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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.578.7820" ID-GBIF-Dataset="0d8819ce-6160-446c-85bc-bc18ceaaf10c" ID-PMC="PMC4829961" ID-Pensoft-Pub="1313-2970-578-45" ID-PubMed="27110196" ID-ZBK="50E7510F61FB444ABF5E1DA830ED4633" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-578-45" ModsDocOrigin="ZooKeys 578" ModsDocTitle="Contribution to the knowledge of Afrotropical Dryinidae, Embolemidae and Sclerogibbidae (Hymenoptera), with description of new species from Central African Republic and Uganda" checkinTime="1460141818630" checkinUser="pensoft" docAuthor="Olmi, Massimo, van Noort, Simon &amp; Guglielmino, Adalgisa" docDate="2016" docId="3BEC542D95C774C523EE89656E948077" docLanguage="en" docName="ZooKeys 578: 45-95" docOrigin="ZooKeys 578" docSource="http://dx.doi.org/10.3897/zookeys.578.7820" docTitle="Anteon ngoyense Olmi" docType="treatment" docVersion="8" lastPageNumber="66" masterDocId="FF8F2C440F19BB7AFFA3991A3B5CFFE6" masterDocTitle="Contribution to the knowledge of Afrotropical Dryinidae, Embolemidae and Sclerogibbidae (Hymenoptera), with description of new species from Central African Republic and Uganda" masterLastPageNumber="95" masterPageNumber="45" pageNumber="65" updateTime="1668162931333" updateUser="ExternalLinkService">
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<mods:title>Contribution to the knowledge of Afrotropical Dryinidae, Embolemidae and Sclerogibbidae (Hymenoptera), with description of new species from Central African Republic and Uganda</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Olmi, Massimo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>van Noort, Simon</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Guglielmino, Adalgisa</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>578</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.578.7820</mods:url>
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<treatment ID-GBIF-Taxon="150811254" LSID="urn:lsid:plazi:treatment:3BEC542D95C774C523EE89656E948077" httpUri="http://treatment.plazi.org/id/3BEC542D95C774C523EE89656E948077" lastPageId="21" lastPageNumber="66" pageId="20" pageNumber="65">
<subSubSection pageId="20" pageNumber="65" type="multiple">
<paragraph pageId="20" pageNumber="65">Taxon classification Animalia Hymenoptera Dryinidae</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="65" type="nomenclature">
<paragraph pageId="20" pageNumber="65">
<taxonomicName authority="Olmi" class="Insecta" family="Dryinidae" genus="Anteon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anteon ngoyense" order="Hymenoptera" pageId="20" pageNumber="65" phylum="Arthropoda" rank="species" species="ngoyense">Anteon ngoyense Olmi</taxonomicName>
</paragraph>
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<paragraph pageId="20" pageNumber="65">
<taxonomicName class="Insecta" family="Dryinidae" genus="Anteon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anteon ngoyense" order="Hymenoptera" pageId="20" pageNumber="65" phylum="Arthropoda" rank="species" species="ngoyense">Anteon ngoyense</taxonomicName>
Olmi, 2009: 451.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="65" type="materials_examined">
<paragraph pageId="20" pageNumber="65">Material examined.</paragraph>
<paragraph pageId="20" pageNumber="65">
Published record.
<bibRefCitation author="Olmi, M" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="46" pageNumber="91" pagination="333 - 380" title="An updated checklist of Dryinidae, Embolemidae and Sclerogibbidae (Hymenoptera) of Kenya and Burundi, with descriptions of thirteen new species." volume="55" year="2015">Olmi et al. (2015</bibRefCitation>
: CENTRAL AFRICAN REPUBLIC:
<normalizedToken originalValue="Sangha-Mbaéré">Sangha-Mbaere</normalizedToken>
Prefecture: Dzanga-Ndoki National Park,
<normalizedToken originalValue="Mabéa">Mabea</normalizedToken>
Bai,21.4 Km 53°NE Bayanga,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="3.033611">03°02'01&quot;N</geoCoordinate>
<geoCoordinate direction="east" orientation="longitude" precision="15" value="16.415833">16°24'57&quot;E</geoCoordinate>
, 510 m, 4-5.V.2001, Malaise trap, lowland rainforest, marsh clearing, S. van Noort leg., 1♂ (SAMC).
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="66" type="hosts">
<paragraph pageId="21" pageNumber="66">
<pageBreakToken pageId="21" pageNumber="66" start="start">Hosts</pageBreakToken>
.
</paragraph>
<paragraph pageId="21" pageNumber="66">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="66" type="distribution">
<paragraph pageId="21" pageNumber="66">Distribution.</paragraph>
<paragraph pageId="21" pageNumber="66">
Central African Republic, South Africa, Uganda (
<bibRefCitation author="Olmi, M" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="46" pageNumber="91" pagination="333 - 380" title="An updated checklist of Dryinidae, Embolemidae and Sclerogibbidae (Hymenoptera) of Kenya and Burundi, with descriptions of thirteen new species." volume="55" year="2015">Olmi et al. 2015</bibRefCitation>
).
</paragraph>
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</treatment>
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<taxonomicName authority="Jordan &amp; Eigenmann, 1889" authorityName="Jordan &amp; Eigenmann" authorityYear="1889" class="Actinopterygii" family="Sciaenidae" genus="Pareques" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pareques umbrosus" order="Perciformes" pageId="0" pageNumber="1100" phylum="Chordata" rank="species" species="umbrosus">Pareques umbrosus (Jordan &amp; Eigenmann, 1889)</taxonomicName>
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<paragraph pageId="0" pageNumber="1100">Materials</paragraph>
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<materialsCitation collectingDate="2007-09-20" collectingMethod="Photosampling" collectionCode="UMDI-SISAL, CIRR" collectorName="Salvador Zarco Perello" country="Mexico" latitude="21.443655" location="Madagascar Reef" longitude="-90.293365" pageId="0" pageNumber="1100" specimenCode="CIRR- 300" specimenCount="5" typeStatus="Other material">
Type status:
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. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1100">CIRR-300</specimenCode>
; recordedBy:
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; individualCount:
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; Location: continent: America; country:
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; stateProvince: Yucatan; locality:
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; verbatimDepth: 16 m; verbatimLatitude: 780535.103072; verbatimLongitude: 2373673.16789; verbatimCoordinateSystem: UTM 15N; verbatimSRS: WGS84; decimalLatitude:
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; Record Level: collectionID: YUC-PEC_239-01-64; institutionCode:
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; collectionCode:
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<paragraph pageId="0" pageNumber="1100">Western Atlantic. Virginia to Brazil. Including Bermuda, Bahamas and throughout the Caribbean Islands.</paragraph>
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</treatment>
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<taxonomicName id="AD57908791AC3783B0DBE4D9D08A35BD" ID-CoL="3NY3R" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29521" class="Insecta" family="Formicidae" genus="Hypoponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Hypoponera nivariana (Santschi)" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nivariana">Hypoponera nivariana (Santschi)</taxonomicName>
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<paragraph id="34E3BFAB380D274CEDC166DCC3DA143B" pageId="75" pageNumber="76">
<taxonomicName id="4DA15FA680CE8099A5BD0B7144018FAA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138865" class="Insecta" family="Formicidae" genus="Ponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Ponera nivariana Santschi" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nivariana">Ponera nivariana Santschi</taxonomicName>
, 1908: 518. Holotype worker, SPAIN: Canary Is, Tenerife, Bajamon, 19.ix.1898 (Cabrera &amp; Diaz) (NHMB) [examined]. [Combination in
<taxonomicName id="98EF8BB6B586A66B997023648B04BC46" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24837" class="Insecta" family="Formicidae" genus="Hypoponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Hypoponera Santschi" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">Hypoponera</taxonomicName>
: Barquin Diez, 1981: 50.]
</paragraph>
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<paragraph id="7DA75BA33AE2AB24C57C71C19A8A2D89" pageId="75" pageNumber="76">WORKER (redescription of holotype). Measurements: HL 0.56, HW 0.46, HS 0.510, SL 0.38, PrW 0.34, WL 0.66, HFL 0.37, PeNL 0.14, PeH 0.30, PeNW 0.22, PeS 0.220. Indices: CI 82, SI 83, PeNI 65, LPeI 47, DPeI 157.</paragraph>
<paragraph id="8087E572506AB5A8165F0732345FEAFF" pageId="75" pageNumber="76">Eyes small but distinct, far forward on side, composed of a low but indeterminate number of partially fused small ommatidia (left eye depigmented). Mid-dorsal longitudinal impression on head terminates immediately behind the frontal lobes and does not extend to the midlength of the vertex or beyond. Apex of scape, when laid straight back from its insertion, fails to reach the midpoint of the posterior margin in full-face view by a distance that is less than the apical scape width; SL/HL 0.68. Cephalic dorsum extremely finely and superficially punctulate. Mesonotal-mesopleural suture absent. Metanotal groove conspicuously incised across dorsum of mesosoma; mesonotum with a well-defined posterior margin. Propodeal declivity bluntly rounded into the sides, without sharp marginations or carinae. Dorsum of mesosoma almost smooth, the minute punctulae that are present are superficial and inconspicuous. Upper half of mesopleuron with very fine superficial sculpture that is weaker than sculpture on side of propodeum; lower half of mesopleuron appears smoother (not clearly visible). Petiole in profile with the anterior and posterior faces of the node almost parallel; faces converge very slightly only at the apex, where they round into the weakly convex dorsum. Subpetiolar process in profile with an obtuse ventral angle. Maximum width of first gastral tergite in dorsal view less than the width of the second tergite at its midlength. Base of cinctus of second gastral tergite smooth and shining in dorsal view, without cross-ribs. Posttergite of second gastral segment, from posterior margin of cinctus to apex, much broader than long. Disc of second gastral tergite minutely superficially punctulate. Full adult colour a uniform dull yellow.</paragraph>
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<subSubSection id="8927D787941D699DA649043A2645460F" pageId="76" pageNumber="77" type="discussion">
<paragraph id="83E715C31476F3B7460C41BDAF3206A6" pageId="76" pageNumber="77">
<pageBreakToken id="7196FD7D0A517BFA28F6B443F6547A81" pageId="76" pageNumber="77" start="start">At</pageBreakToken>
first glance the holotype of
<taxonomicName id="F2394065605D9A29A38FA9BFC8EDB04A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138865" class="Insecta" family="Formicidae" genus="Ponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Ponera nivariana Santschi" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="nivariana">nivariana</taxonomicName>
appears to be a small, depigmented version of
<taxonomicName id="139F758134B67A6E4573B836A58F825C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29472" class="Insecta" family="Formicidae" genus="Hypoponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Hypoponera eduardi (Forel)" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="eduardi">eduardi</taxonomicName>
that has lost most of its sculpture, but whose dimensions fall within the lower end of the
<taxonomicName id="DC71116761BCA6FE9DAECFC70642EDE8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29534" class="Insecta" family="Formicidae" genus="Hypoponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Hypoponera punctatissima (Roger)" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="punctatissima">punctatissima</taxonomicName>
range. It is separated from these two species as follows.
</paragraph>
<paragraph id="F917BFD7F2A1D907EA0D47EB923A9716" pageId="76" pageNumber="77">
It differs from
<taxonomicName id="618460D4722381209749788DC6643CD3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29472" class="Insecta" family="Formicidae" genus="Hypoponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Hypoponera eduardi (Forel)" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="eduardi">eduardi</taxonomicName>
because in
<taxonomicName id="33139E06910A34EE1BE0D1F11B5FE6CA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138865" class="Insecta" family="Formicidae" genus="Ponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Ponera nivariana Santschi" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="nivariana">nivariana</taxonomicName>
colour is dull yellow, dorsum of mesosoma is almost smooth, sculpture of mesopleuron is feeble and superficial, scape is shorter (SI 83, SL/HL 0.68) and fails to reach midpoint of posterior margin of head in full-face view (for comparison, in
<taxonomicName id="F689D46CBE345F5D786027A210F27AA8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29472" class="Insecta" family="Formicidae" genus="Hypoponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Hypoponera eduardi (Forel)" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="eduardi">eduardi</taxonomicName>
SI 86
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93, SL/HL 0.72
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0.78). Overall size of
<taxonomicName id="42674D1CC2A58546A463E3C5EE60F564" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138865" class="Insecta" family="Formicidae" genus="Ponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Ponera nivariana Santschi" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="nivariana">nivariana</taxonomicName>
is smaller (HL 0.56, HW 0.46, SL 0.38) when compared to
<taxonomicName id="8D5B3D04C6A0A4785D32B76D8584EF85" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29472" class="Insecta" family="Formicidae" genus="Hypoponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Hypoponera eduardi (Forel)" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="eduardi">eduardi</taxonomicName>
(HL 0.63
<normalizedToken id="9A9B735AB009215C4B9DD9BD480E9B67" originalValue="–" pageId="76" pageNumber="77">-</normalizedToken>
0.70, HW 0.54
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0.59, SL 0.47
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0.54).
</paragraph>
<paragraph id="BABDB8747FA802BFE5A4E723D2B218B1" pageId="76" pageNumber="77">
It differs from
<taxonomicName id="4170ADE8AA1167CA456C1259ACC84A15" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29534" class="Insecta" family="Formicidae" genus="Hypoponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Hypoponera punctatissima (Roger)" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="punctatissima">punctatissima</taxonomicName>
because in
<taxonomicName id="4467DB208052C7327A4E42F864FF441A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138865" class="Insecta" family="Formicidae" genus="Ponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Ponera nivariana Santschi" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="nivariana">nivariana</taxonomicName>
there is no elongate mid-dorsal cephalic impression, sculpture is present on the mesopleuron, the anterior and posterior faces of the petiole node in profile do not converge towards the apex and the subpetiolar process has an obtuse ventral angle.
</paragraph>
<paragraph id="1A1FF6A4A50A3958468FD7FF4D8A4F1F" pageId="76" pageNumber="77">
<taxonomicName id="2D43C744B1A834FFED822BBCAC81A84F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29521" class="Insecta" family="Formicidae" genus="Hypoponera" genus.bestMatchDistance="1866" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Hypoponera nivariana (Santschi)" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="nivariana">H. nivariana</taxonomicName>
is known from only five collections (Barquin Diez, 1981), all from Tenerife in the Canary Islands. The number of specimens from these five collections totals a mere 13 workers and a single queen, so
<taxonomicName id="02F18EEA637C38D679B3BED2C3AD1C72" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138865" class="Insecta" family="Formicidae" genus="Ponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Ponera nivariana Santschi" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="nivariana">nivariana</taxonomicName>
cannot be considered a common or successful species. It may be a Canaries endemic, or an Afrotropical species that has not yet been discovered on the continent; or it may represent a hybrid between
<taxonomicName id="BD19935B49A036D7D31D1EE33304A2DC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29472" class="Insecta" family="Formicidae" genus="Hypoponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Hypoponera eduardi (Forel)" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="eduardi">eduardi</taxonomicName>
and punctatissima. This last is highly speculative and without proof, though the fact that it possesses a strange mix of characters found in the two other species is suggestive. Its taxonomic history is short and the name
<taxonomicName id="8C7348265EBA07F3ECB7998CDEE9376F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:138865" class="Insecta" family="Formicidae" genus="Ponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Ponera nivariana Santschi" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="nivariana">nivariana</taxonomicName>
has occurred in the literature very few times following
<normalizedToken id="1F02E12DFD029CAC6CC4942F2C8FCEEE" originalValue="Santschi’s" pageId="76" pageNumber="77">Santschi's</normalizedToken>
original description. Wheeler (1927a), Wellenius (1955) and Hohmann, et al. (1993) all merely listed the species as present and endemic in the Canaries. Barquin Diez (1981) redescribed the species, added a few more details of morphology and correctly concluded that it was
<normalizedToken id="93B93D9CBF759036144CB3F62C7728AA" originalValue="“" pageId="76" pageNumber="77">&quot;</normalizedToken>
close to
<taxonomicName id="EA1E0E9B8E2123B05C71F231F9839AF8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:29472" class="Insecta" family="Formicidae" genus="Hypoponera" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Hypoponera eduardi (Forel)" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="eduardi">eduardi</taxonomicName>
<normalizedToken id="4A4C9CD45DDF307CE5B281DBF87F4320" originalValue="”" pageId="76" pageNumber="77">&quot;</normalizedToken>
but differed by its lighter colour, shorter scapes and more feeble dorsal sculpture.
</paragraph>
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</treatment>
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<treatment ID-GBIF-Taxon="182234821" LSID="urn:lsid:plazi:treatment:3BECC0580A995D3D937CC3C8F79D6E17" httpUri="http://treatment.plazi.org/id/3BECC0580A995D3D937CC3C8F79D6E17" lastPageId="10" lastPageNumber="31" pageId="9" pageNumber="30">
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<paragraph pageId="9" pageNumber="30">
<taxonomicName LSID="3BECC058-0A99-5D3D-937C-C3C8F79D6E17" authority="Retezar, 2013" authorityName="Retezar" authorityYear="2013" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. adzinbai" order="Coleoptera" pageId="9" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="adzinbai">
Carabus (Archiplectes) besleticus adzinbai
<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
, 2013
</taxonomicName>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus subspecies constructed using discriminant analysis based on 8 parameters." pageId="9" pageNumber="30">Figs 4</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution map of Carabus (Archiplectes) satyrus, Carabus (Archiplectes) besleticus, and Carabus (Archiplectes) pseudopshuensis in Abkhazia." pageId="9" pageNumber="30">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="31" type="reference_group">
<paragraph pageId="10" pageNumber="31">
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus subsp. adzinbai" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Archiplectes" subSpecies="adzinbai">
<pageBreakToken pageId="10" pageNumber="31" start="start">Carabus</pageBreakToken>
(Archiplectes) satyrus adzinbai
</taxonomicName>
<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
, 2013: 2
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="31" type="comparative material examined">
<paragraph pageId="10" pageNumber="31">Comparative material examined.</paragraph>
<paragraph pageId="10" pageNumber="31">
15 specimens were examined (12 specimens measured, 1 male genitalia preparations studied): 1 male, 2 females, Abkhazia, Bzybian Mt. Range, N slope of Mt. Akibakhu (=Turetskaya shapka), 2000 m, alpine zone, 14.VI.-09.VIII.1986, leg. A. Koval (cZAM); 1 male, 10 females, Abkhazia, Bzybian Mt. Range, N slope of Mt. Akibakhu (=Turetskaya shapka), 2000 m, alpine zone, 7-10.VII.2010, leg. D. Fominykh, A. Bondarenko (cFDD, cSIA); 1 female, Abkhazia, Gudauta distr., S slope Bzybian Mt.R., NW Mt.Akugra,
<geoCoordinate degrees="43" direction="north" minutes="18" orientation="latitude" precision="925" value="43.3">43°18'N</geoCoordinate>
/
<geoCoordinate degrees="40" direction="east" minutes="43" orientation="longitude" precision="925" value="40.716667">40°43'E</geoCoordinate>
, h = 2130-1150 m, alpine zone, 08.08.2014, leg. I. Solodovnikov, E. Tatun (cSIA).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="31" type="description">
<paragraph pageId="10" pageNumber="31">Description.</paragraph>
<paragraph pageId="10" pageNumber="31">
Large form, males 31.0-32.0 mm, according to
<bibRefCitation author="Retezar, I" journalOrPublisher="Folia Entomologica Hungarica" pageId="23" pageNumber="44" pagination="1 - 6" refId="B9" refString="Retezar, I, 2013. Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae). Folia Entomologica Hungarica 74: 1 - 6" title="Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae)." volume="74" year="2013">
<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
(2013)
</bibRefCitation>
32-34 mm long, females 32.0-35.8 mm, according to
<bibRefCitation author="Retezar, I" journalOrPublisher="Folia Entomologica Hungarica" pageId="23" pageNumber="44" pagination="1 - 6" refId="B9" refString="Retezar, I, 2013. Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae). Folia Entomologica Hungarica 74: 1 - 6" title="Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae)." volume="74" year="2013">
<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
(2013)
</bibRefCitation>
33-37 mm long. Underside black, dorsum usually with bright metallic lustre, dark bronze, greenish-bronze, dark blue, reddish-bronze or black, pronotum often greenish; mandibles, palpi, antennae, and legs black. Habitus see
<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
, 2013.
</paragraph>
<paragraph pageId="10" pageNumber="31">
Head normal, frons coarsely, neck moderately wrinkled. Pronotum subquadrate, broadest in anterior one third, PW/PL = 1.14-1.42 (1.25), PW/PB = 1.0-1.26 (1.14), lateral sides of pronotum slightly sinuated before hind angles, the latter strongly protruding backwards and sidewards, pointed apically. Median groove distinct, disk moderately, basal foveae coarsely rugose. Elytra oblong-ovate, broadest behind their middle, EL/EW = 1.61-1.77 (1.69) in males and 1.64-1.85 (1.72) 1.73-1.98 in females. Elytral sculpture somewhat identical in males and females, forming coarse and precise series of links, primary interspaces elevated stronger than secondary ones, regularly interrupted by large foveae. Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric characteristic of males (n = 16) and females (n = 31) of Carabus (Archiplectes) satyrus and males (n = 83) and females (n = 91) of Carabus (Archiplectes) besleticus subspecies." httpUri="http://table.plazi.org/id/E3AA304075ED326595338A0A005BF64B" pageId="10" pageNumber="31" tableUuid="E3AA304075ED326595338A0A005BF64B">2</tableCitation>
lists the main morphometric measurements of one studied population.
</paragraph>
<paragraph pageId="10" pageNumber="31">
The shape of the male genitalia is practically identical to that of the other
<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
taxa.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="31" type="differential diagnosis">
<paragraph pageId="10" pageNumber="31">Differential diagnosis.</paragraph>
<paragraph pageId="10" pageNumber="31">
This form recently described by
<bibRefCitation author="Retezar, I" journalOrPublisher="Folia Entomologica Hungarica" pageId="23" pageNumber="44" pagination="1 - 6" refId="B9" refString="Retezar, I, 2013. Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae). Folia Entomologica Hungarica 74: 1 - 6" title="Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae)." volume="74" year="2013">
<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
(2013)
</bibRefCitation>
, resembles habitually some populations of
<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. mtsaranus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="mtsaranus">Carabus (Archiplectes) besleticus mtsaranus</taxonomicName>
, but unlike them represents somewhat different combination of features. However, the significant and stable difference is not still ascertained, accumulation of the further material from the upper forest belt will apparently make possible unequivocal precision of its taxonomic relationships with other neighboring populations.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="31" type="distribution">
<paragraph pageId="10" pageNumber="31">Distribution.</paragraph>
<paragraph pageId="10" pageNumber="31">Populates the alpine belt of Mt. Khipsta, Mt. Akugra, and Mt. Akibakhu at an altitude of 2000-2300 m.</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="31" type="habitat">
<paragraph pageId="10" pageNumber="31">Habitat.</paragraph>
<paragraph pageId="10" pageNumber="31">
Prefers alpine herb meadows. Activity of imago proceeds from May to August, beetles being active at melting of snow cover.
<taxonomicName authorityName="Zamotajlov &amp; Koval" authorityYear="1989" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Procechenochilus) gusevi" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="gusevi" subGenus="Procechenochilus">Carabus (Procechenochilus) gusevi</taxonomicName>
,
<taxonomicName authority="circassicus" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) circassicus subsp. circassicus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="circassicus" subGenus="Tribax" subSpecies="circassicus">Carabus (Tribax) circassicus circassicus</taxonomicName>
(natio
<taxonomicName class="Insecta" family="Carabidae" form="abasinus" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="form">
<emphasis italics="true" pageId="10" pageNumber="31">abasinus</emphasis>
</taxonomicName>
),
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) constantinovi subsp. otcharensis" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="constantinovi" subGenus="Tribax" subSpecies="otcharensis">Carabus (Tribax) constantinovi otcharensis</taxonomicName>
,
<taxonomicName baseAuthorityName="Ganglbauer" baseAuthorityYear="1889" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) agnatus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="agnatus" subGenus="Tribax">Carabus (Tribax) agnatus</taxonomicName>
,
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Lipaster) stjervalli subsp. humbolti" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="stjervalli" subGenus="Lipaster" subSpecies="humbolti">Carabus (Lipaster) stjervalli humbolti</taxonomicName>
Faldermann, 1835, and
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Pachycarabus) imitator subsp. katharinae" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="imitator" subGenus="Pachycarabus" subSpecies="katharinae">Carabus (Pachycarabus) imitator katharinae</taxonomicName>
Reitter, 1896 occur together with this subspecies.
</paragraph>
</subSubSection>
</treatment>
</document>