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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.37.6499" ID-PMC="PMC4023333" ID-Pensoft-Pub="1314-2003-37-25" ID-Pensoft-UUID="FFE8FFACFF84FFA95573FFFECD03F742" ID-PubMed="24843297" ID-Zenodo-Dep="576215" ModsDocID="1314-2003-37-25" checkinTime="1451251747243" checkinUser="pensoft" docAuthor="Bunwong, Sukhonthip, Chantaranothai, Pranom & Keeley, Sterling C." docDate="2014" docId="3B22DF557A05586DA1A9D058652473A3" docLanguage="en" docName="PhytoKeys 37: 25-101" docOrigin="PhytoKeys 37" docPubDate="2014-05-13" docSource="http://dx.doi.org/10.3897/phytokeys.37.6499" docTitle="Elephantopus scaber subsp. var. var. penicillatus Gagnep., Fl. Indo-Chine [M. H. Lecomte et al.] 3: 503. 1924." docType="treatment" docVersion="3" id="FFE8FFACFF84FFA95573FFFECD03F742" lastPageNumber="69" masterDocId="FFE8FFACFF84FFA95573FFFECD03F742" masterDocTitle="Revisions and key to the Vernonieae (Compositae) of Thailand" masterLastPageNumber="101" masterPageNumber="25" pageNumber="68" updateTime="1668141106797" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revisions and key to the Vernonieae (Compositae) of Thailand</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bunwong, Sukhonthip</mods:namePart>
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<mods:affiliation>Maejo University Phrae Campus, Mae Sai, Rong Kwang, Phrae 54140, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chantaranothai, Pranom</mods:namePart>
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<mods:affiliation>Applied Taxonomic Research Center, Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Keeley, Sterling C.</mods:namePart>
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<mods:affiliation>Department of Botany, University of Hawaii, Honolulu, HI 96816 USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-05-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>37</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>25</mods:start>
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<mods:end>101</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.37.6499</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.37.6499</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-37-25</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFE8FFACFF84FFA95573FFFECD03F742</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576215</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182228629" LSID="urn:lsid:plazi:treatment:3B22DF557A05586DA1A9D058652473A3" httpUri="http://treatment.plazi.org/id/3B22DF557A05586DA1A9D058652473A3" lastPageId="44" lastPageNumber="69" pageId="43" pageNumber="68">
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<subSubSection pageId="43" pageNumber="68" type="nomenclature">
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<paragraph pageId="43" pageNumber="68">
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<taxonomicName LSID="3B22DF55-7A05-586D-A1A9-D058652473A3" authority="Gagnep., Fl. Indo-Chine [M. H. Lecomte et al.] 3: 503. 1924." authorityName="Gagnep., Fl. Indo-Chine [M. H. Lecomte et al.] 3: 503. 1924." class="Dicotyledoneae" family="Asteraceae" genus="Elephantopus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Elephantopus scaber subsp. var. var. penicillatus" order="Campanulales" pageId="43" pageNumber="68" phylum="Angiospermae" rank="variety" species="scaber" subSpecies="var." variety="penicillatus">Elephantopus scaber var. penicillatus Gagnep., Fl. Indo-Chine [M.H. Lecomte et al.] 3: 503. 1924.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="68" type="type">
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<paragraph pageId="43" pageNumber="68">Type.</paragraph>
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<paragraph pageId="43" pageNumber="68">
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Laos, Chedom,
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<emphasis italics="true" pageId="43" pageNumber="68">Thorel</emphasis>
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1407 (holotype: K!).
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Morphology of Vernonieae in Thailand 4. A Elephantopus mollis B-C Elephantopus scaber D Elephantopus scaber var. penicillatus E-F Pseudelephantopus spicatus." httpUri="https://binary.pensoft.net/fig/10649" pageId="43" pageNumber="68">Fig. 8D</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="43" pageNumber="68" type="description">
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<paragraph pageId="43" pageNumber="68">Description.</paragraph>
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<paragraph pageId="43" pageNumber="68">Perennial herbs, 40-100 cm tall. Stems lacking except for inflorescence which is scapose, erect, terete, inconspicuously ribbed and pilose-villose. Leaves in a rosette at base of scape, 6-20 by 6-10 cm, obovate, obovate-lanceolate, elliptic, margin crenate to serrate, apex obtuse, base attenuate, subcoriaceous; upper surface sparsely pilose, without glands; lower surface densely pilose, with filiform cylindrical hairs and capitate glands; lateral veins 7-16-paired; petioles up to 6 cm long. Capitulescences terminal, scapose. Foliose bracts 3, deltoid. Capitula 7-9 mm long. Receptacle flat, ca. 1 mm in diam., glabrous. Involucres tubular, 6.5-8 mm long, 3-4 mm in diam. Phyllaries distichous, decussate, margin entire, outer surface puberulous, glands capitate; the outer ovate-lanceolate, apex acuminate; the inner ones lanceolate or oblong, apex acuminate. Florets 4; corollas salverform, white or purple, zygomorphic, glabrous or rarely hairy; corolla tubes 4-6 mm long; corolla lobes 2.5-3.5 mm long. Anthers ca. 2 mm long, apical appendage acute, base rounded. Styles purple, 6-8 mm long, branches ca. 1 mm long. Achenes 2-2.5 mm long, pubescent with a dense cover of twin hairs, without glands, 10-ribbed. Pappus of 5 bristles ca. 3 mm long, in one series, bristles with dilated bases.</paragraph>
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</subSubSection>
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<subSubSection pageId="44" pageNumber="69" type="distribution">
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<paragraph pageId="44" pageNumber="69">
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<pageBreakToken pageId="44" pageNumber="69" start="start">Distribution</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="44" pageNumber="69">Thailand: Loei, Nong Khai, Sakon Nakhon, Nakhon Phanom, Nakhon Ratchasima, Ubon Ratchathani, Chon Buri. Laos, Vietnam.</paragraph>
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</subSubSection>
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<subSubSection pageId="44" pageNumber="69" type="specimens examined">
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<paragraph pageId="44" pageNumber="69">Specimens examined.</paragraph>
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<paragraph pageId="44" pageNumber="69">
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Thailand. Udon Thani, Ban Phue district, Phu Phrabat historical park,
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<geoCoordinate degrees="17" direction="north" minutes="43.84" orientation="latitude" precision="9" value="17.730667">17°43.84'N</geoCoordinate>
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,
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<geoCoordinate degrees="102" direction="east" minutes="29.65" orientation="longitude" precision="9" value="102.49416">102°29.65'E</geoCoordinate>
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, 29 Sep 2007,
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<emphasis italics="true" pageId="44" pageNumber="69">S. Bunwong</emphasis>
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326 (KKU, USLoei, Phu Kra Dung, 17 Aug 1989,
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<emphasis italics="true" pageId="44" pageNumber="69">Din</emphasis>
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174 (BKF); Phu Kra Dung, 22 Oct 1990,
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<emphasis italics="true" pageId="44" pageNumber="69">Dee</emphasis>
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20 (BKF); Nong Khai, Sang Khom, 16 Dec 1982,
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<emphasis italics="true" pageId="44" pageNumber="69">H. Koyama, H. Terao & Th. Wongprasert</emphasis>
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T-31139 (BKF); Sakon Nakhon, Phu Phan national park, 5 Oct 2007,
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<emphasis italics="true" pageId="44" pageNumber="69">S. Bunwong</emphasis>
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333 (KKU, US); Phu Phan national park, 13 Nov 1984,
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<emphasis italics="true" pageId="44" pageNumber="69">G. Murata, C. Phengklai, S. Mitsuta, T. Yahara, H. Nagamasu & N. Nantasan</emphasis>
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T-50689 (BKF); Nakhon Phanom, 14 Dec 1982,
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<emphasis italics="true" pageId="44" pageNumber="69">H. Koyama, H. Terao & Th. Wongprasert</emphasis>
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T-31019 (BKF); Ubon Ratchathani, Route number 2222, 27 Oct 2007,
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<emphasis italics="true" pageId="44" pageNumber="69">S. Bunwong</emphasis>
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345 (KKU, US); Kalasin, Sahatsakhan, 22 Oct 1975,
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<emphasis italics="true" pageId="44" pageNumber="69">B. Sangkhachand & S. Sutheesorn</emphasis>
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3528 (BK); Nakhon Ratchasima, Bau Yai, 31 Oct 1931,
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<emphasis italics="true" pageId="44" pageNumber="69">Put</emphasis>
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4216 (BM, BK, K, L); Chon Buri, Khao Khiew, 19 Oct 1975,
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<emphasis italics="true" pageId="44" pageNumber="69">J.F. Maxwell</emphasis>
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75-1040 (AAU, BK, BKF).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="44" pageNumber="69" type="diagnostic characters">
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<paragraph pageId="44" pageNumber="69">Diagnostic characters.</paragraph>
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<paragraph pageId="44" pageNumber="69">
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<taxonomicName class="Dicotyledoneae" family="Asteraceae" genus="Elephantopus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Campanulales" pageId="44" pageNumber="69" phylum="Angiospermae" rank="species" species="scaber">
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<emphasis italics="true" pageId="44" pageNumber="69">Elephantopus scaber</emphasis>
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</taxonomicName>
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var.
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<emphasis italics="true" pageId="44" pageNumber="69">penicillatus</emphasis>
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differs from the typical variety in having large capitula and whitish pilose-tomentose leaf surface and involucres.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="44" pageNumber="69" type="ecology">
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<paragraph pageId="44" pageNumber="69">Ecology.</paragraph>
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<paragraph pageId="44" pageNumber="69">Dipterocarp or dry evergreen forest, alt. 100-400 m; flowering August to January.</paragraph>
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</subSubSection>
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<subSubSection pageId="44" pageNumber="69" type="vernacular name">
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<paragraph pageId="44" pageNumber="69">Vernacular name.</paragraph>
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<paragraph pageId="44" pageNumber="69">Doo Mai Ru Lom (โด่ไม่รู้ล้ม), Kee Fai Nok Khum (ขี้ไฟนกคุ้ม).</paragraph>
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</subSubSection>
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</treatment>
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</document>
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