plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity

Add updates up until 2025-03-04 01:01:06

Browse source
This commit is contained in:
ggserver 2025-03-04 01:03:25 +00:00
parent 98c033bbbe
commit 68acd6b219
2 changed files with 437 additions and 0 deletions

279
data/03/DC/87/03DC8796FF8C666195A5F98A074AFDF1.xml Normal file
View file

@ -0,0 +1,279 @@
<document id="F735C22CC361CC70CFBA69834693E522" ID-DOI="10.11646/zootaxa.5588.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14896574" ID-ZooBank="E3697AAB-FB6D-440B-89C7-3AD757157481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1740004092971" checkinUser="plazi" docAuthor="Bao, Liang, Beutel, Rolf Georg, Niu, Kecheng &amp; Bao, Tong" docDate="2025" docId="03DC8796FF8C666195A5F98A074AFDF1" docLanguage="en" docName="zootaxa.5588.3.1.pdf" docOrigin="Zootaxa 5588 (3)" docSource="https://doi.org/10.11646/zootaxa.5588.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tigrivia baii Bao 2023" docType="treatment" docVersion="1" lastPageNumber="407" masterDocId="FFE5FFEEFF8E66679532FFDA0639FF90" masterDocTitle="Redescription of the highly specialized aquatic † Tigrivia and the classification of the Mesozoic † Coptoclavidae (Coleoptera, Adephaga, Dytiscoidea)" masterLastPageNumber="425" masterPageNumber="401" pageNumber="403" updateTime="1741050061580" updateUser="jonas" zenodo-license-document="CLOSED">
<mods:mods id="DDE68E9A84F5EFCE2F2530DC31F1A539" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AC83DFEDD50A739803474E6693A44050">
<mods:title id="81298C3AF7D48F10DCC1AB013AFAD795">Redescription of the highly specialized aquatic † Tigrivia and the classification of the Mesozoic † Coptoclavidae (Coleoptera, Adephaga, Dytiscoidea)</mods:title>
</mods:titleInfo>
<mods:name id="89B34A2D93304112C586CCF77A9618B2" type="personal">
<mods:role id="D94C0E49B579E1257E56C3D9FA5ED904">
<mods:roleTerm id="0CC157B9EB48EE67428B72C28CD9BC22">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="74B3590D7BACEA98CA7D5FCAD58EFFB7">Bao, Liang</mods:namePart>
<mods:affiliation id="7135685F9096C0E83D1562EEE05A0BE1">School of Earth Sciences and Engineering, Guangdong Provincial Key Lab of Geological Processes and Mineral Resources, Sun Yat-sen University, Guangzhou 510275, China. &amp; School of Ecology / State Key Laboratory of Biocontrol, Shenzhen Campus of Sun Yat-sen University, Shenzhen 518107, China.</mods:affiliation>
</mods:name>
<mods:name id="E66F49AC4778D1648168E9C323CF5488" type="personal">
<mods:role id="3648B57664633B12E95908B2CB096D7D">
<mods:roleTerm id="F744F4E8A81768217EAE3D8E8D92F043">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B5078C1597D930C454109AC8CE42F7C9">Beutel, Rolf Georg</mods:namePart>
<mods:affiliation id="6330B213DBFF22891BDF0D973430838C">Institut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität Jena, Jena 07743, Germany.</mods:affiliation>
</mods:name>
<mods:name id="CF80C60038809B53EED394C9732008E1" type="personal">
<mods:role id="4C7B0736890D704C9CDA8927830CD64E">
<mods:roleTerm id="01F59646185FCA631AAA8B8818ACB24C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CD482B6A20DFBB3D9EE94815CB1A4BC4">Niu, Kecheng</mods:namePart>
<mods:affiliation id="5F0CFE9842C3CEA2C9F20154F4FAA82C">State Key Laboratory of Cellular Stress Biology, School of Life Sciences, Xiamen University, Xiamen 361105, China. &amp; Yingliang Stone Natural History Museum, Nan’an 362300, China.</mods:affiliation>
</mods:name>
<mods:name id="50A91465424985D2C39BC0E809C4B816" type="personal">
<mods:role id="400782506EAA3B4D92761F3CCA1BA087">
<mods:roleTerm id="9C3B443FA8F166F3106ECE6F7C3B1486">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B201EC2E1FCF9B8DC8D7783D10044F9B">Bao, Tong</mods:namePart>
<mods:affiliation id="B343CC677C11E023BC2F09CAE024DF39">School of Ecology / State Key Laboratory of Biocontrol, Shenzhen Campus of Sun Yat-sen University, Shenzhen 518107, China. &amp; Yunnan Key Laboratory for Palaeobiology, Yunnan University, Kunming 650091, China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E92A3BC6675E38AB7E373B6A94506E51">text</mods:typeOfResource>
<mods:relatedItem id="FC4E3ACF0D574AFC6327FDED82B13387" type="host">
<mods:titleInfo id="43120FA975480E10791E7A00D0E177BC">
<mods:title id="5256664658F35AFBDB0A79937F19E7F0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A0E821386B40082382B366797A740AB2">
<mods:date id="EA704F0ED7BEF0A61FA9849FCD713E95">2025</mods:date>
<mods:detail id="22CAA7342255EAF2A12CB18908071C6F" type="pubDate">
<mods:number id="36F588A4748C3AFF1168BE0102F6F54D">2025-02-17</mods:number>
</mods:detail>
<mods:detail id="8D16B3FCD43A7DB318876A193435695D" type="volume">
<mods:number id="3136E1B4B59F6453FE7B26FDE4A7C24F">5588</mods:number>
</mods:detail>
<mods:detail id="E0B8E485D924D81B4A870528B965D0DB" type="issue">
<mods:number id="B2AF3F4E6B4347552FE1A0F52D578A6C">3</mods:number>
</mods:detail>
<mods:extent id="EF88F4FCEB5DEE87CEF572B1C9EA892B" unit="page">
<mods:start id="C6C4E0437483E995F329A29A2816DDE1">401</mods:start>
<mods:end id="F5A2102AA169B9F45D24647405025869">425</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="1DFD7DBF6B10B841D9F9E99C000288B7">
<mods:url id="80C5EE7C0FE198D75FBA2BC4E9B63F97">https://doi.org/10.11646/zootaxa.5588.3.1</mods:url>
</mods:location>
<mods:classification id="0BC7FE134742C44628542DC8E67F43BD">journal article</mods:classification>
<mods:identifier id="2EB919CFC3E2C5BC4BBAD33908A344C1" type="DOI">10.11646/zootaxa.5588.3.1</mods:identifier>
<mods:identifier id="B7B9AEB04D3F7ABBAAB11A9A70CAD317" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0BB4EAE4872B00C113DB596175A64933" type="Zenodo-Dep">14896574</mods:identifier>
<mods:identifier id="6B603A5DD44996A56CC51445CDFE1F58" type="ZooBank">E3697AAB-FB6D-440B-89C7-3AD757157481</mods:identifier>
</mods:mods>
<treatment id="03DC8796FF8C666195A5F98A074AFDF1" LSID="urn:lsid:plazi:treatment:03DC8796FF8C666195A5F98A074AFDF1" httpUri="http://treatment.plazi.org/id/03DC8796FF8C666195A5F98A074AFDF1" lastPageId="6" lastPageNumber="407" pageId="2" pageNumber="403">
<subSubSection id="C36F650BFF8C666595A5F98A07C7F9FA" box="[151,510,1615,1642]" pageId="2" pageNumber="403" type="nomenclature">
<paragraph id="8BCA3680FF8C666595A5F98A07C7F9FA" blockId="2.[151,962,1615,1679]" box="[151,510,1615,1642]" pageId="2" pageNumber="403">
<heading id="D08281ECFF8C666595A5F98A07C7F9FA" bold="true" box="[151,510,1615,1642]" fontSize="11" level="1" pageId="2" pageNumber="403" reason="1">
<emphasis id="B901EA92FF8C666595A5F98A07C7F9FA" bold="true" box="[151,510,1615,1642]" pageId="2" pageNumber="403">
†
<taxonomicName id="4C754D03FF8C66659594F98A07C7F9FA" authority="Bao et al., 2023" authorityName="Bao" authorityYear="2023" box="[166,510,1615,1642]" class="Insecta" family="Coptoclavidae" genus="Tigrivia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="baii">
<emphasis id="B901EA92FF8C66659594F98A0703F9FA" bold="true" box="[166,314,1615,1642]" italics="true" pageId="2" pageNumber="403">Tigrivia baii</emphasis>
<bibRefCitation id="EFE44B71FF8C66659473F98A07C7F9FA" author="Bao, L. &amp; Li, L. &amp; Niu, K. &amp; Wang, N. &amp; Kroeck, D. M. &amp; Bao, T." box="[321,510,1615,1642]" pageId="2" pageNumber="403" pagination="1 - E 6" refId="ref7184" refString="Bao, L., Li, L., Niu, K., Wang, N., Kroeck, D. M. &amp; Bao, T. (2023) Retracted: A new aquatic beetle (Adephaga: Coptoclavidae) from the Middle Jurassic Daohugou Biota. The Anatomical Record, 306 (8), E 1-E6. https://doi.org/10.1002/ar.25221" type="journal article" year="2023">
Bao
<emphasis id="B901EA92FF8C6665944BF98A078DF9FA" bold="true" box="[377,436,1615,1642]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
, 2023
</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F650BFF8C666595A5F9AE05FBF91F" box="[151,962,1652,1679]" pageId="2" pageNumber="403" type="description">
<paragraph id="8BCA3680FF8C666595A5F9AE05FBF91F" blockId="2.[151,962,1615,1679]" box="[151,962,1652,1679]" pageId="2" pageNumber="403">
<uri id="FFE43A82FF8C666595A5F9AE05FBF91F" box="[151,962,1652,1679]" pageId="2" pageNumber="403">
urn:lsid:zoobank.org:act:
<uuid id="FFD30C55FF8C66659483F9AE05FBF91F" box="[433,962,1652,1679]" pageId="2" pageNumber="403">19BDD6A1-C747-4995-8526-CC3F6751B1F1</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C36F650BFF8C666595A5F961023DF946" box="[151,1028,1723,1751]" pageId="2" pageNumber="403" type="etymology">
<paragraph id="8BCA3680FF8C666595A5F961023DF946" blockId="2.[151,1436,1723,1966]" box="[151,1028,1723,1751]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595A5F9610720F946" bold="true" box="[151,281,1723,1750]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595A5F961072BF946" bold="true" box="[151,274,1723,1750]" italics="true" pageId="2" pageNumber="403">Etymology</emphasis>
.
</emphasis>
The name paid homage to Bai, a fossil collector (
<bibRefCitation id="EFE44B71FF8C66659675F96605C1F946" author="Bao, L. &amp; Li, L. &amp; Niu, K. &amp; Wang, N. &amp; Kroeck, D. M. &amp; Bao, T." box="[839,1016,1724,1750]" pageId="2" pageNumber="403" pagination="1 - E 6" refId="ref7184" refString="Bao, L., Li, L., Niu, K., Wang, N., Kroeck, D. M. &amp; Bao, T. (2023) Retracted: A new aquatic beetle (Adephaga: Coptoclavidae) from the Middle Jurassic Daohugou Biota. The Anatomical Record, 306 (8), E 1-E6. https://doi.org/10.1002/ar.25221" type="journal article" year="2023">
Bao
<emphasis id="B901EA92FF8C6665964EF967058CF946" box="[892,949,1724,1750]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36F650BFF8C666595F5F93A07EEF96B" box="[199,471,1760,1787]" pageId="2" pageNumber="403" type="diagnosis">
<paragraph id="8BCA3680FF8C666595F5F93A07EEF96B" blockId="2.[151,1436,1723,1966]" box="[199,471,1760,1787]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595F5F93A0778F96A" bold="true" box="[199,321,1760,1786]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595F5F93A0703F96A" bold="true" box="[199,314,1760,1786]" italics="true" pageId="2" pageNumber="403">Diagnosis</emphasis>
.
</emphasis>
as for genus.
</paragraph>
</subSubSection>
<subSubSection id="C36F650BFF8C666595F5F8DE0547F8D2" pageId="2" pageNumber="403" type="materials_examined">
<paragraph id="8BCA3680FF8C666595F5F8DE0547F8D2" blockId="2.[151,1436,1723,1966]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595F5F8DE055BF88E" bold="true" box="[199,866,1795,1822]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595F5F8DE0562F88E" bold="true" box="[199,859,1795,1822]" italics="true" pageId="2" pageNumber="403">Type material used here and in the original description</emphasis>
.
</emphasis>
<materialsCitation id="3B1D3CDDFF8C6665965EF8DE0540F8D2" collectionCode="YLSNHM" pageId="2" pageNumber="403" specimenCode="YLSNHM01120" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54CE8822FF8C6665965EF8DE05E0F88E" box="[876,985,1796,1822]" pageId="2" pageNumber="403" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBD39EFBFF8C666596D7F8DE0298F88E" box="[997,1185,1796,1822]" pageId="2" pageNumber="403">YLSNHM01120</specimenCode>
/
<emphasis id="B901EA92FF8C6665918FF8DF0375F88E" box="[1213,1356,1796,1822]" italics="true" pageId="2" pageNumber="403">Tigrivia baii</emphasis>
;
<specimenCount id="9D73FD09FF8C6665906FF8DE03A1F88E" box="[1373,1432,1796,1822]" pageId="2" pageNumber="403" type="adult">adult</specimenCount>
,
<specimenCount id="9D73FD09FF8C666595A5F8F206DAF8D2" box="[151,227,1832,1858]" pageId="2" pageNumber="403" type="female">female</specimenCount>
(visible female gonocoxosterna), well preserved specimen
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F650BFF8C666595F5F8960548F83E" pageId="2" pageNumber="403" type="biology_ecology">
<paragraph id="8BCA3680FF8C666595F5F89604CEF8F6" blockId="2.[151,1436,1723,1966]" box="[199,759,1867,1894]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595F5F8960411F8F6" bold="true" box="[199,552,1867,1894]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595F5F8960418F8F6" bold="true" box="[199,545,1867,1894]" italics="true" pageId="2" pageNumber="403">
<typeStatus id="54CE8822FF8C666595F5F89606C7F8F6" box="[199,254,1868,1894]" pageId="2" pageNumber="403">Type</typeStatus>
stratum and
<typeStatus id="54CE8822FF8C666594A5F89607FFF8F6" box="[407,454,1868,1894]" pageId="2" pageNumber="403">type</typeStatus>
locality
</emphasis>
.
</emphasis>
Impression fossil.
</paragraph>
<paragraph id="8BCA3680FF8C666595F5F8AA0548F83E" blockId="2.[151,1436,1723,1966]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595F5F8AA0413F81A" bold="true" box="[199,554,1903,1930]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595F5F8AA041AF81A" bold="true" box="[199,547,1903,1930]" italics="true" pageId="2" pageNumber="403">
<typeStatus id="54CE8822FF8C666595F5F8AA06C7F81A" box="[199,254,1904,1930]" pageId="2" pageNumber="403">Type</typeStatus>
stratum and
<typeStatus id="54CE8822FF8C666594ABF8AA07F1F81A" box="[409,456,1904,1930]" pageId="2" pageNumber="403">type</typeStatus>
locality
</emphasis>
.
</emphasis>
<location id="8EAA605BFF8C66659703F8AA0494F81A" LSID="urn:lsid:plazi:treatment:03DC8796FF8C666195A5F98A074AFDF1:8EAA605BFF8C66659703F8AA0494F81A" box="[561,685,1904,1930]" country="China" county="Weichang County" municipality="Daobaliang mountain" name="Impression" pageId="2" pageNumber="403" stateProvince="Hebei">Impression</location>
fossil.
<location id="8EAA605BFF8C66659632F8AA05DEF81A" LSID="urn:lsid:plazi:treatment:03DC8796FF8C666195A5F98A074AFDF1:8EAA605BFF8C66659632F8AA05DEF81A" box="[768,999,1904,1930]" country="China" county="Weichang County" municipality="Daobaliang mountain" name="Jiufotang Formation" pageId="2" pageNumber="403" stateProvince="Hebei">Jiufotang Formation</location>
,
<location id="8EAA605BFF8C666596C1F8AA0209F81A" LSID="urn:lsid:plazi:treatment:03DC8796FF8C666195A5F98A074AFDF1:8EAA605BFF8C666596C1F8AA0209F81A" box="[1011,1072,1904,1930]" country="China" county="Weichang County" municipality="Daobaliang mountain" name="Early" pageId="2" pageNumber="403" stateProvince="Hebei">Early</location>
<taxonomicName id="4C754D03FF8C66659105F8AA03A1F81A" authority="(Chang et al. 2009)" baseAuthorityName="Chang" baseAuthorityYear="2009" box="[1079,1432,1904,1930]" class="Insecta" family="Scarabaeidae" genus="Cretaceous" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
Cretaceous (
<bibRefCitation id="EFE44B71FF8C666591F0F8AA03A9F81A" author="Chang, S. &amp; Zhang, H. &amp; Renne, P. &amp; Fang, Y." box="[1218,1424,1904,1930]" pageId="2" pageNumber="403" pagination="94 - 104" refId="ref8704" refString="Chang, S., Zhang, H., Renne, P. &amp; Fang, Y. (2009) High - precision 40 Ar / 39 Ar age for the Jehol Biota. Palaeogeography, Palaeoclimatology, Palaeoecology, 280 (1 - 2), 94-104. https://doi.org/10.1016/j.palaeo.2009.06.021" type="journal article" year="2009">
Chang
<emphasis id="B901EA92FF8C66659026F8AB0374F81A" box="[1300,1357,1904,1930]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
2009
</bibRefCitation>
)
</taxonomicName>
,
<collectingMunicipality id="6BAEACFAFF8C666595A5F84E07B5F83E" box="[151,396,1940,1966]" pageId="2" pageNumber="403">Daobaliang mountain</collectingMunicipality>
,
<collectingCounty id="62AB4E0CFF8C666594A5F84E0458F83E" box="[407,609,1940,1966]" pageId="2" pageNumber="403">Weichang County</collectingCounty>
,
<collectingRegion id="49B1F862FF8C66659759F84E0522F83E" box="[619,795,1940,1966]" country="China" name="Hebei" pageId="2" pageNumber="403">Hebei Province</collectingRegion>
,
<collectingCountry id="F3627610FF8C66659615F84E0557F83E" box="[807,878,1940,1966]" name="China" pageId="2" pageNumber="403">China</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF0A6608FF8D666495A5F8700346F852" ID-DOI="http://doi.org/10.5281/zenodo.14896576" ID-Zenodo-Dep="14896576" box="[151,1407,1962,1987]" httpUri="https://zenodo.org/record/14896576/files/figure.png" pageId="3" pageNumber="404" startId="3.[151,250,1962,1986]" targetBox="[162,1425,198,1936]" targetPageId="3" targetType="figure">
<paragraph id="8BCA3680FF8D666495A5F8700346F852" blockId="3.[151,1407,1962,1987]" box="[151,1407,1962,1987]" pageId="3" pageNumber="404">
<emphasis id="B901EA92FF8D666495A5F870072AF853" bold="true" box="[151,275,1962,1987]" pageId="3" pageNumber="404">FIGURE 1.</emphasis>
Photographs of
<emphasis id="B901EA92FF8D66649488F871040EF853" box="[442,567,1963,1987]" italics="true" pageId="3" pageNumber="404">Tigrivia baii</emphasis>
, holotype, YLSNHM01120, (A) dorsal view; (B) ventral view. Scale bars 3.5 mm.
</paragraph>
</caption>
<subSubSection id="C36F650BFF8A666195F5FF4C074AFDF1" lastPageId="6" lastPageNumber="407" pageId="4" pageNumber="405" type="description">
<paragraph id="8BCA3680FF8A666395F5FF4C0341FF45" blockId="4.[151,1437,150,1293]" pageId="4" pageNumber="405">
<emphasis id="B901EA92FF8A666395F5FF4C07D1FF20" bold="true" box="[199,488,150,177]" pageId="4" pageNumber="405">
<emphasis id="B901EA92FF8A666395F5FF4C075FFF21" bold="true" box="[199,358,150,177]" italics="true" pageId="4" pageNumber="405">Redescription</emphasis>
(
<figureCitation id="134E2A05FF8A66639446FF4C07E0FF20" box="[372,473,150,176]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1962,1986]" captionStartId-1="4.[151,250,1909,1933]" captionStartId-2="5.[151,250,1114,1138]" captionTargetBox-0="[162,1425,198,1936]" captionTargetBox-1="[286,1303,1320,1869]" captionTargetBox-2="[159,1423,184,1082]" captionTargetId-0="figure-18@3.[162,1425,198,1936]" captionTargetId-1="figure-482@4.[280,1308,1314,1883]" captionTargetId-2="figure-18@5.[151,1436,179,1088]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Photographs of Tigrivia baii, holotype, YLSNHM01120, (A) dorsal view; (B) ventral view. Scale bars 3.5 mm." captionText-1="FIGURE 2. Line drawing of Tigrivia baii, based on holotype, ventral view. Abbreviations: aes1, mesanepisternum; aes3, metanepisternum; cx, coxa; cxpl,coxal plate;ep, episternum;fe, femur;gcxs, gonocoxosterna; gu, gula; kes3, metakatepisternum; lbr, labrum; md, mandible; mt, mentum; pl, propleuron; pmt, prementum; pst, prosternum; pta, protarsi; scu2, mesoscutum; st, sternite; ti1, tibia1; tisp, tibial spur; tr, trochantin; v2, mesoventrite; v3, metaventrite." captionText-2="FIGURE 3. Reconstruction of Tigrivia baii, based on holotype and drawing, (A) ventral view; (B) dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.14896576" figureDoi-1="http://doi.org/10.5281/zenodo.14896578" figureDoi-2="http://doi.org/10.5281/zenodo.14896580" httpUri-0="https://zenodo.org/record/14896576/files/figure.png" httpUri-1="https://zenodo.org/record/14896578/files/figure.png" httpUri-2="https://zenodo.org/record/14896580/files/figure.png" pageId="4" pageNumber="405">Figs 1–3</figureCitation>
).
</emphasis>
Body oval, posteriorly widening. Length
<quantity id="4C8D9B65FF8A6663968BFF4D0233FF21" box="[953,1034,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="405" unit="mm" value="30.0">30 mm</quantity>
; maximum width ca. 12.5 mm; ratio of length to maximum width 2.38. Elytra posteriorly truncated, leaving posterior abdominal structures exposed.
</paragraph>
<paragraph id="8BCA3680FF8A666395F5FF050358FE65" blockId="4.[151,1437,150,1293]" pageId="4" pageNumber="405">
Head prognathous, widening posteriorly, strongly retracted into prothorax, especially dorsally. Length of exposed part 3.7 mm, maximum width
<quantity id="4C8D9B65FF8A66639761FED904A1FE8D" box="[595,664,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="405" unit="mm" value="4.0">4 mm</quantity>
. Laterally placed compound eyes large, round, diameter ca. 2.4 mm, slightly protruding; dorsally and ventrally visible portions of about equal size, not separated by a chitinous bridge, compound eyes thus undivided; hind margin almost adjacent with anterior pronotal margin. Labrum transverse, anterolaterally rounded, medially with shallow, rounded emargination (visible in ventral view). Antennae not recognizable. Mandibles comparatively short and robust, partly exposed, distally slightly curved inwards. Maxillae not clearly recognizable; maxillary palps not visible. Labium with distinct, rounded lateral lobes of mentum enclosing large prementum, labial palps not recognizable. Gula delimited by distinct gular ridges (visible on right side).
</paragraph>
<paragraph id="8BCA3680FF8A666395F5FE250796FA9D" blockId="4.[151,1437,150,1293]" pageId="4" pageNumber="405">Prothorax small in relation, to pterothoracic segments. Pronotum trapezoid, with anteriorly converging, very slightly convex lateral edges; length 4.3 mm, width of anterior margin 5.3 mm, width of posterior margin 6.3 mm; anterior pronotal margin very slightly concave, slightly wider than occipital region of head, distinctly overlapping with it; posterior margin distinctly protruding between elytral bases, rounded posteromedially. Prosternum very short, transverse; length 1.0 mm, width 3.0 mm.Prosternal process almost completely reduced, present only as small triangle projecting slightly between procoxae. Sternopleural suture not visible. Propleura large, slightly widening laterally; suture separating it from pronotal hypomeron not visible; protrochantin also not recognizable. Mesoscutellar shield small but distinctly visible, triangular. Ventral side of thorax strongly disrupted between procoxae and mesoventrite; mesonotum visible in wide gap as exposed skeletal element, with anteromedian prominence with two rounded tips separated by round median incision, pair of distinct lateral processes, and quadrangular posterior part, separated from remaining structure by distinct transverse line. Mesoventrite visible as well-developed transverse and largely flat plate-like sclerite anterior to large mesocoxae; length 3.0 mm, width 7.0 mm; hexagonal groove for reception of prosternal process absent; long median discrimen apparently present, visible as deep median impression; transverse anterolateral delimited areas of mesoventrite distinct, probably concave and functioning as procoxal and protrochanteral rests. Mesopleuron laterally adjacent with mesoventrite; anepisternum roughly triangular; pleural suture recognizable as faintly impressed line; posterior border of mesepimeron only partly visible. Metathorax distinctly larger than mesothorax. Metaventrite short and wide, 3.4 times as wide as long, but not reaching elytral epipleura and anterolateral metacoxal edge laterally; rounded anteromedian process between posterior mesocoxal margins present but short; discrimen of metaventrite largely absent or not recognizable externally, only visible on katepisternal region anterior to metacoxae. Very short katepisternum delimited by faintly visible transverse suture, ending at about 1/3 of metacoxal width. Metanepisternum elongate, fairly narrow, widening anteriorly; metepimeron not visible.</paragraph>
<caption id="DF0A6608FF8A666395A5F8AF05CDF869" ID-DOI="http://doi.org/10.5281/zenodo.14896578" ID-Zenodo-Dep="14896578" httpUri="https://zenodo.org/record/14896578/files/figure.png" pageId="4" pageNumber="405" startId="4.[151,250,1909,1933]" targetBox="[286,1303,1320,1869]" targetPageId="4" targetType="figure">
<paragraph id="8BCA3680FF8A666395A5F8AF05CDF869" blockId="4.[151,1437,1909,2041]" pageId="4" pageNumber="405">
<emphasis id="B901EA92FF8A666395A5F8AF072EF81D" bold="true" box="[151,279,1909,1933]" pageId="4" pageNumber="405">FIGURE 2.</emphasis>
Line drawing of
<emphasis id="B901EA92FF8A666394E4F8AC0461F81D" box="[470,600,1909,1933]" italics="true" pageId="4" pageNumber="405">Tigrivia baii</emphasis>
, based on holotype, ventral view. Abbreviations: aes1, mesanepisternum; aes3, metanepisternum; cx, coxa; cxpl, coxal plate; ep, episternum; fe, femur; gcxs, gonocoxosterna; gu, gula; kes3, metakatepisternum; lbr, labrum; md, mandible; mt, mentum; pl, propleuron; pmt, prementum; pst, prosternum; pta, protarsi; scu2, mesoscutum; st, sternite; ti1, tibia1; tisp, tibial spur; tr, trochantin; v2, mesoventrite; v3, metaventrite.
</paragraph>
</caption>
<caption id="DF0A6608FF8B666295A5FB800339FBE2" ID-DOI="http://doi.org/10.5281/zenodo.14896580" ID-Zenodo-Dep="14896580" box="[151,1280,1114,1138]" httpUri="https://zenodo.org/record/14896580/files/figure.png" pageId="5" pageNumber="406" startId="5.[151,250,1114,1138]" targetBox="[159,1423,184,1082]" targetPageId="5" targetType="figure">
<paragraph id="8BCA3680FF8B666295A5FB800339FBE2" blockId="5.[151,1280,1114,1138]" box="[151,1280,1114,1138]" pageId="5" pageNumber="406">
<emphasis id="B901EA92FF8B666295A5FB80072AFBE2" bold="true" box="[151,275,1114,1138]" pageId="5" pageNumber="406">FIGURE 3.</emphasis>
Reconstruction of
<emphasis id="B901EA92FF8B666294E7FB81046BFBE2" box="[469,594,1114,1138]" italics="true" pageId="5" pageNumber="406">Tigrivia baii</emphasis>
, based on holotype and drawing, (A) ventral view; (B) dorsal view.
</paragraph>
</caption>
<paragraph id="8BCA3680FF8B666295F5FB1E04A5F842" blockId="5.[151,1437,1220,2003]" pageId="5" pageNumber="406">
Tarsal formula 5–5–5 (
<figureCitation id="134E2A05FF8B666294FDFB1E0422FB4E" box="[463,539,1220,1246]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[286,1303,1320,1869]" captionTargetId="figure-482@4.[280,1308,1314,1883]" captionTargetPageId="4" captionText="FIGURE 2. Line drawing of Tigrivia baii, based on holotype, ventral view. Abbreviations: aes1, mesanepisternum; aes3, metanepisternum; cx, coxa; cxpl,coxal plate;ep, episternum;fe, femur;gcxs, gonocoxosterna; gu, gula; kes3, metakatepisternum; lbr, labrum; md, mandible; mt, mentum; pl, propleuron; pmt, prementum; pst, prosternum; pta, protarsi; scu2, mesoscutum; st, sternite; ti1, tibia1; tisp, tibial spur; tr, trochantin; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.14896578" httpUri="https://zenodo.org/record/14896578/files/figure.png" pageId="5" pageNumber="406">Figs 2</figureCitation>
,
<figureCitation id="134E2A05FF8B66629718FB1E0469FB4E" box="[554,592,1220,1246]" captionStart="FIGURE 3" captionStartId="5.[151,250,1114,1138]" captionTargetBox="[159,1423,184,1082]" captionTargetId="figure-18@5.[151,1436,179,1088]" captionTargetPageId="5" captionText="FIGURE 3. Reconstruction of Tigrivia baii, based on holotype and drawing, (A) ventral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14896580" httpUri="https://zenodo.org/record/14896580/files/figure.png" pageId="5" pageNumber="406">3A</figureCitation>
). Forelegs strongly elongated, raptorial. Procoxa moderately sized, wider than long; rounded anteriorly, laterally, and posterolaterally, but with straight, medially adjacent mesal margins. Protrochanter distinctly smaller than meso- and metathoracic equivalents. Profemur elongated, 5.9 mm long, very slightly widening distally. Protibia elongated and slender, slightly narrowed in middle region, 5.8 mm long; with single strongly developed mesal spur on apex. Protarsus elongate; length formula 1.1–1.4–0.75–0.5–1.4 mm; protarsomere 1 moderately widening distally, likely equipped with curved spur (visible on left side); large protarsomere 2 distinctly widening distally; protarsomeres 3 and 4 small; protarsomere 5 elongate and slender; apically one claw visible (possibly an artefact). Middle legs and hind legs long, with distal parts moderately flattened and broadened, modified as paddle-like swimming legs, more distinctly on hind legs (
<figureCitation id="134E2A05FF8B66629155FA3E02FAFA6E" box="[1127,1219,1508,1534]" captionStart="FIGURE 1" captionStartId="3.[151,250,1962,1986]" captionTargetBox="[162,1425,198,1936]" captionTargetId="figure-18@3.[162,1425,198,1936]" captionTargetPageId="3" captionText="FIGURE 1. Photographs of Tigrivia baii, holotype, YLSNHM01120, (A) dorsal view; (B) ventral view. Scale bars 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14896576" httpUri="https://zenodo.org/record/14896576/files/figure.png" pageId="5" pageNumber="406">Figs 1B</figureCitation>
,
<figureCitation id="134E2A05FF8B666291E3FA3E02D8FA6E" box="[1233,1249,1508,1534]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[286,1303,1320,1869]" captionTargetId="figure-482@4.[280,1308,1314,1883]" captionTargetPageId="4" captionText="FIGURE 2. Line drawing of Tigrivia baii, based on holotype, ventral view. Abbreviations: aes1, mesanepisternum; aes3, metanepisternum; cx, coxa; cxpl,coxal plate;ep, episternum;fe, femur;gcxs, gonocoxosterna; gu, gula; kes3, metakatepisternum; lbr, labrum; md, mandible; mt, mentum; pl, propleuron; pmt, prementum; pst, prosternum; pta, protarsi; scu2, mesoscutum; st, sternite; ti1, tibia1; tisp, tibial spur; tr, trochantin; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.14896578" httpUri="https://zenodo.org/record/14896578/files/figure.png" pageId="5" pageNumber="406">2</figureCitation>
,
<figureCitation id="134E2A05FF8B666291DCFA3E032DFA6E" box="[1262,1300,1508,1534]" captionStart="FIGURE 3" captionStartId="5.[151,250,1114,1138]" captionTargetBox="[159,1423,184,1082]" captionTargetId="figure-18@5.[151,1436,179,1088]" captionTargetPageId="5" captionText="FIGURE 3. Reconstruction of Tigrivia baii, based on holotype and drawing, (A) ventral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14896580" httpUri="https://zenodo.org/record/14896580/files/figure.png" pageId="5" pageNumber="406">3A</figureCitation>
). Mesocoxa distinctly larger than procoxa, rounded to triangular, with nearly straight and adjacent mesal walls. Mesotrochantin not recognizable. Mesotrochanter distinctly larger than protrochanter. Mesofemur 6.1 mm long, slightly longer than profemur but distinctly broader, and widening distally. Mesotibiae 5.0 mm long, slightly shorter and narrower than femur, also widening distally. Mesotarsus broadened, especially mesotarsomere 1; length formula 2–0.5–0.7–0.4– 1.2 mm; distal tarsomeres only indistinctly visible on left side. Distal part of hind legs similar to corresponding parts of middle legs, but tarsus distinctly larger, apparently functioning as main propulsive organ. Metacoxa transverse, reaching elytral epipleuron laterally, slightly broader than posterior edge of metaventrite; length
<quantity id="4C8D9B65FF8B666291F1F93A033CF96B" box="[1219,1285,1760,1787]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="406" unit="mm" value="3.0">3 mm</quantity>
, width
<quantity id="4C8D9B65FF8B66629067F93A03A1F96B" box="[1365,1432,1760,1787]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="406" unit="mm" value="5.0">5 mm</quantity>
; posteromedian part covering part of abdominal sternite III (
<figureCitation id="134E2A05FF8B6662960AF8DE05B8F88E" box="[824,897,1796,1822]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[286,1303,1320,1869]" captionTargetId="figure-482@4.[280,1308,1314,1883]" captionTargetPageId="4" captionText="FIGURE 2. Line drawing of Tigrivia baii, based on holotype, ventral view. Abbreviations: aes1, mesanepisternum; aes3, metanepisternum; cx, coxa; cxpl,coxal plate;ep, episternum;fe, femur;gcxs, gonocoxosterna; gu, gula; kes3, metakatepisternum; lbr, labrum; md, mandible; mt, mentum; pl, propleuron; pmt, prementum; pst, prosternum; pta, protarsi; scu2, mesoscutum; st, sternite; ti1, tibia1; tisp, tibial spur; tr, trochantin; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.14896578" httpUri="https://zenodo.org/record/14896578/files/figure.png" pageId="5" pageNumber="406">Figs 2</figureCitation>
,
<figureCitation id="134E2A05FF8B666296BCF8DE058DF88E" box="[910,948,1796,1822]" captionStart="FIGURE 3" captionStartId="5.[151,250,1114,1138]" captionTargetBox="[159,1423,184,1082]" captionTargetId="figure-18@5.[151,1436,179,1088]" captionTargetPageId="5" captionText="FIGURE 3. Reconstruction of Tigrivia baii, based on holotype and drawing, (A) ventral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14896580" httpUri="https://zenodo.org/record/14896580/files/figure.png" pageId="5" pageNumber="406">3A</figureCitation>
); mesal walls extensively fused; metacoxal plates present but small, not forming duplicatures, anteriorly ending at about half width of anterior coxal margin; rounded posteriorly. Metatrochanter exposed, slightly larger than mesotrochanter. Metafemur 5.8 mm long, metatibia 5.4 mm; both similar to corresponding mesothoracic equivalents. Metatarsus large, distinctly broadened, paddle-like; length formula 2.0–1.0–1.0–1.5–2.0 mm; tarsomere 1 largest; tarsomere 5 cylindrical, with double claw (
<figureCitation id="134E2A05FF8B66629058F84E0680F842" captionStart="FIGURE 1" captionStartId="3.[151,250,1962,1986]" captionTargetBox="[162,1425,198,1936]" captionTargetId="figure-18@3.[162,1425,198,1936]" captionTargetPageId="3" captionText="FIGURE 1. Photographs of Tigrivia baii, holotype, YLSNHM01120, (A) dorsal view; (B) ventral view. Scale bars 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14896576" httpUri="https://zenodo.org/record/14896576/files/figure.png" pageId="5" pageNumber="406">Figs 1B</figureCitation>
,
<figureCitation id="134E2A05FF8B666295F4F86206EFF842" box="[198,214,1976,2002]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[286,1303,1320,1869]" captionTargetId="figure-482@4.[280,1308,1314,1883]" captionTargetPageId="4" captionText="FIGURE 2. Line drawing of Tigrivia baii, based on holotype, ventral view. Abbreviations: aes1, mesanepisternum; aes3, metanepisternum; cx, coxa; cxpl,coxal plate;ep, episternum;fe, femur;gcxs, gonocoxosterna; gu, gula; kes3, metakatepisternum; lbr, labrum; md, mandible; mt, mentum; pl, propleuron; pmt, prementum; pst, prosternum; pta, protarsi; scu2, mesoscutum; st, sternite; ti1, tibia1; tisp, tibial spur; tr, trochantin; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.14896578" httpUri="https://zenodo.org/record/14896578/files/figure.png" pageId="5" pageNumber="406">2</figureCitation>
,
<figureCitation id="134E2A05FF8B666295D0F862073EF842" box="[226,263,1976,2002]" captionStart="FIGURE 3" captionStartId="5.[151,250,1114,1138]" captionTargetBox="[159,1423,184,1082]" captionTargetId="figure-18@5.[151,1436,179,1088]" captionTargetPageId="5" captionText="FIGURE 3. Reconstruction of Tigrivia baii, based on holotype and drawing, (A) ventral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14896580" httpUri="https://zenodo.org/record/14896580/files/figure.png" pageId="5" pageNumber="406">3A</figureCitation>
). Swimming hairs not recognizable.
</paragraph>
<paragraph id="8BCA3680FF88666195F5FF4D0235FED1" blockId="6.[151,1437,151,609]" pageId="6" pageNumber="407">
Elytral shoulder region obsolete, pronoto-elytral angle thus indistinct. Elytra
<quantity id="4C8D9B65FF8866619114FF4D024CFF21" box="[1062,1141,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="407" unit="mm" value="20.0">20 mm</quantity>
long and maximum width 6.2 mm, distinctly widening posteriorly, widest at about posterior 1/3 (preserved with distal portion of mesal edges separated); posteriorly truncated, with very small posterolateral protuberance and straight posterior edge. Striae or other surface structures not recognizable, elytra apparently smooth. Epipleura fairly narrow, parallel-sided, slightly narrowing anteriorly, posteriorly reaching posterior elytral margin (
<figureCitation id="134E2A05FF88666196BBFEFD05F4FED1" box="[905,973,295,321]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[286,1303,1320,1869]" captionTargetId="figure-482@4.[280,1308,1314,1883]" captionTargetPageId="4" captionText="FIGURE 2. Line drawing of Tigrivia baii, based on holotype, ventral view. Abbreviations: aes1, mesanepisternum; aes3, metanepisternum; cx, coxa; cxpl,coxal plate;ep, episternum;fe, femur;gcxs, gonocoxosterna; gu, gula; kes3, metakatepisternum; lbr, labrum; md, mandible; mt, mentum; pl, propleuron; pmt, prementum; pst, prosternum; pta, protarsi; scu2, mesoscutum; st, sternite; ti1, tibia1; tisp, tibial spur; tr, trochantin; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.14896578" httpUri="https://zenodo.org/record/14896578/files/figure.png" pageId="6" pageNumber="407">Fig. 2</figureCitation>
,
<figureCitation id="134E2A05FF88666196EBFEFD05C6FED1" box="[985,1023,295,321]" captionStart="FIGURE 3" captionStartId="5.[151,250,1114,1138]" captionTargetBox="[159,1423,184,1082]" captionTargetId="figure-18@5.[151,1436,179,1088]" captionTargetPageId="5" captionText="FIGURE 3. Reconstruction of Tigrivia baii, based on holotype and drawing, (A) ventral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14896580" httpUri="https://zenodo.org/record/14896580/files/figure.png" pageId="6" pageNumber="407">3A</figureCitation>
).
</paragraph>
<paragraph id="8BCA3680FF88666195F5FE91074AFDF1" blockId="6.[151,1437,151,609]" pageId="6" pageNumber="407">
Abdomen with six exposed sternites (II–VII), widest in middle region, narrowing posteriorly. Sternite II (= ventrite 1) completely divided by median part of metacoxae, sternite III partially (
<figureCitation id="134E2A05FF8866619117FEB50254FE19" box="[1061,1133,367,393]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[286,1303,1320,1869]" captionTargetId="figure-482@4.[280,1308,1314,1883]" captionTargetPageId="4" captionText="FIGURE 2. Line drawing of Tigrivia baii, based on holotype, ventral view. Abbreviations: aes1, mesanepisternum; aes3, metanepisternum; cx, coxa; cxpl,coxal plate;ep, episternum;fe, femur;gcxs, gonocoxosterna; gu, gula; kes3, metakatepisternum; lbr, labrum; md, mandible; mt, mentum; pl, propleuron; pmt, prementum; pst, prosternum; pta, protarsi; scu2, mesoscutum; st, sternite; ti1, tibia1; tisp, tibial spur; tr, trochantin; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.14896578" httpUri="https://zenodo.org/record/14896578/files/figure.png" pageId="6" pageNumber="407">Figs 2</figureCitation>
,
<figureCitation id="134E2A05FF886661914AFEB502A7FE19" box="[1144,1182,367,393]" captionStart="FIGURE 3" captionStartId="5.[151,250,1114,1138]" captionTargetBox="[159,1423,184,1082]" captionTargetId="figure-18@5.[151,1436,179,1088]" captionTargetPageId="5" captionText="FIGURE 3. Reconstruction of Tigrivia baii, based on holotype and drawing, (A) ventral view; (B) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14896580" httpUri="https://zenodo.org/record/14896580/files/figure.png" pageId="6" pageNumber="407">3A</figureCitation>
). Sternites II–VI about equally long, with straight posterior margins. Terminal sternite VII about 1.5 times as long as preceding sternites, 2.4 times as wide as long; posterior edge indistinct, apparently sinuate laterally and rounded caudally. Genital complex comprising paired gonocoxosterna and gonocoxae distinctly exposed posterior to sternite VII, tergum VIII and elytra (
<figureCitation id="134E2A05FF8866619424FE250745FD89" box="[278,380,511,537]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1962,1986]" captionStartId-1="4.[151,250,1909,1933]" captionStartId-2="5.[151,250,1114,1138]" captionTargetBox-0="[162,1425,198,1936]" captionTargetBox-1="[286,1303,1320,1869]" captionTargetBox-2="[159,1423,184,1082]" captionTargetId-0="figure-18@3.[162,1425,198,1936]" captionTargetId-1="figure-482@4.[280,1308,1314,1883]" captionTargetId-2="figure-18@5.[151,1436,179,1088]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Photographs of Tigrivia baii, holotype, YLSNHM01120, (A) dorsal view; (B) ventral view. Scale bars 3.5 mm." captionText-1="FIGURE 2. Line drawing of Tigrivia baii, based on holotype, ventral view. Abbreviations: aes1, mesanepisternum; aes3, metanepisternum; cx, coxa; cxpl,coxal plate;ep, episternum;fe, femur;gcxs, gonocoxosterna; gu, gula; kes3, metakatepisternum; lbr, labrum; md, mandible; mt, mentum; pl, propleuron; pmt, prementum; pst, prosternum; pta, protarsi; scu2, mesoscutum; st, sternite; ti1, tibia1; tisp, tibial spur; tr, trochantin; v2, mesoventrite; v3, metaventrite." captionText-2="FIGURE 3. Reconstruction of Tigrivia baii, based on holotype and drawing, (A) ventral view; (B) dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.14896576" figureDoi-1="http://doi.org/10.5281/zenodo.14896578" figureDoi-2="http://doi.org/10.5281/zenodo.14896580" httpUri-0="https://zenodo.org/record/14896576/files/figure.png" httpUri-1="https://zenodo.org/record/14896578/files/figure.png" httpUri-2="https://zenodo.org/record/14896580/files/figure.png" pageId="6" pageNumber="407">Figs 1–3</figureCitation>
). The gonocoxosterna are usually paired structures forming the ventral cover of segment VIII and invaginated in the resting position of the genital apparatus (
<bibRefCitation id="EFE44B71FF886661966AFDF90220FDAD" author="Burmeister, E. G." box="[856,1049,547,573]" pageId="6" pageNumber="407" pagination="165 - 257" refId="ref8521" refString="Burmeister, E. G. (1976) The ovipositor of the Hydradephaga (Coleoptera) and its phylogenetic significance, with special considerations of the Dytiscidae. Zoomorphologie, 85, 165-257. https://doi.org/10.1007/BF00993515" type="journal article" year="1976">Burmeister 1976</bibRefCitation>
: figs 3, 4;
<bibRefCitation id="EFE44B71FF88666191B8FDF903A1FDAD" author="Balke, M. &amp; Hendrich, L." box="[1162,1432,547,573]" pageId="6" pageNumber="407" pagination="118 - 140" refId="ref7097" refString="Balke, M. &amp; Hendrich, L. (2016) Dytiscidae Leach 1915. In: Beutel, R. G. &amp; Leschen, R. A. B. (Eds), Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). Handbook of Zoology Vol. IV, Arthropoda: Insecta, 2 nd ed. Walter De Gruyter, Berlin and New York, pp. 118-140." type="book chapter" year="2016">Balke &amp; Hendrich 2016</bibRefCitation>
; see also
<bibRefCitation id="EFE44B71FF88666195C4FD9D075FFDF1" author="Bils, W." box="[246,358,583,609]" pageId="6" pageNumber="407" pagination="113 - 193" refId="ref8377" refString="Bils, W. (1976) Das Abdomenende weiblicher, terrestrisch lebender Adephaga (Coleoptera) und seine Bedeutung fur die Phylogenie. Zoomorphologie, 84, 113-193. https://doi.org/10.1007/BF00999711" type="journal article" year="1976">Bils 1976</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

158
data/03/DC/87/03DC8796FF8C666595A5FC9804ABFA6E.xml Normal file
View file

@ -0,0 +1,158 @@
<document id="5616944B5426332BB3C438472565C93D" ID-DOI="10.11646/zootaxa.5588.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14896574" ID-ZooBank="E3697AAB-FB6D-440B-89C7-3AD757157481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1740004092971" checkinUser="plazi" docAuthor="Bao, Liang, Beutel, Rolf Georg, Niu, Kecheng &amp; Bao, Tong" docDate="2025" docId="03DC8796FF8C666595A5FC9804ABFA6E" docLanguage="en" docName="zootaxa.5588.3.1.pdf" docOrigin="Zootaxa 5588 (3)" docSource="https://doi.org/10.11646/zootaxa.5588.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tigrivia Bao et al. 2023" docType="treatment" docVersion="1" lastPageNumber="403" masterDocId="FFE5FFEEFF8E66679532FFDA0639FF90" masterDocTitle="Redescription of the highly specialized aquatic † Tigrivia and the classification of the Mesozoic † Coptoclavidae (Coleoptera, Adephaga, Dytiscoidea)" masterLastPageNumber="425" masterPageNumber="401" pageNumber="403" updateTime="1741050061580" updateUser="jonas" zenodo-license-document="CLOSED">
<mods:mods id="A937F2965525EE99209582BFC5F51765" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4CAB87F575D1F43D8FDD741FDD5EAD14">
<mods:title id="7B690E914B4E1C0003324F42F095E2C4">Redescription of the highly specialized aquatic † Tigrivia and the classification of the Mesozoic † Coptoclavidae (Coleoptera, Adephaga, Dytiscoidea)</mods:title>
</mods:titleInfo>
<mods:name id="C429EEA6D6032B116B036BB97E6C436F" type="personal">
<mods:role id="33B1F6B5F3A5977AFB480A02DBF74A66">
<mods:roleTerm id="D9D2B355972FDF3718AA8840890A90D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4825D27FC79E0D8E650AB4D3AB0A53AC">Bao, Liang</mods:namePart>
<mods:affiliation id="5FF45FD5E537202EE61B1AE527FF4312">School of Earth Sciences and Engineering, Guangdong Provincial Key Lab of Geological Processes and Mineral Resources, Sun Yat-sen University, Guangzhou 510275, China. &amp; School of Ecology / State Key Laboratory of Biocontrol, Shenzhen Campus of Sun Yat-sen University, Shenzhen 518107, China.</mods:affiliation>
</mods:name>
<mods:name id="41060EAC2B48D70040B9C05775FFB9F2" type="personal">
<mods:role id="EF37D5EB62C03AB7CC338FBFD5661153">
<mods:roleTerm id="511DD9074005CF8BD0C194605667C068">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7E18F899F4DEFA9C9CC54140BCE85FB1">Beutel, Rolf Georg</mods:namePart>
<mods:affiliation id="EE936C70156899F93CA237CA0FC77282">Institut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität Jena, Jena 07743, Germany.</mods:affiliation>
</mods:name>
<mods:name id="055C78140D5E304C894165AA62DF0FCA" type="personal">
<mods:role id="954AA20E09324E8AF190DCC335881FFE">
<mods:roleTerm id="7A99D57B64BA174EDB967B759154EB6E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1BC57CB96778D0A8B9E0568FEF4B217A">Niu, Kecheng</mods:namePart>
<mods:affiliation id="A9B4F33554B27A83FA66D9A8CBE95660">State Key Laboratory of Cellular Stress Biology, School of Life Sciences, Xiamen University, Xiamen 361105, China. &amp; Yingliang Stone Natural History Museum, Nan’an 362300, China.</mods:affiliation>
</mods:name>
<mods:name id="4388A39C9BED48FEC436BDB7BCA132A0" type="personal">
<mods:role id="B668C70E4481B34411734924CBA3342E">
<mods:roleTerm id="6E060104854A80D6137BC7B0D92BEEE8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4ED50304D7BDE047F752CBE7C39B2F17">Bao, Tong</mods:namePart>
<mods:affiliation id="9F0DFA67BBDE1BD2108D96FC22193DE6">School of Ecology / State Key Laboratory of Biocontrol, Shenzhen Campus of Sun Yat-sen University, Shenzhen 518107, China. &amp; Yunnan Key Laboratory for Palaeobiology, Yunnan University, Kunming 650091, China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="D80EC9C0D0DAF1662967F7446F8CCD0F">text</mods:typeOfResource>
<mods:relatedItem id="35B5F6A1DC48ECF536BF08B29E10BC3B" type="host">
<mods:titleInfo id="1DABC3D5D8516532F72086052D53FFCB">
<mods:title id="04D8275C2747E4031416C48708726E07">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4D652A9F9965D55B35750791A23F1CE8">
<mods:date id="B40FCABB07BD5035164B9441010B47FF">2025</mods:date>
<mods:detail id="47AC9DE4E599AEEE13ADD0F8AD5A6B29" type="pubDate">
<mods:number id="92275A284FCB753BDD92C7A7C1255C11">2025-02-17</mods:number>
</mods:detail>
<mods:detail id="8E4AD923CFBAB365EA6894F9A786B1BF" type="volume">
<mods:number id="422A5B643E16AF9D0E24A9DB76F9B698">5588</mods:number>
</mods:detail>
<mods:detail id="5A069D9AEF9A1F8C1DE847FC328676AD" type="issue">
<mods:number id="EBE2C8C39C30A567D98E4D33A88A392A">3</mods:number>
</mods:detail>
<mods:extent id="A04A65FB3F69953A6CA634FE17E9D941" unit="page">
<mods:start id="D8E3459F8722402BB8C4327A3EE2FFF8">401</mods:start>
<mods:end id="11D2B8DD43B324DF43236877F8D83599">425</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="A71693FBB31E6D580FF890B11AB41EC2">
<mods:url id="5B572ED66F0113E1EE03D629CFCCFE44">https://doi.org/10.11646/zootaxa.5588.3.1</mods:url>
</mods:location>
<mods:classification id="79CE00653B53E1E7B533423B5360C25C">journal article</mods:classification>
<mods:identifier id="31478156F2223CD6C368DEEC852178F0" type="DOI">10.11646/zootaxa.5588.3.1</mods:identifier>
<mods:identifier id="D25EB8FC4EE45E5792B39C81815A03D0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EF7A372B6A1026879C921B6D17964B37" type="Zenodo-Dep">14896574</mods:identifier>
<mods:identifier id="F92E44B84C053001DE10B70CB4FCFA06" type="ZooBank">E3697AAB-FB6D-440B-89C7-3AD757157481</mods:identifier>
</mods:mods>
<treatment id="03DC8796FF8C666595A5FC9804ABFA6E" LSID="urn:lsid:plazi:treatment:03DC8796FF8C666595A5FC9804ABFA6E" httpUri="http://treatment.plazi.org/id/03DC8796FF8C666595A5FC9804ABFA6E" lastPageNumber="403" pageId="2" pageNumber="403">
<subSubSection id="C36F650BFF8C666595A5FC980413FCCC" box="[151,554,834,861]" pageId="2" pageNumber="403" type="nomenclature">
<paragraph id="8BCA3680FF8C666595A5FC980413FCCC" blockId="2.[151,554,834,861]" box="[151,554,834,861]" pageId="2" pageNumber="403">
<heading id="D08281ECFF8C666595A5FC980413FCCC" bold="true" box="[151,554,834,861]" fontSize="11" level="1" pageId="2" pageNumber="403" reason="1">
<emphasis id="B901EA92FF8C666595A5FC980413FCCC" bold="true" box="[151,554,834,861]" pageId="2" pageNumber="403">
Genus: †
<taxonomicName id="4C754D03FF8C66659435FC990413FCCC" authority="Bao et al., 2023" authorityName="Bao et al." authorityYear="2023" box="[263,554,834,861]" class="Insecta" family="Coptoclavidae" genus="Tigrivia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B901EA92FF8C66659435FC99075FFCCD" bold="true" box="[263,358,835,861]" italics="true" pageId="2" pageNumber="403">Tigrivia</emphasis>
<bibRefCitation id="EFE44B71FF8C6665945FFC990413FCCC" author="Bao, L. &amp; Li, L. &amp; Niu, K. &amp; Wang, N. &amp; Kroeck, D. M. &amp; Bao, T." box="[365,554,834,861]" pageId="2" pageNumber="403" pagination="1 - E 6" refId="ref7184" refString="Bao, L., Li, L., Niu, K., Wang, N., Kroeck, D. M. &amp; Bao, T. (2023) Retracted: A new aquatic beetle (Adephaga: Coptoclavidae) from the Middle Jurassic Daohugou Biota. The Anatomical Record, 306 (8), E 1-E6. https://doi.org/10.1002/ar.25221" type="journal article" year="2023">
Bao
<emphasis id="B901EA92FF8C66659497FC9907D8FCCD" bold="true" box="[421,481,834,861]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
, 2023
</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F650BFF8C666595A5FC520460FC0F" box="[151,601,903,928]" pageId="2" pageNumber="403" type="type_taxon">
<paragraph id="8BCA3680FF8C666595A5FC520460FC0F" blockId="2.[151,601,903,928]" box="[151,601,903,928]" pageId="2" pageNumber="403">
<typeStatus id="54CE8822FF8C666595A5FC5206F0FC30" box="[151,201,904,928]" pageId="2" pageNumber="403">Type</typeStatus>
species: †
<taxonomicName id="4C754D03FF8C66659401FC520460FC0F" authority="Bao et al., 2023" authorityName="Bao" authorityYear="2023" box="[307,601,903,927]" class="Insecta" family="Coptoclavidae" genus="Tigrivia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="baii">
<emphasis id="B901EA92FF8C66659401FC520788FC0F" box="[307,433,903,927]" italics="true" pageId="2" pageNumber="403">Tigrivia baii</emphasis>
<bibRefCitation id="EFE44B71FF8C66659485FC520460FC0F" author="Bao, L. &amp; Li, L. &amp; Niu, K. &amp; Wang, N. &amp; Kroeck, D. M. &amp; Bao, T." box="[439,601,903,927]" pageId="2" pageNumber="403" pagination="1 - E 6" refId="ref7184" refString="Bao, L., Li, L., Niu, K., Wang, N., Kroeck, D. M. &amp; Bao, T. (2023) Retracted: A new aquatic beetle (Adephaga: Coptoclavidae) from the Middle Jurassic Daohugou Biota. The Anatomical Record, 306 (8), E 1-E6. https://doi.org/10.1002/ar.25221" type="journal article" year="2023">
Bao
<emphasis id="B901EA92FF8C666594D4FC520423FC0F" box="[486,538,903,927]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
, 2023
</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C36F650BFF8C666595A5FC1D04F8FB96" pageId="2" pageNumber="403" type="etymology">
<paragraph id="8BCA3680FF8C666595A5FC1D04F8FB96" blockId="2.[151,1437,967,1534]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595A5FC1D0720FC72" bold="true" box="[151,281,967,994]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595A5FC1D072BFC72" bold="true" box="[151,274,967,994]" italics="true" pageId="2" pageNumber="403">Etymology</emphasis>
.
</emphasis>
The term “tigrivia” is derived from “tiger” and “via” (=way). This amalgamation succinctly denotes that the species had a specialized predacious lifestyle.
</paragraph>
</subSubSection>
<subSubSection id="C36F650BFF8C666595F5FBD504ABFA6E" pageId="2" pageNumber="403" type="diagnosis">
<paragraph id="8BCA3680FF8C666595F5FBD5045BFB4E" blockId="2.[151,1437,967,1534]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595F5FBD50483FBBA" bold="true" box="[199,698,1039,1066]" pageId="2" pageNumber="403">
<emphasis id="B901EA92FF8C666595F5FBD50410FBBA" bold="true" box="[199,553,1039,1066]" italics="true" pageId="2" pageNumber="403">Revised differential diagnosis</emphasis>
(
<figureCitation id="134E2A05FF8C6665970CFBCA0492FBBA" box="[574,683,1040,1066]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1962,1986]" captionStartId-1="4.[151,250,1909,1933]" captionStartId-2="5.[151,250,1114,1138]" captionTargetBox-0="[162,1425,198,1936]" captionTargetBox-1="[286,1303,1320,1869]" captionTargetBox-2="[159,1423,184,1082]" captionTargetId-0="figure-18@3.[162,1425,198,1936]" captionTargetId-1="figure-482@4.[280,1308,1314,1883]" captionTargetId-2="figure-18@5.[151,1436,179,1088]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Photographs of Tigrivia baii, holotype, YLSNHM01120, (A) dorsal view; (B) ventral view. Scale bars 3.5 mm." captionText-1="FIGURE 2. Line drawing of Tigrivia baii, based on holotype, ventral view. Abbreviations: aes1, mesanepisternum; aes3, metanepisternum; cx, coxa; cxpl,coxal plate;ep, episternum;fe, femur;gcxs, gonocoxosterna; gu, gula; kes3, metakatepisternum; lbr, labrum; md, mandible; mt, mentum; pl, propleuron; pmt, prementum; pst, prosternum; pta, protarsi; scu2, mesoscutum; st, sternite; ti1, tibia1; tisp, tibial spur; tr, trochantin; v2, mesoventrite; v3, metaventrite." captionText-2="FIGURE 3. Reconstruction of Tigrivia baii, based on holotype and drawing, (A) ventral view; (B) dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.14896576" figureDoi-1="http://doi.org/10.5281/zenodo.14896578" figureDoi-2="http://doi.org/10.5281/zenodo.14896580" httpUri-0="https://zenodo.org/record/14896576/files/figure.png" httpUri-1="https://zenodo.org/record/14896578/files/figure.png" httpUri-2="https://zenodo.org/record/14896580/files/figure.png" pageId="2" pageNumber="403">Figs 1–3</figureCitation>
).
</emphasis>
(1) Body distinctly widened posteriorly; (2) elytra truncated, exposing gonocoxosterna and gonocoxae; (3) only one pair of compound eyes; (4) prosternal process very short; (5) mesoventrite fairly extensive, largely flat, lacking a hexagonal groove for reception of the prosternal process; (6) forelegs elongated, raptorial, with distinct medial tibial spur; (7) protarsomeres 2 and 5 longest, 3 and 4 short; (8) middle legs and hind legs long and distally flattened, paddle-like; (8) metacoxal plates small, only with medial portion, leaving metatrochanter exposed.
</paragraph>
<paragraph id="8BCA3680FF8C666595F5FB3204ABFA6E" blockId="2.[151,1437,967,1534]" pageId="2" pageNumber="403">
The structure of the protarsus exhibits a characteristic pattern akin to the raptorial forelegs observed in some †Coptoclavinae (
<bibRefCitation id="EFE44B71FF8C66659467FAD6040DFAB6" author="Ponomarenko, A. G." box="[341,564,1292,1318]" pageId="2" pageNumber="403" pagination="1 - 204" refId="ref10114" refString="Ponomarenko, A. G. (1977) Suborder Adephaga, etc. In: Arnoldi, L. V., Zherikin, V. V., Nikritin, L. M. &amp; Ponomarenko, A. G. (Eds.), Mesozoic Coleoptera. Trudy Paleonthologicheskogo Instituta Akademiya Nauk SSSR, 161, pp. 1-204. [in Russian]" type="book chapter" year="1977">Ponomarenko 1977</bibRefCitation>
: fig. 13;
<bibRefCitation id="EFE44B71FF8C666597A7FAD60577FAB6" author="Zhao, X. &amp; Zhao, X. &amp; Jarzembowski, E. A. &amp; Chen, L. &amp; Wang, B." box="[661,846,1292,1318]" pageId="2" pageNumber="403" pagination="205 - 209" refId="ref11806" refString="Zhao, X., Zhao, X., Jarzembowski, E. A., Chen, L. &amp; Wang, B. (2018) First record of adult Coptoclava longipoda Ping (Coleoptera: Coptoclavidae) from the Lower Cretaceous of Laiyang, China. Cretaceous Research, 92, 205-209. https://doi.org/10.1016/j.cretres.2018.08.013" type="journal article" year="2018">
Zhao
<emphasis id="B901EA92FF8C666597E7FAD70535FAB6" box="[725,780,1292,1318]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
2018
</bibRefCitation>
: fig. 4). Based on two intricate apomorphic traits, the specimen under examination can confidently be classified within this subfamily and considered a close relative of †
<taxonomicName id="4C754D03FF8C66659597FA8E0794FAFE" authority="Ping, 1928" authorityName="Ping" authorityYear="1928" box="[165,429,1364,1390]" class="Insecta" genus="Coptoclava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B901EA92FF8C66659597FA8E071FFAFE" box="[165,294,1364,1390]" italics="true" pageId="2" pageNumber="403">Coptoclava</emphasis>
<bibRefCitation id="EFE44B71FF8C6665941DFA8E0794FAFE" author="Ping, C." box="[303,429,1364,1390]" pageId="2" pageNumber="403" pagination="1 - 47" refId="ref9940" refString="Ping, C. (1928) Study of the Cretaceous fossil insects of China. Palaeontologia Sinica, 13, 1-47." type="journal article" year="1928">Ping, 1928</bibRefCitation>
</taxonomicName>
. These features include (1) elongated raptorial forelegs, displaying a remarkably similar and distinctive pattern of tarsomeres also present in †
<taxonomicName id="4C754D03FF8C666597C9FAA20545FA02" authorityName="Ping" authorityYear="1928" box="[763,892,1400,1426]" class="Insecta" genus="Coptoclava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B901EA92FF8C666597C9FAA20545FA02" box="[763,892,1400,1426]" italics="true" pageId="2" pageNumber="403">Coptoclava</emphasis>
</taxonomicName>
(as noted previously;
<bibRefCitation id="EFE44B71FF8C6665914EFAA202CAFA02" author="Ping, C." box="[1148,1267,1400,1426]" pageId="2" pageNumber="403" pagination="1 - 47" refId="ref9940" refString="Ping, C. (1928) Study of the Cretaceous fossil insects of China. Palaeontologia Sinica, 13, 1-47." type="journal article" year="1928">Ping 1928</bibRefCitation>
;
<bibRefCitation id="EFE44B71FF8C66659033FAA206EBFA26" author="Ponomarenko, A. G." pageId="2" pageNumber="403" pagination="1 - 204" refId="ref10114" refString="Ponomarenko, A. G. (1977) Suborder Adephaga, etc. In: Arnoldi, L. V., Zherikin, V. V., Nikritin, L. M. &amp; Ponomarenko, A. G. (Eds.), Mesozoic Coleoptera. Trudy Paleonthologicheskogo Instituta Akademiya Nauk SSSR, 161, pp. 1-204. [in Russian]" type="book chapter" year="1977">Ponomarenko 1977</bibRefCitation>
;
<bibRefCitation id="EFE44B71FF8C666595EFFA4607AFFA26" author="Zhao, X. &amp; Zhao, X. &amp; Jarzembowski, E. A. &amp; Chen, L. &amp; Wang, B." box="[221,406,1436,1462]" pageId="2" pageNumber="403" pagination="205 - 209" refId="ref11806" refString="Zhao, X., Zhao, X., Jarzembowski, E. A., Chen, L. &amp; Wang, B. (2018) First record of adult Coptoclava longipoda Ping (Coleoptera: Coptoclavidae) from the Lower Cretaceous of Laiyang, China. Cretaceous Research, 92, 205-209. https://doi.org/10.1016/j.cretres.2018.08.013" type="journal article" year="2018">
Zhao
<emphasis id="B901EA92FF8C6665942FFA47076DFA26" box="[285,340,1436,1462]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
2018
</bibRefCitation>
: fig. 4), and (2) large paddle-like swimming legs on the meso- and metathorax, characterized by flattened and moderately broadened tarsi, with the intermediate ones typically heart-shaped (
<bibRefCitation id="EFE44B71FF8C66659028FA1A03AEFA4A" author="Ping, C." box="[1306,1431,1472,1498]" pageId="2" pageNumber="403" pagination="1 - 47" refId="ref9940" refString="Ping, C. (1928) Study of the Cretaceous fossil insects of China. Palaeontologia Sinica, 13, 1-47." type="journal article" year="1928">Ping 1928</bibRefCitation>
;
<bibRefCitation id="EFE44B71FF8C666595A5FA3E074CFA6E" author="Ponomarenko, A. G." box="[151,373,1508,1534]" pageId="2" pageNumber="403" pagination="1 - 204" refId="ref10114" refString="Ponomarenko, A. G. (1977) Suborder Adephaga, etc. In: Arnoldi, L. V., Zherikin, V. V., Nikritin, L. M. &amp; Ponomarenko, A. G. (Eds.), Mesozoic Coleoptera. Trudy Paleonthologicheskogo Instituta Akademiya Nauk SSSR, 161, pp. 1-204. [in Russian]" type="book chapter" year="1977">Ponomarenko 1977</bibRefCitation>
;
<bibRefCitation id="EFE44B71FF8C666594B3FA3E0405FA6E" author="Zhao, X. &amp; Zhao, X. &amp; Jarzembowski, E. A. &amp; Chen, L. &amp; Wang, B." box="[385,572,1508,1534]" pageId="2" pageNumber="403" pagination="205 - 209" refId="ref11806" refString="Zhao, X., Zhao, X., Jarzembowski, E. A., Chen, L. &amp; Wang, B. (2018) First record of adult Coptoclava longipoda Ping (Coleoptera: Coptoclavidae) from the Lower Cretaceous of Laiyang, China. Cretaceous Research, 92, 205-209. https://doi.org/10.1016/j.cretres.2018.08.013" type="journal article" year="2018">
Zhao
<emphasis id="B901EA92FF8C666594F3FA3F07C0FA6E" box="[449,505,1508,1534]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
2018
</bibRefCitation>
: fig. 4).
</paragraph>
</subSubSection>
</treatment>
</document>