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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1024.60393" ID-GBIF-Dataset="16f599ee-42e4-4042-920d-8c9c4623ea0d" ID-PMC="PMC7985133" ID-Pensoft-Pub="1313-2970-1024-31" ID-Pensoft-UUID="713DF2410D3B562C867366B14011E11E" ID-PubMed="33776521" ID-ZooBank="52DD663A1C914E86A8E3A68C33F1A9EF" ModsDocID="1313-2970-1024-31" checkinTime="1615893241554" checkinUser="pensoft" docAuthor="Savage, Jade &amp; Sorokina, Vera S." docDate="2021" docId="AF631217181A557E824CD12BEDF83754" docLanguage="en" docName="ZooKeys 1024: 31-89" docOrigin="ZooKeys 1024" docPubDate="2021-03-15" docSource="http://dx.doi.org/10.3897/zookeys.1024.60393" docTitle="Drymeia ponti Savage &amp; Sorokina 2021, sp. nov." docType="treatment" docUuid="DCDED5B3-721C-442C-A16C-590AD57F5D4D" docUuidSource="ZooBank" docVersion="5" id="713DF2410D3B562C867366B14011E11E" lastPageNumber="31" masterDocId="713DF2410D3B562C867366B14011E11E" masterDocTitle="Review of the North American fauna of Drymeia Meigen (Diptera, Muscidae) and evaluation of DNA barcodes for species-level identification in the genus" masterLastPageNumber="89" masterPageNumber="31" pageNumber="31" updateTime="1668149910883" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the North American fauna of Drymeia Meigen (Diptera, Muscidae) and evaluation of DNA barcodes for species-level identification in the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Savage, Jade</mods:namePart>
<mods:affiliation>Bishop's University, Sherbrooke, Quebec, Canada</mods:affiliation>
<mods:nameIdentifier type="email">jsavage@ubishops.ca</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sorokina, Vera S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3679-9005</mods:nameIdentifier>
<mods:affiliation>Institute of Systematics and Ecology of Animals, Siberian Branch of the Russian Academy of Sciences, Novosibirsk, 630091, Russia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1024</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>31</mods:start>
<mods:end>89</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1024.60393</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1024.60393</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1024-31</mods:identifier>
<mods:identifier type="ZooBank">52DD663A1C914E86A8E3A68C33F1A9EF</mods:identifier>
<mods:identifier type="Pensoft-UUID">713DF2410D3B562C867366B14011E11E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="180583876" LSID="urn:lsid:zoobank.org:act:DCDED5B3-721C-442C-A16C-590AD57F5D4D" httpUri="http://treatment.plazi.org/id/AF631217181A557E824CD12BEDF83754" lastPageNumber="31" pageId="0" pageNumber="31">
<subSubSection pageId="0" pageNumber="31" type="nomenclature">
<paragraph pageId="0" pageNumber="31">
<taxonomicName LSID="http://zoobank.org/DCDED5B3-721C-442C-A16C-590AD57F5D4D" authority="Savage &amp; Sorokina, 2021" authorityName="Savage &amp; Sorokina" authorityYear="2021" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia ponti" order="Diptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="ponti" status="sp. nov.">Drymeia ponti</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="31">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male frons, frontal view A D. ponti B D. aterrima C D. neoborealis D D. vockerothi. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1024.60393.figure1" httpUri="https://binary.pensoft.net/fig/520342" pageId="0" pageNumber="31">Figs 1A</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male hind tibia, dorsal view A D. ponti B D. aterrima C D. groenlandica D D. setibasis E D. similis F D. vockerothi G D. quadrisetosa H D. glacialis I D. hucketti J D. woodorum. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1024.60393.figure2" httpUri="https://binary.pensoft.net/fig/520343" pageId="0" pageNumber="31">, 2A</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female fore tarsomeres 3 - 5 A D. ponti B D. aldrichi C D. similis D D. profrontalis E D. minor. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1024.60393.figure10" httpUri="https://binary.pensoft.net/fig/520351" pageId="0" pageNumber="31">, 10A</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Female frons A D. similis B D. latifrons C D. profrontalis D D. ponti E D. hucketti. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1024.60393.figure11" httpUri="https://binary.pensoft.net/fig/520352" pageId="0" pageNumber="31">, 11D</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Drymeia ponti sp. nov. A male habitus, paratype B male head, frontal view C male mid femur, anterodorsal view D male hind tibia, anterior view. Scale bars: 1 mm (A, B); 0.5 mm (C, D)." figureDoi="10.3897/zookeys.1024.60393.figure13" httpUri="https://binary.pensoft.net/fig/520354" pageId="0" pageNumber="31">, 13</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Drymeia ponti sp. nov., male terminalia A lateral view B posterior view C sternite V." figureDoi="10.3897/zookeys.1024.60393.figure14" httpUri="https://binary.pensoft.net/fig/520355" pageId="0" pageNumber="31">, 14</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Drymeia ponti sp. nov. A female habitus, paratype B female head, frontal view C female tarsomeres 3 - 5. Scale bars: 1 mm (A, B); 0.5 mm (C)." figureDoi="10.3897/zookeys.1024.60393.figure15" httpUri="https://binary.pensoft.net/fig/520356" pageId="0" pageNumber="31">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="type material">
<paragraph pageId="0" pageNumber="31">
<typeStatus>Type</typeStatus>
material.
</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">
<typeStatus>Holotype</typeStatus>
</emphasis>
male labelled &quot;W. Side Cortes Pass/ 1100'
<collectingCountry name="Mexico">Mexico</collectingCountry>
,
<collectingCountry name="Mexico">Mexico</collectingCountry>
/
<date value="1954-08-13">13-VIII-1954</date>
/ J.G. Chillcott&quot;; &quot;
<typeStatus>HOLOTYPE</typeStatus>
/
<taxonomicName authorityName="Savage &amp; Sorokina" authorityYear="2021" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia ponti" order="Diptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="ponti">Drymeia ponti</taxonomicName>
<specimenCount>♂</specimenCount>
/ Savage &amp; Sorokina [red]&quot; (
<collectionCode country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
).
<materialsCitation ID-GBIF-Occurrence="3063620405" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" location="Paratypes" specimenCount="2" typeStatus="holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">
<typeStatus>
<location LSID="urn:lsid:plazi:treatment:AF631217181A557E824CD12BEDF83754:4F1DFB5647656CCBE74B8AF8EBE9A76A" name="Paratypes">Paratypes</location>
</typeStatus>
</emphasis>
: all with &quot;
<typeStatus>PARATYPE</typeStatus>
/
<taxonomicName authorityName="Savage &amp; Sorokina" authorityYear="2021" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia ponti" order="Diptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="ponti">Drymeia ponti</taxonomicName>
/ Savage &amp; Sorokina [yellow]&quot;
</materialsCitation>
:
<specimenCount type="male">2 males</specimenCount>
and
<specimenCount type="female">15 females</specimenCount>
, same as
<typeStatus>holotype</typeStatus>
.
<materialsCitation ID-GBIF-Occurrence="3063620337" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="BUIC" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
<specimenCount type="male">1 male</specimenCount>
and
<specimenCount type="female">2 females</specimenCount>
, same as
<typeStatus>holotype</typeStatus>
except (
<collectionCode>BUIC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063620472" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="CNC" collectorName="D. Kritsch" location="Rio Frio" municipality="Fed. Dist. Of Mex." specimenCount="6" specimenCount-female="5" specimenCount-male="1" typeStatus="holotype">
<specimenCount type="male">1 male</specimenCount>
and
<specimenCount type="female">5 females</specimenCount>
labelled &quot;
<location LSID="urn:lsid:plazi:treatment:AF631217181A557E824CD12BEDF83754:735A522AED4A85C2E0685ACC4ABB6F6B" municipality="Fed. Dist. Of Mex." name="Rio Frio">Rio Frio</location>
/
<collectingMunicipality>Fed. Dist. Of Mex.</collectingMunicipality>
/ I.IX.69/
<collectorName>D. Kritsch</collectorName>
&quot; (
<collectionCode country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063620380" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="BUIC" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount type="male">1 male</specimenCount>
, same as previous except (
<collectionCode>BUIC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063620476" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="CNC" collectorName="D. Kritsch" country="Mexico" location="Rio Frio" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount type="male">1 male</specimenCount>
labelled &quot;
<location LSID="urn:lsid:plazi:treatment:AF631217181A557E824CD12BEDF83754:01E5E054FD5F37373F5122151C6AA428" country="Mexico" name="Rio Frio">Rio Frio</location>
,
<collectingCountry name="Mexico">MEX</collectingCountry>
./ MWX.,27-VIII-69/
<collectorName>D. Kritsch</collectorName>
BL 10,000'&quot; (
<collectionCode country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1024.60393.figure11" httpUri="https://binary.pensoft.net/fig/520352" pageId="0" pageNumber="31" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figure 11.</emphasis>
Female frons
<emphasis bold="true" pageId="0" pageNumber="31">A</emphasis>
<taxonomicName lsidName="D. similis" pageId="0" pageNumber="31" rank="species" species="similis">
<emphasis italics="true" pageId="0" pageNumber="31">D. similis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="31">B</emphasis>
<taxonomicName lsidName="D. latifrons" pageId="0" pageNumber="31" rank="species" species="latifrons">
<emphasis italics="true" pageId="0" pageNumber="31">D. latifrons</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="31">C</emphasis>
<taxonomicName lsidName="D. profrontalis" pageId="0" pageNumber="31" rank="species" species="profrontalis">
<emphasis italics="true" pageId="0" pageNumber="31">D. profrontalis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="31">D</emphasis>
<taxonomicName lsidName="D. ponti" pageId="0" pageNumber="31" rank="species" species="ponti">
<emphasis italics="true" pageId="0" pageNumber="31">D. ponti</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="31">E</emphasis>
<taxonomicName lsidName="D. hucketti" pageId="0" pageNumber="31" rank="species" species="hucketti">
<emphasis italics="true" pageId="0" pageNumber="31">D. hucketti</emphasis>
</taxonomicName>
. Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="etymology">
<paragraph pageId="0" pageNumber="31">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="31">
The species name is a patronym in honour of Adrian C. Pont (
<collectingCountry name="United Kingdom">UK</collectingCountry>
), an exceptional dipterist and mentor to both co-authors.
</paragraph>
<caption doi="10.3897/zookeys.1024.60393.figure12" httpUri="https://binary.pensoft.net/fig/520353" pageId="0" pageNumber="31" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figure 12.</emphasis>
Female arista
<emphasis bold="true" pageId="0" pageNumber="31">A</emphasis>
<taxonomicName lsidName="D. hucketti" pageId="0" pageNumber="31" rank="species" species="hucketti">
<emphasis italics="true" pageId="0" pageNumber="31">D. hucketti</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="31">B</emphasis>
<taxonomicName lsidName="D. aterrima" pageId="0" pageNumber="31" rank="species" species="aterrima">
<emphasis italics="true" pageId="0" pageNumber="31">D. aterrima</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="31">C</emphasis>
<taxonomicName lsidName="D. minor" pageId="0" pageNumber="31" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="31">D. minor</emphasis>
</taxonomicName>
. Scale bars:
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.5" unit="mm" value="0.25">0.25 mm</quantity>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="diagnosis">
<paragraph pageId="0" pageNumber="31">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="31">
Small dark species with strong prealar, 2+3
<emphasis italics="true" pageId="0" pageNumber="31">dc</emphasis>
and strong costal spine. This species is similar to
<taxonomicName baseAuthorityName="Wulp" baseAuthorityYear="1896" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia aterrima" order="Diptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="aterrima">
<emphasis italics="true" pageId="0" pageNumber="31">Drymeia aterrima</emphasis>
</taxonomicName>
(Wulp, 1896), especially in the dark male calypter, but can be distinguished from it in the male by a broad frontal vitta (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Drymeia ponti sp. nov. A male habitus, paratype B male head, frontal view C male mid femur, anterodorsal view D male hind tibia, anterior view. Scale bars: 1 mm (A, B); 0.5 mm (C, D)." figureDoi="10.3897/zookeys.1024.60393.figure13" httpUri="https://binary.pensoft.net/fig/520354" pageId="0" pageNumber="31">13B</figureCitation>
) and a distinct ventral apical process on
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
3 (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Drymeia ponti sp. nov. A male habitus, paratype B male head, frontal view C male mid femur, anterodorsal view D male hind tibia, anterior view. Scale bars: 1 mm (A, B); 0.5 mm (C, D)." figureDoi="10.3897/zookeys.1024.60393.figure13" httpUri="https://binary.pensoft.net/fig/520354" pageId="0" pageNumber="31">13D</figureCitation>
), and in the female by the presence of a flattened fore tarsomere 5 (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Drymeia ponti sp. nov. A female habitus, paratype B female head, frontal view C female tarsomeres 3 - 5. Scale bars: 1 mm (A, B); 0.5 mm (C)." figureDoi="10.3897/zookeys.1024.60393.figure15" httpUri="https://binary.pensoft.net/fig/520356" pageId="0" pageNumber="31">15C</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1024.60393.figure13" httpUri="https://binary.pensoft.net/fig/520354" pageId="0" pageNumber="31" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figure 13.</emphasis>
<taxonomicName authorityName="Savage &amp; Sorokina" authorityYear="2021" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia ponti" order="Diptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="ponti">
<emphasis italics="true" pageId="0" pageNumber="31">Drymeia ponti</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="31">A</emphasis>
male habitus,
<typeStatus>paratype</typeStatus>
<emphasis bold="true" pageId="0" pageNumber="31">B</emphasis>
male head, frontal view
<emphasis bold="true" pageId="0" pageNumber="31">C</emphasis>
male mid femur, anterodorsal view
<emphasis bold="true" pageId="0" pageNumber="31">D</emphasis>
male hind tibia, anterior view. Scale bars:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
(
<emphasis bold="true" pageId="0" pageNumber="31">A, B</emphasis>
);
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
(
<emphasis bold="true" pageId="0" pageNumber="31">C, D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="description">
<paragraph pageId="0" pageNumber="31">Description.</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Male.</emphasis>
Body length:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.7" metricValueMin="4.1" unit="mm" value="4.9" valueMax="5.7" valueMin="4.1">4.1-5.7 mm</quantity>
; wing length:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.2" metricValueMin="3.8" unit="mm" value="4.5" valueMax="5.2" valueMin="3.8">3.8-5.2 mm</quantity>
.
</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">
<emphasis italics="true" pageId="0" pageNumber="31">Head</emphasis>
:
</emphasis>
Ground colour black; eye bare; fronto-orbital plate and parafacial dark brown pruinose; face black, gena and lower occiput dark brown pruinose; frons at narrowest point approximately 1.5
<normalizedToken originalValue="×">x</normalizedToken>
width of ocellar triangle with black frontal vitta exposed (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Drymeia ponti sp. nov. A male habitus, paratype B male head, frontal view C male mid femur, anterodorsal view D male hind tibia, anterior view. Scale bars: 1 mm (A, B); 0.5 mm (C, D)." figureDoi="10.3897/zookeys.1024.60393.figure13" httpUri="https://binary.pensoft.net/fig/520354" pageId="0" pageNumber="31">13B</figureCitation>
); parafacial in lateral view equal to or slightly wider than width of first flagellomere along most of its length (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Drymeia ponti sp. nov. A male habitus, paratype B male head, frontal view C male mid femur, anterodorsal view D male hind tibia, anterior view. Scale bars: 1 mm (A, B); 0.5 mm (C, D)." figureDoi="10.3897/zookeys.1024.60393.figure13" httpUri="https://binary.pensoft.net/fig/520354" pageId="0" pageNumber="31">13A</figureCitation>
); lower margin of the face projecting slightly beyond lower level of frons in lateral view; gena at narrowest point as high as length of first flagellomere, densely setose and without a group of upcurved setae on anterior part of genal dilation; 9-12 frontal setae (including interstitials) reaching to anterior ocellus; antenna black; first flagellomere 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide; arista swollen near base and pubescent, with longest hair as long as basal diameter of arista; palpus black; proboscis long and narrow with prementum approximately 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as palpus, undusted and glossy; labella small.
</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">
<emphasis italics="true" pageId="0" pageNumber="31">Thorax</emphasis>
:
</emphasis>
Ground colour black; scutum, postpronotum, notopleuron, postalar callus and scutellum with dense brown dusting; pleuron dark brown, slightly shiny with brown dusting; katepisternum with small anterior undusted glossy patch, meron entirely dusted; anepimeron and katepimeron bare; notopleuron setulose;
<emphasis italics="true" pageId="0" pageNumber="31">acr</emphasis>
0+1; 2+3
<emphasis italics="true" pageId="0" pageNumber="31">dc</emphasis>
; prealar long and strong, as long as second notopleural.
</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">
<emphasis italics="true" pageId="0" pageNumber="31">Legs</emphasis>
:
</emphasis>
Black;
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
1 with 2
<emphasis italics="true" pageId="0" pageNumber="31">pv</emphasis>
on apical 1/2; F2 straight, with matching rows of long strong
<emphasis italics="true" pageId="0" pageNumber="31">av</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="31">pv</emphasis>
on apical 2/3, these much longer than width of femur (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Drymeia ponti sp. nov. A male habitus, paratype B male head, frontal view C male mid femur, anterodorsal view D male hind tibia, anterior view. Scale bars: 1 mm (A, B); 0.5 mm (C, D)." figureDoi="10.3897/zookeys.1024.60393.figure13" httpUri="https://binary.pensoft.net/fig/520354" pageId="0" pageNumber="31">13A, C</figureCitation>
), a complete row of short
<emphasis italics="true" pageId="0" pageNumber="31">ad</emphasis>
, and 2 or 3 preapical
<emphasis italics="true" pageId="0" pageNumber="31">pd-p</emphasis>
;
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
2 with 1 long and 2 shorter
<emphasis italics="true" pageId="0" pageNumber="31">ad</emphasis>
on apical 1/2, 4
<emphasis italics="true" pageId="0" pageNumber="31">pd</emphasis>
and 1
<emphasis italics="true" pageId="0" pageNumber="31">pv</emphasis>
; F3 with a complete row of
<emphasis italics="true" pageId="0" pageNumber="31">ad</emphasis>
, a row of
<emphasis italics="true" pageId="0" pageNumber="31">av</emphasis>
, longer on apical 1/2, a row of
<emphasis italics="true" pageId="0" pageNumber="31">p</emphasis>
on basal 2/3 and a row of long strong
<emphasis italics="true" pageId="0" pageNumber="31">pv</emphasis>
on apical 1/3;
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
3 with 2
<emphasis italics="true" pageId="0" pageNumber="31">av</emphasis>
, 4
<emphasis italics="true" pageId="0" pageNumber="31">ad</emphasis>
, 3
<emphasis italics="true" pageId="0" pageNumber="31">pd</emphasis>
and with 4 short delicate
<emphasis italics="true" pageId="0" pageNumber="31">pv</emphasis>
on apical 1/2, ventral apical process short but distinct (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Drymeia ponti sp. nov. A male habitus, paratype B male head, frontal view C male mid femur, anterodorsal view D male hind tibia, anterior view. Scale bars: 1 mm (A, B); 0.5 mm (C, D)." figureDoi="10.3897/zookeys.1024.60393.figure13" httpUri="https://binary.pensoft.net/fig/520354" pageId="0" pageNumber="31">13D</figureCitation>
), apical
<emphasis italics="true" pageId="0" pageNumber="31">pv</emphasis>
absent or reduced, no longer than 1/2 the length of apical
<emphasis italics="true" pageId="0" pageNumber="31">av</emphasis>
when visible.
</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">
<emphasis italics="true" pageId="0" pageNumber="31">Wing</emphasis>
:
</emphasis>
Brown, darker at base with dark brown veins; basicosta and tegula black; costal spinules strong, with costal spine 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as costal spinules; calypters with membrane and edges dark brown.
</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">
<emphasis italics="true" pageId="0" pageNumber="31">Abdomen</emphasis>
:
</emphasis>
Conical; ground colour black; brown dusted, subshiny with no median vittae; sternite I bare; sternite
<collectionCode country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
as in Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Drymeia ponti sp. nov., male terminalia A lateral view B posterior view C sternite V." figureDoi="10.3897/zookeys.1024.60393.figure14" httpUri="https://binary.pensoft.net/fig/520355" pageId="0" pageNumber="31">14C</figureCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1024.60393.figure14" httpUri="https://binary.pensoft.net/fig/520355" pageId="0" pageNumber="31" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figure 14.</emphasis>
<taxonomicName authorityName="Savage &amp; Sorokina" authorityYear="2021" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia ponti" order="Diptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="ponti">
<emphasis italics="true" pageId="0" pageNumber="31">Drymeia ponti</emphasis>
</taxonomicName>
sp. nov., male terminalia
<emphasis bold="true" pageId="0" pageNumber="31">A</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="31">B</emphasis>
posterior view
<emphasis bold="true" pageId="0" pageNumber="31">C</emphasis>
sternite V.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">
<emphasis italics="true" pageId="0" pageNumber="31">Terminalia</emphasis>
:
</emphasis>
Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Drymeia ponti sp. nov., male terminalia A lateral view B posterior view C sternite V." figureDoi="10.3897/zookeys.1024.60393.figure14" httpUri="https://binary.pensoft.net/fig/520355" pageId="0" pageNumber="31">14A, B</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Female.</emphasis>
Body length:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" unit="mm" value="5.5" valueMax="6.5" valueMin="4.5">4.5-6.5 mm</quantity>
; wing length: 4.0-
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.5" unit="mm" value="5.5">5.5 mm</quantity>
. Differs from the male as follows:
</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">
<emphasis italics="true" pageId="0" pageNumber="31">Head</emphasis>
:
</emphasis>
Frontal triangle undefined; frontal vitta black; parafacial mostly dusted, with a small narrow shiny patch near base of antenna (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Drymeia ponti sp. nov. A female habitus, paratype B female head, frontal view C female tarsomeres 3 - 5. Scale bars: 1 mm (A, B); 0.5 mm (C)." figureDoi="10.3897/zookeys.1024.60393.figure15" httpUri="https://binary.pensoft.net/fig/520356" pageId="0" pageNumber="31">15A, B</figureCitation>
); frons at midpoint approximately 0.4
<normalizedToken originalValue="×">x</normalizedToken>
as wide as head and approximately 0.9
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide; fronto-orbital plate narrow, approximately as wide as distance between inner margins of posterior ocelli; three or four medioclinate frontal setae and several weaker interstitials, three orbital setae, the upper two reclinate and lateroclinate, the lower one proclinate.
</paragraph>
<caption doi="10.3897/zookeys.1024.60393.figure15" httpUri="https://binary.pensoft.net/fig/520356" pageId="0" pageNumber="31" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figure 15.</emphasis>
<taxonomicName authorityName="Savage &amp; Sorokina" authorityYear="2021" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia ponti" order="Diptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="ponti">
<emphasis italics="true" pageId="0" pageNumber="31">Drymeia ponti</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="31">A</emphasis>
female habitus,
<typeStatus>paratype</typeStatus>
<emphasis bold="true" pageId="0" pageNumber="31">B</emphasis>
female head, frontal view
<emphasis bold="true" pageId="0" pageNumber="31">C</emphasis>
female tarsomeres 3-5. Scale bars:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
(
<emphasis bold="true" pageId="0" pageNumber="31">A, B</emphasis>
);
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
(
<emphasis bold="true" pageId="0" pageNumber="31">C</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">
<emphasis italics="true" pageId="0" pageNumber="31">Thorax</emphasis>
:
</emphasis>
As in male.
</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Legs</emphasis>
(chaetotaxy described in full):
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
1 with 1 or 2
<emphasis italics="true" pageId="0" pageNumber="31">pv</emphasis>
; fore tarsomere 5 distinctively flattened (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female fore tarsomeres 3 - 5 A D. ponti B D. aldrichi C D. similis D D. profrontalis E D. minor. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1024.60393.figure10" httpUri="https://binary.pensoft.net/fig/520351" pageId="0" pageNumber="31">10A</figureCitation>
,
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Drymeia ponti sp. nov. A female habitus, paratype B female head, frontal view C female tarsomeres 3 - 5. Scale bars: 1 mm (A, B); 0.5 mm (C)." figureDoi="10.3897/zookeys.1024.60393.figure15" httpUri="https://binary.pensoft.net/fig/520356" pageId="0" pageNumber="31">15C</figureCitation>
); F2 with 1 or 2 prebasal and 2 or 3 preapical
<emphasis italics="true" pageId="0" pageNumber="31">av</emphasis>
without
<emphasis italics="true" pageId="0" pageNumber="31">pv</emphasis>
;
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
2 with 0 or 1
<emphasis italics="true" pageId="0" pageNumber="31">av</emphasis>
(most without), 2 or 3
<emphasis italics="true" pageId="0" pageNumber="31">ad</emphasis>
, 4 or 5
<emphasis italics="true" pageId="0" pageNumber="31">pd</emphasis>
and 2
<emphasis italics="true" pageId="0" pageNumber="31">pv</emphasis>
; F3 with
<emphasis italics="true" pageId="0" pageNumber="31">av</emphasis>
row complete, without
<emphasis italics="true" pageId="0" pageNumber="31">pv</emphasis>
;
<collectionCode name="Tavera, Department of Geology and Geophysics">T</collectionCode>
3 with 2 or 3
<emphasis italics="true" pageId="0" pageNumber="31">av</emphasis>
, 3 or 4
<emphasis italics="true" pageId="0" pageNumber="31">ad</emphasis>
and 3
<emphasis italics="true" pageId="0" pageNumber="31">pd</emphasis>
, apical
<emphasis italics="true" pageId="0" pageNumber="31">pv</emphasis>
usually absent but if present, then no longer than 1/2 length of apical
<emphasis italics="true" pageId="0" pageNumber="31">av</emphasis>
.
</paragraph>
<caption doi="10.3897/zookeys.1024.60393.figure16" httpUri="https://binary.pensoft.net/fig/520357" pageId="0" pageNumber="31" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figure 16.</emphasis>
<taxonomicName authorityName="Savage &amp; Sorokina" authorityYear="2021" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia hucketti" order="Diptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="hucketti">
<emphasis italics="true" pageId="0" pageNumber="31">Drymeia hucketti</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="31">A</emphasis>
<typeStatus>holotype</typeStatus>
habitus
<emphasis bold="true" pageId="0" pageNumber="31">B</emphasis>
male abdomen, dorsal view
<emphasis bold="true" pageId="0" pageNumber="31">C</emphasis>
male mid femur, anterior view
<emphasis bold="true" pageId="0" pageNumber="31">D</emphasis>
male hind tibia. Scale bars:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
(
<emphasis bold="true" pageId="0" pageNumber="31">A, B</emphasis>
);
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
(
<emphasis bold="true" pageId="0" pageNumber="31">C, D</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">
<emphasis italics="true" pageId="0" pageNumber="31">Wing</emphasis>
:
</emphasis>
Light brown, darker near base; with membrane and edges dark yellow.
</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">
<emphasis italics="true" pageId="0" pageNumber="31">Abdomen</emphasis>
:
</emphasis>
as in male.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="distribution">
<paragraph pageId="0" pageNumber="31">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="31">
Neotropical:
<collectingCountry name="Mexico">Mexico</collectingCountry>
(
<collectingCountry name="Mexico">Mexico</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="dna barcode">
<paragraph pageId="0" pageNumber="31">DNA Barcode.</paragraph>
<paragraph pageId="0" pageNumber="31">None available.</paragraph>
</subSubSection>
</treatment>
</document>

135
data/AF/63/4E/AF634E30201B5FA89D59A15E71291C04.xml Normal file
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@ -0,0 +1,135 @@
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:number>2022-08-25</mods:number>
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<taxonomicName LSID="AF634E30-201B-5FA8-9D59-A15E71291C04" authority="Fischer, 1898" authorityName="Fischer" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina billeti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="billeti">Pupina billeti Fischer, 1898</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">Figs 21S</figureCitation>
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. A Pupina bensoni sp. nov., paratype CUMZ 12046 / 1 from Khao Wong Cave, Uthai Thani B Pupina hungerfordiana, specimen NHMUK 91.3.14.686 - 7 from Hsaddan Koo C Pupina billeti, holotype MNHN-IM- 2000 - 35841 D Pupina verneaui, syntype MNHN-IM- 2000 - 35843 from Ha-Giang, Tonkin E, F Pupina peguensis E syntype of Pupina blanfordi NHMUK 1888.12.4.100 from Pegu and F specimen NHMUK ex. Cuming coll. from Lao Mountains, Camboja. Photo: P. Maestrati, MNHN (C, D)." figureDoi="10.3897/zookeys.1119.85400.figure26" httpUri="https://binary.pensoft.net/fig/735387" pageId="0" pageNumber="1">, 26C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fischer" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina billeti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="billeti">Pupina billeti</taxonomicName>
Fischer, 1898: 333, 334, pl. 18, figs 38-41. Type locality: Rochers calcaires
<normalizedToken originalValue="Déo-Ma-Phuc">Deo-Ma-Phuc</normalizedToken>
[limestone areas around Ma Phuc Pass, Tra Linh District, Cao Bang Province, Vietnam].
<bibRefCitation author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
: 431, Bac-Kan, Tonkin [Bac Kan Province, Vietnam].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fischer" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) billeti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="billeti" subGenus="Tylotoechus">Pupina (Tylotoechus) billeti</taxonomicName>
-
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
: 309.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
MNHN-IM-2000-35841 (Figs
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21S</figureCitation>
,
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. A Pupina bensoni sp. nov., paratype CUMZ 12046 / 1 from Khao Wong Cave, Uthai Thani B Pupina hungerfordiana, specimen NHMUK 91.3.14.686 - 7 from Hsaddan Koo C Pupina billeti, holotype MNHN-IM- 2000 - 35841 D Pupina verneaui, syntype MNHN-IM- 2000 - 35843 from Ha-Giang, Tonkin E, F Pupina peguensis E syntype of Pupina blanfordi NHMUK 1888.12.4.100 from Pegu and F specimen NHMUK ex. Cuming coll. from Lao Mountains, Camboja. Photo: P. Maestrati, MNHN (C, D)." figureDoi="10.3897/zookeys.1119.85400.figure26" httpUri="https://binary.pensoft.net/fig/735387" pageId="0" pageNumber="1">26C</figureCitation>
) from Deo-Ma-Phuc.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell ovate; last whorl ca. 70% of shell height. Apertural lip extremely thickened; with a furrow between inner and outer peristomes; inner peristome thickened and cord-like; parietal callus distinct. Both parietal and columellar teeth extremely thickened; parietal tooth covering posterior canal; columellar tooth next to slit-like anterior canal.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Fischer" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina billeti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="billeti">
<emphasis italics="true" pageId="0" pageNumber="1">Pupina billeti</emphasis>
</taxonomicName>
can be distinguished from all other species in the
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
</taxonomicName>
species group from mainland Southeast Asia by having the thickest parietal and columellar teeth and apertural lip, and a distinct parietal callus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Northern Vietnam (
<bibRefCitation author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
As
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. billeti" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="billeti">
<emphasis italics="true" pageId="0" pageNumber="1">P. billeti</emphasis>
</taxonomicName>
was described based on a single specimen as explicitly stated in the original description, that specimen is the holotype fixed by monotypy (
<bibRefCitation author="Hwang, C-C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" refId="B70" refString="ICZN (1999) International Code of Zoological Nomenclature, 4th Edn.International Trust for Zoological Nomenclature, London, 306 pp." year="2014">ICZN 1999</bibRefCitation>
: Art. 73.1.2).
</paragraph>
</subSubSection>
</treatment>
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<emphasis id="7CE6265D1BF3F11D0C825C0F959C5C00" italics="true">Alchemilla decumbens</emphasis>
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aggr.
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<paragraph id="6C85D8CCF34FF4362DC1289C20C8076D">
<vernacularName id="BAD404E23F662841620B140E120A7468">Niederliegender Frauenmantel</vernacularName>
</paragraph>
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<paragraph id="03FA60C2DAFB035EBADE806FAF5BBB18">
Art ISFS: 13100 Checklist: 1001530
<taxonomicName id="D03E1F2477379CBE1C6783666CD5E2EB" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
<taxonomicName id="34386F5D5D4281B31F4FD12F302FD3BE" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="genus">Alchemilla</taxonomicName>
<taxonomicName id="139E6059585BD2C770A63C93BF6894AC" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="decumbens">Alchemilla decumbens</taxonomicName>
aggr.
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:
<taxonomicName id="53E96D465B41FB11BC2726498668858B" authority="Buser" authorityName="Buser" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="decumbens">Alchemilla decumbens Buser</taxonomicName>
<taxonomicName id="D1F37E53B0434ED51EC46D32C6D012FB" authority="Huegin &amp; S. E. Froehner" authorityName="Huegin &amp; S. E. Froehner" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="pseudodecumbens">
Alchemilla pseudodecumbens
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&amp; S. E.
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<taxonomicName id="21C8F29B7A995F3D346B3CCB6809A584" authority="Buser" authorityName="Buser" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="undulata">Alchemilla undulata Buser</taxonomicName>
</paragraph>
<paragraph id="43E237EA5FC33904F8E997B67652D330">
<heading id="C0A8DF6CC9F7AF4536CBB30AE38EBF9B">Zusammenfassung</heading>
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<subSubSection id="21134CB78FD84447BF6624530CAA68B7" type="description">
<paragraph id="843B596DAD54A214EECF216ECDF29B1F">
Artbeschreibung
<emphasis id="432F601AF04660D287E79EEDF49826DD" italics="true">
(nach
<bibRefCitation id="56B3FB1C780398FE407FFA39C5FA180B" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
:
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hoch, zumindest
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unterer Teil der
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oder Blattstiele abstehend behaart.
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niederliegend
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, die
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nicht
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, diese
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im
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Drittel oder bis
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die Mitte 7-9lappig
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. Abschnitte
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<normalizedToken id="A87159980183AB207D49CE5315F15D39" originalValue="verschmälert">verschmaelert</normalizedToken>
, jederseits mit 4-6 groben
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,
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, nicht zusammenneigend.
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und Kelch kahl
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.
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als die inneren.
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<paragraph id="64918E64466E4F303C028F0B18B0A7DA">
<normalizedToken id="508603933D5F7971663A99A9C9917715" originalValue="Blütezeit">Bluetezeit</normalizedToken>
<emphasis id="64148AB68028FC50ECD65840BEEBD1F0" italics="true">
(nach
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)
</emphasis>
: 7-8
</paragraph>
<paragraph id="0D1D42F968AB3D0132B733B93726F628">
Standort und Verbreitung in der Schweiz
<emphasis id="F16870BC0E3384177F8221090E1BCA4C" italics="true">
(nach
<bibRefCitation id="E471D659E7F4DB71BBB750B421D6C3DB" ID-ISBN="978-3-258-08047-5" author="Lauber, K. &amp; Wagner, G. &amp; Gygax, A." location="Bern" publisher="Haupt Verlag" refString="Lauber, K., Wagner, G., Gygax, A. 2018. Flora Helvetica. Haupt Verlag, Bern. ISBN: 978-3-258-08047-5." title="Flora Helvetica" year="2018">Lauber &amp; al. 2018</bibRefCitation>
)
</emphasis>
: Weiden, Rasen,
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, Hochstaudenfluren / subalpin-alpin / A
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</subSubSection>
<subSubSection id="2B67070B23453F2FDC6A503D234863A6" type="multiple">
<paragraph id="210DA26EC379887050DE1ACF853BFC59">Status</paragraph>
</subSubSection>
<subSubSection id="0AE8313CFC5DD003E41FFF5F8541A388" type="conservation">
<paragraph id="56670586B4BAED97422AB74F4394456F">
<heading id="B6AE0EEE0C8FF0CE29963982AEB0A859">Status IUCN</heading>
: Nicht
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</paragraph>
</subSubSection>
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<paragraph id="2ACFFFCBF26D5148BE397C13126D5A01">
<heading id="1D6B3BD65B620752CDFC5407F7083C4F">
<normalizedToken id="BF5B01B3EAD1EA9E822B6401FCDBD4BF" originalValue="Ökologie">Oekologie</normalizedToken>
</heading>
</paragraph>
<paragraph id="FAEC9053D10911897D653F657552D2F4">
<heading id="C6A577FAE088BC8AD60D99C4946B2FBD">Lebensraum Lebensraum</heading>
nach
<bibRefCitation id="0A239542D547430B409479EDD44D95B9" author="Delarze, R. &amp; Gonseth, Y. &amp; Eggenberger, S. &amp; Vust, M." location="Bad Hersfeld, Germany" publisher="Ott Verlag" refString="Delarze, R., Gonseth, Y., Eggenberger, S., Vust, M. 2015. Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition). Ott Verlag, Bad Hersfeld, Germany, 456 pp." title="Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition)" year="2015">Delarze &amp; al. 2015</bibRefCitation>
</paragraph>
<paragraph id="AA088F57F95229F0EE631622CA26C3CE">
<table id="8CF7771B0424B27AC270E7C37DC529DC" class="table" inLine="true">
<tr id="DC0870149C568D92B10943F143BFF141">
<td id="F937BF5F7594249B9C802DFE90137C06">
<tbody id="A6A537F89EE81660C4A327A7359E26D5">
4.4.1 - Kalkreiches
<normalizedToken id="A1B9BF1E37D11BC7EB407A720CD3C30B" originalValue="Schneetälchen">Schneetaelchen</normalizedToken>
(
<emphasis id="857C8EA86CB1854BA418D4A50A87CF84" italics="true">Arabidion caerulae</emphasis>
)
</tbody>
</td>
</tr>
</table>
</paragraph>
<paragraph id="1FB8EE794D91E772E2FC1BB765C2EFFB">
<tableNote id="84AC5987CC3B59BCBF0A053289EA841A">
<emphasis id="15796610F12BE0C58E2B5E5C38BB888B" bold="true">fett</emphasis>
<emphasis id="5D7953F720E53993F059EA2C6DD41DAF" italics="true">
Dominante Art, welche das Aussehen des Lebensraumes
<normalizedToken id="2335D7651D9741491E5264544BDE28CF" originalValue="mitprägt">mitpraegt</normalizedToken>
</emphasis>
<emphasis id="49B691204B34A1BFD3A7876E998585E2" italics="true">Charakterart</emphasis>
<emphasis id="2746A93D9E5440AD786BEB911AD42946" italics="true">Weniger strikt an den Lebensraum gebundene Art</emphasis>
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="C6FA2C1F88912C70C2DB31E361A961FA" type="biology_ecology">
<paragraph id="22951CE527366699E5CB38DA86422643">
<normalizedToken id="C28BABB1F362F9CD84B104589739F5CC" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="5A4EA17A35113D9912949BCD7652659C" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="E719DE4A848D0850F5475FFA45FBDB81">
<table id="8FDFBA18B851D419C2A3411E816A57BD" class="table" inLine="true">
<tbody id="053433A6B75C89AB31B2CD8B57A96D28">
<tr id="682C1C09CA6EB879E6054801E806B100">
<th id="CF94D8CF30C681410F95D4478CB9FD21" colspan="2">
<emphasis id="5B15E9E0EF475D33F5F54FD176AFD864" bold="true">Bodenfaktoren</emphasis>
</th>
<th id="640F1B910B1F393BE9BC9B867BCD3A57" colspan="2">
<emphasis id="C4393089A3621213C356F2E11F20A892" bold="true">Klimafaktoren</emphasis>
</th>
<th id="5A69DA66A7265B280ABD6C0888C65D53" colspan="2">
<emphasis id="EE000B4F3810FF841FB6E38C598AC037" bold="true">Salztoleranz</emphasis>
</th>
</tr>
<tr id="51DC05D62378B9363E9B65FA0478FB20">
<td id="2FE313D93E71D0F709D2A06C9EF5DEB5">Feuchtezahl F</td>
<td id="3665784BDB7014FF546C00B1E6049B04" style="text-align:left;width:100px">--</td>
<td id="790303100262FB5146A0F4D58DD70A26">Lichtzahl L</td>
<td id="B6D7A9558E5D70E699072913CED86C0B" style="text-align:left;width:60px">--</td>
<td id="DFFAA808D64ACC7D8FDFBA7589BB7BB0">Salzzeichen</td>
<td id="A4B401691027B90C93B213B84A4FB76D" style="text-align:left;width:60px">--</td>
</tr>
<tr id="10F199CF30CFB103C1269130E7198F32">
<td id="36C3D310A455D0DC8DD617C079F8ECBD">Reaktionszahl R</td>
<td id="B304493CDD6C5CBEAC79822EF44F3341" style="text-align:left">--</td>
<td id="67743E886470521EAEB7275A9F8164F5">Temperaturzahl T</td>
<td id="1DDA57A7C0EDC3A07F67C32070B9967F" style="text-align:left">--</td>
</tr>
<tr id="E7FF84DD52D78AB0B42DBAFF35433950">
<td id="F891579DF2CA0F8EE1A00EB342F50F3D">
<normalizedToken id="614A8F73FAEE85C4916A44678675CB00" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="9AE05C872C720049361F473A7F477A76" style="text-align:left">--</td>
<td id="FB97B108BFB02DB97CB47E27F803D8AD">
<normalizedToken id="4988FE3FCF696275A7C858910F1D8F23" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="65EFE52339ABD38835362D0E73083ED2" style="text-align:left">--</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="A706B5A103598AACEA998E2A8AC0E2D8" type="reference_group">
<paragraph id="D6571300455EA8C489C52831A1EABFE7">
<heading id="F180B70AAD0201AADA50D25516219CBE">Nomenklatur</heading>
</paragraph>
<paragraph id="BBC518E8A2F7F92C69B3BCA6F11D7164">
<treatmentCitation id="A341638EE5AF555F18AFD45A61BF5E32">
<heading id="E706AFD13F06CE875FA28BBB51CD13DD">
<normalizedToken id="1ADDA502487930D208A1C201848BAA94" originalValue="Gültiger">Gueltiger</normalizedToken>
Name (
<bibRefCitation id="43ECBAF98CEB5C27E238D8AC196E3EE2" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</heading>
:
<taxonomicName id="69090BA9A5A80C0E44EF0E0FB108D5BA" ID-CoL="BHL4" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="decumbens">
<emphasis id="CE6A2097C748FBDDDE57F476929C9241" italics="true">Alchemilla decumbens</emphasis>
</taxonomicName>
</treatmentCitation>
aggr.
</paragraph>
</subSubSection>
<subSubSection id="3820A707CE63B61DE37AD2E9C607D24B" type="vernacular_names">
<paragraph id="24A932C466120CD08BA069B5D5E4A82A">
Volksname Deutscher Name:
<vernacularName id="6108136D8C68DC23D22AEFE3554DAA3E" language="deu">Niederliegender Frauenmantel</vernacularName>
Nom
<normalizedToken id="92489FAED0225675ED2935800675E75C" originalValue="français">francais</normalizedToken>
:
<vernacularName id="33EFD2B2DB05A5AEFEC67EE141355C91" language="fra">
<normalizedToken id="7433891A7B14C651E56592E98BD85E53" originalValue="Alchémille">Alchemille</normalizedToken>
rampante
</vernacularName>
Nome italiano:
<vernacularName id="DF002E3CBEE9D60ADAC2ED909E98D087" language="ita">Ventaglina sdraiata</vernacularName>
</paragraph>
</subSubSection>
<subSubSection id="1925558423F0CCB0A674274DD8A359F0" type="reference_group">
<paragraph id="05368F2B474FBBF7848269148395F8E2">
<normalizedToken id="8BF8C2A504A73F47850E340220BA804D" originalValue="Übereinstimmung">Uebereinstimmung</normalizedToken>
mit anderen Referenzwerken
</paragraph>
<paragraph id="EA6B7C7C7010BA78162BDAF761888D83">
<table id="D7BBE86FA2F163679D75CBB55B2E7855" class="table table-condensed" inLine="true">
<tbody id="0DB7A41CE592D7A982F42C3BFB17984E">
<tr id="BE90317C59942CC2D366607DC0C7FB40">
<th id="4B0793F7EBA2FEB6ABFBC3AC4771526B" width="80px">Relation</th>
<th id="1C458DF0FD0EFDCBF6123FA8382B6CDA">Nom</th>
<th id="DD0207EDE82EE1DF959D7462D3A30EDA">Referenzwerke</th>
<th id="6DBE65D23CA276A31119D03360AD579C" width="90px">No</th>
</tr>
<tr id="D82BF1CB52E5A995E6595179731E2F5A">
<td id="F84AA02B0AB9DDA1182B1BF6505960A0" class="taxon-relation">=</td>
<td id="3A82D1A74800FC5B35E5D3A73D0CCF6F">
<taxonomicName id="D918A9525245356C05763FEBAE969AEA">Alchemilla decumbens aggr.</taxonomicName>
</td>
<td id="CE1AAE7FC30F546E4824D1336D27097F">
<treatmentCitation id="3E03738973839DD108EDEE5772255552">
<bibRefCitation id="F3327EF6F41A3C54BA6B95C8EDD4A3E1" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
</treatmentCitation>
</td>
<td id="A62D896CB19C3EA250C9354D42F599BF">13100</td>
</tr>
<tr id="F64AD0094FEFF713B3F95CDEA1DA0ECE">
<td id="59A41D6751B1F73DF53C241ECD6EF825" class="taxon-relation">=</td>
<td id="A471A1B0E8916E33E93E87D5E172EAE8">
<taxonomicName id="3BCD2681937D89D69C75960392AAF969">Alchemilla decumbens aggr. sensu Heitz</taxonomicName>
</td>
<td id="475AEBBD15A33A8B10491272ECB1A12B">
<treatmentCitation id="B4604C3407D9CBE761B6B69D5F20C470">
<bibRefCitation id="EAA112FC2B77EDF4E0805CB9E97C1254">Index synonymique 1996</bibRefCitation>
</treatmentCitation>
</td>
<td id="9E71A9C88B7F5F86E1626178C40029F4">13100</td>
</tr>
<tr id="AC8B7E40E68FC0799FD7936E2D45846B">
<td id="A412CA01F11D09E2EA0F79E09298DDCE" class="taxon-relation">=</td>
<td id="66DAC37C37F3E389592D043EF619DAF1">
<taxonomicName id="E6A15CB42E719B282CDD5F1BD62CB06C">Alchemilla decumbens aggr. sensu Heitz</taxonomicName>
</td>
<td id="05647F576FFC960EE10CA5FD154AE331">
<treatmentCitation id="A69A7B50C762785D32ADA28CEAB82908">
<bibRefCitation id="1F9CB0FDC034A6B9EDA6F869B5DDB285">SISF/ISFS 2</bibRefCitation>
</treatmentCitation>
</td>
<td id="9EEDD7AEB2C7055A12A3F349EFB0BE54">13100</td>
</tr>
<tr id="58B05B157EBFC27A3884997D04E54581">
<td id="C07069186349BFEF2DDE02C76C2AA01A" class="taxon-relation">&lt;</td>
<td id="0C966806E983AD01D707F5200AB8A548">
<taxonomicName id="882FC35525010765577ABB0627BA18DB" authority="Buser" authorityName="Buser" class="Magnoliopsida" family="Rosaceae" genus="Alchemilla" kingdom="Plantae" order="Rosales" phylum="Tracheophyta" rank="species" species="decumbens">Alchemilla decumbens Buser</taxonomicName>
</td>
<td id="6C7F9D5E150EE013D222A3C69F180831">
<treatmentCitation id="4CE810E586D4166B852D1A5F5A91415D">
<bibRefCitation id="322071878A0A633AAE19290C52664DF4">SISF/ISFS 2</bibRefCitation>
</treatmentCitation>
</td>
<td id="617D7D9A4CC2272A5D0902F8E1A17B10">13000</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="E8BDEA082D5568397893E2B1907F275F">
<tableNote id="736B4C112B3CDF8AE63AF30BC212FDBD">
= Taxon stimmt mit akzeptiertem Taxon
<normalizedToken id="B86DCFF5BA69E8056D78E5FD2AC0F05C" originalValue="überein">ueberein</normalizedToken>
(
<bibRefCitation id="EF2646DC75329608977ABDDA822FD769" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) &lt;Taxon ist im akzeptierten Taxon (
<bibRefCitation id="64073D795E0A1DDB4B681BF965B7CE04" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) enthalten&gt; Taxon
<normalizedToken id="F315349A039111E22FFDA4FF16DD0361" originalValue="enthält">enthaelt</normalizedToken>
(neben anderen) auch das akzeptierte Taxon (
<bibRefCitation id="7C3186EF9CDB5B1AF1A19710D68FEEDB" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="C472C54BE0081A9CC039B18AEA51053F" type="multiple">
<paragraph id="8A1CA9853BCF7C3026B0E2574C275D4F">
Kommentare aus der
<bibRefCitation id="3638C90D1E61D625965F9B592E63F141" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
Korrektur Autorangabe: Die bisherige Zusatzbezeichnung &quot;sensu AUTOR&quot;
<normalizedToken id="D01A35F07ABC6BED02CB711346F4D294" originalValue="fällt">faellt</normalizedToken>
weg, da das Aggregat
<normalizedToken id="E3B364FA77D32E4A180B3BA50562CDB5" originalValue="über">ueber</normalizedToken>
die Gruppierung in der vorliegenden
<bibRefCitation id="C2314404906FC93A0343C805174A2840" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
und dem
<normalizedToken id="B53440B5B8411A8204DDB33DB8582295" originalValue="Schlüssel">Schluessel</normalizedToken>
der Flora Helvetica definiert wird. Checklist
</paragraph>
</subSubSection>
<subSubSection id="A0C3234D34DF143D19825D8C3B89F5F6" type="conservation">
<paragraph id="3C9DE81ABBA0975A8360B5E7DA2ADDAD">
<heading id="2F05CE37539BBE2A6C6CD39A5D8B6500">Status Indigenat</heading>
: Indigen
</paragraph>
<paragraph id="1712AC637B794FAD3385800FD8AB0F7D">
<emphasis id="BFA88599525794654FA186705C154609" bold="true">
<heading id="791CFB7147ABDD93705F349896B98867">
Liste der
<normalizedToken id="9B59B412DACDCF13AABE61242D719637" originalValue="gefährdeten">gefaehrdeten</normalizedToken>
Pflanzen IUCN
</heading>
(nach
<bibRefCitation id="F0EE42A77FCC2DB7E57A4A3356FE36EB" author="Walter, K.S. &amp; Gillett, H.J." location="Gland, Switzerland and Cambridge, UK" publicationUrl="https://doi.org/10.5962/bhl.title.44833" publisher="IUCN" refString="Walter, K.S. and Gillett, H.J. (editors). 1998. 1997 IUCN Red List of Threatened Plants. IUCN, Gland, Switzerland and Cambridge, UK. DOI: https://doi.org/10.5962/bhl.title.44833" title="IUCN Red List of Threatened Plants" year="1997">Walter &amp; Gillett 1997</bibRefCitation>
):
</emphasis>
Nein
</paragraph>
<paragraph id="C383547F11757972F4B2C79E8080E19D">
<heading id="028E403892A387BD58D81AAC7C9A06D6">Status Rote Liste national 2016</heading>
</paragraph>
<paragraph id="856497B66349A7FA610A1C9905E227A9">
<heading id="B9E108B477C598C9855DAC652B3C36B4">Status IUCN</heading>
: Nicht
<normalizedToken id="2958AB319374BA9029B603D49343F8E7" originalValue="gefährdet">gefaehrdet</normalizedToken>
</paragraph>
<paragraph id="1D7BD813019278A29E350A8161E43D22">
<heading id="576FC5B2C9DED7D31E57D57AF1CACB16">
<normalizedToken id="3B3C7331E8927662C44840C48142DAEA" originalValue="Zusätzliche">Zusaetzliche</normalizedToken>
Informationen
</heading>
Kriterien IUCN: --
</paragraph>
<paragraph id="0A48DED5A39F0E7AE215F7BA68ADC097">
<heading id="C7A31EBBB141B9C3F6AD50296EBC36FD">Status Rote Liste regional 2019</heading>
</paragraph>
<paragraph id="292AB69E865BC9C380EF139854EC6026">
<table id="88F27094CD3205F7471FB129C51B2137" class="table table-condensed" inLine="true">
<tbody id="226611EFB77385FB459659ED5F26F9F5">
<tr id="070F80A9989D2B195B7A38B6C3D5D8FE">
<th id="1EFB5102F831F219F8CE4D9F20A1B708">Biogeografische Regionen</th>
<th id="392A3E431554F331853B74EEEE04F4B6">Status</th>
<th id="A882C4C7A7BEEB67F34AC808486A8FFA">Kriterien IUCN</th>
</tr>
<tr id="2A29EEBE841397CAF803D68089F646FE">
<td id="DA546843AC140049AB182C9A919EA805" class="regional_status">Jura (JU)</td>
<td id="CCD41AC43D3E0D21E8213BBE726CDED0">nicht beurteilt (Not Evaluated)</td>
</tr>
<tr id="7701E2D8A2C26711A58500AC1BD5C96A">
<td id="78CA6EEF69F8AFE39A17B273EA767F6C" class="regional_status">Mittelland (MP)</td>
<td id="7B02F88B1A9F95E8F68B5C84F6B30022">nicht beurteilt (Not Evaluated)</td>
</tr>
<tr id="C93754E67EAA227FAC33855A64F62D16">
<td id="05AECD36D75AE8CBB6C7EA8EF19C8FEE" class="regional_status">Alpennordflanke (NA)</td>
<td id="D64C49ED97E567142319EA5870C5ABAE">nicht beurteilt (Not Evaluated)</td>
</tr>
<tr id="B17E9FB35FF302A018EB69CE008DC2B4">
<td id="028237E91AAB82E40262E6643A7C0C16" class="regional_status">
<normalizedToken id="B489E1FA5917D65907B8E87FC7164C2F" originalValue="Alpensüdflanke">Alpensuedflanke</normalizedToken>
(SA)
</td>
<td id="D49B55D3391E386A7A333C1E54E7A574">nicht beurteilt (Not Evaluated)</td>
</tr>
<tr id="6246D3306C5C8530757C2BEA1A7B3FFC">
<td id="976554A93C8FA4FEF2DF7A29B910173C" class="regional_status">
<normalizedToken id="E79B00ADBD677EF3C613FA738BF8D6EE" originalValue="Östliche">Oestliche</normalizedToken>
Zentralalpen (EA)
</td>
<td id="B84BDDCAB5F2CF097F8A7E242EFB311A">nicht beurteilt (Not Evaluated)</td>
</tr>
<tr id="5A2552F82DF19165FC6F3CD54C2B80C2">
<td id="1D9CA7CAF5BD71D380CF016D8FFDDDA7" class="regional_status">Westliche Zentralalpen (WA)</td>
<td id="73CA8BB0F1B2D3B7949C2B93F89CFF1E">nicht beurteilt (Not Evaluated)</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="EDA6A6B86C1E93E4806E048493BC6CFA">
<heading id="938D597133359B103354FDA0247186ED">
Status nationale
<normalizedToken id="7F841CDDD29FB4F48EFB2E3218CDDC04" originalValue="Priorität">Prioritaet</normalizedToken>
/Verantwortung
</heading>
</paragraph>
<paragraph id="DCA866A6CA82E52B940DCD3CAD402503">
<table id="6EC090E8402CA077AC8F735B1A01F5CA" class="table" inLine="true">
<tr id="8CE6085646BEE6065E8FA7FD2D07E320">
<td id="0AACEE91887B31B805829C85C1476B6A">
Keine nationale
<normalizedToken id="E34571DFCB9FA5AECD0EFF151A486A43" originalValue="Priorität">Prioritaet</normalizedToken>
oder internationale Verantwortung
</td>
</tr>
</table>
</paragraph>
<paragraph id="9A8856B487D996860C287E1649413FF8">
<heading id="2FDD92A883D1DB134BC7ECC5674D1F50">Schutzstatus</heading>
</paragraph>
<paragraph id="2F6863D73C5BDD7BB88DF8459E732532">
<table id="E60402C3A93470FA7B23040DB184F709" class="table" inLine="true">
<tr id="FCDCE82ECD74DCD1773B13BBF116A5FF">
<td id="C9BDA38E640861FC3D5B9F85520F7E61">Kein internationaler, nationaler oder kantonaler Schutz</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

205
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<mods:title>A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini)</mods:title>
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<mods:namePart>Maddison, Wayne P.</mods:namePart>
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<mods:namePart>Beattie, Imara</mods:namePart>
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<mods:namePart>Marathe, Kiran</mods:namePart>
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<mods:namePart>Kanesharatnam, Nilani</mods:namePart>
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<mods:namePart>Benjamin, Suresh P.</mods:namePart>
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<taxonomicName LSID="AF6389BD-82C6-5C13-88F6-91AFD766E090" authority="Simon, 1885" authorityName="Simon" authorityYear="1885" class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus opaciceps" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="opaciceps">Stagetillus opaciceps Simon, 1885</taxonomicName>
<figureCitation captionStart="Figures 4–35" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">Figs 8</figureCitation>
<figureCitation captionStart="Figures 4–35" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">, 15</figureCitation>
<figureCitation captionStart="Figures 4–35" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">, 16</figureCitation>
<figureCitation captionStart="Figures 4–35" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">, 23</figureCitation>
<figureCitation captionStart="Figures 4–35" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">, 31</figureCitation>
<figureCitation captionStart="Figures 143–150" captionStartId="F16" captionText="Figures 143 - 150. Stagetillus opaciceps male, and a female tentatively identified as S. opaciceps. 143 Male left palp, ventral view (specimen JK. 13.02.26.0017) 144 same, retrolateral view of tibia 145 living male from Belait, Brunei (photograph Joseph Koh 2019) 146, 147 male (specimen JK. 08.08.19.0001) 148 female epigyne (specimen MRB 079), ventral 149 vulva, dorsal 150 living female. Scale bars: on genitalia 0.1 mm; on bodies 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures143-150" httpUri="https://binary.pensoft.net/fig/487547" pageId="0" pageNumber="27">, 143-150</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="reference_group">
<paragraph pageId="0" pageNumber="27">
<taxonomicName authorityName="Simon" authorityYear="1885" class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus opaciceps" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="opaciceps">Stagetillus opaciceps</taxonomicName>
Simon, 1885: 32.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="notes">
<paragraph pageId="0" pageNumber="27">Note.</paragraph>
<paragraph pageId="0" pageNumber="27">
<bibRefCitation author="Proszynski, J" journalOrPublisher="Zeszyty Naukowe Wyzszej Szkoly Rolniczo-Pedagogicznej, Siedlcach" pageId="0" pageNumber="27" refId="B38" refString="Proszynski, J, 1987. Atlas rysunkow diagnostycznych mniej znanych Salticidae 2. Zeszyty Naukowe Wyzszej Szkoly Rolniczo-Pedagogicznej, Siedlcach" title="Atlas rysunkow diagnostycznych mniej znanych Salticidae 2." year="1987">
<normalizedToken originalValue="Prószyński’s">Proszynski's</normalizedToken>
(1987)
</bibRefCitation>
illustrations of
<normalizedToken originalValue="Simon’s">Simon's</normalizedToken>
type specimen characterize the male well. Two additional males illustrated here (Figs
<figureCitation captionStart="Figures 143–150" captionStartId="F16" captionText="Figures 143 - 150. Stagetillus opaciceps male, and a female tentatively identified as S. opaciceps. 143 Male left palp, ventral view (specimen JK. 13.02.26.0017) 144 same, retrolateral view of tibia 145 living male from Belait, Brunei (photograph Joseph Koh 2019) 146, 147 male (specimen JK. 08.08.19.0001) 148 female epigyne (specimen MRB 079), ventral 149 vulva, dorsal 150 living female. Scale bars: on genitalia 0.1 mm; on bodies 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures143-150" httpUri="https://binary.pensoft.net/fig/487547" pageId="0" pageNumber="27">143-147</figureCitation>
) are conspecific or at least very close (the male from Brunei, Figs
<figureCitation captionStart="Figures 143–150" captionStartId="F16" captionText="Figures 143 - 150. Stagetillus opaciceps male, and a female tentatively identified as S. opaciceps. 143 Male left palp, ventral view (specimen JK. 13.02.26.0017) 144 same, retrolateral view of tibia 145 living male from Belait, Brunei (photograph Joseph Koh 2019) 146, 147 male (specimen JK. 08.08.19.0001) 148 female epigyne (specimen MRB 079), ventral 149 vulva, dorsal 150 living female. Scale bars: on genitalia 0.1 mm; on bodies 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures143-150" httpUri="https://binary.pensoft.net/fig/487547" pageId="0" pageNumber="27">146</figureCitation>
,
<figureCitation captionStart="Figures 143–150" captionStartId="F16" captionText="Figures 143 - 150. Stagetillus opaciceps male, and a female tentatively identified as S. opaciceps. 143 Male left palp, ventral view (specimen JK. 13.02.26.0017) 144 same, retrolateral view of tibia 145 living male from Belait, Brunei (photograph Joseph Koh 2019) 146, 147 male (specimen JK. 08.08.19.0001) 148 female epigyne (specimen MRB 079), ventral 149 vulva, dorsal 150 living female. Scale bars: on genitalia 0.1 mm; on bodies 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures143-150" httpUri="https://binary.pensoft.net/fig/487547" pageId="0" pageNumber="27">147</figureCitation>
, has a slightly narrower embolus). Females of the species have not been reported, although we now tentatively identify, as the female of
<taxonomicName family="Salticidae" kingdom="Animalia" lsidName="S. opaciceps" order="Araneae" pageId="0" pageNumber="27" rank="species" species="opaciceps">
<emphasis italics="true" pageId="0" pageNumber="27">S. opaciceps</emphasis>
</taxonomicName>
, the specimen MRB079 from Ulu Gombak used by
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.440.7891" author="Maddison, WP" journalOrPublisher="ZooKeys" pageId="0" pageNumber="27" pagination="57 - 87" refId="B30" refString="Maddison, WP, Li, DQ, Bodner, MR, Zhang, JX, Xu, X, Liu, QQ, 2014. The deep phylogeny of jumping spiders (Araneae, Salticidae). ZooKeys 440: 57 - 87, DOI: https://doi.org/10.3897/zookeys.440.7891" title="The deep phylogeny of jumping spiders (Araneae, Salticidae)." url="https://doi.org/10.3897/zookeys.440.7891" volume="440" year="2014">Maddison et al. (2014)</bibRefCitation>
for molecular phylogeny (Figs
<figureCitation captionStart="Figures 143–150" captionStartId="F16" captionText="Figures 143 - 150. Stagetillus opaciceps male, and a female tentatively identified as S. opaciceps. 143 Male left palp, ventral view (specimen JK. 13.02.26.0017) 144 same, retrolateral view of tibia 145 living male from Belait, Brunei (photograph Joseph Koh 2019) 146, 147 male (specimen JK. 08.08.19.0001) 148 female epigyne (specimen MRB 079), ventral 149 vulva, dorsal 150 living female. Scale bars: on genitalia 0.1 mm; on bodies 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures143-150" httpUri="https://binary.pensoft.net/fig/487547" pageId="0" pageNumber="27">148-150</figureCitation>
). That the female is a
<taxonomicName authorityName="Simon" authorityYear="1885" class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Stagetillus</emphasis>
</taxonomicName>
is suggested by its similarities to the male in size and body form, carapace shape, yellow coloration, and visibility of the ocular digestive diverticulum. The modest ECP and epigynal openings are what might be expected from the small simple RTA and embolus of the known male of
<taxonomicName family="Salticidae" kingdom="Animalia" lsidName="S. opaciceps" order="Araneae" pageId="0" pageNumber="27" rank="species" species="opaciceps">
<emphasis italics="true" pageId="0" pageNumber="27">S. opaciceps</emphasis>
</taxonomicName>
. Given that Ulu Gombak is only 70 km from a known male of
<taxonomicName family="Salticidae" kingdom="Animalia" lsidName="S. opaciceps" order="Araneae" pageId="0" pageNumber="27" rank="species" species="opaciceps">
<emphasis italics="true" pageId="0" pageNumber="27">S. opaciceps</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figures 143–150" captionStartId="F16" captionText="Figures 143 - 150. Stagetillus opaciceps male, and a female tentatively identified as S. opaciceps. 143 Male left palp, ventral view (specimen JK. 13.02.26.0017) 144 same, retrolateral view of tibia 145 living male from Belait, Brunei (photograph Joseph Koh 2019) 146, 147 male (specimen JK. 08.08.19.0001) 148 female epigyne (specimen MRB 079), ventral 149 vulva, dorsal 150 living female. Scale bars: on genitalia 0.1 mm; on bodies 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures143-150" httpUri="https://binary.pensoft.net/fig/487547" pageId="0" pageNumber="27">143</figureCitation>
,
<figureCitation captionStart="Figures 143–150" captionStartId="F16" captionText="Figures 143 - 150. Stagetillus opaciceps male, and a female tentatively identified as S. opaciceps. 143 Male left palp, ventral view (specimen JK. 13.02.26.0017) 144 same, retrolateral view of tibia 145 living male from Belait, Brunei (photograph Joseph Koh 2019) 146, 147 male (specimen JK. 08.08.19.0001) 148 female epigyne (specimen MRB 079), ventral 149 vulva, dorsal 150 living female. Scale bars: on genitalia 0.1 mm; on bodies 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures143-150" httpUri="https://binary.pensoft.net/fig/487547" pageId="0" pageNumber="27">144</figureCitation>
), the match of the Ulu Gombak female to
<taxonomicName family="Salticidae" kingdom="Animalia" lsidName="S. opaciceps" order="Araneae" pageId="0" pageNumber="27" rank="species" species="opaciceps">
<emphasis italics="true" pageId="0" pageNumber="27">S. opaciceps</emphasis>
</taxonomicName>
is credible. We therefore label the female, used here for molecular data, as
<taxonomicName family="Salticidae" kingdom="Animalia" lsidName="S. opaciceps" order="Araneae" pageId="0" pageNumber="27" rank="species" species="opaciceps">S. cf. opaciceps</taxonomicName>
(Figs
<figureCitation captionStart="Figures 143–150" captionStartId="F16" captionText="Figures 143 - 150. Stagetillus opaciceps male, and a female tentatively identified as S. opaciceps. 143 Male left palp, ventral view (specimen JK. 13.02.26.0017) 144 same, retrolateral view of tibia 145 living male from Belait, Brunei (photograph Joseph Koh 2019) 146, 147 male (specimen JK. 08.08.19.0001) 148 female epigyne (specimen MRB 079), ventral 149 vulva, dorsal 150 living female. Scale bars: on genitalia 0.1 mm; on bodies 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures143-150" httpUri="https://binary.pensoft.net/fig/487547" pageId="0" pageNumber="27">148-150</figureCitation>
). In
<normalizedToken originalValue="WPM’s">WPM's</normalizedToken>
lab notebooks the informal code for the female was
<normalizedToken originalValue="“STULG”">&quot;STULG&quot;</normalizedToken>
.
</paragraph>
<caption doi="10.3897/zookeys.1004.57526.figures143-150" httpUri="https://binary.pensoft.net/fig/487547" pageId="0" pageNumber="27" start="Figures 143–150" startId="F16">
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Figures 143-150.</emphasis>
<taxonomicName authorityName="Simon" authorityYear="1885" class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus opaciceps" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="opaciceps">
<emphasis italics="true" pageId="0" pageNumber="27">Stagetillus opaciceps</emphasis>
</taxonomicName>
male, and a female tentatively identified as
<taxonomicName family="Salticidae" kingdom="Animalia" lsidName="S. opaciceps" order="Araneae" pageId="0" pageNumber="27" rank="species" species="opaciceps">
<emphasis italics="true" pageId="0" pageNumber="27">S. opaciceps</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="27">143</emphasis>
Male left palp, ventral view (specimen JK.13.02.26.0017)
<emphasis bold="true" pageId="0" pageNumber="27">144</emphasis>
same, retrolateral view of tibia
<emphasis bold="true" pageId="0" pageNumber="27">145</emphasis>
living male from Belait, Brunei (photograph Joseph Koh 2019)
<emphasis bold="true" pageId="0" pageNumber="27">146, 147</emphasis>
male (specimen JK.08.08.19.0001)
<emphasis bold="true" pageId="0" pageNumber="27">148</emphasis>
female epigyne (specimen MRB079), ventral
<emphasis bold="true" pageId="0" pageNumber="27">149</emphasis>
vulva, dorsal
<emphasis bold="true" pageId="0" pageNumber="27">150</emphasis>
living female. Scale bars: on genitalia 0.1 mm; on bodies 1.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="materials_examined">
<paragraph pageId="0" pageNumber="27">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="27">
Male (specimen JK.13.02.26.0017, in LKCNHM) Malaysia: Negeri Sembilan, Hutan Lipur Ulu Bendul,
<geoCoordinate degrees="2.73" direction="north" orientation="latitude" precision="555" value="2.73">2.73°N</geoCoordinate>
,
<geoCoordinate degrees="102.0789" direction="east" orientation="longitude" precision="5" value="102.0789">102.0789°E</geoCoordinate>
, J. K. H. Koh 26 February 2013; Male (JK.08.08.19.0001, in LKCNHM) Brunei: Belait, Disturbed forest off Labi Road,
<geoCoordinate degrees="4.5858" direction="north" orientation="latitude" precision="5" value="4.5858">4.5858°N</geoCoordinate>
,
<geoCoordinate degrees="114.5067" direction="east" orientation="longitude" precision="5" value="114.5067">114.5067°E</geoCoordinate>
, J. K. H. Koh 19 August 2008. Female (specimen MRB079, in UBCZ) Malaysia: Selangor: Ulu Gombak Field Station.
<geoCoordinate degrees="3.325" direction="north" orientation="latitude" precision="55" value="3.325">3.325°N</geoCoordinate>
,
<geoCoordinate degrees="101.753" direction="east" orientation="longitude" precision="55" value="101.753">101.753°E</geoCoordinate>
, 250 m el. 16-19 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang, WPM#05-026.
</paragraph>
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Casey, 1914: 246. Type locality: &quot;near the city [of New York], New York&quot; (original citation). Lectotype (♂), designated by Lindroth (1975: 143), in USNM [# 47999].
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Type status:
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. Occurrence: catalogNumber:
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; Taxon: scientificName: Ophioglossum reticulatum L.; namePublishedIn: Sp. Pl. 2: 1063 (1753); kingdom: Plantae; phylum: Pteridophyta; class: Polypodiopsida; order: Ophioglossales; family: Ophioglossaceae; genus: Ophioglossum; specificEpithet: reticulatum; scientificNameAuthorship: L.; Location: continent: Africa; country:
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; basisOfRecord: Preserved specimen
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</paragraph>
</subSubSection>
<subSubSection id="1C75FA1465C5A4504AC8C58AD9CFBA7B" pageId="0" pageNumber="24137" type="ecological interactions">
<paragraph id="7347A1CEE51D8002840201028BE412E9" pageId="0" pageNumber="24137">Ecological interactions</paragraph>
<subSection id="4DFA28BFF422A1E084F603BA9C3A13C3" pageId="0" pageNumber="24137" type="native status">
<paragraph id="1EAC9F5D5868D2C720E35FD720F89D04" pageId="0" pageNumber="24137">Native status</paragraph>
<paragraph id="649CF581555C005ED28DE6A7BA7D33DD" pageId="0" pageNumber="24137">Native</paragraph>
</subSection>
</subSubSection>
<subSubSection id="235B110224F855123C69DE7F7219AC10" pageId="0" pageNumber="24137" type="distribution">
<paragraph id="D82E09192CC57D0F4766B1E1FDAA1ACD" pageId="0" pageNumber="24137">Distribution</paragraph>
<paragraph id="BE8A3F86D056613DD164F0E569247C15" pageId="0" pageNumber="24137">Zone 3</paragraph>
</subSubSection>
</treatment>
</document>