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<document id="2301F231067F2D71FA91245FDBAEEBA4" ID-DOI="10.15298/rusentj.31.2.06" ID-ISSN="0132-8069" ID-Zenodo-Dep="13179704" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722596705233" checkinUser="felipe" docAuthor="Ponomarenko, A. G., Fedorenko, D. N. & Bashkuev, A. S." docDate="2022" docId="3D3E5E12B11D5C64FECEBA09FC00D3BD" docLanguage="en" docName="RussEntomolJ.31.2.128-131.pdf" docOrigin="Russian Entomological Journal 31 (2)" docSource="http://dx.doi.org/10.15298/rusentj.31.2.06" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Holcoptera rasnitsyni Ponomarenko, Fedorenko & Bashkuev, 2022, sp.n." docType="treatment" docVersion="2" lastPageNumber="131" masterDocId="C107266AB11F5C67FFA5BB32FF85D761" masterDocTitle="A new species of the beetle genus Holcoptera (Coleoptera: Coptoclavidae) from the Upper Triassic of east Ukraine" masterLastPageNumber="131" masterPageNumber="128" pageNumber="130" updateTime="1722637783978" updateUser="ExternalLinkService">
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<mods:title id="A2C48FBC956D38954C7CA9E89BEED24A">A new species of the beetle genus Holcoptera (Coleoptera: Coptoclavidae) from the Upper Triassic of east Ukraine</mods:title>
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<mods:namePart id="CE16C981137DF007BD87D4AA0B8E8038">Ponomarenko, A. G.</mods:namePart>
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<mods:namePart id="1C4DB9726F2ED7902F7AF20118A1783A">Fedorenko, D. N.</mods:namePart>
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<mods:namePart id="4C43A5EDB15360523F11CC6D69EBAB96">Bashkuev, A. S.</mods:namePart>
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<mods:title id="DD36BEB439201F914CB2CD63FE143FF1">Russian Entomological Journal</mods:title>
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<mods:part id="AC72EE5F5573C29EA2E31D81682B21B6">
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<mods:date id="49FFCE39667B801221B6B1301C046E8E">2022</mods:date>
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<mods:number id="4F58C716EC0F3DECF41EE01437A57831">2022-06-30</mods:number>
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<mods:number id="07B0E00E7C1C8CA9C895C0716933004E">31</mods:number>
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<mods:identifier id="A54CE5C485E837FF0293D31DF090319A" type="ISSN">0132-8069</mods:identifier>
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<treatment id="3D3E5E12B11D5C64FECEBA09FC00D3BD" LSID="urn:lsid:plazi:treatment:3D3E5E12B11D5C64FECEBA09FC00D3BD" httpUri="http://treatment.plazi.org/id/3D3E5E12B11D5C64FECEBA09FC00D3BD" lastPageId="3" lastPageNumber="131" pageId="2" pageNumber="130">
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<subSubSection id="FD8DBC8FB11D5C65FECEBA09FD12D610" pageId="2" pageNumber="130" type="nomenclature">
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<paragraph id="B528EF04B11D5C65FECEBA09FDD2D632" blockId="2.[189,776,315,982]" box="[363,599,315,339]" pageId="2" pageNumber="130">
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<heading id="EE605868B11D5C65FECEBA09FDD2D632" box="[363,599,315,339]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="130" reason="8">
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<taxonomicName id="72979487B11D5C65FECEBA09FDD2D632" authorityName="Ponomarenko & Fedorenko & Bashkuev" authorityYear="2022" box="[363,599,315,339]" class="Insecta" family="Coptoclavidae" genus="Holcoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="130" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. n.">
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<emphasis id="87E33316B11D5C65FECEBA09FDD2D632" box="[363,599,315,339]" italics="true" pageId="2" pageNumber="130">Holcoptera rasnitsyni</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="B528EF04B11D5C65FE8EBA6BFD12D610" blockId="2.[189,776,315,982]" box="[299,663,345,369]" pageId="2" pageNumber="130">
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Ponomarenko et Fedorenko,
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<emphasis id="87E33316B11D5C65FDC7BA6BFD12D610" bold="true" box="[610,663,345,369]" pageId="2" pageNumber="130">
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<taxonomicNameLabel id="9CD08E6DB11D5C65FDC7BA6BFD12D610" box="[610,663,345,369]" pageId="2" pageNumber="130" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="FD8DBC8FB11D5C65FE0ABA44FD91D6EF" box="[431,532,374,398]" pageId="2" pageNumber="130" type="description">
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<paragraph id="B528EF04B11D5C65FE0ABA44FD91D6EF" blockId="2.[189,776,315,982]" box="[431,532,374,398]" pageId="2" pageNumber="130">
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<figureCitation id="2DACF381B11D5C65FE0ABA44FD95D6EF" box="[431,528,374,398]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Figs 3–5" captionStartId-0="1.[226,258,2017,2036]" captionStartId-1="2.[226,258,1921,1940]" captionStartId-2="3.[226,263,984,1003]" captionTargetBox-0="[189,1399,1168,1997]" captionTargetBox-1="[189,1398,998,1881]" captionTargetBox-2="[189,1399,314,962]" captionTargetId-0="figure-421@1.[189,1399,1168,1997]" captionTargetId-1="figure-566@2.[189,1398,998,1881]" captionTargetId-2="figure-314@3.[189,1399,314,962]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Holcoptera rasnitsyni, sp.n.: holotype PIN 3320/204a. Scale bar: 2.0 mm. Рис. 1. Holcoptera rasnitsyni, sp.n.: голотип ПИН 3320/204a. МасШтаб: 2,0 мм." captionText-1="Fig. 2. Holcoptera rasnitsyni, sp.n.: holotype PIN 3320/204a, drawing of wings. Abbreviations: 1r — 1st radial cell; AAP — vein AA 3a” +(AA 3b +(AA 4 +AP 1+2 )); cb — costal bar; cbz — costal bending zone; o — oblongum cell; pst — pterostigma; rc — radial cell; x1, x2, y — veins; z — posterobasal margin of folded wing (jugal fold). Scale bar: 1.0 mm. Рис. 2. Holcoptera rasnitsyni, sp.n.: голотип ПИН 3320/204a, прорисовка крыльев. ОбоЗначениЯ: 1r — 1-Я радиальнаЯ Ячейка; AAP — жилка AA 3a” +(AA3b+(AA 4 +AP 1+2 )); cb — костальнаЯ ось; cbz — Зона костального иЗгиба; o — продолговатаЯ Ячейка; pst — птеростигма; rc — радиальнаЯ Ячейка; x1, x2, y — жилки; z — постеробаЗальный край сложенного крыла (югальнаЯ складка). МасШтаб: 1,0 мм." captionText-2="Figs 3–5. Holcoptera rasnitsyni, sp.n.: 3–4 — paratype PIN 3320/113a, photo and drawing (solid line — elytra, dashed line — wings, dotted line — metanotum); 5 — Paratype PIN 3320/206a. Scale bars: 2.0 mm. Рис. 3–5. Holcoptera rasnitsyni, sp.n.: 3–4 — паратип PIN 3320/113a, фотографиЯ и прорисовка (сплоШнаЯ линиЯ — контуры надкрылий, пунктирнаЯ линиЯ — крыльЯ, точечнаЯ линиЯ — Заднеспинка); 5 — паратип PIN 3320/206a. МасШтаб: 2,0 мм." figureDoi-0="http://doi.org/10.5281/zenodo.13179706" figureDoi-1="http://doi.org/10.5281/zenodo.13179708" figureDoi-2="http://doi.org/10.5281/zenodo.13179710" httpUri-0="https://zenodo.org/record/13179706/files/figure.png" httpUri-1="https://zenodo.org/record/13179708/files/figure.png" httpUri-2="https://zenodo.org/record/13179710/files/figure.png" pageId="2" pageNumber="130">Figs 1–5</figureCitation>
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.
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</paragraph>
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||||
</subSubSection>
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||||
<subSubSection id="FD8DBC8FB11D5C65FF47BAA9FE19D5F7" pageId="2" pageNumber="130" type="materials_examined">
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<paragraph id="B528EF04B11D5C65FF47BAA9FD2ED697" blockId="2.[189,776,315,982]" pageId="2" pageNumber="130">
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||||
<materialsCitation id="05FFE559B11D5C65FF47BAA9FD22D697" collectionCode="PIN" pageId="2" pageNumber="130" specimenCode="PIN 3320" specimenCount="2" typeStatus="holotype">
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MATERIAL.
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<typeStatus id="6A2C51A6B11D5C65FEC4BAA9FE30D6CF" box="[353,437,411,430]" pageId="2" pageNumber="130" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="E531477FB11D5C65FE64BAA9FD9FD6CF" box="[449,538,411,430]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="2" pageNumber="130">PIN 3320</specimenCode>
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/204a±, imprint of incomplete beetle with carbonized organic matter, head, thorax, elytra and hind wings.
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<typeStatus id="6A2C51A6B11D5C65FE97BAF9FE0FD6BF" box="[306,394,459,478]" pageId="2" pageNumber="130" type="paratype">Paratypes</typeStatus>
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:
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<specimenCode id="E531477FB11D5C65FE32BAF9FE74D6BF" box="[407,497,459,478]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="2" pageNumber="130">PIN 3320</specimenCode>
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/113a± (
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<figureCitation id="2DACF381B11D5C65FD99BAF9FD14D6BF" box="[572,657,459,478]" captionStart="Figs 3–5" captionStartId="3.[226,263,984,1003]" captionTargetBox="[189,1399,314,962]" captionTargetId="figure-314@3.[189,1399,314,962]" captionTargetPageId="3" captionText="Figs 3–5. Holcoptera rasnitsyni, sp.n.: 3–4 — paratype PIN 3320/113a, photo and drawing (solid line — elytra, dashed line — wings, dotted line — metanotum); 5 — Paratype PIN 3320/206a. Scale bars: 2.0 mm. Рис. 3–5. Holcoptera rasnitsyni, sp.n.: 3–4 — паратип PIN 3320/113a, фотографиЯ и прорисовка (сплоШнаЯ линиЯ — контуры надкрылий, пунктирнаЯ линиЯ — крыльЯ, точечнаЯ линиЯ — Заднеспинка); 5 — паратип PIN 3320/206a. МасШтаб: 2,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179710" httpUri="https://zenodo.org/record/13179710/files/figure.png" pageId="2" pageNumber="130">Figs 3–4</figureCitation>
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), 3320/205a, incomplete beetles; 3320/206a, isolated elytron (
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<figureCitation id="2DACF381B11D5C65FDCCBAD1FD1AD697" box="[617,671,483,502]" captionStart="Figs 3–5" captionStartId="3.[226,263,984,1003]" captionTargetBox="[189,1399,314,962]" captionTargetId="figure-314@3.[189,1399,314,962]" captionTargetPageId="3" captionText="Figs 3–5. Holcoptera rasnitsyni, sp.n.: 3–4 — paratype PIN 3320/113a, photo and drawing (solid line — elytra, dashed line — wings, dotted line — metanotum); 5 — Paratype PIN 3320/206a. Scale bars: 2.0 mm. Рис. 3–5. Holcoptera rasnitsyni, sp.n.: 3–4 — паратип PIN 3320/113a, фотографиЯ и прорисовка (сплоШнаЯ линиЯ — контуры надкрылий, пунктирнаЯ линиЯ — крыльЯ, точечнаЯ линиЯ — Заднеспинка); 5 — паратип PIN 3320/206a. МасШтаб: 2,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179710" httpUri="https://zenodo.org/record/13179710/files/figure.png" pageId="2" pageNumber="130">Fig. 5</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="B528EF04B11D5C65FF47BAC9FE19D5F7" blockId="2.[189,776,315,982]" pageId="2" pageNumber="130">
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LOCALITY AND HORIZON.
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<collectingCountry id="CD80AF94B11D5C65FD81BAC9FDF1D570" box="[548,628,507,529]" name="Ukraine" pageId="2" pageNumber="130">Ukraine</collectingCountry>
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,
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<collectingRegion id="775321E6B11D5C65FD21BAC9FF6FD54A" country="Ukraine" name="Kharkiv" pageId="2" pageNumber="130">Kharkov Region</collectingRegion>
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, Izyum District, the right bank of the Bereka River (tributary of the Seversky Donets River),
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<quantity id="726F42E1B11D5C65FDFDB902FD0ED527" box="[600,651,560,582]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="130" unit="km" value="3.0">3 km</quantity>
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north of the Velikaya Kamyshevakha village, Dolgy Jar ravine, Garazhovka locality; Upper Triassic, upper Carnian–lower Norian, Protopivka Formation.
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</paragraph>
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</subSubSection>
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<subSubSection id="FD8DBC8FB11D5C65FF47B9A9FE1DD587" pageId="2" pageNumber="130" type="diagnosis">
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<paragraph id="B528EF04B11D5C65FF47B9A9FE1DD587" blockId="2.[189,776,315,982]" pageId="2" pageNumber="130">DIAGNOSIS.The new species has the palest elytra among its congeners, with the external pale field occupying about 2/ 5 of the elytron width.</paragraph>
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</subSubSection>
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<subSubSection id="FD8DBC8FB11D5C64FF47B9D9FEF2D2AE" lastPageId="3" lastPageNumber="131" pageId="2" pageNumber="130" type="description">
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<paragraph id="B528EF04B11D5C65FF47B9D9FE6AD40A" blockId="2.[189,776,315,982]" pageId="2" pageNumber="130">
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DESCRIPTION. Body oval, about twice as long as wide. Elytra pale, each with four, dark, longitudinal stripes in inner three fifths; these stripes subequally distant
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<emphasis id="87E33316B11D5C65FDC8B813FD33D457" box="[621,694,801,822]" italics="true" pageId="2" pageNumber="130">inter se</emphasis>
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; sutural stripe curved slightly laterad along apical margin; outer three reaching neither base nor apex.
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</paragraph>
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<paragraph id="B528EF04B11D5C65FF47B842FCEDD60D" blockId="2.[189,776,315,982]" lastBlockId="2.[812,1399,315,977]" pageId="2" pageNumber="130">Head about as long as wide. Eyes small and round. Pronotum wider than long. Profemur 2.3 times as long as wide. Protibia slender, curved, slightly longer than femur. Metanotum heavily sclerotized, twice as wide as long. Elytra subconvex, with apices widely rounded, less so at sutural angle.</paragraph>
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<paragraph id="B528EF04B11D5C65FCF7BA43FC38D59D" blockId="2.[812,1399,315,977]" pageId="2" pageNumber="130">
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<materialsCitation id="05FFE559B11D5C65FCF7BA43FC3DD59D" collectionCode="PIN, R, MP" pageId="2" pageNumber="130" specimenCount="1" typeStatus="holotype">
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Wings,
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<typeStatus id="6A2C51A6B11D5C65FC05BA43FC70D6E6" box="[928,1013,369,391]" pageId="2" pageNumber="130" type="holotype">holotype</typeStatus>
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<collectionCode id="D38677C1B11D5C65FC58BA43FBA3D6E7" box="[1021,1062,369,390]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="2" pageNumber="130">PIN</collectionCode>
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, No. 3320/204a (
|
||||
<figureCitation id="2DACF381B11D5C65FB70BA43FAB0D6E7" box="[1237,1333,369,391]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="1.[226,258,2017,2036]" captionStartId-1="2.[226,258,1921,1940]" captionTargetBox-0="[189,1399,1168,1997]" captionTargetBox-1="[189,1398,998,1881]" captionTargetId-0="figure-421@1.[189,1399,1168,1997]" captionTargetId-1="figure-566@2.[189,1398,998,1881]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Holcoptera rasnitsyni, sp.n.: holotype PIN 3320/204a. Scale bar: 2.0 mm. Рис. 1. Holcoptera rasnitsyni, sp.n.: голотип ПИН 3320/204a. МасШтаб: 2,0 мм." captionText-1="Fig. 2. Holcoptera rasnitsyni, sp.n.: holotype PIN 3320/204a, drawing of wings. Abbreviations: 1r — 1st radial cell; AAP — vein AA 3a” +(AA 3b +(AA 4 +AP 1+2 )); cb — costal bar; cbz — costal bending zone; o — oblongum cell; pst — pterostigma; rc — radial cell; x1, x2, y — veins; z — posterobasal margin of folded wing (jugal fold). Scale bar: 1.0 mm. Рис. 2. Holcoptera rasnitsyni, sp.n.: голотип ПИН 3320/204a, прорисовка крыльев. ОбоЗначениЯ: 1r — 1-Я радиальнаЯ Ячейка; AAP — жилка AA 3a” +(AA3b+(AA 4 +AP 1+2 )); cb — костальнаЯ ось; cbz — Зона костального иЗгиба; o — продолговатаЯ Ячейка; pst — птеростигма; rc — радиальнаЯ Ячейка; x1, x2, y — жилки; z — постеробаЗальный край сложенного крыла (югальнаЯ складка). МасШтаб: 1,0 мм." figureDoi-0="http://doi.org/10.5281/zenodo.13179706" figureDoi-1="http://doi.org/10.5281/zenodo.13179708" httpUri-0="https://zenodo.org/record/13179706/files/figure.png" httpUri-1="https://zenodo.org/record/13179708/files/figure.png" pageId="2" pageNumber="130">Figs. 1–2</figureCitation>
|
||||
). Two almost completely folded hind wings visible, with apex of right wing somewhat unfolded; no distinct folds. Two longitudinal wing axes located in basal part of hind wing. Costal bar (
|
||||
<emphasis id="87E33316B11D5C65FCFCBAE9FCF5D691" box="[857,880,475,496]" italics="true" pageId="2" pageNumber="130">cb</emphasis>
|
||||
= C+ScA+ScP+
|
||||
<collectionCode id="D38677C1B11D5C65FBB4BAE9FBA7D691" box="[1041,1058,475,496]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="130">R</collectionCode>
|
||||
) distinctly and densely cross-striat- ed, consisting of strong vein C+ScA+ScP and barely detectable radial vein (
|
||||
<collectionCode id="D38677C1B11D5C65FC69B923FC5AD547" box="[972,991,529,550]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="130">R</collectionCode>
|
||||
), which is narrow and desclerotized distally. Cubital bar (Cu–CuA) broken into fragments, entire in distal half only, distally extended into large, well-defined, slightly transverse oblongum cell. Cubital joint at respective pivot on CuA proximal in position, so that CuA is abruptly geniculate at considerable distance from oblongum cell in both folded wings. Median bar (M–
|
||||
<collectionCode id="D38677C1B11D5C65FB36B983FB32D5A7" box="[1171,1207,689,710]" country="0" name="Mohonk Preserve, Inc." pageId="2" pageNumber="130">MP</collectionCode>
|
||||
) not visible except small distal section apparently retained in form of vein ‘
|
||||
<emphasis id="87E33316B11D5C65FAEEB9FEFAD3D580" box="[1355,1366,716,737]" italics="true" pageId="2" pageNumber="130">y</emphasis>
|
||||
’ in left hind wing
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="B528EF04B11D5C65FCF7B833FBAAD4FD" blockId="2.[812,1399,315,977]" pageId="2" pageNumber="130">
|
||||
Clavus and jugal lobe with few veins, broken and shifted basad, two vein fragments being only detectable: (1) CuP without apical section and (2) fragment of clavus “skeleton” with remains of first and second anal cells. Distinct line ‘
|
||||
<emphasis id="87E33316B11D5C65FAEDB863FAD4D407" box="[1352,1361,849,870]" italics="true" pageId="2" pageNumber="130">z</emphasis>
|
||||
’ on the imprint apparently represents no vein, but posterobasal wing margin or jugal fold.
|
||||
</paragraph>
|
||||
<paragraph id="B528EF04B11D5C64FCF7B893FEE9D39B" blockId="2.[812,1399,315,977]" lastBlockId="3.[189,776,1118,1754]" lastPageId="3" lastPageNumber="131" pageId="2" pageNumber="130">
|
||||
Venation of central region of wing hardly detectable. Region around first radial cell and probably also inner borders of carpal cell and first radial cell only traceable. This border is represented by vein
|
||||
<emphasis id="87E33316B11C5C64FE7FBF4BFE74D3EF" box="[474,497,1145,1166]" italics="true" pageId="3" pageNumber="131">x1</emphasis>
|
||||
identifiable as either base of RP or complex vein RP–MA. Vein
|
||||
<emphasis id="87E33316B11C5C64FD89BFA6FDC6D3C8" box="[556,579,1172,1193]" italics="true" pageId="3" pageNumber="131">x2</emphasis>
|
||||
looking like direct extension of vein
|
||||
<emphasis id="87E33316B11C5C64FECABF9DFE03D3A5" box="[367,390,1199,1220]" italics="true" pageId="3" pageNumber="131">x1</emphasis>
|
||||
caudad, suggesting that
|
||||
<emphasis id="87E33316B11C5C64FDD8BF9DFD33D3A5" box="[637,694,1199,1220]" italics="true" pageId="3" pageNumber="131">x1–x2</emphasis>
|
||||
may be simple vein. Yet, attribution of
|
||||
<emphasis id="87E33316B11C5C64FE4BBFFBFD80D3BF" box="[494,517,1225,1246]" italics="true" pageId="3" pageNumber="131">x2</emphasis>
|
||||
to left wing CuA
|
||||
<subScript id="2913ED41B11C5C64FD0ABFEAFD33D385" attach="left" box="[687,694,1240,1252]" fontSize="5" pageId="3" pageNumber="131">1</subScript>
|
||||
appears more appropriate.
|
||||
</paragraph>
|
||||
<caption id="E1E8BF8CB11D5C65FF47BCB3FAE0DF71" ID-DOI="http://doi.org/10.5281/zenodo.13179708" ID-Zenodo-Dep="13179708" httpUri="https://zenodo.org/record/13179708/files/figure.png" pageId="2" pageNumber="130" startId="2.[226,258,1921,1940]" targetBox="[189,1398,998,1881]" targetPageId="2" targetType="figure">
|
||||
<paragraph id="B528EF04B11D5C65FF47BCB3FC19D0A4" blockId="2.[188,1398,1921,2064]" pageId="2" pageNumber="130">
|
||||
Fig. 2.
|
||||
<taxonomicName id="72979487B11D5C65FE80BCB3FE66D0F5" authorityName="Ponomarenko & Fedorenko & Bashkuev" authorityYear="2022" box="[293,483,1921,1940]" class="Insecta" family="Coptoclavidae" genus="Holcoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="130" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
|
||||
<emphasis id="87E33316B11D5C65FE80BCB3FE66D0F5" box="[293,483,1921,1940]" italics="true" pageId="2" pageNumber="130">Holcoptera rasnitsyni</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="87E33316B11D5C65FE54BCB3FD99D0F5" bold="true" box="[497,540,1921,1940]" pageId="2" pageNumber="130">
|
||||
<taxonomicNameLabel id="9CD08E6DB11D5C65FE54BCB3FD99D0F5" box="[497,540,1921,1940]" pageId="2" pageNumber="130" rank="species">sp.n.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
: holotype PIN 3320/204a, drawing of wings. Abbreviations: 1r — 1st radial cell; AAP — vein AA
|
||||
<subScript id="2913ED41B11D5C65FF7EBC94FF69D0D3" attach="left" box="[219,236,1958,1970]" fontSize="5" pageId="2" pageNumber="130">3a”</subScript>
|
||||
+(AA
|
||||
<subScript id="2913ED41B11D5C65FEBABC94FEA9D0D3" attach="right" box="[287,300,1958,1970]" fontSize="5" pageId="2" pageNumber="130">3b</subScript>
|
||||
+(AA
|
||||
<subScript id="2913ED41B11D5C65FEFBBC94FEE1D0D3" attach="right" box="[350,356,1958,1970]" fontSize="5" pageId="2" pageNumber="130">4</subScript>
|
||||
+AP
|
||||
<subScript id="2913ED41B11D5C65FE29BC94FE1AD0D3" attach="right" box="[396,415,1958,1970]" fontSize="5" pageId="2" pageNumber="130">1+2</subScript>
|
||||
));
|
||||
<emphasis id="87E33316B11D5C65FE1FBCABFE4BD0CD" box="[442,462,1945,1964]" italics="true" pageId="2" pageNumber="130">cb</emphasis>
|
||||
— costal bar;
|
||||
<emphasis id="87E33316B11D5C65FDF5BCABFDE8D0CD" box="[592,621,1945,1964]" italics="true" pageId="2" pageNumber="130">cbz</emphasis>
|
||||
— costal bending zone;
|
||||
<emphasis id="87E33316B11D5C65FCE3BCA8FCD4D0CD" box="[838,849,1946,1964]" italics="true" pageId="2" pageNumber="130">o</emphasis>
|
||||
— oblongum cell;
|
||||
<emphasis id="87E33316B11D5C65FC5EBCA8FB91D0CD" box="[1019,1044,1946,1964]" italics="true" pageId="2" pageNumber="130">pst</emphasis>
|
||||
— pterostigma;
|
||||
<emphasis id="87E33316B11D5C65FB02BCA8FB3CD0CD" box="[1191,1209,1946,1964]" italics="true" pageId="2" pageNumber="130">rc</emphasis>
|
||||
— radial cell;
|
||||
<emphasis id="87E33316B11D5C65FA98BCABFAD4D0CD" box="[1341,1361,1945,1964]" italics="true" pageId="2" pageNumber="130">x1</emphasis>
|
||||
,
|
||||
<emphasis id="87E33316B11D5C65FAF9BCABFAF4D0CD" box="[1372,1393,1945,1964]" italics="true" pageId="2" pageNumber="130">x2</emphasis>
|
||||
,
|
||||
<emphasis id="87E33316B11D5C65FF19BC80FF40D0A5" box="[188,197,1970,1988]" italics="true" pageId="2" pageNumber="130">y</emphasis>
|
||||
— veins;
|
||||
<emphasis id="87E33316B11D5C65FE87BC80FEC6D0A5" box="[290,323,1970,1988]" italics="true" pageId="2" pageNumber="130">z —</emphasis>
|
||||
posterobasal margin of folded wing (jugal fold). Scale bar: 1.0 mm.
|
||||
</paragraph>
|
||||
<paragraph id="B528EF04B11D5C65FF47BCFFFAE0DF71" blockId="2.[188,1398,1921,2064]" pageId="2" pageNumber="130">
|
||||
Рис. 2.
|
||||
<taxonomicName id="72979487B11D5C65FE86BCFFFE5DD081" authorityName="Ponomarenko & Fedorenko & Bashkuev" authorityYear="2022" box="[291,472,1997,2016]" class="Insecta" family="Coptoclavidae" genus="Holcoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="130" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
|
||||
<emphasis id="87E33316B11D5C65FE86BCFFFE5DD081" box="[291,472,1997,2016]" italics="true" pageId="2" pageNumber="130">Holcoptera rasnitsyni</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="87E33316B11D5C65FE41BCFFFD88D081" bold="true" box="[484,525,1997,2016]" pageId="2" pageNumber="130">
|
||||
<taxonomicNameLabel id="9CD08E6DB11D5C65FE41BCFFFD88D081" box="[484,525,1997,2016]" pageId="2" pageNumber="130" rank="species">sp.n.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
: голотип ПИН 3320/204a, прорисовка крыльев. ОбоЗначениЯ: 1r — 1-Я радиальнаЯ Ячейка; AAP — жилка AA
|
||||
<subScript id="2913ED41B11D5C65FEB2BCC0FEADD09F" attach="left" box="[279,296,2034,2046]" fontSize="5" pageId="2" pageNumber="130">3a”</subScript>
|
||||
+(AA
|
||||
<subScript id="2913ED41B11D5C65FEFFBCC3FEE2D09C" attach="left" box="[346,359,2033,2045]" fontSize="5" pageId="2" pageNumber="130">3b</subScript>
|
||||
+(AA
|
||||
<subScript id="2913ED41B11D5C65FE3DBCC0FE1BD09F" attach="left" box="[408,414,2034,2046]" fontSize="5" pageId="2" pageNumber="130">4</subScript>
|
||||
+AP
|
||||
<subScript id="2913ED41B11D5C65FE60BCC0FE5DD09F" attach="left" box="[453,472,2034,2046]" fontSize="5" pageId="2" pageNumber="130">1+2</subScript>
|
||||
));
|
||||
<emphasis id="87E33316B11D5C65FE54BCD7FD80D099" box="[497,517,2021,2040]" italics="true" pageId="2" pageNumber="130">cb</emphasis>
|
||||
— костальнаЯ ось;
|
||||
<emphasis id="87E33316B11D5C65FD16BCD7FD4AD099" box="[691,719,2021,2040]" italics="true" pageId="2" pageNumber="130">cbz</emphasis>
|
||||
— Зона костального иЗгиба;
|
||||
<emphasis id="87E33316B11D5C65FC74BCD4FC59D099" box="[977,988,2022,2040]" italics="true" pageId="2" pageNumber="130">o</emphasis>
|
||||
— продолговатаЯ Ячейка;
|
||||
<emphasis id="87E33316B11D5C65FB6CBCD4FB67D099" box="[1225,1250,2022,2040]" italics="true" pageId="2" pageNumber="130">pst</emphasis>
|
||||
— птеростигма;
|
||||
<emphasis id="87E33316B11D5C65FF19BCCCFF4BDF71" box="[188,206,2046,2064]" italics="true" pageId="2" pageNumber="130">rc</emphasis>
|
||||
— радиальнаЯ Ячейка;
|
||||
<emphasis id="87E33316B11D5C65FE03BCCFFE3FDF71" box="[422,442,2045,2064]" italics="true" pageId="2" pageNumber="130">x1</emphasis>
|
||||
,
|
||||
<emphasis id="87E33316B11D5C65FE60BCCFFE5FDF71" box="[453,474,2045,2064]" italics="true" pageId="2" pageNumber="130">x2</emphasis>
|
||||
,
|
||||
<emphasis id="87E33316B11D5C65FE40BCCCFE6BDF71" box="[485,494,2046,2064]" italics="true" pageId="2" pageNumber="130">y</emphasis>
|
||||
— жилки;
|
||||
<emphasis id="87E33316B11D5C65FDF2BCCCFDFCDF71" box="[599,633,2046,2064]" italics="true" pageId="2" pageNumber="130">z —</emphasis>
|
||||
постеробаЗальный край сложенного крыла (югальнаЯ складка). МасШтаб: 1,0 мм.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="E1E8BF8CB11C5C64FF47B8EAFAB3D356" ID-DOI="http://doi.org/10.5281/zenodo.13179710" ID-Zenodo-Dep="13179710" httpUri="https://zenodo.org/record/13179710/files/figure.png" pageId="3" pageNumber="131" startId="3.[226,263,984,1003]" targetBox="[189,1399,314,962]" targetPageId="3" targetType="figure">
|
||||
<paragraph id="B528EF04B11C5C64FF47B8EAFCF4D365" blockId="3.[188,1399,984,1079]" pageId="3" pageNumber="131">
|
||||
Figs 3–5.
|
||||
<taxonomicName id="72979487B11C5C64FE9CB8EAFE73D48A" authorityName="Ponomarenko & Fedorenko & Bashkuev" authorityYear="2022" box="[313,502,984,1003]" class="Insecta" family="Coptoclavidae" genus="Holcoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="131" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
|
||||
<emphasis id="87E33316B11C5C64FE9CB8EAFE73D48A" box="[313,502,984,1003]" italics="true" pageId="3" pageNumber="131">Holcoptera rasnitsyni</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="87E33316B11C5C64FDA7B8EAFDA8D48A" bold="true" box="[514,557,984,1003]" pageId="3" pageNumber="131">
|
||||
<taxonomicNameLabel id="9CD08E6DB11C5C64FDA7B8EAFDA8D48A" box="[514,557,984,1003]" pageId="3" pageNumber="131" rank="species">sp.n.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
: 3–4 — paratype PIN 3320/113a, photo and drawing (solid line — elytra, dashed line — wings, dotted line — metanotum); 5 — Paratype PIN 3320/206a. Scale bars: 2.0 mm.
|
||||
</paragraph>
|
||||
<paragraph id="B528EF04B11C5C64FF47BF3EFAB3D356" blockId="3.[188,1399,984,1079]" pageId="3" pageNumber="131">
|
||||
Рис. 3–5.
|
||||
<taxonomicName id="72979487B11C5C64FE98BF3EFE79D37F" authorityName="Ponomarenko & Fedorenko & Bashkuev" authorityYear="2022" box="[317,508,1036,1055]" class="Insecta" family="Coptoclavidae" genus="Holcoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="131" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
|
||||
<emphasis id="87E33316B11C5C64FE98BF3EFE79D37F" box="[317,508,1036,1055]" italics="true" pageId="3" pageNumber="131">Holcoptera rasnitsyni</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="87E33316B11C5C64FDACBF3EFDB1D37E" bold="true" box="[521,564,1036,1055]" pageId="3" pageNumber="131">
|
||||
<taxonomicNameLabel id="9CD08E6DB11C5C64FDACBF3EFDB1D37E" box="[521,564,1036,1055]" pageId="3" pageNumber="131" rank="species">sp.n.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
: 3–4 — паратип PIN 3320/113a, фотографиЯ и прорисовка (сплоШнаЯ линиЯ — контуры надкрылий, пунктирнаЯ линиЯ — крыльЯ, точечнаЯ линиЯ — Заднеспинка); 5 — паратип PIN 3320/206a. МасШтаб: 2,0 мм.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="B528EF04B11C5C64FF47BFCCFE9CD2F8" blockId="3.[189,776,1118,1754]" pageId="3" pageNumber="131">Pterostigma and almost all veins are recognizable in anterior part of unfolded wing apex frontal to longitudinal axial fold. Radial cell apparently partly incorporated in complex pterostigma. Caudal and basal borders of this cell invisible, except for distal portion of radial cell limited by crossvein (r2).</paragraph>
|
||||
<paragraph id="B528EF04B11C5C64FF47BEADFEF2D2AE" blockId="3.[189,776,1118,1754]" pageId="3" pageNumber="131">
|
||||
MEASUREMENTS. Elytron length 5.0–6.0 mm, elytron width
|
||||
<quantity id="726F42E1B11C5C64FF5EBE8BFEF6D2AE" box="[251,371,1465,1487]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="3" pageNumber="131" unit="mm" value="2.1" valueMax="2.5" valueMin="1.7">1.7–2.5 mm</quantity>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FD8DBC8FB11C5C64FF47BEE6FDD3D1DE" pageId="3" pageNumber="131" type="discussion">
|
||||
<paragraph id="B528EF04B11C5C64FF47BEE6FDD3D1DE" blockId="3.[189,776,1118,1754]" pageId="3" pageNumber="131">
|
||||
REMARKS. The described hind wing is extremely similar to another unfolded wing of a coptoclavid beetle [
|
||||
<bibRefCitation id="D10692F5B11C5C64FD41BEDDFED7D17F" author="Fedorenko D. N." pageId="3" pageNumber="131" pagination="605 - 609" refId="ref2552" refString="Fedorenko D. N. 2014. Wings of Coptoclavidae (Coleoptera, Adephaga) from the Keuper (Triassic) of Frankonia, Germany // Paleontological Journal. Vol. 48. No. 6. P. 605 - 609." type="journal article" year="2014">Fedorenko, 2014</bibRefCitation>
|
||||
]. It matches well the wing ground plan of
|
||||
<taxonomicName id="72979487B11C5C64FF18BD16FEA7D15B" authorityName="Schellenberg" authorityYear="1806" box="[189,290,1572,1594]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="129" phylum="Tracheophyta" rank="subOrder" subOrder="Adephaga">Adephaga</taxonomicName>
|
||||
, notably Hydradephaga, virtually in all details. Only two features are worthy of note. Firstly, the large and elliptic oblongum cell is peculiar to nearly all Recent Hydradephaga. Secondly, the cubital joint is proximal, which has been observed in the most primitive representatives (
|
||||
<taxonomicName id="72979487B11C5C64FF66BD9BFEADD1DE" box="[195,296,1705,1727]" class="Insecta" family="Gyrinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="131" phylum="Arthropoda" rank="family">Gyrinidae</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="72979487B11C5C64FE96BD9BFE1CD1DE" box="[307,409,1705,1727]" class="Insecta" family="Haliplidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="131" phylum="Arthropoda" rank="family">Haliplidae</taxonomicName>
|
||||
) of Hydradephaga.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FD8DBC8FB11C5C64FF47BDF6FAA1D3EF" pageId="3" pageNumber="131" type="etymology">
|
||||
<paragraph id="B528EF04B11C5C64FF47BDF6FC8DD1BB" blockId="3.[189,776,1118,1754]" box="[226,776,1732,1754]" pageId="3" pageNumber="131">ETYMOLOGY. Named in honor of Alexandr Rasnitsyn.</paragraph>
|
||||
<paragraph id="B528EF04B11C5C64FF47BDCBFAA1D3EF" blockId="3.[189,774,1785,1860]" lastBlockId="3.[812,1399,1118,1167]" pageId="3" pageNumber="131">
|
||||
<emphasis id="87E33316B11C5C64FF47BDCBFE1AD06E" bold="true" box="[226,415,1785,1807]" pageId="3" pageNumber="131">Acknowledgments</emphasis>
|
||||
. We are grateful to two anonymous reviewers for their helpful comments. Many thanks to Anna Ponomarenko and Roman Rakitov for help in preparing the manuscript. The research was partly supported by the Russian Science Foundation, project No. 21-14-00284.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="FD8DBC8FB11C5C64FCF7BF9EFC00D3BD" pageId="3" pageNumber="131" type="description">
|
||||
<paragraph id="B528EF04B11C5C64FCF7BF9EFC00D3BD" blockId="3.[812,1398,1196,1244]" pageId="3" pageNumber="131">
|
||||
<emphasis id="87E33316B11C5C64FCF7BF9EFBA4D3A3" bold="true" box="[850,1057,1196,1218]" pageId="3" pageNumber="131">Competing interests</emphasis>
|
||||
. The authors declare no competing interests.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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