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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="ED0E607AA38B8BE223D151A1DF360802" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus juninensis Caterino &amp; Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="79" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="78" updateTime="1668155382655" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>271</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>401</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152040857" LSID="urn:lsid:zoobank.org:act:65BEDD1C-1A34-4AA3-8DC2-99D0B8B12BA9" httpUri="http://treatment.plazi.org/id/ED0E607AA38B8BE223D151A1DF360802" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
<subSubSection pageId="77" pageNumber="78" type="nomenclature">
<paragraph pageId="77" pageNumber="78">
<taxonomicName LSID="urn:lsid:zoobank.org:act:65BEDD1C-1A34-4AA3-8DC2-99D0B8B12BA9" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus juninensis" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="juninensis">Operclipygus juninensis</taxonomicName>
<taxonomicNameLabel pageId="77" pageNumber="78">sp. n.</taxonomicNameLabel>
Fig. 20FMap 7
</paragraph>
</subSubSection>
<subSubSection pageId="77" pageNumber="78" type="type locality">
<paragraph pageId="77" pageNumber="78">Type locality.</paragraph>
<paragraph pageId="77" pageNumber="78">
PERU:
<normalizedToken originalValue="Junín">Junin</normalizedToken>
: ca. 1 km N Satipo, Sector San Isidro [
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-11.241834">11°14.51'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-74.649666">74°38.98'W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78" type="type material">
<paragraph pageId="77" pageNumber="78">Type material.</paragraph>
<paragraph lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
Holotype male: &quot;PERU: Depto.
<normalizedToken originalValue="Junín">Junin</normalizedToken>
, ~1km N Satipo, Sector San Isidro.
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-11.241834">11°14.51'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-74.649666">74°38.98'W</geoCoordinate>
730m. Window trap at treefall, 11-12 April 2009. A.V.Petrov&quot; / &quot;Caterino/Tishechkin
<taxonomicName lsidName="" pageId="77" pageNumber="78" rank="tribe" tribe="Exosternini">Exosternini</taxonomicName>
Voucher EXO-01401&quot; (FMNH). Paratypes (31): BRAZIL: Mato Grosso: 1: Querencia, Fazenda
<normalizedToken originalValue="São">Sao</normalizedToken>
Luiz,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-12.646983">12°38.819'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.374866">52°22.492'W</geoCoordinate>
, 560m, vii.2008, FIT, R.Andrade (CEMT); 2:
<normalizedToken originalValue="Cotriguaçu">Cotriguacu</normalizedToken>
, Fazenda
<normalizedToken originalValue="São">Sao</normalizedToken>
Nicolau, Mata Norte,
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-9.819167">9°49.15'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-58.26">58°15.6'W</geoCoordinate>
, 8-14.xii.2010, FIT, F.Z. Vaz-de-Mello (CEMT).
<normalizedToken originalValue="Pará">Para</normalizedToken>
: 1:Jacareacanga, xii.1968, M. Alvarenga (UFPR). PERU: Cusco: 2: Consuelo, Manu rd. km 165, 9.x.1982, 10.x.1982, FIT, L. Watrous &amp; G. Mazurek (FMNH);
<normalizedToken originalValue="Junín">Junin</normalizedToken>
: 1: ~1km N Satipo Sector San Isidro,
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-11.241834">11°14.51'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-74.649666">74°38.98'W</geoCoordinate>
, 730m, 11-12.iv.2009, Window trap, at treefall, A.V. Petrov (AKTC); 2: 11km NE Puerto Ocopa, Los Olivos,
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-11.05">11°3.00'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-74.25867">74°15.52'W</geoCoordinate>
, 1200m, 30-31.iii.2009, FIT, A.K. Tishechkin (AKTC) DNA Extract MSC-2142; 1: 26-28.iii.2009, Window trap, A.V. Petrov (MUSM); 1: 8km NNE Puerto Ocopa, Rio Perene, Cananeden,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-11.1">11°06'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-73.833336">73°50'W</geoCoordinate>
, 1100m, 6.ii.2008, A.V. Petrov (AKTC); 1: 7.5km NE Puerto Ocopa Cananeden,
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-11.081667">11°4.9'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-74.26833">74°16.1'W</geoCoordinate>
, 1180m, 11.i.2007, A.V. Petrov (MUSM);
<pageBreakToken pageId="78" pageNumber="79" start="start">1</pageBreakToken>
: 25km SW Satipo, Kuviriaki, 1100m, 23.i.2007, A.V. Petrov (AKTC); Loreto: 2: Iquitos - Nauta rd., km 58, Rio Itaya,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-4.2623">4°15.738'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-73.46754">73°28.052'W</geoCoordinate>
, 120m, 5-9.v.2009, Window trap, next to entrance of
<taxonomicName class="Insecta" family="Formicidae" genus="Eciton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eciton burchelli" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="burchelli">Eciton burchelli</taxonomicName>
statary bivouac in hollow treee, A.V. Petrov (AKTC); 4: 68km SW Iquitos to Nauta, Rio Itaya,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-4.1833334">4°11'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-73.433334">73°26'W</geoCoordinate>
, 110m, 18-19.i.2008, A.V. Petrov (AKTC, MUSM), 2: 1-3.iii.2008, A.V. Petrov (AKTC), 1: 26-30.ii.2008, A.V. Petrov (AKTC); 2: km 63, rd. Iquitos - Nauta, Rio Itaya,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-4.2534165">4°15.205'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-73.43457">73°26.074'W</geoCoordinate>
, 140m, 8-17.i.2010, A.V. Petrov (AKTC); Madre de Dios: 2: Manu National Park, Manu,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-12.5624">12.5624°S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-70.093">70.0930°W</geoCoordinate>
, 288m, 2-11.i.2007, pitfall, bamboo forest, J. Jacobs (CASC), 1: 20-29.vii.2006, pitfall, bamboo forest, J. Jacobs (CASC); 1: Manu National Park, Manu,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-12.5421">12.5421°S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-70.1435">70.1435°W</geoCoordinate>
, 292m, 20-29.vii.2006, pitfall, bamboo forest, J. Jacobs (CASC); 1: Los Amigos Field Sta.,
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-12.5421">12.5421°S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-70.1435">70.1435°W</geoCoordinate>
, 292m, 31.vii-9.viii.2006, pitfall, terra firma forest, J. Jacobs (CASC); 1: Manu National Park, Pantiacolla Lodge, 5.5km NW El Mirador Trail, Alto Madre de Dios River,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.652778">12°39'10&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-71.257774">71°15'28&quot;W</geoCoordinate>
, 500m, 23-26.x.2000, FIT, R. Brooks (SEMC); 1: Manu National Park, Pakitza Bio. Stn., Reserved Zone, Castanal Trail,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-11.944722">11°56'41&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-71.28333">71°17'0&quot;W</geoCoordinate>
, 317m, 15-16.x.2000, FIT, R. Brooks (SEMC).
</paragraph>
</subSubSection>
<subSubSection pageId="78" pageNumber="79" type="description">
<paragraph pageId="78" pageNumber="79">Diagnostic description.</paragraph>
<paragraph pageId="78" pageNumber="79">
This species is extremely similar to
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus sejunctus" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sejunctus">Operclipygus sejunctus</taxonomicName>
, differing only in a few characters as follows: length: 1.44-1.75 mm, width: 1.28-1.34 mm; body uniformly rufo-piceous; frons depressed, with coarse punctures intermingled with fine ground punctation; supraorbital stria interrupted; pronotal disk with slightly broader band of lateral punctures; elytra with sutural stria frequently longer, present on apical two-thirds to three-fourths; coarser elytral punctures densest at apex, but with few coarse punctures extending up the apical thirds of interstriae (as they do in
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus pecki" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="pecki">Operclipygus pecki</taxonomicName>
); sterna quite variable, but generally similar to
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus pecki" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="pecki">Operclipygus pecki</taxonomicName>
, the mesoventral disk with few coarse punctures at sides; metaventral disk with strong ground punctation, with coarser punctures gradually becoming larger and denser toward metacoxae; secondary lateral metaventral stria longer, extending from mesometaventral suture about halfway to metacoxa; pygidia also as in
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus pecki" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="pecki">Operclipygus pecki</taxonomicName>
, with very dense punctation. Male genitalia (Fig. 20F): as in
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus sejunctus" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sejunctus">Operclipygus sejunctus</taxonomicName>
except S9 shorter, more markedly expanded to base, with basal margin varied from truncate to broadly rounded, lacking apical emargination, lateral flanges wide and strongly recurved at inner corners; tegmen with sides more or less evenly rounded from base to apex, apex subacute, medioventral process short, wide, slightly sinuate, with strong apical point located just basad tegmen midpoint, projecting beneath; median lobe about one-fourth tegmen length.
</paragraph>
</subSubSection>
<subSubSection pageId="78" pageNumber="79" type="remarks">
<paragraph pageId="78" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="78" pageNumber="79">
This species is very similar externally to
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus pecki" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="pecki">Operclipygus pecki</taxonomicName>
and
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus depressus" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="depressus">Operclipygus depressus</taxonomicName>
. It can be separated from the former by the lack of any hint of bicoloration (with the pronotum and elytra identical in coloration). From the latter it can only be confidently separated by the shape of the aedeagus (Fig. 20F).
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus juninensis" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="juninensis">Operclipygus juninensis</taxonomicName>
also generally has the outer lateral metaventral stria longer reaching the midpoint of the metaventrite, whereas it is generally very short in
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus pecki" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="pecki">Operclipygus pecki</taxonomicName>
and
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus depressus" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="depressus">Operclipygus depressus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="78" pageNumber="79" type="etymology">
<paragraph pageId="78" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="78" pageNumber="79">
We name this species for
<normalizedToken originalValue="Junín">Junin</normalizedToken>
department, Peru, where the type specimen was collected.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="12FE2B9CC580AC6875A4924267D03B20">On a taxonomic feature that has been overestimated in classification practice: an integrative taxonomic revision of Stephoblemmus Saussure, 1877 based on morphology and molecular phylogeny (Orthoptera: Grylloidea; Gryllidae; Gryllinae)</mods:title>
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<mods:namePart id="63F58ADFC0BFB76940CA3F35A02D9C14">Zheng, Yan-Na</mods:namePart>
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<mods:affiliation id="F9C1301879157D1B51574B8BAF657643">College of Life Sciences, Shaanxi Normal University, Xi′an, China, 710119</mods:affiliation>
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<mods:namePart id="BB8471061F9D7EFAC9ED7EEF934A5621">Gu, Jun-Jie</mods:namePart>
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<mods:affiliation id="01E81712888D8D0643C12A500FDC2159">College of Agronomy, Sichuan Agricultural University, Chengdu, China, 611130</mods:affiliation>
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<mods:namePart id="BD7F350A5B9EE96E338296A8C20D372B">He, Zhu-Qing</mods:namePart>
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<mods:affiliation id="D2213DC29C2C1865EFE5BCCC7E7BE6F1">School of Life Sciences, East China Normal University, Shanghai, China, 200241</mods:affiliation>
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<mods:namePart id="D68231769D5AEA8E69280B0AC7A165FD">Huang, Huateng</mods:namePart>
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<mods:title id="25F173774D24E5DE11DB97F0C7200667">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:part id="A67635D880388E041CA2971A4B5A0A7A">
<mods:date id="BA190915448DE67230A9BD608F0D73DF">2023</mods:date>
<mods:detail id="E380BFCAF2AF228E886BC0E9071F1AFB" type="pubDate">
<mods:number id="4E6BCF05C4D10D810A460FED5CD4955A">2023-09-13</mods:number>
</mods:detail>
<mods:detail id="0982812AE8A66C714E6D9172D57D5275" type="volume">
<mods:number id="A915B26E5241D58F830975339FE89FC6">81</mods:number>
</mods:detail>
<mods:extent id="3B964A1F082CEF6FA8E30707BA6DFDBF" unit="page">
<mods:start id="15B71829A0523A7FD42CF253D7C60196">761</mods:start>
<mods:end id="01757A8F4C7D3C8E3642A25F8418D9FC">779</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="C87D32C0247A24ABB68C74A0CED96F27">
<mods:url id="E7B41F05531FC3E3C9C34242E98500D2">http://dx.doi.org/10.3897/asp.81.e104772</mods:url>
</mods:location>
<mods:classification id="76C920756AD0059908455E8F7E3C170D">journal article</mods:classification>
<mods:identifier id="F41E521D6D37E87DBD4C87FBFCC49508" type="DOI">http://dx.doi.org/10.3897/asp.81.e104772</mods:identifier>
<mods:identifier id="B273DA53015357604A1F71EBBF87BF3C" type="Pensoft-Pub">1864-8312-81-761</mods:identifier>
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<treatment id="ED0EDFE0C60C5042A2BAE2B6139CECBA" LSID="urn:lsid:plazi:treatment:ED0EDFE0C60C5042A2BAE2B6139CECBA" httpUri="http://treatment.plazi.org/id/ED0EDFE0C60C5042A2BAE2B6139CECBA" lastPageNumber="761" pageId="0" pageNumber="761">
<subSubSection id="D52BBC5CB723F3B3588BB1C38CF36465" pageId="0" pageNumber="761" type="nomenclature">
<paragraph id="B79829C84C271AC9FA53B7BA0E942453" pageId="0" pageNumber="761">
Genus
<taxonomicName id="02996357E469A070EBD8B6A469B5D52A" ID-CoL="8NZK2" LSID="ED0EDFE0-C60C-5042-A2BA-E2B6139CECBA" authority="Saussure, 1877" authorityName="Saussure" authorityYear="1877" class="Insecta" family="Gryllidae" genus="Stephoblemmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephoblemmus" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="genus">Stephoblemmus Saussure, 1877</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E6DA7D2DBEBB971CB03AE99E4948DA86" pageId="0" pageNumber="761" type="reference_group">
<paragraph id="CBD6EFEE22FC093D007660FDEABAD6EA" pageId="0" pageNumber="761">
<taxonomicName id="F9DD70EB78889BA2709254FD3E921996" class="Insecta" family="Gryllidae" genus="Mitius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitius" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="genus">Mitius</taxonomicName>
Gorochov, 1985: 95,
<emphasis id="250FE8F4CB88AF4F7CF7DF4B7668F7B9" bold="true" pageId="0" pageNumber="761">syn. n.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="929DC5DDE2EE04D489EAA82ECD6F2EA3" pageId="0" pageNumber="761" type="type species">
<paragraph id="BDCA9325B071F3F2E825CBD1268F7867" pageId="0" pageNumber="761">Type species.</paragraph>
<paragraph id="929E03F09027842E3F00542B7414EE68" pageId="0" pageNumber="761">
<taxonomicName id="A7ACD6407A6E5AF623DADE94D0BC18FA" authorityName="Saussure" authorityYear="1877" class="Insecta" family="Gryllidae" genus="Stephoblemmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephoblemmus humbertiellus" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="species" species="humbertiellus">
<emphasis id="76D2A6A1591C16F8017CC714D460FA69" italics="true" pageId="0" pageNumber="761">Stephoblemmus humbertiellus</emphasis>
</taxonomicName>
Saussure, 1877.
</paragraph>
</subSubSection>
<subSubSection id="8D153978BEC9C57E38504A34C370226F" pageId="0" pageNumber="761" type="chinese name">
<paragraph id="8425ABC235BE6C25869B382E080BC7E6" pageId="0" pageNumber="761">Chinese name.</paragraph>
<paragraph id="EA93DCC13B0F533A90A73713504F5AC9" pageId="0" pageNumber="761">
<normalizedToken id="651011041C755D5571B09DBD07B77C83" originalValue="素蟋属">素蟋属</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="C0D42B5DE8A196644B399AA5777FD9B5" pageId="0" pageNumber="761" type="emended diagnosis">
<paragraph id="D7E5EA815D7DE6ECAAFE86E7D22E5CF4" pageId="0" pageNumber="761">Emended diagnosis.</paragraph>
<paragraph id="53057F39423A35624D64946CF5FDD3CE" pageId="0" pageNumber="761">
<emphasis id="597F9D71591830BB5742D2C4499959E9" bold="true" pageId="0" pageNumber="761">Male</emphasis>
: Vertex broad and flattened, slightly inclined. Occiput slightly convex. Frons inclined dorsally and ventrally, and nearly two times wider than antennal scape. Antennal scape shield-like. Median ocellus ovoid, lateral ocelli transversely ovoid. Two lateral ocelli located on the upper edge of the antennal scape and distributed in an inverted triangle with the median ocellus. Eyes ovoid. Pronotum broad and flattened. The median groove of the pronotal disc distinct. Posterior margin straight, the middle of the anterior margin concave, and the posterior and anterior margins almost equal in width. In the dorsal field of tegmina, bearing three chord veins, with the apical one connecting to the proximal part of the mirror by two veins; diagonal vein proximally bifurcated, both branches connecting to CuA vein; two oblique veins; mirror shield-like; dividing vein absent. Outer tympanum shaped as an elongate oval, inner tympanum ovoid or absent. Hind tibiae armed with dorsal spurs which almost equal in length. Epiphallus armed with a finger-like median lobe (Fig.
<figureCitation id="82BA00D1B63E093B2E249C9E24F5E643" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pattern of male genitalia of Stephoblemmus. A dorsal view, B lateral view, C ventral view." figureDoi="10.3897/asp.81.e104772.figure4" httpUri="https://binary.pensoft.net/fig/905090" pageId="0" pageNumber="761">4</figureCitation>
).
<emphasis id="C6386D86AD9F9DC0A477C3918EF91163" bold="true" pageId="0" pageNumber="761">Female</emphasis>
: Similar to male in the head, pronotum, and feet features. Tegmina
<normalizedToken id="81005EF6A532F9F428277E20BDBB3F53" originalValue="don’t">don't</normalizedToken>
reach the end of the abdomen, with longitudinal veins interspersed with false veins. Ovipositor smooth, arrow-like.
</paragraph>
<caption id="AABC2853FD605684E51A50BD3EE1B192" doi="10.3897/asp.81.e104772.figure4" httpUri="https://binary.pensoft.net/fig/905090" pageId="0" pageNumber="761" start="Figure 4" startId="F4">
<paragraph id="A028BAE8DD77BA404BFAB3985326F4C1" pageId="0" pageNumber="761">
<emphasis id="236842A2436ED47E4D257B7FD35E0167" bold="true" pageId="0" pageNumber="761">Figure 4.</emphasis>
Pattern of male genitalia of
<taxonomicName id="545746C01956DFF60158C532B9C73CA4" authorityName="Saussure" authorityYear="1877" class="Insecta" family="Gryllidae" genus="Stephoblemmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephoblemmus" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="genus">
<emphasis id="B2B52F1941E315C0E0835ADE9E764F58" italics="true" pageId="0" pageNumber="761">Stephoblemmus</emphasis>
</taxonomicName>
.
<emphasis id="EFAFA601CB554EAD5FE47299CCC90315" bold="true" pageId="0" pageNumber="761">A</emphasis>
dorsal view,
<emphasis id="FD7ECB943F2A67CB382B60E5970DDD96" bold="true" pageId="0" pageNumber="761">B</emphasis>
lateral view,
<emphasis id="7E501DA4B5213970A75BCF59B6855268" bold="true" pageId="0" pageNumber="761">C</emphasis>
ventral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="18D9D76FAC9F5F7640FBF598E1DFFE81" pageId="0" pageNumber="761" type="included species">
<paragraph id="53BEE11272EC2465E39FC0EDC3E37E44" pageId="0" pageNumber="761">Included species.</paragraph>
<paragraph id="D2FB2F1CC958CEACCB1D814A215BA9F7" pageId="0" pageNumber="761">
<taxonomicName id="217B4432BDD747A60DFED78A803491B7" baseAuthorityName="Saussure" baseAuthorityYear="1877" class="Insecta" family="Gryllidae" genus="Mitius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitius blennus" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="species" species="blennus">
<emphasis id="68D3ABA98BA744615F6C6F26BE292E37" italics="true" pageId="0" pageNumber="761">Mitius blennus</emphasis>
</taxonomicName>
(Saussure, 1877)
<taxonomicName id="6B87BE3FC580ADD58D04602866B57B3B" baseAuthorityName="Chopard" baseAuthorityYear="1937" class="Insecta" family="Gryllidae" genus="Mitius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitius castaneus" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="7DA8755916539756FC2972128853D6AC" italics="true" pageId="0" pageNumber="761">Mitius castaneus</emphasis>
</taxonomicName>
(Chapord, 1937),
<taxonomicName id="B6165D4CC016273E46E48AD99578CBF2" authorityName="Zheng &amp; Gu &amp; He &amp; Huang &amp; Ma" authorityYear="2023" baseAuthorityName="Zheng &amp; Gu &amp; He &amp; Huang &amp; Ma" baseAuthorityYear="2023" class="Insecta" family="Gryllidae" genus="Mitius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitius enatus" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="species" species="enatus">
<emphasis id="75AC87F6854780C8E80CC8584F399EFF" italics="true" pageId="0" pageNumber="761">Mitius enatus</emphasis>
</taxonomicName>
(Gorochov, 1994),
<taxonomicName id="6EA943C2C77119C70C6C0CE9E7D8378A" baseAuthorityName="Chopard" baseAuthorityYear="1928" class="Insecta" family="Gryllidae" genus="Mitius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitius flavipes" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="55D932BAEC76972DA91D5056745F663A" italics="true" pageId="0" pageNumber="761">Mitius flavipes</emphasis>
</taxonomicName>
(Chopard, 1928),
<taxonomicName id="7276673E3E250D534253E42A5B6EE8E7" authorityName="Zheng &amp; Gu &amp; He &amp; Huang &amp; Ma" authorityYear="2023" baseAuthorityName="Zheng &amp; Gu &amp; He &amp; Huang &amp; Ma" baseAuthorityYear="2023" class="Insecta" family="Gryllidae" genus="Mitius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitius minor" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B952891905BEBA0A8195FAEE0A51576D" italics="true" pageId="0" pageNumber="761">Mitius minor</emphasis>
</taxonomicName>
(Shiraki, 1911),
<taxonomicName id="3CACC210AFD90E136A21DE945C0D779B" authorityName="Zheng &amp; Gu &amp; He &amp; Huang &amp; Ma" authorityYear="2023" baseAuthorityName="Zheng &amp; Gu &amp; He &amp; Huang &amp; Ma" baseAuthorityYear="2023" class="Insecta" family="Gryllidae" genus="Mitius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitius minutulus" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="species" species="minutulus">
<emphasis id="E86C2E0608030408AD89C8EAC35EAADC" italics="true" pageId="0" pageNumber="761">Mitius minutulus</emphasis>
</taxonomicName>
Yang &amp; Yang, 1995,
<taxonomicName id="01C36DC1E6A09FD668139AEB94B97E4F" baseAuthorityName="Otte &amp; Cowper" baseAuthorityYear="2007" class="Insecta" family="Gryllidae" genus="Mitius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitius vaturu" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="species" species="vaturu">
<emphasis id="3CB6E83113482DFFBD98215DC551E833" italics="true" pageId="0" pageNumber="761">Mitius vaturu</emphasis>
</taxonomicName>
(Otte &amp; Cowper, 2007), and
<taxonomicName id="37C22694B556C25F060F825BC5A55240" authorityName="Saussure" authorityYear="1877" class="Insecta" family="Gryllidae" genus="Stephoblemmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephoblemmus humbertiellus" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="species" species="humbertiellus">
<emphasis id="52087F4CB1C8E3118E8F0106E22A29B8" italics="true" pageId="0" pageNumber="761">Stephoblemmus humbertiellus</emphasis>
</taxonomicName>
Saussure, 1877.
</paragraph>
</subSubSection>
<subSubSection id="8488624CEE1185B65E1982AEB5DCE07C" pageId="0" pageNumber="761" type="remarks">
<paragraph id="0B8AB806F6FA86B433853B221F285BA6" pageId="0" pageNumber="761">Remarks.</paragraph>
<paragraph id="6E4E58FCC4E15A930B86D38E9A43E959" pageId="0" pageNumber="761">
<taxonomicName id="3BB696A97E1FE5CE7C6440922902998F" authorityName="Saussure" authorityYear="1877" class="Insecta" family="Gryllidae" genus="Stephoblemmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephoblemmus" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="genus">
<emphasis id="8FC4051058E9C65A70D3D0BCB45ECD8F" italics="true" pageId="0" pageNumber="761">Stephoblemmus</emphasis>
</taxonomicName>
was established by
<bibRefCitation id="3087356247F7A182C7A5151F374D0943" author="Saussure, H" journalOrPublisher="Systematic Biology" pageId="0" pageNumber="761" refId="B46" refString="Saussure, H, 1877. Melanges orthopterologiques V. fascicule Gryllides. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve 25(1): 169-504 [161-352]." title="Melanges orthopterologiques V. fascicule Gryllides. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve 25 (1): 169 - 504 [161 - 352]." year="1877">Saussure (1877)</bibRefCitation>
and its dominant feature is the flat frons of males (Fig. S5C-D, G-H).
<taxonomicName id="827670A4837567B51ACF0EBCD89E4430" class="Insecta" family="Gryllidae" genus="Mitius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitius" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="genus">
<emphasis id="86760473C1163D709164C0CAA28D7401" italics="true" pageId="0" pageNumber="761">Mitius</emphasis>
</taxonomicName>
was established by Gorochov (1985) and species of the genus possess two types of frons (truncated and rounded) (Fig. S5A-B, E-F, I-P). The phylogenetic relationship reveals that
<taxonomicName id="A4E795DD18B94A3B0AB5C87DA9837E4F" authorityName="Saussure" authorityYear="1877" class="Insecta" family="Gryllidae" genus="Stephoblemmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephoblemmus" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="genus">
<emphasis id="D71B531B0DAE36B93F08ECC0617A23E2" italics="true" pageId="0" pageNumber="761">Stephoblemmus</emphasis>
</taxonomicName>
(
<taxonomicName id="DE87F8BF05F680C1BEC2005D6ACD4C8B" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. humbertiellus" order="Orthoptera" pageId="0" pageNumber="761" rank="species" species="humbertiellus">
<emphasis id="3C51BA2333B4677390AD02EA9784C541" italics="true" pageId="0" pageNumber="761">S. humbertiellus</emphasis>
</taxonomicName>
) and
<taxonomicName id="9BA5FFAE006592A8609F8F094EDE649F" class="Insecta" family="Gryllidae" genus="Mitius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitius" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="genus">
<emphasis id="72449F62F2E0D8BF5A876897C2168262" italics="true" pageId="0" pageNumber="761">Mitius</emphasis>
</taxonomicName>
(
<taxonomicName id="B455C6F54D32542AFF5C34B86770B8B6" family="Gryllidae" kingdom="Animalia" lsidName="M. minor" order="Orthoptera" pageId="0" pageNumber="761" rank="species" species="minor">
<emphasis id="98BDE1DEFD50E7A336348C9F9450E4C4" italics="true" pageId="0" pageNumber="761">M. minor</emphasis>
</taxonomicName>
(=
<taxonomicName id="EF973668D8D34A0BE6235B25E756BBD4" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. minor" order="Orthoptera" pageId="0" pageNumber="761" rank="species" species="minor">
<emphasis id="EFDFED6568FD8587F3BB5C13B6042AA6" italics="true" pageId="0" pageNumber="761">S. minor</emphasis>
</taxonomicName>
<emphasis id="774108783FF82BBD893A94AE837EADF3" bold="true" pageId="0" pageNumber="761">comb. n.</emphasis>
)) are sister group (Fig.
<figureCitation id="E8FD7610FB832D3D06C36B311A89F971" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Time-calibrated phylogeny inferred using BEAST. Posterior probability (PP) values are indicated by node color. Node numbers next to nodes represent the divergent times. Purple bars indicate 95 % credibility intervals of estimated node ages. The left in Roman numbers represents fossil calibration points. The primary species of study for this article are shaded red, including species of Loxoblemmus, Mitius, and Stephoblemmus. The blue letter is the type species of Stephoblemmus. The geological timescale is shown at the bottom (Ma = million years ago)." figureDoi="10.3897/asp.81.e104772.figure2" httpUri="https://binary.pensoft.net/fig/905088" pageId="0" pageNumber="761">2</figureCitation>
) and their male genitalia are also similar. Accordingly,
<taxonomicName id="F29A202A7C5AA21786C353C454611D79" class="Insecta" family="Gryllidae" genus="Mitius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mitius" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="genus">
<emphasis id="7273B5F87FF07F3DD643C3F01405EC82" italics="true" pageId="0" pageNumber="761">Mitius</emphasis>
</taxonomicName>
should be a junior synonym of
<taxonomicName id="00FDA463B106A8E953C99B7BD2A3A164" authorityName="Saussure" authorityYear="1877" class="Insecta" family="Gryllidae" genus="Stephoblemmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stephoblemmus" order="Orthoptera" pageId="0" pageNumber="761" phylum="Arthropoda" rank="genus">
<emphasis id="1AA5458D91F590701205CA4ED5BF59CF" italics="true" pageId="0" pageNumber="761">Stephoblemmus</emphasis>
</taxonomicName>
and we combine them.
</paragraph>
</subSubSection>
</treatment>
</document>