307]" italics="true" pageId="2" pageNumber="412">°</emphasis>
N
</geoCoordinate>
to
<geoCoordinate id="8C8CE5FCDB076F26FCAE1633FC23E168" box="[863,979,1283,1307]" degrees="43.438" direction="north" orientation="latitude" pageId="2" pageNumber="412" precision="55" value="43.438">
43.438
<emphasis id="DBCC5F29DB076F26FC581633FC43E168" bold="true" box="[937,947,1283,1307]" italics="true" pageId="2" pageNumber="412">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="8C8CE5FCDB076F26FC111633FBBFE168" box="[992,1103,1283,1307]" degrees="87.214" direction="east" orientation="longitude" pageId="2" pageNumber="412" precision="55" value="87.214">
87.214
<emphasis id="DBCC5F29DB076F26FBDB1633FBC4E168" bold="true" box="[1066,1076,1283,1307]" italics="true" pageId="2" pageNumber="412">°</emphasis>
E
</geoCoordinate>
to
<geoCoordinate id="8C8CE5FCDB076F26FF421612FED3E149" box="[179,291,1314,1338]" degrees="87.262" direction="east" orientation="longitude" pageId="2" pageNumber="412" precision="55" value="87.262">
87.262
<emphasis id="DBCC5F29DB076F26FF0D1612FEF6E149" bold="true" box="[252,262,1314,1338]" italics="true" pageId="2" pageNumber="412">°</emphasis>
E
</geoCoordinate>
,
<quantity id="2E402EDEDB076F26FEC01612FE36E149" box="[305,454,1314,1338]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="2" pageNumber="412" unit="m" value="1950.0" valueMax="2100.0" valueMin="1800.0">
<elevation id="62956408DB076F26FEC01612FE36E149" box="[305,454,1314,1338]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="2" pageNumber="412" unit="m" value="1950.0" valueMax="2100.0" valueMin="1800.0">1800–2100 m</elevation>
</quantity>
,
<date id="9D06A5FBDB076F26FE221612FD2EE149" box="[467,734,1314,1338]" pageId="2" pageNumber="412" value="2004-05-03" valueMax="2004-06-28" valueMin="2004-05-03">
<collectingDate id="8D425C13DB076F26FE221612FD2EE149" box="[467,734,1314,1338]" pageId="2" pageNumber="412" value="2004-05-03" valueMax="2004-06-28" valueMin="2004-05-03">3 May to 28 June 2004</collectingDate>
</date>
(coll. N.
<collectionCode id="8FA91BFEDB076F26FCB71612FCADE149" box="[838,861,1314,1338]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="412">R</collectionCode>
. Fritzén).
</materialsCitation>
</paragraph>
<paragraph id="E907833BDB076F26FF631643FEFBE1F8" blockId="2.[146,267,1395,1419]" box="[146,267,1395,1419]" pageId="2" pageNumber="412">
<emphasis id="DBCC5F29DB076F26FF631643FEFBE1F8" box="[146,267,1395,1419]" italics="true" pageId="2" pageNumber="412">Etymology</emphasis>
</paragraph>
<paragraph id="E907833BDB076F26FF6316AAFDBDE1A1" blockId="2.[146,1103,1434,1491]" pageId="2" pageNumber="412">
The specific name is an adjective in apposition and reflects the close relationship between the new species and
<taxonomicName id="2EB8F8B8DB076F26FE12168BFDB5E1A0" authorityName="sensu Chen and Song" authorityYear="2002" box="[483,581,1467,1491]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="412" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB076F26FE12168BFDB5E1A0" box="[483,581,1467,1491]" italics="true" pageId="2" pageNumber="412">P. mixta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E907833BDB076F26FF63153BFF0FE250" blockId="2.[146,255,1547,1571]" box="[146,255,1547,1571]" pageId="2" pageNumber="412">
<emphasis id="DBCC5F29DB076F26FF63153BFF0FE250" box="[146,255,1547,1571]" italics="true" pageId="2" pageNumber="412">Diagnosis</emphasis>
</paragraph>
<paragraph id="E907833BDB076F22FF631503FF1CE199" blockId="2.[146,1103,1586,1706]" lastBlockId="6.[146,1103,1231,1514]" lastPageId="6" lastPageNumber="416" pageId="2" pageNumber="412">
<taxonomicName id="2EB8F8B8DB076F26FF631503FE70E238" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[146,384,1587,1611]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="412" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB076F26FF631503FE70E238" box="[146,384,1587,1611]" italics="true" pageId="2" pageNumber="412">Pardosa pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB076F26FE771502FE2FE239" box="[390,479,1586,1610]" pageId="2" pageNumber="412" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to
<taxonomicName id="2EB8F8B8DB076F26FD541503FCF5E238" authorityName="sensu Chen and Song" authorityYear="2002" box="[677,773,1587,1611]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="412" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB076F26FD541503FCF5E238" box="[677,773,1587,1611]" italics="true" pageId="2" pageNumber="412">P. mixta</emphasis>
</taxonomicName>
known from the Alps. Males of the two species can easily be separated by the unmodified tarsi and metatarsi of leg I (
<figureCitation id="71839FBEDB076F26FF241542FE99E2F9" box="[213,361,1650,1674]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="2" pageNumber="412">Figures 1E,F</figureCitation>
;
<figureCitation id="71839FBEDB076F26FE861542FE42E2F9" box="[375,434,1650,1674]" captionStart="Figure 2" captionStartId="4.[146,213,1558,1580]" captionTargetBox="[195,1053,165,1522]" captionTargetId="figure-1@4.[195,1054,165,1522]" captionTargetPageId="4" captionText="Figure 2. Males of Pardosa pseudomixta sp. nov. (A,B,F–J) and Pardosa mixta (C–E). (A,C) leg I, prolateral; (B,D) metatarsus I, prolateral; (E,F,I) palp ventral; (G) palp retrolateral; (H) palp from above; (J) fine pore on metatarsus I (arrowed). Arrows indicate differences in the setae arrangement on the metatarsi between both species and fine pore. Scale 0.1 mm if not otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4651958" httpUri="https://zenodo.org/record/4651958/files/figure.png" pageId="2" pageNumber="412">2A,B</figureCitation>
) (swollen and with pubescence in
<taxonomicName id="2EB8F8B8DB076F26FCCD1542FC6EE2F9" authorityName="sensu Chen and Song" authorityYear="2002" box="[828,926,1650,1674]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="412" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB076F26FCCD1542FC6EE2F9" box="[828,926,1650,1674]" italics="true" pageId="2" pageNumber="412">P. mixta</emphasis>
</taxonomicName>
;
<figureCitation id="71839FBEDB076F26FC5C1542FBB9E2F9" box="[941,1097,1650,1674]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="2" pageNumber="412">Figures 1G,H</figureCitation>
;
<figureCitation id="71839FBEDB076F26FF6315A2FF22E2D9" box="[146,210,1682,1706]" captionStart="Figure 2" captionStartId="4.[146,213,1558,1580]" captionTargetBox="[195,1053,165,1522]" captionTargetId="figure-1@4.[195,1054,165,1522]" captionTargetPageId="4" captionText="Figure 2. Males of Pardosa pseudomixta sp. nov. (A,B,F–J) and Pardosa mixta (C–E). (A,C) leg I, prolateral; (B,D) metatarsus I, prolateral; (E,F,I) palp ventral; (G) palp retrolateral; (H) palp from above; (J) fine pore on metatarsus I (arrowed). Arrows indicate differences in the setae arrangement on the metatarsi between both species and fine pore. Scale 0.1 mm if not otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4651958" httpUri="https://zenodo.org/record/4651958/files/figure.png" pageId="2" pageNumber="412">2C,D</figureCitation>
), the black tip of tarsus I, and details of the embolic division of the pedipalp (compare
<figureCitation id="71839FBEDB036F22FEF917FFFE93E094" box="[264,355,1231,1255]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="6" pageNumber="416">Figure 3</figureCitation>
B–D with
<figureCitation id="71839FBEDB036F22FE1017FFFD77E094" box="[481,647,1231,1255]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="6" pageNumber="416">Figure 3A,E,F</figureCitation>
). In addition, male
<taxonomicName id="2EB8F8B8DB036F22FC8517FFFBD4E094" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[884,1060,1231,1255]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="416" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB036F22FC8517FFFBD4E094" box="[884,1060,1231,1255]" italics="true" pageId="6" pageNumber="416">P. pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB036F22FBDF17FFFF34E174" pageId="6" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
have brown lateral bands on the carapace (
<figureCitation id="71839FBEDB036F22FD4817DFFCDBE174" box="[697,811,1263,1287]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="6" pageNumber="416">Figure 1C</figureCitation>
), whereas
<taxonomicName id="2EB8F8B8DB036F22FC5417C0FBF6E17B" authorityName="sensu Chen and Song" authorityYear="2002" box="[933,1030,1264,1288]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="416" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB036F22FC5417C0FBF6E17B" box="[933,1030,1264,1288]" italics="true" pageId="6" pageNumber="416">P. mixta</emphasis>
</taxonomicName>
males have yellow lateral bands (
<figureCitation id="71839FBEDB036F22FE3C1620FDB2E15B" box="[461,578,1296,1320]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="6" pageNumber="416">Figure 1D</figureCitation>
). Females of these sibling species are almost indistinguishable by the shape of the epigyne, although its apical pockets are almost fused in
<taxonomicName id="2EB8F8B8DB036F22FF031661FEA1E11A" authorityName="sensu Chen and Song" authorityYear="2002" box="[242,337,1361,1385]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="416" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB036F22FF031661FEA1E11A" box="[242,337,1361,1385]" italics="true" pageId="6" pageNumber="416">P. mixta</emphasis>
</taxonomicName>
(
<figureCitation id="71839FBEDB036F22FE911661FDEEE11A" box="[352,542,1361,1385]" captionStart="Figure 4" captionStartId="6.[146,213,1108,1130]" captionTargetBox="[146,1103,166,1071]" captionTargetId="figure-148@6.[146,1103,166,1072]" captionTargetPageId="6" captionText="Figure 4. Pardosa mixta (A,C,D,H,I) and Pardosa pseudomixta sp. nov. (B,E–G,J). (A,B,I,J) epigyne after maceration, ventral; (C) embolic division, ventral; (D–H) epigyne, ventral view. Analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651966" httpUri="https://zenodo.org/record/4651966/files/figure.png" pageId="6" pageNumber="416">Figure 4A,D,H,I</figureCitation>
) while in the new species the inner margins of the apical pockets are clearly separated (
<figureCitation id="71839FBEDB036F22FDC31641FD42E1FA" box="[562,690,1393,1417]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="6" pageNumber="416">Figures 3G</figureCitation>
;
<figureCitation id="71839FBEDB036F22FD4F1641FD10E1FA" box="[702,736,1393,1417]" captionStart="Figure 4" captionStartId="6.[146,213,1108,1130]" captionTargetBox="[146,1103,166,1071]" captionTargetId="figure-148@6.[146,1103,166,1072]" captionTargetPageId="6" captionText="Figure 4. Pardosa mixta (A,C,D,H,I) and Pardosa pseudomixta sp. nov. (B,E–G,J). (A,B,I,J) epigyne after maceration, ventral; (C) embolic division, ventral; (D–H) epigyne, ventral view. Analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651966" httpUri="https://zenodo.org/record/4651966/files/figure.png" pageId="6" pageNumber="416">4B</figureCitation>
,E–G,J). In addition, the median septum in
<taxonomicName id="2EB8F8B8DB036F22FEF616A2FE43E1D9" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[263,435,1426,1450]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="416" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB036F22FEF616A2FE43E1D9" box="[263,435,1426,1450]" italics="true" pageId="6" pageNumber="416">P. pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB036F22FE4816A1FDE2E1DA" box="[441,530,1425,1449]" pageId="6" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
does not have a longitudinal groove as in
<taxonomicName id="2EB8F8B8DB036F22FC1F16A2FBBEE1D9" authorityName="sensu Chen and Song" authorityYear="2002" box="[1006,1102,1426,1450]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="416" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB036F22FC1F16A2FBBEE1D9" box="[1006,1102,1426,1450]" italics="true" pageId="6" pageNumber="416">P. mixta</emphasis>
</taxonomicName>
(
<figureCitation id="71839FBEDB036F22FF6B1682FED2E1B9" box="[154,290,1458,1482]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="6" pageNumber="416">Figure 3H,I</figureCitation>
). The main differences between the two species are summarized in
<tableCitation id="A43AB680DB036F22FF6316E2FF16E199" box="[146,230,1490,1514]" captionStart="Table 1" captionStartId="2.[146,204,178,200]" captionText="Table 1. Main differences between sibling Pardosa pseudomixta sp. nov. and Pardosa mixta" pageId="6" pageNumber="416">Table 1</tableCitation>
.
</paragraph>
<caption id="BDC7D3B3DB066F27FF241577FC3AE2C7" ID-DOI="http://doi.org/10.5281/zenodo.4651954" ID-Zenodo-Dep="4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="3" pageNumber="413" startId="3.[213,280,1607,1629]" targetBox="[213,1170,166,1570]" targetPageId="3">
<paragraph id="E907833BDB066F27FF241577FC3AE2C7" blockId="3.[213,1169,1607,1716]" pageId="3" pageNumber="413">
Figure 1.
<taxonomicName id="2EB8F8B8DB066F27FECC1577FDE4E22F" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[317,532,1607,1628]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="413" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB066F27FECC1577FDE4E22F" box="[317,532,1607,1628]" italics="true" pageId="3" pageNumber="413">Pardosa pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB066F27FDED1577FD9EE22E" box="[540,622,1607,1629]" pageId="3" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
(A–C,E,F) and
<taxonomicName id="2EB8F8B8DB066F27FCE41577FC59E22F" authorityName="sensu Chen and Song" authorityYear="2002" box="[789,937,1607,1628]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="413" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB066F27FCE41577FC59E22F" box="[789,937,1607,1628]" italics="true" pageId="3" pageNumber="413">Pardosa mixta</emphasis>
</taxonomicName>
(D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification.
</paragraph>
</caption>
<caption id="BDC7D3B3DB016F20FF631526FE90E2D2" ID-DOI="http://doi.org/10.5281/zenodo.4651958" ID-Zenodo-Dep="4651958" httpUri="https://zenodo.org/record/4651958/files/figure.png" pageId="4" pageNumber="414" startId="4.[146,213,1558,1580]" targetBox="[195,1053,165,1522]" targetPageId="4">
<paragraph id="E907833BDB016F20FF631526FE90E2D2" blockId="4.[146,1103,1558,1697]" pageId="4" pageNumber="414">
Figure 2. Males of
<taxonomicName id="2EB8F8B8DB016F20FEAC1527FDC3E25F" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[349,563,1559,1580]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="414" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB016F20FEAC1527FDC3E25F" box="[349,563,1559,1580]" italics="true" pageId="4" pageNumber="414">Pardosa pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB016F20FDCA1526FD7DE25F" box="[571,653,1558,1580]" pageId="4" pageNumber="414" rank="species">sp. nov.</taxonomicNameLabel>
(A,B,F–J) and
<taxonomicName id="2EB8F8B8DB016F20FCDE1527FC32E25F" authorityName="sensu Chen and Song" authorityYear="2002" box="[815,962,1559,1580]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="414" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB016F20FCDE1527FC32E25F" box="[815,962,1559,1580]" italics="true" pageId="4" pageNumber="414">Pardosa mixta</emphasis>
</taxonomicName>
(C–E). (A,C) leg I, prolateral; (B,D) metatarsus I, prolateral; (E,F,I) palp ventral; (G) palp retrolateral; (H) palp from above; (J) fine pore on metatarsus I (arrowed). Arrows indicate differences in the setae arrangement on the metatarsi between both species and fine pore. Scale 0.1 mm if not otherwise indicated.
</paragraph>
</caption>
<caption id="BDC7D3B3DB006F21FF241526FB9AE2F7" ID-DOI="http://doi.org/10.5281/zenodo.4651962" ID-Zenodo-Dep="4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="5" pageNumber="415" startId="5.[213,280,1558,1580]" targetBox="[251,1131,165,1522]" targetPageId="5">
<paragraph id="E907833BDB006F21FF241526FB9AE2F7" blockId="5.[213,1169,1558,1668]" pageId="5" pageNumber="415">
Figure 3.
<taxonomicName id="2EB8F8B8DB006F21FEB21527FE2BE25F" authorityName="sensu Chen and Song" authorityYear="2002" box="[323,475,1559,1580]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="415" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB006F21FEB21527FE2BE25F" box="[323,475,1559,1580]" italics="true" pageId="5" pageNumber="415">Pardosa mixta</emphasis>
</taxonomicName>
(A,E,F,H,I) and
<taxonomicName id="2EB8F8B8DB006F21FD511527FC8CE25F" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[672,892,1559,1580]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="415" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB006F21FD511527FC8CE25F" box="[672,892,1559,1580]" italics="true" pageId="5" pageNumber="415">Pardosa pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB006F21FC781526FC10E25F" box="[905,992,1558,1580]" pageId="5" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
(B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification.
</paragraph>
</caption>
<caption id="BDC7D3B3DB036F22FF631764FD43E0D7" ID-DOI="http://doi.org/10.5281/zenodo.4651966" ID-Zenodo-Dep="4651966" httpUri="https://zenodo.org/record/4651966/files/figure.png" pageId="6" pageNumber="416" startId="6.[146,213,1108,1130]" targetBox="[146,1103,166,1071]" targetPageId="6">
<paragraph id="E907833BDB036F22FF631764FD43E0D7" blockId="6.[146,1103,1108,1188]" pageId="6" pageNumber="416">
Figure 4.
<taxonomicName id="2EB8F8B8DB036F22FF0A1764FE7FE01A" authorityName="sensu Chen and Song" authorityYear="2002" box="[251,399,1108,1129]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="416" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB036F22FF0A1764FE7FE01A" box="[251,399,1108,1129]" italics="true" pageId="6" pageNumber="416">Pardosa mixta</emphasis>
</taxonomicName>
(A,C,D,H,I) and
<taxonomicName id="2EB8F8B8DB036F22FDB81764FCD0E01A" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[585,800,1108,1129]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="416" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB036F22FDB81764FCD0E01A" box="[585,800,1108,1129]" italics="true" pageId="6" pageNumber="416">Pardosa pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB036F22FCD91764FC8AE019" box="[808,890,1108,1130]" pageId="6" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
(B,E–G,J). (A,B,I,J) epigyne after maceration, ventral; (C) embolic division, ventral; (D–H) epigyne, ventral view. Analogous figures have same scale or magnification.
</paragraph>
</caption>
<paragraph id="E907833BDB036F22FF631513FEE1E248" blockId="6.[146,273,1571,1595]" box="[146,273,1571,1595]" pageId="6" pageNumber="416">
<emphasis id="DBCC5F29DB036F22FF631513FEE1E248" box="[146,273,1571,1595]" italics="true" pageId="6" pageNumber="416">Description</emphasis>
</paragraph>
<paragraph id="E907833BDB036F23FF63157CFE00E5F2" blockId="6.[146,1103,1612,1701]" lastBlockId="7.[213,1169,168,385]" lastPageId="7" lastPageNumber="417" pageId="6" pageNumber="416">
<emphasis id="DBCC5F29DB036F22FF63157CFF27E217" box="[146,215,1612,1636]" italics="true" pageId="6" pageNumber="416">Males</emphasis>
. Carapace brown-black with narrow yellow median band (
<figureCitation id="71839FBEDB036F22FC8C157CFC00E217" box="[893,1008,1612,1636]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="6" pageNumber="416">Figure 1C</figureCitation>
); lateral light bands brown, darker than median band, and slightly narrower than dark marginal bands; clypeus light brown, lighter than head flanks (
<figureCitation id="71839FBEDB036F22FCA615BDFC39E2D6" box="[855,969,1677,1701]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="6" pageNumber="416">Figure 1B</figureCitation>
). Abdomen similar colour to carapace, with light brown median heart mark. Legs yellow without annulations. Femora basally dark. Metatarsi I with a few scattered small pores (
<figureCitation id="71839FBEDB026F23FF2D13D8FEB9E573" box="[220,329,232,256]" captionStart="Figure 2" captionStartId="4.[146,213,1558,1580]" captionTargetBox="[195,1053,165,1522]" captionTargetId="figure-1@4.[195,1054,165,1522]" captionTargetPageId="4" captionText="Figure 2. Males of Pardosa pseudomixta sp. nov. (A,B,F–J) and Pardosa mixta (C–E). (A,C) leg I, prolateral; (B,D) metatarsus I, prolateral; (E,F,I) palp ventral; (G) palp retrolateral; (H) palp from above; (J) fine pore on metatarsus I (arrowed). Arrows indicate differences in the setae arrangement on the metatarsi between both species and fine pore. Scale 0.1 mm if not otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4651958" httpUri="https://zenodo.org/record/4651958/files/figure.png" pageId="7" pageNumber="417">Figure 2J</figureCitation>
). Tarsi I apically dark (
<figureCitation id="71839FBEDB026F23FD8713D8FCF0E573" box="[630,768,232,256]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="7" pageNumber="417">Figure 1E,F</figureCitation>
). Chelicerae of similar colour to carapace, with longitudinal stripes of similar colour as clypeus. Pedipalps dark brown; patellae slightly lighter than other segments. Palea with smoothly rounded ridge (
<figureCitation id="71839FBEDB026F23FED31279FE5EE512" box="[290,430,329,353]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="7" pageNumber="417">Figure 3B,C</figureCitation>
); terminal apophysis rounded (
<figureCitation id="71839FBEDB026F23FCD51279FC20E512" box="[804,976,329,353]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="7" pageNumber="417">Figure 3A,C,D</figureCitation>
), with distinctly separated mesal portion.
</paragraph>
<paragraph id="E907833BDB026F23FF24128AFE6BE6C1" blockId="7.[213,1169,441,690]" pageId="7" pageNumber="417">
<emphasis id="DBCC5F29DB026F23FF24128AFEC0E5A1" box="[213,304,442,466]" italics="true" pageId="7" pageNumber="417">Females</emphasis>
. Carapace brown with wide yellow median band and distinct cephalic spot (
<figureCitation id="71839FBEDB026F23FF2D12E9FEA0E582" box="[220,336,473,497]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="7" pageNumber="417">Figure 1A</figureCitation>
); lateral light bands twice as wide as marginal dark bands. Abdomen with distinct light heart mark and five pairs of light median spots. Anterior spots separated by heart mark, the posterior spots fused. Legs with dark annulations (
<figureCitation id="71839FBEDB026F23FF2D110AFEA1E621" box="[220,337,570,594]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="7" pageNumber="417">Figure 1A</figureCitation>
). Epigyne with wide median septum, that covers the entire cavity and distinct apical pockets (
<figureCitation id="71839FBEDB026F23FE1D116AFDA3E601" box="[492,595,602,626]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="7" pageNumber="417">Figures 3</figureCitation>
G–I; 4A,H); median septum variable in size and shape, its median part without longitudinal groove (
<figureCitation id="71839FBEDB026F23FCA8114AFC3CE6E1" box="[857,972,634,658]" captionStart="Figure 4" captionStartId="6.[146,213,1108,1130]" captionTargetBox="[146,1103,166,1071]" captionTargetId="figure-148@6.[146,1103,166,1072]" captionTargetPageId="6" captionText="Figure 4. Pardosa mixta (A,C,D,H,I) and Pardosa pseudomixta sp. nov. (B,E–G,J). (A,B,I,J) epigyne after maceration, ventral; (C) embolic division, ventral; (D–H) epigyne, ventral view. Analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651966" httpUri="https://zenodo.org/record/4651966/files/figure.png" pageId="7" pageNumber="417">Figure 4E</figureCitation>
); apical pockets widely separated.
</paragraph>
<paragraph id="E907833BDB026F23FF2411DBFE86E770" blockId="7.[213,374,747,771]" box="[213,374,747,771]" pageId="7" pageNumber="417">
<emphasis id="DBCC5F29DB026F23FF2411DBFE86E770" box="[213,374,747,771]" italics="true" pageId="7" pageNumber="417">Measurements</emphasis>
</paragraph>
<paragraph id="E907833BDB026F23FF241024FD94E738" blockId="7.[213,1169,787,843]" pageId="7" pageNumber="417">
<emphasis id="DBCC5F29DB026F23FF241024FE84E75F" box="[213,372,788,812]" italics="true" pageId="7" pageNumber="417">Males/females</emphasis>
. total length 4.9–5.3/5.7–6.1; carapace length 2.58–2.73/2.83–3.03; carapace width 2.0–2.13/2.28–2.33.
</paragraph>
<paragraph id="E907833BDB026F23FF2410B5FE02E7AF" blockId="7.[213,1169,900,988]" pageId="7" pageNumber="417">
<emphasis id="DBCC5F29DB026F23FF2410B5FE80E7EE" box="[213,368,901,925]" italics="true" pageId="7" pageNumber="417">Length ratios</emphasis>
. tibia/metatarsus I 0.87–0.92/0.92–0.97; carapace/metatarsus I 1.4– 1.49/1.63–1.66; carapace/tarsus IV (females only) 1.66–1.67; carapace/metatarsus IV (females only) 0.78–0.81.
</paragraph>
<paragraph id="E907833BDB026F23FF241725FEABE05E" blockId="7.[213,347,1045,1069]" box="[213,347,1045,1069]" pageId="7" pageNumber="417">
<emphasis id="DBCC5F29DB026F23FF241725FEABE05E" box="[213,347,1045,1069]" italics="true" pageId="7" pageNumber="417">Distribution</emphasis>
</paragraph>
<paragraph id="E907833BDB026F23FF24170EFB78E025" blockId="7.[213,1160,1086,1110]" box="[213,1160,1086,1110]" pageId="7" pageNumber="417">
<taxonomicName id="2EB8F8B8DB026F23FF24170EFE35E025" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[213,453,1086,1110]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="417" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB026F23FF24170EFE35E025" box="[213,453,1086,1110]" italics="true" pageId="7" pageNumber="417">Pardosa pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB026F23FE3F170EFDD9E025" box="[462,553,1086,1110]" pageId="7" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
is currently known from
<collectingRegion id="2B7C4DD9DB026F23FCA3170EFC47E025" box="[850,951,1086,1110]" country="China" name="Xinjiang Uygur" pageId="7" pageNumber="417">Xinjiang</collectingRegion>
,
<collectingCountry id="91AFC3ABDB026F23FC34170EFBFAE025" box="[965,1034,1086,1110]" name="China" pageId="7" pageNumber="417">China</collectingCountry>
(
<figureCitation id="71839FBEDB026F23FBEB170EFB8AE025" box="[1050,1146,1086,1110]" captionStart="Figure 5" captionStartId="8.[146,213,711,733]" captionTargetBox="[146,1104,165,675]" captionTargetId="figure-314@8.[146,1104,165,675]" captionTargetPageId="8" captionText="Figure 5. Records of Pardosa pseudomixta sp. nov. (full circles) and Pardosa mixta (full diamonds). The map also shows erroneous records of P. mixta from Uzbekistan (full squares) and presumed records of P. pseudomixta sp. nov. that were reported as P. mixta from China (open circle)." figureDoi="http://doi.org/10.5281/zenodo.4651968" httpUri="https://zenodo.org/record/4651968/files/figure.png" pageId="7" pageNumber="417">Figure 5</figureCitation>
).
</paragraph>
<paragraph id="E907833BDB026F23FF2417BFFE40E0D4" blockId="7.[213,432,1167,1191]" box="[213,432,1167,1191]" pageId="7" pageNumber="417">
<emphasis id="DBCC5F29DB026F23FF2417BFFE40E0D4" box="[213,432,1167,1191]" italics="true" pageId="7" pageNumber="417">Habitat preferences</emphasis>
</paragraph>
<paragraph id="E907833BDB026F23FF241788FBAAE263" blockId="7.[213,1169,1208,1552]" pageId="7" pageNumber="417">
The type material of
<taxonomicName id="2EB8F8B8DB026F23FE121788FD67E0A3" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[483,663,1208,1232]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="417" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB026F23FE121788FD67E0A3" box="[483,663,1208,1232]" italics="true" pageId="7" pageNumber="417">P. pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB026F23FD571788FCF7E0A3" box="[678,775,1208,1232]" pageId="7" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
was collected in the Tian-Shan Mountains about
<quantity id="2E402EDEDB026F23FE4B17E8FE0EE083" box="[442,510,1240,1264]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="417" unit="km" value="70.0">70 km</quantity>
southwest of Urumqi in the
<collectingRegion id="2B7C4DD9DB026F23FC7E17E8FB97E083" box="[911,1127,1240,1264]" country="China" name="Xinjiang Uygur" pageId="7" pageNumber="417">Xinjiang Province</collectingRegion>
in northwestern
<collectingCountry id="91AFC3ABDB026F23FE8B17C8FE33E163" box="[378,451,1272,1296]" name="China" pageId="7" pageNumber="417">China</collectingCountry>
. Most of the material was collected in an area of about
<quantity id="2E402EDEDB026F23FF241628FEE9E143" box="[213,281,1304,1328]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="417" unit="km" value="10.0">10 km</quantity>
<superScript id="1ECD2E73DB026F23FEE91622FED2E151" attach="left" box="[280,290,1298,1314]" fontSize="7" pageId="7" pageNumber="417">2</superScript>
between the hotel in the village of Nantaizi and a water control station. The Nantaizi area is located on the east side of the Urumqi River and consists of half a dozen moraine ridges with relatively steep slopes at elevations ranging from about
<quantity id="2E402EDEDB026F23FF241648FE74E1E3" box="[213,388,1400,1424]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="7" pageNumber="417" unit="m" value="1950.0" valueMax="2100.0" valueMin="1800.0">1800 to 2100 m</quantity>
. Beneath the slopes and near the river there is a flat area. The north facing slopes are dominated by spruce forests, and the south facing slopes mainly by steep, xeric sandy and stony habitats with relatively sparse vegetation or, in places, with a cover of bushes. The west facing slopes and the lower flat areas are mainly dry meadows. The whole area is heavily grazed by goats, sheep, cattle and horses.
</paragraph>
<paragraph id="E907833BDB026F23FF241579FEC8E212" blockId="7.[213,312,1609,1633]" box="[213,312,1609,1633]" pageId="7" pageNumber="417">
<emphasis id="DBCC5F29DB026F23FF241579FEC8E212" box="[213,312,1609,1633]" italics="true" pageId="7" pageNumber="417">Remarks</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A1A2D0B0DB026F2CFF241542FE0CE1CB" lastPageId="8" lastPageNumber="418" pageId="7" pageNumber="417" type="reference_group">
<paragraph id="E907833BDB026F2CFF241542FE0CE1CB" blockId="7.[213,1169,1650,1706]" lastBlockId="8.[146,1103,856,1464]" lastPageId="8" lastPageNumber="418" pageId="7" pageNumber="417">
<taxonomicName id="2EB8F8B8DB026F23FF241542FE33E2F9" ID-CoL="4DJZF" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[213,451,1650,1674]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="417" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB026F23FF241542FE33E2F9" box="[213,451,1650,1674]" italics="true" pageId="7" pageNumber="417">Pardosa pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB026F23FE381542FDD2E2F9" box="[457,546,1650,1674]" pageId="7" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
was reported as
<taxonomicName id="2EB8F8B8DB026F23FD2E1542FBB0E2F9" ID-CoL="4DJVZ" authority="(Chen and Song 2002)" baseAuthorityName="Chen and Song" baseAuthorityYear="2002" box="[735,1088,1650,1674]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="417" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB026F23FD2E1542FCCEE2F9" box="[735,830,1650,1674]" italics="true" pageId="7" pageNumber="417">P. mixta</emphasis>
(
<bibRefCitation id="8D29FECADB026F23FCBC1542FBC9E2F9" author="Chen J &amp; Song DX" box="[845,1081,1650,1674]" pageId="7" pageNumber="417" pagination="341 - 344" refId="ref4077" refString="Chen J, Song DX. 2002. A new species and a new record of monticola group of the genus Pardosa from China (Araneae, Lycosidae). J Shanxi Univ (Nat Sci). 25: 341 - 344. Chinese." type="journal article" year="2002">Chen and Song 2002</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="2EB8F8B8DB026F23FB891542FD84E2D9" ID-CoL="4DJVZ" authority="sensu Chen and Song (2002)" authorityName="sensu Chen and Song" authorityYear="2002" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="417" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB026F23FB891542FEADE2D9" italics="true" pageId="7" pageNumber="417">P. mixta sensu</emphasis>
<bibRefCitation id="8D29FECADB026F23FE9815A2FD84E2D9" author="Chen J &amp; Song DX" box="[361,628,1682,1706]" pageId="7" pageNumber="417" pagination="341 - 344" refId="ref4077" refString="Chen J, Song DX. 2002. A new species and a new record of monticola group of the genus Pardosa from China (Araneae, Lycosidae). J Shanxi Univ (Nat Sci). 25: 341 - 344. Chinese." type="journal article" year="2002">Chen and Song (2002)</bibRefCitation>
</taxonomicName>
from Xinjiang and seems to be restricted to northern Xinjiang.
<taxonomicName id="2EB8F8B8DB0D6F2CFE811069FDE4E702" ID-CoL="4DJVZ" authorityName="sensu Chen and Song" authorityYear="2002" box="[368,532,857,881]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="418" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB0D6F2CFE811069FDE4E702" box="[368,532,857,881]" italics="true" pageId="8" pageNumber="418">Pardosa mixta</emphasis>
</taxonomicName>
was identified and reported from Central Asia at least twice, first by
<bibRefCitation id="8D29FECADB0D6F2CFE751049FDABE7E2" author="Charitonov DE" box="[388,603,889,913]" pageId="8" pageNumber="418" pagination="59 - 133" refId="ref4038" refString="Charitonov DE. 1969. Materials to the fauna of spiders of the USSR. Uchenyye Zapiski Permskogo Ordena Trudovogo Krasnogo Znameni gosudarstvennogo Universiteta imeni A. M. Gorkogo. 179: 59 - 133. Russian." type="journal article" year="1969">Charitonov (1969)</bibRefCitation>
from
<collectingCountry id="91AFC3ABDB0D6F2CFD5D1049FCC0E7E2" box="[684,816,889,913]" name="Uzbekistan" pageId="8" pageNumber="418">Uzbekistan</collectingCountry>
, then by
<bibRefCitation id="8D29FECADB0D6F2CFC581049FEAEE7C2" author="Bronstein TG &amp; Murtazaev AB" pageId="8" pageNumber="418" pagination="124 - 143" refId="ref3958" refString="Bronstein TG, Murtazaev AB. 1974. Materials to the spider fauna of Samarkand Area. Voprosy Zashchity Rasteniy, Trudy Uzbekskogo Universiteta, Novaya Seriya. 247: 124 - 143. Russian." type="journal article" year="1974">Bronstein and Murtazaev (1974)</bibRefCitation>
from the Samarkand area of
<collectingCountry id="91AFC3ABDB0D6F2CFD4D10A9FCCDE7C2" box="[700,829,921,945]" name="Uzbekistan" pageId="8" pageNumber="418">Uzbekistan</collectingCountry>
(
<figureCitation id="71839FBEDB0D6F2CFCBC10A9FC5DE7C2" box="[845,941,921,945]" captionStart="Figure 5" captionStartId="8.[146,213,711,733]" captionTargetBox="[146,1104,165,675]" captionTargetId="figure-314@8.[146,1104,165,675]" captionTargetPageId="8" captionText="Figure 5. Records of Pardosa pseudomixta sp. nov. (full circles) and Pardosa mixta (full diamonds). The map also shows erroneous records of P. mixta from Uzbekistan (full squares) and presumed records of P. pseudomixta sp. nov. that were reported as P. mixta from China (open circle)." figureDoi="http://doi.org/10.5281/zenodo.4651968" httpUri="https://zenodo.org/record/4651968/files/figure.png" pageId="8" pageNumber="418">Figure 5</figureCitation>
). We checked the records of
<bibRefCitation id="8D29FECADB0D6F2CFEB7108AFDECE7A1" author="Charitonov DE" box="[326,540,954,978]" pageId="8" pageNumber="418" pagination="59 - 133" refId="ref4038" refString="Charitonov DE. 1969. Materials to the fauna of spiders of the USSR. Uchenyye Zapiski Permskogo Ordena Trudovogo Krasnogo Znameni gosudarstvennogo Universiteta imeni A. M. Gorkogo. 179: 59 - 133. Russian." type="journal article" year="1969">Charitonov (1969)</bibRefCitation>
for
<taxonomicName id="2EB8F8B8DB0D6F2CFDA6108AFD4CE7A1" ID-CoL="4DJVZ" authorityName="sensu Chen and Song" authorityYear="2002" box="[599,700,954,978]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="418" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB0D6F2CFDA6108AFD4CE7A1" box="[599,700,954,978]" italics="true" pageId="8" pageNumber="418">P. mixta</emphasis>
</taxonomicName>
from
<collectingCountry id="91AFC3ABDB0D6F2CFCFC108AFC62E7A1" box="[781,914,954,978]" name="Uzbekistan" pageId="8" pageNumber="418">Uzbekistan</collectingCountry>
. His specimens, housed in the Perm State University, belong to another, apparently new species. Likewise, the record from Samarkand may refer to another species of the
<taxonomicName id="2EB8F8B8DB0D6F2CFC2810CBFBB6E060" ID-CoL="4DJW6" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[985,1094,1019,1043]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="418" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="DBCC5F29DB0D6F2CFC2810CBFBB6E060" box="[985,1094,1019,1043]" italics="true" pageId="8" pageNumber="418">monticola</emphasis>
</taxonomicName>
- group because
<bibRefCitation id="8D29FECADB0D6F2CFECD172BFD5FE040" author="Bronstein TG &amp; Murtazaev AB" box="[316,687,1051,1075]" pageId="8" pageNumber="418" pagination="124 - 143" refId="ref3958" refString="Bronstein TG, Murtazaev AB. 1974. Materials to the spider fauna of Samarkand Area. Voprosy Zashchity Rasteniy, Trudy Uzbekskogo Universiteta, Novaya Seriya. 247: 124 - 143. Russian." type="journal article" year="1974">Bronstein and Murtazaev (1974)</bibRefCitation>
, as amateur arachnologists, may not have had the appropriate literature available to identify the species. Investigations of other available specimens belonging to the
<taxonomicName id="2EB8F8B8DB0D6F2CFD60176CFCD2E007" ID-CoL="4DJW6" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[657,802,1116,1140]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="418" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="DBCC5F29DB0D6F2CFD60176CFCD2E007" box="[657,802,1116,1140]" italics="true" pageId="8" pageNumber="418">P. monticola</emphasis>
</taxonomicName>
-group from Central Asia (
<collectingCountry id="91AFC3ABDB0D6F2CFF6B174CFED6E0E7" box="[154,294,1148,1172]" name="Kazakhstan" pageId="8" pageNumber="418">Kazakhstan</collectingCountry>
,
<collectingCountry id="91AFC3ABDB0D6F2CFEC6174CFE4DE0E7" box="[311,445,1148,1172]" name="Kyrgyzstan" pageId="8" pageNumber="418">Kyrgyzstan</collectingCountry>
,
<collectingCountry id="91AFC3ABDB0D6F2CFE3D174CFDBDE0E7" box="[460,589,1148,1172]" name="Uzbekistan" pageId="8" pageNumber="418">Uzbekistan</collectingCountry>
and
<collectingCountry id="91AFC3ABDB0D6F2CFD7E174CFCC0E0E7" box="[655,816,1148,1172]" name="Turkmenistan" pageId="8" pageNumber="418">Turkmenistan</collectingCountry>
) revealed that there are several species, of which at least three are related to
<taxonomicName id="2EB8F8B8DB0D6F2CFD1317ADFCB2E0C6" ID-CoL="4DJVZ" authorityName="sensu Chen and Song" authorityYear="2002" box="[738,834,1181,1205]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="418" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB0D6F2CFD1317ADFCB2E0C6" box="[738,834,1181,1205]" italics="true" pageId="8" pageNumber="418">P. mixta</emphasis>
</taxonomicName>
and
<taxonomicName id="2EB8F8B8DB0D6F2CFC8A17ADFBD7E0C6" ID-CoL="4DJZF" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[891,1063,1181,1205]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="418" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB0D6F2CFC8A17ADFBD7E0C6" box="[891,1063,1181,1205]" italics="true" pageId="8" pageNumber="418">P. pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB0D6F2CFBDF17ACFF34E0A6" pageId="8" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
Males of these species exhibit only minor differences in the shape of the terminal apophysis but much better diagnostic characters in the shape, colour and lack of pubescence on the metatarsi and tarsi of leg I. A separate paper devoted to the study of Central Asian members of
<taxonomicName id="2EB8F8B8DB0D6F2CFDAA162EFD19E145" ID-CoL="4DJW6" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[603,745,1310,1334]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="418" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="DBCC5F29DB0D6F2CFDAA162EFD19E145" box="[603,745,1310,1334]" italics="true" pageId="8" pageNumber="418">P. monticola</emphasis>
</taxonomicName>
-group is forthcoming. Judging from the localized distribution of the species of this group in Central Asia, it seems likely that specimens from Xinjiang and Gansu (
<figureCitation id="71839FBEDB0D6F2CFD49166FFCE7E104" box="[696,791,1375,1399]" captionStart="Figure 5" captionStartId="8.[146,213,711,733]" captionTargetBox="[146,1104,165,675]" captionTargetId="figure-314@8.[146,1104,165,675]" captionTargetPageId="8" captionText="Figure 5. Records of Pardosa pseudomixta sp. nov. (full circles) and Pardosa mixta (full diamonds). The map also shows erroneous records of P. mixta from Uzbekistan (full squares) and presumed records of P. pseudomixta sp. nov. that were reported as P. mixta from China (open circle)." figureDoi="http://doi.org/10.5281/zenodo.4651968" httpUri="https://zenodo.org/record/4651968/files/figure.png" pageId="8" pageNumber="418">Figure 5</figureCitation>
) treated by
<bibRefCitation id="8D29FECADB0D6F2CFC51166FFF29E1E4" author="Chen J &amp; Song DX" pageId="8" pageNumber="418" pagination="341 - 344" refId="ref4077" refString="Chen J, Song DX. 2002. A new species and a new record of monticola group of the genus Pardosa from China (Araneae, Lycosidae). J Shanxi Univ (Nat Sci). 25: 341 - 344. Chinese." type="journal article" year="2002">Chen and Song (2002)</bibRefCitation>
may also belong to a different species. Unfortunately we were not able to restudy specimens from Gansu.
</paragraph>
</subSubSection>
</treatment>
</document>

272
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<paragraph id="E907833BDB0D6F2CFE5F16FEFCC3E195" blockId="8.[335,914,1485,1542]" box="[430,819,1485,1510]" pageId="8" pageNumber="418">
<heading id="B24F3457DB0D6F2CFE5F16FEFCC3E195" box="[430,819,1485,1510]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="418" reason="3">
<taxonomicName id="2EB8F8B8DB0D6F2CFE5F16FEFCC3E195" ID-CoL="4DJVZ" authority="(Kulczynski, 1887)" authorityName="Alps" baseAuthorityName="Kulczynski" baseAuthorityYear="1887" box="[430,819,1485,1510]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="418" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB0D6F2CFE5F16FEFDA3E195" bold="true" box="[430,595,1486,1510]" italics="true" pageId="8" pageNumber="418">Pardosa mixta</emphasis>
(
<bibRefCitation id="8D29FECADB0D6F2CFD9516FDFCDBE195" author="Kulczynski W" box="[612,811,1485,1510]" pageId="8" pageNumber="418" pagination="245 - 356" refId="ref4251" refString="Kulczynski W. 1887. Przyczynek do tyrolskiej fauny pajeczakow. Rozpravy i sprawozdanje wydzyalu matematyczno-przyrodnyczego. Akademia umiejentnoszczi. Krakowie. 16: 245 - 356." type="journal article" year="1887">Kulczyński, 1887</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1A2D0B0DB0D6F2DFEBE16DEFC3DE2D9" lastPageId="9" lastPageNumber="419" pageId="8" pageNumber="418" type="description">
<paragraph id="E907833BDB0D6F2CFEBE16DEFC62E275" blockId="8.[335,914,1485,1542]" box="[335,914,1518,1542]" pageId="8" pageNumber="418">
(
<figureCitation id="71839FBEDB0D6F2CFEA616DEFDE1E275" box="[343,529,1518,1542]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="8" pageNumber="418">Figures 1D,G,H</figureCitation>
;
<figureCitation id="71839FBEDB0D6F2CFDEE16DEFDDDE275" box="[543,557,1518,1542]" captionStart="Figure 2" captionStartId="4.[146,213,1558,1580]" captionTargetBox="[195,1053,165,1522]" captionTargetId="figure-1@4.[195,1054,165,1522]" captionTargetPageId="4" captionText="Figure 2. Males of Pardosa pseudomixta sp. nov. (A,B,F–J) and Pardosa mixta (C–E). (A,C) leg I, prolateral; (B,D) metatarsus I, prolateral; (E,F,I) palp ventral; (G) palp retrolateral; (H) palp from above; (J) fine pore on metatarsus I (arrowed). Arrows indicate differences in the setae arrangement on the metatarsi between both species and fine pore. Scale 0.1 mm if not otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4651958" httpUri="https://zenodo.org/record/4651958/files/figure.png" pageId="8" pageNumber="418">2</figureCitation>
C–E; 3A,E,F,H,I; 4A,C,D,H,I)
</paragraph>
<paragraph id="E907833BDB0D6F2CFF63151AFC79E231" blockId="8.[146,905,1577,1701]" box="[146,905,1577,1602]" pageId="8" pageNumber="418">
<emphasis id="DBCC5F29DB0D6F2CFF63151AFEF6E231" box="[146,262,1578,1602]" italics="true" pageId="8" pageNumber="418">
<taxonomicName id="2EB8F8B8DB0D6F2CFF63151AFF11E231" box="[146,225,1578,1602]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="418" phylum="Arthropoda" rank="genus">Lycosa</taxonomicName>
m.
</emphasis>
<bibRefCitation id="8D29FECADB0D6F2CFEFE1519FE27E231" author="Kulczynski W" box="[271,471,1577,1602]" pageId="8" pageNumber="418" pagination="245 - 356" refId="ref4251" refString="Kulczynski W. 1887. Przyczynek do tyrolskiej fauny pajeczakow. Rozpravy i sprawozdanje wydzyalu matematyczno-przyrodnyczego. Akademia umiejentnoszczi. Krakowie. 16: 245 - 356." type="journal article" year="1887">Kulczyński, 1887</bibRefCitation>
, p. 299, 301, pl. 5, figures 11, 13 (
<emphasis id="DBCC5F29DB0D6F2CFCA8151BFC9BE231" box="[857,875,1579,1602]" italics="true" pageId="8" pageNumber="418">„</emphasis>
ǫ).
</paragraph>
<paragraph id="E907833BDB0D6F2CFF63157BFCBAE210" blockId="8.[146,905,1577,1701]" box="[146,842,1610,1635]" pageId="8" pageNumber="418">
<emphasis id="DBCC5F29DB0D6F2CFF63157BFEF6E210" box="[146,262,1611,1635]" italics="true" pageId="8" pageNumber="418">
<taxonomicName id="2EB8F8B8DB0D6F2CFF63157BFF11E210" box="[146,225,1611,1635]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="418" phylum="Arthropoda" rank="genus">Lycosa</taxonomicName>
m.
</emphasis>
:
<bibRefCitation id="8D29FECADB0D6F2CFEE7157AFE2EE210" author="Kulczynski W" box="[278,478,1610,1635]" pageId="8" pageNumber="418" pagination="427 - 472" refId="ref4279" refString="Kulczynski W. 1909. Fragmenta Arachnologica. XI-XIII. Bull Acad Sci Cracov. 1909: 427 - 472." type="journal article" year="1909">Kulczyński, 1909</bibRefCitation>
, p. 441, pl. 22, figures 9, 15 (
<emphasis id="DBCC5F29DB0D6F2CFCD9157CFCCAE210" box="[808,826,1612,1635]" italics="true" pageId="8" pageNumber="418">„</emphasis>
).
</paragraph>
<paragraph id="E907833BDB0D6F2CFF63155CFCCBE2F7" blockId="8.[146,905,1577,1701]" box="[146,827,1643,1668]" pageId="8" pageNumber="418">
<emphasis id="DBCC5F29DB0D6F2CFF63155CFEF6E2F7" box="[146,262,1644,1668]" italics="true" pageId="8" pageNumber="418">
<taxonomicName id="2EB8F8B8DB0D6F2CFF63155CFF11E2F7" box="[146,225,1644,1668]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="418" phylum="Arthropoda" rank="genus">Lycosa</taxonomicName>
m.
</emphasis>
:
<bibRefCitation id="8D29FECADB0D6F2CFEE7155CFE2AE2F7" author="Kratochvil J" box="[278,474,1644,1668]" pageId="8" pageNumber="418" pagination="10 - 25" refId="ref4178" refString="Kratochvil J. 1935. Araignees nouvelles ou non encore signalees en Yougoslavie. Premiere partie. Folia Zool Hydrobiol. 8: 10 - 25." type="journal article" year="1935">Kratochvil, 1935</bibRefCitation>
, p. 14, 18, figures 2–4, 10 (
<emphasis id="DBCC5F29DB0D6F2CFCFA155CFCEDE2F0" box="[779,797,1644,1667]" italics="true" pageId="8" pageNumber="418">„</emphasis>
ǫ).
</paragraph>
<paragraph id="E907833BDB0D6F2CFF6315BDFC85E2D6" blockId="8.[146,905,1577,1701]" box="[146,885,1676,1701]" pageId="8" pageNumber="418">
<emphasis id="DBCC5F29DB0D6F2CFF6315BDFF20E2D6" box="[146,208,1677,1701]" italics="true" pageId="8" pageNumber="418">P. m.</emphasis>
:
<bibRefCitation id="8D29FECADB0D6F2CFF1115BDFE56E2D6" author="Tongiorgi P" box="[224,422,1677,1701]" pageId="8" pageNumber="418" pagination="275 - 334" refId="ref4541" refString="Tongiorgi P. 1966 a. Italian wolf spiders of the genus Pardosa (Araneae: Lycosidae). Bull Mus Compar Zool Harvard. 134: 275 - 334." type="journal article" year="1966">Tongiorgi, 1966a</bibRefCitation>
, p. 282, figures 80, 81, 93, 104–106 (
<emphasis id="DBCC5F29DB0D6F2CFCB715BDFCA8E2D7" box="[838,856,1677,1700]" italics="true" pageId="8" pageNumber="418">„</emphasis>
ǫ).
</paragraph>
<paragraph id="E907833BDB0C6F2DFF241398FD2EE4B3" blockId="9.[213,993,168,289]" box="[213,734,168,192]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF241398FEE2E4B3" box="[213,274,168,192]" italics="true" pageId="9" pageNumber="419">P. m.</emphasis>
:
<bibRefCitation id="8D29FECADB0C6F2DFED31398FE1AE4B3" author="Tongiorgi P" box="[290,490,168,192]" pageId="9" pageNumber="419" pagination="335 - 359" refId="ref4572" refString="Tongiorgi P. 1966 b. Wolf spiders of the Pardosa monticola group (Araneae: Lycosidae). Bull Mus Compar Zool Harvard. 134: 335 - 359." type="journal article" year="1966">Tongiorgi, 1966b</bibRefCitation>
, p. 342, figure 23 (
<emphasis id="DBCC5F29DB0C6F2DFD4D1399FD3EE4B3" box="[700,718,169,192]" italics="true" pageId="9" pageNumber="419">„</emphasis>
).
</paragraph>
<paragraph id="E907833BDB0C6F2DFF2413F9FC11E493" blockId="9.[213,993,168,289]" box="[213,993,200,225]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF2413F9FEE2E492" box="[213,274,201,225]" italics="true" pageId="9" pageNumber="419">P. m.</emphasis>
:
<bibRefCitation id="8D29FECADB0C6F2DFED313F8FD46E493" author="Fuhn IE &amp; Niculescu-Burlacu F" box="[290,694,200,224]" pageId="9" pageNumber="419" refId="ref4123" refString="Fuhn IE, Niculescu-Burlacu F. 1971. Fauna Republicii Socialiste Romania. Arachnida. Volumul V, Fascicula 3. Fam. Lycosidae. Bucharest: Academia Republicii Socialiste Romania." type="book" year="1971">Fuhn and Niculescu-Burlacu, 1971</bibRefCitation>
, p. 101, figure 43a–d (
<emphasis id="DBCC5F29DB0C6F2DFC4013F9FC33E493" box="[945,963,201,224]" italics="true" pageId="9" pageNumber="419">„</emphasis>
ǫ).
</paragraph>
<paragraph id="E907833BDB0C6F2DFF2413D9FC9BE572" blockId="9.[213,993,168,289]" box="[213,875,232,257]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF2413D9FEE2E572" box="[213,274,233,257]" italics="true" pageId="9" pageNumber="419">P. m.</emphasis>
:
<bibRefCitation id="8D29FECADB0C6F2DFED313D9FDACE572" author="Heimer S &amp; Nentwig W" box="[290,604,233,257]" pageId="9" pageNumber="419" refId="ref4155" refString="Heimer S, Nentwig W. 1991. Spinnen Mitteleuropas: Ein Bestimmungsbuch. Berlin (Germany): Verlag Paul Parey." type="book" year="1991">Heimer and Nentwig, 1991</bibRefCitation>
, p. 324, figure 859 (
<emphasis id="DBCC5F29DB0C6F2DFCCD13D9FCBEE573" box="[828,846,233,256]" italics="true" pageId="9" pageNumber="419">„</emphasis>
ǫ).
</paragraph>
<paragraph id="E907833BDB0C6F2DFF241239FCB3E552" blockId="9.[213,993,168,289]" box="[213,835,265,289]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF241239FEE2E552" box="[213,274,265,289]" italics="true" pageId="9" pageNumber="419">P. m.</emphasis>
:
<bibRefCitation id="8D29FECADB0C6F2DFED31239FD99E552" author="Pirchegger H &amp; Thaler K" box="[290,617,265,289]" pageId="9" pageNumber="419" pagination="47 - 53" refId="ref4359" refString="Pirchegger H, Thaler K. 1999. Zur Unterscheidung der Mannchen von Pardosa blanda (C. L. Koch) und P. torrentum Simon (Araneae, Lycosidae). Mitteil schweiz entomol Gesellsch. 72: 47 - 53." type="journal article" year="1999">Pirchegger and Thaler, 1999</bibRefCitation>
, p. 49, figure 4 (
<emphasis id="DBCC5F29DB0C6F2DFCD0123AFCC3E552" box="[801,819,266,289]" italics="true" pageId="9" pageNumber="419">„</emphasis>
).
</paragraph>
<paragraph id="E907833BDB0C6F2DFF24126AFE59E501" blockId="9.[213,425,346,370]" box="[213,425,346,370]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF24126AFE59E501" box="[213,425,346,370]" italics="true" pageId="9" pageNumber="419">Material examined</emphasis>
</paragraph>
<paragraph id="E907833BDB0C6F2DFF2412B2FDA6E5AA" blockId="9.[213,1169,385,473]" pageId="9" pageNumber="419">
<materialsCitation id="59D08966DB0C6F2DFF2412B2FDA2E5AA" collectionCode="ZMUH" collectorName="P. Palmgren" country="Austria" location="Tyrol" longitude="11.316667" pageId="9" pageNumber="419" specimenCount="1" stateProvince="Tirol">
<collectingCountry id="91AFC3ABDB0C6F2DFF2412B2FEDDE5E9" box="[213,301,386,410]" name="Austria" pageId="9" pageNumber="419">Austria</collectingCountry>
,
<collectingRegion id="2B7C4DD9DB0C6F2DFEB412B2FE78E5E9" box="[325,392,386,410]" country="Austria" name="Tirol" pageId="9" pageNumber="419">Tyrol</collectingRegion>
: numerous
<emphasis id="DBCC5F29DB0C6F2DFDD712B2FDB9E5EA" box="[550,585,386,409]" italics="true" pageId="9" pageNumber="419">„„</emphasis>
and ǫǫ (
<collectionCode id="8FA91BFEDB0C6F2DFD2412B2FCC1E5E9" box="[725,817,386,410]" country="Viet Nam" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoological Museum, University of Hanoi" pageId="9" pageNumber="419">ZMUH</collectionCode>
), Obergurgl [Dorfumgebung (5around village), Rotmoostal, Schönwaisalm-Gurgler Alm],
<emphasis id="DBCC5F29DB0C6F2DFC5C1292FC49E5C9" box="[941,953,418,442]" italics="true" pageId="9" pageNumber="419">c</emphasis>
. 46
<emphasis id="DBCC5F29DB0C6F2DFC171291FC00E5CA" bold="true" box="[998,1008,417,441]" italics="true" pageId="9" pageNumber="419">°</emphasis>
529 N,
<geoCoordinate id="8C8CE5FCDB0C6F2DFBB81291FF19E5AA" degrees="11" direction="east" minutes="019" orientation="longitude" pageId="9" pageNumber="419" precision="925" value="11.316667">
11
<emphasis id="DBCC5F29DB0C6F2DFB921291FB9DE5CA" bold="true" box="[1123,1133,417,441]" italics="true" pageId="9" pageNumber="419">°</emphasis>
019 E
</geoCoordinate>
, 1960–1964 (coll.
<collectorName id="444DE6EDDB0C6F2DFE4812F1FDBAE5AA" box="[441,586,449,473]" pageId="9" pageNumber="419">P. Palmgren</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E907833BDB0C6F2DFF241122FEA4E659" blockId="9.[213,340,530,554]" box="[213,340,530,554]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF241122FEA4E659" box="[213,340,530,554]" italics="true" pageId="9" pageNumber="419">Description</emphasis>
</paragraph>
<paragraph id="E907833BDB0C6F2DFF24110AFE35E6E1" blockId="9.[213,1169,570,658]" pageId="9" pageNumber="419">
<bibRefCitation id="8D29FECADB0C6F2DFF24110AFE51E621" author="Tongiorgi P" box="[213,417,570,594]" pageId="9" pageNumber="419" pagination="275 - 334" refId="ref4541" refString="Tongiorgi P. 1966 a. Italian wolf spiders of the genus Pardosa (Araneae: Lycosidae). Bull Mus Compar Zool Harvard. 134: 275 - 334." type="journal article" year="1966">Tongiorgi (1966a)</bibRefCitation>
provides a detailed redescription of this species. Here, we only list and illustrate the main diagnostic features to distinguish this species from
<taxonomicName id="2EB8F8B8DB0C6F2DFB89116AFE91E6E1" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="419" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB0C6F2DFB89116AFE91E6E1" italics="true" pageId="9" pageNumber="419">P. pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB0C6F2DFE98114AFE35E6E1" box="[361,453,634,658]" pageId="9" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="E907833BDB0C6F2DFF2411FAFE85E712" blockId="9.[213,1169,714,865]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF2411FAFEFFE691" box="[213,271,714,738]" italics="true" pageId="9" pageNumber="419">Male</emphasis>
. Carapace with narrow yellow median band and similarly coloured lateral bands (
<figureCitation id="71839FBEDB0C6F2DFED811DAFE6FE771" box="[297,415,746,770]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="9" pageNumber="419">Figure 1D</figureCitation>
). Metatarsi and tarsi I swollen and covered with dense and long setae (
<figureCitation id="71839FBEDB0C6F2DFED0103AFE4EE751" box="[289,446,778,802]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="9" pageNumber="419">Figures 1G,H</figureCitation>
;
<figureCitation id="71839FBEDB0C6F2DFE21103AFDE1E751" box="[464,529,778,802]" captionStart="Figure 2" captionStartId="4.[146,213,1558,1580]" captionTargetBox="[195,1053,165,1522]" captionTargetId="figure-1@4.[195,1054,165,1522]" captionTargetPageId="4" captionText="Figure 2. Males of Pardosa pseudomixta sp. nov. (A,B,F–J) and Pardosa mixta (C–E). (A,C) leg I, prolateral; (B,D) metatarsus I, prolateral; (E,F,I) palp ventral; (G) palp retrolateral; (H) palp from above; (J) fine pore on metatarsus I (arrowed). Arrows indicate differences in the setae arrangement on the metatarsi between both species and fine pore. Scale 0.1 mm if not otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4651958" httpUri="https://zenodo.org/record/4651958/files/figure.png" pageId="9" pageNumber="419">2C,D</figureCitation>
). Tarsi I uniformly yellow (
<figureCitation id="71839FBEDB0C6F2DFC97103AFC09E751" box="[870,1017,778,802]" captionStart="Figure 1" captionStartId="3.[213,280,1607,1629]" captionTargetBox="[213,1170,166,1570]" captionTargetId="figure-1@3.[213,1170,166,1570]" captionTargetPageId="3" captionText="Figure 1. Pardosa pseudomixta sp. nov. (A–C,E,F) and Pardosa mixta (D,G,H). (A) Female, dorsal; (B) male, frontal; (C,D) male, lateral; (E,G), male leg I, prolateral; (F,H) male leg I, retrolateral. Arrows indicate differences in the coloration of the prosoma (light marginal band) and legs in males. Analogous figures have the same magnification." figureDoi="http://doi.org/10.5281/zenodo.4651954" httpUri="https://zenodo.org/record/4651954/files/figure.png" pageId="9" pageNumber="419">Figure 1G,H</figureCitation>
). Palea with steeply rounded ridge (
<figureCitation id="71839FBEDB0C6F2DFDF21019FD6BE732" box="[515,667,809,833]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="9" pageNumber="419">Figures 3A,E</figureCitation>
;
<figureCitation id="71839FBEDB0C6F2DFD441019FD28E732" box="[693,728,809,833]" captionStart="Figure 4" captionStartId="6.[146,213,1108,1130]" captionTargetBox="[146,1103,166,1071]" captionTargetId="figure-148@6.[146,1103,166,1072]" captionTargetPageId="6" captionText="Figure 4. Pardosa mixta (A,C,D,H,I) and Pardosa pseudomixta sp. nov. (B,E–G,J). (A,B,I,J) epigyne after maceration, ventral; (C) embolic division, ventral; (D–H) epigyne, ventral view. Analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651966" httpUri="https://zenodo.org/record/4651966/files/figure.png" pageId="9" pageNumber="419">4C</figureCitation>
); terminal apophysis not rounded (
<figureCitation id="71839FBEDB0C6F2DFF2D1079FE98E712" box="[220,360,841,865]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="9" pageNumber="419">Figure 3E,F</figureCitation>
).
</paragraph>
<paragraph id="E907833BDB0C6F2DFF2410AAFC02E782" blockId="9.[213,1169,921,1009]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF2410AAFED5E7C1" box="[213,293,922,946]" italics="true" pageId="9" pageNumber="419">Female</emphasis>
. Epigyne variable in shape (
<figureCitation id="71839FBEDB0C6F2DFD8710A9FCF8E7C2" box="[630,776,921,945]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="9" pageNumber="419">Figures 3H,I</figureCitation>
;
<figureCitation id="71839FBEDB0C6F2DFCE610A9FC76E7C2" box="[791,902,921,945]" captionStart="Figure 4" captionStartId="6.[146,213,1108,1130]" captionTargetBox="[146,1103,166,1071]" captionTargetId="figure-148@6.[146,1103,166,1072]" captionTargetPageId="6" captionText="Figure 4. Pardosa mixta (A,C,D,H,I) and Pardosa pseudomixta sp. nov. (B,E–G,J). (A,B,I,J) epigyne after maceration, ventral; (C) embolic division, ventral; (D–H) epigyne, ventral view. Analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651966" httpUri="https://zenodo.org/record/4651966/files/figure.png" pageId="9" pageNumber="419">4A,D,H,I</figureCitation>
), apical pockets fused; however, fusion is clearly visible only in scanning electron micrographs (
<figureCitation id="71839FBEDB0C6F2DFF2D10E9FE93E782" box="[220,355,985,1009]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="9" pageNumber="419">Figure 3H,I</figureCitation>
); median septum with longitudinal groove (
<figureCitation id="71839FBEDB0C6F2DFCAC10E9FC15E782" box="[861,997,985,1009]" captionStart="Figure 3" captionStartId="5.[213,280,1558,1580]" captionTargetBox="[251,1131,165,1522]" captionTargetId="figure-0@5.[251,1131,165,1522]" captionTargetPageId="5" captionText="Figure 3. Pardosa mixta (A,E,F,H,I) and Pardosa pseudomixta sp. nov. (B–D,G). (A,B) embolic division of bulb, ventral; (C,E) embolic division of bulb, apical; (D,F) terminal apophysis; (G–I) epigyne, ventral. Arrows indicate main differences in male palp and epigyne between both species. Scale 0.1 mm, analogous figures have same scale or magnification." figureDoi="http://doi.org/10.5281/zenodo.4651962" httpUri="https://zenodo.org/record/4651962/files/figure.png" pageId="9" pageNumber="419">Figure 3H,I</figureCitation>
).
</paragraph>
<paragraph id="E907833BDB0C6F2DFF24171AFE86E031" blockId="9.[213,374,1066,1090]" box="[213,374,1066,1090]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF24171AFE86E031" box="[213,374,1066,1090]" italics="true" pageId="9" pageNumber="419">Measurements</emphasis>
</paragraph>
<paragraph id="E907833BDB0C6F2DFF241762FCDEE019" blockId="9.[213,814,1106,1130]" box="[213,814,1106,1130]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF241762FE84E019" box="[213,372,1106,1130]" italics="true" pageId="9" pageNumber="419">Males/females</emphasis>
. Carapace length 2.98–3.18/2.93–3.35.
</paragraph>
<paragraph id="E907833BDB0C6F2DFF241792FE7CE089" blockId="9.[213,1169,1186,1274]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF241792FE9DE0C9" box="[213,365,1186,1210]" italics="true" pageId="9" pageNumber="419">Length ratios</emphasis>
. Tibia/metatarsus I 1.03–1.05/0.98; carapace/metatarsus I 1.61–1.65/ 1.74; carapace/tarsus IV (females only) 1.92–2.02; carapace/metatarsus IV (females only) 0.89–0.92.
</paragraph>
<paragraph id="E907833BDB0C6F2DFF241602FEABE139" blockId="9.[213,347,1330,1354]" box="[213,347,1330,1354]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF241602FEABE139" box="[213,347,1330,1354]" italics="true" pageId="9" pageNumber="419">Distribution</emphasis>
</paragraph>
<paragraph id="E907833BDB0C6F2DFF24166AFEBAE1C1" blockId="9.[213,1169,1370,1458]" pageId="9" pageNumber="419">
Central Alps (
<collectingCountry id="91AFC3ABDB0C6F2DFE84166AFE38E101" box="[373,456,1370,1394]" name="France" pageId="9" pageNumber="419">France</collectingCountry>
,
<collectingCountry id="91AFC3ABDB0C6F2DFE25166AFDFCE101" box="[468,524,1370,1394]" name="Italy" pageId="9" pageNumber="419">Italy</collectingCountry>
,
<collectingCountry id="91AFC3ABDB0C6F2DFDE6166AFD6FE101" box="[535,671,1370,1394]" name="Switzerland" pageId="9" pageNumber="419">Switzerland</collectingCountry>
, former
<collectingCountry id="91AFC3ABDB0C6F2DFCF1166AFC73E101" box="[768,899,1370,1394]" name="Serbia" pageId="9" pageNumber="419">Yugoslavia</collectingCountry>
,
<collectingCountry id="91AFC3ABDB0C6F2DFC7E166AFC14E101" box="[911,996,1370,1394]" name="Austria" pageId="9" pageNumber="419">Austria</collectingCountry>
and
<collectingCountry id="91AFC3ABDB0C6F2DFBEC166AFB79E101" box="[1053,1161,1370,1394]" name="Germany" pageId="9" pageNumber="419">Germany</collectingCountry>
) (
<bibRefCitation id="8D29FECADB0C6F2DFF2D164AFE58E1E1" author="Tongiorgi P" box="[220,424,1402,1426]" pageId="9" pageNumber="419" pagination="275 - 334" refId="ref4541" refString="Tongiorgi P. 1966 a. Italian wolf spiders of the genus Pardosa (Araneae: Lycosidae). Bull Mus Compar Zool Harvard. 134: 275 - 334." type="journal article" year="1966">Tongiorgi 1966a</bibRefCitation>
,b;
<bibRefCitation id="8D29FECADB0C6F2DFE20164AFCECE1E1" author="Heimer S &amp; Nentwig W" box="[465,796,1402,1426]" pageId="9" pageNumber="419" refId="ref4155" refString="Heimer S, Nentwig W. 1991. Spinnen Mitteleuropas: Ein Bestimmungsbuch. Berlin (Germany): Verlag Paul Parey." type="book" year="1991">Heimer and Nentwig 1991</bibRefCitation>
;
<bibRefCitation id="8D29FECADB0C6F2DFCC3164AFB79E1E1" author="Pirchegger H &amp; Thaler K" box="[818,1161,1402,1426]" pageId="9" pageNumber="419" pagination="47 - 53" refId="ref4359" refString="Pirchegger H, Thaler K. 1999. Zur Unterscheidung der Mannchen von Pardosa blanda (C. L. Koch) und P. torrentum Simon (Araneae, Lycosidae). Mitteil schweiz entomol Gesellsch. 72: 47 - 53." type="journal article" year="1999">Pirchegger and Thaler 1999</bibRefCitation>
) (
<figureCitation id="71839FBEDB0C6F2DFF2D16AAFECCE1C1" box="[220,316,1434,1458]" captionStart="Figure 5" captionStartId="8.[146,213,711,733]" captionTargetBox="[146,1104,165,675]" captionTargetId="figure-314@8.[146,1104,165,675]" captionTargetPageId="8" captionText="Figure 5. Records of Pardosa pseudomixta sp. nov. (full circles) and Pardosa mixta (full diamonds). The map also shows erroneous records of P. mixta from Uzbekistan (full squares) and presumed records of P. pseudomixta sp. nov. that were reported as P. mixta from China (open circle)." figureDoi="http://doi.org/10.5281/zenodo.4651968" httpUri="https://zenodo.org/record/4651968/files/figure.png" pageId="9" pageNumber="419">Figure 5</figureCitation>
).
</paragraph>
<paragraph id="E907833BDB0C6F2DFF2416DAFEC8E271" blockId="9.[213,312,1514,1538]" box="[213,312,1514,1538]" pageId="9" pageNumber="419">
<emphasis id="DBCC5F29DB0C6F2DFF2416DAFEC8E271" box="[213,312,1514,1538]" italics="true" pageId="9" pageNumber="419">Remarks</emphasis>
</paragraph>
<paragraph id="E907833BDB0C6F2DFF241522FC3DE2D9" blockId="9.[213,1169,1554,1706]" pageId="9" pageNumber="419">
The extensive pubescence on leg I of
<taxonomicName id="2EB8F8B8DB0C6F2DFD891523FD28E258" authorityName="Alps" baseAuthorityName="Kulczynski" baseAuthorityYear="1887" box="[632,728,1555,1579]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="419" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB0C6F2DFD891523FD28E258" box="[632,728,1555,1579]" italics="true" pageId="9" pageNumber="419">P. mixta</emphasis>
</taxonomicName>
males led us to expect that this species might have numerous pores associated with these dense setae similar to the condition in certain other lycosid species (
<bibRefCitation id="8D29FECADB0C6F2DFD901562FC23E219" author="Kronestedt T &amp; Marusik YM" box="[609,979,1618,1642]" pageId="9" pageNumber="419" pagination="63 - 71" refId="ref4205" refString="Kronestedt T, Marusik YM. 2002. On Acantholycosa solituda (Levi &amp; Levi) and A. sterneri (Marusik) (Araneae, Lycosidae), a pair of geographically distant allied species. Acta Arachnol. 51: 63 - 71." type="journal article" year="2002">Kronestedt and Marusik 2002</bibRefCitation>
). However, our scanning electron microscopy studies revealed no distinct pores in
<taxonomicName id="2EB8F8B8DB0C6F2DFC4F1542FBEDE2F9" authorityName="Alps" baseAuthorityName="Kulczynski" baseAuthorityYear="1887" box="[958,1053,1650,1674]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="419" phylum="Arthropoda" rank="species" species="mixta">
<emphasis id="DBCC5F29DB0C6F2DFC4F1542FBEDE2F9" box="[958,1053,1650,1674]" italics="true" pageId="9" pageNumber="419">P. mixta</emphasis>
</taxonomicName>
, although scattered pores were found in
<taxonomicName id="2EB8F8B8DB0C6F2DFDC315A2FD10E2D9" authority="Marusik &amp; Fritzén, 2009" authorityName="Marusik &amp; Fritzén" authorityYear="2009" box="[562,736,1682,1706]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="419" phylum="Arthropoda" rank="species" species="pseudomixta" status="sp. nov.">
<emphasis id="DBCC5F29DB0C6F2DFDC315A2FD10E2D9" box="[562,736,1682,1706]" italics="true" pageId="9" pageNumber="419">P. pseudomixta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0FFE252DB0C6F2DFD1915A2FCB3E2D9" box="[744,835,1682,1706]" pageId="9" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="71839FBEDB0C6F2DFCA515A2FC31E2D9" box="[852,961,1682,1706]" captionStart="Figure 2" captionStartId="4.[146,213,1558,1580]" captionTargetBox="[195,1053,165,1522]" captionTargetId="figure-1@4.[195,1054,165,1522]" captionTargetPageId="4" captionText="Figure 2. Males of Pardosa pseudomixta sp. nov. (A,B,F–J) and Pardosa mixta (C–E). (A,C) leg I, prolateral; (B,D) metatarsus I, prolateral; (E,F,I) palp ventral; (G) palp retrolateral; (H) palp from above; (J) fine pore on metatarsus I (arrowed). Arrows indicate differences in the setae arrangement on the metatarsi between both species and fine pore. Scale 0.1 mm if not otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4651958" httpUri="https://zenodo.org/record/4651958/files/figure.png" pageId="9" pageNumber="419">Figure 2J</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="806600DADDD5F139E60FC0368928257A">Acleistochelys, a New Side-Necked Turtle (Pelomedusoides: Bothremydidae) from the Paleocene of Mali</mods:title>
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<subSubSection id="A1A2651272184D1B575CFEC1FB99FEA4" box="[779,1070,346,370]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="E907369972184D1B575CFEC1FB99FEA4" blockId="3.[779,1070,346,370]" box="[779,1070,346,370]" pageId="3" pageNumber="4">
<heading id="B24F81F572184D1B575CFEC1FB99FEA4" box="[779,1070,346,370]" centered="true" fontSize="36" level="2" pageId="3" pageNumber="4" reason="8">
<taxonomicName id="2EB84D1A72184D1B575CFEC1FC10FEA4" authority="Gaffney &amp; Roberts &amp; Sissoko &amp; Bouaré &amp; Tapanila &amp; O'leary, 2007" authorityName="Gaffney &amp; Roberts &amp; Sissoko &amp; Bouaré &amp; Tapanila &amp; O'leary" authorityYear="2007" box="[779,935,346,370]" class="Reptilia" family="Bothremydidae" genus="Acleistochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="4" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="DBCCEA8B72184D1B575CFEC1FC10FEA4" bold="true" box="[779,935,346,370]" italics="true" pageId="3" pageNumber="4">Acleistochelys</emphasis>
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,
<taxonomicNameLabel id="C0FF57F072184D1B57E0FEC1FB99FEA4" box="[951,1070,346,370]" pageId="3" pageNumber="4" rank="genus">new genus</taxonomicNameLabel>
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<paragraph id="E907369972184D1B56E7FE1DFC96FE6D" blockId="3.[662,1188,390,1268]" pageId="3" pageNumber="4">
<materialsCitation id="59D03CC472184D1B56E7FE1DFCAAFE6D" ID-GBIF-Occurrence="3340940302" collectionCode="SPECIES" pageId="3" pageNumber="4" specimenCount="1" typeStatus="holotype">
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:
<taxonomicName id="2EB84D1A72184D1B572CFE1DFB2BFE48" authority="Gaffney &amp; Roberts &amp; Sissoko &amp; Bouaré &amp; Tapanila &amp; O'leary, 2007" authorityName="Gaffney &amp; Roberts &amp; Sissoko &amp; Bouaré &amp; Tapanila &amp; O'leary" authorityYear="2007" box="[891,1180,390,414]" class="Reptilia" family="Bothremydidae" genus="Acleistochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="maliensis" status="sp. nov.">
<emphasis id="DBCCEA8B72184D1B572CFE1DFB2BFE48" box="[891,1180,390,414]" italics="true" pageId="3" pageNumber="4">Acleistochelys maliensis</emphasis>
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,
<taxonomicNameLabel id="C0FF57F072184D1B56C1FE38FCAAFE6D" box="[662,797,419,443]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
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.
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<paragraph id="E907369972184D1B56E7FE5AFBF7FE0F" blockId="3.[662,1188,390,1268]" box="[688,1088,449,473]" pageId="3" pageNumber="4">
DISTRIBUTION: Paleocene of
<collectingCountry id="91AF760972184D1B5055FE5AFB8BFE0F" box="[1026,1084,449,473]" name="Serbia" pageId="3" pageNumber="4">Mali</collectingCountry>
.
</paragraph>
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<subSubSection id="A1A2651272184D1B56E7FE44FC73FD86" pageId="3" pageNumber="4" type="etymology">
<paragraph id="E907369972184D1B56E7FE44FC73FD86" blockId="3.[662,1188,390,1268]" pageId="3" pageNumber="4">
ETYMOLOGY:
<emphasis id="DBCCEA8B72184D1B570FFE7BFC0FFE2E" box="[856,952,480,504]" italics="true" pageId="3" pageNumber="4">Aklystos</emphasis>
, Greek for ‘‘sheltered’’, or ‘‘hollowed’’, in allusion to the small, paired pits on the triturating surface;
<emphasis id="DBCCEA8B72184D1B56C1FDA3FD6EFD86" box="[662,729,568,592]" italics="true" pageId="3" pageNumber="4">chelys</emphasis>
, Greek for ‘‘turtle’’.
</paragraph>
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<subSubSection id="A1A2651272184D1B56E7FDCDFC9DFB6F" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="E907369972184D1B56E7FDCDFC9DFB6F" blockId="3.[662,1188,390,1268]" pageId="3" pageNumber="4">
DIAGNOSIS: A bothremydid pleurodire with these unique characters among the tribe
<taxonomicName id="2EB84D1A72184D1B56C1FD0AFCF6FD7F" authorityName="GAFFNEY, TONG, AND MEYLAN" authorityYear="2006" box="[662,833,657,681]" class="Reptilia" family="Bothremydidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="4" phylum="Chordata" rank="tribe" tribe="Taphrosphyini">Taphrosphyini</taxonomicName>
: small pit formed by jugal, maxilla, and palatine on triturating surface; jugal exposed on triturating surface; accessory ridge present on anterior triturating surface; wide palatine-basisphenoid contact separating pterygoids on midline; supraoccipital-quadrate contact present; basioccipital narrowly enters condylus occipitalis; palatine-jugal contact in small septum orbitotemporale. Other distinguishing characters are: skull relatively long and narrow; fossa pterygoideus deep and narrow as in
<taxonomicName id="2EB84D1A72184D1B5715FC4FFC0EFC3A" authorityName="Broin" authorityYear="1977" box="[834,953,980,1004]" class="Reptilia" family="Bothremydidae" genus="Nigeremys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
<emphasis id="DBCCEA8B72184D1B5715FC4FFC0EFC3A" box="[834,953,980,1004]" italics="true" pageId="3" pageNumber="4">Nigeremys</emphasis>
</taxonomicName>
but in contrast to
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<emphasis id="DBCCEA8B72184D1B56C1FC6AFC93FBDF" box="[662,804,1009,1033]" italics="true" pageId="3" pageNumber="4">Taphrosphys</emphasis>
</taxonomicName>
; foramen posterius canalis carotici interni formed by pterygoid, basisphenoid, and quadrate in contrast to
<taxonomicName id="2EB84D1A72184D1B57B3FBB7FB2BFB92" box="[996,1180,1068,1092]" class="Reptilia" family="Bothremydidae" genus="Phosphatochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
<emphasis id="DBCCEA8B72184D1B57B3FBB7FB2BFB92" box="[996,1180,1068,1092]" italics="true" pageId="3" pageNumber="4">Phosphatochelys</emphasis>
</taxonomicName>
; small remnant of septum orbitotemporale present consisting of ventral parietal process as in
<taxonomicName id="2EB84D1A72184D1B56BCFB1FFC14FB4A" box="[747,931,1156,1180]" class="Reptilia" family="Bothremydidae" genus="Phosphatochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
<emphasis id="DBCCEA8B72184D1B56BCFB1FFC14FB4A" box="[747,931,1156,1180]" italics="true" pageId="3" pageNumber="4">Phosphatochelys</emphasis>
</taxonomicName>
but in contrast to
<taxonomicName id="2EB84D1A72184D1B56C1FB3AFC93FB6F" authorityName="Cope" authorityYear="1869" box="[662,804,1185,1209]" class="Reptilia" family="Bothremydidae" genus="Taphrosphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
<emphasis id="DBCCEA8B72184D1B56C1FB3AFC93FB6F" box="[662,804,1185,1209]" italics="true" pageId="3" pageNumber="4">Taphrosphys</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A1A2651272184D1B56E7FB24FBF6FB22" pageId="3" pageNumber="4" type="discussion">
<paragraph id="E907369972184D1B56E7FB24FBF6FB22" blockId="3.[662,1188,390,1268]" pageId="3" pageNumber="4">
DISCUSSION: See table 1 for a comparison of genera in the tribe
<taxonomicName id="2EB84D1A72184D1B57C5FB47FB8AFB22" authorityName="GAFFNEY, TONG, AND MEYLAN" authorityYear="2006" box="[914,1085,1244,1268]" class="Reptilia" family="Bothremydidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="4" phylum="Chordata" rank="tribe" tribe="Taphrosphyini">Taphrosphyini</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="6094DCF1B9652752BFA3C7FC5BE8C18A">Acleistochelys, a New Side-Necked Turtle (Pelomedusoides: Bothremydidae) from the Paleocene of Mali</mods:title>
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<emphasis id="DBCCEA8B72184D1B569AFAAEFC6FFA9B" bold="true" box="[717,984,1333,1357]" italics="true" pageId="3" pageNumber="4">Acleistochelys maliensis</emphasis>
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<materialsCitation id="59D03CC472184D1B56E7FAFAFBE4F957" ID-GBIF-Occurrence="3340940303" collectionCode="SPECIMEN, CNRST" pageId="3" pageNumber="4" specimenCode="SUNY 199" specimenCount="1" typeStatus="holotype">
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:
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<specimenCode id="B91E9EE272184D1B5059FAFAFB13FAAF" box="[1038,1188,1377,1401]" collectionCode="SUNY" pageId="3" pageNumber="4">SUNY 199</specimenCode>
(figs. 2–5), partial skull including fragments of the lower jaw, lacking some of the left temporal region, and both posterior parts of the skull roof. Skull measurements are in table 2. Associated shell fragments include the anterior margin of the nuchal (fig. 6), neurals one and four (fig. 7), and peripheral two (fig. 7). A partial cervical vertebra (figs. 8, 9) and some pelvic fragments are present
</materialsCitation>
.
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<materialsCitation id="59D03CC472184D1B56E7F91CFC79F90C" ID-GBIF-Occurrence="3340940301" collectionCode="LOCALITY" country="Serbia" location="Adrar des Iforas Mountains" pageId="3" pageNumber="4" specimenCount="1" typeStatus="holotype">
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:
<collectingCountry id="91AF760972184D1B572AF91CFC04F949" box="[893,947,1671,1695]" name="Serbia" pageId="3" pageNumber="4">Mali</collectingCountry>
17, located south of the
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<collectingCountry id="91AF760972184D1B578BF959FBA1F90C" box="[988,1046,1730,1754]" name="Serbia" pageId="3" pageNumber="4">Mali</collectingCountry>
.
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HORIZON: Middle to upper portion of the Teberemt Formation of
<bibRefCitation id="8D294B68721F4D1C55CAFF6AFF76FEF0" author="Moody, R. T. J. &amp; P. J. C. Sutcliffe" pageId="4" pageNumber="5" pagination="137 - 157" refId="ref10784" refString="Moody, R. T. J., and P. J. C. Sutcliffe. 1991. The Cretaceous deposits of the Iullemmeden Basin of Niger, central West Africa. Cretaceous Research 12: 137 - 157." type="journal article" year="1991">Moody and Sutcliffe (1991</bibRefCitation>
,
<bibRefCitation id="8D294B68721F4D1C5499FE95FEBFFEF0" author="Moody, R. T. J. &amp; P. J. C. Sutcliffe" box="[206,264,270,294]" pageId="4" pageNumber="5" pagination="539 - 554" refId="ref10826" refString="Moody, R. T. J., and P. J. C. Sutcliffe. 1993. The sedimentology and palaeontology of the Upper Cretaceous - Tertiary deposits of central West Africa. Modern Geology 18: 539 - 554." type="journal article" year="1993">1993</bibRefCitation>
).
</paragraph>
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<paragraph id="E9073699721F4D1C54CDFEB6FF61FEB4" blockId="4.[128,654,211,565]" pageId="4" pageNumber="5">DEPOSITIONAL ENVIRONMENT: Near shore marine.</paragraph>
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<subSubSection id="A1A26512721F4D1C54CDFEF2FE7CFE57" box="[154,459,361,385]" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="E9073699721F4D1C54CDFEF2FE7CFE57" blockId="4.[128,654,211,565]" box="[154,459,361,385]" pageId="4" pageNumber="5">DIAGNOSIS: As for genus.</paragraph>
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<subSubSection id="A1A26512721F4D1C54CDFE1CFE58FE0D" pageId="4" pageNumber="5" type="etymology">
<paragraph id="E9073699721F4D1C54CDFE1CFF45FE6B" blockId="4.[128,654,211,565]" pageId="4" pageNumber="5">ETYMOLOGY: Named for the country of discovery.</paragraph>
<paragraph id="E9073699721F4D1C54CDFE58FE58FE0D" blockId="4.[128,654,211,565]" box="[154,495,451,475]" pageId="4" pageNumber="5">REFERRED MATERIAL: None.</paragraph>
</subSubSection>
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<paragraph id="E9073699721F4D1C54CDFE7AFEC1FDE3" blockId="4.[128,654,211,565]" pageId="4" pageNumber="5">
PREVIOUS WORK: CNRST SUNY 199 is included in the phylogenetic analysis of
<bibRefCitation id="8D294B68721F4D1C54D7FD86FEC5FDE3" author="Gaffney, E. S. &amp; H. Tong &amp; P. A. Meylan" box="[128,370,541,565]" pageId="4" pageNumber="5" pagination="1 - 698" refId="ref10646" refString="Gaffney, E. S., H. Tong, and P. A. Meylan. 2006. Evolution of the side-necked turtles: the Families Bothremydidae, Euraxemydidae, and Araripemydidae. Bulletin of the American Museum of Natural History 300: 1 - 698." type="journal article" year="2006">Gaffney et al. (2006)</bibRefCitation>
.
</paragraph>
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