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<document ID-DOI="http://dx.doi.org/10.3897/jor.28.34605" ID-GBIF-Dataset="9a18facb-ad00-4ad1-9112-3cedd2455536" ID-GBIF-Taxon="159387938" ID-Pensoft-Pub="1937-2426-2-167" ID-Pensoft-UUID="1F59236AD82D5DC0B9FE5382623E640C" LSID="urn:lsid:zoobank.org:act:DF0F6588-4DC4-477B-9E4E-F7DC8DD56A76" ModsDocID="1937-2426-2-167" checkinTime="1567634996903" checkinUser="pensoft" docAuthor="Tumbrinck, Josef" docDate="2019" docId="72A2116B73E45EB0B650B133CA6A67F7" docLanguage="en" docName="JourOrthoptRes 28(2): 167-180" docOrigin="Journal of Orthoptera Research 28 (2)" docSource="http://dx.doi.org/10.3897/jor.28.34605" docTitle="Lamellitettigodes novaeguineae Tumbrinck, 2019, sp. nov." docType="treatment" docUuid="DF0F6588-4DC4-477B-9E4E-F7DC8DD56A76" docUuidSource="ZooBank" docVersion="4" id="1F59236AD82D5DC0B9FE5382623E640C" lastPageNumber="175" masterDocId="1F59236AD82D5DC0B9FE5382623E640C" masterDocTitle="Taxonomic and biogeographic revision of the genus Lamellitettigodes (Orthoptera: Tetrigidae) with description of two new species and additional notes on Lamellitettix, Probolotettix, and Scelimena" masterLastPageNumber="180" masterPageNumber="167" pageId="8" pageNumber="175" updateTime="1643481801102" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Taxonomic and biogeographic revision of the genus Lamellitettigodes (Orthoptera: Tetrigidae) with description of two new species and additional notes on Lamellitettix, Probolotettix, and Scelimena</mods:title>
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<mods:title>Journal of Orthoptera Research</mods:title>
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<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>28</mods:number>
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<mods:number>2</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jor.28.34605</mods:identifier>
<mods:identifier type="Pensoft-Pub">1937-2426-2-167</mods:identifier>
<mods:identifier type="ZooBank">CA0FD29D-1EE9-4176-9E1F-2EACBFFB68F1</mods:identifier>
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<treatment ID-GBIF-Taxon="159387938" LSID="urn:lsid:zoobank.org:act:DF0F6588-4DC4-477B-9E4E-F7DC8DD56A76" httpUri="http://treatment.plazi.org/id/72A2116B73E45EB0B650B133CA6A67F7" lastPageNumber="175" pageId="8" pageNumber="175">
<subSubSection pageId="8" pageNumber="175" type="nomenclature">
<paragraph pageId="8" pageNumber="175">
<taxonomicName LSID="72A2116B73E45EB0B650B133CA6A67F7" class="Insecta" family="Tetrigidae" genus="Lamellitettigodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lamellitettigodes novaeguineae" order="Orthoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="novaeguineae">Lamellitettigodes novaeguineae</taxonomicName>
<taxonomicNameLabel pageId="8" pageNumber="175">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Plate 2" captionStartId="F4" captionText="Plate 2. E-H (lateral view). E. Lamellitettigodes palawanicus (SMTD), ♀; F. Lamellitettigodes novaeguineae, ♀ HT; G. Lamellitettigodes novaeguineae, ♂ PT 9 / 12; H. Lamellitettigodes karwinkeli, ♀ HT. Scale bars: 1 mm." figureDoi="10.3897/jor.28.34605.plate2" httpUri="https://binary.pensoft.net/fig/330948" pageId="8" pageNumber="175" tableDoi="10.3897/jor.28.34605.plate2">Plates 2F, G</figureCitation>
,
<figureCitation captionStart="Plate 4" captionStartId="F6" captionText="Plate 4. M-P (dorsal view). M. Lamellitettigodes palawanicus (SMTD), ♀; N. Lamellitettigodes novaeguineae, ♀ HT; O. Lamellitettigodes novaeguineae, ♂ PT 9 / 12; P. Lamellitettigodes karwinkeli, ♀ HT. Scale bars: 1 mm." figureDoi="10.3897/jor.28.34605.plate4" httpUri="https://binary.pensoft.net/fig/330950" pageId="8" pageNumber="175" tableDoi="10.3897/jor.28.34605.plate4">4N, O</figureCitation>
,
<figureCitation captionStart="Plate 5" captionStartId="F7" captionText="Plate 5. Q-X (frontal view). Q. Lamellitettigodes contractus, ♀ HT; R. Lamellitettigodes sagittatus, ♀ HT; S. Lamellitettigodes signatus, ♀ HT; T. Lamellitettigodes cultratus, ♀ HT; U. Lamellitettigodes palawanicus (SMTD), ♀; V. Lamellitettigodes novaeguineae, ♀ HT; W. Lamellitettigodes novaeguineae, ♂ PT 9 / 12; X. Lamellitettigodes karwinkeli, ♀ HT. Scale bars: 1 mm." figureDoi="10.3897/jor.28.34605.plate5" httpUri="https://binary.pensoft.net/fig/330951" pageId="8" pageNumber="175" tableDoi="10.3897/jor.28.34605.plate5">5V, W</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="175" type="materials_examined">
<paragraph pageId="8" pageNumber="175">
<emphasis italics="true" pageId="8" pageNumber="175">Material examined.</emphasis>
-
</paragraph>
<paragraph pageId="8" pageNumber="175">
HT
<taxonomicName class="Insecta" family="Tetrigidae" genus="Lamellitettigodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lamellitettigodes novaeguineae" order="Orthoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="novaeguineae">
<emphasis italics="true" pageId="8" pageNumber="175">Lamellitettigodes novaeguineae</emphasis>
</taxonomicName>
: ♀, Papua New Guinea, Gulf Prov., Lakekamu Basin, Ivimka Res. Station (Malaise Trap),
<geoCoordinate degrees="7" direction="south" minutes="44" orientation="latitude" precision="925" value="-7.7333336">7°44'S</geoCoordinate>
,
<geoCoordinate degrees="146" direction="east" minutes="30" orientation="longitude" precision="925" value="146.5">146°30'E</geoCoordinate>
, 120 m, 4.-6.III.2000, leg. T. A. Sears, BMEC.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="175" type="type material">
<paragraph pageId="8" pageNumber="175">
<emphasis italics="true" pageId="8" pageNumber="175">Type material.</emphasis>
-
</paragraph>
<paragraph pageId="8" pageNumber="175">
PTs
<taxonomicName class="Insecta" family="Tetrigidae" genus="Lamellitettigodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lamellitettigodes novaeguineae" order="Orthoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="species" species="novaeguineae">
<emphasis italics="true" pageId="8" pageNumber="175">Lamellitettigodes novaeguineae</emphasis>
</taxonomicName>
: ♂, ♀ (1/12+2/12), Papua New Guinea, Gulf [Prov.], Lakekamu Basin, Ivimka Res. Station, (M[alaise]T[rap]),
<geoCoordinate degrees="7" direction="south" minutes="44" orientation="latitude" precision="925" value="-7.7333336">7°44'S</geoCoordinate>
,
<geoCoordinate degrees="146" direction="east" minutes="30" orientation="longitude" precision="925" value="146.5">146°30'E</geoCoordinate>
, 120 m, 4.-6.III.2000, leg. T. A. Sears, BMEC (deposited in ZFMK); 3♀♀ (3/12-5/12), Papua New Guinea, Gulf [Prov.], Lakekamu Basin, Ivimka Res. Station,
<geoCoordinate degrees="7" direction="south" minutes="44" orientation="latitude" precision="925" value="-7.7333336">7°44'S</geoCoordinate>
,
<geoCoordinate degrees="146" direction="east" minutes="30" orientation="longitude" precision="925" value="146.5">146°30'E</geoCoordinate>
, 120 m, 5.III.2000, leg. T. A. Sears, BMEC; ♀ (6/12), Papua New Guinea, Gulf [Prov.], Lakekamu Basin, Ivimka Res. Station, (M[alaise]T[rap]),
<geoCoordinate degrees="7" direction="south" minutes="44" orientation="latitude" precision="925" value="-7.7333336">7°44'S</geoCoordinate>
,
<geoCoordinate degrees="146" direction="east" minutes="30" orientation="longitude" precision="925" value="146.5">146°30'E</geoCoordinate>
, 120 m, 19.-23.IV.2000, leg. T. A. Sears, BMEC (deposited in BMNH); ♀ (7/12), Papua New Guinea, Gulf [Prov.], Lakekamu Basin, Ivimka Res. Station,
<geoCoordinate degrees="7" direction="south" minutes="44" orientation="latitude" precision="925" value="-7.7333336">7°44'S</geoCoordinate>
,
<geoCoordinate degrees="146" direction="east" minutes="30" orientation="longitude" precision="925" value="146.5">146°30'E</geoCoordinate>
, 120 m, 15.-20.IV.2000, leg. T. A. Sears, BMEC (deposited in ZMHU); ♀ (8/12), Papua New Guinea, Gulf [Prov.], Lakekamu Basin, Ivimka Res. Station, (M[alaise]T[rap]),
<geoCoordinate degrees="7" direction="south" minutes="44" orientation="latitude" precision="925" value="-7.7333336">7°44'S</geoCoordinate>
,
<geoCoordinate degrees="146" direction="east" minutes="30" orientation="longitude" precision="925" value="146.5">146°30'E</geoCoordinate>
, 120 m, 18.IV.2000, leg. T. A. Sears, BMEC (deposited in RMNH); ♂ (9/12), Papua New Guinea, Gulf [Prov.], Lakekamu Basin, Ivimka Res. Station,
<geoCoordinate degrees="7" direction="south" minutes="44" orientation="latitude" precision="925" value="-7.7333336">7°44'S</geoCoordinate>
,
<geoCoordinate degrees="146" direction="east" minutes="30" orientation="longitude" precision="925" value="146.5">146°30'E</geoCoordinate>
, 120 m, 18.IV.2000, leg. T. A. Sears &amp; binatung brigade, BMEC (deposited in RMNH); ♂ (10/12), Papua New Guinea, Gulf [Prov.], Lakekamu Basin, Ivimka Res. Station, (M[alaise]T[rap]),
<geoCoordinate degrees="7" direction="south" minutes="44" orientation="latitude" precision="925" value="-7.7333336">7°44'S</geoCoordinate>
,
<geoCoordinate degrees="146" direction="east" minutes="30" orientation="longitude" precision="925" value="146.5">146°30'E</geoCoordinate>
, 120 m, 26.III.2000, leg. T. A. Sears &amp; binatung brigade, BMEC; 2♀♀ (11/12+12/12), Papua New Guinea, Fly River, Kiunga, 35 m, VIII.1969, leg. J. &amp; M. Sedlacek, BPBM.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="175" type="diagnosis">
<paragraph pageId="8" pageNumber="175">
<emphasis italics="true" pageId="8" pageNumber="175">Diagnosis.</emphasis>
-
</paragraph>
<paragraph pageId="8" pageNumber="175">
The species is easily recognizable by the raised median carina with broad depression between the shoulders. This depression clearly distinguishes it from
<taxonomicName genus="L." lsidName="L. cultratus" pageId="8" pageNumber="175" rank="species" species="cultratus">
<emphasis italics="true" pageId="8" pageNumber="175">L. cultratus</emphasis>
</taxonomicName>
. It is also slightly raised in Chinese
<taxonomicName genus="L." lsidName="L. karwinkeli" pageId="8" pageNumber="175" rank="species" species="karwinkeli">
<emphasis italics="true" pageId="8" pageNumber="175">L. karwinkeli</emphasis>
</taxonomicName>
but does not reach the same height and is not lamellated. As a further feature, the fastigium tip is only slightly visible in front of the eyes.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="175" type="description">
<paragraph pageId="8" pageNumber="175">
<emphasis italics="true" pageId="8" pageNumber="175">Description.</emphasis>
-
</paragraph>
<paragraph pageId="8" pageNumber="175">Body of moderate size and slender. Head and pronotum smooth. Head and eye in lateral view weakly elevated above the pronotal discus. Dorsal margin of antennal groove slightly above the ventral margin of eye. Eyes small and in lateral view blunt and indistinctly conoidal. Antenna filiform, 14-segmented in male (including scapus and pedicel), while 15-segmented in female, 1.5-2.0 times longer than fore femur. Fastigium of vertex in dorsal view slightly wider than one eye. Fastigium in frontal view between the eyes slightly depressed. Fastigium in dorsal view with well-developed medial and lateral carinae and fossulae. Lateral carina clearly elevated, but in lateral view as high as the dorsal margin of eyes, short, not converging towards the front, bending over at nearly right angles to the medial carina. Transverse carina in dorsal view straight. Frontal costa and facial carina in lateral view minimally visible in front of eye. Tip of fastigium and frontal costa in lateral view slightly, but still clearly, protuberant. Bifurcation of frontal costa into facial carinae between dorsal margin of lateral ocelli and transverse carina, close to transverse carina by less than one diameter of ocellus. Facial carinae in lateral view clearly concave in front of eye and slightly convex in front of antenna. Last segments of maxillary palp not widened. Anterior margin of the pronotum truncated. Pronotum slightly widened between shoulders. Clearly keeled prozonal carinae converging backwards. Median carina distinctly lamellate from anterior margin of the pronotum to the end of tegmen, with a broad depression between shoulders. Interhumeral carinae absent. Internal lateral carinae weak. Pronotum behind shoulders weakly depressed on both sides of median carina, with large black spots like a broad black band (posthumeral). Lateral lobes broadly rounded, more or less close to body. Tegmen almost as long as the fore femur, rather broad, at the end broadly rounded. Macropterous and macropronotal. Alae clearly exceed the pronotum apex. Fore and middle femora short and widened, with wavy margins. Hind femora with very small antegenicular and especially genicular teeth and with some sharply tuberculated raised lobes on the middle outer surface. Posterior tibia distally not or weakly widened, with a few small spines on upper edges. First segment of the hind tarsus not longer than third segment. Pulvilli acute.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="175" type="description">
<paragraph pageId="8" pageNumber="175">
<emphasis italics="true" pageId="8" pageNumber="175">Measurements.</emphasis>
-
</paragraph>
<paragraph pageId="8" pageNumber="175">(in mm) HT: Pronotum length 12.87; pronotum lobe width 3.10; pronotum height 1.85; vertex width 0.52; eye width 0.45; tegmen length 1.50; hind wing length 10.79; postfemur length 4.68; postfemur width 1.45.</paragraph>
<paragraph pageId="8" pageNumber="175">3 PT♂♂: Pronotum length 9.88-10.40, average 10.18; pronotum lobe width 2.35-2.45, average 2.40; pronotum height 1.3-1.55, average 1.42; vertex width 0.43-0.43, average 0.43; eye width 0.39-0.43, average 0.41; tegmen length 1.20-1.25, average 1.23; hind wing length 8.97-9.75, average 9.45; postfemur length 4.00-4.10, average 4.05; postfemur width 1.15-1.20, average 1.16.</paragraph>
<paragraph pageId="8" pageNumber="175">9 PT♀♀+HT: Pronotum length 10.27-12.74, average 11.62; pronotum lobe width 2.75-3.10, average 2.96; pronotum height 1.65-1.90, average 1.82; vertex width 0.43-0.52, average 0.49; eye width 0.41-0.45, average 0.43; tegmen length 1.30-1.65, average 1.51; hind wing length 9.36-11.96, average 10.88; postfemur length 4.30-4.75, average 4.55; postfemur width 1.30-1.50, average 1.38.</paragraph>
<paragraph pageId="8" pageNumber="175">
Pictures of HT and PT are available in higher resolution in OSF (
<bibRefCitation author="Cigliano, MM" journalOrPublisher="University Studies, University of Nebraska" pageId="11" pageNumber="178" publicationUrl="http://Orthoptera.SpeciesFile.org" refId="B8" refString="Cigliano, MM, Braun, H, Eades, DC, Otte, D, 2019. . http://Orthoptera.SpeciesFile.org" url="http://Orthoptera.SpeciesFile.org" year="2019">Cigliano et al. 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="175" type="distribution">
<paragraph pageId="8" pageNumber="175">
<emphasis italics="true" pageId="8" pageNumber="175">Distribution.</emphasis>
-
</paragraph>
<paragraph pageId="8" pageNumber="175">The species is only known from lowlands in the South of New Guinea (surroundings of Fly River and Lakekamu River).</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="175" type="etymology">
<paragraph pageId="8" pageNumber="175">
<emphasis italics="true" pageId="8" pageNumber="175">Etymology.</emphasis>
-
</paragraph>
<paragraph pageId="8" pageNumber="175">
The species is named after the island of New Guinea (
<emphasis italics="true" pageId="8" pageNumber="175">Nova Guinea</emphasis>
) where it is an endemic species of the genus
<taxonomicName class="Insecta" family="Tetrigidae" genus="Lamellitettigodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lamellitettigodes" order="Orthoptera" pageId="8" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="8" pageNumber="175">Lamellitettigodes</emphasis>
</taxonomicName>
. The specific epitheton is made of two words, adjective and noun of the first (A) Latin declension, both in Genitive case (
<emphasis italics="true" pageId="8" pageNumber="175">Novae Guineae</emphasis>
=
<emphasis italics="true" pageId="8" pageNumber="175">novaeguineae</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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Von Heyden, 1826
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Von Heyden, 1826, p. 612.
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<taxonomicName id="398FDC1E81837E8219F4F01D53CDC401" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Nothrus</taxonomicName>
, Berlese, 1883a, fasc. 17 (6); 1885c, pp. 3, 49; 1885d, p. 127; 1896b, pp. 24, 28; 1913a, p. 60.
</paragraph>
</subSubSection>
<subSubSection id="C9EB7396AD50AC8A8E3B0FD3015FDB2D" type="discussion">
<paragraph id="C229750B48D9E88546DFAC026E8AC21E" pageId="64" pageNumber="65">
Von Heyden (1826) created the genus
<taxonomicName id="B9015432F5809A186EB5F9D12AC66B1F" class="Arachnida" family="Camisiidae" genus="Camisia" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Camisia</taxonomicName>
and designated
<taxonomicName id="0328A8200B3B8E99A9FC0FEBCDD863CA" class="Insecta" family="Crambidae" genus="Notaspis" kingdom="Animalia" order="Lepidoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="segnis">Notaspis segnis Hermann</taxonomicName>
as the type. Berlese did not know the name
<taxonomicName id="C1ECA046B757BA9DC135C725B33E1C3F" class="Arachnida" family="Camisiidae" genus="Camisia" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Camisia</taxonomicName>
and used
<taxonomicName id="A3E6CC251753421A15D29FA92767DCCA" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Nothrus</taxonomicName>
instead of it. The name
<taxonomicName id="F6756BC78639C940AA7076ECEAE8FE35" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Nothrus</taxonomicName>
is now used for a different genus as was mentioned above in the chapter on the family
<taxonomicName id="B8718665763321074FC6B962CEE2E24E" class="Arachnida" family="Nothridae" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="family">Nothridae</taxonomicName>
.
</paragraph>
<paragraph id="3C51DFB5EAFCEAE44150AA6A50B6F7C3" pageId="64" pageNumber="65">
Berlese (1885a) published a generic diagnosis and mentioned
<taxonomicName id="1DF840248BA42B37E4A7500B085B46DE" class="Arachnida" family="Nothridae" genus="Nothrus" genus.outerRound="1" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="bicarinatus">N. bicarinatus</taxonomicName>
(correct name:
<taxonomicName id="C4D8EA136E8B4DCAEAB205A244BD5641" class="Arachnida" family="Camisiidae" genus="Camisia" genus.outerRound="1" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="segnis">C. segnis</taxonomicName>
) as type of the genus. In the same year (Berlese, 1885c) he mentioned the following species in a key:
<taxonomicName id="5B93C5B583D804BE6AB7916F61AA3A41" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="spinifer">spinifer</taxonomicName>
,
<taxonomicName id="D79A56AB494E3C7C3B0926693949A34A" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="bicarinatus">bicarinatus</taxonomicName>
(=
<taxonomicName id="2E8E912DCAB198C5388C59AED7FE583C" class="Arachnida" family="Camisiidae" genus="Camisia" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="segnis">segnis</taxonomicName>
),
<taxonomicName id="3FA5EF07FA9C4A6B8E25E0288886123E" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="segnis">segnis</taxonomicName>
(=
<taxonomicName id="19563770DA762EBB93E1D2C06A760FF3" class="Arachnida" family="Camisiidae" genus="Camisia" genus.completionEvidence="missing" genus.completionRound="1" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="biurus">biurus</taxonomicName>
),
<taxonomicName id="CDAB5970C9D99E0F06DB66E684E53AD2" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="angulatus">angulatus</taxonomicName>
(=
<taxonomicName id="26F0DDD0C9BA372F3C0C1D2C0F448FCB" class="Arachnida" family="Camisiidae" genus="Camisia" genus.completionEvidence="missing" genus.completionRound="1" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="horrida">horrida</taxonomicName>
),
<taxonomicName id="41EAFDA408368482F31A12F9E56A484B" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="horridus">horridus</taxonomicName>
(=
<taxonomicName id="527AAA684A1D981AAE1064535BDD808E" class="Arachnida" family="Camisiidae" genus="Camisia" genus.completionEvidence="missing" genus.completionRound="1" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="biverrucata">biverrucata</taxonomicName>
); in another paper (Berlese, 1885d) he gave a short diagnosis only.
</paragraph>
<paragraph id="17AF14A90730FB91B312CAB2E9405841" pageId="64" pageNumber="65">
Berlese (1896b) mentioned a number of species, whilst later (1913a) he gave a long list, among which are species that now belong to the genera
<taxonomicName id="C4577E534E0F90B71D04E5F35A05CA86" class="Arachnida" family="Camisiidae" genus="Heminothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Heminothrus</taxonomicName>
,
<taxonomicName id="B45AE2CC13D63AC88283BF69BF0EE915" authority="Berlese, 1917" authorityName="Berlese" authorityYear="1917" class="Arachnida" family="Camisiidae" genus="Acronothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oribatida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<taxonomicName id="6DA1F8C1769A8FC7B9D5CBEF250292F7">Acronothrus</taxonomicName>
</taxonomicName>
, and
<taxonomicName id="60FD6581C4DF401AA01FDC568346BA3A" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Nothrus</taxonomicName>
(
<taxonomicName id="EC5A54EA5BA0925DE7833E97A9DAEF9C" class="Arachnida" family="Nothridae" genus="Nothrus" genus.outerRound="1" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="quadripilus">N. quadripilus</taxonomicName>
is listed in the same paper with
<taxonomicName id="BF3FE6E5853C9165B0B45D1812980444" authority="Berlese, 1885" authorityName="Berlese" authorityYear="1885" class="Arachnida" family="Lohmanniidae" genus="Angelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oribatida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Angelia</taxonomicName>
as well as with
<taxonomicName id="F8EDC7F90C468E5B0A117B1A70D77A6B" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">Nothrus</taxonomicName>
).
</paragraph>
<paragraph id="F8E8C604D86EA97D59418D1428187C8E" pageId="64" pageNumber="65">Berlese made a number of errors in the nomenclature of the species, which are dealt with below.</paragraph>
<paragraph id="2B3A38FB39A938CE0E886F1895E8025D" pageId="64" pageNumber="65">
I remark that in 1913 Berlese created a subgenus
<taxonomicName id="885A1F56576DC080F87ED072CA6C44C7" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="subGenus" subGenus="Uronothrus">Uronothrus</taxonomicName>
with
<taxonomicName id="9E884DBB1580412EAA844C981E4C4DEA" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="segnis">segnis</taxonomicName>
(read:
<taxonomicName id="677CFB9279FA7BC0AEB597C0E043D2FF" class="Arachnida" family="Camisiidae" genus="Camisia" genus.completionEvidence="missing" genus.completionRound="1" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="biurus">biurus</taxonomicName>
) as type. In the Berlese Collection
<taxonomicName id="29A52373CB25BBD9F056FCB0C1820071" class="Arachnida" family="Camisiidae" genus="Camisia" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="biurus">biurus</taxonomicName>
is indeed labelled as
<taxonomicName id="5D0E6F958A480A38617F25CEECD7D579" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="segnis" subGenus="Uronothrus">Nothrus (Uronothrus) segnis</taxonomicName>
. On the other hand, the real
<taxonomicName id="9DECAA7E0768C522AADEFA7CF20552D8" class="Arachnida" family="Camisiidae" genus="Camisia" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="segnis">segnis</taxonomicName>
and
<taxonomicName id="D4254C6ABC774F0E378EBCF93643B94B" class="Arachnida" family="Camisiidae" genus="Camisia" genus.completionEvidence="missing" genus.completionRound="1" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="horrida">horrida</taxonomicName>
are labelled as representatives of a subgenus
<taxonomicName id="D4EE1D07F3DC6ECEB553E03A0E3A2AD9" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="subGenus" subGenus="Eunothrus">Eunothrus</taxonomicName>
that has never been published. The creation of a subgenus
<taxonomicName id="B82594EE83EC812172D319C739412399" class="Arachnida" family="Nothridae" genus="Nothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="64" pageNumber="65" phylum="Arthropoda" rank="subGenus" subGenus="Uronothrus">Uronothrus</taxonomicName>
appears superfluous.
</paragraph>
</subSubSection>
</treatment>
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Berlese, 1910
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<paragraph id="E50CDB982C530D9FCF78F615E68377BE" pageId="27" pageNumber="28">
<taxonomicName id="960632C4BE6AC635F5C8BBF53A574C94" class="Arachnida" family="Protoplophoridae" genus="Protoplophora" kingdom="Animalia" order="Sarcoptiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Protoplophora</taxonomicName>
Berlese, 1910, p. 217.
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Berlese (1910) placed the genus at the base of the
<taxonomicName id="D52553CEA6AD823D580394FDDF7F3B1F" authority="(Huxley, 1864)" authorityName="(Huxley, 1864)" class="Mammalia" family="Hoplophoridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cingulata" pageId="27" pageNumber="28" phylum="Chordata" rank="family">Hoplophoridae</taxonomicName>
(the family in which at that time all ptychoid Oribatid mites had been classified), but nevertheless pointed to a relationship with the
<taxonomicName id="317E0D212CF80C2180F58FA36789BB3F" class="Arachnida" family="Hypochthoniidae" kingdom="Animalia" order="Sarcoptiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="family">Hypochthoniidae</taxonomicName>
. In the
<pageBreakToken id="82B02A621E72FE88D1B427B7D3FC90EA" pageId="28" pageNumber="29" start="start">diagnosis</pageBreakToken>
he characterizes the notogaster as divided into several parts; he distinctly figures several &quot;coupures&quot; and telescoping tergites. The long palp is remarkable, but Berlese leaves us in doubt as to the structure of rostrum and mandibles. The genus is founded on one species (
<taxonomicName id="F9F0BD05348931D4B4A3AD89AF7A12E3" class="Arachnida" family="Protoplophoridae" genus="Protoplophora" genus.outerRound="1" kingdom="Animalia" order="Sarcoptiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="palpalis">P. palpalis</taxonomicName>
) from Palermo, Sicily.
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Klotzsch ex Benth., Prodr. [A. P. de Candolle] 7(2): 659. 1839 [wfo-0000673283], nom. illeg., non Waitz, Beschreibung der Gattung und Arten der Heiden: 220. 1805 [wfo-0000673282]; Guthrie and Bolus, Fl. Capensis 4,1: 57. 1905.
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<collectingCountry id="7C4130C897F32639DFD22E610399C561" name="South Africa">South Africa</collectingCountry>
. &quot;in Strandweld [Strandveld]&quot;,
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: †B, GDC [https://plants.jstor.org/stable/10.5555/al.ap.specimen.g00465165], HBG [https://plants.jstor.org/stable/10.5555/al.ap.specimen.hbg507932], K, W [https://plants.jstor.org/stable/10.5555/al.ap.specimen.w18890186305, https://plants.jstor.org/stable/10.5555/al.ap.specimen.w18890321973, https://plants.jstor.org/stable/10.5555/al.ap.specimen.w0005951]).
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<taxonomicName class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myrmica jessensis" order="Hymenoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="jessensis">Myrmica jessensis</taxonomicName>
Forel, 1901
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<paragraph pageId="7" pageNumber="92">Doubtful species whose presence in China could not be verified.</paragraph>
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Distribution. China: Gansu (
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), Hebei (
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), Heilongjiang (
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), Hubei (Hua 2006), Hunan (
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,
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), Inner Mongolia (
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), Jilin (
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), Ningxia (Wang 2009), Shaanxi (
<bibRefCitation author="Tie, R" journalOrPublisher="Journal of Ningxia Agricultural College" pageId="39" pageNumber="124" pagination="4 - 9" title="Variety and distribution of ants in Northwest China." volume="25" year="2004">Tie and Xu 2004</bibRefCitation>
), Sichuan (
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), Xizang (
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,
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Map showing the geographic distribution records and extent of occurrence of the lizards in the Caatinga region: Norops brasiliensis, Norops fuscoauratus, Diploglossus lessonae, Ophiodes striatus, Ophiodes sp. 2, Hemidactylus agrius, Hemidactylus brasilianus, Hemidactylus mabouia, Lygodactylus klugei, Lygodactylus sp. 1, Lygodactylus sp. 2. Red circles represent data occurrence data produced in this study; Black circles represent data from literature. Ecorregions: 1: Brejos; 2: Complexo da Chapada Diamantina; 3: Complexo Ibiapaba - Araripe; 4: Depressao Sertaneja Meridional; 5: Depressao Sertaneja Setentrional; 6: Dunas do Sao Francisco; 7: Planalto da Borborema; 8: Raso Catarina and 9: Sao Francisco / Gurgueia." figureDoi="10.3897/vz.72.e78828.figure13" httpUri="https://binary.pensoft.net/fig/726821" pageId="0" pageNumber="599">Fig. 13</figureCitation>
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<paragraph pageId="0" pageNumber="599">Distribution.</paragraph>
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Caatinga endemic species. It is recorded only in the state of Bahia. It shows restricted distribution in the Caatinga and occurs along one ecoregion (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Lizards from the Caatinga. Summarized data about distribution, ecology, and conservation. Caatinga ecoregions (Silva et al. 2017) - B: Brejos, CCD: Complexo Chapada Diamantina, CIA: Complexo Ibiapaba-Araripe, DSM: Depressao Sertaneja Meridional, DSS: Depressao Sertaneja Setentorial, DSF: Dunas do Sao Francisco, PB: Planalto da Borborema, RC: Raso da Catarina, SFG: Sao Francisco - Gurgueia. Distribution Patterns - WD: widespread (EOO greater than 20,000 km 2, see Material and Methods for details), R: restricted (EOO less than 20,000 km 2). Habitat - O: open areas; F: forested areas; Preferred habitat type: AT: ant hills and termite mounds, B: bromeliads, G: general, LL: leaf litter, P: psammophilous, A: anthropic; S: saxicolous and TB: trees and bushes; Activity - D: diurnal, N: nocturnal and C: crepuscular. Conservation status (IUCN 2021; ICMBio 2018) - Critically Endangered (CR), Data Deficient (DD), Endangered (EN), Least Concern (LC), Near Threatened (NT), Not Evaluated (NE) and Vulnerable (VU). Caatinga endemic species ' * '." httpUri="http://table.plazi.org/id/DCA21B21C4ADC68A0D52968CF9844DE2" pageId="0" pageNumber="599" tableUuid="DCA21B21C4ADC68A0D52968CF9844DE2">1</tableCitation>
; Appendix S3). It occurs in high elevation areas (1,105 m a.s.l.), with annual mean temperature of 21°C, and average annual rainfall of 774 mm.
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Arboreal and diurnal
<bibRefCitation author="Vanzolini, PE" journalOrPublisher="Academia Brasileira de Ciencias, Rio de Janeiro" pageId="0" pageNumber="599" refId="B247" refString="Vanzolini, PE, Ramos-Costa, AMM, Vitt, LJ, 1980. Repteis das Caatingas. Academia Brasileira de Ciencias, Rio de Janeiro" title="Repteis das Caatingas." year="1980">(Vanzolini et al. 1980</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2018.06.010" author="Lanna, FM" journalOrPublisher="Molecular phylogenetics and evolution" pageId="0" pageNumber="599" pagination="638 - 645" refId="B105" refString="Lanna, FM, Werneck, FP, Gehara, M, Fonseca, EM, Colli, GR, Sites Jr, JW, Rodrigues, MT, Garda, AA, 2018. The evolutionary history of Lygodactylus lizards in the South American open diagonal. Molecular phylogenetics and evolution 127: 638 - 645, DOI: https://doi.org/10.1016/j.ympev.2018.06.010" title="The evolutionary history of Lygodactylus lizards in the South American open diagonal." url="https://doi.org/10.1016/j.ympev.2018.06.010" volume="127" year="2018">Lanna et al. 2018</bibRefCitation>
). It is restricted to semiarboreal/ bush vegetation in the Quaternary sand dunes of the
<normalizedToken originalValue="São">Sao</normalizedToken>
Francisco River (
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2018.06.010" author="Lanna, FM" journalOrPublisher="Molecular phylogenetics and evolution" pageId="0" pageNumber="599" pagination="638 - 645" refId="B105" refString="Lanna, FM, Werneck, FP, Gehara, M, Fonseca, EM, Colli, GR, Sites Jr, JW, Rodrigues, MT, Garda, AA, 2018. The evolutionary history of Lygodactylus lizards in the South American open diagonal. Molecular phylogenetics and evolution 127: 638 - 645, DOI: https://doi.org/10.1016/j.ympev.2018.06.010" title="The evolutionary history of Lygodactylus lizards in the South American open diagonal." url="https://doi.org/10.1016/j.ympev.2018.06.010" volume="127" year="2018">Lanna et al. 2018</bibRefCitation>
). Diet based mainly on arthropods, no information about the preferred items on the diet of this species is known. Oviparous, no detailed data is known about the number of eggs laid by the species, but it could be similar to other
<taxonomicName class="Reptilia" family="Gekkonidae" genus="Lygodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lygodactylus" order="Squamata" pageId="0" pageNumber="599" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="599">Lygodactylus</emphasis>
</taxonomicName>
species (
<bibRefCitation DOI="https://doi.org/10.1007/978-1-4612-2866-0_10" author="Vitt, LJ" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="599" refId="B253" refString="Vitt, LJ, 1992. Diversity of reproductive strategies among Brazilian lizards and snakes: the significance of lineage and adaptation. In: Hamlett WC (Ed) Reproductive biology of South American vertebrates, Springer, New York, 135-149. https://doi.org/10.1007/978-1-4612-2866-0_10" title="Diversity of reproductive strategies among Brazilian lizards and snakes: the significance of lineage and adaptation. In: Hamlett WC (Ed) Reproductive biology of South American vertebrates, Springer, New York, 135 - 149." url="https://doi.org/10.1007/978-1-4612-2866-0_10" year="1992">Vitt 1992</bibRefCitation>
,
<bibRefCitation author="Vitt, LJ" journalOrPublisher="Occasional Papers of the Oklahoma Museum of Natural History" pageId="0" pageNumber="599" pagination="1 - 29" refId="B254" refString="Vitt, LJ, 1995. The ecology of tropical lizard in the caatinga of northeast Brazil. Occasional Papers of the Oklahoma Museum of Natural History 1: 1 - 29" title="The ecology of tropical lizard in the caatinga of northeast Brazil." volume="1" year="1995">1995</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>