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<document id="2B757EBE762A37C9DC6D7AC56C56AF80" ID-DOI="10.5281/zenodo.212553" ID-GBIF-Dataset="7e49d608-5e2b-4987-b853-64678deae6c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="212553" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460536598469" checkinUser="plazi" docAuthor="Stekolnikov, Alexandr A., Carranza, Salvador &amp; Gomez-Diaz, Elena" docDate="2012" docId="3B330041FFE8FFFD56988FCE9262D641" docLanguage="en" docName="zt03499p080.pdf" docOrigin="Zootaxa 3499" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Arabapolonia Stekolnikov, Carranza &amp; Gomez-Diaz, 2012, gen. nov." docType="treatment" docVersion="8" lastPageNumber="76" masterDocId="C70A7839FFE9FFFF560F89719062D251" masterDocTitle="A new genus and species of Apoloniinae (Acari: Trombiculidae) from Oman" masterLastPageNumber="80" masterPageNumber="74" pageNumber="75" updateTime="1698316509788" updateUser="plazi">
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<mods:title id="B67178B6A5E6354D8BE2FF56E08AE08A">A new genus and species of Apoloniinae (Acari: Trombiculidae) from Oman</mods:title>
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<mods:namePart id="AB72EEA1785C5A20A6C81D50CBE69051">Stekolnikov, Alexandr A.</mods:namePart>
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<mods:namePart id="BE5DAB3E9EF546A36F2ABC18D2336703">Carranza, Salvador</mods:namePart>
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<mods:namePart id="C0AB002FBEF6F87B9CEB0513A2BC1BEA">Gomez-Diaz, Elena</mods:namePart>
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<mods:date id="56673821B65DA088C16B6992ABB3E6C0">2012</mods:date>
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<treatment id="3B330041FFE8FFFD56988FCE9262D641" ID-DOI="http://doi.org/10.5281/zenodo.6168254" ID-GBIF-Taxon="119527718" ID-Zenodo-Dep="6168254" LSID="urn:lsid:zoobank.org:act:FA60E7D8-2A5A-4F28-ADCD-18A474D41229" httpUri="http://treatment.plazi.org/id/3B330041FFE8FFFD56988FCE9262D641" lastPageId="2" lastPageNumber="76" pageId="1" pageNumber="75">
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<paragraph id="B325B157FFE8FFFE56988FCE91D0D488" blockId="1.[151,434,1727,1753]" box="[151,434,1727,1753]" pageId="1" pageNumber="75">
<heading id="E86D063BFFE8FFFE56988FCE91D0D488" bold="true" box="[151,434,1727,1753]" fontSize="11" level="1" pageId="1" pageNumber="75" reason="1">
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<taxonomicName id="749ACAD4FFE8FFFE56988FCE915FD488" box="[151,317,1727,1753]" class="Arachnida" family="Trombiculidae" genus="Arabapolonia" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="81EE6D45FFE8FFFE56988FCE915FD488" bold="true" box="[151,317,1727,1753]" italics="true" pageId="1" pageNumber="75">Arabapolonia</emphasis>
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<taxonomicNameLabel id="9ADDD03EFFE8FFFE574B8FCE91D0D488" box="[324,434,1727,1753]" pageId="1" pageNumber="75" rank="genus">gen. nov.</taxonomicNameLabel>
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<subSubSection id="FB80E2DCFFE8FFFE56988E769349D54D" box="[151,811,1798,1821]" pageId="1" pageNumber="75" type="materials_examined">
<paragraph id="B325B157FFE8FFFE56988E769349D54D" blockId="1.[151,811,1798,1821]" box="[151,811,1798,1821]" pageId="1" pageNumber="75">
<typeStatus id="6C210FF5FFE8FFFE56988E7690AAD54D" box="[151,200,1799,1820]" pageId="1" pageNumber="75">Type</typeStatus>
species:
<taxonomicName id="749ACAD4FFE8FFFE57298E769246D54C" box="[294,548,1799,1821]" class="Arachnida" family="Trombiculidae" genus="Arabapolonia" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="75" phylum="Arthropoda" rank="species" species="omanensis" status="sp. nov.">
<emphasis id="81EE6D45FFE8FFFE57298E769246D54C" box="[294,548,1799,1821]" italics="true" pageId="1" pageNumber="75">Arabapolonia omanensis</emphasis>
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<emphasis id="81EE6D45FFE8FFFE54258E7692E3D54D" bold="true" box="[554,641,1799,1820]" pageId="1" pageNumber="75">
<taxonomicNameLabel id="9ADDD03EFFE8FFFE54258E76921FD54D" box="[554,637,1799,1820]" pageId="1" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
designated here.
</paragraph>
</subSubSection>
<subSubSection id="FB80E2DCFFE8FFFD56988E389418D3D6" lastPageId="2" lastPageNumber="76" pageId="1" pageNumber="75" type="diagnosis">
<paragraph id="B325B157FFE8FFFD56988E389161D335" blockId="1.[151,1436,1865,2033]" lastBlockId="2.[151,1436,151,1040]" lastPageId="2" lastPageNumber="76" pageId="1" pageNumber="75">
<emphasis id="81EE6D45FFE8FFFE56988E389173D533" bold="true" box="[151,273,1865,1890]" pageId="1" pageNumber="75">Diagnosis.</emphasis>
SIF = 6BS-N(b)-1-2110.0000; fsp = 7.7.7; fCx = 1.0.1; fSt = 0.2.4; fPp = B/B/BBB. Cheliceral blade with 1 inconspicuous denticle near its apex; galeal seta nude or having 1 small cilium; palpal claw single; all palpal setae branched; palpal tarsus with 6 non-specialized setae, nude subterminala and tarsala. Tracheae present; spiracles not found. Scutum with projected anterior margin, without nasus, posterior scutal margin slightly prominent, angulated or rounded; two anteromedian setae, two anterolateral and two posterolateral setae situated on scutum; sensilla flagelliform, branched in distal half; eyes 2 + 2. Legs 7-segmented, with 1 pair of claws and claw-like empodium; onychotriches present; leg subterminala and parasubterminala absent; pretarsalae I (?) and II absent; microgenuala II absent; microtarsala I situated distal of tarsala I; microtarsala II situated distal of tarsala II; tarsala II slender, with bulbiform apex; tibiala III and mastitarsalae III absent. Number of setae on leg coxae I–III: 1, 0, 1; sternal setae between coxae I absent; two sternal setae between coxae II and four sternal setae between coxae III.
</paragraph>
<paragraph id="B325B157FFEBFFFD56C888019418D3D6" blockId="2.[151,1436,151,1040]" box="[199,1146,367,392]" pageId="2" pageNumber="76">
<emphasis id="81EE6D45FFEBFFFD56C888019173D3D9" bold="true" box="[199,273,368,392]" pageId="2" pageNumber="76">Hosts.</emphasis>
Geckos of the families
<taxonomicName id="749ACAD4FFEBFFFD5413881E92C4D3D6" box="[540,678,367,391]" class="Reptilia" family="Gekkonidae" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="76" phylum="Chordata" rank="family">Gekkonidae</taxonomicName>
,
<taxonomicName id="749ACAD4FFEBFFFD54BE881E930FD3D6" box="[689,877,367,391]" class="Reptilia" family="Phyllodactylidae" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="76" phylum="Chordata" rank="family">Phyllodactylidae</taxonomicName>
and
<taxonomicName id="749ACAD4FFEBFFFD55AC881E9414D3D6" box="[931,1142,367,391]" class="Reptilia" family="Sphaerodactylidae" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="76" phylum="Chordata" rank="family">Sphaerodactylidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FB80E2DCFFEBFFFD56C888E291D1D3FD" box="[199,435,403,428]" pageId="2" pageNumber="76" type="distribution">
<paragraph id="B325B157FFEBFFFD56C888E291D1D3FD" blockId="2.[151,1436,151,1040]" box="[199,435,403,428]" pageId="2" pageNumber="76">
<emphasis id="81EE6D45FFEBFFFD56C888E29102D3FD" bold="true" box="[199,352,403,428]" pageId="2" pageNumber="76">Distribution.</emphasis>
<collectingCountry id="CB8DF1C7FFEBFFFD576888E591CDD3FD" box="[359,431,404,428]" name="Oman" pageId="2" pageNumber="76">Oman</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="FB80E2DCFFEBFFFD56C888C993A5D3A5" pageId="2" pageNumber="76" type="etymology">
<paragraph id="B325B157FFEBFFFD56C888C993A5D3A5" blockId="2.[151,1436,151,1040]" pageId="2" pageNumber="76">
<emphasis id="81EE6D45FFEBFFFD56C888C9912CD381" bold="true" box="[199,334,440,464]" pageId="2" pageNumber="76">Etymology.</emphasis>
The name of the new genus refers to the Arabian Peninsula, where its
<typeStatus id="6C210FF5FFEBFFFD526C88C994F1D381" box="[1123,1171,440,464]" pageId="2" pageNumber="76">type</typeStatus>
species was found, and to
<taxonomicName id="749ACAD4FFEBFFFD56BB88AD9249D3A5" authority="Torres and Braga, 1939" authorityName="Torres and Braga" authorityYear="1939" box="[180,555,476,501]" class="Arachnida" family="Leeuwenhoekiidae" genus="Apolonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="81EE6D45FFEBFFFD56BB88AD9179D3A5" box="[180,283,476,500]" italics="true" pageId="2" pageNumber="76">Apolonia</emphasis>
<bibRefCitation id="D70BCCA6FFEBFFFD572E88AC9249D3A5" author="Torres" box="[289,555,476,501]" pageId="2" pageNumber="76" refString="Torres, S. &amp; Braga, W. (1939) Apolonia tigipioensis g. e sp. n. (Trombiculinae) parasito de Gallus gallus dom. Chave para determinacao de generos. B 0 ̸ et i m da Secretar i a de Agr i cu ̸ tura, IΠduStr i a e C 0 merc i 0, 4, 37 - 44." type="journal article" year="1939">Torres and Braga, 1939</bibRefCitation>
</taxonomicName>
, the nominate genus of Apoloniinae.
</paragraph>
</subSubSection>
<subSubSection id="FB80E2DCFFEBFFFD56C88B719262D641" pageId="2" pageNumber="76" type="discussion">
<paragraph id="B325B157FFEBFFFD56C88B71914AD145" blockId="2.[151,1436,151,1040]" pageId="2" pageNumber="76">
<emphasis id="81EE6D45FFEBFFFD56C88B719159D049" bold="true" box="[199,315,512,536]" pageId="2" pageNumber="76">Remarks.</emphasis>
<taxonomicName id="749ACAD4FFEBFFFD574B8B719182D049" box="[324,480,512,536]" class="Arachnida" family="Trombiculidae" genus="Arabapolonia" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="81EE6D45FFEBFFFD574B8B719182D049" box="[324,480,512,536]" italics="true" pageId="2" pageNumber="76">Arabapolonia</emphasis>
</taxonomicName>
differs from all other Apoloniinae in a single palpal claw, the absence of coxala II (fCx = 1.0.1 vs. 1.1.1 or 1.2.1) and the absence of the sternal setae between leg coxae I (fSt = 0.2.4). The new genus shares the non-reduced scutum with the genera
<taxonomicName id="749ACAD4FFEBFFFD54D18B39945BD00E" authority="Lawrence, 1949" authorityName="Lawrence" authorityYear="1949" box="[734,1081,583,608]" class="Arachnida" family="Leeuwenhoekiidae" genus="Sauracarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="81EE6D45FFEBFFFD54D18B39930DD031" box="[734,879,584,608]" italics="true" pageId="2" pageNumber="76">Sauracarella</emphasis>
<bibRefCitation id="D70BCCA6FFEBFFFD55728B39945BD00E" author="Lawrence" box="[893,1081,583,607]" pageId="2" pageNumber="76" refString="Lawrence, R. F. (1949) The larval trombiculid mites of South African vertebrates. Annals of the Natal Museum, 11, 405 - 486." type="journal article" year="1949">Lawrence, 1949</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="749ACAD4FFEBFFFD52718B39949CD031" box="[1150,1278,584,608]" class="Arachnida" family="Leeuwenhoekiidae" genus="Afracarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="81EE6D45FFEBFFFD52718B39949CD031" box="[1150,1278,584,608]" italics="true" pageId="2" pageNumber="76">Afracarella</emphasis>
</taxonomicName>
Vercammen-
<bibRefCitation id="D70BCCA6FFEBFFFD56988B1D927AD0D5" author="Grandjean" box="[151,536,620,644]" pageId="2" pageNumber="76" refString="Vercammen-Grandjean, P. H. &amp; Kolebinova, M. (1968) Revision of the subfamily Apoloniinae Wharton, 1947 (Leeuwenhoekiidae: Acarina). Acarologia, 10, 250 - 268." type="journal article" year="1968">Grandjean and Kolebinova, 1968</bibRefCitation>
(all other genera of Apoloniinae have peniscutum, i.e. scutum with reduced posterior angles, with PL setae situated off scutum), but these two genera, unlike
<emphasis id="81EE6D45FFEBFFFD52218BE194B3D0F9" box="[1070,1233,656,680]" italics="true" pageId="2" pageNumber="76">
<taxonomicName id="749ACAD4FFEBFFFD52218BE194AFD0F9" box="[1070,1229,656,680]" class="Arachnida" family="Trombiculidae" genus="Arabapolonia" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">Arabapolonia</taxonomicName>
,
</emphasis>
possess the nasus and clavate sensilla.
<taxonomicName id="749ACAD4FFEBFFFD579A8BC59253D09D" box="[405,561,692,716]" class="Arachnida" family="Trombiculidae" genus="Arabapolonia" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="81EE6D45FFEBFFFD579A8BC59253D09D" box="[405,561,692,716]" italics="true" pageId="2" pageNumber="76">Arabapolonia</emphasis>
</taxonomicName>
is the second known genus of Apoloniinae (after the monotypic genus
<taxonomicName id="749ACAD4FFEBFFFD56988BA9909CD0A1" box="[151,254,728,752]" class="Arachnida" family="Leeuwenhoekiidae" genus="Apolonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="81EE6D45FFEBFFFD56988BA9909CD0A1" box="[151,254,728,752]" italics="true" pageId="2" pageNumber="76">Apolonia</emphasis>
</taxonomicName>
) having onychotriches and the second genus with palpal tarsus 6BS (after the monotypic genus
<taxonomicName id="749ACAD4FFEBFFFD56988B8D9175D145" box="[151,279,764,788]" class="Arachnida" family="Leeuwenhoekiidae" genus="Afracarella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="81EE6D45FFEBFFFD56988B8D9175D145" box="[151,279,764,788]" italics="true" pageId="2" pageNumber="76">Afracarella</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="B325B157FFEBFFFD56C88A5193E5D12E" blockId="2.[151,1436,151,1040]" pageId="2" pageNumber="76">
<taxonomicName id="749ACAD4FFEBFFFD56C88A519101D169" box="[199,355,800,824]" class="Arachnida" family="Trombiculidae" genus="Arabapolonia" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="81EE6D45FFEBFFFD56C88A519101D169" box="[199,355,800,824]" italics="true" pageId="2" pageNumber="76">Arabapolonia</emphasis>
</taxonomicName>
is the first known Apoloniinae genus having tracheae. These structures belong to the duct system of unknown function which was previously recorded in some genera of Leeuwenhoekiinae (
<bibRefCitation id="D70BCCA6FFEBFFFD532F8A3590B1D12E" author="Goff" pageId="2" pageNumber="76" refString="Goff, M. L., Loomis, R. B., Welbourn, W. C., &amp; Wrenn, W. J. (1982) A glossary of chigger terminology (Acari: Trombiculidae). Journal of Medical Entomology, 19, 221 - 238." type="journal article" year="1982">
Goff
<emphasis id="81EE6D45FFEBFFFD536F8A3495FED10D" box="[1376,1436,836,860]" italics="true" pageId="2" pageNumber="76">et al.</emphasis>
1982
</bibRefCitation>
), while in other genera of this subfamily it is certainly absent.
</paragraph>
<paragraph id="B325B157FFEBFFFD56C88AFC9262D641" blockId="2.[151,1436,151,1040]" pageId="2" pageNumber="76">
We were unable to find pretarsala on leg I in the new genus; actually, in the compact group of setae at the tip of the tarsus, one seta may be pretarsala, but its recognition can probably be realized only on new material of better quality or by scanning electron microscope. Tentatively, the absence of this seta can be regarded as another autapomorphy of
<taxonomicName id="749ACAD4FFEBFFFD57518A899198D641" box="[350,506,1016,1040]" class="Arachnida" family="Trombiculidae" genus="Arabapolonia" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="81EE6D45FFEBFFFD57518A899198D641" box="[350,506,1016,1040]" italics="true" pageId="2" pageNumber="76">Arabapolonia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="A7955EAB09A6A3CC742B346CF0B24970">A new genus and species of Apoloniinae (Acari: Trombiculidae) from Oman</mods:title>
</mods:titleInfo>
<mods:name id="E4768E3F5DDD361ABD77F176A2F38726" type="personal">
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</mods:role>
<mods:namePart id="FD5FDFB9B408443B3EFCB739A0249AEE">Stekolnikov, Alexandr A.</mods:namePart>
</mods:name>
<mods:name id="C0262DC93DAEABF0E7ECCBABC9981109" type="personal">
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</mods:role>
<mods:namePart id="FA17E7C3AD854B8C5217717082B2F310">Carranza, Salvador</mods:namePart>
</mods:name>
<mods:name id="80A13206D474C8250E85E00D19229378" type="personal">
<mods:role id="24F41E9B601428DC9E3D396C9CE5F441">
<mods:roleTerm id="D91F5F0D2D06CE84297A457DB6C8990B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C37DEE544D57520453EDFB040762C03B">Gomez-Diaz, Elena</mods:namePart>
</mods:name>
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<mods:title id="37E7626DF2A15C9DEF412A691A44CF5C">Zootaxa</mods:title>
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<mods:part id="6903D82EBDA8BD3C9C7008BBB10BCD34">
<mods:date id="61A5867D855A02F497D7859591C30F92">2012</mods:date>
<mods:detail id="B7E7AB04D2A0A2C5CDF89DFA5FEBA334" type="volume">
<mods:number id="6259758FA722DB692EE33ECE07C28E67">3499</mods:number>
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<mods:identifier id="372B5EFC783D3AF66A8AD5BE35B81359" type="DOI">10.5281/zenodo.212553</mods:identifier>
<mods:identifier id="7BD557BA38D6F389527972E4295A199F" type="GBIF-Dataset">7e49d608-5e2b-4987-b853-64678deae6c3</mods:identifier>
<mods:identifier id="529CEF685080A0D2DFB04E317714B499" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B071E4F554A772B80FE3A30F549DE736" type="Zenodo-Dep">212553</mods:identifier>
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<treatment id="3B330041FFEBFFFA56988D159093D5B0" ID-DOI="http://doi.org/10.5281/zenodo.6168256" ID-GBIF-Taxon="119527719" ID-Zenodo-Dep="6168256" LSID="urn:lsid:zoobank.org:act:E955DE37-7804-4B12-B9BF-3489140803D7" httpUri="http://treatment.plazi.org/id/3B330041FFEBFFFA56988D159093D5B0" lastPageId="5" lastPageNumber="79" pageId="2" pageNumber="76">
<subSubSection id="FB80E2DCFFEBFFFD56988D159170D6CF" pageId="2" pageNumber="76" type="nomenclature">
<paragraph id="B325B157FFEBFFFD56988D159248D62C" blockId="2.[151,554,1123,1182]" box="[151,554,1123,1150]" pageId="2" pageNumber="76">
<heading id="E86D063BFFEBFFFD56988D159248D62C" bold="true" box="[151,554,1123,1150]" fontSize="11" level="1" pageId="2" pageNumber="76" reason="1">
<emphasis id="81EE6D45FFEBFFFD56988D159248D62C" bold="true" box="[151,554,1123,1150]" pageId="2" pageNumber="76">
<taxonomicName id="749ACAD4FFEBFFFD56988D1591A7D62F" box="[151,453,1124,1150]" class="Arachnida" family="Trombiculidae" genus="Arabapolonia" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="76" phylum="Arthropoda" rank="species" species="omanensis" status="sp. nov.">
<emphasis id="81EE6D45FFEBFFFD56988D1591A7D62F" bold="true" box="[151,453,1124,1150]" italics="true" pageId="2" pageNumber="76">Arabapolonia omanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9ADDD03EFFEBFFFD57C38D129248D62C" box="[460,554,1123,1149]" pageId="2" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B325B157FFEBFFFD56988DF79170D6CF" blockId="2.[151,554,1123,1182]" box="[151,274,1158,1182]" pageId="2" pageNumber="76">(Figs. 1, 2)</paragraph>
</subSubSection>
<subSubSection id="FB80E2DCFFEBFFFC56988DBF92CDD588" lastPageId="3" lastPageNumber="77" pageId="2" pageNumber="76" type="diagnosis">
<paragraph id="B325B157FFEBFFFD56988DBF9131D77F" blockId="2.[151,1436,1230,1326]" pageId="2" pageNumber="76">
<emphasis id="81EE6D45FFEBFFFD56988DBF9173D6B6" bold="true" box="[151,273,1230,1255]" pageId="2" pageNumber="76">Diagnosis.</emphasis>
SIF = 6BS-N(b)-1-2110.0000; fsp = 7.7.7; fCx = 1.0.1; fSt = 0.2.4; fPp = B/B/BBB; fSc: PL&gt; AM&gt; AL; Ip = 711–941; fD = 2H-
<date id="C7249797FFEBFFFD57EB8D83924ED75B" box="[484,556,1266,1290]" pageId="2" pageNumber="76" value="2010-08-08">10-8-8</date>
(7)-6-4(2); DS = 35–38; VS = 15–17; NDV = 52–53. Standard measurements given in
<tableCitation id="FE1884ECFFEBFFFD56F68C67912ED77F" box="[249,332,1302,1326]" captionStart="TABLE 1" captionStartId="2.[151,235,1369,1390]" captionText="TABLE 1. Standard measurements of Arabapolonia omanensis gen. et sp. nov." pageId="2" pageNumber="76">Table 1</tableCitation>
.
</paragraph>
<caption id="E7E5E1DFFFEBFFFD56988C2893DCD73E" box="[151,958,1369,1391]" pageId="2" pageNumber="76">
<paragraph id="B325B157FFEBFFFD56988C2893DCD73E" blockId="2.[151,1408,1369,1648]" box="[151,958,1369,1391]" pageId="2" pageNumber="76">
<emphasis id="81EE6D45FFEBFFFD56988C289166D73F" bold="true" box="[151,260,1369,1390]" pageId="2" pageNumber="76">TABLE 1.</emphasis>
Standard measurements of
<taxonomicName id="749ACAD4FFEBFFFD54138C289378D73E" box="[540,794,1369,1391]" class="Arachnida" family="Trombiculidae" genus="Arabapolonia" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="76" phylum="Arthropoda" rank="species" species="omanensis">
<emphasis id="81EE6D45FFEBFFFD54138C289378D73E" box="[540,794,1369,1391]" italics="true" pageId="2" pageNumber="76">Arabapolonia omanensis</emphasis>
</taxonomicName>
gen. et sp. nov.
</paragraph>
</caption>
<paragraph id="B325B157FFEBFFFD56908CFB95E2D43E" blockId="2.[151,1408,1369,1648]" pageId="2" pageNumber="76">
Slide AW PW SB ASB PSB SD P-PL AP AA AM AL PL S H 8423 (
<typeStatus id="6C210FF5FFEBFFFD56D08CC29159D798" box="[223,315,1459,1481]" pageId="2" pageNumber="76" type="holotype">holotype</typeStatus>
) 45 62
<date id="C7249797FFEBFFFD54258CC59280D798" box="[554,738,1460,1481]" pageId="2" pageNumber="76" value="1923-12-23">23 23 12</date>
35
<date id="C7249797FFEBFFFD556B8CC59475D798" box="[868,1047,1459,1481]" pageId="2" pageNumber="76" value="1914-05-15">14 15 5</date>
31 25 36 72 32 8424 (EGD-7) 49 62
<date id="C7249797FFEBFFFD54258CAC92B4D7A2" box="[554,726,1501,1523]" pageId="2" pageNumber="76" value="1926-08-24">26 24 8</date>
32
<date id="C7249797FFEBFFFD556B8CAF9475D7A2" box="[868,1047,1501,1523]" pageId="2" pageNumber="76" value="1912-05-15">12 15 5</date>
29 25 34 - 27 8419 (EGD-1) 42 56
<date id="C7249797FFEBFFFD54258F7692B4D44D" box="[554,726,1543,1564]" pageId="2" pageNumber="76" value="1922-09-21">22 21 9</date>
30
<date id="C7249797FFEBFFFD556B8F769475D44D" box="[868,1047,1542,1564]" pageId="2" pageNumber="76" value="1914-05-11">14 11 5</date>
26 22 29 58 - 8417 (EGD-6) 41 52 22 19
<date id="C7249797FFEBFFFD54C68F41931CD414" box="[713,894,1584,1605]" pageId="2" pageNumber="76" value="1932-12-13">13 32 12</date>
13
<date id="C7249797FFEBFFFD52058F419564D414" box="[1034,1286,1584,1606]" pageId="2" pageNumber="76" value="1922-05-15">5 - 15 22</date>
52 20 8421 (EGD-8) 40 54
<date id="C7249797FFEBFFFD54258F2B9282D43E" box="[554,736,1626,1647]" pageId="2" pageNumber="76" value="1923-11-23">23 23 11</date>
33
<date id="C7249797FFEBFFFD556B8F2B9475D43E" box="[868,1047,1625,1647]" pageId="2" pageNumber="76" value="1914-05-14">14 14 5</date>
26 16 24 54 20
</paragraph>
<paragraph id="B325B157FFEBFFFD56988FE79160D4FD" blockId="2.[151,1398,1686,1967]" box="[151,258,1686,1708]" pageId="2" pageNumber="76">continued.</paragraph>
<paragraph id="B325B157FFEBFFFC56908FB992CDD588" blockId="2.[151,1398,1686,1967]" lastBlockId="3.[151,1437,1890,2009]" lastPageId="3" lastPageNumber="77" pageId="2" pageNumber="76">
Slide D min D max V min V max pa pm pp Ip DS VS NDV TaIII TaW 8423 (
<typeStatus id="6C210FF5FFEBFFFD56D08F829159D558" box="[223,315,1779,1801]" pageId="2" pageNumber="76" type="holotype">holotype</typeStatus>
) 29 37 29 36 315 290 337 941 - - - 95 11 8424 (EGD-7) 29 34 19 32 295 261 304 860 38 15 53 86 11 8419 (EGD-1) 29 34 20 30 283 230 274 787 - - - 77 10 8417 (EGD-6) 20 23 16 24 248 211 252 711 35 17 52 70 10 8421 (EGD-8) 20 23 - - 256 230 268 754 - - - 70 10
<emphasis id="81EE6D45FFEAFFFC56988E159171D528" bold="true" box="[151,275,1892,1915]" pageId="3" pageNumber="77">FIGURE 1.</emphasis>
<taxonomicName id="749ACAD4FFEAFFFC57168E159275D52B" box="[281,535,1892,1914]" class="Arachnida" family="Trombiculidae" genus="Arabapolonia" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="77" phylum="Arthropoda" rank="species" species="omanensis" status="gen. nov.">
<emphasis id="81EE6D45FFEAFFFC57168E159275D52B" box="[281,535,1892,1914]" italics="true" pageId="3" pageNumber="77">Arabapolonia omanensis</emphasis>
</taxonomicName>
<emphasis id="81EE6D45FFEAFFFC54128E149364D528" bold="true" box="[541,774,1892,1914]" pageId="3" pageNumber="77">
<taxonomicNameLabel id="9ADDD03EFFEAFFFC54128E14921ED52B" box="[541,636,1893,1914]" pageId="3" pageNumber="77" rank="genus">gen. nov.</taxonomicNameLabel>
et sp. nov. A
</emphasis>
, scutum and eyes of
<typeStatus id="6C210FF5FFEAFFFC55D28E15945BD52B" box="[989,1081,1892,1914]" pageId="3" pageNumber="77" type="holotype">holotype</typeStatus>
;
<emphasis id="81EE6D45FFEAFFFC524C8E149436D52B" bold="true" box="[1091,1108,1893,1914]" pageId="3" pageNumber="77">B</emphasis>
, dorsal idiosomal seta of 1st row in
<typeStatus id="6C210FF5FFEAFFFC56BA8EF49173D5CA" box="[181,273,1925,1947]" pageId="3" pageNumber="77" type="holotype">holotype</typeStatus>
;
<emphasis id="81EE6D45FFEAFFFC57108EF49153D5CD" bold="true" box="[287,305,1925,1948]" pageId="3" pageNumber="77">C</emphasis>
, dorsal idiosomal seta of 1st row in specimen No. 8421;
<emphasis id="81EE6D45FFEAFFFC55978EF793C8D5CA" bold="true" box="[920,938,1926,1947]" pageId="3" pageNumber="77">D</emphasis>
, ventral idiosomal seta in
<typeStatus id="6C210FF5FFEAFFFC52C58EF49544D5CA" box="[1226,1318,1925,1947]" pageId="3" pageNumber="77" type="holotype">holotype</typeStatus>
;
<emphasis id="81EE6D45FFEAFFFC533B8EF79527D5CA" bold="true" box="[1332,1349,1926,1947]" pageId="3" pageNumber="77">E</emphasis>
, ventral idiosomal seta in specimen No. 8421;
<emphasis id="81EE6D45FFEAFFFC54258ED49255D5EB" bold="true" box="[554,567,1957,1978]" pageId="3" pageNumber="77">F</emphasis>
, arrangement of dorsal idiosomal setae;
<emphasis id="81EE6D45FFEAFFFC55ED8ED59391D5EA" bold="true" box="[994,1011,1956,1979]" pageId="3" pageNumber="77">G</emphasis>
, arrangement of ventral idiosomal setae. Scale bars: 50 μm (
<emphasis id="81EE6D45FFEAFFFC57548EB2910FD589" bold="true" box="[347,365,1987,2008]" pageId="3" pageNumber="77">A</emphasis>
), 20 μm (
<emphasis id="81EE6D45FFEAFFFC57DD8EB29262D589" bold="true" box="[466,512,1987,2008]" pageId="3" pageNumber="77">B–E</emphasis>
), 100 μm (
<emphasis id="81EE6D45FFEAFFFC547D8EB292FFD588" bold="true" box="[626,669,1986,2009]" pageId="3" pageNumber="77">F, G</emphasis>
).
</paragraph>
</subSubSection>
<caption id="E7E5E1DFFFEDFFFB56988EF193E9D5A3" httpUri="https://zenodo.org/record/212554/files/figure.png" pageId="4" pageNumber="78" targetBox="[154,1424,208,1898]" targetPageId="4">
<paragraph id="B325B157FFEDFFFB56988EF193E9D5A3" blockId="4.[151,1436,1920,2034]" pageId="4" pageNumber="78">
<emphasis id="81EE6D45FFEDFFFB56988EF19176D5C4" bold="true" box="[151,276,1920,1943]" pageId="4" pageNumber="78">FIGURE 2.</emphasis>
<taxonomicName id="749ACAD4FFEDFFFB57138EF1927ED5C7" box="[284,540,1920,1942]" class="Arachnida" family="Trombiculidae" genus="Arabapolonia" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="78" phylum="Arthropoda" rank="species" species="omanensis" status="gen. nov.">
<emphasis id="81EE6D45FFEDFFFB57138EF1927ED5C7" box="[284,540,1920,1942]" italics="true" pageId="4" pageNumber="78">Arabapolonia omanensis</emphasis>
</taxonomicName>
<emphasis id="81EE6D45FFEDFFFB542B8EF09376D5C4" bold="true" box="[548,788,1920,1942]" pageId="4" pageNumber="78">
<taxonomicNameLabel id="9ADDD03EFFEDFFFB542B8EF092E6D5C7" box="[548,644,1921,1942]" pageId="4" pageNumber="78" rank="genus">gen. nov.</taxonomicNameLabel>
et sp. nov. A
</emphasis>
, dorsal aspect of gnathosoma in holotype;
<emphasis id="81EE6D45FFEDFFFB52DD8EF09481D5C7" bold="true" box="[1234,1251,1921,1942]" pageId="4" pageNumber="78">B</emphasis>
, ventral aspect of gnathosoma in holotype;
<emphasis id="81EE6D45FFEDFFFB57938EEF91CCD5E4" bold="true" box="[412,430,1950,1973]" pageId="4" pageNumber="78">C</emphasis>
, leg I;
<emphasis id="81EE6D45FFEDFFFB57F48EEE926FD5E5" bold="true" box="[507,525,1951,1972]" pageId="4" pageNumber="78">D</emphasis>
, leg II;
<emphasis id="81EE6D45FFEDFFFB546D8EEE9211D5E5" bold="true" box="[610,627,1951,1972]" pageId="4" pageNumber="78">E</emphasis>
, leg III;
<emphasis id="81EE6D45FFEDFFFB54DF8EEE92BFD5E5" bold="true" box="[720,733,1951,1972]" pageId="4" pageNumber="78">F</emphasis>
, tip of leg tarsus III in holotype showing two onychotriches in both claws;
<emphasis id="81EE6D45FFEDFFFB56D18ECC908DD585" bold="true" box="[222,239,1981,2004]" pageId="4" pageNumber="78">G</emphasis>
, tarsala and microtarsala I in holotype;
<emphasis id="81EE6D45FFEDFFFB54898ECF92F8D582" bold="true" box="[646,666,1982,2003]" pageId="4" pageNumber="78">H</emphasis>
, tarsala and microtarsala II in holotype. Scale bars: 20 μm (
<emphasis id="81EE6D45FFEDFFFB52F08ECC950FD582" bold="true" box="[1279,1389,1981,2003]" pageId="4" pageNumber="78">A, B, F–H</emphasis>
), 50 μm (
<emphasis id="81EE6D45FFEDFFFB56C88EAA9095D5A3" bold="true" box="[199,247,2011,2034]" pageId="4" pageNumber="78">C–E</emphasis>
). Abbreviation: S—subterminala (eupathidium) on palpal tarsus.
</paragraph>
</caption>
<subSubSection id="FB80E2DCFFECFFFA56C889E693C6D1BE" pageId="5" pageNumber="79" type="description">
<paragraph id="B325B157FFECFFFA56C889E6927DD313" blockId="5.[151,1437,151,2017]" pageId="5" pageNumber="79">
<emphasis id="81EE6D45FFECFFFA56C889E691D2D2E1" bold="true" box="[199,432,151,176]" pageId="5" pageNumber="79">Description (larva).</emphasis>
IDIOSOMA. Eyes large, 2 + 2; eyes of anterior and posterior pair separated, ocular plate absent. One or 2 pairs of humeral setae (marginal idiosomal setae of 1st dorsal row may be considered as second pair of humeral setae), 33–36 moderately barbed dorsal idiosomal setae, fD=2H-
<date id="C7249797FFECFFFA5236899394E2D2AB" box="[1081,1152,226,250]" pageId="5" pageNumber="79" value="2010-08-08">10-8-8</date>
(7)-6-4(2); 2 sternal setae between leg coxae II and 4 sternal setae between leg coxae III; 15–17 ventral setae; total number of idiosomal setae excluding coxal and sternal 52–53.
</paragraph>
<paragraph id="B325B157FFECFFFA56C8883E9258D06E" blockId="5.[151,1437,151,2017]" pageId="5" pageNumber="79">GNATHOSOMA. Cheliceral blade with 1 inconspicuous denticle near apex, without tricuspid cap; cheliceral base with few puncta in proximal part; gnathobase with sparse puncta and transverse cuticular striations, with 1 pair of branched setae; tracheae extend from gnathobase to leg coxa I, spiracles not found; palpal femur and genu with few puncta; galeala nude or having one small cilium; palpal claw single; setae on palpal femur and genu branched; ventral and lateral palpal tibial setae branched, dorsal palpal tibial seta heavily barbed in distal half, in shape of tooth-brush; palpal tarsus with 6 non-specialized setae (3 setae definitely branched and 3 setae seem nude), nude subterminala and tarsala.</paragraph>
<paragraph id="B325B157FFECFFFA56C88B3B9346D09E" blockId="5.[151,1437,151,2017]" pageId="5" pageNumber="79">SCUTUM. Nearly hexagonal, with large, rounded anterior projection, without nasus, posterior scutal margin slightly prominent, angulated or rounded, with few cuticular striations; 2 anteromedian setae, 2 anterolateral and 2 posterolateral setae situated on scutum; AM bases at level of ALs; SB slightly posterior to level of PLs; PL&gt; AM&gt; AL; flagelliform sensilla heavily branched in distal half.</paragraph>
<paragraph id="B325B157FFECFFFA56C88BAB93C6D1BE" blockId="5.[151,1437,151,2017]" pageId="5" pageNumber="79">LEGS. All 7-segmented, with 1 pair of claws and claw-like empodium; claws with 1–2 long onychotriches, almost as long as claw itself. Leg I: coxa with 1 non-specialized branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, two genualae, microgenuala; tibia 7B, two tibialae, microtibiala; tarsus 23B, tarsala 16–22 long, with sparse annulation, microtarsala distal of tarsala; subterminala, parasubterminala and pretarsala absent. Leg II: coxa without seta; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala, without microgenuala; tibia 6B, 2 tibialae; tarsus 15B, tarsala 19–26 long, very thin, with sparse annulation and bulbiform apex, microtarsala distal of tarsala, pretarsala absent. Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 4B, tibiala absent; tarsus 15B, mastitarsalae absent.</paragraph>
</subSubSection>
<subSubSection id="FB80E2DCFFECFFFA56C88A8A9261D7B6" pageId="5" pageNumber="79" type="materials_examined">
<paragraph id="B325B157FFECFFFA56C88A8A92B0D7CE" blockId="5.[151,1437,151,2017]" pageId="5" pageNumber="79">
<emphasis id="81EE6D45FFECFFFA56C88A8A9116D642" bold="true" box="[199,372,1018,1043]" pageId="5" pageNumber="79">
<typeStatus id="6C210FF5FFECFFFA56C88A8A9162D642" box="[199,256,1019,1043]" pageId="5" pageNumber="79">Type</typeStatus>
material.
</emphasis>
<typeStatus id="6C210FF5FFECFFFA57718A8B9184D643" box="[382,486,1018,1042]" pageId="5" pageNumber="79" type="holotype">Holotype</typeStatus>
larva (
<collectionCode id="D58B2992FFECFFFA54388A8B9210D643" box="[567,626,1018,1042]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="79">ZISP</collectionCode>
collection number 8423, T-Tr.-56) ex
<taxonomicName id="749ACAD4FFECFFFA52278A8A9559D642" box="[1064,1339,1019,1043]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="79" phylum="Chordata" rank="species" species="lemurinus">
<emphasis id="81EE6D45FFECFFFA52278A8A9559D642" box="[1064,1339,1019,1043]" italics="true" pageId="5" pageNumber="79">Hemidactylus lemurinus</emphasis>
</taxonomicName>
(Sauria:
<taxonomicName id="749ACAD4FFECFFFA56988D6E9142D666" box="[151,288,1055,1079]" class="Reptilia" family="Gekkonidae" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="79" phylum="Chordata" rank="family">Gekkonidae</taxonomicName>
) (mite Id S7609, tube No. EGD-3),
<collectingCountry id="CB8DF1C7FFECFFFA54ED8D6F9359D666" box="[738,827,1054,1079]" name="Oman" pageId="5" pageNumber="79">
<emphasis id="81EE6D45FFECFFFA54ED8D6F9359D666" bold="true" box="[738,827,1054,1079]" pageId="5" pageNumber="79">OMAN</emphasis>
</collectingCountry>
: Dhofar Governorate, Wadi Ayoun (locality No. 1 according to above list),
<geoCoordinate id="D6AED790FFECFFFA57B98D33925DD60B" box="[438,575,1090,1114]" direction="north" orientation="latitude" pageId="5" pageNumber="79" precision="1" value="17.24218">17.24218 N</geoCoordinate>
,
<geoCoordinate id="D6AED790FFECFFFA54438D3392B3D60B" box="[588,721,1090,1114]" direction="east" orientation="longitude" pageId="5" pageNumber="79" precision="1" value="53.89095">53.89095 E</geoCoordinate>
,
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a.s.l.,
<date id="C7249797FFECFFFA55848D33947BD60B" box="[907,1049,1090,1115]" pageId="5" pageNumber="79" value="2011-05-01">1 May 2011</date>
, coll. S. Carranza and E. Gómez- Díaz; 1 larva
<typeStatus id="6C210FF5FFECFFFA57228D1991EFD6D1" box="[301,397,1128,1152]" pageId="5" pageNumber="79" type="paratype">paratype</typeStatus>
(8419) ex
<taxonomicName id="749ACAD4FFECFFFA540A8D1992D7D62E" box="[517,693,1127,1151]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="79" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="81EE6D45FFECFFFA540A8D1992D7D62E" box="[517,693,1127,1151]" italics="true" pageId="5" pageNumber="79">H. homoeolepis</emphasis>
</taxonomicName>
(S7673, EGD-1), Ash Sharqiyah Region,
<quantity id="74621CB2FFECFFFA52848D1694A1D62E" box="[1163,1219,1127,1151]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="79" unit="km" value="2.0">2 km</quantity>
W from Qumaylah (locality No. 2),
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,
<geoCoordinate id="D6AED790FFECFFFA57ED8DFB9235D6F3" box="[482,599,1162,1186]" direction="east" orientation="longitude" pageId="5" pageNumber="79" precision="5" value="59.6082">59.6082 E</geoCoordinate>
,
<quantity id="74621CB2FFECFFFA546D8DFB92F9D6F3" box="[610,667,1162,1186]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="79" unit="m" value="42.0">42 m</quantity>
a.s.l.,
<date id="C7249797FFECFFFA54EC8DFB93C0D6F3" box="[739,930,1162,1186]" pageId="5" pageNumber="79" value="2010-10-10">10 October 2010</date>
, coll. S. Carranza; 1 larva
<typeStatus id="6C210FF5FFECFFFA52C68DFA954BD6F2" box="[1225,1321,1163,1187]" pageId="5" pageNumber="79" type="paratype">paratype</typeStatus>
(8424) ex
<taxonomicName id="749ACAD4FFECFFFA56988DDE91D9D696" box="[151,443,1199,1223]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="79" phylum="Chordata" rank="species" species="hasselquistii">
<emphasis id="81EE6D45FFECFFFA56988DDE91D9D696" box="[151,443,1199,1223]" italics="true" pageId="5" pageNumber="79">Ptyodactylus hasselquistii</emphasis>
</taxonomicName>
(Sauria:
<taxonomicName id="749ACAD4FFECFFFA542E8DDE92BDD696" box="[545,735,1199,1223]" class="Reptilia" family="Phyllodactylidae" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="79" phylum="Chordata" rank="family">Phyllodactylidae</taxonomicName>
) (S7652, EGD-7), Ash Sharqiyah Region, Wadi Tiwi (locality No. 3),
<geoCoordinate id="D6AED790FFECFFFA56FA8DA391E2D6BB" box="[245,384,1234,1258]" direction="north" orientation="latitude" pageId="5" pageNumber="79" precision="1" value="22.75737">22.75737 N</geoCoordinate>
,
<geoCoordinate id="D6AED790FFECFFFA57808DA39274D6BB" box="[399,534,1234,1258]" direction="east" orientation="longitude" pageId="5" pageNumber="79" precision="1" value="59.09399">59.09399 E</geoCoordinate>
,
<quantity id="74621CB2FFECFFFA54298DA392F1D6BB" box="[550,659,1234,1258]" metricMagnitude="3" metricUnit="m" metricValue="1.1965" pageId="5" pageNumber="79" unit="m" value="1196.5">1196.5 m</quantity>
a.s.l.,
<date id="C7249797FFECFFFA54EA8DA29314D6BB" box="[741,886,1234,1259]" pageId="5" pageNumber="79" value="2011-05-07">7 May 2011</date>
, coll. S. Carranza and E. Gómez-Díaz; 2 larva
<typeStatus id="6C210FF5FFECFFFA56988D899160D741" box="[151,258,1272,1296]" pageId="5" pageNumber="79" type="paratype">paratypes</typeStatus>
(8417 and 8418) ex
<taxonomicName id="749ACAD4FFECFFFA57F28D8992B6D75E" box="[509,724,1272,1295]" class="Reptilia" family="Sphaerodactylidae" genus="Pristurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="79" phylum="Chordata" rank="species" species="rupestris">
<emphasis id="81EE6D45FFECFFFA57F28D8992B6D75E" box="[509,724,1272,1295]" italics="true" pageId="5" pageNumber="79">Pristurus rupestris</emphasis>
</taxonomicName>
(Sauria:
<taxonomicName id="749ACAD4FFECFFFA554E8D869470D75E" box="[833,1042,1271,1295]" class="Reptilia" family="Sphaerodactylidae" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="79" phylum="Chordata" rank="family">Sphaerodactylidae</taxonomicName>
) (S7270, EGD-6), Ad Dakhiliyah Governorate,
<quantity id="74621CB2FFECFFFA573D8C6B910FD763" box="[306,365,1306,1330]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="79" unit="km" value="8.0">8 km</quantity>
N of Tanuf, Jebel Akhdar (locality No. 4),
<geoCoordinate id="D6AED790FFECFFFA556C8C6B9389D763" box="[867,1003,1306,1330]" direction="north" orientation="latitude" pageId="5" pageNumber="79" precision="1" value="23.12441">23.12441 N</geoCoordinate>
,
<geoCoordinate id="D6AED790FFECFFFA55F68C6B941CD763" box="[1017,1150,1306,1330]" direction="east" orientation="longitude" pageId="5" pageNumber="79" precision="1" value="57.45664">57.45664 E</geoCoordinate>
,
<quantity id="74621CB2FFECFFFA52848C6B9495D763" box="[1163,1271,1306,1330]" metricMagnitude="3" metricUnit="m" metricValue="1.7427" pageId="5" pageNumber="79" unit="m" value="1742.7">1742.7 m</quantity>
a.s.l.,
<date id="C7249797FFECFFFA534B8C6B90B3D706" pageId="5" pageNumber="79" value="2011-05-10">10 May 2011</date>
, coll. S. Carranza and E. Gómez-Díaz; 2 larva
<typeStatus id="6C210FF5FFECFFFA54F28C31930BD709" box="[765,873,1344,1368]" pageId="5" pageNumber="79" type="paratype">paratypes</typeStatus>
(8421 and 8422) ex
<taxonomicName id="749ACAD4FFECFFFA526C8C319558D706" box="[1123,1338,1344,1367]" class="Reptilia" family="Sphaerodactylidae" genus="Pristurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="79" phylum="Chordata" rank="species" species="rupestris">
<emphasis id="81EE6D45FFECFFFA526C8C319558D706" box="[1123,1338,1344,1367]" italics="true" pageId="5" pageNumber="79">Pristurus rupestris</emphasis>
</taxonomicName>
(S7251, EGD-8), Al Batinah Region, Haat Village, Jebel Akhdar (locality No. 5),
<geoCoordinate id="D6AED790FFECFFFA55C08C139437D72B" box="[975,1109,1378,1402]" direction="north" orientation="latitude" pageId="5" pageNumber="79" precision="1" value="23.17711">23.17711 N</geoCoordinate>
,
<geoCoordinate id="D6AED790FFECFFFA526E8C139486D72B" box="[1121,1252,1378,1402]" direction="east" orientation="longitude" pageId="5" pageNumber="79" precision="1" value="57.41018">57.41018 E</geoCoordinate>
,
<quantity id="74621CB2FFECFFFA52FF8C139538D72B" box="[1264,1370,1378,1402]" metricMagnitude="3" metricUnit="m" metricValue="1.3077" pageId="5" pageNumber="79" unit="m" value="1307.7">1307.7 m</quantity>
a.s.l.,
<date id="C7249797FFECFFFA56988CF69142D7CE" box="[151,288,1415,1440]" pageId="5" pageNumber="79" value="2011-05-08">8 May 2011</date>
, coll. S. Carranza and E. Gómez-Díaz.
</paragraph>
<paragraph id="B325B157FFECFFFA56C88CDA9261D7B6" blockId="5.[151,1437,151,2017]" pageId="5" pageNumber="79">
<emphasis id="81EE6D45FFECFFFA56C88CDA91E6D792" bold="true" box="[199,388,1450,1475]" pageId="5" pageNumber="79">
<typeStatus id="6C210FF5FFECFFFA56C88CDA9160D792" box="[199,258,1451,1475]" pageId="5" pageNumber="79">Type</typeStatus>
deposition
</emphasis>
.
<typeStatus id="6C210FF5FFECFFFA579D8CDB9198D793" box="[402,506,1450,1474]" pageId="5" pageNumber="79" type="holotype">Holotype</typeStatus>
and
<typeStatus id="6C210FF5FFECFFFA543E8CDA92FFD792" box="[561,669,1451,1475]" pageId="5" pageNumber="79" type="paratype">paratypes</typeStatus>
are deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg,
<collectingCountry id="CB8DF1C7FFECFFFA57BE8CBE919DD7B6" box="[433,511,1487,1511]" name="Russia" pageId="5" pageNumber="79">Russia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="FB80E2DCFFECFFFA56C88C829574D45B" box="[199,1302,1522,1547]" pageId="5" pageNumber="79" type="etymology">
<paragraph id="B325B157FFECFFFA56C88C829574D45B" blockId="5.[151,1437,151,2017]" box="[199,1302,1522,1547]" pageId="5" pageNumber="79">
<emphasis id="81EE6D45FFECFFFA56C88C82912CD45A" bold="true" box="[199,334,1523,1547]" pageId="5" pageNumber="79">Etymology.</emphasis>
The species epithet refers to the
<collectingCountry id="CB8DF1C7FFECFFFA54B08C8393F2D45B" box="[703,912,1522,1546]" name="Oman" pageId="5" pageNumber="79">Sultanate of Oman</collectingCountry>
where the
<typeStatus id="6C210FF5FFECFFFA52038C82945ED45A" box="[1036,1084,1523,1547]" pageId="5" pageNumber="79">type</typeStatus>
locality is situated.
</paragraph>
</subSubSection>
<subSubSection id="FB80E2DCFFECFFFA56C88F669093D5B0" pageId="5" pageNumber="79" type="discussion">
<paragraph id="B325B157FFECFFFA56C88F669093D5B0" blockId="5.[151,1437,151,2017]" pageId="5" pageNumber="79">
<emphasis id="81EE6D45FFECFFFA56C88F669159D47E" bold="true" box="[199,315,1559,1583]" pageId="5" pageNumber="79">Remarks.</emphasis>
Size differences within the examined material are very significant. First two specimens in
<tableCitation id="FE1884ECFFECFFFA53478F6695FED47E" box="[1352,1436,1559,1583]" captionStart="TABLE 1" captionStartId="2.[151,235,1369,1390]" captionText="TABLE 1. Standard measurements of Arabapolonia omanensis gen. et sp. nov." pageId="5" pageNumber="79">Table 1</tableCitation>
(Nos 8423 and 8424) have the widest scutum (measurements AW and PW), longest setae (AM–V max) and longest legs (pa–Ip and TaIII). Other two specimens (Nos 8417 and 8421) are the smallest, and specimen No. 8419 occupies the intermediate position. It is remarkable that the dorsal idiosomal setae in the largest specimens are almost twice longer than those in the smallest specimens (compare Fig. 1 B and C) and the legs of the
<typeStatus id="6C210FF5FFECFFFA531E8FD89511D490" box="[1297,1395,1705,1729]" pageId="5" pageNumber="79" type="holotype">holotype</typeStatus>
are 1.3 times longer than the legs of the smallest specimen (Ip = 941 vs. 711). In view of the fact, that non-metric characters are identical in all examined specimens, we consider these differences to be an individual variation. Dependence of size parameters on climatic factors was repeatedly recorded in chigger mites (e.g. Stekolnikov 1998, 2002;
<bibRefCitation id="D70BCCA6FFECFFFA57268E489217D500" author="Stekolnikov" box="[297,629,1849,1873]" pageId="5" pageNumber="79" refString="Stekolnikov, A. A. &amp; Klimov, P. B. (2010) A revision of chiggers of the minuta species-group (Acari: Trombiculidae: Neotrombicula Hirst, 1925) using multivariate morphometrics. Systematic Parasitology, 77, 55 - 69." type="journal article" year="2010">Stekolnikov &amp; Klimov 2010</bibRefCitation>
); revealing of such variability is very natural in generally hot and dry climate of
<collectingCountry id="CB8DF1C7FFECFFFA571E8E2C913BD524" box="[273,345,1885,1909]" name="Oman" pageId="5" pageNumber="79">Oman</collectingCountry>
. It is noteworthy that two smallest specimens (Nos 8417 and 8421) were collected in the places having the same ecological characteristics—localities 4 and 5, respectively. They are situated in the mountains, at elevations higher than in other collection localities (1743 and 1308 vs. 679, 1197, and
<quantity id="74621CB2FFECFFFA52558ED494F4D5EC" box="[1114,1174,1957,1981]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="79" unit="m" value="42.0">42 m</quantity>
), and in more wet local climate.
</paragraph>
</subSubSection>
</treatment>
</document>

152
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.523.6066" ID-GBIF-Dataset="afcfa837-6ba8-4a8d-bf4f-618e7380e8f3" ID-PMC="PMC4602297" ID-Pensoft-Pub="1313-2970-523-63" ID-PubMed="26478699" ID-ZBK="1E0C16D8DD11429BA6BC65862AA7C42B" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-523-63" ModsDocOrigin="ZooKeys 523" ModsDocTitle="Five new cryptic freshwater gastropod species from New Caledonia (Caenogastropoda, Truncatelloidea, Tateidae)" checkinTime="1451243964501" checkinUser="pensoft" docAuthor="Haase, Martin &amp; Zielske, Susan" docDate="2015" docId="3B3307D63110FCFA774403E77ECBF887" docLanguage="en" docName="ZooKeys 523: 63-87" docOrigin="ZooKeys 523" docSource="http://dx.doi.org/10.3897/zookeys.523.6066" docTitle="Leiorhagium adioincola Haase &amp; Zielske, 2015, sp. n." docType="treatment" docUuid="CCC4F863-76C3-44C2-A4AA-CE9DE0B726AB" docUuidSource="ZooBank" docVersion="4" lastPageNumber="68" masterDocId="FF8EE7237E0EFFF13504FFC5FFC9876C" masterDocTitle="Five new cryptic freshwater gastropod species from New Caledonia (Caenogastropoda, Truncatelloidea, Tateidae)" masterLastPageNumber="87" masterPageNumber="63" pageNumber="67" updateTime="1668160744127" updateUser="ExternalLinkService">
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<mods:title>Five new cryptic freshwater gastropod species from New Caledonia (Caenogastropoda, Truncatelloidea, Tateidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Haase, Martin</mods:namePart>
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<mods:role>
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<mods:namePart>Zielske, Susan</mods:namePart>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.523.6066</mods:identifier>
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<treatment ID-GBIF-Taxon="152062498" LSID="urn:lsid:zoobank.org:act:CCC4F863-76C3-44C2-A4AA-CE9DE0B726AB" httpUri="http://treatment.plazi.org/id/3B3307D63110FCFA774403E77ECBF887" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67">
<subSubSection pageId="4" pageNumber="67" type="multiple">
<paragraph pageId="4" pageNumber="67">
<pageBreakToken pageId="4" pageNumber="67" start="start">Taxon</pageBreakToken>
classification Animalia Littorinimorpha Tateidae
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="67" type="nomenclature">
<paragraph pageId="4" pageNumber="67">
<taxonomicName LSID="http://zoobank.org/CCC4F863-76C3-44C2-A4AA-CE9DE0B726AB" class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium adioincola" order="Littorinimorpha" pageId="4" pageNumber="67" phylum="Mollusca" rank="species" species="adioincola">Leiorhagium adioincola</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="67">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="67" type="type material">
<paragraph pageId="4" pageNumber="67">Type material.</paragraph>
<paragraph pageId="4" pageNumber="67">Holotype MNHN IM 2000-27860; paratypes MNHN IM 2000-27861 (29), NHMW 110182 (5).</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="67" type="type locality">
<paragraph pageId="4" pageNumber="67">Type locality.</paragraph>
<paragraph pageId="4" pageNumber="67">
NeCa 49, Poya: Massif
<normalizedToken originalValue="d’Adio">d'Adio</normalizedToken>
, stream flowing into Grotte
<normalizedToken originalValue="d’Adio">d'Adio</normalizedToken>
, open secondary forest,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-21.256779">21°15'24.4&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="165.24623">165°14'46.4&quot;E</geoCoordinate>
, 29 May 2012.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="67" type="materials_examined">
<paragraph pageId="4" pageNumber="67">Other material.</paragraph>
<paragraph pageId="4" pageNumber="67">
NeCa 43, Poya: small stream on W-side of road between
<normalizedToken originalValue="Nétéa">Netea</normalizedToken>
and Goipin, on forest edge,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-21.268333">21°16'06.0&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="165.24223">165°14'32.0&quot;E</geoCoordinate>
, 28 May 2012, MNHN-IM-2012-36075 (23), NHMW 110183 (10).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="67" type="etymology">
<paragraph pageId="4" pageNumber="67">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="67">Adioincola is composed of the name of the area of Adio and the Latin noun incola meaning inhabitant, and thus refers to the type locality of the new species.</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67" type="diagnosis">
<paragraph pageId="4" pageNumber="67">Diagnosis.</paragraph>
<paragraph lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67">
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium adioincola" order="Littorinimorpha" pageId="4" pageNumber="67" phylum="Mollusca" rank="species" species="adioincola">Leiorhagium adioincola</taxonomicName>
sp. n. is very similar to
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium kavuneva" order="Littorinimorpha" pageId="4" pageNumber="67" phylum="Mollusca" rank="species" species="kavuneva">Leiorhagium kavuneva</taxonomicName>
and
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium clandestinum" order="Littorinimorpha" pageId="4" pageNumber="67" phylum="Mollusca" rank="species" species="clandestinum">Leiorhagium clandestinum</taxonomicName>
sp. n. The former pair differs in penial shape, slender vs. basally broad with long terminal filament.
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium adioincola" order="Littorinimorpha" pageId="4" pageNumber="67" phylum="Mollusca" rank="species" species="adioincola">Leiorhagium adioincola</taxonomicName>
sp. n. tends to have fewer radular denticles than
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium kavuneva" order="Littorinimorpha" pageId="4" pageNumber="67" phylum="Mollusca" rank="species" species="kavuneva">Leiorhagium kavuneva</taxonomicName>
. Genetically, these species differed on average at 9.65% of the positions of COI. Due
<pageBreakToken pageId="5" pageNumber="68" start="start">to</pageBreakToken>
the lack of anatomical data, both new species can only be distinguished genetically. Their COI sequences differed on average by 9.5% (p-distance).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="68" type="shell">
<paragraph pageId="5" pageNumber="68">Shell.</paragraph>
<paragraph pageId="5" pageNumber="68">Pupiform, 1.8 times higher than wide, 4.125-4.75 whorls, without colour, transparent; protoconch faintly pitted with c. 1 whorl; palatal denticle a small droplet 1/8 whorl behind outer lip; aperture as high as wide (Figs 2B, 3C, D, 4C, D).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="68" type="operculum">
<paragraph pageId="5" pageNumber="68">Operculum.</paragraph>
<paragraph pageId="5" pageNumber="68">Elongate-ellipsoidal, paucisprial, nucleus submarginal, orange, usually two non-calcareous white pegs, eventually accompanied by a small third one (N=3) (Fig. 5C,D).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="68" type="external features">
<paragraph pageId="5" pageNumber="68">External features.</paragraph>
<paragraph pageId="5" pageNumber="68">Epidermis without pigment, eyes black.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="68" type="mantle cavity">
<paragraph pageId="5" pageNumber="68">Mantle cavity.</paragraph>
<paragraph pageId="5" pageNumber="68">Ctenidium with 18-19 (3 males) or 21-24 (2 females) filaments; osphradium kidney-shaped, behind middle of ctenidium.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="68" type="digestive system">
<paragraph pageId="5" pageNumber="68">Digestive system.</paragraph>
<paragraph pageId="5" pageNumber="68">Radula formula (N=3) (Fig. 6B): R: 4 1 4/2 2, L: 4-5 1 6, M1: 22-27, M2: 21-29; stomach without caecum; rectum close to pallial oviduct in females and to prostate in males.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="68" type="female genitalia">
<paragraph pageId="5" pageNumber="68">Female genitalia.</paragraph>
<paragraph pageId="5" pageNumber="68">Ovary without lobes, proximal end 1.25 whorls below apex, comprising 0.25-0.5 whorls, eventually reaching stomach; anterior capsule gland yellow-orange, posterior capsule gland opaque-white, albumen gland milky-white; proximal loop of renal oviduct bent forward, distal loop short; bursa copulatrix almost cubical, reaching behind albumen gland; bursal duct long, entering anterior; no seminal receptacle (N=2) (Fig. 7B).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="68" type="male genitalia">
<paragraph pageId="5" pageNumber="68">Male genitalia.</paragraph>
<paragraph pageId="5" pageNumber="68">Proximal end of lobate testis 1.25-1.5 whorls below apex, comprising 0.5-0.75 whorls, covering proximal end of stomach; vesicula seminalis arising from anterior half of testis; penis slender, terminal end occasionally forming short filament (N=3) (Fig. 8C).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="68" type="remarks">
<paragraph pageId="5" pageNumber="68">Remarks.</paragraph>
<paragraph pageId="5" pageNumber="68">
This is
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium" order="Littorinimorpha" pageId="5" pageNumber="68" phylum="Mollusca" rank="genus">Leiorhagium</taxonomicName>
sp. n. 4 of
<bibRefCitation author="Zielske, S" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="16" pageNumber="79" pagination="171 - 181" title="Molecular phylogeny and a modified approach of character-based barcoding refining the taxonomy of New Caledonian freshwater gastropods (Caenogastropoda, Truncatelloidea, Tateidae)." url="10.1016/j.ympev.2015.04.020" volume="89" year="2015">Zielske and Haase (2015)</bibRefCitation>
.
<taxonomicName class="Gastropoda" family="Tateidae" genus="Leiorhagium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiorhagium adioincola" order="Littorinimorpha" pageId="5" pageNumber="68" phylum="Mollusca" rank="species" species="adioincola">Leiorhagium adioincola</taxonomicName>
sp. n. occurs in the area between the villages of
<normalizedToken originalValue="Nétéa">Netea</normalizedToken>
and Goipin including the Massif
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<taxonomicName id="E3D436AEB6AE31D755ED2DADA1A1A56C" ID-CoL="5WWHM" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:26708" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus flavocrines Donisthorpe" order="Hymenoptera" pageNumber="11" phylum="Arthropoda" rank="species" species="flavocrines">
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flavocrines Donisthorpe
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<paragraph id="59B170EDEF408EF80BFE8BA563969D06" pageNumber="11">
<taxonomicName id="EB1A62ED964A02A849D6DE128FEFFBD5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:135363" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Camponotus (Myrmophyma) flavocrines Donisthorpe" order="Hymenoptera" pageNumber="11" phylum="Arthropoda" rank="species" species="flavocrines" subGenus="Myrmophyma">Camponotus (Myrmophyma) flavocrines Donisthorpe</taxonomicName>
1941b: 207.
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<typeStatus id="7EE1DB11DE3B9DB9F15279472B8CC689">Holotype</typeStatus>
worker from
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,
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(
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<paragraph id="C8BADE035C102104A7311B4E0332BEAE" pageNumber="11">Comments. This species is so far known only from the single type specimen.</paragraph>
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Subgenus
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<taxonomicName authorityName="Roeschke" authorityYear="1907" class="Insecta" family="Carabidae" genus="Neocychrus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="genus">Neocychrus</taxonomicName>
Roeschke, 1907a: 197. Type species:
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<emphasis italics="true" pageId="169" pageNumber="170">Cychrus angulatus</emphasis>
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Harris, 1839 designated by Lindroth (1961a: 24). Etymology. From the Greek prefix
<emphasis italics="true" pageId="169" pageNumber="170">neo</emphasis>
- (new) and the generic name
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<emphasis italics="true" pageId="169" pageNumber="170">Cychrus</emphasis>
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[
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<paragraph pageId="169" pageNumber="170">Identification.</paragraph>
<paragraph pageId="169" pageNumber="170">Van Dyke (1944) first reviewed the species. Subsequently, Gidaspow (1973) revised them and provided a key for their identification.</paragraph>
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