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<document ID-DOI="http://dx.doi.org/10.3897/dez.70.108096" ID-Pensoft-Pub="1860-1324-2-387" ID-Pensoft-UUID="7AFF3005E27B5133A05FB99D6920E718" ID-ZooBank="95E6EB7594FA4DA79DE4D3324C9ED1CE" ModsDocID="1860-1324-70-2-387" checkinTime="1698104555921" checkinUser="pensoft" docAuthor="Zheng, Xuhongyi &amp; Zhou, Changfa" docDate="2023" docId="3B9F6DEDAB1C5562A8B39CB77D71F50A" docLanguage="en" docName="DeutEntomolZeit 70(2): 387-401" docOrigin="Deutsche Entomologische Zeitschrift 70 (2)" docPubDate="2023-10-23" docSource="http://dx.doi.org/10.3897/dez.70.108096" docTitle="Deuterophlebia pachychaeta Zheng &amp; Zhou 2023, sp. nov." docType="treatment" docUuid="99503019-F78C-49B5-B602-8CE1D53BD96A" docUuidSource="ZooBank" docVersion="1" id="7AFF3005E27B5133A05FB99D6920E718" lastPageNumber="387" masterDocId="7AFF3005E27B5133A05FB99D6920E718" masterDocTitle="Two new species of Deuterophlebia Edwards, 1922 from Southwestern China (Diptera, Deuterophlebiidae)" masterLastPageNumber="401" masterPageNumber="387" pageNumber="387" updateTime="1698104555921" updateUser="pensoft">
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<mods:title id="8ED373A90149DEF3F661782A2CA9AF7E">Two new species of Deuterophlebia Edwards, 1922 from Southwestern China (Diptera, Deuterophlebiidae)</mods:title>
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<mods:namePart id="0AD611D883C1ED30F331EC35B62C0153">Zheng, Xuhongyi</mods:namePart>
<mods:affiliation id="77C0454C770C8E131A3663DD3E0E7767">The Key Laboratory of Jiangsu Biodiversity and Biotechnology, College of Life Sciences, Nanjing Normal University, Nanjing 210023, China</mods:affiliation>
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<mods:namePart id="B56995C50E972D705D19F38FD85DB764">Zhou, Changfa</mods:namePart>
<mods:affiliation id="51FA95DE639E4373031752F79E427D68">The Key Laboratory of Jiangsu Biodiversity and Biotechnology, College of Life Sciences, Nanjing Normal University, Nanjing 210023, China</mods:affiliation>
<mods:nameIdentifier id="6C13B37867CECCECE150056F0EA0BE85" type="email">zhouchangfa@njnu.edu.cn</mods:nameIdentifier>
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<mods:date id="73D824829DABD637EE00677209A84DE5">2023</mods:date>
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<treatment id="3B9F6DEDAB1C5562A8B39CB77D71F50A" LSID="urn:lsid:zoobank.org:act:99503019-F78C-49B5-B602-8CE1D53BD96A" httpUri="http://treatment.plazi.org/id/3B9F6DEDAB1C5562A8B39CB77D71F50A" lastPageNumber="387" pageId="0" pageNumber="387">
<subSubSection id="EBA268409DA14CA7C088EC392077CF96" pageId="0" pageNumber="387" type="nomenclature">
<paragraph id="893A62178DDFD0A8EE231CDF0A52B25E" pageId="0" pageNumber="387">
<taxonomicName id="5BF71C6DF40FBFCCC7BC50724E5AA17D" LSID="https://zoobank.org/99503019-F78C-49B5-B602-8CE1D53BD96A" authority="Zheng &amp; Zhou, 2023" authorityName="Zheng &amp; Zhou" authorityYear="2023" class="Insecta" family="Deuterophlebiidae" genus="Deuterophlebia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deuterophlebia pachychaeta" order="Diptera" pageId="0" pageNumber="387" phylum="Arthropoda" rank="species" species="pachychaeta" status="sp. nov.">Deuterophlebia pachychaeta</taxonomicName>
<taxonomicNameLabel id="9F11A0E7362D0C92A149F4477E7AFE99" pageId="0" pageNumber="387">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B585577ECAFC99ED17ADD785AAF905EA" pageId="0" pageNumber="387" type="description">
<paragraph id="A096CBD87DE003102095186AFF68EB3A" pageId="0" pageNumber="387">Description.</paragraph>
<paragraph id="FCA8299AEEBCF616FB4305F6B86C7C85" pageId="0" pageNumber="387">
<emphasis id="DDD0E0C00F66CE2866B33167F40D7BA1" bold="true" pageId="0" pageNumber="387">Male adults.</emphasis>
Body length ca. 3.0 mm (n = 2), uniformly brownish black. Head brownish black, flattened and nearly trapezoidal, width ca. 0.50 mm (Fig.
<figureCitation id="7CB6B17816BB21FA6B406208AD7C309C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6A</figureCitation>
). Head densely covered with microtrichia. Median clypeal lobe slightly convex, semicircle shaped with around 20 sharp setae (Fig.
<figureCitation id="ADE91E37718D7F831D366F4F5090070E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6A</figureCitation>
). Mouthparts in form of an invaginated tubule, oral region depressed (Fig.
<figureCitation id="BC7B9BA78D779A6555E9F0F8469920DE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6A, B</figureCitation>
). Edges of oral region (or mouth opening) ridged, forming a protruded angle on inner side of each eye (Fig.
<figureCitation id="B41968531EBEAEACC8092B2AB756C961" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6A, B</figureCitation>
). Ventral ridge of oral region convex medially and forming a blunt mental tooth (Fig.
<figureCitation id="C2089033AFCC9DEDE34E9AE37A3A7BDA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6A, B</figureCitation>
). Postgena and oral region with microtrichia but more sparse than other regions, a pair of tentorial pits present on each side of oral region (Fig.
<figureCitation id="17AE427BDDEB58DDB2BC0D32DD45C0B0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6A, B</figureCitation>
). Compound eyes glabrous, width ca. 0.13 mm, distance between eyes ca. 0.30 mm (Fig.
<figureCitation id="A4954A4BA44F7BE5400D39F478247CE6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6A</figureCitation>
).
</paragraph>
<caption id="9DD72C4CE144D05EEAD1082E4C8B823F" doi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387" start="Figure 6" startId="F6">
<paragraph id="110B2D247B96A15E0E07712FBB576FB2" pageId="0" pageNumber="387">
<emphasis id="F099286F0FEE9B62B281D9F88A7E3DE3" bold="true" pageId="0" pageNumber="387">Figure 6.</emphasis>
Male adult of
<taxonomicName id="04AC5CAA3EF886683D898513C6E26E6C" lsidName="D. pachychaeta" pageId="0" pageNumber="387" rank="species" species="pachychaeta">
<emphasis id="5FD131364E1AF62399825CDDC3FC3574" italics="true" pageId="0" pageNumber="387">D. pachychaeta</emphasis>
</taxonomicName>
sp. nov.:
<emphasis id="A949094C3D3788735B8CEF72D57C3E91" bold="true" pageId="0" pageNumber="387">A.</emphasis>
Head (ventral view);
<emphasis id="DD511E6CB80D1ED761905CA836B63E73" bold="true" pageId="0" pageNumber="387">B.</emphasis>
Oral region (ventral view);
<emphasis id="D4B8BA0F3804CE6EAD6E06675D887F93" bold="true" pageId="0" pageNumber="387">C.</emphasis>
Flagellomeres;
<emphasis id="1E8A989438107F5EE44CB066BA27CFA7" bold="true" pageId="0" pageNumber="387">D.</emphasis>
Foreleg;
<emphasis id="6B81498627CE06AA40A28E4C4246254E" bold="true" pageId="0" pageNumber="387">E.</emphasis>
Midleg;
<emphasis id="420C0F2E659CE186A2512D42D498C3CE" bold="true" pageId="0" pageNumber="387">F.</emphasis>
Hindleg;
<emphasis id="39DA9E25F3E6CC1E48AED402B82E0554" bold="true" pageId="0" pageNumber="387">G.</emphasis>
Terminalia (dorsal view). Scale bars: 0.1 mm (
<emphasis id="9D8D4751E15ACA0A15BEE7C3B1265309" bold="true" pageId="0" pageNumber="387">A, C, G</emphasis>
); 0.05 mm (
<emphasis id="BC8C40D8D9040ACA193F72D6351DAA90" bold="true" pageId="0" pageNumber="387">B</emphasis>
); 0.5 mm (
<emphasis id="CA12F50834833114A93DFFF14368B0B9" bold="true" pageId="0" pageNumber="387">D-F</emphasis>
).
</paragraph>
</caption>
<paragraph id="E1EA783637DAE7D77D3F9B6C4610B986" pageId="0" pageNumber="387">
Antennae ca. 13.0 mm (n=5) (Fig.
<figureCitation id="AE828914D3965C30185AF1EF29F65DC7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6A, C</figureCitation>
). Scape oval shaped, pedicel globular, both scape and pedicel covered with microtrichia (Fig.
<figureCitation id="373CC90D673727B54DAA32568DBB2EE5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6A, C</figureCitation>
). Flagella four segmented, flagellomeres I-III slender cylindrical, each with a subapical tubercle on front margin and bearing 8-10 digitiform setae (Fig.
<figureCitation id="91282C8E832C72D6D20A8EC68043D9B3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6C</figureCitation>
). Flagellomere IV flattened and elongated, with curved hair-like setae on the anterior side of basal half, apical half generally glabrous but bearing 4-5 clusters of curved hair-like setae, apex slightly expanded with some curved hair-like setae (Fig.
<figureCitation id="879AFD5E96C09B3F9D0BF746C8E9914B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6C</figureCitation>
). Antennal ratio ca. 4.0: 2.0: 5.0: 3.0: 3.0: 287.0, flagellomere IV about 17
<normalizedToken id="99621381FFE082DFF59C91A0A6CAF5FC" originalValue="×">x</normalizedToken>
combined length of five basal antennal articles or about 4
<normalizedToken id="40CBA4F8D4DA0FBF69A67551D602CB6F" originalValue="×">x</normalizedToken>
body length (Fig.
<figureCitation id="D52F2B6CCBD4766B526EEE461CCBEB5A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6C</figureCitation>
).
</paragraph>
<paragraph id="314712AE0F3B5CE10CBD25EBF5B46F59" pageId="0" pageNumber="387">
Thorax brownish black, densely covered with microtrichia. Mesonotum strongly expanded. Wings ca. 5.5 mm, shape, venation and other features similar to
<taxonomicName id="B672B0C95D6BE6C720D2416EFA97C309" lsidName="D. pseudopoda" pageId="0" pageNumber="387" rank="species" species="pseudopoda">
<emphasis id="490B4EF16A0E86C9A8BFEB17F897FA93" italics="true" pageId="0" pageNumber="387">D. pseudopoda</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="47DB9121C923208134BC6CBCB5406C9C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male adult of D. pseudopoda sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure1" httpUri="https://binary.pensoft.net/fig/925433" pageId="0" pageNumber="387">1</figureCitation>
).
</paragraph>
<paragraph id="7C814513155616DE8A7B687F41726CBC" pageId="0" pageNumber="387">
Legs brownish black, slender, three pairs sharing similar chaetotaxy exhibiting four types of setae: (1) microtrichia, densely covered on all segments; (2) sharp macrotrichia, sparsely on dorsal margin of femora and tibiae; (3) long capitate setae, on ventral side of tarsomere I-IV of each leg, distal half of ventral edge of all tibiae, surrounding the top of fore- and midtibiae, and also densely arranged radially on each empodium; (4) digitiform setae, 1-3 pairs for each tarsomere (Fig.
<figureCitation id="DB67D4FC035F0DF86CC473739F2108BB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6D-F</figureCitation>
). In foreleg, femur: tibia: tarsus = 11.0: 17.0: 16.0; femur slightly flattened, tibia slender cylindrical and gradually broader apically; tarsomere I: II: III: IV: V = 8.5: 2.0: 2.0: 1.5: 0.8, tarsomeres I-IV cylindrical, tarsomere V conical; empodium shell-shaped, length subequal to tarsomere V; claw slender tapered, shorter than empodium (Fig.
<figureCitation id="F0C6D44E0696A9DB9AF3F025BB3E8B17" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male adult of D. pseudopoda sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. hindleg; G. Terminalia (dorsal view). Abbreviations: cp, median clypeal lobe; pd, pedicel; fl, flagellomere; or, oral region; mt, mental tooth; pg, postgena; co, coxa; tr, trochanter; fe, femur; tb, tibia; ts, tarsomere; em, empodium; dp, dorsal plate; gc, gonocoxite; gs, gonostylus; ad, aedeagus. Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure2" httpUri="https://binary.pensoft.net/fig/925434" pageId="0" pageNumber="387">2B</figureCitation>
) (Fig.
<figureCitation id="13845BEE9A63812090C46A561AA14D8A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6D</figureCitation>
). Midleg shortest among all legs, features generally similar to foreleg, femur: tibia: tarsus = 10.0: 12.5: 14.0, tarsomere I: II: III: IV: V = 8.5: 2.0: 2.0: 1.5: 0.8 (Fig.
<figureCitation id="390556A5EC620A24345BAE5299ABCB49" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6E</figureCitation>
). In hindleg, femur: tibia: tarsus of hindleg = 14.0: 17.0: 11.0, tarsomere I: II: III: IV: V = 3.0: 2.0: 2.0: 1.5: 0.8 (Fig.
<figureCitation id="5EE7401BA278E24871D147DBB2AFE806" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6F</figureCitation>
).
</paragraph>
<paragraph id="05A1E7FC4ADC7EB3C3D2325F7EC2C03F" pageId="0" pageNumber="387">
Abdomen brownish black, densely covered with microtrichia, nine segmented, tapering posteriorly. Segment VIII in form of a short chitin ring, sternite IX almost glabrous, connected with dorsal plate and gonocoxite (Fig.
<figureCitation id="EA299D2A093D1242500B3A8EBDBD97A6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6G</figureCitation>
). Gonocoxite with posterior projection ca. 1/2 length of gonostylus (Fig.
<figureCitation id="0817FF1F818C78998D7FCF85339D82CE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6G</figureCitation>
). Gonostylus subequal dorsal plate, flattened, slender oval shaped, flexor surface with numerous curved sharp setae (Fig.
<figureCitation id="F0D995E5D625428C6200C813928D3946" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6G</figureCitation>
). Dorsal plate parallel-sided, posterior margin slightly depressed with a median cleft and some stout setae on margin (Fig.
<figureCitation id="AFF93B39CC4594464366EEDCFA3C15E3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6G</figureCitation>
). Aedeagus in form of a smooth tube, length subequal to gonostylus, longer than dorsal plate (Fig.
<figureCitation id="C78E51CCCF1261F5303BFE58CE54AADC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6G</figureCitation>
).
</paragraph>
<paragraph id="4B50C1B596ACBBD342F713F5473277F5" pageId="0" pageNumber="387">
<emphasis id="C3E8B380ADFD46D23A0B87DEA93140EC" bold="true" pageId="0" pageNumber="387">Female adult.</emphasis>
Body length ca. 2.3 mm (n = 1). Besides sexual differences, generally similar to the males except following features (Fig.
<figureCitation id="4F7A0D79EF48B3558E4C851F802A3AC3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7A-D</figureCitation>
).
</paragraph>
<caption id="CF795091272B982CD61FC8489C35BA48" doi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387" start="Figure 7" startId="F7">
<paragraph id="6783C91454B1FFA8987FB43595F9C6C5" pageId="0" pageNumber="387">
<emphasis id="72E13749ABA0F3659493C58CAA544281" bold="true" pageId="0" pageNumber="387">Figure 7.</emphasis>
Female adult of
<taxonomicName id="C54920A718CBBA1B659114758C30F5C2" lsidName="D. pachychaeta" pageId="0" pageNumber="387" rank="species" species="pachychaeta">
<emphasis id="64B3C936EB2E1C5F2972658FC2FA4C05" italics="true" pageId="0" pageNumber="387">D. pachychaeta</emphasis>
</taxonomicName>
sp. nov.:
<emphasis id="D23E7ED62848C25793E1187C9CE7C909" bold="true" pageId="0" pageNumber="387">A.</emphasis>
Head (ventral view);
<emphasis id="980479B6DB73BE5682BDB6787F85E905" bold="true" pageId="0" pageNumber="387">B.</emphasis>
Foreleg;
<emphasis id="129615EA132401AD412B830E1E3F2FD3" bold="true" pageId="0" pageNumber="387">C.</emphasis>
Midleg;
<emphasis id="28D75DC333D7CB67AD5A3623D5DD6396" bold="true" pageId="0" pageNumber="387">D.</emphasis>
Hindleg. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="FAEC788A432BC64EDD8AEE22CCCED838" pageId="0" pageNumber="387">
Head width ca. 0.40 mm (Fig.
<figureCitation id="4C1E67200E971797CFF2FEF353A745AB" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7A</figureCitation>
). Median clypeal lobe slightly convex with ca. 20 setae (Fig.
<figureCitation id="6BF76A02D7D54E287FDF3E375B798354" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7A</figureCitation>
). Oral region located near anterior margin of head, postgena and oral region with microtrichia but more sparse than other regions (Fig.
<figureCitation id="BC456DF855A2538F6E395E44C268EAD3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7A</figureCitation>
). Compound eyes more prominent than males (Fig.
<figureCitation id="3CED673334E9B42E6B09D7A8280D5CDA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7A</figureCitation>
). Antenna ca. 0.3 mm. Scape slender oval-shaped, pedicel globular, both scape and pedicel covered with microtrichia with several sharp setae (Fig.
<figureCitation id="F9B414920B771045D4C8572ED482E2CA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7A</figureCitation>
). Flagellomere I slender cylindrical, flagellomeres II-III slender oval shaped, flagellomere IV dripping shaped (Fig.
<figureCitation id="D1723F5BDAC94BCE9179A4B3FE39B035" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7A</figureCitation>
). Flagellomeres I-IV bearing 2, 4, 10 and 2 digitiform setae apically respectively, flagellomere III-IV with 1 and 4 sharp setae respectively (Fig.
<figureCitation id="D1502DA9D69D1F58BCE57D3C63EFD79B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7A</figureCitation>
). Antennal ratio = 7.0: 4.0: 10.0: 5.0: 5.0: 4.0 (Fig.
<figureCitation id="49FCA83F204B00666678E923C1575DE5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7A</figureCitation>
).
</paragraph>
<paragraph id="D800A6A88D21F3E3C08A2D481E597881" pageId="0" pageNumber="387">
Legs sharing similar chaetotaxy and exhibiting three types of setae, chaetotaxy similar to males but without capitate setae (Fig.
<figureCitation id="D3A86EB35B2CC1C3D228949E5A04AA24" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7B-D</figureCitation>
). In all legs, coxae about twice longer and broader than trochanters, femora slightly flattened, tibiae and tarsomeres slender cylindrical (Fig.
<figureCitation id="0018B6C7ED3A2CDF1398F57E9B7C228D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7B-D</figureCitation>
). In foreleg and midleg, femur: tibia: tarsus = 1.0: 2.0: 1.5, tarsomere I: II: III: IV: V = 1.2: 0.8: 0.8: 0.8: 2.4 (Fig.
<figureCitation id="DC882491440D469D5D31049884953DD1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7B, C</figureCitation>
). In hindleg, femur: tibia: tarsus = 1.0: 1.7: 1.1, tarsomere I: II: III: IV: V = 1.0: 0.8: 0.8: 0.8: 2.4 (Fig.
<figureCitation id="ADC45411EFECB20137433D7A1AE24010" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7D</figureCitation>
). Claws of all legs similar, paired, stout and curved, with a blunt protrusion in the middle (Fig.
<figureCitation id="31626D0AA315AE1FE92666E85D5C7298" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7B-D</figureCitation>
). Empodium in form of a long and hairy spine, subequal to the length of claw (Fig.
<figureCitation id="EF5A2F4962D20C249CB07F9765F597CC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7B-D</figureCitation>
).
</paragraph>
<paragraph id="49E048609214C9678A9E2E01C7BC4367" pageId="0" pageNumber="387">
<emphasis id="BEB0B7B4B5EFE22E6BC5A457935C04E4" bold="true" pageId="0" pageNumber="387">Male pupae.</emphasis>
Pupae flattened oval shaped, length 2.8 mm (n = 2), width 2.0 mm (Fig.
<figureCitation id="483C0004E148161C59FF47D30D1C7A22" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8A, B</figureCitation>
). Dorsal integument dark brown, divided into 11 segments (Fig.
<figureCitation id="7057E7CB2A9BC381E33550F2A8B85094" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8A, B</figureCitation>
).
</paragraph>
<caption id="B5D2DBCBD07EA8EA06B33A0E57AB26AD" doi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387" start="Figure 8" startId="F8">
<paragraph id="715894176BACB2F34FFF084A50A6F397" pageId="0" pageNumber="387">
<emphasis id="224FCB4E67E7D74B82DAF876B4D712C3" bold="true" pageId="0" pageNumber="387">Figure 8.</emphasis>
Pupae of
<taxonomicName id="8878463B02C096BF5936FB564FEEBC48" lsidName="D. pachychaeta" pageId="0" pageNumber="387" rank="species" species="pachychaeta">
<emphasis id="D7C0732EFDA4D5199067CB9553375DCD" italics="true" pageId="0" pageNumber="387">D. pachychaeta</emphasis>
</taxonomicName>
sp. nov.:
<emphasis id="13A037427BD1E54EB64936E7558F239A" bold="true" pageId="0" pageNumber="387">A.</emphasis>
Male pupa (dorsal view);
<emphasis id="44259E4AE013B44AEE8F8300D0CB7028" bold="true" pageId="0" pageNumber="387">B.</emphasis>
Male pupa (ventral view);
<emphasis id="71260CBD1097D17564B81F6AA231888D" bold="true" pageId="0" pageNumber="387">C.</emphasis>
Female pupa (dorsal view);
<emphasis id="43F72EF08343035AAAFE1D7C9114CB5E" bold="true" pageId="0" pageNumber="387">D.</emphasis>
Female pupa (ventral view). Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="FE46F65D51CF822D27270173EC23E5ED" pageId="0" pageNumber="387">
Prothorax fused with mesothorax, forming a conical segment with a median suture (Fig.
<figureCitation id="DEC938A00E7E7AC5F38F271F1DE81382" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8A, B</figureCitation>
). Mesothoracic lateral margins each with a pair of sharp spines and a gill (Figs
<figureCitation id="A3F1DCF2A03D977D1B8BFCD26F55ADEC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8A, B</figureCitation>
,
<figureCitation id="0F2A3B3627BCCBCA924087CB5764E962" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male pupa of D. pachychaeta sp. nov.: A. Thoracic spine (dorsal view); B. Gill (ventral view). Scale bar: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure9" httpUri="https://binary.pensoft.net/fig/925441" pageId="0" pageNumber="387">9A, B</figureCitation>
). Spines ca. 0.4 mm, expanded basally, slightly curved, dark brown, originated from a round base (Fig.
<figureCitation id="6E2FD206E4168BE45A220BA3C0350618" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male pupa of D. pachychaeta sp. nov.: A. Thoracic spine (dorsal view); B. Gill (ventral view). Scale bar: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure9" httpUri="https://binary.pensoft.net/fig/925441" pageId="0" pageNumber="387">9A</figureCitation>
). Ventral gills light to dark brown, length subequal to the dorsal spines, each including four filaments: posterior filament single and short, pointing backward; anterior three gill filaments on a common base, similar in shape, slender and twisted; second and third filaments fused at base (Fig.
<figureCitation id="7B7D03546B24CDC812B69C849B3D7A0E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male pupa of D. pachychaeta sp. nov.: A. Thoracic spine (dorsal view); B. Gill (ventral view). Scale bar: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure9" httpUri="https://binary.pensoft.net/fig/925441" pageId="0" pageNumber="387">9B</figureCitation>
). Metathorax completely surrounded by mesothorax and first abdominal segment (Fig.
<figureCitation id="59F519914E9EAD6A8954DA1EF75B95F5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8A, B</figureCitation>
).
</paragraph>
<paragraph id="0B28716B749151F1F9080FAF8CF8F034" pageId="0" pageNumber="387">
Abdominal segment I and II similar, each with a pair of anterolateral projections, each projection pointing forward and bearing ca. 18 spines (Fig.
<figureCitation id="82EF4692FEDCBE355CBB7B8D5D847160" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8A, B</figureCitation>
). Lateral margins of segments VI and VII with several spines on both dorsal and ventral sides. Segment VIII shield-shaped, surrounded by segments VII and IX (Fig.
<figureCitation id="66384C219AB28B0096508F0094ED35C1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8A, B</figureCitation>
).
</paragraph>
<paragraph id="963AEB86765D25A334688D6C54E897AC" pageId="0" pageNumber="387">
Adult structures visible on ventral side (Fig.
<figureCitation id="E474BF1EE8B711F84FE2100873299EC8" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8B</figureCitation>
). Head present below mesothorax; antennal sheaths in form of a large elliptic ring, surrounding body 2.0 times (Fig.
<figureCitation id="AF1B9FCA0B12DAE6515A99A97CE0C08F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8B</figureCitation>
). Leg sacs extended to posterior end of antennal ring, strongly expanded apically. Abdominal segments III-V each with a pair of black adhesive discs (Fig.
<figureCitation id="B0F067402395D0E26E3D9465831C56A4" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8B</figureCitation>
).
</paragraph>
<paragraph id="029E5ADDC33872285428C9054CE16FC4" pageId="0" pageNumber="387">
<emphasis id="27070D2EE7E15C8EC8B29E998D1EA86C" bold="true" pageId="0" pageNumber="387">Female pupae.</emphasis>
Length ca. 2.8 mm (n = 1), width ca. 2.0 mm (Fig.
<figureCitation id="FF356688485FF530FADD369EA3C98884" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8C, D</figureCitation>
). Dorsal morphology similar to male except for smaller mesothorax (Fig.
<figureCitation id="ADCF6F522D37865FE2B9C88542CA7BA0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8C, D</figureCitation>
). Gender can be identified through the absence of antennal ring, apex of female leg sheaths not expanded (Fig.
<figureCitation id="C54068E8DE7D708B530D759A0886D9AF" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pupae of D. pachychaeta sp. nov.: A. Male pupa (dorsal view); B. Male pupa (ventral view); C. Female pupa (dorsal view); D. Female pupa (ventral view). Scale bars: 1.0 mm." figureDoi="10.3897/dez.70.108096.figure8" httpUri="https://binary.pensoft.net/fig/925440" pageId="0" pageNumber="387">8C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9700BC33BC3C2F1450002C87C3299E1C" pageId="0" pageNumber="387" type="materials_examined">
<paragraph id="1D94475D8C185E39D51B19FA99075BC8" pageId="0" pageNumber="387">Material examined.</paragraph>
<paragraph id="C6529E3B6434A39674289239C3B0B455" pageId="0" pageNumber="387">
<materialsCitation id="6A9402B7A40B120CC213E1103CFC11A9" collectingDate="2023-02-04" collectorName="Xuhongyi Zheng" country="China" county="Gongshan County" elevation="1470" latitude="27.837267" location="Dulongjiang Township" longLatPrecision="1" longitude="98.33172" municipality="Nujiang Lisu Autonomous Prefecture" specimenCount="1" stateProvince="Yunnan Province" typeStatus="Holotype">
<emphasis id="52EBE35EE08656DF1DB3558BE48C19AF" bold="true" italics="true" pageId="0" pageNumber="387">
<typeStatus id="79085DF180C261850C4B483DA699754E">Holotype</typeStatus>
</emphasis>
: male adult,
<collectingCountry id="B3BFA0257364D39639C64C1B8E0E01C0" name="China">China</collectingCountry>
:
<collectingRegion id="3879D51572BCD5236DA4839DEFD03302" country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="C0DE41988E1EC425A6F94B70F45D0B99">Nujiang Lisu Autonomous Prefecture</collectingMunicipality>
,
<collectingCounty id="5DC754150FE8BE85D8E89867E094AE1D">Gongshan County</collectingCounty>
,
<location id="1815FC3BDACF46F09E0DC8B3D36434BD" LSID="urn:lsid:plazi:treatment:3B9F6DEDAB1C5562A8B39CB77D71F50A:1815FC3BDACF46F09E0DC8B3D36434BD" country="China" county="Gongshan County" latitude="27.837267" longLatPrecision="1" longitude="98.33172" municipality="Nujiang Lisu Autonomous Prefecture" name="Dulongjiang Township" stateProvince="Yunnan Province">Dulongjiang Township</location>
,
<location id="5DDDE34C4824BF7C89DF8666F3F99087" LSID="urn:lsid:plazi:treatment:3B9F6DEDAB1C5562A8B39CB77D71F50A:5DDDE34C4824BF7C89DF8666F3F99087" country="China" county="Gongshan County" latitude="27.837267" longLatPrecision="1" longitude="98.33172" municipality="Nujiang Lisu Autonomous Prefecture" name="Dulongjiang River" stateProvince="Yunnan Province">Dulongjiang River</location>
,
<geoCoordinate id="5280A873383480CD036B2543FBD9C4FD" degrees="27" direction="north" minutes="50" orientation="latitude" precision="1" seconds="14.16" value="27.837267">27°50'14.16&quot;N</geoCoordinate>
,
<geoCoordinate id="02619405163CF2DAC809794D1839E17C" degrees="98" direction="east" minutes="19" orientation="longitude" precision="1" seconds="54.2" value="98.33172">98°19'54.2&quot;E</geoCoordinate>
,
<elevation id="1003B4B653A06B138EF6111ABE618388" metricMagnitude="3" metricUnit="m" metricValue="1.47" unit="m" value="1470.0">
<quantity id="4D867696A23B191648B90A7D58D4E04D" metricMagnitude="3" metricUnit="m" metricValue="1.47" unit="m" value="1470.0">1470 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="340C189E1274127E5F77485F73323E6F" value="2023-02-04">4.II.2023</collectingDate>
,
<collectorName id="C5606D54810D7BF784E9D89172A5741C">Xuhongyi Zheng</collectorName>
leg.
</materialsCitation>
<emphasis id="2F7BBF0A839BBB8167E0115EAB16B3EA" bold="true" italics="true" pageId="0" pageNumber="387">
<typeStatus id="3A6B1ED772B9AFC3996DB2674FCBC6AD">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="4CB681C002244F957CA553738D0529CD" type="male">1 male</specimenCount>
adult,
<materialsCitation id="7029A55E8ED494FDAAF8AA7FAB3D374B" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Xuhongyi Zheng" country="China" county="Gongshan County" elevation="1470" latitude="27.837267" location="Dulongjiang Township" longLatPrecision="1" longitude="98.33172" municipality="Nujiang Lisu Autonomous Prefecture" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="7AC2C6B87004D342AB6BC66D4619F271" type="male">2 male</specimenCount>
pupae, same locality and data as
<typeStatus id="1F541FDE5EE25D08B8A9597ED66182B2">holotype</typeStatus>
</materialsCitation>
;
<materialsCitation id="FEE242EF7DF5914BCF9B6EA59C4D70D3" collectingDate="2022-07-03" collectorName="Xuhongyi Zheng, Zhenxing Ma, Zhiteng Chen, Pengxu Mu." country="China" county="Fugong County" elevation="1265" latitude="26.954811" location="Nujiang Lisu Autonomous Prefecture" longLatPrecision="1" longitude="98.867516" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount id="8F363683A95E4B4E214102042657BFD6" type="female">1 female</specimenCount>
adult dissected from a pupa,
<collectingRegion id="8F0EE60BB270F13BFA6F0583C457EED2" country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<location id="A75BF0681D9724BE2ECE4D9752D5B1CA" LSID="urn:lsid:plazi:treatment:3B9F6DEDAB1C5562A8B39CB77D71F50A:A75BF0681D9724BE2ECE4D9752D5B1CA" country="China" county="Fugong County" latitude="26.954811" longLatPrecision="1" longitude="98.867516" name="Nujiang Lisu Autonomous Prefecture" stateProvince="Yunnan Province">Nujiang Lisu Autonomous Prefecture</location>
,
<collectingCounty id="2BCF7396D1D60495C6F069CC87A03C2C">Fugong County</collectingCounty>
,
<geoCoordinate id="F11F72F020C9A895FC4337CA650F7AE9" degrees="26" direction="north" minutes="57" orientation="latitude" precision="1" seconds="17.32" value="26.954811">26°57'17.32&quot;N</geoCoordinate>
,
<geoCoordinate id="5300B9EA03965E733CF0E8FCAB676391" degrees="98" direction="east" minutes="52" orientation="longitude" precision="1" seconds="3.04" value="98.867516">98°52'3.04&quot;E</geoCoordinate>
,
<elevation id="763C7E2FE13F75F0BB95645EC8DA405F" metricMagnitude="3" metricUnit="m" metricValue="1.265" unit="m" value="1265.0">
<quantity id="D6A1D1152C7CBB05F93C89650D10287F" metricMagnitude="3" metricUnit="m" metricValue="1.265" unit="m" value="1265.0">1265 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="5B4EFEC9E5EF67C585F72CC8910D3319" value="2022-07-03">3. VII. 2022</collectingDate>
, leg.
<collectorName id="DBA0A49DFEB0147E782749CA72D236D6">Xuhongyi Zheng</collectorName>
,
<collectorName id="07C09F08EC78383F2790F9BA34A137FF">Zhenxing Ma</collectorName>
,
<collectorName id="9F76A83DB599A900418BF2467137A532">Zhiteng Chen</collectorName>
&amp;
<collectorName id="9BD6C4CD96AB1DEE3D48D87B64111C73">Pengxu Mu.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C62872E5856B8848165E7C57DEA14316" pageId="0" pageNumber="387" type="diagnosis">
<paragraph id="FAF838D6032246AF81F1B5DC6BAED4DD" pageId="0" pageNumber="387">Diagnosis.</paragraph>
<paragraph id="0552543D316822D4C894CBE9DF76AC84" pageId="0" pageNumber="387">
Male adults of
<taxonomicName id="337282F9CEE43F21A4E0BFE5C25F2C9E" authorityName="Zheng &amp; Zhou" authorityYear="2023" class="Insecta" family="Deuterophlebiidae" genus="Deuterophlebia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Deuterophlebia pachychaeta" order="Diptera" pageId="0" pageNumber="387" phylum="Arthropoda" rank="species" species="pachychaeta">
<emphasis id="5C78C87ECF7AB2ADC8B63F70C0C3AE0B" italics="true" pageId="0" pageNumber="387">Deuterophlebia pachychaeta</emphasis>
</taxonomicName>
sp. nov. can be identified by their head shape and chaetotaxy of antennae (Fig.
<figureCitation id="053094BD3426CF9B490F43D0CED9ED5B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6A, C</figureCitation>
). Following the keys of
<bibRefCitation id="68005426677A7042BF2BEF7C5769A694" DOI="https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" author="Courtney, GW" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="387" pagination="1 - 24" refId="B6" refString="Courtney, GW, 1994. Revision of Palaearctic mountain midges (Diptera: Deuterophlebiidae), with phylogenetic and biogeographic analyses of world species. Systematic Entomology 19 (1): 1 - 24, DOI: https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" title="Revision of Palaearctic mountain midges (Diptera: Deuterophlebiidae), with phylogenetic and biogeographic analyses of world species." url="https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" volume="19" year="1994">Courtney (1994)</bibRefCitation>
and
<bibRefCitation id="40FE5041C9161C0748C767B1086FF788" DOI="https://doi.org/10.3390/insects13070593" author="Zheng, XHY" journalOrPublisher="Insects" pageId="0" pageNumber="387" pagination=": 593" refId="B21" refString="Zheng, XHY, Chen, ZT, Mu, PX, Ma, ZX, Zhou, CF, 2022. Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae). Insects 13 (7): 593, DOI: https://doi.org/10.3390/insects13070593" title="Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae)." url="https://doi.org/10.3390/insects13070593" volume="13" year="2022">Zheng et al. (2022)</bibRefCitation>
, males of this species, which are similar to
<taxonomicName id="0731FFE13C5B5ABB492B7FE2C35876E5" lsidName="D. mirabilis" pageId="0" pageNumber="387" rank="species" species="mirabilis">
<emphasis id="6433F14C78BAFBA312C97368114CC86A" italics="true" pageId="0" pageNumber="387">D. mirabilis</emphasis>
</taxonomicName>
Edwards, 1922,
<taxonomicName id="155DE4CAD55851C06071ABC763EBE968" lsidName="D. blepharis" pageId="0" pageNumber="387" rank="species" species="blepharis">
<emphasis id="912D52FE95E1369902AE0C67FDF54A5F" italics="true" pageId="0" pageNumber="387">D. blepharis</emphasis>
</taxonomicName>
Courtney, 1994 and
<taxonomicName id="B363B8BAEAEC646C9DF3BA969EDFFE89" lsidName="D. sinensis" pageId="0" pageNumber="387" rank="species" species="sinensis">
<emphasis id="471BF5491094737926F0E69B1389471F" italics="true" pageId="0" pageNumber="387">D. sinensis</emphasis>
</taxonomicName>
<bibRefCitation id="290A086B17D81257E4328757A1CAEEA0" DOI="https://doi.org/10.3390/insects13070593" author="Zheng, XHY" journalOrPublisher="Insects" pageId="0" pageNumber="387" pagination=": 593" refId="B21" refString="Zheng, XHY, Chen, ZT, Mu, PX, Ma, ZX, Zhou, CF, 2022. Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae). Insects 13 (7): 593, DOI: https://doi.org/10.3390/insects13070593" title="Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae)." url="https://doi.org/10.3390/insects13070593" volume="13" year="2022">Zheng et al., 2022</bibRefCitation>
, can be separated from other Asian species by their relatively long antennae (longer than 8.0 mm), and postgena covered with microtrichia (Fig.
<figureCitation id="7660CFB7912DD4C7E9B3C2D75679FC2C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6B</figureCitation>
). Chaetotaxy on flagellomeres can be used to differentiate those four similar species: in the new species, there were ca. 8-10 digitiform setae on each of flagellomeres I-III (Fig.
<figureCitation id="609F7E5EDDE544081453EF911BAA87EE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Oral region (ventral view); C. Flagellomeres; D. Foreleg; E. Midleg; F. Hindleg; G. Terminalia (dorsal view). Scale bars: 0.1 mm (A, C, G); 0.05 mm (B); 0.5 mm (D-F)." figureDoi="10.3897/dez.70.108096.figure6" httpUri="https://binary.pensoft.net/fig/925438" pageId="0" pageNumber="387">6A, C</figureCitation>
) (
<bibRefCitation id="06E08EFF35582F8DA0752396928071C6" DOI="https://doi.org/10.1080/00222939000770071" author="Courtney, GW" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="387" pagination="81 - 118" refId="B4" refString="Courtney, GW, 1990. Revision of Nearctic mountain midges (Diptera: Deuterophlebiidae). Journal of Natural History 24 (1): 81 - 118, DOI: https://doi.org/10.1080/00222939000770071" title="Revision of Nearctic mountain midges (Diptera: Deuterophlebiidae)." url="https://doi.org/10.1080/00222939000770071" volume="24" year="1990">Courtney 1990</bibRefCitation>
,
<bibRefCitation id="5001261B898ED7586D86ABA998BC2E39" DOI="https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" author="Courtney, GW" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="387" pagination="1 - 24" refId="B6" refString="Courtney, GW, 1994. Revision of Palaearctic mountain midges (Diptera: Deuterophlebiidae), with phylogenetic and biogeographic analyses of world species. Systematic Entomology 19 (1): 1 - 24, DOI: https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" title="Revision of Palaearctic mountain midges (Diptera: Deuterophlebiidae), with phylogenetic and biogeographic analyses of world species." url="https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" volume="19" year="1994">1994</bibRefCitation>
;
<bibRefCitation id="E29279F19336584DD2EF5008E15136C1" DOI="https://doi.org/10.3390/insects13070593" author="Zheng, XHY" journalOrPublisher="Insects" pageId="0" pageNumber="387" pagination=": 593" refId="B21" refString="Zheng, XHY, Chen, ZT, Mu, PX, Ma, ZX, Zhou, CF, 2022. Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae). Insects 13 (7): 593, DOI: https://doi.org/10.3390/insects13070593" title="Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae)." url="https://doi.org/10.3390/insects13070593" volume="13" year="2022">Zheng et al. 2022</bibRefCitation>
).
</paragraph>
<paragraph id="A7F765BFBD80B327733B6F291F2960E3" pageId="0" pageNumber="387">
Female adults of
<taxonomicName id="826D25A5D0ED40A4C87CFC9D94C633B7" lsidName="D. pachychaeta" pageId="0" pageNumber="387" rank="species" species="pachychaeta">
<emphasis id="C49CFD0406E0F87F6C052B1BECCF9BBC" italics="true" pageId="0" pageNumber="387">D. pachychaeta</emphasis>
</taxonomicName>
sp. nov. can be recognized through a combination of the smooth median clypeal lobe, chaetotaxy of antennae, and shape of flagella (Fig.
<figureCitation id="68004221DB3D0EE86983BB4C99147802" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7A</figureCitation>
). Its smooth and convex median clypeal lobe is similar to
<taxonomicName id="F01654E28567FE48CFC5380705E48237" lsidName="D. mirabilis" pageId="0" pageNumber="387" rank="species" species="mirabilis">
<emphasis id="6888A91389FBBD579E270C83FFC5E168" italics="true" pageId="0" pageNumber="387">D. mirabilis</emphasis>
</taxonomicName>
Edwards, 1922,
<taxonomicName id="A66CF92B2958C5C57CA58E2EFB9C07A2" lsidName="D. blepharis" pageId="0" pageNumber="387" rank="species" species="blepharis">
<emphasis id="3DC6E34A6A0A4577592D1D2AC72AB337" italics="true" pageId="0" pageNumber="387">D. blepharis</emphasis>
</taxonomicName>
Courtney, 1994,
<taxonomicName id="3656419A7113B3E29122E1FB70442EB7" lsidName="D. sinensis" pageId="0" pageNumber="387" rank="species" species="sinensis">
<emphasis id="7F3E172DED2DD2E210CD2F29271494B4" italics="true" pageId="0" pageNumber="387">D. sinensis</emphasis>
</taxonomicName>
<bibRefCitation id="17124FC348B73B3FC0EADA1C0A67282C" DOI="https://doi.org/10.3390/insects13070593" author="Zheng, XHY" journalOrPublisher="Insects" pageId="0" pageNumber="387" pagination=": 593" refId="B21" refString="Zheng, XHY, Chen, ZT, Mu, PX, Ma, ZX, Zhou, CF, 2022. Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae). Insects 13 (7): 593, DOI: https://doi.org/10.3390/insects13070593" title="Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae)." url="https://doi.org/10.3390/insects13070593" volume="13" year="2022">Zheng et al., 2022</bibRefCitation>
,
<taxonomicName id="6A8DD976CDC0757C931C57FF8F9E1CA1" lsidName="D. yunnanensis" pageId="0" pageNumber="387" rank="species" species="yunnanensis">
<emphasis id="4399CEDEE6A31457DE002953445D8A7A" italics="true" pageId="0" pageNumber="387">D. yunnanensis</emphasis>
</taxonomicName>
<bibRefCitation id="7412BB1A6C72027EA1FB5520A7D72F6B" DOI="https://doi.org/10.3390/insects13070593" author="Zheng, XHY" journalOrPublisher="Insects" pageId="0" pageNumber="387" pagination=": 593" refId="B21" refString="Zheng, XHY, Chen, ZT, Mu, PX, Ma, ZX, Zhou, CF, 2022. Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae). Insects 13 (7): 593, DOI: https://doi.org/10.3390/insects13070593" title="Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae)." url="https://doi.org/10.3390/insects13070593" volume="13" year="2022">Zheng et al., 2022</bibRefCitation>
and
<taxonomicName id="5F44901E27F6AF35FFED7D75830A0F76" lsidName="D. wuyiensis" pageId="0" pageNumber="387" rank="species" species="wuyiensis">
<emphasis id="3C90D456EA7723FF839D7CB6CD8A824A" italics="true" pageId="0" pageNumber="387">D. wuyiensis</emphasis>
</taxonomicName>
<bibRefCitation id="69B5D33758B0663EB4AFDC8F936CC2AF" DOI="https://doi.org/10.3390/insects13070593" author="Zheng, XHY" journalOrPublisher="Insects" pageId="0" pageNumber="387" pagination=": 593" refId="B21" refString="Zheng, XHY, Chen, ZT, Mu, PX, Ma, ZX, Zhou, CF, 2022. Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae). Insects 13 (7): 593, DOI: https://doi.org/10.3390/insects13070593" title="Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae)." url="https://doi.org/10.3390/insects13070593" volume="13" year="2022">Zheng et al., 2022</bibRefCitation>
, but can be differentiated from them by its antenna: flagellomeres I-IV bearing ca. 2, 4, 10 and 2 digitiform setae respectively, flagellomere IV with additionally 4 sharp setae; antennal ratio = 7.0: 4.0: 10.0: 5.0: 5.0: 4.0 (Fig.
<figureCitation id="AEBB6BBDD0D00D6931D2434E1F89B0F3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female adult of D. pachychaeta sp. nov.: A. Head (ventral view); B. Foreleg; C. Midleg; D. Hindleg. Scale bars: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure7" httpUri="https://binary.pensoft.net/fig/925439" pageId="0" pageNumber="387">7A</figureCitation>
) (
<bibRefCitation id="A986948CCD964C69E575EFA5619C4AF6" DOI="https://doi.org/10.1080/00222939000770071" author="Courtney, GW" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="387" pagination="81 - 118" refId="B4" refString="Courtney, GW, 1990. Revision of Nearctic mountain midges (Diptera: Deuterophlebiidae). Journal of Natural History 24 (1): 81 - 118, DOI: https://doi.org/10.1080/00222939000770071" title="Revision of Nearctic mountain midges (Diptera: Deuterophlebiidae)." url="https://doi.org/10.1080/00222939000770071" volume="24" year="1990">Courtney 1990</bibRefCitation>
,
<bibRefCitation id="71989B933BA2759AD4CD90CAD2FE1720" DOI="https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" author="Courtney, GW" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="387" pagination="1 - 24" refId="B6" refString="Courtney, GW, 1994. Revision of Palaearctic mountain midges (Diptera: Deuterophlebiidae), with phylogenetic and biogeographic analyses of world species. Systematic Entomology 19 (1): 1 - 24, DOI: https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" title="Revision of Palaearctic mountain midges (Diptera: Deuterophlebiidae), with phylogenetic and biogeographic analyses of world species." url="https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" volume="19" year="1994">1994</bibRefCitation>
;
<bibRefCitation id="535D398BEEC5E2EDCA7067B76AE0B1CC" DOI="https://doi.org/10.3390/insects13070593" author="Zheng, XHY" journalOrPublisher="Insects" pageId="0" pageNumber="387" pagination=": 593" refId="B21" refString="Zheng, XHY, Chen, ZT, Mu, PX, Ma, ZX, Zhou, CF, 2022. Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae). Insects 13 (7): 593, DOI: https://doi.org/10.3390/insects13070593" title="Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae)." url="https://doi.org/10.3390/insects13070593" volume="13" year="2022">Zheng et al. 2022</bibRefCitation>
).
</paragraph>
<paragraph id="15FB8BCC5AEFE7D1D37C724076B88CD1" pageId="0" pageNumber="387">
Pupae can be identified by their mesothoracic spines: there are two spines on each side, expanded basally (Fig.
<figureCitation id="8CA595485775A925AC8D401BE2A6B37D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male pupa of D. pachychaeta sp. nov.: A. Thoracic spine (dorsal view); B. Gill (ventral view). Scale bar: 0.1 mm." figureDoi="10.3897/dez.70.108096.figure9" httpUri="https://binary.pensoft.net/fig/925441" pageId="0" pageNumber="387">9A, B</figureCitation>
). Compared with the 15 species with a clear pupal stage (
<bibRefCitation id="93D94FC85C37EE2D9B90AB109EE4A325" DOI="https://doi.org/10.1080/00222939000770071" author="Courtney, GW" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="387" pagination="81 - 118" refId="B4" refString="Courtney, GW, 1990. Revision of Nearctic mountain midges (Diptera: Deuterophlebiidae). Journal of Natural History 24 (1): 81 - 118, DOI: https://doi.org/10.1080/00222939000770071" title="Revision of Nearctic mountain midges (Diptera: Deuterophlebiidae)." url="https://doi.org/10.1080/00222939000770071" volume="24" year="1990">Courtney 1990</bibRefCitation>
,
<bibRefCitation id="7E3498DF5D56A9AB0AB2D1EA989C7889" DOI="https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" author="Courtney, GW" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="387" pagination="1 - 24" refId="B6" refString="Courtney, GW, 1994. Revision of Palaearctic mountain midges (Diptera: Deuterophlebiidae), with phylogenetic and biogeographic analyses of world species. Systematic Entomology 19 (1): 1 - 24, DOI: https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" title="Revision of Palaearctic mountain midges (Diptera: Deuterophlebiidae), with phylogenetic and biogeographic analyses of world species." url="https://doi.org/10.1111/j.1365-3113.1994.tb00576.x" volume="19" year="1994">1994</bibRefCitation>
;
<bibRefCitation id="FFC59017876DEE94B80363CA6DC4AAAF" DOI="https://doi.org/10.3390/insects13070593" author="Zheng, XHY" journalOrPublisher="Insects" pageId="0" pageNumber="387" pagination=": 593" refId="B21" refString="Zheng, XHY, Chen, ZT, Mu, PX, Ma, ZX, Zhou, CF, 2022. Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae). Insects 13 (7): 593, DOI: https://doi.org/10.3390/insects13070593" title="Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae)." url="https://doi.org/10.3390/insects13070593" volume="13" year="2022">Zheng et al. 2022</bibRefCitation>
), only
<taxonomicName id="E60C38AAD7E6B1113CDB2CA27252527F" lsidName="D. acutirhina" pageId="0" pageNumber="387" rank="species" species="acutirhina">
<emphasis id="DA0C4E5BA3A24020748980039CCD4BF4" italics="true" pageId="0" pageNumber="387">D. acutirhina</emphasis>
</taxonomicName>
<bibRefCitation id="F1DC0E593AE626A205F5868D0DC18B88" DOI="https://doi.org/10.3390/insects13070593" author="Zheng, XHY" journalOrPublisher="Insects" pageId="0" pageNumber="387" pagination=": 593" refId="B21" refString="Zheng, XHY, Chen, ZT, Mu, PX, Ma, ZX, Zhou, CF, 2022. Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae). Insects 13 (7): 593, DOI: https://doi.org/10.3390/insects13070593" title="Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae)." url="https://doi.org/10.3390/insects13070593" volume="13" year="2022">Zheng et al., 2022</bibRefCitation>
has similar basally expanded spines, but less expanded than in the new species. They can also be distinguished by the absence of abdominal dots (
<bibRefCitation id="3E84FD5978D616B6EC9A790C70AD30DE" DOI="https://doi.org/10.3390/insects13070593" author="Zheng, XHY" journalOrPublisher="Insects" pageId="0" pageNumber="387" pagination=": 593" refId="B21" refString="Zheng, XHY, Chen, ZT, Mu, PX, Ma, ZX, Zhou, CF, 2022. Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae). Insects 13 (7): 593, DOI: https://doi.org/10.3390/insects13070593" title="Descriptions and barcoding of five new Chinese Deuterophlebia species revealing this genus in both Holarctic and Oriental realms (Diptera: Deuterophlebiidae)." url="https://doi.org/10.3390/insects13070593" volume="13" year="2022">Zheng et al. 2022</bibRefCitation>
).
</paragraph>
<caption id="8C736FA32DC524741C8DA66AB1709734" doi="10.3897/dez.70.108096.figure9" httpUri="https://binary.pensoft.net/fig/925441" pageId="0" pageNumber="387" start="Figure 9" startId="F9">
<paragraph id="590B3202028B3E1234A887BE1BF1658E" pageId="0" pageNumber="387">
<emphasis id="7D2930E28C9CDF590248A7BE4EB3ED9E" bold="true" pageId="0" pageNumber="387">Figure 9.</emphasis>
Male pupa of
<taxonomicName id="560E411C8483B7981F4BA17BABEDC466" lsidName="D. pachychaeta" pageId="0" pageNumber="387" rank="species" species="pachychaeta">
<emphasis id="9CA6BFB536E7C75A676F38E95BE5A257" italics="true" pageId="0" pageNumber="387">D. pachychaeta</emphasis>
</taxonomicName>
sp. nov.:
<emphasis id="7F9ABEC0676F293BC9F88062C5CF42B5" bold="true" pageId="0" pageNumber="387">A.</emphasis>
Thoracic spine (dorsal view);
<emphasis id="F519140F4A95B6C9166296FBB72E2A22" bold="true" pageId="0" pageNumber="387">B.</emphasis>
Gill (ventral view). Scale bar: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="CBFC18884A0F1391C2525B8A461D8D22" pageId="0" pageNumber="387" type="etymology">
<paragraph id="70C30730018621E38B4A04DFF455E775" pageId="0" pageNumber="387">Etymology.</paragraph>
<paragraph id="D9F0D1FE51F1CFC8D51B5EB4F681D5D1" pageId="0" pageNumber="387">
The
<normalizedToken id="E3239530FEE8E385C1D49783138F544A" originalValue="specific">specific</normalizedToken>
epithet &quot;
<emphasis id="52FE4F666E50838E7547A42A43FDE0D6" italics="true" pageId="0" pageNumber="387">pachychaeta</emphasis>
&quot; is a combination of the words
<emphasis id="CAAC925D8EF575649C51CE6880170556" italics="true" pageId="0" pageNumber="387">pachy</emphasis>
and
<emphasis id="B7B051B2CB9FAAA6B8D26853C6BE07C2" italics="true" pageId="0" pageNumber="387">chaeta</emphasis>
, which refers to its thickened spines on pupal mesothorax.
</paragraph>
</subSubSection>
<subSubSection id="22493DD7A9DA6BD41F5556AD07ABC8DD" pageId="0" pageNumber="387" type="distribution">
<paragraph id="0455268F62677A69D942EA0D245BC1F1" pageId="0" pageNumber="387">Distribution.</paragraph>
<paragraph id="FCDBDC0EBD96FFFC3C6DFE8A76838726" pageId="0" pageNumber="387">China (Yunnan Province).</paragraph>
</subSubSection>
</treatment>
</document>