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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.225.99356" ID-Pensoft-Pub="1314-2003-225-115" ID-Pensoft-UUID="AB75E2E9609C5A7583F34B48EA61B976" ModsDocID="1314-2003-225-115" checkinTime="1682624511249" checkinUser="pensoft" docAuthor="Khamar, Hamid, Oualidi, Jalal El, Touhami, Amina Ouazzani &amp; Civeyrel, Laure" docDate="2023" docId="EDC48B9092E4524F82367F8F85EBC897" docLanguage="en" docName="PhytoKeys 225: 115-152" docOrigin="PhytoKeys 225" docPubDate="2023-04-27" docSource="http://dx.doi.org/10.3897/phytokeys.225.99356" docTitle="Verbascum erosum Cav., Elench. Pl. Hort. Matrit.: 38. 1803." docType="treatment" docVersion="1" id="AB75E2E9609C5A7583F34B48EA61B976" lastPageNumber="115" masterDocId="AB75E2E9609C5A7583F34B48EA61B976" masterDocTitle="Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa" masterLastPageNumber="152" masterPageNumber="115" pageNumber="115" updateTime="1682624511249" updateUser="pensoft">
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<mods:title>Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Khamar, Hamid</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6008-3264</mods:nameIdentifier>
<mods:affiliation>Department of Botany and Plant Ecology, Scientific Institute, University Mohammed V in Rabat, B. P. 703, Rabat 10106, Morocco</mods:affiliation>
<mods:nameIdentifier type="email">khamarhamid90@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oualidi, Jalal El</mods:namePart>
<mods:affiliation>Department of Botany and Plant Ecology, Scientific Institute, University Mohammed V in Rabat, B. P. 703, Rabat 10106, Morocco</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Touhami, Amina Ouazzani</mods:namePart>
<mods:affiliation>Plant, Animal Productions and Agro-industry Laboratory, Department of Biology, Faculty of Sciences, Ibn Tofail University, B. P. 133 14000, Kenitra, Morocco</mods:affiliation>
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<mods:namePart>Civeyrel, Laure</mods:namePart>
<mods:affiliation>Laboratoire Evolution &amp; Diversite Biologique (EDB UMR 5174), Universite de Toulouse, CNRS, IRD, UPS. 118 route de Narbonne, Bat 4 R 1, 31062 Toulouse cedex 9, France</mods:affiliation>
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<mods:date>2023</mods:date>
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<mods:number>2023-04-27</mods:number>
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<mods:number>225</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.225.99356</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-225-115</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:EDC48B9092E4524F82367F8F85EBC897" httpUri="http://treatment.plazi.org/id/EDC48B9092E4524F82367F8F85EBC897" lastPageNumber="115" pageId="0" pageNumber="115">
<subSubSection pageId="0" pageNumber="115" type="nomenclature">
<paragraph pageId="0" pageNumber="115">
<taxonomicName LSID="EDC48B90-92E4-524F-8236-7F8F85EBC897" authority="Cav., Elench. Pl. Hort. Matrit.: 38. 1803." authorityName="Cav., Elench. Pl. Hort. Matrit.: 38. 1803." class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Verbascum erosum" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="erosum">Verbascum erosum Cav., Elench. Pl. Hort. Matrit.: 38. 1803.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="reference_group">
<paragraph pageId="0" pageNumber="115">
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia sinuata" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="sinuata">Celsia sinuata</taxonomicName>
=
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia sinuata" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="sinuata">Celsia sinuata</taxonomicName>
Cav., in Anal. Cienc. Nat. 3: 68. 1801.
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia laciniata" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="laciniata">Celsia laciniata</taxonomicName>
=
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia laciniata" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="laciniata">Celsia laciniata</taxonomicName>
Poir., Lam., in Encycl., Suppl. 2: 147. 1811.
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia barnadesii var. baetica" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="variety" species="barnadesii" variety="baetica">Celsia barnadesii baetica</taxonomicName>
=
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia barnadesii subsp. var. var. baetica" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="variety" species="barnadesii" subSpecies="var." variety="baetica">
Celsia barnadesii (Vahl) G.Don
<normalizedToken originalValue="fìl">fil</normalizedToken>
. var. baetica
</taxonomicName>
Willk., in Ill. Fl. Hispan. 2(14): 55. 1888.
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia jeriaensis" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="jeriaensis">Celsia jeriaensis</taxonomicName>
=
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia jeriaensis" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="jeriaensis">Celsia jeriaensis</taxonomicName>
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
Lara, in Willk. Ill. FI. Hispan. 2(14): 56. 1888, nom. nud.
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Verbascum laciniatum" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="laciniatum">Verbascum laciniatum</taxonomicName>
=
<taxonomicName class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Verbascum laciniatum" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="laciniatum">Verbascum laciniatum</taxonomicName>
(Poir.) Kuntze, in Rev. Gen. 469. 1891.
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia baetica" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="baetica">Celsia baetica</taxonomicName>
=
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia baetica" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="baetica">Celsia baetica</taxonomicName>
(Willk.) Murb., in Lunds Univ. Arsskrift, n. s. 17 (9): 3. 1921.
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia baetica" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="baetica">Celsia baetica</taxonomicName>
=
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia baetica" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="baetica">Celsia baetica</taxonomicName>
(Willk.) Font Quer, in Butll. Inst. Catalana Hist. Nat. 26: 56. 1926. comb. nov. superfl.
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia lyrata var. sinuata" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="variety" species="lyrata" variety="sinuata">Celsia lyrata sinuata</taxonomicName>
=
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia lyrata subsp. var. var. sinuata" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="variety" species="lyrata" subSpecies="var." variety="sinuata">Celsia lyrata var. sinuata</taxonomicName>
(Cav.) Maire, in Jahand. &amp; Maire, Cat. Pl. Maroc: 668. 1934.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="materials_examined">
<paragraph pageId="0" pageNumber="115">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="115">
[
<collectingCountry name="Morocco">Morocco</collectingCountry>
]:
<normalizedToken originalValue="Tánger">Tanger</normalizedToken>
vicinii, [Without date],
<emphasis italics="true" pageId="0" pageNumber="115">Broussonet, s.n</emphasis>
. (
<typeStatus>lectotype</typeStatus>
, designated by
<bibRefCitation author="Benedi, C" journalOrPublisher="Collectanea Botanica" pageId="0" pageNumber="115" pagination="101 - 112" refId="B16" refString="Benedi, C, Montserrat, JM, 1985. Taxonomic and nomenclatural notes on the genus Verbascum L. (Celsia L.) in the Iberian Peninsula and the Balearic Islands. Collectanea Botanica 16 (1): 101 - 112" title="Taxonomic and nomenclatural notes on the genus Verbascum L. (Celsia L.) in the Iberian Peninsula and the Balearic Islands." volume="16" year="1985">
<normalizedToken originalValue="Benedí">Benedi</normalizedToken>
and
<collectingCountry name="Montserrat">Montserrat</collectingCountry>
1985
</bibRefCitation>
, pg.104):
<materialsCitation accessionNumber="MA108913" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Montserrat" specimenCount="1" typeStatus="holotype">
MA [
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MA108913">MA108913</accessionNumber>
]
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="notes">
<paragraph pageId="0" pageNumber="115">Notes.</paragraph>
<paragraph pageId="0" pageNumber="115">
This species was originally described as
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia sinuata" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="sinuata">
<emphasis italics="true" pageId="0" pageNumber="115">Celsia sinuata</emphasis>
</taxonomicName>
by
<bibRefCitation author="Cavanilles, AJ" journalOrPublisher="Anales de Ciencias Naturales" pageId="0" pageNumber="115" pagination="68 - 69" refId="B24" refString="Cavanilles, AJ, 1801. De las plantas que el Ciudadano Augusto Broussonet colecto en las costas septentrionales de la Africa y en las islas Canarias. Anales de Ciencias Naturales 3: 68 - 69" title="De las plantas que el Ciudadano Augusto Broussonet colecto en las costas septentrionales de la Africa y en las islas Canarias." volume="3" year="1801">Cavanilles (1801</bibRefCitation>
: 68) based on material collected by Broussonet in Tangier, Morocco (
<bibRefCitation author="Benedi, C" journalOrPublisher="Anales del Jardin Botanico de Madrid" pageId="0" pageNumber="115" pagination="182 - 183" refId="B18" refString="Benedi, C, Montserrat, JM, 1998. Verbascum erosum Cav. (Verbascoideae - Scrophulariaceae): Endemismo betico-rifeno. Anales del Jardin Botanico de Madrid 56 (1): 182 - 183" title="Verbascum erosum Cav. (Verbascoideae - Scrophulariaceae): Endemismo betico-rifeno." volume="56" year="1998">
<normalizedToken originalValue="Benedí">Benedi</normalizedToken>
and Montserrat 1998
</bibRefCitation>
). When the genus
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Celsia</emphasis>
</taxonomicName>
was subsumed in the genus
<taxonomicName class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Verbascum" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Verbascum</emphasis>
</taxonomicName>
the epithet
<emphasis italics="true" pageId="0" pageNumber="115">sinuata</emphasis>
could not be used for this species as it already exists in the combination
<taxonomicName class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Verbascum sinuatum" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="sinuatum">
<emphasis italics="true" pageId="0" pageNumber="115">Verbascum sinuatum</emphasis>
</taxonomicName>
L. for another species. We concur with the conclusion of
<bibRefCitation author="Benedi, C" journalOrPublisher="Collectanea Botanica" pageId="0" pageNumber="115" pagination="101 - 112" refId="B16" refString="Benedi, C, Montserrat, JM, 1985. Taxonomic and nomenclatural notes on the genus Verbascum L. (Celsia L.) in the Iberian Peninsula and the Balearic Islands. Collectanea Botanica 16 (1): 101 - 112" title="Taxonomic and nomenclatural notes on the genus Verbascum L. (Celsia L.) in the Iberian Peninsula and the Balearic Islands." volume="16" year="1985">
<normalizedToken originalValue="Benedí">Benedi</normalizedToken>
and Montserrat (1985)
</bibRefCitation>
that the correct name for this plant in question is
<taxonomicName family="Scrophulariaceae" kingdom="Plantae" lsidName="V. erosum" order="Lamiales" pageId="0" pageNumber="115" rank="species" species="erosum">
<emphasis italics="true" pageId="0" pageNumber="115">V. erosum</emphasis>
</taxonomicName>
Cav.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="A176EFCED0A3B55D05B65DCFF011CDEC" ID-CoL="773LJ" authority="Spencer 1897" authorityName="Spencer" authorityYear="1897" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageNumber="40" phylum="Chordata" rank="species" species="eremiana">
<emphasis id="3FA44D77C99A5774863E000AE17C7ADD" italics="true" pageNumber="40">Perameles eremiana</emphasis>
Spencer 1897
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<treatmentCitation id="876534A8DD74AE0AC4134892B4051535" author="Spencer" authority="Spencer 1897" authorityName="Spencer" authorityYear="1897" class="Mammalia" family="Peramelidae" genus="Perameles" journal="Proc. R. Soc. Victoria, n. s." kingdom="Animalia" order="Peramelemorphia" page="9" pageNumber="40" phylum="Chordata" rank="species" species="eremiana" title="Perameles eremiana" volume="9" year="1897">
<taxonomicName id="7EDF41F230C3B45D03B58059B38966D4" ID-CoL="773LJ" authority="Spencer 1897" authorityName="Spencer" authorityYear="1897" class="Mammalia" family="Peramelidae" genus="Perameles" kingdom="Animalia" order="Peramelemorphia" pageNumber="40" phylum="Chordata" rank="species" species="eremiana">
<emphasis id="B87F9ECF19D0C03C8C6AE68BF1F62E78" italics="true" pageNumber="40">Perameles eremiana</emphasis>
Spencer 1897
</taxonomicName>
,
<bibCitation id="A96943821416B9374D5C07950D1FB7A3" author="Spencer" journal="Proc. R. Soc. Victoria, n. s." pageNumber="40" pagination="9" title="Perameles eremiana" volume="9" year="1897">
Proc.
<collectionCode id="645CE528D20A2B438C812CC7C9F82961" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
. Soc.
<collectingRegion id="63A765A16C52E768D6EEC1FD70004DE1" country="Australia" name="Victoria">Victoria</collectingRegion>
, n. s., 9: 9
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A30F542FA524969DC10AD8750013FA6E" pageNumber="40" type="materials_examined">
<paragraph id="F2DAFB1F61BD723E624AF73DB6412218" pageNumber="40">
<emphasis id="E2BCBB53FDCF59B04C2DFDC6AA5E7543" inLineHeading="true" pageNumber="40">Type Locality:</emphasis>
<materialsCitation id="9D0532F499A1708BE6519067BBDBA304" ID-GBIF-Occurrence="4867347311" country="Australia" pageNumber="40" stateProvince="Northern Territory">
<collectingCountry id="25B817B2A6D4A41547099AB87A34B47D" name="Australia">Australia</collectingCountry>
,
<collectingRegion id="2BA6F6A91E235A4D28A718D34A315840" country="Australia" name="Northern Territory">Northern Territory</collectingRegion>
, Burt Plain (N of Alice Springs).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BD7170486ADB65B2BD1C8FC49A452EBC" pageNumber="40" type="vernacular_names">
<paragraph id="35BC561E2D0C9B92FF3A8F3E0A482069" pageNumber="40">
<emphasis id="E45E7FCED788A896162298BC3E577845" inLineHeading="true" pageNumber="40">Vernacular Names:</emphasis>
<vernacularName id="BC36303A30CF82521389D5F6FB3C25C2" language="eng" pageNumber="40">Desert Bandicoot</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="A4311A87F9C8BB1439FDEE6F47176B4F" pageNumber="40" type="distribution">
<paragraph id="46513E63958FB6D4BB12C8B5EF88A96E" pageNumber="40">
<emphasis id="DD94A5134499A1D1E1DCE4B84A018503" inLineHeading="true" pageNumber="40">Distribution:</emphasis>
N
<collectingRegion id="9A414C4CC41D4B7F3832700D828584CC" country="Australia" name="South Australia">South Australia</collectingRegion>
, S
<collectingRegion id="F05D551A65ACE80780A56BB500D78DE6" country="Australia" name="Northern Territory">Northern Territory</collectingRegion>
, Great
<collectingRegion id="D1D5BCE5CD532DB541F9B659A10C3736" country="Australia" name="Victoria">Victoria</collectingRegion>
Desert (
<collectingRegion id="8EB7AFF2E451B66BAC1783E6DD759489" country="Australia" name="Western Australia">Western Australia</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="B29387AC446E4035449AB42D2F923DEC" pageNumber="40" type="conservation">
<paragraph id="5EF64A1E6742639B44DA2D615BA2D446" pageNumber="40">
<emphasis id="1FF3C208011FF051197053ED035F4BEC" inLineHeading="true" pageNumber="40">Conservation:</emphasis>
<collectingCountry id="B47BC8FE84681B8585B397D195353956" name="United States of America">U.S.</collectingCountry>
ESA – Endangered;
<collectionCode id="376A708018C2DF9352BF4DD26F9EA4BF">IUCN</collectionCode>
– Extinct.
</paragraph>
</subSubSection>
<subSubSection id="789173695A7AE1B2DA029A2CB1E0F719" pageNumber="40" type="discussion">
<paragraph id="245EC8AE325AA9F38815CD67334A3236" pageNumber="40">
<emphasis id="E3B9CEBF3C6C9F2FDDD1B6149F626444" inLineHeading="true" pageNumber="40">Discussion:</emphasis>
Possibly extinct; see
<bibRefCitation id="224AB5AB7AF75940F7F3D7C213898044" author="Ride, W. D. L." refId="ref305072" refString="Ride, W. D. L. 1970. A guide to the native mammals of Australia. Oxford University Press, Melbourne, 249 pp." year="1970">Ride (1970:200)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>