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<mods:title id="F01FE810090880B676AD4B0564A0902B">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<treatment ID-GBIF-Taxon="161776991" LSID="urn:lsid:zoobank.org:act:5845FFBC-72D4-43A6-AF1D-11C57445217F" httpUri="http://treatment.plazi.org/id/FE9EE26351DF4A7FD85F6AB8757CD313" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/5845FFBC-72D4-43A6-AF1D-11C57445217F" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana" status="sp. nov.">Pheidole ankerana</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">Figs 71A-F</figureCitation>
|
||||
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84D</figureCitation>
|
||||
<figureCitation captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86D</figureCitation>
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||||
<treatment id="FE9EE26351DF4A7FD85F6AB8757CD313" ID-GBIF-Taxon="161776991" LSID="urn:lsid:zoobank.org:act:5845FFBC-72D4-43A6-AF1D-11C57445217F" httpUri="http://treatment.plazi.org/id/FE9EE26351DF4A7FD85F6AB8757CD313" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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||||
<subSubSection id="8828591AEE246AAABB7E60566AAF5A3F" pageId="0" pageNumber="1" type="nomenclature">
|
||||
<paragraph id="0AF87A46FC5DA7FE92A73A095243B865" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="61912BAC717191190785C455A3AE5F24" ID-CoL="CMKV3" LSID="http://zoobank.org/5845FFBC-72D4-43A6-AF1D-11C57445217F" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana" status="sp. nov.">Pheidole ankerana</taxonomicName>
|
||||
<taxonomicNameLabel id="BA368B8A6736FA90AA1FB3957EB7A305" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
|
||||
<figureCitation id="F29D3283E692E41335890B06B795C40B" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">Figs 71A-F</figureCitation>
|
||||
<figureCitation id="53F4A5475BF1360E8716DF8F4B60B9CD" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84D</figureCitation>
|
||||
<figureCitation id="E0FFABA1BD8E773D4A5D26244A4B2854" captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86D</figureCitation>
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="type material">
|
||||
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||||
<subSubSection id="4368F6C3F68EDAD058DACFA482BECE22" pageId="0" pageNumber="1" type="type material">
|
||||
<paragraph id="436B0D2E6C32FAD9AC5232DED7D44AFF" pageId="0" pageNumber="1">Type material.</paragraph>
|
||||
<paragraph id="3CAC724A085C2A74DC0EA5E27E22E555" pageId="0" pageNumber="1">
|
||||
<emphasis id="448E7CA9F2F5A5958C56BE12009FB9A7" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="2E97010AF628E9C7F3CD55DBA599D907" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||||
.
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</emphasis>
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Madagascar. •1 major worker; Toamasina; Ankerana;
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||||
<geoCoordinate degrees="18.4104" direction="south" orientation="latitude" precision="5" value="-18.4104">-18.4104</geoCoordinate>
|
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<geoCoordinate id="00E1ECD29B9411BD761481D020D93B29" degrees="18.4104" direction="south" orientation="latitude" precision="5" value="-18.4104">-18.4104</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="48.8189" direction="east" orientation="longitude" precision="5" value="48.8189">48.8189</geoCoordinate>
|
||||
<geoCoordinate id="41B43B09CC3DE623744D746E0CBBC3E2" degrees="48.8189" direction="east" orientation="longitude" precision="5" value="48.8189">48.8189</geoCoordinate>
|
||||
; alt. 855 m; 27 Jan 2012; B.L. Fisher et al. leg.; BLF28148, CASENT0923171 (CASC).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
|
||||
<emphasis id="496A23F5329FD70FE48CB7C48FB75DB7" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="A666E432FB2DC473176594C88910929E" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
|
||||
.
|
||||
</emphasis>
|
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Madagascar. •2w., 1q.; same data as for holotype; CASENT0274002, CASENT0274003, CASENT0872185 (CASC).
|
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</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="49BFBD89609C745E538CDFC9E3810AA9" pageId="0" pageNumber="1" type="materials_examined">
|
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<paragraph id="CAF991987D22EDDEF685B1EC6FC1240C" pageId="0" pageNumber="1">Other material.</paragraph>
|
||||
<paragraph id="A0C02992DE5852F933BE2E8DCD7F6018" pageId="0" pageNumber="1">
|
||||
Madagascar. -
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
|
||||
<emphasis id="9CAB8061AECF9B0DEFF9FF1B6E2B2B63" bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
|
||||
: •3w.; Ankerana;
|
||||
<geoCoordinate degrees="18.4061" direction="south" orientation="latitude" precision="5" value="-18.4061">-18.4061</geoCoordinate>
|
||||
<geoCoordinate id="40000E7EC25611985479FAD6D881FF6A" degrees="18.4061" direction="south" orientation="latitude" precision="5" value="-18.4061">-18.4061</geoCoordinate>
|
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,
|
||||
<geoCoordinate degrees="48.82029" direction="east" orientation="longitude" precision="1" value="48.82029">48.82029</geoCoordinate>
|
||||
<geoCoordinate id="DCAED65971B29FD70703704024AB872D" degrees="48.82029" direction="east" orientation="longitude" precision="1" value="48.82029">48.82029</geoCoordinate>
|
||||
; alt. 725 m; 20 Jan 2012; B.L. Fisher et al. leg.; CASENT0274747, CASENT0274762 (CASC). •3w., 1m.; Ankerana;
|
||||
<geoCoordinate degrees="18.40829" direction="south" orientation="latitude" precision="1" value="-18.40829">-18.40829</geoCoordinate>
|
||||
<geoCoordinate id="93101F3F2C389024FE582C0490FC0EDB" degrees="18.40829" direction="south" orientation="latitude" precision="1" value="-18.40829">-18.40829</geoCoordinate>
|
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,
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||||
<geoCoordinate degrees="48.82107" direction="east" orientation="longitude" precision="1" value="48.82107">48.82107</geoCoordinate>
|
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<geoCoordinate id="82AB2D3E4D20F443E246350CA1167239" degrees="48.82107" direction="east" orientation="longitude" precision="1" value="48.82107">48.82107</geoCoordinate>
|
||||
; alt. 750 m; 21 Jan 2012; B.L. Fisher et al. leg.; CASENT0275260, CASENT0275261 (CASC). •2w.; Ankerana;
|
||||
<geoCoordinate degrees="18.4104" direction="south" orientation="latitude" precision="5" value="-18.4104">-18.4104</geoCoordinate>
|
||||
<geoCoordinate id="AEF714E059DAD9AD4F2BAB3A9A180946" degrees="18.4104" direction="south" orientation="latitude" precision="5" value="-18.4104">-18.4104</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="48.8189" direction="east" orientation="longitude" precision="5" value="48.8189">48.8189</geoCoordinate>
|
||||
<geoCoordinate id="FDD440CDEAC88194A4DE00D96B6854D8" degrees="48.8189" direction="east" orientation="longitude" precision="5" value="48.8189">48.8189</geoCoordinate>
|
||||
; alt. 855 m; 22 Jan 2012; B.L. Fisher et al. leg.; CASENT0274408, CASENT0275713 (CASC). •2w.; Ankerana;
|
||||
<geoCoordinate degrees="18.40062" direction="south" orientation="latitude" precision="1" value="-18.40062">-18.40062</geoCoordinate>
|
||||
<geoCoordinate id="05DED4B11022C152B59A9550FB447F0B" degrees="18.40062" direction="south" orientation="latitude" precision="1" value="-18.40062">-18.40062</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="48.81311" direction="east" orientation="longitude" precision="1" value="48.81311">48.81311</geoCoordinate>
|
||||
<geoCoordinate id="0DE0094551BA2254394261ADFDC37C15" degrees="48.81311" direction="east" orientation="longitude" precision="1" value="48.81311">48.81311</geoCoordinate>
|
||||
; alt. 865 m; 17 Jan 2012; B.L. Fisher et al. leg.; CASENT0274909 (CASC). •1w., 1s.; Parc National de Zahamena, Besaky River;
|
||||
<geoCoordinate degrees="17.75244" direction="south" orientation="latitude" precision="1" value="-17.75244">-17.75244</geoCoordinate>
|
||||
<geoCoordinate id="B6FE70AEDA687DB845C42ED0D35FD160" degrees="17.75244" direction="south" orientation="latitude" precision="1" value="-17.75244">-17.75244</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="48.85321" direction="east" orientation="longitude" precision="1" value="48.85321">48.85321</geoCoordinate>
|
||||
<geoCoordinate id="AB6C0D96DA555D07B1863C3BCA392A03" degrees="48.85321" direction="east" orientation="longitude" precision="1" value="48.85321">48.85321</geoCoordinate>
|
||||
; alt. 760 m; 22 Feb 2009; B.L. Fisher et al. leg.; CASENT0149769 (CASC).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||||
<subSubSection id="4420328281D0F9F4F67E69622A25D0FE" pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph id="C51A590FD526404102BBD477F37E2737" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph id="8554689F3242474BBA8DBECE3FF2B8C3" pageId="0" pageNumber="1">
|
||||
<emphasis id="85BB1509B9584E31F3D5CD1C97B40B8D" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="62DF07DF54DA4FF9984435F62E027C31" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Head in full-face view rectangular, with lateral sides relatively straight, only posteriormost part slightly convex; sides of the head with dense, long, suberect pilosity; occipital lobes and genae shiny, with thick, sparse, irregular rugae; frons with sparse, thick, and longitudinal rugae, interspaces smooth; inner hypostomal teeth distinct, small, and low, closely spaced, lobe-like, with rounded apex and wide base; outer hypostomal teeth bigger and wider than inner hypostomal teeth, triangular; inner and outer teeth closely spaced and connected by concavity; propodeal spines moderately long, triangular, with acute apex; gaster smooth; body reddish brown.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||||
<emphasis id="7D4CE41DA52C38ECA634AE1A29C61964" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="3FE3C46ADD4E13DE9DD1549FCEFA25E1" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Head foveolate with additional thin, sparse rugae on the whole surface; promesonotum low, short, slightly convex, with relatively steep posterior declivity; propodeal spines minute, triangular; mesosoma foveolate with additional thin, sparse rugae, katepisternum smooth.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||||
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||||
<subSubSection id="05F8264E03A6C0E6E78D0E0ACC012FF3" pageId="0" pageNumber="1" type="description">
|
||||
<paragraph id="CB4658168C37BC2EAD15D48F5493B6EF" pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph id="401FCA43CBFED8E6D22D431DA1DB484D" pageId="0" pageNumber="1">
|
||||
<emphasis id="144918B55747998E373207E3E5EDD71E" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||||
Measurements (
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
<emphasis id="4B763AF9F722FC0B332DFA6CBEB6D214" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
= 5): HL: 1.42-1.52 (1.48); HW: 1.34-1.43 (1.38); SL: 0.61-0.65 (0.63); EL: 0.16-0.2 (0.18); WL: 1.07-1.16 (1.12); PSL: 0.19-0.21 (0.2); MTL: 0.65-0.68 (0.66); PNW: 0.62-0.67 (0.64); PTW: 0.15-0.17 (0.16); PPW: 0.47-0.51 (0.49); CI: 92.1-94.1 (93.3); SI: 44.7-46.2 (45.5); PSLI: 12.1-14.6 (13.4); PPI: 28.9-32.7 (31.6); PNI: 45.7-47.5 (46.6); MTI: 47.4-48.8 (48.3).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
<emphasis id="36640E64845E254A3FA5489CBB04255D" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="3BE157E6A0A1556724AE8D02924DE702" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
In full-face view rectangular, with lateral sides relatively straight, only posteriormost part slightly convex (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71B</figureCitation>
|
||||
<figureCitation id="F56B6EC2F889F2938FDBA8684651E594" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71B</figureCitation>
|
||||
). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming shallow transverse depression between frons and occipital lobes; inner hypostomal teeth invisible. Sides of the head with dense, long, suberect pilosity; whole head with dense, moderately long, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, with dense, thick, and irregular rugoreticulation. Occipital lobes and genae shiny, with thick, sparse, irregular rugae; frons with sparse, thick, and longitudinal rugae, interspaces smooth; malar area with dense, thick, and longitudinal rugae, interspaces smooth to finely rugulose. Centre of clypeus smooth and shiny, lateral sides with longitudinal rugae; median notch present, narrow and shallow to moderate; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, slightly exceeding the midlength of head; pilosity decumbent to erect (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71B, D</figureCitation>
|
||||
<figureCitation id="4316FC8D73F64630E50DE79583C1B247" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71B, D</figureCitation>
|
||||
). Inner hypostomal teeth distinct, small and low, closely spaced, lobe-like, with rounded apex and wide base; outer hypostomal teeth bigger and wider than inner hypostomal teeth, triangular; inner and outer teeth closely spaced and connected by concavity (Fig.
|
||||
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84D</figureCitation>
|
||||
<figureCitation id="E2EEA2277BDBBCE1445055913D8608D2" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84D</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
<emphasis id="C805DC8E473B1785342B19FBAC4CA8B6" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="E5E337C6F335005B1C326A2D9274CCBF" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
In lateral view, promesonotum short, angular, and low, posterior mesonotum relatively steep, with indistinct tubercle-like projection; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, triangular, with acute apex; humeral area laterally weakly produced (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D</figureCitation>
|
||||
<figureCitation id="6503B924043EE4C4BFA9E34BE2956ED1" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D</figureCitation>
|
||||
). Surface shiny, with fine to thin, dense rugoreticulation, sculpture weakening on dorsum, katepisternum smooth. Pilosity moderately sparse, long, and erect (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
|
||||
<figureCitation id="3D946DC8D0ECC95C34851A77AD156F73" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="05B5351440F70B746733CFB887AB8999" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="58CC32593D51A80817CE776777B2B494" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Shiny; peduncle shagreened, short, without horizontal lobes on its basal part; node smooth, relatively high and thin, triangular, with rounded apex, in rear view node straight; pilosity moderately sparse and erect (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
|
||||
<figureCitation id="9DC7CC1C286BADCC61CDBD1BC6A53742" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
<emphasis id="1A52C2A8B824F4F44B3F3A730E3938CF" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="3161B2C794E614BD8D20DB3337E00CAF" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Finely shagreened, shiny; in dorsal view sides with moderately short, acute, and horn-like projections; pilosity long, moderately sparse and erect (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
|
||||
<figureCitation id="63984CDD33E6D3E6C2684F8E21AFD0A0" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="2860A3C17CA6FEA16FCE7C9C18EB34CC" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="FEC23A05EC0FA1A7E270125260C86F74" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Smooth; pilosity moderately dense, long, and erect (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
|
||||
<figureCitation id="2556DF9811E02776B7CC4D758A0A6EB0" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
<emphasis id="40DD04EA772AB9E8FDCA736676A1541C" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="D94EE3F5372F97A93DB8C4BD826A27A0" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Unicolourous, reddish brown; legs dark yellow (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
|
||||
<figureCitation id="FD6DC2E8D5D4C23A71777B1170A67790" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1" start="Figure 71" startId="F71">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 71.</emphasis>
|
||||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ankerana</emphasis>
|
||||
<caption id="A42DC3B37F9CE71FCCC4DE079C8F7B0D" doi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1" start="Figure 71" startId="F71">
|
||||
<paragraph id="8D4D9F4629678680D640E5636BEE4DDC" pageId="0" pageNumber="1">
|
||||
<emphasis id="5AA625DC7FC5DD0E7A80A40B97ABB946" bold="true" pageId="0" pageNumber="1">Figure 71.</emphasis>
|
||||
<taxonomicName id="4AFE2D83E38CA6C1906C8687FBFD46D8" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana">
|
||||
<emphasis id="A221ED0EAED1B82CAC9299682F8D9284" italics="true" pageId="0" pageNumber="1">Pheidole ankerana</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., full-face view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<emphasis id="6F3DF0D21B46B6B445503A5417C63782" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
), profile (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<emphasis id="8BAF79266BCC3759E3CFDCDCC252068A" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
), and dorsal view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<emphasis id="D5F9A9EB8943A6F5F68A48F9511798E5" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
) of paratype minor worker (CASENT0274003) and full-face view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<emphasis id="A245A6940CBC77DCCD3E594B955CCEF9" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
), profile (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<emphasis id="6C06F414BDF12AB9CC013CA317F3C125" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
), and dorsal view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
<emphasis id="0ECC5C4CB6F3CEE20133AD9692B950ED" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
) of holotype major worker (CASENT0923171).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||||
<paragraph id="C8D55398111E34A2C0931A841824ABAB" pageId="0" pageNumber="1">
|
||||
<emphasis id="334534ED487095051A72AF168623FFF6" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||||
Measurements (
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
<emphasis id="43FC0C2A44B2A7D03229877887CEF00F" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
= 5): HL: 0.61-0.69 (0.66); HW: 0.55-0.63 (0.59); SL: 0.58-0.64 (0.61); EL: 0.11-0.13 (0.12); WL: 0.75-0.85 (0.8); PSL: 0.09-0.11 (0.1); MTL: 0.48-0.55 (0.51); PNW: 0.37-0.43 (0.4); PTW: 0.06-0.08 (0.07); PPW: 0.13-0.2 (0.16); CI: 88.5-93.8 (90.3); SI: 99.2-105.1 (102.4); PSLI: 14.4-17.3 (15.5); PPI: 38.2-52.7 (45.5); PNI: 66.1-68.6 (67.6); MTI: 79.5-90.1 (86.9).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
<emphasis id="1D733188C202BDA5EB5954EA97AE7CF2" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="E921AC53A4B37994219362632A963ED7" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Occipital margin straight or indistinctly concave; occipital carina indistinct, weakly developed (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71A</figureCitation>
|
||||
<figureCitation id="39462E5B9E9B3280A2633756695DC158" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71A</figureCitation>
|
||||
). Pilosity moderately dense, long, erect. Whole head foveolate with additional thin, sparse rugae on the whole surface. Clypeus shiny, with sparse, irregular rugae; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity erect (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71A, C</figureCitation>
|
||||
<figureCitation id="77257CC26E2CDA2D6DF20DDCA5E9C27E" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71A, C</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
<emphasis id="B309460AEA0B84D956AEF0DD065EE552" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="C398B8A1FDF551A3F4A29DF3B76721DF" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
In lateral view, promesonotum low, short, slightly convex, with relatively steep posterior declivity; promesonotal groove absent; metanotal groove absent; propodeal spines minute, triangular, apex acute (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C</figureCitation>
|
||||
<figureCitation id="0257A840B81E813F0586361154E3DDA5" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C</figureCitation>
|
||||
). Sculpture foveolate with additional thin, sparse rugae on the whole surface, katepisternum smooth. Pilosity sparse, long, and erect (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
||||
<figureCitation id="1B67B3D28868FDA9BEA15F431E7320E8" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="56CDA8CDDD1972D949F9B90CFBABCE72" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="79DB54235AED5DA3A48A6C2449EB1CA1" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Peduncle short and thin; node moderately high, triangular and small; with few long, erect setae (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
||||
<figureCitation id="DF890C3708A4235457662DACEC22CFD2" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
<emphasis id="AED2827EE87759B21202451B1AFADF94" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="2666D6BBA40EE45E00140374022E4135" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Short, low, and slightly convex; with few short, erect setae (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
||||
<figureCitation id="CD4278FD21DC71DCD1B8A1804E178B78" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="F80122C514FF2FB0D0D3AA2F6A9E08A5" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="7AFA1304DB1342490ADFED505B289539" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
With moderately sparse, erect pilosity (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
||||
<figureCitation id="917BC3DC4586AABC8A97763DC1385AD7" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
<emphasis id="FAAE84423E5DC989D9302939DDC11057" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="0D08BAFF928D9AD9831B79961775E90B" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Unicolourous, yellow (Fig.
|
||||
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
||||
<figureCitation id="09421BA88E431DFD287021DAD68EC690" captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
|
||||
<subSubSection id="463DC17D81B8171AB58CE68554D591CA" pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph id="E75BF75DB7897F4BBC7A333056E1D8E6" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph id="AF206F36DF9A84A58C33D114A1E1F172" pageId="0" pageNumber="1">From the type locality.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">The species was collected between 725-865 m in elevation, in rainforest. Nests were located in rotten logs.</paragraph>
|
||||
<subSubSection id="2832CE9973F9F0E48ECD0D45675EE6F4" pageId="0" pageNumber="1" type="biology_ecology">
|
||||
<paragraph id="CF01C15D275B2AACAC24642BCECB6F91" pageId="0" pageNumber="1">Biology.</paragraph>
|
||||
<paragraph id="1EB1367EFF17CAA9EA9A2F013AD4F99A" pageId="0" pageNumber="1">The species was collected between 725-865 m in elevation, in rainforest. Nests were located in rotten logs.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="comments">
|
||||
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||||
<subSubSection id="641406A054BFEAF44BF1DF64E6B8A623" pageId="0" pageNumber="1" type="comments">
|
||||
<paragraph id="01C906DF9DB9EC94F0600C3EFE30ADFA" pageId="0" pageNumber="1">Comments.</paragraph>
|
||||
<paragraph id="60CE75D381876BEFDE65F29560A23F93" pageId="0" pageNumber="1">
|
||||
<emphasis id="2118C3FF7980828769FF1409D69E5DCD" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="3E0AF402EF47416C1A3B4C0C9EF0A89E" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ankerana</emphasis>
|
||||
<taxonomicName id="5AEECEFD5E92E0DEA7E06E7DAE9F32FE" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana">
|
||||
<emphasis id="C2CC32439139851F18580C19DEC4420A" italics="true" pageId="0" pageNumber="1">Pheidole ankerana</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. is most similar to
|
||||
<taxonomicName lsidName="P. vatovavensis" pageId="0" pageNumber="1" rank="species" species="vatovavensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">P. vatovavensis</emphasis>
|
||||
<taxonomicName id="F78354E42FFBAE33375E20BDCBCC7998" lsidName="P. vatovavensis" pageId="0" pageNumber="1" rank="species" species="vatovavensis">
|
||||
<emphasis id="531D783916008B09961AF65E65C5B7FC" italics="true" pageId="0" pageNumber="1">P. vatovavensis</emphasis>
|
||||
</taxonomicName>
|
||||
. It differs from
|
||||
<taxonomicName lsidName="P. vatovavensis" pageId="0" pageNumber="1" rank="species" species="vatovavensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">P. vatovavensis</emphasis>
|
||||
<taxonomicName id="AD17DEABC343DB9A37967E135EF6CFA2" lsidName="P. vatovavensis" pageId="0" pageNumber="1" rank="species" species="vatovavensis">
|
||||
<emphasis id="9F069DCE7C0CCA4809B3A951D78867AE" italics="true" pageId="0" pageNumber="1">P. vatovavensis</emphasis>
|
||||
</taxonomicName>
|
||||
in denser and longer pilosity on the sides of head, smooth sculpture of interspaces on frons, sculpture of scrobes never foveolate, lower promesonotum and lobe-like shape of inner hypostomal teeth.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||||
<emphasis id="F192268BDB74AE3A914BFD7FB7D195A2" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="C12E26B713D7BCF0B8DF93F170B100AB" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ankerana</emphasis>
|
||||
<taxonomicName id="D36CAA5C96D6744DDBCAE5222D914865" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana">
|
||||
<emphasis id="72505C37E4CC6165C96FADC212BEC00B" italics="true" pageId="0" pageNumber="1">Pheidole ankerana</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. differs from
|
||||
<taxonomicName lsidName="P. vatovavensis" pageId="0" pageNumber="1" rank="species" species="vatovavensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">P. vatovavensis</emphasis>
|
||||
<taxonomicName id="941E5C2EACBE8EEE5308F2ED7AF14937" lsidName="P. vatovavensis" pageId="0" pageNumber="1" rank="species" species="vatovavensis">
|
||||
<emphasis id="DC6FFA0DB8203C5A2974C7FB90D59ECF" italics="true" pageId="0" pageNumber="1">P. vatovavensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. by presence of additional thin, sparse rugae on the whole surface of head and mesosoma and minute size of propodeal spines.
|
||||
</paragraph>
|
||||
|
|
|
|||
Loading…
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Reference in a new issue