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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1019.60364" ID-GBIF-Dataset="cc04dabd-7ac1-4177-b2ed-6024ebed4dee" ID-PMC="PMC7921072" ID-Pensoft-Pub="1313-2970-1019-35" ID-Pensoft-UUID="8D799427736453288CA3E1DB9933AAC6" ID-PubMed="33679171" ID-ZooBank="EE1BAF875BD74E189DF929E2C9CC7DAC" ModsDocID="1313-2970-1019-35" checkinTime="1614092287051" checkinUser="pensoft" docAuthor="Hansson, Christer" docDate="2021" docId="EC283969F287567FAB77CAF4C24773BD" docLanguage="en" docName="ZooKeys 1019: 35-91" docOrigin="ZooKeys 1019" docPubDate="2021-02-22" docSource="http://dx.doi.org/10.3897/zookeys.1019.60364" docTitle="Eriastichus vestis Hansson 2021, sp. nov." docType="treatment" docUuid="59E0A806-AAE9-4C16-B373-23BBE4FE3BDC" docUuidSource="ZooBank" docVersion="4" id="8D799427736453288CA3E1DB9933AAC6" lastPageNumber="35" masterDocId="8D799427736453288CA3E1DB9933AAC6" masterDocTitle="The genus Eriastichus La Salle (Hymenoptera, Eulophidae, Tetrastichinae) in the Neotropical region, introducing 48 new species" masterLastPageNumber="91" masterPageNumber="35" pageNumber="35" updateTime="1668149819104" updateUser="ExternalLinkService">
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<mods:title>The genus Eriastichus La Salle (Hymenoptera, Eulophidae, Tetrastichinae) in the Neotropical region, introducing 48 new species</mods:title>
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<mods:title id="6A13983E0688D9B346C2915DD929C320">The genus Eriastichus La Salle (Hymenoptera, Eulophidae, Tetrastichinae) in the Neotropical region, introducing 48 new species</mods:title>
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<mods:namePart>Hansson, Christer</mods:namePart>
<mods:affiliation>Scientific Associate Biological Museum (Entomology), Lund University, Soelvegatan 37, SE- 22362 Lund, Sweden &amp; Natural History Museum, Life Sciences, Cromwell Road, London, UK</mods:affiliation>
<mods:nameIdentifier type="email">christer.hansson@biol.lu.se</mods:nameIdentifier>
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<mods:affiliation id="A181956A3C301A3A23B823604ADC05E6">Scientific Associate Biological Museum (Entomology), Lund University, Soelvegatan 37, SE- 22362 Lund, Sweden &amp; Natural History Museum, Life Sciences, Cromwell Road, London, UK</mods:affiliation>
<mods:nameIdentifier id="9B7672E6519A29048B6B78EFC6422923" type="email">christer.hansson@biol.lu.se</mods:nameIdentifier>
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<subSection pageId="0" pageNumber="35" type="taxonomic accounts">
<treatment ID-GBIF-Taxon="179904964" LSID="urn:lsid:zoobank.org:act:59E0A806-AAE9-4C16-B373-23BBE4FE3BDC" httpUri="http://treatment.plazi.org/id/EC283969F287567FAB77CAF4C24773BD" lastPageNumber="35" pageId="0" pageNumber="35">
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<taxonomicName LSID="http://zoobank.org/59E0A806-AAE9-4C16-B373-23BBE4FE3BDC" authority="Hansson, 2021" authorityName="Hansson" authorityYear="2021" class="Insecta" family="Eulophidae" genus="Eriastichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriastichus vestis" order="Hymenoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="vestis" status="sp. nov.">Eriastichus vestis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="35">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 5760" captionStartId="F9" captionText="Figures 57 - 60. Eriastichus spp. 57, 58 head and antenna in lateral view, male holotypes: 57 E. tonioazofeifai 58 E. vestis; 59, 60 head in frontal view, male holotypes: 59 E. pallidops 60 E. denotatis." figureDoi="10.3897/zookeys.1019.60364.figures57-60" httpUri="https://binary.pensoft.net/fig/512586" pageId="0" pageNumber="35">Figure 58</figureCitation>
<subSection id="20062145CDF1606E7939BB76157712D6" pageId="0" pageNumber="35" type="taxonomic accounts">
<treatment id="EC283969F287567FAB77CAF4C24773BD" ID-GBIF-Taxon="179904964" LSID="urn:lsid:zoobank.org:act:59E0A806-AAE9-4C16-B373-23BBE4FE3BDC" httpUri="http://treatment.plazi.org/id/EC283969F287567FAB77CAF4C24773BD" lastPageNumber="35" pageId="0" pageNumber="35" scope_family="Eulophidae" scope_order="Hymenoptera">
<subSubSection id="FA1483140FE3CA1A3E1BE4F798024A05" pageId="0" pageNumber="35" type="nomenclature">
<paragraph id="6E527A4A1E42C783D422436B3EBE0BB5" pageId="0" pageNumber="35">
<taxonomicName id="594045C2A9B3BC22E01BA18B4B095405" ID-CoL="C9NHF" LSID="http://zoobank.org/59E0A806-AAE9-4C16-B373-23BBE4FE3BDC" authority="Hansson, 2021" authorityName="Hansson" authorityYear="2021" class="Insecta" family="Eulophidae" genus="Eriastichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriastichus vestis" order="Hymenoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="vestis" status="sp. nov.">Eriastichus vestis</taxonomicName>
<taxonomicNameLabel id="41C8037F7BEBF5DAE981EC8C31D30E74" pageId="0" pageNumber="35">sp. nov.</taxonomicNameLabel>
<figureCitation id="FCB46594532D66FB3D5AE9EF8476F45F" captionStart="Figures 5760" captionStartId="F9" captionText="Figures 57 - 60. Eriastichus spp. 57, 58 head and antenna in lateral view, male holotypes: 57 E. tonioazofeifai 58 E. vestis; 59, 60 head in frontal view, male holotypes: 59 E. pallidops 60 E. denotatis." figureDoi="10.3897/zookeys.1019.60364.figures57-60" httpUri="https://binary.pensoft.net/fig/512586" pageId="0" pageNumber="35">Figure 58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="type locality">
<paragraph pageId="0" pageNumber="35">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="35">
<subSubSection id="75CE544027B311FBA995AEA0FE074BFC" pageId="0" pageNumber="35" type="type locality">
<paragraph id="39DA97F8EF7025F86D5B35985E60C86C" pageId="0" pageNumber="35">Type locality.</paragraph>
<paragraph id="9A31E8307FAF1C23D59661CCDBAA04B8" pageId="0" pageNumber="35">
Costa Rica, Puntarenas, San Vito,
<normalizedToken originalValue="Estación">Estacion</normalizedToken>
<normalizedToken originalValue="Biológica">Biologica</normalizedToken>
<normalizedToken id="AEB192BDCCD6C226F8043F798AFB5E01" originalValue="Estación">Estacion</normalizedToken>
<normalizedToken id="F95D8D1A290656183230383E36A96AC4" originalValue="Biológica">Biologica</normalizedToken>
Las Alturas, 1500 m,
<geoCoordinate degrees="8" direction="north" minutes="57" orientation="latitude" precision="925" value="8.95">8°57'N</geoCoordinate>
<geoCoordinate id="FBC1390402D11DDF0701E13D301A3FD9" degrees="8" direction="north" minutes="57" orientation="latitude" precision="925" value="8.95">8°57'N</geoCoordinate>
,
<geoCoordinate degrees="82" direction="west" minutes="50" orientation="longitude" precision="925" value="-82.833336">82°50'W</geoCoordinate>
<geoCoordinate id="5331D8C3AA92CE95CBDA27DBA8A86814" degrees="82" direction="west" minutes="50" orientation="longitude" precision="925" value="-82.833336">82°50'W</geoCoordinate>
, 17-18.ii.2012, J.S. Noyes leg.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="type specimen">
<paragraph pageId="0" pageNumber="35">Type specimen.</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
<subSubSection id="3F33FF30CB17885B766625CC7255A1BE" pageId="0" pageNumber="35" type="type specimen">
<paragraph id="75868031CC8064548473685E572F06CA" pageId="0" pageNumber="35">Type specimen.</paragraph>
<paragraph id="4E8921460E21F0BE39F7377DAB003449" pageId="0" pageNumber="35">
<emphasis id="E6B00C81109A581AF5EC9477CF9F2E23" bold="true" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
male dried and glued to a paper card. Original labels: &quot;COSTA RICA, Puntarenas, San Vito, E.B. Las Alturas, 1500 m,
<geoCoordinate degrees="8" direction="north" minutes="57" orientation="latitude" precision="925" value="8.95">8°57'N</geoCoordinate>
<geoCoordinate id="92539D3B06A96A4490CB78B49E6D3A8A" degrees="8" direction="north" minutes="57" orientation="latitude" precision="925" value="8.95">8°57'N</geoCoordinate>
,
<geoCoordinate degrees="82" direction="west" minutes="50" orientation="longitude" precision="925" value="-82.833336">82°50'W</geoCoordinate>
<geoCoordinate id="BA7DA28BD90A023492019C8F37A69828" degrees="82" direction="west" minutes="50" orientation="longitude" precision="925" value="-82.833336">82°50'W</geoCoordinate>
, 17-18.ii.2012, J.S. Noyes, NHM (Ent) 2012-91&quot;, &quot;HOLOTYPE
<taxonomicName authorityName="Hansson" authorityYear="2021" class="Insecta" family="Eulophidae" genus="Eriastichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriastichus vestis" order="Hymenoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="vestis">Eriastichus vestis</taxonomicName>
<taxonomicName id="6FBF2821CF4C063835CB93DEB210724B" authorityName="Hansson" authorityYear="2021" class="Insecta" family="Eulophidae" genus="Eriastichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriastichus vestis" order="Hymenoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="vestis">Eriastichus vestis</taxonomicName>
Hansson&quot; [red printed label], (NHMUK014431074).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="diagnosis">
<paragraph pageId="0" pageNumber="35">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" pageId="0" pageNumber="35">(male).</emphasis>
<subSubSection id="56222FF7EE19BEC16AFC83472D47987A" pageId="0" pageNumber="35" type="diagnosis">
<paragraph id="C3D1BCAAB064BCAF3884044A58D688BF" pageId="0" pageNumber="35">Diagnosis</paragraph>
<paragraph id="A6B7A03CE32D9AF4400443C85CDF4817" pageId="0" pageNumber="35">
<emphasis id="49A50DFE1894F6F26C32D6E469984CB4" bold="true" pageId="0" pageNumber="35">(male).</emphasis>
Head dark brown, scape and pedicel pale brown, flagellum brown; ventral plaque on scape ca. 0.3
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="B27995870234B2A73D1D97FE98882C06" originalValue="×">x</normalizedToken>
as long as scape (Fig.
<figureCitation captionStart="Figures 5760" captionStartId="F9" captionText="Figures 57 - 60. Eriastichus spp. 57, 58 head and antenna in lateral view, male holotypes: 57 E. tonioazofeifai 58 E. vestis; 59, 60 head in frontal view, male holotypes: 59 E. pallidops 60 E. denotatis." figureDoi="10.3897/zookeys.1019.60364.figures57-60" httpUri="https://binary.pensoft.net/fig/512586" pageId="0" pageNumber="35">58</figureCitation>
<figureCitation id="2C2C4A32D34463A5971A98891432D223" captionStart="Figures 5760" captionStartId="F9" captionText="Figures 57 - 60. Eriastichus spp. 57, 58 head and antenna in lateral view, male holotypes: 57 E. tonioazofeifai 58 E. vestis; 59, 60 head in frontal view, male holotypes: 59 E. pallidops 60 E. denotatis." figureDoi="10.3897/zookeys.1019.60364.figures57-60" httpUri="https://binary.pensoft.net/fig/512586" pageId="0" pageNumber="35">58</figureCitation>
), antenna with dorsobasal setae on F1 0.9
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="5A2EAF348897F39C2642B36404E82847" originalValue="×">x</normalizedToken>
as long as F1; gaster with lateral tufts of pale and flattened setae on Gt6.
</paragraph>
<caption doi="10.3897/zookeys.1019.60364.figures57-60" httpUri="https://binary.pensoft.net/fig/512586" pageId="0" pageNumber="35" start="Figures 5760" startId="F9">
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" pageId="0" pageNumber="35">Figures 57-60.</emphasis>
<taxonomicName authorityName="La Salle" authorityYear="1994" class="Insecta" family="Eulophidae" genus="Eriastichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriastichus" order="Hymenoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="35">Eriastichus</emphasis>
<caption id="9C9FA6BA6630E9810D3D1A8C59CF3EDC" doi="10.3897/zookeys.1019.60364.figures57-60" httpUri="https://binary.pensoft.net/fig/512586" pageId="0" pageNumber="35" start="Figures 5760" startId="F9">
<paragraph id="D5E31C66826A8C30B996D36CFC608506" pageId="0" pageNumber="35">
<emphasis id="AFF38E64A7882C97FEB6EFBB413D2BA4" bold="true" pageId="0" pageNumber="35">Figures 57-60.</emphasis>
<taxonomicName id="1B592A8835D9281C55366071FB106F61" authorityName="La Salle" authorityYear="1994" class="Insecta" family="Eulophidae" genus="Eriastichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriastichus" order="Hymenoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="ED8EDA01A18F5158D3BC13CC17DA6952" italics="true" pageId="0" pageNumber="35">Eriastichus</emphasis>
</taxonomicName>
spp.
<emphasis bold="true" pageId="0" pageNumber="35">57, 58</emphasis>
<emphasis id="E7066AD6AB02387B95C22AC29CE24A6D" bold="true" pageId="0" pageNumber="35">57, 58</emphasis>
head and antenna in lateral view, male holotypes:
<emphasis bold="true" pageId="0" pageNumber="35">57</emphasis>
<taxonomicName lsidName="E. tonioazofeifai" pageId="0" pageNumber="35" rank="species" species="tonioazofeifai">
<emphasis italics="true" pageId="0" pageNumber="35">E. tonioazofeifai</emphasis>
<emphasis id="D7C994CF1399C3ACB2B1D123FF39A93C" bold="true" pageId="0" pageNumber="35">57</emphasis>
<taxonomicName id="A9D7E752289A7846DFC5EB66D5A24E75" lsidName="E. tonioazofeifai" pageId="0" pageNumber="35" rank="species" species="tonioazofeifai">
<emphasis id="ADF50901334EFB8ACE10F73BADF0EB76" italics="true" pageId="0" pageNumber="35">E. tonioazofeifai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="35">58</emphasis>
<taxonomicName lsidName="E. vestis" pageId="0" pageNumber="35" rank="species" species="vestis">
<emphasis italics="true" pageId="0" pageNumber="35">E. vestis</emphasis>
<emphasis id="1AEBA29651357EA57402A991CEF141CF" bold="true" pageId="0" pageNumber="35">58</emphasis>
<taxonomicName id="6F7E0C39D30A35BE34567CB718AE25EE" lsidName="E. vestis" pageId="0" pageNumber="35" rank="species" species="vestis">
<emphasis id="104DC860446F09F73A66501D63C02A18" italics="true" pageId="0" pageNumber="35">E. vestis</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="35">59, 60</emphasis>
<emphasis id="C9F8A1DFF62221B60D3D3AD06C3C0E0D" bold="true" pageId="0" pageNumber="35">59, 60</emphasis>
head in frontal view, male holotypes:
<emphasis bold="true" pageId="0" pageNumber="35">59</emphasis>
<taxonomicName lsidName="E. pallidops" pageId="0" pageNumber="35" rank="species" species="pallidops">
<emphasis italics="true" pageId="0" pageNumber="35">E. pallidops</emphasis>
<emphasis id="EEB2377C87466E425FA23C2F1B3BD76C" bold="true" pageId="0" pageNumber="35">59</emphasis>
<taxonomicName id="679F71A27A01A55C7400A3A08881F002" lsidName="E. pallidops" pageId="0" pageNumber="35" rank="species" species="pallidops">
<emphasis id="116839E0BFE9486B1D24D284A692A0F5" italics="true" pageId="0" pageNumber="35">E. pallidops</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="35">60</emphasis>
<taxonomicName lsidName="E. denotatis" pageId="0" pageNumber="35" rank="species" species="denotatis">
<emphasis italics="true" pageId="0" pageNumber="35">E. denotatis</emphasis>
<emphasis id="4519E54245B4A774B38B721B7FC5031B" bold="true" pageId="0" pageNumber="35">60</emphasis>
<taxonomicName id="7525B4D331F2EB914A1413F9ABA3C062" lsidName="E. denotatis" pageId="0" pageNumber="35" rank="species" species="denotatis">
<emphasis id="10FC715A8FB95937524F9D27082F24FE" italics="true" pageId="0" pageNumber="35">E. denotatis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="35" type="description">
<paragraph pageId="0" pageNumber="35">Description</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" pageId="0" pageNumber="35">(male holotype NHMUK014431074).</emphasis>
<subSubSection id="59236BD211DC7705F6B0CF358BD1E088" pageId="0" pageNumber="35" type="description">
<paragraph id="5D8AF84BC31A3205AC4F7D03C4A05120" pageId="0" pageNumber="35">Description</paragraph>
<paragraph id="E0D9703F718A579B38D0193952D6674D" pageId="0" pageNumber="35">
<emphasis id="6454BC40538C2FEBBE55113D3D965281" bold="true" pageId="0" pageNumber="35">(male holotype NHMUK014431074).</emphasis>
Length of body 1.5 mm. Head dark brown, scape and pedicel pale brown, flagellum brown, ventral plaque dark brown. Mesoscutum, mesoscutellum and propodeum black with metallic tinges, dorsellum dark brown. Legs with coxae and femora dark brown, remaining parts of legs yellowish brown. Gaster dark brown.
</paragraph>
<paragraph pageId="0" pageNumber="35">
<emphasis bold="true" pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Head</emphasis>
<paragraph id="101841F4D7CB642546E105FD97508D42" pageId="0" pageNumber="35">
<emphasis id="149C55E01F0B2776AC2625145C31AF70" bold="true" pageId="0" pageNumber="35">
<emphasis id="B0B70CA83A946CE5630C02366D784690" italics="true" pageId="0" pageNumber="35">Head</emphasis>
.
</emphasis>
Length/width in frontal view 0.8; width/length in dorsal view 2.4; POL/OOL 1.9; WM/MS 1.5; MS/HE 0.6; HE/head length in frontal view 0.5; widths head/mesoscutum 1.0.
<emphasis bold="true" pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Antenna</emphasis>
<emphasis id="07CD58691569E5FA3618D484884A0C34" bold="true" pageId="0" pageNumber="35">
<emphasis id="0CF29EE6C9B460586733FFC0663A10C4" italics="true" pageId="0" pageNumber="35">Antenna</emphasis>
.
</emphasis>
Pedicel + flagellum length/mesoscutum width 1.8; pedicel + flagellum length/head width 1.8; lengths scape/ventral plaque 2.9; ventral plaque located above the middle of scape; scape length/width 2.9; lengths scape/head (dorsal view) 0.5; scape length/HE 1.0; length/width F1, F2, F3, F4, clava: 3.0, 3.2, 3.0, 3.0, 7.4; length dorsobasal setae on F1/length F1 0.9.
<emphasis bold="true" pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Mesosoma</emphasis>
<emphasis id="C927FDB8270EFAA6868F1DA12CB2370F" bold="true" pageId="0" pageNumber="35">
<emphasis id="2C0141B8598D936E899E4F9EEFD2AB8C" italics="true" pageId="0" pageNumber="35">Mesosoma</emphasis>
.
</emphasis>
Length/width 1.5; mesoscutum length/width 0.6; mesoscutellum length/width 0.7; widths SMG/SLG 1.5; enclosed space between SMG length/width 1.7; lengths mesoscutum/mesoscutellum 1.4; lengths mesoscutellum/dorsellum 2.5; lengths mesosoma/gaster 0.9.
<emphasis bold="true" pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Wings</emphasis>
<emphasis id="675B2F40F4C8C6CE52547DFE548DD82C" bold="true" pageId="0" pageNumber="35">
<emphasis id="8FF2191E2136CA26E43B4D084D13B414" italics="true" pageId="0" pageNumber="35">Wings</emphasis>
.
</emphasis>
CC length/width 16.0; lengths CC/MV 1.0; lengths MV/ST 2.0; lengths MV/PM 5.3; lengths PM/ST 0.4; submarginal vein with five setae on dorsal surface.
<emphasis bold="true" pageId="0" pageNumber="35">
<emphasis italics="true" pageId="0" pageNumber="35">Gaster</emphasis>
<emphasis id="DDF2EC374D8E771AF9FDFD9881CC34B3" bold="true" pageId="0" pageNumber="35">
<emphasis id="71E6C642911CE1938B75227C386C1405" italics="true" pageId="0" pageNumber="35">Gaster</emphasis>
.
</emphasis>
With lateral tufts of pale and flattened setae on Gt6.

View file

@ -1,354 +1,354 @@
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<treatment ID-GBIF-Taxon="161777038" LSID="urn:lsid:zoobank.org:act:0E6A56CB-6415-4402-A3C9-80FFD4F86BF7" httpUri="http://treatment.plazi.org/id/ECDAF5B877D124C1802B904D5509173E" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/0E6A56CB-6415-4402-A3C9-80FFD4F86BF7" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina" status="sp. nov.">Pheidole maizina</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">Figs 27A-F</figureCitation>
<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85C</figureCitation>
<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87G</figureCitation>
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<paragraph id="F5587B5469A0C1A348C5BAD992E22A4F" pageId="0" pageNumber="1">
<taxonomicName id="1558B67E0464B96EFD130A3A2AB8D40F" ID-CoL="CMLFT" LSID="http://zoobank.org/0E6A56CB-6415-4402-A3C9-80FFD4F86BF7" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina" status="sp. nov.">Pheidole maizina</taxonomicName>
<taxonomicNameLabel id="A7C2B12DCB6F1E2729B45B9714CF5830" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="1FED6A2ACB7A2534A0B65422E651A303" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">Figs 27A-F</figureCitation>
<figureCitation id="A4F90C67C6B79D240B95ABC6DF6EF47D" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85C</figureCitation>
<figureCitation id="BF927FD7BBE60476B10E811F36B29710" captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
<subSubSection id="D58D0C409CFDF72E766DD1E105059F43" pageId="0" pageNumber="1" type="type material">
<paragraph id="C3ECB16F68C45FFEC4867C31B2ACB761" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="23DC03030A6C5254360CD8D36A1B52B3" pageId="0" pageNumber="1">
<emphasis id="5C7C9A2812373ED53C2B7D4A73B55748" bold="true" pageId="0" pageNumber="1">
<emphasis id="4B7B5309DCA428FED2A6CC26C41E56C5" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Madagascar. •1 major worker; Toamasina;
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken id="52DF1ED5BFCF35323E3A1A31D9BD9103" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="F6AC8B2FF3C7EF79E6329963FEC689DE" originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato River;
<geoCoordinate degrees="16.80561" direction="south" orientation="latitude" precision="1" value="-16.80561">-16.80561</geoCoordinate>
<geoCoordinate id="394064CD6C8149A6099C8468F341C850" degrees="16.80561" direction="south" orientation="latitude" precision="1" value="-16.80561">-16.80561</geoCoordinate>
,
<geoCoordinate degrees="49.29507" direction="east" orientation="longitude" precision="1" value="49.29507">49.29507</geoCoordinate>
<geoCoordinate id="4C68E98506D4FDC1CC10BAD4B415F55B" degrees="49.29507" direction="east" orientation="longitude" precision="1" value="49.29507">49.29507</geoCoordinate>
; alt. 480 m; 27 Feb 2010; B.L. Fisher et al. leg.; BLF25039, CASENT0162231 (CASC).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
<emphasis id="32CD660FA7F21109E1688D5F54B54A5D" bold="true" pageId="0" pageNumber="1">
<emphasis id="452C74304A4902BBEE6E8938BE0824F1" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
.
</emphasis>
Madagascar. •2w., 1q.; same data as for holotype; CASENT0162230, CASENT0923223, CASENT0872222 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="1BE4647D6F69C5FBDC88F491623A0CCF" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="8754A8CB5B0524B142FE928DCE695318" pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph id="059D91A5C0D663476BE57119FE5844AC" pageId="0" pageNumber="1">
Madagascar. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
<emphasis id="EA3D5CD994C48391CB13BDAB73814E86" bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
: •3w., 1s.; 6.5 km SSW Befingotra,
<normalizedToken originalValue="Rés">Res</normalizedToken>
<normalizedToken id="971B433D56B598C955E65027E8301007" originalValue="Rés">Res</normalizedToken>
. Anjanaharibe-Sud; -14.75, 49.5; alt. 875 m; 19 Oct 1994; B.L. Fisher leg.; CASENT0198389, CASENT0198390 (CASC). •7w., 4s., 1q.; 9.2 km WSW Befingotra,
<normalizedToken originalValue="Rés">Res</normalizedToken>
<normalizedToken id="6B51EE98236854FCB2385C6B2A21D9E5" originalValue="Rés">Res</normalizedToken>
. Anjanaharibe-Sud; -14.75, 49.46667; alt. 1200 m; 8 Nov 1994; B.L. Fisher leg.; CASENT0198391, CASENT0198393, CASENT0198394, CASENT0235113, CASENT0235114 (CASC). -
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
<emphasis id="36D3A5E76D831BE52CB5EE1B1AB96CAF" bold="true" italics="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
: •1w.; Parc National de Ranomafana, Vatoharanana River, 4.1 km 231°SW Ranomafana; -21.29, 47.43333; alt. 1100 m; 27 Mar 2003; B.L. Fisher et al. leg.; CASENT0497619 (CASC). -
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mahajanga</emphasis>
<emphasis id="79ACEF4D54150C3E139BEF087B98F12D" bold="true" italics="true" pageId="0" pageNumber="1">Mahajanga</emphasis>
: •3w., 2s.;
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken id="7D300A5D4070ED92ADFEA2819F1AAFB3" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="251988074DB79B6325F61466775B350C" originalValue="Spéciale">Speciale</normalizedToken>
Marotandrano, Marotandrano 48.3 km S Mandritsara;
<geoCoordinate degrees="16.28322" direction="south" orientation="latitude" precision="1" value="-16.28322">-16.28322</geoCoordinate>
<geoCoordinate id="D3609B27C9BD465185B72A4152886B7B" degrees="16.28322" direction="south" orientation="latitude" precision="1" value="-16.28322">-16.28322</geoCoordinate>
,
<geoCoordinate degrees="48.81443" direction="east" orientation="longitude" precision="1" value="48.81443">48.81443</geoCoordinate>
<geoCoordinate id="EF097638061F20CCBDB42DF6E9035D24" degrees="48.81443" direction="east" orientation="longitude" precision="1" value="48.81443">48.81443</geoCoordinate>
; alt. 865 m; 7 Dec 2007; B.L. Fisher et al. leg.; CASENT0134242, CASENT0134279, CASENT0235121 (CASC). -
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
<emphasis id="43685CE54CE7ED141F607EE48A576A33" bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
: •1w.; Ambanizana, Parc National Masoala;
<geoCoordinate degrees="15.57222" direction="south" orientation="latitude" precision="1" value="-15.57222">-15.57222</geoCoordinate>
<geoCoordinate id="7DF4F29AB26A5D66A30D5DFCEFC4BA07" degrees="15.57222" direction="south" orientation="latitude" precision="1" value="-15.57222">-15.57222</geoCoordinate>
,
<geoCoordinate degrees="50.00694" direction="east" orientation="longitude" precision="1" value="50.00694">50.00694</geoCoordinate>
<geoCoordinate id="CCD272FEFA3D77DD6DC3E076C9EFD144" degrees="50.00694" direction="east" orientation="longitude" precision="1" value="50.00694">50.00694</geoCoordinate>
; alt. 930-1110 m; 2 Mar 2003; Andriamalala et al. leg.; CASENT0073482 (CASC). •1w.; F.C. Andriantantely;
<geoCoordinate degrees="18.695" direction="south" orientation="latitude" precision="55" value="-18.695">-18.695</geoCoordinate>
<geoCoordinate id="26612C1046214A86020ED74888BECDB8" degrees="18.695" direction="south" orientation="latitude" precision="55" value="-18.695">-18.695</geoCoordinate>
,
<geoCoordinate degrees="48.81333" direction="east" orientation="longitude" precision="1" value="48.81333">48.81333</geoCoordinate>
<geoCoordinate id="F12D685F6F01A399D71AF39DF09359F2" degrees="48.81333" direction="east" orientation="longitude" precision="1" value="48.81333">48.81333</geoCoordinate>
; alt. 530 m; 4 Dec 1998; Ratsirarson leg.; CASENT0198237 (CASC). •19w., 1s.; Montagne
<normalizedToken originalValue="dAkirindro">d'Akirindro</normalizedToken>
<normalizedToken id="55A77DBBB9DC2BB1B92DE35A9AF3490C" originalValue="dAkirindro">d'Akirindro</normalizedToken>
7.6 km 341°NNW Ambinanitelo;
<geoCoordinate degrees="15.28833" direction="south" orientation="latitude" precision="1" value="-15.28833">-15.28833</geoCoordinate>
<geoCoordinate id="C1A50111E33809F11F142013C1F412B9" degrees="15.28833" direction="south" orientation="latitude" precision="1" value="-15.28833">-15.28833</geoCoordinate>
,
<geoCoordinate degrees="49.54833" direction="east" orientation="longitude" precision="1" value="49.54833">49.54833</geoCoordinate>
<geoCoordinate id="4769FEE79D6BABBE7259C194C6AFB03C" degrees="49.54833" direction="east" orientation="longitude" precision="1" value="49.54833">49.54833</geoCoordinate>
; alt. 600 m; 17 Mar 2003; B.L. Fisher et al. leg.; CASENT0039054, CASENT0039064, CASENT0039103, CASENT0039188, CASENT0039200, CASENT0039209, CASENT0039219, CASENT0235138, CASENT0496323, CASENT0496345, CASENT0496346, CASENT0496347 (CASC). •15w., 4s.; Montagne
<normalizedToken originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
<normalizedToken id="B7761BA99ABD55450D1A201F2E05234C" originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
, 18.0 km 21°NNE Ambinanitelo;
<geoCoordinate degrees="15.18833" direction="south" orientation="latitude" precision="1" value="-15.18833">-15.18833</geoCoordinate>
<geoCoordinate id="44D89A28EF7686DECC18463329A9F735" degrees="15.18833" direction="south" orientation="latitude" precision="1" value="-15.18833">-15.18833</geoCoordinate>
,
<geoCoordinate degrees="49.615" direction="east" orientation="longitude" precision="55" value="49.615">49.615</geoCoordinate>
<geoCoordinate id="D0313B664E2B57DF7ABC98862E1794B3" degrees="49.615" direction="east" orientation="longitude" precision="55" value="49.615">49.615</geoCoordinate>
; alt. 470 m; 8 Mar 2003; B.L. Fisher et al. leg.; CASENT0037623, CASENT0037626, CASENT0048951, CASENT0495382, CASENT0495383, CASENT0495384, CASENT0495480, CASENT0495481, CASENT0495482 (CASC). •1w.;
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken id="C54A256C4EC52F9D60D0CD68B500056F" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="111C61933EF043634F078F398CD3F1FD" originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato River;
<geoCoordinate degrees="16.7633" direction="south" orientation="latitude" precision="5" value="-16.7633">-16.7633</geoCoordinate>
<geoCoordinate id="F5E5B2447CADC1DDC2EB954F7127004B" degrees="16.7633" direction="south" orientation="latitude" precision="5" value="-16.7633">-16.7633</geoCoordinate>
,
<geoCoordinate degrees="49.26692" direction="east" orientation="longitude" precision="1" value="49.26692">49.26692</geoCoordinate>
<geoCoordinate id="73D392A7B0946EB457F5D0827F5E61E3" degrees="49.26692" direction="east" orientation="longitude" precision="1" value="49.26692">49.26692</geoCoordinate>
; alt. 520 m; 22 Feb 2010; B.L. Fisher et al. leg.; CASENT0160480 (CASC). •2w.;
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken id="9AF4CCBC7397989A22DDF149294D5ABA" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="AA8CE0F510757C51EF98B5B1D24921CB" originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato River;
<geoCoordinate degrees="16.7702" direction="south" orientation="latitude" precision="5" value="-16.7702">-16.7702</geoCoordinate>
<geoCoordinate id="AD998C038A3D9C5178F4CDEEAD1F4157" degrees="16.7702" direction="south" orientation="latitude" precision="5" value="-16.7702">-16.7702</geoCoordinate>
,
<geoCoordinate degrees="49.26638" direction="east" orientation="longitude" precision="1" value="49.26638">49.26638</geoCoordinate>
<geoCoordinate id="2A42999D06E6A8FD760B510BD02D77A8" degrees="49.26638" direction="east" orientation="longitude" precision="1" value="49.26638">49.26638</geoCoordinate>
; alt. 470 m; 23 Feb 2010; B.L. Fisher et al. leg.; CASENT0161455 (CASC). •1w.;
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken id="ED1EED3A367874C2AFC4BC7166449A42" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="B0E54C4D3C4F00260C6CDB8DFF826DF4" originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato River;
<geoCoordinate degrees="16.8162" direction="south" orientation="latitude" precision="5" value="-16.8162">-16.8162</geoCoordinate>
<geoCoordinate id="E3E11E01E474B6009D728EC8AE277288" degrees="16.8162" direction="south" orientation="latitude" precision="5" value="-16.8162">-16.8162</geoCoordinate>
,
<geoCoordinate degrees="49.29202" direction="east" orientation="longitude" precision="1" value="49.29202">49.29202</geoCoordinate>
<geoCoordinate id="71E85703490A64F81442DB57B2325C51" degrees="49.29202" direction="east" orientation="longitude" precision="1" value="49.29202">49.29202</geoCoordinate>
; alt. 425 m; 25 Feb 2010; B.L. Fisher et al. leg.; CASENT0161854 (CASC). •2w.;
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken id="F82D4544A8C747A911606EDE36EA6FA1" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="C5F78383DC6DE549FFB17DC6262669E1" originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato River;
<geoCoordinate degrees="16.81745" direction="south" orientation="latitude" precision="1" value="-16.81745">-16.81745</geoCoordinate>
<geoCoordinate id="9F5C1B07E2D509B3EB40E53FC26923D7" degrees="16.81745" direction="south" orientation="latitude" precision="1" value="-16.81745">-16.81745</geoCoordinate>
,
<geoCoordinate degrees="49.2925" direction="east" orientation="longitude" precision="5" value="49.2925">49.2925</geoCoordinate>
<geoCoordinate id="4E2B7B6B1EA76772D3FDAE4DDBC0B098" degrees="49.2925" direction="east" orientation="longitude" precision="5" value="49.2925">49.2925</geoCoordinate>
; alt. 400 m; 26 Feb 2010; B.L. Fisher et al. leg.; CASENT0162160 (CASC). •4w., 2s., 1q.;
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken id="768F5EFFE1FD388CA3E2147438C7EC57" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="2E9EA1026F9E8B4E1175D55E5275F212" originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato River;
<geoCoordinate degrees="16.77274" direction="south" orientation="latitude" precision="1" value="-16.77274">-16.77274</geoCoordinate>
<geoCoordinate id="A12EBB41864635EB371F3A16716CC93F" degrees="16.77274" direction="south" orientation="latitude" precision="1" value="-16.77274">-16.77274</geoCoordinate>
,
<geoCoordinate degrees="49.26551" direction="east" orientation="longitude" precision="1" value="49.26551">49.26551</geoCoordinate>
<geoCoordinate id="46C804CCD19E376210AB592F6B0A5371" degrees="49.26551" direction="east" orientation="longitude" precision="1" value="49.26551">49.26551</geoCoordinate>
; alt. 450 m; 20 Feb 2010; B.L. Fisher et al. leg.; CASENT0162675, CASENT0162686, CASENT0162688, CASENT0162689 (CASC). •4w., 4s., 3q.;
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken id="6B910CCABCEF751E69CAD715CE63FD3C" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="885EB6C0746580E7608299B2628AED5A" originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato River;
<geoCoordinate degrees="16.80561" direction="south" orientation="latitude" precision="1" value="-16.80561">-16.80561</geoCoordinate>
<geoCoordinate id="660D6B59B81B9EAFE3880F5203049D5F" degrees="16.80561" direction="south" orientation="latitude" precision="1" value="-16.80561">-16.80561</geoCoordinate>
,
<geoCoordinate degrees="49.29507" direction="east" orientation="longitude" precision="1" value="49.29507">49.29507</geoCoordinate>
<geoCoordinate id="20E5127619EFE11D49B9D1FCB50552A7" degrees="49.29507" direction="east" orientation="longitude" precision="1" value="49.29507">49.29507</geoCoordinate>
; alt. 480 m; 27 Feb 2010; B.L. Fisher et al. leg.; CASENT0163009, CASENT0163019, CASENT0163021, CASENT0163027, CASENT0163032, CASENT0163040, CASENT0163073 (CASC). •2w.;
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken id="9A02201DB07B15DE83DA3E66FF7866CF" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="5F42203BA7B809EBEBAD32FB5BAD1B72" originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato River;
<geoCoordinate degrees="16.7633" direction="south" orientation="latitude" precision="5" value="-16.7633">-16.7633</geoCoordinate>
<geoCoordinate id="D5A1C690AB0BD56322F4445AE267975C" degrees="16.7633" direction="south" orientation="latitude" precision="5" value="-16.7633">-16.7633</geoCoordinate>
,
<geoCoordinate degrees="49.26692" direction="east" orientation="longitude" precision="1" value="49.26692">49.26692</geoCoordinate>
<geoCoordinate id="4AEEEE88E3FC2E2327842481D0F18557" degrees="49.26692" direction="east" orientation="longitude" precision="1" value="49.26692">49.26692</geoCoordinate>
; alt. 520 m; 22 Feb 2010; B.L. Fisher et al. leg.; CASENT0163856, CASENT0163945 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
<subSubSection id="23C67CEDD78330CEDC97B3D91AF4BC3A" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="BA6675D90A5026970490B291A88BEAD4" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="524DDE41E71D7B56BEF8E83B2416249A" pageId="0" pageNumber="1">
<emphasis id="782CABCFE36462EC97B9134F0A3D3C3D" bold="true" pageId="0" pageNumber="1">
<emphasis id="C920D11442E111902FD4BC1FFA5605C5" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
Large species: HL: 2.38-2.71 (2.55), HW: 2.17-2.54 (2.4), WL: 1.6-1.95 (1.76); propodeal spines very long (PSL: 0.48-0.59 (0.54)); head in full-face view trapezoid, widened posteriorly; sides of the head with sparse, relatively long, erect pilosity; frons with longitudinal, sparse, and directed outward rugae, interspaces with dense and fine rugulae, sculpture weakening posteriorly; first gastral tergite shagreened, at least on its basal part; body dark brown to black; inner hypostomal teeth distinct, triangular and thick, with rounded apex, closely spaced; outer hypostomal teeth distinct, low, lobe-like, with wide base and tops directed outward, smaller and thinner than inner hypostomal teeth.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
<emphasis id="2214DF7AFB0077601705F4A9BCF2A129" bold="true" pageId="0" pageNumber="1">
<emphasis id="2704D541811397B17E5CE59A75BEF0FF" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
Large species: HL: 0.85-1.03 (0.93), HW: 0.8-1.0 (0.89), WL: 1.21-1.47 (1.36); propodeal spines very long (PSL: 0.34-0.44 (0.39)); scape, when laid back, surpassing posterior head margin by one-third of its length; head sculpture rugoreticulate; mesosoma with dense rugoreticulation, sometimes dorsum with weaker sculpture, but never smooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
<subSubSection id="75763D74E3E02D2BBA5682E58966C3E9" pageId="0" pageNumber="1" type="description">
<paragraph id="2DB6DB034384B0DB271DC9D3568AB099" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="9EB6536174AFA7C9E5DC9984FF0D0576" pageId="0" pageNumber="1">
<emphasis id="C88BB0645EB54FB316658E392524765A" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
<emphasis id="19F21C386DFF6100A49346E9AE0B63DE" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 10): HL: 2.38-2.71 (2.55); HW: 2.17-2.54 (2.4); SL: 0.95-1.17 (1.08); EL: 0.24-0.3 (0.27); WL: 1.6-1.95 (1.76); PSL: 0.48-0.59 (0.54); MTL: 1.16-1.28 (1.23); PNW: 0.8-0.95 (0.9); PTW: 0.24-0.31 (0.28); PPW: 0.7-0.92 (0.83); CI: 91.3-98.5 (94.0); SI: 43.6-46.3 (45.2); PSLI: 19.8-22.6 (21.2); PPI: 31.0-36.0 (34.1); PNI: 35.5-38.8 (37.4); MTI: 49.5-53.8 (51.2).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
<emphasis id="455DEDA546D7A1117D1D23EC5247B136" bold="true" pageId="0" pageNumber="1">
<emphasis id="4A67F604BE986D7601C8DCC601C51E8C" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
In full-face view trapezoid, widened posteriorly (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27B</figureCitation>
<figureCitation id="D8B4059403B53DC0E1324260E84F075F" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27B</figureCitation>
). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with sparse, relatively long, erect pilosity; whole head with moderately sparse, long, erect setae. Antennal scrobes absent. Occipital lobes shiny, and smooth at posterior part, anterior part with fine and dense rugulae; genae shiny, with fine and dense rugulae, sometimes posterior part smooth or with reduced sculpture; malar area and frons with longitudinal, sparse, and directed outward rugae, interspaces with dense and fine rugulae; lateral sides of head shiny, with dense and fine rugulae and thin, indistinct, longitudinal rugae. Centre of clypeus smooth and shiny, lateral sides with longitudinal rugae; median notch present, shallow, and wide; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, reaching mid-length of head; pilosity suberect to erect (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27B, D</figureCitation>
<figureCitation id="01A1C615C73456DE0A9F22302D20355D" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27B, D</figureCitation>
). Inner hypostomal teeth distinct, triangular, and thick, with rounded apex, closely spaced; outer hypostomal teeth distinct, low, lobe-like, with wide base and tops directed outward, smaller and thinner than inner hypostomal teeth (Fig.
<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85C</figureCitation>
<figureCitation id="C63A5ED92C7C9055528086285E9B31A7" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
<emphasis id="6F6FD0AA3E9C18B680A07C68D95A1F91" bold="true" pageId="0" pageNumber="1">
<emphasis id="C66E9B3CAC6C9C1AED3929046025B538" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum short, low, and evenly convex, with relatively steep posterior declivity; promesonotal groove absent; metanotal groove absent; propodeal spines very long, massive basally, with acute apex; humeral area laterally weakly produced (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D</figureCitation>
<figureCitation id="339A090C97280C9C86462365888F578B" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D</figureCitation>
). Surface shiny, with dense rugoreticulation, mesosomal dorsum and propodeum with weaker sculpture, but never smooth. Pilosity relatively dense, long, and erect (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
<figureCitation id="829A71D7516A920812E108169632B89D" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="8A5BDF1A06A4D337CE91B6FDDCDE9863" bold="true" pageId="0" pageNumber="1">
<emphasis id="4014C13C6578E0DD02B9ED52C82A7123" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shagreened; peduncle relatively long and thin; node triangular with rounded apex, in rear view node dorsoventrally slightly depressed; pilosity moderately sparse and erect (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
<figureCitation id="CCA0CF51919A86B118A1C9D73535EA51" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
<emphasis id="13BEBE9BD410AD3FBAC15122E197E589" bold="true" pageId="0" pageNumber="1">
<emphasis id="BD0A3A3FBAFFC3C7004E370050BDDAEA" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Shagreened; in dorsal view sides with acute, and short to moderately short angular projections; pilosity long, moderately sparse, and erect (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
<figureCitation id="36C566CA9C62DEF74A2E0938AF410309" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="5AB44873B945566D6CCB573FE9EC7E30" bold="true" pageId="0" pageNumber="1">
<emphasis id="55B1965191783040FE74999E647641B2" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
First gastral tergite shagreened, at least on its basal part; pilosity sparse, long and erect (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
<figureCitation id="32BFDFD851CC970826466B1B7A3B1578" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
<emphasis id="F2AD6D60A93EEC2413C31CEE396F5518" bold="true" pageId="0" pageNumber="1">
<emphasis id="69A2EF8ABBC0C5345B93CE8C97847920" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, dark brown to black (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
<figureCitation id="341AE4BE5B6B48FAD218F8CBB3DDA655" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27D, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1" start="Figure 27" startId="F27">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 27.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole maizina</emphasis>
<caption id="B861B8A8D1508BE63FC3D3816A9C9460" doi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1" start="Figure 27" startId="F27">
<paragraph id="F537971E2B53ED2C20D3CB729BF9A21F" pageId="0" pageNumber="1">
<emphasis id="C099EE5CE66024B6302A1FAC89CCF065" bold="true" pageId="0" pageNumber="1">Figure 27.</emphasis>
<taxonomicName id="B2B8726F036D2CB0756D8BF15B720B0E" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina">
<emphasis id="480D0EA8E78D627FD2807FE55582B7BA" italics="true" pageId="0" pageNumber="1">Pheidole maizina</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<emphasis id="C6DB4D44F784C7820F2FA22C07FDF552" bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<emphasis id="F5216DF83BE43A17A4DA3B8F0F8443E9" bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<emphasis id="3853414C88C81AECEA7841DA309A73D4" bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0923223) and full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<emphasis id="D4ABE1B6F4630A8CD6B6F1D7179E63AA" bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<emphasis id="41E9069B3A1B09CBB54269EC5412DECE" bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<emphasis id="4A02AB61FFD494A08264D09C77ED9822" bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0162231).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
<paragraph id="278D3618FA4B3EE1746744BC4B8287E8" pageId="0" pageNumber="1">
<emphasis id="212825B68B1935CF7349F3D578C40566" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
<emphasis id="B77CA2FA655DAEB41B67FA8CE6F0AE39" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 10): HL: 0.85-1.03 (0.93); HW: 0.8-1.0 (0.89); SL: 1.07-1.2 (1.16); EL: 0.16-0.21 (0.18); WL: 1.21-1.47 (1.36); PSL: 0.34-0.44 (0.39); MTL: 0.91-1.1 (1.01); PNW: 0.55-0.66 (0.61); PTW: 0.1-0.16 (0.13); PPW: 0.19-0.28 (0.23); CI: 93.3-101.0 (95.5); SI: 122.0-135.6 (130.7); PSLI: 40.0-44.9 (42.3); PPI: 50.5-60.2 (55.3); PNI: 66.3-70.4 (68.7); MTI: 111.2-117.0 (114.1).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
<emphasis id="DBA9CA73C01158030C08A20C5DCFCE58" bold="true" pageId="0" pageNumber="1">
<emphasis id="CC2D29499F7A57BEAF1524CBEE1C853A" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Occipital margin convex; occipital carina narrow, weakly developed (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27A</figureCitation>
<figureCitation id="2694B1B5C95C3F1E5B4E64220AD855A8" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27A</figureCitation>
). Pilosity sparse, long, and erect. Sculpture variable, finely to strongly rugoreticulate on the whole surface; only centre of frons sometimes with sculpture reduced or smooth; antennal sockets with sparse carinae curved outward. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae present. Scape, when laid back, surpassing posterior head margin by one-third of its length; pilosity suberect to erect (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27A, C</figureCitation>
<figureCitation id="74905F5509C1A0A5AECBDF9335722715" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27A, C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
<emphasis id="FB35DEC58882A32A43A9CDBE9D2F00AA" bold="true" pageId="0" pageNumber="1">
<emphasis id="DB87CF2011FC018E6D1D6E521824EF41" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum low, long, and slightly convex; promesonotal groove absent; metanotal groove very indistinct; propodeal spines very long, massive basally, with acute apex (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C</figureCitation>
<figureCitation id="A3909F5F5F7764117F5D8B7D6AB29901" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C</figureCitation>
). Surface shiny, with dense rugoreticulation, sometimes dorsum with weaker sculpture, but never smooth. Pilosity sparse, long, and erect (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
<figureCitation id="47A79CB89E41E5884111A8B3E863FA73" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="1BA98266AE8BE3ED89418BC8177CAA17" bold="true" pageId="0" pageNumber="1">
<emphasis id="CC3FBE8BD56DF0FEB8E4A4AF26C2C8EC" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Finely shagreened; peduncle long and thin; node triangular with rounded apex; with few long, erect setae (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
<figureCitation id="8886AB0FE7BCD4ECB39E5766B3357772" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
<emphasis id="849E2CAB45871A4B374152D891BC400C" bold="true" pageId="0" pageNumber="1">
<emphasis id="BDFE8584586987C8F614EEC00F0D68D5" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Moderately shagreened; moderately short, low, and convex; with few long, erect setae at the anterior edge (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
<figureCitation id="729EAC09A9084157A40F01D95E558845" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="0C4121F40A6C07D43B7EC0DDBD50093D" bold="true" pageId="0" pageNumber="1">
<emphasis id="59791AA1CC06335319B8FE292AEC845C" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
With few long, erect setae (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
<figureCitation id="CE33BB1C5D96919938E291076BA6F5AB" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
<emphasis id="2DDC619A5243477BB7F4C2ED7D2ADF02" bold="true" pageId="0" pageNumber="1">
<emphasis id="717B2D3F3A94310D165E603488FA58C4" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, dark brown to black (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
<figureCitation id="F43E725E5F2101FA051E7119DC16EBBD" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Pheidole maizina sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923223) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0162231)." figureDoi="10.3897/zookeys.905.39592.figure27" httpUri="https://binary.pensoft.net/fig/373074" pageId="0" pageNumber="1">27C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Malagasy for dark, in reference to dark body colouration.</paragraph>
<subSubSection id="F162629C002D9458731AAF1367F7D04B" pageId="0" pageNumber="1" type="etymology">
<paragraph id="267CF3A2607669DA870881B8D74AD826" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="FD71FA498BF260328D44ADD74F3BD962" pageId="0" pageNumber="1">Malagasy for dark, in reference to dark body colouration.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was collected at between 400-1200 m in elevation, in rainforest, montane rainforest, and transitional humid forest. Nests were located in in rotten logs and stumps, and in rotten sticks on the ground.</paragraph>
<subSubSection id="F9BE83647D4CFF087C55E8718C53EA7F" pageId="0" pageNumber="1" type="biology_ecology">
<paragraph id="44D1AE93FF4892D197082EB6281F20AE" pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph id="1D96B2D969797D70DF571B3F7B31CA97" pageId="0" pageNumber="1">The species was collected at between 400-1200 m in elevation, in rainforest, montane rainforest, and transitional humid forest. Nests were located in in rotten logs and stumps, and in rotten sticks on the ground.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="9F753FC221D8C26F81917015BD2D5344" pageId="0" pageNumber="1" type="comments">
<paragraph id="CD6D215A9F2CDA2B1B6FB456BAF71205" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="0E02E85B8266445EC70C136EB9304D6D" pageId="0" pageNumber="1">
This species is most similar to
<taxonomicName lsidName="P. scabrata" pageId="0" pageNumber="1" rank="species" species="scabrata">
<emphasis italics="true" pageId="0" pageNumber="1">P. scabrata</emphasis>
<taxonomicName id="C97F9FD5B70889BAFCB0D5E9D7F8C443" lsidName="P. scabrata" pageId="0" pageNumber="1" rank="species" species="scabrata">
<emphasis id="4D2C5B333F4927BF1DF4A348DF97A0B9" italics="true" pageId="0" pageNumber="1">P. scabrata</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
<emphasis id="D97EFFC89EA6C5C72367E4E99A30F507" bold="true" pageId="0" pageNumber="1">
<emphasis id="F9C06758BE9FF91280C9BEC809DDB5BF" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole maizina</emphasis>
<taxonomicName id="9854DDA62FDB844307596311EB148897" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina">
<emphasis id="8C3654F6B96EBBD618183469A06213D3" italics="true" pageId="0" pageNumber="1">Pheidole maizina</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName lsidName="P. scabrata" pageId="0" pageNumber="1" rank="species" species="scabrata">
<emphasis italics="true" pageId="0" pageNumber="1">P. scabrata</emphasis>
<taxonomicName id="0C4F74C2D792F8A1B37A75D9EC51CC79" lsidName="P. scabrata" pageId="0" pageNumber="1" rank="species" species="scabrata">
<emphasis id="EBBF4068964C058CC42CA512F03DFE1D" italics="true" pageId="0" pageNumber="1">P. scabrata</emphasis>
</taxonomicName>
by having distinct rugulae on at least half of the genae, tips of outer hypostomal teeth directed outward, never smooth katepisternum, densely shagreened surface of first gastral tergite, and darker body colouration.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
<emphasis id="48F7D036A11A35ADC20FB84FA22A5E65" bold="true" pageId="0" pageNumber="1">
<emphasis id="52E64333C3CA389468C821025D51775B" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole maizina</emphasis>
<taxonomicName id="F9F8051C3E64E14E26236CB6C9F5EEAB" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole maizina" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="maizina">
<emphasis id="F68BB74BA6823723FF8FA70072DDE5E2" italics="true" pageId="0" pageNumber="1">Pheidole maizina</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName lsidName="P. scabrata" pageId="0" pageNumber="1" rank="species" species="scabrata">
<emphasis italics="true" pageId="0" pageNumber="1">P. scabrata</emphasis>
<taxonomicName id="AC5A3A91EC3BA70B92C48E3423927B31" lsidName="P. scabrata" pageId="0" pageNumber="1" rank="species" species="scabrata">
<emphasis id="E32573B63B414824CEBA7AB2BF797EAC" italics="true" pageId="0" pageNumber="1">P. scabrata</emphasis>
</taxonomicName>
by rugoreticulate head sculpture with never smooth genae.
</paragraph>