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<mods:title id="649C9C602A0F32DB0CA07DB803FAEE4D">A synopsis of Actenosigynes Moure, Graf &amp; Urban, 1999 (Hymenoptera: Colletidae) — new species, possible oligolecty and biogeographic comments</mods:title>
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<treatment id="03D887E7FFA95D2BC4C3FBCE55CAB75C" ID-DOI="http://doi.org/10.5281/zenodo.6222423" ID-GBIF-Taxon="119626585" ID-Zenodo-Dep="6222423" LSID="urn:lsid:plazi:treatment:03D887E7FFA95D2BC4C3FBCE55CAB75C" httpUri="http://treatment.plazi.org/id/03D887E7FFA95D2BC4C3FBCE55CAB75C" lastPageId="6" lastPageNumber="21" pageId="2" pageNumber="17">
<subSubSection id="C36B657AFFA95D2FC4C3FBCE505DB2DC" pageId="2" pageNumber="17" type="nomenclature">
<paragraph id="8BCE36F1FFA95D2FC4C3FBCE536EB2F8" blockId="2.[151,625,1128,1190]" box="[151,625,1128,1155]" pageId="2" pageNumber="17">
<heading id="D086819DFFA95D2FC4C3FBCE536EB2F8" bold="true" box="[151,625,1128,1155]" fontSize="11" level="1" pageId="2" pageNumber="17" reason="1">
<emphasis id="B905EAE3FFA95D2FC4C3FBCE536EB2F8" bold="true" box="[151,625,1128,1155]" pageId="2" pageNumber="17">
<taxonomicName id="4C714D72FFA95D2FC4C3FBCE5312B2F9" box="[151,525,1129,1155]" class="Magnoliopsida" family="Loasaceae" genus="Actenosigynes" kingdom="Plantae" order="Cornales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="mantiqueirensis" status="sp. nov.">
<taxonomicName id="4C714D72FFA95D2FC4C3FBCE5312B2F9" box="[151,525,1129,1155]" class="Insecta" family="Colletidae" genus="Actenosigynes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="mantiqueirensis" status="sp. nov.">
<emphasis id="B905EAE3FFA95D2FC4C3FBCE5312B2F9" bold="true" box="[151,525,1129,1155]" italics="true" pageId="2" pageNumber="17">Actenosigynes mantiqueirensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2365798FFA95D2FC640FBCF536EB2F8" box="[532,625,1128,1154]" pageId="2" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
@ -45,29 +46,29 @@
</paragraph>
<paragraph id="8BCE36F1FFA95D2FC4C3FB2B505DB2DC" blockId="2.[151,625,1128,1190]" box="[151,322,1164,1190]" pageId="2" pageNumber="17">
(
<figureCitation id="134A2A74FFA95D2FC4CBFB2B51EEB2DC" box="[159,241,1164,1190]" captionStart="FIGURE 2" captionStartId="4.[151,255,1965,1989]" captionTargetBox="[263,1326,215,1953]" captionTargetId="figure@4.[260,1327,194,1953]" captionTargetPageId="4" captionText="FIGURE 2. Female of Actenosigynes mantiqueirensis sp. nov., holotype— a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the shiny integument of sternal discs); d) habitus, lateral view." httpUri="https://zenodo.org/record/191459/files/figure.png" pageId="2" pageNumber="17">Figs. 2</figureCitation>
<figureCitation id="134A2A74FFA95D2FC4CBFB2B51EEB2DC" box="[159,241,1164,1190]" captionStart="FIGURE 2" captionStartId="4.[151,255,1965,1989]" captionTargetBox="[263,1326,215,1953]" captionTargetId="figure@4.[260,1327,194,1953]" captionTargetPageId="4" captionText="FIGURE 2. Female of Actenosigynes mantiqueirensis sp. nov., holotype— a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the shiny integument of sternal discs); d) habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.191459" httpUri="https://zenodo.org/record/191459/files/figure.png" pageId="2" pageNumber="17">Figs. 2</figureCitation>
ad, 3)
</paragraph>
</subSubSection>
<subSubSection id="C36B657AFFA95D2FC4C3FB685321B3D1" pageId="2" pageNumber="17" type="diagnosis">
<paragraph id="8BCE36F1FFA95D2FC4C3FB6851FBB3FE" blockId="2.[151,1438,1231,2033]" pageId="2" pageNumber="17">
<emphasis id="B905EAE3FFA95D2FC4C3FB685004B293" bold="true" box="[151,283,1231,1257]" pageId="2" pageNumber="17">Diagnosis.</emphasis>
<taxonomicName id="4C714D72FFA95D2FC577FB775386B293" box="[291,665,1232,1257]" class="Magnoliopsida" family="Loasaceae" genus="Actenosigynes" kingdom="Plantae" order="Cornales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="mantiqueirensis">
<taxonomicName id="4C714D72FFA95D2FC577FB775386B293" box="[291,665,1232,1257]" class="Insecta" family="Colletidae" genus="Actenosigynes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="mantiqueirensis">
<emphasis id="B905EAE3FFA95D2FC577FB775386B293" box="[291,665,1232,1257]" italics="true" pageId="2" pageNumber="17">Actenosigynes mantiqueirensis</emphasis>
</taxonomicName>
can be readily distinguished from the only other species in the genus by its entirely black-pubescent mesosoma (
<figureCitation id="134A2A74FFA95D2FC68AFB51523AB36A" box="[734,805,1270,1296]" captionStart="FIGURE 2" captionStartId="4.[151,255,1965,1989]" captionTargetBox="[263,1326,215,1953]" captionTargetId="figure@4.[260,1327,194,1953]" captionTargetPageId="4" captionText="FIGURE 2. Female of Actenosigynes mantiqueirensis sp. nov., holotype— a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the shiny integument of sternal discs); d) habitus, lateral view." httpUri="https://zenodo.org/record/191459/files/figure.png" pageId="2" pageNumber="17">Fig. 2</figureCitation>
<figureCitation id="134A2A74FFA95D2FC68AFB51523AB36A" box="[734,805,1270,1296]" captionStart="FIGURE 2" captionStartId="4.[151,255,1965,1989]" captionTargetBox="[263,1326,215,1953]" captionTargetId="figure@4.[260,1327,194,1953]" captionTargetPageId="4" captionText="FIGURE 2. Female of Actenosigynes mantiqueirensis sp. nov., holotype— a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the shiny integument of sternal discs); d) habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.191459" httpUri="https://zenodo.org/record/191459/files/figure.png" pageId="2" pageNumber="17">Fig. 2</figureCitation>
a,d) (ferruginous dorsally in
<taxonomicName id="4C714D72FFA95D2FC020FB505412B36A" box="[1140,1293,1271,1296]" class="Magnoliopsida" family="Loasaceae" genus="Actenosigynes" kingdom="Plantae" order="Cornales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="fulvoniger">
<taxonomicName id="4C714D72FFA95D2FC020FB505412B36A" box="[1140,1293,1271,1296]" class="Insecta" family="Colletidae" genus="Actenosigynes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="fulvoniger">
<emphasis id="B905EAE3FFA95D2FC020FB505412B36A" box="[1140,1293,1271,1296]" italics="true" pageId="2" pageNumber="17">A. fulvoniger</emphasis>
</taxonomicName>
<figureCitation id="134A2A74FFA95D2FC179FB515473B36A" box="[1325,1388,1270,1296]" captionStart="FIGURE 1" captionStartId="3.[151,256,1974,1998]" captionTargetBox="[181,1402,195,1932]" captionTargetId="figure@3.[181,1403,195,1949]" captionTargetPageId="3" captionText="FIGURE 1. Female of Actenosigynes fulvoniger (Michener, 1989) — a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the dull integument of sternal discs); d) habitus, lateral view." httpUri="https://zenodo.org/record/191458/files/figure.png" pageId="2" pageNumber="17">Fig. 1</figureCitation>
<figureCitation id="134A2A74FFA95D2FC179FB515473B36A" box="[1325,1388,1270,1296]" captionStart="FIGURE 1" captionStartId="3.[151,256,1974,1998]" captionTargetBox="[181,1402,195,1932]" captionTargetId="figure@3.[181,1403,195,1949]" captionTargetPageId="3" captionText="FIGURE 1. Female of Actenosigynes fulvoniger (Michener, 1989) — a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the dull integument of sternal discs); d) habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.191458" httpUri="https://zenodo.org/record/191458/files/figure.png" pageId="2" pageNumber="17">Fig. 1</figureCitation>
a,d) and by the light-yellow pilosity on the terga (
<figureCitation id="134A2A74FFA95D2FC694FABA5215B34D" box="[704,778,1309,1335]" captionStart="FIGURE 2" captionStartId="4.[151,255,1965,1989]" captionTargetBox="[263,1326,215,1953]" captionTargetId="figure@4.[260,1327,194,1953]" captionTargetPageId="4" captionText="FIGURE 2. Female of Actenosigynes mantiqueirensis sp. nov., holotype— a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the shiny integument of sternal discs); d) habitus, lateral view." httpUri="https://zenodo.org/record/191459/files/figure.png" pageId="2" pageNumber="17">Fig. 2</figureCitation>
<figureCitation id="134A2A74FFA95D2FC694FABA5215B34D" box="[704,778,1309,1335]" captionStart="FIGURE 2" captionStartId="4.[151,255,1965,1989]" captionTargetBox="[263,1326,215,1953]" captionTargetId="figure@4.[260,1327,194,1953]" captionTargetPageId="4" captionText="FIGURE 2. Female of Actenosigynes mantiqueirensis sp. nov., holotype— a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the shiny integument of sternal discs); d) habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.191459" httpUri="https://zenodo.org/record/191459/files/figure.png" pageId="2" pageNumber="17">Fig. 2</figureCitation>
a,d) (entirely black in
<emphasis id="B905EAE3FFA95D2FC04CFABA55CDB34C" box="[1048,1234,1309,1334]" italics="true" pageId="2" pageNumber="17">A. fulvoniger—</emphasis>
<figureCitation id="134A2A74FFA95D2FC086FABA540AB34D" box="[1234,1301,1309,1335]" captionStart="FIGURE 1" captionStartId="3.[151,256,1974,1998]" captionTargetBox="[181,1402,195,1932]" captionTargetId="figure@3.[181,1403,195,1949]" captionTargetPageId="3" captionText="FIGURE 1. Female of Actenosigynes fulvoniger (Michener, 1989) — a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the dull integument of sternal discs); d) habitus, lateral view." httpUri="https://zenodo.org/record/191458/files/figure.png" pageId="2" pageNumber="17">Fig. 1</figureCitation>
<figureCitation id="134A2A74FFA95D2FC086FABA540AB34D" box="[1234,1301,1309,1335]" captionStart="FIGURE 1" captionStartId="3.[151,256,1974,1998]" captionTargetBox="[181,1402,195,1932]" captionTargetId="figure@3.[181,1403,195,1949]" captionTargetPageId="3" captionText="FIGURE 1. Female of Actenosigynes fulvoniger (Michener, 1989) — a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the dull integument of sternal discs); d) habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.191458" httpUri="https://zenodo.org/record/191458/files/figure.png" pageId="2" pageNumber="17">Fig. 1</figureCitation>
a,d). Other characters distinguishing these species are given in the identification key and (in italics) in the description, below.
</paragraph>
<paragraph id="8BCE36F1FFA95D2FC492FA365321B3D1" blockId="2.[151,1438,1231,2033]" box="[198,574,1425,1451]" pageId="2" pageNumber="17">
@ -94,11 +95,11 @@ body black except apical half of mandible, dark ferruginous (interrupted by blac
except Sc+R, brown;
<emphasis id="B905EAE3FFA95D2FC0B4F81652B6B18B" italics="true" pageId="2" pageNumber="17">wing membrane hyaline with yellowish hue especially basally and on marginal cell.</emphasis>
</paragraph>
<caption id="DF0E6679FFA85D2EC4C3F811556BB189" httpUri="https://zenodo.org/record/191458/files/figure.png" pageId="3" pageNumber="18" targetBox="[181,1402,195,1932]" targetPageId="3">
<caption id="DF0E6679FFA85D2EC4C3F811556BB189" ID-DOI="http://doi.org/10.5281/zenodo.191458" httpUri="https://zenodo.org/record/191458/files/figure.png" pageId="3" pageNumber="18" targetBox="[181,1402,195,1932]" targetPageId="3">
<paragraph id="8BCE36F1FFA85D2EC4C3F811556BB189" blockId="3.[151,1436,1974,2035]" pageId="3" pageNumber="18">
<emphasis id="B905EAE3FFA85D2EC4C3F8115004B1B4" bold="true" box="[151,283,1974,1998]" pageId="3" pageNumber="18">FIGURE 1.</emphasis>
Female of
<taxonomicName id="4C714D72FFA85D2EC5CCF8115273B1B7" authority="Michener, 1989" authorityName="Michener" authorityYear="1989" box="[408,876,1974,1997]" class="Magnoliopsida" family="Loasaceae" genus="Actenosigynes" kingdom="Plantae" order="Cornales" pageId="3" pageNumber="18" phylum="Tracheophyta" rank="species" species="fulvoniger">
<taxonomicName id="4C714D72FFA85D2EC5CCF8115273B1B7" authority="Michener, 1989" authorityName="Michener" authorityYear="1989" box="[408,876,1974,1997]" class="Insecta" family="Colletidae" genus="Actenosigynes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="fulvoniger">
<emphasis id="B905EAE3FFA85D2EC5CCF81153B4B1B7" box="[408,683,1974,1997]" italics="true" pageId="3" pageNumber="18">Actenosigynes fulvoniger</emphasis>
(Michener, 1989)
</taxonomicName>
@ -113,11 +114,11 @@ metasoma, ventral view (notice the dull integument of sternal discs);
habitus, lateral view.
</paragraph>
</caption>
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<caption id="DF0E6679FFAF5D29C4C3F80A55BFB190" ID-DOI="http://doi.org/10.5281/zenodo.191459" httpUri="https://zenodo.org/record/191459/files/figure.png" pageId="4" pageNumber="19" targetBox="[263,1326,215,1953]" targetPageId="4">
<paragraph id="8BCE36F1FFAF5D29C4C3F80A55BFB190" blockId="4.[151,1436,1965,2027]" pageId="4" pageNumber="19">
<emphasis id="B905EAE3FFAF5D29C4C3F80A5006B1BF" bold="true" box="[151,281,1965,1989]" pageId="4" pageNumber="19">FIGURE 2.</emphasis>
Female of
<taxonomicName id="4C714D72FFAF5D29C5C5F80953C6B1BF" box="[401,729,1966,1989]" class="Magnoliopsida" family="Loasaceae" genus="Actenosigynes" kingdom="Plantae" order="Cornales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="mantiqueirensis" status="sp. nov.">
<taxonomicName id="4C714D72FFAF5D29C5C5F80953C6B1BF" box="[401,729,1966,1989]" class="Insecta" family="Colletidae" genus="Actenosigynes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="mantiqueirensis" status="sp. nov.">
<emphasis id="B905EAE3FFAF5D29C5C5F80953C6B1BF" box="[401,729,1966,1989]" italics="true" pageId="4" pageNumber="19">Actenosigynes mantiqueirensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2365798FFAF5D29C68BF80A522BB1BE" box="[735,820,1965,1988]" pageId="4" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
@ -174,7 +175,7 @@ Faz[enda]. Tião Loreano; APA Fernão Dias; S[erra]. Mantiqueira; 13441-39731”
<date id="FFCF1031FFAE5D28C582F989537FB032" box="[470,608,1582,1608]" pageId="5" pageNumber="20" value="2006-10-25">25/10/2006</date>
; F. A. Silveira” and “
<emphasis id="B905EAE3FFAE5D28C708F98855D1B032" box="[860,1230,1583,1608]" italics="true" pageId="5" pageNumber="20">
<taxonomicName id="4C714D72FFAE5D28C708F98855D7B032" box="[860,1224,1583,1608]" class="Magnoliopsida" family="Loasaceae" genus="Actenosigynes" kingdom="Plantae" order="Cornales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="mantiqueirensis">Actenosigynes mantiqueirensis</taxonomicName>
<taxonomicName id="4C714D72FFAE5D28C708F98855D7B032" box="[860,1224,1583,1608]" class="Insecta" family="Colletidae" genus="Actenosigynes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="mantiqueirensis">Actenosigynes mantiqueirensis</taxonomicName>
;
</emphasis>
Silveira sp. nov.;
@ -233,11 +234,11 @@ The arrangement of sternal pilosity on both species of
as dense, wide, marginal bands, and the way the pollen is held in the
<typeStatus id="54CA8853FFAD5D2BC74EFF19529DB6A2" box="[794,898,190,216]" pageId="6" pageNumber="21" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C714D72FFAD5D2BC7FFFF185595B6A2" box="[939,1162,191,216]" class="Magnoliopsida" family="Loasaceae" genus="Actenosigynes" kingdom="Plantae" order="Cornales" pageId="6" pageNumber="21" phylum="Tracheophyta" rank="species" species="mantiqueirensis">
<taxonomicName id="4C714D72FFAD5D2BC7FFFF185595B6A2" box="[939,1162,191,216]" class="Insecta" family="Colletidae" genus="Actenosigynes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="mantiqueirensis">
<emphasis id="B905EAE3FFAD5D2BC7FFFF185595B6A2" box="[939,1162,191,216]" italics="true" pageId="6" pageNumber="21">A. mantiqueirensis</emphasis>
</taxonomicName>
(
<figureCitation id="134A2A74FFAD5D2BC0CFFF1955FBB6A2" box="[1179,1252,190,216]" captionStart="FIGURE 2" captionStartId="4.[151,255,1965,1989]" captionTargetBox="[263,1326,215,1953]" captionTargetId="figure@4.[260,1327,194,1953]" captionTargetPageId="4" captionText="FIGURE 2. Female of Actenosigynes mantiqueirensis sp. nov., holotype— a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the shiny integument of sternal discs); d) habitus, lateral view." httpUri="https://zenodo.org/record/191459/files/figure.png" pageId="6" pageNumber="21">Fig. 2</figureCitation>
<figureCitation id="134A2A74FFAD5D2BC0CFFF1955FBB6A2" box="[1179,1252,190,216]" captionStart="FIGURE 2" captionStartId="4.[151,255,1965,1989]" captionTargetBox="[263,1326,215,1953]" captionTargetId="figure@4.[260,1327,194,1953]" captionTargetPageId="4" captionText="FIGURE 2. Female of Actenosigynes mantiqueirensis sp. nov., holotype— a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the shiny integument of sternal discs); d) habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.191459" httpUri="https://zenodo.org/record/191459/files/figure.png" pageId="6" pageNumber="21">Fig. 2</figureCitation>
c) suggests that females may actively use them to collect and/or carry pollen from flowers, despite the fact that the scopae in the hind leg appear to be normal, comparatively to other
<taxonomicName id="4C714D72FFAD5D2BC765FEAB52C5B75C" box="[817,986,268,294]" class="Insecta" family="Colletidae" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="subFamily" subFamily="Paracolletinae">Paracolletinae</taxonomicName>
, and fully functional.

View file

@ -1,41 +1,42 @@
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<taxonomicName id="4C714D72FFA95D2FC4C3FF3053BDB6CB" authority="Michener, 1989" authorityName="Michener" authorityYear="1989" box="[151,674,151,177]" class="Magnoliopsida" family="Loasaceae" genus="Actenosigynes" kingdom="Plantae" order="Cornales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="fulvoniger">
<taxonomicName id="4C714D72FFA95D2FC4C3FF3053BDB6CB" authority="Michener, 1989" authorityName="Michener" authorityYear="1989" box="[151,674,151,177]" class="Insecta" family="Colletidae" genus="Actenosigynes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="fulvoniger">
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<emphasis id="B905EAE3FFA95D2FC4C3FF3050D7B6CB" bold="true" box="[151,456,151,177]" italics="true" pageId="2" pageNumber="17">Actenosigynes fulvoniger</emphasis>
(
@ -47,7 +48,7 @@
</paragraph>
<paragraph id="8BCE36F1FFA95D2FC4C3FF1D505DB6AE" blockId="2.[151,674,151,212]" box="[151,322,186,212]" pageId="2" pageNumber="17">
(
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<figureCitation id="134A2A74FFA95D2FC4CBFF1D51F3B6AE" box="[159,236,186,212]" captionStart="FIGURE 1" captionStartId="3.[151,256,1974,1998]" captionTargetBox="[181,1402,195,1932]" captionTargetId="figure@3.[181,1403,195,1949]" captionTargetPageId="3" captionText="FIGURE 1. Female of Actenosigynes fulvoniger (Michener, 1989) — a) habitus, dorsal view; b) head, frontal view; c) metasoma, ventral view (notice the dull integument of sternal discs); d) habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.191458" httpUri="https://zenodo.org/record/191458/files/figure.png" pageId="2" pageNumber="17">Figs. 1</figureCitation>
ad, 3)
</paragraph>
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@ -67,7 +68,7 @@ ad, 3)
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<taxonomicName id="4C714D72FFA95D2FC4C3FE8F50B9B745" box="[151,422,296,319]" class="Insecta" family="Colletidae" genus="Actenosigynes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="fulvoniger">Actenosigynes fulvoniger</taxonomicName>
;
</emphasis>
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@ -142,7 +143,7 @@ is active during the rainy season, between October and December.
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<emphasis id="B905EAE3FFA95D2FC492FD64504BB4A7" bold="true" box="[198,340,707,733]" pageId="2" pageNumber="17">Comments.</emphasis>
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<emphasis id="B905EAE3FFA95D2FC50AFD635397B4A7" box="[350,648,708,733]" italics="true" pageId="2" pageNumber="17">Actenosigynes fulvoniger</emphasis>
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is only known from the southernmost, temperate Brazilian states (
@ -154,7 +155,7 @@ Moure
,
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). References to the biology of
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<emphasis id="B905EAE3FFA95D2FC760FD4D52CFB579" box="[820,976,746,771]" italics="true" pageId="2" pageNumber="17">A. fulvoniger</emphasis>
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in the literature are meager.

View file

@ -1,63 +1,64 @@
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<mods:title id="72A9F3AE614DE23984808D126655FD15">Systematics, phylogeny and biology of a new genus of Lithocolletinae (Lepidoptera: Gracillariidae) associated with Cistaceae</mods:title>
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Key to the species of
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<figureCitation id="FACE33B29D200473DB20663F7FFBBA1F" box="[843,917,783,805]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246619" httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="19" pageNumber="220">Figs 16</figureCitation>
,
<figureCitation id="FACE33B29D200473DBF4663F7FB3BA1F" box="[927,989,783,805]" captionStart="FIGURES 22 25" captionStartId="9.[151,264,1917,1940]" captionTargetBox="[172,1397,481,1900]" captionTargetId="figure@9.[172,1408,481,1900]" captionTargetPageId="9" captionText="FIGURES 22 25. Female genitalia of Triberta species. 22, Triberta cistifoliella, Greece, prep. TRB 3559 ♀. Scale bar 200 µm. 23, same preparation, signum on corpus bursae, prep. TRB 3559 ♀. Scale bar 100 µm. 24, Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. DP 3807 ♀. Scale bar 200 µm. 25, same preparation, signum on corpus bursae, prep. DP 3807 ♀. Scale bar 100 µm." httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="19" pageNumber="220">22, 23</figureCitation>
<figureCitation id="FACE33B29D200473DBF4663F7FB3BA1F" box="[927,989,783,805]" captionStart="FIGURES 22 25" captionStartId="9.[151,264,1917,1940]" captionTargetBox="[172,1397,481,1900]" captionTargetId="figure@9.[172,1408,481,1900]" captionTargetPageId="9" captionText="FIGURES 22 25. Female genitalia of Triberta species. 22, Triberta cistifoliella, Greece, prep. TRB 3559 ♀. Scale bar 200 µm. 23, same preparation, signum on corpus bursae, prep. TRB 3559 ♀. Scale bar 100 µm. 24, Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. DP 3807 ♀. Scale bar 200 µm. 25, same preparation, signum on corpus bursae, prep. DP 3807 ♀. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246620" httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="19" pageNumber="220">22, 23</figureCitation>
,
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).....................
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<figureCitation id="FACE33B29D200473DBCC66577857BA47" box="[935,1081,871,893]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="19" pageNumber="220">Figs 1719, 21</figureCitation>
<figureCitation id="FACE33B29D200473DBCC66577857BA47" box="[935,1081,871,893]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246619" httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="19" pageNumber="220">Figs 1719, 21</figureCitation>
; 24, 25, 61).....
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<taxonomicName id="A5F554B49D210472D8FC67787E80BB58" authority="Herrich-Schaffer, 1861" authorityName="Herrich-Schaffer" authorityYear="1861" box="[151,750,584,610]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="18" pageNumber="219" phylum="Tracheophyta" rank="species" species="helianthemella" status="comb. nov.">
<taxonomicName id="A5F554B49D210472D8FC67787E80BB58" ID-CoL="8VLV8" authority="Herrich-Schaffer, 1861" authorityName="Herrich-Schaffer" authorityYear="1861" box="[151,750,584,610]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="helianthemella" status="comb. nov.">
<emphasis id="5081F3259D210472D8FC67787DDBBB58" bold="true" box="[151,437,584,610]" italics="true" pageId="18" pageNumber="219">Triberta helianthemella</emphasis>
(Herrich-Schäffer, 1861)
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@ -168,11 +169,11 @@ and
and were collected and reared in Regensburg without indication which specimen is the name bearing
<typeStatus id="BD4E91959D210472DBC462087F8DBE6A" box="[943,995,1848,1872]" pageId="18" pageNumber="219">type</typeStatus>
. We believe that these five specimens were originally collected by Gottlieb August Wilhelm Herrich-Schäffer and sold to Otto Staudinger (
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<figureCitation id="FACE33B29D210472DD66626C79E1BE4F" box="[1293,1423,1884,1909]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246618" httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="18" pageNumber="219">Figs 1115</figureCitation>
). The phenotype of the male specimen bearing the label of the genitalia slide TRB3935♂ most closely resembles the original description. Therefore, we designate it as the
<typeStatus id="BD4E91959D210472DA9962947F36BE87" box="[754,856,1956,1981]" pageId="18" pageNumber="219" type="lectotype">lectotype</typeStatus>
(
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<figureCitation id="FACE33B29D210472DB0362947FD7BE86" box="[872,953,1956,1981]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246618" httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="18" pageNumber="219">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="624A2F379D200473D8FC65A87CB9B9EF" blockId="19.[151,1436,151,213]" pageId="19" pageNumber="220">
@ -187,9 +188,9 @@ Herrich-Schäffer, 1861
</paragraph>
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♂, [1]Rgsbg.[handwritten]; [2]ex coll. Staudinger; [3] genitalia slide TRB3935♂, in ZMHB (
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<figureCitation id="FACE33B29D200473DCBA64347949B826" box="[1233,1319,260,285]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246618" httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="19" pageNumber="220">Figs 11</figureCitation>
,
<figureCitation id="FACE33B29D200473DD586434793CB826" box="[1331,1362,260,284]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="19" pageNumber="220">17</figureCitation>
<figureCitation id="FACE33B29D200473DD586434793CB826" box="[1331,1362,260,284]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246619" httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="19" pageNumber="220">17</figureCitation>
).
<typeStatus id="BD4E91959D200473D8FC64177D51B87A" box="[151,319,295,320]" pageId="19" pageNumber="220" type="paralectotype">
<emphasis id="5081F3259D200473D8FC64177D51B87A" box="[151,319,295,320]" italics="true" pageId="19" pageNumber="220">Paralectotypes</emphasis>
@ -200,11 +201,11 @@ Herrich-Schäffer, 1861
</paragraph>
<paragraph id="624A2F379D200473D8FC647D786DB8B2" blockId="19.[151,1436,260,536]" pageId="19" pageNumber="220">
♂, [1] Regensburg [handwritten]; [2] ex coll. Staudinger; [3] genitalia slide TRB3945♂, in ZMHB (
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<figureCitation id="FACE33B29D200473DD7E647C7905B85E" box="[1301,1387,332,357]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246618" httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="19" pageNumber="220">Figs 12</figureCitation>
,
<figureCitation id="FACE33B29D200473DD1D647C79FDB85E" box="[1398,1427,332,356]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="19" pageNumber="220">18</figureCitation>
<figureCitation id="FACE33B29D200473DD1D647C79FDB85E" box="[1398,1427,332,356]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246619" httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="19" pageNumber="220">18</figureCitation>
) ♂, [1] Regensb. [handwritten]; [2] ex coll. Staudinger, in ZMHB (
<figureCitation id="FACE33B29D200473DBC9645F7F9BB8B2" box="[930,1013,367,392]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="19" pageNumber="220">Fig. 13</figureCitation>
<figureCitation id="FACE33B29D200473DBC9645F7F9BB8B2" box="[930,1013,367,392]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246618" httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="19" pageNumber="220">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
@ -214,14 +215,14 @@ Herrich-Schäffer, 1861
the sub-triangular shape of the distal half of valva in the male genitalia is variable within the
<typeStatus id="BD4E91959D200473DD4E64A5793BB897" box="[1317,1365,405,429]" pageId="19" pageNumber="220">type</typeStatus>
series (
<figureCitation id="FACE33B29D200473D8F464877D4EB8EA" box="[159,288,439,464]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="19" pageNumber="220">Figs 17, 18</figureCitation>
<figureCitation id="FACE33B29D200473D8F464877D4EB8EA" box="[159,288,439,464]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246619" httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="19" pageNumber="220">Figs 17, 18</figureCitation>
).
</paragraph>
<paragraph id="624A2F379D200473D8FC64ED78B7BB22" blockId="19.[151,1436,260,536]" pageId="19" pageNumber="220">
♀, [1] Regensburg [handwritten]; [2] ex coll. Staudinger, in ZMHB (
<figureCitation id="FACE33B29D200473DBAB64EC787DB8CE" box="[960,1043,476,501]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="19" pageNumber="220">Fig. 14</figureCitation>
<figureCitation id="FACE33B29D200473DBAB64EC787DB8CE" box="[960,1043,476,501]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246618" httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="19" pageNumber="220">Fig. 14</figureCitation>
). ♀, [1] Regensburg [handwritten]; [2] ex coll. Staudinger, [3] Coll. Led., in ZMHB (
<figureCitation id="FACE33B29D200473DC1264CF78A2BB22" box="[1145,1228,511,536]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="19" pageNumber="220">Fig. 15</figureCitation>
<figureCitation id="FACE33B29D200473DC1264CF78A2BB22" box="[1145,1228,511,536]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246618" httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="19" pageNumber="220">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,70 +1,71 @@
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<taxonomicName id="A5F554B49D230470D8FC63AF7E0CBF83" authority="Groschke, 1944" authorityName="Groschke" authorityYear="1944" box="[151,610,1695,1721]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="217" phylum="Tracheophyta" rank="species" species="cistifoliella" status="comb. nov.">
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(Groschke, 1944)
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@ -259,7 +260,7 @@ were lost during the last years of the second world war or the years after the w
under the conditions specified by the ICZN Art. 75: (1) the
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is designated with the express purpose to clarify the taxonomic status of closely related species within the new genus
<taxonomicName id="A5F554B49D220471DC7B634D7802BFAC" box="[1040,1132,1661,1686]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="17" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
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<emphasis id="5081F3259D220471DC7B634D7802BFAC" box="[1040,1132,1661,1686]" italics="true" pageId="17" pageNumber="218">Triberta</emphasis>
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and to restore the species
@ -320,9 +321,9 @@ Groschke, 1944
♂, [1] Sicilia, Palermo, S. Martino d. Scale,
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, J. Klimesch; [2] genitalia slide TRB3554♂, in the ZSM (
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,
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<figureCitation id="FACE33B29D210472D929645F7D0FB8B2" box="[322,353,367,392]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246619" httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="18" pageNumber="219">16</figureCitation>
).
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<treatment id="EA5C9E219D300470D8FC677579E8BF2A" ID-DOI="http://doi.org/10.5281/zenodo.6157653" ID-GBIF-Taxon="119583676" ID-Zenodo-Dep="6157653" LSID="urn:lsid:plazi:treatment:EA5C9E219D300470D8FC677579E8BF2A" httpUri="http://treatment.plazi.org/id/EA5C9E219D300470D8FC677579E8BF2A" lastPageId="16" lastPageNumber="217" pageId="3" pageNumber="204">
<subSubSection id="2AEF7CBC9D300463D8FC6775796BBA15" pageId="3" pageNumber="204" type="nomenclature">
<paragraph id="624A2F379D300463D8FC67757D8FBB65" blockId="3.[151,481,581,607]" box="[151,481,581,607]" pageId="3" pageNumber="204">
<heading id="3902985B9D300463D8FC67757D8FBB65" bold="true" box="[151,481,581,607]" fontSize="11" level="1" pageId="3" pageNumber="204" reason="1">
<emphasis id="5081F3259D300463D8FC67757D8FBB65" bold="true" box="[151,481,581,607]" pageId="3" pageNumber="204">
<taxonomicName id="A5F554B49D300463D8FC67757D02BB65" authority="De Prins" authorityName="De Prins" box="[151,364,581,607]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="204" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463D8FC67757D02BB65" ID-CoL="8VL84" authority="De Prins" authorityName="De Prins" box="[151,364,581,607]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463D8FC67757C97BB65" bold="true" box="[151,249,581,607]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
De Prins
</taxonomicName>
@ -82,7 +83,7 @@ Groschke, 1944
, by present designation.
</paragraph>
<paragraph id="624A2F379D300463D8FC67FE796BBA15" blockId="3.[151,1436,718,2009]" pageId="3" pageNumber="204">
<taxonomicName id="A5F554B49D300463D8FC67FE7C9DBBDD" box="[151,243,718,743]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463D8FC67FE7C9DBBDD" box="[151,243,718,743]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463D8FC67FE7C9DBBDD" box="[151,243,718,743]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
is assigned to the subfamily
@ -94,7 +95,7 @@ on the basis of eleven nuclear genes and the following putative morphological sy
<paragraph id="624A2F379D300464D8AC660A7F6FBD6D" blockId="3.[151,1436,718,2009]" lastBlockId="4.[151,1436,151,2025]" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204">
<emphasis id="5081F3259D300463D8AC660A7D54BA69" bold="true" box="[199,314,826,851]" pageId="3" pageNumber="204">Diagnosis</emphasis>
. According to the phenotypic, biological and ecological characters
<taxonomicName id="A5F554B49D300463DC5E660A78FFBA69" box="[1077,1169,826,851]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463DC5E660A78FFBA69" box="[1077,1169,826,851]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463DC5E660A78FFBA69" box="[1077,1169,826,851]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
resembles other genera of
@ -105,11 +106,11 @@ resembles other genera of
Braun, 1929
</taxonomicName>
. We define
<taxonomicName id="A5F554B49D300463DA1D66B27EBCBAA1" box="[630,722,898,923]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463DA1D66B27EBCBAA1" box="[630,722,898,923]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463DA1D66B27EBCBAA1" box="[630,722,898,923]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
as a genus-group taxon of the clade
<taxonomicName id="A5F554B49D300463DC1F66B278BEBAA1" box="[1140,1232,898,923]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463DC1F66B278BEBAA1" box="[1140,1232,898,923]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463DC1F66B278BEBAA1" box="[1140,1232,898,923]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
+
@ -125,7 +126,7 @@ as a genus-group taxon of the clade
<emphasis id="5081F3259D300463D92E66967DAEBA85" box="[325,448,934,959]" italics="true" pageId="3" pageNumber="204">Chrysaster</emphasis>
</taxonomicName>
) (Fig. 01). The ecological trait defining
<taxonomicName id="A5F554B49D300463DBEA66967FB3BA85" box="[897,989,934,959]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463DBEA66967FB3BA85" box="[897,989,934,959]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463DBEA66967FB3BA85" box="[897,989,934,959]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
is that pupation occurs outside the mine (a trait shared with
@ -137,7 +138,7 @@ and
<emphasis id="5081F3259D300463DA4E66FA7EC0BAD9" box="[549,686,970,995]" italics="true" pageId="3" pageNumber="204">Leucanthiza</emphasis>
</taxonomicName>
) in a transparent oval-shaped and slightly raised cocoon (
<figureCitation id="FACE33B29D300463DD5066FA79FEBAD9" box="[1339,1424,970,995]" captionStart="FIGURES 56 59" captionStartId="15.[151,264,1478,1501]" captionTargetBox="[151,1436,373,1456]" captionTargetId="figure@15.[151,1436,373,1457]" captionTargetPageId="15" captionText="FIGURES 56 59. Biology of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 56, host plant Helianthemum nummularium. 57, mine, adaxial view. 58, excrements of larva in mine, adaxial view. 59, transparent cocoon outside the mine. Scale bar 1 mm." httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="3" pageNumber="204">Fig. 59</figureCitation>
<figureCitation id="FACE33B29D300463DD5066FA79FEBAD9" box="[1339,1424,970,995]" captionStart="FIGURES 56 59" captionStartId="15.[151,264,1478,1501]" captionTargetBox="[151,1436,373,1456]" captionTargetId="figure@15.[151,1436,373,1457]" captionTargetPageId="15" captionText="FIGURES 56 59. Biology of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 56, host plant Helianthemum nummularium. 57, mine, adaxial view. 58, excrements of larva in mine, adaxial view. 59, transparent cocoon outside the mine. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246625" httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="3" pageNumber="204">Fig. 59</figureCitation>
). In contrast the pupation of
<taxonomicName id="A5F554B49D300463D9AE66DF7E2EBD3C" box="[453,576,1007,1030]" class="Insecta" family="Gracillariidae" genus="Cameraria" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="5081F3259D300463D9AE66DF7E2EBD3C" box="[453,576,1007,1030]" italics="true" pageId="3" pageNumber="204">Cameraria</emphasis>
@ -155,7 +156,7 @@ spins an ellipsoid silken cocoon at the edge of a contracted leaf (Kumata 1961)
<emphasis id="5081F3259D300463D8FC616A7D4EBD49" box="[151,288,1114,1139]" italics="true" pageId="3" pageNumber="204">Leucanthiza</emphasis>
</taxonomicName>
weaves its cocoon on some substance on the surface of the ground (Clemens 1859). The shape and structure of this cocoon spinning clearly separates the new genus
<taxonomicName id="A5F554B49D300463DBE2614E7F8BBDAD" box="[905,997,1150,1175]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463DBE2614E7F8BBDAD" box="[905,997,1150,1175]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463DBE2614E7F8BBDAD" box="[905,997,1150,1175]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
from the latter two genera
@ -167,11 +168,11 @@ and
<emphasis id="5081F3259D300463D8A261927D3CBD81" box="[201,338,1186,1211]" italics="true" pageId="3" pageNumber="204">Leucanthiza</emphasis>
</taxonomicName>
. In addition to the ecological and molecular characters,
<taxonomicName id="A5F554B49D300463DBB561927854BD81" box="[990,1082,1186,1211]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463DBB561927854BD81" box="[990,1082,1186,1211]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463DBB561927854BD81" box="[990,1082,1186,1211]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
may be distinguished from all remaining lithocolletine genera by the combination of the following morphological diagnostic traits: the forewing pattern with three white fasciae or chevrons strongly angulated at midline, or rows edged basally and apically (a trait easily differentiating
<taxonomicName id="A5F554B49D300463D9A2603E7E4BBC1D" box="[457,549,1294,1319]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463D9A2603E7E4BBC1D" box="[457,549,1294,1319]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463D9A2603E7E4BBC1D" box="[457,549,1294,1319]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
from
@ -193,7 +194,7 @@ and
<emphasis id="5081F3259D300463D8FC60667D4EBC55" box="[151,288,1366,1391]" italics="true" pageId="3" pageNumber="204">Leucanthiza</emphasis>
</taxonomicName>
where vertex is smooth (
<figureCitation id="FACE33B29D300463DA5260667ED4BC55" box="[569,698,1366,1391]" captionStart="FIGURES 60 61" captionStartId="16.[151,264,858,881]" captionTargetBox="[151,1436,193,837]" captionTargetId="figure@16.[151,1436,193,837]" captionTargetPageId="16" captionText="FIGURES 60 61. Diagnostic characters in Triberta head vestiture of adults. 60, vertex of the neotype of Lithocolletis cistifoliella Groschke, 1944. 61, vertex of the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861. Scale bar 100 µm." httpUri="https://zenodo.org/record/246626/files/figure.png" pageId="3" pageNumber="204">Figs 60, 61</figureCitation>
<figureCitation id="FACE33B29D300463DA5260667ED4BC55" box="[569,698,1366,1391]" captionStart="FIGURES 60 61" captionStartId="16.[151,264,858,881]" captionTargetBox="[151,1436,193,837]" captionTargetId="figure@16.[151,1436,193,837]" captionTargetPageId="16" captionText="FIGURES 60 61. Diagnostic characters in Triberta head vestiture of adults. 60, vertex of the neotype of Lithocolletis cistifoliella Groschke, 1944. 61, vertex of the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246626" httpUri="https://zenodo.org/record/246626/files/figure.png" pageId="3" pageNumber="204">Figs 60, 61</figureCitation>
). Wing venation is similar in that of
<taxonomicName id="A5F554B49D300463DC396066797BBC55" box="[1106,1301,1366,1391]" class="Insecta" family="Gracillariidae" genus="Protolithocolletis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="5081F3259D300463DC396066797BBC55" box="[1106,1301,1366,1391]" italics="true" pageId="3" pageNumber="204">Protolithocolletis</emphasis>
@ -264,7 +265,7 @@ and
,
</emphasis>
a trait easily differentiating
<taxonomicName id="A5F554B49D300463DD2B630479F2BF77" box="[1344,1436,1588,1613]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463DD2B630479F2BF77" box="[1344,1436,1588,1613]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463DD2B630479F2BF77" box="[1344,1436,1588,1613]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
from the latter genera. In
@ -272,7 +273,7 @@ from the latter genera. In
<emphasis id="5081F3259D300463D9D063697E58BF4A" box="[443,566,1625,1648]" italics="true" pageId="3" pageNumber="204">Cameraria</emphasis>
</taxonomicName>
sternum VIII often bifid caudally, while in
<taxonomicName id="A5F554B49D300463DC42636878EBBF4B" box="[1065,1157,1624,1649]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463DC42636878EBBF4B" box="[1065,1157,1624,1649]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463DC42636878EBBF4B" box="[1065,1157,1624,1649]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
it is shaped like a broad triangle (Figs 0 4, 0 6, 21). Uncus fused with tegumen to form a single sclerite (Kuznetzov 1981) (Fig. 08). The apical part of tegumen in the male genitalia with two apical setae like in
@ -284,7 +285,7 @@ it is shaped like a broad triangle (Figs 0 4, 0 6, 21). Uncus fused with tegumen
<emphasis id="5081F3259D300463DB5B63F47FB8BFE7" box="[816,982,1732,1757]" italics="true" pageId="3" pageNumber="204">Phyllonorycter</emphasis>
</taxonomicName>
. Ventral surface of valvae in
<taxonomicName id="A5F554B49D300463DD2B63F479F2BFE7" box="[1344,1436,1732,1757]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463DD2B63F479F2BFE7" box="[1344,1436,1732,1757]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463DD2B63F479F2BFE7" box="[1344,1436,1732,1757]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
covered with long hair-like setae (Fig. 06), anellus strongly developed, tubular in many species of
@ -308,11 +309,11 @@ and
<emphasis id="5081F3259D300463DBC8623C7808BE1F" box="[931,1126,1804,1829]" italics="true" pageId="3" pageNumber="204">Protolithocolletis</emphasis>
</taxonomicName>
saccus in being simple, not prominent; aedoeagus is rather short, ca. 2/3 of valval length, bifurcate at coecum, a trait easily differentiating
<taxonomicName id="A5F554B49D300463D8FC62647C9DBE57" box="[151,243,1876,1901]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463D8FC62647C9DBE57" box="[151,243,1876,1901]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463D8FC62647C9DBE57" box="[151,243,1876,1901]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
from all remaining lithocolletine genera; transtilla in
<taxonomicName id="A5F554B49D300463DB0262647FABBE57" box="[873,965,1876,1901]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D300463DB0262647FABBE57" box="[873,965,1876,1901]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D300463DB0262647FABBE57" box="[873,965,1876,1901]" italics="true" pageId="3" pageNumber="204">Triberta</emphasis>
</taxonomicName>
incomplete like in the great majority of
@ -324,7 +325,7 @@ species and
<emphasis id="5081F3259D300463D9CC62487E76BEAB" box="[423,536,1912,1937]" italics="true" pageId="3" pageNumber="204">Hyloconis</emphasis>
</taxonomicName>
(Figs 0609, 1619). In the female genitalia (
<figureCitation id="FACE33B29D300463DC45624878DCBEAB" box="[1070,1202,1912,1937]" captionStart="FIGURES 22 25" captionStartId="9.[151,264,1917,1940]" captionTargetBox="[172,1397,481,1900]" captionTargetId="figure@9.[172,1408,481,1900]" captionTargetPageId="9" captionText="FIGURES 22 25. Female genitalia of Triberta species. 22, Triberta cistifoliella, Greece, prep. TRB 3559 ♀. Scale bar 200 µm. 23, same preparation, signum on corpus bursae, prep. TRB 3559 ♀. Scale bar 100 µm. 24, Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. DP 3807 ♀. Scale bar 200 µm. 25, same preparation, signum on corpus bursae, prep. DP 3807 ♀. Scale bar 100 µm." httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="3" pageNumber="204">Figs 2225</figureCitation>
<figureCitation id="FACE33B29D300463DC45624878DCBEAB" box="[1070,1202,1912,1937]" captionStart="FIGURES 22 25" captionStartId="9.[151,264,1917,1940]" captionTargetBox="[172,1397,481,1900]" captionTargetId="figure@9.[172,1408,481,1900]" captionTargetPageId="9" captionText="FIGURES 22 25. Female genitalia of Triberta species. 22, Triberta cistifoliella, Greece, prep. TRB 3559 ♀. Scale bar 200 µm. 23, same preparation, signum on corpus bursae, prep. TRB 3559 ♀. Scale bar 100 µm. 24, Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. DP 3807 ♀. Scale bar 200 µm. 25, same preparation, signum on corpus bursae, prep. DP 3807 ♀. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246620" httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="3" pageNumber="204">Figs 2225</figureCitation>
), papillae anales are semi-circular, not flattened like as in
<taxonomicName id="A5F554B49D300463DA2B62AD7ED5BE8E" box="[576,699,1949,1972]" class="Insecta" family="Gracillariidae" genus="Cameraria" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="5081F3259D300463DA2B62AD7ED5BE8E" box="[576,699,1949,1972]" italics="true" pageId="3" pageNumber="204">Cameraria</emphasis>
@ -383,9 +384,9 @@ and
<emphasis id="5081F3259D370464D908645F7E7AB8B2" box="[355,532,367,392]" italics="true" pageId="4" pageNumber="205">Neolithocolletis</emphasis>
</taxonomicName>
, where the signum area consists of numerous microscopic spicules scattered on the wall of corpus bursae or arranged in linear series on subcaudal part of corpus bursae. Larva constructs an abaxial mine (
<figureCitation id="FACE33B29D370464D92A64877DA5B8EA" box="[321,459,439,464]" captionStart="FIGURES 56 59" captionStartId="15.[151,264,1478,1501]" captionTargetBox="[151,1436,373,1456]" captionTargetId="figure@15.[151,1436,373,1457]" captionTargetPageId="15" captionText="FIGURES 56 59. Biology of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 56, host plant Helianthemum nummularium. 57, mine, adaxial view. 58, excrements of larva in mine, adaxial view. 59, transparent cocoon outside the mine. Scale bar 1 mm." httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="4" pageNumber="205">Figs 5658</figureCitation>
<figureCitation id="FACE33B29D370464D92A64877DA5B8EA" box="[321,459,439,464]" captionStart="FIGURES 56 59" captionStartId="15.[151,264,1478,1501]" captionTargetBox="[151,1436,373,1456]" captionTargetId="figure@15.[151,1436,373,1457]" captionTargetPageId="15" captionText="FIGURES 56 59. Biology of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 56, host plant Helianthemum nummularium. 57, mine, adaxial view. 58, excrements of larva in mine, adaxial view. 59, transparent cocoon outside the mine. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246625" httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="4" pageNumber="205">Figs 5658</figureCitation>
). Larva of
<taxonomicName id="A5F554B49D370464DA3D64877EDCB8EA" box="[598,690,439,464]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D370464DA3D64877EDCB8EA" box="[598,690,439,464]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D370464DA3D64877EDCB8EA" box="[598,690,439,464]" italics="true" pageId="4" pageNumber="205">Triberta</emphasis>
</taxonomicName>
is of cylindrical-
@ -410,13 +411,13 @@ group (Chambers 1878; Chapman 1902; Braun 1908). As was noted by Stehr (1987) in
bears a transverse series of six, relatively large, spinose, digitate lobes (Davis
<emphasis id="5081F3259D370464DD75675D7920BBBF" box="[1310,1358,620,645]" italics="true" pageId="4" pageNumber="205">et al</emphasis>
. 2013:
<figureCitation id="FACE33B29D370464D8FC67BF7D79BB92" box="[151,279,655,680]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="4" pageNumber="205">Figs 1315</figureCitation>
<figureCitation id="FACE33B29D370464D8FC67BF7D79BB92" box="[151,279,655,680]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246618" httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="4" pageNumber="205">Figs 1315</figureCitation>
). The hypopharynx of tissue-feeding
<taxonomicName id="A5F554B49D370464DAD767BF7F0ABB92" box="[700,868,655,680]" class="Insecta" family="Gracillariidae" genus="Phyllonorycter" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="5081F3259D370464DAD767BF7F0ABB92" box="[700,868,655,680]" italics="true" pageId="4" pageNumber="205">Phyllonorycter</emphasis>
</taxonomicName>
larvae possesses a series of over 20 much smaller, curved, smooth spines (Davis &amp; Deschka 2001:
<figureCitation id="FACE33B29D370464DAB967847F36BBF7" box="[722,856,692,717]" captionStart="FIGURES 56 59" captionStartId="15.[151,264,1478,1501]" captionTargetBox="[151,1436,373,1456]" captionTargetId="figure@15.[151,1436,373,1457]" captionTargetPageId="15" captionText="FIGURES 56 59. Biology of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 56, host plant Helianthemum nummularium. 57, mine, adaxial view. 58, excrements of larva in mine, adaxial view. 59, transparent cocoon outside the mine. Scale bar 1 mm." httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="4" pageNumber="205">Figs 5658</figureCitation>
<figureCitation id="FACE33B29D370464DAB967847F36BBF7" box="[722,856,692,717]" captionStart="FIGURES 56 59" captionStartId="15.[151,264,1478,1501]" captionTargetBox="[151,1436,373,1456]" captionTargetId="figure@15.[151,1436,373,1457]" captionTargetPageId="15" captionText="FIGURES 56 59. Biology of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 56, host plant Helianthemum nummularium. 57, mine, adaxial view. 58, excrements of larva in mine, adaxial view. 59, transparent cocoon outside the mine. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246625" httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="4" pageNumber="205">Figs 5658</figureCitation>
). The hypopharynx of the two non-feeding larval instars in
<taxonomicName id="A5F554B49D370464D96867E87D10BBD5" box="[259,382,728,751]" class="Insecta" family="Gracillariidae" genus="Cameraria" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="5081F3259D370464D96867E87D10BBD5" box="[259,382,728,751]" italics="true" pageId="4" pageNumber="205">Cameraria</emphasis>
@ -424,11 +425,11 @@ larvae possesses a series of over 20 much smaller, curved, smooth spines (Davis
is mostly smooth and without anterior spines (De Prins
<emphasis id="5081F3259D370464DB8667E97870BBCA" box="[1005,1054,727,752]" italics="true" pageId="4" pageNumber="205">et al</emphasis>
. 2003:
<figureCitation id="FACE33B29D370464DC1A67E778ADBBCA" box="[1137,1219,727,752]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="4" pageNumber="205">Fig. 10</figureCitation>
<figureCitation id="FACE33B29D370464DC1A67E778ADBBCA" box="[1137,1219,727,752]" captionStart="FIGURES 10 15" captionStartId="7.[151,264,1564,1587]" captionTargetBox="[151,1413,178,1538]" captionTargetId="figure@7.[151,1436,155,1538]" captionTargetPageId="7" captionText="FIGURES 10 15. The primary types of Triberta species. 10, the neotype of Lithocolletis cistifoliella Groschke, 1944, male, in ZSM. 11, the lectotype of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 12, the paralectotype 1 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 13, the paralectotype 2 of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, in ZMHB. 14, the paralectotype 3 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. 15, the paralectotype 4 of Lithocolletis helianthemella Herrich-Schäffer, 1861, female, in ZMHB. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246618" httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="4" pageNumber="205">Fig. 10</figureCitation>
; Davis
<emphasis id="5081F3259D370464DD7D67E97928BBCA" box="[1302,1350,727,752]" italics="true" pageId="4" pageNumber="205">et al</emphasis>
. 2013). The ventral surface of the last instar larva of
<taxonomicName id="A5F554B49D370464DAB667CC7F57BA2F" box="[733,825,764,789]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D370464DAB667CC7F57BA2F" box="[733,825,764,789]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D370464DAB667CC7F57BA2F" box="[733,825,764,789]" italics="true" pageId="4" pageNumber="205">Triberta</emphasis>
</taxonomicName>
is covered with hypopharyngeal striations, which differentiates the latter genus from
@ -444,7 +445,7 @@ and
<emphasis id="5081F3259D370464DBD4662F780BBA02" box="[959,1125,799,824]" italics="true" pageId="4" pageNumber="205">Phyllonorycter</emphasis>
</taxonomicName>
. Pupa without cremaster in
<taxonomicName id="A5F554B49D370464D8FC66747C9DBA67" box="[151,243,836,861]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D370464D8FC66747C9DBA67" box="[151,243,836,861]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D370464D8FC66747C9DBA67" box="[151,243,836,861]" italics="true" pageId="4" pageNumber="205">Triberta</emphasis>
</taxonomicName>
and remaining eight lithocolletine genera, except in
@ -466,7 +467,7 @@ and reduced, consisting of four little hooks, in
</taxonomicName>
. A thickened sclerotized ridge present along the lateral sides of pupal A18, in
<emphasis id="5081F3259D370464DC8166BC7922BA9F" box="[1258,1356,908,933]" italics="true" pageId="4" pageNumber="205">
<taxonomicName id="A5F554B49D370464DC8166BC7926BA9F" box="[1258,1352,908,933]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">Triberta</taxonomicName>
<taxonomicName id="A5F554B49D370464DC8166BC7926BA9F" box="[1258,1352,908,933]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">Triberta</taxonomicName>
,
</emphasis>
which differentiates the new genus from
@ -487,13 +488,13 @@ and
;
</emphasis>
and in a transparent oval-shaped and slightly raised cocoon. Members of the new genus
<taxonomicName id="A5F554B49D370464DD7D66C7791CBD2A" box="[1302,1394,1015,1040]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D370464DD7D66C7791CBD2A" box="[1302,1394,1015,1040]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D370464DD7D66C7791CBD2A" box="[1302,1394,1015,1040]" italics="true" pageId="4" pageNumber="205">Triberta</emphasis>
</taxonomicName>
are known to feed exclusively on
<taxonomicName id="A5F554B49D370464DA71612C7EE7BD0F" box="[538,649,1052,1077]" class="Magnoliopsida" family="Cistaceae" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="205" phylum="Tracheophyta" rank="family">Cistaceae</taxonomicName>
.
<taxonomicName id="A5F554B49D370464DAF6612C7E97BD0F" box="[669,761,1052,1077]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D370464DAF6612C7E97BD0F" box="[669,761,1052,1077]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D370464DAF6612C7E97BD0F" box="[669,761,1052,1077]" italics="true" pageId="4" pageNumber="205">Triberta</emphasis>
</taxonomicName>
also differs from
@ -508,7 +509,7 @@ in a concatenated dataset of eleven nuclear gene sequences.
</paragraph>
<paragraph id="624A2F379D370464D8AC61557FCCBC40" blockId="4.[151,1436,151,2025]" pageId="4" pageNumber="205">
To facilitate the assignment of lithocolletine moths to
<taxonomicName id="A5F554B49D370464DB2161547FC8BD47" box="[842,934,1124,1149]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D370464DB2161547FC8BD47" box="[842,934,1124,1149]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D370464DB2161547FC8BD47" box="[842,934,1124,1149]" italics="true" pageId="4" pageNumber="205">Triberta</emphasis>
</taxonomicName>
, a summary of the foregoing diagnoses is provided:
@ -531,7 +532,7 @@ hind wing like in
: 5) pupation outside the mine; pupa with lateral cuticle sclerotization and without cremaster.
<emphasis id="5081F3259D370464DB42600E7FCABC6D" bold="true" box="[809,932,1342,1367]" pageId="4" pageNumber="205">Molecular</emphasis>
: 6)
<taxonomicName id="A5F554B49D370464DBB3600E785ABC6D" box="[984,1076,1342,1367]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D370464DBB3600E785ABC6D" box="[984,1076,1342,1367]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D370464DBB3600E785ABC6D" box="[984,1076,1342,1367]" italics="true" pageId="4" pageNumber="205">Triberta</emphasis>
</taxonomicName>
is not nested within two wellsupported clades
@ -570,7 +571,7 @@ Maximum Likelihood tree of Lithocolletiinae. The numbers above nodes denote the
<paragraph id="624A2F379D360465D8FC624C797CBE88" blockId="5.[151,1435,1916,1970]" pageId="5" pageNumber="206">
<emphasis id="5081F3259D360465D8FC624C7D38BEA8" bold="true" box="[151,342,1916,1939]" pageId="5" pageNumber="206">FIGURES 0203.</emphasis>
Adult morphology of
<taxonomicName id="A5F554B49D360465DA2E624D7F59BEA9" box="[581,823,1917,1939]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="206" phylum="Tracheophyta" rank="species" species="helianthemella">
<taxonomicName id="A5F554B49D360465DA2E624D7F59BEA9" box="[581,823,1917,1939]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="helianthemella">
<emphasis id="5081F3259D360465DA2E624D7F59BEA9" box="[581,823,1917,1939]" italics="true" pageId="5" pageNumber="206">Triberta helianthemella</emphasis>
</taxonomicName>
,
@ -588,7 +589,7 @@ Male genitalia of
<emphasis id="5081F3259D350466DA5F63887EC1BFF5" box="[564,687,1720,1743]" italics="true" pageId="6" pageNumber="207">Cameraria</emphasis>
</taxonomicName>
and
<taxonomicName id="A5F554B49D350466DA8E63877F2FBFEA" box="[741,833,1719,1744]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="6" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D350466DA8E63877F2FBFEA" box="[741,833,1719,1744]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D350466DA8E63877F2FBFEA" box="[741,833,1719,1744]" italics="true" pageId="6" pageNumber="207">Triberta</emphasis>
</taxonomicName>
. 0 4, Male genitalia of
@ -600,7 +601,7 @@ and
, Tervuren,
<date id="164B09F79D350466D96A63E97D00BFD5" box="[257,366,1753,1775]" pageId="6" pageNumber="207" value="2012-05-11">11.v. 2012</date>
, leg. E. De Coninck. Scale bar as indicated. 0 5, same preparation, tegumen with a pair of apical setae. Scale bar as indicated. 0 6, male genitalia of
<taxonomicName id="A5F554B49D350466DA3D63C77F2BBE37" box="[598,837,1783,1805]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="6" pageNumber="207" phylum="Tracheophyta" rank="species" species="helianthemella">
<taxonomicName id="A5F554B49D350466DA3D63C77F2BBE37" box="[598,837,1783,1805]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="helianthemella">
<emphasis id="5081F3259D350466DA3D63C77F2BBE37" box="[598,837,1783,1805]" italics="true" pageId="6" pageNumber="207">Triberta helianthemella</emphasis>
</taxonomicName>
,
@ -609,11 +610,11 @@ and
<date id="164B09F79D350466DC4163C778F6BE37" box="[1066,1176,1783,1805]" pageId="6" pageNumber="207" value="1964-11-07">07.xi.1964</date>
; prep. DP 3826♂; arrows indicate the modified sternum VIII. 0 7, same preparation, aedoeagus. 0 8, same preparation, tegumen. 0 9, same preparation, apex of tegumen; arrows indicate a pair of apical setae. Scale bar as indicated.
</paragraph>
<caption id="368A7FBF9D340467D8FC632C7CB7BFF1" httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="7" pageNumber="208" targetBox="[151,1413,178,1538]" targetPageId="7">
<caption id="368A7FBF9D340467D8FC632C7CB7BFF1" ID-DOI="http://doi.org/10.5281/zenodo.246618" httpUri="https://zenodo.org/record/246618/files/figure.png" pageId="7" pageNumber="208" targetBox="[151,1413,178,1538]" targetPageId="7">
<paragraph id="624A2F379D340467D8FC632C7CB7BFF1" blockId="7.[151,1436,1564,1739]" pageId="7" pageNumber="208">
<emphasis id="5081F3259D340467D8FC632C7D3DBF0B" bold="true" box="[151,339,1564,1587]" pageId="7" pageNumber="208">FIGURES 1015.</emphasis>
The primary types of
<taxonomicName id="A5F554B49D340467DA5F632C7EE9BF08" box="[564,647,1564,1586]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="7" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D340467DA5F632C7EE9BF08" box="[564,647,1564,1586]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D340467DA5F632C7EE9BF08" box="[564,647,1564,1586]" italics="true" pageId="7" pageNumber="208">Triberta</emphasis>
</taxonomicName>
species. 10, the neotype of
@ -656,14 +657,14 @@ Herrich-Schäffer, 1861
<paragraph id="624A2F379D3B0468D8AC65A77EF4B87A" blockId="8.[151,1436,151,320]" pageId="8" pageNumber="209">
<emphasis id="5081F3259D3B0468D8AC65A77D1EB98A" box="[199,368,151,176]" italics="true" pageId="8" pageNumber="209">Male genitalia</emphasis>
(
<figureCitation id="FACE33B29D3B0468D9E965A77E68B98A" box="[386,518,151,176]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="8" pageNumber="209">Figs 1619</figureCitation>
<figureCitation id="FACE33B29D3B0468D9E965A77E68B98A" box="[386,518,151,176]" captionStart="FIGURES 16 21" captionStartId="8.[151,264,1819,1842]" captionTargetBox="[151,1436,357,1798]" captionTargetId="figure@8.[151,1436,355,1799]" captionTargetPageId="8" captionText="FIGURES 16 21. Male genitalia of Triberta species. 16, Lithocolletis cistifoliella Groschke, 1944, male, neotype, prep. TRB 3554 ♂, in ZSM. 17, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. 18, Lithocolletis helianthemella Herrich-Schäffer, 1861, male, paralectotype, prep. TRB 3945 ♂, in ZMHB. 19, Valva of Triberta helianthemella, male, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. TRB 3552 ♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 200 µm. 21, sternum VIII of Lithocolletis helianthemella Herrich-Schäffer, 1861, male, lectotype, prep. TRB 3935 ♂, in ZMHB. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246619" httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="8" pageNumber="209">Figs 1619</figureCitation>
). Tegumen rather long, with a pair of subapical setae. Tuba analis sometimes significantly protruded. Valvae symmetrical, setose. Transtilla incomplete. Vinculum developed, crescent or trapezium shaped, without prominent saccus. Anellus developed, strongly sclerotized, tubular, broader at base; juxta present. Aedoeagus ca. ¼ shorter than valva, relatively complex, with bifurcated coecal end and gradually tapering towards slender apex; cornuti absent.
</paragraph>
<caption id="368A7FBF9D3B0468D8FC622B7DC8BED0" httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="8" pageNumber="209" targetBox="[151,1436,357,1798]" targetPageId="8">
<caption id="368A7FBF9D3B0468D8FC622B7DC8BED0" ID-DOI="http://doi.org/10.5281/zenodo.246619" httpUri="https://zenodo.org/record/246619/files/figure.png" pageId="8" pageNumber="209" targetBox="[151,1436,357,1798]" targetPageId="8">
<paragraph id="624A2F379D3B0468D8FC622B7DC8BED0" blockId="8.[151,1437,1819,2026]" pageId="8" pageNumber="209">
<emphasis id="5081F3259D3B0468D8FC622B7D36BE0B" bold="true" box="[151,344,1819,1842]" pageId="8" pageNumber="209">FIGURES 1621.</emphasis>
Male genitalia of
<taxonomicName id="A5F554B49D3B0468DA4F622C7E19BE08" box="[548,631,1820,1842]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D3B0468DA4F622C7E19BE08" box="[548,631,1820,1842]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D3B0468DA4F622C7E19BE08" box="[548,631,1820,1842]" italics="true" pageId="8" pageNumber="209">Triberta</emphasis>
</taxonomicName>
species. 16,
@ -682,7 +683,7 @@ Herrich-Schäffer, 1861
Herrich-Schäffer, 1861
</taxonomicName>
, male, paralectotype, prep. TRB3945♂, in ZMHB. 19, Valva of
<taxonomicName id="A5F554B49D3B0468D8FC62487DE8BEB4" box="[151,390,1912,1934]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="8" pageNumber="209" phylum="Tracheophyta" rank="species" species="helianthemella">
<taxonomicName id="A5F554B49D3B0468D8FC62487DE8BEB4" box="[151,390,1912,1934]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="helianthemella">
<emphasis id="5081F3259D3B0468D8FC62487DE8BEB4" box="[151,390,1912,1934]" italics="true" pageId="8" pageNumber="209">Triberta helianthemella</emphasis>
</taxonomicName>
, male, Italy, Verona, 25/30.vi.2010, leg. P. Triberti, prep. TRB3552♂, in MSNV. Scale bar 100 µm. 20, abdomen opening of
@ -701,36 +702,36 @@ Herrich-Schäffer, 1861
<paragraph id="624A2F379D3A0469D8AC65A77EE5B897" blockId="9.[151,1436,151,429]" pageId="9" pageNumber="210">
<emphasis id="5081F3259D3A0469D8AC65A77DE9B98A" box="[199,391,151,176]" italics="true" pageId="9" pageNumber="210">Female genitalia</emphasis>
(
<figureCitation id="FACE33B29D3A0469D9FD65A77E74B98A" box="[406,538,151,176]" captionStart="FIGURES 22 25" captionStartId="9.[151,264,1917,1940]" captionTargetBox="[172,1397,481,1900]" captionTargetId="figure@9.[172,1408,481,1900]" captionTargetPageId="9" captionText="FIGURES 22 25. Female genitalia of Triberta species. 22, Triberta cistifoliella, Greece, prep. TRB 3559 ♀. Scale bar 200 µm. 23, same preparation, signum on corpus bursae, prep. TRB 3559 ♀. Scale bar 100 µm. 24, Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. DP 3807 ♀. Scale bar 200 µm. 25, same preparation, signum on corpus bursae, prep. DP 3807 ♀. Scale bar 100 µm." httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="9" pageNumber="210">Figs 2225</figureCitation>
<figureCitation id="FACE33B29D3A0469D9FD65A77E74B98A" box="[406,538,151,176]" captionStart="FIGURES 22 25" captionStartId="9.[151,264,1917,1940]" captionTargetBox="[172,1397,481,1900]" captionTargetId="figure@9.[172,1408,481,1900]" captionTargetPageId="9" captionText="FIGURES 22 25. Female genitalia of Triberta species. 22, Triberta cistifoliella, Greece, prep. TRB 3559 ♀. Scale bar 200 µm. 23, same preparation, signum on corpus bursae, prep. TRB 3559 ♀. Scale bar 100 µm. 24, Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. DP 3807 ♀. Scale bar 200 µm. 25, same preparation, signum on corpus bursae, prep. DP 3807 ♀. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246620" httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="9" pageNumber="210">Figs 2225</figureCitation>
). Papillae anales rounded caudally, not connected laterally. Segment VIII short, well connected to segment VII. Posterior apophyses without basal plate, but thickened basally and gradually acute apically; anterior apophyses originating from sclerotized basal plate of segment VIII, similar in form to posterior apophyses. Ostium bursae opens at the subanterior region of segment VIII. Ductus bursae very narrow, long with sclerotized antrum; ductus seminalis arises ca. midway along ductus bursae (
<figureCitation id="FACE33B29D3A0469DC68641778E7B87A" box="[1027,1161,295,320]" captionStart="FIGURES 22 25" captionStartId="9.[151,264,1917,1940]" captionTargetBox="[172,1397,481,1900]" captionTargetId="figure@9.[172,1408,481,1900]" captionTargetPageId="9" captionText="FIGURES 22 25. Female genitalia of Triberta species. 22, Triberta cistifoliella, Greece, prep. TRB 3559 ♀. Scale bar 200 µm. 23, same preparation, signum on corpus bursae, prep. TRB 3559 ♀. Scale bar 100 µm. 24, Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. DP 3807 ♀. Scale bar 200 µm. 25, same preparation, signum on corpus bursae, prep. DP 3807 ♀. Scale bar 100 µm." httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="9" pageNumber="210">Figs 22, 24</figureCitation>
<figureCitation id="FACE33B29D3A0469DC68641778E7B87A" box="[1027,1161,295,320]" captionStart="FIGURES 22 25" captionStartId="9.[151,264,1917,1940]" captionTargetBox="[172,1397,481,1900]" captionTargetId="figure@9.[172,1408,481,1900]" captionTargetPageId="9" captionText="FIGURES 22 25. Female genitalia of Triberta species. 22, Triberta cistifoliella, Greece, prep. TRB 3559 ♀. Scale bar 200 µm. 23, same preparation, signum on corpus bursae, prep. TRB 3559 ♀. Scale bar 100 µm. 24, Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. DP 3807 ♀. Scale bar 200 µm. 25, same preparation, signum on corpus bursae, prep. DP 3807 ♀. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246620" httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="9" pageNumber="210">Figs 22, 24</figureCitation>
). Corpus bursae distinct from ductus bursae, globular, with two signa regions: a relatively large speculate-sculptured disc along lateral wall of corpus bursae and a strongly sclerotized stellate signum covered with small sharp barbs about midway along the opposite wall of corpus bursae (
<figureCitation id="FACE33B29D3A0469D99764A47E10B897" box="[508,638,404,429]" captionStart="FIGURES 22 25" captionStartId="9.[151,264,1917,1940]" captionTargetBox="[172,1397,481,1900]" captionTargetId="figure@9.[172,1408,481,1900]" captionTargetPageId="9" captionText="FIGURES 22 25. Female genitalia of Triberta species. 22, Triberta cistifoliella, Greece, prep. TRB 3559 ♀. Scale bar 200 µm. 23, same preparation, signum on corpus bursae, prep. TRB 3559 ♀. Scale bar 100 µm. 24, Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. DP 3807 ♀. Scale bar 200 µm. 25, same preparation, signum on corpus bursae, prep. DP 3807 ♀. Scale bar 100 µm." httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="9" pageNumber="210">Figs 23, 25</figureCitation>
<figureCitation id="FACE33B29D3A0469D99764A47E10B897" box="[508,638,404,429]" captionStart="FIGURES 22 25" captionStartId="9.[151,264,1917,1940]" captionTargetBox="[172,1397,481,1900]" captionTargetId="figure@9.[172,1408,481,1900]" captionTargetPageId="9" captionText="FIGURES 22 25. Female genitalia of Triberta species. 22, Triberta cistifoliella, Greece, prep. TRB 3559 ♀. Scale bar 200 µm. 23, same preparation, signum on corpus bursae, prep. TRB 3559 ♀. Scale bar 100 µm. 24, Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti, prep. DP 3807 ♀. Scale bar 200 µm. 25, same preparation, signum on corpus bursae, prep. DP 3807 ♀. Scale bar 100 µm." figureDoi="http://doi.org/10.5281/zenodo.246620" httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="9" pageNumber="210">Figs 23, 25</figureCitation>
).
</paragraph>
<caption id="368A7FBF9D3A0469D8FC624D7DADBECA" httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="9" pageNumber="210" targetBox="[172,1397,481,1900]" targetPageId="9">
<caption id="368A7FBF9D3A0469D8FC624D7DADBECA" ID-DOI="http://doi.org/10.5281/zenodo.246620" httpUri="https://zenodo.org/record/246620/files/figure.png" pageId="9" pageNumber="210" targetBox="[172,1397,481,1900]" targetPageId="9">
<paragraph id="624A2F379D3A0469D8FC624D7DADBECA" blockId="9.[151,1436,1917,2032]" pageId="9" pageNumber="210">
<emphasis id="5081F3259D3A0469D8FC624D7D3ABEA9" bold="true" box="[151,340,1917,1940]" pageId="9" pageNumber="210">FIGURES 2225.</emphasis>
Female genitalia of
<taxonomicName id="A5F554B49D3A0469DA4F624E7E19BEAE" box="[548,631,1918,1940]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D3A0469DA4F624E7E19BEAE" box="[548,631,1918,1940]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D3A0469DA4F624E7E19BEAE" box="[548,631,1918,1940]" italics="true" pageId="9" pageNumber="210">Triberta</emphasis>
</taxonomicName>
species. 22,
<taxonomicName id="A5F554B49D3A0469DA92624E7FABBEAE" box="[761,965,1918,1940]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="210" phylum="Tracheophyta" rank="species" species="cistifoliella">
<taxonomicName id="A5F554B49D3A0469DA92624E7FABBEAE" box="[761,965,1918,1940]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="cistifoliella">
<emphasis id="5081F3259D3A0469DA92624E7FABBEAE" box="[761,965,1918,1940]" italics="true" pageId="9" pageNumber="210">Triberta cistifoliella</emphasis>
</taxonomicName>
, Greece, prep. TRB3559♀. Scale bar 200 µm. 23, same preparation, signum on corpus bursae, prep. TRB3559♀. Scale bar 100 µm. 24,
<taxonomicName id="A5F554B49D3A0469DC7162AD7967BE89" box="[1050,1289,1949,1971]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="9" pageNumber="210" phylum="Tracheophyta" rank="species" species="helianthemella">
<taxonomicName id="A5F554B49D3A0469DC7162AD7967BE89" box="[1050,1289,1949,1971]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="helianthemella">
<emphasis id="5081F3259D3A0469DC7162AD7967BE89" box="[1050,1289,1949,1971]" italics="true" pageId="9" pageNumber="210">Triberta helianthemella</emphasis>
</taxonomicName>
, Italy, Verona, 25/30.vi.2010, leg. P. Triberti, prep. DP3807♀. Scale bar 200 µm. 25, same preparation, signum on corpus bursae, prep. DP3807♀. Scale bar 100 µm.
</paragraph>
</caption>
<caption id="368A7FBF9D39046AD8FC63117DA7BF4F" httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211" targetBox="[175,1410,151,1545]" targetPageId="10">
<caption id="368A7FBF9D39046AD8FC63117DA7BF4F" ID-DOI="http://doi.org/10.5281/zenodo.246621" httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211" targetBox="[175,1410,151,1545]" targetPageId="10">
<paragraph id="624A2F379D39046AD8FC63117DA7BF4F" blockId="10.[151,1435,1569,1653]" pageId="10" pageNumber="211">
<emphasis id="5081F3259D39046AD8FC63117D3ABF0D" bold="true" box="[151,340,1569,1592]" pageId="10" pageNumber="211">FIGURES 2631.</emphasis>
Head of late instar larva of
<taxonomicName id="A5F554B49D39046ADA1B63127F0EBF02" box="[624,864,1570,1592]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="211" phylum="Tracheophyta" rank="species" species="helianthemella">
<taxonomicName id="A5F554B49D39046ADA1B63127F0EBF02" box="[624,864,1570,1592]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="helianthemella">
<emphasis id="5081F3259D39046ADA1B63127F0EBF02" box="[624,864,1570,1592]" italics="true" pageId="10" pageNumber="211">Triberta helianthemella</emphasis>
</taxonomicName>
, Italy, Verona, 25/30.vi.2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated.
@ -739,37 +740,37 @@ Head of late instar larva of
<paragraph id="624A2F379D39046BD8AC63AC7935B8EE" blockId="10.[151,1436,1692,2004]" lastBlockId="11.[151,1437,151,469]" lastPageId="11" lastPageNumber="212" pageId="10" pageNumber="211">
<emphasis id="5081F3259D39046AD8AC63AC7D7EBF8F" bold="true" box="[199,272,1692,1717]" pageId="10" pageNumber="211">Larva</emphasis>
. Last instar larva ca. 3.6 mm long. Head of late instars approximately round with full complement of mouthparts (
<figureCitation id="FACE33B29D39046AD94E63F07DBFBFE3" box="[293,465,1728,1753]" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211">Fig. 26, 2931</figureCitation>
<figureCitation id="FACE33B29D39046AD94E63F07DBFBFE3" box="[293,465,1728,1753]" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246621" httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211">Fig. 26, 2931</figureCitation>
); 3 pairs of stemmata arranged per pair: stemmata 1 and
<quantity id="A50D82D29D39046ADC3063F178E9BFE3" box="[1115,1159,1728,1753]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="211" unit="in" value="2.0">2 in</quantity>
a horizontal row distant from each other; stemmata 3 and 4 close to antenna in a vertical row at short distance close to each other; stemmata 5 and
<quantity id="A50D82D29D39046AD8B662377D67BE1A" box="[221,265,1799,1824]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="211" unit="in" value="6.0">6 in</quantity>
a vertical row touching each other (
<figureCitation id="FACE33B29D39046ADACA62377F4BBE25" box="[673,805,1799,1824]" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211">Figs 27, 28</figureCitation>
<figureCitation id="FACE33B29D39046ADACA62377F4BBE25" box="[673,805,1799,1824]" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246621" httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211">Figs 27, 28</figureCitation>
). Nine setae surround the stemmata of which S3 is very short; antenna 3-segmented with second segment moderately long; sensilla as shown in
<figureCitation id="FACE33B29D39046ADCED621A78B4BE79" box="[1158,1242,1834,1859]" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211">Fig. 31</figureCitation>
<figureCitation id="FACE33B29D39046ADCED621A78B4BE79" box="[1158,1242,1834,1859]" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246621" httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211">Fig. 31</figureCitation>
; labrum strongly bilobed with raised median part (Fig. 33); M1 and M4 present; inner perimeter of labrum without spines; the dorsal surface of the hypopharynx of the last tissue feeding larval instar is covered with hypopharyngeal striations (
<figureCitation id="FACE33B29D39046ADD0162447C8FBE8A" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211">Figs 2931</figureCitation>
<figureCitation id="FACE33B29D39046ADD0162447C8FBE8A" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246621" httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211">Figs 2931</figureCitation>
); ventral surface of head covered with numerous short rough spines; maxillary palpus as shown in
<figureCitation id="FACE33B29D39046ADD5162A77CD8BEEE" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211">Figs 29 31</figureCitation>
<figureCitation id="FACE33B29D39046ADD5162A77CD8BEEE" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246621" httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="10" pageNumber="211">Figs 29 31</figureCitation>
; median lobe of the maxilla with numerous sensory setae of different lengths. Labial palpus bears two sensory setae: one short seta, ca. 2× longer than apical palpomere and one long seta ca.
<date id="164B09F79D38046BDC5065A87839B98A" box="[1083,1111,152,176]" pageId="11" pageNumber="212">4x</date>
length of apical palpomere. Spinneret a relatively long striate tube (
<figureCitation id="FACE33B29D38046BDA3A658B7ECBB9EE" box="[593,677,187,212]" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="11" pageNumber="212">Fig. 29</figureCitation>
<figureCitation id="FACE33B29D38046BDA3A658B7ECBB9EE" box="[593,677,187,212]" captionStart="FIGURES 26 31" captionStartId="10.[151,264,1569,1592]" captionTargetBox="[175,1410,151,1545]" captionTargetId="figure@10.[175,1411,151,1546]" captionTargetPageId="10" captionText="FIGURES 26 31. Head of late instar larva of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 26, frontal view. 27, lateral view. 28, mouthparts. 29, mandibles and hypopharynx, ventral view. 30, ventral surface of head. 31, maxillary palpus. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246621" httpUri="https://zenodo.org/record/246621/files/figure.png" pageId="11" pageNumber="212">Fig. 29</figureCitation>
). Thorax with setae XD1 and XD2, SD1 long, immediately ventral to XD2; SD2 absent on T1; L group of setae long, L2 longer than L1; dorsal surface of T1 covered with both longer and shorter protrubances (Fig. 34); T2 and T3 with long D1 and shorter D2; dorsal surface of thoracic segments covered with tiny spines (Fig. 35). Legs fully developed, coxae with 4 coxal setae; femur and tibia with very long setae; tarsal claw with a sharp spine (Figs 36, 37). Abdomen with bisetose D group on A18, unisetose on A9 (
<figureCitation id="FACE33B29D38046BDD50646179FEB850" box="[1339,1424,337,362]" captionStart="FIGURES 38 43" captionStartId="12.[151,264,1662,1685]" captionTargetBox="[151,1436,193,1640]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURES 38 43. Prolegs of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 38, ventral view. 39, crochets in rest position, ventral view. 40, crochets in open position, ventral view. 41, caudal segments. 42, abdominal segment A 9, dorsal view. 43, caudal prolegs, ventral view. Scale bar as indicated." httpUri="https://zenodo.org/record/246622/files/figure.png" pageId="11" pageNumber="212">Fig. 42</figureCitation>
<figureCitation id="FACE33B29D38046BDD50646179FEB850" box="[1339,1424,337,362]" captionStart="FIGURES 38 43" captionStartId="12.[151,264,1662,1685]" captionTargetBox="[151,1436,193,1640]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURES 38 43. Prolegs of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 38, ventral view. 39, crochets in rest position, ventral view. 40, crochets in open position, ventral view. 41, caudal segments. 42, abdominal segment A 9, dorsal view. 43, caudal prolegs, ventral view. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246622" httpUri="https://zenodo.org/record/246622/files/figure.png" pageId="11" pageNumber="212">Fig. 42</figureCitation>
). Prolegs present on A35, 10; crochets consisting of 16 larger hooks arranged in biordinal semicircle and four small spines arranged in a row opposite the larger hooks (
<figureCitation id="FACE33B29D38046BDABC64A87F34B88B" box="[727,858,408,433]" captionStart="FIGURES 38 43" captionStartId="12.[151,264,1662,1685]" captionTargetBox="[151,1436,193,1640]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURES 38 43. Prolegs of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 38, ventral view. 39, crochets in rest position, ventral view. 40, crochets in open position, ventral view. 41, caudal segments. 42, abdominal segment A 9, dorsal view. 43, caudal prolegs, ventral view. Scale bar as indicated." httpUri="https://zenodo.org/record/246622/files/figure.png" pageId="11" pageNumber="212">Figs 3840</figureCitation>
<figureCitation id="FACE33B29D38046BDABC64A87F34B88B" box="[727,858,408,433]" captionStart="FIGURES 38 43" captionStartId="12.[151,264,1662,1685]" captionTargetBox="[151,1436,193,1640]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURES 38 43. Prolegs of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 38, ventral view. 39, crochets in rest position, ventral view. 40, crochets in open position, ventral view. 41, caudal segments. 42, abdominal segment A 9, dorsal view. 43, caudal prolegs, ventral view. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246622" httpUri="https://zenodo.org/record/246622/files/figure.png" pageId="11" pageNumber="212">Figs 3840</figureCitation>
); anal prolegs with crochets consisting of ca. 1012 rather large hooks arranged in a uniordinal semicircle (
<figureCitation id="FACE33B29D38046BDA96648C7F3FB8EE" box="[765,849,444,469]" captionStart="FIGURES 38 43" captionStartId="12.[151,264,1662,1685]" captionTargetBox="[151,1436,193,1640]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURES 38 43. Prolegs of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 38, ventral view. 39, crochets in rest position, ventral view. 40, crochets in open position, ventral view. 41, caudal segments. 42, abdominal segment A 9, dorsal view. 43, caudal prolegs, ventral view. Scale bar as indicated." httpUri="https://zenodo.org/record/246622/files/figure.png" pageId="11" pageNumber="212">Fig. 43</figureCitation>
<figureCitation id="FACE33B29D38046BDA96648C7F3FB8EE" box="[765,849,444,469]" captionStart="FIGURES 38 43" captionStartId="12.[151,264,1662,1685]" captionTargetBox="[151,1436,193,1640]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURES 38 43. Prolegs of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 38, ventral view. 39, crochets in rest position, ventral view. 40, crochets in open position, ventral view. 41, caudal segments. 42, abdominal segment A 9, dorsal view. 43, caudal prolegs, ventral view. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246622" httpUri="https://zenodo.org/record/246622/files/figure.png" pageId="11" pageNumber="212">Fig. 43</figureCitation>
). Anal plate with three pairs of setae (
<figureCitation id="FACE33B29D38046BDC91648C7920B8EE" box="[1274,1358,444,469]" captionStart="FIGURES 38 43" captionStartId="12.[151,264,1662,1685]" captionTargetBox="[151,1436,193,1640]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURES 38 43. Prolegs of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 38, ventral view. 39, crochets in rest position, ventral view. 40, crochets in open position, ventral view. 41, caudal segments. 42, abdominal segment A 9, dorsal view. 43, caudal prolegs, ventral view. Scale bar as indicated." httpUri="https://zenodo.org/record/246622/files/figure.png" pageId="11" pageNumber="212">Fig. 41</figureCitation>
<figureCitation id="FACE33B29D38046BDC91648C7920B8EE" box="[1274,1358,444,469]" captionStart="FIGURES 38 43" captionStartId="12.[151,264,1662,1685]" captionTargetBox="[151,1436,193,1640]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURES 38 43. Prolegs of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 38, ventral view. 39, crochets in rest position, ventral view. 40, crochets in open position, ventral view. 41, caudal segments. 42, abdominal segment A 9, dorsal view. 43, caudal prolegs, ventral view. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246622" httpUri="https://zenodo.org/record/246622/files/figure.png" pageId="11" pageNumber="212">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="624A2F379D38046BD8FC625B7C95BEE7" blockId="11.[151,1436,1897,2013]" pageId="11" pageNumber="212">
<emphasis id="5081F3259D38046BD8FC625B7D38BEBB" bold="true" box="[151,342,1899,1922]" pageId="11" pageNumber="212">FIGURES 3237.</emphasis>
Late instar larva of
<taxonomicName id="A5F554B49D38046BDA40625C7F72BEB8" box="[555,796,1900,1922]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="helianthemella">
<taxonomicName id="A5F554B49D38046BDA40625C7F72BEB8" box="[555,796,1900,1922]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="helianthemella">
<emphasis id="5081F3259D38046BDA40625C7F72BEB8" box="[555,796,1900,1922]" italics="true" pageId="11" pageNumber="212">Triberta helianthemella</emphasis>
</taxonomicName>
,
@ -778,11 +779,11 @@ Late instar larva of
<date id="164B09F79D38046BDBB5625C7824BEB8" box="[990,1098,1900,1922]" pageId="11" pageNumber="212" value="2010-06-30">30.vi.2010</date>
, leg. P. Triberti. 32, head and 1st thoracic segment of last instar larva, dorsal view. 33, same preparation, proximal part of head. 34, dorsal surface of T1. 35, dorsal surface of T3. 36, legs of late instar larva, latero-ventral view. 37, median thoracic leg, lateral view. Scale bar as indicated.
</paragraph>
<caption id="368A7FBF9D3F046CD8FC634E7F1ABFE8" httpUri="https://zenodo.org/record/246622/files/figure.png" pageId="12" pageNumber="213" targetBox="[151,1436,193,1640]" targetPageId="12">
<caption id="368A7FBF9D3F046CD8FC634E7F1ABFE8" ID-DOI="http://doi.org/10.5281/zenodo.246622" httpUri="https://zenodo.org/record/246622/files/figure.png" pageId="12" pageNumber="213" targetBox="[151,1436,193,1640]" targetPageId="12">
<paragraph id="624A2F379D3F046CD8FC634E7F1ABFE8" blockId="12.[151,1436,1662,1746]" pageId="12" pageNumber="213">
<emphasis id="5081F3259D3F046CD8FC634E7D36BFAE" bold="true" box="[151,344,1662,1685]" pageId="12" pageNumber="213">FIGURES 3843.</emphasis>
Prolegs of
<taxonomicName id="A5F554B49D3F046CD9B1634F7EA0BFAF" box="[474,718,1663,1685]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="helianthemella">
<taxonomicName id="A5F554B49D3F046CD9B1634F7EA0BFAF" box="[474,718,1663,1685]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="helianthemella">
<emphasis id="5081F3259D3F046CD9B1634F7EA0BFAF" box="[474,718,1663,1685]" italics="true" pageId="12" pageNumber="213">Triberta helianthemella</emphasis>
</taxonomicName>
, Italy, Verona, 25/30.vi.2010, leg. P. Triberti. 38, ventral view. 39, crochets in rest position, ventral view. 40, crochets in open position, ventral view. 41, caudal segments. 42, abdominal segment A9, dorsal view. 43, caudal prolegs, ventral view. Scale bar as indicated.
@ -791,45 +792,45 @@ Prolegs of
<paragraph id="624A2F379D3F046DD8AC63CE7EE4B897" blockId="12.[151,1437,1790,2031]" lastBlockId="13.[151,1436,151,429]" lastPageId="13" lastPageNumber="214" pageId="12" pageNumber="213">
<emphasis id="5081F3259D3F046CD8AC63CE7D6BBE2D" bold="true" box="[199,261,1790,1815]" pageId="12" pageNumber="213">Pupa</emphasis>
. Maximum length ca. 2.5 mm, width ca. 0.8 mm; body elongate, compact, relatively flat, narrower over last 4 segments (
<figureCitation id="FACE33B29D3F046CD93A62137DBABE06" box="[337,468,1827,1852]" captionStart="FIGURES 44 49" captionStartId="13.[151,264,1951,1974]" captionTargetBox="[151,1436,481,1929]" captionTargetId="figure@13.[151,1436,481,1930]" captionTargetPageId="13" captionText="FIGURES 44 49. Pupa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 44, ventral view. 45, lateral view. 46, pupal head. 47, A pair of frontal sensillae on pupal head. 48, sclerotized ridge on pupal segments A 2. 49, sclerotized ridge on pupal segments A 6 - 8. Scale bar as indicated." httpUri="https://zenodo.org/record/246623/files/figure.png" pageId="12" pageNumber="213">Figs 4445</figureCitation>
<figureCitation id="FACE33B29D3F046CD93A62137DBABE06" box="[337,468,1827,1852]" captionStart="FIGURES 44 49" captionStartId="13.[151,264,1951,1974]" captionTargetBox="[151,1436,481,1929]" captionTargetId="figure@13.[151,1436,481,1930]" captionTargetPageId="13" captionText="FIGURES 44 49. Pupa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 44, ventral view. 45, lateral view. 46, pupal head. 47, A pair of frontal sensillae on pupal head. 48, sclerotized ridge on pupal segments A 2. 49, sclerotized ridge on pupal segments A 6 - 8. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246623" httpUri="https://zenodo.org/record/246623/files/figure.png" pageId="12" pageNumber="213">Figs 4445</figureCitation>
); color a bright shade of brown. A thickened sclerotized ridge present along the lateral sides of A18. Head smooth, with two pairs of sockets bearing sensillae (
<figureCitation id="FACE33B29D3F046CDB9962767818BE65" box="[1010,1142,1862,1887]" captionStart="FIGURES 44 49" captionStartId="13.[151,264,1951,1974]" captionTargetBox="[151,1436,481,1929]" captionTargetId="figure@13.[151,1436,481,1930]" captionTargetPageId="13" captionText="FIGURES 44 49. Pupa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 44, ventral view. 45, lateral view. 46, pupal head. 47, A pair of frontal sensillae on pupal head. 48, sclerotized ridge on pupal segments A 2. 49, sclerotized ridge on pupal segments A 6 - 8. Scale bar as indicated." httpUri="https://zenodo.org/record/246623/files/figure.png" pageId="12" pageNumber="213">Fig. 46, 47</figureCitation>
<figureCitation id="FACE33B29D3F046CDB9962767818BE65" box="[1010,1142,1862,1887]" captionStart="FIGURES 44 49" captionStartId="13.[151,264,1951,1974]" captionTargetBox="[151,1436,481,1929]" captionTargetId="figure@13.[151,1436,481,1930]" captionTargetPageId="13" captionText="FIGURES 44 49. Pupa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 44, ventral view. 45, lateral view. 46, pupal head. 47, A pair of frontal sensillae on pupal head. 48, sclerotized ridge on pupal segments A 2. 49, sclerotized ridge on pupal segments A 6 - 8. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246623" httpUri="https://zenodo.org/record/246623/files/figure.png" pageId="12" pageNumber="213">Fig. 46, 47</figureCitation>
). Vertex furnished with a frontal process (cocoon cutter), which is relatively short, broadly triangular, with slightly wrinkled median surface (
<figureCitation id="FACE33B29D3F046CD8F462BE7C98BE9D" box="[159,246,1934,1959]" captionStart="FIGURES 44 49" captionStartId="13.[151,264,1951,1974]" captionTargetBox="[151,1436,481,1929]" captionTargetId="figure@13.[151,1436,481,1930]" captionTargetPageId="13" captionText="FIGURES 44 49. Pupa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 44, ventral view. 45, lateral view. 46, pupal head. 47, A pair of frontal sensillae on pupal head. 48, sclerotized ridge on pupal segments A 2. 49, sclerotized ridge on pupal segments A 6 - 8. Scale bar as indicated." httpUri="https://zenodo.org/record/246623/files/figure.png" pageId="12" pageNumber="213">Fig. 46</figureCitation>
<figureCitation id="FACE33B29D3F046CD8F462BE7C98BE9D" box="[159,246,1934,1959]" captionStart="FIGURES 44 49" captionStartId="13.[151,264,1951,1974]" captionTargetBox="[151,1436,481,1929]" captionTargetId="figure@13.[151,1436,481,1930]" captionTargetPageId="13" captionText="FIGURES 44 49. Pupa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 44, ventral view. 45, lateral view. 46, pupal head. 47, A pair of frontal sensillae on pupal head. 48, sclerotized ridge on pupal segments A 2. 49, sclerotized ridge on pupal segments A 6 - 8. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246623" httpUri="https://zenodo.org/record/246623/files/figure.png" pageId="12" pageNumber="213">Fig. 46</figureCitation>
). Forewings long, extending to the posterior margin of abdominal segment A6, and contiguous at their distal ends (
<figureCitation id="FACE33B29D3F046CD97562837D18BEF1" box="[286,374,1971,1996]" captionStart="FIGURES 44 49" captionStartId="13.[151,264,1951,1974]" captionTargetBox="[151,1436,481,1929]" captionTargetId="figure@13.[151,1436,481,1930]" captionTargetPageId="13" captionText="FIGURES 44 49. Pupa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 44, ventral view. 45, lateral view. 46, pupal head. 47, A pair of frontal sensillae on pupal head. 48, sclerotized ridge on pupal segments A 2. 49, sclerotized ridge on pupal segments A 6 - 8. Scale bar as indicated." httpUri="https://zenodo.org/record/246623/files/figure.png" pageId="12" pageNumber="213">Figs 44</figureCitation>
<figureCitation id="FACE33B29D3F046CD97562837D18BEF1" box="[286,374,1971,1996]" captionStart="FIGURES 44 49" captionStartId="13.[151,264,1951,1974]" captionTargetBox="[151,1436,481,1929]" captionTargetId="figure@13.[151,1436,481,1930]" captionTargetPageId="13" captionText="FIGURES 44 49. Pupa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 44, ventral view. 45, lateral view. 46, pupal head. 47, A pair of frontal sensillae on pupal head. 48, sclerotized ridge on pupal segments A 2. 49, sclerotized ridge on pupal segments A 6 - 8. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246623" httpUri="https://zenodo.org/record/246623/files/figure.png" pageId="12" pageNumber="213">Figs 44</figureCitation>
,
<figureCitation id="FACE33B29D3F046CD9E962837DCCBEF6" box="[386,418,1971,1996]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="12" pageNumber="213">54</figureCitation>
<figureCitation id="FACE33B29D3F046CD9E962837DCCBEF6" box="[386,418,1971,1996]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246624" httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="12" pageNumber="213">54</figureCitation>
). The appendages of antennae longer than forewings and extend up to caudal end of pupa. Labium rather long, ca. 1.7× shorter than proboscis. The hind leg sheaths are slightly shorter than antennal appendages (
<figureCitation id="FACE33B29D3E046DD94265A77D14B98A" box="[297,378,151,176]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Fig. 54</figureCitation>
<figureCitation id="FACE33B29D3E046DD94265A77D14B98A" box="[297,378,151,176]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246624" httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Fig. 54</figureCitation>
). The distance between the apices of the mesothoracic and metathoracic legs is approximately 1.46× the distance between the apices of the prothoracic and mesothoracic legs. One pair of dorsal setae is present on segments TIITIII. Abdominal segments A15 of more or less equal size, A68 strongly decreasing in size. Abdominal segments A38 covered with dense, minute spines dorsally (
<figureCitation id="FACE33B29D3E046DDBD46434787CB827" box="[959,1042,260,285]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Fig. 51</figureCitation>
<figureCitation id="FACE33B29D3E046DDBD46434787CB827" box="[959,1042,260,285]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246624" httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Fig. 51</figureCitation>
); the posterior margin of A1 with a wrinkled band (
<figureCitation id="FACE33B29D3E046DD92564177DC8B87A" box="[334,422,295,320]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Fig. 50</figureCitation>
<figureCitation id="FACE33B29D3E046DD92564177DC8B87A" box="[334,422,295,320]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246624" httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Fig. 50</figureCitation>
); dorsal surface of each abdominal segment carries a pair of setae, except segment A9 which lacks setae (
<figureCitation id="FACE33B29D3E046DD91C647C7DBEB85F" box="[375,464,332,357]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Fig. 52</figureCitation>
<figureCitation id="FACE33B29D3E046DD91C647C7DBEB85F" box="[375,464,332,357]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246624" httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Fig. 52</figureCitation>
). Terminal segment A10 fully covered with small spinaculae (
<figureCitation id="FACE33B29D3E046DDCC4647C7954B85F" box="[1199,1338,332,357]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Figs 53, 54</figureCitation>
<figureCitation id="FACE33B29D3E046DDCC4647C7954B85F" box="[1199,1338,332,357]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246624" httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Figs 53, 54</figureCitation>
); caudal surface of A10 without cremaster but with eight tiny protuberances each carrying a tiny nodule (
<figureCitation id="FACE33B29D3E046DDCAC645F7976B8B2" box="[1223,1304,367,392]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Fig. 55</figureCitation>
<figureCitation id="FACE33B29D3E046DDCAC645F7976B8B2" box="[1223,1304,367,392]" captionStart="FIGURES 50 55. P" captionStartId="14.[151,264,1638,1661]" captionTargetBox="[161,1424,187,1614]" captionTargetId="figure@14.[161,1425,187,1615]" captionTargetPageId="14" captionText="FIGURES 50 55. P upa of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 50, dorsal surface of segment A 1. 51, dorsal surface of segment A 8. 52, segments A 8 - 9, lateral view. 53, caudal segment, dorsal view. 54, A 10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated." figureDoi="http://doi.org/10.5281/zenodo.246624" httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="13" pageNumber="214">Fig. 55</figureCitation>
) (Gregor &amp; Patočka 2001: 34; Patočka &amp; Turčáni 2005).
</paragraph>
</subSubSection>
<caption id="368A7FBF9D3E046DD8FC62AF7ED6BEC8" httpUri="https://zenodo.org/record/246623/files/figure.png" pageId="13" pageNumber="214" targetBox="[151,1436,481,1929]" targetPageId="13">
<caption id="368A7FBF9D3E046DD8FC62AF7ED6BEC8" ID-DOI="http://doi.org/10.5281/zenodo.246623" httpUri="https://zenodo.org/record/246623/files/figure.png" pageId="13" pageNumber="214" targetBox="[151,1436,481,1929]" targetPageId="13">
<paragraph id="624A2F379D3E046DD8FC62AF7ED6BEC8" blockId="13.[151,1435,1951,2034]" pageId="13" pageNumber="214">
<emphasis id="5081F3259D3E046DD8FC62AF7D3BBE8E" bold="true" box="[151,341,1951,1974]" pageId="13" pageNumber="214">FIGURES 4449.</emphasis>
Pupa of
<taxonomicName id="A5F554B49D3E046DD9DF62AF7ECBBE8F" box="[436,677,1951,1973]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="13" pageNumber="214" phylum="Tracheophyta" rank="species" species="helianthemella">
<taxonomicName id="A5F554B49D3E046DD9DF62AF7ECBBE8F" box="[436,677,1951,1973]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="helianthemella">
<emphasis id="5081F3259D3E046DD9DF62AF7ECBBE8F" box="[436,677,1951,1973]" italics="true" pageId="13" pageNumber="214">Triberta helianthemella</emphasis>
</taxonomicName>
, Italy, Verona, 25/30.vi.2010, leg. P. Triberti. 44, ventral view. 45, lateral view. 46, pupal head. 47, A pair of frontal sensillae on pupal head. 48, sclerotized ridge on pupal segments A2. 49, sclerotized ridge on pupal segments A6-8. Scale bar as indicated.
</paragraph>
</caption>
<caption id="368A7FBF9D3D046ED8FC63567E8ABF80" httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="14" pageNumber="215" targetBox="[161,1424,187,1614]" targetPageId="14">
<caption id="368A7FBF9D3D046ED8FC63567E8ABF80" ID-DOI="http://doi.org/10.5281/zenodo.246624" httpUri="https://zenodo.org/record/246624/files/figure.png" pageId="14" pageNumber="215" targetBox="[161,1424,187,1614]" targetPageId="14">
<paragraph id="624A2F379D3D046ED8FC63567E8ABF80" blockId="14.[151,1436,1638,1722]" pageId="14" pageNumber="215">
<emphasis id="5081F3259D3D046ED8FC63567D07BF46" bold="true" box="[151,361,1638,1661]" pageId="14" pageNumber="215">FIGURES 5055. P</emphasis>
upa of
<taxonomicName id="A5F554B49D3D046ED9DB63567ECEBF46" box="[432,672,1638,1660]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="14" pageNumber="215" phylum="Tracheophyta" rank="species" species="helianthemella">
<taxonomicName id="A5F554B49D3D046ED9DB63567ECEBF46" box="[432,672,1638,1660]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="helianthemella">
<emphasis id="5081F3259D3D046ED9DB63567ECEBF46" box="[432,672,1638,1660]" italics="true" pageId="14" pageNumber="215">Triberta helianthemella</emphasis>
</taxonomicName>
, Italy, Verona, 25/30.vi.2010, leg. P. Triberti. 50, dorsal surface of segment A1. 51, dorsal surface of segment A8. 52, segments A8-9, lateral view. 53, caudal segment, dorsal view. 54, A10, caudal view. 55, nodule on caudal protuberance. Scale bar as indicated.
@ -841,13 +842,13 @@ upa of
. Larva solitary, feeding exclusively on
<taxonomicName id="A5F554B49D3D046EDAB063D17F24BFC0" box="[731,842,1761,1786]" class="Magnoliopsida" family="Cistaceae" kingdom="Plantae" order="Malvales" pageId="14" pageNumber="215" phylum="Tracheophyta" rank="family">Cistaceae</taxonomicName>
. Abaxial mine begins as a small corridor that closely follows a thick vein of the plant. It continues into a contracted blotch mine that may occupy the entire leaf; mine becomes transparent when it is fully grown. The tissue-feeding instars remove mesophyll layers of the leaf, and though the mine is formed under the abaxial epidermial layer, from adaxial leaf surface it is visible as a large, contiguous brownish blotch (
<figureCitation id="FACE33B29D3D046ED9B562417E0FBEB0" box="[478,609,1905,1930]" captionStart="FIGURES 56 59" captionStartId="15.[151,264,1478,1501]" captionTargetBox="[151,1436,373,1456]" captionTargetId="figure@15.[151,1436,373,1457]" captionTargetPageId="15" captionText="FIGURES 56 59. Biology of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 56, host plant Helianthemum nummularium. 57, mine, adaxial view. 58, excrements of larva in mine, adaxial view. 59, transparent cocoon outside the mine. Scale bar 1 mm." httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="14" pageNumber="215">Figs 57, 58</figureCitation>
<figureCitation id="FACE33B29D3D046ED9B562417E0FBEB0" box="[478,609,1905,1930]" captionStart="FIGURES 56 59" captionStartId="15.[151,264,1478,1501]" captionTargetBox="[151,1436,373,1456]" captionTargetId="figure@15.[151,1436,373,1457]" captionTargetPageId="15" captionText="FIGURES 56 59. Biology of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 56, host plant Helianthemum nummularium. 57, mine, adaxial view. 58, excrements of larva in mine, adaxial view. 59, transparent cocoon outside the mine. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246625" httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="14" pageNumber="215">Figs 57, 58</figureCitation>
). However, in some cases the mine can be formed on adaxial surface (see the original description of
<taxonomicName id="A5F554B49D3D046ED9BF62A57E84BE94" box="[468,746,1941,1966]" class="Insecta" family="Elachistidae" genus="Lithocolletis" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="cistifoliella">
<emphasis id="5081F3259D3D046ED9BF62A57E84BE94" box="[468,746,1941,1966]" italics="true" pageId="14" pageNumber="215">Lithocolletis cistifoliella</emphasis>
</taxonomicName>
here below). Frass distributed either in a row while larva crosses the vein or loose in small accumulations. Pupation occurs outside the mine in a transparent cocoon on a clean background without excrements (
<figureCitation id="FACE33B29D3D046EDA2462ED7ECABECC" box="[591,676,2013,2038]" captionStart="FIGURES 56 59" captionStartId="15.[151,264,1478,1501]" captionTargetBox="[151,1436,373,1456]" captionTargetId="figure@15.[151,1436,373,1457]" captionTargetPageId="15" captionText="FIGURES 56 59. Biology of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 56, host plant Helianthemum nummularium. 57, mine, adaxial view. 58, excrements of larva in mine, adaxial view. 59, transparent cocoon outside the mine. Scale bar 1 mm." httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="14" pageNumber="215">Fig. 59</figureCitation>
<figureCitation id="FACE33B29D3D046EDA2462ED7ECABECC" box="[591,676,2013,2038]" captionStart="FIGURES 56 59" captionStartId="15.[151,264,1478,1501]" captionTargetBox="[151,1436,373,1456]" captionTargetId="figure@15.[151,1436,373,1457]" captionTargetPageId="15" captionText="FIGURES 56 59. Biology of Triberta helianthemella, Italy, Verona, 25 / 30. vi. 2010, leg. P. Triberti. 56, host plant Helianthemum nummularium. 57, mine, adaxial view. 58, excrements of larva in mine, adaxial view. 59, transparent cocoon outside the mine. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.246625" httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="14" pageNumber="215">Fig. 59</figureCitation>
). Biology bivoltine with hibernation as adults is reported.
</paragraph>
<paragraph id="624A2F379D3C046FD8AC65A778D3B87A" blockId="15.[151,1436,151,320]" pageId="15" pageNumber="216">
@ -897,11 +898,11 @@ Gaertn. (Herrich-Schaffer 1861: 20)
(L.) Fourr. (Roüast 1884: 138).
</paragraph>
</subSubSection>
<caption id="368A7FBF9D3C046FD8FC60F67D9EBF20" httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="15" pageNumber="216" targetBox="[151,1436,373,1456]" targetPageId="15">
<caption id="368A7FBF9D3C046FD8FC60F67D9EBF20" ID-DOI="http://doi.org/10.5281/zenodo.246625" httpUri="https://zenodo.org/record/246625/files/figure.png" pageId="15" pageNumber="216" targetBox="[151,1436,373,1456]" targetPageId="15">
<paragraph id="624A2F379D3C046FD8FC60F67D9EBF20" blockId="15.[151,1436,1478,1562]" pageId="15" pageNumber="216">
<emphasis id="5081F3259D3C046FD8FC60F67D33BCE6" bold="true" box="[151,349,1478,1501]" pageId="15" pageNumber="216">FIGURES 5659.</emphasis>
Biology of
<taxonomicName id="A5F554B49D3C046FD99860F67E82BCE6" box="[499,748,1478,1500]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="15" pageNumber="216" phylum="Tracheophyta" rank="species" species="helianthemella">
<taxonomicName id="A5F554B49D3C046FD99860F67E82BCE6" box="[499,748,1478,1500]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="helianthemella">
<emphasis id="5081F3259D3C046FD99860F67E82BCE6" box="[499,748,1478,1500]" italics="true" pageId="15" pageNumber="216">Triberta helianthemella</emphasis>
</taxonomicName>
, Italy, Verona, 25/30.vi.2010, leg. P. Triberti. 56, host plant
@ -941,7 +942,7 @@ Bouček, 1958 (Bouček 1959: 131)
<subSubSection id="2AEF7CBC9D3C0470D8AC63827823BC9E" lastPageId="16" lastPageNumber="217" pageId="15" pageNumber="216" type="distribution">
<paragraph id="624A2F379D3C046FD8AC63827D9ABFD5" blockId="15.[151,1436,1606,1991]" pageId="15" pageNumber="216">
<emphasis id="5081F3259D3C046FD8AC63827D0EBFF1" bold="true" box="[199,352,1714,1739]" pageId="15" pageNumber="216">Distribution.</emphasis>
<taxonomicName id="A5F554B49D3C046FD90263827DABBFF1" box="[361,453,1714,1739]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="15" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D3C046FD90263827DABBFF1" box="[361,453,1714,1739]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D3C046FD90263827DABBFF1" box="[361,453,1714,1739]" italics="true" pageId="15" pageNumber="216">Triberta</emphasis>
</taxonomicName>
is a genus of xerothermic habitats in central and southern Europe, the Mediterranean region, and the Canary Islands.
@ -980,11 +981,11 @@ is a genus of xerothermic habitats in central and southern Europe, the Mediterra
<collectingCountry id="1AE26FA79D3C046FDD2762BA79F2BE99" box="[1356,1436,1930,1955]" name="Turkey" pageId="15" pageNumber="216">Turkey</collectingCountry>
(Stainton 1867: 35).
</paragraph>
<caption id="368A7FBF9D230470D8FC666A7CAEBA94" httpUri="https://zenodo.org/record/246626/files/figure.png" pageId="16" pageNumber="217" targetBox="[151,1436,193,837]" targetPageId="16">
<caption id="368A7FBF9D230470D8FC666A7CAEBA94" ID-DOI="http://doi.org/10.5281/zenodo.246626" httpUri="https://zenodo.org/record/246626/files/figure.png" pageId="16" pageNumber="217" targetBox="[151,1436,193,837]" targetPageId="16">
<paragraph id="624A2F379D230470D8FC666A7CAEBA94" blockId="16.[151,1436,858,942]" pageId="16" pageNumber="217">
<emphasis id="5081F3259D230470D8FC666A7D37BA4A" bold="true" box="[151,345,858,881]" pageId="16" pageNumber="217">FIGURES 6061.</emphasis>
Diagnostic characters in
<taxonomicName id="A5F554B49D230470DA1B666B7EADBA4B" box="[624,707,859,881]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D230470DA1B666B7EADBA4B" box="[624,707,859,881]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D230470DA1B666B7EADBA4B" box="[624,707,859,881]" italics="true" pageId="16" pageNumber="217">Triberta</emphasis>
</taxonomicName>
head vestiture of adults. 60, vertex of the neotype of
@ -1003,7 +1004,7 @@ Herrich-Schäffer, 1861
<paragraph id="624A2F379D230470D8AC66EB7F09BC2E" blockId="16.[151,1436,987,1552]" pageId="16" pageNumber="217">
<emphasis id="5081F3259D230470D8AC66EB7EA5BACE" bold="true" box="[199,715,987,1012]" pageId="16" pageNumber="217">Generic relationships and species diversity.</emphasis>
Based on taxa included in our phylogenetic analysis
<taxonomicName id="A5F554B49D230470DD4D66EB79ECBACE" box="[1318,1410,987,1012]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D230470DD4D66EB79ECBACE" box="[1318,1410,987,1012]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D230470DD4D66EB79ECBACE" box="[1318,1410,987,1012]" italics="true" pageId="16" pageNumber="217">Triberta</emphasis>
</taxonomicName>
is the sister lineage to the clade
@ -1017,7 +1018,7 @@ is the sister lineage to the clade
</emphasis>
. The ecological / morphological traits shared by these latter genera are as follows:
<emphasis id="5081F3259D230470DAF561137FFABD06" box="[670,916,1059,1084]" italics="true" pageId="16" pageNumber="217">
<taxonomicName id="A5F554B49D230470DAF561137E93BD06" box="[670,765,1059,1084]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">Triberta</taxonomicName>
<taxonomicName id="A5F554B49D230470DAF561137E93BD06" box="[670,765,1059,1084]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">Triberta</taxonomicName>
,
<taxonomicName id="A5F554B49D230470DB6061137FFABD06" box="[779,916,1059,1084]" class="Insecta" family="Gracillariidae" genus="Leucanthiza" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">Leucanthiza</taxonomicName>
</emphasis>
@ -1026,7 +1027,7 @@ and
<emphasis id="5081F3259D230470DBBB61137825BD06" box="[976,1099,1059,1084]" italics="true" pageId="16" pageNumber="217">Chrysaster</emphasis>
</taxonomicName>
pupate outside the mine, the mine of
<taxonomicName id="A5F554B49D230470D89361777D3ABD5A" box="[248,340,1095,1120]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D230470D89361777D3ABD5A" box="[248,340,1095,1120]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D230470D89361777D3ABD5A" box="[248,340,1095,1120]" italics="true" pageId="16" pageNumber="217">Triberta</emphasis>
</taxonomicName>
and
@ -1042,15 +1043,15 @@ and
<emphasis id="5081F3259D230470DC6861777810BD5A" box="[1027,1150,1095,1120]" italics="true" pageId="16" pageNumber="217">Chrysaster</emphasis>
</taxonomicName>
construct adaxial mines; pupa in
<taxonomicName id="A5F554B49D230470D893615B7D3ABDBE" box="[248,340,1131,1156]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D230470D893615B7D3ABDBE" box="[248,340,1131,1156]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D230470D893615B7D3ABDBE" box="[248,340,1131,1156]" italics="true" pageId="16" pageNumber="217">Triberta</emphasis>
</taxonomicName>
without cremaster like in other members of the clade; in the male genitalia, apex of tegumen carries one pair of setae, valvae symmetrical, densely setose; sternum VIII in
<taxonomicName id="A5F554B49D230470DC7161BF7818BD92" box="[1050,1142,1167,1192]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D230470DC7161BF7818BD92" box="[1050,1142,1167,1192]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D230470DC7161BF7818BD92" box="[1050,1142,1167,1192]" italics="true" pageId="16" pageNumber="217">Triberta</emphasis>
</taxonomicName>
is caudally extended, the character that separates
<taxonomicName id="A5F554B49D230470D9CC61837E6CBDF6" box="[423,514,1203,1228]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D230470D9CC61837E6CBDF6" box="[423,514,1203,1228]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D230470D9CC61837E6CBDF6" box="[423,514,1203,1228]" italics="true" pageId="16" pageNumber="217">Triberta</emphasis>
</taxonomicName>
from the other genera of the clade
@ -1069,7 +1070,7 @@ and
</paragraph>
<paragraph id="624A2F379D230470D8AC601078D1BC61" blockId="16.[151,1436,987,1552]" pageId="16" pageNumber="217">
In addition to the morphological characters mentioned above, a molecular phylogeny based on eleven nuclear genes places
<taxonomicName id="A5F554B49D230470D94060737DE9BC66" box="[299,391,1347,1372]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D230470D94060737DE9BC66" box="[299,391,1347,1372]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D230470D94060737DE9BC66" box="[299,391,1347,1372]" italics="true" pageId="16" pageNumber="217">Triberta</emphasis>
</taxonomicName>
apart from
@ -1088,11 +1089,11 @@ apart from
</paragraph>
<paragraph id="624A2F379D230470D8AC60587823BC9E" blockId="16.[151,1436,987,1552]" pageId="16" pageNumber="217">
Two species, both Palaearctic, are currently recognized in the new genus
<taxonomicName id="A5F554B49D230470DC56605778F7BCBA" box="[1085,1177,1383,1408]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D230470DC56605778F7BCBA" box="[1085,1177,1383,1408]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D230470DC56605778F7BCBA" box="[1085,1177,1383,1408]" italics="true" pageId="16" pageNumber="217">Triberta</emphasis>
</taxonomicName>
. The circumscription, delimitation and delineation of
<taxonomicName id="A5F554B49D230470D99D60BB7E3CBC9E" box="[502,594,1419,1444]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D230470D99D60BB7E3CBC9E" box="[502,594,1419,1444]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D230470D99D60BB7E3CBC9E" box="[502,594,1419,1444]" italics="true" pageId="16" pageNumber="217">Triberta</emphasis>
</taxonomicName>
species will be presented in a separate study.
@ -1102,7 +1103,7 @@ species will be presented in a separate study.
<paragraph id="624A2F379D230470D8AC609F79E8BF2A" blockId="16.[151,1436,987,1552]" pageId="16" pageNumber="217">
<emphasis id="5081F3259D230470D8AC609F7D28BCF2" bold="true" box="[199,326,1455,1480]" pageId="16" pageNumber="217">Etymology</emphasis>
. The name
<taxonomicName id="A5F554B49D230470D9A1609F7E48BCF2" box="[458,550,1455,1480]" class="Magnoliopsida" family="Cistaceae" genus="Triberta" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="201" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<taxonomicName id="A5F554B49D230470D9A1609F7E48BCF2" box="[458,550,1455,1480]" class="Insecta" family="Gracillariidae" genus="Triberta" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="201" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5081F3259D230470D9A1609F7E48BCF2" box="[458,550,1455,1480]" italics="true" pageId="16" pageNumber="217">Triberta</emphasis>
</taxonomicName>
is used as a noun in the nominative singular and feminine gender. This name is a patronym in honour of Paolo Triberti, a lepidopterist from the Museo Civico di Storia Naturale, Verona,