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<document id="A71356A8758DEA3B7FFFB5FA0A0607A3" ID-DOI="10.11646/zootaxa.5537.4.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="14240290" ID-ZooBank="4963FF97-53E7-4A0C-BED7-966A6AD41A1D" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1732802264253" checkinUser="plazi" docAuthor="Nastasi, Louis F. &amp; Deans, Andrew R." docDate="2024" docId="1C2087D0FFE54130FF69F8EF016FD611" docLanguage="en" docName="zootaxa.5537.4.4.pdf" docOrigin="Zootaxa 5537 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5537.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aulacideini Nastasi &amp; Deans 2024" docType="key" docVersion="1" lastPageNumber="524" masterDocId="E019FFA8FFE9413DFFFEFFC20554D44D" masterDocTitle="New herb gall wasps (Hymenoptera: Cynipidae: Aulacideini) from Kyrgyzstan, with description of a new genus and a review of the genus Panteliella Kieffer, 1901" masterLastPageNumber="526" masterPageNumber="511" pageNumber="523" updateTime="1733169968947" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:titleInfo id="249A8782FE95B28112101C7AFCC53D8D">
<mods:title id="89A5EB7A74A849F46B8A527DB712EA9C">New herb gall wasps (Hymenoptera: Cynipidae: Aulacideini) from Kyrgyzstan, with description of a new genus and a review of the genus Panteliella Kieffer, 1901</mods:title>
</mods:titleInfo>
<mods:name id="CE2F766F804358242F25DD8F3635C583" type="personal">
<mods:role id="6764B1DC3F8DA080D861110AA050B0FD">
<mods:roleTerm id="35B57B74810AF15665DFF63541CC9E25">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02B48B24BDB564DA8AF6DF0BC2D492FA">Nastasi, Louis F.</mods:namePart>
<mods:nameIdentifier id="D142E0A6E684039F4C86849E84D75A99" type="ORCID">0000-0001-7825-480X</mods:nameIdentifier>
<mods:affiliation id="C2AAE95333D6FC1A2A258C09664979DB">Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA, 16802.</mods:affiliation>
</mods:name>
<mods:name id="5A64C54F9EC5C2FE30200700F820E74E" type="personal">
<mods:role id="999631430D57587FB1ED476E65BA31BF">
<mods:roleTerm id="C295402784029ED5F7FC27C3ED3A34B9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="93F43EE0E0062690DF17BBC5CB8236BD">Deans, Andrew R.</mods:namePart>
<mods:nameIdentifier id="71E9D1BBB78C158C42AF381BEC545117" type="ORCID">0000-0002-2119-4663</mods:nameIdentifier>
<mods:affiliation id="441199F8FD96D5910ECFC0E3844B4D88">Frost Entomological Museum, Department of Entomology, The Pennsylvania State University, University Park, PA, 16802.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C4E677F6F11D0806018DB9FEF7A7BA36">text</mods:typeOfResource>
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<mods:title id="228B8CC97DA19A6CDDD5545988BCDAD7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="23EC1205F77C4B1BC098FD92A19B928D">
<mods:date id="5130A812125459EF3128CCA2EB77B8DA">2024</mods:date>
<mods:detail id="74FA083DCE326970C67ABC3557C08F71" type="pubDate">
<mods:number id="E657E620AA2D6D68A9F4BE1A26EB498C">2024-11-12</mods:number>
</mods:detail>
<mods:detail id="79E836F15BA9D437C800F5EFE7053019" type="volume">
<mods:number id="4A9A4F2D1B31E05D59AF3A7B062E376C">5537</mods:number>
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<mods:number id="250089D0DACDA82D4ACEC666A559334F">4</mods:number>
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<mods:url id="38A16ACC4298D0A068F0B2CBE6777BCC">http://dx.doi.org/10.11646/zootaxa.5537.4.4</mods:url>
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<mods:classification id="A98639C831121AE7B75B189E8C51C202">journal article</mods:classification>
<mods:identifier id="F6B504E5ABA524956FB93B3B734AED3E" type="DOI">10.11646/zootaxa.5537.4.4</mods:identifier>
<mods:identifier id="4CDC72929001D7340702725593002D54" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="665B1831F142727ABD4122E169906A46" type="Zenodo-Dep">14240290</mods:identifier>
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<treatment id="1C2087D0FFE54130FF69F8EF016FD611" LSID="urn:lsid:plazi:treatment:1C2087D0FFE54130FF69F8EF016FD611" httpUri="http://treatment.plazi.org/id/1C2087D0FFE54130FF69F8EF016FD611" lastPageId="13" lastPageNumber="524" pageId="12" pageNumber="523">
<subSubSection id="DC93654DFFE54131FF69F8EF0795D30A" box="[151,705,1836,1863]" pageId="12" pageNumber="523" type="nomenclature">
<paragraph id="943636C6FFE54131FF69F8EF0795D30A" blockId="12.[151,705,1836,1863]" box="[151,705,1836,1863]" pageId="12" pageNumber="523">
<heading id="CF7E81AAFFE54131FF69F8EF0795D30A" bold="true" box="[151,705,1836,1863]" fontSize="11" level="1" pageId="12" pageNumber="523" reason="1">
<emphasis id="A6FDEAD4FFE54131FF69F8EF0795D30A" bold="true" box="[151,705,1836,1863]" pageId="12" pageNumber="523">
Key to the
<taxonomicName id="53894D45FFE54131FEE0F8EE04F9D30B" authorityName="Nastasi &amp; Deans" authorityYear="2024" box="[286,429,1836,1862]" class="Insecta" family="Cynipidae" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="tribe" tribe="Aulacideini">Aulacideini</taxonomicName>
genera of
<collectingCountry id="EC9E7656FFE54131FDCEF8EF0795D30A" box="[560,705,1837,1863]" name="Kyrgyzstan" pageId="12" pageNumber="523">Kyrgyzstan</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DC93654DFFE54131FF69F8B7060FD3B6" pageId="12" pageNumber="523" type="discussion">
<paragraph id="943636C6FFE54131FF69F8B7060FD3B6" blockId="12.[151,1437,1908,2043]" pageId="12" pageNumber="523">
We provide the following key to allow for the separation of all
<taxonomicName id="53894D45FFE54131FC6BF8B6014CD3C3" authorityName="Nastasi &amp; Deans" authorityYear="2024" box="[917,1048,1908,1934]" class="Insecta" family="Cynipidae" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="tribe" tribe="Aulacideini">Aulacideini</taxonomicName>
genera known from
<collectingCountry id="EC9E7656FFE54131FAEBF8B700CDD3C2" box="[1301,1433,1909,1935]" name="Kyrgyzstan" pageId="12" pageNumber="523">Kyrgyzstan</collectingCountry>
. Specimens may be identified as
<taxonomicName id="53894D45FFE54131FDFEF85A07D7D3FF" authorityName="Nastasi &amp; Deans" authorityYear="2024" box="[512,643,1944,1970]" class="Insecta" family="Cynipidae" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="tribe" tribe="Aulacideini">Aulacideini</taxonomicName>
using the identification key in
<bibRefCitation id="F0184B37FFE54131FC1EF85A01B8D3FE" author="Buffington, M. L. &amp; Forshage, M. &amp; Liljeblad, J. &amp; Tang, C. T. &amp; van Noort, S." box="[992,1260,1944,1971]" pageId="12" pageNumber="523" refId="ref9290" refString="Buffington, M. L., Forshage, M., Liljeblad, J., Tang, C. T. &amp; van Noort, S. (2020) World Cynipoidea (Hymenoptera): A key to higher-level groups. Insect Systematics and Diversity, 4, 4, 1. https: // doi. org / 10.1093 / isd / ixaa 003" type="journal volume" year="2020">Buffington et al. (2020)</bibRefCitation>
, although some updated diagnostic characters were provided by
<bibRefCitation id="F0184B37FFE54131FD53F87E06C3D39A" author="Nastasi, L. F. &amp; Buffington, M. L. &amp; Davis, C. K. &amp; Deans, A. R." box="[685,919,1980,2007]" pageId="12" pageNumber="523" pagination="177 - 207" refId="ref9809" refString="Nastasi, L. F., Buffington, M. L., Davis, C. K. &amp; Deans, A. R. (2024 a) Key to the North American tribes and genera of herb, rose, bramble, and inquiline gall wasps (Hymenoptera, Cynipoidea, Cynipidae sensu lato). ZooKeys, 1196, 177 - 207. https: // doi. org / 10.3897 / zookeys. 1196.118460" type="journal article" year="2024">Nastasi et al. (2024a)</bibRefCitation>
to prevent the exclusion of taxa with reticulate sculpture on the mesopleuron (
<emphasis id="A6FDEAD4FFE54131FE0CF8230775D3B7" box="[498,545,2017,2042]" italics="true" pageId="12" pageNumber="523">e.g.,</emphasis>
<taxonomicName id="53894D45FFE54131FDD6F823078ED3B7" authority="Nastasi" authorityName="Nastasi &amp; Deans" authorityYear="2024" box="[552,730,2016,2043]" class="Insecta" family="Cynipidae" genus="Soparia" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A6FDEAD4FFE54131FDD6F82307D5D3B6" box="[552,641,2017,2043]" italics="true" pageId="12" pageNumber="523">Soparia</emphasis>
Nastasi
</taxonomicName>
<emphasis id="A6FDEAD4FFE54131FD1CF823061FD3B6" bold="true" box="[738,843,2017,2043]" pageId="12" pageNumber="523">
<taxonomicNameLabel id="BDCE57AFFFE54131FD1CF823061FD3B6" box="[738,843,2017,2043]" pageId="12" pageNumber="523" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="DC93654DFFE44130FF69FF5500C8D5A2" pageId="13" pageNumber="524" type="key">
<keyStep id="2F7D2E63FFE44130FF69FF5500C8D487" pageId="13" pageNumber="524">
<paragraph id="943636C6FFE44130FF69FF5500C8D4E1" blockId="13.[151,1436,151,203]" box="[151,1436,151,173]" pageId="13" pageNumber="524">
<keyLead id="2F7895F3FFE44130FF69FF5500C8D4E1" box="[151,1436,151,173]" pageId="13" pageNumber="524">
<emphasis id="A6FDEAD4FFE44130FF69FF5505FED4E1" bold="true" box="[151,170,151,172]" pageId="13" pageNumber="524">1.</emphasis>
Fore wing marginal cell closed (
<figureCitation id="0CB22A43FFE44130FDE6FF550707D4E0" box="[536,595,151,173]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1734,1758]" captionTargetBox="[152,1436,181,1710]" captionTargetId="figure-22@2.[152,1436,181,1710]" captionTargetPageId="2" captionText="FIGURES 1–6. Soparia oshensis Nastasi gen. and sp. nov. All images show holotype female INHS Insect Collection 288148. (1) Lateral habitus. (2) Head in anterior view; dotted line indicates ventral margin of clypeus. (3) Mesoscutum in dorsal view. (4) Scutellum in dorsal view. (5) Mesopleuron in lateral view. (6) Wings." figureDoi="http://doi.org/10.5281/zenodo.14240292" httpUri="https://zenodo.org/record/14240292/files/figure.png" pageId="13" pageNumber="524">Fig. 6</figureCitation>
)................................ .....................................
<emphasis id="A6FDEAD4FFE44130FA6EFF5500C8D4E1" bold="true" box="[1424,1436,151,172]" pageId="13" pageNumber="524">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="943636C6FFE44130FF69FF7700C8D487" blockId="13.[151,1436,151,203]" box="[151,1436,181,203]" pageId="13" pageNumber="524">
<keyLead id="2F7895F3FFE44130FF69FF7700C8D487" box="[151,1436,181,203]" pageId="13" pageNumber="524">
- Fore wing marginal cell open (
<figureCitation id="0CB22A43FFE44130FDF7FF770705D487" box="[521,593,181,203]" captionStart="FIGURES 9–14" captionStartId="10.[151,264,1667,1691]" captionTargetBox="[180,1407,181,1644]" captionTargetId="figure-22@10.[180,1407,181,1644]" captionTargetPageId="10" captionText="FIGURES 9–14. Panteliella rugosa Nastasi sp. nov. All images show holotype female INHS Insect Collection 288137. (9) Lateral habitus. (10) Head in anterior view; dotted line indicates ventral margin of clypeus. (11) Mesoscutum in dorsal view. (12) Scutellum in dorsal view. (13) Mesopleuron in lateral view. (14) Fore wing." figureDoi="http://doi.org/10.5281/zenodo.14240300" httpUri="https://zenodo.org/record/14240300/files/figure.png" pageId="13" pageNumber="524">Fig. 14</figureCitation>
).....................................................................
<emphasis id="A6FDEAD4FFE44130FA6EFF7700C8D487" bold="true" box="[1424,1436,181,202]" pageId="13" pageNumber="524">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2F7D2E63FFE44130FF69FF2D00C8D511" pageId="13" pageNumber="524">
<paragraph id="943636C6FFE44130FF69FF2D00C8D56F" blockId="13.[151,1436,239,349]" pageId="13" pageNumber="524">
<keyLead id="2F7895F3FFE44130FF69FF2D00C8D56F" pageId="13" pageNumber="524">
<emphasis id="A6FDEAD4FFE44130FF69FF2D05FDD549" bold="true" box="[151,169,239,260]" pageId="13" pageNumber="524">2.</emphasis>
Mesopleuron sculpture entirely striate-reticulate (
<figureCitation id="0CB22A43FFE44130FD35FF2D065ED548" box="[715,778,239,261]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1734,1758]" captionTargetBox="[152,1436,181,1710]" captionTargetId="figure-22@2.[152,1436,181,1710]" captionTargetPageId="2" captionText="FIGURES 1–6. Soparia oshensis Nastasi gen. and sp. nov. All images show holotype female INHS Insect Collection 288148. (1) Lateral habitus. (2) Head in anterior view; dotted line indicates ventral margin of clypeus. (3) Mesoscutum in dorsal view. (4) Scutellum in dorsal view. (5) Mesopleuron in lateral view. (6) Wings." figureDoi="http://doi.org/10.5281/zenodo.14240292" httpUri="https://zenodo.org/record/14240292/files/figure.png" pageId="13" pageNumber="524">Fig. 5</figureCitation>
). Notauli incomplete and weakly impressed, most conspicuous in posterior half of mesoscutum (
<figureCitation id="0CB22A43FFE44130FDF6FECF0715D56F" box="[520,577,269,291]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1734,1758]" captionTargetBox="[152,1436,181,1710]" captionTargetId="figure-22@2.[152,1436,181,1710]" captionTargetPageId="2" captionText="FIGURES 1–6. Soparia oshensis Nastasi gen. and sp. nov. All images show holotype female INHS Insect Collection 288148. (1) Lateral habitus. (2) Head in anterior view; dotted line indicates ventral margin of clypeus. (3) Mesoscutum in dorsal view. (4) Scutellum in dorsal view. (5) Mesopleuron in lateral view. (6) Wings." figureDoi="http://doi.org/10.5281/zenodo.14240292" httpUri="https://zenodo.org/record/14240292/files/figure.png" pageId="13" pageNumber="524">Fig. 3</figureCitation>
)..................................................
<emphasis id="A6FDEAD4FFE44130FB60FECF00C8D56F" bold="true" box="[1182,1436,269,291]" pageId="13" pageNumber="524">
<taxonomicName id="53894D45FFE44130FB60FECF0069D56E" authority="Nastasi" authorityName="Nastasi &amp; Deans" authorityYear="2024" box="[1182,1341,269,291]" class="Insecta" family="Cynipidae" genus="Soparia" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="524" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A6FDEAD4FFE44130FB60FECF01BFD56F" bold="true" box="[1182,1259,269,290]" italics="true" pageId="13" pageNumber="524">Soparia</emphasis>
Nastasi
</taxonomicName>
<taxonomicNameLabel id="BDCE57AFFFE44130FABCFECF00C8D56F" box="[1346,1436,269,290]" pageId="13" pageNumber="524" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="943636C6FFE44130FF69FEE900C8D511" blockId="13.[151,1436,239,349]" pageId="13" pageNumber="524">
<keyLead id="2F7895F3FFE44130FF69FEE900C8D511" pageId="13" pageNumber="524">
- Mesopleuron sculpture mostly or entirely striate; at most with some reticulate sculpture apparent in ventral third. Notauli complete, deeply impressed across entire mesoscutum*..................................
<taxonomicName id="53894D45FFE44130FB68FE8500C8D511" ID-CoL="35MW" authority="Ashmead, 1897" authorityName="Ashmead" authorityYear="1897" box="[1174,1436,327,349]" class="Insecta" family="Cynipidae" genus="Aulacidea" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFE44130FB68FE8500C8D511" bold="true" box="[1174,1436,327,349]" pageId="13" pageNumber="524">
<emphasis id="A6FDEAD4FFE44130FB68FE8501AED510" bold="true" box="[1174,1274,327,349]" italics="true" pageId="13" pageNumber="524">Aulacidea</emphasis>
<bibRefCitation id="F0184B37FFE44130FAFEFE8500C8D511" author="Ashmead, W. H." box="[1280,1436,327,348]" pageId="13" pageNumber="524" pagination="67 - 69" refId="ref9208" refString="Ashmead, W. H. (1897) Description of Some New Genera in the Family Cynipidae. Psyche, 8, 67 - 69. https: // doi. org / 10.1155 / 1897 / 60546" type="journal article" year="1897">Ashmead, 1897</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2F7D2E63FFE44130FF69FE4000C8D5A2" pageId="13" pageNumber="524">
<paragraph id="943636C6FFE44130FF69FE4000C8D5F9" blockId="13.[151,1437,386,496]" pageId="13" pageNumber="524">
<keyLead id="2F7895F3FFE44130FF69FE4000C8D5F9" pageId="13" pageNumber="524">
<emphasis id="A6FDEAD4FFE44130FF69FE4005FDD5DA" bold="true" box="[151,169,386,407]" pageId="13" pageNumber="524">3.</emphasis>
Notauli conspicuous and strongly impressed (
<figureCitation id="0CB22A43FFE44130FD67FE4007B5D5DA" box="[665,737,386,408]" captionStart="FIGURES 15–21" captionStartId="12.[151,264,1597,1621]" captionTargetBox="[210,1377,181,1573]" captionTargetId="figure-22@12.[210,1377,181,1573]" captionTargetPageId="12" captionText="FIGURES 15–21. Isocolus barakus Nastasi sp. nov. All images show holotype female INHS Insect Collection 288135. (9) Lateral habitus. (10) Head in anterior view; dotted line indicates ventral margin of clypeus. (11) Pronotum in anterior view; arrows indicate conspicuously defined range of pronotal lateral suture. (12) Proximal antennomeres. (13) Mesopleuron in lateral view. (14) Mesoscutum in dorsal view. (15) Scutellum in dorsal view; posterior scutellum (partially covered by wings and thus clouded posteriorly in image)." figureDoi="http://doi.org/10.5281/zenodo.14240302" httpUri="https://zenodo.org/record/14240302/files/figure.png" pageId="13" pageNumber="524">Fig. 20</figureCitation>
). Clypeus only slightly projecting ventrally over base of mandibles (
<figureCitation id="0CB22A43FFE44130FA8BFE4005A9D5F8" captionStart="FIGURES 15–21" captionStartId="12.[151,264,1597,1621]" captionTargetBox="[210,1377,181,1573]" captionTargetId="figure-22@12.[210,1377,181,1573]" captionTargetPageId="12" captionText="FIGURES 15–21. Isocolus barakus Nastasi sp. nov. All images show holotype female INHS Insect Collection 288135. (9) Lateral habitus. (10) Head in anterior view; dotted line indicates ventral margin of clypeus. (11) Pronotum in anterior view; arrows indicate conspicuously defined range of pronotal lateral suture. (12) Proximal antennomeres. (13) Mesopleuron in lateral view. (14) Mesoscutum in dorsal view. (15) Scutellum in dorsal view; posterior scutellum (partially covered by wings and thus clouded posteriorly in image)." figureDoi="http://doi.org/10.5281/zenodo.14240302" httpUri="https://zenodo.org/record/14240302/files/figure.png" pageId="13" pageNumber="524">Fig. 16</figureCitation>
)................................................................................
<taxonomicName id="53894D45FFE44130FB40FE5D00C8D5F9" ID-CoL="9CL3P" authority="Forster, 1869" authorityName="Forster" authorityYear="1869" box="[1214,1436,415,437]" class="Insecta" family="Cynipidae" genus="Isocolus" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFE44130FB40FE5D00C8D5F9" bold="true" box="[1214,1436,415,437]" pageId="13" pageNumber="524">
<emphasis id="A6FDEAD4FFE44130FB40FE5D0045D5F8" bold="true" box="[1214,1297,415,437]" italics="true" pageId="13" pageNumber="524">Isocolus</emphasis>
<bibRefCitation id="F0184B37FFE44130FAE8FE6200C8D5F9" author="Forster, A." box="[1302,1436,415,437]" pageId="13" pageNumber="524" pagination="325 - 370" refId="ref9503" refString="Forster, A. (1869) Ueber die Gallwespen. Verhandlungen der k. k. Zoologisch-botanischen Gesellschaft in Wien, 19, 325 - 370." type="journal article" year="1869">Förster, 1869</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="943636C6FFE44130FF69FE7F00C8D5A2" blockId="13.[151,1437,386,496]" pageId="13" pageNumber="524">
<keyLead id="2F7895F3FFE44130FF69FE7F00C8D5A2" pageId="13" pageNumber="524">
- Notauli inconspicuous to weakly impressed (
<figureCitation id="0CB22A43FFE44130FD6CFE7F078CD59F" box="[658,728,445,467]" captionStart="FIGURES 9–14" captionStartId="10.[151,264,1667,1691]" captionTargetBox="[180,1407,181,1644]" captionTargetId="figure-22@10.[180,1407,181,1644]" captionTargetPageId="10" captionText="FIGURES 9–14. Panteliella rugosa Nastasi sp. nov. All images show holotype female INHS Insect Collection 288137. (9) Lateral habitus. (10) Head in anterior view; dotted line indicates ventral margin of clypeus. (11) Mesoscutum in dorsal view. (12) Scutellum in dorsal view. (13) Mesopleuron in lateral view. (14) Fore wing." figureDoi="http://doi.org/10.5281/zenodo.14240300" httpUri="https://zenodo.org/record/14240300/files/figure.png" pageId="13" pageNumber="524">Fig. 11</figureCitation>
). Clypeus broadly projecting ventrally over base of mandibles (
<figureCitation id="0CB22A43FFE44130FAC3FE7F00D1D59F" box="[1341,1413,445,467]" captionStart="FIGURES 9–14" captionStartId="10.[151,264,1667,1691]" captionTargetBox="[180,1407,181,1644]" captionTargetId="figure-22@10.[180,1407,181,1644]" captionTargetPageId="10" captionText="FIGURES 9–14. Panteliella rugosa Nastasi sp. nov. All images show holotype female INHS Insect Collection 288137. (9) Lateral habitus. (10) Head in anterior view; dotted line indicates ventral margin of clypeus. (11) Mesoscutum in dorsal view. (12) Scutellum in dorsal view. (13) Mesopleuron in lateral view. (14) Fore wing." figureDoi="http://doi.org/10.5281/zenodo.14240300" httpUri="https://zenodo.org/record/14240300/files/figure.png" pageId="13" pageNumber="524">Fig. 10</figureCitation>
)..................................................................................
<taxonomicName id="53894D45FFE44130FB56FE1800C8D5A2" authority="Kieffer, 1901" authorityName="Kieffer" authorityYear="1901" box="[1192,1436,474,496]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="524" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFE44130FB56FE1800C8D5A2" bold="true" box="[1192,1436,474,496]" pageId="13" pageNumber="524">
<emphasis id="A6FDEAD4FFE44130FB56FE180047D5BD" bold="true" box="[1192,1299,474,496]" italics="true" pageId="13" pageNumber="524">Panteliella</emphasis>
Kieffer, 1901
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<subSubSection id="DC93654DFFE44130FF69FDE2016FD611" pageId="13" pageNumber="524" type="multiple">
<paragraph id="943636C6FFE44130FF69FDE2016FD611" blockId="13.[151,1436,544,604]" pageId="13" pageNumber="524">
* = Notauli complete in both species known from
<collectingCountry id="EC9E7656FFE44130FD53FDE30675D674" box="[685,801,545,569]" name="Kyrgyzstan" pageId="13" pageNumber="524">Kyrgyzstan</collectingCountry>
(
<taxonomicName id="53894D45FFE44130FCCDFDE2018BD675" authority="Tyurebaev, 1972" authorityName="Tyurebaev" authorityYear="1972" box="[819,1247,544,568]" class="Insecta" family="Cynipidae" genus="Aulacidea" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="524" phylum="Arthropoda" rank="species" species="acroptilonica">
<emphasis id="A6FDEAD4FFE44130FCCDFDE2017FD675" box="[819,1067,544,568]" italics="true" pageId="13" pageNumber="524">Aulacidea acroptilonica</emphasis>
<bibRefCitation id="F0184B37FFE44130FBCDFDE2018BD675" author="Tyurebaev, S. S." box="[1075,1247,544,568]" pageId="13" pageNumber="524" pagination="52 - 53" refId="ref10509" refString="Tyurebaev, S. S. (1972) Gall-forming insects attacking Russian knapweed in the middle reaches of the R. Irtysh. Vestnik Sel'skokhozyaistvennoi Nauki Kazakhskoi SSR, 15 (6), 52 - 53." type="journal article" year="1972">Tyurebaev, 1972</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="53894D45FFE44130FAEBFDE30431D611" authority="Diakontschuk, 1984" authorityName="Diakontschuk" authorityYear="1984" class="Insecta" family="Cynipidae" genus="Aulacidea" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="524" phylum="Arthropoda" rank="species" species="serratulae">
<emphasis id="A6FDEAD4FFE44130FAEBFDE300C8D675" box="[1301,1436,544,568]" italics="true" pageId="13" pageNumber="524">A. serratulae</emphasis>
<bibRefCitation id="F0184B37FFE44130FF69FD860431D611" author="Diakontschuk, L. A." box="[151,357,580,604]" pageId="13" pageNumber="524" pagination="74 - 77" refId="ref9469" refString="Diakontschuk, L. A. (1984) New species of Cynipidae (Cynipidae, Hymenoptera) from Georgia. Vestnik Zoologii, 3, 74 - 77. [in Russian]" type="journal article" year="1984">Diakontschuk, 1984</bibRefCitation>
</taxonomicName>
) but incomplete in some species known from other geographic regions.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="68791C571E828ABC813BF60C21E4A116">New herb gall wasps (Hymenoptera: Cynipidae: Aulacideini) from Kyrgyzstan, with description of a new genus and a review of the genus Panteliella Kieffer, 1901</mods:title>
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<paragraph id="943636C6FFEC4138FF69FF550495D4FC" blockId="5.[151,449,151,213]" box="[151,449,151,178]" pageId="5" pageNumber="516">
<heading id="CF7E81AAFFEC4138FF69FF550495D4FC" bold="true" box="[151,449,151,178]" fontSize="11" level="1" pageId="5" pageNumber="516" reason="1">
<taxonomicName id="53894D45FFEC4138FF69FF550495D4FC" authority="Kieffer, 1901" authorityName="Kieffer" authorityYear="1901" box="[151,449,151,178]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FF69FF550495D4FC" bold="true" box="[151,449,151,178]" pageId="5" pageNumber="516">
<emphasis id="A6FDEAD4FFEC4138FF69FF55044DD4FF" bold="true" box="[151,281,151,178]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
Kieffer, 1901
</emphasis>
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<subSubSection id="DC93654DFFEC4138FF69FF790474D498" box="[151,288,187,213]" pageId="5" pageNumber="516" type="description">
<paragraph id="943636C6FFEC4138FF69FF790474D498" blockId="5.[151,449,151,213]" box="[151,288,187,213]" pageId="5" pageNumber="516">
(
<figureCitation id="0CB22A43FFEC4138FF61FF790443D498" box="[159,279,187,213]" captionStart-0="FIGURES 7–8" captionStart-1="FIGURES 9–14" captionStartId-0="4.[151,264,1951,1975]" captionStartId-1="10.[151,264,1667,1691]" captionTargetBox-0="[213,1375,181,1927]" captionTargetBox-1="[180,1407,181,1644]" captionTargetId-0="figure-22@4.[213,1375,181,1927]" captionTargetId-1="figure-22@10.[180,1407,181,1644]" captionTargetPageId-0="4" captionTargetPageId-1="10" captionText-0="FIGURES 7–8. Panteliella fedtschenkoi (Rübsaamen, 1896), female USNMENT 01823223. (7) Lateral habitus. (8) Dorsal mesosoma." captionText-1="FIGURES 9–14. Panteliella rugosa Nastasi sp. nov. All images show holotype female INHS Insect Collection 288137. (9) Lateral habitus. (10) Head in anterior view; dotted line indicates ventral margin of clypeus. (11) Mesoscutum in dorsal view. (12) Scutellum in dorsal view. (13) Mesopleuron in lateral view. (14) Fore wing." figureDoi-0="http://doi.org/10.5281/zenodo.14240294" figureDoi-1="http://doi.org/10.5281/zenodo.14240300" httpUri-0="https://zenodo.org/record/14240294/files/figure.png" httpUri-1="https://zenodo.org/record/14240300/files/figure.png" pageId="5" pageNumber="516">Figs. 7–14</figureCitation>
)
</paragraph>
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<subSubSection id="DC93654DFFEC4138FF69FF380777D628" pageId="5" pageNumber="516" type="reference_group">
<paragraph id="943636C6FFEC4138FF69FF3804D1D55F" blockId="5.[151,548,250,366]" box="[151,389,250,274]" pageId="5" pageNumber="516">
<taxonomicName id="53894D45FFEC4138FF69FF3804D1D55F" ID-CoL="8NWNQ" authority="Kieffer, 1901 a" authorityName="Kieffer" authorityYear="1901" box="[151,389,250,274]" class="Insecta" family="Cynipidae" genus="Pantelia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FF69FF3805B9D55F" box="[151,237,250,274]" italics="true" pageId="5" pageNumber="516">Pantelia</emphasis>
<bibRefCitation id="F0184B37FFEC4138FF0DFF3804D1D55F" author="Kieffer, J. J." box="[243,389,250,274]" pageId="5" pageNumber="516" refId="ref9612" refString="Kieffer, J. J. (1901 a) Monographie des Cynipides d'Europe et d'Algerie: Les Cynipides. Froment-Dubosclard, Paris, 687 pp." type="book" year="1901">Kieffer, 1901a</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="943636C6FFEC4138FF69FEDB04F4D57C" blockId="5.[151,548,250,366]" box="[151,416,281,305]" pageId="5" pageNumber="516">
<taxonomicName id="53894D45FFEC4138FF69FEDB04F4D57C" authority="Kieffer, 1901 b" authorityName="Kieffer" authorityYear="1901" box="[151,416,281,305]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FF69FEDB0452D57C" box="[151,262,281,305]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
<bibRefCitation id="F0184B37FFEC4138FEF3FEDB04F4D57C" author="Kieffer, J. J." box="[269,416,281,305]" pageId="5" pageNumber="516" pagination="233 - 579" refId="ref9639" refString="Kieffer, J. J. (1901 b) Synopsis de Zoocecidies d'Europe. Annalles de la Societe Entomologique de France, 1901, 233 - 579. https: // doi. org / 10.1080 / 21686351.1901.12279879" type="journal article" year="1901">Kieffer, 1901b</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="943636C6FFEC4138FF69FEFA0423D502" blockId="5.[151,548,250,366]" box="[151,375,311,336]" pageId="5" pageNumber="516">
<taxonomicName id="53894D45FFEC4138FF69FEFA0423D502" authority="Belizin, 1959" authorityName="Belizin" authorityYear="1959" box="[151,375,311,336]" class="Insecta" family="Cynipidae" genus="Vetustia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FF69FEFA05B3D502" box="[151,231,312,335]" italics="true" pageId="5" pageNumber="516">Vetustia</emphasis>
<bibRefCitation id="F0184B37FFEC4138FF10FEFA0423D502" author="Belizin, V. I." box="[238,375,311,336]" pageId="5" pageNumber="516" pagination="662 - 674" refId="ref9247" refString="Belizin, V. I. (1959) Gall wasps of the tribe Aylaxini (Hymenoptera, Cynipoidea) new for the fauna of the U. S. S. R. Entomologicheskoe Obozrenie, 38, 662 - 674." type="book chapter" year="1959">Belizin, 1959</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="943636C6FFEC4138FF69FE940770D523" blockId="5.[151,548,250,366]" box="[151,548,342,366]" pageId="5" pageNumber="516">
<taxonomicName id="53894D45FFEC4138FF69FE940770D523" authority="Ionescu &amp; Roman, 1960" authorityName="Ionescu &amp; Roman" authorityYear="1960" box="[151,548,342,366]" class="Insecta" family="Cynipidae" genus="Endocaulonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FF69FE940471D523" box="[151,293,342,366]" italics="true" pageId="5" pageNumber="516">Endocaulonia</emphasis>
<bibRefCitation id="F0184B37FFEC4138FED2FE950770D523" author="Ionescu, M. A. &amp; Roman, N." box="[300,548,342,366]" pageId="5" pageNumber="516" pagination="221 - 225" refId="ref9557" refString="Ionescu, M. A. &amp; Roman, N. (1960) Un genre nouveau de Cynipides gallicoles Endocaulonia bicolor n. g., n. sp. (Hymenoptera, Cynipoidea). Revue de Biologie Academiei Republicii Populare Romine Bucaresti, 5 (3), 221 - 225." type="journal article" year="1960">Ionescu &amp; Roman, 1960</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="943636C6FFEC4138FF69FE550777D628" blockId="5.[151,1437,407,2018]" pageId="5" pageNumber="516">
<emphasis id="A6FDEAD4FFEC4138FF69FE5504D3D5FC" bold="true" box="[151,391,407,433]" pageId="5" pageNumber="516">Historical overview.</emphasis>
<taxonomicName id="53894D45FFEC4138FE6EFE55075FD5FC" authorityName="Kieffer" authorityYear="1901" box="[400,523,407,433]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FE6EFE55075FD5FC" box="[400,523,407,433]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
</taxonomicName>
previously included two valid species:
<taxonomicName id="53894D45FFEC4138FC2EFE5A001AD5FC" authority="Vyrzhikovskaya, 1962" authorityName="Vyrzhikovskaya" authorityYear="1962" box="[976,1358,407,433]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="bianchii">
<emphasis id="A6FDEAD4FFEC4138FC2EFE5A011ED5FC" box="[976,1098,407,433]" italics="true" pageId="5" pageNumber="516">P. bianchii</emphasis>
<bibRefCitation id="F0184B37FFEC4138FBACFE55001AD5FC" author="Vyrzhikovskaya, A. V." box="[1106,1358,407,433]" pageId="5" pageNumber="516" pagination="138 - 171" refId="ref10548" refString="Vyrzhikovskaya, A. V. (1962) Gall-wasps (Hymenoptera, Cynipoidea, Cynipinae) of the Leningrad Region. Proceedings of the Zoological Institute of the USSR Academy of Sciences, XXXI, 138 - 171. [in Russian]" type="book chapter" year="1962">Vyrzhikovskaya, 1962</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="53894D45FFEC4138FA76FE5A075DD598" authority="(Rubsaamen, 1896)" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEC4138FA76FE5A0477D598" italics="true" pageId="5" pageNumber="516">P. fedtschenkoi</emphasis>
(
<bibRefCitation id="F0184B37FFEC4138FECCFE790754D598" author="Rubsaamen, E. H." box="[306,512,443,469]" pageId="5" pageNumber="516" pagination="396 - 481" refId="ref10418" refString="Rubsaamen, E. H. (1896) Uber russische Zoocecidien und deren Erzeuger. Bulletin de la Societe imperiale des naturalistes de Moscou, 3, 396 - 481." type="journal article" year="1896">Rübsaamen, 1896</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="53894D45FFEC4138FDEAFE7907DBD598" authorityName="Kieffer" authorityYear="1901" box="[532,655,443,469]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FDEAFE7907DBD598" box="[532,655,443,469]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
</taxonomicName>
was first described to accommodate
<taxonomicName id="53894D45FFEC4138FBD0FE79058FD5B4" authority="(Rubsaamen, 1896)" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" class="Magnoliopsida" family="Proteaceae" genus="Aulax" kingdom="Plantae" order="Proteales" pageId="5" pageNumber="516" phylum="Tracheophyta" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEC4138FBD0FE790056D598" box="[1070,1282,443,469]" italics="true" pageId="5" pageNumber="516">Aulax fedtschenkoi</emphasis>
(
<bibRefCitation id="F0184B37FFEC4138FAEFFE790587D5B4" author="Rubsaamen, E. H." pageId="5" pageNumber="516" pagination="396 - 481" refId="ref10418" refString="Rubsaamen, E. H. (1896) Uber russische Zoocecidien und deren Erzeuger. Bulletin de la Societe imperiale des naturalistes de Moscou, 3, 396 - 481." type="journal article" year="1896">Rübsaamen, 1896</bibRefCitation>
)
</taxonomicName>
, a species inducing small galls on leaves of
<taxonomicName id="53894D45FFEC4138FD19FE1D0682D5B4" ID-CoL="4GJ33" authority="(L.) Moench" box="[743,982,479,505]" class="Magnoliopsida" family="Lamiaceae" genus="Phlomoides" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="516" phylum="Tracheophyta" rank="species" species="tuberosa">
<emphasis id="A6FDEAD4FFEC4138FD19FE1D0682D5B4" box="[743,982,479,505]" italics="true" pageId="5" pageNumber="516">Phlomoides tuberosa</emphasis>
</taxonomicName>
(L.) Moench (
<taxonomicName id="53894D45FFEC4138FB7DFE1D0056D5B4" ID-CoL="BR9" box="[1155,1282,479,505]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="516" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
:
<taxonomicName id="53894D45FFEC4138FAEEFE1D00CCD5B4" authorityName="Harley" authorityYear="2003" box="[1296,1432,479,505]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="516" phylum="Tracheophyta" rank="subFamily" subFamily="Lamioideae">Lamioideae</taxonomicName>
; synonym
<taxonomicName id="53894D45FFEC4138FEF8FDC1049FD650" ID-CoL="77BVK" authority="L." box="[262,459,515,541]" class="Magnoliopsida" family="Lamiaceae" genus="Phlomis" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="516" phylum="Tracheophyta" rank="species" species="tuberosa">
<emphasis id="A6FDEAD4FFEC4138FEF8FDC1049FD650" box="[262,459,515,541]" italics="true" pageId="5" pageNumber="516">Phlomis tuberosa</emphasis>
</taxonomicName>
L.) (
<bibRefCitation id="F0184B37FFEC4138FDFAFDC107F2D650" author="Kieffer, J. J." box="[516,678,515,541]" pageId="5" pageNumber="516" refId="ref9612" refString="Kieffer, J. J. (1901 a) Monographie des Cynipides d'Europe et d'Algerie: Les Cynipides. Froment-Dubosclard, Paris, 687 pp." type="book" year="1901">Kieffer 1901a</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEC4138FD4DFDC107A9D650" author="Kieffer, J. J." box="[691,765,515,541]" pageId="5" pageNumber="516" pagination="233 - 579" refId="ref9639" refString="Kieffer, J. J. (1901 b) Synopsis de Zoocecidies d'Europe. Annalles de la Societe Entomologique de France, 1901, 233 - 579. https: // doi. org / 10.1080 / 21686351.1901.12279879" type="journal article" year="1901">1901b</bibRefCitation>
).
<bibRefCitation id="F0184B37FFEC4138FCECFDC10695D650" author="Kieffer, J. J." box="[786,961,515,541]" pageId="5" pageNumber="516" refId="ref9612" refString="Kieffer, J. J. (1901 a) Monographie des Cynipides d'Europe et d'Algerie: Les Cynipides. Froment-Dubosclard, Paris, 687 pp." type="book" year="1901">Kieffer (1901a)</bibRefCitation>
first described this genus under the name
<taxonomicName id="53894D45FFEC4138FF69FDE505A2D60C" ID-CoL="8NWNQ" authorityName="Kieffer" authorityYear="1901" box="[151,246,551,577]" class="Insecta" family="Cynipidae" genus="Pantelia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FF69FDE505A2D60C" box="[151,246,551,577]" italics="true" pageId="5" pageNumber="516">Pantelia</emphasis>
</taxonomicName>
, but this name was preoccupied by an orthopteran genus;
<bibRefCitation id="F0184B37FFEC4138FC66FDE5011FD60C" author="Kieffer, J. J." box="[920,1099,551,577]" pageId="5" pageNumber="516" pagination="233 - 579" refId="ref9639" refString="Kieffer, J. J. (1901 b) Synopsis de Zoocecidies d'Europe. Annalles de la Societe Entomologique de France, 1901, 233 - 579. https: // doi. org / 10.1080 / 21686351.1901.12279879" type="journal article" year="1901">Kieffer (1901b)</bibRefCitation>
thus applied the replacement name
<taxonomicName id="53894D45FFEC4138FF25FD890402D628" authorityName="Kieffer" authorityYear="1901" box="[219,342,587,613]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FF25FD890402D628" box="[219,342,587,613]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
</taxonomicName>
shortly thereafter.
</paragraph>
</subSubSection>
<subSubSection id="DC93654DFFEC413BFF39FDB20077D59C" lastPageId="6" lastPageNumber="517" pageId="5" pageNumber="516" type="description">
<paragraph id="943636C6FFEC4138FF39FDB207B9D783" blockId="5.[151,1437,407,2018]" pageId="5" pageNumber="516">
In the following century, several authors described additional genera and species of herb gall wasps associated with
<taxonomicName id="53894D45FFEC4138FF2FFD5104EAD6E0" box="[209,446,659,685]" class="Magnoliopsida" family="Lamiaceae" genus="Phlomoides" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="516" phylum="Tracheophyta" rank="species" species="tuberosa">
<emphasis id="A6FDEAD4FFEC4138FF2FFD5104EAD6E0" box="[209,446,659,685]" italics="true" pageId="5" pageNumber="516">Phlomoides tuberosa</emphasis>
</taxonomicName>
(L.) Moench, all of which have since been synonymized with
<taxonomicName id="53894D45FFEC4138FB78FD5100C1D6E0" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[1158,1429,659,685]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEC4138FB78FD5100C1D6E0" box="[1158,1429,659,685]" italics="true" pageId="5" pageNumber="516">Panteliella fedtschenkoi</emphasis>
</taxonomicName>
.
<bibRefCitation id="F0184B37FFEC4138FF69FD750468D69F" author="Belizin, V. I." box="[151,316,695,722]" pageId="5" pageNumber="516" pagination="662 - 674" refId="ref9247" refString="Belizin, V. I. (1959) Gall wasps of the tribe Aylaxini (Hymenoptera, Cynipoidea) new for the fauna of the U. S. S. R. Entomologicheskoe Obozrenie, 38, 662 - 674." type="book chapter" year="1959">Belizin (1959)</bibRefCitation>
described a morphologically similar species,
<taxonomicName id="53894D45FFEC4138FCB4FD7A0173D69C" authorityName="Belizin" authorityYear="1959" box="[842,1063,696,721]" class="Insecta" family="Cynipidae" genus="Vetustia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="investigata">
<emphasis id="A6FDEAD4FFEC4138FCB4FD7A0173D69C" box="[842,1063,696,721]" italics="true" pageId="5" pageNumber="516">Vetustia investigata</emphasis>
</taxonomicName>
, from galls in the inflorescence.
<bibRefCitation id="F0184B37FFEC4138FF69FD1E049CD6B8" author="Ionescu, M. A. &amp; Roman, N." box="[151,456,731,758]" pageId="5" pageNumber="516" pagination="221 - 225" refId="ref9557" refString="Ionescu, M. A. &amp; Roman, N. (1960) Un genre nouveau de Cynipides gallicoles Endocaulonia bicolor n. g., n. sp. (Hymenoptera, Cynipoidea). Revue de Biologie Academiei Republicii Populare Romine Bucaresti, 5 (3), 221 - 225." type="journal article" year="1960">Ionescu &amp; Roman (1960)</bibRefCitation>
described another similar species,
<taxonomicName id="53894D45FFEC4138FC91FD19013FD6B8" authorityName="Ionescu &amp; Roman" authorityYear="1960" box="[879,1131,731,757]" class="Insecta" family="Cynipidae" genus="Endocaulonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A6FDEAD4FFEC4138FC91FD19013FD6B8" box="[879,1131,731,757]" italics="true" pageId="5" pageNumber="516">Endocaulonia bicolor</emphasis>
</taxonomicName>
, from cryptic stem galls.
<bibRefCitation id="F0184B37FFEC4138FF69FD3D0417D754" author="Quinlan, J." box="[151,323,767,793]" pageId="5" pageNumber="516" pagination="275 - 286" refId="ref10265" refString="Quinlan, J. (1968) Cynipinae (Hymenoptera) occurring on Phlomis L. Transactions of the Royal Entomological Society of London, 120, 275 - 286. https: // doi. org / 10.1111 / j. 1365 - 2311.1968. tb 00339. x" type="journal article" year="1968">Quinlan (1968)</bibRefCitation>
examined
<taxonomicName id="53894D45FFEC4138FE3EFD3D076FD754" authorityName="Kieffer" authorityYear="1901" box="[448,571,767,793]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FE3EFD3D076FD754" box="[448,571,767,793]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
</taxonomicName>
along with other cynipids then associated with
<taxonomicName id="53894D45FFEC4138FBABFD3D01E5D754" box="[1109,1201,767,793]" class="Magnoliopsida" family="Lamiaceae" genus="Phlomis" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="516" phylum="Tracheophyta" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FBABFD3D01E5D754" box="[1109,1201,767,793]" italics="true" pageId="5" pageNumber="516">Phlomis</emphasis>
</taxonomicName>
L., providing a brief redescription of
<taxonomicName id="53894D45FFEC4138FEB5FCE10703D770" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[331,599,803,829]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEC4138FEB5FCE10703D770" box="[331,599,803,829]" italics="true" pageId="5" pageNumber="516">Panteliella fedtschenkoi</emphasis>
</taxonomicName>
and treating
<taxonomicName id="53894D45FFEC4138FD1BFCE606E9D770" authorityName="Belizin" authorityYear="1959" box="[741,957,804,829]" class="Insecta" family="Cynipidae" genus="Vetustia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="investigata">
<emphasis id="A6FDEAD4FFEC4138FD1BFCE606E9D770" box="[741,957,804,829]" italics="true" pageId="5" pageNumber="516">Vetustia investigata</emphasis>
</taxonomicName>
, but did not include
<taxonomicName id="53894D45FFEC4138FB62FCE1006DD770" authorityName="Ionescu &amp; Roman" authorityYear="1960" box="[1180,1337,803,829]" class="Insecta" family="Cynipidae" genus="Endocaulonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FB62FCE1006DD770" box="[1180,1337,803,829]" italics="true" pageId="5" pageNumber="516">Endocaulonia</emphasis>
</taxonomicName>
.
<bibRefCitation id="F0184B37FFEC4138FABBFCE1046DD72C" author="Nieves-Aldrey, J. L." pageId="5" pageNumber="516" pagination="175 - 206" refId="ref10135" refString="Nieves-Aldrey, J. L. (1994) Revision of West-European Genera of the Tribe Aylacini Ashmead (Hymenoptera, Cynipidae). Journal of Hymenoptera Research, 3, 175 - 206." type="journal article" year="1994">Nieves-Aldrey (1994)</bibRefCitation>
synonymized
<taxonomicName id="53894D45FFEC4138FE1CFC85072BD72C" authorityName="Ionescu &amp; Roman" authorityYear="1960" box="[482,639,839,865]" class="Insecta" family="Cynipidae" genus="Endocaulonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FE1CFC85072BD72C" box="[482,639,839,865]" italics="true" pageId="5" pageNumber="516">Endocaulonia</emphasis>
</taxonomicName>
with
<emphasis id="A6FDEAD4FFEC4138FD3BFC850614D72C" box="[709,832,839,865]" italics="true" pageId="5" pageNumber="516">Panteleilla</emphasis>
, without establishing synonyms at the species level.
<bibRefCitation id="F0184B37FFEC4138FF69FCA9046DD7C8" author="Melika, G." box="[151,313,875,901]" pageId="5" pageNumber="516" refId="ref9772" refString="Melika, G. (2006) Gall wasps of Ukraine. Vol. 1. Vestnik zoologii, The Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev, 300 pp." type="book" year="2006">Melika (2006)</bibRefCitation>
synonymized
<taxonomicName id="53894D45FFEC4138FE20FCAE07EDD7C8" authorityName="Belizin" authorityYear="1959" box="[478,697,876,901]" class="Insecta" family="Cynipidae" genus="Vetustia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="investigata">
<emphasis id="A6FDEAD4FFEC4138FE20FCAE07EDD7C8" box="[478,697,876,901]" italics="true" pageId="5" pageNumber="516">Vetustia investigata</emphasis>
</taxonomicName>
and
<taxonomicName id="53894D45FFEC4138FD0FFCA90691D7C8" authorityName="Nieves-Aldrey" authorityYear="1994" baseAuthorityName="Ionescu &amp; Roman" baseAuthorityYear="1960" box="[753,965,875,901]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A6FDEAD4FFEC4138FD0FFCA90691D7C8" box="[753,965,875,901]" italics="true" pageId="5" pageNumber="516">Panteliella bicolor</emphasis>
</taxonomicName>
with
<emphasis id="A6FDEAD4FFEC4138FBF8FCAE07B2D780" italics="true" pageId="5" pageNumber="516">
<taxonomicName id="53894D45FFEC4138FBF8FCAE01E5D7C8" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[1030,1201,875,901]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">P. fedtschenkoi</taxonomicName>
; the latter synonymy was not discussed beyond mentioning
<taxonomicName id="53894D45FFEC4138FDB1FC5207E9D7E4" authorityName="Nieves-Aldrey" authorityYear="1994" baseAuthorityName="Ionescu &amp; Roman" baseAuthorityYear="1960" box="[591,701,911,937]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="bicolor">P. bicolor</taxonomicName>
as a synonym of
<taxonomicName id="53894D45FFEC4138FC74FC520162D7E4" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[906,1078,911,937]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">P. fedtschenkoi</taxonomicName>
, although some discussion was given regarding the synonymy of
<taxonomicName id="53894D45FFEC4138FDF5FC7607B2D780" authorityName="Belizin" authorityYear="1959" box="[523,742,948,973]" class="Insecta" family="Cynipidae" genus="Vetustia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="investigata">Vetustia investigata</taxonomicName>
</emphasis>
.
</paragraph>
<paragraph id="943636C6FFEC4138FF39FC1A014DD15C" blockId="5.[151,1437,407,2018]" pageId="5" pageNumber="516">
No comprehensive revision has yet addressed all names included in
<taxonomicName id="53894D45FFEC4138FC31FC15011ED7BC" authorityName="Kieffer" authorityYear="1901" box="[975,1098,983,1009]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FC31FC15011ED7BC" box="[975,1098,983,1009]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
</taxonomicName>
. After synonymies by
<bibRefCitation id="F0184B37FFEC4138FAB3FC1505B6D058" author="Melika, G." pageId="5" pageNumber="516" refId="ref9772" refString="Melika, G. (2006) Gall wasps of Ukraine. Vol. 1. Vestnik zoologii, The Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev, 300 pp." type="book" year="2006">Melika (2006)</bibRefCitation>
and
<bibRefCitation id="F0184B37FFEC4138FEDFFC39074AD058" author="Nieves-Aldrey, J. L." box="[289,542,1019,1045]" pageId="5" pageNumber="516" pagination="175 - 206" refId="ref10135" refString="Nieves-Aldrey, J. L. (1994) Revision of West-European Genera of the Tribe Aylacini Ashmead (Hymenoptera, Cynipidae). Journal of Hymenoptera Research, 3, 175 - 206." type="journal article" year="1994">Nieves-Aldrey (1994)</bibRefCitation>
, only the species
<taxonomicName id="53894D45FFEC4138FD08FC3E0626D058" authorityName="Vyrzhikovskaya" authorityYear="1962" box="[758,882,1019,1045]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="bianchii">
<emphasis id="A6FDEAD4FFEC4138FD08FC3E0626D058" box="[758,882,1019,1045]" italics="true" pageId="5" pageNumber="516">P. bianchii</emphasis>
</taxonomicName>
and
<taxonomicName id="53894D45FFEC4138FC4FFC3E0109D058" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[945,1117,1019,1045]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEC4138FC4FFC3E0109D058" box="[945,1117,1019,1045]" italics="true" pageId="5" pageNumber="516">P. fedtschenkoi</emphasis>
</taxonomicName>
remain valid, although the former species has not been mentioned in the literature aside from its original description (
<bibRefCitation id="F0184B37FFEC4138FB71FBDD00C4D074" author="Vyrzhikovskaya, A. V." box="[1167,1424,1055,1081]" pageId="5" pageNumber="516" pagination="138 - 171" refId="ref10548" refString="Vyrzhikovskaya, A. V. (1962) Gall-wasps (Hymenoptera, Cynipoidea, Cynipinae) of the Leningrad Region. Proceedings of the Zoological Institute of the USSR Academy of Sciences, XXXI, 138 - 171. [in Russian]" type="book chapter" year="1962">Vyrzhikovskaya, 1962</bibRefCitation>
). Considering the biologies of synonymized species,
<taxonomicName id="53894D45FFEC4138FD24FB8606D6D010" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[730,898,1091,1117]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEC4138FD24FB8606D6D010" box="[730,898,1091,1117]" italics="true" pageId="5" pageNumber="516">P. fedtschenkoi</emphasis>
</taxonomicName>
is currently hypothesized to induce galls on the leaves and in the stems and inflorescences (
<bibRefCitation id="F0184B37FFEC4138FD7CFBA50642D0CC" author="Melika, G." box="[642,790,1127,1153]" pageId="5" pageNumber="516" refId="ref9772" refString="Melika, G. (2006) Gall wasps of Ukraine. Vol. 1. Vestnik zoologii, The Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev, 300 pp." type="book" year="2006">Melika 2006</bibRefCitation>
), which is highly atypical for a gall wasp species. Melika mentions several adult morphological characters that seem to differ depending on the plant organ from which the specimen was reared, particularly the body coloration and the distinction of the notauli. Melika also states that the number of antennomeres in the female varies from
<quantity id="53719B23FFEC4138FD21FB110608D0A0" box="[735,860,1235,1262]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.556" metricValueMin="3.302" pageId="5" pageNumber="516" unit="in" value="13.5" valueMax="14.0" valueMin="13.0">13 to 14 in</quantity>
some specimens, and other characters, such as the dimensions of the marginal cell (there termed the radial cell), vary considerably.
</paragraph>
<paragraph id="943636C6FFEC4138FF39FAD90718D240" blockId="5.[151,1437,407,2018]" pageId="5" pageNumber="516">
Further complicating the taxonomy of
<taxonomicName id="53894D45FFEC4138FD8AFAD907BBD178" authorityName="Kieffer" authorityYear="1901" box="[628,751,1307,1333]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FD8AFAD907BBD178" box="[628,751,1307,1333]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
</taxonomicName>
is differential treatments of synonymous taxa in phylogenetic studies.
<bibRefCitation id="F0184B37FFEC4138FF0BFAFD076DD114" author="Liljeblad, J. &amp; Ronquist, F." box="[245,569,1343,1369]" pageId="5" pageNumber="516" pagination="229 - 252" refId="ref9718" refString="Liljeblad, J. &amp; Ronquist, F. (1998) A phylogenetic analysis of higher-level gall wasp relationships (Hymenoptera: Cynipidae). Systematic Entomology, 23 (3), 229 - 252. https: // doi. org / 10.1046 / j. 1365 - 3113.1998.00053. x" type="journal article" year="1998">Liljeblad &amp; Ronquist (1998)</bibRefCitation>
included
<taxonomicName id="53894D45FFEC4138FD55FAFD06D4D114" authorityName="Nieves-Aldrey" authorityYear="1994" baseAuthorityName="Ionescu &amp; Roman" baseAuthorityYear="1960" box="[683,896,1343,1369]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A6FDEAD4FFEC4138FD55FAFD06D4D114" box="[683,896,1343,1369]" italics="true" pageId="5" pageNumber="516">Panteliella bicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="53894D45FFEC4138FC47FA8201C2D114" authorityName="Belizin" authorityYear="1959" box="[953,1174,1344,1369]" class="Insecta" family="Cynipidae" genus="Vetustia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="investigata">
<emphasis id="A6FDEAD4FFEC4138FC47FA8201C2D114" box="[953,1174,1344,1369]" italics="true" pageId="5" pageNumber="516">Vetustia investigata</emphasis>
</taxonomicName>
in their morphological phylogenetic analysis, but they did not treat the
<typeStatus id="4B328864FFEC4138FD39FAA607A3D133" box="[711,759,1380,1406]" pageId="5" pageNumber="516">type</typeStatus>
species
<taxonomicName id="53894D45FFEC4138FCA2FAA60152D130" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[860,1030,1379,1405]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEC4138FCA2FAA60152D130" box="[860,1030,1379,1405]" italics="true" pageId="5" pageNumber="516">P. fedtschenkoi</emphasis>
</taxonomicName>
.
<bibRefCitation id="F0184B37FFEC4138FBE9FAA10042D133" author="Ronquist, F. &amp; Nieves-Aldrey, J. L. &amp; Buffington, M. L. &amp; Liu, Z. &amp; Liljeblad, J. &amp; Nylander, J. A." box="[1047,1302,1379,1406]" pageId="5" pageNumber="516" refId="ref10342" refString="Ronquist, F., Nieves-Aldrey, J. L., Buffington, M. L., Liu, Z., Liljeblad, J. &amp; Nylander, J. A. (2015) Phylogeny, evolution and classification of gall wasps: the plot thickens. PLoS ONE, 10 (5), e 0123301. https: // doi. org / 10.1371 / journal. pone. 0123301" type="journal volume" year="2015">
Ronquist
<emphasis id="A6FDEAD4FFEC4138FB78FAA60195D130" box="[1158,1217,1379,1405]" italics="true" pageId="5" pageNumber="516">et al.</emphasis>
(2015)
</bibRefCitation>
based their morphological analysis on Liljeblad &amp; Ronquist’s character set, apparently without considering the synonymies published by
<bibRefCitation id="F0184B37FFEC4138FECFFA690483D188" author="Melika, G." box="[305,471,1451,1477]" pageId="5" pageNumber="516" refId="ref9772" refString="Melika, G. (2006) Gall wasps of Ukraine. Vol. 1. Vestnik zoologii, The Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev, 300 pp." type="book" year="2006">Melika (2006)</bibRefCitation>
. The “
<taxonomicName id="53894D45FFEC4138FDDDFA6E07C6D188" authorityName="Nieves-Aldrey" authorityYear="1994" baseAuthorityName="Ionescu &amp; Roman" baseAuthorityYear="1960" box="[547,658,1451,1477]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A6FDEAD4FFEC4138FDDDFA6E07C6D188" box="[547,658,1451,1477]" italics="true" pageId="5" pageNumber="516">P. bicolor</emphasis>
</taxonomicName>
” specimens studied in these analyses were reared from stems of the known host plant species in
<collectingCountry id="EC9E7656FFEC4138FE21FA120711D1A7" box="[479,581,1488,1514]" name="Hungary" pageId="5" pageNumber="516">Hungary</collectingCountry>
; the same stems apparently also yielded specimens of
<taxonomicName id="53894D45FFEC4138FB47FA0D047AD243" authority="Belizin, 1959" authorityName="Belizin" authorityYear="1959" class="Insecta" family="Cynipidae" genus="Aulacidea" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="phlomica">
<emphasis id="A6FDEAD4FFEC4138FB47FA0D00C8D1A4" box="[1209,1436,1487,1513]" italics="true" pageId="5" pageNumber="516">Aulacidea phlomica</emphasis>
<bibRefCitation id="F0184B37FFEC4138FF69FA31047AD243" author="Belizin, V. I." box="[151,302,1523,1550]" pageId="5" pageNumber="516" pagination="662 - 674" refId="ref9247" refString="Belizin, V. I. (1959) Gall wasps of the tribe Aylaxini (Hymenoptera, Cynipoidea) new for the fauna of the U. S. S. R. Entomologicheskoe Obozrenie, 38, 662 - 674." type="book chapter" year="1959">Belizin, 1959</bibRefCitation>
</taxonomicName>
(Ronquist, pers. comm.).
</paragraph>
<paragraph id="943636C6FFEC413BFF39F9D50035D4B4" blockId="5.[151,1437,407,2018]" lastBlockId="6.[151,1436,151,465]" lastPageId="6" lastPageNumber="517" pageId="5" pageNumber="516">
Altogether,
<taxonomicName id="53894D45FFEC4138FEB5F9DA04A6D27C" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[331,498,1559,1585]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEC4138FEB5F9DA04A6D27C" box="[331,498,1559,1585]" italics="true" pageId="5" pageNumber="516">P. fedtschenkoi</emphasis>
</taxonomicName>
requires further revision, to determine the conspecificity of species currently treated as synonyms, and we presently expect that multiple valid species will be identified among the broad concept of
<taxonomicName id="53894D45FFEC4138FF69F9A20415D234" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[151,321,1631,1657]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEC4138FF69F9A20415D234" box="[151,321,1631,1657]" italics="true" pageId="5" pageNumber="516">P. fedtschenkoi</emphasis>
</taxonomicName>
employed in prior studies (
<emphasis id="A6FDEAD4FFEC4138FD7DF9A207F0D234" box="[643,676,1632,1657]" italics="true" pageId="5" pageNumber="516">e.g</emphasis>
.,
<bibRefCitation id="F0184B37FFEC4138FD42F99D0606D234" author="Melika, G." box="[700,850,1631,1657]" pageId="5" pageNumber="516" refId="ref9772" refString="Melika, G. (2006) Gall wasps of Ukraine. Vol. 1. Vestnik zoologii, The Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev, 300 pp." type="book" year="2006">Melika 2006</bibRefCitation>
). The synonymy of
<taxonomicName id="53894D45FFEC4138FBC5F99D01E2D234" authorityName="Kieffer" authorityYear="1901" box="[1083,1206,1631,1657]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FBC5F99D01E2D234" box="[1083,1206,1631,1657]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
</taxonomicName>
,
<taxonomicName id="53894D45FFEC4138FB38F99D0037D234" authorityName="Ionescu &amp; Roman" authorityYear="1960" box="[1222,1379,1631,1657]" class="Insecta" family="Cynipidae" genus="Endocaulonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FB38F99D0037D234" box="[1222,1379,1631,1657]" italics="true" pageId="5" pageNumber="516">Endocaulonia</emphasis>
</taxonomicName>
, and
<taxonomicName id="53894D45FFEC4138FF69F94605A4D2D0" authorityName="Belizin" authorityYear="1959" box="[151,240,1668,1693]" class="Insecta" family="Cynipidae" genus="Vetustia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FF69F94605A4D2D0" box="[151,240,1668,1693]" italics="true" pageId="5" pageNumber="516">Vetustia</emphasis>
</taxonomicName>
should also be verified through further study. To facilitate continuing work on
<taxonomicName id="53894D45FFEC4138FBAAF941019BD2D0" authorityName="Kieffer" authorityYear="1901" box="[1108,1231,1667,1693]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FBAAF941019BD2D0" box="[1108,1231,1667,1693]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
</taxonomicName>
, we redescribe the genus below, and also provide a redescription of
<taxonomicName id="53894D45FFEC4138FD4EF96A0602D28C" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[688,854,1703,1729]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEC4138FD4EF96A0602D28C" box="[688,854,1703,1729]" italics="true" pageId="5" pageNumber="516">P. fedtschenkoi</emphasis>
</taxonomicName>
. The specimen collected in
<collectingCountry id="EC9E7656FFEC4138FB77F96A005ED28F" box="[1161,1290,1704,1730]" name="Kyrgyzstan" pageId="5" pageNumber="516">Kyrgyzstan</collectingCountry>
substantially differs from the
<taxonomicName id="53894D45FFEC4138FEB5F90E04A6D2A8" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[331,498,1739,1765]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEC4138FEB5F90E04A6D2A8" box="[331,498,1739,1765]" italics="true" pageId="5" pageNumber="516">P. fedtschenkoi</emphasis>
</taxonomicName>
we examined, justifying its establishment as a new species. We describe it here as
<taxonomicName id="53894D45FFEC4138FA76F90E046AD344" authority="Nastasi" authorityName="Nastasi &amp; Deans" authorityYear="2024" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="rugosa" status="sp. nov.">
<emphasis id="A6FDEAD4FFEC4138FA76F90E05B1D344" italics="true" pageId="5" pageNumber="516">P. rugosa</emphasis>
Nastasi
</taxonomicName>
<emphasis id="A6FDEAD4FFEC4138FEBBF92D04CAD344" bold="true" box="[325,414,1775,1801]" pageId="5" pageNumber="516">
<taxonomicNameLabel id="BDCE57AFFFEC4138FEBBF92D04CAD344" box="[325,414,1775,1801]" pageId="5" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Additionally, the species
<taxonomicName id="53894D45FFEC4138FD42F9320667D344" authorityName="Vyrzhikovskaya" authorityYear="1962" box="[700,819,1775,1801]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="bianchii">
<emphasis id="A6FDEAD4FFEC4138FD42F9320667D344" box="[700,819,1775,1801]" italics="true" pageId="5" pageNumber="516">P. bianchii</emphasis>
</taxonomicName>
has only been mentioned in its original description and has not been treated by any further studies despite several subsequent works concerning
<taxonomicName id="53894D45FFEC4138FB77F8D10050D360" authorityName="Kieffer" authorityYear="1901" box="[1161,1284,1811,1837]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FB77F8D10050D360" box="[1161,1284,1811,1837]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
</taxonomicName>
(
<emphasis id="A6FDEAD4FFEC4138FAEBF8D60010D360" box="[1301,1348,1812,1837]" italics="true" pageId="5" pageNumber="516">e.g.,</emphasis>
<bibRefCitation id="F0184B37FFEC4138FAB3F8D10586D31C" author="Melika, G." pageId="5" pageNumber="516" refId="ref9772" refString="Melika, G. (2006) Gall wasps of Ukraine. Vol. 1. Vestnik zoologii, The Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev, 300 pp." type="book" year="2006">Melika 2006</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEC4138FF25F8F50496D31C" author="Nieves-Aldrey, J. L." box="[219,450,1847,1873]" pageId="5" pageNumber="516" pagination="393 - 413" refId="ref10170" refString="Nieves-Aldrey, J. L. (2022) Description of Fumariphilus Nieves-Aldrey, gen. nov., a new genus of herb gall wasps, with a key to genera of the tribe Aulacideini (Hymenoptera: Cynipidae). Zootaxa, 5155 (3), 393 - 413. https: // doi. org / 10.11646 / zootaxa. 5155.3.5" type="journal article" year="2022">Nieves-Aldrey 2022</bibRefCitation>
). We have not examined the
<typeStatus id="4B328864FFEC4138FD09F8FA0673D31F" box="[759,807,1848,1874]" pageId="5" pageNumber="516">type</typeStatus>
material, but based on its original description,
<taxonomicName id="53894D45FFEC4138FAD9F8FA00C8D31C" authorityName="Vyrzhikovskaya" authorityYear="1962" box="[1319,1436,1847,1873]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="bianchii">
<emphasis id="A6FDEAD4FFEC4138FAD9F8FA00C8D31C" box="[1319,1436,1847,1873]" italics="true" pageId="5" pageNumber="516">P. bianchii</emphasis>
</taxonomicName>
appears to represent a valid species. To assist in resolving taxonomic issues in
<taxonomicName id="53894D45FFEC4138FBF5F89901D2D338" authorityName="Kieffer" authorityYear="1901" box="[1035,1158,1883,1909]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEC4138FBF5F89901D2D338" box="[1035,1158,1883,1909]" italics="true" pageId="5" pageNumber="516">Panteliella</emphasis>
</taxonomicName>
, we provide a key to the three known species, including
<taxonomicName id="53894D45FFEC4138FE03F8420721D3D4" authorityName="Vyrzhikovskaya" authorityYear="1962" box="[509,629,1919,1945]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="bianchii">
<emphasis id="A6FDEAD4FFEC4138FE03F8420721D3D4" box="[509,629,1919,1945]" italics="true" pageId="5" pageNumber="516">P. bianchii</emphasis>
</taxonomicName>
based on characters present in its original description. We also provide a translation of the original description of
<emphasis id="A6FDEAD4FFEC4138FD87F8660113D3F0" box="[633,1095,1955,1981]" italics="true" pageId="5" pageNumber="516">
<taxonomicName id="53894D45FFEC4138FD87F86607A6D3F0" authorityName="Vyrzhikovskaya" authorityYear="1962" box="[633,754,1955,1981]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="516" phylum="Arthropoda" rank="species" species="bianchii">P. bianchii</taxonomicName>
(see below species treatment)
</emphasis>
. Lastly, we provide a table of diagnostic characters for the three valid species we recognize as well as the two species presently synonymized with
<taxonomicName id="53894D45FFEF413BFF31FF550422D4FC" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[207,374,151,177]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEF413BFF31FF550422D4FC" box="[207,374,151,177]" italics="true" pageId="6" pageNumber="517">P. fedtschenkoi</emphasis>
</taxonomicName>
(
<tableCitation id="D90B037DFFEF413BFE7DFF550482D4FC" box="[387,470,151,177]" captionStart="TABLE 1" captionStartId="6.[152,244,512,536]" captionTargetPageId="6" captionText="TABLE 1. Major characters of valid Panteliella species and synonyms of P. fedtschenkoi. *: Material not examined here; characters based on treatments by Belizin (1959), Ionescu &amp; Roman (1960), Vyrzhikovskaya (1962), Quinlan (1968), Nieves-Aldrey (1994), Liljeblad &amp; Ronquist (1998), Melika (2006)." pageId="6" pageNumber="517">Table 1</tableCitation>
). Given the high degree of morphological divergence of
<taxonomicName id="53894D45FFEF413BFBB0FF55007CD4FD" authorityName="Belizin" authorityYear="1959" box="[1102,1320,151,176]" class="Insecta" family="Cynipidae" genus="Vetustia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="species" species="investigata">
<emphasis id="A6FDEAD4FFEF413BFBB0FF55007CD4FD" box="[1102,1320,151,176]" italics="true" pageId="6" pageNumber="517">Vetustia investigata</emphasis>
</taxonomicName>
(currently placed as a synonym of
<taxonomicName id="53894D45FFEF413BFE59FF79071AD498" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[423,590,187,213]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEF413BFE59FF79071AD498" box="[423,590,187,213]" italics="true" pageId="6" pageNumber="517">P. fedtschenkoi</emphasis>
</taxonomicName>
; see below species treatment and
<tableCitation id="D90B037DFFEF413BFC33FF79014BD498" box="[973,1055,187,213]" captionStart="TABLE 1" captionStartId="6.[152,244,512,536]" captionTargetPageId="6" captionText="TABLE 1. Major characters of valid Panteliella species and synonyms of P. fedtschenkoi. *: Material not examined here; characters based on treatments by Belizin (1959), Ionescu &amp; Roman (1960), Vyrzhikovskaya (1962), Quinlan (1968), Nieves-Aldrey (1994), Liljeblad &amp; Ronquist (1998), Melika (2006)." pageId="6" pageNumber="517">Table 1</tableCitation>
), it is evident that our description and diagnosis for
<taxonomicName id="53894D45FFEF413BFEA1FF1D048ED4B4" authorityName="Kieffer" authorityYear="1901" box="[351,474,223,249]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFEA1FF1D048ED4B4" box="[351,474,223,249]" italics="true" pageId="6" pageNumber="517">Panteliella</emphasis>
</taxonomicName>
will likely require revision after
<typeStatus id="4B328864FFEF413BFCB2FF1D0628D4B4" box="[844,892,223,249]" pageId="6" pageNumber="517">type</typeStatus>
material of these species can be examined.
</paragraph>
<paragraph id="943636C6FFEF413BFF39FEC10077D59C" blockId="6.[151,1436,151,465]" pageId="6" pageNumber="517">
More broadly, the diagnostic characters for
<taxonomicName id="53894D45FFEF413BFD50FEC1067DD550" authorityName="Kieffer" authorityYear="1901" box="[686,809,259,285]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFD50FEC1067DD550" box="[686,809,259,285]" italics="true" pageId="6" pageNumber="517">Panteliella</emphasis>
</taxonomicName>
given in the literature vary substantially and complicate the recognition of the genus. For instance, in the last published generic key (
<bibRefCitation id="F0184B37FFEF413BFBEEFEE501A9D50C" author="Nieves-Aldrey, J. L." box="[1040,1277,295,321]" pageId="6" pageNumber="517" pagination="393 - 413" refId="ref10170" refString="Nieves-Aldrey, J. L. (2022) Description of Fumariphilus Nieves-Aldrey, gen. nov., a new genus of herb gall wasps, with a key to genera of the tribe Aulacideini (Hymenoptera: Cynipidae). Zootaxa, 5155 (3), 393 - 413. https: // doi. org / 10.11646 / zootaxa. 5155.3.5" type="journal article" year="2022">Nieves-Aldrey 2022</bibRefCitation>
), a diagnostic character for
<taxonomicName id="53894D45FFEF413BFED1FE8904FED528" authorityName="Kieffer" authorityYear="1901" box="[303,426,331,357]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFED1FE8904FED528" box="[303,426,331,357]" italics="true" pageId="6" pageNumber="517">Panteliella</emphasis>
</taxonomicName>
is the lack of defined lateral propodeal carinae. However, in all specimens we examined here, there are distinct lateral propodeal carinae. To address these conflicting elements, we provide a redescription, based on the material we examined in this study, particularly isotypic specimens of
<taxonomicName id="53894D45FFEF413BFBBEFE5101BCD5E0" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[1088,1256,403,429]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEF413BFBBEFE5101BCD5E0" box="[1088,1256,403,429]" italics="true" pageId="6" pageNumber="517">P. fedtschenkoi</emphasis>
</taxonomicName>
(see below). As mentioned above, further studies will be needed to resolve the generic and specific limits of
<taxonomicName id="53894D45FFEF413BFB5FFE750048D59C" authorityName="Kieffer" authorityYear="1901" box="[1185,1308,439,465]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFB5FFE750048D59C" box="[1185,1308,439,465]" italics="true" pageId="6" pageNumber="517">Panteliella</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="C0F6664EFFEF413BFF66FDC20696D62D" pageId="6" pageNumber="517" startId="6.[152,244,512,536]" targetBox="[163,1374,624,963]" targetIsTable="true" targetPageId="6" targetType="table">
<paragraph id="943636C6FFEF413BFF66FDC20696D62D" blockId="6.[151,1437,512,609]" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFF66FDC201DCD654" bold="true" box="[152,1160,512,537]" pageId="6" pageNumber="517">
TABLE 1. Major characters of valid
<taxonomicName id="53894D45FFEF413BFDBFFDC20659D655" box="[577,781,512,536]" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFDBFFDC207ECD655" bold="true" box="[577,696,512,536]" italics="true" pageId="6" pageNumber="517">Panteliella</emphasis>
species
</taxonomicName>
and synonyms of
<taxonomicName id="53894D45FFEF413BFC25FDC301D4D655" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[987,1152,512,536]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEF413BFC25FDC301D4D655" bold="true" box="[987,1152,512,536]" italics="true" pageId="6" pageNumber="517">P. fedtschenkoi</emphasis>
</taxonomicName>
.
</emphasis>
*: Material not examined here; characters based on treatments by
<bibRefCitation id="F0184B37FFEF413BFDA4FDE607AFD671" author="Belizin, V. I." box="[602,763,548,572]" pageId="6" pageNumber="517" pagination="662 - 674" refId="ref9247" refString="Belizin, V. I. (1959) Gall wasps of the tribe Aylaxini (Hymenoptera, Cynipoidea) new for the fauna of the U. S. S. R. Entomologicheskoe Obozrenie, 38, 662 - 674." type="book chapter" year="1959">Belizin (1959)</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEF413BFCF4FDE7017DD671" author="Ionescu, M. A. &amp; Roman, N." box="[778,1065,548,573]" pageId="6" pageNumber="517" pagination="221 - 225" refId="ref9557" refString="Ionescu, M. A. &amp; Roman, N. (1960) Un genre nouveau de Cynipides gallicoles Endocaulonia bicolor n. g., n. sp. (Hymenoptera, Cynipoidea). Revue de Biologie Academiei Republicii Populare Romine Bucaresti, 5 (3), 221 - 225." type="journal article" year="1960">Ionescu &amp; Roman (1960)</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEF413BFBC9FDE60063D671" author="Vyrzhikovskaya, A. V." box="[1079,1335,548,572]" pageId="6" pageNumber="517" pagination="138 - 171" refId="ref10548" refString="Vyrzhikovskaya, A. V. (1962) Gall-wasps (Hymenoptera, Cynipoidea, Cynipinae) of the Leningrad Region. Proceedings of the Zoological Institute of the USSR Academy of Sciences, XXXI, 138 - 171. [in Russian]" type="book chapter" year="1962">Vyrzhikovskaya (1962)</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEF413BFAB8FDE605B5D62D" author="Quinlan, J." pageId="6" pageNumber="517" pagination="275 - 286" refId="ref10265" refString="Quinlan, J. (1968) Cynipinae (Hymenoptera) occurring on Phlomis L. Transactions of the Royal Entomological Society of London, 120, 275 - 286. https: // doi. org / 10.1111 / j. 1365 - 2311.1968. tb 00339. x" type="journal article" year="1968">Quinlan (1968)</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEF413BFF12FD8A048DD62C" author="Nieves-Aldrey, J. L." box="[236,473,584,609]" pageId="6" pageNumber="517" pagination="175 - 206" refId="ref10135" refString="Nieves-Aldrey, J. L. (1994) Revision of West-European Genera of the Tribe Aylacini Ashmead (Hymenoptera, Cynipidae). Journal of Hymenoptera Research, 3, 175 - 206." type="journal article" year="1994">Nieves-Aldrey (1994)</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEF413BFE1AFD8A0642D62C" author="Liljeblad, J. &amp; Ronquist, F." box="[484,790,584,609]" pageId="6" pageNumber="517" pagination="229 - 252" refId="ref9718" refString="Liljeblad, J. &amp; Ronquist, F. (1998) A phylogenetic analysis of higher-level gall wasp relationships (Hymenoptera: Cynipidae). Systematic Entomology, 23 (3), 229 - 252. https: // doi. org / 10.1046 / j. 1365 - 3113.1998.00053. x" type="journal article" year="1998">Liljeblad &amp; Ronquist (1998)</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEF413BFCDFFD8A06EAD62D" author="Melika, G." box="[801,958,584,608]" pageId="6" pageNumber="517" refId="ref9772" refString="Melika, G. (2006) Gall wasps of Ukraine. Vol. 1. Vestnik zoologii, The Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev, 300 pp." type="book" year="2006">Melika (2006)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="943636C6FFEF413BFF5DFDB2000AD78E" pageId="6" pageNumber="517">
<table id="E689C466FFEFBEC2FF5DFDB2000AD78E" box="[163,1374,624,963]" gridcols="6" gridrows="6" pageId="6" pageNumber="517">
<tr id="2AB93484FFEFBEC2FF5DFDB2000AD6E1" box="[163,1374,624,684]" gridrow="0" pageId="6" pageNumber="517">
<th id="69685DF8FFEFBEC2FF5DFDB20421D6E1" box="[163,373,624,684]" gridcol="0" gridrow="0" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFF5DFDB205A6D6C5" bold="true" box="[163,242,624,648]" pageId="6" pageNumber="517">Species</emphasis>
</th>
<th id="69685DF8FFEFBEC2FE6FFDB20701D6E1" box="[401,597,624,684]" gridcol="1" gridrow="0" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFE6FFDB00701D6C5" bold="true" box="[401,597,624,651]" pageId="6" pageNumber="517">♀ # antennomeres</emphasis>
</th>
<th id="69685DF8FFEFBEC2FD8AFDB207ABD6E1" box="[628,767,624,684]" gridcol="2" gridrow="0" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFD8AFDB207ABD6C5" bold="true" box="[628,767,624,648]" pageId="6" pageNumber="517">F2:F1 length</emphasis>
</th>
<th id="69685DF8FFEFBEC2FCE0FDB206F8D6E1" box="[798,940,624,684]" gridcol="3" gridrow="0" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFCE0FDB2063AD6C5" bold="true" box="[798,878,624,648]" pageId="6" pageNumber="517">Notauli</emphasis>
</th>
<th id="69685DF8FFEFBEC2FBF9FDB201DBD6E1" box="[1031,1167,624,684]" gridcol="4" gridrow="0" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFBF9FDB20111D6E1" bold="true" box="[1031,1167,624,684]" pageId="6" pageNumber="517">Mesoscutum rugae</emphasis>
</th>
<th id="69685DF8FFEFBEC2FB28FDB2000AD6E1" box="[1238,1374,624,684]" gridcol="5" gridrow="0" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFB28FDB2001ED6C5" bold="true" box="[1238,1354,624,648]" pageId="6" pageNumber="517">Gall organ</emphasis>
</th>
</tr>
<tr id="2AB93484FFEFBEC2FF5DFD7E000AD698" box="[163,1374,700,725]" gridrow="1" pageId="6" pageNumber="517">
<th id="69685DF8FFEFBEC2FF5DFD7E0421D698" box="[163,373,700,725]" gridcol="0" gridrow="1" pageId="6" pageNumber="517">
<taxonomicName id="53894D45FFEF413BFF5DFD7F045AD698" authorityName="Vyrzhikovskaya" authorityYear="1962" box="[163,270,701,725]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="species" species="bianchii">
<emphasis id="A6FDEAD4FFEF413BFF5DFD7F045AD698" box="[163,270,701,725]" italics="true" pageId="6" pageNumber="517">P. bianchii</emphasis>
</taxonomicName>
*
</th>
<td id="69685DF8FFEFBEC2FE6FFD7E0701D698" box="[401,597,700,725]" gridcol="1" gridrow="1" pageId="6" pageNumber="517">14</td>
<td id="69685DF8FFEFBEC2FD8AFD7E07ABD698" box="[628,767,700,725]" gridcol="2" gridrow="1" pageId="6" pageNumber="517">F1 longer</td>
<td id="69685DF8FFEFBEC2FCE0FD7E06F8D698" box="[798,940,700,725]" gridcol="3" gridrow="1" pageId="6" pageNumber="517">Incomplete(?)</td>
<td id="69685DF8FFEFBEC2FBF9FD7E01DBD698" box="[1031,1167,700,725]" gridcol="4" gridrow="1" pageId="6" pageNumber="517">Absent</td>
<td id="69685DF8FFEFBEC2FB28FD7E000AD698" box="[1238,1374,700,725]" gridcol="5" gridrow="1" pageId="6" pageNumber="517">Unknown</td>
</tr>
<tr id="2AB93484FFEFBEC2FF5DFD31000AD746" box="[163,1374,755,779]" gridrow="2" pageId="6" pageNumber="517">
<th id="69685DF8FFEFBEC2FF5DFD310421D746" box="[163,373,755,779]" gridcol="0" gridrow="2" pageId="6" pageNumber="517">
<taxonomicName id="53894D45FFEF413BFF5DFD36046ED746" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[163,314,755,779]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEF413BFF5DFD36046ED746" box="[163,314,755,779]" italics="true" pageId="6" pageNumber="517">P. fedtschenkoi</emphasis>
</taxonomicName>
</th>
<td id="69685DF8FFEFBEC2FE6FFD310701D746" box="[401,597,755,779]" gridcol="1" gridrow="2" pageId="6" pageNumber="517">14</td>
<td id="69685DF8FFEFBEC2FD8AFD3107ABD746" box="[628,767,755,779]" gridcol="2" gridrow="2" pageId="6" pageNumber="517">Subequal</td>
<td id="69685DF8FFEFBEC2FCE0FD3106F8D746" box="[798,940,755,779]" gridcol="3" gridrow="2" pageId="6" pageNumber="517">Incomplete</td>
<td id="69685DF8FFEFBEC2FBF9FD3101DBD746" box="[1031,1167,755,779]" gridcol="4" gridrow="2" pageId="6" pageNumber="517">Absent</td>
<td id="69685DF8FFEFBEC2FB28FD31000AD746" box="[1238,1374,755,779]" gridcol="5" gridrow="2" pageId="6" pageNumber="517">Leaf</td>
</tr>
<tr id="2AB93484FFEFBEC2FF5DFCEB000AD70C" box="[163,1374,809,833]" gridrow="3" pageId="6" pageNumber="517">
<th id="69685DF8FFEFBEC2FF5DFCEB0421D70C" box="[163,373,809,833]" gridcol="0" gridrow="3" pageId="6" pageNumber="517">
<taxonomicName id="53894D45FFEF413BFF5DFCE80455D70C" authorityName="Nastasi &amp; Deans" authorityYear="2024" box="[163,257,810,833]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="A6FDEAD4FFEF413BFF5DFCE80455D70C" box="[163,257,810,833]" italics="true" pageId="6" pageNumber="517">P. rugosa</emphasis>
</taxonomicName>
</th>
<td id="69685DF8FFEFBEC2FE6FFCEB0701D70C" box="[401,597,809,833]" gridcol="1" gridrow="3" pageId="6" pageNumber="517">13</td>
<td id="69685DF8FFEFBEC2FD8AFCEB07ABD70C" box="[628,767,809,833]" gridcol="2" gridrow="3" pageId="6" pageNumber="517">Subequal</td>
<td id="69685DF8FFEFBEC2FCE0FCEB06F8D70C" box="[798,940,809,833]" gridcol="3" gridrow="3" pageId="6" pageNumber="517">Incomplete</td>
<td id="69685DF8FFEFBEC2FBF9FCEB01DBD70C" box="[1031,1167,809,833]" gridcol="4" gridrow="3" pageId="6" pageNumber="517">Present</td>
<td id="69685DF8FFEFBEC2FB28FCEB000AD70C" box="[1238,1374,809,833]" gridcol="5" gridrow="3" pageId="6" pageNumber="517">Unknown</td>
</tr>
<tr id="2AB93484FFEFBEC2FF5DFC9D000AD7D6" box="[163,1374,863,923]" gridrow="4" pageId="6" pageNumber="517">
<th id="69685DF8FFEFBEC2FF5DFC9D0421D7D6" box="[163,373,863,923]" gridcol="0" gridrow="4" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFF5DFC9D05ACD7D6" box="[163,305,863,923]" italics="true" pageId="6" pageNumber="517">
<taxonomicName id="53894D45FFEF413BFF5DFC9D05BBD7D6" authorityName="Ionescu &amp; Roman" authorityYear="1960" box="[163,305,863,923]" class="Insecta" family="Cynipidae" genus="Endocaulonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="species" species="bicolor">Endocaulonia bicolor</taxonomicName>
*
</emphasis>
</th>
<td id="69685DF8FFEFBEC2FE6FFC9D0701D7D6" box="[401,597,863,923]" gridcol="1" gridrow="4" pageId="6" pageNumber="517">14</td>
<td id="69685DF8FFEFBEC2FD8AFC9D07ABD7D6" box="[628,767,863,923]" gridcol="2" gridrow="4" pageId="6" pageNumber="517">Subequal</td>
<td id="69685DF8FFEFBEC2FCE0FC9D06F8D7D6" box="[798,940,863,923]" gridcol="3" gridrow="4" pageId="6" pageNumber="517">Incomplete</td>
<td id="69685DF8FFEFBEC2FBF9FC9D01DBD7D6" box="[1031,1167,863,923]" gridcol="4" gridrow="4" pageId="6" pageNumber="517">Absent</td>
<td id="69685DF8FFEFBEC2FB28FC9D000AD7D6" box="[1238,1374,863,923]" gridcol="5" gridrow="4" pageId="6" pageNumber="517">Stem</td>
</tr>
<tr id="2AB93484FFEFBEC2FF5DFC69000AD78E" box="[163,1374,939,963]" gridrow="5" pageId="6" pageNumber="517">
<th id="69685DF8FFEFBEC2FF5DFC690421D78E" box="[163,373,939,963]" gridcol="0" gridrow="5" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFF5DFC6E0421D78F" box="[163,373,939,963]" italics="true" pageId="6" pageNumber="517">
<taxonomicName id="53894D45FFEF413BFF5DFC6E043FD78F" authorityName="Belizin" authorityYear="1959" box="[163,363,939,963]" class="Insecta" family="Cynipidae" genus="Vetustia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="species" species="investigata">Vetustia investigata</taxonomicName>
*
</emphasis>
</th>
<td id="69685DF8FFEFBEC2FE6FFC690701D78E" box="[401,597,939,963]" gridcol="1" gridrow="5" pageId="6" pageNumber="517">12</td>
<td id="69685DF8FFEFBEC2FD8AFC6907ABD78E" box="[628,767,939,963]" gridcol="2" gridrow="5" pageId="6" pageNumber="517">F1 longer</td>
<td id="69685DF8FFEFBEC2FCE0FC6906F8D78E" box="[798,940,939,963]" gridcol="3" gridrow="5" pageId="6" pageNumber="517">Complete</td>
<td id="69685DF8FFEFBEC2FBF9FC6901DBD78E" box="[1031,1167,939,963]" gridcol="4" gridrow="5" pageId="6" pageNumber="517">Absent</td>
<td id="69685DF8FFEFBEC2FB28FC69000AD78E" box="[1238,1374,939,963]" gridcol="5" gridrow="5" pageId="6" pageNumber="517">Inflorescence</td>
</tr>
</table>
</paragraph>
<subSubSection id="DC93654DFFEF413BFF39FBD40176D1F0" pageId="6" pageNumber="517" type="diagnosis">
<paragraph id="943636C6FFEF413BFF39FBD404E6D0A8" blockId="6.[151,1437,1046,2009]" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFF39FBD40416D07D" bold="true" box="[199,322,1046,1072]" pageId="6" pageNumber="517">Diagnosis.</emphasis>
<taxonomicName id="53894D45FFEF413BFEB7FBD40490D07D" authorityName="Kieffer" authorityYear="1901" box="[329,452,1046,1072]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFEB7FBD40490D07D" box="[329,452,1046,1072]" italics="true" pageId="6" pageNumber="517">Panteliella</emphasis>
</taxonomicName>
, as presently circumscribed, can be easily distinguished from other
<taxonomicName id="53894D45FFEF413BFB39FBD4001ED07D" authorityName="Nastasi &amp; Deans" authorityYear="2024" box="[1223,1354,1046,1072]" class="Insecta" family="Cynipidae" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="tribe" tribe="Aulacideini">Aulacideini</taxonomicName>
genera by the following combination of characters: Fore wing with marginal cell open; mesopleuron sculpture almost entirely striate (
<figureCitation id="0CB22A43FFEF413BFEAEFB9C04F2D034" box="[336,422,1118,1145]" captionStart="FIGURES 9–14" captionStartId="10.[151,264,1667,1691]" captionTargetBox="[180,1407,181,1644]" captionTargetId="figure-22@10.[180,1407,181,1644]" captionTargetPageId="10" captionText="FIGURES 9–14. Panteliella rugosa Nastasi sp. nov. All images show holotype female INHS Insect Collection 288137. (9) Lateral habitus. (10) Head in anterior view; dotted line indicates ventral margin of clypeus. (11) Mesoscutum in dorsal view. (12) Scutellum in dorsal view. (13) Mesopleuron in lateral view. (14) Fore wing." figureDoi="http://doi.org/10.5281/zenodo.14240300" httpUri="https://zenodo.org/record/14240300/files/figure.png" pageId="6" pageNumber="517">Fig. 13</figureCitation>
); second metasomal tergite without conspicuous patch of setae (
<figureCitation id="0CB22A43FFEF413BFB65FB9C01B7D034" box="[1179,1251,1118,1145]" captionStart="FIGURES 9–14" captionStartId="10.[151,264,1667,1691]" captionTargetBox="[180,1407,181,1644]" captionTargetId="figure-22@10.[180,1407,181,1644]" captionTargetPageId="10" captionText="FIGURES 9–14. Panteliella rugosa Nastasi sp. nov. All images show holotype female INHS Insect Collection 288137. (9) Lateral habitus. (10) Head in anterior view; dotted line indicates ventral margin of clypeus. (11) Mesoscutum in dorsal view. (12) Scutellum in dorsal view. (13) Mesopleuron in lateral view. (14) Fore wing." figureDoi="http://doi.org/10.5281/zenodo.14240300" httpUri="https://zenodo.org/record/14240300/files/figure.png" pageId="6" pageNumber="517">Fig. 9</figureCitation>
); clypeus large, broadly projecting over base of mandibles and with clypeo-pleurostomal lines strongly divergent ventrally (
<figureCitation id="0CB22A43FFEF413BFAC2FB4000DAD0D0" box="[1340,1422,1154,1181]" captionStart="FIGURES 9–14" captionStartId="10.[151,264,1667,1691]" captionTargetBox="[180,1407,181,1644]" captionTargetId="figure-22@10.[180,1407,181,1644]" captionTargetPageId="10" captionText="FIGURES 9–14. Panteliella rugosa Nastasi sp. nov. All images show holotype female INHS Insect Collection 288137. (9) Lateral habitus. (10) Head in anterior view; dotted line indicates ventral margin of clypeus. (11) Mesoscutum in dorsal view. (12) Scutellum in dorsal view. (13) Mesopleuron in lateral view. (14) Fore wing." figureDoi="http://doi.org/10.5281/zenodo.14240300" httpUri="https://zenodo.org/record/14240300/files/figure.png" pageId="6" pageNumber="517">Fig. 10</figureCitation>
); notauli incomplete, poorly impressed, more or less indistinct (
<figureCitation id="0CB22A43FFEF413BFCB7FB6406C3D08C" box="[841,919,1190,1217]" captionStart="FIGURES 7–8" captionStartId="4.[151,264,1951,1975]" captionTargetBox="[213,1375,181,1927]" captionTargetId="figure-22@4.[213,1375,181,1927]" captionTargetPageId="4" captionText="FIGURES 7–8. Panteliella fedtschenkoi (Rübsaamen, 1896), female USNMENT 01823223. (7) Lateral habitus. (8) Dorsal mesosoma." figureDoi="http://doi.org/10.5281/zenodo.14240294" httpUri="https://zenodo.org/record/14240294/files/figure.png" pageId="6" pageNumber="517">Figs. 8</figureCitation>
;
<figureCitation id="0CB22A43FFEF413BFC5AFB650695D08C" box="[932,961,1191,1217]" captionStart="FIGURES 9–14" captionStartId="10.[151,264,1667,1691]" captionTargetBox="[180,1407,181,1644]" captionTargetId="figure-22@10.[180,1407,181,1644]" captionTargetPageId="10" captionText="FIGURES 9–14. Panteliella rugosa Nastasi sp. nov. All images show holotype female INHS Insect Collection 288137. (9) Lateral habitus. (10) Head in anterior view; dotted line indicates ventral margin of clypeus. (11) Mesoscutum in dorsal view. (12) Scutellum in dorsal view. (13) Mesopleuron in lateral view. (14) Fore wing." figureDoi="http://doi.org/10.5281/zenodo.14240300" httpUri="https://zenodo.org/record/14240300/files/figure.png" pageId="6" pageNumber="517">11</figureCitation>
), apparent only as short longitudinal rugae in posterior mesoscutum.
</paragraph>
<paragraph id="943636C6FFEF413BFF39FB2C0176D1F0" blockId="6.[151,1437,1046,2009]" pageId="6" pageNumber="517">
<taxonomicName id="53894D45FFEF413BFF39FB2C0416D145" authorityName="Kieffer" authorityYear="1901" box="[199,322,1262,1288]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFF39FB2C0416D145" box="[199,322,1262,1288]" italics="true" pageId="6" pageNumber="517">Panteliella</emphasis>
</taxonomicName>
is closest morphologically to
<taxonomicName id="53894D45FFEF413BFD77FB2C06F2D145" authority="Forster, 1869" authorityName="Forster" authorityYear="1869" box="[649,934,1262,1289]" class="Insecta" family="Cynipidae" genus="Liposthenes" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFD77FB2C065BD145" box="[649,783,1262,1288]" italics="true" pageId="6" pageNumber="517">Liposthenes</emphasis>
<bibRefCitation id="F0184B37FFEF413BFCEAFB2D06F2D145" author="Forster, A." box="[788,934,1262,1289]" pageId="6" pageNumber="517" pagination="325 - 370" refId="ref9503" refString="Forster, A. (1869) Ueber die Gallwespen. Verhandlungen der k. k. Zoologisch-botanischen Gesellschaft in Wien, 19, 325 - 370." type="journal article" year="1869">Förster, 1869</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="F0184B37FFEF413BFC4CFB2C01CED144" author="Nieves-Aldrey, J. L." box="[946,1178,1262,1289]" pageId="6" pageNumber="517" pagination="393 - 413" refId="ref10170" refString="Nieves-Aldrey, J. L. (2022) Description of Fumariphilus Nieves-Aldrey, gen. nov., a new genus of herb gall wasps, with a key to genera of the tribe Aulacideini (Hymenoptera: Cynipidae). Zootaxa, 5155 (3), 393 - 413. https: // doi. org / 10.11646 / zootaxa. 5155.3.5" type="journal article" year="2022">Nieves-Aldrey 2022</bibRefCitation>
), but is easily separated by the notauli (complete and distinctly impressed in
<taxonomicName id="53894D45FFEF413BFD08FAD00628D161" authorityName="Forster" authorityYear="1869" box="[758,892,1298,1324]" class="Insecta" family="Cynipidae" genus="Liposthenes" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFD08FAD00628D161" box="[758,892,1298,1324]" italics="true" pageId="6" pageNumber="517">Liposthenes</emphasis>
</taxonomicName>
).
<taxonomicName id="53894D45FFEF413BFC68FAD00145D161" authorityName="Kieffer" authorityYear="1901" box="[918,1041,1298,1324]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFC68FAD00145D161" box="[918,1041,1298,1324]" italics="true" pageId="6" pageNumber="517">Panteliella</emphasis>
</taxonomicName>
is also morphologically similar to some
<taxonomicName id="53894D45FFEF413BFF24FAF40495D11D" box="[218,449,1334,1360]" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFF24FAF4043CD11D" box="[218,360,1334,1360]" italics="true" pageId="6" pageNumber="517">Antistrophus</emphasis>
species
</taxonomicName>
based on the state of the notauli, open marginal cell, and lack of a distinct setose patch on the second metasomal tergite (
<bibRefCitation id="F0184B37FFEF413BFDF2FA9807B8D138" author="Nastasi, L. F. &amp; Buffington, M. L. &amp; Davis, C. K. &amp; Deans, A. R." box="[524,748,1370,1397]" pageId="6" pageNumber="517" pagination="177 - 207" refId="ref9809" refString="Nastasi, L. F., Buffington, M. L., Davis, C. K. &amp; Deans, A. R. (2024 a) Key to the North American tribes and genera of herb, rose, bramble, and inquiline gall wasps (Hymenoptera, Cynipoidea, Cynipidae sensu lato). ZooKeys, 1196, 177 - 207. https: // doi. org / 10.3897 / zookeys. 1196.118460" type="journal article" year="2024">
Nastasi
<emphasis id="A6FDEAD4FFEF413BFD99FA9907CBD139" box="[615,671,1370,1396]" italics="true" pageId="6" pageNumber="517">et al.</emphasis>
2024a
</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEF413BFD06FA980616D139" author="Nastasi, L. F. &amp; Tooker, J. F. &amp; Davis, C. K. &amp; Smith, C. N. &amp; Frey, T. S. &amp; Hatfield, M. J. &amp; Presnall, T. M. &amp; Hines, H. M. &amp; Deans, A. R." box="[760,834,1370,1396]" pageId="6" pageNumber="517" pagination="399 - 439" refId="ref10028" refString="Nastasi, L. F., Tooker, J. F., Davis, C. K., Smith, C. N., Frey, T. S., Hatfield, M. J., Presnall, T. M., Hines, H. M. &amp; Deans, A. R. (2024 b) Cryptic or underworked? Taxonomic revision of the Antistrophus rufus species complex (Cynipoidea, Aulacideini). Journal of Hymenoptera Research, 97, 399 - 439. https: // doi. org / 10.3897 / jhr. 97.121918" type="journal article" year="2024">2024b</bibRefCitation>
). However, the two genera are easily differentiated by the mesopleuron sculpture (always with some amount of perceptible reticulate sculpture in
<taxonomicName id="53894D45FFEF413BFB72FABC004ED1D5" authorityName="Walsh" authorityYear="1869" box="[1164,1306,1406,1432]" class="Insecta" family="Cynipidae" genus="Antistrophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFB72FABC004ED1D5" box="[1164,1306,1406,1432]" italics="true" pageId="6" pageNumber="517">Antistrophus</emphasis>
</taxonomicName>
) and by the clypeus, which is smaller and does not strongly project ventrally in
<taxonomicName id="53894D45FFEF413BFC73FA60014FD1F1" authorityName="Walsh" authorityYear="1869" box="[909,1051,1442,1468]" class="Insecta" family="Cynipidae" genus="Antistrophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFC73FA60014FD1F1" box="[909,1051,1442,1468]" italics="true" pageId="6" pageNumber="517">Antistrophus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DC93654DFFEF413AFF39FA040415D4B4" lastPageId="7" lastPageNumber="518" pageId="6" pageNumber="517" type="description">
<paragraph id="943636C6FFEF413AFF39FA040415D4B4" blockId="6.[151,1437,1046,2009]" lastBlockId="7.[151,1436,151,501]" lastPageId="7" lastPageNumber="518" pageId="6" pageNumber="517">
<emphasis id="A6FDEAD4FFEF413BFF39FA040421D1AD" bold="true" box="[199,373,1478,1504]" pageId="6" pageNumber="517">Redescription.</emphasis>
Head mostly orangish brown or entirely black, mesosoma black, and metasoma dark brown to black. Head subtrapezoidal in anterior view, conspicuously wider than tall. Facial radiating striae always poorly impressed and incomplete, only reaching slightly beyond clypeus. POL much longer than OOL. Malar space much shorter than height of compound eye. Clypeus large, broadly projecting over base of mandibles, and with clypeo-pleurostomal lines strongly divergent ventrally. Female antennae with 13 or 14 antennomeres; males (not examined here) reportedly with 14 or 15. Female F1 equal to F2 (apparently also in males). Mesosoma in lateral view convex to conspicuously angled posteriorly. Lateral pronotum without rugose sculpture (strong rugose sculpture often present in other
<taxonomicName id="53894D45FFEF413BFEB3F9000484D291" authorityName="Nastasi &amp; Deans" authorityYear="2024" box="[333,464,1730,1756]" class="Insecta" family="Cynipidae" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="tribe" tribe="Aulacideini">Aulacideini</taxonomicName>
such as some
<taxonomicName id="53894D45FFEF413BFD8BF9000657D291" authorityName="Walsh" authorityYear="1869" box="[629,771,1730,1756]" class="Insecta" family="Cynipidae" genus="Antistrophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFD8BF9000657D291" box="[629,771,1730,1756]" italics="true" pageId="6" pageNumber="517">Antistrophus</emphasis>
</taxonomicName>
or
<taxonomicName id="53894D45FFEF413BFCD4F90006E4D291" authorityName="Forster" authorityYear="1869" box="[810,944,1730,1756]" class="Insecta" family="Cynipidae" genus="Liposthenes" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEF413BFCD4F90006E4D291" box="[810,944,1730,1756]" italics="true" pageId="6" pageNumber="517">Liposthenes</emphasis>
</taxonomicName>
). Pronotal plate incomplete but with lateral sutures terminating only shortly before reaching mesoscutum. Mesopleuron almost entirely striate, with a very small ventral area of reticulate sculpture. Mesoscutum coriaceous to reticulate, occasionally with some degree of perceptible rugose sculpture. Notauli incomplete, narrow, and poorly impressed, most apparent in posterior third of mesoscutum. Median mesoscutal impression not apparent. Mesoscutellar foveae ovate to subquadrate, about one quarter as long as mesoscutellar disc and with posterior margins poorly defined. Propodeum with or without a conspicuous pair of lateral carinae (lateral carinae absent in specimens examined by
<bibRefCitation id="F0184B37FFEF413BFBC4F8580066D3F8" author="Nieves-Aldrey, J. L." box="[1082,1330,1946,1973]" pageId="6" pageNumber="517" pagination="393 - 413" refId="ref10170" refString="Nieves-Aldrey, J. L. (2022) Description of Fumariphilus Nieves-Aldrey, gen. nov., a new genus of herb gall wasps, with a key to genera of the tribe Aulacideini (Hymenoptera: Cynipidae). Zootaxa, 5155 (3), 393 - 413. https: // doi. org / 10.11646 / zootaxa. 5155.3.5" type="journal article" year="2022">Nieves-Aldrey [2022]</bibRefCitation>
, in which only a strong median carina is apparent). Fore wing with marginal cell open, with vein R1 indistinctly reaching fore wing margin, and with conspicuous marginal setae. Areolet absent. Metasoma with conspicuous micropunctation at least on third and following metasomal tergites. Second metasomal tergite without setose patch, at most with a few scattered setae.
</paragraph>
</subSubSection>
<subSubSection id="DC93654DFFEE413AFF39FEC00172D5C4" pageId="7" pageNumber="518" type="biology_ecology">
<paragraph id="943636C6FFEE413AFF39FEC00172D5C4" blockId="7.[151,1436,151,501]" pageId="7" pageNumber="518">
<emphasis id="A6FDEAD4FFEE413AFF39FEC00473D551" bold="true" box="[199,295,258,284]" pageId="7" pageNumber="518">Biology.</emphasis>
<taxonomicName id="53894D45FFEE413AFED3FEC1076ED550" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[301,570,259,285]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEE413AFED3FEC1076ED550" box="[301,570,259,285]" italics="true" pageId="7" pageNumber="518">Panteliella fedtschenkoi</emphasis>
</taxonomicName>
induces galls on
<taxonomicName id="53894D45FFEE413AFD04FEC106B1D550" box="[762,997,259,285]" class="Magnoliopsida" family="Lamiaceae" genus="Phlomoides" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="518" phylum="Tracheophyta" rank="species" species="tuberosa">
<emphasis id="A6FDEAD4FFEE413AFD04FEC106B1D550" box="[762,997,259,285]" italics="true" pageId="7" pageNumber="518">Phlomoides tuberosa</emphasis>
</taxonomicName>
(
<taxonomicName id="53894D45FFEE413AFC0DFEC10127D550" box="[1011,1139,259,285]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="518" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
). The genus was originally described for a species inducing monothalamous galls on the leaves, but genera currently synonymized with
<taxonomicName id="53894D45FFEE413AFF69FE890446D528" authorityName="Kieffer" authorityYear="1901" box="[151,274,331,357]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEE413AFF69FE890446D528" box="[151,274,331,357]" italics="true" pageId="7" pageNumber="518">Panteliella</emphasis>
</taxonomicName>
were described from galls in the inflorescence and cryptic galls in the stems (
<bibRefCitation id="F0184B37FFEE413AFB78FE89004FD528" author="Belizin, V. I." box="[1158,1307,330,357]" pageId="7" pageNumber="518" pagination="662 - 674" refId="ref9247" refString="Belizin, V. I. (1959) Gall wasps of the tribe Aylaxini (Hymenoptera, Cynipoidea) new for the fauna of the U. S. S. R. Entomologicheskoe Obozrenie, 38, 662 - 674." type="book chapter" year="1959">Belizin 1959</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEE413AFAD9FE89047FD5C4" author="Ionescu, M. A. &amp; Roman, N." pageId="7" pageNumber="518" pagination="221 - 225" refId="ref9557" refString="Ionescu, M. A. &amp; Roman, N. (1960) Un genre nouveau de Cynipides gallicoles Endocaulonia bicolor n. g., n. sp. (Hymenoptera, Cynipoidea). Revue de Biologie Academiei Republicii Populare Romine Bucaresti, 5 (3), 221 - 225." type="journal article" year="1960">Ionescu &amp; Roman 1960</bibRefCitation>
). The host plants of
<taxonomicName id="53894D45FFEE413AFDF0FEAD07D2D5C4" authorityName="Vyrzhikovskaya" authorityYear="1962" box="[526,646,367,393]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="bianchii">
<emphasis id="A6FDEAD4FFEE413AFDF0FEAD07D2D5C4" box="[526,646,367,393]" italics="true" pageId="7" pageNumber="518">P. bianchii</emphasis>
</taxonomicName>
and
<taxonomicName id="53894D45FFEE413AFD42FEAD0671D5C5" authorityName="Nastasi &amp; Deans" authorityYear="2024" box="[700,805,367,392]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="rugosa" status="sp. nov.">
<emphasis id="A6FDEAD4FFEE413AFD42FEAD0671D5C5" box="[700,805,367,392]" italics="true" pageId="7" pageNumber="518">P. rugosa</emphasis>
</taxonomicName>
<emphasis id="A6FDEAD4FFEE413AFCD2FEAD06D2D5C4" bold="true" box="[812,902,367,393]" pageId="7" pageNumber="518">
<taxonomicNameLabel id="BDCE57AFFFEE413AFCD2FEAD06D2D5C4" box="[812,902,367,393]" pageId="7" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are unknown.
</paragraph>
</subSubSection>
<subSubSection id="DC93654DFFEE413AFF39FE500613D5B8" pageId="7" pageNumber="518" type="distribution">
<paragraph id="943636C6FFEE413AFF39FE500613D5B8" blockId="7.[151,1436,151,501]" pageId="7" pageNumber="518">
<emphasis id="A6FDEAD4FFEE413AFF39FE500435D5E1" bold="true" box="[199,353,402,428]" pageId="7" pageNumber="518">Distribution.</emphasis>
<taxonomicName id="53894D45FFEE413AFE9BFE5104B4D5E0" authorityName="Kieffer" authorityYear="1901" box="[357,480,403,429]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFEE413AFE9BFE5104B4D5E0" box="[357,480,403,429]" italics="true" pageId="7" pageNumber="518">Panteliella</emphasis>
</taxonomicName>
has been recorded throughout Europe and parts of continental Asia including
<collectingCountry id="EC9E7656FFEE413AFAD1FE5100C8D5E0" box="[1327,1436,403,429]" name="Mongolia" pageId="7" pageNumber="518">Mongolia</collectingCountry>
(
<bibRefCitation id="F0184B37FFEE413AFF61FE7504D3D59C" author="Nieves-Aldrey, J. L." box="[159,391,439,465]" pageId="7" pageNumber="518" pagination="175 - 206" refId="ref10135" refString="Nieves-Aldrey, J. L. (1994) Revision of West-European Genera of the Tribe Aylacini Ashmead (Hymenoptera, Cynipidae). Journal of Hymenoptera Research, 3, 175 - 206." type="journal article" year="1994">Nieves-Aldrey 1994</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEE413AFE6CFE750771D59C" author="Belizin, V. I." box="[402,549,438,465]" pageId="7" pageNumber="518" pagination="662 - 674" refId="ref9247" refString="Belizin, V. I. (1959) Gall wasps of the tribe Aylaxini (Hymenoptera, Cynipoidea) new for the fauna of the U. S. S. R. Entomologicheskoe Obozrenie, 38, 662 - 674." type="book chapter" year="1959">Belizin 1959</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEE413AFDCEFE750798D59C" author="Quinlan, J." box="[560,716,439,465]" pageId="7" pageNumber="518" pagination="275 - 286" refId="ref10265" refString="Quinlan, J. (1968) Cynipinae (Hymenoptera) occurring on Phlomis L. Transactions of the Royal Entomological Society of London, 120, 275 - 286. https: // doi. org / 10.1111 / j. 1365 - 2311.1968. tb 00339. x" type="journal article" year="1968">Quinlan 1968</bibRefCitation>
,
<bibRefCitation id="F0184B37FFEE413AFD29FE75063ED59C" author="Melika, G." box="[727,874,439,465]" pageId="7" pageNumber="518" refId="ref9772" refString="Melika, G. (2006) Gall wasps of Ukraine. Vol. 1. Vestnik zoologii, The Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev, 300 pp." type="book" year="2006">Melika 2006</bibRefCitation>
). We document the genus from
<collectingCountry id="EC9E7656FFEE413AFB35FE750018D59C" box="[1227,1356,439,465]" name="Kyrgyzstan" pageId="7" pageNumber="518">Kyrgyzstan</collectingCountry>
for the first time (see treatment of
<taxonomicName id="53894D45FFEE413AFE39FE1907DDD5B8" authority="Nastasi" authorityName="Nastasi &amp; Deans" authorityYear="2024" box="[455,649,475,501]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="rugosa" status="sp. nov.">
<emphasis id="A6FDEAD4FFEE413AFE39FE190765D5B9" box="[455,561,475,500]" italics="true" pageId="7" pageNumber="518">P. rugosa</emphasis>
Nastasi
</taxonomicName>
<emphasis id="A6FDEAD4FFEE413AFD6FFE1907BED5B8" bold="true" box="[657,746,475,501]" pageId="7" pageNumber="518">
<taxonomicNameLabel id="BDCE57AFFFEE413AFD6FFE1907BED5B8" box="[657,746,475,501]" pageId="7" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
below).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="3CB89D98D05381A950B9AA1B2A8A1A22">New herb gall wasps (Hymenoptera: Cynipidae: Aulacideini) from Kyrgyzstan, with description of a new genus and a review of the genus Panteliella Kieffer, 1901</mods:title>
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<mods:namePart id="A82E5D04466FD54AFA6E48A1E05CAF91">Nastasi, Louis F.</mods:namePart>
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<paragraph id="943636C6FFEE413AFF69FBA507E6D0CF" blockId="7.[151,690,1127,1190]" box="[151,690,1127,1154]" pageId="7" pageNumber="518">
<heading id="CF7E81AAFFEE413AFF69FBA507E6D0CF" bold="true" box="[151,690,1127,1154]" fontSize="11" level="1" pageId="7" pageNumber="518" reason="1">
<taxonomicName id="53894D45FFEE413AFF69FBA507E6D0CF" authority="(Rubsaamen, 1896)" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[151,690,1127,1154]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEE413AFF69FBA507E6D0CF" bold="true" box="[151,690,1127,1154]" pageId="7" pageNumber="518">
<emphasis id="A6FDEAD4FFEE413AFF69FBA504EED0CF" bold="true" box="[151,442,1127,1154]" italics="true" pageId="7" pageNumber="518">Panteliella fedtschenkoi</emphasis>
(
<bibRefCitation id="F0184B37FFEE413AFE37FBAA07FED0CF" author="Rubsaamen, E. H." box="[457,682,1128,1154]" pageId="7" pageNumber="518" pagination="396 - 481" refId="ref10418" refString="Rubsaamen, E. H. (1896) Uber russische Zoocecidien und deren Erzeuger. Bulletin de la Societe imperiale des naturalistes de Moscou, 3, 396 - 481." type="journal article" year="1896">Rübsaamen, 1896</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="DC93654DFFEE413AFF69FB4E0446D0EB" box="[151,274,1164,1190]" pageId="7" pageNumber="518" type="description">
<paragraph id="943636C6FFEE413AFF69FB4E0446D0EB" blockId="7.[151,690,1127,1190]" box="[151,274,1164,1190]" pageId="7" pageNumber="518">
(
<figureCitation id="0CB22A43FFEE413AFF61FB4E045ED0EB" box="[159,266,1164,1190]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–8" captionStartId-0="2.[151,264,1734,1758]" captionStartId-1="4.[151,264,1951,1975]" captionTargetBox-0="[152,1436,181,1710]" captionTargetBox-1="[213,1375,181,1927]" captionTargetId-0="figure-22@2.[152,1436,181,1710]" captionTargetId-1="figure-22@4.[213,1375,181,1927]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 1–6. Soparia oshensis Nastasi gen. and sp. nov. All images show holotype female INHS Insect Collection 288148. (1) Lateral habitus. (2) Head in anterior view; dotted line indicates ventral margin of clypeus. (3) Mesoscutum in dorsal view. (4) Scutellum in dorsal view. (5) Mesopleuron in lateral view. (6) Wings." captionText-1="FIGURES 7–8. Panteliella fedtschenkoi (Rübsaamen, 1896), female USNMENT 01823223. (7) Lateral habitus. (8) Dorsal mesosoma." figureDoi-0="http://doi.org/10.5281/zenodo.14240292" figureDoi-1="http://doi.org/10.5281/zenodo.14240294" httpUri-0="https://zenodo.org/record/14240292/files/figure.png" httpUri-1="https://zenodo.org/record/14240294/files/figure.png" pageId="7" pageNumber="518">Figs. 6–7</figureCitation>
)
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<subSubSection id="DC93654DFFEE413AFF69FB1204B8D1CF" pageId="7" pageNumber="518" type="reference_group">
<paragraph id="943636C6FFEE413AFF69FB120747D0A5" blockId="7.[151,767,1232,1410]" box="[151,531,1232,1256]" pageId="7" pageNumber="518">
<taxonomicName id="53894D45FFEE413AFF69FB120747D0A5" authority="Rubsaamen, 1896" authorityName="Rubsaamen" authorityYear="1896" box="[151,531,1232,1256]" class="Insecta" family="Cynipidae" genus="Aulax" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEE413AFF69FB12040CD0A5" box="[151,344,1232,1256]" italics="true" pageId="7" pageNumber="518">Aulax fedtschenkoi</emphasis>
<bibRefCitation id="F0184B37FFEE413AFEA0FB120747D0A5" author="Rubsaamen, E. H." box="[350,531,1232,1256]" pageId="7" pageNumber="518" pagination="396 - 481" refId="ref10418" refString="Rubsaamen, E. H. (1896) Uber russische Zoocecidien und deren Erzeuger. Bulletin de la Societe imperiale des naturalistes de Moscou, 3, 396 - 481." type="journal article" year="1896">Rübsaamen, 1896</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="943636C6FFEE413AFF69FB2D07C3D14A" blockId="7.[151,767,1232,1410]" box="[151,663,1263,1287]" pageId="7" pageNumber="518">
<taxonomicName id="53894D45FFEE413AFF69FB2D07C3D14A" authority="(Rubsaamen) Kieffer, 1901 a" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" box="[151,663,1263,1287]" class="Insecta" family="Cynipidae" genus="Pantelia" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEE413AFF69FB2D0426D14A" box="[151,370,1263,1287]" italics="true" pageId="7" pageNumber="518">Pantelia fedtschenkoi</emphasis>
(Rübsaamen)
<bibRefCitation id="F0184B37FFEE413AFDF8FB2D07C3D14A" author="Kieffer, J. J." box="[518,663,1263,1287]" pageId="7" pageNumber="518" refId="ref9612" refString="Kieffer, J. J. (1901 a) Monographie des Cynipides d'Europe et d'Algerie: Les Cynipides. Froment-Dubosclard, Paris, 687 pp." type="book" year="1901">Kieffer, 1901a</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="943636C6FFEE413AFF69FACC07E7D16B" blockId="7.[151,767,1232,1410]" box="[151,691,1294,1318]" pageId="7" pageNumber="518">
<taxonomicName id="53894D45FFEE413AFF69FACC07E7D16B" authority="(Rubsaamen) Kieffer, 1901 b" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[151,691,1294,1318]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEE413AFF69FACC04D8D16B" box="[151,396,1294,1318]" italics="true" pageId="7" pageNumber="518">Panteliella fedtschenkoi</emphasis>
(Rübsaamen)
<bibRefCitation id="F0184B37FFEE413AFDE1FACC07E7D16B" author="Kieffer, J. J." box="[543,691,1294,1318]" pageId="7" pageNumber="518" pagination="233 - 579" refId="ref9639" refString="Kieffer, J. J. (1901 b) Synopsis de Zoocecidies d'Europe. Annalles de la Societe Entomologique de France, 1901, 233 - 579. https: // doi. org / 10.1080 / 21686351.1901.12279879" type="journal article" year="1901">Kieffer, 1901b</bibRefCitation>
</taxonomicName>
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<paragraph id="943636C6FFEE413AFF69FAEE0727D109" blockId="7.[151,767,1232,1410]" box="[151,627,1324,1349]" pageId="7" pageNumber="518">
<taxonomicName id="53894D45FFEE413AFF69FAEE0727D109" authority="Ionescu &amp; Roman, 1960" authorityName="Ionescu &amp; Roman" authorityYear="1960" box="[151,627,1324,1349]" class="Insecta" family="Cynipidae" genus="Endocaulonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A6FDEAD4FFEE413AFF69FAEE0421D109" box="[151,373,1324,1348]" italics="true" pageId="7" pageNumber="518">Endocaulonia bicolor</emphasis>
<bibRefCitation id="F0184B37FFEE413AFE85FAEF0727D109" author="Ionescu, M. A. &amp; Roman, N." box="[379,627,1324,1349]" pageId="7" pageNumber="518" pagination="221 - 225" refId="ref9557" refString="Ionescu, M. A. &amp; Roman, N. (1960) Un genre nouveau de Cynipides gallicoles Endocaulonia bicolor n. g., n. sp. (Hymenoptera, Cynipoidea). Revue de Biologie Academiei Republicii Populare Romine Bucaresti, 5 (3), 221 - 225." type="journal article" year="1960">Ionescu &amp; Roman, 1960</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="943636C6FFEE413AFF69FA8907ABD12E" blockId="7.[151,767,1232,1410]" box="[151,767,1355,1379]" pageId="7" pageNumber="518">
<taxonomicName id="53894D45FFEE413AFF69FA8907ABD12E" authority="(Ionescu &amp; Roman) Nieves-Aldrey, 1994" authorityName="Nieves-Aldrey" authorityYear="1994" baseAuthorityName="Ionescu &amp; Roman" baseAuthorityYear="1960" box="[151,767,1355,1379]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="A6FDEAD4FFEE413AFF69FA890402D12E" box="[151,342,1355,1379]" italics="true" pageId="7" pageNumber="518">Panteliella bicolor</emphasis>
(Ionescu &amp; Roman)
<bibRefCitation id="F0184B37FFEE413AFDD2FA8907ABD12E" author="Nieves-Aldrey, J. L." box="[556,767,1355,1379]" pageId="7" pageNumber="518" pagination="175 - 206" refId="ref10135" refString="Nieves-Aldrey, J. L. (1994) Revision of West-European Genera of the Tribe Aylacini Ashmead (Hymenoptera, Cynipidae). Journal of Hymenoptera Research, 3, 175 - 206." type="journal article" year="1994">Nieves-Aldrey, 1994</bibRefCitation>
</taxonomicName>
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<paragraph id="943636C6FFEE413AFF69FAA804B8D1CF" blockId="7.[151,767,1232,1410]" box="[151,492,1386,1410]" pageId="7" pageNumber="518">
<taxonomicName id="53894D45FFEE413AFF69FAA804B8D1CF" authority="Belizin, 1959" authorityName="Belizin" authorityYear="1959" box="[151,492,1386,1410]" class="Insecta" family="Cynipidae" genus="Vetustia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="investigata">
<emphasis id="A6FDEAD4FFEE413AFF69FAA80409D1CC" box="[151,349,1386,1409]" italics="true" pageId="7" pageNumber="518">Vetustia investigata</emphasis>
<bibRefCitation id="F0184B37FFEE413AFE9DFAA804B8D1CF" author="Belizin, V. I." box="[355,492,1386,1410]" pageId="7" pageNumber="518" pagination="662 - 674" refId="ref9247" refString="Belizin, V. I. (1959) Gall wasps of the tribe Aylaxini (Hymenoptera, Cynipoidea) new for the fauna of the U. S. S. R. Entomologicheskoe Obozrenie, 38, 662 - 674." type="book chapter" year="1959">Belizin, 1959</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="DC93654DFFEE413AFF69FA7200DCD2EF" pageId="7" pageNumber="518" type="materials_examined">
<paragraph id="943636C6FFEE413AFF69FA7200DCD2EF" blockId="7.[151,1437,1456,2023]" pageId="7" pageNumber="518">
<emphasis id="A6FDEAD4FFEE413AFF69FA7204D5D187" bold="true" box="[151,385,1456,1482]" pageId="7" pageNumber="518">Material examined.</emphasis>
<materialsCitation id="24E13C9BFFEE413AFE77FA7306BDD233" collectionCode="USNM" county="Bijouk-Onlar" location="Oktyabrsk" municipality="Bijuk-Onlar" pageId="7" pageNumber="518" specimenCode="USNMENT 01823222" specimenCount="2" specimenCount-female="2" stateProvince="Crimea">
<specimenCount id="828FFD4FFFEE413AFE77FA730748D187" box="[393,540,1456,1483]" count="2" pageId="7" pageNumber="518" type="female">Two females</specimenCount>
,
<specimenCode id="C42F9EBDFFEE413AFDD6FA72067ED187" box="[552,810,1456,1482]" pageId="7" pageNumber="518">USNMENT 01823222</specimenCode>
, 01823223, both with same labels given verbatim here: “Bijouk-Onlar, Krim.” / “Cut out of type galls by Weld 1929” / “From Vienna Museum, 1929” / “
<taxonomicName id="53894D45FFEE413AFADFFA1604E6D25F" authority="(Rubsaam.)" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="518" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFEE413AFADFFA160477D25F" italics="true" pageId="7" pageNumber="518">Panteliella fedtschenkoi</emphasis>
(Rübsaam.)
</taxonomicName>
” /
<collectionCode id="F298AE03FFEE413AFE26FA3A077DD25F" box="[472,553,1528,1554]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="518" type="Museum">USNM</collectionCode>
barcode label. The labels report the locality only as “Bijouk-Onlar, Krim.”, which likely refers to Bijuk-Onlar, a former name of the settlement
<location id="9156601DFFEE413AFC7CF9DE06A3D27B" LSID="urn:lsid:plazi:treatment:1C2087D0FFEE4135FF69FBA50617D18C:9156601DFFEE413AFC7CF9DE06A3D27B" box="[898,1015,1564,1590]" county="Bijouk-Onlar" municipality="Bijuk-Onlar" name="Oktyabrsk" pageId="7" pageNumber="518" stateProvince="Crimea">Oktyabrsk</location>
in the Crimean Peninsula. Labels that were evidently added later correspond to
<bibRefCitation id="F0184B37FFEE413AFD9FF98207ABD217" author="Weld, L. H." box="[609,767,1600,1626]" pageId="7" pageNumber="518" pagination="137 - 144" refId="ref10592" refString="Weld, L. H. (1930) Notes on types (Hymenoptera: Cynipidae). Proceedings of the Entomological Society of Washington, 32 (8), 137 - 144." type="journal article" year="1930">Weld’s (1930)</bibRefCitation>
observations and indicate that the specimens were dissected from galls collected and preserved alongside the type series of this species.
</materialsCitation>
<location id="9156601DFFEE413AFC0EF9A60172D233" LSID="urn:lsid:plazi:treatment:1C2087D0FFEE4135FF69FBA50617D18C:9156601DFFEE413AFC0EF9A60172D233" box="[1008,1062,1636,1662]" county="Bijouk-Onlar" municipality="Bijuk-Onlar" name="Both" pageId="7" pageNumber="518" stateProvince="Crimea">Both</location>
specimens are in relatively good condition, with
<specimenCount id="828FFD4FFFEE413AFEB6F94B04B5D2EF" box="[328,481,1672,1699]" count="1" pageId="7" pageNumber="518" type="generic">one specimen</specimenCount>
missing only an antenna and the other mostly intact except for undeveloped wings.
</paragraph>
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<emphasis id="A6FDEAD4FFEE413AFF39F96E070DD28B" bold="true" box="[199,601,1708,1734]" pageId="7" pageNumber="518">
Redescription of female (
<figureCitation id="0CB22A43FFEE413AFDFEF96E071FD28B" box="[512,587,1708,1734]" captionStart="FIGURES 7–8" captionStartId="4.[151,264,1951,1975]" captionTargetBox="[213,1375,181,1927]" captionTargetId="figure-22@4.[213,1375,181,1927]" captionTargetPageId="4" captionText="FIGURES 7–8. Panteliella fedtschenkoi (Rübsaamen, 1896), female USNMENT 01823223. (7) Lateral habitus. (8) Dorsal mesosoma." figureDoi="http://doi.org/10.5281/zenodo.14240294" httpUri="https://zenodo.org/record/14240294/files/figure.png" pageId="7" pageNumber="518">Fig. 7</figureCitation>
).
</emphasis>
Body
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in length (n = 2). Coloration clearly faded since initial preservation, but coloration of head much lighter brownish than that of mesosoma and metasoma, especially on face. Antennae reddish yellow. Wing veins pale brown. Legs more or less orangish throughout.
</paragraph>
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<emphasis id="A6FDEAD4FFEE413AFF39F8DA0450D37F" box="[199,260,1816,1842]" italics="true" pageId="7" pageNumber="518">Head</emphasis>
. In dorsal view 1.9× as wide as long; in anterior view subtrapezoidal and 1.4× as wide as tall. Sculpture excluding weak facial radiating striae entirely coriaceous. POL:LOL:OOL:DLO in
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measuring 15:7:7:4. Toruli situated about mid-height of eyes. Malar space 0.6× as long as compound eye in anterior view. Facial radiating striae incomplete, very poorly impressed, apparent as indistinct rugae emanating for a short distance from the clypeus. Clypeus large, broadly projecting ventrally over base of mandibles, and with clypeo-pleurostomal lines strongly divergent ventrally. Genae only slightly expanded posterior to eyes.
</paragraph>
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<emphasis id="A6FDEAD4FFE14135FF39FF550465D4FC" box="[199,305,151,177]" italics="true" pageId="8" pageNumber="519">Antennae</emphasis>
(
<figureCitation id="0CB22A43FFE14135FEBDFF5504DED4FC" box="[323,394,151,177]" captionStart="FIGURES 7–8" captionStartId="4.[151,264,1951,1975]" captionTargetBox="[213,1375,181,1927]" captionTargetId="figure-22@4.[213,1375,181,1927]" captionTargetPageId="4" captionText="FIGURES 7–8. Panteliella fedtschenkoi (Rübsaamen, 1896), female USNMENT 01823223. (7) Lateral habitus. (8) Dorsal mesosoma." figureDoi="http://doi.org/10.5281/zenodo.14240294" httpUri="https://zenodo.org/record/14240294/files/figure.png" pageId="8" pageNumber="519">Fig. 7</figureCitation>
). With 14 antennomeres, about two thirds as long as body length. Conspicuous placodeal sensilla present on F2 and following flagellomeres. Pedicel about as long as F1. F1 2.0× as long as wide. F2 1.8× as long as wide; about 1.0× as long as F1. Apical flagellomere 2.1× as long as wide.
</paragraph>
<paragraph id="943636C6FFE14135FF39FEC10157D6BC" blockId="8.[151,1437,151,1473]" pageId="8" pageNumber="519">
<emphasis id="A6FDEAD4FFE14135FF39FEC1046BD551" box="[199,319,259,284]" italics="true" pageId="8" pageNumber="519">Mesosoma</emphasis>
(
<figureCitation id="0CB22A43FFE14135FEAEFEC104EAD550" box="[336,446,259,285]" captionStart="FIGURES 7–8" captionStartId="4.[151,264,1951,1975]" captionTargetBox="[213,1375,181,1927]" captionTargetId="figure-22@4.[213,1375,181,1927]" captionTargetPageId="4" captionText="FIGURES 7–8. Panteliella fedtschenkoi (Rübsaamen, 1896), female USNMENT 01823223. (7) Lateral habitus. (8) Dorsal mesosoma." figureDoi="http://doi.org/10.5281/zenodo.14240294" httpUri="https://zenodo.org/record/14240294/files/figure.png" pageId="8" pageNumber="519">Figs. 7–8</figureCitation>
). Strongly convex in lateral view. Pronotum coriaceous, with medial height about 0.3× height of lateral margin. Submedial pronotal pits transverse, deep, and clearly separated. Pronotal plate incomplete, with lateral sutures terminating only shortly before reaching mesoscutum. Mesopleuron almost entirely striate, with a very small ventral area of reticulate sculpture. Mesopleural triangle conspicuously impressed, with only a few scattered setae anteriorly. Mesoscutum mostly weakly coriaceous, with a few conspicuous longitudinal rugae apparent for a short distance anterior to scutellar margin, with only a few sparse, scattered setae. Notauli incomplete, inconspicuously impressed; apparent only as short longitudinal rugae in posterior third of mesoscutum. Median mesoscutal impression not apparent but mesoscutum very slightly impressed posteromedially. Mesoscutellar disc mostly reticulate, with conspicuous longitudinal rugae anteriorly and rugose-reticulate toward outer margins. Mesoscutellar foveae ovate; relatively short, about one quarter as long as mesoscutellar disc; posterior margins poorly defined but strongly divergent; without setae; rugose within margins; separated by a broad subtriangular area. Metapleural sulcus meeting mesopleuron in dorsal third. Metapleuron coriaceous to rugose, more or less moderately setose throughout. Propodeal carinae conspicuous, apparent as paired subparallel lateral carinae, with strong sculpture between them producing a faint indication of a medial carina.
</paragraph>
<paragraph id="943636C6FFE14135FF39FD390042D774" blockId="8.[151,1437,151,1473]" pageId="8" pageNumber="519">
<emphasis id="A6FDEAD4FFE14135FF39FD390468D759" box="[199,316,763,788]" italics="true" pageId="8" pageNumber="519">Fore wing</emphasis>
(
<figureCitation id="0CB22A43FFE14135FEB1FD3904C1D758" box="[335,405,763,789]" captionStart="FIGURES 7–8" captionStartId="4.[151,264,1951,1975]" captionTargetBox="[213,1375,181,1927]" captionTargetId="figure-22@4.[213,1375,181,1927]" captionTargetPageId="4" captionText="FIGURES 7–8. Panteliella fedtschenkoi (Rübsaamen, 1896), female USNMENT 01823223. (7) Lateral habitus. (8) Dorsal mesosoma." figureDoi="http://doi.org/10.5281/zenodo.14240294" httpUri="https://zenodo.org/record/14240294/files/figure.png" pageId="8" pageNumber="519">Fig. 7</figureCitation>
). Hyaline and setose throughout. Marginal cell open, 2.9× as long as wide, with vein R1 indistinctly reaching fore wing margin. Areolet absent. Distal margin with conspicuous marginal setae.
</paragraph>
<paragraph id="943636C6FFE14135FF39FC81044ED7E8" blockId="8.[151,1437,151,1473]" pageId="8" pageNumber="519">
<emphasis id="A6FDEAD4FFE14135FF39FC810417D711" box="[199,323,835,860]" italics="true" pageId="8" pageNumber="519">Metasoma.</emphasis>
Metasoma slightly shorter than combined length of head and mesosoma. Second metasomal tergite without conspicuous anteromedial setose patch. Third and following metasomal tergites with sparse micropunctation throughout.
</paragraph>
<paragraph id="943636C6FFE14135FF39FC6C0787D014" blockId="8.[151,1437,151,1473]" pageId="8" pageNumber="519">
<emphasis id="A6FDEAD4FFE14135FF39FC6C049ED784" bold="true" box="[199,458,942,969]" pageId="8" pageNumber="519">Redescription of gall.</emphasis>
On the adaxial leaves of
<taxonomicName id="53894D45FFE14135FD0AFC6D06B6D784" box="[756,994,943,969]" class="Magnoliopsida" family="Lamiaceae" genus="Phlomoides" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="519" phylum="Tracheophyta" rank="species" species="tuberosa">
<emphasis id="A6FDEAD4FFE14135FD0AFC6D06B6D784" box="[756,994,943,969]" italics="true" pageId="8" pageNumber="519">Phlomoides tuberosa</emphasis>
</taxonomicName>
. Conspicuous, solitary, semideciduous outgrowths of the leaf, 2.0–4.0 mm in diameter. Monothalamous. Shape spherical. Surface slightly rugose, with dense, moderately long pilosity. Color green when fresh to pink or brown when mature or dry. While
<taxonomicName id="53894D45FFE14135FADFFC350477D078" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="519" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFE14135FADFFC350477D078" italics="true" pageId="8" pageNumber="519">Panteliella fedtschenkoi</emphasis>
</taxonomicName>
has been mentioned to induce galls in the inflorescence or stems, these records almost certainly correspond to distinct species (see remarks below).
</paragraph>
</subSubSection>
<subSubSection id="DC93654DFFE14135FF39FBA10617D18C" pageId="8" pageNumber="519" type="discussion">
<paragraph id="943636C6FFE14135FF39FBA10617D18C" blockId="8.[151,1437,151,1473]" pageId="8" pageNumber="519">
<emphasis id="A6FDEAD4FFE14135FF39FBA1046FD030" bold="true" box="[199,315,1123,1149]" pageId="8" pageNumber="519">Remarks.</emphasis>
Several authors have treated
<taxonomicName id="53894D45FFE14135FD64FBA106F8D030" authorityName="Kieffer" authorityYear="1901" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[666,940,1123,1149]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="519" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="A6FDEAD4FFE14135FD64FBA106F8D030" box="[666,940,1123,1149]" italics="true" pageId="8" pageNumber="519">Panteliella fedtschenkoi</emphasis>
</taxonomicName>
since its description, often disagreeing on diagnostic characters (see remarks for
<taxonomicName id="53894D45FFE14135FDB1FB45079ED0EC" authorityName="Kieffer" authorityYear="1901" box="[591,714,1159,1185]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFE14135FDB1FB45079ED0EC" box="[591,714,1159,1185]" italics="true" pageId="8" pageNumber="519">Panteliella</emphasis>
</taxonomicName>
above).
<bibRefCitation id="F0184B37FFE14135FCD0FB4506EFD0EC" author="Weld, L. H." box="[814,955,1159,1185]" pageId="8" pageNumber="519" pagination="137 - 144" refId="ref10592" refString="Weld, L. H. (1930) Notes on types (Hymenoptera: Cynipidae). Proceedings of the Entomological Society of Washington, 32 (8), 137 - 144." type="journal article" year="1930">Weld (1930)</bibRefCitation>
redescribed
<taxonomicName id="53894D45FFE14135FBB0FB450009D0EC" baseAuthorityName="Rubsaamen" baseAuthorityYear="1896" box="[1102,1373,1159,1185]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="519" phylum="Arthropoda" rank="species" species="fedtschenoki">
<emphasis id="A6FDEAD4FFE14135FBB0FB450009D0EC" box="[1102,1373,1159,1185]" italics="true" pageId="8" pageNumber="519">Panteliella fedtschenoki</emphasis>
</taxonomicName>
from
<specimenCount id="828FFD4FFFE14135FF69FB690414D088" box="[151,320,1195,1221]" count="2" pageId="8" pageNumber="519" type="generic">two specimens</specimenCount>
dissected from galls of the
<typeStatus id="4B328864FFE14135FD7BFB6907E1D088" box="[645,693,1195,1221]" pageId="8" pageNumber="519">type</typeStatus>
series as the original description was insufficient. We examined Weld’s material (see above) and redescribe the species in line with characters included in our description of
<taxonomicName id="53894D45FFE14135FA76FB0D046BD140" authority="Nastasi" authorityName="Nastasi &amp; Deans" authorityYear="2024" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="519" phylum="Arthropoda" rank="species" species="rugosa" status="sp. nov.">
<emphasis id="A6FDEAD4FFE14135FA76FB0D05B1D141" italics="true" pageId="8" pageNumber="519">P. rugosa</emphasis>
Nastasi
</taxonomicName>
<emphasis id="A6FDEAD4FFE14135FEB6FB3104F7D140" bold="true" box="[328,419,1267,1293]" pageId="8" pageNumber="519">
<taxonomicNameLabel id="BDCE57AFFFE14135FEB6FB3104F7D140" box="[328,419,1267,1293]" pageId="8" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(below). As mentioned above, further study of material of this species is badly needed to better understand its limits;
<bibRefCitation id="F0184B37FFE14135FE24FAD507C4D17C" author="Melika, G." box="[474,656,1303,1329]" pageId="8" pageNumber="519" refId="ref9772" refString="Melika, G. (2006) Gall wasps of Ukraine. Vol. 1. Vestnik zoologii, The Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev, 300 pp." type="book" year="2006">Melika’s (2006)</bibRefCitation>
treatment of
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<emphasis id="A6FDEAD4FFE14135FCD3FAD50683D17C" box="[813,983,1303,1329]" italics="true" pageId="8" pageNumber="519">P. fedtschenkoi</emphasis>
</taxonomicName>
probably encapsulates multiple species (see remarks for
<taxonomicName id="53894D45FFE14135FEA7FAF90480D118" authorityName="Kieffer" authorityYear="1901" box="[345,468,1339,1365]" class="Insecta" family="Cynipidae" genus="Panteliella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="519" phylum="Arthropoda" rank="genus">
<emphasis id="A6FDEAD4FFE14135FEA7FAF90480D118" box="[345,468,1339,1365]" italics="true" pageId="8" pageNumber="519">Panteliella</emphasis>
</taxonomicName>
above). Given that the material we examined here was dissected by Weld from galls preserved alongside Rübsaamen’s
<typeStatus id="4B328864FFE14135FDDCFA9D0706D134" box="[546,594,1375,1401]" pageId="8" pageNumber="519">type</typeStatus>
material (
<bibRefCitation id="F0184B37FFE14135FD34FA9D0618D134" author="Weld, L. H." box="[714,844,1375,1401]" pageId="8" pageNumber="519" pagination="137 - 144" refId="ref10592" refString="Weld, L. H. (1930) Notes on types (Hymenoptera: Cynipidae). Proceedings of the Entomological Society of Washington, 32 (8), 137 - 144." type="journal article" year="1930">Weld 1930</bibRefCitation>
), and that the known adult
<typeStatus id="4B328864FFE14135FB73FA9D019CD134" box="[1165,1224,1375,1401]" pageId="8" pageNumber="519">types</typeStatus>
of the species are incomplete and mounted on slides (
<bibRefCitation id="F0184B37FFE14135FDCEFA410784D1D0" author="Quinlan, J." box="[560,720,1411,1437]" pageId="8" pageNumber="519" pagination="275 - 286" refId="ref10265" refString="Quinlan, J. (1968) Cynipinae (Hymenoptera) occurring on Phlomis L. Transactions of the Royal Entomological Society of London, 120, 275 - 286. https: // doi. org / 10.1111 / j. 1365 - 2311.1968. tb 00339. x" type="journal article" year="1968">Quinlan 1968</bibRefCitation>
,
<bibRefCitation id="F0184B37FFE14135FD23FA410693D1D0" author="Nieves-Aldrey, J. L." box="[733,967,1411,1437]" pageId="8" pageNumber="519" pagination="175 - 206" refId="ref10135" refString="Nieves-Aldrey, J. L. (1994) Revision of West-European Genera of the Tribe Aylacini Ashmead (Hymenoptera, Cynipidae). Journal of Hymenoptera Research, 3, 175 - 206." type="journal article" year="1994">Nieves-Aldrey 1994</bibRefCitation>
), we believe that our treatment here best represents the species hypothesis as intended by Rübsaamen.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -127,5 +127,20 @@ Nastasi
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* = We include
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<emphasis id="A6FDEAD4FFEE413AFEC1FC18075ED7BF" box="[319,522,986,1010]" italics="true" pageId="7" pageNumber="518">Panteliella bianchii</emphasis>
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in the key based on characters given in the original description (see translation below). Examination of the
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<emphasis id="A6FDEAD4FFEE413AFE53FC3D074DD05B" box="[429,537,1022,1046]" italics="true" pageId="7" pageNumber="518">P. bianchii</emphasis>
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will be needed to confirm the diagnostic characters given here.
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<emphasis id="918A492ECD09FFFE83CCFE69D9EB31B6" bold="true" box="[151,459,432,459]" italics="true" pageId="1" pageNumber="246">Eremobothynus cornutus</emphasis>
(
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)
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(
<figureCitation id="3BC589B9CD09FFFE83C4FE0FD9313193" box="[159,273,470,494]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2‒12" captionStartId-0="2.[151,250,1818,1842]" captionStartId-1="3.[151,264,1716,1740]" captionTargetBox-0="[172,1414,320,1781]" captionTargetBox-1="[180,1415,194,1676]" captionTargetId-0="figure-90@2.[160,1427,307,1794]" captionTargetId-1="figure-19@3.[151,1436,181,1693]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Records of Eremobothynus cornutus. Green star: type locality of Eremobothynus cornutus, yellow star: type locality of Aphonides turbator, solid red circles: material examined with locality data, open red circles: material examined without precise locality data, black circles: previous bibliographic records." captionText-1="FIGURES 2‒12. Eremobothynus cornutus (female specimens). 2, dorsal view of Aphonides turbator holotype (ZMB); 3, ventral view of Aphonides turbator holotype (ZMB); 4, lateral view of Aphonides turbator holotype (ZMB); 5, labels of Aphonides turbator holotype; 6, dorsal view of Aphonides turbator paratype (ZMB); 7, ventral view of Aphonides turbator paratype (ZMB); 8, lateral view of Aphonides turbator paratype (ZMB); 9, labels of Aphonides turbator paratype; 10, dorsal view of a specimen from Córdoba Province, Argentina (LEBA); 11, ventral view of a specimen from Córdoba Province, Argentina (LEBA); 12, lateral view of a specimen from Córdoba Province, Argentina (LEBA). Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14240866" figureDoi-1="http://doi.org/10.5281/zenodo.14240868" httpUri-0="https://zenodo.org/record/14240866/files/figure.png" httpUri-1="https://zenodo.org/record/14240868/files/figure.png" pageId="1" pageNumber="246">Figs. 1‒12</figureCitation>
)
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(original combination).
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<emphasis id="918A492ECD09FFFE83CCFDCAD9B53256" box="[151,405,531,555]" italics="true" pageId="1" pageNumber="246">Eremobothynus cornutus</emphasis>
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:
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<treatmentCitation id="225FB32DCD09FFFE83CCFDEBDA0B3237" author="Endrodi, S." box="[151,555,562,586]" page="199" pageId="1" pageNumber="246" year="1981">
<taxonomicName id="64FEEEBFCD09FFFE83CCFDEBDA0B3237" ID-CoL="FJH9" authority="Endrodi, 1981: 199" authorityName="Endrodi" authorityPageNumber="199" authorityYear="1981" box="[151,555,562,586]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD09FFFE83CCFDEBD97C3237" box="[151,348,562,586]" italics="true" pageId="1" pageNumber="246">Aphonides turbator</emphasis>
<bibRefCitation id="C76FE8CDCD09FFFE8238FDEBDA0B3237" author="Endrodi, S." box="[355,555,562,586]" pageId="1" pageNumber="246" pagination="197 - 202" refId="ref3414" refString="Endrodi, S. (1981) Neue und seltene Dynastinen aus Sudamerika und eine synonymische Bemerkung (Coleoptera, Melolonthidae). Annales Historico-Naturales Musei Nationalis Hungarici, 73, 197 - 202." type="journal article" year="1981">Endrödi, 1981: 199</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<emphasis id="918A492ECD09FFFE816EFDEBDAE83237" bold="true" box="[565,712,562,586]" pageId="1" pageNumber="246">new synonym</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A341953CCD09FFFE83CCFD88DAB43214" blockId="1.[151,831,432,617]" box="[151,660,593,617]" pageId="1" pageNumber="246">
<treatmentCitationGroup id="83EEB212CD09FFFE83CCFD88DAB43214" box="[151,660,593,617]" pageId="1" pageNumber="246">
<taxonomicName id="64FEEEBFCD09FFFE83CCFD88D9A13214" authority="(Endrodi, 1981)" baseAuthorityName="Endrodi" baseAuthorityYear="1981" box="[151,385,593,617]" class="Insecta" family="Scarabaeidae" genus="Anoplognatho" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD09FFFE83CCFD88D9A13214" box="[151,385,593,617]" italics="true" pageId="1" pageNumber="246">Anoplognatho turbator</emphasis>
</taxonomicName>
:
<treatmentCitation id="225FB32DCD09FFFE82D4FD88DAAF3214" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Le Tirant, S." box="[399,655,593,617]" page="417" pageId="1" pageNumber="246" year="2023">
<bibRefCitation id="C76FE8CDCD09FFFE82D4FD88DAAF3214" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Le Tirant, S." box="[399,655,593,617]" pageId="1" pageNumber="246" pagination="1 - 486" refId="ref3715" refString="Ratcliffe, B. C., Cave, R. D. &amp; Le Tirant, S. (2023) The dynastine scarab beetles of Argentina, Paraguay, and Uruguay (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of University of Nebraska State Museum, 34, 1 - 486." type="journal article" year="2023">
Ratcliffe
<emphasis id="918A492ECD09FFFE82B6FD8BDA013214" box="[493,545,593,617]" italics="true" pageId="1" pageNumber="246">et al.</emphasis>
2023: 417
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="EBE4C6B7CD09FFFE83CCFD43DAD63770" pageId="1" pageNumber="246" type="materials_examined">
<paragraph id="A341953CCD09FFFE83CCFD43DC2C32A8" blockId="1.[151,1437,665,2021]" pageId="1" pageNumber="246">
<emphasis id="918A492ECD09FFFE83CCFD43D91832CC" box="[151,312,665,689]" italics="true" pageId="1" pageNumber="246">
<typeStatus id="7C452B9ECD09FFFE83CCFD43D8E932CC" box="[151,201,666,689]" pageId="1" pageNumber="246">Type</typeStatus>
localities:
</emphasis>
Bahía Blanca [
<geoCoordinate id="C6CAF3FBCD09FFFE828DFD40DA1132CC" box="[470,561,665,689]" degrees="38" direction="south" minutes="43" orientation="latitude" pageId="1" pageNumber="246" precision="925" value="-38.716667">38°43’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE8160FD40DA8132CC" box="[571,673,665,689]" degrees="62" direction="west" minutes="16" orientation="longitude" pageId="1" pageNumber="246" precision="925" value="-62.266666">62°16’W</geoCoordinate>
],
<collectingRegion id="613A5BDECD09FFFE81EAFD40DB8532CC" box="[689,933,665,689]" country="Argentina" name="Buenos Aires" pageId="1" pageNumber="246">Buenos Aires Province</collectingRegion>
,
<collectingCountry id="DBE9D5ACCD09FFFE80F0FD40DC3532CC" box="[939,1045,665,689]" name="Argentina" pageId="1" pageNumber="246">Argentina</collectingCountry>
(
<taxonomicName id="64FEEEBFCD09FFFE8778FD40DD0B32CC" baseAuthorityName="Steinheil" baseAuthorityYear="1874" box="[1059,1323,665,689]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="918A492ECD09FFFE8778FD40DD0B32CC" box="[1059,1323,665,689]" italics="true" pageId="1" pageNumber="246">Eremobothynus cornutus</emphasis>
</taxonomicName>
); Estación Crespo [
<geoCoordinate id="C6CAF3FBCD09FFFE83ABFD64D96C32A8" box="[240,332,701,725]" degrees="32" direction="south" minutes="02" orientation="latitude" pageId="1" pageNumber="246" precision="925" value="-32.033333">32°02’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE820CFD64D99D32A8" box="[343,445,701,725]" degrees="60" direction="west" minutes="19" orientation="longitude" pageId="1" pageNumber="246" precision="925" value="-60.316666">60°19’W</geoCoordinate>
],
<collectingRegion id="613A5BDECD09FFFE828BFD64DA8A32A8" box="[464,682,701,725]" country="Argentina" name="Entre Rios" pageId="1" pageNumber="246">Entre Ríos Province</collectingRegion>
,
<collectingCountry id="DBE9D5ACCD09FFFE81E9FD64DB3C32A8" box="[690,796,701,725]" name="Argentina" pageId="1" pageNumber="246">Argentina</collectingCountry>
(
<taxonomicName id="64FEEEBFCD09FFFE8077FD64DBDC32A8" authorityName="Endrodi" authorityYear="1981" box="[812,1020,701,725]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD09FFFE8077FD64DBDC32A8" box="[812,1020,701,725]" italics="true" pageId="1" pageNumber="246">Aphonides turbator</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="A341953CCD09FFFE839CFD3BDB9533F4" blockId="1.[151,1437,665,2021]" pageId="1" pageNumber="246">
<emphasis id="918A492ECD09FFFE839CFD3BDA3B3284" box="[199,539,737,761]" italics="true" pageId="1" pageNumber="246">Previous bibliographic records:</emphasis>
<collectingCountry id="DBE9D5ACCD09FFFE8179FD38DAE23284" box="[546,706,737,761]" name="Argentina" pageId="1" pageNumber="246">ARGENTINA</collectingCountry>
:
<collectingRegion id="613A5BDECD09FFFE8190FD38DB9F3284" box="[715,959,737,761]" country="Argentina" name="Buenos Aires" pageId="1" pageNumber="246">Buenos Aires Province</collectingRegion>
(
<bibRefCitation id="C76FE8CDCD09FFFE8097FD38DC4C3284" author="Steinheil, E." box="[972,1132,737,761]" pageId="1" pageNumber="246" pagination="554 - 578" refId="ref3840" refString="Steinheil, E. (1874) Symbolae ad historiam Coleopterorum Argentinae meridionalis, ossia enumerazione dei coleotteri raccolti dal Prof. Strobel, durante il suo soggiorno in Buenos Aires e nei viaggi de la intrapresi a Mendoza e nal Chili, indi a Bahia Blanca ed al Carmen de los Patagones; e descrizione delle specie nuove. Atti Della Societa Italiana di Scienze Naturali, 15, 554 - 578. [Dated 1872]." type="journal article" year="1874">Steinheil 1874</bibRefCitation>
;
<bibRefCitation id="C76FE8CDCD09FFFE872DFD38DCD93284" author="Ohaus, F." box="[1142,1273,737,761]" pageId="1" pageNumber="246" pagination="173 - 186" refId="ref3669" refString="Ohaus, F. (1910) Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift, 1910, 173 - 186." type="journal article" year="1910">Ohaus 1910</bibRefCitation>
,
<bibRefCitation id="C76FE8CDCD09FFFE8658FD38DD1C3284" author="Ohaus, F." box="[1283,1340,737,761]" pageId="1" pageNumber="246" pagination="299 - 304" refId="ref3692" refString="Ohaus, F. (1914) Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift, 1914, 299 - 304." type="journal article" year="1914">1914</bibRefCitation>
;
<bibRefCitation id="C76FE8CDCD09FFFE861DFD38D8EF3360" author="Endrodi, S." pageId="1" pageNumber="246" pagination="1 - 145" refId="ref3349" refString="Endrodi, S. (1969) Monographie der Dynastinae, 4. Tribus, amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden, 37, 1 - 145." type="journal article" year="1969">Endrödi 1969</bibRefCitation>
,
<bibRefCitation id="C76FE8CDCD09FFFE8386FCDCD9353360" author="Endrodi, S." box="[221,277,773,797]" pageId="1" pageNumber="246" refId="ref3449" refString="Endrodi, S. (1985) The Dynastinae of the World. Dr. W. Junk, Dordrecht, Netherlands, 800 pp." type="book" year="1985">1985</bibRefCitation>
;
<bibRefCitation id="C76FE8CDCD09FFFE827FFCDCDA273360" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Le Tirant, S." box="[292,519,773,797]" pageId="1" pageNumber="246" pagination="1 - 486" refId="ref3715" refString="Ratcliffe, B. C., Cave, R. D. &amp; Le Tirant, S. (2023) The dynastine scarab beetles of Argentina, Paraguay, and Uruguay (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of University of Nebraska State Museum, 34, 1 - 486." type="journal article" year="2023">
Ratcliffe
<emphasis id="918A492ECD09FFFE82D1FCDFD9E33360" box="[394,451,773,797]" italics="true" pageId="1" pageNumber="246">et al.</emphasis>
2023
</bibRefCitation>
);
<collectingRegion id="613A5BDECD09FFFE8147FCDCDAC13360" box="[540,737,773,797]" country="Argentina" name="Cordoba" pageId="1" pageNumber="246">Córdoba Province</collectingRegion>
(
<bibRefCitation id="C76FE8CDCD09FFFE81A9FCDCDB5B3360" author="Ohaus, F." box="[754,891,773,797]" pageId="1" pageNumber="246" pagination="299 - 304" refId="ref3692" refString="Ohaus, F. (1914) Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift, 1914, 299 - 304." type="journal article" year="1914">Ohaus 1914</bibRefCitation>
;
<bibRefCitation id="C76FE8CDCD09FFFE80D3FCDCDC003360" author="Endrodi, S." box="[904,1056,773,797]" pageId="1" pageNumber="246" pagination="1 - 145" refId="ref3349" refString="Endrodi, S. (1969) Monographie der Dynastinae, 4. Tribus, amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden, 37, 1 - 145." type="journal article" year="1969">Endrödi 1969</bibRefCitation>
;
<bibRefCitation id="C76FE8CDCD09FFFE8774FCDCDD323360" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Le Tirant, S." box="[1071,1298,773,797]" pageId="1" pageNumber="246" pagination="1 - 486" refId="ref3715" refString="Ratcliffe, B. C., Cave, R. D. &amp; Le Tirant, S. (2023) The dynastine scarab beetles of Argentina, Paraguay, and Uruguay (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of University of Nebraska State Museum, 34, 1 - 486." type="journal article" year="2023">
Ratcliffe
<emphasis id="918A492ECD09FFFE87CDFCDFDCEF3360" box="[1174,1231,773,797]" italics="true" pageId="1" pageNumber="246">et al.</emphasis>
2023
</bibRefCitation>
);
<collectingRegion id="613A5BDECD09FFFE8673FCDCD8D6333C" country="Argentina" name="Entre Rios" pageId="1" pageNumber="246">Entre Ríos Province</collectingRegion>
(
<bibRefCitation id="C76FE8CDCD09FFFE8258FCF0D9B9333C" author="Endrodi, S." box="[259,409,809,833]" pageId="1" pageNumber="246" pagination="197 - 202" refId="ref3414" refString="Endrodi, S. (1981) Neue und seltene Dynastinen aus Sudamerika und eine synonymische Bemerkung (Coleoptera, Melolonthidae). Annales Historico-Naturales Musei Nationalis Hungarici, 73, 197 - 202." type="journal article" year="1981">Endrödi 1981</bibRefCitation>
;
<bibRefCitation id="C76FE8CDCD09FFFE82FFFCF0DA5C333C" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Le Tirant, S." box="[420,636,809,833]" pageId="1" pageNumber="246" pagination="1 - 486" refId="ref3715" refString="Ratcliffe, B. C., Cave, R. D. &amp; Le Tirant, S. (2023) The dynastine scarab beetles of Argentina, Paraguay, and Uruguay (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of University of Nebraska State Museum, 34, 1 - 486." type="journal article" year="2023">
Ratcliffe
<emphasis id="918A492ECD09FFFE815CFCF3DA1C333C" box="[519,572,809,833]" italics="true" pageId="1" pageNumber="246">et al.</emphasis>
2023
</bibRefCitation>
);
<collectingRegion id="613A5BDECD09FFFE81D5FCF0DB0D333C" box="[654,813,809,833]" country="Argentina" name="Jujuy" pageId="1" pageNumber="246">Jujuy Province</collectingRegion>
(
<bibRefCitation id="C76FE8CDCD09FFFE8061FCF0DC34333C" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Le Tirant, S." box="[826,1044,809,833]" pageId="1" pageNumber="246" pagination="1 - 486" refId="ref3715" refString="Ratcliffe, B. C., Cave, R. D. &amp; Le Tirant, S. (2023) The dynastine scarab beetles of Argentina, Paraguay, and Uruguay (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of University of Nebraska State Museum, 34, 1 - 486." type="journal article" year="2023">
Ratcliffe
<emphasis id="918A492ECD09FFFE80C4FCF3DBF4333C" box="[927,980,809,833]" italics="true" pageId="1" pageNumber="246">et al.</emphasis>
2023
</bibRefCitation>
);
<collectingRegion id="613A5BDECD09FFFE877EFCF0DCD6333C" box="[1061,1270,809,833]" country="Argentina" name="La Pampa" pageId="1" pageNumber="246">La Pampa Province</collectingRegion>
(
<bibRefCitation id="C76FE8CDCD09FFFE8659FCF0D8F13318" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Le Tirant, S." pageId="1" pageNumber="246" pagination="1 - 486" refId="ref3715" refString="Ratcliffe, B. C., Cave, R. D. &amp; Le Tirant, S. (2023) The dynastine scarab beetles of Argentina, Paraguay, and Uruguay (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of University of Nebraska State Museum, 34, 1 - 486." type="journal article" year="2023">
Ratcliffe
<emphasis id="918A492ECD09FFFE863CFCF3DDBD333C" box="[1383,1437,809,833]" italics="true" pageId="1" pageNumber="246">et al.</emphasis>
2023
</bibRefCitation>
);
<collectingRegion id="613A5BDECD09FFFE83BEFC94D9A13318" box="[229,385,845,869]" country="Argentina" name="Salta" pageId="1" pageNumber="246">Salta Province</collectingRegion>
(
<bibRefCitation id="C76FE8CDCD09FFFE82D4FC94DA363318" author="Ohaus, F." box="[399,534,845,869]" pageId="1" pageNumber="246" pagination="173 - 186" refId="ref3669" refString="Ohaus, F. (1910) Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift, 1910, 173 - 186." type="journal article" year="1910">Ohaus 1910</bibRefCitation>
;
<bibRefCitation id="C76FE8CDCD09FFFE817AFC94DADE3318" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Le Tirant, S." box="[545,766,845,869]" pageId="1" pageNumber="246" pagination="1 - 486" refId="ref3715" refString="Ratcliffe, B. C., Cave, R. D. &amp; Le Tirant, S. (2023) The dynastine scarab beetles of Argentina, Paraguay, and Uruguay (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of University of Nebraska State Museum, 34, 1 - 486." type="journal article" year="2023">
Ratcliffe
<emphasis id="918A492ECD09FFFE81DDFC97DA9D3318" box="[646,701,845,869]" italics="true" pageId="1" pageNumber="246">et al.</emphasis>
2023
</bibRefCitation>
);
<collectingRegion id="613A5BDECD09FFFE8049FC94DBF83318" box="[786,984,845,869]" country="Argentina" name="San Luis" pageId="1" pageNumber="246">San Luis Province</collectingRegion>
(
<bibRefCitation id="C76FE8CDCD09FFFE80BDFC94DCE53318" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Le Tirant, S." box="[998,1221,845,869]" pageId="1" pageNumber="246" pagination="1 - 486" refId="ref3715" refString="Ratcliffe, B. C., Cave, R. D. &amp; Le Tirant, S. (2023) The dynastine scarab beetles of Argentina, Paraguay, and Uruguay (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of University of Nebraska State Museum, 34, 1 - 486." type="journal article" year="2023">
Ratcliffe
<emphasis id="918A492ECD09FFFE8717FC97DCA33318" box="[1100,1155,845,869]" italics="true" pageId="1" pageNumber="246">et al.</emphasis>
2023
</bibRefCitation>
);
<collectingRegion id="613A5BDECD09FFFE8783FC94DDBC3318" box="[1240,1436,845,869]" country="Argentina" name="Santa Fe" pageId="1" pageNumber="246">Santa Fe Province</collectingRegion>
(
<bibRefCitation id="C76FE8CDCD09FFFE83C5FCA8D95A33F4" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Le Tirant, S." box="[158,378,881,905]" pageId="1" pageNumber="246" pagination="1 - 486" refId="ref3715" refString="Ratcliffe, B. C., Cave, R. D. &amp; Le Tirant, S. (2023) The dynastine scarab beetles of Argentina, Paraguay, and Uruguay (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of University of Nebraska State Museum, 34, 1 - 486." type="journal article" year="2023">
Ratcliffe
<emphasis id="918A492ECD09FFFE825FFCABD91933F4" box="[260,313,881,905]" italics="true" pageId="1" pageNumber="246">et al.</emphasis>
2023
</bibRefCitation>
);
<collectingRegion id="613A5BDECD09FFFE82D7FCA8DAE133F4" box="[396,705,881,905]" country="Argentina" name="Santiago del Estero" pageId="1" pageNumber="246">Santiago del Estero Province</collectingRegion>
(
<bibRefCitation id="C76FE8CDCD09FFFE8195FCA8DB8A33F4" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Le Tirant, S." box="[718,938,881,905]" pageId="1" pageNumber="246" pagination="1 - 486" refId="ref3715" refString="Ratcliffe, B. C., Cave, R. D. &amp; Le Tirant, S. (2023) The dynastine scarab beetles of Argentina, Paraguay, and Uruguay (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of University of Nebraska State Museum, 34, 1 - 486." type="journal article" year="2023">
Ratcliffe
<emphasis id="918A492ECD09FFFE806FFCABDB4933F4" box="[820,873,881,905]" italics="true" pageId="1" pageNumber="246">et al.</emphasis>
2023
</bibRefCitation>
).
</paragraph>
<paragraph id="A341953CCD09FFFE839CFC4CDAD63770" blockId="1.[151,1437,665,2021]" pageId="1" pageNumber="246">
<emphasis id="918A492ECD09FFFE839CFC4CD9BA33D0" box="[199,410,917,941]" italics="true" pageId="1" pageNumber="246">Material examined:</emphasis>
Photographs of
<typeStatus id="7C452B9ECD09FFFE811CFC4CDA8433D0" box="[583,676,917,941]" pageId="1" pageNumber="246" type="holotype">holotype</typeStatus>
(
<figureCitation id="3BC589B9CD09FFFE81E9FC4CDB3733D0" box="[690,791,917,941]" captionStart="FIGURES 2‒12" captionStartId="3.[151,264,1716,1740]" captionTargetBox="[180,1415,194,1676]" captionTargetId="figure-19@3.[151,1436,181,1693]" captionTargetPageId="3" captionText="FIGURES 2‒12. Eremobothynus cornutus (female specimens). 2, dorsal view of Aphonides turbator holotype (ZMB); 3, ventral view of Aphonides turbator holotype (ZMB); 4, lateral view of Aphonides turbator holotype (ZMB); 5, labels of Aphonides turbator holotype; 6, dorsal view of Aphonides turbator paratype (ZMB); 7, ventral view of Aphonides turbator paratype (ZMB); 8, lateral view of Aphonides turbator paratype (ZMB); 9, labels of Aphonides turbator paratype; 10, dorsal view of a specimen from Córdoba Province, Argentina (LEBA); 11, ventral view of a specimen from Córdoba Province, Argentina (LEBA); 12, lateral view of a specimen from Córdoba Province, Argentina (LEBA). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14240868" httpUri="https://zenodo.org/record/14240868/files/figure.png" pageId="1" pageNumber="246">Figs. 2‒4</figureCitation>
) and
<specimenCount id="B5F85EB5CD09FFFE800BFC4CDBF933D0" box="[848,985,917,941]" count="1" pageId="1" pageNumber="246" type="generic" typeStatus="paratype">one paratype</specimenCount>
(
<figureCitation id="3BC589B9CD09FFFE80BDFC4CDC6B33D0" box="[998,1099,917,941]" captionStart="FIGURES 2‒12" captionStartId="3.[151,264,1716,1740]" captionTargetBox="[180,1415,194,1676]" captionTargetId="figure-19@3.[151,1436,181,1693]" captionTargetPageId="3" captionText="FIGURES 2‒12. Eremobothynus cornutus (female specimens). 2, dorsal view of Aphonides turbator holotype (ZMB); 3, ventral view of Aphonides turbator holotype (ZMB); 4, lateral view of Aphonides turbator holotype (ZMB); 5, labels of Aphonides turbator holotype; 6, dorsal view of Aphonides turbator paratype (ZMB); 7, ventral view of Aphonides turbator paratype (ZMB); 8, lateral view of Aphonides turbator paratype (ZMB); 9, labels of Aphonides turbator paratype; 10, dorsal view of a specimen from Córdoba Province, Argentina (LEBA); 11, ventral view of a specimen from Córdoba Province, Argentina (LEBA); 12, lateral view of a specimen from Córdoba Province, Argentina (LEBA). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.14240868" httpUri="https://zenodo.org/record/14240868/files/figure.png" pageId="1" pageNumber="246">Figs. 6‒8</figureCitation>
) of
<taxonomicName id="64FEEEBFCD09FFFE872EFC4CDDB933D0" authority="(ZMB)" baseAuthorityName="ZMB" box="[1141,1433,917,941]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD09FFFE872EFC4CDD6433D0" box="[1141,1348,917,941]" italics="true" pageId="1" pageNumber="246">Aphonides turbator</emphasis>
(ZMB)
</taxonomicName>
.
<materialsCitation id="13969F61CD09FFFE83CCFC60DB033388" collectingDate="1946-11-09" collectingDateMax="1946-11-12" collectingDateMin="1946-11-09" collectionCode="FML, MLP" collectorName="K. Hayward &amp; A. Willink &amp; C. Bruch" country="Argentina" latitude="-38.008335" location="Bahia Blanca" longLatPrecision="117" longitude="-62.816666" municipality="Bahia Blanca" pageId="1" pageNumber="246" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Buenos Aires Province">
<collectingCountry id="DBE9D5ACCD09FFFE83CCFC60D91733AC" box="[151,311,953,977]" name="Argentina" pageId="1" pageNumber="246">ARGENTINA</collectingCountry>
:
<emphasis id="918A492ECD09FFFE8264FC60DA6333AC" bold="true" box="[319,579,953,977]" pageId="1" pageNumber="246">
<collectingRegion id="613A5BDECD09FFFE8264FC60DA1D33AC" box="[319,573,953,977]" country="Argentina" name="Buenos Aires" pageId="1" pageNumber="246">Buenos Aires Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="43250F46CD09FFFE811CFC60DAF333AC" box="[583,723,953,977]" pageId="1" pageNumber="246">Bahía Blanca</collectingMunicipality>
[
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,
<geoCoordinate id="C6CAF3FBCD09FFFE800CFC60DBF233AC" box="[855,978,953,977]" degrees="62" direction="west" minutes="16.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-62.266666">62°16.0’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE80BAFC60DC5133AC" box="[993,1137,953,977]" pageId="1" pageNumber="246" value="1946-11-09" valueMax="1946-11-12" valueMin="1946-11-09">
<collectingDate id="C7044A14CD09FFFE80BAFC60DC5133AC" box="[993,1137,953,977]" pageId="1" pageNumber="246" value="1946-11-09" valueMax="1946-11-12" valueMin="1946-11-09">9–12.xi.1946</collectingDate>
</date>
,
<collectorName id="0E0BF0EACD09FFFE8722FC60DCDA33AC" box="[1145,1274,953,977]" pageId="1" pageNumber="246">K. Hayward</collectorName>
and
<collectorName id="0E0BF0EACD09FFFE867CFC60DDB833AC" box="[1319,1432,953,977]" pageId="1" pageNumber="246">A. Willink</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE83CCFC04D89E3389" box="[151,190,989,1013]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
and
<specimenCount id="B5F85EB5CD09FFFE83ABFC04D9383389" box="[240,280,989,1013]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE827EFC04D97F3388" box="[293,351,989,1013]" country="Argentina" httpUri="http://grbio.org/cool/sggj-9jsz" name="Fundacion Miguel Lillo" pageId="1" pageNumber="246">FML</collectionCode>
) and [without date],
<collectorName id="0E0BF0EACD09FFFE8161FC04DABC3388" box="[570,668,989,1013]" pageId="1" pageNumber="246">C. Bruch</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE81FDFC04DAED3389" box="[678,717,989,1013]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE8180FC04DB373388" box="[731,791,989,1013]" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="246">MLP</collectionCode>
);
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE8072FC04D9983464" collectingDate="1994-11-13" collectingDateMax="1994-11-13" collectingDateMin="1941-12" collectionCode="MMPE" collectorName="J. Farina" country="Argentina" latitude="-38.008335" location="Estafeta Chapadmalal" longLatPrecision="117" longitude="-62.816666" municipality="Bahia Blanca" pageId="1" pageNumber="246" specimenCount="2" specimenCount-female="2" specimenCount-male="2" stateProvince="Buenos Aires">
<location id="A621C3E7CD09FFFE8072FC04DC313388" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE8072FC04DC313388" box="[809,1041,989,1013]" country="Argentina" latitude="-38.008335" longLatPrecision="117" longitude="-62.816666" municipality="Bahia Blanca" name="Estafeta Chapadmalal" pageId="1" pageNumber="246" stateProvince="Buenos Aires">Estafeta Chapadmalal</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE8745FC04DCAD3388" box="[1054,1165,989,1013]" degrees="38" direction="south" minutes="10.1" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-38.168335">38°10.1’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE87C3FC04DD333388" box="[1176,1299,989,1013]" degrees="57" direction="west" minutes="38.2" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-57.636665">57°38.2’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE8678FC04DDB83388" box="[1315,1432,989,1013]" pageId="1" pageNumber="246" value="1994-11-13">
<collectingDate id="C7044A14CD09FFFE8678FC04DDB83388" box="[1315,1432,989,1013]" pageId="1" pageNumber="246" value="1994-11-13">13.xi.1994</collectingDate>
</date>
,
<collectorName id="0E0BF0EACD09FFFE83CCFBD8D8D93464" box="[151,249,1025,1049]" pageId="1" pageNumber="246">J. Farina</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE825CFBD8D9673465" box="[263,327,1025,1049]" count="2" pageId="1" pageNumber="246" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE8203FBD8D98D3464" box="[344,429,1025,1049]" pageId="1" pageNumber="246">MMPE</collectionCode>
);
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE8299FBD8DD483464" box="[450,1384,1025,1049]" collectingDate="1941-12" collectingDateMax="1994-11-13" collectingDateMin="1941-12" collectionCode="FML, MLP, MMPE" collectorName="A. Martinez" country="Argentina" latitude="-38.008335" location="Felipe Sola" longLatPrecision="117" longitude="-62.816666" municipality="Bahia Blanca" pageId="1" pageNumber="246" specimenCode="MACN-En 40338" specimenCount="1" specimenCount-female="1" specimenCount-male="2" stateProvince="Buenos Aires">
<location id="A621C3E7CD09FFFE8299FBD8DA1E3464" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE8299FBD8DA1E3464" box="[450,574,1025,1049]" country="Argentina" latitude="-38.008335" longLatPrecision="117" longitude="-62.816666" municipality="Bahia Blanca" name="Felipe Sola" pageId="1" pageNumber="246" stateProvince="Buenos Aires">Felipe Solá</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE8114FBD8DAE03464" box="[591,704,1025,1049]" degrees="38" direction="south" minutes="00.5" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-38.008335">38°00.5’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE8195FBD8DB683464" box="[718,840,1025,1049]" degrees="62" direction="west" minutes="49.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-62.816666">62°49.0’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE8006FBD8DB983464" box="[861,952,1025,1049]" pageId="1" pageNumber="246" value="1941-12">
<collectingDate id="C7044A14CD09FFFE8006FBD8DB983464" box="[861,952,1025,1049]" pageId="1" pageNumber="246" value="1941-12">xii.1941</collectingDate>
</date>
,
<collectorName id="0E0BF0EACD09FFFE809EFBD8DC6C3464" box="[965,1100,1025,1049]" pageId="1" pageNumber="246">A. Martínez</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE8701FBD8DCA53465" box="[1114,1157,1025,1049]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(
<specimenCode id="F3583D47CD09FFFE87CDFBD8DD403464" box="[1174,1376,1025,1049]" pageId="1" pageNumber="246">MACN-En 40338</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="13969F61CD09FFFE862DFBD8DC393440" collectingDate="1998-11-23" collectingDateMax="1998-11-25" collectingDateMin="1998-11-23" collectionCode="MMPE" collectorName="J. Farina" country="Argentina" latitude="-37.695" location="Las Grutas Necochea" longLatPrecision="117" longitude="-62.29" municipality="Las Grutas Necochea" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Buenos Aires">
<collectingMunicipality id="43250F46CD09FFFE862DFBD8D9703440" pageId="1" pageNumber="246">Las Grutas Necochea</collectingMunicipality>
[
<geoCoordinate id="C6CAF3FBCD09FFFE8204FBFCD9EF3440" box="[351,463,1061,1085]" degrees="38" direction="south" minutes="36.5" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-38.608334">38°36.5’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE8287FBFCDA773440" box="[476,599,1061,1085]" degrees="58" direction="west" minutes="48.5" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-58.808334">58°48.5’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE8131FBFCDB263440" box="[618,774,1061,1085]" pageId="1" pageNumber="246" value="1998-11-23" valueMax="1998-11-25" valueMin="1998-11-23">
<collectingDate id="C7044A14CD09FFFE8131FBFCDB263440" box="[618,774,1061,1085]" pageId="1" pageNumber="246" value="1998-11-23" valueMax="1998-11-25" valueMin="1998-11-23">23–25.xi.1998</collectingDate>
</date>
,
<collectorName id="0E0BF0EACD09FFFE8048FBFCDB533440" box="[787,883,1061,1085]" pageId="1" pageNumber="246">J. Farina</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE8024FBFCDB893441" box="[895,937,1061,1085]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE80E2FBFCDC2E3440" box="[953,1038,1061,1085]" pageId="1" pageNumber="246">MMPE</collectionCode>
);
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE8779FBFCD980341C" collectingDate="2013-11-07" collectingDateMax="2013-11-07" collectingDateMin="1998-11-23" collectionCode="MMPE" collectorName="J. Farina" country="Argentina" latitude="-37.695" location="Necochea" longLatPrecision="117" longitude="-62.29" municipality="Las Grutas Necochea" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Buenos Aires">
<location id="A621C3E7CD09FFFE8779FBFCDCAB3440" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE8779FBFCDCAB3440" box="[1058,1163,1061,1085]" country="Argentina" latitude="-37.695" longLatPrecision="117" longitude="-62.29" municipality="Las Grutas Necochea" name="Necochea" pageId="1" pageNumber="246" stateProvince="Buenos Aires">Necochea</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE87C1FBFCDD2A3440" box="[1178,1290,1061,1085]" degrees="38" direction="south" minutes="34.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-38.566666">38°34.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE864CFBFCDDB23440" box="[1303,1426,1061,1085]" degrees="58" direction="west" minutes="45.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-58.75">58°45.0’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE83CCFB90D8DE341C" box="[151,254,1097,1121]" pageId="1" pageNumber="246" value="2013-11-07">
<collectingDate id="C7044A14CD09FFFE83CCFB90D8DE341C" box="[151,254,1097,1121]" pageId="1" pageNumber="246" value="2013-11-07">7.xi.2013</collectingDate>
</date>
,
<specimenCount id="B5F85EB5CD09FFFE8251FB90D912341D" box="[266,306,1097,1121]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE821BFB90D9B5341C" box="[320,405,1097,1121]" pageId="1" pageNumber="246">MMPE</collectionCode>
);
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE82FCFB90DDBC341C" box="[423,1436,1097,1121]" collectingDate="2021-10" collectingDateMax="2021-10" collectingDateMin="1998-11-23" collectionCode="LEBA" collectorName="S. Ferrer" country="Argentina" latitude="-37.695" location="Parque los Robles" longLatPrecision="117" longitude="-62.29" municipality="Las Grutas Necochea" pageId="1" pageNumber="246" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Buenos Aires">
<location id="A621C3E7CD09FFFE82FCFB90DA4B341C" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE82FCFB90DA4B341C" box="[423,619,1097,1121]" country="Argentina" latitude="-37.695" longLatPrecision="117" longitude="-62.29" municipality="Las Grutas Necochea" name="Parque los Robles" pageId="1" pageNumber="246" stateProvince="Buenos Aires">Parque los Robles</location>
, Olavarría [
<geoCoordinate id="C6CAF3FBCD09FFFE81B3FB90DB78341C" box="[744,856,1097,1121]" degrees="36" direction="south" minutes="54.4" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-36.906666">36°54.4’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE803FFB90DBFF341C" box="[868,991,1097,1121]" degrees="60" direction="west" minutes="20.5" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-60.341667">60°20.5’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE80ABFB90DC1C341C" box="[1008,1084,1097,1121]" pageId="1" pageNumber="246" value="2021-10">
<collectingDate id="C7044A14CD09FFFE80ABFB90DC1C341C" box="[1008,1084,1097,1121]" pageId="1" pageNumber="246" value="2021-10">x.2021</collectingDate>
</date>
,
<collectorName id="0E0BF0EACD09FFFE871CFB90DC87341D" box="[1095,1191,1097,1121]" pageId="1" pageNumber="246">S. Ferrer</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE87EAFB90DCFA341D" box="[1201,1242,1097,1121]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
and
<specimenCount id="B5F85EB5CD09FFFE8655FB90DD16341D" box="[1294,1334,1097,1121]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE861EFB90DDB0341C" box="[1349,1424,1097,1121]" pageId="1" pageNumber="246">LEBA</collectionCode>
);
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE83CCFBB4DD3C34F8" box="[151,1308,1133,1157]" collectingDate="1998-11-23" collectingDateMax="2021-10" collectingDateMin="1998-11-23" collectionCode="MMPE, LEBA" country="Argentina" latitude="-37.695" location="Sierra de Curamalal" longLatPrecision="117" longitude="-62.29" municipality="Las Grutas Necochea" pageId="1" pageNumber="246" specimenCode="MACN-En 40331, MACN-En 40332" specimenCount="2" specimenCount-female="2" specimenCount-male="2" stateProvince="Buenos Aires">
<location id="A621C3E7CD09FFFE83CCFBB4D94934F8" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE83CCFBB4D94934F8" box="[151,361,1133,1157]" country="Argentina" latitude="-37.695" longLatPrecision="117" longitude="-62.29" municipality="Las Grutas Necochea" name="Sierra de Curamalal" pageId="1" pageNumber="246" stateProvince="Buenos Aires">Sierra de Curamalal</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE822EFBB4D9C634F8" box="[373,486,1133,1157]" degrees="37" direction="south" minutes="41.7" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-37.695">37°41.7’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE82B4FBB4DA4934F8" box="[495,617,1133,1157]" degrees="62" direction="west" minutes="17.4" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-62.29">62°17.4’W</geoCoordinate>
], [without date],
<specimenCount id="B5F85EB5CD09FFFE8041FBB4DB7534F9" box="[794,853,1133,1157]" count="2" pageId="1" pageNumber="246" type="male">2 ♂♂</specimenCount>
(
<specimenCode id="F3583D47CD09FFFE803AFBB4DC0234F8" box="[865,1058,1133,1157]" pageId="1" pageNumber="246">MACN-En 40331</specimenCode>
and
<specimenCode id="F3583D47CD09FFFE8709FBB4DD3534F8" box="[1106,1301,1133,1157]" pageId="1" pageNumber="246">MACN-En 40332</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="13969F61CD09FFFE867DFBB4DC2234D4" collectingDate="1927-11" collectingDateMax="1928-10" collectingDateMin="1927-11" collectionCode="MLP" collectorName="M. Richter" country="Argentina" county="Sierra de la Ventana" latitude="-27.5" location="Sierra de la Ventana" longLatPrecision="123" longitude="-66.95" municipality="Tres Arroyos" pageId="1" pageNumber="246" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Catamarca">
<location id="A621C3E7CD09FFFE867DFBB4D8CD34D5" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE867DFBB4D8CD34D5" country="Argentina" county="Sierra de la Ventana" latitude="-27.5" longLatPrecision="123" longitude="-66.95" municipality="Tres Arroyos" name="Sierra de la Ventana" pageId="1" pageNumber="246" stateProvince="Catamarca">Sierra de la Ventana</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE83A2FB48D94934D4" box="[249,361,1169,1193]" degrees="38" direction="south" minutes="08.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-38.133335">38°08.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE8228FB48D9CD34D4" box="[371,493,1169,1193]" degrees="61" direction="west" minutes="47.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-61.783333">61°47.0’W</geoCoordinate>
], [without date],
<collectorName id="0E0BF0EACD09FFFE81C5FB48DB3234D4" box="[670,786,1169,1193]" pageId="1" pageNumber="246">M. Richter</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE8040FB48DB7634D5" box="[795,854,1169,1193]" count="2" pageId="1" pageNumber="246" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="B5F85EB5CD09FFFE80DCFB48DB8D34D5" box="[903,941,1169,1193]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE80E1FB48DBD634D4" box="[954,1014,1169,1193]" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="246">MLP</collectionCode>
);
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE875CFB48D9CF34B0" collectingDate="1928-10" collectingDateMax="1928-10" collectingDateMin="1927-11" collectionCode="MLP" country="Argentina" county="Sierra de la Ventana" latitude="-27.5" location="Tres Arroyos" longLatPrecision="123" longitude="-66.95" municipality="Tres Arroyos" pageId="1" pageNumber="246" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Catamarca">
<collectingMunicipality id="43250F46CD09FFFE875CFB48DCB134D4" box="[1031,1169,1169,1193]" pageId="1" pageNumber="246">Tres Arroyos</collectingMunicipality>
[
<geoCoordinate id="C6CAF3FBCD09FFFE87C6FB48DD2D34D4" box="[1181,1293,1169,1193]" degrees="38" direction="south" minutes="22.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-38.366665">38°22.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE864CFB48DDB134D4" box="[1303,1425,1169,1193]" degrees="60" direction="west" minutes="17.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-60.283333">60°17.0’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE83CCFB6CD8C334B0" box="[151,227,1205,1229]" pageId="1" pageNumber="246" value="1928-10">
<collectingDate id="C7044A14CD09FFFE83CCFB6CD8C334B0" box="[151,227,1205,1229]" pageId="1" pageNumber="246" value="1928-10">x.1928</collectingDate>
</date>
,
<specimenCount id="B5F85EB5CD09FFFE83AAFB6CD93C34B1" box="[241,284,1205,1229]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
and
<specimenCount id="B5F85EB5CD09FFFE820DFB6CD9B534B1" box="[342,405,1205,1229]" count="2" pageId="1" pageNumber="246" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE82FCFB6CD9C334B0" box="[423,483,1205,1229]" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="246">MLP</collectionCode>
);
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE82A2FB6CDCEF34B0" box="[505,1231,1205,1229]" collectingDate="1927-11" collectingDateMax="1928-10" collectingDateMin="1927-11" collectionCode="MLP" collectorName="C. Bruch" country="Argentina" county="Sierra de la Ventana" latitude="-27.5" location="La Cienaga" longLatPrecision="123" longitude="-66.95" municipality="Tres Arroyos" pageId="1" pageNumber="246" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Catamarca">
[without locality], [without date],
<collectorName id="0E0BF0EACD09FFFE802AFB6CDBF734B0" box="[881,983,1205,1229]" pageId="1" pageNumber="246">C. Bruch</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE80BFFB6CDC3034B1" box="[996,1040,1205,1229]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
and
<specimenCount id="B5F85EB5CD09FFFE8711FB6CDC5534B1" box="[1098,1141,1205,1229]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE87DDFB6CDCE234B0" box="[1158,1218,1205,1229]" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="246">MLP</collectionCode>
);
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE8782FB6CDA33348C" collectingDate="1927-11" collectingDateMax="1928-10" collectingDateMin="1927-11" collectionCode="MLP" collectorName="J. Boso" country="Argentina" county="Sierra de la Ventana" latitude="-27.5" location="La Cienaga" longLatPrecision="123" longitude="-66.95" municipality="Tres Arroyos" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="6" specimenCount-male="1" stateProvince="Catamarca">
[without locality], [without date],
<collectorName id="0E0BF0EACD09FFFE8261FB00D9AA348D" box="[314,394,1241,1264]" pageId="1" pageNumber="246">J. Boso</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE82CEFB00D99D348D" box="[405,445,1241,1265]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE8291FB00DA27348C" box="[458,519,1241,1265]" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="246">MLP</collectionCode>
).
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE8143FB00DB253568" collectingDate="1927-11" collectingDateMax="1928-10" collectingDateMin="1927-11" collectionCode="MLP" collectorName="W. Weiser &amp; F. Wolters" country="Argentina" county="Sierra de la Ventana" latitude="-27.5" location="La Cienaga" longLatPrecision="123" longitude="-66.95" municipality="Tres Arroyos" pageId="1" pageNumber="246" specimenCount="2" specimenCount-female="2" specimenCount-male="5" stateProvince="Catamarca Province">
<emphasis id="918A492ECD09FFFE8143FB00DB2B348C" bold="true" box="[536,779,1241,1265]" pageId="1" pageNumber="246">
<collectingRegion id="613A5BDECD09FFFE8143FB00DB25348C" box="[536,773,1241,1265]" country="Argentina" name="Catamarca" pageId="1" pageNumber="246">Catamarca Province</collectingRegion>
:
</emphasis>
<location id="A621C3E7CD09FFFE804AFB00DBAA348C" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE804AFB00DBAA348C" box="[785,906,1241,1265]" country="Argentina" county="Sierra de la Ventana" latitude="-27.5" longLatPrecision="123" longitude="-66.95" municipality="Tres Arroyos" name="La Cienaga" pageId="1" pageNumber="246" stateProvince="Catamarca Province">La Ciénaga</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE80CCFB00DC27348C" box="[919,1031,1241,1265]" degrees="27" direction="south" minutes="30.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-27.5">27°30.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE8749FB00DCAD348C" box="[1042,1165,1241,1265]" degrees="66" direction="west" minutes="57.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-66.95">66°57.0’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE87C5FB00DCD1348C" box="[1182,1265,1241,1265]" pageId="1" pageNumber="246" value="1927-11">
<collectingDate id="C7044A14CD09FFFE87C5FB00DCD1348C" box="[1182,1265,1241,1265]" pageId="1" pageNumber="246" value="1927-11">xi.1927</collectingDate>
</date>
,
<collectorName id="0E0BF0EACD09FFFE87A0FB00DD47348C" box="[1275,1383,1241,1265]" pageId="1" pageNumber="246">W. Weiser</collectorName>
&amp;
<collectorName id="0E0BF0EACD09FFFE86D2FB00D8CC3568" pageId="1" pageNumber="246">F. Wolters</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE83ACFB24D9143569" box="[247,308,1277,1301]" count="2" pageId="1" pageNumber="246" type="female">2 ♀♀</specimenCount>
(MACN-En 40343 and MACN-En 40344)
</materialsCitation>
;
<materialsCitation id="13969F61CD09FFFE804AFB24D8C63544" collectorName="L. Luciat" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="1">
[without locality], [without date],
<collectorName id="0E0BF0EACD09FFFE8726FB24DCC23568" box="[1149,1250,1277,1301]" pageId="1" pageNumber="246">L. Luciat</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE87B6FB24DD363569" box="[1261,1302,1277,1301]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(MACN-En 40334)
</materialsCitation>
.
<materialsCitation id="13969F61CD09FFFE83A9FAF9D9073520" collectingDate="1970-11-14" collectingDateMax="1970-11-14" collectingDateMin="1934-12" collectionCode="LEBA" collectorName="Martinez" country="Argentina" county="Aeropuerto" latitude="-32.066666" location="Aeropuerto" longLatPrecision="121" longitude="-65.03333" municipality="La Buena Parada" pageId="1" pageNumber="246" specimenCount="24" specimenCount-female="14" specimenCount-male="10" stateProvince="Cordoba Province">
<emphasis id="918A492ECD09FFFE83A9FAF9D9EC3544" bold="true" box="[242,460,1312,1337]" pageId="1" pageNumber="246">
<collectingRegion id="613A5BDECD09FFFE83A9FAF9D9E73544" box="[242,455,1312,1337]" country="Argentina" name="Cordoba" pageId="1" pageNumber="246">Córdoba Province</collectingRegion>
:
</emphasis>
<collectingCounty id="4A20EDB0CD09FFFE828FFAF8DA6D3544" box="[468,589,1313,1337]" pageId="1" pageNumber="246">Aeropuerto</collectingCounty>
de
<collectingRegion id="613A5BDECD09FFFE8123FAF8DAF43544" box="[632,724,1313,1337]" country="Argentina" name="Cordoba" pageId="1" pageNumber="246">Córdoba</collectingRegion>
[
<geoCoordinate id="C6CAF3FBCD09FFFE81B8FAF8DB733544" box="[739,851,1313,1337]" degrees="31" direction="south" minutes="19.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-31.316668">31°19.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE803BFAF8DBFA3544" box="[864,986,1313,1337]" degrees="64" direction="west" minutes="12.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-64.2">64°12.0’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE80B5FAF8DC423544" box="[1006,1122,1313,1337]" pageId="1" pageNumber="246" value="1970-11-14">
<collectingDate id="C7044A14CD09FFFE80B5FAF8DC423544" box="[1006,1122,1313,1337]" pageId="1" pageNumber="246" value="1970-11-14">14.xi.1970</collectingDate>
</date>
, A.
<collectorName id="0E0BF0EACD09FFFE87CBFAF8DCD33544" box="[1168,1267,1313,1337]" pageId="1" pageNumber="246">Martínez</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE87A4FAF8DD6A3545" box="[1279,1354,1313,1337]" count="10" pageId="1" pageNumber="246" type="male">10 ♂♂</specimenCount>
and
<specimenCount id="B5F85EB5CD09FFFE86DAFAF8D89F3521" count="14" pageId="1" pageNumber="246" type="female">14 ♀♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE838BFA9CD93B3520" box="[208,283,1349,1373]" pageId="1" pageNumber="246">LEBA</collectionCode>
);
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE826BFA9CDBD13520" box="[304,1009,1349,1373]" collectingDate="1934-12" collectingDateMax="1970-11-14" collectingDateMin="1934-12" collectionCode="MLP" collectorName="Martinez &amp; A. Bachmann" country="Argentina" county="Aeropuerto" latitude="-32.066666" location="La Buena Parada" longLatPrecision="121" longitude="-65.03333" municipality="La Buena Parada" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="15" specimenCount-male="1" stateProvince="Cordoba">
<collectingMunicipality id="43250F46CD09FFFE826BFA9CD9CB3520" box="[304,491,1349,1373]" pageId="1" pageNumber="246">La Buena Parada</collectingMunicipality>
[
<geoCoordinate id="C6CAF3FBCD09FFFE82A1FA9CDA4A3520" box="[506,618,1349,1373]" degrees="31" direction="south" minutes="10.3" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-31.171667">31°10.3’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE8123FA9CDAD23520" box="[632,754,1349,1373]" degrees="63" direction="west" minutes="36.4" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-63.606667">63°36.4’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE805DFA9CDB413520" box="[774,865,1349,1373]" pageId="1" pageNumber="246" value="1934-12">
<collectingDate id="C7044A14CD09FFFE805DFA9CDB413520" box="[774,865,1349,1373]" pageId="1" pageNumber="246" value="1934-12">xii.1934</collectingDate>
</date>
,
<specimenCount id="B5F85EB5CD09FFFE8035FA9CDBB83521" box="[878,920,1349,1373]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE80F2FA9CDBC53520" box="[937,997,1349,1373]" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="246">MLP</collectionCode>
);
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE80A1FA9CDA9635FC" collectingDate="1946-01-10" collectingDateMax="1970-11-14" collectingDateMin="1934-12" collectionCode="LEBA, MLP" collectorName="A. Bachmann" country="Argentina" county="Aeropuerto" latitude="-32.066666" location="La Poblacion" longLatPrecision="121" longitude="-65.03333" municipality="La Buena Parada" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="1" specimenCount-male="11" stateProvince="Cordoba">
<location id="A621C3E7CD09FFFE80A1FA9CDCAA3520" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE80A1FA9CDCAA3520" box="[1018,1162,1349,1373]" country="Argentina" county="Aeropuerto" latitude="-32.066666" longLatPrecision="121" longitude="-65.03333" municipality="La Buena Parada" name="La Poblacion" pageId="1" pageNumber="246" stateProvince="Cordoba">La Población</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE87C2FA9CDD293520" box="[1177,1289,1349,1373]" degrees="32" direction="south" minutes="04.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-32.066666">32°04.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE864CFA9CDDB23520" box="[1303,1426,1349,1373]" degrees="65" direction="west" minutes="02.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-65.03333">65°02.0’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE83CCFAB0D8DE35FC" box="[151,254,1385,1409]" pageId="1" pageNumber="246" value="1946-01-10">
<collectingDate id="C7044A14CD09FFFE83CCFAB0D8DE35FC" box="[151,254,1385,1409]" pageId="1" pageNumber="246" value="1946-01-10">10.i.1946</collectingDate>
</date>
,
<collectorName id="0E0BF0EACD09FFFE8251FAB0D98135FC" box="[266,417,1385,1409]" pageId="1" pageNumber="246">A. Bachmann</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE82F6FAB0D9F735FD" box="[429,471,1385,1409]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(MACN-En 40339)
</materialsCitation>
;
<materialsCitation id="13969F61CD09FFFE8198FAB0DC2935D8" collectingDate="1938-11" collectorName="G. Pellerano" country="Argentina" latitude="-31.183332" location="Rio Ceballos" longLatPrecision="121" longitude="-64.316666" pageId="1" pageNumber="246" specimenCount="4" specimenCount-male="4" stateProvince="Cordoba">
<location id="A621C3E7CD09FFFE8198FAB0DB6F35FC" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE8198FAB0DB6F35FC" box="[707,847,1385,1409]" country="Argentina" latitude="-31.183332" longLatPrecision="121" longitude="-64.316666" name="Rio Ceballos" pageId="1" pageNumber="246" stateProvince="Cordoba">Río Ceballos</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE8005FAB0DBED35FC" box="[862,973,1385,1409]" degrees="31" direction="south" minutes="11.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-31.183332">31°11.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE8081FAB0DC7535FC" box="[986,1109,1385,1409]" degrees="64" direction="west" minutes="19.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-64.316666">64°19.0’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE8733FAB0DC9B35FC" box="[1128,1211,1385,1409]" pageId="1" pageNumber="246" value="1938-11">
<collectingDate id="C7044A14CD09FFFE8733FAB0DC9B35FC" box="[1128,1211,1385,1409]" pageId="1" pageNumber="246" value="1938-11">xi.1938</collectingDate>
</date>
,
<collectorName id="0E0BF0EACD09FFFE8793FAB0DD7235FC" box="[1224,1362,1385,1409]" pageId="1" pageNumber="246">G. Pellerano</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE8605FAB0DDBC35FD" box="[1374,1436,1385,1409]" count="4" pageId="1" pageNumber="246" type="male">4 ♂♂</specimenCount>
(MACN-En 40327, MACN-En 40328, MACN-En 40329, and MACN-En 40330)
</materialsCitation>
;
<materialsCitation id="13969F61CD09FFFE874EFA54DCA035B4" country="Argentina" latitude="-33.116665" location="Rio Cuarto" longLatPrecision="120" longitude="-64.35" pageId="1" pageNumber="246" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Cordoba">
<location id="A621C3E7CD09FFFE874EFA54DCAC35D8" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE874EFA54DCAC35D8" box="[1045,1164,1421,1445]" country="Argentina" latitude="-33.116665" longLatPrecision="120" longitude="-64.35" name="Rio Cuarto" pageId="1" pageNumber="246" stateProvince="Cordoba">Río Cuarto</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE87C1FA54DD2B35D8" box="[1178,1291,1421,1445]" degrees="33" direction="south" minutes="07.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-33.116665">33°07.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE864CFA54DDB235D8" box="[1303,1426,1421,1445]" degrees="64" direction="west" minutes="21.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-64.35">64°21.0’W</geoCoordinate>
], [without date],
<specimenCount id="B5F85EB5CD09FFFE8267FA68D95835B5" box="[316,376,1457,1481]" count="2" pageId="1" pageNumber="246" type="male">2 ♂♂</specimenCount>
(MACN-En 40323 and MACN-En 40324) and
<specimenCount id="B5F85EB5CD09FFFE8020FA68DB8335B5" box="[891,931,1457,1481]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(MACN-En 40325)
</materialsCitation>
;
<materialsCitation id="13969F61CD09FFFE87D0FA68DC403590" collectingDate="1927-09" collectingDateMax="1932-12" collectingDateMin="1927-09" collectionCode="MLP" country="Argentina" latitude="-31.95" location="Villa Dolores" longLatPrecision="121" longitude="-65.183334" pageId="1" pageNumber="246" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cordoba">
<location id="A621C3E7CD09FFFE87D0FA68DD3935B4" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE87D0FA68DD3935B4" box="[1163,1305,1457,1481]" country="Argentina" latitude="-31.95" longLatPrecision="121" longitude="-65.183334" name="Villa Dolores" pageId="1" pageNumber="246" stateProvince="Cordoba">Villa Dolores</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE8673FA68DDB835B4" box="[1320,1432,1457,1481]" degrees="31" direction="south" minutes="57.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-31.95">31°57.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE83CCFA0CD9313590" box="[151,273,1493,1517]" degrees="65" direction="west" minutes="11.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-65.183334">65°11.0’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE827DFA0CD9A13590" box="[294,385,1493,1517]" pageId="1" pageNumber="246" value="1932-12">
<collectingDate id="C7044A14CD09FFFE827DFA0CD9A13590" box="[294,385,1493,1517]" pageId="1" pageNumber="246" value="1932-12">xii.1932</collectingDate>
</date>
,
<specimenCount id="B5F85EB5CD09FFFE82CBFA0CD99B3591" box="[400,443,1493,1517]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE8296FA0CDA293590" box="[461,521,1493,1517]" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="246">MLP</collectionCode>
); [without locality],
<date id="D740B3FCCD09FFFE81B4FA0CDB623590" box="[751,834,1493,1517]" pageId="1" pageNumber="246" value="1927-09">
<collectingDate id="C7044A14CD09FFFE81B4FA0CDB623590" box="[751,834,1493,1517]" pageId="1" pageNumber="246" value="1927-09">ix.1927</collectingDate>
</date>
,
<specimenCount id="B5F85EB5CD09FFFE800AFA0CDB5D3591" box="[849,893,1493,1517]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
(MACN-En 40326)
</materialsCitation>
.
<materialsCitation id="13969F61CD09FFFE8735FA0CDB11366C" collectingDate="1967-09" collectionCode="MLP" collectorName="P. Kohler" country="Argentina" county="El Volcan" latitude="-35.658333" location="El Volcan" longLatPrecision="119" longitude="-63.766666" municipality="General Pico" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Jujuy Province">
<emphasis id="918A492ECD09FFFE8735FA0CDD073590" bold="true" box="[1134,1319,1493,1517]" pageId="1" pageNumber="246">
<collectingRegion id="613A5BDECD09FFFE8735FA0CDD013590" box="[1134,1313,1493,1517]" country="Argentina" name="Jujuy" pageId="1" pageNumber="246">Jujuy Province</collectingRegion>
:
</emphasis>
<collectingCounty id="4A20EDB0CD09FFFE866AFA0CDDBD3590" box="[1329,1437,1493,1517]" pageId="1" pageNumber="246">El Volcán</collectingCounty>
[
<geoCoordinate id="C6CAF3FBCD09FFFE83C5FA20D92E366C" box="[158,270,1529,1553]" degrees="23" direction="south" minutes="55.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-23.916666">23°55.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE8243FA20D9B3366C" box="[280,403,1529,1553]" degrees="65" direction="west" minutes="28.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-65.46667">65°28.0’W</geoCoordinate>
], [without date],
<collectorName id="0E0BF0EACD09FFFE811DFA20DA8A366C" box="[582,682,1529,1553]" pageId="1" pageNumber="246">P. Kohler</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE81EFFA20DAFB366D" box="[692,731,1529,1553]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE81B3FA20DB05366C" box="[744,805,1529,1553]" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="246">MLP</collectionCode>
).
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE806DFA20D9463624" collectingDate="1967-09" collectionCode="MLP" collectorName="J. Williamson" country="Argentina" county="El Volcan" latitude="-35.658333" location="Instituto de Patologia Vegetal" longLatPrecision="119" longitude="-63.766666" municipality="General Pico" pageId="1" pageNumber="246" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="La Pampa Province">
<emphasis id="918A492ECD09FFFE806DFA20DC3E366C" bold="true" box="[822,1054,1529,1553]" pageId="1" pageNumber="246">
<collectingRegion id="613A5BDECD09FFFE806DFA20DC39366C" box="[822,1049,1529,1553]" country="Argentina" name="La Pampa" pageId="1" pageNumber="246">La Pampa Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="43250F46CD09FFFE877FFA20DC90366C" box="[1060,1200,1529,1553]" pageId="1" pageNumber="246">General Pico</collectingMunicipality>
,
<location id="A621C3E7CD09FFFE87E1FA20D8C73648" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE87E1FA20D8C73648" country="Argentina" county="El Volcan" latitude="-35.658333" longLatPrecision="119" longitude="-63.766666" municipality="General Pico" name="Instituto de Patologia Vegetal" pageId="1" pageNumber="246" stateProvince="La Pampa Province">Instituto de Patología Vegetal</location>
INTA [
<geoCoordinate id="C6CAF3FBCD09FFFE826DF9C4D9863648" box="[310,422,1565,1589]" degrees="35" direction="south" minutes="39.5" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-35.658333">35°39.5’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE82EBF9C4DA0B3648" box="[432,555,1565,1589]" degrees="63" direction="west" minutes="46.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-63.766666">63°46.0’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE8160F9C4DAAF3648" box="[571,655,1565,1589]" pageId="1" pageNumber="246" value="1967-09">
<collectingDate id="C7044A14CD09FFFE8160F9C4DAAF3648" box="[571,655,1565,1589]" pageId="1" pageNumber="246" value="1967-09">ix.1967</collectingDate>
</date>
,
<collectorName id="0E0BF0EACD09FFFE81C2F9C4DB0D3648" box="[665,813,1565,1589]" pageId="1" pageNumber="246">J. Williamson</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE806CF9C4DB7E3649" box="[823,862,1565,1589]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
(MACN-En 40335) and
<specimenCount id="B5F85EB5CD09FFFE873CF9C4DC823649" box="[1127,1186,1565,1589]" count="2" pageId="1" pageNumber="246" type="female">2 ♀♀</specimenCount>
(MACN-En 40336 and MACN-En 40337)
</materialsCitation>
;
<materialsCitation id="13969F61CD09FFFE8229F998DB5C3624" box="[370,892,1601,1625]" collectingDate="1920-01" country="Argentina" location="Pampa Central" pageId="1" pageNumber="246" specimenCount="1" specimenCount-male="1" stateProvince="Jujuy">
<location id="A621C3E7CD09FFFE8229F998DA343624" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE8229F998DA343624" box="[370,532,1601,1625]" country="Argentina" name="Pampa Central" pageId="1" pageNumber="246" stateProvince="Jujuy">Pampa Central</location>
,
<date id="D740B3FCCD09FFFE817BF998DA463624" box="[544,614,1601,1625]" pageId="1" pageNumber="246" value="1920-01">
<collectingDate id="C7044A14CD09FFFE817BF998DA463624" box="[544,614,1601,1625]" pageId="1" pageNumber="246" value="1920-01">i.1920</collectingDate>
</date>
,
<specimenCount id="B5F85EB5CD09FFFE8128F998DABD3625" box="[627,669,1601,1625]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
(MACN-En 40342)
</materialsCitation>
;
<materialsCitation id="13969F61CD09FFFE80D2F998DD0E3624" box="[905,1326,1601,1625]" collectingDate="1926-10" collectionCode="MLP" collectorName="Breyer" country="Argentina" location="La Rioja Province" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="1" stateProvince="La Rioja">
[without locality],
<date id="D740B3FCCD09FFFE870FF998DC803624" box="[1108,1184,1601,1625]" pageId="1" pageNumber="246" value="1926-10">
<collectingDate id="C7044A14CD09FFFE870FF998DC803624" box="[1108,1184,1601,1625]" pageId="1" pageNumber="246" value="1926-10">x.1926</collectingDate>
</date>
,
<specimenCount id="B5F85EB5CD09FFFE87F7F998DCF63625" box="[1196,1238,1601,1625]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE87BDF998DD033624" box="[1254,1315,1601,1625]" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="246">MLP</collectionCode>
).
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE866DF998DBD23600" collectingDate="1926-10" collectionCode="MLP" collectorName="Breyer" country="Argentina" location="La Rioja Province" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="1" stateProvince="La Rioja Province">
<emphasis id="918A492ECD09FFFE866DF998D9253600" bold="true" pageId="1" pageNumber="246">
<collectingRegion id="613A5BDECD09FFFE866DF998D8DF3600" country="Argentina" name="La Rioja" pageId="1" pageNumber="246">La Rioja Province</collectingRegion>
:
</emphasis>
[without locality], [without date], A.
<collectorName id="0E0BF0EACD09FFFE81CCF9BCDAC13600" box="[663,737,1637,1661]" pageId="1" pageNumber="246">Breyer</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE81B7F9BCDB353601" box="[748,789,1637,1661]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(MACN-En 40333)
</materialsCitation>
.
<materialsCitation id="13969F61CD09FFFE80A0F9BCDA5936DC" collectingDate="1993-10-05" collectingDateMax="1993-10-08" collectingDateMin="1993-10-05" collectionCode="MLP" collectorName="Wolffhugel" country="Argentina" county="Misiones Province" latitude="-33.8" location="Misiones Province" longLatPrecision="119" longitude="-65.55" municipality="Chimpay" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Rio Negro">
<emphasis id="918A492ECD09FFFE80A0F9BCDCF53600" bold="true" box="[1019,1237,1637,1661]" pageId="1" pageNumber="246">
<collectingCounty id="4A20EDB0CD09FFFE80A0F9BCDCEF3600" box="[1019,1231,1637,1661]" pageId="1" pageNumber="246">Misiones Province</collectingCounty>
:
</emphasis>
[without locality], [without date], K.
<collectorName id="0E0BF0EACD09FFFE8233F950D9C336DC" box="[360,483,1673,1697]" pageId="1" pageNumber="246">Wolffhügel</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE82A9F950DA3E36DD" box="[498,542,1673,1697]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE816AF950DA4E36DC" box="[561,622,1673,1697]" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="246">MLP</collectionCode>
).
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE81D8F951D9A236B8" collectingDate="1993-10-05" collectingDateMax="1993-10-08" collectingDateMin="1993-10-05" collectionCode="MMPE" collectorName="J. Farina" country="Argentina" county="Misiones Province" latitude="-33.8" location="Chimpay" longLatPrecision="119" longitude="-65.55" municipality="Chimpay" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Rio Negro Province">
<emphasis id="918A492ECD09FFFE81D8F951DB5336DC" bold="true" box="[643,883,1672,1697]" pageId="1" pageNumber="246">
<collectingRegion id="613A5BDECD09FFFE81D8F951DB4E36DC" box="[643,878,1672,1697]" country="Argentina" name="Rio Negro" pageId="1" pageNumber="246">Río Negro Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="43250F46CD09FFFE8025F950DBC036DC" box="[894,992,1673,1697]" pageId="1" pageNumber="246">Chimpay</collectingMunicipality>
[
<geoCoordinate id="C6CAF3FBCD09FFFE80A9F950DC4236DC" box="[1010,1122,1673,1697]" degrees="39" direction="south" minutes="10.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-39.166668">39°10.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE872AF950DCCC36DC" box="[1137,1260,1673,1697]" degrees="66" direction="west" minutes="09.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-66.15">66°09.0’W</geoCoordinate>
],
<date id="D740B3FCCD09FFFE8659F950DD5C36DC" box="[1282,1404,1673,1697]" pageId="1" pageNumber="246" value="1993-10-05" valueMax="1993-10-08" valueMin="1993-10-05">
<collectingDate id="C7044A14CD09FFFE8659F950DD5C36DC" box="[1282,1404,1673,1697]" pageId="1" pageNumber="246" value="1993-10-05" valueMax="1993-10-08" valueMin="1993-10-05">5–8.x.1993</collectingDate>
</date>
,
<collectorName id="0E0BF0EACD09FFFE86D0F950D8FE36B8" pageId="1" pageNumber="246">J. Farina</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE83B1F974D93336B9" box="[234,275,1709,1733]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE8279F974D95736B8" box="[290,375,1709,1733]" pageId="1" pageNumber="246">MMPE</collectionCode>
).
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE82D2F974DD1636B8" box="[393,1334,1709,1733]" collectingDate="1993-10-05" collectingDateMax="1993-10-08" collectingDateMin="1993-10-05" collectionCode="MLP, MMPE" collectorName="Wolffhugel &amp; J. Farina" country="Argentina" county="Misiones Province" latitude="-33.8" location="Las Isletas" longLatPrecision="119" longitude="-65.55" municipality="Chimpay" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="2" specimenCount-male="1" stateProvince="San Luis">
<emphasis id="918A492ECD09FFFE82D2F974D9D736B8" bold="true" box="[393,503,1709,1733]" pageId="1" pageNumber="246">
<collectingRegion id="613A5BDECD09FFFE82D2F974D9D236B8" box="[393,498,1709,1733]" country="Argentina" name="San Luis" pageId="1" pageNumber="246">San Luis</collectingRegion>
:
</emphasis>
<location id="A621C3E7CD09FFFE82A4F974DA5136B8" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE82A4F974DA5136B8" box="[511,625,1709,1733]" country="Argentina" county="Misiones Province" latitude="-33.8" longLatPrecision="119" longitude="-65.55" municipality="Chimpay" name="Las Isletas" pageId="1" pageNumber="246" stateProvince="San Luis">Las Isletas</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE81DBF974DACF36B8" box="[640,751,1709,1733]" degrees="33" direction="south" minutes="48.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-33.8">33°48.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE81A7F974DB5736B8" box="[764,887,1709,1733]" degrees="65" direction="west" minutes="33.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-65.55">65°33.0’W</geoCoordinate>
], [without date],
<specimenCount id="B5F85EB5CD09FFFE8774F974DC7836B9" box="[1071,1112,1709,1733]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
(MACN-En 40340)
</materialsCitation>
;
<materialsCitation id="13969F61CD09FFFE8619F974DAB93694" collectionCode="MLP" collectorName="Richter" country="Paraguay" latitude="-21.3" location="Puerto Guarani" longLatPrecision="126" longitude="-57.916668" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Alto Paraguay">
[without locality], [without date], M.
<collectorName id="0E0BF0EACD09FFFE8299F908DA323694" box="[450,530,1745,1769]" pageId="1" pageNumber="246">Richter</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE8147F908DA633695" box="[540,579,1745,1769]" count="1" pageId="1" pageNumber="246" type="male">1 ♂</specimenCount>
(
<collectionCode id="C5EF0DF9CD09FFFE810BF908DAAD3694" box="[592,653,1745,1769]" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="1" pageNumber="246">MLP</collectionCode>
).
</materialsCitation>
<materialsCitation id="13969F61CD09FFFE81C5F908DAD23770" collectionCode="MLP" collectorName="J. Daguerre" country="Paraguay" latitude="-21.3" location="Puerto Guarani" longLatPrecision="126" longitude="-57.916668" pageId="1" pageNumber="246" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Alto Paraguay Department">
<collectingCountry id="DBE9D5ACCD09FFFE81C5F908DB0F3694" box="[670,815,1745,1769]" name="Paraguay" pageId="1" pageNumber="246">PARAGUAY</collectingCountry>
:
<emphasis id="918A492ECD09FFFE8063F908DC553694" bold="true" box="[824,1141,1745,1769]" pageId="1" pageNumber="246">
<collectingRegion id="613A5BDECD09FFFE8063F908DC503694" box="[824,1136,1745,1769]" country="Paraguay" name="Alto Paraguay" pageId="1" pageNumber="246">Alto Paraguay Department</collectingRegion>
:
</emphasis>
<location id="A621C3E7CD09FFFE8721F908DD3B3694" LSID="urn:lsid:plazi:treatment:2B57242ACD09FFFB83CCFE69DCC2342A:A621C3E7CD09FFFE8721F908DD3B3694" box="[1146,1307,1745,1769]" country="Paraguay" latitude="-21.3" longLatPrecision="126" longitude="-57.916668" name="Puerto Guarani" pageId="1" pageNumber="246" stateProvince="Alto Paraguay Department">Puerto Guaraní</location>
[
<geoCoordinate id="C6CAF3FBCD09FFFE8673F908DDB83694" box="[1320,1432,1745,1769]" degrees="21" direction="south" minutes="18.0" orientation="latitude" pageId="1" pageNumber="246" precision="92" value="-21.3">21°18.0’S</geoCoordinate>
,
<geoCoordinate id="C6CAF3FBCD09FFFE83CCF92CD9323770" box="[151,274,1781,1805]" degrees="57" direction="west" minutes="55.0" orientation="longitude" pageId="1" pageNumber="246" precision="92" value="-57.916668">57°55.0’W</geoCoordinate>
], 1920,
<collectorName id="0E0BF0EACD09FFFE823DF92CD9C43770" box="[358,484,1781,1805]" pageId="1" pageNumber="246">J. Daguerre</collectorName>
,
<specimenCount id="B5F85EB5CD09FFFE82B5F92CDA363771" box="[494,534,1781,1805]" count="1" pageId="1" pageNumber="246" type="female">1 ♀</specimenCount>
(MACN-En 40341)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EBE4C6B7CD09FFFD839CF8C3DC8E3166" lastPageId="2" lastPageNumber="247" pageId="1" pageNumber="246" type="diagnosis">
<paragraph id="A341953CCD09FFFD839CF8C3DC8E3166" blockId="1.[151,1437,665,2021]" lastBlockId="2.[151,1437,151,283]" lastPageId="2" lastPageNumber="247" pageId="1" pageNumber="246">
<emphasis id="918A492ECD09FFFE839CF8C3D91C374C" box="[199,316,1818,1841]" italics="true" pageId="1" pageNumber="246">Diagnosis:</emphasis>
Length:
<quantity id="640638D9CD09FFFE82C1F8C0DA13374D" box="[410,563,1817,1841]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.25" metricValueMin="1.65" pageId="1" pageNumber="246" unit="mm" value="19.5" valueMax="22.5" valueMin="16.5">16.5‒22.5 mm</quantity>
(females,
<emphasis id="918A492ECD09FFFE81FBF8C3DA8D374C" box="[672,685,1818,1841]" italics="true" pageId="1" pageNumber="246">n</emphasis>
= 22),
<quantity id="640638D9CD09FFFE81ADF8C0DBAF374D" box="[758,911,1817,1841]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.05" metricValueMin="1.45" pageId="1" pageNumber="246" unit="mm" value="17.5" valueMax="20.5" valueMin="14.5">14.5‒20.5 mm</quantity>
(males,
<emphasis id="918A492ECD09FFFE80B3F8C3DBD5374C" box="[1000,1013,1818,1841]" italics="true" pageId="1" pageNumber="246">n</emphasis>
= 22); greatest pronotal width:
<quantity id="640638D9CD09FFFE8664F8C0D8E03729" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.5" metricValueMin="7.5" pageId="1" pageNumber="246" unit="mm" value="9.0" valueMax="10.5" valueMin="7.5">7.5‒10.5 mm</quantity>
(females,
<emphasis id="918A492ECD09FFFE826AF8E7D91E3728" box="[305,318,1854,1877]" italics="true" pageId="1" pageNumber="246">n</emphasis>
= 22), 7.5‒11.0 mm (males,
<emphasis id="918A492ECD09FFFE8128F8E7DAA03728" box="[627,640,1854,1877]" italics="true" pageId="1" pageNumber="246">n</emphasis>
= 22). Colour reddish brown, shiny. Body oval and convex, dorsal surface glabrous, ventrally setose. Head triangular, ending in a rounded tip. In females, with a conical, weak tubercle; in males, frons armed with a small, erect horn. Mentum strongly convex. Antennae with nine antennomeres. Pronotum in females, entirely convex, finely rugo-punctate on anterior portion of disc, appearing smooth on middle and posterior portions, without declivity. In males, globose, with a vertical declivity in the anterior third. Scutellum broad, approximately onequarter wider than long, punctate. Elytron shiny, surface with weak punctate striae. Epipleuron entire, more pronounced on the anterior third. Protibiae with three well-defined teeth, equidistant, with black margins. Meso- and metatibiae with two transverse carinae in middle and apex with two pointed spurs on mesotibiae and rounded spatulate shape on metatibiae. Pygidium mostly smooth, with some small setigerous punctures scattered on the sides.
</paragraph>
</subSubSection>
<caption id="F781C5B4CD0AFFFD83CCF8C3DB183707" ID-DOI="http://doi.org/10.5281/zenodo.14240866" ID-Zenodo-Dep="14240866" httpUri="https://zenodo.org/record/14240866/files/figure.png" pageId="2" pageNumber="247" startId="2.[151,250,1818,1842]" targetBox="[172,1414,320,1781]" targetPageId="2" targetType="figure">
<paragraph id="A341953CCD0AFFFD83CCF8C3DB183707" blockId="2.[151,1436,1818,1914]" pageId="2" pageNumber="247">
<emphasis id="918A492ECD0AFFFD83CCF8C3D932374F" bold="true" box="[151,274,1818,1842]" pageId="2" pageNumber="247">FIGURE 1.</emphasis>
Records of
<taxonomicName id="64FEEEBFCD0AFFFD82D2F8C3DAA5374F" baseAuthorityName="Steinheil" baseAuthorityYear="1874" box="[393,645,1818,1842]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="247" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="918A492ECD0AFFFD82D2F8C3DAA5374F" box="[393,645,1818,1842]" italics="true" pageId="2" pageNumber="247">Eremobothynus cornutus</emphasis>
</taxonomicName>
. Green star: type locality of
<taxonomicName id="64FEEEBFCD0AFFFD80C5F8C3DCBA374F" baseAuthorityName="Steinheil" baseAuthorityYear="1874" box="[926,1178,1818,1842]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="247" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="918A492ECD0AFFFD80C5F8C3DCBA374F" box="[926,1178,1818,1842]" italics="true" pageId="2" pageNumber="247">Eremobothynus cornutus</emphasis>
</taxonomicName>
, yellow star: type locality of
<taxonomicName id="64FEEEBFCD0AFFFD83EEF8E7D95E372B" box="[181,382,1854,1878]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="247" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0AFFFD83EEF8E7D95E372B" box="[181,382,1854,1878]" italics="true" pageId="2" pageNumber="247">Aphonides turbator</emphasis>
</taxonomicName>
, solid red circles: material examined with locality data, open red circles: material examined without precise locality data, black circles: previous bibliographic records.
</paragraph>
</caption>
<caption id="F781C5B4CD0BFFFC83CCF96DDC8037FD" ID-DOI="http://doi.org/10.5281/zenodo.14240868" ID-Zenodo-Dep="14240868" httpUri="https://zenodo.org/record/14240868/files/figure.png" pageId="3" pageNumber="248" startId="3.[151,264,1716,1740]" targetBox="[180,1415,194,1676]" targetPageId="3" targetType="figure">
<paragraph id="A341953CCD0BFFFC83CCF96DDC8037FD" blockId="3.[151,1436,1716,1921]" pageId="3" pageNumber="248">
<emphasis id="918A492ECD0BFFFC83CCF96DD96636B1" bold="true" box="[151,326,1716,1740]" pageId="3" pageNumber="248">FIGURES 2‒12.</emphasis>
<taxonomicName id="64FEEEBFCD0BFFFC8211F96CDA6636B1" baseAuthorityName="Steinheil" baseAuthorityYear="1874" box="[330,582,1717,1741]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="918A492ECD0BFFFC8211F96CDA6636B1" box="[330,582,1717,1741]" italics="true" pageId="3" pageNumber="248">Eremobothynus cornutus</emphasis>
</taxonomicName>
(female specimens).
<emphasis id="918A492ECD0BFFFC8041F96DDB0C36B1" bold="true" box="[794,812,1716,1740]" pageId="3" pageNumber="248">2,</emphasis>
dorsal view of
<taxonomicName id="64FEEEBFCD0BFFFC809FF96CDCA836B0" box="[964,1160,1717,1741]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0BFFFC809FF96CDCA836B0" box="[964,1160,1717,1741]" italics="true" pageId="3" pageNumber="248">Aphonides turbator</emphasis>
</taxonomicName>
holotype (ZMB);
<emphasis id="918A492ECD0BFFFC8665F96DDD7136B1" bold="true" box="[1342,1361,1716,1740]" pageId="3" pageNumber="248">3,</emphasis>
ventral view of
<taxonomicName id="64FEEEBFCD0BFFFC83B5F900D995368C" box="[238,437,1753,1777]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0BFFFC83B5F900D995368C" box="[238,437,1753,1777]" italics="true" pageId="3" pageNumber="248">Aphonides turbator</emphasis>
</taxonomicName>
holotype (ZMB);
<emphasis id="918A492ECD0BFFFC812EF901DAA8368D" bold="true" box="[629,648,1752,1776]" pageId="3" pageNumber="248">4,</emphasis>
lateral view of
<taxonomicName id="64FEEEBFCD0BFFFC8075F900DBD5368C" box="[814,1013,1753,1777]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0BFFFC8075F900DBD5368C" box="[814,1013,1753,1777]" italics="true" pageId="3" pageNumber="248">Aphonides turbator</emphasis>
</taxonomicName>
holotype (ZMB);
<emphasis id="918A492ECD0BFFFC87EDF900DCE9368C" bold="true" box="[1206,1225,1753,1777]" pageId="3" pageNumber="248">5,</emphasis>
labels of
<taxonomicName id="64FEEEBFCD0BFFFC866AF900D8CB3768" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0BFFFC866AF900D8CB3768" italics="true" pageId="3" pageNumber="248">Aphonides turbator</emphasis>
</taxonomicName>
holotype;
<emphasis id="918A492ECD0BFFFC8204F925D9533769" bold="true" box="[351,371,1788,1812]" pageId="3" pageNumber="248">6,</emphasis>
dorsal view of
<taxonomicName id="64FEEEBFCD0BFFFC8145F924DAC73768" box="[542,743,1789,1813]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0BFFFC8145F924DAC73768" box="[542,743,1789,1813]" italics="true" pageId="3" pageNumber="248">Aphonides turbator</emphasis>
</taxonomicName>
paratype (ZMB);
<emphasis id="918A492ECD0BFFFC80F0F924DB9D3768" bold="true" box="[939,957,1789,1813]" pageId="3" pageNumber="248">7,</emphasis>
ventral view of
<taxonomicName id="64FEEEBFCD0BFFFC8729F924DD1B3768" box="[1138,1339,1789,1813]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0BFFFC8729F924DD1B3768" box="[1138,1339,1789,1813]" italics="true" pageId="3" pageNumber="248">Aphonides turbator</emphasis>
</taxonomicName>
paratype (ZMB);
<emphasis id="918A492ECD0BFFFC83B5F8F9D9223745" bold="true" box="[238,258,1824,1848]" pageId="3" pageNumber="248">8,</emphasis>
lateral view of
<taxonomicName id="64FEEEBFCD0BFFFC82F3F8F8DA4F3744" box="[424,623,1825,1849]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0BFFFC82F3F8F8DA4F3744" box="[424,623,1825,1849]" italics="true" pageId="3" pageNumber="248">Aphonides turbator</emphasis>
</taxonomicName>
paratype (ZMB);
<emphasis id="918A492ECD0BFFFC8075F8F9DB613745" bold="true" box="[814,833,1824,1848]" pageId="3" pageNumber="248">9,</emphasis>
labels of
<taxonomicName id="64FEEEBFCD0BFFFC80F1F8F8DC513744" box="[938,1137,1825,1849]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="248" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0BFFFC80F1F8F8DC513744" box="[938,1137,1825,1849]" italics="true" pageId="3" pageNumber="248">Aphonides turbator</emphasis>
</taxonomicName>
paratype;
<emphasis id="918A492ECD0BFFFC87BBF8F9DCDF3745" bold="true" box="[1248,1279,1824,1848]" pageId="3" pageNumber="248">10,</emphasis>
dorsal view of a specimen from Córdoba Province, Argentina (LEBA);
<emphasis id="918A492ECD0BFFFC81B3F89DDB263721" bold="true" box="[744,774,1860,1884]" pageId="3" pageNumber="248">11,</emphasis>
ventral view of a specimen from Córdoba Province, Argentina (LEBA);
<emphasis id="918A492ECD0BFFFC83A3F8B1D93737FD" bold="true" box="[248,279,1896,1920]" pageId="3" pageNumber="248">12,</emphasis>
lateral view of a specimen from Córdoba Province, Argentina (LEBA). Scale bar: 5 mm.
</paragraph>
</caption>
<subSubSection id="EBE4C6B7CD0CFFFB839CFF4EDCC2342A" pageId="4" pageNumber="249" type="discussion">
<paragraph id="A341953CCD0CFFFB839CFF4EDD26308A" blockId="4.[151,1437,151,1111]" pageId="4" pageNumber="249">
<emphasis id="918A492ECD0CFFFB839CFF4ED90E30D2" box="[199,302,151,175]" italics="true" pageId="4" pageNumber="249">Remarks:</emphasis>
<bibRefCitation id="C76FE8CDCD0CFFFB826FFF4ED9F830CD" author="Endrodi, S." box="[308,472,151,176]" pageId="4" pageNumber="249" pagination="197 - 202" refId="ref3414" refString="Endrodi, S. (1981) Neue und seltene Dynastinen aus Sudamerika und eine synonymische Bemerkung (Coleoptera, Melolonthidae). Annales Historico-Naturales Musei Nationalis Hungarici, 73, 197 - 202." type="journal article" year="1981">Endrödi (1981)</bibRefCitation>
differentiated
<taxonomicName id="64FEEEBFCD0CFFFB812FFF4EDB6430D2" box="[628,836,151,175]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0CFFFB812FFF4EDB6430D2" box="[628,836,151,175]" italics="true" pageId="4" pageNumber="249">Aphonides turbator</emphasis>
</taxonomicName>
from
<taxonomicName id="64FEEEBFCD0CFFFB80DEFF4EDCB130D2" authorityName="Rivers" authorityYear="1888" box="[901,1169,151,175]" class="Insecta" family="Scarabaeidae" genus="Anoplognatho" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="dunnianus">
<emphasis id="918A492ECD0CFFFB80DEFF4EDCB130D2" box="[901,1169,151,175]" italics="true" pageId="4" pageNumber="249">Anoplognatho dunnianus</emphasis>
</taxonomicName>
(the
<typeStatus id="7C452B9ECD0CFFFB8793FF41DCD630CD" box="[1224,1270,152,176]" pageId="4" pageNumber="249">type</typeStatus>
species of both generic names) by the presence of a conical, weak tubercle on the head. In this study, all examined female specimens of
<taxonomicName id="64FEEEBFCD0CFFFB83CCFF39D932308A" baseAuthorityName="Steinheil" baseAuthorityYear="1874" box="[151,274,224,247]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="918A492ECD0CFFFB83CCFF39D932308A" box="[151,274,224,247]" italics="true" pageId="4" pageNumber="249">E. cornutus</emphasis>
</taxonomicName>
, as well as the
<typeStatus id="7C452B9ECD0CFFFB82EEFF39D9C33085" box="[437,483,224,248]" pageId="4" pageNumber="249">type</typeStatus>
material of
<taxonomicName id="64FEEEBFCD0CFFFB813EFF06DB15308A" box="[613,821,223,247]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0CFFFB813EFF06DB15308A" box="[613,821,223,247]" italics="true" pageId="4" pageNumber="249">Aphonides turbator</emphasis>
</taxonomicName>
, have a conical, weak tubercle on the head.
</paragraph>
<paragraph id="A341953CCD0CFFFB839CFEDADB963222" blockId="4.[151,1437,151,1111]" pageId="4" pageNumber="249">
The type material of
<taxonomicName id="64FEEEBFCD0CFFFB82EBFEDADAA23166" box="[432,642,259,283]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0CFFFB82EBFEDADAA23166" box="[432,642,259,283]" italics="true" pageId="4" pageNumber="249">Aphonides turbator</emphasis>
</taxonomicName>
differs from the other two valid species of
<taxonomicName id="64FEEEBFCD0CFFFB8738FEDADD293166" authorityName="Ohaus" authorityYear="1910" box="[1123,1289,259,283]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="918A492ECD0CFFFB8738FEDADD293166" box="[1123,1289,259,283]" italics="true" pageId="4" pageNumber="249">Eremobothynus</emphasis>
</taxonomicName>
based on the variation in the morphological characters outlined below.
<bibRefCitation id="C76FE8CDCD0CFFFB8048FEFEDB9A313D" author="Endrodi, S." box="[787,954,295,320]" pageId="4" pageNumber="249" pagination="105 - 108" refId="ref3382" refString="Endrodi, S. (1970) Drei neue Dynastinen (Coleoptera, Melolonthidae) aus Amerika. Mitteilungen aus dem Zoologischen Museum in Berlin, 46, 105 - 108." type="journal article" year="1970">Endrödi (1970)</bibRefCitation>
identified a double tubercle on the front as the main diagnostic character for distinguishing
<taxonomicName id="64FEEEBFCD0CFFFB81C5FE95DB33311E" authorityName="Endrodi" authorityYear="1970" box="[670,787,331,355]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="918A492ECD0CFFFB81C5FE95DB33311E" box="[670,787,331,355]" italics="true" pageId="4" pageNumber="249">E. bicuspis</emphasis>
</taxonomicName>
from
<taxonomicName id="64FEEEBFCD0CFFFB800EFE95DBF1311E" baseAuthorityName="Steinheil" baseAuthorityYear="1874" box="[853,977,332,355]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="918A492ECD0CFFFB800EFE95DBF1311E" box="[853,977,332,355]" italics="true" pageId="4" pageNumber="249">E. cornutus</emphasis>
</taxonomicName>
. This same tubercle is characteristic of the females of
<taxonomicName id="64FEEEBFCD0CFFFB8249FEA9D9A931FA" authorityName="Endrodi" authorityYear="1970" box="[274,393,367,391]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="918A492ECD0CFFFB8249FEA9D9A931FA" box="[274,393,367,391]" italics="true" pageId="4" pageNumber="249">E. bicuspis</emphasis>
</taxonomicName>
, allowing them to be differentiated from the type material of
<taxonomicName id="64FEEEBFCD0CFFFB876AFEB6DD2331FA" box="[1073,1283,367,391]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0CFFFB876AFEB6DD2331FA" box="[1073,1283,367,391]" italics="true" pageId="4" pageNumber="249">Aphonides turbator</emphasis>
</taxonomicName>
. On the other hand,
<bibRefCitation id="C76FE8CDCD0CFFFB8383FE4AD9F731D1" author="Grossi, P. C. &amp; Abadie, E. I." box="[216,471,403,428]" pageId="4" pageNumber="249" pagination="57 - 60" refId="ref3475" refString="Grossi, P. C. &amp; Abadie, E. I. (2008) A new species of Eremobothynus Ohaus (Coleoptera: Scarabaeidae: Dynastinae) from Argentina, with notes on Eremobothynus cornutus (Steinheil). Zootaxa, 1733 (1), 57 - 60. https: // doi. org / 10.11646 / zootaxa. 1733.1.3" type="journal article" year="2008">Grossi &amp; Abadie (2008)</bibRefCitation>
described
<taxonomicName id="64FEEEBFCD0CFFFB8113FE4DDA9B31D6" authorityName="Grossi &amp; Abadie" authorityYear="2008" box="[584,699,403,427]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="918A492ECD0CFFFB8113FE4DDA9B31D6" box="[584,699,403,427]" italics="true" pageId="4" pageNumber="249">E. hirsutus</emphasis>
</taxonomicName>
based solely on
<specimenCount id="B5F85EB5CD0CFFFB8032FE4ADBF931D6" box="[873,985,403,427]" count="3" pageId="4" pageNumber="249" type="male">three male</specimenCount>
specimens. The diagnostic characters they mentioned include a short head with a square clypeus, a flattened mentum, and an almost absent epipleuron. However, these traits do not align with those observed in the
<typeStatus id="7C452B9ECD0CFFFB81E7FE02DB39318E" box="[700,793,475,499]" pageId="4" pageNumber="249" type="holotype">holotype</typeStatus>
of
<taxonomicName id="64FEEEBFCD0CFFFB8065FE02DC2F318E" box="[830,1039,475,499]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0CFFFB8065FE02DC2F318E" box="[830,1039,475,499]" italics="true" pageId="4" pageNumber="249">Aphonides turbator</emphasis>
</taxonomicName>
, which features a longer head with a rounded clypeus, a strongly convex mentum, and an entire epipleuron more pronounced on the anterior third. Although the female of
<taxonomicName id="64FEEEBFCD0CFFFB826BFDFDD9863246" authorityName="Grossi &amp; Abadie" authorityYear="2008" box="[304,422,547,571]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="918A492ECD0CFFFB826BFDFDD9863246" box="[304,422,547,571]" italics="true" pageId="4" pageNumber="249">E. hirsutus</emphasis>
</taxonomicName>
has not been documented, the lack of sexual dimorphism in the mentioned characters within
<taxonomicName id="64FEEEBFCD0CFFFB83CCFD9ED93F3222" authorityName="Mulsant" authorityYear="1842" box="[151,287,583,607]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="tribe" tribe="Pentodontini">Pentodontini</taxonomicName>
suggests that these traits may be consistent across both sexes.
</paragraph>
<paragraph id="A341953CCD0CFFFB839CFDB5DAA93396" blockId="4.[151,1437,151,1111]" pageId="4" pageNumber="249">
In this contribution,
<taxonomicName id="64FEEEBFCD0CFFFB82C6FDB2DA4C32FE" box="[413,620,619,643]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0CFFFB82C6FDB2DA4C32FE" box="[413,620,619,643]" italics="true" pageId="4" pageNumber="249">Aphonides turbator</emphasis>
</taxonomicName>
is proposed as a junior synonym of
<taxonomicName id="64FEEEBFCD0CFFFB80BFFDB2DCCD32FE" baseAuthorityName="Steinheil" baseAuthorityYear="1874" box="[996,1261,619,643]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="918A492ECD0CFFFB80BFFDB2DCCD32FE" box="[996,1261,619,643]" italics="true" pageId="4" pageNumber="249">Eremobothynus cornutus</emphasis>
</taxonomicName>
. This synonymy is established based on the analysis of the original descriptions of genera and species (
<bibRefCitation id="C76FE8CDCD0CFFFB8704FD56DD2332D5" author="Steinheil, E." box="[1119,1283,655,680]" pageId="4" pageNumber="249" pagination="554 - 578" refId="ref3840" refString="Steinheil, E. (1874) Symbolae ad historiam Coleopterorum Argentinae meridionalis, ossia enumerazione dei coleotteri raccolti dal Prof. Strobel, durante il suo soggiorno in Buenos Aires e nei viaggi de la intrapresi a Mendoza e nal Chili, indi a Bahia Blanca ed al Carmen de los Patagones; e descrizione delle specie nuove. Atti Della Societa Italiana di Scienze Naturali, 15, 554 - 578. [Dated 1872]." type="journal article" year="1874">Steinheil 1874</bibRefCitation>
;
<bibRefCitation id="C76FE8CDCD0CFFFB864AFD56DDB832D5" author="Ohaus, F." box="[1297,1432,655,680]" pageId="4" pageNumber="249" pagination="173 - 186" refId="ref3669" refString="Ohaus, F. (1910) Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift, 1910, 173 - 186." type="journal article" year="1910">Ohaus 1910</bibRefCitation>
;
<bibRefCitation id="C76FE8CDCD0CFFFB83CCFD6AD90D32B1" author="Endrodi, S." box="[151,301,691,716]" pageId="4" pageNumber="249" pagination="197 - 202" refId="ref3414" refString="Endrodi, S. (1981) Neue und seltene Dynastinen aus Sudamerika und eine synonymische Bemerkung (Coleoptera, Melolonthidae). Annales Historico-Naturales Musei Nationalis Hungarici, 73, 197 - 202." type="journal article" year="1981">Endrödi 1981</bibRefCitation>
) and the observation of the
<typeStatus id="7C452B9ECD0CFFFB8131FD6DDAB832B1" box="[618,664,692,716]" pageId="4" pageNumber="249">type</typeStatus>
specimens of
<taxonomicName id="64FEEEBFCD0CFFFB8061FD6ADC2E32B6" box="[826,1038,691,715]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0CFFFB8061FD6ADC2E32B6" box="[826,1038,691,715]" italics="true" pageId="4" pageNumber="249">Aphonides turbator</emphasis>
</taxonomicName>
and specimens of
<taxonomicName id="64FEEEBFCD0CFFFB87BAFD6DDD7E32B6" baseAuthorityName="Steinheil" baseAuthorityYear="1874" box="[1249,1374,692,715]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="918A492ECD0CFFFB87BAFD6DDD7E32B6" box="[1249,1374,692,715]" italics="true" pageId="4" pageNumber="249">E. cornutus</emphasis>
</taxonomicName>
from various localities. The morphological characters that support the synonymy are: (1) oval and convex body; (2) shiny reddish-brown color; (3) head front armed with a weak tubercle (females) or a small erect horn (males); (4) triangular clypeus ending in a rounded point; (5) mentum strongly convex; (6) antennae with nine antennomeres; (7) pronotum notably curved, wider than long, with rounded posterior corners and more noticeable punctation on the anterior half; (8) broad triangular scutellum; (9) convex elytra finely punctate, with entire epipleuron; (10) protibiae with three strong teeth; (11) mesotibiae and metatibiae with two oblique carinae; (12) convex, smooth, and shiny pygidium, with isolated punctures and a higher density of punctures in the lateral corners. In all these characters, the
<typeStatus id="7C452B9ECD0CFFFB8720FC69DC8933B5" box="[1147,1193,944,968]" pageId="4" pageNumber="249">type</typeStatus>
material of
<taxonomicName id="64FEEEBFCD0CFFFB8677FC76D8D03396" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0CFFFB8677FC76D8D03396" italics="true" pageId="4" pageNumber="249">Aphonides turbator</emphasis>
</taxonomicName>
is indistinguishable from
<taxonomicName id="64FEEEBFCD0CFFFB815CFC0DDAA13396" baseAuthorityName="Steinheil" baseAuthorityYear="1874" box="[519,641,980,1003]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="918A492ECD0CFFFB815CFC0DDAA13396" box="[519,641,980,1003]" italics="true" pageId="4" pageNumber="249">E. cornutus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A341953CCD0CFFFB839CFC21DCC2342A" blockId="4.[151,1437,151,1111]" pageId="4" pageNumber="249">
In conclusion, the results of this study provide a solid basis for the synonymy of
<taxonomicName id="64FEEEBFCD0CFFFB8749FC2EDCC13472" box="[1042,1249,1015,1039]" class="Insecta" family="Scarabaeidae" genus="Aphonides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="turbator">
<emphasis id="918A492ECD0CFFFB8749FC2EDCC13472" box="[1042,1249,1015,1039]" italics="true" pageId="4" pageNumber="249">Aphonides turbator</emphasis>
</taxonomicName>
with
<taxonomicName id="64FEEEBFCD0CFFFB8647FC21DDB53472" baseAuthorityName="Steinheil" baseAuthorityYear="1874" box="[1308,1429,1016,1039]" class="Insecta" family="Scarabaeidae" genus="Eremobothynus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="918A492ECD0CFFFB8647FC21DDB53472" box="[1308,1429,1016,1039]" italics="true" pageId="4" pageNumber="249">E. cornutus</emphasis>
</taxonomicName>
, supported by a detailed evaluation of multiple morphological characters. The integration of original descriptions and observations of specimens from various localities has facilitated a clearer understanding of the species.
</paragraph>
</subSubSection>
</treatment>
</document>