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classification Animalia Coleoptera Leiodidae
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classification Animalia
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<taxonomicName LSID="http://zoobank.org/D518FD86-9AD9-458D-972C-121655AF9A0D" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="11" pageNumber="54" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae">Ptomaphagus (s. str.) tingtingae</taxonomicName>
<taxonomicNameLabel pageId="11" pageNumber="54">sp. n.</taxonomicNameLabel>
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<taxonomicNameLabel id="AFBAA91D5101D7C231C39DCC385ED013" pageId="11" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
Figs 3
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; 9B, D, F, I, J; 10C, D; 11
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<pageBreakToken pageId="12" pageNumber="55" start="start">Type</pageBreakToken>
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locality.
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<paragraph id="00E7BEA86E04EADC335C3C3CD2DAA6E7" pageId="12" pageNumber="55">
Taiwan, Fushan [福山, ca.
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,
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<geoCoordinate id="73E2E51D847BD3A7933935C95C4A588A" direction="east" orientation="longitude" precision="925" value="121.5">121°30'E</geoCoordinate>
].
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<paragraph pageId="12" pageNumber="55">Type material.</paragraph>
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Holotype: ♂, CHINA, Taiwan, Fushan [福山, ca.
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<geoCoordinate id="F0475CAB41937CD34EC682327835BFAD" direction="north" orientation="latitude" precision="925" value="24.766666">24°46'N</geoCoordinate>
,
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], mouse carcass bait, III.2007, Wen-Bo Huang leg. (5#) (NTUC). Paratypes: 6♂♂7♀♀, same data as holotype (1♂1♀ in CCBW, 1♂1♀ in CJRZ, 1♂1♀ in CMNE, 1♂1♀ in CMPR and 2♂♂3♀♀ in NTUC); 7♀♀, same data as holotype ex
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: IV.2007, (1#) (NTUC); 2♂♂1♀, Taiwan: Tai Pei Co. / Noi Dong [?Neidong=内洞, ca.
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,
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] Logging Road, / 850 m, 19.ii.2004, / Flight intercept trap. / leg. Chun Lin Li (ZMHB); 2♀♀, Taiwan: Tai Chun Co., / An Ma Shan [鞍馬山, ca.
<geoCoordinate direction="north" orientation="latitude" precision="925" value="24.266666">24°16'N</geoCoordinate>
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,
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], 2 km, / 24.-26.vi.2003 / Flight intercept trap. / leg. Chun Lin Li (ZMHB).
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<paragraph pageId="13" pageNumber="56">Description.</paragraph>
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<paragraph id="0BF003919BFD3F6E8D5A7EC20EE953EF" pageId="13" pageNumber="56">Description.</paragraph>
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Male. EBL: 4.0-4.3 mm (4.1 mm in holotype). Length of different body parts: HL: AL: PL: ELL = 0.62: 1.22: 1.00: 2.27 mm; width: HW: EW: PW: ELW = 1.00: 0.11: 1.55: 1.68 mm. Proportion of antennomeres from base to tip in
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width): 180
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132.
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<paragraph pageId="14" pageNumber="57">Habitus (Fig. 3C, E) elongated oval, regularly convex and sublustrous. Well pigmented: mostly dark brown, head darker; mouthparts, basal three antennomeres and apical half of ultimate antennomere, protarsi, and apex of meso- and metatarsi somewhat yellowish. Dorsum continuously clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.</paragraph>
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<paragraph id="9E0BA50527AA65E66F0F2CCBC40E2557" pageId="14" pageNumber="57">Habitus (Fig. 3C, E) elongated oval, regularly convex and sublustrous. Well pigmented: mostly dark brown, head darker; mouthparts, basal three antennomeres and apical half of ultimate antennomere, protarsi, and apex of meso- and metatarsi somewhat yellowish. Dorsum continuously clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.</paragraph>
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transverse, HW/HL = 1.6. Clypeofrontal suture absent. Clypeus with anterior margin slightly rounded. Compound eyes well developed, EW/HW = 0.1. Antennae (Fig. 11A) slender, AL/HW = 1.2; antennomere III a little shorter than II; VI with length/width = 0.5; XI pear-shaped.
</paragraph>
<paragraph pageId="15" pageNumber="58">Pronotum (Fig. 11B) transverse, widest just before hind angles, PW/PL = 1.6. Sides gently arched, gradually narrowing from posterior to anterior; hind angles projected backwards and acute. Posterior margin widely protruding in the middle part, distinctly emarginate near hind angles.</paragraph>
<paragraph pageId="16" pageNumber="59">
<pageBreakToken pageId="16" pageNumber="59" start="start">Elytra</pageBreakToken>
<paragraph id="5F0C4067A9BAA2ADC2A66B5E09B44AB3" pageId="15" pageNumber="58">Pronotum (Fig. 11B) transverse, widest just before hind angles, PW/PL = 1.6. Sides gently arched, gradually narrowing from posterior to anterior; hind angles projected backwards and acute. Posterior margin widely protruding in the middle part, distinctly emarginate near hind angles.</paragraph>
<paragraph id="73FE9C00C5049D8920E6A8902059FF1B" pageId="16" pageNumber="59">
<pageBreakToken id="7CC09F4494EB47A1DCA88A3C90118AE4" pageId="16" pageNumber="59" start="start">Elytra</pageBreakToken>
oval, widest at about basal 1/3, ELL/EW = 1.4. Sides weakly arched, gradually narrowing from widest part to apices, which narrowly rounded (Fig. 11G). Sutural striae present. Metathoracic wings fully developed.
</paragraph>
<paragraph pageId="16" pageNumber="59">Prolegs robust, with basal three protarsomeres (Fig. 11C) strongly expanded: TW/BTW = 1.2. Protibiae (Fig. 11C) distinctly expanded towards apex. Profemora rather broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender and straight.</paragraph>
<paragraph pageId="16" pageNumber="59">Abdominal ventrite VIII (Fig. 11I) narrowly round at posterior edge, though an inconspicuous median notch at the median. Genital segment (Fig. 8J) with very slender spiculum gastrale, protruding about 3/8 of its length beyond anterior edge of epipleurite IX.</paragraph>
<paragraph lastPageId="17" lastPageNumber="60" pageId="16" pageNumber="59">
<paragraph id="32192FE3CA2B0055F94C4EA9BA1B4818" pageId="16" pageNumber="59">Prolegs robust, with basal three protarsomeres (Fig. 11C) strongly expanded: TW/BTW = 1.2. Protibiae (Fig. 11C) distinctly expanded towards apex. Profemora rather broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender and straight.</paragraph>
<paragraph id="728DFF4650D0F94448485D0256A5DDF2" pageId="16" pageNumber="59">Abdominal ventrite VIII (Fig. 11I) narrowly round at posterior edge, though an inconspicuous median notch at the median. Genital segment (Fig. 8J) with very slender spiculum gastrale, protruding about 3/8 of its length beyond anterior edge of epipleurite IX.</paragraph>
<paragraph id="95A24FA97A9B81F71BBB571D61A2C142" lastPageId="17" lastPageNumber="60" pageId="16" pageNumber="59">
Aedeagus (Fig. 9B) long and rather slender, with median lobe gradually narrowing towards narrowly lanceolate apical part and terminated by round knob in dorsal view; opening of genital orifice situated on dorsal surface, deeply cut inwards on preapical
<pageBreakToken pageId="17" pageNumber="60" start="start">left</pageBreakToken>
<pageBreakToken id="560884C6CA89D3500863633F2940E924" pageId="17" pageNumber="60" start="start">left</pageBreakToken>
margin of median lobe. Ventral surface of the apex of the median lobe (Fig. 9J) inserted with a row of 6 ventrally-oriented setae on the left side and a row of 5 ventrally-oriented setae on the right side. Parameres narrow, reaching about apical 1/6 of median lobe, each with 1 apical and 2 preapical setae, the apical one very shorter (Fig. 9F). In lateral view (Fig. 9D), median lobe regularly bent ventrad, gradually tapering apically. Endophallus with stylus quite slender, a cheliform complex just below the base of stylus, and a circular complex in basal region.
</paragraph>
<paragraph pageId="17" pageNumber="60">Female. Similar to male in general appearance (Fig. 3D), including elytral apices (Fig. 11H), but distinguished by the following characteristics: protarsi (Fig. 11D) simply linear; protibiae (Fig. 11F) only slightly narrower; abdominal ventrite VIII (Fig. 10D) regularly rounded at posterior edge; genital segment as shown in Fig. 10C: spermatheca (Fig. 10C) curved in distal part and coiled in proximal part.</paragraph>
<paragraph id="D3A568FEF281D3BC5803B6E92F0E50F0" pageId="17" pageNumber="60">Female. Similar to male in general appearance (Fig. 3D), including elytral apices (Fig. 11H), but distinguished by the following characteristics: protarsi (Fig. 11D) simply linear; protibiae (Fig. 11F) only slightly narrower; abdominal ventrite VIII (Fig. 10D) regularly rounded at posterior edge; genital segment as shown in Fig. 10C: spermatheca (Fig. 10C) curved in distal part and coiled in proximal part.</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="60" type="diagnosis">
<paragraph pageId="17" pageNumber="60">Diagnosis.</paragraph>
<paragraph pageId="17" pageNumber="60">
<subSubSection id="6E7F5004D4228D2A0D1AABABCCB62075" pageId="17" pageNumber="60" type="diagnosis">
<paragraph id="7B96801195B94DC4990EE6F0CD689375" pageId="17" pageNumber="60">Diagnosis.</paragraph>
<paragraph id="14968A991C56AC71A5B55DDE6CE1F967" pageId="17" pageNumber="60">
See under
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="17" pageNumber="60" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi">Ptomaphagus (s. str.) chenggongi</taxonomicName>
sp. n. above.
<taxonomicName id="83737D847C0DEA12933F3E2C6D1C6678" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="17" pageNumber="60" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi" status="sp. nov.">Ptomaphagus (s. str.) chenggongi</taxonomicName>
<taxonomicNameLabel id="61451655F891C74D86CF4081AE93ED78" rank="species">sp. n.</taxonomicNameLabel>
above.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="60" type="etymology">
<paragraph pageId="17" pageNumber="60">Etymology.</paragraph>
<paragraph pageId="17" pageNumber="60">
<subSubSection id="635BBEFEFD68946BF943B8D01B60B603" pageId="17" pageNumber="60" type="etymology">
<paragraph id="6771B90D7CF83A72A6A857C37BAE0343" pageId="17" pageNumber="60">Etymology.</paragraph>
<paragraph id="B33C560A1800C18FBCAC1A666A620847" pageId="17" pageNumber="60">
The specific epithet is dedicated to Miss Ting-Ting Song, the first
<normalizedToken originalValue="authors">author's</normalizedToken>
<normalizedToken id="195B4995E88B955E7F3188C164A2A27C" originalValue="authors">author's</normalizedToken>
former colleague (Institute of Zoology, Chinese Academy of Sciences, Beijing, China), who did important primary work on Chinese
<taxonomicName class="Insecta" family="Leiodidae" genus="Leiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiodes" order="Coleoptera" pageId="17" pageNumber="60" phylum="Arthropoda" rank="genus">Leiodes</taxonomicName>
Latreille, 1796 (
<taxonomicName family="Leiodidae" lsidName="" pageId="17" pageNumber="60" rank="family">Leiodidae</taxonomicName>
<taxonomicName id="AEE35585A359F88FE1851BA25E99DCA9" authority="Latreille, 1796" authorityName="Latreille" authorityYear="1796" class="Insecta" family="Leiodidae" genus="Leiodes" kingdom="Animalia" lsidName="Leiodes" order="Coleoptera" pageId="17" pageNumber="60" phylum="Arthropoda" rank="genus">Leiodes Latreille, 1796</taxonomicName>
(
<taxonomicName id="CF90247ABEF6F8D1B66B59EF4DE09FB5" class="Insecta" family="Leiodidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="60" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
:
<taxonomicName lsidName="" pageId="17" pageNumber="60" rank="subFamily" subFamily="Leiodinae">Leiodinae</taxonomicName>
<taxonomicName id="05C88A98E79DAB5AC171F8A77D90FB08" class="Insecta" family="Leiodidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="60" phylum="Arthropoda" rank="subFamily" subFamily="Leiodinae">Leiodinae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="60" type="distribution">
<paragraph pageId="17" pageNumber="60">Distribution.</paragraph>
<paragraph pageId="17" pageNumber="60">China (Taiwan) (Fig. 12).</paragraph>
<subSubSection id="BEDD048AE6A5C2F8D3EA4F17DC48783B" pageId="17" pageNumber="60" type="distribution">
<paragraph id="ACAE208B62D0BBFE112D0689E69110DE" pageId="17" pageNumber="60">Distribution.</paragraph>
<paragraph id="08F762C65F804BA5F22920D57E5CB68E" pageId="17" pageNumber="60">China (Taiwan) (Fig. 12).</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,170 +1,177 @@
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<paragraph pageId="6" pageNumber="49">Taxon classification Animalia Coleoptera Leiodidae</paragraph>
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Taxon classification Animalia
<taxonomicName id="D09D01CA7A5DC502807A3B3C9F43CDC0" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="B1A4EF99CD6BA9A4212ADF023B4479F5" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
</paragraph>
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<subSubSection pageId="6" pageNumber="49" type="nomenclature">
<paragraph pageId="6" pageNumber="49">
<taxonomicName LSID="http://zoobank.org/D6994368-3822-4571-8B04-7C3AAF0BC100" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi">Ptomaphagus (s. str.) chenggongi</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="49">sp. n.</taxonomicNameLabel>
<subSubSection id="7D1344B097E5D8CEBF6BDD1397BA17E7" pageId="6" pageNumber="49" type="nomenclature">
<paragraph id="0BE7A4A2A6E685F4565467EA1E1E6A2B" pageId="6" pageNumber="49">
<taxonomicName id="AA255F1537D55BE90FE2755725C5CB39" ID-CoL="6WLPY" LSID="http://zoobank.org/D6994368-3822-4571-8B04-7C3AAF0BC100" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi" status="sp. nov.">Ptomaphagus (s. str.) chenggongi</taxonomicName>
<taxonomicNameLabel id="6C01A1CAE2C00D54FFA66843F5897DE0" pageId="6" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
Figs 3; 8; 9A, C, E, G, H; 10
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="49" type="type locality">
<paragraph pageId="6" pageNumber="49">Type locality.</paragraph>
<paragraph pageId="6" pageNumber="49">
<subSubSection id="9A3A4741DE2E5A3470D4D79B6A0B5DD0" pageId="6" pageNumber="49" type="type locality">
<paragraph id="09BB210CC3DCFAAB4BB114C9F2DFEA6B" pageId="6" pageNumber="49">Type locality.</paragraph>
<paragraph id="EA8901321C00E936DE1D55A4A2562E2D" pageId="6" pageNumber="49">
Central Taiwan, Nantou Hsien, Tsuifeng [翠峰, ca.
<geoCoordinate direction="north" orientation="latitude" precision="925" value="24.1">24°06'N</geoCoordinate>
<geoCoordinate id="249F4F6AD15021A146D8AE567857BE9A" direction="north" orientation="latitude" precision="925" value="24.1">24°06'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="121.183334">121°11'E</geoCoordinate>
<geoCoordinate id="46474AA950A05D6F8DDD02AB77BDE261" direction="east" orientation="longitude" precision="925" value="121.183334">121°11'E</geoCoordinate>
], 2,300 m.
</paragraph>
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<subSubSection pageId="6" pageNumber="49" type="type material">
<paragraph pageId="6" pageNumber="49">Type material.</paragraph>
<paragraph pageId="6" pageNumber="49">
<subSubSection id="143AAFCFA5F73CC92A0ACA4C1643AFA1" pageId="6" pageNumber="49" type="type material">
<paragraph id="D42CEC9A80423C17A99D4BE2DDA1CA9D" pageId="6" pageNumber="49">Type material.</paragraph>
<paragraph id="F1390FBA90A6D32662B79ECE1A85437A" pageId="6" pageNumber="49">
Holotype: ♂, [Taiwan] Tsuifeng [翠峰, ca.
<geoCoordinate direction="north" orientation="latitude" precision="925" value="24.1">24°06'N</geoCoordinate>
<geoCoordinate id="3F959A20E521366155DFA96566A30890" direction="north" orientation="latitude" precision="925" value="24.1">24°06'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="121.183334">121°11'E</geoCoordinate>
<geoCoordinate id="A7314287C6B9F6E4AA7BF0B8032E5782" direction="east" orientation="longitude" precision="925" value="121.183334">121°11'E</geoCoordinate>
], 2,300 m / FIT: in shady Forest / Nantou Hsien // Central Taiwan / 7~14-VIII-2003 / Wataru Suzuki leg. (NSMT). Paratypes: 22♂♂24♀♀, same data as holotype (22♂♂22♀♀ in CMNE, 2♀♀ in NSMT); 2♂♂, same data as holotype except: FIT: Forest edge (CMNE); 10♂♂10♀♀, same data as holotype except: 2,200 m / FIT: shady natural forest (CMNE); 2♂♂1♀, same data as holotype except: 2,200 m (CMNE); 1♀, TAIWAN, Nantou / Hsien, Meifeng [梅峰, ca.
<geoCoordinate direction="north" orientation="latitude" precision="925" value="24.083334">24°05'N</geoCoordinate>
<geoCoordinate id="4D4912E4D156BCA3A6B6364C8B09F512" direction="north" orientation="latitude" precision="925" value="24.083334">24°05'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="121.166664">121°10'E</geoCoordinate>
<geoCoordinate id="D4A7025C05D8BB9ADAD57B30C44C7FF2" direction="east" orientation="longitude" precision="925" value="121.166664">121°10'E</geoCoordinate>
] / 2130 m 10-17.VII.[19]93 / yellow pan traps / A. Smetana [leg.] (T147) (CJRZ).
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="52" pageId="6" pageNumber="49" type="description">
<paragraph pageId="6" pageNumber="49">Description.</paragraph>
<paragraph lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49">
<subSubSection id="CBA4FAD63CDFB4607B058BA1B2E46E3F" lastPageId="9" lastPageNumber="52" pageId="6" pageNumber="49" type="description">
<paragraph id="7E638B6ED58BF440480A5EFFBE3804CB" pageId="6" pageNumber="49">Description.</paragraph>
<paragraph id="D7C57DB90C20D819FFD69FEE8089DBC2" lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49">
Male. EBL: 4.1-4.2 mm (4.2 mm in holotype). Length of different body parts: HL: AL: PL: ELL = 0.67: 1.26: 1.00: 2.32 mm; width: HW: EW: PW: ELW = 1.04: 0.09: 1.49: 1.64 mm. Proportion of antennomeres from base to tip in
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<normalizedToken originalValue="μm">μm</normalizedToken>
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<normalizedToken id="B82E1D378791A5B9AF46FB3FE1FB718B" originalValue="μm">μm</normalizedToken>
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(length
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width): 179
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77, 123
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68, 105
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<normalizedToken id="DB442F8C96263EC7CD744CD28EEDB7C4" originalValue="×">x</normalizedToken>
74, 75
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="935DAC6EDF730F66AAF0AE54FB1EF8B0" originalValue="×">x</normalizedToken>
82, 75
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="990E4B4A37C0F25653DB2F4194D291BF" originalValue="×">x</normalizedToken>
99, 53
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113, 104
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<normalizedToken id="5FDF58D05872FEE90F2FA7B2CB7A742F" originalValue="×">x</normalizedToken>
135, 53
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132, 102
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138, 117
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137, 217
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116.
</paragraph>
<paragraph pageId="7" pageNumber="50">Habitus (Fig. 3A) elongated oval, regularly convex and sublustrous. Well pigmented: mostly brown; mouthparts, basal four or five antennomeres and apical half of ultimate antennomere, protarsi, and apex of meso- and metatarsi yellowish. Dorsum continuously clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.</paragraph>
<paragraph pageId="7" pageNumber="50">Head transverse, HW/HL = 1.5. Clypeofrontal suture absent. Clypeus with anterior margin slightly rounded. Compound eyes well developed, EW/HW = 0.1. Antennae (Fig. 8A) slender, AL/HW = 1.3; antennomere III a little shorter than II; VI with length/width = 0.5; XI elongated pear-shaped.</paragraph>
<paragraph pageId="7" pageNumber="50">Pronotum (Fig. 8B) transverse, widest just before hind angles, PW/PL = 1.5. Sides gently arched, narrowing from posterior to anterior, and slightly constricted before hind angles, which projected backwards and subacute. Posterior margin widely protruding in the middle part, distinctly emarginate near hind angles.</paragraph>
<paragraph pageId="7" pageNumber="50">Elytra oval, widest at about basal 2/7, ELL/EW = 1.4. Sides weakly arched, gradually narrowing from widest part to apices, which narrowly rounded (Fig. 8G). Sutural striae present. Metathoracic wings fully developed.</paragraph>
<paragraph pageId="7" pageNumber="50">Prolegs robust, with basal three protarsomeres (Fig. 8C) strongly expanded: TW/BTW = 1.0. Protibiae (Fig. 8E) expanded towards apex. Profemora rather broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender and straight.</paragraph>
<paragraph pageId="7" pageNumber="50">Abdominal ventrite VIII (Fig. 8I) round at posterior edge, though an inconspicuous median notch at the median. Genital segment (Fig. 8J) with spiculum gastrale protruding about 3/8 of its length beyond anterior edge of epipleurite IX.</paragraph>
<paragraph lastPageId="8" lastPageNumber="51" pageId="7" pageNumber="50">
<paragraph id="2164185155ED122F5675227A08248225" pageId="7" pageNumber="50">Habitus (Fig. 3A) elongated oval, regularly convex and sublustrous. Well pigmented: mostly brown; mouthparts, basal four or five antennomeres and apical half of ultimate antennomere, protarsi, and apex of meso- and metatarsi yellowish. Dorsum continuously clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.</paragraph>
<paragraph id="308FE9403629ACAF1C6C705F82CE56EB" pageId="7" pageNumber="50">Head transverse, HW/HL = 1.5. Clypeofrontal suture absent. Clypeus with anterior margin slightly rounded. Compound eyes well developed, EW/HW = 0.1. Antennae (Fig. 8A) slender, AL/HW = 1.3; antennomere III a little shorter than II; VI with length/width = 0.5; XI elongated pear-shaped.</paragraph>
<paragraph id="40F17AC0C893EF9D095181C0D8D6DABB" pageId="7" pageNumber="50">Pronotum (Fig. 8B) transverse, widest just before hind angles, PW/PL = 1.5. Sides gently arched, narrowing from posterior to anterior, and slightly constricted before hind angles, which projected backwards and subacute. Posterior margin widely protruding in the middle part, distinctly emarginate near hind angles.</paragraph>
<paragraph id="2EA4DE8D0AC3B708FD25A3A2B651C8CF" pageId="7" pageNumber="50">Elytra oval, widest at about basal 2/7, ELL/EW = 1.4. Sides weakly arched, gradually narrowing from widest part to apices, which narrowly rounded (Fig. 8G). Sutural striae present. Metathoracic wings fully developed.</paragraph>
<paragraph id="3367B07D3DC44515843BDE10568090D7" pageId="7" pageNumber="50">Prolegs robust, with basal three protarsomeres (Fig. 8C) strongly expanded: TW/BTW = 1.0. Protibiae (Fig. 8E) expanded towards apex. Profemora rather broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender and straight.</paragraph>
<paragraph id="7611BE99D82BA81A15054FD6775AC8E8" pageId="7" pageNumber="50">Abdominal ventrite VIII (Fig. 8I) round at posterior edge, though an inconspicuous median notch at the median. Genital segment (Fig. 8J) with spiculum gastrale protruding about 3/8 of its length beyond anterior edge of epipleurite IX.</paragraph>
<paragraph id="AEF62ABE775A2E8B9B4630438CC3219B" lastPageId="8" lastPageNumber="51" pageId="7" pageNumber="50">
Aedeagus (Fig. 9A) long and slender, with median lobe gradually narrowing towards lanceolate apical part and terminated by round knob in dorsal view; opening of genital orifice situated on dorsal surface, deeply cut inwards on preapical left margin of median lobe. Ventral surface of the apex of the median lobe (Fig. 9G) inserted with
<pageBreakToken pageId="8" pageNumber="51" start="start">a</pageBreakToken>
<pageBreakToken id="E2296EB9E0E6E1A271A4FFEA645BF580" pageId="8" pageNumber="51" start="start">a</pageBreakToken>
row of 6 ventrally oriented setae (the bottom one is very short) on the left side and a row of 4 ventrally oriented setae on the right side. Parameres narrow, reaching about apical 1/5 of median lobe, each with 1 apical and 2 preapical setae, the apical one slightly shorter (Fig. 9E). In lateral view (Fig. 9C), median lobe regularly bent ventrad, gradually tapering apically. Endophallus with stylus quite slender, a cheliform complex just below the base of stylus, and a circular complex in basal region.
</paragraph>
<paragraph lastPageId="9" lastPageNumber="52" pageId="8" pageNumber="51">
<paragraph id="70A0EBF3651A3FCC2AA8B16EAB00CCCD" lastPageId="9" lastPageNumber="52" pageId="8" pageNumber="51">
Female. Similar to male in general appearance (Fig. 3B), including elytral apices (Fig. 8H), but distinguished by the following characteristics: protarsi (Fig. 8D)
<pageBreakToken pageId="9" pageNumber="52" start="start">simply</pageBreakToken>
<pageBreakToken id="0145EF9333A7AFDECA11D34504E76E4F" pageId="9" pageNumber="52" start="start">simply</pageBreakToken>
linear; protibiae (Fig. 8F) only slightly narrower; abdominal ventrite VIII (Fig. 9A) almost regularly rounded at posterior edge; genital segment as shown in Fig. 9B: spermatheca (Fig. 9B) curved in distal part, coiled and extended leftwards in proximal part.
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="53" pageId="9" pageNumber="52" type="diagnosis">
<paragraph pageId="9" pageNumber="52">Diagnosis.</paragraph>
<paragraph lastPageId="10" lastPageNumber="53" pageId="9" pageNumber="52">
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi">Ptomaphagus (s. str.) chenggongi</taxonomicName>
sp. n. has very similar aedeagus to
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae">Ptomaphagus (s. str.) tingtingae</taxonomicName>
sp. n., but can be distinguished from the latter by the following
<pageBreakToken pageId="10" pageNumber="53" start="start">characters</pageBreakToken>
<subSubSection id="02DE54337374660B4F9E0391A96056A7" lastPageId="10" lastPageNumber="53" pageId="9" pageNumber="52" type="diagnosis">
<paragraph id="C7E62A0C2E8DB39DB42F36EFB33AAEC7" pageId="9" pageNumber="52">Diagnosis.</paragraph>
<paragraph id="7D3F0EB06344FA76BA4E9B4E96D11AAD" lastPageId="10" lastPageNumber="53" pageId="9" pageNumber="52">
<taxonomicName id="7CD31D3ECB66BD51331D1FCB6EB6F96D" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi" status="sp. nov.">Ptomaphagus (s. str.) chenggongi</taxonomicName>
<taxonomicNameLabel id="0ED58AB7CA503B6DA535CA1814BEF34D" rank="species">sp. n.</taxonomicNameLabel>
has very similar aedeagus to
<taxonomicName id="7FED53DE25194C01E348E004927B1590" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae" status="sp. nov.">Ptomaphagus (s. str.) tingtingae</taxonomicName>
<taxonomicNameLabel id="77011BAE22478B52963071EEF3D3D098" rank="species">sp. n.</taxonomicNameLabel>
, but can be distinguished from the latter by the following
<pageBreakToken id="C28FF92A01A6EBF58584BA45E8A60147" pageId="10" pageNumber="53" start="start">characters</pageBreakToken>
: in
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="10" pageNumber="53" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi">Ptomaphagus (s. str.) chenggongi</taxonomicName>
sp. n., antennomere XI with length/width = 1.9, hind angles of pronotum subacute, spiculum gastrale of genital segment with ordinary width, right apicoventral piece of median lobe broad (Fig. 9H), apical seta of parameres slightly shorter than preapical setae, and spermatheca coiled and extended leftwards in proximal part; while in
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="10" pageNumber="53" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae">Ptomaphagus (s. str.) tingtingae</taxonomicName>
sp. n., antennomere XI with length/width = 1.3, hind angles of pronotum acute, spiculum gastrale of genital segment very narrow, right apicoventral piece of median lobe rather small (Fig. 9I), apical seta of parameres very shorter than preapical setae, and spermatheca coiled but not extended leftwards in proximal part.
<taxonomicName id="DC1465B9503862E3F56086D087B0ECD6" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="10" pageNumber="53" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi" status="sp. nov.">Ptomaphagus (s. str.) chenggongi</taxonomicName>
<taxonomicNameLabel id="3470E1B299E105CE65D3965538EA0995" rank="species">sp. n.</taxonomicNameLabel>
, antennomere XI with length/width = 1.9, hind angles of pronotum subacute, spiculum gastrale of genital segment with ordinary width, right apicoventral piece of median lobe broad (Fig. 9H), apical seta of parameres slightly shorter than preapical setae, and spermatheca coiled and extended leftwards in proximal part; while in
<taxonomicName id="44B78E39021D4C8495D3C41593014563" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="10" pageNumber="53" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae" status="sp. nov.">Ptomaphagus (s. str.) tingtingae</taxonomicName>
<taxonomicNameLabel id="9F65215A5FCA82B547E04C27E1DD68C5" rank="species">sp. n.</taxonomicNameLabel>
, antennomere XI with length/width = 1.3, hind angles of pronotum acute, spiculum gastrale of genital segment very narrow, right apicoventral piece of median lobe rather small (Fig. 9I), apical seta of parameres very shorter than preapical setae, and spermatheca coiled but not extended leftwards in proximal part.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="53" type="etymology">
<paragraph pageId="10" pageNumber="53">Etymology.</paragraph>
<paragraph pageId="10" pageNumber="53">The specific epithet is dedicated to Cheng-Gong Zheng (1624-1662), a military leader at the end of the Chinese Ming Dynasty, for his feats in 1662 when he defeated the forces of the Dutch East India Company and claimed Taiwan, bringing it under Chinese Han rule.</paragraph>
<subSubSection id="63B7F37E466D7B6EBCFDF1A8BD7BFE35" pageId="10" pageNumber="53" type="etymology">
<paragraph id="B835D18C8212FA43575E858145BD1EC9" pageId="10" pageNumber="53">Etymology.</paragraph>
<paragraph id="3B27F5F07FDCE1A5AC94564ACAB33460" pageId="10" pageNumber="53">The specific epithet is dedicated to Cheng-Gong Zheng (1624-1662), a military leader at the end of the Chinese Ming Dynasty, for his feats in 1662 when he defeated the forces of the Dutch East India Company and claimed Taiwan, bringing it under Chinese Han rule.</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="53" type="distribution">
<paragraph pageId="10" pageNumber="53">Distribution.</paragraph>
<paragraph pageId="10" pageNumber="53">China (Taiwan) (Fig. 12).</paragraph>
<subSubSection id="6C93435B463005F35CA2D7EFE3B35B6D" pageId="10" pageNumber="53" type="distribution">
<paragraph id="95FCD0E1E45C39B90ED20E2CC193D335" pageId="10" pageNumber="53">Distribution.</paragraph>
<paragraph id="7F8F3A8E2DE76BC8D43627C0DA682EAA" pageId="10" pageNumber="53">China (Taiwan) (Fig. 12).</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="8AD3EFC9A12D160F9851EA2E0DFA2B01" ID-GBIF-Taxon="127887148" LSID="urn:lsid:plazi:treatment:8AD3EFC9A12D160F9851EA2E0DFA2B01" httpUri="http://treatment.plazi.org/id/8AD3EFC9A12D160F9851EA2E0DFA2B01" lastPageNumber="46" pageId="3" pageNumber="46">
<subSubSection id="2AFFFD6CDD65C88B972C640CD3841797" pageId="3" pageNumber="46" type="multiple">
<paragraph id="E07E84EFE39DA9D21E0936AAD7FB0C47" pageId="3" pageNumber="46">Taxon classification Animalia Coleoptera Leiodidae</paragraph>
<paragraph id="E07E84EFE39DA9D21E0936AAD7FB0C47" pageId="3" pageNumber="46">
Taxon classification Animalia
<taxonomicName id="015975230BD03BD118E0C5BE8DD002E4" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="25646B79E9B0828BF913B451B72CA1A2" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="746C991C445A56772E90E3BFF391496C" pageId="3" pageNumber="46" type="nomenclature">
<paragraph id="2FEE2360979BB2B2B579C632A11FAF30" pageId="3" pageNumber="46">
<taxonomicName id="CE7B6AF209E9C447BB1B5B9B8F6071B8" ID-CoL="4Q9F7" authority="Sokolowski, 1957" authorityName="Sokolowski" authorityYear="1957" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="kuntzeni">Ptomaphagus (s. str.) kuntzeni Sokolowski, 1957</taxonomicName>
<taxonomicName id="CE7B6AF209E9C447BB1B5B9B8F6071B8" ID-CoL="4Q9F7" authority="Sokolowski, 1957" authorityName="Sokolowski" authorityYear="1957" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="kuntzeni">Ptomaphagus (s. str.) kuntzeni Sokolowski, 1957</taxonomicName>
Figs 1; 4
</paragraph>
</subSubSection>
<subSubSection id="E52634F3DFD11E6AE90111172E05D5F4" pageId="3" pageNumber="46" type="reference_group">
<paragraph id="6654D6D19D8475FD1460F019B2D9A706" pageId="3" pageNumber="46">
<taxonomicName id="F797F763CACE4EED130C45DFC2172808" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="kuntzeni">Ptomaphagus (s. str.) kuntzeni</taxonomicName>
<taxonomicName id="72632C6E50D84716969D1C5B596E12AA" authority="Sokolowski 1957: 140" authorityName="Sokolowski" authorityPageNumber="140" authorityYear="1957" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="kuntzeni">
Ptomaphagus (s. str.) kuntzeni
<bibRefCitation id="16DEAB0298EFA74A1504FB2035C1FA2D" author="Sokolowski, K" journalOrPublisher="Deutsche entomologische Zeitschrift (N. F.)" pageId="18" pageNumber="61" pagination="140 - 142" title="Zwei neue japanische Catopiden (Col. Catopidae) (Catopiden-Studien 6)." volume="4" year="1957">Sokolowski 1957</bibRefCitation>
: 140 (
<taxonomicName id="D947AC1EF4E5674DFF56A649CF45C9AF" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
: 140
</taxonomicName>
(
<taxonomicName id="D947AC1EF4E5674DFF56A649CF45C9AF" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
; type locality: [JAPAN] Hagi (? Landschaft Jamagutshi, Honshiu); ZMHB);
<bibRefCitation id="DBF1052D77E4782140244D5E3C0ABF94" author="Szymczakowski, W" journalOrPublisher="Acta Zoologica Cracoviensia" pageId="18" pageNumber="61" pagination="55 - 289" title="Analyse systematique et zoogeographique des Catopidae (Coleoptera) de la region orientale." volume="9" year="1964">Szymczakowski 1964</bibRefCitation>
: 63 (
<taxonomicName id="FF69A2C20D73124734576F1D6BC9625F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="FF69A2C20D73124734576F1D6BC9625F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
; female description; taxonomic remarks);
<bibRefCitation id="BABC7F8BD6DA2F862F289168DA83C90E" author="Nishikawa, M" journalOrPublisher="Coleopterists' Association of Japan, Tokyo" pageId="18" pageNumber="61" pagination="1 - 6" title="Check-list of Coleoptera of Japan" year="1983">Nishikawa 1983</bibRefCitation>
: 1 (
<taxonomicName id="D509ACED989EE7B5AB6CAC19772857AF" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="D509ACED989EE7B5AB6CAC19772857AF" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in check-list);
<bibRefCitation id="D4E778FA6C1288120B9C0A13B4C23E77" author="Harusawa, K" journalOrPublisher="Nature of Odamiyama" pageId="17" pageNumber="60" pagination="241 - 247" title="Choleved [sic] beetles (Coleoptera) collected from Odamiyama, Ehime Prefecture." volume="2" year="2000">Harusawa and Yamamoto 2000</bibRefCitation>
: 242 (
<taxonomicName id="4C48DCE98F7C4F17D09DEFCF08870914" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="4C48DCE98F7C4F17D09DEFCF08870914" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
; distribution);
<bibRefCitation id="FAF92F82EFC15DF28970D80C2EFF18BA" author="Hayashi, Y" journalOrPublisher="Entomological Review of Japan" pageId="17" pageNumber="60" pagination="189 - 190" title="Cholevine beetles (Coleoptera: Leiodidae) occurring on Toku-no-shima Island, the Ryukyus, Southwest Japan." volume="65" year="2010">Hayashi and Nishikawa 2010</bibRefCitation>
: 190 (
<taxonomicName id="27BC2690E48908F8107BA888B6886625" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="27BC2690E48908F8107BA888B6886625" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
; distribution);
<bibRefCitation id="9BF00271BB8528ECB8D49F8F5EA27836" author="Perreau, M" journalOrPublisher="Memoires de la Societe entomologique de France" pageId="18" pageNumber="61" pagination="1 - 460" title="Catalogue des Coleopteres Leiodidae, Cholevinae et Platypsyllinae." volume="4" year="2000">Perreau 2000</bibRefCitation>
: 363 (
<taxonomicName id="24F790C1F0B008753F6A5CED999352B5" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="24F790C1F0B008753F6A5CED999352B5" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.); in catalogue);
<bibRefCitation id="92015D021A32D39698B21670E419AC8B" author="Perreau, M" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="18" pageNumber="61" pagination="133 - 203" title="Family Leiodidae Fleming, 1821." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 2: Hydrophiloidea - Histeroidea - Staphylinoidea" year="2004">Perreau 2004</bibRefCitation>
: 178 (
<taxonomicName id="044EE1201F2FDA2FB691FCBA19C0114F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="044EE1201F2FDA2FB691FCBA19C0114F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in catalogue);
<bibRefCitation id="CFED114F57DD01629938EAAE0FB87280" author="Nishikawa, M" journalOrPublisher="Special Publication of the Japanese Society of Scarabaeoidology" pageId="18" pageNumber="61" pagination="97 - 102" title="A new Ptomaphagus (Coleoptera: Leiodidae: Cholevinae) from Northwest Thailand." volume="1" year="2011">Nishikawa 2011</bibRefCitation>
: 100 (
<taxonomicName id="C9A427C98450B1ADABA4BFCD0EFAC870" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="C9A427C98450B1ADABA4BFCD0EFAC870" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; distribution; notes);
<bibRefCitation id="736692B9CFD00CDAA76C130BF4ECCE02" author="Nishikawa, M" journalOrPublisher="Tokyo" pageId="18" pageNumber="61" pagination="267 - 278" title="The underground fauna of Agyrtidae and the subfamily Cholevinae of Leiodidae (Coleoptera) in eastern Shikoku, Southwest Japan, with a summary of the habitat diversity of some Japanese cholevines." volumeTitle="Elytra" year="2012">Nishikawa et al. 2012</bibRefCitation>
: 274 (
<taxonomicName id="975E5B0D7890F370269B11B846B37ED6" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="975E5B0D7890F370269B11B846B37ED6" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; distribution);
<bibRefCitation id="38593BF288A211FA6EE5ED7EA3EC109B" author="Perreau, M" editor="Loebl, I" journalOrPublisher="Brill, Leiden &amp; Boston" pageId="18" pageNumber="61" pagination="180 - 290" title="Family Leiodidae Fleming, 1821." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 2 / 1. Hydrophiloidea - Staphylinoidea" year="2015">Perreau 2015</bibRefCitation>
: 249 (
<taxonomicName id="976D4CC33E8567B72398C2E855C450C3" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="976D4CC33E8567B72398C2E855C450C3" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in catalogue);
<bibRefCitation id="2A696192F20478A4CFFB122CC392D485" pageId="3" pageNumber="46">Wang et al. in press</bibRefCitation>
(
<taxonomicName id="A6900A3405F5ED3B888EC1865DBB2FB7" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="A6900A3405F5ED3B888EC1865DBB2FB7" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.); redescription; distribution; remarks).
</paragraph>
<paragraph id="8B74D9CEBECAC3A00F5784F6BC04B340" pageId="3" pageNumber="46">
<taxonomicName id="89387054CD09B6E80C0AF9A641A0C6E9" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus amamianus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="amamianus">Ptomaphagus (s. str.) amamianus</taxonomicName>
<taxonomicName id="A62E45D3FE86F7F687A80E3FF7CFBCD6" authority="Nakane 1963: 42" authorityName="Nakane" authorityPageNumber="42" authorityYear="1963" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus amamianus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="amamianus">
Ptomaphagus (s. str.) amamianus
<bibRefCitation id="EC9E4B575460533831B0E32642FEB223" author="Nakane, T" journalOrPublisher="Fragmenta Coleopterologica Kyoto" pageId="18" pageNumber="61" pagination="40 - 42" title="New or little-known Coleoptera from Japan and its adjacent regions. XXI." volume="10" year="1963">Nakane 1963</bibRefCitation>
: 42 (
<taxonomicName id="DC7F9C360BBC44B35CD379DE5FA96271" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
: 42
</taxonomicName>
(
<taxonomicName id="DC7F9C360BBC44B35CD379DE5FA96271" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
; type locality: [JAPAN] Naze, Amami-Oshima);
<bibRefCitation id="5227470D1773EBCDD33D72A26F643409" author="Hayashi, Y" journalOrPublisher="Entomological Review of Japan" pageId="17" pageNumber="60" pagination="1 - 6" title="Catopidae from Amami-Ohshima Is., Japan (Col.)." volume="12" year="1969">Hayashi 1969</bibRefCitation>
: 2 (
<taxonomicName id="A8F0EA5FA49826B3C90CFAD753FAC7FC" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="A8F0EA5FA49826B3C90CFAD753FAC7FC" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
; characteristic figures; distribution);
<bibRefCitation id="28F199D8274EAD516F76327034417A73" author="Nishikawa, M" journalOrPublisher="Coleopterists' Association of Japan, Tokyo" pageId="18" pageNumber="61" pagination="1 - 6" title="Check-list of Coleoptera of Japan" year="1983">Nishikawa 1983</bibRefCitation>
: 1 (
<taxonomicName id="55C6E0652874E762340FE994126E844B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="55C6E0652874E762340FE994126E844B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in check-list);
<bibRefCitation id="3103FB4F2E4357BBEF938BDCD7EA57D8" author="Perreau, M" journalOrPublisher="Revue Suisse de Zoologie" pageId="18" pageNumber="61" pagination="283 - 297" title="Contribution a la connaissance des Cholevidae du Japon et de Taiwan (Coleoptera)." url="10.5962/bhl.part.79946" volume="103" year="1996">Perreau 1996</bibRefCitation>
: 284 (
<taxonomicName id="ECCBA6981AB51F568E2071BA06E5A0A2" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="ECCBA6981AB51F568E2071BA06E5A0A2" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
; distribution);
<bibRefCitation id="1D3583CF93549EF2A053A8A6CAF60D65" author="Perreau, M" journalOrPublisher="Memoires de la Societe entomologique de France" pageId="18" pageNumber="61" pagination="1 - 460" title="Catalogue des Coleopteres Leiodidae, Cholevinae et Platypsyllinae." volume="4" year="2000">Perreau 2000</bibRefCitation>
: 362 (
<taxonomicName id="34CEA6874B4B5DFC89332AEB4878C1A6" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="34CEA6874B4B5DFC89332AEB4878C1A6" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.); in catalogue);
<bibRefCitation id="831F3E19314EAE5FCA8A15B963EE2207" author="Perreau, M" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="18" pageNumber="61" pagination="133 - 203" title="Family Leiodidae Fleming, 1821." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 2: Hydrophiloidea - Histeroidea - Staphylinoidea" year="2004">Perreau 2004</bibRefCitation>
: 178 (
<taxonomicName id="762463A5E061B6D09FA0B5E5506E9B8B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="762463A5E061B6D09FA0B5E5506E9B8B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in catalogue);
<bibRefCitation id="5CB7DA6FA8AE5D8B84ED5CF7DB2D0FD0" author="Hayashi, Y" journalOrPublisher="Entomological Review of Japan" pageId="17" pageNumber="60" pagination="189 - 190" title="Cholevine beetles (Coleoptera: Leiodidae) occurring on Toku-no-shima Island, the Ryukyus, Southwest Japan." volume="65" year="2010">Hayashi and Nishikawa 2010</bibRefCitation>
: 190 (
<taxonomicName id="1A7799F56358E91C28902148B2A4413E" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="1A7799F56358E91C28902148B2A4413E" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
; distribution);
<bibRefCitation id="0F1E5309208BFA5ED0B311B2DC379BC5" author="Perreau, M" editor="Loebl, I" journalOrPublisher="Brill, Leiden &amp; Boston" pageId="18" pageNumber="61" pagination="180 - 290" title="Family Leiodidae Fleming, 1821." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 2 / 1. Hydrophiloidea - Staphylinoidea" year="2015">Perreau 2015</bibRefCitation>
: 249 (
<taxonomicName id="74C869FC7478E80F85074BC9474ED782" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="74C869FC7478E80F85074BC9474ED782" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in catalogue);
<bibRefCitation id="45AFF2C29E68EF392CDEA7C2E66FA671" pageId="3" pageNumber="46">Wang et al. in press</bibRefCitation>
(
<taxonomicName id="C3C65B9E89B286384D47BF2196462EC9" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="C3C65B9E89B286384D47BF2196462EC9" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.); synonymy with
<taxonomicName id="121E4E84BFCCD54DAF5E18EEB86EB8DA" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="kuntzeni">Ptomaphagus kuntzeni</taxonomicName>
<taxonomicName id="121E4E84BFCCD54DAF5E18EEB86EB8DA" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="kuntzeni">Ptomaphagus kuntzeni</taxonomicName>
).
</paragraph>
</subSubSection>
@ -172,7 +182,9 @@ Figs 1; 4
<geoCoordinate id="50F15B655C5FB9742246A332A99E2187" direction="north" orientation="latitude" precision="925" value="24.266666">24°16'N</geoCoordinate>
,
<geoCoordinate id="337942533FD10CCC7B0FE2215FFE7811" direction="east" orientation="longitude" precision="925" value="121.0">121°00'E</geoCoordinate>
] / 2230 m 30.IV.-4.V.[19]90 / A. Smetana [leg.] (T32) // PTOMAPHAGUS / sp. (SMNS).
] / 2230 m 30.IV.-4.V.[19]90 / A. Smetana [leg.] (T32) //
<taxonomicName id="43B9102F6F47AC6A730072BEA73BBC8E" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">PTOMAPHAGUS</taxonomicName>
/ sp. (SMNS).
</paragraph>
</subSubSection>
<subSubSection id="1989C3E95FD71AB3B4299102B7426385" pageId="3" pageNumber="46" type="remarks">
@ -180,7 +192,7 @@ Figs 1; 4
<paragraph id="A6D40ABF29B05143AAD31B9ED6ADF08D" pageId="3" pageNumber="46">
<bibRefCitation id="E8631D7AC66C3685ABD8F4D6FD672820" author="Perreau, M" journalOrPublisher="Revue Suisse de Zoologie" pageId="18" pageNumber="61" pagination="283 - 297" title="Contribution a la connaissance des Cholevidae du Japon et de Taiwan (Coleoptera)." url="10.5962/bhl.part.79946" volume="103" year="1996">Perreau (1996)</bibRefCitation>
recorded the species from Taiwan Island under the name
<taxonomicName id="181CE74E35919FE18AFDD7D288E3332A" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus amamianus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="amamianus">Ptomaphagus (s. str.) amamianus</taxonomicName>
<taxonomicName id="181CE74E35919FE18AFDD7D288E3332A" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus amamianus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="amamianus">Ptomaphagus (s. str.) amamianus</taxonomicName>
. We re-examined the specimens concerned and found that they have a partly different aedeagus from that of Japanese specimens (Fig. 4A, C, E): in specimens from Taiwan, the right apicoventral piece of median lobe is slender and subtriangular (Fig. 4B, D); while in specimens from Japan, it is shorter and subround (Fig. 4F). However, we consider that it is an intraspecific geographic variation because they have the basically identical shape of aedeagi and spermathecae, and no distinct differences in their external morphology.
</paragraph>
</subSubSection>

View file

@ -1,115 +1,123 @@
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<treatment id="D92546A747598AA6B5FA9FD4F4A0FDD9" ID-GBIF-Taxon="127887146" LSID="urn:lsid:plazi:treatment:D92546A747598AA6B5FA9FD4F4A0FDD9" httpUri="http://treatment.plazi.org/id/D92546A747598AA6B5FA9FD4F4A0FDD9" lastPageId="6" lastPageNumber="49" pageId="3" pageNumber="46">
<subSubSection id="6792FA13B57D3A13A7E2E914CC3D7CF8" pageId="3" pageNumber="46" type="multiple">
<paragraph id="F81496574F3EB3D8FBE1DF72F94E0495" pageId="3" pageNumber="46">Taxon classification Animalia Coleoptera Leiodidae</paragraph>
<paragraph id="F81496574F3EB3D8FBE1DF72F94E0495" pageId="3" pageNumber="46">
Taxon classification Animalia
<taxonomicName id="814A7D92199BF80B5B9110A883893282" class="Insecta" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="6C8737929F3BB1BB26712D56B68C1914" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="581D5CEC5110F6958F9E26A6F29C2DCA" pageId="3" pageNumber="46" type="nomenclature">
<paragraph id="DBDD16FC2CE31A2E88EC821D129D363B" pageId="3" pageNumber="46">
<taxonomicName id="FAC416EB7EA66B75DC1E9491CAC1AD7C" ID-CoL="6WM2B" authority="Nishikawa, 1993" authorityName="Nishikawa" authorityYear="1993" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">Ptomaphagus (s. str.) yasutoshii Nishikawa, 1993</taxonomicName>
<taxonomicName id="FAC416EB7EA66B75DC1E9491CAC1AD7C" ID-CoL="6WM2B" authority="Nishikawa, 1993" authorityName="Nishikawa" authorityYear="1993" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">Ptomaphagus (s. str.) yasutoshii Nishikawa, 1993</taxonomicName>
Figs 2; 5, 6, 7
</paragraph>
</subSubSection>
<subSubSection id="64B35401DE2009A41EB5A3CFBD34165C" lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47" type="reference_group">
<paragraph id="76E03B604179CB55B422EB4AE1B52632" pageId="4" pageNumber="47">
<taxonomicName id="51F4151E8C9430736C22B6325CE9B76F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">
<pageBreakToken id="CFFFFC5D69555229DDB4ABF7E1B249BD" pageId="4" pageNumber="47" start="start">Ptomaphagus</pageBreakToken>
(s. str.) yasutoshii
</taxonomicName>
<taxonomicName id="890A90966DF9F4711BDD676D8A51CB54" authority="Nishikawa 1993: 123" authorityName="Nishikawa" authorityPageNumber="123" authorityYear="1993" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">
<pageBreakToken id="2BCD7688F99EC507C38334EDF3A03114" pageId="4" pageNumber="47" start="start">Ptomaphagus</pageBreakToken>
(s. str.) yasutoshii
<bibRefCitation id="A271AA627CE672952FEFCB6DFC7023C9" author="Nishikawa, M" journalOrPublisher="Elytra (Tokyo)" pageId="18" pageNumber="61" pagination="123 - 128" title="Occurrence of a new apterous species of Ptomaphagus (Coleoptera, Cholevidae) in Taiwan." volume="21" year="1993">Nishikawa 1993</bibRefCitation>
: 123 (
<taxonomicName id="866B483CD05A790C29996E5E9503640A" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
: 123
</taxonomicName>
(
<taxonomicName id="866B483CD05A790C29996E5E9503640A" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.); type locality: Taiwan, Nantou Hsien, near Tsuifeng, 2200 m; NSMT);
<bibRefCitation id="AFE7EB1D2DA1CED3263D796D18E0A500" author="Perreau, M" journalOrPublisher="Memoires de la Societe entomologique de France" pageId="18" pageNumber="61" pagination="1 - 460" title="Catalogue des Coleopteres Leiodidae, Cholevinae et Platypsyllinae." volume="4" year="2000">Perreau 2000</bibRefCitation>
: 367 (
<taxonomicName id="328DF564FBBFF6D9F63D2948131F9ECA" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="328DF564FBBFF6D9F63D2948131F9ECA" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.); in catalogue);
<bibRefCitation id="8E3A5C4878EE98143E5D95C4C06AB305" author="Perreau, M" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="18" pageNumber="61" pagination="133 - 203" title="Family Leiodidae Fleming, 1821." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 2: Hydrophiloidea - Histeroidea - Staphylinoidea" year="2004">Perreau 2004</bibRefCitation>
: 178 (
<taxonomicName id="3C353066F0583BAB596F9FD35C2994F2" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="3C353066F0583BAB596F9FD35C2994F2" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in catalogue);
<bibRefCitation id="2F11D79F9955F65D13BD5747CC9FE5ED" author="Perreau, M" editor="Loebl, I" journalOrPublisher="Brill, Leiden &amp; Boston" pageId="18" pageNumber="61" pagination="180 - 290" title="Family Leiodidae Fleming, 1821." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 2 / 1. Hydrophiloidea - Staphylinoidea" year="2015">Perreau 2015</bibRefCitation>
: 250 (
<taxonomicName id="B8094B191D4CB0589C2FDF565A05E1A4" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="B8094B191D4CB0589C2FDF565A05E1A4" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in catalogue).
</paragraph>
<paragraph id="232F2EF376D4889402D2AACC3E22B15A" lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47">
<taxonomicName id="F4812FB2C672A78010826953E705F2A5" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="smetanai">Ptomaphagus (s. str.) smetanai</taxonomicName>
<taxonomicName id="28388192F4276D3A17C67606FC12095F" authority="Perreau 1996: 285" authorityName="Perreau" authorityPageNumber="285" authorityYear="1996" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="smetanai">
Ptomaphagus (s. str.) smetanai
<bibRefCitation id="5E52AB7590BAD492F50E8438F43E2116" author="Perreau, M" journalOrPublisher="Revue Suisse de Zoologie" pageId="18" pageNumber="61" pagination="283 - 297" title="Contribution a la connaissance des Cholevidae du Japon et de Taiwan (Coleoptera)." url="10.5962/bhl.part.79946" volume="103" year="1996">Perreau 1996</bibRefCitation>
: 285 (
<taxonomicName id="B33D73245FD68901A5D952E7048F706B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
: 285
</taxonomicName>
(
<taxonomicName id="B33D73245FD68901A5D952E7048F706B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.); type locality: Taiwan, Hualien Hsien, Taroko, N. P. Duodyatunshan, 2660 m; MHNG);
<bibRefCitation id="A5B509EF2BF5575A314A97DF0F83FA1A" author="Perreau, M" journalOrPublisher="Memoires de la Societe entomologique de France" pageId="18" pageNumber="61" pagination="1 - 460" title="Catalogue des Coleopteres Leiodidae, Cholevinae et Platypsyllinae." volume="4" year="2000">Perreau 2000</bibRefCitation>
: 364 (
<taxonomicName id="C20C62A97C56C603B15A11BD1F8F61E7" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="C20C62A97C56C603B15A11BD1F8F61E7" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.); in catalogue);
<bibRefCitation id="F00082B1BDDD792653E512F5628208AE" author="Perreau, M" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="18" pageNumber="61" pagination="133 - 203" title="Family Leiodidae Fleming, 1821." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 2: Hydrophiloidea - Histeroidea - Staphylinoidea" year="2004">Perreau 2004</bibRefCitation>
: 178 (
<taxonomicName id="F68FF3AD5D3CEC9F9613B1087DA001F8" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">
<taxonomicName id="F68FF3AD5D3CEC9F9613B1087DA001F8" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">
<pageBreakToken id="9361141D2411A96DBABD29A25EBD90B1" pageId="5" pageNumber="48" start="start">Ptomaphagus</pageBreakToken>
(Ptomaphagus)
</taxonomicName>
; in catalogue);
<bibRefCitation id="4814220F2F614BBB17DC07E72E0BABF4" author="Perreau, M" editor="Loebl, I" journalOrPublisher="Brill, Leiden &amp; Boston" pageId="18" pageNumber="61" pagination="180 - 290" title="Family Leiodidae Fleming, 1821." volumeTitle="Catalogue of Palaearctic Coleoptera. Volume 2 / 1. Hydrophiloidea - Staphylinoidea" year="2015">Perreau 2015</bibRefCitation>
: 249 (
<taxonomicName id="E244FD17F20104882541F982C944B6F7" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
<taxonomicName id="E244FD17F20104882541F982C944B6F7" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
; in catalogue). Syn. n.
</paragraph>
</subSubSection>
@ -117,34 +125,34 @@ Figs 2; 5, 6, 7
<paragraph id="7D65CC34CEE7E9F2669857F075823C68" pageId="5" pageNumber="48">Material examined.</paragraph>
<paragraph id="6931696A1A16D5323E3DF50AE87DAE88" pageId="5" pageNumber="48">
Type material. Holotype of
<taxonomicName id="5FDBF356BB0DBD82D3A71A1721395546" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="yasutoshii">Ptomaphagus yasutoshii</taxonomicName>
<taxonomicName id="5FDBF356BB0DBD82D3A71A1721395546" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="yasutoshii">Ptomaphagus yasutoshii</taxonomicName>
: ♂, (Near TSIFENG [翠峰, ca.
<geoCoordinate id="F460112E9ECF16C6250367D448A3365E" direction="north" orientation="latitude" precision="925" value="24.1">24°06'N</geoCoordinate>
,
<geoCoordinate id="B843165688AF5DE6B2425BF80CFA6E72" direction="east" orientation="longitude" precision="925" value="121.183334">121°11'E</geoCoordinate>
]) / Nantou - Hsien, / TAIWAN (2200 m) / Aug. 25th, 1974 / Coll. Y. Shibata // HOLOTYPE /
<taxonomicName id="1DA3D9DE8A6C9B07EFDA899259ADEC6B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="1DA3D9DE8A6C9B07EFDA899259ADEC6B" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.) /
<taxonomicName id="9E5256B11365B8CBE02373310E4F1CD1" lsidName="yasutoshii" pageId="5" pageNumber="48" rank="species" species="yasutoshii">yasutoshii</taxonomicName>
<taxonomicName id="9E5256B11365B8CBE02373310E4F1CD1" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="yasutoshii" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="yasutoshii">yasutoshii</taxonomicName>
/ M. Nishikawa, 1993 (NSMT). Allotype of
<taxonomicName id="8F382617E94D7FB81CF3D9E00C78B7CE" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="yasutoshii">Ptomaphagus yasutoshii</taxonomicName>
<taxonomicName id="8F382617E94D7FB81CF3D9E00C78B7CE" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="yasutoshii">Ptomaphagus yasutoshii</taxonomicName>
: ♀, (Near TSIFENG [翠峰, ca.
<geoCoordinate id="8EA922CF972749267904002D286007F5" direction="north" orientation="latitude" precision="925" value="24.1">24°06'N</geoCoordinate>
,
<geoCoordinate id="AF3225E43CD6A664088D64D0D99E8FE5" direction="east" orientation="longitude" precision="925" value="121.183334">121°11'E</geoCoordinate>
]) / Nantou - Hsien, / TAIWAN (2200 m) / July 27th, 1974 / Coll. Y. Shibata // ALLOTYPE /
<taxonomicName id="A3CF0B3AFE0D60975AD1CA819A98FB00" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(
<taxonomicName id="BDA905C7DF2115BB0E57364473B6991D" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
) /
<taxonomicName id="22537E686026C69266161D2003AF8A44" lsidName="yasutoshii" pageId="5" pageNumber="48" rank="species" species="yasutoshii">yasutoshii</taxonomicName>
M. Nishikawa, / 1993 / Design. M. Nishikawa, 1993 / # MNIC 124929Ch2S ♀ // ♀ (CMNE). Holotype of
<taxonomicName id="9DD23B02C3D005330C62F0E7B22C5EC8" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="smetanai">Ptomaphagus smetanai</taxonomicName>
<taxonomicName id="A1BC815E45D4F2D9764A6BC64FFDF706" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
/
<taxonomicName id="34CC5BB85E12E98D21E58E444840FD34" authority="M. Nishikawa" authorityName="M. Nishikawa" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="yasutoshii" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="yasutoshii" subGenus="Ptomaphagus">yasutoshii M. Nishikawa</taxonomicName>
, / 1993 / Design. M. Nishikawa, 1993 / # MNIC 124929Ch2S ♀ // ♀ (CMNE). Holotype of
<taxonomicName id="9DD23B02C3D005330C62F0E7B22C5EC8" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="smetanai">Ptomaphagus smetanai</taxonomicName>
: ♂, TAIWAN Hualien / Hsien, Taroko, N. P / [Mt.] Duodyatunshan [多加屯山, ca.
<geoCoordinate id="8E9C3D5305EFE9970E8D1EC7A7029588" direction="north" orientation="latitude" precision="925" value="24.3">24°18'N</geoCoordinate>
,
<geoCoordinate id="8CB562FA9C0BDDC08DE1474F66266DA9" direction="east" orientation="longitude" precision="925" value="121.5">121°30'E</geoCoordinate>
] / 2650 m, 8.-13.V.[19]90 / A. Smetana [leg.] (T57) // HOLOTYPE / PTOMAPHAGUS SMETANAI / M. Perreau det. 1994 // MHNG / ENTO / 00003352 (MHNG).
] / 2650 m, 8.-13.V.[19]90 / A. Smetana [leg.] (T57) // HOLOTYPE /
<taxonomicName id="F7E84D1926F8CFC95C5D6355C74C7711" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="smetanai">PTOMAPHAGUS SMETANAI</taxonomicName>
/ M. Perreau det. 1994 // MHNG / ENTO / 00003352 (MHNG).
</paragraph>
</subSubSection>
<subSubSection id="BF95DD2DA6B78EE75AFEA14AB393B21C" pageId="5" pageNumber="48" type="materials_examined">
@ -155,9 +163,9 @@ M. Nishikawa, / 1993 / Design. M. Nishikawa, 1993 / # MNIC 124929Ch2S ♀ // ♀
,
<geoCoordinate id="8552F5B09D83026E781A6BBB2C972B48" direction="east" orientation="longitude" precision="925" value="120.95">120°57'E</geoCoordinate>
] / TAIWAN / 20.V.1981 / N. ITO [leg.] //
<taxonomicName id="D7CC967A1E33AB639C4911DFD34631EF" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
/ (s. str.)
<taxonomicName id="FBE55700AF60CA3F273A7588DE251CA0" lsidName="yasutoshii" pageId="5" pageNumber="48" rank="species" species="yasutoshii">yasutoshii</taxonomicName>
<taxonomicName id="D7CC967A1E33AB639C4911DFD34631EF" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
/
<taxonomicName id="187349712506AC49DAC26BF450AE6D6F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="yasutoshii" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="yasutoshii">(s. str.) yasutoshii</taxonomicName>
/ MS / Exs. M. NISHIKAWA, 1992 (CYHK).
</paragraph>
</subSubSection>
@ -165,9 +173,9 @@ M. Nishikawa, / 1993 / Design. M. Nishikawa, 1993 / # MNIC 124929Ch2S ♀ // ♀
<paragraph id="29D55890BCC3931AF419FAEB5DD50F2A" pageId="5" pageNumber="48">Redescription.</paragraph>
<paragraph id="36B61E2008CCFAC8D25307116D3C5B43" pageId="5" pageNumber="48">
Male. EBL: 4.3 mm in holotype of
<taxonomicName id="967234F1582D0450373609EA360B2902" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="yasutoshii">Ptomaphagus yasutoshii</taxonomicName>
<taxonomicName id="967234F1582D0450373609EA360B2902" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="yasutoshii">Ptomaphagus yasutoshii</taxonomicName>
and 4.9 mm in holotype of
<taxonomicName id="56C17C04C67F5666D6195E4CF9F9EC53" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="smetanai">Ptomaphagus smetanai</taxonomicName>
<taxonomicName id="56C17C04C67F5666D6195E4CF9F9EC53" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="smetanai">Ptomaphagus smetanai</taxonomicName>
). Length of different body parts: HL: AL: PL: ELL = 0.78: 1.51: 1.08: 2.25 mm; width: HW: EW: PW: ELW = 1.22: 0.10: 1.79: 1.90 mm. Proportion of antennomeres from base to tip in
<normalizedToken id="8DBD85370D71BB609C4E3024ED4957EF" originalValue="μm">μm</normalizedToken>
(length
@ -216,20 +224,20 @@ Female. Similar to male in general appearance (Fig. 2B, D), including elytral ap
<paragraph id="82CED539974619453FBC764B5A85B510" pageId="6" pageNumber="49">Remarks.</paragraph>
<paragraph id="76E98CFFA2C4D695FC861C200D2CCFB2" pageId="6" pageNumber="49">
This species is exceptional in the genus
<taxonomicName id="F3471329FDD491CA15342E2B38B2039D" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="F3471329FDD491CA15342E2B38B2039D" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
in the following characters: metathoracic wings absent; aedeagus strongly asymmetrical, median lobe turning rightwards in apical part; spermatheca spiral-shaped, discoidal in distal part.
</paragraph>
<paragraph id="149B5485E42A766B3AAD5B66BCCF1F70" pageId="6" pageNumber="49">
In addition, the holotypes of
<taxonomicName id="F5E2658B63A3FEAEC4B70A26D673D0CE" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="yasutoshii">Ptomaphagus yasutoshii</taxonomicName>
<taxonomicName id="F5E2658B63A3FEAEC4B70A26D673D0CE" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="yasutoshii">Ptomaphagus yasutoshii</taxonomicName>
and
<taxonomicName id="730CFBC175F2983ADB34DAB19F513277" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="smetanai">Ptomaphagus smetanai</taxonomicName>
<taxonomicName id="730CFBC175F2983ADB34DAB19F513277" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="smetanai">Ptomaphagus smetanai</taxonomicName>
have almost identical aedeagal shape (Fig. 6
<normalizedToken id="5EF2EF99D0C2FB6374D208ED038171B4" originalValue="AD">A-D</normalizedToken>
) and no distinct differences in the external morphology, except some variations exist in the shape of aedeagal apex: the right apicoventral piece of median lobe of
<taxonomicName id="60A426BDB9F1AD61F8DC3BA6AAFDEE3F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="yasutoshii">Ptomaphagus yasutoshii</taxonomicName>
<taxonomicName id="60A426BDB9F1AD61F8DC3BA6AAFDEE3F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="yasutoshii">Ptomaphagus yasutoshii</taxonomicName>
(Fig. 6H) is somewhat wider than that of
<taxonomicName id="84DC3F6AC9B091EDA3C00592FD9E411E" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="smetanai">Ptomaphagus smetanai</taxonomicName>
<taxonomicName id="84DC3F6AC9B091EDA3C00592FD9E411E" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="smetanai">Ptomaphagus smetanai</taxonomicName>
(Fig. 6I). However, such differences, which fall within intraspecific variability, does not prevent us from synonymizing the two species.
</paragraph>
</subSubSection>

View file

@ -1,229 +1,244 @@
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<paragraph pageId="2" pageNumber="45">Taxon classification Animalia Coleoptera Leiodidae</paragraph>
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Taxon classification Animalia
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<taxonomicName id="99FE64BA067DAC5CB6DD9A0E4D592F1C" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
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<paragraph pageId="2" pageNumber="45">
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Subgenus
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Ptomaphagus
<taxonomicNameLabel id="9DFADD3ACE6C3265717CD568C19EF489" sensu="stricto">s. str.</taxonomicNameLabel>
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<paragraph pageId="2" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="2" pageNumber="45">Palaearctic, north Oriental.</paragraph>
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<paragraph id="8CE21C59F56550FFBA38F012F970E409" pageId="2" pageNumber="45">Distribution.</paragraph>
<paragraph id="361445CD6D510AA3166B7B9C70A45FA4" pageId="2" pageNumber="45">Palaearctic, north Oriental.</paragraph>
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Key to species of
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Hellwig from Taiwan Island
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from Taiwan Island
</paragraph>
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6A, B7
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<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">Ptomaphagus (s. str.) yasutoshii</taxonomicName>
Nishikawa
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<td colspan="1" pageId="2" pageNumber="45" rowspan="1">2</td>
<tr id="18686950DEC2635462E39489BAB541C0" pageId="2" pageNumber="45">
<td id="9BA9ED134E64322FFD0072CA4B214F0C" colspan="1" pageId="2" pageNumber="45" rowspan="1">2</td>
</tr>
<tr pageId="2" pageNumber="45">
<td colspan="1" pageId="2" pageNumber="45" rowspan="1">
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="kuntzeni">Ptomaphagus (s. str.) kuntzeni</taxonomicName>
Sokolowski
<tr id="B3F784D17B18B098EBF98F806F32D338" pageId="2" pageNumber="45">
<td id="8683A3C8CFA45F5AAB9A9A5C63EC6A7E" colspan="1" pageId="2" pageNumber="45" rowspan="1">
<taxonomicName id="8F4E424FD9D23C5D8455D8CA539158DF" authority="Sokolowski" authorityName="Sokolowski" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="kuntzeni">Ptomaphagus (s. str.) kuntzeni Sokolowski</taxonomicName>
</td>
</tr>
<tr pageId="3" pageNumber="46">
<td colspan="1" pageId="3" pageNumber="46" rowspan="1">
<pageBreakToken pageId="3" pageNumber="46" start="start">8</pageBreakToken>
<tr id="36E953CB65C55D06FCDC4218C5FA8340" pageId="3" pageNumber="46">
<td id="2241BD3E1FE9674FF806F64F818981DF" colspan="1" pageId="3" pageNumber="46" rowspan="1">
<pageBreakToken id="8C2E7457473AC29D2F6E29FC3B418842" pageId="3" pageNumber="46" start="start">8</pageBreakToken>
G, H11G, H9
<normalizedToken originalValue="AD">A-D</normalizedToken>
<normalizedToken id="7189DB78EA1960CBBF561EE3BD1EBE4C" originalValue="AD">A-D</normalizedToken>
10B, C
</td>
<td colspan="1" pageId="3" pageNumber="46" rowspan="1">3</td>
<td id="7F0B59B3920A115C039B6ED9FC054155" colspan="1" pageId="3" pageNumber="46" rowspan="1">3</td>
</tr>
<tr pageId="3" pageNumber="46">
<td colspan="1" pageId="3" pageNumber="46" rowspan="1">8A9H10B</td>
<td colspan="1" pageId="3" pageNumber="46" rowspan="1">
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi">Ptomaphagus (s. str.) chenggongi</taxonomicName>
sp. n.
<tr id="278A8E839BFEB59E028B86E1CF5A0EF6" pageId="3" pageNumber="46">
<td id="394FF06B5B930340FB27C854C2D84459" colspan="1" pageId="3" pageNumber="46" rowspan="1">8A9H10B</td>
<td id="F97687E437BC88D80F12C0B8E6400BBB" colspan="1" pageId="3" pageNumber="46" rowspan="1">
<taxonomicName id="2FF8921D0EB2A86023EDDFE1BD35504D" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi" status="sp. nov.">Ptomaphagus (s. str.) chenggongi</taxonomicName>
<taxonomicNameLabel id="03529C25A31776DF727EEFBF214D8BC5" rank="species">sp. n.</taxonomicNameLabel>
</td>
</tr>
<tr pageId="3" pageNumber="46">
<td colspan="1" pageId="3" pageNumber="46" rowspan="1">11A9I10C</td>
<td colspan="1" pageId="3" pageNumber="46" rowspan="1">
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae">Ptomaphagus (s. str.) tingtingae</taxonomicName>
sp. n.
<tr id="76EDDD51B46642D1050E1050A00492FD" pageId="3" pageNumber="46">
<td id="11B4525040FE4CCA586D875FC10B065A" colspan="1" pageId="3" pageNumber="46" rowspan="1">11A9I10C</td>
<td id="7DB68DA02D8ED88ED5690C28B88AB060" colspan="1" pageId="3" pageNumber="46" rowspan="1">
<taxonomicName id="13177BF06ACA31457E71F6C42185419C" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae" status="sp. nov.">Ptomaphagus (s. str.) tingtingae</taxonomicName>
<taxonomicNameLabel id="606BAFE3E00E4B4830A4304612A29DDB" rank="species">sp. n.</taxonomicNameLabel>
</td>
</tr>
</table>
</paragraph>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="F48627FA3890ABB69F5C3C8A554A2E01" pageId="3" pageNumber="46">
<paragraph id="CE90895E8E5D24300098D187B5731838" pageId="3" pageNumber="46">
Figure 1. Habitus of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="kuntzeni">Ptomaphagus (s. str.) kuntzeni</taxonomicName>
Sokolowski, 1957 from Taiwan (dorsal view). A ♂ (Fenchihu) B ♀ (Meifeng). Scale bar 1 mm.
<taxonomicName id="E7C4314BF9FB4119E7D44BCE5B1710D4" authority="Sokolowski, 1957" authorityName="Sokolowski" authorityYear="1957" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="kuntzeni">Ptomaphagus (s. str.) kuntzeni Sokolowski, 1957</taxonomicName>
from Taiwan (dorsal view). A ♂ (Fenchihu) B ♀ (Meifeng). Scale bar 1 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="4CDEC93982209FA474FBB9B4C436B895" pageId="3" pageNumber="46">
<paragraph id="0F6E2E87BE7EAC85AF1F39A2901B14D5" pageId="3" pageNumber="46">
Figure 2. Habitus of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">Ptomaphagus (s. str.) yasutoshii</taxonomicName>
Nishikawa, 1993 (dorsal view). A ♂ (holotype) B ♀ (allotype) C ♂ (holotype of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="smetanai">Ptomaphagus (s. str.) smetanai</taxonomicName>
Perreau, 1996, syn. n.) D ♀ (Yushan). Scale bar 1 mm.
<taxonomicName id="987FE07AA838A8CBFFCDDA62A5EC9493" authority="Nishikawa, 1993" authorityName="Nishikawa" authorityYear="1993" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">Ptomaphagus (s. str.) yasutoshii Nishikawa, 1993</taxonomicName>
(dorsal view). A ♂ (holotype) B ♀ (allotype) C ♂ (holotype of
<taxonomicName id="EE3384EFEE97D0241907B5D0E19674CA" authority="Perreau, 1996" authorityName="Perreau" authorityYear="1996" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="smetanai" status="syn. nov.">Ptomaphagus (s. str.) smetanai Perreau, 1996</taxonomicName>
,
<taxonomicNameLabel id="8F08EF136CC3EA1EB715F82653D65E47">syn. n.</taxonomicNameLabel>
) D ♀ (Yushan). Scale bar 1 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="580DB02C9F9AB7E9CA506547C0D12310" pageId="3" pageNumber="46">
<paragraph id="C0ED5510D92C69225D3FA41B16B34F90" pageId="3" pageNumber="46">
Figure 3. Habitus of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="5AD9A6344230D022791C128363CCAF44" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.) spp. (dorsal view). A, B
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi">Ptomaphagus (s. str.) chenggongi</taxonomicName>
sp. n.
<normalizedToken originalValue="CE">C-E</normalizedToken>
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae">Ptomaphagus (s. str.) tingtingae</taxonomicName>
sp. n. A, C ♂ (holotypes) B, D ♀ (paratypes) E ♂ (line-art drawing). Scale bar 1 mm.
<taxonomicName id="58E53E28DD09959322467ABFCDCBFF30" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi" status="sp. nov.">Ptomaphagus (s. str.) chenggongi</taxonomicName>
<taxonomicNameLabel id="766569B89D5F685FD7303503239AB0C2" rank="species">sp. n.</taxonomicNameLabel>
<normalizedToken id="0562CDB327408CEF4D328FEF20126E37" originalValue="CE">C-E</normalizedToken>
<taxonomicName id="74263F24A036487EF753CA1E47F4D4CD" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae" status="sp. nov.">Ptomaphagus (s. str.) tingtingae</taxonomicName>
<taxonomicNameLabel id="12CA65929FE2386A2E21A17E27D3F146" rank="species">sp. n.</taxonomicNameLabel>
A, C ♂ (holotypes) B, D ♀ (paratypes) E ♂ (line-art drawing). Scale bar 1 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="7CD9A4DB4628E45FC94805B246F37156" pageId="3" pageNumber="46">
<paragraph id="95EB7ACB55CCDEFCCD20C623C71FE3EF" pageId="3" pageNumber="46">
Figure 4. Aedeagal apices of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="kuntzeni">Ptomaphagus (s. str.) kuntzeni</taxonomicName>
Sokolowski, 1957 (ventral view).
<normalizedToken originalValue="AD">A-D</normalizedToken>
<taxonomicName id="D2212FAAB899233DB3C7FD8C42D526B8" authority="Sokolowski, 1957" authorityName="Sokolowski" authorityYear="1957" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus kuntzeni" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="kuntzeni">Ptomaphagus (s. str.) kuntzeni Sokolowski, 1957</taxonomicName>
(ventral view).
<normalizedToken id="E2A9D407833DBAF6C879557D93655B6C" originalValue="AD">A-D</normalizedToken>
Fenchihu, Taiwan Island E, F
<normalizedToken originalValue="Amami-Ôshima">Amami-Oshima</normalizedToken>
<normalizedToken id="52FF3132FDFDA93C1E1A71C8FCC03AC9" originalValue="Amami-Ôshima">Amami-Oshima</normalizedToken>
Island, Japan B, D, F right apicoventral piece of median lobe. Scale bars 0.1 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="0FD195DA32FF7FC52266DC80CBEEF690" pageId="3" pageNumber="46">
<paragraph id="70E9DB16D99EC567556673B98DB44025" pageId="3" pageNumber="46">
Figure 5.
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">Ptomaphagus (s. str.) yasutoshii</taxonomicName>
Nishikawa, 1993 (♂ holotype; ♀ allotype). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scale bars 0.1 mm.
<taxonomicName id="0A2CB26B7688AD1B39C0E325BB6CC6DF" authority="Nishikawa, 1993" authorityName="Nishikawa" authorityYear="1993" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">Ptomaphagus (s. str.) yasutoshii Nishikawa, 1993</taxonomicName>
(♂ holotype; ♀ allotype). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scale bars 0.1 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="2063E137BB4A7F8492B44C78EAC38546" pageId="3" pageNumber="46">
<paragraph id="7E215297D262F247CA3CA65B1F1A5DFA" pageId="3" pageNumber="46">
Figure 6. Aedeagi of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">Ptomaphagus (s. str.) yasutoshii</taxonomicName>
Nishikawa, 1993. A, C, F, H holotype B, D, E, G, I holotype of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="smetanai">Ptomaphagus (s. str.) smetanai</taxonomicName>
Perreau, 1996, syn. n. A, B aedeagi (dorsal view) C, D aedeagi (lateral view) E paramere apex (lateral view) F, G aedeagal apices (ventral view) H, I right apicoventral piece of median lobe (ventral view). Scale bars 0.1 mm.
<taxonomicName id="1BF002865A29B032B69C2B54542735A9" authority="Nishikawa, 1993" authorityName="Nishikawa" authorityYear="1993" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">Ptomaphagus (s. str.) yasutoshii Nishikawa, 1993</taxonomicName>
. A, C, F, H holotype B, D, E, G, I holotype of
<taxonomicName id="B83404BB04AB1E474F93E22DF76249C4" authority="Perreau, 1996" authorityName="Perreau" authorityYear="1996" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus smetanai" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="smetanai" status="syn. nov.">Ptomaphagus (s. str.) smetanai Perreau, 1996</taxonomicName>
,
<taxonomicNameLabel id="7354B8F0DC1BFE827B9BB62BB3E955ED">syn. n.</taxonomicNameLabel>
A, B aedeagi (dorsal view) C, D aedeagi (lateral view) E paramere apex (lateral view) F, G aedeagal apices (ventral view) H, I right apicoventral piece of median lobe (ventral view). Scale bars 0.1 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="4C2EB997D7B9B55722CE7680C78A3AD2" pageId="3" pageNumber="46">
<paragraph id="D91E8D99AC8EAFCB98C2A2F1F07EF4AC" pageId="3" pageNumber="46">
Figure 7.
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">Ptomaphagus (s. str.) yasutoshii</taxonomicName>
Nishikawa, 1993 (♀ Yushan). A spermatheca and genital segment (ventral view) B spermatheca (lateral view) C spermatheca (dorsal view) D ventrite VIII ♀ (ventral view). Scale bars 0.1 mm.
<taxonomicName id="BADEF10F97E47BF2F306929F5E24CD0E" authority="Nishikawa, 1993" authorityName="Nishikawa" authorityYear="1993" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus yasutoshii" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="yasutoshii">Ptomaphagus (s. str.) yasutoshii Nishikawa, 1993</taxonomicName>
(♀ Yushan). A spermatheca and genital segment (ventral view) B spermatheca (lateral view) C spermatheca (dorsal view) D ventrite VIII ♀ (ventral view). Scale bars 0.1 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="6D56796FF2D3C5395A3FABFC4E3DE758" pageId="3" pageNumber="46">
<paragraph id="7BD003B3C8B77B13E804006C0A64CAB4" pageId="3" pageNumber="46">
Figure 8.
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi">Ptomaphagus (s. str.) chenggongi</taxonomicName>
sp. n. (paratypes). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scale bars 0.1 mm.
<taxonomicName id="47B4A67A15572EA6A2D3EDCAAA9C7911" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi" status="sp. nov.">Ptomaphagus (s. str.) chenggongi</taxonomicName>
<taxonomicNameLabel id="1EF882F767F4F68DA370C1BD3365AE12" rank="species">sp. n.</taxonomicNameLabel>
(paratypes). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scale bars 0.1 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="9B80789283152B00F029C24D5C5A6BED" pageId="3" pageNumber="46">
<paragraph id="AFC63D64BEA2E858F3ED7B48D2BD08CC" pageId="3" pageNumber="46">
Figure 9. Aedeagi of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="D23877D50E5880D9B43C79E7B16D96A5" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
(s. str.) spp. A, C, E, G, H
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi">Ptomaphagus (s. str.) chenggongi</taxonomicName>
sp. n. (paratype) B, D, F, I, J
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae">Ptomaphagus (s. str.) tingtingae</taxonomicName>
sp. n. (paratype) A, B aedeagi (dorsal view) C, D aedeagi (lateral view) E, F paramere apices (lateral view) G, J aedeagal apices (ventral view) H, I right apicoventral piece of median lobe (ventral view). Scale bars 0.1 mm.
<taxonomicName id="04A477E502437A6D2B7A05C1FA172E7F" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi" status="sp. nov.">Ptomaphagus (s. str.) chenggongi</taxonomicName>
<taxonomicNameLabel id="456E887DE4A9E3AF9F3DF321C0AD8875" rank="species">sp. n.</taxonomicNameLabel>
(paratype) B, D, F, I, J
<taxonomicName id="C19D2CDB7AF6C17A9EE02E141E9AA743" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae" status="sp. nov.">Ptomaphagus (s. str.) tingtingae</taxonomicName>
<taxonomicNameLabel id="2F4534C89EF7435A168F02DFFC1168E6" rank="species">sp. n.</taxonomicNameLabel>
(paratype) A, B aedeagi (dorsal view) C, D aedeagi (lateral view) E, F paramere apices (lateral view) G, J aedeagal apices (ventral view) H, I right apicoventral piece of median lobe (ventral view). Scale bars 0.1 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="572FDAD30DF3B2A901C4BE752CE2F974" pageId="3" pageNumber="46">
<paragraph id="D840001FCA3E3009D870D897B121764F" pageId="3" pageNumber="46">
Figure 10. A, B
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi">Ptomaphagus (s. str.) chenggongi</taxonomicName>
sp. n. (paratype) C, D
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae">Ptomaphagus (s. str.) tingtingae</taxonomicName>
sp. n. (paratype) A, D ventrites VIII (ventral view) B, C spermathecae and genital segments (ventral view). Scale bars 0.1 mm.
<taxonomicName id="B033EDEC3A517C97823FDB5045DE2D9A" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus chenggongi" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="chenggongi" status="sp. nov.">Ptomaphagus (s. str.) chenggongi</taxonomicName>
<taxonomicNameLabel id="2C5B82A546886A4416B93B75FE79E374" rank="species">sp. n.</taxonomicNameLabel>
(paratype) C, D
<taxonomicName id="6235AB3C8AD7246BCD749DED9740A2BC" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae" status="sp. nov.">Ptomaphagus (s. str.) tingtingae</taxonomicName>
<taxonomicNameLabel id="0021E38927F0B235FF578653FE592F2E" rank="species">sp. n.</taxonomicNameLabel>
(paratype) A, D ventrites VIII (ventral view) B, C spermathecae and genital segments (ventral view). Scale bars 0.1 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="10A830E7A630B30E7DE17BF38CF4F8ED" pageId="3" pageNumber="46">
<paragraph id="6B387035C0E9E8A49FDEF33752C75E0F" pageId="3" pageNumber="46">
Figure 11.
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae">Ptomaphagus (s. str.) tingtingae</taxonomicName>
sp. n. (paratypes). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (dorsal view) F protibia and profemur ♀ (dorsal view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scale bars 0.1 mm.
<taxonomicName id="42CE5BF51BC3CCFD0AACF96BF0B91F6C" class="Insecta" family="Leiodidae" genus="Ptomaphagus" isSensuStricto="true" kingdom="Animalia" lsidName="Ptomaphagus tingtingae" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="tingtingae" status="sp. nov.">Ptomaphagus (s. str.) tingtingae</taxonomicName>
<taxonomicNameLabel id="DE5F5B9E9C0416AD3D62927010A39316" rank="species">sp. n.</taxonomicNameLabel>
(paratypes). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (dorsal view) F protibia and profemur ♀ (dorsal view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scale bars 0.1 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="46">
<paragraph pageId="3" pageNumber="46">
<caption id="8D093D6DFDEEF6055CEAFD32B1048D69" pageId="3" pageNumber="46">
<paragraph id="09194E54BD6837D478FD0999BBA33396" pageId="3" pageNumber="46">
Figure 12. Distribution map of
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
<taxonomicName id="354BA016C39FB95C14DC866F7D7C1DAE" class="Insecta" family="Leiodidae" genus="Ptomaphagus" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
species from Taiwan Island.
</paragraph>
</caption>