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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.245.3416" ID-PMC="PMC3577090" ID-Pensoft-Pub="1313-2970-245-1" ID-Pensoft-UUID="FFFF52503A0AFF882450FFB66D45FF8E" ID-PubMed="23431087" ID-Zenodo-Dep="578462" ModsDocID="1313-2970-245-1" checkinTime="1425642919252" checkinUser="pensoft" docAuthor="Bousquet, Yves" docDate="2012" docId="098F1B15147788C9A3969EFD305E21EB" docLanguage="en" docName="ZooKeys 245: 1-1722" docOrigin="ZooKeys 245" docPubDate="2012-11-28" docSource="http://dx.doi.org/10.3897/zookeys.245.3416" docTitle="Pterostichus coracinus" docType="treatment" docVersion="10" id="FFFF52503A0AFF882450FFB66D45FF8E" lastPageNumber="518" masterDocId="FFFF52503A0AFF882450FFB66D45FF8E" masterDocTitle="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico" masterLastPageNumber="1722" masterPageNumber="1" pageNumber="517" updateTime="1668154792637" updateUser="ExternalLinkService">
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<mods:title>Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bousquet, Yves</mods:namePart>
<mods:affiliation>Agriculture and Agri-Food Canada, Central Experimental Farm, Ottawa, Ontario, Canada</mods:affiliation>
<mods:nameIdentifier type="email">bousquety1@yahoo.com</mods:nameIdentifier>
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<mods:date>2012</mods:date>
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<mods:number>2012-11-28</mods:number>
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<mods:number>245</mods:number>
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<treatment ID-GBIF-Taxon="127868281" LSID="urn:lsid:plazi:treatment:098F1B15147788C9A3969EFD305E21EB" httpUri="http://treatment.plazi.org/id/098F1B15147788C9A3969EFD305E21EB" lastPageId="517" lastPageNumber="518" pageId="516" pageNumber="517">
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<taxonomicName LSID="098F1B15-1477-88C9-A396-9EFD305E21EB" authority="(Newman, 1838)" baseAuthorityName="Newman" baseAuthorityYear="1838" class="Hexapoda" family="Carabidae" genus="Pterostichus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pterostichus coracinus" order="Coleoptera" pageId="516" pageNumber="517" phylum="Arthropoda" rank="species" species="coracinus">Pterostichus coracinus (Newman, 1838)</taxonomicName>
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<subSubSection lastPageId="517" lastPageNumber="518" pageId="516" pageNumber="517" type="reference_group">
<paragraph pageId="516" pageNumber="517">
<taxonomicName authorityName="Newman" authorityYear="1838" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="516" pageNumber="517" phylum="Arthropoda" rank="species" species="coracina">Feronia coracina</taxonomicName>
Newman, 1838a: 386. Type locality: &quot;northern states of America&quot; (original citation, see page 388), restricted to &quot;Rumney [Grafton County], N[ew] H[ampshire]&quot; by Lindroth (1966: 494). Syntype(s) lost (Lindroth 1966: 494).
</paragraph>
<paragraph pageId="516" pageNumber="517">
<taxonomicName authorityName="Newman" authorityYear="1838" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="516" pageNumber="517" phylum="Arthropoda" rank="species" species="monedula">Feronia monedula</taxonomicName>
Newman, 1838a: 386 [
<emphasis italics="true" pageId="516" pageNumber="517">nomen dubium</emphasis>
]. Type locality: &quot;northern states of America&quot; (original citation, see page 388). Syntype(s) probably lost (Lindroth 1966: 494). Synonymy established with doubt by LeConte (1873a: 317).
</paragraph>
<paragraph pageId="516" pageNumber="517">
<taxonomicName authorityName="Newman" authorityYear="1838" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="516" pageNumber="517" phylum="Arthropoda" rank="species" species="moerens">Feronia moerens</taxonomicName>
Newman, 1838a: 387. Type locality: &quot;northern states of America&quot; (original citation, see page 388). Lectotype (♂), designated by Lindroth (1966: 494), in BMNH. Synonymy established by LeConte (1870: 399), confirmed by Lindroth (1966: 494).
</paragraph>
<paragraph pageId="516" pageNumber="517">
<taxonomicName authorityName="LeConte" authorityYear="1853" class="Hexapoda" family="Carabidae" genus="Pterostichus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="516" pageNumber="517" phylum="Arthropoda" rank="species" species="adiunctus">Pterostichus adiunctus</taxonomicName>
LeConte, 1853a: 245. Type locality: &quot;Lake Superior&quot; (original citation). Lectotype (♀), designated by Bousquet (1999: 138), in MCZ [# 5631]. Synonymy established by LeConte (1870: 399), confirmed by Lindroth (1966: 494).
</paragraph>
<paragraph pageId="516" pageNumber="517">
<taxonomicName class="Hexapoda" family="Carabidae" genus="Pterostichus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="516" pageNumber="517" phylum="Arthropoda" rank="species" species="flebilis">Pterostichus flebilis</taxonomicName>
LeConte, 1853a: 245. Type locality: &quot;Lake Superior&quot; (original citation). Holotype [by monotypy] (♂) in MCZ [# 5632]. Synonymy established by LeConte (1870: 399), confirmed by Lindroth (1966: 494).
</paragraph>
<paragraph pageId="516" pageNumber="517">
<taxonomicName authorityName="Casey" authorityYear="1913" class="Hexapoda" family="Carabidae" genus="Pterostichus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="516" pageNumber="517" phylum="Arthropoda" rank="species" species="erebeus">Pterostichus erebeus</taxonomicName>
Casey, 1913: 134. Type locality: &quot;Bayfield [Bayfield County], Wisconsin&quot; (original citation). Lectotype (♀), designated by Lindroth (1975: 124), in USNM [# 47063]. Synonymy established by Lindroth (1966: 494).
</paragraph>
<paragraph pageId="517" pageNumber="518">
<taxonomicName authorityName="Casey" authorityYear="1920" class="Insecta" family="Carabidae" genus="Euferonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="517" pageNumber="518" phylum="Arthropoda" rank="species" species="roanica">
<pageBreakToken pageId="517" pageNumber="518" start="start">Euferonia</pageBreakToken>
roanica
</taxonomicName>
Casey, 1920: 188. Type locality: &quot;Roan Mountain, North Carolina&quot; (original citation). Lectotype (♂), designated by Lindroth (1975: 124), in USNM [# 47059]. Synonymy established by Lindroth (1966: 494).
</paragraph>
<paragraph pageId="517" pageNumber="518">
<taxonomicName class="Insecta" family="Carabidae" genus="Euferonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="517" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="coracina" subSpecies="venator">Euferonia coracina venator</taxonomicName>
Casey, 1920: 189. Type locality: &quot;Danville [Montour County], Pennsylvania&quot; (original citation). Lectotype (♂), designated by Lindroth (1975: 124), in USNM [# 47060]. Synonymy established by Nicolay and Weiss (1934: 204), confirmed by Lindroth (1966: 494).
</paragraph>
<paragraph pageId="517" pageNumber="518">
<taxonomicName class="Insecta" family="Carabidae" genus="Euferonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="517" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="coracina" subSpecies="ludibunda">Euferonia coracina ludibunda</taxonomicName>
Casey, 1920: 189. Type locality: &quot;Buena Vista Spring, Franklin Co[unty], Pennsylvania&quot; (original citation). Lectotype (♀), designated by Lindroth (1975: 124), in USNM [# 47061]. Synonymy established, under the name
<taxonomicName class="Hexapoda" family="Carabidae" genus="Pterostichus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="517" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="coracinus" subSpecies="erebeus">
<emphasis italics="true" pageId="517" pageNumber="518">Pterostichus coracinus erebeus</emphasis>
</taxonomicName>
Casey, by Nicolay and Weiss (1934: 206), confirmed by Lindroth (1966: 494).
</paragraph>
<paragraph pageId="517" pageNumber="518">
<taxonomicName authorityName="Casey" authorityYear="1924" class="Insecta" family="Carabidae" genus="Euferonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="517" pageNumber="518" phylum="Arthropoda" rank="species" species="lacustris">Euferonia lacustris</taxonomicName>
Casey, 1924: 71. Type locality: &quot;near Chicago, Illinois&quot; (original citation). Lectotype (♂), designated by Lindroth (1975: 124), in USNM [# 47062]. Synonymy established by Nicolay and Weiss (1934: 204), confirmed by Lindroth (1966: 494).
</paragraph>
<paragraph pageId="517" pageNumber="518">
<taxonomicName authorityName="Casey" authorityYear="1924" class="Insecta" family="Carabidae" genus="Euferonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="517" pageNumber="518" phylum="Arthropoda" rank="species" species="strigosula">Euferonia strigosula</taxonomicName>
Casey, 1924: 72. Type locality: &quot;Hagerstown [Washington County], Maryland&quot; (original citation). Lectotype (♂), designated by Lindroth (1975: 124), in USNM [# 47056]. Synonymy established, under the name
<taxonomicName class="Hexapoda" family="Carabidae" genus="Pterostichus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="517" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="coracinus" subSpecies="roanicus">
<emphasis italics="true" pageId="517" pageNumber="518">Pterostichus coracinus roanicus</emphasis>
</taxonomicName>
(Casey), by Nicolay and Weiss (1934: 205), confirmed by Lindroth (1966: 494).
</paragraph>
<paragraph pageId="517" pageNumber="518">
<taxonomicName authorityName="Nicolay &amp; Weiss" authorityYear="1934" class="Insecta" family="Carabidae" genus="Euferonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="517" pageNumber="518" phylum="Arthropoda" rank="species" species="washingtonensis">Euferonia washingtonensis</taxonomicName>
Nicolay and Weiss, 1934: 203. Type locality: &quot;slopes of M[oun]t Madison [Coos County], New Hampshire&quot; (original citation). Holotype (♂) in MCZ [# 22985]. Synonymy established by Lindroth (1966: 494).
</paragraph>
<paragraph pageId="517" pageNumber="518">
<taxonomicName authorityName="Nicolay &amp; Weiss" authorityYear="1934" class="Insecta" family="Carabidae" genus="Euferonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="517" pageNumber="518" phylum="Arthropoda" rank="species" species="washingtonensis">Euferonia washingtonensis</taxonomicName>
var.
<emphasis italics="true" pageId="517" pageNumber="518">rufitarsis</emphasis>
Nicolay and Weiss, 1934: 204. Type locality: &quot;Black Mountain, North Carolina&quot; (original citation). Holotype (♂) in MCZ [# 22986]. Synonymy established by Lindroth (1966: 494).
</paragraph>
</subSubSection>
<subSubSection pageId="517" pageNumber="518" type="distribution">
<paragraph pageId="517" pageNumber="518">Distribution.</paragraph>
<paragraph pageId="517" pageNumber="518">
This species is found from Newfoundland (Lindroth 1955a: 86) to eastern South Dakota (French et al. 2004: 557), north to the James Bay area in Quebec (Larochelle 1975: 86), south along the Appalachian Mountains to eastern Tennessee (Nicolay and Weiss 1934: 206, as
<taxonomicName class="Insecta" family="Carabidae" genus="Euferonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="517" pageNumber="518" phylum="Arthropoda" rank="subSpecies" species="coracina" subSpecies="roanica">
<emphasis italics="true" pageId="517" pageNumber="518">Euferonia coracina roanica</emphasis>
</taxonomicName>
) and northern Georgia (Fattig 1949: 24). The record from the Black Hills in southwestern South Dakota (Stockton 1954: 15) is doubtful.
</paragraph>
</subSubSection>
<subSubSection pageId="517" pageNumber="518" type="records">
<paragraph pageId="517" pageNumber="518">Records.</paragraph>
<paragraph pageId="517" pageNumber="518">
<emphasis bold="true" pageId="517" pageNumber="518">FRA</emphasis>
: PM
<emphasis bold="true" pageId="517" pageNumber="518">CAN</emphasis>
: LB, NB, NF, NS (CBI), ON, PE, QC
<emphasis bold="true" pageId="517" pageNumber="518">USA</emphasis>
: CT, DC, GA, IA, IL, IN, KY, MA, MD, ME, MI, MN, NC, NH, NJ, NY, OH, PA, RI, SC, SD, TN, VA, VT, WI, WV
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart id="E560152F4D8D00891EE44CEB0F6F21A1">Zazzo, Antoine</mods:namePart>
<mods:affiliation id="D4FD86E00DDCDDE463E4CC745C72F007">Archeozoologie, Archeobotanique, Societes, Pratiques et Environnements (AASPE), UMR 7209, Museum national d'Histoire naturelle, CNRS, Paris, France</mods:affiliation>
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<mods:namePart id="49FED7B773ED2216C44F14CC6581080D">Sanchez-Villagra, Marcelo Ricardo</mods:namePart>
<mods:affiliation id="E1FD387EABAC5285B962E545C6D0010D">Universitaet Zuerich, Palaeontologisches Institut, Karl-Schmid-Strasse 4, 8006 Zuerich, Switzerland</mods:affiliation>
<mods:nameIdentifier id="55219628F318D438BDBA915B1D9EE7F3" type="email">m.sanchez@pim.uzh.ch</mods:nameIdentifier>
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<mods:typeOfResource id="9482A0CDAEC286CD0FB39ABC7270C86A">text</mods:typeOfResource>
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<mods:titleInfo id="5382187EE5B4F474F79C05A8B9DCB08D">
<mods:title id="6761DEAEC85D97590C2190EB1E74CCEE">Fossil Record</mods:title>
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<mods:part id="C8437283E11DC78EB7E1B5220A91D479">
<mods:date id="AEA0CD9B2F8E51A2801D4AA32117A303">2024</mods:date>
<mods:detail id="E396BE2F8811B7F3E2C61725D302FF05" type="pubDate">
<mods:number id="A7E4651BA6165895E57E91769209BEF7">2024-04-11</mods:number>
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<mods:detail id="4CE438D2FAD469BCBAD9D99F91D77DF7" type="volume">
<mods:number id="2753E14594C361D68955645991166504">27</mods:number>
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<mods:number id="97F4ED51877108FEA8377609ACF4E634">1</mods:number>
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<mods:start id="86ACE73FFA013A2E7C15B6C8FEEBC535">187</mods:start>
<mods:end id="087837364EEEC332DA43B855F49BAC50">207</mods:end>
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<mods:location id="2189A349A3CFAC0BAFAC162DB82BFCD3">
<mods:url id="2814369AE4EF373AD5911ED3494F4A24">http://dx.doi.org/10.3897/fr.27.e119967</mods:url>
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<mods:classification id="68652812C69D0A19C48185E8F5D71915">journal article</mods:classification>
<mods:identifier id="7FAE930452B8180EF18203987B314609" type="DOI">http://dx.doi.org/10.3897/fr.27.e119967</mods:identifier>
<mods:identifier id="0BE11CA245E7B07B38748697F0C81402" type="Pensoft-Pub">2193-0074-1-187</mods:identifier>
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<treatment id="098FA39DC1BD5A82B83A10F86FF25E38" LSID="urn:lsid:plazi:treatment:098FA39DC1BD5A82B83A10F86FF25E38" httpUri="http://treatment.plazi.org/id/098FA39DC1BD5A82B83A10F86FF25E38" lastPageNumber="187" pageId="0" pageNumber="187">
<subSubSection id="469D03AF2B7A6EB920BC0874EB1E6DB1" pageId="0" pageNumber="187" type="nomenclature">
<paragraph id="F93D223BA7D3A7E63862DFC0FA0EE097" pageId="0" pageNumber="187">
†cf.
<taxonomicName id="34FA9BF7FAE32A90BD1EB87F9C167F41" LSID="098FA39D-C1BD-5A82-B83A-10F86FF25E38" authority="(Lund, 1842)" baseAuthorityName="Lund" baseAuthorityYear="1842" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium laurillardi" order="Phyllophaga" pageId="0" pageNumber="187" phylum="Chordata" rank="species" species="laurillardi">Eremotherium laurillardi (Lund, 1842)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="5BD2FD6CBA734F32F681ED90144EC10D" pageId="0" pageNumber="187" type="description">
<paragraph id="384F7B7C5C4CE2AED19420F66E1604A8" pageId="0" pageNumber="187">
<figureCitation id="B611398FCD960D58F82BEE2B1B96AD69" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megatheres (A 1 - G) and mylodontids (H 1, H 2) from the Cauca site. A 1 - A 4. Skull of cf. Eremotherium laurillardi (AMU-CURS- 1268 a) in right lateral, ventral, posterior and anteroventral view; B 1, B 2. Right hemi-mandible (AMU-CURS- 1268 b) in right lateral view; C 1 - C 3. Right radius (AMU-CURS- 1268 d) in proximal, dorsal and distal view; D 1, D 2. Right clavicle (AMU-CURS- 1268 c); E 1, E 2. Thoracic vertebra (AMU-CURS- 1268 j) in anterior and left lateral view; F. Thoracic or lumbar vertebral centrum (AMU-CURS- 1268 k) in anterior view; G. Left IV metatarsal (AMU-CURS- 1268 g); H 1, H 2. cf. Mylodontidae, proximal and dorsal right radius (AMU-CURS- 1363). Abbreviations: (alv.) alveolus, (fm) foramen magnum, (mf) lower molariform, (Mf) upper molariform, (oc) occipital condyle." figureDoi="10.3897/fr.27.e119967.figure5" httpUri="https://binary.pensoft.net/fig/1023340" pageId="0" pageNumber="187">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8F3C3FDA1EF2C71A274F342AAAB92CAD" pageId="0" pageNumber="187" type="materials_examined">
<paragraph id="7A814707053C0FDB2C59756B567FF7B5" pageId="0" pageNumber="187">Referred material.</paragraph>
<paragraph id="3210B331397F261B0E2A52AE56DB2526" pageId="0" pageNumber="187">
Remains of
<taxonomicName id="985E235A2DD5B2E6B922832BF2A598AE" authorityName="Spillmann" authorityYear="1948" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="4506BAB2C8A9A211615538E144E6F16C" italics="true" pageId="0" pageNumber="187">Eremotherium</emphasis>
</taxonomicName>
cf.
<taxonomicName id="457C1CE94350B501242FCCCCCCF20F81" lsidName="E. laurillardi" pageId="0" pageNumber="187" rank="species" species="laurillardi">
<emphasis id="E3C0109B2169C85D3D7B74504A66AAFE" italics="true" pageId="0" pageNumber="187">E. laurillardi</emphasis>
</taxonomicName>
have been recovered at the Cauca site in bone groupings 2-6, with only groupings 3-5 being excavated (Fig.
<figureCitation id="F9C68D308E6A877327CCEBE1B04ADE47" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographic location of the Cauca site (A) and the excavation area (B). The different bone groupings are referred to as &quot; S &quot; and the excavations as &quot; A-E &quot;. The M 2 isolated from a juvenile Notiomastodon platensis (AMU-CURS- 1045) is referred to with a triangle and *. Lithic artefacts referred to here are illustrated in Fig. 9." figureDoi="10.3897/fr.27.e119967.figure1" httpUri="https://binary.pensoft.net/fig/1023336" pageId="0" pageNumber="187">1B</figureCitation>
). Fourteen cranial and postcranial remains that include a right hemi-mandible (with mf 1 to 4), skull (with right Mf 1 to 5 and left Mf 2 to 4), right clavicle, right radius, presumably left navicular, cuneiform, IV left metatarsal, a worn element that we presume corresponds to a scaphoid, fragment of pelvis with acetabulum and part of the pubic ramus and five incomplete vertebrae, from excavation Cauca
<normalizedToken id="2D49EE837B2BF1AD72ACF0793925B9B1" originalValue="“B”">&quot;B&quot;</normalizedToken>
(Fig.
<figureCitation id="F0EE0C93257EA9E60C554F9909C93C2D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Excavations at the Cauca site. Cauca &quot; A &quot; (A), Cauca &quot; B &quot; (B), and Cauca &quot; E &quot; (C), showing the defence of Notiomastodon platensis (AMU-CURS- 1359) in situ. Mandible of N. platensis in plaster jacket in Cauca &quot; A &quot; (D). Cauca &quot; D &quot; with remains (tibia and ribs) of cf. Eremotherium laurillardi (E)." figureDoi="10.3897/fr.27.e119967.figure4" httpUri="https://binary.pensoft.net/fig/1023339" pageId="0" pageNumber="187">4B</figureCitation>
) and assigned to catalogue number AMU-CURS-1268. A probable right tibia (AMU-CURS-sn) and seven rib fragments were found in Cauca
<normalizedToken id="6F4AE9368153255F5DF7BB79D44A8C1B" originalValue="“D”">&quot;D&quot;</normalizedToken>
. The other cranial and postcranial elements identified in Cauca
<normalizedToken id="1EBE8F479EF17B0EE447367410FD19CA" originalValue="“C”">&quot;C&quot;</normalizedToken>
and unexcavated groupings 2 and 6 were not collected due to their poor state of preservation, being in some cases disintegrated (Fig.
<figureCitation id="8C2D03DDFD18F8D680BB4862FB20B7EB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Remains of fossil vertebrates emerging on the surface at the Cauca site. Skull (A), pelvis (B) and appendicular elements (C) of megatheriid Eremotherium. A, C. correspond to unexcavated S 2 (see Fig. 1 B). Upper molars (D) of gomphoteriid Notiomastodon platensis, with the root completely eroded, associated with the mandible AMU-CURS- 1269 from Cauca &quot; A &quot; (Fig. 1 B)." figureDoi="10.3897/fr.27.e119967.figure3" httpUri="https://binary.pensoft.net/fig/1023338" pageId="0" pageNumber="187">3A-C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9A51F80C3B8A603E754CE605B3D93324" pageId="0" pageNumber="187" type="description">
<paragraph id="E27C6E7ED4FF1BC09CE72EDF6543F98D" pageId="0" pageNumber="187">Descriptions.</paragraph>
<paragraph id="41A9EBC9FBB622FC7A38672701740908" pageId="0" pageNumber="187">
The 14 disarticulated cranial and postcranial elements recovered in Cauca
<normalizedToken id="0EE59581910143FAD389C25E83CCE76C" originalValue="“B”">&quot;B&quot;</normalizedToken>
most probably belong to the same individual (AMU-CURS-1268). The cranial material is composed of only the skull (AMU-CURS-1268a) (Fig.
<figureCitation id="72BF66A41656F30B67CDF828EEDAA9B4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megatheres (A 1 - G) and mylodontids (H 1, H 2) from the Cauca site. A 1 - A 4. Skull of cf. Eremotherium laurillardi (AMU-CURS- 1268 a) in right lateral, ventral, posterior and anteroventral view; B 1, B 2. Right hemi-mandible (AMU-CURS- 1268 b) in right lateral view; C 1 - C 3. Right radius (AMU-CURS- 1268 d) in proximal, dorsal and distal view; D 1, D 2. Right clavicle (AMU-CURS- 1268 c); E 1, E 2. Thoracic vertebra (AMU-CURS- 1268 j) in anterior and left lateral view; F. Thoracic or lumbar vertebral centrum (AMU-CURS- 1268 k) in anterior view; G. Left IV metatarsal (AMU-CURS- 1268 g); H 1, H 2. cf. Mylodontidae, proximal and dorsal right radius (AMU-CURS- 1363). Abbreviations: (alv.) alveolus, (fm) foramen magnum, (mf) lower molariform, (Mf) upper molariform, (oc) occipital condyle." figureDoi="10.3897/fr.27.e119967.figure5" httpUri="https://binary.pensoft.net/fig/1023340" pageId="0" pageNumber="187">5A1-A4</figureCitation>
) and the right hemi-mandible (AMU-CURS-1268b) (Fig.
<figureCitation id="2E704004E070BC9B9A87C02D4398EE3C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megatheres (A 1 - G) and mylodontids (H 1, H 2) from the Cauca site. A 1 - A 4. Skull of cf. Eremotherium laurillardi (AMU-CURS- 1268 a) in right lateral, ventral, posterior and anteroventral view; B 1, B 2. Right hemi-mandible (AMU-CURS- 1268 b) in right lateral view; C 1 - C 3. Right radius (AMU-CURS- 1268 d) in proximal, dorsal and distal view; D 1, D 2. Right clavicle (AMU-CURS- 1268 c); E 1, E 2. Thoracic vertebra (AMU-CURS- 1268 j) in anterior and left lateral view; F. Thoracic or lumbar vertebral centrum (AMU-CURS- 1268 k) in anterior view; G. Left IV metatarsal (AMU-CURS- 1268 g); H 1, H 2. cf. Mylodontidae, proximal and dorsal right radius (AMU-CURS- 1363). Abbreviations: (alv.) alveolus, (fm) foramen magnum, (mf) lower molariform, (Mf) upper molariform, (oc) occipital condyle." figureDoi="10.3897/fr.27.e119967.figure5" httpUri="https://binary.pensoft.net/fig/1023340" pageId="0" pageNumber="187">5B1, B2</figureCitation>
). The skull has a Tl of 580 mm and preserves only the palatal section, since part of the cranial vault, nasal bones and part of the maxilla bones were destroyed by erosive processes. The palatal section is relatively well preserved, with both the right Mf1-Mf5 and the left series Mf2-4 and the alveolus of left Mf1 and Mf5. The hemi-mandible has a Tl of 450 mm, preserving the four molariforms mf1-mf4 (Fig.
<figureCitation id="F6721ABA097989BBE7E9311D902AB402" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megatheres (A 1 - G) and mylodontids (H 1, H 2) from the Cauca site. A 1 - A 4. Skull of cf. Eremotherium laurillardi (AMU-CURS- 1268 a) in right lateral, ventral, posterior and anteroventral view; B 1, B 2. Right hemi-mandible (AMU-CURS- 1268 b) in right lateral view; C 1 - C 3. Right radius (AMU-CURS- 1268 d) in proximal, dorsal and distal view; D 1, D 2. Right clavicle (AMU-CURS- 1268 c); E 1, E 2. Thoracic vertebra (AMU-CURS- 1268 j) in anterior and left lateral view; F. Thoracic or lumbar vertebral centrum (AMU-CURS- 1268 k) in anterior view; G. Left IV metatarsal (AMU-CURS- 1268 g); H 1, H 2. cf. Mylodontidae, proximal and dorsal right radius (AMU-CURS- 1363). Abbreviations: (alv.) alveolus, (fm) foramen magnum, (mf) lower molariform, (Mf) upper molariform, (oc) occipital condyle." figureDoi="10.3897/fr.27.e119967.figure5" httpUri="https://binary.pensoft.net/fig/1023340" pageId="0" pageNumber="187">5B2</figureCitation>
). This hemi-mandible has a convex ventral margin and projects downwards at the level of the first molars, forming a moderate mandibular protuberance. The molariforms preserved in the skull and hemi-mandible are of quadrangular section with crowns characterised by two transverse and parallel ridges, separated by a
<normalizedToken id="52E2EC1B2705E706759B8012AEAFAA02" originalValue="“V”">&quot;V&quot;</normalizedToken>
-shaped valley. Measurements of the molariforms are presented in Table
<tableCitation id="0DC81E341ED9A164EC38A8FAFCED61CA" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Dental measurements of cf. Eremotherium laurillardi (AMU-CURS- 1269 a, b) collected at the Cauca site. Abbreviations: upper molariform (Mf), lower molariform (mf), right (d), left (i). (*): incomplete or missing pieces. Measures in millimeters." httpUri="http://table.plazi.org/id/2FEA4BA083D75AE14B8ADBC26E232DBD" pageId="0" pageNumber="187" tableUuid="2FEA4BA083D75AE14B8ADBC26E232DBD">2</tableCitation>
. Associated with these cranial remains were recovered the right clavicle with a Tl of 240 mm (AMU-CURS-1268c; Fig.
<figureCitation id="D622493ECDD54A3E068EDCC48294F31C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megatheres (A 1 - G) and mylodontids (H 1, H 2) from the Cauca site. A 1 - A 4. Skull of cf. Eremotherium laurillardi (AMU-CURS- 1268 a) in right lateral, ventral, posterior and anteroventral view; B 1, B 2. Right hemi-mandible (AMU-CURS- 1268 b) in right lateral view; C 1 - C 3. Right radius (AMU-CURS- 1268 d) in proximal, dorsal and distal view; D 1, D 2. Right clavicle (AMU-CURS- 1268 c); E 1, E 2. Thoracic vertebra (AMU-CURS- 1268 j) in anterior and left lateral view; F. Thoracic or lumbar vertebral centrum (AMU-CURS- 1268 k) in anterior view; G. Left IV metatarsal (AMU-CURS- 1268 g); H 1, H 2. cf. Mylodontidae, proximal and dorsal right radius (AMU-CURS- 1363). Abbreviations: (alv.) alveolus, (fm) foramen magnum, (mf) lower molariform, (Mf) upper molariform, (oc) occipital condyle." figureDoi="10.3897/fr.27.e119967.figure5" httpUri="https://binary.pensoft.net/fig/1023340" pageId="0" pageNumber="187">5D1, D2</figureCitation>
), complete right radius with a Tl of 700 mm (AMU-CURS-1268d; Fig.
<figureCitation id="08967AD0926F1141833D036F51ACD583" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megatheres (A 1 - G) and mylodontids (H 1, H 2) from the Cauca site. A 1 - A 4. Skull of cf. Eremotherium laurillardi (AMU-CURS- 1268 a) in right lateral, ventral, posterior and anteroventral view; B 1, B 2. Right hemi-mandible (AMU-CURS- 1268 b) in right lateral view; C 1 - C 3. Right radius (AMU-CURS- 1268 d) in proximal, dorsal and distal view; D 1, D 2. Right clavicle (AMU-CURS- 1268 c); E 1, E 2. Thoracic vertebra (AMU-CURS- 1268 j) in anterior and left lateral view; F. Thoracic or lumbar vertebral centrum (AMU-CURS- 1268 k) in anterior view; G. Left IV metatarsal (AMU-CURS- 1268 g); H 1, H 2. cf. Mylodontidae, proximal and dorsal right radius (AMU-CURS- 1363). Abbreviations: (alv.) alveolus, (fm) foramen magnum, (mf) lower molariform, (Mf) upper molariform, (oc) occipital condyle." figureDoi="10.3897/fr.27.e119967.figure5" httpUri="https://binary.pensoft.net/fig/1023340" pageId="0" pageNumber="187">5C1-C3</figureCitation>
), presumably left navicular (AMU-CURS-1268e), cuneiform (AMU-CURS-1268f), IV left metatarsal (AMU-CURS-1268g; Fig.
<figureCitation id="0634C054B890E41C3720ABF711A1BC42" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megatheres (A 1 - G) and mylodontids (H 1, H 2) from the Cauca site. A 1 - A 4. Skull of cf. Eremotherium laurillardi (AMU-CURS- 1268 a) in right lateral, ventral, posterior and anteroventral view; B 1, B 2. Right hemi-mandible (AMU-CURS- 1268 b) in right lateral view; C 1 - C 3. Right radius (AMU-CURS- 1268 d) in proximal, dorsal and distal view; D 1, D 2. Right clavicle (AMU-CURS- 1268 c); E 1, E 2. Thoracic vertebra (AMU-CURS- 1268 j) in anterior and left lateral view; F. Thoracic or lumbar vertebral centrum (AMU-CURS- 1268 k) in anterior view; G. Left IV metatarsal (AMU-CURS- 1268 g); H 1, H 2. cf. Mylodontidae, proximal and dorsal right radius (AMU-CURS- 1363). Abbreviations: (alv.) alveolus, (fm) foramen magnum, (mf) lower molariform, (Mf) upper molariform, (oc) occipital condyle." figureDoi="10.3897/fr.27.e119967.figure5" httpUri="https://binary.pensoft.net/fig/1023340" pageId="0" pageNumber="187">5G</figureCitation>
), a worn element that we presume corresponds to a scaphoid (AMU-CURS-1268h), fragment of pelvis with the acetabulum and part of the pubic ramus (AMU-CURS-1268i) and five incomplete vertebrae (AMU-CURS-1268j-n; Fig.
<figureCitation id="CB3949EC2CFFA290C0CCCEB97C09D650" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Megatheres (A 1 - G) and mylodontids (H 1, H 2) from the Cauca site. A 1 - A 4. Skull of cf. Eremotherium laurillardi (AMU-CURS- 1268 a) in right lateral, ventral, posterior and anteroventral view; B 1, B 2. Right hemi-mandible (AMU-CURS- 1268 b) in right lateral view; C 1 - C 3. Right radius (AMU-CURS- 1268 d) in proximal, dorsal and distal view; D 1, D 2. Right clavicle (AMU-CURS- 1268 c); E 1, E 2. Thoracic vertebra (AMU-CURS- 1268 j) in anterior and left lateral view; F. Thoracic or lumbar vertebral centrum (AMU-CURS- 1268 k) in anterior view; G. Left IV metatarsal (AMU-CURS- 1268 g); H 1, H 2. cf. Mylodontidae, proximal and dorsal right radius (AMU-CURS- 1363). Abbreviations: (alv.) alveolus, (fm) foramen magnum, (mf) lower molariform, (Mf) upper molariform, (oc) occipital condyle." figureDoi="10.3897/fr.27.e119967.figure5" httpUri="https://binary.pensoft.net/fig/1023340" pageId="0" pageNumber="187">5E1-F</figureCitation>
). Other small, fragmented and indeterminate bone elements were observed during the excavation and presumably could be associated with the same individual.
</paragraph>
<caption id="2FEA4BA083D75AE14B8ADBC26E232DBD" ID-Table-UUID="2FEA4BA083D75AE14B8ADBC26E232DBD" httpUri="http://table.plazi.org/id/2FEA4BA083D75AE14B8ADBC26E232DBD" pageId="0" pageNumber="187" start="Table 2" startId="T2">
<paragraph id="E5D61EA5FC0645239EAEB79FFD0AC196" pageId="0" pageNumber="187">
<emphasis id="F10AD29750F56B009CF48A8FB4C86022" bold="true" pageId="0" pageNumber="187">Table 2.</emphasis>
Dental measurements of cf.
<taxonomicName id="56DB94D89A7D124ED7C187E5F82AAE8C" baseAuthorityName="Lund" baseAuthorityYear="1842" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium laurillardi" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="species" species="laurillardi">
<emphasis id="7CBE0D4E5342BAADB1A1AFA70A71B1AC" italics="true" pageId="0" pageNumber="187">Eremotherium laurillardi</emphasis>
</taxonomicName>
(AMU-CURS-1269a, b) collected at the Cauca site. Abbreviations: upper molariform (Mf), lower molariform (mf), right (d), left (i). (*): incomplete or missing pieces. Measures in millimeters.
</paragraph>
</caption>
<paragraph id="47F0A9DAE8942C0D87BC7F1C75915E29" pageId="0" pageNumber="187">
<table id="F7EC00F6270122D715A75E3AA34FDAB3" pageId="0" pageNumber="187">
<tr id="E6432110B2AB27D29EC1A4BB8C3CF7EC" pageId="0" pageNumber="187">
<th id="34412406D9917B90889ED000E389965A" colspan="1" isEmpty="true" pageId="0" pageNumber="187" rowspan="1">-</th>
<th id="8EC7E056F79D2209A7EB957E0668764E" colspan="1" pageId="0" pageNumber="187" rowspan="1">Mfr1</th>
<th id="B53BA13B0B0A974174427CE11A39B5F7" colspan="1" pageId="0" pageNumber="187" rowspan="1">Mfr2</th>
<th id="C3B1A37D275AEF7F39E883782F7AFFAE" colspan="1" pageId="0" pageNumber="187" rowspan="1">Mfr3</th>
<th id="92FC5822654E82101C63DD454B21E322" colspan="1" pageId="0" pageNumber="187" rowspan="1">Mfr4</th>
<th id="7E6EEEA4FFB60A095222F07DC635367A" colspan="1" pageId="0" pageNumber="187" rowspan="1">Mfr5*</th>
<th id="72A335BC670772B42EA832211CEB25A2" colspan="1" pageId="0" pageNumber="187" rowspan="1">Mfl1</th>
<th id="8ECFEB318633D0831AC8E7B87AAB5558" colspan="1" pageId="0" pageNumber="187" rowspan="1">Mfl2</th>
<th id="90498429AB2F7AC82CE69761CCBAE6BC" colspan="1" pageId="0" pageNumber="187" rowspan="1">Mfl3</th>
<th id="B66450F68618D584AD551CB0E970377A" colspan="1" pageId="0" pageNumber="187" rowspan="1">Mfl4</th>
<th id="CACAA05A79EE1E117534EE19663BF1C0" colspan="1" pageId="0" pageNumber="187" rowspan="1">Mfl5*</th>
<th id="80EC2FB40A50787CC0751BE10ACE5EA9" colspan="1" pageId="0" pageNumber="187" rowspan="1">mfr1</th>
<th id="6D2FC69F89B68EC0BDE5F6C86C41F4CA" colspan="1" pageId="0" pageNumber="187" rowspan="1">mfr2</th>
<th id="6AF655EB4C0C4E52E25919F21F05CBFF" colspan="1" pageId="0" pageNumber="187" rowspan="1">mfr3</th>
<th id="66B852FF89AEB39FC66CC8B81AE4F6B9" colspan="1" pageId="0" pageNumber="187" rowspan="1">mfr4</th>
</tr>
<tr id="27A05203CF2D1E6A20632E756087C057" pageId="0" pageNumber="187">
<td id="E293126C4E565D383E1D1366AC207759" colspan="1" pageId="0" pageNumber="187" rowspan="1">Long</td>
<td id="167A4AC4A56979283D3B073E0540B972" colspan="1" pageId="0" pageNumber="187" rowspan="1">25</td>
<td id="703281C6B9744EF021D0D70547FB2EC4" colspan="1" pageId="0" pageNumber="187" rowspan="1">35</td>
<td id="3A8F2C28113E2CDFD64A615357D740D1" colspan="1" pageId="0" pageNumber="187" rowspan="1">35</td>
<td id="2D4117B3B6450110117281178930A0F0" colspan="1" pageId="0" pageNumber="187" rowspan="1">35</td>
<td id="4390E57B01A31E3C1999D468F202613F" colspan="1" pageId="0" pageNumber="187" rowspan="1">20</td>
<td id="A245F651E2F440CC5E709A13BF5AFA78" colspan="1" pageId="0" pageNumber="187" rowspan="1">25</td>
<td id="7A26B5D41F804921ED0BBDF260CCD812" colspan="1" pageId="0" pageNumber="187" rowspan="1">35</td>
<td id="42E4A1AFC04E1332D40D712657AAEB96" colspan="1" pageId="0" pageNumber="187" rowspan="1">35</td>
<td id="A065C3E065E3EEF2A80B58D3BBF2571E" colspan="1" pageId="0" pageNumber="187" rowspan="1">37</td>
<td id="C2C1982980C3801A7A97730015187651" colspan="1" pageId="0" pageNumber="187" rowspan="1">20</td>
<td id="17627DFD7E09375BC08D4E94F9326C62" colspan="1" pageId="0" pageNumber="187" rowspan="1">40</td>
<td id="B011E98BCFA429FED68218C0011EE4E0" colspan="1" pageId="0" pageNumber="187" rowspan="1">40</td>
<td id="DAF47FB35814C8850C814F3304ADE45E" colspan="1" pageId="0" pageNumber="187" rowspan="1">40</td>
<td id="04BE2A1229A7D7CD64699BEA3DF65D3E" colspan="1" pageId="0" pageNumber="187" rowspan="1">30</td>
</tr>
<tr id="248FF973B99C1BC378A07C88E745BC28" pageId="0" pageNumber="187">
<td id="9F445E3B9FB155B252EEDC75235DC6F8" colspan="1" pageId="0" pageNumber="187" rowspan="1">Width</td>
<td id="C64AD7BABAF493B155A6B63E83FDF82E" colspan="1" pageId="0" pageNumber="187" rowspan="1">36</td>
<td id="D41EBC691CE551CABB8525EE4ED5F827" colspan="1" pageId="0" pageNumber="187" rowspan="1">50</td>
<td id="8F0A267C6C3C98C69CE4135EB565F4AF" colspan="1" pageId="0" pageNumber="187" rowspan="1">47</td>
<td id="88E4615B88ED19D4DB1BC0AE90725EA8" colspan="1" pageId="0" pageNumber="187" rowspan="1">40</td>
<td id="48C86DE31B2D415ECD42166DD40404E7" colspan="1" pageId="0" pageNumber="187" rowspan="1">25</td>
<td id="8A0C547C5F975ED20BA87C21DD259A7B" colspan="1" pageId="0" pageNumber="187" rowspan="1">35</td>
<td id="2205056EF2BA4A3141A26924EEC775F1" colspan="1" pageId="0" pageNumber="187" rowspan="1">45</td>
<td id="0C45790296A9639E7FA19F27AF8FBE63" colspan="1" pageId="0" pageNumber="187" rowspan="1">47</td>
<td id="93609A55ED606448219E2CB33E744A57" colspan="1" pageId="0" pageNumber="187" rowspan="1">40</td>
<td id="8A8348E5EAFF3B71004EEB8580582BB4" colspan="1" pageId="0" pageNumber="187" rowspan="1">15</td>
<td id="1249664A534B1D5B4DB72BA09D254362" colspan="1" pageId="0" pageNumber="187" rowspan="1">44</td>
<td id="71A776A3EDE9FD653009A16A3F053E53" colspan="1" pageId="0" pageNumber="187" rowspan="1">45</td>
<td id="21E0D38F3D38E7ACE7E17A360ABF0D0C" colspan="1" pageId="0" pageNumber="187" rowspan="1">42</td>
<td id="4ACDD2E26BD511E31157B00AF4573DEA" colspan="1" pageId="0" pageNumber="187" rowspan="1">40</td>
</tr>
<tr id="92F58B56D51A064F597252898774A1B9" pageId="0" pageNumber="187">
<td id="553D12DF0530FF08D3F0804AF72E495B" colspan="1" pageId="0" pageNumber="187" rowspan="1">Height</td>
<td id="042B2B824D8262EEF778390CCD473A99" colspan="1" pageId="0" pageNumber="187" rowspan="1">40</td>
<td id="C10354B3129B590028A306835CEE3155" colspan="1" pageId="0" pageNumber="187" rowspan="1">32</td>
<td id="F34120B3C8D491F7A674B6C0B19261BD" colspan="1" pageId="0" pageNumber="187" rowspan="1">22</td>
<td id="FB2A7E692D54BB539D25085A8E66772C" colspan="1" pageId="0" pageNumber="187" rowspan="1">10</td>
<td id="F08061A9A9368A461E7335F13793FAE6" colspan="1" pageId="0" pageNumber="187" rowspan="1">5</td>
<td id="BC96D4F0F4F0F025C0BBC3A52A42E4DB" colspan="1" pageId="0" pageNumber="187" rowspan="1">10</td>
<td id="C54508FC0244D543B095CE8D9CFA5808" colspan="1" pageId="0" pageNumber="187" rowspan="1">10</td>
<td id="CC8867D7EB50AA6E12D0636B9DF97343" colspan="1" pageId="0" pageNumber="187" rowspan="1">27</td>
<td id="9EBB0A2625903C6BAEEB0F07E980AE77" colspan="1" pageId="0" pageNumber="187" rowspan="1">20</td>
<td id="A7C72EE57FE17820FC6DEFF91E447397" colspan="1" pageId="0" pageNumber="187" rowspan="1">-</td>
<td id="8DB27EDF82DC3958D7C60ED7CA7881D8" colspan="1" pageId="0" pageNumber="187" rowspan="1">40</td>
<td id="7D66F680A317D1465591A702E81730FE" colspan="1" pageId="0" pageNumber="187" rowspan="1">30</td>
<td id="EC9949D7551FAD3865615788C372D9B7" colspan="1" pageId="0" pageNumber="187" rowspan="1">10</td>
<td id="323377EB7B3969183773E8DEF3207F68" colspan="1" pageId="0" pageNumber="187" rowspan="1">10</td>
</tr>
</table>
</paragraph>
<caption id="5BCA3A57136425F81CA227D47EC6EC9D" doi="10.3897/fr.27.e119967.figure5" httpUri="https://binary.pensoft.net/fig/1023340" pageId="0" pageNumber="187" start="Figure 5" startId="F5">
<paragraph id="7C9D0BB5D54AA4B6A2F2715EFCD656D8" pageId="0" pageNumber="187">
<emphasis id="4AF3A776EBD72D5C2821208839877D3F" bold="true" pageId="0" pageNumber="187">Figure 5.</emphasis>
Megatheres (
<emphasis id="644D19C51DD865779011867F22B08875" bold="true" pageId="0" pageNumber="187">A1-G</emphasis>
) and mylodontids (
<emphasis id="AF3AD31C9A0E81631BDF9FF5BC91C7E6" bold="true" pageId="0" pageNumber="187">H1, H2</emphasis>
) from the Cauca site.
<emphasis id="47B890C2228D7137EBFB12414BD07952" bold="true" pageId="0" pageNumber="187">A1</emphasis>
-
<emphasis id="F40A71AE94240EFF1FA466AA27CA5B26" bold="true" pageId="0" pageNumber="187">A4</emphasis>
. Skull of cf.
<taxonomicName id="7AA2B394C5062016C41244BD9C8C6ADF" baseAuthorityName="Lund" baseAuthorityYear="1842" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium laurillardi" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="species" species="laurillardi">
<emphasis id="9BE521F3B30149D6E9B8C6CF4EEED1BD" italics="true" pageId="0" pageNumber="187">Eremotherium laurillardi</emphasis>
</taxonomicName>
(AMU-CURS-1268a) in right lateral, ventral, posterior and anteroventral view;
<emphasis id="F1A9461550AC79E5E8AC7D80CB68529B" bold="true" pageId="0" pageNumber="187">B1, B2.</emphasis>
Right hemi-mandible (AMU-CURS-1268b) in right lateral view;
<emphasis id="3F94BC150C02F85EA2B0FF5EBFC0536B" bold="true" pageId="0" pageNumber="187">C1-C3.</emphasis>
Right radius (AMU-CURS-1268d) in proximal, dorsal and distal view;
<emphasis id="11257E5180C1DEA786D881973DBB34B1" bold="true" pageId="0" pageNumber="187">D1, D2.</emphasis>
Right clavicle (AMU-CURS-1268c);
<emphasis id="C2E1E5F6A8EF6DCD9B0BCC0A64C5B083" bold="true" pageId="0" pageNumber="187">E1, E2.</emphasis>
Thoracic vertebra (AMU-CURS-1268j) in anterior and left lateral view;
<emphasis id="87A61D3E1ED9EA57669E8900C7AF3555" bold="true" pageId="0" pageNumber="187">F.</emphasis>
Thoracic or lumbar vertebral centrum (AMU-CURS-1268k) in anterior view;
<emphasis id="473776AE065845D2EB607F421FF018C7" bold="true" pageId="0" pageNumber="187">G.</emphasis>
Left IV metatarsal (AMU-CURS-1268g);
<emphasis id="7E9F453403E171F55F2CB7D9829AF0C8" bold="true" pageId="0" pageNumber="187">H1, H2.</emphasis>
cf.
<taxonomicName id="325FD207E63C36AC7434DFEAF6EED922" family="Mylodontidae" kingdom="Animalia" lsidName="" order="Phyllophaga" pageId="0" pageNumber="187" rank="family">Mylodontidae</taxonomicName>
, proximal and dorsal right radius (AMU-CURS-1363). Abbreviations: (alv.) alveolus, (fm) foramen magnum, (mf) lower molariform, (Mf) upper molariform, (oc) occipital condyle.
</paragraph>
</caption>
<paragraph id="28DC88197D738234BE2E42CD15598889" pageId="0" pageNumber="187">
In Cauca
<normalizedToken id="C4650B7B3A5987412EF00925E584C504" originalValue="“C”">&quot;C&quot;</normalizedToken>
, only a few remains of postcranial elements of an
<taxonomicName id="6B9A8DE897E18CC7CC7378A1B789A404" authorityName="Spillmann" authorityYear="1948" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="A898FADC5EAE0BE254AACE5DF4B74FE4" italics="true" pageId="0" pageNumber="187">Eremotherium</emphasis>
</taxonomicName>
, were found emerging on the surface. These were identified as vertebrae, ribs, pelvis fragments and other small indeterminate fragments. However, the poor state of preservation and disintegration in some cases, did not allow their recovery. Due to the proximity of these materials to the individual collected in Cauca
<normalizedToken id="5B6CB38A87EB01E75992788549489335" originalValue="“B”">&quot;B&quot;</normalizedToken>
, a possible association amongst them is not ruled out here. In Cauca
<normalizedToken id="082B47601A0FF238DB9A706552F56549" originalValue="“D”">&quot;D&quot;</normalizedToken>
, an association of seven fragments of ribs and a right tibia were recovered, also in a poor state of preservation (Fig.
<figureCitation id="38BE245B6E6A3C9F806A7B999F638672" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Excavations at the Cauca site. Cauca &quot; A &quot; (A), Cauca &quot; B &quot; (B), and Cauca &quot; E &quot; (C), showing the defence of Notiomastodon platensis (AMU-CURS- 1359) in situ. Mandible of N. platensis in plaster jacket in Cauca &quot; A &quot; (D). Cauca &quot; D &quot; with remains (tibia and ribs) of cf. Eremotherium laurillardi (E)." figureDoi="10.3897/fr.27.e119967.figure4" httpUri="https://binary.pensoft.net/fig/1023339" pageId="0" pageNumber="187">4E</figureCitation>
). This bone accumulation is located about 8 m north of the Cauca
<normalizedToken id="1E49907918950CD9FA7E4028D344B89D" originalValue="“B”">&quot;B&quot;</normalizedToken>
individual, so a possible association with the latter would not be unlikely.
</paragraph>
<paragraph id="EC16E7BB50D2DE52E1F685FAEEF0317B" pageId="0" pageNumber="187">
Bone grouping 2 (Fig.
<figureCitation id="409CAA8927AE26A5AFE0E98C9C2910F9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographic location of the Cauca site (A) and the excavation area (B). The different bone groupings are referred to as &quot; S &quot; and the excavations as &quot; A-E &quot;. The M 2 isolated from a juvenile Notiomastodon platensis (AMU-CURS- 1045) is referred to with a triangle and *. Lithic artefacts referred to here are illustrated in Fig. 9." figureDoi="10.3897/fr.27.e119967.figure1" httpUri="https://binary.pensoft.net/fig/1023336" pageId="0" pageNumber="187">1B</figureCitation>
) was not excavated due to the partial or total disintegration in very little scraps of bones of the outcropping materials. However, a skull in a state of fragmentation (Fig.
<figureCitation id="98AA71F400A848344A66DE1492969856" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Remains of fossil vertebrates emerging on the surface at the Cauca site. Skull (A), pelvis (B) and appendicular elements (C) of megatheriid Eremotherium. A, C. correspond to unexcavated S 2 (see Fig. 1 B). Upper molars (D) of gomphoteriid Notiomastodon platensis, with the root completely eroded, associated with the mandible AMU-CURS- 1269 from Cauca &quot; A &quot; (Fig. 1 B)." figureDoi="10.3897/fr.27.e119967.figure3" httpUri="https://binary.pensoft.net/fig/1023338" pageId="0" pageNumber="187">3A</figureCitation>
) still offers evidence that allows us to identify this association as a second specimen of
<taxonomicName id="E52217D7337F7740403DB30CC8983131" authorityName="Spillmann" authorityYear="1948" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="7FC9D8583C89CDB14E8F97B0EE579933" italics="true" pageId="0" pageNumber="187">Eremotherium</emphasis>
</taxonomicName>
. In bone grouping 6 (not excavated) to the northwest of Cauca
<normalizedToken id="47093DBAC56BEFA3F6DE2A4B00B16788" originalValue="“D”">&quot;D&quot;</normalizedToken>
, a humerus of
<taxonomicName id="BC57287294EE68B30720EFC5AE0FA43A" authorityName="Spillmann" authorityYear="1948" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="F33F087F01BC5040DC5E19DD9FA7C844" italics="true" pageId="0" pageNumber="187">Eremotherium</emphasis>
</taxonomicName>
was also identified, but its poor state of preservation prevented collection or conservation
<emphasis id="770FBEB9967F67C5776AB4B9D16CABB4" italics="true" pageId="0" pageNumber="187">in situ</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="36A7D0A77004E83F57A53FBAD23003ED" pageId="0" pageNumber="187" type="remarks">
<paragraph id="EBF38421831109C66E8C1A6C91A54E78" pageId="0" pageNumber="187">Remarks.</paragraph>
<paragraph id="D63AC7C496D14865E4E64A7AE066699F" pageId="0" pageNumber="187">
The ground sloth
<taxonomicName id="25D6F6A3716B931EDFE2DEFB3536E05D" baseAuthorityName="Lund" baseAuthorityYear="1842" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium laurillardi" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="species" species="laurillardi">
<emphasis id="D6494D5E8BDD1010919FC037C974E251" italics="true" pageId="0" pageNumber="187">Eremotherium laurillardi</emphasis>
</taxonomicName>
distributed from south-eastern USA to Brazil, is the only species of the genus known from the Late Pleistocene in the Americas (
<bibRefCitation id="12609C23373634883CF796D0EDAC9443" author="Cartelle, C" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="0" pageNumber="187" pagination="830 - 841" publicationUrl="https://www.jstor.org/stable/4523673" refId="B26" refString="Cartelle, C, De Iuliis, G, 1995. Eremotherium laurillardi: the panamerican Late Pleistocene megatheriid sloth. Journal of Vertebrate Paleontology 15 (4): 830 - 841, https://www.jstor.org/stable/4523673" title="Eremotherium laurillardi: the panamerican Late Pleistocene megatheriid sloth." url="https://www.jstor.org/stable/4523673" volume="15" year="1995">Cartelle and De Iuliis 1995</bibRefCitation>
). The taxonomic validity of
<taxonomicName id="500FBC28F8C2DAC55FA1ED9BC25911BC" baseAuthorityName="Schaub" baseAuthorityYear="1935" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium rusconi" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="species" species="rusconi">
<emphasis id="2EADE5950D2156343A9C908174E10750" italics="true" pageId="0" pageNumber="187">Eremotherium rusconi</emphasis>
</taxonomicName>
Schaub, 1935, another species from the Late Pleistocene of South America, is questionable (
<bibRefCitation id="007A28E6FAEFAB280BDB53B910EC1DBB" author="Cartelle, C" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="0" pageNumber="187" pagination="830 - 841" publicationUrl="https://www.jstor.org/stable/4523673" refId="B26" refString="Cartelle, C, De Iuliis, G, 1995. Eremotherium laurillardi: the panamerican Late Pleistocene megatheriid sloth. Journal of Vertebrate Paleontology 15 (4): 830 - 841, https://www.jstor.org/stable/4523673" title="Eremotherium laurillardi: the panamerican Late Pleistocene megatheriid sloth." url="https://www.jstor.org/stable/4523673" volume="15" year="1995">Cartelle and De Iuliis 1995</bibRefCitation>
;
<bibRefCitation id="11724547D63E649C90B8EBC9113E5D5C" DOI="https://doi.org/10.1016/j.crpv.2013.11.001" author="Faure, M" journalOrPublisher="Comptes Rendus Palevol" pageId="0" pageNumber="187" pagination="259 - 266" refId="B41" refString="Faure, M, Guerin, C, Parenti, F, 2014. Sur l'existence de deux especes d' Eremotherium E. rusconii (Schaub, 1935) et E. laurillardi (Lund, 1842) dans le Pleistocene superieur du Bresil intertropical. Comptes Rendus Palevol 13 (4): 259 - 266, DOI: https://doi.org/10.1016/j.crpv.2013.11.001" title="Sur l'existence de deux especes d' Eremotherium E. rusconii (Schaub, 1935) et E. laurillardi (Lund, 1842) dans le Pleistocene superieur du Bresil intertropical." url="https://doi.org/10.1016/j.crpv.2013.11.001" volume="13" year="2014">Faure et al. 2014</bibRefCitation>
;
<bibRefCitation id="BDD538505B234C63FB8F3F850726A5D7" DOI="https://doi.org/10.1016/j.crpv.2014.09.002" author="Cartelle, C" journalOrPublisher="Comptes Rendus Palevol" pageId="0" pageNumber="187" pagination="15 - 23" refId="B27" refString="Cartelle, C, De Iuliis, G, Pujos, F, 2015. Eremotherium laurillardi (Lund, 1842) (Xenarthra, Megatheriinae) is the only valid megatheriine sloth species in the Pleistocene of intertropical Brazil: A response to Faure et al., 2014. Comptes Rendus Palevol 14 (1): 15 - 23, DOI: https://doi.org/10.1016/j.crpv.2014.09.002" title="Eremotherium laurillardi (Lund, 1842) (Xenarthra, Megatheriinae) is the only valid megatheriine sloth species in the Pleistocene of intertropical Brazil: A response to Faure et al., 2014." url="https://doi.org/10.1016/j.crpv.2014.09.002" volume="14" year="2015">Cartelle et al. 2015</bibRefCitation>
).
</paragraph>
<paragraph id="02867624A7F2F6BF649D8ED4FABABEDE" pageId="0" pageNumber="187">
Remains referable to
<taxonomicName id="B2306DBE757B8FA8B64B914B65F535FB" authorityName="Spillmann" authorityYear="1948" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="8B80029F8389AEDD0BCCF093B017B757" italics="true" pageId="0" pageNumber="187">Eremotherium</emphasis>
</taxonomicName>
are common and widely referred to in the Pleistocene fossil record of Venezuela (e.g.
<bibRefCitation id="2214562EAB099C82D50EFE1DD7F1D952" author="Schaub, S." journalOrPublisher="Abhandlungen der Schweizerischen Palaeontologischen Gesellschaft" pageId="0" pageNumber="187" pagination="1 - 21" refId="B112" refString="Schaub, S., 1935. Saeugetierfunde aus Venezuela und Trinidad. Abhandlungen der Schweizerischen Palaeontologischen Gesellschaft 55: 1 - 21" title="Saeugetierfunde aus Venezuela und Trinidad." volume="55" year="1935">Schaub (1935)</bibRefCitation>
;
<bibRefCitation id="E97136C0F1DC0D5CB560DE80149ED727" author="Simpson, G" journalOrPublisher="Wolfgang Walther, Erlangen" pageId="0" pageNumber="187" refId="B115" refString="Simpson, G, 1939. Estudio sobre vertebrados fosiles en Venezuela. Revista de Fomento Ano II(8): 275-283." title="Estudio sobre vertebrados fosiles en Venezuela. Revista de Fomento Ano II (8): 275 - 283." year="1939">Simpson (1939)</bibRefCitation>
;
<bibRefCitation id="392C217BA1586FAE97A51990AA56C0FD" author="Nectario Maria, H" journalOrPublisher="Revista Geografica Americana" pageId="0" pageNumber="187" pagination=": 6" refId="B85" refString="Nectario Maria, H, 1941. El Megatherium larensis de Venezuela. Revista Geografica Americana 16 (98): 6" title="El Megatherium larensis de Venezuela." volume="16" year="1941">Nectario Maria (1941)</bibRefCitation>
;
<bibRefCitation id="FA395292FD9CF8105FCBD595B0626E44" author="Aguilera, OA" journalOrPublisher="Ministerio de la Cultura, Instituto de Patrimonio Cultural, Editorial Arte, Caracas" pageId="0" pageNumber="187" refId="B2" refString="Aguilera, OA, 2006. Tesoros Paleontologicos de Venezuela. El Cuaternario del Estado Falcon. Ministerio de la Cultura, Instituto de Patrimonio Cultural, Editorial Arte, Caracas" title="Tesoros Paleontologicos de Venezuela. El Cuaternario del Estado Falcon." year="2006">Aguilera (2006)</bibRefCitation>
;
<bibRefCitation id="8303792A12A9ECCD20A78CAF8B83E245" author="Carrillo-Briceno, JD" journalOrPublisher="Rio Verde, Caracas" pageId="0" pageNumber="187" refId="B20" refString="Carrillo-Briceno, JD, 2015. Bestias Prehistoricas de Venezuela &quot;Colosos de la Edad de Hielo&quot;. Rio Verde, Caracas" title="Bestias Prehistoricas de Venezuela &quot; Colosos de la Edad de Hielo &quot;." year="2015">
<normalizedToken id="5E17E3E8245B760C0EAE0F8129887C15" originalValue="Carrillo-Briceño">Carrillo-Briceno</normalizedToken>
(2015)
</bibRefCitation>
;
<bibRefCitation id="AEFA8B9F15672ABEEDF2306831A47D08" DOI="https://doi.org/10.5194/fr-20-21-2016" author="Carrillo-Briceno, JD" journalOrPublisher="Fossil Record" pageId="0" pageNumber="187" pagination="21 - 36" refId="B23" refString="Carrillo-Briceno, JD, Amson, E, Zurita, A, Sanchez-Villagra, MR, 2016. Hermann Karsten (1817-1908): a German naturalist in the Neotropics and the significance of his paleovertebrate collection. Fossil Record 20 (1): 21 - 36, DOI: https://doi.org/10.5194/fr-20-21-2016" title="Hermann Karsten (1817 - 1908): a German naturalist in the Neotropics and the significance of his paleovertebrate collection." url="https://doi.org/10.5194/fr-20-21-2016" volume="20" year="2016">
<normalizedToken id="92F49E6D4CBFBD959DDE7183E3E5DD2D" originalValue="Carrillo-Briceño">Carrillo-Briceno</normalizedToken>
et al. (2016)
</bibRefCitation>
;
<bibRefCitation id="6408E61A03819746C6497A416AA78107" author="Chavez-Aponte, EO" journalOrPublisher="Fundacion de Educacion Ambiental - Editorial Amalivaca, Caracas" pageId="0" pageNumber="187" refId="B29" refString="Chavez-Aponte, EO, 2022. Los Megamamiferos de Venezuela: la megafauna del Pleistoceno en el Sur del estado Cojedes. Fundacion de Educacion Ambiental - Editorial Amalivaca, Caracas" title="Los Megamamiferos de Venezuela: la megafauna del Pleistoceno en el Sur del estado Cojedes." year="2022">
<normalizedToken id="A754B083AC3548E437E6D54E21CF457E" originalValue="Chávez-Aponte">Chavez-Aponte</normalizedToken>
(2022)
</bibRefCitation>
and references therein); specific taxonomic assignments of many of these specimens are still uncertain and should be clarified.
</paragraph>
<paragraph id="C2B9DDEE01AEFC859C1A634AB4FAD369" pageId="0" pageNumber="187">
Remains assigned to cf.
<taxonomicName id="263A7F3F5584091525E305A8E1A9C036" baseAuthorityName="Lund" baseAuthorityYear="1842" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium laurillardi" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="species" species="laurillardi">
<emphasis id="540DEDB8982B883EBE8D0A5CD8DF0BD7" italics="true" pageId="0" pageNumber="187">Eremotherium laurillardi</emphasis>
</taxonomicName>
have been reported in Venezuelan territory in palaeontological and archaeological sites ranging from sea level (e.g. Muaco, Taima-Taima and
<normalizedToken id="28DEB01791A81B77BED309B763F5D2A5" originalValue="Cucuruchú">Cucuruchu</normalizedToken>
; see
<bibRefCitation id="F32B5DF45F6A0EE88DCFF86DD25170FD" author="Aguilera, OA" journalOrPublisher="Ministerio de la Cultura, Instituto de Patrimonio Cultural, Editorial Arte, Caracas" pageId="0" pageNumber="187" refId="B2" refString="Aguilera, OA, 2006. Tesoros Paleontologicos de Venezuela. El Cuaternario del Estado Falcon. Ministerio de la Cultura, Instituto de Patrimonio Cultural, Editorial Arte, Caracas" title="Tesoros Paleontologicos de Venezuela. El Cuaternario del Estado Falcon." year="2006">Aguilera (2006)</bibRefCitation>
;
<bibRefCitation id="F24CB1ABD5C46B5772420006BDFCF6D9" author="Carrillo-Briceno, JD" journalOrPublisher="Rio Verde, Caracas" pageId="0" pageNumber="187" refId="B20" refString="Carrillo-Briceno, JD, 2015. Bestias Prehistoricas de Venezuela &quot;Colosos de la Edad de Hielo&quot;. Rio Verde, Caracas" title="Bestias Prehistoricas de Venezuela &quot; Colosos de la Edad de Hielo &quot;." year="2015">
<normalizedToken id="1321D50F303F3B17AEBBBCB261A90266" originalValue="Carrillo-Briceño">Carrillo-Briceno</normalizedToken>
(2015)
</bibRefCitation>
; J.D.C.B pers. obs. (2022)), to mountain areas above 1200 m (e.g. El Vano;
<bibRefCitation id="6B26B58F21C5359E4EDEE936ED8BF4CC" author="Jaimes, A" journalOrPublisher="Boletin de Antropologia Americana" pageId="0" pageNumber="187" refId="B59" refString="Jaimes, A, Sanchez-Villagra, MR, Carrillo-Briceno, JD, 2024b. Fosiles y artefactos en el sitio de matanza de megafauna El Vano, Pleistoceno tardio de Venezuela. In: Sanchez-Villagra MR, Carrillo-Briceno JD, Jaimes A, Arvelo L (Eds) Contribuciones en Venezuela Arqueologica. Scidinge Hall, Tuebingen." title="Fosiles y artefactos en el sitio de matanza de megafauna El Vano, Pleistoceno tardio de Venezuela. In: Sanchez-Villagra MR, Carrillo-Briceno JD, Jaimes A, Arvelo L (Eds) Contribuciones en Venezuela Arqueologica. Scidinge Hall, Tuebingen." year="2024 b">Jaimes et al. (2024b)</bibRefCitation>
).
</paragraph>
<paragraph id="216D710F3A992AC66602BF13659EA550" pageId="0" pageNumber="187">
The cranial and postcranial materials of
<taxonomicName id="9BC6844D2B39B5481F6D3EB3972C6A8D" authorityName="Spillmann" authorityYear="1948" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="B3DBC589DBDBD4F77DC84759BD2E2AC0" italics="true" pageId="0" pageNumber="187">Eremotherium</emphasis>
</taxonomicName>
collected from Cauca likely belong to an adult individual. The assignment of these specimens to the genus
<taxonomicName id="EE5E3CD2A69D345B0F837983F70AD294" authorityName="Spillmann" authorityYear="1948" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="242DD6C1D92D99556D3C71840725ABAD" italics="true" pageId="0" pageNumber="187">Eremotherium</emphasis>
</taxonomicName>
is based on the Hypsodontic index (HI) known for ground sloths. According to
<bibRefCitation id="293AD2D6461D7911670D0218A0E7D889" author="Bargo, MS" journalOrPublisher="Acta Palaeontologica Polonica" pageId="0" pageNumber="187" pagination="53 - 61" refId="B5" refString="Bargo, MS, De Iullis, G, Vizcaino, S, 2006. Hypsodonty in Pleistocene ground sloth. Acta Palaeontologica Polonica 51 (1): 53 - 61" title="Hypsodonty in Pleistocene ground sloth." volume="51" year="2006">Bargo et al. (2006)</bibRefCitation>
, HI is an element that is quantified by using the depth of the mandible (DM) (below the third molariform tooth) divided by the length of the molariform tooth row (LTR). In the right hemi-mandible collected in Cauca
<normalizedToken id="5CB000BFEA0CBF7B0EB9169BE1192301" originalValue="“B”">&quot;B&quot;</normalizedToken>
(AMU-CURS-1268b), the DM is 140 mm and the LTR is 180 mm, with a resulting IH equal to 0.77, a value that coincides with
<taxonomicName id="D7FD58F1C587FB77301CAAF15BE8F196" authorityName="Spillmann" authorityYear="1948" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="E8C08CE78563F0272E86DEFD9426E06E" italics="true" pageId="0" pageNumber="187">Eremotherium</emphasis>
</taxonomicName>
whose range oscillates between 0.73 and 0.83 (
<bibRefCitation id="7CEEA99D40656D94664324870EDD1E96" author="De Iuliis, G" journalOrPublisher="CIPICS / South American Quaternary Documentation Program, Saarbrucken, Germany" pageId="0" pageNumber="187" refId="B38" refString="De Iuliis, G, 1996. A Systematic Review of the Megatheriinae (Mammalia: Xenarthra: Megatheriidae). PhD Thesis, University of Toronto, Toronto, Canada." title="A Systematic Review of the Megatheriinae (Mammalia: Xenarthra: Megatheriidae). PhD Thesis, University of Toronto, Toronto, Canada." year="1996">De Iuliis 1996</bibRefCitation>
;
<bibRefCitation id="B83AE4596BEFCFF32F135A0251D5BFFF" author="Bargo, MS" journalOrPublisher="Acta Palaeontologica Polonica" pageId="0" pageNumber="187" pagination="53 - 61" refId="B5" refString="Bargo, MS, De Iullis, G, Vizcaino, S, 2006. Hypsodonty in Pleistocene ground sloth. Acta Palaeontologica Polonica 51 (1): 53 - 61" title="Hypsodonty in Pleistocene ground sloth." volume="51" year="2006">Bargo et al. 2006</bibRefCitation>
). Based on the taxonomic validity of the Late Pleistocene
<taxonomicName id="9369BED0600B5D17E82247F2EF9C935D" authorityName="Spillmann" authorityYear="1948" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="A44AA0E8E34078C3EC22FE09AD6FDCDF" italics="true" pageId="0" pageNumber="187">Eremotherium</emphasis>
</taxonomicName>
species suggested by
<bibRefCitation id="6FCF566D5387D24FBD7B904D8E12A085" author="Cartelle, C" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="0" pageNumber="187" pagination="830 - 841" publicationUrl="https://www.jstor.org/stable/4523673" refId="B26" refString="Cartelle, C, De Iuliis, G, 1995. Eremotherium laurillardi: the panamerican Late Pleistocene megatheriid sloth. Journal of Vertebrate Paleontology 15 (4): 830 - 841, https://www.jstor.org/stable/4523673" title="Eremotherium laurillardi: the panamerican Late Pleistocene megatheriid sloth." url="https://www.jstor.org/stable/4523673" volume="15" year="1995">Cartelle and De Iuliis (1995)</bibRefCitation>
and
<bibRefCitation id="6F6297B4F850A37D7BFAED487B187EC9" DOI="https://doi.org/10.1016/j.crpv.2014.09.002" author="Cartelle, C" journalOrPublisher="Comptes Rendus Palevol" pageId="0" pageNumber="187" pagination="15 - 23" refId="B27" refString="Cartelle, C, De Iuliis, G, Pujos, F, 2015. Eremotherium laurillardi (Lund, 1842) (Xenarthra, Megatheriinae) is the only valid megatheriine sloth species in the Pleistocene of intertropical Brazil: A response to Faure et al., 2014. Comptes Rendus Palevol 14 (1): 15 - 23, DOI: https://doi.org/10.1016/j.crpv.2014.09.002" title="Eremotherium laurillardi (Lund, 1842) (Xenarthra, Megatheriinae) is the only valid megatheriine sloth species in the Pleistocene of intertropical Brazil: A response to Faure et al., 2014." url="https://doi.org/10.1016/j.crpv.2014.09.002" volume="14" year="2015">Cartelle et al. (2015)</bibRefCitation>
, we tentatively suggest assigning the specimens collected at the Cauca site as cf.
<taxonomicName id="2967DB29A0B62228D3A092290C651C97" baseAuthorityName="Lund" baseAuthorityYear="1842" class="Mammalia" family="Megatheriidae" genus="Eremotherium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremotherium laurillardi" order="Carnivora" pageId="0" pageNumber="187" phylum="Chordata" rank="species" species="laurillardi">
<emphasis id="23CB5D743D98B077EA4B03E8E26F6777" italics="true" pageId="0" pageNumber="187">Eremotherium laurillardi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9A851F71F0AA95642AC0A74D9F446969" pageId="0" pageNumber="187">
<emphasis id="CF0F651A18729C38A2977CEB71825F2A" bold="true" pageId="0" pageNumber="187">
<taxonomicName id="97DE22633EB5CCEF38FC7B6633D904D6" family="Mylodontidae" kingdom="Animalia" lsidName="" order="Phyllophaga" pageId="0" pageNumber="187" rank="family">Mylodontidae</taxonomicName>
Gill, 1872
</emphasis>
</paragraph>
</subSubSection>
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@ -0,0 +1,102 @@
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<mods:role>
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<mods:namePart>Stojkoska, Emilija</mods:namePart>
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<paragraph pageId="0" pageNumber="977">Palearctic.</paragraph>
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<subSubSection pageId="0" pageNumber="977" type="notes">
<paragraph pageId="0" pageNumber="977">Notes</paragraph>
<paragraph pageId="0" pageNumber="977">
Previously recorded from unspecified locality between Resen and Ohrid (
<bibRefCitation author="Drensky, P." journalOrPublisher="Spisanie na Bulgarska Akademiya na Naukite" pageId="0" pageNumber="977" pagination="1 - 76" title="Spinnen (Araneae) aus Mittel und Sued-West Mazedonien" volume="39" year="1929">Drensky 1929</bibRefCitation>
,
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).
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<mods:affiliation>Department of Integrative Biology, University of Guelph and Biodiversity Institute of Ontario, Guelph, Canada</mods:affiliation>
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</mods:name>
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</mods:role>
<mods:namePart>Hebert, Paul</mods:namePart>
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</mods:name>
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</mods:role>
<mods:namePart>Shaw, Scott R.</mods:namePart>
<mods:affiliation>Department of Ecosystem Science, University of Wyoming, 1000 East University Avenue, Laramie, Wyoming 82071, USA</mods:affiliation>
</mods:name>
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<mods:namePart>Kittel, Rebecca N.</mods:namePart>
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<mods:affiliation>Museum Wiesbaden, Hessisches Landesmuseum fuer Kunst und Natur, Friedrich-Ebert-Allee 2, 65185 Wiesbaden, Germany</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart>Solis, M. Alma</mods:namePart>
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<mods:affiliation>Systematic Entomology Laboratory, Beltsville Agriculture Research Center, Agricultural Research Service, U. S. Department of Agriculture, c / o National Museum Natural History, MRC 168, Smithsonian Institution, P. O. Box 37012, Washington, DC, 20013 - 7012, USA</mods:affiliation>
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<mods:namePart>Metz, Mark A.</mods:namePart>
<mods:affiliation>Systematic Entomology Laboratory, Beltsville Agriculture Research Center, Agricultural Research Service, U. S. Department of Agriculture, c / o National Museum Natural History, MRC 168, Smithsonian Institution, P. O. Box 37012, Washington, DC, 20013 - 7012, USA</mods:affiliation>
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<mods:namePart>Goldstein, Paul Z.</mods:namePart>
<mods:affiliation>Systematic Entomology Laboratory, Beltsville Agriculture Research Center, Agricultural Research Service, U. S. Department of Agriculture, c / o National Museum Natural History, MRC 168, Smithsonian Institution, P. O. Box 37012, Washington, DC, 20013 - 7012, USA</mods:affiliation>
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<mods:namePart>Brown, John W.</mods:namePart>
<mods:affiliation>Division of Entomology, PO Box 37012 12. National Museum of Natural History E 515 MRC 127, Washington, DC 20013 - 7012, USA</mods:affiliation>
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<mods:namePart>Quicke, Donald L. J.</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Life Sciences, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart>Achterberg, C. van</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, Postbus 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:namePart>Brown, Brian V.</mods:namePart>
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<mods:affiliation>Department of Entomology, Natural History Museum of Los Angeles County, 900 Exposition Boulevard, Los Angeles, CA, 90007, USA</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burns, John M.</mods:namePart>
<mods:affiliation>Division of Entomology, PO Box 37012 12. National Museum of Natural History E 515 MRC 127, Washington, DC 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1013</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>665</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1013.55600</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1013.55600</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1013-1</mods:identifier>
<mods:identifier type="ZooBank">CFDCEFBB523040339D46E302F66E9886</mods:identifier>
<mods:identifier type="Pensoft-UUID">E4329863A39E5EEBA395938413BDD579</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="177205944" LSID="urn:lsid:zoobank.org:act:AD8DF9AB-82C3-43F2-9CC1-BE14B63D688A" httpUri="http://treatment.plazi.org/id/098FF1D035FCC2FDC34681F887E657D2" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/AD8DF9AB-82C3-43F2-9CC1-BE14B63D688A" authority="Sharkey &amp; van Achterberg" authorityName="Sharkey &amp; van Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Hymenochaonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymenochaonia kateperezae" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kateperezae" status="sp. nov.">Hymenochaonia kateperezae Sharkey &amp; van Achterberg</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 286" captionStartId="F286" captionText="Figure 286. Hymenochaonia kateperezae, holotype." figureDoi="10.3897/zookeys.1013.55600.figure286" httpUri="https://binary.pensoft.net/fig/503741" pageId="0" pageNumber="1">Figure 286</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnostics">
<paragraph pageId="0" pageNumber="1">Diagnostics.</paragraph>
<paragraph pageId="0" pageNumber="1">
BOLD:AAA7061. Consensus barcode. GGTTTTATATTTTATATTTGGGTTATGGGCGGGTATAATTGGKATATCAATAAGATTAGTTATTCGTTTAGAATTAGGTCAATCAGGGGTATTAATTGGTAATGAYCAAATTTATAATAGATTTGTAACTGCCCATGCTTTTGTTATAATTTTTTTTATAGTTATACCTATTATAATTGGGGGGTTTGGTAATTGATTAATTCCTTTAATATTAGGAAGGGTTGATATAGCATTTCCTCGGATAAATAATATGAGATTTTGATTATTAATTCCTTCAATTAATTTATTAGTTTTTAGGGGATTTATAAATATTGGGGTAGGTACTGGGTGAACTGTTTATCCCCCTTTATCATTAAATATTAGACATATAGGTATTTCTGTGGATATAGCAATTTTTTCTTTACATTTAGCAGGGGTTTCTTCTATTATAGGAGCTGTAAATTTTATTGTAACTATTATAAATATACGAAATAATAAAGTATTTATAGATAAAATTAGATTATTATGTTGATCTATTTTAATTACTGYAATTTTATTATTATTGTCTTTACCAGTTTTAGCYGGGGCAATTACAATATTATTAACTGATCGTAATTTAAATACATCTTTTTTTGACCCAGCAGGTGGAGGGGATCCTGTTTTATACCAGCATTTATTT. The long vein 2-SR+M of forewing (ca. as long as vein m-cu), ovipositor sheath ca. 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as body and simple tarsal claws are shared with the
<taxonomicName lsidName="H. texana" pageId="0" pageNumber="1" rank="species" species="texana">
<emphasis italics="true" pageId="0" pageNumber="1">H. texana</emphasis>
</taxonomicName>
(Muesebeck, 1932) from USA and Mexico. However, the new species has vein 1Cua distinctly wider than vein 2-Cu1 (similar in
<taxonomicName lsidName="H. texana" pageId="0" pageNumber="1" rank="species" species="texana">
<emphasis italics="true" pageId="0" pageNumber="1">H. texana</emphasis>
</taxonomicName>
), POL ca. equal to diameter of posterior ocellus (ca. 2
<normalizedToken originalValue="×">x</normalizedToken>
diameter of posterior ocellus in
<taxonomicName lsidName="H. texana" pageId="0" pageNumber="1" rank="species" species="texana">
<emphasis italics="true" pageId="0" pageNumber="1">H. texana</emphasis>
</taxonomicName>
), vertex laterally pale yellowish (dark brown in
<taxonomicName lsidName="H. texana" pageId="0" pageNumber="1" rank="species" species="texana">
<emphasis italics="true" pageId="0" pageNumber="1">H. texana</emphasis>
</taxonomicName>
) and vein cu-a of forewing inclivous (vertical in
<taxonomicName lsidName="H. texana" pageId="0" pageNumber="1" rank="species" species="texana">
<emphasis italics="true" pageId="0" pageNumber="1">H. texana</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="holotype ♀">
<paragraph pageId="0" pageNumber="1">Holotype ♀.</paragraph>
<paragraph pageId="0" pageNumber="1">
Guanacaste, Sector San Cristobal, Tajo Angeles,
<geoCoordinate degrees="10.86472" direction="north" orientation="latitude" precision="1" value="10.86472">10.86472</geoCoordinate>
,
<geoCoordinate degrees="85.41531" direction="west" orientation="longitude" precision="1" value="-85.41531">-85.41531</geoCoordinate>
, 540 meters, caterpillar collection date: 23/vii/2011, wasp eclosion date: 15/viii/2011. Depository: CNC.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Host data</emphasis>
.
</emphasis>
elachBioLep01 BioLep754 (
<taxonomicName authorityName="Meyrick" authorityYear="1883" class="Insecta" family="Depressariidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Depressariidae</taxonomicName>
) feeding on
<taxonomicName class="Dicotyledoneae" family="Anacardiaceae" genus="Tapirira" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Tapirira brenesii" order="Sapindales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="brenesii">
<emphasis italics="true" pageId="0" pageNumber="1">Tapirira brenesii</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="Aublet" authorityYear="1775" class="Dicotyledoneae" family="Anacardiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Sapindales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="family">Anacardiaceae</taxonomicName>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Caterpillar and holotype voucher codes</emphasis>
.
</emphasis>
11-SRNP-2902, DHJPAR0045062.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="paratypes">
<paragraph pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="1">
Host =
<taxonomicName authorityName="Zeller" authorityYear="1839" class="Insecta" family="Depressariidae" genus="Stenoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stenoma" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Stenoma</emphasis>
</taxonomicName>
BioLep86: DHJPAR0035531, DHJPAR0035533. Depository: CNC.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sharkey &amp; van Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Hymenochaonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymenochaonia kateperezae" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kateperezae">
<emphasis italics="true" pageId="0" pageNumber="1">Hymenochaonia kateperezae</emphasis>
</taxonomicName>
is named to honor Ms. Kate Perez for her many years laboring in execution and administration of the field and laboratory actions performed by the Centre for Biodiversity Genomics to rapidly and accurately DNA barcode tens of thousands of Costa Rican insects from Malaise traps installed by ACG and BioAlfa.
</paragraph>
<caption doi="10.3897/zookeys.1013.55600.figure286" httpUri="https://binary.pensoft.net/fig/503741" pageId="0" pageNumber="1" start="Figure 286" startId="F286">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 286.</emphasis>
<taxonomicName authorityName="Sharkey &amp; van Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Hymenochaonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymenochaonia kateperezae" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kateperezae">
<emphasis italics="true" pageId="0" pageNumber="1">Hymenochaonia kateperezae</emphasis>
</taxonomicName>
, holotype.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>