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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.210.3042" ID-PMC="PMC3406452" ID-Pensoft-Pub="1313-2970-210-19" ID-Pensoft-UUID="BD1FF0348E7DFFF87D580B20FFF06438" ID-PubMed="22859895" ID-Zenodo-Dep="577401" ID-ZooBank="DF32BC38E8394DC48EA0FB6508EC7B2B" ModsDocID="1313-2970-210-19" checkinTime="1451248849322" checkinUser="pensoft" docAuthor="Valdes, Pavel" docDate="2012" docId="1F3C6F01123A15FC03B3DA6623DE5867" docLanguage="en" docName="ZooKeys 210: 19-67" docOrigin="ZooKeys 210" docPubDate="2012-07-24" docSource="http://dx.doi.org/10.3897/zookeys.210.3042" docTitle="Semiardistomis maindroni" docType="treatment" docVersion="5" id="BD1FF0348E7DFFF87D580B20FFF06438" lastPageNumber="37" masterDocId="BD1FF0348E7DFFF87D580B20FFF06438" masterDocTitle="Notes about morphological features of the Western Hemisphere subtribe Ardistomina, and revision of genus Semiardistomis Kult (Coleoptera, Carabidae, Scaritinae, Clivinini)" masterLastPageNumber="67" masterPageNumber="19" pageNumber="36" updateTime="1668154243518" updateUser="ExternalLinkService">
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<mods:title>Notes about morphological features of the Western Hemisphere subtribe Ardistomina, and revision of genus Semiardistomis Kult (Coleoptera, Carabidae, Scaritinae, Clivinini)</mods:title>
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<mods:namePart>Valdes, Pavel</mods:namePart>
<mods:affiliation>Gertrudis 365 apto 5 e / D'Strampes y Goicuria, Cp 10500, Habana, Cuba</mods:affiliation>
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<taxonomicName LSID="1F3C6F01-123A-15FC-03B3-DA6623DE5867" authority="(Kult, 1950)" baseAuthorityName="Kult" baseAuthorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Semiardistomis maindroni" order="Coleoptera" pageId="17" pageNumber="36" phylum="Arthropoda" rank="species" species="maindroni">Semiardistomis maindroni (Kult, 1950)</taxonomicName>
<figureCitation captionStart="Figures 4546" captionStartId="F12" captionText="Figures 45 - 46. 45 Semiardistomis jedlickai (Kult) 46 Semiardistomis maindroni (Kult). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19670" pageId="17" pageNumber="36">Figs 46</figureCitation>
<figureCitation captionStart="Figure 57" captionStartId="F19" captionText="Figure 57. Distribution map of members of the puncticollis species-group." httpUri="https://binary.pensoft.net/fig/19677" pageId="17" pageNumber="36">57</figureCitation>
</paragraph>
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<paragraph pageId="17" pageNumber="36">
<taxonomicName authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="36" phylum="Arthropoda" rank="species" species="maindroni" subGenus="Semiardistomis">Ardistomis (Semiardistomis) maindroni</taxonomicName>
Kult, 1950: 312.
</paragraph>
<paragraph pageId="17" pageNumber="36">
<taxonomicName baseAuthorityName="Kult" baseAuthorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="17" pageNumber="36" phylum="Arthropoda" rank="species" species="maindroni">Semiardistomis maindroni</taxonomicName>
(Kult, 1950):
<bibRefCitation author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae). W. Lorenz, Tutzing, Germany" pageId="34" pageNumber="53" refId="B27" refString="Lorenz, W, 1998. Systematic list of extant ground beetles of the world (Insecta, Coleoptera &quot;Geadephaga&quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). W. Lorenz, Tutzing, Germany, 502 pp." title="Systematic list of extant ground beetles of the world (Insecta, Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl." year="1998">Lorenz 1998</bibRefCitation>
: 136.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="36" type="type material">
<paragraph pageId="17" pageNumber="36">Type material.</paragraph>
<paragraph pageId="17" pageNumber="36">
Holotype at ADVA, glued on cardboard pinned, genitalia attached in microvial, labeled: handwritten &quot;Ardistomis maindroni Kt. Det. K. Kult 1948&quot;/ &quot;maindroni Kt. det. K.Kult&quot;/ printed in red paper
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
/ printed &quot;Cuyaba Matt. Grosso&quot;.
</paragraph>
<paragraph pageId="17" pageNumber="36">Paratypes. two exemplars labeled as holotype (not checked) designated by Kult in &quot;collection of National Museum of Paris (ex Maindron- Babault)&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="37" type="type locality">
<paragraph pageId="18" pageNumber="37">
<pageBreakToken pageId="18" pageNumber="37" start="start">Type</pageBreakToken>
locality.
</paragraph>
<paragraph pageId="18" pageNumber="37">
Given by
<bibRefCitation author="Kult, K" journalOrPublisher="Arthropoda" pageId="34" pageNumber="53" pagination="299 - 325" refId="B20" refString="Kult, K, 1950. New Neotropical species of the group Ardistominina (Carabidae, Col.). Arthropoda 1: 299 - 325" title="New Neotropical species of the group Ardistominina (Carabidae, Col.)." volume="1" year="1950">Kult (1950</bibRefCitation>
: 312) as &quot;Brasilia, Matto Grosso, Cuyaba&quot;, Brazil.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="37" type="diagnosis">
<paragraph pageId="18" pageNumber="37">Diagnosis.</paragraph>
<paragraph pageId="18" pageNumber="37">Body piceous with green reflections. Elytral surface covered with isodiametric mesh pattern, striae obliterated in apical half, impunctate, 4 setiferous punctures on interval 3, stria 2 present. Profemur glabrous. Abdominal sternum VII with 4+4 setiferous punctures.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="37" type="habitus">
<paragraph pageId="18" pageNumber="37">Habitus.</paragraph>
<paragraph pageId="18" pageNumber="37">
dorsal aspect, as in
<figureCitation captionStart="Figures 4546" captionStartId="F12" captionText="Figures 45 - 46. 45 Semiardistomis jedlickai (Kult) 46 Semiardistomis maindroni (Kult). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19670" pageId="18" pageNumber="37">Fig. 46</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="37" type="measurements and variation">
<paragraph pageId="18" pageNumber="37">Measurements and variation.</paragraph>
<paragraph pageId="18" pageNumber="37">
Variation of measurements (mm) and ratios for
<taxonomicName baseAuthorityName="Kult" baseAuthorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="18" pageNumber="37" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis italics="true" pageId="18" pageNumber="37">Semiardistomis maindroni</emphasis>
</taxonomicName>
(n=9) are: HL = 0,63-0,66-0,71; PL = 1,18-1,25-1,32; PW = 1,31-1,44-1,57; EL = 3,10-3,38-3,64; EW = 2,02-2,22-2,40;
<emphasis bold="true" pageId="18" pageNumber="37">SBL =</emphasis>
4,93-5,30-5,60; PW/EW = 0,65; PW/PL = 1,15; PL/EL = 0,37; EW/EL = 0,67.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="37" type="geographical distribution">
<paragraph pageId="18" pageNumber="37">Geographical distribution</paragraph>
<paragraph pageId="18" pageNumber="37">
<emphasis bold="true" pageId="18" pageNumber="37">
(
<figureCitation captionStart="Figure 57" captionStartId="F19" captionText="Figure 57. Distribution map of members of the puncticollis species-group." httpUri="https://binary.pensoft.net/fig/19677" pageId="18" pageNumber="37">Fig. 57</figureCitation>
).
</emphasis>
The known range of this species extends from central Brazil southward of the Tropic of Capricorn to central Argentina.
</paragraph>
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<subSubSection pageId="18" pageNumber="37" type="material examined">
<paragraph pageId="18" pageNumber="37">Material examined.</paragraph>
<paragraph pageId="18" pageNumber="37">
ARGENTINA. Cordoba Arguello Nov 1958 (MHNP, 5); Tucuman Nov 1951 (MHNP, 6); Chaco prov.
<normalizedToken originalValue="Capitán">Capitan</normalizedToken>
Solari env. 3-7 Feb. 2004 (PBPC, 2). BRAZIL. Porta da Bocca am Lagua de Jigua Pernambuco Jul 1937 (BMNH, 4).
</paragraph>
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</document>

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<treatment ID-GBIF-Taxon="152044711" LSID="urn:lsid:plazi:treatment:1F3C90FBA59EDB9A7133D26E9C2D2367" httpUri="http://treatment.plazi.org/id/1F3C90FBA59EDB9A7133D26E9C2D2367" lastPageNumber="48" pageId="15" pageNumber="48">
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<paragraph pageId="15" pageNumber="48">
<taxonomicName LSID="http://coreoidea.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=2516" authority="Fabricius" class="Insecta" family="Coreidae" genus="Zoreva" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoreva dentipes" order="Hemiptera" pageId="15" pageNumber="48" phylum="Arthropoda" rank="species" species="dentipes">
<pageBreakToken pageId="15" pageNumber="48" start="start">Zoreva</pageBreakToken>
dentipes Fabricius
</taxonomicName>
Figs 29-33, 45
</paragraph>
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<subSubSection pageId="15" pageNumber="48" type="reference_group">
<paragraph pageId="15" pageNumber="48">
<taxonomicName class="Insecta" family="Coreidae" genus="Coreus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coreus dentipes" order="Hemiptera" pageId="15" pageNumber="48" phylum="Arthropoda" rank="species" species="dentipes">Coreus dentipes</taxonomicName>
<bibRefCitation pageId="15" pageNumber="48">Fabricius 1803</bibRefCitation>
: 196.
</paragraph>
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<subSubSection pageId="15" pageNumber="48" type="description">
<paragraph pageId="15" pageNumber="48">Redescription.</paragraph>
<paragraph pageId="15" pageNumber="48">Holotype. Male. n=1. Total body length: 9.8; head length: 1.1; head width: 1.6; eye width: 0.3; interocular space: 0.7; preocellar distance: 0.2; interocellar space: 0.3. Rostrum: (missing). Antennal segments length: I 2.6, II 2.4, III 1.4, IV 1.8. Pronotal length: 1.8; width: 2.8. Scutellar length: 1.3; width 1.1. Length of abdomen with hemelytra: 8.5; length of abdomen with hemelytra: 8.4; abdomen width: 2.4. Dorsal coloration: Head light brown, except anterior region of ocelli and post ocular region dark brown, with many short hairs. Antennal segments 1-2 longer than 3-4, 4 longer than 3, segment 1-3 light brown, segment 4 dark brown except base, and segment 5 light brown. Pronotum are light brown except anterior margin dark brown. Scutellum brown except edges and posterior process dark brown. Connexival: two lateral segment light brown except lateral projection dark brown. Ventral coloration: light brown except area next to conexivum dark brown to brown, light brown not uniformy red. Legs: fore and middle femora dark brown, tarsi light brown with light brown spots. Mid femur with one spine distally. Fore femur dark brown with spines, basally very long, medially short and distally long. Structure: pronotum granulose; frontal angles granulose and projecting as acute spines reaching ocular tubercles; humeral angles with two rounded projections and shorter spines; scutellum granulose with pilosity. Hemelytra as long as the abdomen, brown with punctuations and pilosity, membrane dark brown.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="48" type="specimen examined">
<paragraph pageId="15" pageNumber="48">Specimen examined.</paragraph>
<paragraph pageId="15" pageNumber="48">
Argentina: Formosa: 1♂ Laguna Oca (
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-26.232292">26°13'56.25&quot;S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-58.21801">58°13'04.84&quot;W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="48" type="distribution">
<paragraph pageId="15" pageNumber="48">Distribution.</paragraph>
<paragraph pageId="15" pageNumber="48">Argentina: Formosa: Laguna Oca.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="48" type="remarks">
<paragraph pageId="15" pageNumber="48">Remarks.</paragraph>
<paragraph pageId="15" pageNumber="48">This is the first record of this species for Argentina</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="1AAC2DC6E45607E3D0872F9EE0BC1327" italics="true" pageNumber="966">Euphrasia hirtella</emphasis>
Reut.
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<subSubSection id="1083A09BAA96DAC05264E9B8F0650018" pageNumber="966" type="description">
<paragraph id="72FAA68C735752606C6656130848A3DA" pageNumber="966">
Artbeschreibung:
<quantity id="BD7B33FE783DFB08C4FC4A91DBAE6DB2" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" pageNumber="952" unit="cm" value="15.0" valueMax="25.0" valueMin="5.0">5-25 cm</quantity>
hoch,
<emphasis id="FF765788902677E84D2F8AD174F3F069" bold="true" pageNumber="966">meist unverzweigt</emphasis>
, seltener unten mit wenigen kurzen
<normalizedToken id="F2EF2D85EF0D607F5E8C0B9C97D88AD9" originalValue="Ästen" pageNumber="952">Aesten</normalizedToken>
,
<emphasis id="F7E32A9AAFB178E1A63FFEF05D64CB00" bold="true" pageNumber="966">
oben reichlich mit
<normalizedToken id="A6FB2A7E23FF1C78E1F6C29DC78919D5" originalValue="drüsenlosen" pageNumber="952">druesenlosen</normalizedToken>
und meist auch
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Haaren.
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meist etwa so lang wie breit
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, die mittleren und oberen jederseits mit 3-6 breiten und spitzen, unbegrannten
<normalizedToken id="4DB9F8732591A6C1103B30F86BD56B67" originalValue="Zähnen" pageNumber="952">Zaehnen</normalizedToken>
.
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Krone
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lang, weiss
</emphasis>
, oft mit
<normalizedToken id="C90CC53C9531BB8D0B5AC0B477D329AC" originalValue="bläulicher" pageNumber="952">blaeulicher</normalizedToken>
Oberlippe und gelbem Schlundfleck. Frucht so lang wie der Kelch oder
<normalizedToken id="B5E176C8EBE3E2C97F5FD5C07CC26D7D" originalValue="länger" pageNumber="952">laenger</normalizedToken>
, behaart, kaum doppelt so lang wie breit.
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<paragraph id="62A6C97EC858931C71D1C0179D41D8EA" pageNumber="966">
<normalizedToken id="748D3074F0A7752F17BE88A4B1E62149" originalValue="Blütezeit" pageNumber="952">Bluetezeit</normalizedToken>
: 6-9
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Standort und Verbreitung in der Schweiz: Weiden, trockene
<normalizedToken id="479022C1E477ECC5E6BDC7466000B408" originalValue="Böden" pageNumber="952">Boeden</normalizedToken>
in
<normalizedToken id="765B6FB3B88509DD6526FC13DE55A8C2" originalValue="wärmeren" pageNumber="952">waermeren</normalizedToken>
Lagen / subalpin(-alpin) / A
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<paragraph id="14731D163755E66377DE8C653BECAECB" pageNumber="966">Verbreitung global: Arktisch-alpin</paragraph>
</subSubSection>
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<paragraph id="9F2138E2689D8F39BD137BC033C850AE" pageNumber="966">
<normalizedToken id="96581192A5CFE9391C30F191E3A662F1" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="DC0FECB2517615FADDEBC3896CBB7957" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." pageNumber="966" publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
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<tr id="480EB511A9ADA747838129A3D658FE90" pageNumber="966">
<th id="C58E9A52CA9F61145EBAF68625E40463" colspan="2" pageNumber="966">
<emphasis id="0ACEAAAB84614456B95FC6D2CE2017F7" bold="true" pageNumber="966">Bodenfaktoren</emphasis>
</th>
<th id="589EFFD55D16FF9F2616EE95D5B4C945" colspan="2" pageNumber="966">
<emphasis id="ABA176299CB37377DA43B06983884CC2" bold="true" pageNumber="966">Klimafaktoren</emphasis>
</th>
<th id="3B365A9078D9491A79988FCFBBF808DB" colspan="2" pageNumber="966">
<emphasis id="4EE0E7F98F8B9A16521B6B0D9130FCC9" bold="true" pageNumber="966">Salztoleranz</emphasis>
</th>
</tr>
<tr id="2FD279DB679891CC3ED9F60A2EF00B16" pageNumber="966">
<td id="478DE3440A94267526D21DF53B6D6A5C" pageNumber="966">Feuchtezahl F</td>
<td id="EFB4EC7291AD833A74F353AEA01B7F1A" pageNumber="966" style="text-align:left;width:100px">
<normalizedToken id="1F74CBBD3091380A47CA2F35BB1EE52B" originalValue="mässig">maessig</normalizedToken>
trocken
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<td id="128792FDABCC8163F51EB9386E7729A6" pageNumber="966">Lichtzahl L</td>
<td id="7EAA4B11B8C89AC5398BB4B43E382399" pageNumber="966" style="text-align:left;width:60px">hell</td>
<td id="BC292A42129613EF0B0F79AFF3756044" pageNumber="966">Salzzeichen</td>
<td id="43D196EA82BCB8B86E14AFB9549A83F5" pageNumber="966" style="text-align:left;width:60px">--</td>
</tr>
<tr id="3566D12DAF746A2875477B8A39586E32" pageNumber="966">
<td id="0C406D068152E8EF7248D64564CA8328" pageNumber="966">Reaktionszahl R</td>
<td id="5D509F041F42BBD913EB4C7E4E8ED8EF" pageNumber="966" style="text-align:left">schwach sauer bis neutral (pH 4.5-7.5)</td>
<td id="5CB00F8B243DCDF3A8CEF52C3787AD6E" pageNumber="966">Temperaturzahl T</td>
<td id="C41BC2D01F1364500C4BAE9DE88591EC" pageNumber="966" style="text-align:left">
subalpin (
<normalizedToken id="EDD15A50B0E8804BB1A8E86944BADD3B" originalValue="Fichtenwälder">Fichtenwaelder</normalizedToken>
ohne Buchen bis zur Obergrenze der Fichte)
</td>
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<td id="6C8E3B51659B12232C36FCCE9604F138" pageNumber="966">
<normalizedToken id="3DA3CCD94BCCD6761CC37962C4023F04" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
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<normalizedToken id="85202F7CA984431042275D9159230EAF" originalValue="nährstoffarm">naehrstoffarm</normalizedToken>
</td>
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<normalizedToken id="CD51AD516BE4016A544E1569A7ED91BF" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="39098EEC4B297BA7349A4DF6883C484D" pageNumber="966" style="text-align:left">subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter)</td>
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</table>
</paragraph>
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<paragraph id="C46440933E4AF9680C62DA213D4A2B8B" pageNumber="966">
Volksname Deutscher Name:
<vernacularName id="695B6AFAAB037EB32CBF015C8837750B" language="deu" pageNumber="966">Zottiger Augentrost</vernacularName>
Nom
<normalizedToken id="55850125D18922437DDA0220E4AF4251" originalValue="français">francais</normalizedToken>
:
<vernacularName id="283274633FBF709633506DD345E373CD" language="fra" pageNumber="966">
Euphraise
<normalizedToken id="209FB561723A4308836BA4757CE14E70" originalValue="hérissée">herissee</normalizedToken>
</vernacularName>
Nome italiano:
<vernacularName id="E7D6BD66E62952AA7765AA30D725CBE2" language="ita" pageNumber="966">Eufrasia pelosetta</vernacularName>
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<mods:url id="824999E2EEC100020B79F25CAD65ECCE">http://dx.doi.org/10.3897/zookeys.390.6661</mods:url>
</mods:location>
<mods:classification id="04D44F36C93719735568391A846E62B4">journal article</mods:classification>
<mods:identifier id="DAD9217ECA17B3C27F1A09D6246B3CD0" type="DOI">http://dx.doi.org/10.3897/zookeys.390.6661</mods:identifier>
<mods:identifier id="A38CDE6B08F4137CF6B125E0749294B9" type="Pensoft-Pub">1313-2970-390-1</mods:identifier>
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</mods:mods>
<treatment id="1F3CD9205D63CE8D2F878134647D9933" ID-GBIF-Taxon="152051690" LSID="urn:lsid:plazi:treatment:1F3CD9205D63CE8D2F878134647D9933" httpUri="http://treatment.plazi.org/id/1F3CD9205D63CE8D2F878134647D9933" lastPageId="36" lastPageNumber="37" pageId="32" pageNumber="33">
<subSubSection id="C1F6F2AA1471C50E84DD3FE10F11CA8F" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="A625CEB268F95A1F49A47276F100737B" pageId="32" pageNumber="33">
<taxonomicName id="CFADBCE8BF674510959EB03373DE1D12" ID-CoL="3VNVK" LSID="http://species-id.net/wiki/Liturgusa_lichenalis" authority="Gerstaecker, 1889" authorityName="Gerstaecker" authorityYear="1889" class="Insecta" family="Liturgusidae" genus="Liturgusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liturgusa lichenalis" order="Mantodea" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lichenalis">Liturgusa lichenalis Gerstaecker, 1889</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="053FD29D8E4046378A57F7D6869AC525" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="503A1F5B65974CD3E49B476BBB8CD823" pageId="32" pageNumber="33">
<taxonomicName id="5F6CE97F7FA25B3D510313A8864E1B6A" class="Insecta" family="Liturgusidae" genus="Liturgusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liturgusa lichenalis" order="Mantodea" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lichenalis">Liturgusa lichenalis</taxonomicName>
:
<bibRefCitation id="C0135057BE22909C6F4A2261DD52DA1C" author="Gerstaecker, KEA" journalOrPublisher="Mittheilunger aus dem Naturwissenschaftlichen Verein Fuer Neu-Vorpommern und Ruegen in Greifswald" pageId="148" pageNumber="149" pagination="43 - 100" title="Charakteristik einer Reihe bemerkenswerther Orthopteren." volume="20" year="1889">Gerstaecker 1889</bibRefCitation>
: 52-53;
<bibRefCitation id="118EFC6F4DA3C5212B38A4136FADC13F" author="Bertkau, P" journalOrPublisher="Archiv fuer Naturgeschichte" pageId="147" pageNumber="148" pagination="1 - 264" title="Bericht ueber die wissenschaftlichen Leistungen im Gebiet der Entomologie waehrend des Jahres 1888." volume="55" year="1889">Bertkau 1889</bibRefCitation>
: 87;
<bibRefCitation id="1DC2E23AC18EE3EA42BF3BC9108A498C" author="Terra, PS" journalOrPublisher="Revista Brasileira de Zoologia" pageId="152" pageNumber="153" pagination="13 - 94" title="Revisao Sistematica Dos Generos De Louva-A-Deus Da Regiao Neotropical (Mantodea)." volume="39" year="1995">Terra 1995</bibRefCitation>
: 54;
<bibRefCitation id="0A292E7CA033B7EAC74217621D0A62C4" author="Ehrmann, R" journalOrPublisher="Natur und Tier-Verlag GmbH, Muenster" pageId="148" pageNumber="149" title="Mantodea: Gottesanbeterinnen der Welt." year="2002">Ehrmann 2002</bibRefCitation>
: 207;
<bibRefCitation id="36A0799B43C518F8CF21D51E619CEDA3" author="Otte, D" journalOrPublisher="Association of Insect Diversity, Philadelphia" pageId="150" pageNumber="151" title="Mantida Species File" volumeTitle="Catalog of the Mantids of the World" year="2005">Otte and Spearman 2005</bibRefCitation>
: 133;
<bibRefCitation id="9EDF9EB8354FA5E6CC66338843CBD166" author="Agudelo, RAA" journalOrPublisher="Biota Colombiana" pageId="147" pageNumber="148" pagination="105 - 158" title="Checklist of the Neotropical mantids (Insecta, Dictyoptera, Mantodea)." volume="8" year="2007">Agudelo et al. 2007</bibRefCitation>
: 116.
</paragraph>
<paragraph id="D3ADD3A2669678AB3DB5C8AE3578B9DF" pageId="32" pageNumber="33">
<taxonomicName id="E0BEDE98EFF381AAAE693DB3B693CB22" class="Insecta" family="Liturgusidae" genus="Liturgousa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Liturgousa lichenalis" order="Mantodea" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lichenalis">Liturgousa lichenalis</taxonomicName>
:
<bibRefCitation id="B45386E4372177063AE7CBB16A3FD7C6" author="Westwood, JO" journalOrPublisher="Gurney and Jackson, London" pageId="152" pageNumber="153" title="Revisio Insectorum Familiae Mantidarum, speciebus novis aut minus cognitis descriptis et delineatis." year="1889">Westwood 1889</bibRefCitation>
: 5, 51;
<bibRefCitation id="DD829D49A3FE850A0E22F6EC046FE6B7" author="Kirby, WF" journalOrPublisher="British Museum, Nat. Hist., London" pageId="150" pageNumber="151" title="A synonymic Catalogue of Orthoptera. I. Orthoptera Euplexoptera, Cursoria et Gressoria)" year="1904">Kirby 1904</bibRefCitation>
: 271;
<bibRefCitation id="B637393E48412CA98D57C56414F215A0" author="Caudell, A" journalOrPublisher="Insecutor Inscitiae Menstruus" pageId="148" pageNumber="149" pagination="1 - 70" title="On a collection of Orthoptera (Exclusive of the Locustidae) made in Central Peru by N. Iconnicoff and C. SCHUNKE." volume="6" year="1918">Caudell 1918</bibRefCitation>
: 5;
<bibRefCitation id="4DE7080B1F21FD1126FBEDADF4FD24E5" author="Rehn, JAG" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="150" pageNumber="151" pagination="167 - 271" title="The Orthoptera of Costa Rica. Part I. - Mantidae." volume="87" year="1935">Rehn 1935</bibRefCitation>
: 199;
<bibRefCitation id="A9FCCAABC9F6154CF9DEBB245C1A3A33" author="Rehn, JAG" journalOrPublisher="Transactions of the American Entomological Society" pageId="151" pageNumber="152" pagination="175 - 184" title="On Some Gerstaecker Types of Neotropical Orthoptera. I. Manteidae." volume="80" year="1954">Rehn 1954</bibRefCitation>
: 179, pl. 1, fig. 1.
</paragraph>
<paragraph id="30ED5F8DEEC69AFFC980A47A6199A6A4" pageId="32" pageNumber="33">
<taxonomicName id="5512F5239DCE9335F49CD187EF6EBD79" class="Insecta" family="Liturgusidae" genus="Liturgusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liturgusa annulipes" order="Mantodea" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="annulipes">Liturgusa annulipes</taxonomicName>
(partim):
<bibRefCitation id="D426424562FACA4062E80D80EB830862" author="Giglio-Tos, E" journalOrPublisher="Walter de Gruyter &amp; Co., Berlin" pageId="148" pageNumber="149" title="Das Tierreich" year="1927">Giglio-Tos 1927</bibRefCitation>
: 294;
<bibRefCitation id="207FE16FE851B7B39F03E2759C82665F" pageId="32" pageNumber="33">Beier 1935</bibRefCitation>
: 11;
<bibRefCitation id="C8FE2BCA4E588B6791A0DB2A0F0DE237" author="Jantsch, LJ" journalOrPublisher="PUCRS, Doutorado, Porto Alegre" pageId="149" pageNumber="150" title="Estudos filogene ́ ticos em Manto ́ deos americanos (Insecta: Pterygota: Mantodea)." year="1999">Jantsch 1999</bibRefCitation>
: 48.
</paragraph>
</subSubSection>
<subSubSection id="F4A9310C9F67D09B92DC2843806F4D7A" pageId="32" pageNumber="33" type="type">
<paragraph id="4705A92F02F35426C6D225C013C1ED24" pageId="32" pageNumber="33">Type.</paragraph>
<paragraph id="C21CFCFA95C93ABEFA577DCF567EC4C5" pageId="32" pageNumber="33">
Holotype Female.
<normalizedToken id="EE40A966B4E039216AAB9BB65A74118C" originalValue="Ernst-Moritz-Arndt-Universität">Ernst-Moritz-Arndt-Universitaet</normalizedToken>
Greifswald, Germany.
</paragraph>
</subSubSection>
<subSubSection id="7B3AC25D0C40744D74F3E5FA59904D4B" pageId="32" pageNumber="33" type="type locality">
<paragraph id="375C49A08362F965750EB39763D0A458" pageId="32" pageNumber="33">Type locality.</paragraph>
<paragraph id="4455186C25FF7FBE4B4AC4A082F8812B" pageId="32" pageNumber="33">Peru: Jurimaguas (Lat. -5.900438, Long. -76.125298).</paragraph>
</subSubSection>
<subSubSection id="23A76F9F8AC092CD4A1779D3800A7C24" pageId="32" pageNumber="33" type="material examined">
<paragraph id="DC0F38FDF0263D6CFE7ABF24FC2D75D5" pageId="32" pageNumber="33">Material examined.</paragraph>
<paragraph id="06C4F9AFBB7DB19C67075F91BAFD2AAB" pageId="32" pageNumber="33">
<taxonomicName id="ED6EA697DFCB9B521E0E21385F49402B" class="Insecta" family="Liturgusidae" genus="Liturgusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liturgusa lichenalis" order="Mantodea" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="lichenalis">Liturgusa lichenalis</taxonomicName>
Gerstaecker, 1889
</paragraph>
<paragraph id="4DCB777911A60B60AC0066CE1F5258BF" pageId="32" pageNumber="33">
<table id="980F15FBED0D276D770353683570FB1A" pageId="32" pageNumber="33">
<tr id="E06F60857DFF8DB7834EFEBC94FD4AD0" pageId="32" pageNumber="33">
<th id="864C49AF2433C651CE90AC03CFF9F3A1" colspan="1" pageId="32" pageNumber="33" rowspan="1">Sex</th>
<th id="0D35AB810E939B23666D05461CA904F9" colspan="1" pageId="32" pageNumber="33" rowspan="1">Type</th>
<th id="8DA7A5C947B4A9600B81878A85CA8516" colspan="1" pageId="32" pageNumber="33" rowspan="1">Country</th>
<th id="717F448615804592F6E8CFC5CBBB7DF9" colspan="1" pageId="32" pageNumber="33" rowspan="1">Label</th>
<th id="59445DAEC8AC7B1AE242FAE4885219D9" colspan="1" pageId="32" pageNumber="33" rowspan="1">Latitude Longitude</th>
<th id="2425B2D1A63138D154E5703C372BFB24" colspan="1" pageId="32" pageNumber="33" rowspan="1">Code</th>
</tr>
<tr id="2FA8F3814FA817AFEB4A533F2764DD1B" pageId="32" pageNumber="33">
<td id="240C7D9F4FAB68928DA9198C9BD1C9FB" colspan="1" pageId="32" pageNumber="33" rowspan="1">-5.900438, -76.125298</td>
</tr>
<tr id="69672787B87B2766CC296C0E4D8D7C62" pageId="32" pageNumber="33">
<td id="C5E7928E4DEF716D97E94582A33BBAF3" colspan="1" pageId="32" pageNumber="33" rowspan="1">-9.314153, -76.006745</td>
</tr>
<tr id="5638F7B30EACD7A8907961E80E6F6C1F" pageId="32" pageNumber="33">
<td id="66AD62D9AA63A3BE98C44FFD80E895D0" colspan="1" pageId="32" pageNumber="33" rowspan="1">-9.314153, -76.006745</td>
</tr>
<tr id="7F389E5AC768F5897C029CE01BEA9AA6" pageId="32" pageNumber="33">
<td id="CB5DDD43C2B513B11131DD25FD9B57A6" colspan="1" pageId="32" pageNumber="33" rowspan="1">-9.314153, -76.006745</td>
</tr>
<tr id="AF8ACFC7DEC78B1D61D32FDE7EE798CC" pageId="32" pageNumber="33">
<td id="DE5750E9F55FE763E5213B450EF7F284" colspan="1" pageId="32" pageNumber="33" rowspan="1">-11.135912, -75.348800</td>
</tr>
<tr id="794AC9A5720FC250F9BFFE4BBD03F8D9" pageId="32" pageNumber="33">
<td id="192DF2EA128A57BAD55F0A358B4F0DE6" colspan="1" pageId="32" pageNumber="33" rowspan="1">-1.197385, -72.937475</td>
</tr>
<tr id="F6139A233ACD46ABCFF8DBECA59796EE" pageId="32" pageNumber="33">
<td id="3706F0C88ABDD5244083D1F401C6F598" colspan="1" pageId="32" pageNumber="33" rowspan="1">-3.617778, -73.262222</td>
</tr>
<tr id="AA1F5DC841BCD51395B545FDFD1259E1" pageId="32" pageNumber="33">
<td id="D6998E757263BF50B1F6E778ADC40607" colspan="1" pageId="32" pageNumber="33" rowspan="1">-0.133333, -77.866667</td>
</tr>
<tr id="B0F272FD6085E878D9F394727DA23B89" pageId="32" pageNumber="33">
<td id="F2B30F1DB12B5EDB060E3EB0BCBD2514" colspan="1" pageId="32" pageNumber="33" rowspan="1">3.198590, -65.555990</td>
</tr>
<tr id="9AA1E24A6955B27B7145C275E439A783" pageId="32" pageNumber="33">
<td id="585E679BF4320FF42A3A08BE5D531649" colspan="1" pageId="32" pageNumber="33" rowspan="1">-11.050000, -75.283333</td>
</tr>
<tr id="A0AA7A4A0B40CF0413B92D1600E17F67" pageId="32" pageNumber="33">
<td id="8191F9AB3A0BCCE9C0443D614EC0B992" colspan="1" pageId="32" pageNumber="33" rowspan="1">-12.833333, -69.283333</td>
</tr>
<tr id="195E5E4BCA7E61D1F7D4E72D01635372" pageId="32" pageNumber="33">
<td id="7B77D6C80F8F87B72BA86E93E7A28364" colspan="1" pageId="32" pageNumber="33" rowspan="1">-12.833333, -69.283333</td>
</tr>
<tr id="50E306ACA37A8CF471A181736D459EA7" pageId="32" pageNumber="33">
<td id="BF4C49C0D4900180CFFF855FCB0EBD51" colspan="1" pageId="32" pageNumber="33" rowspan="1">-12.833333, -69.283333</td>
</tr>
<tr id="2EC6F1EA2FB56F49DCCF65CA8A1D155D" pageId="32" pageNumber="33">
<td id="24D1A6059F4F0F02877A46E8C3015E1C" colspan="1" pageId="32" pageNumber="33" rowspan="1">-12.833333, -69.283333</td>
</tr>
<tr id="4F258560637C19DF58E924C11F32554B" pageId="32" pageNumber="33">
<td id="55489BA195D878F59877C0A2771D883B" colspan="1" pageId="32" pageNumber="33" rowspan="1">-12.833333, -69.283333</td>
</tr>
<tr id="766ADCBD3B712B541EA35447CF7885FC" pageId="32" pageNumber="33">
<td id="66138EEDD423E768B122FAC4F6C4FDCB" colspan="1" pageId="32" pageNumber="33" rowspan="1">-12.833333, -69.283333</td>
</tr>
<tr id="869A872CF0F000237533DE8D80DCB8C5" pageId="32" pageNumber="33">
<td id="81DCB2F73611870AB0368FEB4D7B18FB" colspan="1" pageId="32" pageNumber="33" rowspan="1">-12.833333, -69.283333</td>
</tr>
<tr id="5D3B71358C3CA6EB8EC3E02887E5171F" pageId="32" pageNumber="33">
<td id="C757E5989C39576F41C508F198AA17EE" colspan="1" pageId="32" pageNumber="33" rowspan="1">-12.833333, -69.283333</td>
</tr>
<tr id="40C87A4BEB8907A9D2FC745C8EA82E85" pageId="32" pageNumber="33">
<td id="C4938C42828F513F971E1BF855AACB07" colspan="1" pageId="32" pageNumber="33" rowspan="1">-12.833333, -69.283333</td>
</tr>
<tr id="C4C4751495D6ECDF5E9C0EA46E12E8BA" pageId="32" pageNumber="33">
<td id="35E7F085FFD6D364C0C02DABDF30F4B5" colspan="1" pageId="32" pageNumber="33" rowspan="1">-12.833333, -69.283333</td>
</tr>
<tr id="4B52915FF09CDDBBBF2440637087FF89" pageId="32" pageNumber="33">
<td id="EF67DB4184EF06217165B3B050FEE9AC" colspan="1" pageId="32" pageNumber="33" rowspan="1">-2.753512, -77.444899</td>
</tr>
<tr id="8B6D6ED6AF51F826319B09D6B4CCFAF7" pageId="32" pageNumber="33">
<td id="17FD6D90F2F723253120574E48F74092" colspan="1" pageId="32" pageNumber="33" rowspan="1">-3.894566, -71.971608</td>
</tr>
<tr id="C2B2575C93BF39CFEB65152DA85608EA" pageId="32" pageNumber="33">
<td id="1D037665DF36D1924790F74C3C7BA022" colspan="1" pageId="32" pageNumber="33" rowspan="1">-5.509011, -74.377603</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="79B76024D62EE72834297AE6CFDA3D41" pageId="33" pageNumber="34" type="taxonomic history">
<paragraph id="0582C3DB1BEE04F0E423E8015920FAC4" pageId="33" pageNumber="34">
<pageBreakToken id="FA826D50AA304895F5B959BDB4796893" pageId="33" pageNumber="34" start="start">Taxonomic</pageBreakToken>
history.
</paragraph>
<paragraph id="01FD7078CC66FF5457D3EE4E553D0966" pageId="33" pageNumber="34">
Described in 1889 by Gerstaecker, the species has been largely ignored other than inclusion in taxonomic lists and some regional studies.
<bibRefCitation id="0BFD2F3CDCEC0E1F4B74108F27B5C695" author="Giglio-Tos, E" journalOrPublisher="Walter de Gruyter &amp; Co., Berlin" pageId="148" pageNumber="149" title="Das Tierreich" year="1927">Giglio-Tos (1927)</bibRefCitation>
considered
<taxonomicName id="4C02343532F8CD150127830831697675" class="Insecta" family="Liturgusidae" genus="Liturgusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liturgusa lichenalis" order="Mantodea" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="lichenalis">Liturgusa lichenalis</taxonomicName>
as the synonym of
<taxonomicName id="14776D3AA67DEC0EB7A6713E11D8FA50" class="Insecta" family="Liturgusidae" genus="Fuga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fuga annulipes" order="Mantodea" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="annulipes">Fuga annulipes</taxonomicName>
, but this treatment is erroneous as it is apparent that no previous work had an accurate concept of
<taxonomicName id="A5CB62BA0EFFCD3110B6C46868A800E7" class="Insecta" family="Liturgusidae" genus="Fuga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fuga annulipes" order="Mantodea" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="annulipes">Fuga annulipes</taxonomicName>
.
<bibRefCitation id="464948F9625A02AF913538636647BA95" author="Rehn, JAG" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="150" pageNumber="151" pagination="167 - 271" title="The Orthoptera of Costa Rica. Part I. - Mantidae." volume="87" year="1935">Rehn (1935)</bibRefCitation>
corrected this erroneous synonym by comparing the types of both species.
</paragraph>
</subSubSection>
<subSubSection id="379D08145883DC81815CEE0E0C090BF6" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="5F1597DFB9383347A23577EBE0AC0C34" pageId="33" pageNumber="34">Diagnosis.</paragraph>
<paragraph id="B58E4C94B8A4DD356D449E1C1E5ADBDA" pageId="33" pageNumber="34">
Almost identical to
<taxonomicName id="D41AF15E07FA7A12D36A56B39F1E213D" class="Insecta" family="Liturgusidae" genus="Liturgusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liturgusa cayennensis" order="Mantodea" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cayennensis">Liturgusa cayennensis</taxonomicName>
, but primarily distributed in the western Amazon basin rather than in the Guyanas in the northeastern coastal region of South America. In addition, the abdomens of males and females are broader than
<taxonomicName id="3255CDA87F008351AFF0AF500CC8B783" class="Insecta" family="Liturgusidae" genus="Liturgusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liturgusa cayennensis" order="Mantodea" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cayennensis">Liturgusa cayennensis</taxonomicName>
and have posterolateral tergal projections in the distal half. Another distinguishing external character for
<taxonomicName id="EB76F2548396CD3964D5A51E194C5D31" class="Insecta" family="Liturgusidae" genus="Liturgusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liturgusa lichenalis" order="Mantodea" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="lichenalis">Liturgusa lichenalis</taxonomicName>
comparison to
<taxonomicName id="1F0D6BB39423E6245543281AC48F6F55" class="Insecta" family="Liturgusidae" genus="Liturgusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liturgusa cayennensis" order="Mantodea" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cayennensis">Liturgusa cayennensis</taxonomicName>
is that the first segment of the mesotarsi is obviously shorter than the remaining segments combined while this segment is the same length as the remaining segments combined in
<taxonomicName id="427735629A0500CF333FC6B2F4A9FE42" class="Insecta" family="Liturgusidae" genus="Liturgusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liturgusa cayennensis" order="Mantodea" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cayennensis">Liturgusa cayennensis</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F8050CEBE8EC56C2558DBEBBE5A1C8DF" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34" type="description">
<paragraph id="40DD804710282FC98AC270B23593EB8B" pageId="33" pageNumber="34">Description.</paragraph>
<paragraph id="1E85D52DE2F7FB811E94725D92534F96" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
Male. (Fig. 5A) N=12: Body length 18.59-24.79 (21.45); forewing length 12.62-16.54 (14.17); hindwing length 10.27-11.75 (11.01); pronotum length 5.03-6.59 (5.74); prozone length 1.55-2.07 (1.78); pronotum width 2.33-3.18
<pageBreakToken id="F92CA6395CB429FA0653BDB64D7E284B" pageId="34" pageNumber="35" start="start">(</pageBreakToken>
2.68); pronotum narrow width 1.78-2.47 (2.07); head width 4.54-5.75 (5.03); head vertex to clypeus 1.81-2.03 (1.91); frons width 1.66-1.93 (1.78); frons height 0.59-0.75 (0.67); prothoracic femur length 5.60-7.14 (6.19); mesothoracic femur length 6.37-8.50 (7.38); mesothoracic tibia length 4.78-6.27 (5.52); mesothoracic tarsus length 4.77-6.25 (5.32); metathoracic femur length 6.71-8.92 (7.55); metathoracic tibia length 6.82-8.68 (7.64); metathoracic tarsus length 6.41-8.40 (7.34); pronotal elongation measure 0.30-0.33 (0.31); pronotal shape measure 0.44-0.49 (0.47); head shape measure 0.38-0.40 (0.39); frons shape measure 0.35-0.40 (0.38); anteroventral femoral spine count 13-17 (15); anteroventral tibial spine count 10; posteroventral tibial spine count 7.
</paragraph>
<caption id="5F9E52E0460CB06E699EB92E5C040A91" pageId="34" pageNumber="35">
<paragraph id="F5F29F40C01B670BFC4A1646861F6C1E" pageId="34" pageNumber="35">
Figure 5.
<taxonomicName id="A7EC20C31A84ACCDDE7AC377CCC8973F" class="Insecta" family="Liturgusidae" genus="Liturgusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liturgusa lichenalis" order="Mantodea" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lichenalis">Liturgusa lichenalis</taxonomicName>
Gerstaecker, 1889, dorsal habitus: A male from Tingo Maria, Peru (CAS 010) B female from Ecuador (MNHN 203).
</paragraph>
</caption>
<paragraph id="740F54909977FE75F7F06E7449956037" pageId="34" pageNumber="35">Head (Fig. 40C): Transverse, the juxta-ocular protuberances small, but pronounced, the apex slightly in the lateral half; the vertex is straight, even with the dorsal margin of the eyes. Frontal suture with a medial carina forming a continuous arc, most pronounced medially, the region just ventral depressed. Ocelli small and protruding on small cuticular mounds, but the region between all three slightly raised; the lateral ocelli oriented outward. The carina on the frons present, but not highly pronounced, the medial region just ventral to the carina depressed. Clypeus transverse, the upper margin convex, the lower margin slightly concave; the central carina strongly pronounced and straight. Antennae pale at the base, the flagellum fading gradually to dark brown around a third of the way from the base. Black band extending straight over the medial carina of the frontal suture, the medial portion of the carina pale; a branch of the black band extends ventro-laterally between the eye and antennal insertion at a forty five degree angle; area around parietal sutures pale or dark brown. Lower region of frons darkened; the clypeus is pale; the mandibles and labrum mostly pale, but with some brown marks; the vertex and juxta-ocular protuberances mostly pale; the area immediately adjacent to lateral ocelli black. Palpi are pale.</paragraph>
<paragraph id="64687CD310B46D8C038D53BA0E132926" pageId="34" pageNumber="35">Pronotum (Fig. 47C): Short and squat with a poorly defined supra-coxal bulge; dorsal surface entirely smooth or with few blunt tubercles in the posterior half. Prozone squat with near parallel margins before tapering anteriorly to a rounded anterior margin; the margins smooth. Metazone with barely concave lateral margins, mostly tapering consistently from the supra-coxal bulge posteriorly until a very slight widening in the posterior quarter leading to the posterior margin; margins smooth; posterior margin with a slight medial emargination; the dorsal surface of the posterior half of the metazone depressed; two small and elongate bulges barely present on the dorsal surface near the posterior margin and positioned laterally, sometimes absent. Pale with strong black marks across the surface, two prominent black marks laterally just posterior to the supra-coxal sulcus.</paragraph>
<paragraph id="8A05A1E07875247DC0075878C792019A" pageId="35" pageNumber="36">
<pageBreakToken id="102C541EE15964C4FAD7ABC92C62EE99" pageId="35" pageNumber="36" start="start">Prothoracic</pageBreakToken>
Legs: Femur squat and robust with a slightly concave dorsal margin; strongly defined pale to dark banding on posterior (external) surface; anterior (internal) surface with a very thin black band running medially from the base to terminus, a small black mark dorsal to the band at the midpoint, and a dark mark dorsal to the band near the femoral brush; the ventral surface pale. Posterior surface of femur with few tubercles. A shallow femoral pit to accommodate terminal posteroventral tibial spine positioned slightly distal to the first proximal posteroventral spine and in line with the most distal discoidal spine; pit is pigmented darkly or pale. Posterior prothoracic femoral genicular spine much smaller than posteroventral spines, originating distal to the beginning of the genicular lobe. Prothoracic tibial posteroventral spines with the first (proximal) smallest and the fourth through sixth of similar length, the second and third are slightly longer. Prothoracic coxae smooth, the anterior surface pale.
</paragraph>
<paragraph id="BC3C0BCB6C3CAC2E6A536C447788A857" pageId="35" pageNumber="36">Meso- and Metathoracic Legs: Femora with ventral (posterior) carina; dorsal (anterior) carina pronounced. Mesotarsi with first segment shorter than the remaining segments combined.</paragraph>
<paragraph id="6F42439C2B01EE2F8C030EE077E45A72" pageId="35" pageNumber="36">Wings: Forewings mottled with brown, pale and greenish coloration; the costal region without strongly defined banding, green and brown proximally with some low contrasting bands developing distally; veins not contrasting from surrounding coloration; two pale spots are positioned in the proximal quarter of the discoidal region just posterior to the first radial vein; a large pale spot is positioned centrally. Forewings often, but not always asymmetrically colored; one being mottled as described the other is darkened significantly with a rust tone, the mottled pattern still visible; extending just beyond or as long as the abdomen. Hindwings with discoidal region opaque and colored brown or rust; the anal region smoky and translucent; the terminus of the discoidal region projecting beyond the distal margin of anal region, the wing appearing slightly elongate.</paragraph>
<paragraph id="7E4EAB220F590E4FFC258427F7D033C4" pageId="35" pageNumber="36">Abdomen: Broad, widening until the fifth tergite before a gradual posterior narrowing; a smooth, brown and black colored dorsal surface. Tergites with small posterolateral tergal projections beginning on the sixth segment. Supra-anal plate slightly transverse, margins tapering gradually to a broadly rounded terminus. Subgenital plate irregularly rounded and without styli.</paragraph>
<paragraph id="F66A0E0F4C7408A3FB6B58C38228D2CF" pageId="35" pageNumber="36">Genital Complex (Fig. 51B.1-B.4): The main body of ventral left sclerite (L4A) with rounded terminus, but with a short, laterally positioned distal process (pda) that is rounded and sometimes, but not always, projecting at an angle towards the medial axis of the L4A that can create a strongly angled transition from the terminal margin of the L4A to the medial margin of the pda; sometimes a depression on the lateral half is present. The apofisis falloid (afa) of the main body of dorsal left sclerite (L4B) very short, broad, tapered to the point and heavily sclerotized, not curved; the apical process (paa) short, cylindrical and curved, the terminus with a rounded end. The right dorsal phallomere (fda) of the first sclerite of right phallomere (R1) tapers to a broad, rounded, membranous terminus, the lateral margin often folded over; the ventral plate (pia) long, broadened proximally and with strongly defined grooves; the ventral process (pva) smooth and tapering to a point distally, one edge straight and the other convex, tooth-like in appearance, but the proximal half c-shaped.</paragraph>
</subSubSection>
<subSubSection id="2154CD4AD0AA756217E54E5EC9D5DBB5" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="redescription">
<paragraph id="DFE506D8BC1D548A2E2829D338C144CA" pageId="35" pageNumber="36">Redescription.</paragraph>
<paragraph id="AF2CC998F9464E9A379EB3B56B4026B7" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
Female. (Figs 4C, 5B) N=8: Body length 25.68-32.17 (28.36); forewing length 16.25-19.99 (18.46); hindwing length 13.46-16.50 (14.96); pronotum length 6.68-8.26 (7.48); prozone length 2.12-2.64 (2.36); pronotum width 3.29-3.99 (3.59); pronotum narrow width 2.49-2.82 (2.71); head width 5.96-7.01 (6.48); head vertex to clypeus 2.55-3.05 (2.81); frons width 2.36-2.87 (2.62); frons height 0.93-1.13 (1.01); prothoracic femur length 7.40-9.15 (8.11);
<pageBreakToken id="6D40F1D45C79D7609B16282A9CF41D34" pageId="36" pageNumber="37" start="start">mesothoracic</pageBreakToken>
femur length 8.30-9.76 (8.85); mesothoracic tibia length 6.39-8.13 (7.04); mesothoracic tarsus length 5.67-6.75 (6.19); metathoracic femur length 8.18-9.91 (8.75); metathoracic tibia length 8.59-11.05 (9.60); metathoracic tarsus length 7.73-9.35 (8.51); pronotal elongation measure 0.31-0.33 (0.32); pronotal shape measure 0.47-0.51 (0.48); head shape measure 0.42-0.44 (0.43); frons shape measure 0.36-0.41 (0.39); anteroventral femoral spine count 15-16 (15); anteroventral tibial spine count 10; posteroventral tibial spine count 7.
</paragraph>
<paragraph id="32C48BA5EF66DFD9F482931C4C1E854D" pageId="36" pageNumber="37">Head (Fig. 40D): As long as wide, the juxta-ocular protuberances very large, the apex in the middle; the vertex is slightly concave, above the dorsal margin of the eyes. Ocelli small and protruding on a continuous carina connecting the three and extending slightly laterally; region between all three raised slightly. Black marking present over the medial carina of the frontal suture; the vertex and juxta-ocular protuberances with dark markings.</paragraph>
<paragraph id="D61ED8314D7C042988E4D5B79864264C" pageId="36" pageNumber="37">Pronotum (Fig. 47D): Metazone with barely concave lateral margins, tapering rapidly in the first third from the supra-coxal bulge, the middle third near parallel before widening to the posterior margin; margins smooth or with few small tubercles; posterior margin with a medial emargination; few small tubercles present.</paragraph>
<paragraph id="ED6552AF365362A6DB7B5244220E1224" pageId="36" pageNumber="37">Prothoracic Legs: Anterior (internal) surface of femur with a very thin black band running medially from the base to terminus that is often interrupted, a small black mark dorsal to the band at the midpoint, and a dark mark dorsal to the band near the femoral brush.</paragraph>
<paragraph id="64005250BAFC88F019187B536EB5CA7E" pageId="36" pageNumber="37">Meso- and Metathoracic Legs: As described for males.</paragraph>
<paragraph id="F8FC25EDAB4D242A4EB1C3381867F753" pageId="36" pageNumber="37">Wings: Forewings mottled with contrasting brown, pale and greenish coloration; veins pale brown. Forewings often, but not always asymmetrically colored; one being mottled as described the other is darkened with a rust or black tone, the mottled pattern still visible; extending to the terminus of the abdomen. Hindwings with discoidal region opaque and colored brown, rust or dark yellow; the anal region smoky and translucent.</paragraph>
<paragraph id="3B13C697281D613AA01EA8E14C3D3C1A" pageId="36" pageNumber="37">Abdomen: Very broad, widening until the fifth tergite before a gradual posterior narrowing, the abdomen almost circular; a smooth, brown and black colored dorsal surface. Tergites with large posterolateral tergal projections beginning on the fourth segment. Supra-anal plate slightly transverse, margins tapering gradually to a rounded terminus.</paragraph>
</subSubSection>
</treatment>
</document>