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<mods:title id="830D662EB1905FB28DA619D279E9813D">The description of the first rock-dwelling species of butterfly lizard Leiolepis Cuvier, 1829 (Squamata, Agamidae) from the Khorat Plateau in northeastern Thailand</mods:title>
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<mods:namePart id="A75B1EA6CAE92B729C6DEEB1768261D6">Wanchai, Pratyaporn</mods:namePart>
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<mods:affiliation id="313B05BB97DF651800F8177E54C7D797">Department of Biological Science, Faculty of Science, Ubon Ratchathani University, Warin Chamrap District, Ubon Ratchathani 34190, Thailand</mods:affiliation>
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<mods:namePart id="3A3DB27EC95BC15970919C8A97E68C63">Rujirawan, Attapol</mods:namePart>
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<mods:namePart id="715A718F2BBC48F0C94064310C05B1B8">Murdoch, Matthew L.</mods:namePart>
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<mods:namePart id="B8115F32D14C5CF9ECB570827A074337">Aksornneam, Akrachai</mods:namePart>
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<mods:namePart id="FD7E8D3CDF232D62CE3A2C602FD2ABDD">Promnun, Pattarapon</mods:namePart>
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<mods:affiliation id="6DFFD7F24EB30D7768131A8FD905CDC2">Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92505, USA</mods:affiliation>
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<mods:namePart id="D66CAAC81946BCADA7EA3BC72D5D66C8">Kaatz, Amanda</mods:namePart>
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<mods:namePart id="F38AD761384F4E061B2C4762AFF5BCC7">Gregory, Jeren J.</mods:namePart>
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<mods:namePart id="AFF7C053D032A49A481FD3BFE4B68822">Nguyen, Eddie</mods:namePart>
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<mods:affiliation id="87302D99478F0B438D7060B221573000">Biodiversity Center, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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<mods:namePart id="AD768BFB3CABC6510892749EFF4F8284">Iderstein, William Van</mods:namePart>
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<mods:affiliation id="A2BC7AB5E56A57CA6E33517B6BEA665A">Biodiversity Center, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
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<mods:namePart id="AE7EA571CA93A98DFC5EA226F6564DAE">Quah, Evan S. H.</mods:namePart>
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<mods:affiliation id="4CE78C5697FFF1C32184CC2D2F27D8C4">Geneus Genetics Co., Ltd., 37 / 1 Sukhumvit 101 / 1, Bang Chak, Phra Kanong District, Bangkok 10260, Thailand & Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400, Kota Kinabalu, Sabah, Malaysia & Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Singapore</mods:affiliation>
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<mods:namePart id="6045E5F4C2F91959851BD0230FABF7D8">Grismer, L. Lee</mods:namePart>
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<mods:affiliation id="B4882FC8EC3B17D853A456B599A1C70C">Biodiversity Center, Kasetsart University, Bangkok 10900, Thailand & Geneus Genetics Co., Ltd., 37 / 1 Sukhumvit 101 / 1, Bang Chak, Phra Kanong District, Bangkok 10260, Thailand & School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia</mods:affiliation>
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<mods:namePart id="8A305B8498B9B99F9B5C8B74EBBDE1E8">Grismer, Jesse L.</mods:namePart>
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<mods:namePart id="7AE39441F3D3F87AB03727EB03088183">Aowphol, Anchalee</mods:namePart>
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<mods:title id="E47C0F9B155FC6BFEF5FFD7E2EE325DE">ZooKeys</mods:title>
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<mods:part id="4FEA55D6A89353190A6F059EEC382F8B">
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<mods:date id="3BC5AC448AB121C12D687A69EF87B999">2024</mods:date>
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<mods:number id="72A6F5C2EFCB357F3B200F4A7F8FF533">2024-08-27</mods:number>
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<mods:number id="9C078C710C8B7A61A06CFBCD28BEAAA4">1210</mods:number>
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<mods:identifier id="11AEAA885F027085C84713BADD6C271A" type="DOI">10.3897/zookeys.1210.127557</mods:identifier>
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<treatment id="0E624381AF835F10996C1D54D8DA7038" ID-arpha="0E624381-AF83-5F10-996C-1D54D8DA7038" ID-zoobank="https://zoobank.org/22E2BD02-9672-434B-B4DF-43AD636677B7" LSID="urn:lsid:zoobank.org:act:22E2BD02-9672-434B-B4DF-43AD636677B7" LSID-ZBK="urn:lsid:zoobank.org:act:22E2BD02-9672-434B-B4DF-43AD636677B7" httpUri="http://treatment.plazi.org/id/0E624381AF835F10996C1D54D8DA7038">
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<subSubSection id="CCC0C2DBA22CCE376E66AD2531CD8673" type="nomenclature">
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<paragraph id="7F0DF7BD49B4EF5B9877C5F86A10E931">
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<taxonomicName id="5363AAFE31EF7676458EA765A8DF268A" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
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<emphasis id="E230D2C590A5792737CF0470A547D5B2" italics="true">Leiolepis glaurung</emphasis>
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</taxonomicName>
|
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<taxonomicNameLabel id="79EB929D8000C8EACFFC82266AB4D0C2" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="1E66FB8BDA58AC809A34204CF23C6DEA" type="description">
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<paragraph id="873B58E61CB0D3266184E4765802C9AA">
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<figureCitation id="E4E33D3E416B3F6147DE0CDD238C9032" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal and ventral views of the holotype and paratypes of Leiolepis glaurung sp. nov. A male holotype THNHM 30909 B female paratype THNHM 30908 C male paratype THNHM 30910, and D male paratype THNHM 30911." figureDoi="10.3897/zookeys.1210.127557.figure5" httpUri="https://binary.pensoft.net/fig/1122035">Figs 5</figureCitation>
|
||||
,
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||||
<figureCitation id="A65F94A0ABBF55B7E072DD2CF43DD328" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsal and ventral views of paratypes THNHM 30906, THNHM 30907, THNHM 30904, THNHM 30902, THNHM 30905, and THNHM 30903 Leiolepis glaurung sp. nov. Specimen tags bear field numbers." figureDoi="10.3897/zookeys.1210.127557.figure6" httpUri="https://binary.pensoft.net/fig/1122036">6</figureCitation>
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||||
,
|
||||
<figureCitation id="D610F29D6E3A51B05B9F13262361276F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Female paratype THNHM 30908 B male paratype THNHM 30911 of Leiolepis glaurung sp. nov." figureDoi="10.3897/zookeys.1210.127557.figure9" httpUri="https://binary.pensoft.net/fig/1122039">9</figureCitation>
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,
|
||||
<figureCitation id="EB4DA0C120B5AA60E2E5855F780C9624" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Male paratype THNHM 30911 displaying the dorsal and lateral display colors and its rocky habitat in the background B adult male Leiolepis rubritaeniata from southeastern Cambodia C male paratype THNHM 30911 of Leiolepis glaurung sp. nov." figureDoi="10.3897/zookeys.1210.127557.figure11" httpUri="https://binary.pensoft.net/fig/1122041">11</figureCitation>
|
||||
;
|
||||
<tableCitation id="4D5E826D6621165769628B8443FD6021" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Results from the PERMANOVA analysis comparing pairs of sexual species of Leiolepis bearing statistical mean morphospatial differences at the p. adjusted threshold based on the load scores of Dim 1 – Dim 5 of the MFA. * = significant difference." httpUri="http://table.plazi.org/id/CCA4A8C41C36082CA6FCC650E583B249" tableUuid="CCA4A8C41C36082CA6FCC650E583B249">Table 1</tableCitation>
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<xref id="06ED25C1F0DD87D78692737A16962728" ref-type="supplementary-material" rid="S1">; Suppl. materials 1</xref>
|
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<xref id="435A10F587C55FC1A2EC668110519703" ref-type="supplementary-material" rid="S2">
|
||||
, 2 Suggested English name:
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||||
<collectingRegion id="FAA75707ED0C8CC5BB947BA9321D5CCF" country="Thailand" name="Nakhon Ratchasima">Khorat</collectingRegion>
|
||||
Plateau Butterfly Lizard
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||||
</xref>
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||||
</paragraph>
|
||||
</subSubSection>
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<subSubSection id="SECID0E1YAG" type="type material">
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<heading id="2F3C6AC6AB48E5347BE715520FE216D0" reason="title">Type material.</heading>
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<paragraph id="F89E37E6D3CC0D657F9527D4E6690117">
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<materialsCitation id="3157EC3A3B54DCF1A185D25FF27276E1" collectingDate="2023-03-20" collectionCode="THNHM" collectorName="Pratyaporn Wanchai & Anchalee Aowphol & Attapol Rujirawan & Akrachai Aksornneam & Jesse L. Grismer, L. & Lee Grismer & Evan S. H. Quah & Matthew L. Murdoch" country="Thailand" county="Kut Khaopun District" latitude="15.802973" location="Wat Phu Noi" longLatPrecision="1" longitude="105.15692" municipality="Kaeng Kheng Subdistrict" specimenCode="THNHM 30909" specimenCount="1" specimenCount-male="1" stateProvince="Ubon Ratchathani Province" typeStatus="holotype">
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<emphasis id="18C8AF4732FEEC48E9C2A1B9619DDC24" bold="true">
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<emphasis id="7C8A68CF6A9F37AA24F825641FB861D6" bold="true" italics="true">
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<typeStatus id="6D7C4D6A0D824173B9EACBED5159CCDF" type="holotype">Holotype</typeStatus>
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||||
</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<specimenCount id="4B988CB8088616A4033A4220A23C76F9" count="1" type="male">Adult male</specimenCount>
|
||||
(
|
||||
<specimenCode id="CE22D2EE4210AF2673193A6FC0D277BC">THNHM 30909</specimenCode>
|
||||
; Fig.
|
||||
<figureCitation id="258565F7B8DC8297C95A8917058B6058" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal and ventral views of the holotype and paratypes of Leiolepis glaurung sp. nov. A male holotype THNHM 30909 B female paratype THNHM 30908 C male paratype THNHM 30910, and D male paratype THNHM 30911." figureDoi="10.3897/zookeys.1210.127557.figure5" httpUri="https://binary.pensoft.net/fig/1122035">5 A</figureCitation>
|
||||
) collected from just northeast of
|
||||
<location id="DE214BD0C86596A31A60C0454FA0C953" LSID="urn:lsid:plazi:treatment:0E624381AF835F10996C1D54D8DA7038:DE214BD0C86596A31A60C0454FA0C953" country="Thailand" county="Kut Khaopun District" latitude="15.802973" longLatPrecision="1" longitude="105.15692" municipality="Kaeng Kheng Subdistrict" name="Wat Phu Noi" stateProvince="Ubon Ratchathani Province">Wat Phu Noi</location>
|
||||
in
|
||||
<collectingMunicipality id="46266CC4691DCE7E5A48C52A9A733155">Kaeng Kheng Subdistrict</collectingMunicipality>
|
||||
,
|
||||
<collectingCounty id="DB3C2F1D15E2C4B00183ECBA0F5496E3">Kut Khaopun District</collectingCounty>
|
||||
,
|
||||
<collectingRegion id="62DCEAA3654B7EBFB3BC7D6DD90E8683" country="Thailand" name="Ubon Ratchathani">Ubon Ratchathani Province</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="024A1797D2634066B585A84241AEF549" name="Thailand">Thailand</collectingCountry>
|
||||
(
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMZAG" specific-use="{"type":"Point","coordinates":[105.156917,15.802972]}">
|
||||
<geoCoordinate id="5311E1F544F156967EA4BBF6CA295223" degrees="15" direction="north" minutes="48" orientation="latitude" precision="1" seconds="10.7" value="15.802973">15 ° 48 ' 10.7 " N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="B20A3D53A67F197545A4C325C9B860D0" degrees="105" direction="east" minutes="09" orientation="longitude" precision="1" seconds="24.9" value="105.15692">105 ° 09 ' 24.9 " E</geoCoordinate>
|
||||
</named-content>
|
||||
) on
|
||||
<date id="4AE52F891CDB854A4FD43D222FCB441A" value="2023-03-20">
|
||||
<collectingDate id="8E3E660A2D2E25EC2BAD6B0BEBC79AAA" value="2023-03-20">20 March 2023</collectingDate>
|
||||
</date>
|
||||
, at 10: 00 am by
|
||||
<collectorName id="3362F9B1FE46AED758DE622CBFA70881">Pratyaporn Wanchai</collectorName>
|
||||
,
|
||||
<collectorName id="308979794B5475DA191AA66A0A78B187">Anchalee Aowphol</collectorName>
|
||||
,
|
||||
<collectorName id="EACB728B6FCFEDBE57EBF6D5911C71B2">Attapol Rujirawan</collectorName>
|
||||
,
|
||||
<collectorName id="B37FD85D550DB5BFC8FDA606207DE1BA">Akrachai Aksornneam</collectorName>
|
||||
,
|
||||
<collectorName id="8768210CFE55316A9D13F88DF2B3ECFD">Jesse L. Grismer, L.</collectorName>
|
||||
<collectorName id="42734796396ED87FA8E136A96A9A9797">Lee Grismer</collectorName>
|
||||
,
|
||||
<collectorName id="20491133D0B97145FA8041C959A906E0">Evan S. H. Quah</collectorName>
|
||||
, and
|
||||
<collectorName id="8EA3EDE51D35E2CB1858DE457DDBEBA6">Matthew L. Murdoch</collectorName>
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="0FD2AACA9D0A0C16E0055462AA509195" collectingDate="2023-03-20" collectionCode="THNHM" collectorName="Pratyaporn Wanchai & Anchalee Aowphol & Attapol Rujirawan & Akrachai Aksornneam & Jesse L. Grismer, L. & Lee Grismer & Evan S. H. Quah & Matthew L. Murdoch" country="Thailand" county="Kut Khaopun District" latitude="15.802973" location="Wat Phu Noi" longLatPrecision="1" longitude="105.15692" municipality="Kaeng Kheng Subdistrict" specimenCode="THNHM 30908, THNHM 30910" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Ubon Ratchathani Province" typeStatus="paratype">
|
||||
<emphasis id="9007E5206B92013BFB15C223230BA072" bold="true">
|
||||
<emphasis id="BB952125E707A96479EFB29AA0395FB0" bold="true" italics="true">
|
||||
<typeStatus id="483CF7808D4E857FCEB8519204B1371C" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<specimenCount id="6C624D0C5C0207DB6942058D4DD2ED15" count="1" type="female">Adult female</specimenCount>
|
||||
(
|
||||
<specimenCode id="47FA42A13ACA0E4E8F7C2DCF6C81F78E">THNHM 30908</specimenCode>
|
||||
; Fig.
|
||||
<figureCitation id="C2EC6D26662B9D8C60F7881045BDED4E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal and ventral views of the holotype and paratypes of Leiolepis glaurung sp. nov. A male holotype THNHM 30909 B female paratype THNHM 30908 C male paratype THNHM 30910, and D male paratype THNHM 30911." figureDoi="10.3897/zookeys.1210.127557.figure5" httpUri="https://binary.pensoft.net/fig/1122035">5 B</figureCitation>
|
||||
) and
|
||||
<specimenCount id="6E632F6ED34BB70774E93CEF27789843" count="2" type="male">two adult males</specimenCount>
|
||||
(
|
||||
<specimenCode id="BC51648AF1AC48819C70CCAE4E2B8320">THNHM 30910</specimenCode>
|
||||
–30911; Fig.
|
||||
<figureCitation id="BC0FC2B6DF6E3750A4532E093DF3B09A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal and ventral views of the holotype and paratypes of Leiolepis glaurung sp. nov. A male holotype THNHM 30909 B female paratype THNHM 30908 C male paratype THNHM 30910, and D male paratype THNHM 30911." figureDoi="10.3897/zookeys.1210.127557.figure5" httpUri="https://binary.pensoft.net/fig/1122035">5 C, D</figureCitation>
|
||||
) bear the same locality and collectors as the holotype
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="E8789B22895E20187BE82DAB7BAB5E65" collectingDate="2020-07-04" collectingDateMax="2020-07-10" collectingDateMin="2020-07-04" collectionCode="THNHM" collectorName="Pratyaporn Wanchai" country="Thailand" county="Kut Khaopun District" latitude="15.802973" location="Wat Phu Noi" longLatPrecision="1" longitude="105.15692" municipality="Kaeng Kheng Subdistrict" specimenCode="THNHM 30902, THNHM 30907, THNHM 30904" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Ubon Ratchathani Province">
|
||||
<specimenCount id="F51CF8251CE5E8710312ED3352B3FAF3" count="3" type="male">Three adult males</specimenCount>
|
||||
(
|
||||
<specimenCode id="BA9DB5B8A7F96C3BDF0AD04C63CB8DBD">THNHM 30902</specimenCode>
|
||||
–30903,
|
||||
<specimenCode id="B3E371188678C13D2F48BFD37A41EDC5">THNHM 30907</specimenCode>
|
||||
) and
|
||||
<specimenCount id="C3381D55543A2AFEAB6EFB352E585883" count="3" type="female">three adult females</specimenCount>
|
||||
(
|
||||
<specimenCode id="8F54CBF8E9D83BEBF93583F7D6A60B98">THNHM 30904</specimenCode>
|
||||
–30906; Fig.
|
||||
<figureCitation id="88BACBFEA2361834A260AE58DC22C421" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsal and ventral views of paratypes THNHM 30906, THNHM 30907, THNHM 30904, THNHM 30902, THNHM 30905, and THNHM 30903 Leiolepis glaurung sp. nov. Specimen tags bear field numbers." figureDoi="10.3897/zookeys.1210.127557.figure6" httpUri="https://binary.pensoft.net/fig/1122036">6</figureCitation>
|
||||
) were collected by
|
||||
<collectorName id="DC072B2B52D77758060203E9DEAA564D">Pratyaporn Wanchai</collectorName>
|
||||
between
|
||||
<date id="84695DCCF3A2D843D6FE6B81D574F7F4" value="2020-07-04" valueMax="2020-07-10" valueMin="2020-07-04">
|
||||
<collectingDate id="941ED2021340286B475FDD664B0B013D" value="2020-07-04" valueMax="2020-07-10" valueMin="2020-07-04">4–10 July 2020</collectingDate>
|
||||
</date>
|
||||
from the same locality as the holotype
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="4A8A5C65AEAF53A3C90314F9403575CB" ID-DOI="10.3897/zookeys.1210.127557.figure5" ID-arpha="ADC6A32D-986D-55FC-8B60-01326BD22BBD" httpUri="https://binary.pensoft.net/fig/1122035" startId="F5">
|
||||
<paragraph id="A78BE9946C50A446B2E6CD0762EA99C9">
|
||||
<label id="65C3172DF1C484D1DD391796ADB1D410">Figure 5.</label>
|
||||
</paragraph>
|
||||
<paragraph id="A04A878F7EB55B7AEB94C42AA24E8FE2">
|
||||
Dorsal and ventral views of the holotype and paratypes of
|
||||
<taxonomicName id="B110B6D4DBC5BAA22A66EF070BA7FDFE" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="05AC1C18A606984935DF01A265CCB987" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="7FC55502ED21C078F04F48A1A15A1AD9" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="7DBD880FC22497C1D8FA0A3172D503AF" bold="true">A</emphasis>
|
||||
male holotype THNHM 30909
|
||||
<emphasis id="D14967EF29DAE7E7A30520BFB87FCD64" bold="true">B</emphasis>
|
||||
female paratype THNHM 30908
|
||||
<emphasis id="9C978986601A28C01B02595A9FA0CE5A" bold="true">C</emphasis>
|
||||
male paratype THNHM 30910, and
|
||||
<emphasis id="5A4E5458B0D983EE67D0AA48A08217AC" bold="true">D</emphasis>
|
||||
male paratype THNHM 30911.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="10095DFBED1EEFDBD45FCF59615FEFC7" ID-DOI="10.3897/zookeys.1210.127557.figure6" ID-arpha="5A8AFC16-A5ED-5598-B45D-5BC5B08719D2" httpUri="https://binary.pensoft.net/fig/1122036" startId="F6">
|
||||
<paragraph id="E29583C36CDBE8760F2A56C5A05F4031">
|
||||
<label id="3FDCE41480C9D1FEC3EA6BAF7DE0E62D">Figure 6.</label>
|
||||
</paragraph>
|
||||
<paragraph id="00165E54F84DAB8BE7A9F40410923053">
|
||||
Dorsal and ventral views of paratypes THNHM 30906, THNHM 30907, THNHM 30904, THNHM 30902, THNHM 30905, and THNHM 30903
|
||||
<taxonomicName id="E925710D3EA6BCC5C6181F9AA80D5792" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="D384E65FF3EA2963240A17B312E14921" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="BB539AEE6F5D6073BE23919BE71BA9D4" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
Specimen tags bear field numbers.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EZ2AG" type="diagnosis">
|
||||
<paragraph id="01631DCB1EF93A00F6E950A6A5B69CBC">
|
||||
<heading id="2502E7B30181467D3D834E146692A594" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="A672C5FFFE4E03524372A9E991B42908">
|
||||
<taxonomicName id="8BE563364FB796BA5EB22D912D9CCC18" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="22240C2CB6BA890E3046142DFFE5B716" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="878C851A1BAC8584DA06E6A2D1E049DC" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
can be diagnosed from all sexual species of
|
||||
<taxonomicName id="B78E4F644D8645CD1695539570BF0FA4" authorityName="Cuvier" authorityYear="1829" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="genus">
|
||||
<emphasis id="1D3AC6449B707884A18C0A9DBCC26E32" italics="true">Leiolepis</emphasis>
|
||||
</taxonomicName>
|
||||
by having a black gular region with a wide medial yellow stripe, a yellow ventrum with black mottling, bright red to orange ventral tail coloration, having reduced to no expandable flanks, and having a maximum of one black transverse bar on the flank (Suppl. material
|
||||
<xref id="4EC443864B1553593BEC058862A28FEF" ref-type="supplementary-material" rid="S1">1</xref>
|
||||
).
|
||||
<taxonomicName id="AA309C4700D5D3273142E77807E3EF75" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="5021E64713790F07100EF193ACB77000" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A93478B7B380BBEE2C8FB904F7DEE144" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
can be further diagnosed from all other sexual species by having the combination of a
|
||||
<abbrev id="ABBRID0EA4AG" xlink_title="axilla–groin length">AG</abbrev>
|
||||
of 61.0–88.0 mm;
|
||||
<abbrev id="ABBRID0EE4AG" xlink_title="brachium length">BL</abbrev>
|
||||
of 11.0–
|
||||
<quantity id="353B7FAEBE81C0EB9C865EA72F1DBB91" metricMagnitude="-2" metricUnit="m" metricValue="1.68" unit="mm" value="16.8">16.8 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0EI4AG" xlink_title="chest width">CW</abbrev>
|
||||
of
|
||||
<quantity id="212C8E32356F9DB9E82F9001133FF2AE" metricMagnitude="-2" metricUnit="m" metricValue="2.77" metricValueMax="3.31" metricValueMin="2.23" unit="mm" value="27.7" valueMax="33.1" valueMin="22.3">22.3–33.1 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0EM4AG" xlink_title="distance between nares">DN</abbrev>
|
||||
of
|
||||
<quantity id="579994CB4FDE19E4E873AA3E91369FA2" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.7" metricValueMin="4.3" unit="mm" value="5.5" valueMax="6.7" valueMin="4.3">4.3–6.7 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0EQ4AG" xlink_title="forearm length">FA</abbrev>
|
||||
of 14.4–19.0 mm;
|
||||
<abbrev id="ABBRID0EU4AG" xlink_title="height of ear">HE</abbrev>
|
||||
of
|
||||
<quantity id="D5A7AF437BCCB60E66FAF14FEF9DAEB8" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="7.1" metricValueMin="5.6" unit="mm" value="6.35" valueMax="7.1" valueMin="5.6">5.6–7.1 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0EY4AG" xlink_title="head height">HH</abbrev>
|
||||
of 14.0–
|
||||
<quantity id="B8393FC727AF39B7E0DA47D3BE247E6B" metricMagnitude="-2" metricUnit="m" metricValue="2.13" unit="mm" value="21.3">21.3 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0E34AG" xlink_title="head length">HL</abbrev>
|
||||
of
|
||||
<quantity id="A4BC2D8125C1B8270FDF68E052EB3552" metricMagnitude="-2" metricUnit="m" metricValue="3.535" metricValueMax="4.1899999999999995" metricValueMin="2.88" unit="mm" value="35.35" valueMax="41.9" valueMin="28.8">28.8–41.9 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0EA5AG" xlink_title="head width">HW</abbrev>
|
||||
of
|
||||
<quantity id="520806EFAB2B4BFBA351E1D57C386994" metricMagnitude="-2" metricUnit="m" metricValue="2.445" metricValueMax="2.9699999999999998" metricValueMin="1.92" unit="mm" value="24.45" valueMax="29.7" valueMin="19.2">19.2–29.7 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0EE5AG" xlink_title="pelvis width">PW</abbrev>
|
||||
of 10.5–17.0 mm;
|
||||
<abbrev id="ABBRID0EI5AG" xlink_title="tibia length">TIB</abbrev>
|
||||
of 24.34–29.0 mm;
|
||||
<abbrev id="ABBRID0EM5AG" xlink_title="third toe length">TE</abbrev>
|
||||
of 14.0–18.0 mm,
|
||||
<abbrev id="ABBRID0EQ5AG" xlink_title="rostrum height">RH</abbrev>
|
||||
of
|
||||
<quantity id="455B729ED02153795B81350C14F1DEDA" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.43" metricValueMin="0.97" unit="mm" value="12.0" valueMax="14.3" valueMin="9.7">9.7–14.3 mm</quantity>
|
||||
; three dorsal stripes; 28–29 ventral scales; 21–24 dorsal caudal scales; 19–26 non-pore bearing scales between the pore-bearing femoral scales across the pelvis; maximum number of seven keeled scale rows across the forearm; 17–20 femoral pores per side; 6–8 scales across the frontal bone; maximum of nine subtibial scales; seven scales long the dorsal surface of the first toe; three enlarged subdigital lamellae on the third toe; and 32–34 scales along the dorsal surface of the fourth toe (Figs
|
||||
<figureCitation id="F9C2ACBAAE428F4F8AD98F871E06A114" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Comparisons of the normalized morphometric characters showing the ranges, frequencies, mean (white dots), and 50 % quartiles represented by black bars." figureDoi="10.3897/zookeys.1210.127557.figure7" httpUri="https://binary.pensoft.net/fig/1122037">7</figureCitation>
|
||||
,
|
||||
<figureCitation id="F78AAD6D7E398AF8BC7131E1792FF635" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Comparisons of statistically different meristic characters among well-supported clades of Leiolepis. White circles are means and the black horizontal bars are medians." figureDoi="10.3897/zookeys.1210.127557.figure8" httpUri="https://binary.pensoft.net/fig/1122038">8</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="5DD2E932F10107DAD0D4638B2C19DDE3" ID-DOI="10.3897/zookeys.1210.127557.figure7" ID-arpha="6F0E67A9-167E-5A21-ABF9-183DF23AA3AB" httpUri="https://binary.pensoft.net/fig/1122037" startId="F7">
|
||||
<paragraph id="624677033732EEC8B9AE21A3B0915A8A">
|
||||
<label id="8233E73E88F1A8CF8D65A61466A864D6">Figure 7.</label>
|
||||
</paragraph>
|
||||
<paragraph id="6291941E9A7285D05A1FE80AC98EA5BF">Comparisons of the normalized morphometric characters showing the ranges, frequencies, mean (white dots), and 50 % quartiles represented by black bars.</paragraph>
|
||||
</caption>
|
||||
<caption id="41464DAD30D03F945DA0BDE475E83B1A" ID-DOI="10.3897/zookeys.1210.127557.figure8" ID-arpha="D4F9FF63-16E9-564A-9FA3-1EAA292FD6A2" httpUri="https://binary.pensoft.net/fig/1122038" startId="F8">
|
||||
<paragraph id="B92F3B3C363C4A5EC78E460126A78CEA">
|
||||
<label id="29FB6C9D6DD66567E1E147E4D90E7137">Figure 8.</label>
|
||||
</paragraph>
|
||||
<paragraph id="54D2AEA0E9C1EBAB70142F3BEFE49E3D">
|
||||
Comparisons of statistically different meristic characters among well-supported clades of
|
||||
<taxonomicName id="5452E10F7BBA337B98C244E21A8D961A" authorityName="Cuvier" authorityYear="1829" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="genus">
|
||||
<emphasis id="F47D2007CD25D246CB6C804313B144CD" italics="true">Leiolepis</emphasis>
|
||||
</taxonomicName>
|
||||
. White circles are means and the black horizontal bars are medians.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E36AG" type="description">
|
||||
<paragraph id="5AA3AB81A67A7B91028D33F7DCE98E8A">
|
||||
<heading id="385749C7DD53FF79D37FD8CD066C64C6" reason="title">
|
||||
Description of
|
||||
<typeStatus id="F8EB3747E0E697D8BADC69357DCEEA12" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="6CA15F1789E4518464060FFA04C2231F">
|
||||
Head large, (
|
||||
<abbrev id="ABBRID0ECABG" xlink_title="head length">HL</abbrev>
|
||||
<quantity id="4C08924F22CB4D2AD1F375E8DED3FB0F" metricMagnitude="-2" metricUnit="m" metricValue="4.1899999999999995" unit="mm" value="41.9">41.9 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0EGABG" xlink_title="head length">HL</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EKABG" xlink_title="Snout–vent length">SVL</abbrev>
|
||||
0.25) obtusely rounded in lateral profile, triangular in dorsal profile (
|
||||
<abbrev id="ABBRID0EOABG" xlink_title="head width">HW</abbrev>
|
||||
<quantity id="2B7B69832C17F7A283B11A29B3F6D317" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" unit="mm" value="29.7">29.7 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0ESABG" xlink_title="head width">HW</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EWABG" xlink_title="head length">HL</abbrev>
|
||||
0.71); interorbital, frontal region, and rostrum, convex (
|
||||
<abbrev id="ABBRID0E1ABG" xlink_title="head height">HH</abbrev>
|
||||
<quantity id="9BD859A888DA56A616B5DE98F7043617" metricMagnitude="-2" metricUnit="m" metricValue="2.13" unit="mm" value="21.3">21.3 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0E5ABG" xlink_title="rostrum height">RH</abbrev>
|
||||
<quantity id="DC914D5D2872A210110B756C59009B4B" metricMagnitude="-2" metricUnit="m" metricValue="1.43" unit="mm" value="14.3">14.3 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0ECBBG" xlink_title="rostrum height">RH</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EGBBG" xlink_title="head height">HH</abbrev>
|
||||
0.67), sloped anteriorly, covered with small, undifferentiated keeled scales; occipital and supraorbital regions covered with 19 keeled, granular scale rows half the size as dorsal head scales; canthus rostralis short, rounded; dorsal head scales strongly keeled, eight keeled scales across the frontal bone between supraorbital regions; rostral large, triangular (wider than long), bordered posteriorly by six smaller scales; external nares large, set wide apart (
|
||||
<abbrev id="ABBRID0EKBBG" xlink_title="distance between nares">DN</abbrev>
|
||||
<quantity id="815F4B31B425A70E71FF512B71F19BEF" metricMagnitude="-3" metricUnit="m" metricValue="6.7" unit="mm" value="6.7">6.7 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0EOBBG" xlink_title="distance between nares">DN</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0ESBBG" xlink_title="head width">HW</abbrev>
|
||||
0.23) rounded, directed laterally, set in single, oval, nasal scale surrounded by several small scales; elongate, keeled, large fused, suborbitals (on five right side; five on the left) extend from anterior margin of eye to posterior margin of eye; superciliary scales elongate, keeled, imbricate, continuous with canthal scales; eyelid scales granular; tympanum naked, deeply set, surrounded by granular scales; temporal scales keeled, small, slightly raised; nine rectangular supralabials whose contact with one another produces an distinct labial margin, bordered ventrally by small granular scales; mental longer than wide, pointed posteriorly, larger than adjacent infralabials; two large postmentals in contact medially, being first of a series of 14 enlarged scales along the angle of jaw (left side: right side was damaged); 10 rectangular infralabials; gular scales small, rounded, granular; two distinct anterior and posterior gular folds present; dewlap absent; antehumeral fold continuous with posterior gular fold (Figs
|
||||
<figureCitation id="55D01BA1EAE3C84388132A9BE092944F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal and ventral views of the holotype and paratypes of Leiolepis glaurung sp. nov. A male holotype THNHM 30909 B female paratype THNHM 30908 C male paratype THNHM 30910, and D male paratype THNHM 30911." figureDoi="10.3897/zookeys.1210.127557.figure5" httpUri="https://binary.pensoft.net/fig/1122035">5 A</figureCitation>
|
||||
,
|
||||
<figureCitation id="41EB5FDEFEE70BCA321143FDFDF3EB9B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsal and ventral views of paratypes THNHM 30906, THNHM 30907, THNHM 30904, THNHM 30902, THNHM 30905, and THNHM 30903 Leiolepis glaurung sp. nov. Specimen tags bear field numbers." figureDoi="10.3897/zookeys.1210.127557.figure6" httpUri="https://binary.pensoft.net/fig/1122036">6 A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="745766583B1612597AA02F161399186A">
|
||||
Body elongate (
|
||||
<abbrev id="ABBRID0EACBG" xlink_title="axilla–groin length">AG</abbrev>
|
||||
88.0 mm;
|
||||
<abbrev id="ABBRID0EECBG" xlink_title="axilla–groin length">AG</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EICBG" xlink_title="Snout–vent length">SVL</abbrev>
|
||||
0.52) somewhat dorsoventrally compressed; expandable flanks reduced to absent; body scales small, granular, slightly keeled; 43 scales between dorsolateral stripes; 20 scales between vertebral stripe and dorsolateral stripes at widest point of trunk; scales of flanks abruptly transition into much larger, flat scales of belly and pectoral region; 28 scales across middle of belly contacting apex of the umbilical scar; precloacals smooth and much smaller than ventral scales; forelimbs short, robust (
|
||||
<abbrev id="ABBRID0EMCBG" xlink_title="forearm length">FA</abbrev>
|
||||
19.0 mm;
|
||||
<abbrev id="ABBRID0EQCBG" xlink_title="brachium length">BL</abbrev>
|
||||
<quantity id="CDA0A4C33780C800F36926158EF01A2A" metricMagnitude="-2" metricUnit="m" metricValue="1.68" unit="mm" value="16.8">16.8 mm</quantity>
|
||||
); dorsal surface of forelimbs and posterior surface of brachia covered with large, keeled, imbricate scales; six rows of enlarged, keeled scales across forearm; ventral surface of forelimbs covered with granular scales; plantar scales small, granular; subdigital lamellae of fingers are composed of a single wide transversely elongated scales; claws long; hind limbs relatively long (FM
|
||||
<quantity id="E35DAC567C40856A61F0A396A302C5FB" metricMagnitude="-2" metricUnit="m" metricValue="3.36" unit="mm" value="33.6">33.6 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0EUCBG" xlink_title="tibia length">TIB</abbrev>
|
||||
<quantity id="C3FB3F3A230DF02AF1928EF7E1303053" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" unit="mm" value="27.2">27.2 mm</quantity>
|
||||
); dorsal scales on hind limbs small, weakly keeled; scales on anterior surface of thighs large, flat, weakly keeled, imbricate; those on forelegs slightly enlarged, keeled; postfemoral scales small, granular; eight longitudinal rows of large, smooth, flat, imbricate subtibial scales; 36 total femoral pores; each pore set in larger scale; 25 non-pore-bearing scales between pore-bearing-femoral scales across the pelvis; plantar scales small, raised; subdigital lamellae of toes bicarinate, 32 enlarged, plate-like scales along dorsal surface of the fourth toe; three enlarged, triangular scales on posterior surface at base of third toe (
|
||||
<abbrev id="ABBRID0EYCBG" xlink_title="length of the first enlarged, subdigital lamella on the third toe">ESL</abbrev>
|
||||
<quantity id="0ED820B17ABA265E74B6C128B58CE4C9" metricMagnitude="-3" metricUnit="m" metricValue="2.6" unit="mm" value="2.6">2.6 mm</quantity>
|
||||
;
|
||||
<abbrev id="ABBRID0E3CBG" xlink_title="length of the first enlarged, subdigital lamella on the third toe">ESL</abbrev>
|
||||
/
|
||||
<abbrev id="ABBRID0EADBG" xlink_title="third toe length">TE</abbrev>
|
||||
0.16); seven enlarged, plate-like scales along length of first toe; tail dorsoventrally compressed, noticeably wider at base, constricted at its contact point with body, covered dorsally with small, keeled scales grading ventrally into larger, flat, weakly keeled, subcaudals; caudal scales in transverse rows encircle the tail; and the last
|
||||
<quantity id="B74E7B0036677DE5E1F69F9C82FF6E05" metricMagnitude="-1" metricUnit="m" metricValue="1.1" unit="mm" value="110.0">110 mm</quantity>
|
||||
of the tail is regenerated (Fig.
|
||||
<figureCitation id="5A70ECB098D8D9186CAB4196B3AD71DD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal and ventral views of the holotype and paratypes of Leiolepis glaurung sp. nov. A male holotype THNHM 30909 B female paratype THNHM 30908 C male paratype THNHM 30910, and D male paratype THNHM 30911." figureDoi="10.3897/zookeys.1210.127557.figure5" httpUri="https://binary.pensoft.net/fig/1122035">5 A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EIDBG" type="Coloration in life">
|
||||
<paragraph id="9A834ED7A0F6F250E0BB7EE79BF02BE1">
|
||||
<heading id="B43D085631DC7D57DC3355F503DF7496" reason="title">Coloration in life.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="7E914330C282E2E7BB74A77BAEC07D6D">
|
||||
Dorsal ground color of head, body, limbs, and tail is grey to almost black (when animal is cold base color is black); anterior portion of the head pale grey with no pattern; three white lines radiate from the posterior region of the orbit with one extending posteriorly to the parietal region, one to the tympanic region, one to the corner of the mouth; four yellow stripes extending posteriorly from the parietal region of the head, the two central stripes connect on the nape of the neck forming a yellow Y-shape and extend posteriorly as a vertebral stripe that terminating at the anterior margin of the pelvis, the two lateral stripes extend dorsolaterally from the parietal region running the entire length of the body terminating at the anterior margin of pelvis; dorsal pattern is composed of three yellow dorsal stripes (mentioned above) separated by distinct darker regions with yellow spots; the base color of the flanks are yellowish orange with one black transverse bar in the axillary region, followed by pale yellow transverse bands composed of small yellow spots, three (L) and four (R); the forearms have prominent yellow and white spots; the hind limbs have small white ocelli with diffuse edges; the dorsal caudal coloration is composed of the same small ocelli as on the hind limbs and the coloration turns into dark transverse caudal bars approximately 30 % down the length of the tail; the gular region is black with a wide medial yellow strip; pectoral region is yellow with black mottling; the limb ventral color is yellow; subcaudal coloration is bright red to orange and extends laterally to the dorsolateral coloration of the tail (Figs
|
||||
<figureCitation id="9BD99D82E8E188039B46F3152028716E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal and ventral views of the holotype and paratypes of Leiolepis glaurung sp. nov. A male holotype THNHM 30909 B female paratype THNHM 30908 C male paratype THNHM 30910, and D male paratype THNHM 30911." figureDoi="10.3897/zookeys.1210.127557.figure5" httpUri="https://binary.pensoft.net/fig/1122035">5 A</figureCitation>
|
||||
,
|
||||
<figureCitation id="CCDF48E2BBFFFFEAC46ED9A0580AD05E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsal and ventral views of paratypes THNHM 30906, THNHM 30907, THNHM 30904, THNHM 30902, THNHM 30905, and THNHM 30903 Leiolepis glaurung sp. nov. Specimen tags bear field numbers." figureDoi="10.3897/zookeys.1210.127557.figure6" httpUri="https://binary.pensoft.net/fig/1122036">6</figureCitation>
|
||||
,
|
||||
<figureCitation id="0F2DC1185B08259A0D3E415B829CE779" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Female paratype THNHM 30908 B male paratype THNHM 30911 of Leiolepis glaurung sp. nov." figureDoi="10.3897/zookeys.1210.127557.figure9" httpUri="https://binary.pensoft.net/fig/1122039">9</figureCitation>
|
||||
,
|
||||
<figureCitation id="153332221E19FA58699F9547832A8F0C" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Male paratype THNHM 30911 displaying the dorsal and lateral display colors and its rocky habitat in the background B adult male Leiolepis rubritaeniata from southeastern Cambodia C male paratype THNHM 30911 of Leiolepis glaurung sp. nov." figureDoi="10.3897/zookeys.1210.127557.figure11" httpUri="https://binary.pensoft.net/fig/1122041">11</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
<caption id="E2D7AF780B82701648F79BB2424097F4" ID-DOI="10.3897/zookeys.1210.127557.figure9" ID-arpha="D2539228-2B41-5508-BD55-E5EF99485E89" httpUri="https://binary.pensoft.net/fig/1122039" startId="F9">
|
||||
<paragraph id="65BBD70374BDB136B0CF6461A41A86D1">
|
||||
<label id="3CA486584330993CFCB008BAE85B8EF4">Figure 9.</label>
|
||||
</paragraph>
|
||||
<paragraph id="0CF5768FDA7A71B8F95B0F5779EEFDDC">
|
||||
<emphasis id="21D2E791471F3531E34BBD6886760422" bold="true">A</emphasis>
|
||||
Female paratype THNHM 30908
|
||||
<emphasis id="D804B0FE85E38F3A075252B11C2AB360" bold="true">B</emphasis>
|
||||
male paratype THNHM 30911 of
|
||||
<taxonomicName id="1F3DF538FFCAB84F2B36841F4C3CF154" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="67EC526462E6897ABF5AA4224F72AA3B" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="6A45E16D518884EBFB7932BD9FAD8742" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E1EBG" type="etymology">
|
||||
<paragraph id="03BAEB13C0428E2CA83C5B2B5A71012F">
|
||||
<heading id="2E9215EF516AB0ED93D49446F7AB1DA8" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="ED4629B1E0115668F12684072E7E0682">
|
||||
The specific epithet
|
||||
<taxonomicName id="33D50D1E6368D68C59124DD000D0CC42" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung">
|
||||
<emphasis id="BA1667FF9389A049684B558E16C5588E" italics="true">glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
is in reference to the large, terrestrial, golden-colored, non-winged dragon, Glaurung in Middle-earth – a character created by J. R. R. Tolkien in
|
||||
<emphasis id="EBE462C59EFE347213AB514480C87378" italics="true">The Silmarillion</emphasis>
|
||||
(1977). Glaurung the Golden is the father of all dragons and tunneled into the sides of mountains forming burrows. The reduced expandable lateral flanks, yellow ventral and dorsal colors, with the construction of burrows beneath rocky outcrops is similar to the descriptions of Glaurung mentioned above, from ‘ The Silmarillion’ and ‘ The Children of Húrin’ (
|
||||
<bibRefCitation id="4D1D390A83EBEDF1C9F05AFCBCC8E1E2" author="Tolkien" firstAuthor="Tolkien" journalOrPublisher="George Allen and Unwin Ltd., Great Britain" refId="B52" refString="Tolkien JRR (1977) The Silmarillion. George Allen and Unwin Ltd., Great Britain, 365 pp." title="The Silmarillion." year="1977">Tolkien 1977</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="7F88FD502C86B0DFF8C8E523E67BC7EE" author="Tolkien and Tolkien" firstAuthor="Tolkien" journalOrPublisher="Boston MA, USA" refId="B53" refString="Tolkien JRR, Tolkien C (2007) The Children of Húrin. Houghton Mifflin. Boston MA, USA, 320 pp." title="The Children of Húrin. Houghton Mifflin." year="2007">Tolkien and Tolkien 2007</bibRefCitation>
|
||||
). Additionally, in
|
||||
<collectingCountry id="9808505ACC6BE6711FC50D7E09F0F5C4" name="Thailand">Thailand</collectingCountry>
|
||||
the word “ Yae ” is used to refer to
|
||||
<taxonomicName id="0A07F18916AF99B5A83FCD6DF20F48DF" baseAuthorityName="Hardwicke & Gray" baseAuthorityYear="1827" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="belliana">
|
||||
<emphasis id="9859BD010D1C73A53BCC8F57626380A4" italics="true">Leiolepis belliana</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="26A4C276534547C6514B1D59B5AF2851" authorityName="Peters" authorityYear="1971" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="ocellata">
|
||||
<emphasis id="C849E14C029726907B5FEE5AA75F5808" italics="true">L. ocellata</emphasis>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="761C51D4434FFE0B4A38A514EB5DBF21" authorityName="Mertens" authorityYear="1961" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="rubritaeniata">
|
||||
<emphasis id="BE2C97CCA8F9EE254A32057DFFF71644" italics="true">L. rubritaeniata</emphasis>
|
||||
</taxonomicName>
|
||||
. However, on the
|
||||
<collectingRegion id="1BA12720D2BEC647B0B7CFFCCB6A5A0A" country="Thailand" name="Nakhon Ratchasima">Khorat</collectingRegion>
|
||||
Plateau the populations of
|
||||
<taxonomicName id="3EA88782DDF5FCDE08ED781FB1F2B2A7" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="B5E0619A10FB3EE625CEEAA7969AF22E" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="63324F606E6DF31BB908294C2ACC3FDD" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
are called “ Yarb ”.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EAHBG" type="distribution">
|
||||
<paragraph id="0511E0EB92E2AB3D7CD20D87FD643352">
|
||||
<heading id="CEE9AF8E278E5D8208BFE418A2282115" reason="title">Distribution.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="8CAEEF8EB5C1A4F7D478F8A2C72B3606">
|
||||
<taxonomicName id="B0FE53C092D9EBCB75AEABAFD157616A" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="0BE18B79C6A4BE5BCF82C811713E8113" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="D9C921782672041ADD348C1F36A2EDCB" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is currently known from three locations on the Khorat Plateau (Fig.
|
||||
<figureCitation id="3ABEC7157F7699BDF3D17A23643C74A7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The distribution of all Leiolepis species and the localities of specimens used in this study. Question marks indicate an area where new populations could possibly occur. The different shades of the green circles for Leiolepis peguensis correspond to their phylogenetic positions in Fig. 2." figureDoi="10.3897/zookeys.1210.127557.figure1" httpUri="https://binary.pensoft.net/fig/1122031">1</figureCitation>
|
||||
). We have only collected specimens from the type locality in Ubon Ratchathani Province (Fig.
|
||||
<figureCitation id="1C8C0381EFB95E5CDD43A90766FF089F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The distribution of all Leiolepis species and the localities of specimens used in this study. Question marks indicate an area where new populations could possibly occur. The different shades of the green circles for Leiolepis peguensis correspond to their phylogenetic positions in Fig. 2." figureDoi="10.3897/zookeys.1210.127557.figure1" httpUri="https://binary.pensoft.net/fig/1122031">1</figureCitation>
|
||||
) from the Phu Phan Formation (
|
||||
<bibRefCitation id="5EA58F1C9CCF6851E07A6DE6DE5A2819" author="Jin-geng and Meesook" firstAuthor="Jin-geng" refId="B23" refString="Jin-geng S, Meesook A (2013) Non-Marine Cretaceous Bivalve Biostratigraphy of Thailand and Southern Lao PDR. In The 2 nd Lao-Thai Technical Conference on Geology and Mineral Resources, Vientiane, Lao PDR. 17–18 January 2013, 47–69." year="2013">Jin-geng and Meesook 2013</bibRefCitation>
|
||||
), however a population from Chaiyaphum Province (
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EAIBG" specific-use="{"type":"Point","coordinates":[102.164000,15.902200]}">
|
||||
<geoCoordinate id="069EFB61B14CABB8E7AF613F0B6C01E1" degrees="15" direction="north" minutes="54" orientation="latitude" precision="1" seconds="7.92" value="15.9022">15 ° 54 ' 7.92 " N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="5B6221DB74E6A65908D8060C809A0788" degrees="102" direction="east" minutes="9" orientation="longitude" precision="1" seconds="50.4" value="102.164">102 ° 9 ' 50.4 " E</geoCoordinate>
|
||||
</named-content>
|
||||
) in the western region of the Khorat Plateau was observed and photographed, but no specimens were collected (AK, AR, and PP pers. obs.), and from southern extent of the plateau on the border of
|
||||
<collectingCountry id="2B58E7877C90B862C01AA261AE125F2B" name="Cambodia">Cambodia</collectingCountry>
|
||||
(
|
||||
<abbrev id="ABBRID0EFIBG" xlink_title="pelvis width">PW</abbrev>
|
||||
pers. obs.) in Ubon Ratchathani Province (Fig.
|
||||
<figureCitation id="87178AC88160E0D0A99A4C2D416A47E1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The distribution of all Leiolepis species and the localities of specimens used in this study. Question marks indicate an area where new populations could possibly occur. The different shades of the green circles for Leiolepis peguensis correspond to their phylogenetic positions in Fig. 2." figureDoi="10.3897/zookeys.1210.127557.figure1" httpUri="https://binary.pensoft.net/fig/1122031">1</figureCitation>
|
||||
). Lizards from both populations are very similar in appearance to those from the type locality.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ENIBG" type="variation">
|
||||
<paragraph id="27F8AC5DD82C3C5D2E13BDBDA32B8BB7">
|
||||
<heading id="72E14AC73A3E5286B13027F2A2C8259E" reason="title">Variation.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="53BB89AD7F3194CCB849C50A0F4144C7">
|
||||
Differences in scale counts and measurements are presented in Suppl. material
|
||||
<xref id="101EC1C2F74D95338EE82F353B815526" ref-type="supplementary-material" rid="S1">1</xref>
|
||||
. The coloration of the males THNHM 30911 and THNHM 30910 are similar to the
|
||||
<typeStatus id="9ED54E7B871916CC7FEA73A04D3AF693" type="holotype">holotype</typeStatus>
|
||||
but their subcaudal coloration is a much more vibrant coral-red and they have complete original tails (Fig.
|
||||
<figureCitation id="7E7F8B2C957F88959B7D09C56C68759A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dorsal and ventral views of the holotype and paratypes of Leiolepis glaurung sp. nov. A male holotype THNHM 30909 B female paratype THNHM 30908 C male paratype THNHM 30910, and D male paratype THNHM 30911." figureDoi="10.3897/zookeys.1210.127557.figure5" httpUri="https://binary.pensoft.net/fig/1122035">5 C, D</figureCitation>
|
||||
). The coloration of the yellow stripes and spots that composed the dorsal pattern, and the flank coloration are not as pronounced in the female specimens THNHM 30904–30906 and 30908 as they are in the males (Figs
|
||||
<figureCitation id="991D6FB8858A34BC5AA6518147E86887" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsal and ventral views of paratypes THNHM 30906, THNHM 30907, THNHM 30904, THNHM 30902, THNHM 30905, and THNHM 30903 Leiolepis glaurung sp. nov. Specimen tags bear field numbers." figureDoi="10.3897/zookeys.1210.127557.figure6" httpUri="https://binary.pensoft.net/fig/1122036">6</figureCitation>
|
||||
,
|
||||
<figureCitation id="2D7F9FE0CEFFE84419F7412DFEE4B147" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Female paratype THNHM 30908 B male paratype THNHM 30911 of Leiolepis glaurung sp. nov." figureDoi="10.3897/zookeys.1210.127557.figure9" httpUri="https://binary.pensoft.net/fig/1122039">9</figureCitation>
|
||||
; Suppl. material
|
||||
<xref id="52D8EE3267901CCECCF7E9664645CFF4" ref-type="supplementary-material" rid="S1">1</xref>
|
||||
). Additionally, the dorsal caudal pattern is only faintly visible in the female specimens (Figs
|
||||
<figureCitation id="F663012D91712E79D26CCF61DE2AE5D6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsal and ventral views of paratypes THNHM 30906, THNHM 30907, THNHM 30904, THNHM 30902, THNHM 30905, and THNHM 30903 Leiolepis glaurung sp. nov. Specimen tags bear field numbers." figureDoi="10.3897/zookeys.1210.127557.figure6" httpUri="https://binary.pensoft.net/fig/1122036">6</figureCitation>
|
||||
,
|
||||
<figureCitation id="9AE183DA016EAB2F377B84170F437E94" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Female paratype THNHM 30908 B male paratype THNHM 30911 of Leiolepis glaurung sp. nov." figureDoi="10.3897/zookeys.1210.127557.figure9" httpUri="https://binary.pensoft.net/fig/1122039">9</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EPJBG" type="natural history">
|
||||
<paragraph id="AB6FF7883B3F83D37701DB6E91167EBD">
|
||||
<heading id="8ADABA25D1CA23CE6FCFAFFB37134251" reason="title">Natural history.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="1F54B3076281DB5C289C0F8F54D1CD58">
|
||||
Similar to other species of
|
||||
<taxonomicName id="C5DE7247E005EE14D4426AB71C4E505F" authorityName="Cuvier" authorityYear="1829" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="genus">
|
||||
<emphasis id="CBBF12B2727A3BFEF10FBDE9FD88537A" italics="true">Leiolepis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="33DD79A5E8779A33975F41722C6FD71E" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="C793C3D8DB12163A09458C0EA4767ECC" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="7AECF07998BD41467434549BF728F43C" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
constructs subterranean tunnels. However, given that they live exclusively in rocky habitats, individuals make compressed and shallow burrows in patches of loose soil underneath rocks or rockpiles (Figs
|
||||
<figureCitation id="D8369AB8B41D61DFCACBA7840695FAD5" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. The rocky habitat of the Khorat Plateau from the type locality." figureDoi="10.3897/zookeys.1210.127557.figure10" httpUri="https://binary.pensoft.net/fig/1122040">10</figureCitation>
|
||||
–
|
||||
<figureCitation id="CE14C17C24D23EEE6D6F08462F7694EA" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A Two burrow entrances going beneath a large rock B multiple small burrow entrances in the soil underneath and around a rock pile C a single entrance hole to a burrow beneath a large rock D the rock from panel C lifted to show the shallow subterranean burrow of an individual Leiolepis glaurung sp. nov." figureDoi="10.3897/zookeys.1210.127557.figure12" httpUri="https://binary.pensoft.net/fig/1122042">12</figureCitation>
|
||||
). Males tend to forage during the hottest part of the day and the feces we found appeared to be mostly composed of vegetation and arthropods.
|
||||
<taxonomicName id="493094B4B6CFC5479A0CB2BB72E41C08" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="8968385CA54F4396B64B2083FFC4A4AD" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="E84DD349B7E69682BF336BD32E293A30" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is a food source for local people in the area (as many
|
||||
<taxonomicName id="3B2A2EA78D28F967F1E203676E77A3E3">
|
||||
<emphasis id="F534D63D4AB3D7B6F9EE234171AAB703" italics="true">Leiolepis</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
are across Indochina;
|
||||
<bibRefCitation id="84E08ADC93B52AB77D4436F5A1F9488D" DOI="10.11646/zootaxa.2433.1.3" author="Grismer and Grismer" firstAuthor="Grismer" issue="1" journalOrPublisher="Zootaxa" pagination="47-61" refId="B17" refString="Grismer JL, Grismer LL (2010) Who’s your mommy? Identifying maternal ancestors of unisexual species of Leiolepis Cuvier, 1829 and the description of a new endemic species of unisexual Leiolepis Cuvier, 1829 from southern Vietnam. Zootaxa 2433 (1): 47–61. https://doi.org/10.11646/zootaxa.2433.1.3" title="Who’s your mommy? Identifying maternal ancestors of unisexual species of Leiolepis Cuvier, 1829 and the description of a new endemic species of unisexual Leiolepis Cuvier, 1829 from southern Vietnam." volume="2433" year="2010">Grismer and Grismer 2010</bibRefCitation>
|
||||
) and talking with local collectors, they report that individuals are highly philopatric. The collectors say that if the burrow is open that means they are out foraging and if the burrow is plugged with soil they are inside. Additionally, local people say that
|
||||
<taxonomicName id="7D78B1D08EE1A4B5D2AB923E923E6AB4" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="1B7D1BA811CD7EDEA490AA7E7FD0C88A" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="6B7DC862C87A416B015EBD827E77ABEA" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
is mostly active during the dry season (November – March) and estivates during the rainy season (April – November) but will come out of their burrows after rains to eat arthropods. Lastly, all individuals observed and collected (specimens THNHM 30908–30911) were found during the day at 36 ° C and 77 % humidity. Other species of reptiles observed sharing the same habitat at the type locality included
|
||||
<taxonomicName id="47A24676ECBDFD2B4B0D1AB5B703EAAB" authority="(Daudin, 1802)" baseAuthorityName="Daudin" baseAuthorityYear="1802" class="Squamata" family="Agamidae" genus="Calotes" kingdom="Animalia" phylum="Chordata" rank="species" species="versicolor">
|
||||
<emphasis id="76ECD99CEB1258B96BB22261598621BD" italics="true">Calotes versicolor</emphasis>
|
||||
(Daudin, 1802)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="92B7761CC6F3F10AA73ADF00CAEEB94C" authority="(Boulenger, 1899)" baseAuthorityName="Boulenger" baseAuthorityYear="1899" class="Squamata" family="Gekkonidae" genus="Dixonius" kingdom="Animalia" phylum="Chordata" rank="species" species="siamensis">
|
||||
<emphasis id="381F8274FD6BCCE30D6FBD7BCF0A0AE5" italics="true">Dixonius siamensis</emphasis>
|
||||
(Boulenger, 1899)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="70DC09085AFBD7980DD1EDEE3EC9F744" authority="Taylor, 1962" authorityName="Taylor" authorityYear="1962" class="Squamata" family="Gekkonidae" genus="Gekko" kingdom="Animalia" phylum="Chordata" rank="species" species="petricolus">
|
||||
<emphasis id="C9AF8C9C1C8A27D2839549690CC12A97" italics="true">Gekko petricolus</emphasis>
|
||||
Taylor, 1962
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="5008FBA9EE5A7FBFC1923E6F4ADAB5F9" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Squamata" family="Gekkonidae" genus="Gekko" kingdom="Animalia" phylum="Chordata" rank="species" species="gecko">
|
||||
<emphasis id="03C9DC5ECC32B6C36894733F572CC431" italics="true">G. gecko</emphasis>
|
||||
(Linnaeus, 1758)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="706DC272C358BB17B8F8123461033C5E" authority="(Boulenger, 1887)" baseAuthorityName="Boulenger" baseAuthorityYear="1887" class="Squamata" family="Scincidae" genus="Scincella" kingdom="Animalia" phylum="Chordata" rank="species" species="melanosticta">
|
||||
<emphasis id="3A99EA75A93B9EDF94A6653CBE52C5CF" italics="true">Scincella melanosticta</emphasis>
|
||||
(Boulenger, 1887)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="41C64E5785E44CAEFCCF5D5507584B14" authority="(Smith, 1916)" baseAuthorityName="Smith" baseAuthorityYear="1916" class="Squamata" family="Scincidae" genus="Scincella" kingdom="Animalia" phylum="Chordata" rank="species" species="rupicola">
|
||||
<emphasis id="1B0C37A72298D7D877C57CF1E4C855D0" italics="true">S. rupicola</emphasis>
|
||||
(Smith, 1916)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="57BC5286C14939240372ABBF9D38B21B" authority="(Blyth, 1853)" baseAuthorityName="Blyth" baseAuthorityYear="1853" class="Squamata" family="Scincidae" genus="Eutropis" kingdom="Animalia" phylum="Chordata" rank="species" species="macularia">
|
||||
<emphasis id="FB278BDC4D1535EA56D5CF85B817586B" italics="true">Eutropis macularia</emphasis>
|
||||
(Blyth, 1853)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="182698C8A138D9D575D48B33B9412F03" authority="(Shaw, 1802)" baseAuthorityName="Shaw" baseAuthorityYear="1802" class="Squamata" family="Colubridae" genus="Chrysopelea" kingdom="Animalia" phylum="Chordata" rank="species" species="ornata">
|
||||
<emphasis id="74686B663503D3F72C225E63C7348F25" italics="true">Chrysopelea ornata</emphasis>
|
||||
(Shaw, 1802)
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="8AAE0E93BB8257524770BDB0E28BACE8" authority="Gunther, 1864" authorityName="Gunther" authorityYear="1864" class="Squamata" family="Colubridae" genus="Lycodon" kingdom="Animalia" phylum="Chordata" rank="species" species="laoensis">
|
||||
<emphasis id="171CD9EB7F78DB08D5AE762213EEB9CA" italics="true">Lycodon laoensis</emphasis>
|
||||
Günther, 1864
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="EA9BF56C3C219AB67702FE107A09CB48" authority="(Kuhl, 1824)" baseAuthorityName="Kuhl" baseAuthorityYear="1824" class="Squamata" family="Viperidae" genus="Calloselasma" kingdom="Animalia" phylum="Chordata" rank="species" species="rhodostoma">
|
||||
<emphasis id="3AC46587D40DF7B8A5EBFEE198430603" italics="true">Calloselasma rhodostoma</emphasis>
|
||||
(Kuhl, 1824)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="F4F6AE98701D0F49D5168BDFE8CE343E" ID-DOI="10.3897/zookeys.1210.127557.figure10" ID-arpha="EE5DC932-5E61-5E66-A4F4-46A3EC5AD193" httpUri="https://binary.pensoft.net/fig/1122040" startId="F10">
|
||||
<paragraph id="BEDAA16EA1A8AAFA38451C54FF9D5C8D">
|
||||
<label id="EE2A4B11A065C6453DA79C7C0F74EE36">Figure 10.</label>
|
||||
</paragraph>
|
||||
<paragraph id="2AE98ED3FB5E3727CDAB0C1DA4253A6C">The rocky habitat of the Khorat Plateau from the type locality.</paragraph>
|
||||
</caption>
|
||||
<caption id="1FC58A30ECEC8AF3C7DF24AD64AB3979" ID-DOI="10.3897/zookeys.1210.127557.figure11" ID-arpha="2BE8B181-F737-584F-8052-60B8CCDE926A" httpUri="https://binary.pensoft.net/fig/1122041" startId="F11">
|
||||
<paragraph id="7BC7F7B5888652634BA28387ED9F68D8">
|
||||
<label id="3ED34C309181B79B95557F0F4F21D622">Figure 11.</label>
|
||||
</paragraph>
|
||||
<paragraph id="A0C3FADAD4833F4B044E33B3E3DB61B0">
|
||||
<emphasis id="54726F024CE78EBA7F938DD71313C27D" bold="true">A</emphasis>
|
||||
Male paratype THNHM 30911 displaying the dorsal and lateral display colors and its rocky habitat in the background
|
||||
<emphasis id="C140C891E5B98B038946092C9B378334" bold="true">B</emphasis>
|
||||
adult male
|
||||
<taxonomicName id="4450F69A7ABC2FA0CD7E77AC31AF1346" authorityName="Mertens" authorityYear="1961" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="rubritaeniata">
|
||||
<emphasis id="50ECA080023191BB285DA977542C4C97" italics="true">Leiolepis rubritaeniata</emphasis>
|
||||
</taxonomicName>
|
||||
from southeastern Cambodia
|
||||
<emphasis id="51E2D1407AD45F1075A0630AF4783240" bold="true">C</emphasis>
|
||||
male paratype THNHM 30911 of
|
||||
<taxonomicName id="2C41D9FB8EA4E6A7F9A3D07C06ED7985" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="6D7BFBA941F7D6FED0F202CEB71ECA26" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="80AEEF429DA685D3DC1DF64C83294E53" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="F4DBFBC02E1B507686DCA773C8EE5C2A" ID-DOI="10.3897/zookeys.1210.127557.figure12" ID-arpha="FC42E27C-B5C1-5628-8221-7E0DA23139A0" httpUri="https://binary.pensoft.net/fig/1122042" startId="F12">
|
||||
<paragraph id="B1C5F6E4E3FBECF3584CB5070DA4888E">
|
||||
<label id="390C02F4FEEBE7736E692131449F3FE1">Figure 12.</label>
|
||||
</paragraph>
|
||||
<paragraph id="FCBD9E0DFBD516478BBCD8A7FDF65E03">
|
||||
<emphasis id="2824C17F3A1A1CBA8114F40370043E4C" bold="true">A</emphasis>
|
||||
Two burrow entrances going beneath a large rock
|
||||
<emphasis id="EA2F79F103ABB7F1E65426657B531DCB" bold="true">B</emphasis>
|
||||
multiple small burrow entrances in the soil underneath and around a rock pile
|
||||
<emphasis id="4C04D2C2770C11FA87BEAB7DE57EC410" bold="true">C</emphasis>
|
||||
a single entrance hole to a burrow beneath a large rock
|
||||
<emphasis id="37E4D24F671A72DCBB4A9B4D54E8834F" bold="true">D</emphasis>
|
||||
the rock from panel C lifted to show the shallow subterranean burrow of an individual
|
||||
<taxonomicName id="39ED76C7F1807F8DB9CDC9C8110B0937" authorityName="Wanchai & Rujirawan & Murdoch & Aksornneam & Promnun & Kaatz & Gregory & Nguyen & Iderstein & Quah & Grismer & Grismer & Aowphol" authorityYear="2024" class="Squamata" family="Agamidae" genus="Leiolepis" kingdom="Animalia" phylum="Chordata" rank="species" species="glaurung" status="sp. nov.">
|
||||
<emphasis id="62D7A2827830512D938D30A16C8CFC6C" italics="true">Leiolepis glaurung</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="AEA3870A2780B0771C477EFA9A2175EA" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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