From 695e887ac2755a437595da058c54dbf2ac3f4a5f Mon Sep 17 00:00:00 2001
From: ggserver <ggserver@srv1.plazi.org>
Date: Wed, 25 Sep 2024 17:22:17 +0000
Subject: [PATCH] Add updates up until 2024-09-25 17:16:13

---
 .../CB/29DCCB12F4F85E91AFC577CE36A02436.xml   | 1330 +++++++++++++++++
 1 file changed, 1330 insertions(+)
 create mode 100644 data/29/DC/CB/29DCCB12F4F85E91AFC577CE36A02436.xml

diff --git a/data/29/DC/CB/29DCCB12F4F85E91AFC577CE36A02436.xml b/data/29/DC/CB/29DCCB12F4F85E91AFC577CE36A02436.xml
new file mode 100644
index 00000000000..0c4a4f32482
--- /dev/null
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@@ -0,0 +1,1330 @@
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+<mods:mods id="CE4F6595CA3CB8FF2AEB10118492DF42" xmlns:mods="http://www.loc.gov/mods/v3">
+<mods:titleInfo id="C2BF449FF45F5116A0C8745D2A9FE9D1">
+<mods:title id="49C849AFEF4570E014F63E6D2B286D93">Integrative taxonomy revealed a new species of Lefua (Teleostei, Nemacheilidae) from Fukui Prefecture, Japan</mods:title>
+</mods:titleInfo>
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+<mods:namePart id="A0AD816D5878C83D7B70841D9976DCBC">Katayama, Yuta</mods:namePart>
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+<mods:affiliation id="56E71B36AAD9D4DDBCB7B1D01D544D85">Department of Zoology, Graduate School of Science, Kyoto University, Kitashirakawa-oiwakecho, Sakyo-ku, Kyoto-shi, Kyoto, 606 - 8502 Japan &amp; Department of Biological Sciences, School of Science, The University of Tokyo, 7 - 3 - 1, Hongo, Bunkyo-ku, Tokyo 113 - 0033, Japan</mods:affiliation>
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+<mods:date id="4B4B81A08E7534C3DA7C5ED9D0284939">2024</mods:date>
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+<subSubSection id="8EE9A0F6BF643335B1F2046750A5941B" type="nomenclature">
+<paragraph id="903CEE2629808E840A04F6CF230341E4">
+<taxonomicName id="F67489EF20C0D6953066769EACC1537E" authority="Katayama" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai" status="sp. nov.">
+<emphasis id="A107B9BAB28B20B0F1B62781968AD835" italics="true">Lefua nishimurai</emphasis>
+Katayama
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+<taxonomicNameLabel id="FE9B7E8F137CD2CBC8EEE5F5A296CAA8" rank="species">sp. nov.</taxonomicNameLabel>
+</paragraph>
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+<subSubSection id="1EFD8B89D5BC21717B93D63553F89E04" type="description">
+<paragraph id="0509E8F2AE8499FE32B553D161688D03">
+<tableCitation id="4826AC3B9CFF07D6DAA0C52D2157959D" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Meristics and morphometrics of Lefua torrentis, L. tokaiensis., and L. nishimurai sp. nov. Measurements indicate minimum – maximum values (mean ± SD). Data of the holotypes of L. torrentis and L. tokaiensis were obtained from Hosoya et al. (2018) and Ito et al. (2019), respectively." httpUri="http://table.plazi.org/id/B7857D6A081FBE45190B6B271D4B321C" tableUuid="B7857D6A081FBE45190B6B271D4B321C">Table 4</tableCitation>
+, 
+<figureCitation id="67F597C8DE06BFA9FD04AE2508B0956D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body morphology and coloration of three Lefua species. A. Kii-Shikoku Population of L. torrentis (FAKU 211486); B. Sanyo Population of L. torrentis (FAKU 211492); C. Nihonkai Population of L. torrentis (FAKU 211504); D. L. tokaiensis (FAKU 211510); E. L. nishimurai sp. nov. (LBM 1210059189). Scale bar: 10 mm." httpUri="https://binary.pensoft.net/fig/1141333">Figs 5 E</figureCitation>
+, 
+<figureCitation id="556DD3AB454373257DF594261C09F929" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of dorsal view of the head of three Lefua species. A. Kii-Shikoku Population of L. torrentis (FAKU 211486); B. Sanyo Population of L. torrentis (FAKU 211492); C. Nihonkai Population of L. torrentis (FAKU 211504); D L. tokaiensis (FAKU 211510); E. L. nishimurai sp. nov. (LBM 1210059179). Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1141334">6 E</figureCitation>
+, 
+<figureCitation id="561AD482871B0C8272581828116389D2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Body coloration of the holotype of Lefua nishimurai sp. nov (LBM 1210059189). A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm." httpUri="https://binary.pensoft.net/fig/1141335">7</figureCitation>
+, 
+<figureCitation id="A744323CE0C2F95FB85D33494531DF1E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Detailed morphology of the holotype of Lefua nishimurai sp. nov (LBM 1210059189). A. Head; B. Lateral view of head; C. Mouth; D. Caudal fin base. An arrow in the lateral view of head (B) indicates narrow conspicuous black stripe between base of outer rostral barbel and eye. Arrows in the caudal fin base (D) indicate black spots on ventral and dorsal surfaces. Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1141336">8</figureCitation>
+, 
+<figureCitation id="3A0AA1B20B98AFE1C566E1083E66C981" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Body coloration of atypical specimen of Lefua nishimurai sp. nov. (LBM 1210059188), showing no distinct dark brown spots scattered over the body and obscure dark band in the skin of the caudal fin base. Scale bar: 10 mm." httpUri="https://binary.pensoft.net/fig/1141337">9 New Standard Japanese name: Reihoku-nagare-hotoke-dojyô</figureCitation>
+</paragraph>
+</subSubSection>
+<subSubSection id="E89A79B6AB5BD7D7CB75A414E11DB2AA" type="reference_group">
+<paragraph id="DAD570FA44AF6E09347734A6607D7C35">
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+<treatmentCitation id="84406B2D377B80A175B2DC6664458ACC">
+<taxonomicName id="EEA0DA1277F4BCF3E7EAE418A0C00B81" authorityName=" Hosoya, Ito &amp; Miyazaki" authorityYear="2018" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">
+<emphasis id="E8C75DD61004065F077A213933E39C7D" italics="true">Lefua torrentis</emphasis>
+</taxonomicName>
+<comment id="C61812EE5C7A4600D5EEA087AACC039C">
+: 
+<treatmentCitation id="C19CA2C0DDBC73179FF6D7C1EDCE9A0E" author="" page="110" year="2016">
+<bibRefCitation id="B5B3BC74F5DB752A9CF3DD53B8C2E46A" journalOrPublisher="Fukui Prefecture, Fukui" refId="B37" refString="Natural Environment Division, Department of Safety and Environment, Fukui Prefecture (2016) Threatened Wildlife of Fukui Pref. Fukui Red Data Book. Revised Edition. Fukui Prefecture, Fukui, 497 pp." title="Threatened Wildlife of Fukui Pref. Fukui Red Data Book. Revised Edition." year="2016">
+Natural Environment Division, Department of Safety and Environment, 
+<collectingRegion id="64B7D6019BAACA9C36E0104D13F1B506" country="Japan" name="Fukui">Fukui Prefecture</collectingRegion>
+2016
+</bibRefCitation>
+: 110
+</treatmentCitation>
+(part); 
+<treatmentCitation id="EC72FE06DBA6B9B7C9E47BC0A46D4359" author="Nakajima J &amp; Nishimura T &amp; Ito T &amp; Miyazaki J &amp; Ooi K &amp; Hirakawa S" page="35" year="2021">
+<bibRefCitation id="FDC6F149663B70DEC15872EF62AC7282" DOI="10.34583/ichthy.6.0_33" author="Nakajima J &amp; Nishimura T &amp; Ito T &amp; Miyazaki J &amp; Ooi K &amp; Hirakawa S" etAl="et al." firstAuthor="Nakajima" journalOrPublisher="Ichthy, Natural History of Japan" page="35" pagination="33-37" refId="B35" refString="Nakajima J, Nishimura T, Ito T, Miyazaki J, Ooi K, Hirakawa S (2021) Report of a genetic population of Lefua torrentis (Nemacheilidae) from Reihoku District, Fukui Prefecture, Honshu, Japan. Ichthy, Natural History of Japan 6: 33–37. https://doi.org/10.34583/ichthy.6.0_33" title="Report of a genetic population of Lefua torrentis (Nemacheilidae) from Reihoku District, Fukui Prefecture, Honshu, Japan." volume="6" year="2021">Nakajima et al. 2021: 35</bibRefCitation>
+</treatmentCitation>
+, fig. 1; 
+<treatmentCitation id="ACD509973BC4D24455E51A515F08F64A" author="Katayama Y" page="54" year="2021">
+<bibRefCitation id="C5BB63994C0B22FA152C71351AD50F64" author="Katayama Y" firstAuthor="Katayama" journalOrPublisher="Bulletin of the Fukui City Museum of Natural History" page="54" pagination="53-56" refId="B20" refString="Katayama Y (2021) A survey of the distribution of the Reihoku population of the Lefua torrentis (Nemacheilidae) with a first record in Fukui city. Bulletin of the Fukui City Museum of Natural History 68: 53–56." title="A survey of the distribution of the Reihoku population of the Lefua torrentis (Nemacheilidae) with a first record in Fukui city." volume="68" year="2021">Katayama 2021: 54</bibRefCitation>
+</treatmentCitation>
+, fig. 5.
+</comment>
+</treatmentCitation>
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+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EJDAI" type="material">
+<paragraph id="3C36EFFCB32754DD4B27519292B36014">
+<heading id="C4163C5C3FF5FE06A6B7FE6AED2D8D2B" reason="title">Material examined.</heading>
+</paragraph>
+<paragraph id="4AB6098E327E6F494D50C48181D1CE6D">
+<materialsCitation id="856289BC412E233DC6114AB8AAD935ED" collectingDate="2021-09-06" collectionCode="LBM" collectorName="Yuta Katayama" country="Japan" county="Fukui" latitude="36.0621" location="Kuzuryu River system" longLatPrecision="6" longitude="136.1304" specimenCode="LBM 1210059189" specimenCount="1" stateProvince="Fukui Prefecture" typeStatus="holotype">
+<emphasis id="EB0F3479666015E6A5F35D7069DF1A38" bold="true">
+<emphasis id="8410A78C66B1FE74EB8C14F206B8B3B3" bold="true" italics="true">
+<typeStatus id="465C7261156DA0C8F76916BF2DD076DD" type="holotype">Holotype</typeStatus>
+</emphasis>
+.
+</emphasis>
+• 
+<specimenCode id="00001EE2217CDFA44EB8B58A0920A426" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" type="">
+<named-content id="E753AF1F919A250F0BF95BF7EF5A1794" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/lake-biwa-museum" xlink_title="Lake Biwa Museum">LBM</named-content>
+1210059189
+</specimenCode>
+; 
+<quantity id="CAB7E0380B0CDA4FD5FAD1377C447878" metricMagnitude="-2" metricUnit="m" metricValue="5.12" unit="mm" value="51.2">51.2 mm</quantity>
+SL; collected from 
+<location id="F6CA2DFAF25A04A47BD60EAF44447DE7" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:F6CA2DFAF25A04A47BD60EAF44447DE7" country="Japan" county="Fukui" latitude="36.0621" longLatPrecision="6" longitude="136.1304" name="Kuzuryu River system" stateProvince="Fukui Prefecture">Kuzuryu River system</location>
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+<collectingCounty id="5F02AFD68741A51B86490618B8E30F5D">Fukui</collectingCounty>
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+<collectingRegion id="87AD3D60190FA4EF87A227709F08580E" country="Japan" name="Fukui">Fukui Prefecture</collectingRegion>
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+<collectingCountry id="4C125ECED649199054651C68707B545D" name="Japan">Japan</collectingCountry>
+; 
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+<geoCoordinate id="08B520F6430F2C2A73743150A995DD5A" degrees="36.0621" direction="north" orientation="latitude" precision="5" value="36.0621">36.0621 ° N</geoCoordinate>
+, 
+<geoCoordinate id="F06A73AF4CF61C848FFFE93585E5A28A" degrees="136.1304" direction="east" orientation="longitude" precision="5" value="136.1304">136.1304 ° E</geoCoordinate>
+</named-content>
+; on 
+<date id="B4E99A15C0D468EC1FE9E223E8F81421" value="2021-09-06">
+<collectingDate id="10A3DE9C3C63CFC01A743CEFB03566D9" value="2021-09-06">6 Sep. 2021</collectingDate>
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+by 
+<collectorName id="AFE87AE3F901D291EB592425118BE992">Yuta Katayama</collectorName>
+.
+</materialsCitation>
+</paragraph>
+<paragraph id="54E7C7CCBC5563C2BDD46CCB0DDC828F">
+<emphasis id="F4841D7ECEC11C0011417C57BAA31A30" bold="true">
+<emphasis id="F333DE0D84C029AAE4AFCAB81724A818" bold="true" italics="true">
+<typeStatus id="8D13D22677AE720C740434B1B2E1342B" type="paratype">Paratypes</typeStatus>
+</emphasis>
+.
+</emphasis>
+<specimenCount id="1BFB677A770BD1CB08905DD4E3E4FE85" count="18" type="generic">Eighteen specimens</specimenCount>
+collected from 
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+in 
+<collectingRegion id="61CFF221CED459BF96993A95927434F2" country="Japan" name="Fukui">Fukui Prefecture</collectingRegion>
+, 
+<collectingCountry id="688001FC0441B3D91D8D591B1A1C8BD5" name="Japan">Japan</collectingCountry>
+by 
+<collectorName id="7A24E746E1E81EAF90856568738176EF">Y. Katayama</collectorName>
+. • 
+<materialsCitation id="E47E74A13312801D3223EC2C0890FAD0" collectingDate="2021-07-17" collectionCode="LBM" collectorName="Y. Katayama" country="Japan" location="Echizen" specimenCode="LBM 1210059177" specimenCount="1" stateProvince="Fukui" typeStatus="paratype">
+<specimenCode id="90A4F854DB5F42819675A6C298BFA6A9" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" type="">
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+</specimenCode>
+; 
+<quantity id="4ACC23AF82C740304AE832A814345047" metricMagnitude="-2" metricUnit="m" metricValue="4.4399999999999995" unit="mm" value="44.4">44.4 mm</quantity>
+SL from 
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+on 
+<date id="EBEC8D5BDD0D6E44C41FC71CE122EAFF" value="2021-07-17">
+<collectingDate id="32DC46AA454BDF8CB9AF4B79AE0CCB99" value="2021-07-17">17 Jul. 2021</collectingDate>
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+</materialsCitation>
+; • 
+<materialsCitation id="ACCA0822E164F4C41A30F42167E7A02D" collectionCode="LBM" collectorName="Y. Katayama" country="Japan" location="Kuzuryu River system" specimenCode="LBM 1210059178" specimenCount="1" stateProvince="Fukui" typeStatus="paratype">
+<specimenCode id="ED63140055323E5E843E869F1E62DEF3" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" type="">
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+, 
+<quantity id="F1F989E13B9AF17B755867C832E6E2FD" metricMagnitude="-2" metricUnit="m" metricValue="5.88" unit="mm" value="58.8">58.8 mm</quantity>
+SL
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+; • 
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+</materialsCitation>
+; • 
+<materialsCitation id="C0D0A436B170190C587733F0FE53A6DE" collectingDate="2023-10-01" collectionCode="FAKU" collectorName="Y. Katayama" country="Japan" location="Fukui" specimenCode="FAKU 211460" specimenCount="1" stateProvince="Fukui">
+<specimenCode id="2B3AE665D52358240DF1990C8CEF3352" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="9FF920D15BC5E462E4D98CFA64AB8A8D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211460
+</specimenCode>
+, 
+<quantity id="37CDDE0AF9FE77D241E2E0F47B11A341" metricMagnitude="-2" metricUnit="m" metricValue="4.45" unit="mm" value="44.5">44.5 mm</quantity>
+SL from 
+<location id="336294A89F7B05342BE6570282A68098" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:336294A89F7B05342BE6570282A68098" country="Japan" name="Fukui" stateProvince="Fukui">Fukui</location>
+on 
+<date id="2BD7A990B0D9DA9EC8FC9A3EC36DD30B" value="2023-10-01">
+<collectingDate id="4CC313255793AAE5272ABD5B60C60821" value="2023-10-01">1 Oct. 2023</collectingDate>
+</date>
+</materialsCitation>
+.
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EGIAI" type="diagnosis">
+<paragraph id="319D617AC1315B71D0DC30066852632F">
+<heading id="02A5135CFFF8133E4200AFFA832DD96B" reason="title">Diagnosis.</heading>
+</paragraph>
+<paragraph id="25D307CA21D1C7EEF48F13A2A1A0B940">
+<taxonomicName id="D1FC6B85844682C2F80C236713DB8CE8" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai" status="sp. nov.">
+<emphasis id="0EA83CECEE1B371BD6174B339F7B4B55" italics="true">Lefua nishimurai</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="A69752661E287A26B2A6E8C85B04131F" rank="species">sp. nov.</taxonomicNameLabel>
+can distinguished from all other species of 
+<taxonomicName id="5EC22FA9F3C418DF22C36DC0D7DEB759" authorityName="Herzenstein" authorityYear="1888" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="genus">
+<emphasis id="E8D842BCE6587E65E8645DCC3142D043" italics="true">Lefua</emphasis>
+</taxonomicName>
+by combing following features: absence of rhomboid or triangular dark blotches on middle of caudal fin base; absence of black longitudinal stripe on both body sides in mature males; absence of dusky cross bars on dorsal area of body; absence of dusky bar beside dorsal fin base; eyes located dorsally on head; narrow conspicuous longitudinal mark between base of outer rostral barbel and eye; small dorsal fin; small orbit diameter (6.3–11.2 % of head length); small value of interorbital width relative to body width (28.1–39.4 %); dark spots dorsally and ventrally on caudal fin base; small dark brown spots from snout to caudal peduncle; small dark spots on dorsal and caudal fins (approximately same size as eyes).
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0E5IAI" type="description">
+<paragraph id="59E64D5071607BE5C4B233CB49523AA7">
+<heading id="4D8F0F30299126085A5D15829AD7F374" reason="title">Description.</heading>
+</paragraph>
+<paragraph id="FC0DD058A6BC542FC9AEF0ED7301D812">
+Measurements and counts listed in Table 
+<tableCitation id="D3C2CBCD9B3D289BAA1C7BC9813C97E2" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Meristics and morphometrics of Lefua torrentis, L. tokaiensis., and L. nishimurai sp. nov. Measurements indicate minimum – maximum values (mean ± SD). Data of the holotypes of L. torrentis and L. tokaiensis were obtained from Hosoya et al. (2018) and Ito et al. (2019), respectively." httpUri="http://table.plazi.org/id/B7857D6A081FBE45190B6B271D4B321C" tableUuid="B7857D6A081FBE45190B6B271D4B321C">4</tableCitation>
+. Body elongated and cylindrical, slightly longitudinally flattened, and caudal peduncle laterally flattened. Dorsal and ventral sides of caudal peduncle keeled, and depth of caudal peduncle, including keel, about same as body depth. Head small (less than one-fourth of SL), slightly dorsoventrally compressed. Snout relatively long, comprising 38.7 % in 
+<typeStatus id="86165E1F2FABF650ED56E0256FF0AEE9" type="holotype">holotype</typeStatus>
+(32.3–46.4 % in 
+<typeStatus id="AB1AEB235BCEC6FC6DCCF22CB1FBF632" type="paratype">paratypes</typeStatus>
+and additional specimens) of head length, with tip rounded in dorsal view. Eyes positioned on dorsolateral surface of head (Figs 
+<figureCitation id="6D09A2F83098B6C1EE9E04FC1715D09C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of dorsal view of the head of three Lefua species. A. Kii-Shikoku Population of L. torrentis (FAKU 211486); B. Sanyo Population of L. torrentis (FAKU 211492); C. Nihonkai Population of L. torrentis (FAKU 211504); D L. tokaiensis (FAKU 211510); E. L. nishimurai sp. nov. (LBM 1210059179). Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1141334">6</figureCitation>
+– 
+<figureCitation id="996F9ACCA396D716A18F5423726D15E5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Detailed morphology of the holotype of Lefua nishimurai sp. nov (LBM 1210059189). A. Head; B. Lateral view of head; C. Mouth; D. Caudal fin base. An arrow in the lateral view of head (B) indicates narrow conspicuous black stripe between base of outer rostral barbel and eye. Arrows in the caudal fin base (D) indicate black spots on ventral and dorsal surfaces. Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1141336">8</figureCitation>
+), very small. Interorbital width comprises 51.5 % (42.9–59.0 %) of body width, and orbit diameter comprises 6.6 % (6.3–11.2 %) of head length (Figs 
+<figureCitation id="313F5EE770363BCD8BF6D98EFAC976F2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of dorsal view of the head of three Lefua species. A. Kii-Shikoku Population of L. torrentis (FAKU 211486); B. Sanyo Population of L. torrentis (FAKU 211492); C. Nihonkai Population of L. torrentis (FAKU 211504); D L. tokaiensis (FAKU 211510); E. L. nishimurai sp. nov. (LBM 1210059179). Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1141334">6</figureCitation>
+– 
+<figureCitation id="2023A1E29ECAAF74E4FF56F670576C16" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Detailed morphology of the holotype of Lefua nishimurai sp. nov (LBM 1210059189). A. Head; B. Lateral view of head; C. Mouth; D. Caudal fin base. An arrow in the lateral view of head (B) indicates narrow conspicuous black stripe between base of outer rostral barbel and eye. Arrows in the caudal fin base (D) indicate black spots on ventral and dorsal surfaces. Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1141336">8</figureCitation>
+). Mouth subterminal; small and inferior, slightly arched. Upper and lower lips fleshly and smooth (Figs 
+<figureCitation id="ACE954A4A0BD55C5F52544E4FB428F06" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Body coloration of the holotype of Lefua nishimurai sp. nov (LBM 1210059189). A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm." httpUri="https://binary.pensoft.net/fig/1141335">7</figureCitation>
+, 
+<figureCitation id="D40BFC2088BA148B05D21870DB06CF93" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Detailed morphology of the holotype of Lefua nishimurai sp. nov (LBM 1210059189). A. Head; B. Lateral view of head; C. Mouth; D. Caudal fin base. An arrow in the lateral view of head (B) indicates narrow conspicuous black stripe between base of outer rostral barbel and eye. Arrows in the caudal fin base (D) indicate black spots on ventral and dorsal surfaces. Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1141336">8</figureCitation>
+). No median incision in upper lip. One median notch in lower lip. Three pairs of barbels, two pairs of rostral barbels, and one pair of maxillary barbels. Inner rostral barbel relatively short, reaching to or slightly past vertical through anterior margin of eye when extended horizontally, outer rostral barbel longest, slightly past vertical through posterior margin of eye when extended horizontally, maxillary barbel past vertical through posterior margin of eye when extended horizontally. Nostrils separated from each other, anterior nostril formed as 1 long nasal barbel, posterior nostril larger than anterior nostril, anterior nostril at anterior side of 1 nasal barbel, close to base. Nasal barbel extended horizontally past posterior border of eye. Dorsal and ventral keels on caudal peduncle; dorsal keel starting slightly posterior to extremity of dorsal fin base, ventral keel starting posterior to extremity of anal fin base. Length of caudal peduncle 1.0 (1.0–1.3) times depth of caudal peduncle (depth including keel). Scales embedded on body. Lateral line absent. Dorsal fin rounded. Origins of dorsal fin nearer to caudal fin base than to tip of snout. Pectoral fin horizontal. Pelvic fin anterior to dorsal fin. Anal fin rounded. Anus positioned slightly anterior to anal fin. Caudal fin rounded. Dorsal fin with 4 (3–4) simple and 5 (5–6) branched rays. Second branched ray longest. Anal fin with 4 (3–4) simple and 5 (5–6) branched rays. Pectoral fin with 1 simple and 11 (10–11) branched rays. Second branched ray longest. Pelvic fin with 1 simple and 5 branched rays. No pelvic axillary lobe. Caudal fin with 2 (1–2) simple, 6 (5–7) branched, 7 (6–7) branched and 1 (1–2) simple principal rays. Total vertebrae 41 (39–43); abdominal vertebrae 23 (21–23), caudal vertebrae 18 (16–21).
+</paragraph>
+<caption id="C4B17EAA080DB3DFEC939A61C6A2DD2B" ID-arpha="1F857322-FF8C-5552-8CA1-13F69DB41EA4" httpUri="https://binary.pensoft.net/fig/1141335" startId="F7">
+<paragraph id="AA078B42CEF076C25477F40007D3C4B2">
+<label id="73D4CBE606DB6A084231CF130ACC84BF">Figure 7.</label>
+</paragraph>
+<paragraph id="45D4EF6915D3DAF6A569DCC2A918B7AD">
+Body coloration of the holotype of 
+<taxonomicName id="A11D9BCD758FA2C298FF4AFFF0D29865" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai" status="sp. nov.">
+<emphasis id="47D2535FCE5D8D21823445F43E52E662" italics="true">Lefua nishimurai</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="64DFC0CCECE0E97423B988FF1F49A25F" rank="species">sp. nov</taxonomicNameLabel>
+(
+<named-content id="B5C724E8D1D56968E057B9E57A2E901D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/lake-biwa-museum" xlink_title="Lake Biwa Museum">LBM</named-content>
+1210059189). 
+<emphasis id="D6630F27CA09469003B340CAF09DA166" bold="true">A.</emphasis>
+Lateral view; 
+<emphasis id="E580BE2D71DC4535E23ABA8A80AAB47F" bold="true">B.</emphasis>
+Dorsal view; 
+<emphasis id="8EEBF25C9D95FB3EBC4E2FBE06B026C0" bold="true">C.</emphasis>
+Ventral view. Scale bar: 10 mm.
+</paragraph>
+</caption>
+<caption id="BB19D5BAE6594EAE8EF9ADAC82BBA6C8" ID-arpha="626E29B6-98F0-5779-8FD6-B70120F999DA" httpUri="https://binary.pensoft.net/fig/1141336" startId="F8">
+<paragraph id="35B38C4B30F431C1EBB1568389A07A05">
+<label id="A3DEA361BBC40E22B62D3434037047F2">Figure 8.</label>
+</paragraph>
+<paragraph id="047CDB363D03296574E85E72E36E3A3D">
+Detailed morphology of the holotype of 
+<taxonomicName id="269079519DDC1002DE9CEE2FE8AAAB60" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai" status="sp. nov.">
+<emphasis id="90AF2081EECE9B61530C5517594297EF" italics="true">Lefua nishimurai</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="9CDB03D25BE964B3AB4E13D7DAE14AC3" rank="species">sp. nov</taxonomicNameLabel>
+(
+<named-content id="579A5F7FAD000E9A1A6E5F062DFD9A68" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/lake-biwa-museum" xlink_title="Lake Biwa Museum">LBM</named-content>
+1210059189). 
+<emphasis id="AFDEACFD9281271B44E8F65A86F5294A" bold="true">A.</emphasis>
+Head; 
+<emphasis id="C6FB984CA26648D1296205694FD70792" bold="true">B.</emphasis>
+Lateral view of head; 
+<emphasis id="C7F4EFF5321D156474AE65765BB58D68" bold="true">C.</emphasis>
+Mouth; 
+<emphasis id="32EA4F2D790C3A58EB5E03074AA4AFD2" bold="true">D.</emphasis>
+Caudal fin base. An arrow in the lateral view of head (
+<emphasis id="75DFB3409FF952763D68E02168AC6E73" bold="true">B</emphasis>
+) indicates narrow conspicuous black stripe between base of outer rostral barbel and eye. Arrows in the caudal fin base (
+<emphasis id="1AD8E5B0E27F15C9D9CE2F7DD6E68873" bold="true">D</emphasis>
+) indicate black spots on ventral and dorsal surfaces. Scale bars: 5 mm.
+</paragraph>
+</caption>
+<caption id="B7857D6A081FBE45190B6B271D4B321C" ID-Table-UUID="B7857D6A081FBE45190B6B271D4B321C" httpUri="http://table.plazi.org/id/B7857D6A081FBE45190B6B271D4B321C" startId="T4" targetIsTable="true">
+<paragraph id="4951F79FA27B6E6A7D1357D9BC2D8D29">
+<label id="5CE5DD1E5190F71FD431B0999C947521">Table 4.</label>
+</paragraph>
+<paragraph id="54ECBCA10602F8EA2541ECBA8BFC94C2">
+Meristics and morphometrics of 
+<taxonomicName id="A0B7329D3354FA624F8746235507C25B" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">
+<emphasis id="B4E51796B8A8527469BE6EB0F6AED28E" italics="true">Lefua torrentis</emphasis>
+</taxonomicName>
+, 
+<taxonomicName id="D9757861FC7CB4F051610CD5C8D61492" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="tokaiensis">
+<emphasis id="AA9639C0793EC6BEB0A70884F93DB375" italics="true">L. tokaiensis</emphasis>
+</taxonomicName>
+., and 
+<taxonomicName id="A0F0AC6BE429116297A1818FA34190A6" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai" status="sp. nov.">
+<emphasis id="D7468777AEF85F0C2CEA3906E3CAA4DB" italics="true">L. nishimurai</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="904F537EEF45A88A6B3FF470E1A6DF95" rank="species">sp. nov.</taxonomicNameLabel>
+Measurements indicate minimum – maximum values (mean ± SD). Data of the holotypes of 
+<taxonomicName id="99ABF30A47817B4C906F6E5934BA464E" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">
+<emphasis id="5241023CDA4C61623B93B92938254617" italics="true">L. torrentis</emphasis>
+</taxonomicName>
+and 
+<taxonomicName id="076A8753070275B7CE94EB97DD7D377D" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="tokaiensis">
+<emphasis id="2A1F34E53AF509A6918115CF7F6B71C3" italics="true">L. tokaiensis</emphasis>
+</taxonomicName>
+were obtained from 
+<bibRefCitation id="E02D3BC4013C060653AC4E50CA19CDAE" DOI="10.23788/IEF-1078" author="Hosoya K &amp; Ito T &amp; Miyazaki J" etAl="et al." firstAuthor="Hosoya" journalOrPublisher="Ichthyological Exploration of Freshwaters" pagination="193-201" refId="B11" refString="Hosoya K, Ito T, Miyazaki J (2018) Lefua torrentis, a new species of loach from western Japan (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters 28: 193–201. http://doi.org/10.23788/IEF-1078" title="Lefua torrentis, a new species of loach from western Japan (Teleostei: Nemacheilidae)." volume="28" year="2018">Hosoya et al. (2018)</bibRefCitation>
+and 
+<bibRefCitation id="CA6871104432679745766A194BDB41DE" DOI="10.1007/s10228-019-00690-0" author="Ito T &amp; Hosoya K &amp; Miyazaki J" etAl="et al." firstAuthor="Ito" refId="B15" refString="Ito T, Hosoya K, Miyazaki J (2019) Lefua tokaiensis, a new species of nemacheilid loach from central Japan (Teleostei: Nemacheilidae). Ichthyological Research 66: 479–487. https://doi.org/10.1007/s10228-019-00690-0" year="2019">Ito et al. (2019)</bibRefCitation>
+, respectively.
+</paragraph>
+</caption>
+<paragraph id="6FB3A114C6A923DC373A45C586315B00">
+<table id="TID0EYEBI" rules="all">
+<tbody id="0CF2798982F6620B9507B516F44E85FD">
+<tr id="63F1790405F6CFE967BA40D1CD23D02B">
+<th id="C4B7E8154AFD424515DD9BDD09F63D9F" colspan="1" rowspan="2">Character</th>
+<emphasis id="BB8E9A316E2D6E4F689DF23B4DCDA813" italics="true">
+<th id="3151635E52A595011B5201801F16AE5A" colspan="2" rowspan="1">
+<taxonomicName id="6608373FDCE838B492C6F81FABDDC12A" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">L. torrentis</taxonomicName>
+</th>
+<th id="EE1C583007622A9B9AD0E6D254EBD752" colspan="2" rowspan="1">
+<taxonomicName id="0741EA20B749677298C5783C44AACE46" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="tokaiensis">L. tokaiensis</taxonomicName>
+</th>
+<th id="351FEAF2EA6F7F33104BDE16A0059D28" colspan="3" rowspan="1">
+<taxonomicName id="38E18D9B97EDB57CFDDCFAA21B2D45E8" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai" status="sp. nov.">L. nishimurai</taxonomicName>
+<taxonomicNameLabel id="4E4A5BF78A68F6999AB93C003E4B0121" rank="species">sp. nov.</taxonomicNameLabel>
+</th>
+</emphasis>
+</tr>
+<tr id="BDEB64BFF695C29DAFC4E36B6F45CAD0">
+<th id="C8FA08D2E8F17500E341110BA9F8B731" colspan="1" rowspan="1">Holotype</th>
+<th id="46196B721D18C6C22946F2EFE06890B9" colspan="1" rowspan="1">Non-types (n = 46)</th>
+<th id="5CC1966BA74983916C4EC78236472951" colspan="1" rowspan="1">Holotype</th>
+<th id="1EE064F5D139E4B04781F1427E32AE7A" colspan="1" rowspan="1">Non-types (n = 21)</th>
+<th id="C5A99F657D1F2184D5BA2283B141BE27" colspan="1" rowspan="1">Holotype</th>
+<th id="D2411F3BF379937B005165A9315D2298" colspan="1" rowspan="1">Paratypes (n = 18)</th>
+<th id="9B61D53B7525A98E636D34B9A86D4D28" colspan="1" rowspan="1">Non-types (n = 6)</th>
+</tr>
+<tr id="6C987F3FCBE3F20BAA7C8AC2354DF909">
+<td id="8F3310CF267B9F5B148B766142408C8F" colspan="1" rowspan="1">Dorsal fin</td>
+<td id="927D9A43E936A522D6AF4356B7215B91" colspan="1" rowspan="1">ⅳ, 6</td>
+<td id="4155E9E0009FC55DB3482A053F701910" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="EED484175F3259451C73F81AEC2BACAD" colspan="1" rowspan="1">ⅳ, 6</td>
+<td id="35F59700F7235B4857F0F8856B0AC6D3" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="2BAD1BAF13EBC73D0332A922676DE232" colspan="1" rowspan="1">ⅳ, 5</td>
+<td id="35D0B3966B7FDCF3BD7156BD5F2B6D99" colspan="1" rowspan="1">ⅲ – ⅳ, 5–6</td>
+<td id="092F9B17A0F1E75D452D44506BB88CB8" colspan="1" isEmpty="true" rowspan="1">-</td>
+</tr>
+<tr id="7C724F0D0090069F50D673855C89D7AF">
+<td id="B2483D2C4938B79F72B5227414EA79C4" colspan="1" rowspan="1">Anal fin</td>
+<td id="3BCB4850B7896A511E6306292C05AC4F" colspan="1" rowspan="1">ⅳ, 5</td>
+<td id="E32456E8BF6C7829D44EF244338D399B" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="4C63BAD0540BCD97D022869A516F724C" colspan="1" rowspan="1">ⅳ, 5</td>
+<td id="9D5414817A9733F640E22A124BB72BB6" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="B19C467650DE63EA0CCABBFE0F4F8C5D" colspan="1" rowspan="1">ⅳ, 5</td>
+<td id="7BFFA65A91EF9B7BCC967F0E303DC6C4" colspan="1" rowspan="1">ⅲ – ⅳ, 5–6</td>
+<td id="DFBA1865F4F0F74B418B76E9F21393EA" colspan="1" isEmpty="true" rowspan="1">-</td>
+</tr>
+<tr id="03411E9A4370B4A7EFB2BB3B007FE1CE">
+<td id="897EC9FC58CE7EE38B13AD9CE55E4FA2" colspan="1" rowspan="1">Pectral fin</td>
+<td id="C6ACF8E7B427B3B83EC74B0596389772" colspan="1" rowspan="1">ⅰ, 9</td>
+<td id="0B9B72A13E6E81524DEBBD8833DA6FD3" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="12CD33868D1E7C28366CF68D86B7FE8D" colspan="1" rowspan="1">ⅰ, 7</td>
+<td id="0623A3EF96C892A53259F9CBEBD6CE59" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="9BA387A1E8704DD73B81AF3C86DCE11C" colspan="1" rowspan="1">ⅰ, 11</td>
+<td id="5558873F2D02AC1761540F2413EE4EBD" colspan="1" rowspan="1">ⅰ, 10–11</td>
+<td id="3D589AD10A6A055335044D6257F9F1DB" colspan="1" isEmpty="true" rowspan="1">-</td>
+</tr>
+<tr id="8BE0CE1F35AFFE726D069E58A7DFACBA">
+<td id="2B5EF689C65720B61B3F0A5B8FB0C727" colspan="1" rowspan="1">Pelvic fin</td>
+<td id="92136521C2D1F229B2A3803F67F3B608" colspan="1" rowspan="1">ⅰ, 5</td>
+<td id="221C8B095A24892275FC9599102DF5B6" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="181D1FD3E44E70D0C287E7EF42CBDCD3" colspan="1" rowspan="1">ⅰ, 5</td>
+<td id="7BA0A41E87D52D58F2F06A1614320244" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="226A0991EA3CC9C745D907A7D765810C" colspan="1" rowspan="1">ⅰ, 5</td>
+<td id="16FB0D490618A9E79D093022A1CA1754" colspan="1" rowspan="1">ⅰ, 5</td>
+<td id="CCE4AADBE4EB73C5E6EB1CFE48B70E63" colspan="1" isEmpty="true" rowspan="1">-</td>
+</tr>
+<tr id="A543FD881A7679762B3FB50ECA2F1730">
+<td id="C0E926FBC2C022F97E4E95E49FD70E6D" colspan="1" rowspan="1">Caudal fin</td>
+<td id="ADD4B62FD5EB892669105845EB1B5842" colspan="1" rowspan="1">ⅱ, 6, 8, ⅰ</td>
+<td id="7367FD2083F71F4C6A21924DD5680CD2" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="C8CED6AF26243EE1BF49CC9187A3DB63" colspan="1" rowspan="1">ⅰ, 7, 8, ⅱ</td>
+<td id="D3ECD4E1B63D9DBC9B58783A75AB09B9" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="EB9C867E89A8E112A35AF483CEF365B0" colspan="1" rowspan="1">ⅱ, 6, 7, ⅰ</td>
+<td id="A112D2D095527A68D98E7E5D86057699" colspan="1" rowspan="1">ⅰ – ⅱ, 5–7, 6–7, ⅰ – ⅱ</td>
+<td id="E44780B7B37A955939073D2CF7B5653B" colspan="1" isEmpty="true" rowspan="1">-</td>
+</tr>
+<tr id="A6DBA6DD1B7236389E7AAD0D33AB6529">
+<td id="0EBCB71284BC8749676599FB6773AE30" colspan="1" rowspan="1">Abdominal vertebrae</td>
+<td id="59196DD979FBA0B69235F3280F11525D" colspan="1" rowspan="1">19</td>
+<td id="DC8DA65A577177242E864A9AF96DA698" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="1F4E8368A95A387F2ADD6AFB7A2CC9CB" colspan="1" rowspan="1">19</td>
+<td id="DE8CF31E2FEE801EFAD1D368963F72E7" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="F236BB13945FD45AE2F271B6EED450E1" colspan="1" rowspan="1">23</td>
+<td id="470A22C64985C5FC89A386F35AC6F9A0" colspan="1" rowspan="1">21–23</td>
+<td id="B7BB704678DFEB48497AA2FB66C52775" colspan="1" isEmpty="true" rowspan="1">-</td>
+</tr>
+<tr id="22A6327BED7489C9C697CC6E4B56AECE">
+<td id="D0710DE5F7722173CB8C876F0A3AD3F8" colspan="1" rowspan="1">Caudal vertebrae</td>
+<td id="CD1EAA7E6B3879789CD761631EBA181C" colspan="1" rowspan="1">18</td>
+<td id="6E5F84406F7E3E3C5A650AA874650EB4" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="956C1EE8EB405E847B5C258F8F69AB6C" colspan="1" rowspan="1">18</td>
+<td id="665F60216D9B63977890AA2D639C22CD" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="210E12FDFDA8C0F33EC5DB65A9197399" colspan="1" rowspan="1">18</td>
+<td id="8F1C1784B39E416DB958F9CB16CB7CED" colspan="1" rowspan="1">16–21</td>
+<td id="DA750F2582825A3A2A74FBCD64D28DC5" colspan="1" isEmpty="true" rowspan="1">-</td>
+</tr>
+<tr id="62C31ED6DA97919BF653360D7D5AFBEA">
+<td id="64150AA2A9993193E0548F22196785BF" colspan="1" rowspan="1">Total vertebrae</td>
+<td id="349DC6AF8A6B2200FBADDACC66BD656C" colspan="1" rowspan="1">37</td>
+<td id="8331B11C3D927EDF0214554810344421" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="6891F35BE6DD02C696F18012F93D717C" colspan="1" rowspan="1">37</td>
+<td id="7456FAEF83CE2881EE5EBE16B03360F4" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="900BE450514561F2364EF566BE5F8866" colspan="1" rowspan="1">41</td>
+<td id="8275235D81F88990F07CE1A141EACFBB" colspan="1" rowspan="1">39–43</td>
+<td id="75223B6960415D7E864AA9073F63A998" colspan="1" isEmpty="true" rowspan="1">-</td>
+</tr>
+<tr id="D97003E969A34C0008BAEE11161ED2BD">
+<td id="A08869651EB7853D2AD418C196FA1772" colspan="1" rowspan="1">Standard length (mm)</td>
+<td id="B8546CBFA39E2B70F8ABC90E3A884CA8" colspan="1" rowspan="1">44.9</td>
+<td id="D31036CB5F26CB3D2AD5541DC8C1E253" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="22F735CF3074E5CBE47626148D7B1D6A" colspan="1" rowspan="1">44.2</td>
+<td id="3C9E8FC207797F05CB4A675A0D89DBB9" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="C15AC775C4257F538F115F9B52CB7E97" colspan="1" rowspan="1">51.2</td>
+<td id="0D92B37453A741A4CF4A0E1ED73404AC" colspan="1" rowspan="1">40.0–63.2</td>
+<td id="07B0EC4CE8116A5D939357315BC0D7F1" colspan="1" isEmpty="true" rowspan="1">-</td>
+</tr>
+<tr id="52863A4DEC9A15FF66895AC9CC399638">
+<td id="67772B55DCFBAD8E4BA9F3AD071501BC" colspan="8" rowspan="1">In % of standard length</td>
+</tr>
+<tr id="CDCAA98FC0867A4698AAA1247F94C2D3">
+<td id="AD31ADEBD9760F11868ED24DBCE06E9C" colspan="1" rowspan="1">Head length</td>
+<td id="648347C6DC10BDAF25C248A4F6A10F43" colspan="1" rowspan="1">20.9</td>
+<td id="9F048009F9F4C8378FADE966D6C9C846" colspan="1" rowspan="1">18.4–23.4 (21.0 ± 1.2)</td>
+<td id="613B14385FCB456524DC6D0D1B881EA8" colspan="1" rowspan="1">21.3</td>
+<td id="4F899AD6BA6472AE648D73D8C794893B" colspan="1" rowspan="1">19.1–26.4 (22.0 ± 1.9)</td>
+<td id="56EDDAEFF1E4BDAD8DE4C0FBE5D5B6F2" colspan="1" rowspan="1">20.7</td>
+<td id="ED29387B156FAFB9691090846E2204EC" colspan="1" rowspan="1">19.1–23.6 (20.6 ± 0.9)</td>
+<td id="D42F53053F81FAB2EEEE18D09878ED10" colspan="1" rowspan="1">19.6–22.0 (20.8 ± 1.0)</td>
+</tr>
+<tr id="7D54B481FAC0029D08E1C7DB929C97BB">
+<td id="62579CF295C5C69FCC6FF69EFAE54AF0" colspan="1" rowspan="1">Body depth</td>
+<td id="4BCDDC664A3FE1B5D3B239FB06627F5C" colspan="1" rowspan="1">14.4</td>
+<td id="FB5BEF54D77522AFEBADEA7B5BFF393D" colspan="1" rowspan="1">8.8–15.2 (12.1 ± 1.2)</td>
+<td id="D4FC0BBA30559E5C4EBC44DF2EED0364" colspan="1" rowspan="1">14</td>
+<td id="FFE556EF68A7D2FD7BB8B612B58828C4" colspan="1" rowspan="1">10.8–17.1 (13.6 ± 1.5)</td>
+<td id="BA0AB2469F028AD8D09D9F4BC89571D1" colspan="1" rowspan="1">12.7</td>
+<td id="252BE00311EA771837BE2C2CC540C3B6" colspan="1" rowspan="1">10.7–14.2 (12.4 ± 1.0)</td>
+<td id="6A606A771B7474DF90C8DE61FB3E34EA" colspan="1" rowspan="1">11.8–14.3 (13.1 ± 0.9)</td>
+</tr>
+<tr id="51A1DB18AF57EF187E2DA9F5912BC69E">
+<td id="A7D3C87ED94624E80F9372EBA23D2B79" colspan="1" rowspan="1">Body width</td>
+<td id="75A3CEBB4D83DCB7E1EE8A413281592B" colspan="1" rowspan="1">13.9</td>
+<td id="0D23E4D61B83E7196FBD73912CC56804" colspan="1" rowspan="1">11.2–14.7 (12.7 ± 0.8)</td>
+<td id="989A2187452A7C6DFC9E8037A8FB62AF" colspan="1" rowspan="1">11.7</td>
+<td id="9C15B2D53FE217097C3FBD15E2EDCD79" colspan="1" rowspan="1">12.3–17.2 (14.2 ± 1.4)</td>
+<td id="4030CB5B4363FCC6D873B5D5A492EA47" colspan="1" rowspan="1">13.3</td>
+<td id="23905C0C4AFABD394F799FBA1A3FF72E" colspan="1" rowspan="1">12.5–14.6 (13.6 ± 0.6)</td>
+<td id="5D53821CFE4EDA45917D05CE3DEC8FED" colspan="1" rowspan="1">12.9–14.2 (13.6 ± 0.4)</td>
+</tr>
+<tr id="A1E2C29D7956C543821ED1092B08E221">
+<td id="69B9D330900FD26E60DAB0AF9EFBA2C5" colspan="1" rowspan="1">Head width</td>
+<td id="3FC4F67EF36D47F34A1BBAC84D9BA8AE" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="147C0AE76B1431609B0DB64D91ABA1C6" colspan="1" rowspan="1">11.2–15.9 (13.2 ± 1.0)</td>
+<td id="B22E939C78942EC2A9AFE275A262CD9E" colspan="1" isEmpty="true" rowspan="1">-</td>
+<td id="01E48C843C46B53C934EE49948D902B4" colspan="1" rowspan="1">13.0–17.6 (15.2 ± 1.4)</td>
+<td id="9E7044EA55B20E87EE106E5549C2A996" colspan="1" rowspan="1">13.1</td>
+<td id="CC0571A8B2BA5C34049CA771209E3D03" colspan="1" rowspan="1">12.5–15.2 (13.5 ± 0.6)</td>
+<td id="4036A206DB5AE924F979CB44D34B1F5E" colspan="1" rowspan="1">12.9–14.4 (13.7 ± 0.5)</td>
+</tr>
+<tr id="E7F178D4A408B45141CE1FE85CA68900">
+<td id="8AC23572FD02BF2CA065A80BE054629F" colspan="1" rowspan="1">Depth of caudal peduncle</td>
+<td id="A38BDB525E54DA3890C62C7D9736D422" colspan="1" rowspan="1">12.4</td>
+<td id="39CEAA341A564F38F65C516A8625663A" colspan="1" rowspan="1">10.8–13.3 (12.1 ± 0.7)</td>
+<td id="738720E20BC414337843E790C9979C1A" colspan="1" rowspan="1">13.2</td>
+<td id="778DA4F212AB6BA671C7969D1A24D505" colspan="1" rowspan="1">11.0–17.1 (13.7 ± 1.4)</td>
+<td id="6F6C04CE89B71247F61A96F34CBD1C9B" colspan="1" rowspan="1">12.5</td>
+<td id="D1388FCF329BC24F1A59BF37238F8B8E" colspan="1" rowspan="1">10.7–13.5 (12.0 ± 0.7)</td>
+<td id="1F3A14B16AF60FA89B31A09429CC281A" colspan="1" rowspan="1">11.6–13.0 (12.2 ± 0.5)</td>
+</tr>
+<tr id="7B78FEAF23B496934FB870405B53B3AD">
+<td id="8DE866E25D069BBE0C950A9D428AD8E8" colspan="1" rowspan="1">Length of caudal peduncle</td>
+<td id="2BB6F4C27001A9424D7C2958728BBB56" colspan="1" rowspan="1">14.8</td>
+<td id="7E70B6E90323A91DBEECACAC5848FE9B" colspan="1" rowspan="1">11.7–16.8 (14.4 ± 1.1)</td>
+<td id="62CB011372943F19201519F236E6362F" colspan="1" rowspan="1">16.1</td>
+<td id="BCAAF699C55B30D7A91B174D8B63B4AD" colspan="1" rowspan="1">11.3–116.2 (13.7 ± 1.1)</td>
+<td id="A1C42A69EB7DF0699236662F4C9D7B2E" colspan="1" rowspan="1">12.7</td>
+<td id="7AD25DF7EEE01A51AAD38AAE3CC20CF7" colspan="1" rowspan="1">11.7–15.2 (13.5 ± 0.9)</td>
+<td id="13D082504D62FD73B34D927674D83EE5" colspan="1" rowspan="1">11.9–14.9 (13.3 ± 0.9)</td>
+</tr>
+<tr id="94820AAEB0078175B45043909D66845B">
+<td id="9584686F53C23C094A67F442E408517B" colspan="1" rowspan="1">Predorsal length</td>
+<td id="DC3BB8F9B656D78CD7D02A063C176097" colspan="1" rowspan="1">67.2</td>
+<td id="31331464F60D68B381798EC19FBC31A4" colspan="1" rowspan="1">61.5–69.1 (64.8 ± 1.6)</td>
+<td id="FE65E895C08DC9F2BFF3F76FA20053B9" colspan="1" rowspan="1">62.9</td>
+<td id="D6471E073177F5499931BC186469ADD6" colspan="1" rowspan="1">62.8–76.9 (66.3 ± 3.1)</td>
+<td id="265F626440B2ECFE289E571C08E4A249" colspan="1" rowspan="1">67.2</td>
+<td id="0339E7300324FB2D63C7D124DC9C8A6E" colspan="1" rowspan="1">63.2–69.3 (66.9 ± 1.3)</td>
+<td id="D4ED739D281175D865371F6406C716E9" colspan="1" rowspan="1">65.3–68.1 (67.1 ± 0.9)</td>
+</tr>
+<tr id="EC8A1E9290837C1A8A5B4E29A826AF29">
+<td id="04917EDF5BC237ED7B060F5D127D0BE5" colspan="1" rowspan="1">Preanal length</td>
+<td id="FE7BD60F75B7F69AF4E5651F975477F7" colspan="1" rowspan="1">76</td>
+<td id="7719EC952B889910CB7CB6B70CC2B809" colspan="1" rowspan="1">71.2–80.8 (75.0 ± 1.6)</td>
+<td id="1B2AE92B048D38DBFEE477DB15ECFCA5" colspan="1" rowspan="1">74.8</td>
+<td id="4B672D588432537C798C85288D64C45F" colspan="1" rowspan="1">72.3–87.0 (75.9 ± 3.1)</td>
+<td id="CD0D7B4301016FA401DD0882AC004E90" colspan="1" rowspan="1">77.9</td>
+<td id="67D04EC57775694B3DBF27EDA0F6AE26" colspan="1" rowspan="1">76.6–81.4 (79.3 ± 1.3)</td>
+<td id="0D1060DFBA65E09443C34E088D3413FB" colspan="1" rowspan="1">74.3–82.2 (77.1 ± 2.4)</td>
+</tr>
+<tr id="869F90C3C80566C094A8F78D11D79BB6">
+<td id="91F42A6B15418AE4BDDA7FA9A3F6AB2B" colspan="1" rowspan="1">Prepelvic length</td>
+<td id="5E34E1809B5C5713A8C82F93B6AA19BD" colspan="1" rowspan="1">56.1</td>
+<td id="EABFEFABB49EDA61923F8D37DB404C08" colspan="1" rowspan="1">53.3–65.2 (57.0 ± 1.9)</td>
+<td id="E6F46DF9CAE769D98A7B83D3689BC61C" colspan="1" rowspan="1">53.6</td>
+<td id="0B33EBD53D3C1AF9ACECD180542B9B51" colspan="1" rowspan="1">56.5–74.0 (60.1 ± 4.2)</td>
+<td id="4D3A2F30F14FF3ED6B0DA9A31A167437" colspan="1" rowspan="1">56.8</td>
+<td id="8D497E3E79509FFBD7EBBBF2A28329C5" colspan="1" rowspan="1">53.6–60.9 (57.6 ± 1.7)</td>
+<td id="7F8E47443CCF8A18B342157BDFE0F770" colspan="1" rowspan="1">56.8–60.5 (58.5 ± 1.1)</td>
+</tr>
+<tr id="761CE5C8B10925CA43ED33EE5F97D08D">
+<td id="9F91C9D7C7D7641895CC551C6637A1E3" colspan="1" rowspan="1">Height of dorsal fin</td>
+<td id="E7F4FE5A755E276D2BE3AA3ADC19CDBE" colspan="1" rowspan="1">10.9</td>
+<td id="B315C8490947355B9CC886E3C726A814" colspan="1" rowspan="1">6.6–11.7 (9.2 ± 1.1)</td>
+<td id="A0D621F887955765F46B670E8B79D61B" colspan="1" rowspan="1">11.3</td>
+<td id="36069284592D28FD18B8AAB3FD4ABF33" colspan="1" rowspan="1">6.1–11.8 (8.8 ± 1.3)</td>
+<td id="8522CB99092692843E73E70393AFBFC1" colspan="1" rowspan="1">8.6</td>
+<td id="E3901656A78BBBCB7DE529D1B0E15793" colspan="1" rowspan="1">5.6–10.0 (7.9 ± 1.1)</td>
+<td id="893C5F6715C930F6559155B9D3FE6632" colspan="1" rowspan="1">7.0–9.6 (8.5 ± 1.0)</td>
+</tr>
+<tr id="B90A4C67DC21BE6A9CB3BAF88A185204">
+<td id="49CC46A20B529A12F96B142318CC43AF" colspan="1" rowspan="1">Length of dorsal fin base</td>
+<td id="F47AFED08B892A675D2DAD0C4FC89077" colspan="1" rowspan="1">8.1</td>
+<td id="0DB69D2D4B746D3BA8EAF5D6B9660B25" colspan="1" rowspan="1">6.7–10.1 (8.6 ± 0.7)</td>
+<td id="3D0C635CF49E3E49C7AD5B10C54D0244" colspan="1" rowspan="1">8.6</td>
+<td id="3CA0E8441DCC12D4FEBD6389D9462650" colspan="1" rowspan="1">7.1–9.9 (8.5 ± 0.8)</td>
+<td id="5F800AC0C7757F29BB039D6FC52F2370" colspan="1" rowspan="1">8.2</td>
+<td id="FFE7A14459ADA4991B8BBE60B57343BB" colspan="1" rowspan="1">7.5–9.7 (8.6 ± 0.6)</td>
+<td id="6244E7DBCD8AF8B8E69D52CD5874101B" colspan="1" rowspan="1">7.2–9.0 (8.4 ± 0.6)</td>
+</tr>
+<tr id="8CD42D7B9121771446C88DC0EDC9EEB1">
+<td id="16186CA359EDF17D575CEE12E68812A4" colspan="1" rowspan="1">Height of anal fin</td>
+<td id="F403E18399B2FB806FB559C5188DBD43" colspan="1" rowspan="1">12.5</td>
+<td id="06B23188C062B95A0FFE251ADB5397CE" colspan="1" rowspan="1">5.5–10.7 (7.7 ± 1.1)</td>
+<td id="7D3B127394EFBD967713C9521C4E00AE" colspan="1" rowspan="1">10.3</td>
+<td id="4287454D8533977F178D9AB692034BA1" colspan="1" rowspan="1">7.0–11.3 (8.7 ± 1.0)</td>
+<td id="6324767DCA54BAD81A7690B325815DBA" colspan="1" rowspan="1">9.4</td>
+<td id="F0798BC70675D1B2371D1846B93E4CB5" colspan="1" rowspan="1">6.5–9.3 (7.9 ± 0.9)</td>
+<td id="12AFD94DC8B46C9FAB813EF6920831E1" colspan="1" rowspan="1">5.5–9.0 (7.7 ± 1.1)</td>
+</tr>
+<tr id="E703D22C035FC6D5BB0A50837D9A3F1F">
+<td id="207DDE29F788D063A8DDCBEC6FA69FD7" colspan="1" rowspan="1">Length of analfin base</td>
+<td id="5B14EA34B11F3301BBCED0DF1E59022B" colspan="1" rowspan="1">8.1</td>
+<td id="5ACD723215CF4547ABE3F7E9D39D46D5" colspan="1" rowspan="1">6.8–9.9 (8.9 ± 0.6)</td>
+<td id="2529874EF8A887A28120D40C825941A6" colspan="1" rowspan="1">6.6</td>
+<td id="E44FC3D88B0FCAA505D30F11A594EE91" colspan="1" rowspan="1">6.8–11.9 (8.2 ± 1.0)</td>
+<td id="764FA3F3FEBD8F4CDDFC2E8DB1D25725" colspan="1" rowspan="1">7.2</td>
+<td id="56D6315E4F9D33029DE809E7E001C3F5" colspan="1" rowspan="1">5.8–9.1 (7.8 ± 0.8)</td>
+<td id="8B23EEF8DF8A978B3431FB7691AA57C7" colspan="1" rowspan="1">7.0–8.3 (7.6 ± 0.4)</td>
+</tr>
+<tr id="F18F93D1C32A8CD3546CB31CE7BBD0B0">
+<td id="68A6EF2ED0481CA9F862F5B55E4ABF8C" colspan="1" rowspan="1">Pectral fin length</td>
+<td id="12244E9C20EEC9F40CE3B13ACDBC3D2B" colspan="1" rowspan="1">13.6</td>
+<td id="61FC06AF6C63A163885C51B071FAC8F6" colspan="1" rowspan="1">11.6–17.0 (14.1 ± 1.4)</td>
+<td id="3C5603E9BF1FD24D5489BBAF66830A49" colspan="1" rowspan="1">12.4</td>
+<td id="2586B8AA2F494F632DB3D4E2D448877B" colspan="1" rowspan="1">13.0–18.6 (15.1 ± 1.4)</td>
+<td id="3F5FB15436AB5F93B286EC1A1E72D622" colspan="1" rowspan="1">12.9</td>
+<td id="14649AC4151C06487C0D7F84A969F38A" colspan="1" rowspan="1">11.2–16.5 (13.1 ± 1.4)</td>
+<td id="0DE7B185EF90D207EAA843D01077F45F" colspan="1" rowspan="1">12.5–15.1 (13.5 ± 0.9)</td>
+</tr>
+<tr id="A1AFC62CD833903E2C25936CE744235F">
+<td id="68511137E29486B4CCA761DF9537C600" colspan="8" rowspan="1">In % of head length</td>
+</tr>
+<tr id="622457EEEFE533AB8D0E3F29A5C97C28">
+<td id="5DEE86AD2221EE12363344123C620653" colspan="1" rowspan="1">Snout length</td>
+<td id="CE56162D4DAB3E2C53F20F340B24018E" colspan="1" rowspan="1">34</td>
+<td id="FBD9FBFE418B8D8234F23DD0A4BA62F4" colspan="1" rowspan="1">41.6–53.8 (48.9 ± 3.3)</td>
+<td id="7D4B2D2D88C27139CC858E705DE4A25F" colspan="1" rowspan="1">39.7</td>
+<td id="6D04CCEC06E0D7F35998A2D7D3922920" colspan="1" rowspan="1">44.5–55.5 (49.4 ± 3.4)</td>
+<td id="FDFDAA167D7650CB157A4E84C767AEFA" colspan="1" rowspan="1">38.7</td>
+<td id="056B2585F0DD003B85CE17599D9BFBDB" colspan="1" rowspan="1">32.3–45.7 (40.1 ± 3.4)</td>
+<td id="402FCEA83BF64257A4972DBFAFE7506B" colspan="1" rowspan="1">42.2–46.4 (44.6 ± 1.5)</td>
+</tr>
+<tr id="73320D791462A1393BEA84C3E5813273">
+<td id="6EBED6E2E523319344388F82049CE579" colspan="1" rowspan="1">Orbit diameter</td>
+<td id="DCECC87BC32BC6BC6C196CE1F8EF40A7" colspan="1" rowspan="1">13</td>
+<td id="54599D7EC6C38CE71A05CE34FB424738" colspan="1" rowspan="1">6.9–15.9 (10.5 ± 1.7)</td>
+<td id="6E0E8169FD32BF2D2F7DA5A94F944583" colspan="1" rowspan="1">13.6</td>
+<td id="D9CE1C22108913DE7139EBF27376E5AC" colspan="1" rowspan="1">7.3–13.3 (9.5 ± 1.4)</td>
+<td id="135769C420BFC4D905DE255C15D2DFB6" colspan="1" rowspan="1">6.6</td>
+<td id="2C3D70F8E366E0CA8CC1FF0B60BDA352" colspan="1" rowspan="1">6.3–11.2 (8.1 ± 1.5)</td>
+<td id="1F6345116B8232E2F8FA5151E9E0DCC9" colspan="1" rowspan="1">7.5–9.4 (8.5 ± 0.6)</td>
+</tr>
+<tr id="50244930CA0333712AF55F9B82E8551A">
+<td id="720BE53492A8CD1E17E57F230D051E19" colspan="1" rowspan="1">Interorbital width</td>
+<td id="87114F037EA5CDAB5B6FD5BD8993382D" colspan="1" rowspan="1">39</td>
+<td id="B467D0B299D6BBC28914B5C2852E9564" colspan="1" rowspan="1">32.2–47.3 (39.6 ± 3.8)</td>
+<td id="0BBBC8DB7DC5DFDB1F1EC2285D016F8F" colspan="1" rowspan="1">36.3</td>
+<td id="A43D337B6E26A23C52EA273FA4E620C2" colspan="1" rowspan="1">39.3–48.9 (43.8 ± 2.7)</td>
+<td id="80E7C28A81653E35022B583EF6788E74" colspan="1" rowspan="1">33.0</td>
+<td id="A1674E4BE3F583616C1F7A56B432BF02" colspan="1" rowspan="1">28.1–39.4 (34.5 ± 3.2)</td>
+<td id="8E0998B0B3860AEE7E068139F54A99D4" colspan="1" rowspan="1">33.3–39.1 (36.8 ± 1.8)</td>
+</tr>
+<tr id="EBB7BD9A60095B0F0D5744183C14293E">
+<td id="FA7DC17CB8281603095923BDCC7D4115" colspan="1" rowspan="1">Ratio of interorbital width to body width</td>
+<td id="5697FBFE5DE1450AE445E5C096C8278D" colspan="1" rowspan="1">58.6</td>
+<td id="A635B4B03CF76B6BDF8E52A6256132BF" colspan="1" rowspan="1">55.9–82.1 (65.6 ± 6.0)</td>
+<td id="1A42D80580E19C5437BCEC9505863848" colspan="1" rowspan="1">66.1</td>
+<td id="0A7AFD8682121F24FF0891283F2A561E" colspan="1" rowspan="1">56.3–76.4 (68.0 ± 6.2)</td>
+<td id="92DFD5911D008FCE21431D981C512A9C" colspan="1" rowspan="1">51.5</td>
+<td id="72EDC50F3CC008F6A83A3381534B5438" colspan="1" rowspan="1">42.9–58.2 (52.2 ± 4.3)</td>
+<td id="A9C5E986A0968ECEE251B3E63C540E00" colspan="1" rowspan="1">51.6–59.0 (56.1 ± 3.0)</td>
+</tr>
+</tbody>
+</table>
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EKGBI" type="Coloration">
+<paragraph id="CDE8D6D919088C0ECE14BCFDB091994B">
+<heading id="CFBA63BF279B5BC3EECA09C2E5A1FBFA" reason="title">Coloration.</heading>
+</paragraph>
+<paragraph id="31784044D5E967DBECC811ADC6FDB112">
+Live specimens, body and head light brown with strong yellowish tinge except for whitish gray ventral surface (Fig. 
+<figureCitation id="758681096E0951E1BD49E98EF5254B80" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Body coloration of the holotype of Lefua nishimurai sp. nov (LBM 1210059189). A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm." httpUri="https://binary.pensoft.net/fig/1141335">7</figureCitation>
+). One narrow conspicuous black stripe between base of outer rostral barbel and eye (Fig. 
+<figureCitation id="197A8779CB0769BA21021677C0B5EA08" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Detailed morphology of the holotype of Lefua nishimurai sp. nov (LBM 1210059189). A. Head; B. Lateral view of head; C. Mouth; D. Caudal fin base. An arrow in the lateral view of head (B) indicates narrow conspicuous black stripe between base of outer rostral barbel and eye. Arrows in the caudal fin base (D) indicate black spots on ventral and dorsal surfaces. Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1141336">8</figureCitation>
+). Entire body from snout to caudal peduncle scattered dark brown spots. Largest of spots slightly larger than orbit diameter. Both lips white, with one pair of small brown spots inside lower lip tissue (Fig. 
+<figureCitation id="F4141760B123EA28CEF5D0A6E1C9A266" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Detailed morphology of the holotype of Lefua nishimurai sp. nov (LBM 1210059189). A. Head; B. Lateral view of head; C. Mouth; D. Caudal fin base. An arrow in the lateral view of head (B) indicates narrow conspicuous black stripe between base of outer rostral barbel and eye. Arrows in the caudal fin base (D) indicate black spots on ventral and dorsal surfaces. Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1141336">8</figureCitation>
+). All fins pale to translucent. Pectoral fin yellowish brown, with dark spots at the base and anterior half. Dorsal fin yellowish brown with dark spots from base to middle. Anal fin with a few dark spots on basal half (sometimes absent). No spots on pelvic fin. Caudal fin yellowish brown with dark spots from base to a little closer to tip of middle, but translucent and without spots on margins. Dorsal and ventral surfaces of caudal fin base with dark brown spots (rarely absent) (Figs 
+<figureCitation id="F79A2E7B9DA8F248B8B2B6F8FEE1BE1B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Detailed morphology of the holotype of Lefua nishimurai sp. nov (LBM 1210059189). A. Head; B. Lateral view of head; C. Mouth; D. Caudal fin base. An arrow in the lateral view of head (B) indicates narrow conspicuous black stripe between base of outer rostral barbel and eye. Arrows in the caudal fin base (D) indicate black spots on ventral and dorsal surfaces. Scale bars: 5 mm." httpUri="https://binary.pensoft.net/fig/1141336">8</figureCitation>
+, 
+<figureCitation id="DB45A743A444CA39B0887F85D3AB5649" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Body coloration of atypical specimen of Lefua nishimurai sp. nov. (LBM 1210059188), showing no distinct dark brown spots scattered over the body and obscure dark band in the skin of the caudal fin base. Scale bar: 10 mm." httpUri="https://binary.pensoft.net/fig/1141337">9</figureCitation>
+). No obscure dark band in skin of caudal fin base (rarely present) (Figs 
+<figureCitation id="4360B4B5B2D13F2D7BC6700E5D14CEB2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body morphology and coloration of three Lefua species. A. Kii-Shikoku Population of L. torrentis (FAKU 211486); B. Sanyo Population of L. torrentis (FAKU 211492); C. Nihonkai Population of L. torrentis (FAKU 211504); D. L. tokaiensis (FAKU 211510); E. L. nishimurai sp. nov. (LBM 1210059189). Scale bar: 10 mm." httpUri="https://binary.pensoft.net/fig/1141333">5</figureCitation>
+, 
+<figureCitation id="C86F8D13C1BD1C1FAA2846EC2FD2F1AC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Body coloration of the holotype of Lefua nishimurai sp. nov (LBM 1210059189). A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm." httpUri="https://binary.pensoft.net/fig/1141335">7</figureCitation>
+, 
+<figureCitation id="A5A0512DDB198E59F52CB32C8270FEB0" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Body coloration of atypical specimen of Lefua nishimurai sp. nov. (LBM 1210059188), showing no distinct dark brown spots scattered over the body and obscure dark band in the skin of the caudal fin base. Scale bar: 10 mm." httpUri="https://binary.pensoft.net/fig/1141337">9</figureCitation>
+).
+</paragraph>
+<caption id="3EDF6452C7191E7CE5B39A7B0461535E" ID-arpha="5F620C52-76C6-5604-928B-44266A9F5143" httpUri="https://binary.pensoft.net/fig/1141337" startId="F9">
+<paragraph id="C7C39156105925C06A6ECEF0A80B3585">
+<label id="C58D30315046BCF508CC2A94B44F82DA">Figure 9.</label>
+</paragraph>
+<paragraph id="5C94BBBEDF94812A3858F61BBD149D99">
+Body coloration of atypical specimen of 
+<taxonomicName id="3BC9613760B15973E551D3008977663C" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai" status="sp. nov.">
+<emphasis id="E5EA9859299F10997C8B0FC6A93719B1" italics="true">Lefua nishimurai</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="C52D9BA15CE231F539B6C6148EB870F1" rank="species">sp. nov.</taxonomicNameLabel>
+(
+<named-content id="4F75D475B5206442676A1A4851BA5B3B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/lake-biwa-museum" xlink_title="Lake Biwa Museum">LBM</named-content>
+1210059188), showing no distinct dark brown spots scattered over the body and obscure dark band in the skin of the caudal fin base. Scale bar: 10 mm.
+</paragraph>
+</caption>
+</subSubSection>
+<subSubSection id="SECID0ENIBI" type="etymology">
+<paragraph id="1328EDB274CC3E4D36A39A1F79BC2E80">
+<heading id="24E5AA42902D8758F8D67211397E5B59" reason="title">Etymology.</heading>
+</paragraph>
+<paragraph id="2ABF9580E0E148EF3DA8F57CCA0CD5B2">The specific name is dedicated to Toshiaki Nishimura, who first morphologically distinguished this new species.</paragraph>
+</subSubSection>
+<subSubSection id="SECID0ESIBI" type="distribution">
+<paragraph id="FDE8B9F389F7ECCF173717274B62EB70">
+<heading id="47B3CF201484065C4FF4A825346EA1CB" reason="title">Distribution.</heading>
+</paragraph>
+<paragraph id="686006822BD48327CFD4F51960F7A340">
+This new species was recorded in the Kuzuryu River drainage system of the Reihoku Region in 
+<collectingRegion id="41F82280C1163EA2C283071E9EDA9FE3" country="Japan" name="Fukui">Fukui Prefecture</collectingRegion>
+, central 
+<collectingCountry id="9B2F0F8A8F1FDB7104B03FDA09F77F52" name="Japan">Japan</collectingCountry>
+(
+<bibRefCitation id="6BBDC44999550A834C66CF1590999D9B" author="Katayama" firstAuthor="Katayama" journalOrPublisher="Bulletin of the Fukui City Museum of Natural History" pagination="53-56" refId="B20" refString="Katayama Y (2021) A survey of the distribution of the Reihoku population of the Lefua torrentis (Nemacheilidae) with a first record in Fukui city. Bulletin of the Fukui City Museum of Natural History 68: 53–56." title="A survey of the distribution of the Reihoku population of the Lefua torrentis (Nemacheilidae) with a first record in Fukui city." volume="68" year="2021">Katayama 2021</bibRefCitation>
+; Fig. 
+<figureCitation id="FEE5F3EFA9CF642A88E754B173AFF85A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of central Japan showing the distribution of three Lefua species. Orange, L. torrentis; red, L. tokaiensis; purple, L. nishimurai sp. nov. The distribution of L. torrentis and L. tokaiensis were in accordance with Yamada (2015) and Nakajima and Uchiyama (2017)." httpUri="https://binary.pensoft.net/fig/1141329">1</figureCitation>
+).
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0EAJBI" type="biology|habitat">
+<paragraph id="B42E8E57929BC38F2093D33951A31288">
+<heading id="CC37CB5A7ED4CCA70BD5A744AAFBE3F9" reason="title">Habitat and biology.</heading>
+</paragraph>
+<paragraph id="B8939DC0E900A638414873453DA81656">
+<taxonomicName id="5441720D031BDC67FCA0340578359FE2" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai" status="sp. nov.">
+<emphasis id="EF0E85CB2237828C1E0E18E7E7DF3760" italics="true">Lefua nishimurai</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="8652AD381A91C8DA74F50A4100DA0D71" rank="species">sp. nov.</taxonomicNameLabel>
+inhabits small mountain streams surrounded by natural or planted forests, and prefers sandy or gravelly bottoms (Fig. 
+<figureCitation id="8C9EFD8FD70B8DE8EF4A1878A81DDEC9" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Typical habitat of Lefua nishimurai sp. nov. in the Kuzuryu River drainage system in Fukui Prefecture, Japan." httpUri="https://binary.pensoft.net/fig/1141338">10</figureCitation>
+). The new species usually hides under rocks and stones at the river edge or under sediments, such as fallen leaves. It has been estimated that spawning of this species occurs from late April to mid-June (
+<bibRefCitation id="DEFA3CD508CF9802EB96668BCB814AAE" author="Katayama" firstAuthor="Katayama" journalOrPublisher="Bulletin of the Fukui City Museum of Natural History" pagination="53-56" refId="B20" refString="Katayama Y (2021) A survey of the distribution of the Reihoku population of the Lefua torrentis (Nemacheilidae) with a first record in Fukui city. Bulletin of the Fukui City Museum of Natural History 68: 53–56." title="A survey of the distribution of the Reihoku population of the Lefua torrentis (Nemacheilidae) with a first record in Fukui city." volume="68" year="2021">Katayama 2021</bibRefCitation>
+).
+</paragraph>
+<caption id="28AF3F146AB45CDABBC866CBE6C9A6C6" ID-arpha="FFC76EA7-EF8D-55B5-B0B4-75EBF5DD2F84" httpUri="https://binary.pensoft.net/fig/1141338" startId="F10">
+<paragraph id="3AE17FF6F74DBE676225976ADB8CE7F4">
+<label id="67CB3B26362FEE7F6E793D2B4E40FFE8">Figure 10.</label>
+</paragraph>
+<paragraph id="44A0E4923E7557893B45A465795E73C0">
+Typical habitat of 
+<taxonomicName id="26769D365E85B01417BDDE2E4DC29DCC" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai" status="sp. nov.">
+<emphasis id="5B8B19D2D1156935BCA0B187C74B1B85" italics="true">Lefua nishimurai</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="35678DBD45F4AA759E8DF7C67C5AA5C2" rank="species">sp. nov.</taxonomicNameLabel>
+in the Kuzuryu River drainage system in Fukui Prefecture, Japan.
+</paragraph>
+</caption>
+</subSubSection>
+<subSubSection id="SECID0ERKBI" type="remarks">
+<paragraph id="4B7A5193B153943CB83098CCEDFDBD8C">
+<heading id="29C530F6F052C28A7A9836D0A329461D" reason="title">Remarks.</heading>
+</paragraph>
+<paragraph id="57EE0113BED83CABBA0629EE39099F0F">
+The new species is similar to 
+<taxonomicName id="CACA8DF4346795C9C64A224CF47C99CC" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">
+<emphasis id="88E1459AE077AAE9ABEB18234E0E1C90" italics="true">L. torrentis</emphasis>
+</taxonomicName>
+and 
+<taxonomicName id="1273228E70BE757E8E1C69A6ADFAE223" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="tokaiensis">
+<emphasis id="2AF454AED8D8CA9038B84C28C4F159D1" italics="true">L. tokaiensis</emphasis>
+</taxonomicName>
+with a narrow conspicuous longitudinal mark between the base of the outer rostral barbel and eye. However, 
+<taxonomicName id="596E49BA20D090E43BA64ACEB25F51F5" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai" status="sp. nov.">
+<emphasis id="60E154CCF4BA90AF3010F03ABE965F12" italics="true">L. nishimurai</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="6D00340154093A489198C949330B6002" rank="species">sp. nov.</taxonomicNameLabel>
+can be distinguished from 
+<taxonomicName id="118589608D75490E55FD4BC42415EE3B" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">
+<emphasis id="92009978F2ED5B4295FA136950536F13" italics="true">L. torrentis</emphasis>
+</taxonomicName>
+based on the following characteristics: eyes located dorsally on head (more lateral in 
+<taxonomicName id="9E3972D19B85B95ABD7F8B1DC8F7C5AB" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">
+<emphasis id="A40C4E509035C85BB0E5BD15D552B630" italics="true">L. torrentis</emphasis>
+</taxonomicName>
+), small orbit diameter (larger in 
+<taxonomicName id="C4FEBE10D0D59A8CE98E1F6AC32736AA" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">
+<emphasis id="00832527DD637DECD535363C77C71769" italics="true">L. torrentis</emphasis>
+</taxonomicName>
+), dark spots on the caudal fin base (rarely absent in the new species; usually absent in 
+<taxonomicName id="52D66C416B0D85D2B073F34B0D8EA846" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">
+<emphasis id="C55E1DD9AF6F5036D9D27A93AB7D4E0C" italics="true">L. torrentis</emphasis>
+</taxonomicName>
+), small dark spots on dorsal and caudal fins, and distinct dark brown spots scattered over the body (rarely absent in the new species; entirely absent in 
+<taxonomicName id="EA07C3947CF2416ED15B3DB71CB1BB29" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">
+<emphasis id="BF795B13CFCB45D97509A5ECD2280DA1" italics="true">L. torrentis</emphasis>
+</taxonomicName>
+). The new species and 
+<taxonomicName id="A2356E5A39FE9D90D7DEEE60BDB0CD71" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="tokaiensis">
+<emphasis id="1224D7362EA454863EE7450B1B9F1C27" italics="true">L. tokaiensis</emphasis>
+</taxonomicName>
+can also be discriminated by eyes located dorsally on the head (more lateral in 
+<taxonomicName id="3D3AEEC6BF3C33C55B8C5BF95BF4FD49" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="tokaiensis">
+<emphasis id="C98D01BFD15DD04B3C35884F3A7EE656" italics="true">L. tokaiensis</emphasis>
+</taxonomicName>
+), small orbit diameter (larger in 
+<taxonomicName id="3D1644D8C8AA08690C9D3678BB094B2E" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="tokaiensis">
+<emphasis id="BE3073BA1F44E5969FACE0350B3B8529" italics="true">L. tokaiensis</emphasis>
+</taxonomicName>
+), dark spots dorsally and ventrally on the caudal fin base (rarely absent in the new species; entirely absent in 
+<taxonomicName id="202FB7913553FF5F55742BA4C0F23544" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="tokaiensis">
+<emphasis id="4D72597B0E2F0484B644B72F6A5B7A3D" italics="true">L. tokaiensis</emphasis>
+</taxonomicName>
+), distinct dark brown spots scattered over the body (rarely absent in the new species; entirely absent in 
+<taxonomicName id="A5E7E6D24273E299428F25C48AFB2FAF" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="tokaiensis">
+<emphasis id="B759B2475732D33E81CB85F3660BFE8A" italics="true">L. tokaiensis</emphasis>
+</taxonomicName>
+), and fan-shaped caudal fin of 
+<taxonomicName id="34502B822D5F8A2B58B82E26FA95D60C" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai">
+<emphasis id="29CEBCFC00ABD1E5680CE79F2A6B40B3" italics="true">L. nishimurai</emphasis>
+</taxonomicName>
+(squared-off apex in 
+<taxonomicName id="7BCB07EFBF6ED10535C079C61D4C7E36" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="tokaiensis">
+<emphasis id="AFFDDF6BFFC3CF26C9DBB4E92DE20FA2" italics="true">L. tokaiensis</emphasis>
+</taxonomicName>
+). Both the new species and 
+<taxonomicName id="5B7FD7F53CCD98BC8CDA8B1DCA4F9533" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">
+<emphasis id="914040A8D566D4E7AF9736A48F740D64" italics="true">L. torrentis</emphasis>
+</taxonomicName>
+inhabiting 
+<collectingRegion id="E0F6EDC15CFD5606FB2A4DCC96194054" country="Japan" name="Wakayama">Wakayama Prefecture</collectingRegion>
+, and Shikoku and Awaji Islands possess dark brown spots of varying sizes on the body, dorsal side, and caudal peduncle (
+<bibRefCitation id="4FCC496B09BE90358D4347FA99FB31D4" DOI="10.23788/IEF-1078" author="Hosoya" etAl="et al." firstAuthor="Hosoya" journalOrPublisher="Ichthyological Exploration of Freshwaters" pagination="193-201" refId="B11" refString="Hosoya K, Ito T, Miyazaki J (2018) Lefua torrentis, a new species of loach from western Japan (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters 28: 193–201. http://doi.org/10.23788/IEF-1078" title="Lefua torrentis, a new species of loach from western Japan (Teleostei: Nemacheilidae)." volume="28" year="2018">Hosoya et al. 2018</bibRefCitation>
+). However, 
+<taxonomicName id="05EAFE400C11D34F24FFCDF4AE4B382E" authorityName="Katayama" authorityYear="2024" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="nishimurai" status="sp. nov.">
+<emphasis id="728AF04BF612B355D00B0BE974CDCEAD" italics="true">L. nishimurai</emphasis>
+</taxonomicName>
+<taxonomicNameLabel id="18301ED6B67986C1ABDBE2384ED2E963" rank="species">sp. nov.</taxonomicNameLabel>
+represents smaller spots than the other species on the entire body and dorsal surfaces as well as on the dorsal and caudal fins (approximately the same size as the eyes) (Fig. 
+<figureCitation id="34EA69B677376D4C189920D440E02196" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of body morphology and coloration of three Lefua species. A. Kii-Shikoku Population of L. torrentis (FAKU 211486); B. Sanyo Population of L. torrentis (FAKU 211492); C. Nihonkai Population of L. torrentis (FAKU 211504); D. L. tokaiensis (FAKU 211510); E. L. nishimurai sp. nov. (LBM 1210059189). Scale bar: 10 mm." httpUri="https://binary.pensoft.net/fig/1141333">5</figureCitation>
+).
+</paragraph>
+</subSubSection>
+<subSubSection id="SECID0E1QBI" type="material">
+<paragraph id="EC01EC7D19A0F1CF538D3999CAA5F762">
+<heading id="71F2E6A21C22AF0E499848137B9AFF33" reason="title">Comparative material examined.</heading>
+</paragraph>
+<paragraph id="84C70AB813193297C89E38B9F185DB3B">
+<taxonomicName id="E76D2FFD4203DF7CE053E30ED101FCD6" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="torrentis">
+<emphasis id="F8E8271F3FBDAAA6068F32B23386F6FC" italics="true">Lefua torrentis</emphasis>
+</taxonomicName>
+, 
+<specimenCount id="FA24958FCA6D435C3825704C5630E9C5" count="46" type="generic">46 specimens</specimenCount>
+(
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+SL). 
+<emphasis id="5A557AA9C9B710FE44592DF942F526D5" bold="true" italics="true">
+<typeStatus id="6C3AF8748E520CB166A8DE1C79D6B998" type="holotype">Holotype</typeStatus>
+</emphasis>
+: • KUN-P 45408; 
+<quantity id="32CD23BB214CC1D3DB66B966C6144941" metricMagnitude="-2" metricUnit="m" metricValue="4.49" unit="mm" value="44.9">44.9 mm</quantity>
+SL collected from 
+<collectingCounty id="D95CE44EE1AA3C03781499DE517029F1">Yura River</collectingCounty>
+system, 
+<collectingMunicipality id="41780DE276AF3259E106685865BBB100">Kasuga</collectingMunicipality>
+, 
+<location id="A36227A7D78C7A60E9A0A86EB855BC4D" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:A36227A7D78C7A60E9A0A86EB855BC4D" county="Yura River" municipality="Kasuga" name="Tanba" stateProvince="Hyogo">Tanba</location>
+, 
+<collectingRegion id="FF59754CAAA1AE5232B4565491E56776" country="Japan" name="Hyogo">Hyogo</collectingRegion>
+; • 
+<specimenCode id="578A22FC1D2F0708F854420AB65DBBA7" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="C51DF66BBB6F7467C3D34109A6D18498" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211461–211464
+</specimenCode>
+, 
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+, 
+<quantity id="400643CFE484B7066767432AF5A605E3" metricMagnitude="-2" metricUnit="m" metricValue="4.13" metricValueMax="4.9399999999999995" metricValueMin="3.32" unit="mm" value="41.3" valueMax="49.4" valueMin="33.2">33.2–49.4 mm</quantity>
+SL from 
+<location id="A5292FAC525FD4FAC5BFEC219B09D93F" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:A5292FAC525FD4FAC5BFEC219B09D93F" county="Yura River" municipality="Kasuga" name="Arida River" stateProvince="Hyogo">Arida River</location>
+system, 
+<location id="FB7F54ED46796EC8261463352A132A6C" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:FB7F54ED46796EC8261463352A132A6C" county="Yura River" municipality="Kasuga" name="Aridagawa" stateProvince="Hyogo">Aridagawa</location>
+, 
+<collectingRegion id="B7F6868538D57D0B042944569B41385E" country="Japan" name="Wakayama">Wakayama</collectingRegion>
+; • 
+<specimenCode id="33889440D194FCABE3F987299B6F4946" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="E516341CD9BA7E2C4D841BD1CC729F6F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211465–211468
+</specimenCode>
+, 
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+, 
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+SL, from 
+<location id="CB16F71C6693CE7E2FF12CF4E45CE7B1" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:CB16F71C6693CE7E2FF12CF4E45CE7B1" county="Yura River" municipality="Kasuga" name="Hidaka River" stateProvince="Hyogo">Hidaka River</location>
+system, 
+<location id="289E2C542FB1A71BE7961F5826BF6230" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:289E2C542FB1A71BE7961F5826BF6230" county="Yura River" municipality="Kasuga" name="Hidaka" stateProvince="Hyogo">Hidaka</location>
+, 
+<collectingRegion id="CF6669326097A1714836E0EF5B5B87DC" country="Japan" name="Wakayama">Wakayama</collectingRegion>
+; • 
+<specimenCode id="2559846237FA8A01D8FF1ED294E65620" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="A474BFA50F70305683DAA1573F06BE83" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211469–211477
+</specimenCode>
+, 
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+, 
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+SL from 
+<location id="051476386A41A3A52FAE77D252AECAE8" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:051476386A41A3A52FAE77D252AECAE8" county="Yura River" municipality="Kasuga" name="Orino River" stateProvince="Hyogo">Orino River</location>
+system, 
+<location id="80814FEDBAE91E728BC5AA614BE00638" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:80814FEDBAE91E728BC5AA614BE00638" county="Yura River" municipality="Kasuga" name="Kitanada" stateProvince="Hyogo">Kitanada</location>
+, 
+<collectingRegion id="E429C880D9B4B1873D1C4C8561C79899" country="Japan" name="Tokushima">Tokushima</collectingRegion>
+; • 
+<specimenCode id="4A1BF3B0519994C11BFD125A3473F186" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="D2D0B29491DDECA2625D1C3BE81C7BC1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211478–211480
+</specimenCode>
+, 
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+, 
+<quantity id="3C9731DF147F3FFE935BAFE46C204DC7" metricMagnitude="-2" metricUnit="m" metricValue="4.92" metricValueMax="5.55" metricValueMin="4.29" unit="mm" value="49.2" valueMax="55.5" valueMin="42.9">42.9–55.5 mm</quantity>
+SL from 
+<location id="BE4A4988ABF1F3CED6BA3BF1306EA9A1" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:BE4A4988ABF1F3CED6BA3BF1306EA9A1" county="Yura River" municipality="Kasuga" name="Yoshino River" stateProvince="Hyogo">Yoshino River</location>
+system, 
+<location id="AB2C98880629615CA7BE1A1853CAD2FE" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:AB2C98880629615CA7BE1A1853CAD2FE" county="Yura River" municipality="Kasuga" name="Gomyo" stateProvince="Hyogo">Gomyo</location>
+, 
+<collectingRegion id="2C145FC5EC1C8CBC85C87A4FABA13CBE" country="Japan" name="Kagawa">Kagawa</collectingRegion>
+; • 
+<specimenCode id="EE86049C8C34B5D5B618ED7C8DA84CD2" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="73FBB7FA0C68D94F804C36A79372C3A1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211481–211484
+</specimenCode>
+, 
+<specimenCount id="68E240156B9720DC46FD96B0AB9B22DF" count="4" type="generic">4 specimens</specimenCount>
+, 
+<quantity id="87DBB6089DB45A571E6CE60ED95010E5" metricMagnitude="-2" metricUnit="m" metricValue="5.2" metricValueMax="6.36" metricValueMin="4.04" unit="mm" value="52.0" valueMax="63.6" valueMin="40.4">40.4–63.6 mm</quantity>
+SL from 
+<location id="1B4C91CEBCA14CB1B30834E63E4089D7" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:1B4C91CEBCA14CB1B30834E63E4089D7" county="Yura River" municipality="Kasuga" name="Sumoto River" stateProvince="Hyogo">Sumoto River</location>
+system, 
+<location id="FC55B680913B57D379FAE471069ABC2D" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:FC55B680913B57D379FAE471069ABC2D" county="Yura River" municipality="Kasuga" name="Ayuya" stateProvince="Hyogo">Ayuya</location>
+, 
+<collectingRegion id="FF303CF05976F24B6F8B3A3EBFDAD6F3" country="Japan" name="Hyogo">Hyogo</collectingRegion>
+; • 
+<specimenCode id="4D889C9F0A9510ABB18D75CC45C6EBE7" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="B72AC141CB2B3A35911AE1A9B0E4BD40" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211485–211490
+</specimenCode>
+, 
+<specimenCount id="0E51115FB7A33C9A16AD7371BD5BCD5B" count="6" type="generic">6 specimens</specimenCount>
+, 
+<quantity id="4CD26D5A2F3B176D196B48886AD62371" metricMagnitude="-2" metricUnit="m" metricValue="5.705" metricValueMax="6.99" metricValueMin="4.42" unit="mm" value="57.05" valueMax="69.9" valueMin="44.2">44.2–69.9 mm</quantity>
+SL from 
+<location id="CBB805EB56EB93E89B83F65703534C9D" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:CBB805EB56EB93E89B83F65703534C9D" county="Yura River" municipality="Kasuga" name="Dainichi River" stateProvince="Hyogo">Dainichi River</location>
+system, 
+<location id="1C15B0F29AD634FA8D3E703DF23EAFBF" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:1C15B0F29AD634FA8D3E703DF23EAFBF" county="Yura River" municipality="Kasuga" name="Jindaiurakabe" stateProvince="Hyogo">Jindaiurakabe</location>
+, 
+<collectingRegion id="5B1204B2107A4A133DA2792EA4A4E19B" country="Japan" name="Hyogo">Hyogo</collectingRegion>
+; • 
+<specimenCode id="42B9927324C160B8FD4ABC6F6BA5A390" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="0D549DCC58638608827362C1387D65C3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211491
+</specimenCode>
+, 
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+, 
+<quantity id="A0728BF36D6402AE2D2057D7F82FB8B4" metricMagnitude="-2" metricUnit="m" metricValue="4.96" unit="mm" value="49.6">49.6 mm</quantity>
+SL from 
+<location id="122AB059571D8A83CF408B302648DB43" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:122AB059571D8A83CF408B302648DB43" county="Yura River" municipality="Kasuga" name="Muko River" stateProvince="Hyogo">Muko River</location>
+system, 
+<location id="6C4C4D3676D2525C9C266B96986DC2F9" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:6C4C4D3676D2525C9C266B96986DC2F9" county="Yura River" municipality="Kasuga" name="Dojyo" stateProvince="Hyogo">Dojyo</location>
+, 
+<collectingRegion id="054B6FA295E26FB2054CE43323D38038" country="Japan" name="Hyogo">Hyogo</collectingRegion>
+; • 
+<specimenCode id="2B73211AC3DCB3177242E8E864031878" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="68FD1A1181A164F2B74408267C92C01D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211492
+</specimenCode>
+, 
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+, 40.0 mm SL from 
+<location id="96EA5552BAE84093058D088B9886A1EA" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:96EA5552BAE84093058D088B9886A1EA" county="Yura River" municipality="Kasuga" name="Yoshii River" stateProvince="Hyogo">Yoshii River</location>
+system, 
+<location id="FDD1B6B80B12CA4C1CB9149CE7566DBC" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:FDD1B6B80B12CA4C1CB9149CE7566DBC" county="Yura River" municipality="Kasuga" name="Nagi" stateProvince="Hyogo">Nagi</location>
+, 
+<collectingRegion id="AD04BBBF4B320520B3B073FB6FD7B7DD" country="Japan" name="Okayama">Okayama</collectingRegion>
+; • 
+<specimenCode id="114637E13BC640FCCEEEDC41B3CBB82A" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="66347515306819B393C8261DB6B245B1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211493–211499
+</specimenCode>
+, 
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+, 39.0–60.0 mm SL from 
+<location id="DC183F91ADE896512D0E826E1C06E142" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:DC183F91ADE896512D0E826E1C06E142" county="Yura River" municipality="Kasuga" name="Maruyama River" stateProvince="Hyogo">Maruyama River</location>
+system, 
+<location id="B87402A6F9939E32FBC2E46D30D371BF" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:B87402A6F9939E32FBC2E46D30D371BF" county="Yura River" municipality="Kasuga" name="Santo" stateProvince="Hyogo">Santo</location>
+, 
+<collectingRegion id="CDA1246074C9EE88556412963EF1B4EE" country="Japan" name="Hyogo">Hyogo</collectingRegion>
+; • 
+<specimenCode id="1B49BD2C2644D4F3935337E2E5F8262F" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="69F4B22772C5CE5E3408C9DADD0852CA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211500
+</specimenCode>
+, 
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+, 
+<quantity id="D52B99A167F2B524431666D1448FE042" metricMagnitude="-2" metricUnit="m" metricValue="4.49" unit="mm" value="44.9">44.9 mm</quantity>
+SL from 
+<location id="C3DC05285C59720BDE45E9C9E6C721F7" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:C3DC05285C59720BDE45E9C9E6C721F7" county="Yura River" municipality="Kasuga" name="Saburi River" stateProvince="Hyogo">Saburi River</location>
+system, 
+<location id="666D0A8399F4CB08E9B5ADBCA6F4352F" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:666D0A8399F4CB08E9B5ADBCA6F4352F" county="Yura River" municipality="Kasuga" name="Yamada" stateProvince="Hyogo">Yamada</location>
+, 
+<collectingRegion id="2043569710980466CDB463BCAD1B7EA3" country="Japan" name="Fukui">Fukui</collectingRegion>
+; • 
+<specimenCode id="2839996122860D05F499E9C8B53ECC7E" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="29AB50F2E0DDD9CE87F4FEE5B8F0DA8B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211501–211506
+</specimenCode>
+, 
+<specimenCount id="4A69491B918386DF5EB512327CFE5B19" count="6" type="generic">6 specimens</specimenCount>
+, 
+<quantity id="3A567BB3CCAA9D957BB57060D19337CF" metricMagnitude="-2" metricUnit="m" metricValue="4.65" metricValueMax="5.74" metricValueMin="3.56" unit="mm" value="46.5" valueMax="57.4" valueMin="35.6">35.6–57.4 mm</quantity>
+SL from 
+<location id="F8FAB69A343B5ABCF50A6EA5DF39F6F1" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:F8FAB69A343B5ABCF50A6EA5DF39F6F1" county="Yura River" municipality="Kasuga" name="Onyu River" stateProvince="Hyogo">Onyu River</location>
+system, 
+<location id="2A187B8A05185B1560D727334D9D1A1B" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:2A187B8A05185B1560D727334D9D1A1B" county="Yura River" municipality="Kasuga" name="Taneji" stateProvince="Hyogo">Taneji</location>
+, 
+<collectingRegion id="400A2D98B9F5D47790CCFCD4B048D173" country="Japan" name="Kyoto">Kyoto</collectingRegion>
+.
+</paragraph>
+<paragraph id="E583711810DCDB85117A2184E36D0CF4">
+<taxonomicName id="AA18250793DCFEDBE0B6F6E4CAB4CE0C" family="Nemacheilidae" genus="Lefua" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="tokaiensis">
+<emphasis id="643BEEE9F9A47050F72812F63BE81B31" italics="true">L. tokaiensis</emphasis>
+</taxonomicName>
+, 
+<specimenCount id="5F0C61880491013141C07046D8DF861F" count="22" type="generic">22 specimens</specimenCount>
+(
+<quantity id="B1B9E0DC8F2A02176BC058F3276D8940" metricMagnitude="-2" metricUnit="m" metricValue="4.685" metricValueMax="6.09" metricValueMin="3.28" unit="mm" value="46.849999999999994" valueMax="60.9" valueMin="32.8">32.8–60.9 mm</quantity>
+SL). 
+<emphasis id="A4F1AD7B81D724F727BFACE71E430040" bold="true" italics="true">
+<typeStatus id="D48F63970F52E9B9B1F145F8D6101FF1" type="holotype">Holotype</typeStatus>
+</emphasis>
+: • 
+<specimenCode id="1813A67FFCF7434DB819B68ABA203CD7" collectionCode="NSMT-P">NSMT-P 132821</specimenCode>
+, 
+<quantity id="434128688EF96DCF62514C2DE0FD1A68" metricMagnitude="-2" metricUnit="m" metricValue="4.42" unit="mm" value="44.2">44.2 mm</quantity>
+SL from 
+<collectingCounty id="8584CB15994BBC0617BA16D76E525412">Toyo River</collectingCounty>
+, 
+<collectingMunicipality id="4ADF0F5EB71816BC7423FAAB757231C8">Kadoya</collectingMunicipality>
+, 
+<collectingRegion id="088634D04BA99E62E0790125B7F721E7" country="Japan" name="Aichi">Aichi</collectingRegion>
+; • 
+<specimenCode id="DD0A07B9B79B2735EA8A7F9F2EFE99EF" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="F52137A09C0160E20DEF00213925ADD5" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211507–211511
+</specimenCode>
+, 
+<specimenCount id="87633D519CC3D6C6A374EE0FB731E449" count="5" type="generic">5 specimens</specimenCount>
+, 
+<quantity id="6368A5D88A699EC14874D9DE41A61FE0" metricMagnitude="-2" metricUnit="m" metricValue="4.205" metricValueMax="5.13" metricValueMin="3.28" unit="mm" value="42.05" valueMax="51.3" valueMin="32.8">32.8–51.3 mm</quantity>
+SL from 
+<location id="57D742E1C3CBD97B6A886FB54E00A08B" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:57D742E1C3CBD97B6A886FB54E00A08B" county="Toyo River" municipality="Kadoya" name="Yahagi River" stateProvince="Aichi">Yahagi River</location>
+system, 
+<location id="8FE7F89CC5C62D24C182EE9756CAE41E" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:8FE7F89CC5C62D24C182EE9756CAE41E" county="Toyo River" municipality="Kadoya" name="Myoken" stateProvince="Aichi">Myoken</location>
+, 
+<collectingRegion id="1D5F14744D4BDDE402485D3FCD963B4B" country="Japan" name="Aichi">Aichi</collectingRegion>
+; • 
+<specimenCode id="76A43100207B5FF1324E597D82467EC0" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="CED3CB7CE925A5A9A66365700DD4CB49" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211512–211518
+</specimenCode>
+, 
+<specimenCount id="1C6466C59973EE2D095EFEFD0BEAA351" count="7" type="generic">7 specimens</specimenCount>
+, 
+<quantity id="0905ADA069A0F843A723718DD457AF79" metricMagnitude="-2" metricUnit="m" metricValue="4.409999999999999" metricValueMax="5.4399999999999995" metricValueMin="3.38" unit="mm" value="44.099999999999994" valueMax="54.4" valueMin="33.8">33.8–54.4 mm</quantity>
+SL from 
+<location id="DF763D847191C9F8038775E53A281BD2" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:DF763D847191C9F8038775E53A281BD2" county="Toyo River" municipality="Kadoya" name="Toyo River" stateProvince="Aichi">Toyo River</location>
+system, 
+<location id="9B6946A849EEE26424CA40BC4A80B018" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:9B6946A849EEE26424CA40BC4A80B018" county="Toyo River" municipality="Kadoya" name="Kadoya" stateProvince="Aichi">Kadoya</location>
+, 
+<collectingRegion id="3EBFC1F62D938DA6562CEE34316E8487" country="Japan" name="Aichi">Aichi</collectingRegion>
+; • 
+<specimenCode id="EB54C1217245B20135B53F3B7F69CECC" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="DA1EDB3AAB692FFB677107EB5E8092A9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211519–211526
+</specimenCode>
+, 
+<specimenCount id="0287B3788360005CB048B085EC3611F2" count="8" type="generic">8 specimens</specimenCount>
+, 40.7–60.0 mm SL from 
+<location id="1B7E2805729E52F207CDCB2128D715D5" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:1B7E2805729E52F207CDCB2128D715D5" county="Toyo River" municipality="Kadoya" name="Miyakoda River" stateProvince="Aichi">Miyakoda River</location>
+system, 
+<location id="E243F6743C986B3663E868C0DF3555A4" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:E243F6743C986B3663E868C0DF3555A4" county="Toyo River" municipality="Kadoya" name="Hosoe" stateProvince="Aichi">Hosoe</location>
+, 
+<collectingRegion id="99C76BABB81304ACDC87AC8F0CE3FD32" country="Japan" name="Shizuoka">Shizuoka</collectingRegion>
+; • 
+<specimenCode id="BFFB2D1DE9D494F8C0AAF2CF65148F76" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" type="">
+<named-content id="C02FB20C0B18A882FF1D0DCE337C50BF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kyoto-university" xlink_title="Department of Fisheries, Faculty of Agriculture, Kyoto University">FAKU</named-content>
+211527
+</specimenCode>
+, 
+<specimenCount id="09FC9966F61E34EE005E834E3030244F" count="1" type="generic">1 specimen</specimenCount>
+, 
+<quantity id="05CC0AA135C51A585D117CFEEE660F5D" metricMagnitude="-2" metricUnit="m" metricValue="3.6799999999999997" unit="mm" value="36.8">36.8 mm</quantity>
+SL from 
+<location id="93306318552CC4FD6D69C613559FDAFD" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:93306318552CC4FD6D69C613559FDAFD" county="Toyo River" municipality="Kadoya" name="Ota River" stateProvince="Aichi">Ota River</location>
+system, 
+<location id="88A8B4C911542910FE65610E6DC929CB" LSID="urn:lsid:plazi:treatment:29DCCB12F4F85E91AFC577CE36A02436:88A8B4C911542910FE65610E6DC929CB" county="Toyo River" municipality="Kadoya" name="Ota" stateProvince="Aichi">Ota</location>
+, 
+<collectingRegion id="58E71BDD5FF807C51908D5F84B0654AB" country="Japan" name="Shizuoka">Shizuoka</collectingRegion>
+.
+</paragraph>
+</subSubSection>
+</treatment>
+</document>
\ No newline at end of file