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<document ID-DOI="http://dx.doi.org/10.3897/jor.31.73800" ID-GBIF-Dataset="34ac9f69-6c5b-42a3-a3f6-37c26a0f52bf" ID-GBIF-Taxon="193418338" ID-Pensoft-Pub="1937-2426-1-9" ID-Pensoft-UUID="DF53616E0253557E849F8362F9D7BA1E" ID-ZooBank="3906D11118494F9B87FDF70673B1B60E" ModsDocID="1937-2426-31-1-9" checkinTime="1645432308379" checkinUser="pensoft" docAuthor="Tan, Ming Kai &amp; Robillard, Tony" docDate="2022" docId="09E91DE8334954E88A93081E0D7A58CD" docLanguage="en" docName="JourOrthoptRes 31(1): 9-40" docOrigin="Journal of Orthoptera Research 31 (1)" docPubDate="2022-02-21" docSource="http://dx.doi.org/10.3897/jor.31.73800" docTitle="Rugabinthus leopoldi Tan &amp; Robillard 2022, comb. nov." docType="treatment" docVersion="4" id="DF53616E0253557E849F8362F9D7BA1E" lastPageNumber="9" masterDocId="DF53616E0253557E849F8362F9D7BA1E" masterDocTitle="Rugabinthus, a new genus of Lebinthina (Orthoptera, Gryllidae, Eneopterinae) from New Guinea" masterLastPageNumber="40" masterPageNumber="9" pageId="0" pageNumber="9" updateTime="1645436839305" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Rugabinthus, a new genus of Lebinthina (Orthoptera, Gryllidae, Eneopterinae) from New Guinea</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tan, Ming Kai</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4324-6305</mods:nameIdentifier>
<mods:affiliation>Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum national d'Histoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
<mods:nameIdentifier type="email">orthoptera.mingkai@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Robillard, Tony</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2177-9549</mods:nameIdentifier>
<mods:affiliation>Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum national d'Histoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Orthoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-02-21</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>31</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>9</mods:start>
<mods:end>40</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/jor.31.73800</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jor.31.73800</mods:identifier>
<mods:identifier type="Pensoft-Pub">1937-2426-1-9</mods:identifier>
<mods:identifier type="ZooBank">3906D11118494F9B87FDF70673B1B60E</mods:identifier>
<mods:identifier type="Pensoft-UUID">DF53616E0253557E849F8362F9D7BA1E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="193418338" LSID="urn:lsid:plazi:treatment:09E91DE8334954E88A93081E0D7A58CD" httpUri="http://treatment.plazi.org/id/09E91DE8334954E88A93081E0D7A58CD" lastPageNumber="9" pageId="0" pageNumber="9">
<subSubSection pageId="0" pageNumber="9" type="nomenclature">
<paragraph pageId="0" pageNumber="9">
<taxonomicName LSID="09E91DE8-3349-54E8-8A93-081E0D7A58CD" authority="(Chopard, 1931)" authorityName="Tan &amp; Robillard" authorityYear="2022" baseAuthorityName="Chopard" baseAuthorityYear="1931" family="Gryllidae" genus="Rugabinthus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rugabinthus leopoldi" order="Orthoptera" pageId="0" pageNumber="9" rank="species" species="leopoldi" status="comb. nov.">Rugabinthus leopoldi (Chopard, 1931)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="9">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9" type="description">
<paragraph pageId="0" pageNumber="9">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. R. leopoldi (Chopard, 1931) comb. nov. male (A, B) and female (C, D) habitus in dorsal (A, C) and lateral (B, D) views. Paratype labels (E). Scale bar: 10 mm." figureDoi="10.3897/jor.31.73800.figure1" httpUri="https://binary.pensoft.net/fig/647922" pageId="0" pageNumber="9">(Figs 1</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A. Map of New Guinea island showing the distribution of Rugabinthus species; B. Female Rugabinthus species in their natural habitats in Acemo, South Manokwari (https: // www. inaturalist. org / observations / 73044465) and C. Malagufuk (https: // www. inaturalist. org / observations / 72638321). Photo credit: Benoit Segerer." figureDoi="10.3897/jor.31.73800.figure3" httpUri="https://binary.pensoft.net/fig/647924" pageId="0" pageNumber="9">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Head and pronotum in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 5 mm." figureDoi="10.3897/jor.31.73800.figure4" httpUri="https://binary.pensoft.net/fig/647925" pageId="0" pageNumber="9">, 4A</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Face: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 2 mm." figureDoi="10.3897/jor.31.73800.figure5" httpUri="https://binary.pensoft.net/fig/647926" pageId="0" pageNumber="9">, 5A</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and pronotum in lateral views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 5 mm." figureDoi="10.3897/jor.31.73800.figure6" httpUri="https://binary.pensoft.net/fig/647927" pageId="0" pageNumber="9">, 6A</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male FW in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. karimui sp. nov. Scale bar: 2 mm." figureDoi="10.3897/jor.31.73800.figure7" httpUri="https://binary.pensoft.net/fig/647928" pageId="0" pageNumber="9">, 7A</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male genitalia in dorsal view; posterior apex of pseudepiphallus: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. karimui sp. nov. Scale bar: 0.5 mm." figureDoi="10.3897/jor.31.73800.figure8" httpUri="https://binary.pensoft.net/fig/647929" pageId="0" pageNumber="9">, 8A</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female FW in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. faowi sp. nov.; D. R. kencana sp. nov.; E. R. manokwari sp. nov.; F. R. nabire sp. nov.; G. R. tariku sp. nov.; H. R. albatros sp. nov.; I. R. baduri sp. nov.; J. R. karimui sp. nov.; K. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 2 mm." figureDoi="10.3897/jor.31.73800.figure9" httpUri="https://binary.pensoft.net/fig/647930" pageId="0" pageNumber="9">, 9A</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female copulatory papilla in ventral (all except I, K) and lateral (I, K) views. A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. faowi sp. nov.; D. R. kencana sp. nov.; E. R. manokwari sp. nov.; F. R. nabire sp. nov.; G. R. tariku sp. nov.; H, I. R. albatros sp. nov.; J. R. baduri sp. nov.; K, L. R. karimui sp. nov.; M. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 0.5 mm." figureDoi="10.3897/jor.31.73800.figure10" httpUri="https://binary.pensoft.net/fig/647931" pageId="0" pageNumber="9">, 10A</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male genitalia ventral (A, C, E) and lateral (B, D, F) views: A, B. R. leopoldi (Chopard, 1931) comb. nov.; C, D. R. mamberamo sp. nov.; E, F. R. yayukae sp. nov. Scale bar: 1 mm." figureDoi="10.3897/jor.31.73800.figure11" httpUri="https://binary.pensoft.net/fig/647932" pageId="0" pageNumber="9">, 11A</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male genitalia ventral (A, C, E) and lateral (B, D, F) views: A, B. R. leopoldi (Chopard, 1931) comb. nov.; C, D. R. mamberamo sp. nov.; E, F. R. yayukae sp. nov. Scale bar: 1 mm." figureDoi="10.3897/jor.31.73800.figure11" httpUri="https://binary.pensoft.net/fig/647932" pageId="0" pageNumber="9">, 11B)</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9" type="reference_group">
<paragraph pageId="0" pageNumber="9">
<taxonomicName authorityName="Tan &amp; Robillard" authorityYear="2022" baseAuthorityName="Tan &amp; Robillard" baseAuthorityYear="2022" class="Insecta" family="Gryllidae" genus="Lebinthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lebinthus leopoldi" order="Orthoptera" pageId="0" pageNumber="9" phylum="Arthropoda" rank="species" species="leopoldi" status="comb. nov.">Lebinthus leopoldi</taxonomicName>
Chopard, 1931: 7; 1968: 354;
<bibRefCitation author="Cigliano, MM" journalOrPublisher="Volume 12. Uitgeverij Dr. W. Junk N. V. ' s, Gravenhage" pageId="0" pageNumber="9" publicationUrl="http://Orthoptera.SpeciesFile.org" refId="B8" refString="Cigliano, MM, Braun, H, Eades, DC, Otte, D, 2021. Orthoptera Species File. Version 5.0/5.0 (01/01/2021). http://Orthoptera.SpeciesFile.org" title="Orthoptera Species File. Version 5.0 / 5.0 (01 / 01 / 2021)." url="http://Orthoptera.SpeciesFile.org" year="2021">Cigliano et al. 2021</bibRefCitation>
(
<taxonomicName authorityName="Tan &amp; Robillard" authorityYear="2022" baseAuthorityName="Tan &amp; Robillard" baseAuthorityYear="2022" class="Insecta" family="Gryllidae" genus="Lebinthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="9" phylum="Arthropoda" rank="order" status="comb. nov.">Orthoptera</taxonomicName>
species file online).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9" type="materials_examined">
<paragraph pageId="0" pageNumber="9">
<emphasis italics="true" pageId="0" pageNumber="9">Material examined.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="9">
<materialsCitation collectingDate="1929-03-02" collectorName="Prince Leopold" country="INDONESIA" location="Manoi" specimenCount="♂" stateProvince="West Papua" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="9">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
<collectingCountry name="Indonesia">INDONESIA</collectingCountry>
•
<specimenCount>♂</specimenCount>
;
<collectingRegion country="Indonesia" name="Papua Barat">West Papua</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:09E91DE8334954E88A93081E0D7A58CD:44A0991416E4F011D21A104E60086414" country="INDONESIA" name="Manoi" stateProvince="West Papua">Manoi</location>
[Sorong Manoi];
<collectingDate value="1929-03-02">2 March 1929</collectingDate>
;
<collectorName>Prince Leopold</collectorName>
leg.; RBINS
</materialsCitation>
.
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="INDONESIA" location="INDONESIA" specimenCount="♀" typeStatus="Allotype">
<emphasis bold="true" pageId="0" pageNumber="9">
<typeStatus>Allotype</typeStatus>
</emphasis>
:
<collectingCountry name="Indonesia">INDONESIA</collectingCountry>
•
<specimenCount>♀</specimenCount>
; same information as
<typeStatus>holotype</typeStatus>
; RBINS
</materialsCitation>
.
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="INDONESIA" location="INDONESIA" specimenCount="7" specimenCount-female="4" specimenCount-male="3" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="9">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
<collectingCountry name="Indonesia">INDONESIA</collectingCountry>
•
<specimenCount type="male">3♂</specimenCount>
,
<specimenCount type="female">4♀</specimenCount>
; same information as
<typeStatus>holotype</typeStatus>
; RBINS
</materialsCitation>
•
<materialsCitation collectingDate="1929-03-02" collectorName="Prince Leopold" country="Indonesia" location="Manoi" specimenCount="1" specimenCount-male="1" stateProvince="West Papua" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
;
<collectingRegion country="Indonesia" name="Papua Barat">West Papua</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:09E91DE8334954E88A93081E0D7A58CD:AA3470F995D422C2EB5ACEC2609F9EAA" country="Indonesia" name="Manoi" stateProvince="West Papua">Manoi</location>
[Sorong Manoi];
<collectingDate value="1929-03-02">2 March 1929</collectingDate>
;
<collectorName>Prince Leopold</collectorName>
leg.; MNHN-EO-ENSIF1441
</materialsCitation>
•
<specimenCount type="female">1♀</specimenCount>
; same information as
<typeStatus>holotype</typeStatus>
; molecular sample L173; MNHN-EO-ENSIF1443 •
<specimenCount type="female">1♀</specimenCount>
; same information as
<typeStatus>holotype</typeStatus>
; MNHN-EO-ENSIF1442.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9" type="type locality">
<paragraph pageId="0" pageNumber="9">
<emphasis italics="true" pageId="0" pageNumber="9">Type locality.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="9">INDONESIA: West Papua: Manoi</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9" type="diagnosis">
<paragraph pageId="0" pageNumber="9">
<emphasis italics="true" pageId="0" pageNumber="9">Diagnosis.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="9">This species differs from all congeners by male genitalia with pseudepiphallus more rectangular, very elongate, its posterior part curved posteriorly, its apex truncated with small paired apical lophi; lophi triangular with obtuse apex.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9" type="redescription">
<paragraph pageId="0" pageNumber="9">
<emphasis italics="true" pageId="0" pageNumber="9">Redescription.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="9">
Average size among congeners (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. R. leopoldi (Chopard, 1931) comb. nov. male (A, B) and female (C, D) habitus in dorsal (A, C) and lateral (B, D) views. Paratype labels (E). Scale bar: 10 mm." figureDoi="10.3897/jor.31.73800.figure1" httpUri="https://binary.pensoft.net/fig/647922" pageId="0" pageNumber="9">1</figureCitation>
). Dorsum of head with broad red brown bands barely separated (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Head and pronotum in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 5 mm." figureDoi="10.3897/jor.31.73800.figure4" httpUri="https://binary.pensoft.net/fig/647925" pageId="0" pageNumber="9">4A</figureCitation>
). Fastigium red brown (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Head and pronotum in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 5 mm." figureDoi="10.3897/jor.31.73800.figure4" httpUri="https://binary.pensoft.net/fig/647925" pageId="0" pageNumber="9">4A</figureCitation>
). Scapes red brown with some dark patches. Fastigium verticis brown with two vertical yellow brown stripes diverging ventrad, frons brown with two yellow brown spots ventral of stripes; clypeus sometimes darkened, mouthparts yellow brown to red brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Face: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 2 mm." figureDoi="10.3897/jor.31.73800.figure5" httpUri="https://binary.pensoft.net/fig/647926" pageId="0" pageNumber="9">5A</figureCitation>
). Pronotal disk red brown with lighter lateral ovular spots near anterior half and some faint lighter spots along posterior margin (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Head and pronotum in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 5 mm." figureDoi="10.3897/jor.31.73800.figure4" httpUri="https://binary.pensoft.net/fig/647925" pageId="0" pageNumber="9">4A</figureCitation>
). Lateral lobes of pronotum dark brown, with yellow spot at anterior ventral corner (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head and pronotum in lateral views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. baduri sp. nov.; M. R. karimui sp. nov.; N. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 5 mm." figureDoi="10.3897/jor.31.73800.figure6" httpUri="https://binary.pensoft.net/fig/647927" pageId="0" pageNumber="9">6A</figureCitation>
). FIs and FIIs generally yellow brown with brown spots and rings; TIs and TIIs brown with two yellow brown rings. FIIIs brown, knees slightly darker.
</paragraph>
<paragraph pageId="0" pageNumber="9">
<emphasis bold="true" pageId="0" pageNumber="9">Male.</emphasis>
FW reaching apex of third tergite; apex rounded. FW coloration (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male FW in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. karimui sp. nov. Scale bar: 2 mm." figureDoi="10.3897/jor.31.73800.figure7" httpUri="https://binary.pensoft.net/fig/647928" pageId="0" pageNumber="9">7A</figureCitation>
): Dorsal field cells and veins mostly brown; area between M and R yellow brown; basal area with a wide cream-colored spot on external corner. Lateral field dark brown. FW venation typical of genus; 1A slightly bisinuate anterior to angle; oblique vein trifurcated, posterior branch straight and transverse.
</paragraph>
<paragraph pageId="0" pageNumber="9">
<emphasis italics="true" pageId="0" pageNumber="9">Male genitalia</emphasis>
: (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male genitalia in dorsal view; posterior apex of pseudepiphallus: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. karimui sp. nov. Scale bar: 0.5 mm." figureDoi="10.3897/jor.31.73800.figure8" httpUri="https://binary.pensoft.net/fig/647929" pageId="0" pageNumber="9">8A</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male genitalia ventral (A, C, E) and lateral (B, D, F) views: A, B. R. leopoldi (Chopard, 1931) comb. nov.; C, D. R. mamberamo sp. nov.; E, F. R. yayukae sp. nov. Scale bar: 1 mm." figureDoi="10.3897/jor.31.73800.figure11" httpUri="https://binary.pensoft.net/fig/647932" pageId="0" pageNumber="9">11A</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male genitalia ventral (A, C, E) and lateral (B, D, F) views: A, B. R. leopoldi (Chopard, 1931) comb. nov.; C, D. R. mamberamo sp. nov.; E, F. R. yayukae sp. nov. Scale bar: 1 mm." figureDoi="10.3897/jor.31.73800.figure11" httpUri="https://binary.pensoft.net/fig/647932" pageId="0" pageNumber="9">11B</figureCitation>
) Pseudepiphallus forming a very elongate rectangle, its basal margin slightly indented in the middle, lateral margins widened basally, forming wide shoulders carrying base of rami; posterior part of pseudepiphallus narrow, distinctly curved dorsally posterior to pseudepiphallic parameres, with two short dorsal pre-apical expansions (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male genitalia in dorsal view; posterior apex of pseudepiphallus: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. yayukae sp. nov.; D. R. faowi sp. nov.; E. R. kencana sp. nov.; F. R. manokwari sp. nov.; G. R. maoke sp. nov.; H. R. biakis sp. nov.; I. R. nabire sp. nov.; J. R. tariku sp. nov.; K. R. albatros sp. nov.; L. R. karimui sp. nov. Scale bar: 0.5 mm." figureDoi="10.3897/jor.31.73800.figure8" httpUri="https://binary.pensoft.net/fig/647929" pageId="0" pageNumber="9">8A</figureCitation>
); posterior apex truncated, posterior apex with small paired lophi, triangular with obtuse apex. Rami wide, very short, way shorter than half the pseudepiphallus length, diverging anteriorly, with posterior end protruding externally. Pseudepiphallic parameres strongly bent in basal half (~90°), with posterior apex enlarged, bean shaped. Ectophallic apodemes thin, parallel, and long, reaching beyond anterior margin of pseudepiphallic sclerite. Endophallic sclerite with anterior region very short and not reaching anterior margin of pseudepiphallic sclerite, with lateral arms elongated, tongue-shaped, and longer than median expansion.
</paragraph>
<paragraph pageId="0" pageNumber="9">
<emphasis bold="true" pageId="0" pageNumber="9">Female.</emphasis>
FW reaching base of third tergite, with basal area with a cream-colored oblong spot without clearly defined margin, forming two indistinct spots near base and apex; lateral field dark brown (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female FW in dorsal views: A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. faowi sp. nov.; D. R. kencana sp. nov.; E. R. manokwari sp. nov.; F. R. nabire sp. nov.; G. R. tariku sp. nov.; H. R. albatros sp. nov.; I. R. baduri sp. nov.; J. R. karimui sp. nov.; K. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 2 mm." figureDoi="10.3897/jor.31.73800.figure9" httpUri="https://binary.pensoft.net/fig/647930" pageId="0" pageNumber="9">9A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="9">
<emphasis italics="true" pageId="0" pageNumber="9">Female genitalia</emphasis>
: Ovipositor about as long as FIII. Copulatory papilla very small, rounded, apex folded ventrally, short, pointed; dorsal face with a sclerotized area; ventro-anterior end base forming an oval ring (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female copulatory papilla in ventral (all except I, K) and lateral (I, K) views. A. R. leopoldi (Chopard, 1931) comb. nov.; B. R. mamberamo sp. nov.; C. R. faowi sp. nov.; D. R. kencana sp. nov.; E. R. manokwari sp. nov.; F. R. nabire sp. nov.; G. R. tariku sp. nov.; H, I. R. albatros sp. nov.; J. R. baduri sp. nov.; K, L. R. karimui sp. nov.; M. R. newguineae (Bhowmik, 1981) comb. nov. Scale bar: 0.5 mm." figureDoi="10.3897/jor.31.73800.figure10" httpUri="https://binary.pensoft.net/fig/647931" pageId="0" pageNumber="9">10A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="9" type="description">
<paragraph pageId="0" pageNumber="9">
<emphasis italics="true" pageId="0" pageNumber="9">Measurements.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="9">
See Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements (in mm)." httpUri="http://table.plazi.org/id/1CD01F98C3B7E3BF17EF858517C029E3" pageId="0" pageNumber="9" tableUuid="1CD01F98C3B7E3BF17EF858517C029E3">1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

107
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<taxonomicName id="9048F64F6F9438FAD26164B3296B19DA" ID-CoL="99HXJ" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Asteraceae" genus="Filago" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="DEDC7C5E4E0F199DF30AF64706ED0682" bold="true" pageNumber="520">Filago montana</emphasis>
Linnaeus
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<emphasis id="C2068482D0D5B81C94212D8FA05DC8BF" italics="true" pageNumber="520">Species Plantarum</emphasis>
2
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: 857, 1230. 1753
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.
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<paragraph id="D6ED185AF8FF4EF1E9786B0231AB69C5" pageNumber="520">&quot;Habitat in Europae collibus sabulosis.&quot; RCN: 6707.</paragraph>
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<paragraph id="BDCECF2434C7216F05F55D49090348F2" pageNumber="520">Type not designated.</paragraph>
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Original material:
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Herb. Burser XV(1): 10 (
<collectionCode id="5F086820C7F387F14EBE810AA575C898">UPS</collectionCode>
)
</materialsCitation>
;
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Herb. Linn. No. 1041.8 (
<collectionCode id="91770A8A588B21F1B656C85984F27F78">LINN</collectionCode>
)
</materialsCitation>
; [icon] in Bauhin &amp; Cherler, Hist. Pl. Univ. 3(1): 159. 1651; [icon] in Plantin, Pl. Stirp. Icon.: 481. 1581.
</paragraph>
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<paragraph id="30F2E1B1675EA99C26C38AD1D8D5699E" pageNumber="521">
Current name:
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<emphasis id="773A9B07CDDE2347D656A3CFE7638805" italics="true" pageNumber="521">
<emphasis id="90F2617036E5014FE7BC98088FF7317A" bold="true" italics="true" pageNumber="521">Filago arvensis</emphasis>
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L.
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<normalizedToken id="1B95B062EF52545B758093D882372132" originalValue="×">x</normalizedToken>
<emphasis id="70DF5911D1ED16B1E07952C8D9300C8A" italics="true" pageNumber="521">
<emphasis id="CFE2DB3851A2178FA36F22222711B795" bold="true" italics="true" pageNumber="521">minima</emphasis>
</emphasis>
(Sm.) Pers. (
<taxonomicName id="CBDCC7BFA90BB3D4657B216B8F7C6192" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
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<paragraph id="73A0ECE1165BE20E24AA388F12F43AB1" pageNumber="521">
<emphasis id="A9503E2E4A8B0A33FC50D6838E2E1C65" italics="true" pageNumber="521">Note:</emphasis>
See comments by Greuter (in
<emphasis id="4AF97EDCC42CBAEDE5F8CF57262BB6D1" italics="true" pageNumber="521">Boissiera</emphasis>
13: 137. 1967).
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<taxonomicName LSID="http://zoobank.org/D28FBB73-6436-4F61-91BF-B300A901764C" authority="Brunke" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus temburong" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="temburong">Bolitogyrus temburong Brunke</taxonomicName>
<taxonomicNameLabel pageId="21" pageNumber="22">sp. n.</taxonomicNameLabel>
Figs 10
<normalizedToken originalValue="H–J">H-J</normalizedToken>
, 20A (map)
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="type material">
<paragraph pageId="22" pageNumber="23">
<pageBreakToken pageId="22" pageNumber="23" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph pageId="22" pageNumber="23">
Holotype (♂, BMNH): BRUNEI, E
<geoCoordinate direction="north" orientation="latitude" precision="925" value="115.11667">115°7' N</geoCoordinate>
4°34', Kuala Belalong FSC, Dipterocarp forest, BM(NH) 1991-173 [printed] / Ground Malaise, 20A, 770m alt, 7.ii.92, N. Mawdaley NM302 [printed] / Holotype ♂,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Cyrtothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtothorax temburong" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="temburong">Cyrtothorax temburong</taxonomicName>
Brunke, des. A. Brunke 2017 [red printed label] / AJB0000401 [identifier label].
</paragraph>
<paragraph pageId="22" pageNumber="23">Paratypes (1 ♂ 1 ♀, BMNH): same as holotype except, 440 m, ground malaise #13, 8.II.1992, AJB0000499; same as holotype except, 610 m, ground malaise #18, AJB0000500.</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="diagnosis">
<paragraph pageId="22" pageNumber="23">Diagnosis.</paragraph>
<paragraph pageId="22" pageNumber="23">Among the members of the Caesareus Group: pronotum not entirely pale; elytra without dark spot (Fig. 1D); first two visible abdominal segments with distinct darkened area (Fig. 1D); median lobe in parameral view dilated to blunt apex (Fig. 10H).</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="description">
<paragraph pageId="23" pageNumber="24">
<pageBreakToken pageId="23" pageNumber="24" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="23" pageNumber="24">Measurements ♂ (n = 2): HW/HL 1.37-1.44; PW/PL 1.43-1.52; EW/ EL 1.29-1.31; ESut/PL 0.93-0.94; PW/HW 1.13-1.13; forebody length 5.7-6.2 mm.</paragraph>
<paragraph pageId="23" pageNumber="24">Measurements ♀ (n = 1): HW/HL 1.33; PW/PL 1.64; EW/ EL 1.30; ESut/PL 0.95; PW/HW 1.16; forebody length 6.0 mm.</paragraph>
<paragraph pageId="23" pageNumber="24">
Almost identical to
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus proximus" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="proximus">Bolitogyrus proximus</taxonomicName>
except: pronotum entirely black or with anterior angles orange, though variation similar to
<taxonomicName lsidName="B. proximus" pageId="23" pageNumber="24" rank="species" species="proximus">B. proximus</taxonomicName>
may occur with more specimens; elytra slightly wider; elytra slightly longer proportionally than pronotum; median lobe in lateral view with apical portion thicker and flexed ventrad, median tooth situated more distally (Fig. 10I); median lobe in parameral view widened to a rounded, obtuse apex with outline bearing median tooth (Fig. 10H); paramere with more elongate and blunt apex (Fig. 10J).
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="distribution">
<paragraph pageId="23" pageNumber="24">Distribution.</paragraph>
<paragraph pageId="23" pageNumber="24">Figure 20A. Currently known only from Brunei but probably more widely distributed at medium-low elevations in Borneo.</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="bionomics">
<paragraph pageId="23" pageNumber="24">Bionomics.</paragraph>
<paragraph pageId="23" pageNumber="24">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus temburong" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="temburong">Bolitogyrus temburong</taxonomicName>
has been collected in February from lower montane forests at elevations ranging from 440-770m.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="etymology">
<paragraph pageId="23" pageNumber="24">Etymology.</paragraph>
<paragraph pageId="23" pageNumber="24">The species epithet recognizes Ulu Temburong National Park in Brunei and its conservation achievements.</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="comments">
<paragraph pageId="23" pageNumber="24">Comments.</paragraph>
<paragraph pageId="23" pageNumber="24">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus temburong" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="temburong">Bolitogyrus temburong</taxonomicName>
likely represents the first of many undescribed
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Bolitogyrus</taxonomicName>
species at medium elevations in Borneo. Based on the samples seen by the author, very limited micro-fogging of fungusy logs has been conducted on Borneo.
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Klug, 1817
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Added by
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.
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