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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1067.72779" ID-GBIF-Dataset="39ded5c1-e041-4aa3-bf7e-5efb47e0aee2" ID-PMC="PMC8571249" ID-Pensoft-Pub="1313-2970-1067-57" ID-Pensoft-UUID="921145FDEC60583E831A6266908B4BCD" ID-PubMed="34759719" ID-ZooBank="8DB9C8FE9DF7438D92BA4B40E81F5DE1" ModsDocID="1313-2970-1067-57" checkinTime="1635566822117" checkinUser="pensoft" docAuthor="Kwanboon, Sedtawut, Sartori, Michel &amp; Boonsoong, Boonsatien" docDate="2021" docId="3BD887F49F025DB68BA8297F43999C37" docLanguage="en" docName="ZooKeys 1067: 57-82" docOrigin="ZooKeys 1067" docPubDate="2021-10-29" docSource="http://dx.doi.org/10.3897/zookeys.1067.72779" docTitle="Rhoenanthus (Potamanthindus) magnificus Ulmer 1920" docType="treatment" docVersion="4" id="921145FDEC60583E831A6266908B4BCD" lastPageNumber="57" masterDocId="921145FDEC60583E831A6266908B4BCD" masterDocTitle="Behningiidae and Potamanthidae (Insecta, Ephemeroptera) in Thailand" masterLastPageNumber="82" masterPageNumber="57" pageNumber="57" updateTime="1668151044797" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Behningiidae and Potamanthidae (Insecta, Ephemeroptera) in Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kwanboon, Sedtawut</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3858-9522</mods:nameIdentifier>
<mods:affiliation>Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sartori, Michel</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3397-3397</mods:nameIdentifier>
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Boonsoong, Boonsatien</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8166-0021</mods:nameIdentifier>
<mods:affiliation>Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">fscibtb@ku.ac.th</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-10-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1067</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>57</mods:start>
<mods:end>82</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1067.72779</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1067.72779</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1067-57</mods:identifier>
<mods:identifier type="ZooBank">8DB9C8FE9DF7438D92BA4B40E81F5DE1</mods:identifier>
<mods:identifier type="Pensoft-UUID">921145FDEC60583E831A6266908B4BCD</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="189635843" LSID="urn:lsid:plazi:treatment:3BD887F49F025DB68BA8297F43999C37" httpUri="http://treatment.plazi.org/id/3BD887F49F025DB68BA8297F43999C37" lastPageNumber="57" pageId="0" pageNumber="57">
<subSubSection pageId="0" pageNumber="57" type="nomenclature">
<paragraph pageId="0" pageNumber="57">
<taxonomicName LSID="3BD887F4-9F02-5DB6-8BA8-297F43999C37" authority="Ulmer, 1920" authorityName="Ulmer" authorityYear="1920" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus (Potamanthindus) magnificus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="magnificus" subGenus="Potamanthindus">Rhoenanthus (Potamanthindus) magnificus Ulmer, 1920</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="description">
<paragraph pageId="0" pageNumber="57">
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larval morphology (head and mandibular tusk) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D angle measurement of mandibular tusk E R. distafurcus Bae &amp; McCafferty, 1991 F R. speciosus Eaton, 1881." figureDoi="10.3897/zookeys.1067.72779.figure10" httpUri="https://binary.pensoft.net/fig/604591" pageId="0" pageNumber="57">Figures 10B, D</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larval morphology (foreleg) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D R. distafurcus Bae &amp; McCafferty, 1991 E R. speciosus Eaton, 1881. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1067.72779.figure11" httpUri="https://binary.pensoft.net/fig/604592" pageId="0" pageNumber="57">, 11B</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of larva of Rhoenanthus magnificus Ulmer, 1920." figureDoi="10.3897/zookeys.1067.72779.figure12" httpUri="https://binary.pensoft.net/fig/604593" pageId="0" pageNumber="57">, 12</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Distribution map of the family Potamanthidae in Thailand." figureDoi="10.3897/zookeys.1067.72779.figure14" httpUri="https://binary.pensoft.net/fig/604595" pageId="0" pageNumber="57">, 14</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. The COI phylogenetic construction based on the maximum likelihood (ML) analysis of family Potamanthidae in Thailand." figureDoi="10.3897/zookeys.1067.72779.figure15" httpUri="https://binary.pensoft.net/fig/604596" pageId="0" pageNumber="57">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="materials_examined">
<paragraph pageId="0" pageNumber="57">Materials examined.</paragraph>
<paragraph pageId="0" pageNumber="57">
<materialsCitation collectingDate="2016-03-11" collectorName="B. Boonsoong" country="Thailand" elevation="742" latitude="19.320301" location="Mae Na" longLatPrecision="1" longitude="98.89055" municipality="Chiang Dao" specimenCount="5" specimenCount-larva="5" stateProvince="Chiang Mai province">
<specimenCount type="larva">5 larvae</specimenCount>
,
<collectingCountry name="Thailand">Thailand</collectingCountry>
,
<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai province</collectingRegion>
,
<collectingMunicipality>Chiang Dao</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3BD887F49F025DB68BA8297F43999C37:8B484946B76B2CAB0CDAB672C419011E" country="Thailand" latitude="19.320301" longLatPrecision="1" longitude="98.89055" municipality="Chiang Dao" name="Mae Na" stateProvince="Chiang Mai province">Mae Na</location>
,
<geoCoordinate degrees="19" direction="north" minutes="19" orientation="latitude" precision="1" seconds="13.08" value="19.320301">19°19'13.08&quot;N</geoCoordinate>
,
<geoCoordinate degrees="98" direction="east" minutes="53" orientation="longitude" precision="1" seconds="25.98" value="98.89055">98°53'25.98&quot;E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.42" unit="m" value="742.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.42" unit="m" value="742.0">742 m</elevation>
</quantity>
,
<collectingDate value="2016-03-11">11.III.2016</collectingDate>
,
<collectorName>B. Boonsoong</collectorName>
leg. (ZMKU)
</materialsCitation>
;
<materialsCitation collectingDate="2016-03-20" collectorName="B. Boonsoong" country="Thailand" elevation="338" latitude="17.26014" location="Nam Thob" longLatPrecision="1" longitude="101.58136" municipality="Loel province" specimenCount="8" specimenCount-larva="8" stateProvince="Chiang Mai">
<specimenCount type="larva">8 larvae</specimenCount>
,
<collectingMunicipality>Loel province</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3BD887F49F025DB68BA8297F43999C37:884673183B64751CAA5FA1C4B4EA0089" country="Thailand" latitude="17.26014" longLatPrecision="1" longitude="101.58136" municipality="Loel province" name="Nam Thob" stateProvince="Chiang Mai">Nam Thob</location>
ranger station,
<geoCoordinate degrees="17" direction="north" minutes="15" orientation="latitude" precision="1" seconds="36.5" value="17.26014">17°15'36.5&quot;N</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="34" orientation="longitude" precision="1" seconds="52.9" value="101.58136">101°34'52.9&quot;E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.38" unit="m" value="338.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.38" unit="m" value="338.0">338 m</elevation>
</quantity>
,
<collectingDate value="2016-03-20">20.III.2016</collectingDate>
,
<collectorName>B. Boonsoong</collectorName>
leg. (ZMKU)
</materialsCitation>
;
<materialsCitation collectingDate="2020-11-28" collectorName="B. Boonsoong" country="Thailand" elevation="800" latitude="19.190498" location="Sapan" longLatPrecision="1" longitude="101.198975" municipality="Bo Kluea district" specimenCount="1" specimenCount-larva="1" stateProvince="Nan province">
<specimenCount type="larva">1 larva</specimenCount>
,
<collectingRegion country="Thailand" name="Nan">Nan province</collectingRegion>
,
<collectingMunicipality>Bo Kluea district</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3BD887F49F025DB68BA8297F43999C37:DC6F9B3BAB5628B69DB300830318FB3F" country="Thailand" latitude="19.190498" longLatPrecision="1" longitude="101.198975" municipality="Bo Kluea district" name="Sapan" stateProvince="Nan province">Sapan</location>
waterfall,
<geoCoordinate degrees="19" direction="north" minutes="11" orientation="latitude" precision="1" seconds="25.8" value="19.190498">19°11'25.8&quot;N</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="11" orientation="longitude" precision="1" seconds="56.3" value="101.198975">101°11'56.3&quot;E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<collectingDate value="2020-11-28">28.XI.2020</collectingDate>
,
<collectorName>B. Boonsoong</collectorName>
leg
</materialsCitation>
;
<materialsCitation collectingDate="2020-11-28" collectorName="S. Kwanboon" country="Thailand" latitude="19.152445" location="Lamer" longLatPrecision="1" longitude="101.154724" municipality="Bo Kluea district" specimenCount="2" specimenCount-larva="2" stateProvince="Nan province">
<specimenCount type="larva">2 larvae</specimenCount>
,
<collectingRegion country="Thailand" name="Nan">Nan province</collectingRegion>
,
<collectingMunicipality>Bo Kluea district</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3BD887F49F025DB68BA8297F43999C37:68FF07821D264C48EA139F8E43ACC69C" country="Thailand" latitude="19.152445" longLatPrecision="1" longitude="101.154724" municipality="Bo Kluea district" name="Lamer" stateProvince="Nan province">Lamer</location>
resort,
<geoCoordinate degrees="19" direction="north" minutes="09" orientation="latitude" precision="1" seconds="08.8" value="19.152445">19°09'08.8&quot;N</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="09" orientation="longitude" precision="1" seconds="17.0" value="101.154724">101°09'17.0&quot;E</geoCoordinate>
,
<collectingDate value="2020-11-28">28.XI.2020</collectingDate>
,
<collectorName>S. Kwanboon</collectorName>
leg
</materialsCitation>
;
<materialsCitation collectingDate="2019-11-26" collectorName="B. Boonsoong" country="Thailand" elevation="848" latitude="19.272943" location="Mae Nam Wa" longLatPrecision="1" longitude="101.18005" municipality="Bo Kluea district" specimenCount="3" specimenCount-larva="3" stateProvince="Nan province">
<specimenCount type="larva">3 larvae</specimenCount>
,
<collectingRegion country="Thailand" name="Nan">Nan province</collectingRegion>
,
<collectingMunicipality>Bo Kluea district</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3BD887F49F025DB68BA8297F43999C37:F2B9DD9546EBBEA9FF41827F29A4EEE0" country="Thailand" latitude="19.272943" longLatPrecision="1" longitude="101.18005" municipality="Bo Kluea district" name="Mae Nam Wa" stateProvince="Nan province">Mae Nam Wa</location>
stream,
<geoCoordinate degrees="19" direction="north" minutes="16" orientation="latitude" precision="1" seconds="22.6" value="19.272943">19°16'22.6&quot; N</geoCoordinate>
<geoCoordinate degrees="101" direction="east" minutes="10" orientation="longitude" precision="1" seconds="48.2" value="101.18005">101°10'48.2&quot; E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.48" unit="m" value="848.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.48" unit="m" value="848.0">848 m</elevation>
</quantity>
,
<collectingDate value="2019-11-26">26.XI.2019</collectingDate>
,
<collectorName>B. Boonsoong</collectorName>
leg
</materialsCitation>
;
<materialsCitation collectingDate="2021-03-06" collectorName="S. Kwanboon" country="Thailand" elevation="534" latitude="19.862806" location="Mae Kon" longLatPrecision="1" longitude="99.651306" municipality="Mueang district" specimenCount="7" specimenCount-larva="7" stateProvince="Chiang Rai province">
<specimenCount type="larva">7 larvae</specimenCount>
,
<collectingRegion country="Thailand" name="Chiang Rai">Chiang Rai province</collectingRegion>
,
<collectingMunicipality>Mueang district</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3BD887F49F025DB68BA8297F43999C37:7615167FBB0866F8529B1E16494314F3" country="Thailand" latitude="19.862806" longLatPrecision="1" longitude="99.651306" municipality="Mueang district" name="Mae Kon" stateProvince="Chiang Rai province">Mae Kon</location>
stream,
<geoCoordinate degrees="19" direction="north" minutes="51" orientation="latitude" precision="1" seconds="46.1" value="19.862806">19°51'46.1&quot;N</geoCoordinate>
,
<geoCoordinate degrees="99" direction="east" minutes="39" orientation="longitude" precision="1" seconds="04.7" value="99.651306">99°39'04.7&quot;E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.34" unit="m" value="534.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.34" unit="m" value="534.0">534 m</elevation>
</quantity>
,
<collectingDate value="2021-03-06">6.III.2021</collectingDate>
,
<collectorName>S. Kwanboon</collectorName>
leg
</materialsCitation>
;
<materialsCitation collectingDate="2021-03-07" collectorName="S. Kwanboon" country="Thailand" county="Mueang district" elevation="470" latitude="20.011612" location="Pong Phrabat" longLatPrecision="1" longitude="99.8042" municipality="Mae Kon" specimenCount="2" specimenCount-larva="2" stateProvince="Chiang Rai province">
<specimenCount type="larva">2 larvae</specimenCount>
,
<collectingRegion country="Thailand" name="Chiang Rai">Chiang Rai province</collectingRegion>
,
<collectingCounty>Mueang district</collectingCounty>
,
<collectingMunicipality>Mae Kon</collectingMunicipality>
stream,
<location LSID="urn:lsid:plazi:treatment:3BD887F49F025DB68BA8297F43999C37:1E00A3349AF6A575B0084FEF8950951A" country="Thailand" county="Mueang district" latitude="20.011612" longLatPrecision="1" longitude="99.8042" municipality="Mae Kon" name="Pong Phrabat" stateProvince="Chiang Rai province">Pong Phrabat</location>
waterfall,
<geoCoordinate degrees="20" direction="north" minutes="00" orientation="latitude" precision="1" seconds="41.8" value="20.011612">20°00'41.8&quot;N</geoCoordinate>
,
<geoCoordinate degrees="99" direction="east" minutes="48" orientation="longitude" precision="1" seconds="15.1" value="99.8042">99°48'15.1&quot;E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">470 m</elevation>
</quantity>
,
<collectingDate value="2021-03-07">7.III.2021</collectingDate>
,
<collectorName>S. Kwanboon</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="diagnosis">
<paragraph pageId="0" pageNumber="57">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="57">
The larvae of
<taxonomicName authorityName="Ulmer" authorityYear="1920" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus magnificus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="magnificus">
<emphasis italics="true" pageId="0" pageNumber="57">Rhoenanthus magnificus</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of larva of Rhoenanthus magnificus Ulmer, 1920." figureDoi="10.3897/zookeys.1067.72779.figure12" httpUri="https://binary.pensoft.net/fig/604593" pageId="0" pageNumber="57">12</figureCitation>
) can be distinguished from those of other
<taxonomicName authority="" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus (Potamanthindus)" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="subGenus" subGenus="Potamanthindus">Rhoenanthus (Potamanthindus)</taxonomicName>
species based on the following characteristics: i) large body size (18-21 mm), ii) mandibular tusks arched inward about 33-34° (angle measurement as shown in Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larval morphology (head and mandibular tusk) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D angle measurement of mandibular tusk E R. distafurcus Bae &amp; McCafferty, 1991 F R. speciosus Eaton, 1881." figureDoi="10.3897/zookeys.1067.72779.figure10" httpUri="https://binary.pensoft.net/fig/604591" pageId="0" pageNumber="57">10D</figureCitation>
), iii) length of the mandibular tusks ca 1.4
<normalizedToken originalValue="×">x</normalizedToken>
length of head, and iv) length of the foretibiae ca 1.5
<normalizedToken originalValue="×">x</normalizedToken>
length of the forefemora and about 2.9
<normalizedToken originalValue="×">x</normalizedToken>
length of the foretarsi (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larval morphology (foreleg) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D R. distafurcus Bae &amp; McCafferty, 1991 E R. speciosus Eaton, 1881. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1067.72779.figure11" httpUri="https://binary.pensoft.net/fig/604592" pageId="0" pageNumber="57">11B</figureCitation>
) (
<bibRefCitation author="Nguyen, VV" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="57" pagination="9 - 17" refId="B15" refString="Nguyen, VV, Bae, YJ, 2004. Descriptions of Rhoenanthus sapa, new species, and larval stage of R. magnificus Ulmer (Ephemeroptera: Potamanthidae) from Vietnam. Aquatic Insects 26: 9 - 17" title="Descriptions of Rhoenanthus sapa, new species, and larval stage of R. magnificus Ulmer (Ephemeroptera: Potamanthidae) from Vietnam." volume="26" year="2004">Nguyen and Bae 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="distribution">
<paragraph pageId="0" pageNumber="57">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="57">Chiang Mai, Chiang Rai, Loei, and Nan provinces.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="remarks">
<paragraph pageId="0" pageNumber="57">Remark.</paragraph>
<paragraph pageId="0" pageNumber="57">
The larva of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="R. magnificus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="magnificus">
<emphasis italics="true" pageId="0" pageNumber="57">R. magnificus</emphasis>
</taxonomicName>
was originally described by
<bibRefCitation author="Nguyen, VV" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="57" pagination="9 - 17" refId="B15" refString="Nguyen, VV, Bae, YJ, 2004. Descriptions of Rhoenanthus sapa, new species, and larval stage of R. magnificus Ulmer (Ephemeroptera: Potamanthidae) from Vietnam. Aquatic Insects 26: 9 - 17" title="Descriptions of Rhoenanthus sapa, new species, and larval stage of R. magnificus Ulmer (Ephemeroptera: Potamanthidae) from Vietnam." volume="26" year="2004">Nguyen and Bae (2004)</bibRefCitation>
from material collected in northern and central Vietnam. The species is known from southern China and Vietnam. In the present study, we found this species in streams of several provinces (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Distribution map of the family Potamanthidae in Thailand." figureDoi="10.3897/zookeys.1067.72779.figure14" httpUri="https://binary.pensoft.net/fig/604595" pageId="0" pageNumber="57">14</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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data/3B/D8/BA/3BD8BABDEF722A4BF0CED1583FF20FD1.xml Normal file
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.411.7258" ID-PMC="PMC4042817" ID-Pensoft-Pub="1313-2970-411-1" ID-Pensoft-UUID="F95EFFFAFFF4FFDC0944FF9D2104C057" ID-PubMed="24899854" ID-Zenodo-Dep="578282" ModsDocID="1313-2970-411-1" checkinTime="1451245864541" checkinUser="pensoft" docAuthor="Nantarat, Nattawadee, Sutcharit, Chirasak, Tongkerd, Piyoros, Ablett, Jonathan, Naggs, Fred &amp; Panha, Somsak" docDate="2014" docId="3BD8BABDEF722A4BF0CED1583FF20FD1" docLanguage="en" docName="ZooKeys 411: 1-56" docOrigin="ZooKeys 411" docPubDate="2014-05-23" docSource="http://dx.doi.org/10.3897/zookeys.411.7258" docTitle="Cyclophorus malayanus" docType="treatment" docVersion="5" id="F95EFFFAFFF4FFDC0944FF9D2104C057" lastPageNumber="16" masterDocId="F95EFFFAFFF4FFDC0944FF9D2104C057" masterDocTitle="An annotated catalogue of type specimens of the land snail genus Cyclophorus Monfort, 1810 (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London" masterLastPageNumber="56" masterPageNumber="1" pageNumber="16" updateTime="1668158533112" updateUser="ExternalLinkService">
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<mods:title>An annotated catalogue of type specimens of the land snail genus Cyclophorus Monfort, 1810 (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nantarat, Nattawadee</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Biological Sciences Program, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tongkerd, Piyoros</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ablett, Jonathan</mods:namePart>
<mods:affiliation>Division of Higher Invertebrates, Natural History Museums, London, SW 7 5 BD, United Kingdom</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Naggs, Fred</mods:namePart>
<mods:affiliation>Division of Higher Invertebrates, Natural History Museums, London, SW 7 5 BD, United Kingdom</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2014</mods:date>
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<mods:number>2014-05-23</mods:number>
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<mods:detail type="volume">
<mods:number>411</mods:number>
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<mods:start>1</mods:start>
<mods:end>56</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.411.7258</mods:url>
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<treatment ID-GBIF-Taxon="152053159" LSID="urn:lsid:plazi:treatment:3BD8BABDEF722A4BF0CED1583FF20FD1" httpUri="http://treatment.plazi.org/id/3BD8BABDEF722A4BF0CED1583FF20FD1" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection pageId="15" pageNumber="16" type="nomenclature">
<paragraph pageId="15" pageNumber="16">
<taxonomicName LSID="3BD8BABD-EF72-2A4B-F0CE-D1583FF20FD1" authority="(Benson, 1852)" baseAuthorityName="Benson" baseAuthorityYear="1852" class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Cyclophorus malayanus" order="Architaenioglossa" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="malayanus">Cyclophorus malayanus (Benson, 1852)</taxonomicName>
</paragraph>
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<subSubSection pageId="15" pageNumber="16" type="reference_group">
<paragraph pageId="15" pageNumber="16">
<taxonomicName authorityName="Benson" authorityYear="1852" class="Gastropoda" family="Assimineidae" genus="Cyclostoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="malayanum">Cyclostoma malayanum</taxonomicName>
Benson, 1852: 269.
</paragraph>
<paragraph pageId="15" pageNumber="16">
<taxonomicName baseAuthorityName="Benson" baseAuthorityYear="1852" class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="malayanus">Cyclophorus malayanus</taxonomicName>
-
<bibRefCitation author="Reeve, L" journalOrPublisher="Conchologia Iconica: Illustrations of the shells of molluscous animals" pageId="31" pageNumber="32" refId="B58" refString="Reeve, L, 1861. . Conchologia Iconica: Illustrations of the shells of molluscous animals Volume 13" volume="Volume 13" year="1861">Reeve 1861</bibRefCitation>
: sp. 2.
<bibRefCitation author="Kobelt, W" journalOrPublisher="Das Tierreich" pageId="30" pageNumber="31" pagination="1 - 662" refId="B31" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich 16: 1 - 662" title="Cyclophoridae." volume="16" year="1902">Kobelt 1902</bibRefCitation>
: 130.
<bibRefCitation author="Kobelt, W" journalOrPublisher="Abbildungen nach der Natur mit Beschreibungen. Dritte Abteilung. Cyclophoridae I" pageId="30" pageNumber="31" pagination="401 - 608" refId="B32" refString="Kobelt, W, 1907-1908 1907 1908. . Abbildungen nach der Natur mit Beschreibungen. Dritte Abteilung. Cyclophoridae I 1 (19): 401 - 608" volume="1" year="1907 - 1908">Kobelt 1908</bibRefCitation>
: 658.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="type locality">
<paragraph pageId="15" pageNumber="16">Type locality.</paragraph>
<paragraph pageId="15" pageNumber="16">In montibus vallibusque Insularum Penang et Lancavi, necnon in Peninsula Malayana [In the mountains, the valleys of the islands of Penang and Langkawi, as well as the Peninsula Malaysia]</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="type material">
<paragraph pageId="15" pageNumber="16">Type material.</paragraph>
<paragraph pageId="15" pageNumber="16">
Syntypes NHMUK 20130089 (2 shells;
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Types of Cyclophorus species. A Cyclophorus malayanus (Benson, 1852), syntype NHMUK 20130089 B, C Cyclophorus monachus (Morelet, 1866) B lectotype NHMUK 1893.2.4.499, and C paralectotype NHMUK 1893.2.4.500." httpUri="https://binary.pensoft.net/fig/29288" pageId="15" pageNumber="16">Fig. 13A; D</figureCitation>
=43.5 mm, H=32.1 mm, W=5; D=47.0 mm, H=37.3 mm, W=5)
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="remarks">
<paragraph pageId="15" pageNumber="16">Remarks.</paragraph>
<paragraph pageId="15" pageNumber="16">
The original description did not include an illustration but
<bibRefCitation author="Reeve, L" journalOrPublisher="Conchologia Iconica: Illustrations of the shells of molluscous animals" pageId="31" pageNumber="32" refId="B58" refString="Reeve, L, 1861. . Conchologia Iconica: Illustrations of the shells of molluscous animals Volume 13" volume="Volume 13" year="1861">Reeve (1861</bibRefCitation>
: sp. 2) subsequently re-published the description with illustrations of a specimen from the Cuming collection. The NHM collection contains a lot of three
<normalizedToken originalValue="specimens’">specimens'</normalizedToken>
from the Cuming collection labelled &quot;Malay Peninsula&quot;. A label reads &quot;Mr. Benson has also sent me his
<taxonomicName authorityName="Backer" authorityYear="1924" genus="Malayanum" lsidName="" pageId="15" pageNumber="16" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="16">Malayanum</emphasis>
</taxonomicName>
and the true
<emphasis italics="true" pageId="15" pageNumber="16">volvulus</emphasis>
for comparison
<normalizedToken originalValue="….”">....&quot;</normalizedToken>
. Two of the three shells are close to the measurements and description in the original description and the label, presumably written by Cuming, indicates that these are the three specimens sent to Cuming by Benson, of which two without opercula are syntypes. The third specimen with an operculum being what Benson considered to be '
<emphasis italics="true" pageId="15" pageNumber="16">volvulus</emphasis>
' and not a member of the type series of '
<emphasis italics="true" pageId="15" pageNumber="16">malayanus</emphasis>
'. There is a further lot from &quot;Pulo Penang&quot; housed in the University Museum of Zoology Cambridge with original Benson labels including one specimen labelled by Benson as
<normalizedToken originalValue="‘Type’">'Type'</normalizedToken>
.
</paragraph>
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</treatment>
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