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<document id="A767C265C9F86FED71222B296B1FF3FB" ID-DOI="10.15298/rusentj.30.3.05" ID-ISSN="0132-8069" ID-Zenodo-Dep="13179717" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722596800136" checkinUser="felipe" docAuthor="Lyubarsky, G. Yu. &amp; Perkovsky, E. E." docDate="2021" docId="FA7A878A2543FFD31359F8601FCBFCA6" docLanguage="en" docName="RussEntomolJ.30.3.282-287.pdf" docOrigin="Russian Entomological Journal 30 (3)" docSource="http://dx.doi.org/10.15298/rusentj.30.3.05" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Atomaria groehni Perkovsky et Lyubarsky 2014" docType="treatment" docVersion="2" lastPageNumber="286" masterDocId="0643FFF22542FFD71028FFD41D09FFB1" masterDocTitle="New findings of Cryptophagidae (Coleoptera: Clavicornia) from Baltic amber in the unbiased collection of the Paleontological Institute of RAS" masterLastPageNumber="287" masterPageNumber="282" pageNumber="283" updateTime="1722637898107" updateUser="ExternalLinkService">
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<mods:title id="8CA4CAFE1E732DA720E28FCEFF26F9F5">New findings of Cryptophagidae (Coleoptera: Clavicornia) from Baltic amber in the unbiased collection of the Paleontological Institute of RAS</mods:title>
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<mods:namePart id="42DAA46F50BCBCEC03F359204CA2CABB">Lyubarsky, G. Yu.</mods:namePart>
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<mods:namePart id="FA8603C9C21ADD07698FE25771815EB5">Perkovsky, E. E.</mods:namePart>
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<mods:title id="B867176A3B5AE5A68D8488D4077C62EA">Russian Entomological Journal</mods:title>
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<mods:date id="4FD91C77024A1943DCDBF6A7E5867721">2021</mods:date>
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<subSubSection id="3AC965172543FFD61359F860198DF859" pageId="1" pageNumber="283" type="nomenclature">
<paragraph id="726C369C2543FFD61359F860198DF859" blockId="1.[850,1398,1971,2057]" pageId="1" pageNumber="283">
<heading id="292481F02543FFD61359F860198DF859" centered="true" fontSize="10" level="2" pageId="1" pageNumber="283" reason="8">
<taxonomicName id="B5D34D1F2543FFD61359F860183BF87A" authority="Perkovsky et Lyubarsky, 2014" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[881,1330,1971,1995]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="40A7EA8E2543FFD61359F8601EEBF87A" box="[881,994,1971,1995]" italics="true" pageId="1" pageNumber="283">A. groehni</emphasis>
Perkovsky et
<bibRefCitation id="16424B6D2543FFD61453F867183BF87A" author="Lyubarsky G. Yu. &amp; Perkovsky E. E." box="[1147,1330,1971,1995]" pageId="1" pageNumber="283" pagination="191 - 194" refId="ref4132" refString="Lyubarsky G. Yu., Perkovsky E. E. 2014. New species of the genus Nganasania from Upper Cretaceous of Taymyr (Coleoptera: Cryptophagidae) // Russian Entomol. J. Vol. 23. No. 3. P. 191 - 194." type="journal article" year="2014">Lyubarsky, 2014</bibRefCitation>
</taxonomicName>
<figureCitation id="EAE82A192543FFD61437F8051989F859" box="[1055,1152,2000,2025]" captionStart="Figs 5–6" captionStartId="3.[226,263,2014,2033]" captionTargetBox="[189,1399,474,2000]" captionTargetId="figure-133@3.[189,1399,474,2000]" captionTargetPageId="3" captionText="Figs 5–6. Photo of a new specimen of A. groehni from Baltic amber: 5 — dorsal view; 6 — dorsolateral view. Рис. 5–6. ВнеШний вид A. groehni иЗ балтийского ЯнтарЯ: 5 — сверху; 6 — сверху и сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179723" httpUri="https://zenodo.org/record/13179723/files/figure.png" pageId="1" pageNumber="283">Figs 5–6</figureCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3AC965172543FFD3137AF8221FC3FE36" lastPageId="4" lastPageNumber="286" pageId="1" pageNumber="283" type="materials_examined">
<paragraph id="726C369C2543FFD5137AF8221C1CFED7" blockId="1.[850,1398,1971,2057]" lastBlockId="2.[189,776,315,599]" lastPageId="2" lastPageNumber="284" pageId="1" pageNumber="283">
MATERIAL.
<collectionCode id="14C2AE592543FFD613E5F8221EF9F7B8" box="[973,1008,2038,2057]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="1" pageNumber="283">PIN</collectionCode>
964/1325, Baltic amber, Late Eocene. Specimen is deposited in the amber collection of the
<collectionCode id="14C2AE592540FFD51254FEE81F96FEFE" box="[636,671,316,335]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="2" pageNumber="284">PIN</collectionCode>
RAS. Sex: unknown.
</paragraph>
<paragraph id="726C369C2540FFD510CAFEBF1F16FE60" blockId="2.[189,776,315,599]" pageId="2" pageNumber="284">
RE-DESCRIPTION. Length of body 1.0–
<quantity id="B52B9B792540FFD51257FEBF1FC2FE30" box="[639,715,363,385]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="284" unit="mm" value="1.1">1.1 mm</quantity>
, body elongate (
<figureCitation id="EAE82A192540FFD51133FE521C7BFE2D" box="[283,370,390,412]" captionStart="Figs 5–6" captionStartId="3.[226,263,2014,2033]" captionTargetBox="[189,1399,474,2000]" captionTargetId="figure-133@3.[189,1399,474,2000]" captionTargetPageId="3" captionText="Figs 5–6. Photo of a new specimen of A. groehni from Baltic amber: 5 — dorsal view; 6 — dorsolateral view. Рис. 5–6. ВнеШний вид A. groehni иЗ балтийского ЯнтарЯ: 5 — сверху; 6 — сверху и сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179723" httpUri="https://zenodo.org/record/13179723/files/figure.png" pageId="2" pageNumber="284">Figs 5–6</figureCitation>
), moderately arched, head, pronotum, and elytra light-brown.Elytra slightly convex, covered with slightly curved long elevated pubescence.
</paragraph>
<paragraph id="726C369C2540FFD510CAFE0219ECFE0A" blockId="2.[189,776,315,599]" lastBlockId="2.[812,1399,315,604]" pageId="2" pageNumber="284">
Head transverse, of normal size, with hemispherical facetted eyes, strongly punctured. Antennae moderately long, with club reaching beyond base of pronotum, flagellomeres slightly elongate, antennomeres 1
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, 2
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about 1.5 times as long as broad, 3
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slightly elongate, 4
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and 6
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antennomeres as long as broad, 5
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1.5 times as long as broad, 7
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, 8
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subquadrate, 9
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slightly transverse, 10
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strongly transverse, 11
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obliquely oval, antennomeres 9–11 equal in width. Antennae inserted under the lateral margin of the forehead, widely separated basally, antennal insertions close to the eyes.
</paragraph>
<paragraph id="726C369C2540FFD4137AFE151C50FE0D" blockId="2.[812,1399,315,604]" lastBlockId="3.[189,775,315,444]" lastPageId="3" lastPageNumber="285" pageId="2" pageNumber="284">Pronotum convex, not narrowed basally, slightly transverse, 1.2–1.5 times broader than long, broadest before the middle, moderately strongly and very densely punctured (distance between punctures equal to their diameter), a single puncture smaller than the diameter of the eye facet. Pubescence long and elevated. Sides finely margined, anterior edge weakly sinuate, without excision. Pronotum broadest before middle of its length. Side borders visible from above along the entire length. Lateral margin without callosity. Posterior corners right angled, basal edge lobed, strongly sinuate, basal groove shallow.</paragraph>
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<paragraph id="726C369C2540FFD510CAF87A1CE1F868" blockId="2.[189,1399,1966,2061]" pageId="2" pageNumber="284">
Figs 1–4. Photo of a new specimen of
<taxonomicName id="B5D34D1F2540FFD51206F87A1F92F871" authorityName="Lyubarsky et Perkovsky" authorityYear="2010" box="[558,667,1966,1984]" class="Insecta" family="Cryptophagidae" genus="Micrambe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="284" phylum="Arthropoda" rank="species" species="sarnensis">
<emphasis id="40A7EA8E2540FFD51206F87A1F92F871" box="[558,667,1966,1984]" italics="true" pageId="2" pageNumber="284">M. sarnensis</emphasis>
</taxonomicName>
from Baltic amber: 1 — dorsal view; 2 — lateral view; 3 — ventral view; 4 — head and pronotum, ventrolateral view.
</paragraph>
<paragraph id="726C369C2540FFD510CAF8361F3AF7BC" blockId="2.[189,1399,1966,2061]" pageId="2" pageNumber="284">
Рис. 1–4. ВнеШний вид
<taxonomicName id="B5D34D1F2540FFD51196F8361F24F845" authorityName="Lyubarsky et Perkovsky" authorityYear="2010" box="[446,557,2018,2036]" class="Insecta" family="Cryptophagidae" genus="Micrambe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="284" phylum="Arthropoda" rank="species" species="sarnensis">
<emphasis id="40A7EA8E2540FFD51196F8361F24F845" box="[446,557,2018,2036]" italics="true" pageId="2" pageNumber="284">M. sarnensis</emphasis>
</taxonomicName>
иЗ балтийского ЯнтарЯ: 1 — габитус, сверху; 2 — габитус, сбоку; 3 — габитус,сниЗу; 4 — голова и переднеспинка, сниЗу и сбоку.
</paragraph>
</caption>
<paragraph id="726C369C2541FFD3137AFEEF1FC3FE36" blockId="3.[812,1398,315,444]" lastBlockId="4.[189,776,315,791]" lastPageId="4" lastPageNumber="286" pageId="3" pageNumber="285">Scutellum small, transverse. Elytra short-oval, moderately convex, humeral corners rounded, maximum breadth of elytra in first third of their length, 1.6–1.7 times as long as wide and 2.5–2.7 times as long as thorax, punctuation slightly stronger and sparser than that on pronotum. Punctures in the basal part slightly stronger than those on the pronotal disk, and approximately 1.5–2 diameters apart from their lateral neighbours on an average; humeri not toothed.</paragraph>
</subSubSection>
<caption id="26AC66142541FFD410CAF80A193AF7BC" ID-DOI="http://doi.org/10.5281/zenodo.13179723" ID-Zenodo-Dep="13179723" httpUri="https://zenodo.org/record/13179723/files/figure.png" pageId="3" pageNumber="285" startId="3.[226,263,2014,2033]" targetBox="[189,1399,474,2000]" targetPageId="3" targetType="figure">
<paragraph id="726C369C2541FFD410CAF80A19A2F840" blockId="3.[226,1195,2013,2061]" box="[226,1195,2013,2033]" pageId="3" pageNumber="285">
Figs 5–6. Photo of a new specimen of
<taxonomicName id="B5D34D1F2541FFD4121DF80A1F99F840" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[565,656,2014,2033]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="285" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="40A7EA8E2541FFD4121DF80A1F99F840" box="[565,656,2014,2033]" italics="true" pageId="3" pageNumber="285">A. groehni</emphasis>
</taxonomicName>
from Baltic amber: 5 — dorsal view; 6 — dorsolateral view.
</paragraph>
<paragraph id="726C369C2541FFD410CAF82E193AF7BC" blockId="3.[226,1195,2013,2061]" box="[226,1075,2041,2061]" pageId="3" pageNumber="285">
Рис. 5–6. ВнеШний вид
<taxonomicName id="B5D34D1F2541FFD41197F82E1F12F7BD" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[447,539,2041,2061]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="285" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="40A7EA8E2541FFD41197F82E1F12F7BD" box="[447,539,2041,2061]" italics="true" pageId="3" pageNumber="285">A. groehni</emphasis>
</taxonomicName>
иЗ балтийского ЯнтарЯ: 5 — сверху; 6 — сверху и сбоку.
</paragraph>
</caption>
<subSubSection id="3AC965172546FFD310CAFE5F1FCBFCA6" pageId="4" pageNumber="286" type="discussion">
<paragraph id="726C369C2546FFD310CAFE5F1F2AFE67" blockId="4.[189,776,315,791]" pageId="4" pageNumber="286">
REMARKS. Representatives of
<taxonomicName id="B5D34D1F2546FFD3120EFE581F88FE10" authorityName="Stephens" authorityYear="1829" box="[550,641,396,417]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="40A7EA8E2546FFD3120EFE581F88FE10" box="[550,641,396,417]" italics="true" pageId="4" pageNumber="286">Atomaria</emphasis>
</taxonomicName>
are found in all biogeographic realms including 127 species in the Palaearctic Region [
<bibRefCitation id="16424B6D2546FFD31164FE151F11FE67" author="Johnson C. &amp; Otero J. C. &amp; Leschen R. A. B." box="[332,536,449,471]" pageId="4" pageNumber="286" pagination="513 - 531" refId="ref3479" refString="Johnson C., Otero J. C., Leschen R. A. B. 2007. Family Cryptophagidae // Lobl I., Smetana A. (eds.). A catalogue of Palaearctic Coleoptera. Vol. 4. Steenstrup: Apollo Books. P. 513 - 531." type="book chapter" year="2007">Johnson et al., 2007</bibRefCitation>
].
</paragraph>
<paragraph id="726C369C2546FFD310CAFE081E01FD1D" blockId="4.[189,776,315,791]" pageId="4" pageNumber="286">
<taxonomicName id="B5D34D1F2546FFD310CAFE081C84FE41" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[226,397,475,497]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="40A7EA8E2546FFD310CAFE081C84FE41" box="[226,397,475,497]" italics="true" pageId="4" pageNumber="286">Atomaria groehni</emphasis>
</taxonomicName>
differs from similar species with lateral margin of pronotum visible from above, such as
<taxonomicName id="B5D34D1F2546FFD31287FE221C40FD97" authority="Kolenati, 1846" authorityName="Kolenati" authorityYear="1846" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="40A7EA8E2546FFD31287FE221E0EFDBD" box="[687,775,502,524]" italics="true" pageId="4" pageNumber="286">A. morio</emphasis>
Kolenati, 1846
</taxonomicName>
and
<taxonomicName id="B5D34D1F2546FFD3115DFDC51F8DFD97" authority="Erichson, 1846" authorityName="Erichson" authorityYear="1846" box="[373,644,528,551]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="versicolor">
<emphasis id="40A7EA8E2546FFD3115DFDC51CE7FD97" box="[373,494,529,550]" italics="true" pageId="4" pageNumber="286">A. versicolor</emphasis>
Erichson, 1846
</taxonomicName>
, by small size and long elevated pubescence.
<taxonomicName id="B5D34D1F2546FFD311CAFDF81F4FFDF1" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[482,582,555,577]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="40A7EA8E2546FFD311CAFDF81F4FFDF1" box="[482,582,555,577]" italics="true" pageId="4" pageNumber="286">A. groehni</emphasis>
</taxonomicName>
differs from species similar to
<taxonomicName id="B5D34D1F2546FFD3110DFD921F3BFDEA" authority="Stephens, 1830" authorityName="Stephens" authorityYear="1830" box="[293,562,582,604]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="testacea">
<emphasis id="40A7EA8E2546FFD3110DFD921C98FDED" box="[293,401,582,604]" italics="true" pageId="4" pageNumber="286">A. testacea</emphasis>
Stephens,
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in pronotum slightly narrowed basally and small size.
<taxonomicName id="B5D34D1F2546FFD311DFFDB51F55FDC7" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[503,604,609,630]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="40A7EA8E2546FFD311DFFDB51F55FDC7" box="[503,604,609,630]" italics="true" pageId="4" pageNumber="286">A. groehni</emphasis>
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differs from most species of the genus in the structure of the antenna: the segments of the flagellum are subquadrate or slightly elongate.
</paragraph>
<paragraph id="726C369C2546FFD310CAFD651FCBFCA6" blockId="4.[189,776,315,791]" pageId="4" pageNumber="286">Among the species of amber, it differs in a combination of characters: pronotum broadest before the middle of its length; elytra covered by long elevated pubescence; side borders of pronotum visible from above along the entire length.</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="726C369C2543FFD610E5F8861F1DF839" blockId="1.[189,775,1874,2066]" pageId="1" pageNumber="283">
<heading id="292481F02543FFD610E5F8861F1DF839" centered="true" fontSize="10" level="2" pageId="1" pageNumber="283" reason="8">
<taxonomicName id="B5D34D1F2543FFD610E5F8861FFEF8DB" authority="Lyubarsky et Perkovsky, 2010" authorityName="Lyubarsky et Perkovsky" authorityYear="2010" box="[205,759,1874,1898]" class="Insecta" family="Cryptophagidae" genus="Micrambe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="species" species="sarnensis">
<emphasis id="40A7EA8E2543FFD610E5F8861CAFF8DB" box="[205,422,1874,1898]" italics="true" pageId="1" pageNumber="283">Micrambe sarnensis</emphasis>
<bibRefCitation id="16424B6D2543FFD61185F8861FFEF8DB" author="Lyubarsky G. Yu. &amp; Perkovsky E. E." box="[429,759,1874,1898]" pageId="1" pageNumber="283" pagination="275 - 279" refId="ref4031" refString="Lyubarsky G. Yu., Perkovsky E. E. 2010. The first Eocene species of the genus Micrambe (Coleoptera, Clavicornia, Cryptophagidae) // Vestnik zoologii. Vol. 44. No. 3. P. 275 - 279." type="book chapter" year="2010">Lyubarsky et Perkovsky, 2010</bibRefCitation>
</taxonomicName>
<figureCitation id="EAE82A192543FFD61187F8BB1F19F839" box="[431,528,1903,1928]" captionStart="Figs 1–4" captionStartId="2.[226,263,1966,1985]" captionTargetBox="[189,1399,634,1954]" captionTargetId="figure-76@2.[189,1399,634,1954]" captionTargetPageId="2" captionText="Figs 1–4. Photo of a new specimen of M. sarnensis from Baltic amber: 1 — dorsal view; 2 — lateral view; 3 — ventral view; 4 — head and pronotum, ventrolateral view. Рис. 1–4. ВнеШний вид M. sarnensis иЗ балтийского ЯнтарЯ: 1 — габитус, сверху; 2 — габитус, сбоку; 3 — габитус,сниЗу; 4 — голова и переднеспинка, сниЗу и сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179721" httpUri="https://zenodo.org/record/13179721/files/figure.png" pageId="1" pageNumber="283">Figs 1–4</figureCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3AC965172543FFD610CAF8411913FA9A" pageId="1" pageNumber="283" type="materials_examined">
<paragraph id="726C369C2543FFD610CAF8411C21F7A3" blockId="1.[189,775,1874,2066]" pageId="1" pageNumber="283">
<materialsCitation id="C2BB3CC12543FFD610CAF8411C2DF7A3" collectionCode="PIN" pageId="1" pageNumber="283" specimenCode="PIN 964" specimenCount="1" specimenCount-male="1">
MATERIAL.
<specimenCode id="22759EE72543FFD61172F8411CABF816" box="[346,418,1940,1960]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="1" pageNumber="283">PIN 964</specimenCode>
/1324, Baltic amber, Late Eocene. Sex:
<specimenCount id="64D5FD152543FFD612C5F8431E08F816" box="[749,769,1943,1959]" count="1" pageId="1" pageNumber="283" type="male">♂</specimenCount>
. A new specimen of the recently described species
<taxonomicName id="B5D34D1F2543FFD61095F8131C81F86C" authorityName="Lyubarsky et Perkovsky" authorityYear="2010" box="[189,392,1991,2013]" class="Insecta" family="Cryptophagidae" genus="Micrambe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="species" species="sarnensis">
<emphasis id="40A7EA8E2543FFD61095F8131C81F86C" box="[189,392,1991,2013]" italics="true" pageId="1" pageNumber="283">Micrambe sarnensis</emphasis>
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was found. This allows for a more accurate description of the species, taking into account the variability
</materialsCitation>
.
</paragraph>
<paragraph id="726C369C2543FFD6137AFEEF1862FE36" blockId="1.[812,1399,315,1804]" pageId="1" pageNumber="283">
RE-DESCRIPTION.Body broadly elongate (
<figureCitation id="EAE82A192543FFD614DDFEEF1820FEE1" box="[1269,1321,315,337]" captionStart="Figs 1–4" captionStartId="2.[226,263,1966,1985]" captionTargetBox="[189,1399,634,1954]" captionTargetId="figure-76@2.[189,1399,634,1954]" captionTargetPageId="2" captionText="Figs 1–4. Photo of a new specimen of M. sarnensis from Baltic amber: 1 — dorsal view; 2 — lateral view; 3 — ventral view; 4 — head and pronotum, ventrolateral view. Рис. 1–4. ВнеШний вид M. sarnensis иЗ балтийского ЯнтарЯ: 1 — габитус, сверху; 2 — габитус, сбоку; 3 — габитус,сниЗу; 4 — голова и переднеспинка, сниЗу и сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179721" httpUri="https://zenodo.org/record/13179721/files/figure.png" pageId="1" pageNumber="283">Fig.1</figureCitation>
), slightly convex; head, pronotum, and elytra reddish brown. Elytra slightly convex, covered with almost appressed pubescence.
</paragraph>
<paragraph id="726C369C2543FFD6137AFE5F1841FD20" blockId="1.[812,1399,315,1804]" pageId="1" pageNumber="283">
Head transverse, of normal size, with prominent, hemispherical, somewhat coarsely facetted eyes (
<figureCitation id="EAE82A192543FFD614F1FE72181CFE0A" box="[1241,1301,422,444]" captionStart="Figs 1–4" captionStartId="2.[226,263,1966,1985]" captionTargetBox="[189,1399,634,1954]" captionTargetId="figure-76@2.[189,1399,634,1954]" captionTargetPageId="2" captionText="Figs 1–4. Photo of a new specimen of M. sarnensis from Baltic amber: 1 — dorsal view; 2 — lateral view; 3 — ventral view; 4 — head and pronotum, ventrolateral view. Рис. 1–4. ВнеШний вид M. sarnensis иЗ балтийского ЯнтарЯ: 1 — габитус, сверху; 2 — габитус, сбоку; 3 — габитус,сниЗу; 4 — голова и переднеспинка, сниЗу и сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179721" httpUri="https://zenodo.org/record/13179721/files/figure.png" pageId="1" pageNumber="283">Fig. 4</figureCitation>
), strongly and sparsely punctured. Antennae 11-segmented, long, slen- der, with 3-segmented club, with club reaching beyond the base of pronotum, antennomeres 1–2 elongate, twice as long as wide, 3
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slender, elongate, twice as long as wide, joint 4
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subquadrate, antennomeres 5–6 1.5 times as long as wide, joint 7
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almost equal in length, antennomere 8 slightly transverse, 9
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and 10
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transverse, 10
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1.2 times as long as wide, 11
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obliquely oval, antennomeres 9–11 equal in width.
</paragraph>
<paragraph id="726C369C2543FFD6137AFD421967FCFD" blockId="1.[812,1399,315,1804]" pageId="1" pageNumber="283">Pronotum distinctly transverse, barely 1.8 times broader than long, moderately strongly and densely punctured (0.5–1 diameter apart), an individual puncture almost equal to facette diameter. Pronotum without sublateral line, somewhat convex, slightly rounded at sides, weakly crenulate. Sides finely margined, anterior edge weakly sinuate, posterior edge with broad median lobe, emarginated.</paragraph>
<paragraph id="726C369C2543FFD6137AFC851868FC5A" blockId="1.[812,1399,315,1804]" pageId="1" pageNumber="283">Anterior angles of pronotum with callosity, callosity occupying about one-third or one-fourth of side margin, with a small, elongate-oval patch of bare surface invisible from above; caudolateral corner acute angular, with tip. Lateral edge crenulated, with 7–9 small teeth. Posterior corners obtuse, base round, slightly sinuate, basal groove narrow.</paragraph>
<paragraph id="726C369C2543FFD6137AFC25197EFB90" blockId="1.[812,1399,315,1804]" pageId="1" pageNumber="283">
Tarsal formula 5–5–4, claw simple, without tooth (
<figureCitation id="EAE82A192543FFD61563FC251E5DFB91" captionStart="Figs 1–4" captionStartId="2.[226,263,1966,1985]" captionTargetBox="[189,1399,634,1954]" captionTargetId="figure-76@2.[189,1399,634,1954]" captionTargetPageId="2" captionText="Figs 1–4. Photo of a new specimen of M. sarnensis from Baltic amber: 1 — dorsal view; 2 — lateral view; 3 — ventral view; 4 — head and pronotum, ventrolateral view. Рис. 1–4. ВнеШний вид M. sarnensis иЗ балтийского ЯнтарЯ: 1 — габитус, сверху; 2 — габитус, сбоку; 3 — габитус,сниЗу; 4 — голова и переднеспинка, сниЗу и сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179721" httpUri="https://zenodo.org/record/13179721/files/figure.png" pageId="1" pageNumber="283">Figs 2–3</figureCitation>
). Last of tarsal joints longest.
</paragraph>
<paragraph id="726C369C2543FFD6137AFBF21ED7FAA0" blockId="1.[812,1399,315,1804]" pageId="1" pageNumber="283">
Scutellum small, transverse. Elytra oval, punctuation confused, humeral corners rounded, shoulders a little broader than maximum breadth of pronotum, 1.4–1.5 times as long as wide and 2.8–2.9 times as long as thorax, moderately convex, with moderately strongly rounded sides and a narrowly round- ed apex, puncturation as strong as, yet more sparse than, that on pronotum. Epipleura reduced, present beyond level of posterior margin of 1
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ventrite. Ventrite 1 longer than remaining ventrites.
</paragraph>
<paragraph id="726C369C2543FFD6137AFAC21913FA9A" blockId="1.[812,1399,315,1804]" box="[850,1050,1302,1324]" pageId="1" pageNumber="283">
Length
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.
</paragraph>
</subSubSection>
<subSubSection id="3AC965172543FFD6137AFAE51969F8BD" pageId="1" pageNumber="283" type="discussion">
<paragraph id="726C369C2543FFD6137AFAE51995F9E0" blockId="1.[812,1399,315,1804]" pageId="1" pageNumber="283">
REMARKS.
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<emphasis id="40A7EA8E2543FFD613F0FAE519ABFAF7" box="[984,1186,1329,1350]" italics="true" pageId="1" pageNumber="283">Micrambe sarnensis</emphasis>
</taxonomicName>
was described from
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amber and now found in Baltic amber. The new specimen belongs to this species. The
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<emphasis id="40A7EA8E2543FFD6145AFAB219DAFACA" box="[1138,1235,1382,1403]" italics="true" pageId="1" pageNumber="283">Micrambe</emphasis>
species
</taxonomicName>
are very variable, the most important characters (in addition to the structure of the genitalia) are the structure of the antennae, the structure of the callosity, the relative length and width of the pronotum, and size and number of teeth on the lateral edge. This specimen slightly differs from the
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of
<emphasis id="40A7EA8E2543FFD61527FA381E42F9AD" italics="true" pageId="1" pageNumber="283">
<taxonomicName id="B5D34D1F2543FFD61527FA381E4EF9AD" authorityName="Lyubarsky et Perkovsky" authorityYear="2010" class="Insecta" family="Cryptophagidae" genus="Micrambe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="species" species="sarnensis">M. sarnensis</taxonomicName>
,
</emphasis>
and is within the range of variability that can be observed, for example, in the very widespread
<taxonomicName id="B5D34D1F2543FFD614A0F9F5187BF987" authority="(Payk., 1798)" baseAuthorityName="Payk." baseAuthorityYear="1798" box="[1160,1394,1569,1591]" class="Insecta" family="Cryptophagidae" genus="Micrambe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="species" species="abietis">
<emphasis id="40A7EA8E2543FFD614A0F9F51995F987" box="[1160,1180,1569,1590]" italics="true" pageId="1" pageNumber="283">M</emphasis>
.
<emphasis id="40A7EA8E2543FFD61480F9F519EEF987" box="[1192,1255,1569,1590]" italics="true" pageId="1" pageNumber="283">abietis</emphasis>
(Payk., 1798)
</taxonomicName>
, distributed throughout the Palaearctic.
</paragraph>
<paragraph id="726C369C2543FFD6137AF9821841F967" blockId="1.[812,1399,315,1804]" pageId="1" pageNumber="283">
The most noticeable differences between specimens:slight differences in the structure of antenna, antennomeres 1–2 elongate not transverse; callosity larger than in the
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of
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<emphasis id="40A7EA8E2543FFD61360F9731EB4F90D" box="[840,957,1703,1724]" italics="true" pageId="1" pageNumber="283">M.sarnensis</emphasis>
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, occupying 1/3 of lateral edge. Size of body smaller, length of the
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<emphasis id="40A7EA8E2543FFD61452F91519E6F967" box="[1146,1263,1729,1750]" italics="true" pageId="1" pageNumber="283">M.sarnensis</emphasis>
</taxonomicName>
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.
</paragraph>
<paragraph id="726C369C2543FFD6137AF90F1969F8BD" blockId="1.[812,1399,315,1804]" pageId="1" pageNumber="283">
<taxonomicName id="B5D34D1F2543FFD6137AF90F191CF940" authorityName="Lyubarsky et Perkovsky" authorityYear="2010" box="[850,1045,1755,1777]" class="Insecta" family="Cryptophagidae" genus="Micrambe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="species" species="sarnensis">
<emphasis id="40A7EA8E2543FFD6137AF90F191CF940" box="[850,1045,1755,1777]" italics="true" pageId="1" pageNumber="283">Micrambe sarnensis</emphasis>
</taxonomicName>
is widespread in the late Eocene and in the
<collectingRegion id="B017F87E2543FFD61342F9221EA3F8BA" box="[874,938,1782,1803]" country="Ukraine" name="Rivne" pageId="1" pageNumber="283">Rovno</collectingRegion>
and Baltic amber.
</paragraph>
</subSubSection>
</treatment>
</document>