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<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e83035" ID-Pensoft-Pub="1864-8312-80-439" ID-Pensoft-UUID="3F4E21B524465471831D09738C2304A6" ID-ZooBank="62FFB94CEBF441639F22881435EFC37C" ModsDocID="1864-8312-80-439" checkinTime="1663021027595" checkinUser="pensoft" docAuthor="Santos, Daubian, Santos, Rodrigo dos Reis &amp; Ribeiro, Guilherme Cunha" docDate="2022" docId="AF9A65F292B7551DA2CC86D95B2D4D8A" docLanguage="en" docName="ArthropodSystPhyl 80: 439-494" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-09-12" docSource="http://dx.doi.org/10.3897/asp.80.e83035" docTitle="Amphineurus (Rhamphoneurus) nonnullus Alexander 1967" docType="treatment" docVersion="1" id="3F4E21B524465471831D09738C2304A6" lastPageNumber="439" masterDocId="3F4E21B524465471831D09738C2304A6" masterDocTitle="Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)" masterLastPageNumber="494" masterPageNumber="439" pageNumber="439" updateTime="1663021027595" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Santos, Daubian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1220-1267</mods:nameIdentifier>
<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">daubians@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Santos, Rodrigo dos Reis</mods:namePart>
<mods:affiliation>Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ribeiro, Guilherme Cunha</mods:namePart>
<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>439</mods:start>
<mods:end>494</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.80.e83035</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e83035</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-80-439</mods:identifier>
<mods:identifier type="ZooBank">62FFB94CEBF441639F22881435EFC37C</mods:identifier>
<mods:identifier type="Pensoft-UUID">3F4E21B524465471831D09738C2304A6</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:AF9A65F292B7551DA2CC86D95B2D4D8A" httpUri="http://treatment.plazi.org/id/AF9A65F292B7551DA2CC86D95B2D4D8A" lastPageNumber="439" pageId="0" pageNumber="439">
<subSubSection pageId="0" pageNumber="439" type="nomenclature">
<paragraph pageId="0" pageNumber="439">
3.3.17.
<taxonomicName LSID="AF9A65F2-92B7-551D-A2CC-86D95B2D4D8A" authority="Alexander, 1967" authorityName="Alexander" authorityYear="1967" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) nonnullus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="nonnullus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) nonnullus Alexander, 1967</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="reference_group">
<paragraph pageId="0" pageNumber="439">
<taxonomicName authorityName="Alexander" authorityYear="1967" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) nonnullus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="nonnullus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) nonnullus</taxonomicName>
Alexander, 1967: 491 (description); Alexander, 1968: fig. 17 (male terminalia).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="materials_examined">
<paragraph pageId="0" pageNumber="439">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="439">
<emphasis bold="true" pageId="0" pageNumber="439">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
<specimenCount></specimenCount>
, Chile,
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is., Aucar [
<geoCoordinate degrees="42" direction="south" minutes="9" orientation="latitude" precision="925" value="-42.15">42°9S</geoCoordinate>
<geoCoordinate degrees="73" direction="west" minutes="30" orientation="longitude" precision="925" value="-73.5">73°30W</geoCoordinate>
], 6/
<date value="1952-01-15">15-I-1952</date>
,
<normalizedToken originalValue="Peña">Pena</normalizedToken>
,
<typeStatus>Holotype</typeStatus>
9619 (USNM) [part of specimen pinned and part in slide prepared by C.P. Alexander]. -
<emphasis bold="true" pageId="0" pageNumber="439">Additional material</emphasis>
:
<materialsCitation collectingDate="1952-02-03" collectingDateMax="1954-02-08" collectingDateMin="1952-02-03" collectorName="Chiloe Is." country="CHILE" latitude="-42.033333" location="Chaiten" longLatPrecision="1152" longitude="-74.01667" specimenCount="1">
<emphasis bold="true" pageId="0" pageNumber="439">
<collectingCountry name="Chile">CHILE</collectingCountry>
.
</emphasis>
1 [sex unknown],
<collectorName>
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is.
</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:AF9A65F292B7551DA2CC86D95B2D4D8A:45F7D50E29DE3DC49D26C50228AA25E6" country="CHILE" latitude="-42.033333" longLatPrecision="1152" longitude="-74.01667" name="Chaiten">Chaiten</location>
[
<geoCoordinate degrees="42" direction="south" minutes="58" orientation="latitude" precision="925" value="-42.966667">42°58S</geoCoordinate>
<geoCoordinate degrees="72" direction="west" minutes="32" orientation="longitude" precision="925" value="-72.53333">72°32W</geoCoordinate>
], 5/
<collectingDate value="1954-02-08">8-II-1954</collectingDate>
,
<normalizedToken originalValue="Peña">Pena</normalizedToken>
(USNM)*; 1 [sex unknown],
<collectorName>
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is.
</collectorName>
, Chepu [
<geoCoordinate degrees="42" direction="south" minutes="2" orientation="latitude" precision="925" value="-42.033333">42°2S</geoCoordinate>
<geoCoordinate degrees="74" direction="west" minutes="1" orientation="longitude" precision="925" value="-74.01667">74°1W</geoCoordinate>
], 1/
<collectingDate value="1952-02-03">3-II-1952</collectingDate>
,
<normalizedToken originalValue="Peña">Pena</normalizedToken>
(USNM)
</materialsCitation>
*.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="diagnosis">
<paragraph pageId="0" pageNumber="439">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="439">This species is distinguished by R2 without a dark spot and the beginning of M1 desclerotized. Furthermore, the species has a distinctive male terminalia with a curved sheath of aedeagus with the branches of the clasper of different sizes.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="redescription">
<paragraph pageId="0" pageNumber="439">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="439">
Wing length 5.58 mm, width 1.71 mm. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Coloration</emphasis>
: General coloration brownish-yellow. Rostrum and palpus dark brown. First segment of antenna yellow, remaining segments brownish-yellow. Head brown. Eyes grayish black. Thorax brownish yellow, pleura grayish-yellow. Scutum with one brownish-black stripe. Halter light yellow. Coxae light yellow; legs yellowish-brown. Abdomen yellowish brown. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Head</emphasis>
(dorsal view Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Amphineurus (Rhamphoneurus) nonnullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure22" httpUri="https://binary.pensoft.net/fig/740575" pageId="0" pageNumber="439">22B</figureCitation>
, ventral view Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Amphineurus (Rhamphoneurus) nonnullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure22" httpUri="https://binary.pensoft.net/fig/740575" pageId="0" pageNumber="439">22A</figureCitation>
): Rostrum bifurcated with short appendices; first palpal segment shorter than terminal segment; scape thinner than pedicel, terminal flagellomere as long as penultimate. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Thorax</emphasis>
(lateral view Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Amphineurus (Rhamphoneurus) nonnullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure22" httpUri="https://binary.pensoft.net/fig/740575" pageId="0" pageNumber="439">22E</figureCitation>
, dorsal view Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Amphineurus (Rhamphoneurus) nonnullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure22" httpUri="https://binary.pensoft.net/fig/740575" pageId="0" pageNumber="439">22F</figureCitation>
). Laterotergite almost triangular, anatergite taller than katatergite. Halter with dilated knob. Wing (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Amphineurus (Rhamphoneurus) nonnullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure22" httpUri="https://binary.pensoft.net/fig/740575" pageId="0" pageNumber="439">22D</figureCitation>
) without distinctive markings; veins near fork of bM (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Amphineurus (Rhamphoneurus) nonnullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure22" httpUri="https://binary.pensoft.net/fig/740575" pageId="0" pageNumber="439">22C</figureCitation>
) faded; m-cu inserted distal to fork of bM; M1+2 shorter than basal deflection of M1; R2+3 longer than R2+3+4. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Male terminalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Amphineurus (Rhamphoneurus) nonnullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure22" httpUri="https://binary.pensoft.net/fig/740575" pageId="0" pageNumber="439">22G</figureCitation>
): Male tergite IX slightly darkened on distal margin, with wide median U-shaped notch; posterolateral lobes rounded. Dorsal branch of gonocoxite half-length of ventral branch; both branches setose. Lobe of gonostylus with rounded lobule; lobule about as long as half-length of lobe of gonostylus, narrower than stem. Clasper of gonostylus with differently shaped branches: lateral branch straight; medial branch curved, club-shaped. Mesal lobes of gonocoxite asymmetrical; both mesal lobes with sharp, 90° turn on posterior end; right mesal lobe significantly longer than left. Sheath of aedeagus curved, darkened only at distal portion.
</paragraph>
<caption doi="10.3897/asp.80.e83035.figure22" httpUri="https://binary.pensoft.net/fig/740575" pageId="0" pageNumber="439" start="Figure 22" startId="F22">
<paragraph pageId="0" pageNumber="439">
<emphasis bold="true" pageId="0" pageNumber="439">Figure 22.</emphasis>
<taxonomicName authorityName="Alexander" authorityYear="1967" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) nonnullus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="nonnullus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) nonnullus</taxonomicName>
Alexander.
<emphasis bold="true" pageId="0" pageNumber="439">A</emphasis>
head (ventral view);
<emphasis bold="true" pageId="0" pageNumber="439">B</emphasis>
head (dorsal view);
<emphasis bold="true" pageId="0" pageNumber="439">C</emphasis>
detail of wing;
<emphasis bold="true" pageId="0" pageNumber="439">D</emphasis>
wing;
<emphasis bold="true" pageId="0" pageNumber="439">E</emphasis>
thorax (lateral view);
<emphasis bold="true" pageId="0" pageNumber="439">F</emphasis>
thorax (dorsal view);
<emphasis bold="true" pageId="0" pageNumber="439">G</emphasis>
male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations:
<emphasis bold="true" pageId="0" pageNumber="439">aed sh</emphasis>
, sheath of aedeagus;
<emphasis bold="true" pageId="0" pageNumber="439">anatg</emphasis>
, anatergite;
<emphasis bold="true" pageId="0" pageNumber="439">anepm</emphasis>
, anepimeron;
<emphasis bold="true" pageId="0" pageNumber="439">anepst</emphasis>
, anepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">aprn</emphasis>
, anteropronotum;
<emphasis bold="true" pageId="0" pageNumber="439">cgonst</emphasis>
, clasper of gonostylus;
<emphasis bold="true" pageId="0" pageNumber="439">cx</emphasis>
, coxa;
<emphasis bold="true" pageId="0" pageNumber="439">goncx</emphasis>
, gonocoxite;
<emphasis bold="true" pageId="0" pageNumber="439">hlt</emphasis>
, halter;
<emphasis bold="true" pageId="0" pageNumber="439">k</emphasis>
, knob of halter;
<emphasis bold="true" pageId="0" pageNumber="439">kepm</emphasis>
, katepimeron;
<emphasis bold="true" pageId="0" pageNumber="439">kepst</emphasis>
, katepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">ktg</emphasis>
, katatergite;
<emphasis bold="true" pageId="0" pageNumber="439">l ms</emphasis>
, left mesal lobe of gonocoxite;
<emphasis bold="true" pageId="0" pageNumber="439">lgonst</emphasis>
; lobe of gonostylus;
<emphasis bold="true" pageId="0" pageNumber="439">mr</emphasis>
, meron;
<emphasis bold="true" pageId="0" pageNumber="439">mtanepst</emphasis>
, metanepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">mtepm</emphasis>
, metepimeron;
<emphasis bold="true" pageId="0" pageNumber="439">mtg</emphasis>
, mediotergite;
<emphasis bold="true" pageId="0" pageNumber="439">mtkepst</emphasis>
, metakatepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">mtn</emphasis>
, metanotum;
<emphasis bold="true" pageId="0" pageNumber="439">p</emphasis>
, posterior basalare;
<emphasis bold="true" pageId="0" pageNumber="439">patg</emphasis>
, paratergite;
<emphasis bold="true" pageId="0" pageNumber="439">pprn</emphasis>
, postpronotum;
<emphasis bold="true" pageId="0" pageNumber="439">presct</emphasis>
, prescutum;
<emphasis bold="true" pageId="0" pageNumber="439">r ms</emphasis>
, right mesal lobe of gonocoxite;
<emphasis bold="true" pageId="0" pageNumber="439">sct</emphasis>
, scutum;
<emphasis bold="true" pageId="0" pageNumber="439">sctl</emphasis>
, scutellum;
<emphasis bold="true" pageId="0" pageNumber="439">t9</emphasis>
, male tergite IX.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="remarks">
<paragraph pageId="0" pageNumber="439">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="439">
This species resembles
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) apiculatus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="apiculatus" subGenus="R.">A. (R.) apiculatus</taxonomicName>
Alexander but differs mainly in the fork of bM, shape of male tergite IX, clasper of gonostylus and mesal lobe.
</paragraph>
</subSubSection>
</treatment>
</document>