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<mods:title id="D76135575137B8BF3094ACFBF0067C9D">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<treatment ID-GBIF-Taxon="161777035" LSID="urn:lsid:zoobank.org:act:88804AF2-4F5F-4A67-B2BD-9012E4F43AE9" httpUri="http://treatment.plazi.org/id/D3723344842638A6D42382F7CD46EBAD" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/88804AF2-4F5F-4A67-B2BD-9012E4F43AE9" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole longipilosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longipilosa" status="sp. nov.">Pheidole longipilosa</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">Figs 39A-F</figureCitation>
|
||||
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84X</figureCitation>
|
||||
<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87B</figureCitation>
|
||||
<treatment id="D3723344842638A6D42382F7CD46EBAD" ID-GBIF-Taxon="161777035" LSID="urn:lsid:zoobank.org:act:88804AF2-4F5F-4A67-B2BD-9012E4F43AE9" httpUri="http://treatment.plazi.org/id/D3723344842638A6D42382F7CD46EBAD" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="E41A6D3D0B86E5B22F5B407E97DE13B1" pageId="0" pageNumber="1" type="nomenclature">
|
||||
<paragraph id="8011F53769B6494175ED5691B9FD137E" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="3FB735F9E41575EDF41F55BDF2837DA5" ID-CoL="CMLF3" LSID="http://zoobank.org/88804AF2-4F5F-4A67-B2BD-9012E4F43AE9" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole longipilosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longipilosa" status="sp. nov.">Pheidole longipilosa</taxonomicName>
|
||||
<taxonomicNameLabel id="8794B79F6CAAED517314568111EB563D" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
|
||||
<figureCitation id="99B90BD15A19A65242B23A64AD8A0EEE" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">Figs 39A-F</figureCitation>
|
||||
<figureCitation id="7AFBEEFD3333C8255312770692901907" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84X</figureCitation>
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||||
<figureCitation id="A986DA22B56B026147EE87BB86690E87" captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87B</figureCitation>
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||||
</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="type material">
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||||
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||||
<subSubSection id="E517DC104EE07A05EB9283F2A24B4B27" pageId="0" pageNumber="1" type="type material">
|
||||
<paragraph id="56E05ED43A71F5EF802063532A91E0FC" pageId="0" pageNumber="1">Type material.</paragraph>
|
||||
<paragraph id="BCED85C5ECB7E301751C72A7F02F9145" pageId="0" pageNumber="1">
|
||||
<emphasis id="05998A580DCEBB24E4DE585714398E30" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="07E35931160CC7B441EB568E43C5CBB1" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Toliara;
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken originalValue="Classée">Classee</normalizedToken>
|
||||
<normalizedToken originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||||
<normalizedToken id="124BAEF859FA933E0664250ECABB410A" originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="F2D31A9AF8C88A8B06559EED0E30D5C0" originalValue="Classée">Classee</normalizedToken>
|
||||
<normalizedToken id="B86F541B837EF53BCEE8837B4998DF87" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||||
, 29.2 km 343°NNW Mahaboboka; -22.675, 44.19; alt. 1100 m; 18 Feb 2003; B.L. Fisher et al. leg.; CASENT0498298 (CASC).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
|
||||
<emphasis id="B570484D9CB669F2DBF1B5AC19C5BBF6" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="48557DE95FF79BCBD6F9441F6C161E2D" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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Madagascar. •9w.; same data as for holotype; CASENT0235043, CASENT0498295-CASENT0498297, CASENT0872154-CASENT0872158 (CASC).
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</paragraph>
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</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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||||
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="038F3C1AC7A9E400F3E873F22374824B" pageId="0" pageNumber="1" type="materials_examined">
|
||||
<paragraph id="F9F9868ADBAB824FA5E8472CE8D30A81" pageId="0" pageNumber="1">Other material.</paragraph>
|
||||
<paragraph id="24F89F3AA10013736A5E6A3899A68FD6" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
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||||
<emphasis id="F2EB4D97C3CC298398E6FF7E3A61406E" bold="true" italics="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
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: •1w.; Parc National
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<normalizedToken originalValue="d’Isalo">d'Isalo</normalizedToken>
|
||||
<normalizedToken id="3828BA7865AA652E18EED8F35A2B4828" originalValue="d’Isalo">d'Isalo</normalizedToken>
|
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, Sahanafa River, 29.2 km 351°N Ranohira;
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<geoCoordinate degrees="22.31333" direction="south" orientation="latitude" precision="1" value="-22.31333">-22.31333</geoCoordinate>
|
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<geoCoordinate id="D3475C88E4B4043A5113214D743E3FDA" degrees="22.31333" direction="south" orientation="latitude" precision="1" value="-22.31333">-22.31333</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="45.29167" direction="east" orientation="longitude" precision="1" value="45.29167">45.29167</geoCoordinate>
|
||||
<geoCoordinate id="2CE86AD7BEFC0D8335033F8FC64870A1" degrees="45.29167" direction="east" orientation="longitude" precision="1" value="45.29167">45.29167</geoCoordinate>
|
||||
; alt. 500 m; 10 Feb 2003; Fisher et al. leg.; CASENT0031767 (CASC). -
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toliara</emphasis>
|
||||
<emphasis id="2DC83900DC9DEF3E589F4EADD83EE104" bold="true" italics="true" pageId="0" pageNumber="1">Toliara</emphasis>
|
||||
: •7w., 4s., 3q.;
|
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken originalValue="Classée">Classee</normalizedToken>
|
||||
<normalizedToken originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||||
<normalizedToken id="4210075314B9B5053EB48D88660F2473" originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="DC74E87AF23D18B0C9D61FFC192E8ECA" originalValue="Classée">Classee</normalizedToken>
|
||||
<normalizedToken id="046257DE4580BCC4EACEE9EEEF344D07" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||||
, 29.2 km 343°NNW Mahaboboka; -22.675, 44.19; alt. 1100 m; 18 Feb 2003; Fisher et al. leg.; CASENT0031011, CASENT0031099, CASENT0049501, CASENT0073507, CASENT0235043, CASENT0496936, CASENT0496937, CASENT0496938 (CASC). •1w., 5s.;
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken originalValue="Classée">Classee</normalizedToken>
|
||||
<normalizedToken originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||||
<normalizedToken id="016140602B96821EB331D6F9F9EBECE8" originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="8253A12D5F42D88757BCEAE279768B49" originalValue="Classée">Classee</normalizedToken>
|
||||
<normalizedToken id="A9F82C85C45C985573BE385F0A38EBD8" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||||
, 29.4 km 343°NNW Mahaboboka;
|
||||
<geoCoordinate degrees="22.675" direction="south" orientation="latitude" precision="55" value="-22.675">-22.675</geoCoordinate>
|
||||
<geoCoordinate id="CD1DCF1F32D4440329B8A75649B7FB94" degrees="22.675" direction="south" orientation="latitude" precision="55" value="-22.675">-22.675</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="44.18667" direction="east" orientation="longitude" precision="1" value="44.18667">44.18667</geoCoordinate>
|
||||
<geoCoordinate id="00963321A539EC87A6A0CAD512EADCFC" degrees="44.18667" direction="east" orientation="longitude" precision="1" value="44.18667">44.18667</geoCoordinate>
|
||||
; alt. 1050 m; 21 Feb 2003; Fisher et al. leg.; CASENT0023935, CASENT0023938, CASENT0023943, CASENT0023946, CASENT0023947, CASENT0024032 (CASC).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||||
<subSubSection id="3F7C83100AB2BC87071257797B11A0DB" pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph id="E3B2F4B0663867F9DCD4AD2D59FD736F" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph id="02EC153C6674B057B2A71D31FC2FCBDA" pageId="0" pageNumber="1">
|
||||
<emphasis id="AB61AFFB9982F2CFA8D319E637F304E6" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="33A200022A5CE53511E6FDE16CBC12C7" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Head in full-face view sub-oval; sides of the head with dense, long, erect pilosity; occipital lobes shiny, with thick, irregular rugae, interspaces with distinct, irregular rugulae not fading posteriorly; inner hypostomal teeth distinct, high, closely spaced, triangular, with rounded apex directed inward, and wide base; outer hypostomal teeth taller and wider than inner hypostomal teeth, lobe-like, directed outward; gaster smooth.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||||
<emphasis id="0000A83E7B20D9A13FFA3EEBB3123054" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="B9B88286455B05723D3219E272823BE0" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Head shiny, foveolate, with additional short, longitudinal, thick rugae on frons, genae with fading sculpture and smooth notch; scape, when laid back, surpassing the posterior head margin by one-fifth of its length; promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum; mesosoma foveolate, katepisternum smooth; propodeal spines short, triangular.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||||
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||||
<subSubSection id="62049B4575DB927328C07DD24713FC0C" pageId="0" pageNumber="1" type="description">
|
||||
<paragraph id="1B46FABD83999E1A25D1BBC80DFB7626" pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph id="BB085002939D10244622E830679E58F6" pageId="0" pageNumber="1">
|
||||
<emphasis id="377D2D1656025EF99E1EE9BCFB174011" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||||
Measurements (
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
<emphasis id="CBAA46D82A84ED9FBCC2E92C4DC33D0A" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
= 10): HL: 1.05-1.11 (1.09); HW: 0.98-1.04 (1.0); SL: 0.46-0.52 (0.48); EL: 0.13-0.16 (0.13); WL: 0.84-0.97 (0.88); PSL: 0.14-0.17 (0.15); MTL: 0.47-0.5 (0.49); PNW: 0.52-0.58 (0.54); PTW: 0.12-0.14 (0.13); PPW: 0.35-0.4 (0.37); CI: 90.0-93.9 (91.7); SI: 46.5-51.8 (48.1); PSLI: 12.5-15.4 (13.7); PPI: 31.9-39.0 (34.9); PNI: 53.1-55.5 (54.3); MTI: 45.8 -50.9 (48.8).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
<emphasis id="3D8716A3C526AC3415BE347C0FA5A51C" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="09A44ED83DE5091F8877AF3A53919B26" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
In full-face view longer than wide, anterior of eyes slightly convex, posterior of eyes convex (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39B</figureCitation>
|
||||
<figureCitation id="5829CEAC308D0E7F12E3991D48662F1A" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39B</figureCitation>
|
||||
). In lateral view sub-oval; ventral and dorsal faces convex; dorsal face not depressed posteriorly; inner hypostomal teeth visible. Sides of the head with dense, very long, erect pilosity; whole head with moderately dense, long, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, with thick, longitudinal, and long rugae; interspaces distinctly rugulose. Occipital lobes shiny, with thick, irregular rugae, interspaces with distinct, irregular rugulae not fading posteriorly; frons with moderately dense, thick, longitudinal rugae, interspaces smooth, with indistinct rugulae weakening anteriorly; genae shiny, with dense and thin, irregular rugulae, central part with smooth notch; malar area with thin, longitudinal, dense rugae, interspaces with distinct rugulae. Centre of clypeus shiny and smooth, lateral sides with longitudinal rugulae; median notch present, narrow, and moderately deep; median longitudinal carina present, indistinct; lateral longitudinal carinae present. Scape, when laid back, reaching the midlength of head; pilosity erect (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39B, D</figureCitation>
|
||||
<figureCitation id="873B24F05421BC8BEB74CB17AEDE352F" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39B, D</figureCitation>
|
||||
). Inner hypostomal teeth distinct, high, closely spaced, triangular, with rounded apex directed inward, and wide base; outer hypostomal teeth taller and wider than inner hypostomal teeth, lobe-like, directed outward (Fig.
|
||||
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84X</figureCitation>
|
||||
<figureCitation id="C7283CBBBB634EDDB1818A7B75B52871" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84X</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
<emphasis id="BCDD8214C2711FE51FE1B8F29C238593" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="5F78A81E87904AF9B19D775BB8E90C15" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
In lateral view, promesonotum relatively high and arched, posterior mesonotum relatively steep, with tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines small to moderately long, triangular, with sharp apex and wide base; humeral area laterally weakly produced (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D</figureCitation>
|
||||
<figureCitation id="4E173D55AF8802C6F50748C94A893D14" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D</figureCitation>
|
||||
). Surface shiny, promesonotum foveolate with additional indistinct, sparse rugoreticulations, sometimes sculpture fading on dorsal surface and lower parts of lateral sides; katepisternum rugoreticulate, with smooth notch; anepisternum and lateral sides of propodeum with distinct rugoreticulation; dorsoventral side of propodeum smooth or with indistinct sculpture. Pilosity moderately dense, very long, and erect (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
|
||||
<figureCitation id="AB2565D82032CF0FDD8A624443464069" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="22BF66EC4E2089FC40F71AFD1A1511EA" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="22082A09B4770271D13C600E07A079E6" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Shiny; peduncle short, finely foveolate, with indistinct horizontal lobes on its basal part; node with fading sculpture, relatively low, triangular, with rounded apex, in rear view node slightly convex; pilosity long and erect (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
|
||||
<figureCitation id="6D6C814110679E2C4FB6A615FCF5DD3C" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
<emphasis id="343FC1C803DE33976B94121D4F99629F" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="782D3CDDD5559896B8F27ABD0CF843EA" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Shiny, finely shagreened, with dorsum at least partially smooth; in dorsal view sides with moderately short, acute, and triangular projections; pilosity long, long and erect (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
|
||||
<figureCitation id="1F663A741CE110C672B4715B658B6395" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="50855C5250E2DAE24535BAF2A62167AC" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="C794BB784EF89A5034597EDCC412ACDF" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Shiny and smooth, only basal part of first gastral tergite finely shagreened; pilosity dense, long, and erect (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
|
||||
<figureCitation id="ABAEA51C301F3F0ACB9FF96497695D84" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
<emphasis id="0D229780F10A5B612B33AF3D7C37D9A8" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="9C1C026DBA378C513125F60D3A9B6685" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Reddish brown to bright brown (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
|
||||
<figureCitation id="582E5A24DD3DB662F24525B2C342CCFE" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1" start="Figure 39" startId="F39">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 39.</emphasis>
|
||||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole longipilosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longipilosa">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole longipilosa</emphasis>
|
||||
<caption id="52E5D70CC32951F30F4E13CFD936A47A" doi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1" start="Figure 39" startId="F39">
|
||||
<paragraph id="F1453EFAEF545E51F24FECF9845927EF" pageId="0" pageNumber="1">
|
||||
<emphasis id="5AAFFAAD66612B1DE0836C495D435F9E" bold="true" pageId="0" pageNumber="1">Figure 39.</emphasis>
|
||||
<taxonomicName id="A73FC1556AB9557AFBCAE661D548D1C9" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole longipilosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longipilosa">
|
||||
<emphasis id="0B957F7DB174F3755D27D2F9B8E2DDB7" italics="true" pageId="0" pageNumber="1">Pheidole longipilosa</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., full-face view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<emphasis id="8F6282D5C253BDF106E185F6D2D7B883" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
), profile (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<emphasis id="0C0410F88821572B0D42846E998FE228" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
), and dorsal view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<emphasis id="7EEFD4ED0F4D7C78B3F164D274B85228" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
) of paratype minor worker (CASENT0235043) and full-face view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<emphasis id="D8718A7D1B8E246E180B552099C82B15" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
), profile (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<emphasis id="9FCFA945E0617032791C01AA23353C74" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
), and dorsal view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
<emphasis id="44EFD410BB910417044593E48E9BC0BF" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
) of holotype major worker (CASENT0498298).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||||
<paragraph id="399188C2C8194FB66D789CB6C825681C" pageId="0" pageNumber="1">
|
||||
<emphasis id="2646087E78E2FE389A0B728646CF7A1A" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||||
Measurements (
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
<emphasis id="3F4EFADF34D764C913FC8A99EB410A0E" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
= 10): HL: 0.49-0.55 (0.51); HW: 0.44-0.47 (0.45); SL: 0.46-0.47 (0.46); EL: 0.09-0.11 (0.1); WL: 0.57-0.6 (0.58); PSL: 0.09-0.11 (0.1); MTL: 0.33-0.36 (0.34); PNW: 0.29-0.32 (0.31); PTW: 0.06-0.08 (0.07); PPW: 0.1-0.12 (0.11); CI: 85.7-90.4 (88.6); SI: 100.2-105.6 (103.0); PSLI: 18.1-21.8 (20.0); PPI: 56.5-64.5 (60.8); PNI: 65.9-70.1 (68.2); MTI: 73.5-80.1 (76.1).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
<emphasis id="3617028DC05DCA78B9ACFC85093352EC" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="841454701A29D1F32200ADC541BA50B8" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Occipital margin straight or indistinctly concave; occipital carina absent (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39A</figureCitation>
|
||||
<figureCitation id="D407665653ED22DB790D9AB89AE396DF" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39A</figureCitation>
|
||||
). Pilosity moderately sparse, moderately long, suberect to erect. Head shiny, foveolate, with additional short, longitudinal, thick rugae on frons; genae with fading sculpture and smooth notch. Clypeus smooth, with basal part finely rugulose; median longitudinal carina present; two lateral longitudinal carinae indistinct. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity suberect to erect (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39A, C</figureCitation>
|
||||
<figureCitation id="1F14F4D1D981ADAF2DCD51948F713421" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39A, C</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
<emphasis id="AB2CF3918B528A8CA592FB1BA7B82DDD" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="789776920B5680264E1E3C148AD8BDD5" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
In lateral view, promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum; promesonotal groove absent; metanotal groove absent; propodeal spines short, triangular, with acute apex (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C</figureCitation>
|
||||
<figureCitation id="D4C845D79DD3B5D3D392D1D95DD906CF" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C</figureCitation>
|
||||
). Sculpture foveolate; katepisternum smooth; anepisternum and mesonotum with fading sculpture and sometimes smooth. Pilosity moderately sparse, long, and erect (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
|
||||
<figureCitation id="B8B1989009A2F086F3B90B85B47165A6" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="513D67D9F5D1000A644228844D6ADDA3" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="90BDE4835CC5AEE785FC349352F66040" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Peduncle short and thin with ventral face slightly convex; with few short, erect setae (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
|
||||
<figureCitation id="F04DDE84CF3C9CB42B2B3082DD68D9A9" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
<emphasis id="989D58046BD4E7CB8B7936360522FC94" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="01951126A1F370A09C29CDB6E6A6C14D" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Short, low, and convex; with few short, erect setae (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
|
||||
<figureCitation id="B8BA95041141CE2A678ED8579A216D13" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="6658337918FDC025AAE24E4D45E7C016" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="59CE5EB3F4ED0918F6068A1C219E99BE" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
With moderately sparse, erect pilosity (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
|
||||
<figureCitation id="DD8516A997AF1F6DC4CB04D07629FC05" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
<emphasis id="536A67E6CCA011FB8DBF34EA647FD80E" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="A30086538DB451D98EAFB95E840B7745" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Yellow (Fig.
|
||||
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
|
||||
<figureCitation id="D249B7E7E3DED059334E1923373AA5AB" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">Latin for long pilosity, in reference to the very long pilosity on sides of head in major workers.</paragraph>
|
||||
<subSubSection id="BB39166B1AF505784EB42720EB56FB28" pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph id="AF602AB98C27003504AA898C56D4C2F7" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph id="83FC8721C46D978C2BECFAF02FB10986" pageId="0" pageNumber="1">Latin for long pilosity, in reference to the very long pilosity on sides of head in major workers.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">The species was collected between 500-1100 m in elevation, in montane and gallery forests. Nests were located in rotten logs.</paragraph>
|
||||
<subSubSection id="F0B3D1DA5697300626DF9A74C1864A71" pageId="0" pageNumber="1" type="biology_ecology">
|
||||
<paragraph id="638C23B63AD64F2CAC66082F4172115F" pageId="0" pageNumber="1">Biology.</paragraph>
|
||||
<paragraph id="6CF63A6E37E9633DBEC21984BBFFA1EA" pageId="0" pageNumber="1">The species was collected between 500-1100 m in elevation, in montane and gallery forests. Nests were located in rotten logs.</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,185 +1,185 @@
|
|||
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<mods:affiliation id="E557455454968E815FD7D4DB402A30A6">The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:nameIdentifier id="3B2B6D07F03425A4890CFBDB5C7BBD6F" type="email">s.knapp@nhm.ac.uk</mods:nameIdentifier>
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<mods:title id="645F13CE0DC1FEA750F19EC21200D95A">PhytoKeys</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-01</mods:number>
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<mods:date id="D9916CCBED90D83FEDAA8C4888C5411A">2022</mods:date>
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<mods:number id="80657D234E75CA8F8C3FB3DB102ECDB4">2022-06-01</mods:number>
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<mods:detail type="volume">
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<mods:number>198</mods:number>
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<mods:number id="681B6C77C4D8C1106EE4996967D774F8">198</mods:number>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.198.79514</mods:url>
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<mods:url id="4BCDEE05634126E35CFF2778764BDBEC">http://dx.doi.org/10.3897/phytokeys.198.79514</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.198.79514</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-198-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">486F1F1B4F5854D2831AAA341B9A322C</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:D3D37CA3CFCE32A0851061B10E277C7C" httpUri="http://treatment.plazi.org/id/D3D37CA3CFCE32A0851061B10E277C7C" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<treatment id="D3D37CA3CFCE32A0851061B10E277C7C" ID-GBIF-Taxon="195723712" LSID="urn:lsid:plazi:treatment:D3D37CA3CFCE32A0851061B10E277C7C" httpUri="http://treatment.plazi.org/id/D3D37CA3CFCE32A0851061B10E277C7C" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Solanaceae">
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<subSubSection id="647064ABCD8C188E25686DC925ED1103" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="0CD25CD51A93B0C393C9365718DF1568" pageId="0" pageNumber="1">
|
||||
16.
|
||||
<taxonomicName authority="Dunal, Encycl. [J. Lamarck & al.] Suppl. 3: 763. 1814." authorityName="Dunal, Encycl. [J. Lamarck & al.] Suppl. 3: 763. 1814." class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">Solanum graciliflorum Dunal, Encycl. [J. Lamarck & al.] Suppl. 3: 763. 1814.</taxonomicName>
|
||||
<taxonomicName id="CFE1B5F3AD71A21998AB2EE64FDED45E" ID-CoL="99TH3" authority="Dunal, Encycl. [J. Lamarck & al.] Suppl. 3: 763. 1814." authorityName="Dunal, Encycl. [J. Lamarck & al.] Suppl. 3: 763. 1814." class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">Solanum graciliflorum Dunal, Encycl. [J. Lamarck & al.] Suppl. 3: 763. 1814.</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
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<subSubSection pageId="0" pageNumber="1" type="description">
|
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<paragraph pageId="0" pageNumber="1">
|
||||
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Solanum graciliflorum Dunal Herbarium specimen (epitype) collected in Indonesia (Horsfield s. n., BM 000886121). Photograph credit: CC-BY, © copyright The Trustees of the Natural History Museum, London." figureDoi="10.3897/phytokeys.198.79514.figure27" httpUri="https://binary.pensoft.net/fig/689720" pageId="0" pageNumber="1">Fig. 27</figureCitation>
|
||||
<subSubSection id="B7362CC553311E4CC352BDF7EEED102D" pageId="0" pageNumber="1" type="description">
|
||||
<paragraph id="8CCC6FBA5C64E3653B7536E701113BD4" pageId="0" pageNumber="1">
|
||||
<figureCitation id="46D863F643C1668A1D1EDD339C24704B" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Solanum graciliflorum Dunal Herbarium specimen (epitype) collected in Indonesia (Horsfield s. n., BM 000886121). Photograph credit: CC-BY, © copyright The Trustees of the Natural History Museum, London." figureDoi="10.3897/phytokeys.198.79514.figure27" httpUri="https://binary.pensoft.net/fig/689720" pageId="0" pageNumber="1">Fig. 27</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum athroanthum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="athroanthum">Solanum athroanthum</taxonomicName>
|
||||
<subSubSection id="436FD9790E2633C700BF28E583DAE89F" pageId="0" pageNumber="1" type="reference_group">
|
||||
<paragraph id="782D6FDA13FF0A88A8E78C205BF2B3C7" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="79A3C7972A4F24780F0F3D567A26D40C" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum athroanthum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="athroanthum">Solanum athroanthum</taxonomicName>
|
||||
Dunal, Prodr. [A. P. de Candolle] 13(1): 208. 1852. Type. Indonesia. Java: [Prov. Banjinwanyne] "in sylvis prope Sukaradja" [Sukaraja], 1846,
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">H. Zollinger 2907</emphasis>
|
||||
<emphasis id="AF323134AB89143D06461BE1E9005965" italics="true" pageId="0" pageNumber="1">H. Zollinger 2907</emphasis>
|
||||
(holotype: G-DC [G00145833]; isotypes: BM [BM000778325], G [G00301684, G003043306], MPU [MPU012648], P [P00368939, P00368940, P00368941]).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<typeStatus>Type</typeStatus>
|
||||
<subSubSection id="BACA9F0CE45DB0EEC07955B66C329216" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="755BA801B0650304DAE0D74DB056BBB3" pageId="0" pageNumber="1">
|
||||
<typeStatus id="477FFBC54F2FA3E8BE8C2696F699FED5">Type</typeStatus>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<paragraph id="914DE659D0B34BD8422545C9E3DEA9BD" pageId="0" pageNumber="1">
|
||||
Based on an unpublished illustration of Leschenault collection kept in the
|
||||
<normalizedToken originalValue="Node-Véran">Node-Veran</normalizedToken>
|
||||
<normalizedToken id="B786CCFD8A416839635E5F6664278E81" originalValue="Node-Véran">Node-Veran</normalizedToken>
|
||||
collection in Montpellier (
|
||||
<typeStatus>lectotype</typeStatus>
|
||||
<typeStatus id="7D14B71CDF9A0EAA2ECDFD6C9B41EB1B">lectotype</typeStatus>
|
||||
, designated by
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.70.9758" author="Aubriot, X" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="97 - 110" refId="B9" refString="Aubriot, X, Loup, C, Knapp, S, 2016b. Confirming the identity of two enigmatic "spiny solanums" (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java. PhytoKeys 70: 97 - 110, DOI: https://doi.org/10.3897/phytokeys.70.9758" title="Confirming the identity of two enigmatic " spiny solanums " (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java." url="https://doi.org/10.3897/phytokeys.70.9758" volume="70" year="2016 b">Aubriot et al. 2016b</bibRefCitation>
|
||||
<bibRefCitation id="5D94889EE9BC561831B8CF07F93A8C61" DOI="https://doi.org/10.3897/phytokeys.70.9758" author="Aubriot, X" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="97 - 110" refId="B9" refString="Aubriot, X, Loup, C, Knapp, S, 2016b. Confirming the identity of two enigmatic "spiny solanums" (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java. PhytoKeys 70: 97 - 110, DOI: https://doi.org/10.3897/phytokeys.70.9758" title="Confirming the identity of two enigmatic " spiny solanums " (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java." url="https://doi.org/10.3897/phytokeys.70.9758" volume="70" year="2016 b">Aubriot et al. 2016b</bibRefCitation>
|
||||
, pg. 100: Sol. Tab. 47 [MPU028534]).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/phytokeys.198.79514.figure27" httpUri="https://binary.pensoft.net/fig/689720" pageId="0" pageNumber="1" start="Figure 27" startId="F27">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 27.</emphasis>
|
||||
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Solanum graciliflorum</emphasis>
|
||||
<caption id="0CFBF8CEE29586A2341540B55EB38240" doi="10.3897/phytokeys.198.79514.figure27" httpUri="https://binary.pensoft.net/fig/689720" pageId="0" pageNumber="1" start="Figure 27" startId="F27">
|
||||
<paragraph id="8D440455A5A775E003C67793F77C59FA" pageId="0" pageNumber="1">
|
||||
<emphasis id="4CA18009B906C30BA9F0119DE2890974" bold="true" pageId="0" pageNumber="1">Figure 27.</emphasis>
|
||||
<taxonomicName id="1F71A1FA4081ADBD4594E4E44F1C9EA8" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">
|
||||
<emphasis id="F92E37E5DD91D9780668DF07A915D42B" italics="true" pageId="0" pageNumber="1">Solanum graciliflorum</emphasis>
|
||||
</taxonomicName>
|
||||
Dunal Herbarium specimen (epitype) collected in
|
||||
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
|
||||
<collectingCountry id="956D56F286CF0F8D57347BF2438AD7A9" name="Indonesia">Indonesia</collectingCountry>
|
||||
(
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Horsfield s.n.</emphasis>
|
||||
<emphasis id="C0D55E18512813683CD4BC20A54B5BF5" italics="true" pageId="0" pageNumber="1">Horsfield s.n.</emphasis>
|
||||
, BM000886121). Photograph credit: CC-BY, © copyright The Trustees of the Natural History Museum, London.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||||
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">Scandent shrub to 2 m tall, armed. Stems apparently erect to decumbent, prickly and very sparsely stellate-pubescent; prickles to 7 mm long, to 8 mm wide at the base, curved, deltate, laterally flattened, pale yellow, glabrous; trichomes porrect-stellate, sessile to subsessile, the rays (4-)5-8, 0.1-0.25 mm long, the midpoints to 0.15 mm long; new growth sparsely stellate-pubescent; bark of older stems dark brownish grey, glabrescent. Sympodial units difoliate, the leaves geminate, usually similar in size. Leaves simple, more or less deeply lobed, the blades (4.5-)7-11 cm long, (1.5-)3-5 cm wide, ca. 2 times longer than wide, elliptic to ovate, chartaceous, slightly discolorous, usually prickly with 1-10(-12) prickles per leaf side, mostly along the midvein, to 9 mm long, to 2 mm wide at the base, straight or slightly curved at the tip, awl-shaped, conical, pale yellow, glabrous; adaxial and abaxial surfaces sparsely to very sparsely stellate-pubescent with sessile to subsessile porrect-stellate trichomes, the rays 6-8, 0.1-0.25 mm long, the midpoint to 0.25 mm long, usually as long as the rays; major veins 3-4 pairs drying dark; base attenuate to truncate; margins shallowly to deeply lobed, the lobes 1-3 on each side, 0.5-2.5 cm long, broadly deltate, apically rounded, the sinuses extending up to 2/3 to the midrib; apex rounded to acute; petiole 0.5-1.8 cm long, 1/10-1/6 of the leaf blade length, unarmed or prickly with 1-2 prickles like those of the blades, sparsely stellate-pubescent with porrect, subsessile trichomes denser at the very base. Inflorescences 2-4 cm long, leaf-opposed or apparently lateral and borne between leaf pairs, unbranched to up to 6 times branched, with 15-50+ flowers, many flowers open at any one time, sparsely to very sparsely stellate-pubescent with trichomes like those of the stems, unarmed; peduncle 1-2(-2.5) cm long, with 0-1 prickles like those of the leaves and stems; pedicels 4-7 mm long, ca. 0.5 mm in diameter at the base, ca. 1 mm in diameter at the apex, erect, unarmed and very sparsely stellate-pubescent with trichomes like those of the stems, articulated at the base; pedicel scars spaced 1-5 mm apart. Buds ellipsoid, strongly exserted from the calyx before anthesis. Flowers 5-merous, apparently all perfect. Calyx with the tube 1-1.5 mm long, campanulate, the lobes 0.25-0.75 mm long, 0.25-0.75 mm wide, deltate, apically acute, unarmed and pubescent abaxially with sessile porrect-stellate trichomes like those of the stems. Corolla 0.5-1 cm in diameter, white to pale lilac, stellate, lobed nearly to the base, the lobes 4-5 mm long, ca. 1 mm wide, narrowly deltate to linear, reflexed at anthesis, glabrous adaxially, densely stellate-pubescent abaxially with sessile porrect trichomes where exposed in bud. Stamens slightly unequal; anthers unequal, three of the five 4.5-5 mm long and two 3-4 mm long, all 0.5-0.75 mm wide, somewhat connivent, tapering, yellow, glabrous, poricidal at the tips, the pores not elongating to slits with drying; filament tube <0.5 mm long, glabrous; free portion of the filaments almost equal, 0.5-1.25 mm long, glabrous. Ovary conical, minutely glandular-puberulent; style ca. 5.5 mm long, slender, curved at the apex, glabrous; stigma capitate, the surfaces minutely papillate. Fruit a globose berry, several to many per infructescence, 0.3-0.5 cm in diameter, red when mature, the pericarp shiny, glabrous; fruiting pedicels 0.8-1.2 cm long, ca. 0.5 mm in diameter at the base, tapering to a slightly enlarged apex, 0.75-1 mm in diameter at the apex, somewhat woody, spreading, unarmed; fruiting calyx lobes slightly expanding to 1.5 mm long, ca. 1/5 the length of the mature fruit, deltate to lanceolate, unarmed. Seeds 6-9 per berry, 3.5-4 mm long, 3-3.5 mm wide, flattened-reniform, orange-brown, the surface minutely pitted, the testal cells pentagonal in outline. Chromosome number: not known.</paragraph>
|
||||
<caption doi="10.3897/phytokeys.198.79514.figure28" httpUri="https://binary.pensoft.net/fig/689721" pageId="0" pageNumber="1" start="Figure 28" startId="F28">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 28.</emphasis>
|
||||
<subSubSection id="F5B4628B5B11AE3496C410F41F42B775" pageId="0" pageNumber="1" type="description">
|
||||
<paragraph id="5480B333C658FC24B7361C84764D1DE6" pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph id="82BDD85BFE9AE5E177D96351CAF78AF7" pageId="0" pageNumber="1">Scandent shrub to 2 m tall, armed. Stems apparently erect to decumbent, prickly and very sparsely stellate-pubescent; prickles to 7 mm long, to 8 mm wide at the base, curved, deltate, laterally flattened, pale yellow, glabrous; trichomes porrect-stellate, sessile to subsessile, the rays (4-)5-8, 0.1-0.25 mm long, the midpoints to 0.15 mm long; new growth sparsely stellate-pubescent; bark of older stems dark brownish grey, glabrescent. Sympodial units difoliate, the leaves geminate, usually similar in size. Leaves simple, more or less deeply lobed, the blades (4.5-)7-11 cm long, (1.5-)3-5 cm wide, ca. 2 times longer than wide, elliptic to ovate, chartaceous, slightly discolorous, usually prickly with 1-10(-12) prickles per leaf side, mostly along the midvein, to 9 mm long, to 2 mm wide at the base, straight or slightly curved at the tip, awl-shaped, conical, pale yellow, glabrous; adaxial and abaxial surfaces sparsely to very sparsely stellate-pubescent with sessile to subsessile porrect-stellate trichomes, the rays 6-8, 0.1-0.25 mm long, the midpoint to 0.25 mm long, usually as long as the rays; major veins 3-4 pairs drying dark; base attenuate to truncate; margins shallowly to deeply lobed, the lobes 1-3 on each side, 0.5-2.5 cm long, broadly deltate, apically rounded, the sinuses extending up to 2/3 to the midrib; apex rounded to acute; petiole 0.5-1.8 cm long, 1/10-1/6 of the leaf blade length, unarmed or prickly with 1-2 prickles like those of the blades, sparsely stellate-pubescent with porrect, subsessile trichomes denser at the very base. Inflorescences 2-4 cm long, leaf-opposed or apparently lateral and borne between leaf pairs, unbranched to up to 6 times branched, with 15-50+ flowers, many flowers open at any one time, sparsely to very sparsely stellate-pubescent with trichomes like those of the stems, unarmed; peduncle 1-2(-2.5) cm long, with 0-1 prickles like those of the leaves and stems; pedicels 4-7 mm long, ca. 0.5 mm in diameter at the base, ca. 1 mm in diameter at the apex, erect, unarmed and very sparsely stellate-pubescent with trichomes like those of the stems, articulated at the base; pedicel scars spaced 1-5 mm apart. Buds ellipsoid, strongly exserted from the calyx before anthesis. Flowers 5-merous, apparently all perfect. Calyx with the tube 1-1.5 mm long, campanulate, the lobes 0.25-0.75 mm long, 0.25-0.75 mm wide, deltate, apically acute, unarmed and pubescent abaxially with sessile porrect-stellate trichomes like those of the stems. Corolla 0.5-1 cm in diameter, white to pale lilac, stellate, lobed nearly to the base, the lobes 4-5 mm long, ca. 1 mm wide, narrowly deltate to linear, reflexed at anthesis, glabrous adaxially, densely stellate-pubescent abaxially with sessile porrect trichomes where exposed in bud. Stamens slightly unequal; anthers unequal, three of the five 4.5-5 mm long and two 3-4 mm long, all 0.5-0.75 mm wide, somewhat connivent, tapering, yellow, glabrous, poricidal at the tips, the pores not elongating to slits with drying; filament tube <0.5 mm long, glabrous; free portion of the filaments almost equal, 0.5-1.25 mm long, glabrous. Ovary conical, minutely glandular-puberulent; style ca. 5.5 mm long, slender, curved at the apex, glabrous; stigma capitate, the surfaces minutely papillate. Fruit a globose berry, several to many per infructescence, 0.3-0.5 cm in diameter, red when mature, the pericarp shiny, glabrous; fruiting pedicels 0.8-1.2 cm long, ca. 0.5 mm in diameter at the base, tapering to a slightly enlarged apex, 0.75-1 mm in diameter at the apex, somewhat woody, spreading, unarmed; fruiting calyx lobes slightly expanding to 1.5 mm long, ca. 1/5 the length of the mature fruit, deltate to lanceolate, unarmed. Seeds 6-9 per berry, 3.5-4 mm long, 3-3.5 mm wide, flattened-reniform, orange-brown, the surface minutely pitted, the testal cells pentagonal in outline. Chromosome number: not known.</paragraph>
|
||||
<caption id="C9AA9854421E24370554512F829D9A38" doi="10.3897/phytokeys.198.79514.figure28" httpUri="https://binary.pensoft.net/fig/689721" pageId="0" pageNumber="1" start="Figure 28" startId="F28">
|
||||
<paragraph id="165562FC7558EA5B2A68FDE2C5035295" pageId="0" pageNumber="1">
|
||||
<emphasis id="65BA3D74FDB61C9EB71FB474C1A24CE3" bold="true" pageId="0" pageNumber="1">Figure 28.</emphasis>
|
||||
Distribution of
|
||||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. graciliflorum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="graciliflorum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. graciliflorum</emphasis>
|
||||
<taxonomicName id="5D6783811A31C4EF561704C900FDE366" class="Magnoliopsida" family="Solanaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">
|
||||
<emphasis id="97CFE3C08B56A2A9F368427EDCB4B33C" italics="true" pageId="0" pageNumber="1">S. graciliflorum</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="1">Distribution</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<subSubSection id="FDAB899783762BAF19B3F29DDDD5A365" pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph id="DFAF7F2EF48D21B261F8CCCB47BB482C" pageId="0" pageNumber="1">Distribution</paragraph>
|
||||
<paragraph id="19A7688B9590E0012C342A77C1B09965" pageId="0" pageNumber="1">
|
||||
<emphasis id="65F2E97673397F8D3CC716A52580CC63" bold="true" pageId="0" pageNumber="1">
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of S. graciliflorum." figureDoi="10.3897/phytokeys.198.79514.figure28" httpUri="https://binary.pensoft.net/fig/689721" pageId="0" pageNumber="1">28</figureCitation>
|
||||
<figureCitation id="1571E2B8BCCAE6A47509C29AD24B54FA" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of S. graciliflorum." figureDoi="10.3897/phytokeys.198.79514.figure28" httpUri="https://binary.pensoft.net/fig/689721" pageId="0" pageNumber="1">28</figureCitation>
|
||||
).
|
||||
</emphasis>
|
||||
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Solanum graciliflorum</emphasis>
|
||||
<taxonomicName id="F21CA177BBAC8E0A77B87E6F72056349" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">
|
||||
<emphasis id="35B0E699709437E0C9DC70E9AEAB73F6" italics="true" pageId="0" pageNumber="1">Solanum graciliflorum</emphasis>
|
||||
</taxonomicName>
|
||||
is known from only a few collections from the islands of Java, Bali, Sulawesi and Ambon (Indonesia).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="ecology">
|
||||
<paragraph pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Solanum graciliflorum</emphasis>
|
||||
<subSubSection id="B4CF8AD3914C2129F6EDDFC48051D6FE" pageId="0" pageNumber="1" type="ecology">
|
||||
<paragraph id="C291A2FD824799276BB7F50CB49D5559" pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
|
||||
<paragraph id="5CDDC3E62D9DC727DCF75A35A79D4F6E" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="62438DE25D1E0C6E2841412DA772BB8C" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">
|
||||
<emphasis id="9A73C324D568726CF6C059931E075B76" italics="true" pageId="0" pageNumber="1">Solanum graciliflorum</emphasis>
|
||||
</taxonomicName>
|
||||
is a plant of tropical rainforest understory; elevation not recorded.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="common names and uses">
|
||||
<paragraph pageId="0" pageNumber="1">Common names and uses.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">None recorded.</paragraph>
|
||||
<subSubSection id="0B54A298C0410E772F7D93197AFA5540" pageId="0" pageNumber="1" type="common names and uses">
|
||||
<paragraph id="EB1794295762F989B07C8CB7F73DBBD9" pageId="0" pageNumber="1">Common names and uses.</paragraph>
|
||||
<paragraph id="E1C6E834F40C645945A9B3ED4E356008" pageId="0" pageNumber="1">None recorded.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="preliminary conservation status">
|
||||
<paragraph pageId="0" pageNumber="1">Preliminary conservation status</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<subSubSection id="94334935C4CB35C90140D962EE9BA1EB" pageId="0" pageNumber="1" type="preliminary conservation status">
|
||||
<paragraph id="4FE061116502F6E68FAF1F303D709A81" pageId="0" pageNumber="1">Preliminary conservation status</paragraph>
|
||||
<paragraph id="212096E386EA7E090864D14CD075D0F0" pageId="0" pageNumber="1">
|
||||
<emphasis id="4DDD3A0AA369D5131FC54704CC693D95" bold="true" pageId="0" pageNumber="1">
|
||||
(
|
||||
<bibRefCitation author="Hutchings, A" journalOrPublisher="AGRIVITA Journal of Agricultural Science" pageId="0" pageNumber="1" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B105" refString="2019. . http://www.iucnredlist.org/documents/RedListGuidelines.pdf" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2019">IUCN 2019</bibRefCitation>
|
||||
<bibRefCitation id="604C9E2BAE7652C01F169A5E487DD105" author="Hutchings, A" journalOrPublisher="AGRIVITA Journal of Agricultural Science" pageId="0" pageNumber="1" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B105" refString="2019. . http://www.iucnredlist.org/documents/RedListGuidelines.pdf" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2019">IUCN 2019</bibRefCitation>
|
||||
).
|
||||
</emphasis>
|
||||
Near Threatened (NT). EOO (131,936 km2); AOO (16 km2).
|
||||
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Solanum graciliflorum</emphasis>
|
||||
<taxonomicName id="7CF7D790898912F9367EADB9A39A7A90" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">
|
||||
<emphasis id="59F9B00D1E221D4C087F0A639B9F975F" italics="true" pageId="0" pageNumber="1">Solanum graciliflorum</emphasis>
|
||||
</taxonomicName>
|
||||
is known only from seven collections, several of which are of uncertain localities. It has not been re-collected since the first half of the 20th century, indicating it is certainly of conservation concern, thus we suggest a threat status of Near Threatened to reflect this. Additional collection of this species and exploration of the type locality are priorities.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="discussion">
|
||||
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Solanum graciliflorum</emphasis>
|
||||
<subSubSection id="680BC28C7D4D37EFA74F90B71F268A06" pageId="0" pageNumber="1" type="discussion">
|
||||
<paragraph id="24F4746834A36B9F0BA67A45D016800D" pageId="0" pageNumber="1">Discussion.</paragraph>
|
||||
<paragraph id="A95E3EAE8D5CC64EA7CE0A94561BEE16" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="B3C4CE4739ACB34C36153FFFEDF2CA4A" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum graciliflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="graciliflorum">
|
||||
<emphasis id="AD4740DAD79CAFFA563EE89DC8F0F8A7" italics="true" pageId="0" pageNumber="1">Solanum graciliflorum</emphasis>
|
||||
</taxonomicName>
|
||||
is morphologically similar to the sympatric
|
||||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. cyanocarphium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cyanocarphium">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. cyanocarphium</emphasis>
|
||||
<taxonomicName id="05CC0DDC0B2A8C9610B5AB2ED02A8A49" class="Magnoliopsida" family="Solanaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. cyanocarphium" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cyanocarphium">
|
||||
<emphasis id="3889B2F570ABE2F5E8D65F6780C39D82" italics="true" pageId="0" pageNumber="1">S. cyanocarphium</emphasis>
|
||||
</taxonomicName>
|
||||
in being a weak, scrambling plant with hooked prickles, but differs from it in its branched inflorescences, smaller flowers and fruits and calyx that is not accrescent in fruit. The inflorescence of
|
||||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. cyanocarphium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cyanocarphium">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. cyanocarphium</emphasis>
|
||||
<taxonomicName id="AE85C404F2C9953CAF582EAA7317900D" class="Magnoliopsida" family="Solanaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. cyanocarphium" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cyanocarphium">
|
||||
<emphasis id="415996E0680680AC8714B3E7DCDEEFA1" italics="true" pageId="0" pageNumber="1">S. cyanocarphium</emphasis>
|
||||
</taxonomicName>
|
||||
is unbranched and the calyx strongly accrescent and covers the berry in fruit.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.70.9758" author="Aubriot, X" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="97 - 110" refId="B9" refString="Aubriot, X, Loup, C, Knapp, S, 2016b. Confirming the identity of two enigmatic "spiny solanums" (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java. PhytoKeys 70: 97 - 110, DOI: https://doi.org/10.3897/phytokeys.70.9758" title="Confirming the identity of two enigmatic " spiny solanums " (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java." url="https://doi.org/10.3897/phytokeys.70.9758" volume="70" year="2016 b">Aubriot et al. (2016b)</bibRefCitation>
|
||||
<paragraph id="844A87CBD06C605C0578FD5543F2CBCA" pageId="0" pageNumber="1">
|
||||
<bibRefCitation id="63C0BFC46FDF9F6565CC459C6913682B" DOI="https://doi.org/10.3897/phytokeys.70.9758" author="Aubriot, X" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="97 - 110" refId="B9" refString="Aubriot, X, Loup, C, Knapp, S, 2016b. Confirming the identity of two enigmatic "spiny solanums" (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java. PhytoKeys 70: 97 - 110, DOI: https://doi.org/10.3897/phytokeys.70.9758" title="Confirming the identity of two enigmatic " spiny solanums " (Solanum subgenus Leptostemonum, Solanaceae) collected by Jean-Baptiste Leschenault in Java." url="https://doi.org/10.3897/phytokeys.70.9758" volume="70" year="2016 b">Aubriot et al. (2016b)</bibRefCitation>
|
||||
superfluously lectotypified
|
||||
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. athroanthum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="athroanthum">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">S. athroanthum</emphasis>
|
||||
<taxonomicName id="F5E5BC7E06344D7F71CE1D4D7D8164A8" class="Magnoliopsida" family="Solanaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="S. athroanthum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="athroanthum">
|
||||
<emphasis id="A91957FF1BD4596D07C87AA8729AEE45" italics="true" pageId="0" pageNumber="1">S. athroanthum</emphasis>
|
||||
</taxonomicName>
|
||||
with the specimen in G-DC (G00145833) but
|
||||
<bibRefCitation author="Dunal, M-F" journalOrPublisher=") Prodromus systematis naturalis regni vegetabilis" pageId="0" pageNumber="1" pagination="1 - 690" refId="B73" refString="Dunal, M-F, 1852. Solanaceae. In: Candolle AP de (Ed.) Prodromus systematis naturalis regni vegetabilis 13 (1): 1 - 690" title="Solanaceae. In: Candolle AP de (Ed." volume="13" year="1852">Dunal (1852)</bibRefCitation>
|
||||
<bibRefCitation id="2C788959760122F96F4AFB636A324265" author="Dunal, M-F" journalOrPublisher=") Prodromus systematis naturalis regni vegetabilis" pageId="0" pageNumber="1" pagination="1 - 690" refId="B73" refString="Dunal, M-F, 1852. Solanaceae. In: Candolle AP de (Ed.) Prodromus systematis naturalis regni vegetabilis 13 (1): 1 - 690" title="Solanaceae. In: Candolle AP de (Ed." volume="13" year="1852">Dunal (1852)</bibRefCitation>
|
||||
clearly cited only this specimen in the protologue, thus making it the holotype.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">See Suppl. materials 1-3.</paragraph>
|
||||
<subSubSection id="E41E8798D7401C38C9CC0687C6D016C7" pageId="0" pageNumber="1" type="materials_examined">
|
||||
<paragraph id="0C7584CA4598B303FF61E98C068C8C63" pageId="0" pageNumber="1">Specimens examined.</paragraph>
|
||||
<paragraph id="3B8721C65A586E99A29BAB483B562164" pageId="0" pageNumber="1">See Suppl. materials 1-3.</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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