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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:title id="4DABE2B30E97C803F89428ECC44134EB">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<treatment ID-GBIF-Taxon="161777032" LSID="urn:lsid:zoobank.org:act:CB6E1BA0-DC84-4613-85DD-53DC22FCDF80" httpUri="http://treatment.plazi.org/id/C5FF7EB3DB2DFC9F7C1750C659A0D940" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/CB6E1BA0-DC84-4613-85DD-53DC22FCDF80" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole bemarivoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemarivoensis" status="sp. nov.">Pheidole bemarivoensis</taxonomicName>
|
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">Figs 60A-F</figureCitation>
|
||||
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84H</figureCitation>
|
||||
<figureCitation captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86H</figureCitation>
|
||||
<treatment id="C5FF7EB3DB2DFC9F7C1750C659A0D940" ID-GBIF-Taxon="161777032" LSID="urn:lsid:zoobank.org:act:CB6E1BA0-DC84-4613-85DD-53DC22FCDF80" httpUri="http://treatment.plazi.org/id/C5FF7EB3DB2DFC9F7C1750C659A0D940" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="B3972FB186525F95F14EFAFCBEAE7CAE" pageId="0" pageNumber="1" type="nomenclature">
|
||||
<paragraph id="02BEF91DCBE8641A396BDCE54F60A15B" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="3CA5DBB5372A850EACFE560936895BEA" ID-CoL="CMKWP" LSID="http://zoobank.org/CB6E1BA0-DC84-4613-85DD-53DC22FCDF80" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole bemarivoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemarivoensis" status="sp. nov.">Pheidole bemarivoensis</taxonomicName>
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||||
<taxonomicNameLabel id="3D58CCB4D8CAF57163FAE8B2EDC25608" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
|
||||
<figureCitation id="3AC671125B9FD6A59C121F1215173BEC" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">Figs 60A-F</figureCitation>
|
||||
<figureCitation id="83D690A171319593087ACB134A8F601F" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84H</figureCitation>
|
||||
<figureCitation id="C5B1A33939C6D0BEF68F7F66DDFA0096" captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86H</figureCitation>
|
||||
</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="type material">
|
||||
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||||
<subSubSection id="2D9781A3A3AEB7CC0235E1EDC98F46CE" pageId="0" pageNumber="1" type="type material">
|
||||
<paragraph id="DB48A5474FB3CAA6AA76EDAC36279940" pageId="0" pageNumber="1">Type material.</paragraph>
|
||||
<paragraph id="6C138BF0C57669574F8FEEA0459A30A0" pageId="0" pageNumber="1">
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||||
<emphasis id="CD55DB145FBF1716A481DF84AB814105" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="CD77214D7B3968392A3A08D3A2EA3047" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Mahajanga;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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<normalizedToken id="C03DA2AA59A1346EB30B02B04F285B2A" originalValue="Spéciale">Speciale</normalizedToken>
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de Bemarivo, 23.8 km 223°SW Besalampy;
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<geoCoordinate degrees="16.925" direction="south" orientation="latitude" precision="55" value="-16.925">-16.925</geoCoordinate>
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||||
<geoCoordinate id="295342CA83E8914E13B500398C4981A4" degrees="16.925" direction="south" orientation="latitude" precision="55" value="-16.925">-16.925</geoCoordinate>
|
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,
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<geoCoordinate degrees="44.36833" direction="east" orientation="longitude" precision="1" value="44.36833">44.36833</geoCoordinate>
|
||||
<geoCoordinate id="7E019E5716FFA6EBA198E41CC8CF6115" degrees="44.36833" direction="east" orientation="longitude" precision="1" value="44.36833">44.36833</geoCoordinate>
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; alt. 30 m; 19 Nov 2002; B.L. Fisher et al. leg.; BLF06797, CASENT0489439 (CASC).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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<emphasis id="F7AC19B071FC68E567B4A99820982CCC" bold="true" pageId="0" pageNumber="1">
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<emphasis id="EC29DF73DA1E1D7961289C93DD272E61" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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||||
.
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</emphasis>
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Madagascar. •8w., 2s., 6m.; same data as for holotype; CASENT0923160, CASENT0235042, CASENT0489441, CASENT0489440, CASENT0489442, CASENT0489438, CASENT0489437, CASENT0872160-CASENT0872168 (CASC).
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</paragraph>
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</subSubSection>
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||||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="649535CD44D950F92B3F03C241397C1C" pageId="0" pageNumber="1" type="materials_examined">
|
||||
<paragraph id="4DF8FB3D2A89BF25FEC6F83FE18D4052" pageId="0" pageNumber="1">Other material.</paragraph>
|
||||
<paragraph id="F89CA8A090A3D40752695E6E5BA76CDB" pageId="0" pageNumber="1">
|
||||
Madagascar. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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<emphasis id="A99BA181F8CDA7A71BD4AAB8091B6D04" bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: •4w., 3s., 1q.; Ampasindava, Andranomatavy Forest;
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||||
<geoCoordinate degrees="13.6648" direction="south" orientation="latitude" precision="5" value="-13.6648">-13.6648</geoCoordinate>
|
||||
<geoCoordinate id="AB20998EDBCD414035AF593748D90DFA" degrees="13.6648" direction="south" orientation="latitude" precision="5" value="-13.6648">-13.6648</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="47.98702" direction="east" orientation="longitude" precision="1" value="47.98702">47.98702</geoCoordinate>
|
||||
<geoCoordinate id="EF3F5A0CB03B1F95A8175D6079B9D29B" degrees="47.98702" direction="east" orientation="longitude" precision="1" value="47.98702">47.98702</geoCoordinate>
|
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; alt. 275 m; 7 Oct 2013; B.L. Fisher et al. leg.; CASENT0369320, CASENT0369321, CASENT0369333, CASENT0370561 (CASC). •9w., 6s.; Ampasindava,
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken originalValue="d’Ambilanivy">d'Ambilanivy</normalizedToken>
|
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|
||||
, 3.9 km 181°S Ambaliha;
|
||||
<geoCoordinate degrees="13.79861" direction="south" orientation="latitude" precision="1" value="-13.79861">-13.79861</geoCoordinate>
|
||||
<geoCoordinate id="8FDFDFB73CF48F24133CD68625D71F48" degrees="13.79861" direction="south" orientation="latitude" precision="1" value="-13.79861">-13.79861</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="48.16167" direction="east" orientation="longitude" precision="1" value="48.16167">48.16167</geoCoordinate>
|
||||
<geoCoordinate id="DCA8B31F017CE2F0DEA894099B798CAE" degrees="48.16167" direction="east" orientation="longitude" precision="1" value="48.16167">48.16167</geoCoordinate>
|
||||
; alt. 600 m; 4 Mar 2001; B.L. Fisher et al. leg.; CASENT0420033-CASENT0420035, CASENT0420037, CASENT0420038, CASENT0421224, CASENT0427706, CASENT0427707, CASENT0427894 (CASC). •2w., 2s.;
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="90817EF757C1E42F3AC5818D3D2C315C" originalValue="Forêt">Foret</normalizedToken>
|
||||
Ambato, 26.6 km 33° Ambanja;
|
||||
<geoCoordinate degrees="13.4645" direction="south" orientation="latitude" precision="5" value="-13.4645">-13.4645</geoCoordinate>
|
||||
<geoCoordinate id="FD1FB45E6503FFB323336C5A3C903B58" degrees="13.4645" direction="south" orientation="latitude" precision="5" value="-13.4645">-13.4645</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="48.55167" direction="east" orientation="longitude" precision="1" value="48.55167">48.55167</geoCoordinate>
|
||||
<geoCoordinate id="13A7D1E0F019EA5D8E7C0AEF7BCCAC7A" degrees="48.55167" direction="east" orientation="longitude" precision="1" value="48.55167">48.55167</geoCoordinate>
|
||||
; alt. 150 m; 10 Dec 2004; B.L. Fisher leg.; CASENT0107077, CASENT0107715 (CASC). •4w., 2s.; Galoko chain, Mont Galoko;
|
||||
<geoCoordinate degrees="13.58745" direction="south" orientation="latitude" precision="1" value="-13.58745">-13.58745</geoCoordinate>
|
||||
<geoCoordinate id="08EA31B375A4DCD8F7F5B9CFD179D957" degrees="13.58745" direction="south" orientation="latitude" precision="1" value="-13.58745">-13.58745</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="48.71419" direction="east" orientation="longitude" precision="1" value="48.71419">48.71419</geoCoordinate>
|
||||
<geoCoordinate id="E2F6434F9BBC621A2DD7D14140EEDFE0" degrees="48.71419" direction="east" orientation="longitude" precision="1" value="48.71419">48.71419</geoCoordinate>
|
||||
; alt. 380 m; 23 Feb 2013; B.L. Fisher et al. leg.; CASENT0302986, CASENT0302993, CASENT0302995, CASENT0303012, CASENT0303013, CASENT0303015 (CASC). •2w., 1s.; Galoko chain, Mont Galoko;
|
||||
<geoCoordinate degrees="13.58487" direction="south" orientation="latitude" precision="1" value="-13.58487">-13.58487</geoCoordinate>
|
||||
<geoCoordinate id="E3A04302E7455FAE1CC47F6E9D861469" degrees="13.58487" direction="south" orientation="latitude" precision="1" value="-13.58487">-13.58487</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="48.71818" direction="east" orientation="longitude" precision="1" value="48.71818">48.71818</geoCoordinate>
|
||||
<geoCoordinate id="CA02F36A2778C036590948FBDC6AA98D" degrees="48.71818" direction="east" orientation="longitude" precision="1" value="48.71818">48.71818</geoCoordinate>
|
||||
; alt. 520 m; 19 Feb 2013; B.L. Fisher et al. leg.; CASENT0302958, CASENT0303564 (CASC). •5w., 4s.; Nosy Be,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="422B1E2C692100E62E32DF29209D5752" originalValue="Réserve">Reserve</normalizedToken>
|
||||
Naturelle
|
||||
<normalizedToken originalValue="Intégrale">Integrale</normalizedToken>
|
||||
<normalizedToken id="32B36C088F347677E6F1024EB2CAE4FC" originalValue="Intégrale">Integrale</normalizedToken>
|
||||
de Lokobe, 6.3 km 112°ESE Hellville;
|
||||
<geoCoordinate degrees="13.41933" direction="south" orientation="latitude" precision="1" value="-13.41933">-13.41933</geoCoordinate>
|
||||
<geoCoordinate id="7B20B49EE0B3A7A13C9FFAE266AF6891" degrees="13.41933" direction="south" orientation="latitude" precision="1" value="-13.41933">-13.41933</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="48.33117" direction="east" orientation="longitude" precision="1" value="48.33117">48.33117</geoCoordinate>
|
||||
<geoCoordinate id="060FABFF2032B7104175C51D8D8825F2" degrees="48.33117" direction="east" orientation="longitude" precision="1" value="48.33117">48.33117</geoCoordinate>
|
||||
; alt. 30 m; 19 Mar 2001; B.L. Fisher et al. leg.; CASENT0421454, CASENT0421455, CASENT0427817, CASENT0427823, CASENT0427829, CASENT0466223, CASENT0466279 (CASC). •1w., 3s.; R.S. Manongarivo, 10.8 km 229°SW Antanambao;
|
||||
<geoCoordinate degrees="13.96167" direction="south" orientation="latitude" precision="1" value="-13.96167">-13.96167</geoCoordinate>
|
||||
<geoCoordinate id="702A62CBF49DEB99FBA5DCE3F88FB6E9" degrees="13.96167" direction="south" orientation="latitude" precision="1" value="-13.96167">-13.96167</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="48.43333" direction="east" orientation="longitude" precision="1" value="48.43333">48.43333</geoCoordinate>
|
||||
<geoCoordinate id="F6251CEBA17A10741B8E548A24662E05" degrees="48.43333" direction="east" orientation="longitude" precision="1" value="48.43333">48.43333</geoCoordinate>
|
||||
; alt. 400 m; 8 Nov 1998; B.L. Fisher leg.; CASENT0198891, CASENT0198892 (CASC). •2w., 6s.; Sahamalaza Peninsula,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken originalValue="d’Anabohazo">d'Anabohazo</normalizedToken>
|
||||
<normalizedToken id="A1139439BC70ECE94D996FDEF0344EDE" originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="C17825FB0F292B99342622C7B00F7750" originalValue="d’Anabohazo">d'Anabohazo</normalizedToken>
|
||||
, 21.6 km 247°WSW Maromandia;
|
||||
<geoCoordinate degrees="14.30889" direction="south" orientation="latitude" precision="1" value="-14.30889">-14.30889</geoCoordinate>
|
||||
<geoCoordinate id="77152C9685FE923B2BFCCE52CE320E34" degrees="14.30889" direction="south" orientation="latitude" precision="1" value="-14.30889">-14.30889</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="47.91433" direction="east" orientation="longitude" precision="1" value="47.91433">47.91433</geoCoordinate>
|
||||
<geoCoordinate id="8EBDB2C799C4BE4354E14EA476B3F4D9" degrees="47.91433" direction="east" orientation="longitude" precision="1" value="47.91433">47.91433</geoCoordinate>
|
||||
; alt. 120 m; 11 Mar 2001; B.L. Fisher et al. leg.; CASENT0406442, CASENT0406444, CASENT0406447, CASENT0406454, CASENT0458224, CASENT0484564 (CASC). -
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mahajanga</emphasis>
|
||||
<emphasis id="0D8FAE0196BFE1B1CF39969FA3EE81BC" bold="true" italics="true" pageId="0" pageNumber="1">Mahajanga</emphasis>
|
||||
: •4s.; Parc National de Namoroka, 16.9 km 317°NW Vilanandro; -16.40667, 45.31; alt. 100 m; 12 Nov 2002; B.L. Fisher et al. leg.; CASENT0038950, CASENT0038953, CASENT0486454 (CASC). •2w., 1s.; Parc National Tsingy de Bemaraha, 10.6 km ESE 123° Antsalova;
|
||||
<geoCoordinate degrees="18.70944" direction="south" orientation="latitude" precision="1" value="-18.70944">-18.70944</geoCoordinate>
|
||||
<geoCoordinate id="7CB58F9E54661E404738B98FFDFBF4BF" degrees="18.70944" direction="south" orientation="latitude" precision="1" value="-18.70944">-18.70944</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="44.71817" direction="east" orientation="longitude" precision="1" value="44.71817">44.71817</geoCoordinate>
|
||||
<geoCoordinate id="5634C119CF03ADB49B207695B7087D0E" degrees="44.71817" direction="east" orientation="longitude" precision="1" value="44.71817">44.71817</geoCoordinate>
|
||||
; alt. 150 m; 16 Nov 2001; B.L. Fisher et al. leg.; CASENT0437624, CASENT0437625, CASENT0437632 (CASC). •6w., 3s., 1m.; Parc National Tsingy de Bemaraha, 2.5 km 62°ENE Bekopaka, Ankidrodroa River;
|
||||
<geoCoordinate degrees="19.13222" direction="south" orientation="latitude" precision="1" value="-19.13222">-19.13222</geoCoordinate>
|
||||
<geoCoordinate id="289A11DDF1291BDE55494CFE38B93097" degrees="19.13222" direction="south" orientation="latitude" precision="1" value="-19.13222">-19.13222</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="44.81467" direction="east" orientation="longitude" precision="1" value="44.81467">44.81467</geoCoordinate>
|
||||
<geoCoordinate id="A303C4E684AE2A29FD33CA539FF0830F" degrees="44.81467" direction="east" orientation="longitude" precision="1" value="44.81467">44.81467</geoCoordinate>
|
||||
; alt. 100 m; 11 Nov 2001; B.L. Fisher et al. leg.; CASENT0425317, CASENT0425320, CASENT0425323, CASENT0443937, CASENT0443958, CASENT0443966, CASENT0443973, CASENT0444152, CASENT0444155 (CASC). •1w.;
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
|
||||
<normalizedToken id="FDDE7FCC2D946EF661356B8A73A3580E" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="0ED4B5A291FB7DDBFA6EAB958F44A326" originalValue="forestière">forestiere</normalizedToken>
|
||||
Beanka, 53.6 km E Maintirano;
|
||||
<geoCoordinate degrees="18.04014" direction="south" orientation="latitude" precision="1" value="-18.04014">-18.04014</geoCoordinate>
|
||||
<geoCoordinate id="957FDE561E737C49A9EE6BBDECD64B16" degrees="18.04014" direction="south" orientation="latitude" precision="1" value="-18.04014">-18.04014</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="44.53394" direction="east" orientation="longitude" precision="1" value="44.53394">44.53394</geoCoordinate>
|
||||
<geoCoordinate id="14B5C4C7E420DC95AF4A5251A38AB928" degrees="44.53394" direction="east" orientation="longitude" precision="1" value="44.53394">44.53394</geoCoordinate>
|
||||
; alt. 272 m; 25 Oct 2009; B.L. Fisher et al. leg.; CASENT0156697 (CASC). •6w., 8s.;
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="C9673E2D277F9446AA3219D5716ED299" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="4F9DDF2C8211765D711C5CF81AB2EF38" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
de Bemarivo, 23.8 km 223°SW Besalampy;
|
||||
<geoCoordinate degrees="16.925" direction="south" orientation="latitude" precision="55" value="-16.925">-16.925</geoCoordinate>
|
||||
<geoCoordinate id="590D39C6593F0C3F21AB819ADEB64CC1" degrees="16.925" direction="south" orientation="latitude" precision="55" value="-16.925">-16.925</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="44.36833" direction="east" orientation="longitude" precision="1" value="44.36833">44.36833</geoCoordinate>
|
||||
<geoCoordinate id="80AD136C8865B4783AF586E990D6726E" degrees="44.36833" direction="east" orientation="longitude" precision="1" value="44.36833">44.36833</geoCoordinate>
|
||||
; alt. 30 m; 19 Nov 2002; B.L. Fisher et al. leg.; CASENT0491028, CASENT0491029, CASENT0491089, CASENT0491090, CASENT0491091 (CASC).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||||
<subSubSection id="8D3EFC9C8F3EE17377DEFCC992D037AA" pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph id="46279341C8997B8DE546D5621F167918" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph id="225112B205C3030CB1B8E04F1AB65B61" pageId="0" pageNumber="1">
|
||||
<emphasis id="81B5BAF504EC86A19A68727756686E2B" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="48FCC0363F4BDDA714C708CBAB9402D7" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Small species: HL: HL: 1.01-1.07 (1.04); HW: 0.94-1.02 (0.99), WL: 0.75-0.81 (0.78); head in full-face view square, anterior and posterior sides of eyes slightly convex; in lateral view sub-oval; ventral and dorsal faces convex; dorsal face not depressed posteriorly; sides of the head with sparse, long, erect pilosity; frons with dense, thick, and longitudinal rugae, interspaces superficially foveolate; inner hypostomal teeth distinct, approximately the same size, closely spaced, dentate, with rounded apex and wide base; outer hypostomal teeth with base slightly wider than inner hypostomal teeth; gaster indistinctly shagreened; body brown to dark brown.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||||
<emphasis id="B6ACE303E330C2726FFE926A1D3B21C4" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="273AE5F47E25388E5A2681F46347B293" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Head and mesosoma foveolate; promesonotum low, long, slightly convex, with relatively steep posterior declivity; propodeal spines very small.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||||
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||||
<subSubSection id="7EC8640A875A497DDB2FB60BB081CF9A" pageId="0" pageNumber="1" type="description">
|
||||
<paragraph id="37A6CE7F1CAA378E63B8804D89B365D0" pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph id="24EDC3F26EEAC3E66228F15412B853D7" pageId="0" pageNumber="1">
|
||||
<emphasis id="D0BC860E8C5B09E0ECE417884C7933F2" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||||
Measurements (
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
<emphasis id="FDEF0E21725EBF4874730DC25D1ADE12" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
= 10): HL: 1.01-1.07 (1.04); HW: 0.94-1.02 (0.99); SL: 0.43-0.51 (0.48); EL: 0.14-0.15 (0.14); WL: 0.75-0.81 (0.78); PSL: 0.14-0.16 (0.15); MTL: 0.44-0.49 (0.47); PNW: 0.41-0.51 (0.45); PTW: 0.1-0.13 (0.12); PPW: 0.33-0.39 (0.36); CI: 92.8-98.1 (95.1); SI: 46.0-51.0 (48.4); PSLI: 13.0-15.5 (14.2); PPI: 29.0-34.3 (32.3); PNI: 43.5-50.1 (45.9); MTI: 44.9-49.3 (47.1).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
<emphasis id="3D60244B91DE5E261CD18E1C21DA7846" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="B70EE8253BFADA18515F3C66F86A9D26" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
In full-face view square, anterior and posterior sides of eyes slightly convex (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60B</figureCitation>
|
||||
<figureCitation id="3FA36D1DBEE2087163270312E3B9B925" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60B</figureCitation>
|
||||
). In lateral view sub-oval; ventral and dorsal faces convex; dorsal face not depressed posteriorly; inner hypostomal teeth visible. Sides of the head with sparse, long, erect pilosity; whole head with dense, short, suberect to erect pilosity. Antennal scrobes very indistinct and not delimited by carinulae; scrobe surface shiny, finely foveolate with sparse, thick, longitudinal to irregular rugulae in posterior part. Occipital lobes shiny, with fine foveolae and indistinct, sparse, irregular rugae, sculpture weakening posteriorly; frons and malar area with dense, thick, and longitudinal rugae, interspaces superficially foveolate; genae shiny, with fine to indistinct foveolae. Centre of clypeus smooth and shiny, lateral sides with longitudinal rugae; median notch present, narrow, and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, slightly exceeding the midlength of head; pilosity suberect to erect (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60B, D</figureCitation>
|
||||
<figureCitation id="934B67D75504CF9B855CACAAEE7340D0" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60B, D</figureCitation>
|
||||
). Inner and outer hypostomal teeth distinct, approximately the same size, closely spaced, dentate, with rounded apex and wide base; outer hypostomal teeth with base slightly wider than inner hypostomal teeth (Fig.
|
||||
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84H</figureCitation>
|
||||
<figureCitation id="16D26D5D6AE6876C48545B01F3C6CB5E" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84H</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
<emphasis id="D070EFE92A5BB177C7659CCAC5186E2D" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="61C5F888ABE5558A1A58A93FF9C6EEFA" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
In lateral view, promesonotum moderately short, relatively low, and convex, posterior mesonotum steep, with small tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, triangular, with rounded apex; humeral area laterally well produced (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D</figureCitation>
|
||||
<figureCitation id="ACE497F4A974DBCE3F6DB8FD941A8402" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D</figureCitation>
|
||||
). Surface shiny, with fine but distinct foveolae, additional sparse, fine, transverse to irregular rugae on promesonotal dorsum and sometimes propodeum, sculpture slightly weakening on dorsal surface. Pilosity moderately sparse, very long, and erect (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D, F</figureCitation>
|
||||
<figureCitation id="76752C5A0FE133549F118A4A0D09E250" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="C745E4EB56BB70F37ECDDE5A46754BE2" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="07D55F68185CD7C6350B246159F62877" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Shiny and with fine foveolae; peduncle short, with short horizontal lobes on its basal part; node relatively high, triangular, with rounded apex, in rear view node straight or dorsoventrally slightly concave; pilosity moderately sparse and erect (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D, F</figureCitation>
|
||||
<figureCitation id="D0BCD713FE5769B3B3B7966938697521" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
<emphasis id="32EB1FABB8AAB3C10BDAC35394CBFFFC" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="CB22926160548730E0B2E06CA322F699" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Shiny, with fine and sparse foveolae, smooth on dorsum; in dorsal view sides with moderately short, acute, and triangular projections; pilosity moderately long, dense, and erect (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D, F</figureCitation>
|
||||
<figureCitation id="4C3425F55830B5905CEF4480CA74C6FC" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="4BDF26527933B351210D1D5B8709B44C" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="064A636C197F3983CDF7C812608CC987" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Shiny and indistinctly shagreened; pilosity moderately dense, short, and erect (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D, F</figureCitation>
|
||||
<figureCitation id="7C913A3747D69CC54368339605B205D4" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D, F</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
<emphasis id="CAC88DEA675D949A65BF5C635583B75A" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="D277A2F531D69B9D43313F485C75FAA3" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Unicolourous, brown to dark brown; legs yellow (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D, F</figureCitation>
|
||||
<figureCitation id="5E77D05608A4E64DBC9598A011E53DF3" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60D, F</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1" start="Figure 60" startId="F60">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 60.</emphasis>
|
||||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole bemarivoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemarivoensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole bemarivoensis</emphasis>
|
||||
<caption id="29C583F53C131E5951365B975BC483EC" doi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1" start="Figure 60" startId="F60">
|
||||
<paragraph id="B5A0273667412B756A2F647EF0A87923" pageId="0" pageNumber="1">
|
||||
<emphasis id="90DEF714E6B08835230A97F61E7B959E" bold="true" pageId="0" pageNumber="1">Figure 60.</emphasis>
|
||||
<taxonomicName id="037A9C3002CF3A66C5E1739FBB7DBDFF" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole bemarivoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemarivoensis">
|
||||
<emphasis id="BE7017A69CFE63B22B3A23C23D4B82BB" italics="true" pageId="0" pageNumber="1">Pheidole bemarivoensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., full-face view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<emphasis id="30A11C0A45BE7F1EEF34047419D2056C" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
), profile (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<emphasis id="50F0A028F2CFBF60F6A8FA17B8A1AA9D" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
), and dorsal view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<emphasis id="C1611F326DB2E8697BB1954DF8A76E6F" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
) of paratype minor worker (CASENT0923160) and full-face view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<emphasis id="3CDCFD442954EF642955930E203D3C3C" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
), profile (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<emphasis id="4A3707885AFC4ACF0612CC7539ED7240" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
), and dorsal view (
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
<emphasis id="4212277FE68C25030EB07B71847BEDD1" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||||
) of holotype major worker (CASENT0489439).
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||||
<paragraph id="B76D232D9B35FCB515A9DF2DA2A17DB8" pageId="0" pageNumber="1">
|
||||
<emphasis id="A3B64F9FAAABF00A430AD23DB4C704BA" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||||
Measurements (
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
<emphasis id="3F709E6CA75C43D15D0C5BCCCD31FFB4" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||||
= 10): HL: 0.43-0.49 (0.47); HW: 0.39-0.44 (0.42); SL: 0.4-0.45 (0.42); EL: 0.09-0.11 (0.1); WL: 0.5-0.58 (0.54); PSL: 0.07-0.1 (0.09); MTL: 0.29-0.35 (0.32); PNW: 0.25-0.29 (0.27); PTW: 0.06-0.08 (0.07); PPW: 0.11-0.14 (0.12); CI: 85.9-92.1 (89.8); SI: 96.5-105.9 (100.3); PSLI: 15.2-20.1 (18.1); PPI: 51.3-62.2 (56.9); PNI: 62.4-67.6 (64.3); MTI: 73.3-79.5 (77.1).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
<emphasis id="C0BE1A1BC02E760115D76469FD6C73E0" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="5162B8A462B94FA0F7167E2D91FE831A" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Occipital margin straight or indistinctly concave; occipital carina absent (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60A</figureCitation>
|
||||
<figureCitation id="6E096661E719F78C285C1F4CEF8F0545" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60A</figureCitation>
|
||||
). Pilosity moderately sparse, long, suberect to erect. Head foveolate. Clypeus with fine and sometimes reduced foveolae; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity suberect to erect (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60A, C</figureCitation>
|
||||
<figureCitation id="88EF4F2D6AB1FCF4397DC557EB168F2A" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60A, C</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
<emphasis id="3A32B6928998AD5D4E5E36BF64D34061" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="8064232C3875CF8C85DE7F5964028B2E" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
In lateral view, promesonotum low, long, slightly convex, with relatively steep posterior declivity; promesonotal groove absent; metanotal groove indistinct; propodeal spines very small, triangular, apex acute (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C</figureCitation>
|
||||
<figureCitation id="942C3CB59B0609BA39D453F4EFA1EA5B" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C</figureCitation>
|
||||
). Sculpture foveolate. Pilosity sparse, moderately long, and erect (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C, E</figureCitation>
|
||||
<figureCitation id="9A0D7F5C559DCECCC253405366BD2278" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="0DA0D2558250AB9DBAA061FB290D3D11" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="19D522CB249F327B629079BFF6D6E5B4" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Shiny and with fine foveolae; peduncle short and thin with ventral face slightly convex; node moderately high, triangular, and small; with few short, erect setae (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C, E</figureCitation>
|
||||
<figureCitation id="320F5F9BB0CB90639B49CCFE8D6931F1" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
<emphasis id="282A958BFE330CA62A7EA6E4ABAB268E" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="ABBC76D9C65D510C63BD4936D8767DB0" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Moderately long, low, and slightly convex; with few short, erect setae (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C, E</figureCitation>
|
||||
<figureCitation id="97284C5AA4905D86135CC37EFACF4AAE" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
<emphasis id="AF512989641778178B14768A123719C9" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="830140374D9A3E86FC6701A223921306" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
With sparse, erect pilosity (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C, E</figureCitation>
|
||||
<figureCitation id="E7535BC10E2A3CEDE154F5EBBD6E78DB" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C, E</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
<emphasis id="26B4DEF4C4E511570FFFF3CB4CAB9477" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="58501B18BE4A31DF3A42C55CD8BBC2D1" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
Unicolourous, yellow (Fig.
|
||||
<figureCitation captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C, E</figureCitation>
|
||||
<figureCitation id="7DF979B466AEE01B065BA024A50DDEC4" captionStart="Figure 60" captionStartId="F60" captionText="Figure 60. Pheidole bemarivoensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923160) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0489439)." figureDoi="10.3897/zookeys.905.39592.figure60" httpUri="https://binary.pensoft.net/fig/373107" pageId="0" pageNumber="1">60C, E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
|
||||
<subSubSection id="F6D9BE070769B1BCA22DF7B8A359B5E6" pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph id="8D839660F4A35BA4AEB731FE98D58EE1" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph id="E83BD0B1EBDF19F6A5C51D22568C0979" pageId="0" pageNumber="1">From the type locality.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">The species was collected between 30-600 m in elevation, in rainforest, tropical dry forest, and disturbed dry forest. Nests were located in rotten logs, branches on ground, and soil.</paragraph>
|
||||
<subSubSection id="6A76C91485A424C268073E846B67C98E" pageId="0" pageNumber="1" type="biology_ecology">
|
||||
<paragraph id="A6FD142AC6F31E6175F1F12F44A26FB5" pageId="0" pageNumber="1">Biology.</paragraph>
|
||||
<paragraph id="5A2AFA7811574DAA7236D199C7A75B4C" pageId="0" pageNumber="1">The species was collected between 30-600 m in elevation, in rainforest, tropical dry forest, and disturbed dry forest. Nests were located in rotten logs, branches on ground, and soil.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="comments">
|
||||
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole bemarivoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemarivoensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole bemarivoensis</emphasis>
|
||||
<subSubSection id="84251530EAC7C381F0A923C20EA89310" pageId="0" pageNumber="1" type="comments">
|
||||
<paragraph id="A32D401458AB407540DFB7F0175BFAA4" pageId="0" pageNumber="1">Comments.</paragraph>
|
||||
<paragraph id="3E3B1F53D3ABCC6FD7D9519DE1B9AC97" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="A717919611D1A94FBF4281A3ECF6901A" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole bemarivoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemarivoensis">
|
||||
<emphasis id="E892AB952C3877A53893BB57B2ADE9C1" italics="true" pageId="0" pageNumber="1">Pheidole bemarivoensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. is most similar to
|
||||
<taxonomicName lsidName="P. bemarahaensis" pageId="0" pageNumber="1" rank="species" species="bemarahaensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">P. bemarahaensis</emphasis>
|
||||
<taxonomicName id="7F2099AEF106C38C135F8EF78ACBB3B1" lsidName="P. bemarahaensis" pageId="0" pageNumber="1" rank="species" species="bemarahaensis">
|
||||
<emphasis id="B6D32BF9884AA94BCB3C232FA2E3070E" italics="true" pageId="0" pageNumber="1">P. bemarahaensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||||
<emphasis id="8B00A63A518077464FEA0B6617FD7FDB" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="3B6C95B70B108280297E6E00DC65DFD5" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole bemarivoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemarivoensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole bemarivoensis</emphasis>
|
||||
<taxonomicName id="5958FF1D5CF5657A6CBA5E40DE0060E1" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole bemarivoensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bemarivoensis">
|
||||
<emphasis id="7728E45806D337CDB965B991C18E9B2D" italics="true" pageId="0" pageNumber="1">Pheidole bemarivoensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. differs from
|
||||
<taxonomicName lsidName="P. bemarahaensis" pageId="0" pageNumber="1" rank="species" species="bemarahaensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">P. bemarahaensis</emphasis>
|
||||
<taxonomicName id="A6C0F031C0FEF810AD13F0CE04BEA842" lsidName="P. bemarahaensis" pageId="0" pageNumber="1" rank="species" species="bemarahaensis">
|
||||
<emphasis id="EA4D8F78F58CD67DB71ADD214A73CAC7" italics="true" pageId="0" pageNumber="1">P. bemarahaensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. in foveolae covering the whole surface of frons, and weaker sculpture on genae and dorsal surface of promesonotum.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||||
<emphasis id="AD193FD6E100B1E4A5D8EC7C5EA88249" bold="true" pageId="0" pageNumber="1">
|
||||
<emphasis id="E19D82F11627FBC7ED7ECA6E1F4CBE4D" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<taxonomicName lsidName="P. bemarivoensis" pageId="0" pageNumber="1" rank="species" species="bemarivoensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">P. bemarivoensis</emphasis>
|
||||
<taxonomicName id="756E5DC8FD2FA545061B37619DED1AE6" lsidName="P. bemarivoensis" pageId="0" pageNumber="1" rank="species" species="bemarivoensis">
|
||||
<emphasis id="15DE697BDB3D631F4B5D1F0B1AA238FA" italics="true" pageId="0" pageNumber="1">P. bemarivoensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. differs from
|
||||
<taxonomicName lsidName="P. bemarivoensis" pageId="0" pageNumber="1" rank="species" species="bemarivoensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">P. bemarivoensis</emphasis>
|
||||
<taxonomicName id="14D29F7BE2071B618C96EDB314741017" lsidName="P. bemarivoensis" pageId="0" pageNumber="1" rank="species" species="bemarivoensis">
|
||||
<emphasis id="41F813EDC4CB2BC9F016FF811C70803C" italics="true" pageId="0" pageNumber="1">P. bemarivoensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. in minute and short propodeal spines.
|
||||
</paragraph>
|
||||
|
|
|
|||
Loading…
Add table
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Reference in a new issue