diff --git a/data/03/CD/6B/03CD6B03FFB1CE6BFF28423FFA25FB38.xml b/data/03/CD/6B/03CD6B03FFB1CE6BFF28423FFA25FB38.xml index c44927f9c0f..0eff60b48df 100644 --- a/data/03/CD/6B/03CD6B03FFB1CE6BFF28423FFA25FB38.xml +++ b/data/03/CD/6B/03CD6B03FFB1CE6BFF28423FFA25FB38.xml @@ -1,54 +1,54 @@ - - - -Hemimyzon luae, a new species of balitorid fish (Teleostei: Baltioridae) from the Tzengwen River basin, southern Taiwan + + + +Hemimyzon luae, a new species of balitorid fish (Teleostei: Baltioridae) from the Tzengwen River basin, southern Taiwan - - -Author + + +Author -Chen, I-Shiung -Institute of Marine Biology, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. & Center for Excellence of the Oceans, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. +Chen, I-Shiung +Institute of Marine Biology, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. & Center for Excellence of the Oceans, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. - - -Author + + +Author -Yeh, Ming-Fon -Biodiversity Research Center, Taiwan, R. O. C. +Yeh, Ming-Fon +Biodiversity Research Center, Taiwan, R. O. C. -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-12-10 + +2024 + +2024-12-10 - -5550 + +5550 - -1 + +1 - -278 -286 + +278 +286 - -https://doi.org/10.11646/zootaxa.5550.1.28 + +https://doi.org/10.11646/zootaxa.5550.1.28 -journal article -10.11646/zootaxa.5550.1.28 -1175-5326 -14389219 -96E8FBAF-E0D2-4B99-BDE0-DC4761F49B32 +journal article +10.11646/zootaxa.5550.1.28 +1175-5326 +14389219 +96E8FBAF-E0D2-4B99-BDE0-DC4761F49B32 - + @@ -63,38 +63,50 @@ Chen, Harefa, Chang, Han - + Holotype -.—NTOUP-2021-12-325, +.— +NTOUP-2021-12-325 +, 55.3 mm -SL, Nar-Mar-Shar County, Nan-Tsi-Shien River, -Kaoping River -basin, coll. +SL, +Nar-Mar-Shar County +, +Nan-Tsi-Shien River +, +Kaoping River basin +, coll. C.C. Han , July, 20, 2021, -Kaohsiung City +Kaohsiung City , Taiwan , ROC. - + Paratypes -.—NTOUP-2021-12-326, +.— +NTOUP-2021-12-326 +, 8 specimens , 51.9–66.6 mm -SL, Nar-Mar-Shar County, Nan-Tsi-Shien River, -Kaoping River -basin, -Coll. C.C. Han +SL, +Nar-Mar-Shar County +, +Nan-Tsi-Shien River +, +Kaoping River basin +, Coll. +C.C. Han , July, 20, 2021, -Kaohsiung City +Kaohsiung City , Taiwan , ROC diff --git a/data/03/CD/6B/03CD6B03FFB1CE6BFF2844B3FA9DFCE8.xml b/data/03/CD/6B/03CD6B03FFB1CE6BFF2844B3FA9DFCE8.xml index 5ee64505918..4ef4e4816c2 100644 --- a/data/03/CD/6B/03CD6B03FFB1CE6BFF2844B3FA9DFCE8.xml +++ b/data/03/CD/6B/03CD6B03FFB1CE6BFF2844B3FA9DFCE8.xml @@ -1,52 +1,52 @@ - - - -Hemimyzon luae, a new species of balitorid fish (Teleostei: Baltioridae) from the Tzengwen River basin, southern Taiwan + + + +Hemimyzon luae, a new species of balitorid fish (Teleostei: Baltioridae) from the Tzengwen River basin, southern Taiwan - - -Author + + +Author -Chen, I-Shiung -Institute of Marine Biology, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. & Center for Excellence of the Oceans, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. +Chen, I-Shiung +Institute of Marine Biology, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. & Center for Excellence of the Oceans, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. - - -Author + + +Author -Yeh, Ming-Fon -Biodiversity Research Center, Taiwan, R. O. C. +Yeh, Ming-Fon +Biodiversity Research Center, Taiwan, R. O. C. -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-12-10 + +2024 + +2024-12-10 - -5550 + +5550 - -1 + +1 - -278 -286 + +278 +286 - -https://doi.org/10.11646/zootaxa.5550.1.28 + +https://doi.org/10.11646/zootaxa.5550.1.28 -journal article -10.11646/zootaxa.5550.1.28 -1175-5326 -14389219 -96E8FBAF-E0D2-4B99-BDE0-DC4761F49B32 +journal article +10.11646/zootaxa.5550.1.28 +1175-5326 +14389219 +96E8FBAF-E0D2-4B99-BDE0-DC4761F49B32 @@ -62,19 +62,21 @@ Chen & Fang - - + + Holotype .— -NTOU -P-2007-07-077, +NTOU P-2007-07-077 +, 50.1 mm SL, small tributary ( -3 km + +3 km + south to -Yi-Ting mountain +Yi-Ting mountain ) in upper reaches of the Tar-Ju River, Tar-Ren Village , @@ -88,14 +90,16 @@ south to July 11, 1993 -. - +. + + + Paratypes .— -NTOU -P- 2007-07-078, +NTOU P- 2007-07-078 +, 2 specimens , 11.2–30.6 mm diff --git a/data/03/CD/6B/03CD6B03FFB8CE62FF2844AEFF7DFCC2.xml b/data/03/CD/6B/03CD6B03FFB8CE62FF2844AEFF7DFCC2.xml deleted file mode 100644 index 4de87e3dccb..00000000000 --- a/data/03/CD/6B/03CD6B03FFB8CE62FF2844AEFF7DFCC2.xml +++ /dev/null @@ -1,77 +0,0 @@ - - - -Hemimyzon luae, a new species of balitorid fish (Teleostei: Baltioridae) from the Tzengwen River basin, southern Taiwan - - - -Author - -Chen, I-Shiung -Institute of Marine Biology, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. & Center for Excellence of the Oceans, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. - - - -Author - -Yeh, Ming-Fon -Biodiversity Research Center, Taiwan, R. O. C. - -text - - -Zootaxa - - -2024 - -2024-12-10 - - -5550 - - -1 - - -278 -286 - - - - -https://doi.org/10.11646/zootaxa.5550.1.28 - -journal article -10.11646/zootaxa.5550.1.28 -1175-5326 -14389219 -96E8FBAF-E0D2-4B99-BDE0-DC4761F49B32 - - - - - - - -Hemymizon luae - -new species - - - - -( -ṘOEOiȒẵẘ -) - - - - -( -Figs. 1–2 -) - - - - \ No newline at end of file diff --git a/data/03/CD/6B/03CD6B03FFB8CE65FF2844AEFD44FADF.xml b/data/03/CD/6B/03CD6B03FFB8CE65FF2844AEFD44FADF.xml new file mode 100644 index 00000000000..b6525082aab --- /dev/null +++ b/data/03/CD/6B/03CD6B03FFB8CE65FF2844AEFD44FADF.xml @@ -0,0 +1,1509 @@ + + + +Hemimyzon luae, a new species of balitorid fish (Teleostei: Baltioridae) from the Tzengwen River basin, southern Taiwan + + + +Author + +Chen, I-Shiung +Institute of Marine Biology, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. & Center for Excellence of the Oceans, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. + + + +Author + +Yeh, Ming-Fon +Biodiversity Research Center, Taiwan, R. O. C. + +text + + +Zootaxa + + +2024 + +2024-12-10 + + +5550 + + +1 + + +278 +286 + + + + +https://doi.org/10.11646/zootaxa.5550.1.28 + +journal article +10.11646/zootaxa.5550.1.28 +1175-5326 +14389219 +96E8FBAF-E0D2-4B99-BDE0-DC4761F49B32 + + + + + + + +Hemymizon luae + +new species + + + + + + +( +ṘOEOiȒẵẘ +) + + + + +( +Figs. 1–2 +) + + + + + +Materials examined + + + + + +Holotype + +.— +NTOUP-2022-10-361 +, +72.4 mm +SL, coll. +I-S. Chen +, coll. + +Oct. 22 2022 + +, +Alishan District +, +Tzengwen River basin +, +Tainan City +, +Taiwan + +. + + + + +Paratypes + +.— +NTOUP-2022-10-362 +, +9 specimens +, +57.8–71.6 mm +SL, the collection data same as holotype + +. + + + + +Diagnosis + + +The new species can be well distinguished from other congeneric species by following combination of features: (1) dorsal fin rays 3 + 8; pectoral fin rays 11–12 + 9–11 (total 20–22; modally 22); (2) pelvic fin moderate large, extending to rear vertical of dorsal fin, pelvic simple rays modally 3 which fewer than any other endemic ones of +Taiwan +; (3) lateral-line scales 68–72 (modally 70); predorsal scales 39–42 (modally 40); (4) the position of anus with much larger distance of pelvic rear tip to anus about 2.4–3.2 times of that of anus to anal fin origin; and (5) specific coloration: dorsum of body and head olive brown to greenish brown; predrorsal region deep brown with several rounded or oblong cloudy deep brown patches; lateral body uniformly olive brown to deep brown; and pectoral, pelvic and anal fins pale white with deep brown rays. + + + + +Description + + +The morphometrics of this new species as percentages of standard length are listed in +Table 1 +and meristic features are listed in +Table 2 +. Head and body very strongly depressed with flat ventral side anteriorly. Posterior trunk from anus to caudal peduncle rather compressed. See +Table 1 +for morphometric characters. Head with a few tiny tubercles. Upper lip with 9–12 small papillae; no distinct papillae on lower lip except a pair of somewhat crescent projections on inner side. Four rostral barbels and two barbels at both corners of mouth which anterior one similar to length of posterior one. The length of anterior barbels smaller than that of the eye diameter. Interorbital region rather wide. Gill-opening small and very restricted, merely extending above anterior origin of pectoral fin. The location of anus is closer to anal fin origin, with large distance of rear tip of pelvic fin to anus about 2.4–3.2 times to that of anus to anal fin origin. + +Dorsal fin 3+8; anal fin 2+5; pectoral fin 11–12 + 9–11 (total rays 20–22; modally 22); pelvic fin 3–4 + 8–9 (simple rays modally 3; branched rays modally 8; total rays 11–13 modally 11). Origin of dorsal fin behind origin of pelvic fin origin. Pectoral fin rather large, its rear margin extending beyond origin of pelvic fin. Pelvic fin well separate, the gap between the attachment of their innermost rays about 2.0–2.5 times of eye diameter; its rear margin extending to the rear tip of dorsal fin when depressed. Caudal fin forked, its lower lobe always longer than upper one. + + +FIGURE 1. + +Hemimyzon luae + +, holotype, NTOUP-2022-12-361, 72.4 mm SL, upper: dorsal view; middle: lateral view; lower: ventral view; Alishan, Tzengwen River basin, Tainan City, Taiwan, ROC. + + + + +TABLE 1. +Morphometry of + +Hemimyzon luae + +from southern Taiwan. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
TypeHPPPP
SL (mm)72.468.871.270.071.6
Lateral head length23.5%23.4%22.1%23.8%21.3%
Dorsal head length21.7%21.2%21.5%21.9%21.0%
Ventral head length12.3%14.2%12.9%13.9%12.9%
Head width18.7%20.2%20.3%19.5%19.6%
Snout length13.1%13.9%13.8%13.5%13.0%
Eye diameter3.0%4.4%2.8%3.2%3.0%
Interorbital length10.6%11.2%11.0%10.1%10.1%
Mouth width10.8%10.0%10.1%9.9%10.7%
Body width at pectoral fin origin18.9%19.8%18.7%19.9%19.3%
Body width at pelvic fin origin22.9%26.2%26.5%25.6%24.6%
Body depth at dorsal fin origin14.2%17.7%16.3%14.8%16.6%
Caudal peduncle length12.5%13.5%14.8%11.0%10.8%
Caudal peduncle depth9.5%11.0%9.9%10.0%10.8%
Length of last simple pectoral fin rays16.0%18.9%17.9%19.2%14.2%
Pectoral fin length33.0%36.4%32.8%35.2%32.6%
Pelvic fin length27.2%28.0%26.9%27.4%24.8%
Length of upper caudal fin lobe23.7%26.6%23.1%22.7%22.8%
Length of lower caudal fin lobe28.1%27.6%24.3%27.6%24.6%
Distance between pelvic fin bases8.8%8.5%7.9%8.0%8.6%
Predorsal length48.9%55.3%50.7%51.1%51.0%
Prepectoral length16.1%17.0%16.5%16.7%16.9%
Distance between pelvic and pectoral fins20.8%31.5%29.6%31.6%29.4%
Anal fin base6.0%7.4%7.1%6.8%7.7%
Anus to A origin3.6%3.8%3.7%3.6%3.7%
Anus to Rear V8.9%9.0%10.6%9.0%12.0%
Anus to Rear V / Anus to A origin2.452.352.882.493.23
+ + + +H: +Holotype +; P: +Paratype +. + + +Dorsal part of body with very small cycloid scales. Larger specimens with reduced, smaller size of predorsal scales. Ventral region between the paired fins naked. Body scales slightly larger posteriorly. Lateral-line scales 69–72 (modally 70) and predorsal scales 39–32 (modally 40). + + +Coloration in fresh + + +Dorsum of body and head olive brown to greenish brown. Predrorsal region olive brown to deep brown with several rounded or oblong cloudy deep brown patches with lighter margin. Lateral body uniformly olive brown to deep brown. Ventral side unique pale creamy white. Dorsal fin pale white with deep brown rays with about 2–3 rows of indistinct deep brown spots. Pectoral, pelvic and anal fins pale white with deep brown rays. Caudal fin with broad black outer margin with 3-4 oblique, zigzag creamy white streaks. + + + +TABLE 2 +Comparison of distribution frequency of meristic features from five endemic nominal species of + +Hemimyzon + +in Taiwan. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
SpeciesPectoral fin
Simple raysBranched raystotal rays
1011121314M910111213M20212223242526M
+ +H. luae +n. sp. + + +1 + +10 + +3 + +- + +- + +11.1 + +1 + +7 + +6 + +- + +- + +10.4 + +1 + +5 + +8 + +- + +- + +- + +- + +21.5 +
+ +H. formosanus + +11912--11.37187--10.011912----21.3
+ +H. sehni + +---3-13.0--5--11.0----3--24.0
+ +H. taitungensis + +--17413.3--18312.2-----7525.4
+ +H. yushanensis + +-1172-11.64151--9.9--137---22.4
+ + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
SpeciesDorsal finPelvic fin
Branched raysSimple raysBranched raystotal rays
78M34567M891011M11121314151617M
+ +H. luae +n. sp. + +-78.0122---3.195--8.4932- +- + +- + +- + +11.5 +
+ +H. formosanus + +16-7.0-1616--4.5-2012-9.4--4271--13.9
+ +H. sehni + +-38.0-3---4.0--3-10.0---3---14.0
+ +H. taitungensis + +-68.0---866.5--6610.5------1217.0
+ +H. yushanensis + +-108.0-1631-4.3-614-9.7--2171--14.0
+ + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
SpeciesLateral line scales
686970717273747576777879808182838485868788M
+ +H. luae +n. sp. + + +2 + +3 + +4 + +2 + +3 + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +70.1 +
+ +H. formosanus + +1896521--------------70.5
+ +H. sehni + +----------121--------79.0
+ +H. taitungensis + +---------------12422185.4
+ +H. yushanensis + +-5645----------------70.5
+ + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
SpeciesPredorsal scales
0-25262728293031323334353637383940414142M
+ +H. luae +n. sp. + + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +- + +1 + +3 + +1 + +1 + +1 + +40.7 +
+ +H. formosanus + +-------------673-----36.8
+ +H. sehni + +--------11-----------31.5
+ +H. taitungensis + +6--------------------0
+ +H. yushanensis + +--133111-------------27.1
+ + + +Distribution + + +The new species + +Hemimyzon luae + +is merely found from the +type +locality of upper reaches of Tzengwen River basin, +Taiwan +. It may be only found in the Tzegnwen River basin, not occurring in others. + + + + +Etymology + + +The specific name, + +luae + +is named after the President of Yushan National Park - S.F. Lu, for the honor of her kindly supporting the ecological conservation and biodiversity research and also for all her great contribution for the National Park. + + + + +FIGURE 2. +Ventral view of mouth structure of + +Hemimyzon luae + +, holotype, 72.4 mm SL, NTOUP-2022-10-361, Alishan District, Tzengwen River basin, Tainan City. + + + + +FIGURE 3. +Alive specimen of +Hemimyzon luae +. + + + + + +Discussion + + + +In the taxonomy of + +Hemimyzon + +, four nonimal species were documented in mainland +China +( +Yue 2000 +) while recorded two endemic species of +Taiwan +(Tzeng & Chen 1982; +Shen 1993 +) before 2000. After then, +Chen and Fang (2009) +described + +Hemimyzon sheni + +from eastern +Taiwan +added the third endemic species to +Taiwan +. More recently, +Chen, Harefa, Chang & Han (2022) +described the fourth species— + +Hemimyzon yushanensis + +from southern +Taiwan +. After the current new species accounted, the island river basins with 5 endemic species of +Taiwan +which is top diversity rather than others of mainland +China +. Even the disjunction of geographical distribution of +Taiwan +and mainland +China +, we still consider the Chinese species may be convergent adaption to fast running waters but not directly belonging to true defined + +Hemimyzon +species + +of +Taiwan +. The more detailed morphological evidence might be needed to clarify the whole generic limitation of the genus. + + +The new species + +Hemimyzon luae + +would be the new discovery of species boundary geographically separating the + +Hemimyzon formosanus + +and + +Hemimyzon yushanensis +. + +Since their high endemicity in Taiwanese waters, the new species + +H. luae + +established, this species can be merely found from hill tributaries of the Tzengwen River basin in +Tainan +City. + +H. yushanensis + +can be found at least from Kaoping River basins in +Kaohsiung City +as well as +Pingtung County +. Therefore, the update, true range of + +H. formosanus + +would be more restricted to further northern regions as all river basins from the Ilan County, +Taipei +City, +New Taipei City +, +Taoyuan City +, Shinchu County, then southward to Charyi City which mainly in the western slope of Central Mountain Ridge. + +H. taitungensis + +can be seen from 3 main larger river basins from +Hualian +to +Taitung County +. + +H. sheni + +can only be seen in Tarchu river basin of +Taitung County +. Among them, all of them share allopatric distribution pattern without range overlapping. + + +Among the four congeneric, endemic species of +Taiwan +, about the differentiation of dorsal fin rays, there are apparently two groups of + +Hemimyzon + +in +Taiwan +. One group is dorsal fin rays 3+7 which now merely seen in wide distributed species, + +H. formosanus + +; another group is dorsal fin rays 3+8 which can be seen in the following three species as + +H. taitungensis + +, + +H. sheni + +, + +H. yushanensis + +as well as + +H. yushanensis + +. The 3+8 group only can be seen in eastern +Taiwan +as both + +H. taitungensis + +and + +H. sheni + +. + +H. yushanensis + +can be only found from the Kaoping River basin originating from southern slope of the Yushan mountain Ridge in southern +Taiwan +. + +H. luae + +can be merely regarded endemic in only one, Tzengwen River basin. + + + +H. luae + +shares the same dorsal fin ray formula, 3+8 with + +H. sheni + +and + +H. yushanensis + +as well as + +H. taitungensis + +. However, it can be well distinguished from + +H. taitungensis + +by the pectoral fin rays 20–22 vs. 25–26; pelvic fin total rays 11 vs. 17. It also can be well distinguished from both + +H. yushanensis + +and + +H. sheni + +by the lower counts of pelvic fin simple rays modally 3 vs. 4 and its total rays 11 vs. 14. The limited distribution of current species is needed for further formal concern of its ecological conservation issue. + + + + \ No newline at end of file diff --git a/data/03/CD/6B/03CD6B03FFBFCE65FF28435FFBF3F913.xml b/data/03/CD/6B/03CD6B03FFBFCE65FF28435FFBF3F913.xml new file mode 100644 index 00000000000..259dd2e1964 --- /dev/null +++ b/data/03/CD/6B/03CD6B03FFBFCE65FF28435FFBF3F913.xml @@ -0,0 +1,132 @@ + + + +Hemimyzon luae, a new species of balitorid fish (Teleostei: Baltioridae) from the Tzengwen River basin, southern Taiwan + + + +Author + +Chen, I-Shiung +Institute of Marine Biology, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. & Center for Excellence of the Oceans, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. + + + +Author + +Yeh, Ming-Fon +Biodiversity Research Center, Taiwan, R. O. C. + +text + + +Zootaxa + + +2024 + +2024-12-10 + + +5550 + + +1 + + +278 +286 + + + + +https://doi.org/10.11646/zootaxa.5550.1.28 + +journal article +10.11646/zootaxa.5550.1.28 +1175-5326 +14389219 +96E8FBAF-E0D2-4B99-BDE0-DC4761F49B32 + + + + + + +An artificial key to 5 endemic species + +Hemimyzon + +in +Taiwan +: + + + + + + + + + +1a Dorsal fin rays 3+7......................................................................... + +H. formosanus + + + + +1b Dorsal fin rays 3+8.................................................................................... 2 + + + + + +2a Pelvic fin rays 17; inner gap between pelvic fins very narrow....................................... + +H. taitungensis + + + + +2b Pelvic fin rays no more than 15; inner gap between pelvic fins wider than......................................... 3 + + + + + +3a Pelvic fin simple rays modally 3, total rays always 11.............................................. + + +H. luae + +n. sp. + + + + +3b Pelvic fin simple rays modally 4, total rays always 14........................................................ 4 + + + + + +4a Pectoral fin rays 24; lateral-line scales 78–80; the distance of pelvic rear tip to anus about 3 times to that of anus to anal fins.............................................................................................. + +H. sheni + + + + + +4b Pectoral fin rays modally 22; lateral-line scales 69–72; the distance of pelvic rear tip to anus about 1.2–1.8 times to that of anus to anal fins............................................................................... + +H. yushanensis + + + + + + + + \ No newline at end of file