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<treatment ID-GBIF-Taxon="161776999" LSID="urn:lsid:zoobank.org:act:C9CB164B-0A45-49AE-99AB-9AB732539425" httpUri="http://treatment.plazi.org/id/16346E3D69641A7BF5785C4940299067" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/C9CB164B-0A45-49AE-99AB-9AB732539425" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole makirovana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makirovana" status="sp. nov.">Pheidole makirovana</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">Figs 54A-F</figureCitation>
<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85E</figureCitation>
<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87I</figureCitation>
<treatment id="16346E3D69641A7BF5785C4940299067" ID-GBIF-Taxon="161776999" LSID="urn:lsid:zoobank.org:act:C9CB164B-0A45-49AE-99AB-9AB732539425" httpUri="http://treatment.plazi.org/id/16346E3D69641A7BF5785C4940299067" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="9A0C719170570A8E05DB9D666BFE88C5" pageId="0" pageNumber="1" type="nomenclature">
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<taxonomicName id="33F71CBC94E04605AE553FA755857E19" ID-CoL="CMLFW" LSID="http://zoobank.org/C9CB164B-0A45-49AE-99AB-9AB732539425" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole makirovana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makirovana" status="sp. nov.">Pheidole makirovana</taxonomicName>
<taxonomicNameLabel id="C8ABC5CFFBE2F9D700A6854A6355E238" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="B496907857397E62DDED9964FFB2BAB8" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">Figs 54A-F</figureCitation>
<figureCitation id="05F2FB84BF0A5C131520331DA8A3BB5C" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85E</figureCitation>
<figureCitation id="50B44C2CECA8C6FFE7699D8B1A43B30E" captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87I</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
<subSubSection id="C3548DB6B8D612885E21BC8B51FCDA63" pageId="0" pageNumber="1" type="type material">
<paragraph id="F77924175BE93EFEF709E9182C55BFBD" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="24FC35D10225A7FE30949608752B0192" pageId="0" pageNumber="1">
<emphasis id="2DC895180A2F70C203EE7DE8FC55CDC9" bold="true" pageId="0" pageNumber="1">
<emphasis id="7966EBCE5DC7F8ED9EB9AA0EBEE55E3B" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Madagascar. •1 major worker; Antsiranana; Makirovana Forest;
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<geoCoordinate id="61A6AA1EE0445EFA2C986811A02D72F3" degrees="14.16506" direction="south" orientation="latitude" precision="1" value="-14.16506">-14.16506</geoCoordinate>
,
<geoCoordinate degrees="49.9477" direction="east" orientation="longitude" precision="5" value="49.9477">49.9477</geoCoordinate>
<geoCoordinate id="7A25102BB91998FF4539C6B323F18B0A" degrees="49.9477" direction="east" orientation="longitude" precision="5" value="49.9477">49.9477</geoCoordinate>
; alt. 900 m; 30 Apr 2011; B.L. Fisher et al. leg.; BLF26745, CASENT0231059 (CASC).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
<emphasis id="EF5FD8F3CF7E66BFC57FB2C89751E1EE" bold="true" pageId="0" pageNumber="1">
<emphasis id="BFCDC7BDA5F21A0027FAA0C34C2DB8D9" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
.
</emphasis>
Madagascar. •1w.; same data as for holotype; CASENT0923174 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="08C9C20C476C91995EDB8E6CBD3B7773" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="BD82049143674CEC45015EEC2D8C3F6C" pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph id="831CAE1942EA65C5E22BA34C48E4B099" pageId="0" pageNumber="1">
Madagascar. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
<emphasis id="5E0DDF7189B38E884E0C23F0A1CD2D58" bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
: •2w., 1s., 2q.; 6.5 km SSW Befingotra,
<normalizedToken originalValue="Rés">Res</normalizedToken>
<normalizedToken id="EFBAA2ABECA070C0EA672C34A63012AA" originalValue="Rés">Res</normalizedToken>
. Anjanaharibe-Sud; -14.75, 49.5; alt. 875 m; 23 Oct 1994; B.L. Fisher leg.; CASENT0198415, CASENT0198572, CASENT0235044, CASENT0235045 (CASC). •8w., 4s., 1q.; 9.2 km WSW Befingotra,
<normalizedToken originalValue="Rés">Res</normalizedToken>
<normalizedToken id="9584DA973CFDF5C6B5FB938434D96BAB" originalValue="Rés">Res</normalizedToken>
. Anjanaharibe-Sud; -14.75, 49.46667; alt. 1200 m; 11 Nov 1994; B.L. Fisher leg.; CASENT0198570, CASENT0198571, CASENT0198574, CASENT0235107 (CASC). •2w., 2q.; Makirovana Forest;
<geoCoordinate degrees="14.16506" direction="south" orientation="latitude" precision="1" value="-14.16506">-14.16506</geoCoordinate>
<geoCoordinate id="23B2F9284D78A396A9660666AB221733" degrees="14.16506" direction="south" orientation="latitude" precision="1" value="-14.16506">-14.16506</geoCoordinate>
,
<geoCoordinate degrees="49.9477" direction="east" orientation="longitude" precision="5" value="49.9477">49.9477</geoCoordinate>
<geoCoordinate id="B87601D7CED1FF78465D3CD06E4CF83D" degrees="49.9477" direction="east" orientation="longitude" precision="5" value="49.9477">49.9477</geoCoordinate>
; alt. 900 m; 30 Apr 2011; B.L. Fisher et al. leg.; CASENT0231036, CASENT0231037, CASENT0923175 (CASC). •6w., 3s.; Parc National de Marojejy, Manantenina River, 27.6 km 35°NE Andapa, 9.6 km 327°NNW Manantenina; -14.435, 49.76; alt. 775 m; 15 Nov 2003; B.L. Fisher et al. leg.; CASENT0495015, CASENT0495016, CASENT0495019 (CASC). •3w.; Parc National de Marojejy, Manantenina River, 28.0 km 38°NE Andapa, 8.2 km 333°NNW Manantenina;
<geoCoordinate degrees="14.43667" direction="south" orientation="latitude" precision="1" value="-14.43667">-14.43667</geoCoordinate>
<geoCoordinate id="1CFF12F18F0DB9816AF3D924B26ABE01" degrees="14.43667" direction="south" orientation="latitude" precision="1" value="-14.43667">-14.43667</geoCoordinate>
,
<geoCoordinate degrees="49.775" direction="east" orientation="longitude" precision="55" value="49.775">49.775</geoCoordinate>
<geoCoordinate id="8ED75DBC13DCD58032F1328EC9263B67" degrees="49.775" direction="east" orientation="longitude" precision="55" value="49.775">49.775</geoCoordinate>
; alt. 450 m; 12 Nov 2003; B.L. Fisher et al. leg.; CASENT0077212 (CASC). -
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mahajanga</emphasis>
<emphasis id="A2EEF7E7D7B7E46471CDF01C68AFB701" bold="true" italics="true" pageId="0" pageNumber="1">Mahajanga</emphasis>
: •1w., 1s.;
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken id="E333FA8A0BBAA2884C6EBA29E3932C12" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="A8DEC6D0F3C6CC8F1B939202F386FADB" originalValue="Spéciale">Speciale</normalizedToken>
Marotandrano, Marotandrano 48.3 km S Mandritsara;
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<geoCoordinate id="5081C3027CE203A719F8B70D15751261" degrees="16.28322" direction="south" orientation="latitude" precision="1" value="-16.28322">-16.28322</geoCoordinate>
,
<geoCoordinate degrees="48.81443" direction="east" orientation="longitude" precision="1" value="48.81443">48.81443</geoCoordinate>
<geoCoordinate id="A848444E8F4D5B90EE712F3AAF436862" degrees="48.81443" direction="east" orientation="longitude" precision="1" value="48.81443">48.81443</geoCoordinate>
; alt. 865 m; 7 Dec 2007; B.L. Fisher et al. leg.; CASENT0134300 (CASC). -
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
<emphasis id="01F93778687937AE5906443C2C678C48" bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
: •2w., 2s.; F.C. Didy;
<geoCoordinate degrees="18.19833" direction="south" orientation="latitude" precision="1" value="-18.19833">-18.19833</geoCoordinate>
<geoCoordinate id="3F5B7294ABC87D0C3930D2D7174CAD49" degrees="18.19833" direction="south" orientation="latitude" precision="1" value="-18.19833">-18.19833</geoCoordinate>
,
<geoCoordinate degrees="48.57833" direction="east" orientation="longitude" precision="1" value="48.57833">48.57833</geoCoordinate>
<geoCoordinate id="4F1A48C2EDDC843E52CAD921CA50FFB9" degrees="48.57833" direction="east" orientation="longitude" precision="1" value="48.57833">48.57833</geoCoordinate>
; alt. 960 m; 16 Dec 1998; Ratsirarson leg.; CASENT0198558, CASENT0198898 (CASC). •5w., 2s.; Montagne
<normalizedToken originalValue="dAkirindro">d'Akirindro</normalizedToken>
<normalizedToken id="8100D0FE11DCDE2682D90E4301943C37" originalValue="dAkirindro">d'Akirindro</normalizedToken>
7.6 km 341°NNW Ambinanitelo;
<geoCoordinate degrees="15.28833" direction="south" orientation="latitude" precision="1" value="-15.28833">-15.28833</geoCoordinate>
<geoCoordinate id="329683CAB634B3B8CBBDC73469224356" degrees="15.28833" direction="south" orientation="latitude" precision="1" value="-15.28833">-15.28833</geoCoordinate>
,
<geoCoordinate degrees="49.54833" direction="east" orientation="longitude" precision="1" value="49.54833">49.54833</geoCoordinate>
<geoCoordinate id="E2A65525EB5A949080EDFE11996B6E68" degrees="49.54833" direction="east" orientation="longitude" precision="1" value="49.54833">49.54833</geoCoordinate>
; alt. 600 m; 17 Mar 2003; B.L. Fisher et al. leg.; CASENT0038919, CASENT0496319, CASENT0496320 (CASC). •2s.; Montagne
<normalizedToken originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
<normalizedToken id="D05DB3B698B54068997002DD63466975" originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
, 19.5 km 27°NNE Ambinanitelo;
<geoCoordinate degrees="15.17833" direction="south" orientation="latitude" precision="1" value="-15.17833">-15.17833</geoCoordinate>
<geoCoordinate id="9B163FEA4ECF91B4B21EF106B904EDD6" degrees="15.17833" direction="south" orientation="latitude" precision="1" value="-15.17833">-15.17833</geoCoordinate>
,
<geoCoordinate degrees="49.635" direction="east" orientation="longitude" precision="55" value="49.635">49.635</geoCoordinate>
<geoCoordinate id="67CA0BED5EA2C0E3B461FC8AFC433F75" degrees="49.635" direction="east" orientation="longitude" precision="55" value="49.635">49.635</geoCoordinate>
; alt. 1100 m; 12 Mar 2003; B.L. Fisher et al. leg.; CASENT0038295, CASENT0038311 (CASC). •3w., 1s., 1m.; Parc National de Zahamena, Tetezambatana forest, near junction of Nosivola and Manakambahiny Rivers;
<geoCoordinate degrees="17.74298" direction="south" orientation="latitude" precision="1" value="-17.74298">-17.74298</geoCoordinate>
<geoCoordinate id="A0DEFD865DEFA11C66A50CD395B10C28" degrees="17.74298" direction="south" orientation="latitude" precision="1" value="-17.74298">-17.74298</geoCoordinate>
,
<geoCoordinate degrees="48.72936" direction="east" orientation="longitude" precision="1" value="48.72936">48.72936</geoCoordinate>
<geoCoordinate id="6D24EBFB48DA136962DC22AC13E61C51" degrees="48.72936" direction="east" orientation="longitude" precision="1" value="48.72936">48.72936</geoCoordinate>
; alt. 860 m; 18 Feb 2009; B.L. Fisher et al. leg.; CASENT0150555, CASENT0150556, CASENT0151067, CASENT0235021 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
<subSubSection id="442C89CD20B5CC1BEA79088529820B06" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="717079635286608FF08B325010029446" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="0C6572A17F9F3D55F3E7CFE6437C04EC" pageId="0" pageNumber="1">
<emphasis id="EE1BAA8ED3C5ED1F3D614FDDC959E96C" bold="true" pageId="0" pageNumber="1">
<emphasis id="7D7F5AD7758EEB3C414C6D661F006FFF" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
Head in full face view rectangular, slightly widening posteriorly; sides of the head with moderately dense, long, erect pilosity; frons with thick, sparse to moderately sparse, longitudinal rugae, interspaces smooth or with indistinct but dense rugulae; inner hypostomal teeth distinct, low, thick and triangular, with rounded apex; outer hypostomal teeth higher and slightly wider than inner hypostomal teeth, thick, with rounded tips, and wide base; inner and outer teeth closely spaced and connected by indistinct concavity; propodeal spines moderately long, narrow, with base slightly wider than top; first gastral tergite finely shagreened.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
<emphasis id="B25AB84A2AF76D28357CAB8F75358978" bold="true" pageId="0" pageNumber="1">
<emphasis id="827EF4D9DED12012601EFC9B5661D9E5" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
Whole head foveolate; thick, sparse, longitudinal to irregular rugae overlie foveolate sculpture on frons and malar area, upper part of frons with rugae curved outward; sparse, irregular to arcuate, thick rugae overlie foveolate sculpture on vertex and genae; promesonotum low, short, flat or slightly convex, with steep posterior declivity; mesosoma foveolate, with sparse, thick and irregular rugae overlying foveolae; anepisternum, katepisternum, and mesosoma smooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
<subSubSection id="FF8FEA0712B45A6838E9D6F59B7EBEAC" pageId="0" pageNumber="1" type="description">
<paragraph id="23DBFBEF5B4386937244E08889A408EC" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="CD35E0561EF6396B4466E88B70884C49" pageId="0" pageNumber="1">
<emphasis id="ACC23862F1C57C08E0921F06285ACB6E" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
<emphasis id="032456ED12584E08C6F458A05BDC1CF7" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 10): HL: 1.7-1.92 (1.8); HW: 1.57-1.7 (1.62); SL: 0.64-0.71 (0.68); EL: 0.18-0.2 (0.19); WL: 1.09-1.27 (1.18); PSL: 0.24-0.27 (0.26); MTL: 0.68-0.73 (0.7); PNW: 0.77-0.8 (0.78); PTW: 0.23-0.28 (0.25); PPW: 0.62-0.73 (0.68); CI: 87.2-95.0 (90.1); SI: 39.0-44.0 (41.7); PSLI: 13.4-15.9 (14.4); PPI: 32.4-39.7 (36.2); PNI: 46.4-50.0 (48.2); MTI: 41.1-44.4 (43.4).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
<emphasis id="D88F209CBF984A67D5BDB46FA19D9C7E" bold="true" pageId="0" pageNumber="1">
<emphasis id="008EAE1C57CBA6744C7A28F605A99D03" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
In full-face view rectangular, slightly widening posteriorly, anterior of eyes relatively straight, posterior of eyes slightly convex (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54B</figureCitation>
<figureCitation id="A1993C3A749A7796AADC546DCB2E1AF3" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54B</figureCitation>
). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming shallow transverse depression between frons and occipital lobes; inner hypostomal teeth invisible. Sides of the head with moderately dense, long, erect pilosity; whole head with dense, moderately long, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae. Occipital lobes shiny, with thick, sparse, irregular rugae, interspaces smooth to finely rugulose; frons with thick, sparse to moderately sparse, longitudinal rugae, interspaces smooth or with indistinct but dense rugulae; lateral sides of head with thin, dense, irregularly rugoreticulate; malar area with dense and thin longitudinal rugulae; genae with fine, thin, dense rugulae. Clypeus shiny and smooth, with thin, longitudinal rugulae on lateral sides; median notch present, narrow and moderately deep; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, reaching midlength of head; pilosity decumbent to erect (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54B, D</figureCitation>
<figureCitation id="28C800B981A149E1465B6C0FDBD15D70" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54B, D</figureCitation>
). Inner hypostomal teeth distinct, low, thick, and triangular, with rounded apex; outer hypostomal teeth higher and slightly wider than inner hypostomal teeth, thick, with rounded tips, and wide base; inner and outer teeth closely spaced and connected by indistinct concavity (Fig.
<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85E</figureCitation>
<figureCitation id="8A33AE119D6C96AFAE684D0890BC8FBE" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
<emphasis id="6ED5CB82EF0697B87B2381343FA4DCDA" bold="true" pageId="0" pageNumber="1">
<emphasis id="78FF8A26FC784868EEC380E23C4800A2" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum short, angular, and relatively low, posterior mesonotum steep, with moderately large, tubercle-like projections; promesonotal groove absent; metanotal groove absent or indistinct; propodeal spines moderately long, narrow, with wide base, apex rounded; humeral area with small and flat tubercles (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D</figureCitation>
<figureCitation id="18B46AECF8D92466E6289A2BCD34932F" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D</figureCitation>
). Surface shiny, pronotum, mesonotum, anepisternum, and katepisternum smooth, with very sparse, thick to thin, irregular rugae, lateral sides of propodeum with few thick, longitudinal rugae; sometimes surface with additional indistinct and sparse rugulae on promesonotum. Pilosity moderately sparse, long, and erect (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D, F</figureCitation>
<figureCitation id="817ADC9D72DB5789F140ADABE8B58C2A" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="3373F46B46E7A0BA10395FA46D5C9D3F" bold="true" pageId="0" pageNumber="1">
<emphasis id="31D486983A361E727EB073CAEF760C0C" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny, finely shagreened to smooth; peduncle short, with small, rounded horizontal lobes on its basal part; node moderately high and narrow, with convex apex, in rear view node relatively straight; pilosity moderately dense and erect (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D, F</figureCitation>
<figureCitation id="710530332E8C98DA2A4BC95A2358A51A" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
<emphasis id="3F06560F7060A2BCEE76D42F62613ACB" bold="true" pageId="0" pageNumber="1">
<emphasis id="68DCB85F3BBA17C9732F7EE7C713E443" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Shiny and finely shagreened; short and rounded; in dorsal view sides with relatively long, acute, triangular projections; pilosity long, moderately dense and erect (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D, F</figureCitation>
<figureCitation id="A82EFD9835FD13E8991B196AE2ABD9BB" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="7449829945627C85B149564D1D0E5CBF" bold="true" pageId="0" pageNumber="1">
<emphasis id="AA34D124E0F79A285B6860EB65FF9452" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
First gastral tergite shiny and finely shagreened; pilosity dense, long and erect (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D, F</figureCitation>
<figureCitation id="9F9D5932AE3F0DB7C94BC530FCECA5EB" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
<emphasis id="1CAA6CB203E3A23753033972BAC39DCE" bold="true" pageId="0" pageNumber="1">
<emphasis id="691108ACB30918F3BAF359C977A8EA31" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, brown to dark brown; legs dark yellow (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D, F</figureCitation>
<figureCitation id="10F6A9F04B3D13473D910BC5DC784631" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54D, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1" start="Figure 54" startId="F54">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 54.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole makirovana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makirovana">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole makirovana</emphasis>
<caption id="CCC089DB71951C04235D428369440390" doi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1" start="Figure 54" startId="F54">
<paragraph id="467146B6E708AA948CD290882E69F28B" pageId="0" pageNumber="1">
<emphasis id="2A8797A740102B28F0119B6E4AFB5CDD" bold="true" pageId="0" pageNumber="1">Figure 54.</emphasis>
<taxonomicName id="8353B75E52DB6F6BA53EF01ACAF05A21" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole makirovana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makirovana">
<emphasis id="78A0F747CB9583A6367D663DC99EACB6" italics="true" pageId="0" pageNumber="1">Pheidole makirovana</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<emphasis id="BFB89D272749B25BB0B56A732F73E222" bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<emphasis id="FD90DDE81818DCF680704A80DB4E04AE" bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<emphasis id="CEC694CFE3BB5987F8B22D3D307445D9" bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0923174) and full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<emphasis id="217E76AB274164C501E5CDCC46B61486" bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<emphasis id="10D0DC2418A9AE846DBA5732BA6562C8" bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<emphasis id="18D93C0D0514FF5DACDEF1073436FF92" bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0231059).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
<paragraph id="D4CA1CDADC1C90578A670CC2FD524F49" pageId="0" pageNumber="1">
<emphasis id="B762FC0D0DF246ABD056B7345D647AD8" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
<emphasis id="264FDDD054338ABEF4EE58FEB695C621" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 10): HL: 0.57-0.62 (0.59); HW: 0.53-0.57 (0.54); SL: 0.56-0.61 (0.59); EL: 0.11-0.12 (0.11); WL: 0.69-0.79 (0.75); PSL: 0.11-0.14 (0.12); MTL: 0.44-0.48 (0.47); PNW: 0.36-0.4 (0.37); PTW: 0.06-0.09 (0.07); PPW: 0.11-0.13 (0.12); CI: 88.5-94.6 (91.8); SI: 103.2-112.8 (107.8); PSLI: 17.9-23.6 (21.0); PPI: 54.6-67.2 (61.4); PNI: 66.8-71.8 (69.2); MTI: 80.0-90.7 (86.0).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
<emphasis id="2151922557845EEE74C9200011C33FE4" bold="true" pageId="0" pageNumber="1">
<emphasis id="2632404D8665AFCB49E6C319137B666F" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Occipital margin straight or indistinctly concave; occipital carina indistinct, weakly developed (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54A</figureCitation>
<figureCitation id="52E2745DDBD694C8041770BB205DADC2" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54A</figureCitation>
). Pilosity moderately dense, long, erect. Whole head foveolate; thick, sparse, longitudinal to irregular rugae overlie foveolate sculpture on frons and malar area, upper part of frons with rugae curved outward; sparse, irregular to arcuate, thick rugae overlie foveolate sculpture on vertex and genae. Clypeus with median longitudinal carina present; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity erect (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54A, C</figureCitation>
<figureCitation id="DF4ABE27C6B05D9F638E8019CE140CEB" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54A, C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
<emphasis id="E8A8EBD4DCD6D5D8FA70EC049BFFD9CB" bold="true" pageId="0" pageNumber="1">
<emphasis id="30870DED87A31D3446A1B5BA55224384" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum low, short, flat or slightly convex, with steep posterior declivity; promesonotal groove absent; metanotal groove absent or indistinct; propodeal spines moderately long, triangular, apex acute (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C</figureCitation>
<figureCitation id="E1A4BDB66130D0F2E9450D2B6B1EAD46" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C</figureCitation>
). Sculpture foveolate, with sparse, thick, and irregular rugae overlying foveolae; anepisternum, katepisternum, and mesosoma smooth. Pilosity sparse, long, and erect (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C, E</figureCitation>
<figureCitation id="A54F91F68C5707341D9F838E102C0441" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="542249B2EB99CAE4553C9C42D45EECDC" bold="true" pageId="0" pageNumber="1">
<emphasis id="5516F76BBD415BF4EBABC74810B44C8A" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Peduncle moderately long and thin; node moderately high, triangular, and small; with few long, erect setae (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C, E</figureCitation>
<figureCitation id="4E5362DACD7374D438252FE878B646E1" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
<emphasis id="2340EA4720F13B73DFDFD13EC19A3DB4" bold="true" pageId="0" pageNumber="1">
<emphasis id="5BB267F8B5DD512E2FF8F1C56C44C570" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Short, low, and convex; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C, E</figureCitation>
<figureCitation id="F8450347B9AEAA79722D7E226593C7EA" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="682C60605E24742BA3D4BABA5EBF415F" bold="true" pageId="0" pageNumber="1">
<emphasis id="CD1B54CC48FA51BEB2BD6AE775D61C67" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
With sparse, erect pilosity (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C, E</figureCitation>
<figureCitation id="FA0822277495ACE36800DBC40489B301" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
<emphasis id="A1E33BBB39FAA43474D637E831DC6408" bold="true" pageId="0" pageNumber="1">
<emphasis id="FB96C6DE26BC63FFA9F92897316B7DBD" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, yellowish brown to brown; legs yellow (Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C, E</figureCitation>
<figureCitation id="B8D6594633C159573E441030F1537728" captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Pheidole makirovana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923174) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0231059)." figureDoi="10.3897/zookeys.905.39592.figure54" httpUri="https://binary.pensoft.net/fig/373101" pageId="0" pageNumber="1">54C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
<subSubSection id="8DA3F3B4EBFABCF52D7518E655F61380" pageId="0" pageNumber="1" type="etymology">
<paragraph id="AE7DDAAC147C6898244CEA08092893E0" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="9BAEBA13718BC6DE237A156A507E8CFA" pageId="0" pageNumber="1">From the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was collected between 450-1200 m in elevation, in transitional humid forest, rainforest, and montane rainforest. Nests were located in rotten logs and rotten twigs on the ground.</paragraph>
<subSubSection id="482973F2568BF2DCFF1335B054C04F8A" pageId="0" pageNumber="1" type="biology_ecology">
<paragraph id="2AE315B0C8EBC9DC246C458FC2C813B4" pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph id="11ADB2892073C70D9CA0EF48B44D40A3" pageId="0" pageNumber="1">The species was collected between 450-1200 m in elevation, in transitional humid forest, rainforest, and montane rainforest. Nests were located in rotten logs and rotten twigs on the ground.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole makirovana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makirovana">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole makirovana</emphasis>
<subSubSection id="9CDC4D0E399AFE15162A72E25D513AF9" pageId="0" pageNumber="1" type="comments">
<paragraph id="23628CF080B96EC1DC4A48FA114B1DFE" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="5F1DB975574F39480723D72AD138A492" pageId="0" pageNumber="1">
<taxonomicName id="C064BB5C3D885FEB477DEA3959D78E56" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole makirovana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makirovana">
<emphasis id="A96B543F96A6EFF0DBC0624CBA21504B" italics="true" pageId="0" pageNumber="1">Pheidole makirovana</emphasis>
</taxonomicName>
sp. nov. is most similar to
<taxonomicName lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
<taxonomicName id="10FA7F2E2B76F2D9103AEAFC709DBD42" lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
<emphasis id="A265721C6B0A2A8BD8E78526713D8139" italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
<emphasis id="6D49FFBE1F5F100D3D11B5DB1ABE1AD0" bold="true" pageId="0" pageNumber="1">
<emphasis id="7052C48865DE4DC5F12DE6DCE5CE975C" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole makirovana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makirovana">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole makirovana</emphasis>
<taxonomicName id="7DAD250A25DF5078329DC03FA87B8FB2" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole makirovana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makirovana">
<emphasis id="A1A9A9416F89AEBE4C37F4FB864A8372" italics="true" pageId="0" pageNumber="1">Pheidole makirovana</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
<taxonomicName id="699380364547361B2AADA36802964397" lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
<emphasis id="C61DE633138DD80CACB0A047BA0C279F" italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
</taxonomicName>
sp. nov. by pronotum, mesonotum, anepisternum, and katepisternum with smooth notches and outer hypostomal teeth wider than inner hypostomal teeth.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
<emphasis id="13957E0369E7CC32F0FE4F638F4938BA" bold="true" pageId="0" pageNumber="1">
<emphasis id="56D2D352289C61B870DB17162EA2C4E0" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole makirovana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makirovana">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole makirovana</emphasis>
<taxonomicName id="49E75F842CDCDAB5D421215BE6CDBD59" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole makirovana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makirovana">
<emphasis id="184DE9185FEEECEE951865FF3C56F4F6" italics="true" pageId="0" pageNumber="1">Pheidole makirovana</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
<taxonomicName id="944A5E1AEC48431CCA97A98C6AC26DD9" lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
<emphasis id="244288F9FDCF082881C1F0592D546C0B" italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
</taxonomicName>
sp. nov. by rugae on upper part of frons curved outward and smooth mesosoma.
</paragraph>

View file

@ -1,254 +1,254 @@
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:title id="D6ED4264ABA1D1479748A8C6C8A26BC8">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<treatment ID-GBIF-Taxon="161777000" LSID="urn:lsid:zoobank.org:act:D3D0EC8A-6586-4D6A-B972-D9AFDE718683" httpUri="http://treatment.plazi.org/id/16A8E0EA117393477442362ACB83A687" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/D3D0EC8A-6586-4D6A-B972-D9AFDE718683" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ehazoara" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ehazoara" status="sp. nov.">Pheidole ehazoara</taxonomicName>
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<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">Figs 49A-F</figureCitation>
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84N</figureCitation>
<figureCitation captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86N</figureCitation>
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<taxonomicName id="E0713342AAAC88AC5B2E560B07858D46" ID-CoL="CML4T" LSID="http://zoobank.org/D3D0EC8A-6586-4D6A-B972-D9AFDE718683" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ehazoara" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ehazoara" status="sp. nov.">Pheidole ehazoara</taxonomicName>
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<figureCitation id="4E7A8A139877A6649930865021AA530F" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">Figs 49A-F</figureCitation>
<figureCitation id="AE0E2067FADB32D4ED41ABE587EBE10A" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84N</figureCitation>
<figureCitation id="7A3804EBF493F3CBD198EF62EB467DC2" captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86N</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
<subSubSection id="51946E11CAA6F43AF445626BC797C056" pageId="0" pageNumber="1" type="type material">
<paragraph id="B8A992BC6B76058C455533385C5F6FBA" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="7FCAEA5AC2CBE5AF38F22170BAC51B2E" pageId="0" pageNumber="1">
<emphasis id="C3041A5E816D545BA2ED82B8CFE67596" bold="true" pageId="0" pageNumber="1">
<emphasis id="77DA0D878D5C43194A9D6FFD90B59CB5" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Madagascar. •1 major worker; Toliara; Ehazoara Canyon, 26 km E Betioky;
<geoCoordinate degrees="23.68333" direction="south" orientation="latitude" precision="1" value="-23.68333">-23.68333</geoCoordinate>
<geoCoordinate id="6E25427388A72001AB0FE8A3F3382995" degrees="23.68333" direction="south" orientation="latitude" precision="1" value="-23.68333">-23.68333</geoCoordinate>
,
<geoCoordinate degrees="44.63333" direction="east" orientation="longitude" precision="1" value="44.63333">44.63333</geoCoordinate>
<geoCoordinate id="D10C2A80F1288B00ABF703EB2AE8CC3B" degrees="44.63333" direction="east" orientation="longitude" precision="1" value="44.63333">44.63333</geoCoordinate>
; alt. 175 m; 27 Apr 1997; Fisher et al. leg.; BLF01518, CASENT0872059, middle specimen (CASC).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
<emphasis id="10217B6CF294DE0981B3D0CB8CE7C198" bold="true" pageId="0" pageNumber="1">
<emphasis id="A6DAB0B030B0CFB5FEF680AAEC1BA943" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
.
</emphasis>
Madagascar. •2w., 2s.; same data as for holotype; CASENT0198555, CASENT0923186, CASENT0872218, CASENT0872219 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
<subSubSection id="26D8F967B86CFC93868096EAF0F57A90" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="1302AD26DF27FF4D5D8A1F7A84164269" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="D28822D7D191DDB54FF36C124808B9BB" pageId="0" pageNumber="1">
<emphasis id="6F8BC9241D395E21A6116C9F404BB80B" bold="true" pageId="0" pageNumber="1">
<emphasis id="CB3661037CC6705CA36D5DB7C4FD47C9" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
Head in full-face view elongated; sides of the head with dense, very long, erect pilosity; occipital lobes with thick, sparse, short, irregular, and arcuate rugae, interspaces indistinctly rugulose; inner hypostomal teeth distinct, narrow, and triangular, closely spaced, with narrow and rounded apex directed outward; outer hypostomal teeth bigger and wider than inner hypostomal teeth, dentate.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
<emphasis id="8542F40B105A54D15B971763E0B2B8B5" bold="true" pageId="0" pageNumber="1">
<emphasis id="C55EEB7135A109F75E8D85C4A4227DFE" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
Head foveolate, foveolate, with additional longitudinal rugae on frons and malar area; genae with fading sculpture; scape, when laid back, reaching head margin; promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum; mesosoma foveolate; propodeal spines short, triangular.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
<subSubSection id="CCDE9FE84589CD31BA60BC5DF1C5FEE4" pageId="0" pageNumber="1" type="description">
<paragraph id="B7ADD5BB97B2299D1408A412C19CC205" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="1B7ADBDD4E06249425A27B1FE39784A7" pageId="0" pageNumber="1">
<emphasis id="447D6FCE950C94C8C47D3D92D575E09A" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
<emphasis id="A75E1B7F37ED963DADB89419570D1C92" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 3): HL: 1.07-1.1 (1.08); HW: 0.85-0.88 (0.87); SL: 0.41-0.44 (0.42); EL: 0.14-0.15 (0.14); WL: 0.82-0.86 (0.84); PSL: 0.13-0.14 (0.14); MTL: 0.44-0.46 (0.45); PNW: 0.49-0.51 (0.5); PTW: 0.15-0.16 (0.16); PPW: 0.38-0.42 (0.4); CI: 81.5-83.6 (82.3); SI: 41.8-49.0 (45.9); PSLI: 13.5-15.0 (14.09); PPI: 34.8-37.3 (36.2); PNI: 54.9-55.7 (55.4); MTI: 47.2-52.5 (50.4).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
<emphasis id="C01B7DA6A1315AA8BC5016240645A407" bold="true" pageId="0" pageNumber="1">
<emphasis id="FDF9D024A3B64ACFD7E483AD72446D04" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
In full-face view longitudinal, longer than wide, anterior of eyes straight, posterior of eyes slightly convex, occipital margin of lobes straight, inclining towards centre (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49B</figureCitation>
<figureCitation id="8CCC0B72B1658B04DF8E1A2BC38A4372" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49B</figureCitation>
). In lateral view sub-oval; ventral and dorsal faces convex; dorsal face finely depressed posteriorly, forming indistinct transverse depression between frons and occipital lobes; inner hypostomal teeth visible. Sides of the head with dense, very long, erect pilosity; whole head with dense, moderately long, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, with thick, moderately sparse, longitudinal rugae; interspaces rugulose. Occipital lobes shiny, with thick, sparse, short, irregular, and arcuate rugae, interspaces indistinctly rugulose; frons with moderately sparse, thick, longitudinal rugae, on the anterior part of frons interspaces with fine, short, and longitudinal rugulae, on the posterior part of frons interspaces with fine and irregular rugulae; genae shiny, with fine, dense and longitudinal rugulae, and behind the eyes smooth; malar area with thick, sparse, longitudinal rugae, interspaces smooth to indistinctly rugulose. Centre of clypeus shiny and smooth, lateral sides with longitudinal rugae; median notch indistinct; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, not reaching the midlength of head; pilosity erect (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49B, D</figureCitation>
<figureCitation id="C432392C779A79DF10914F7D4D118AC2" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49B, D</figureCitation>
). Inner hypostomal teeth distinct, narrow, and triangular, closely spaced, with narrow and rounded apex directed outward; outer hypostomal teeth bigger and wider than inner hypostomal teeth, dentate (Fig.
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84N</figureCitation>
<figureCitation id="C594E33B335FADB6ECCB0CD218988808" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84N</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
<emphasis id="59F0103771D44A977115FAE5C6ACAFE8" bold="true" pageId="0" pageNumber="1">
<emphasis id="70CDE8F35B6947B0800E9001E935C4EF" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum relatively low and arched, posterior mesonotum relatively convex, with small, tubercle-like projections; promesonotal groove absent; metanotal groove indistinct; propodeal spines short, triangular, with sharp apex and very wide base; humeral area laterally weakly produced (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D</figureCitation>
<figureCitation id="B9EBF6581AD36069C1FD57B15578CC27" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D</figureCitation>
). Surface foveolate; promesonotum with additional indistinct and thin, transverse rugulae; its dorsal surface with fading sculpture; propodeum with additional longitudinal rugae, its dorsoventral side with fading sculpture. Pilosity moderately dense, long, suberect to erect (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D, F</figureCitation>
<figureCitation id="B372E1FE6AD5FAA3DEBB5B0EC8DCFF15" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="2EC4C72ED84440DBE2DBA804A1F7EB61" bold="true" pageId="0" pageNumber="1">
<emphasis id="BE386D2CCFE50C2BD581B5175D97E1D5" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny; peduncle moderately short, finely foveolate, with distinct, short, horizontal lobes on its basal part; node with fading sculpture, relatively high, triangular, with rounded apex, in rear view node slightly convex; pilosity long and erect (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D, F</figureCitation>
<figureCitation id="B65737012209FF610096C66BFDDAAF8E" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
<emphasis id="6D693028400B61A13151881D7F8A5D66" bold="true" pageId="0" pageNumber="1">
<emphasis id="2C6060C61A0DDC574341C93608405B03" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Shiny and finely shagreened, dorsum with fading sculpture; in dorsal view sides with moderately short, very wide, acute, and triangular projections; pilosity long and erect (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D, F</figureCitation>
<figureCitation id="D4827F5E2D963D6D9C51A1C8861C0C25" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="9553292007FFC3FCDC7155F2100827FD" bold="true" pageId="0" pageNumber="1">
<emphasis id="2E98054F39BC4AFAE7DCF6E473C7BBB6" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny and finely shagreened on the basal part of first gastral tergite; pilosity dense, long, and erect (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D, F</figureCitation>
<figureCitation id="22889B5079F4DA959F231A22055BEA8B" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
<emphasis id="EC792E60840BC63C0C7FFE93F184B6F0" bold="true" pageId="0" pageNumber="1">
<emphasis id="C89DA5170AE982958A121E2B0B4A39C9" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Dark yellow; lower part of malar area brown; gaster yellowish brown to brown (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D, F</figureCitation>
<figureCitation id="D99D5E15F842CBC568FF7A2F4AEAADA3" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49D, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1" start="Figure 49" startId="F49">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 49.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ehazoara" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ehazoara">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ehazoara</emphasis>
<caption id="90CF32C8FB861E6F095B3EE5DBE05734" doi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1" start="Figure 49" startId="F49">
<paragraph id="8129D696102689E2EDA6E76FFA989D6F" pageId="0" pageNumber="1">
<emphasis id="90A60B17AE8F6B2916599EA366A57CB3" bold="true" pageId="0" pageNumber="1">Figure 49.</emphasis>
<taxonomicName id="69AC1FCC18D1D04BA4A6AF53F580CB81" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ehazoara" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ehazoara">
<emphasis id="B9A45CCE1A00EFA0FEDED5FFA1002D03" italics="true" pageId="0" pageNumber="1">Pheidole ehazoara</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<emphasis id="347B472BF8BF676E5856ADCDFA483A40" bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<emphasis id="D1497A713BCF37946E2ACF1A8F9CB284" bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<emphasis id="6D4E86D2086B8BDF73EF4A3137600DD9" bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0923186) and full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<emphasis id="76245883DEECE24316EC240EFCE7F59E" bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<emphasis id="EDA5A0E4C0111D0D8B2A5FDC9DBAB1B0" bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<emphasis id="3457E6404D407E377ED5BDB1D733DCC6" bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0872059).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
<paragraph id="3F7F41CE2ED299083BFA79C165548580" pageId="0" pageNumber="1">
<emphasis id="8B5D4621804EF937805ABEBE4ECFF522" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
<emphasis id="DA67E3645CB42C1017354D7FD887D5F3" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 2): HL: 0.52-0.51; HW: 0.47-0.46; SL: 0.42-0.4; EL: 0.12-0.1; WL: 0.6-0.59; PSL: 0.1-0.09; MTL: 0.37-0.34; PNW: 0.33-0.32; PTW: 0.09-0.09; PPW: 0.17-0.15; CI: 83.6-81.5; SI: 41.8-49.0; PSLI: 15.0-13.9; PPI: 36.3-34.8; PNI: 54.9-55.7; MTI: 47.2-52.5.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
<emphasis id="5A9FA21807322D580D575331C10EDCBB" bold="true" pageId="0" pageNumber="1">
<emphasis id="1A42383B96FCF6DC6BBC869932DF69DE" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Occipital margin straight or indistinctly concave; occipital carina indistinct (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49A</figureCitation>
<figureCitation id="CD7F4FFB7F0916F7C1976C3B3253D000" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49A</figureCitation>
). Pilosity moderately dense, short, suberect. Head shiny, foveolate, with additional longitudinal rugae on frons and malar area; genae with fading sculpture. Clypeus with fine, short, and longitudinal rugulae; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, reaching head margin; pilosity suberect to erect (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49A, C</figureCitation>
<figureCitation id="C6E2526EC4F0BF7C6AA210622A9C0F4A" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49A, C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
<emphasis id="ACAE2BE53C40B3C26AE39D66A33DDD9A" bold="true" pageId="0" pageNumber="1">
<emphasis id="E5594ADAA5F0D9CDED64F39F15CEC556" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum; promesonotal groove absent; metanotal groove present; propodeal spines small, triangular, with acute apex (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C</figureCitation>
<figureCitation id="EDC88D8D25CAF7D2EF90CFF051FEFD99" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C</figureCitation>
). Sculpture foveolate. Pilosity moderately sparse, moderately short, and erect (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C, E</figureCitation>
<figureCitation id="CC1C6DCD1A00548167161CD8DC225FEF" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="9A297D8B033485E23C9BC15C38D288F1" bold="true" pageId="0" pageNumber="1">
<emphasis id="B2D83228D5B1B7B7C99CA0622649C254" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny; peduncle finely foveolate; short and thin; node moderately low, globular, and small; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C, E</figureCitation>
<figureCitation id="66150F9DBEA11792243C822F5A9DCE50" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
<emphasis id="371A7FFA226EE23067E7511F7C040D75" bold="true" pageId="0" pageNumber="1">
<emphasis id="911A06D05BABC2C0560869E9AEA9C142" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Moderately long, low, and slightly convex; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C, E</figureCitation>
<figureCitation id="8EB50B868C2AD2131E34BEE641DA5F25" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="E8B628ABA42088F9D9903532BC8900AE" bold="true" pageId="0" pageNumber="1">
<emphasis id="60EDC9C79D78663D494EF1B0C76BFF1D" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
With moderately sparse, erect pilosity (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C, E</figureCitation>
<figureCitation id="0647E445CBB3DDF8B48843353927B1A0" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
<emphasis id="7F61C3620B05653855D8766AD76D16D5" bold="true" pageId="0" pageNumber="1">
<emphasis id="213A32558B7D123CA56F273135E6EB73" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Dark yellow (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C, E</figureCitation>
<figureCitation id="F7C5CB0BEF7156B1F09692867A465132" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Pheidole ehazoara sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0872059)." figureDoi="10.3897/zookeys.905.39592.figure49" httpUri="https://binary.pensoft.net/fig/373096" pageId="0" pageNumber="1">49C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">After the locus typicus.</paragraph>
<subSubSection id="0E81D3307673CEB41BC10344B2FDB243" pageId="0" pageNumber="1" type="etymology">
<paragraph id="3753DC63F894FC9220BFB337DC53FF05" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="46494FC487B60E7BD6A06161A5E3BCB8" pageId="0" pageNumber="1">After the locus typicus.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was collected at 175 m in elevation, in tropical dry rainforest. Nest was located under a stone.</paragraph>
<subSubSection id="39062848A72278A7D5506D6888C89D62" pageId="0" pageNumber="1" type="biology_ecology">
<paragraph id="08F8627A2BC843A9D3147F82C4EB8227" pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph id="F79C139301B36BDCAE62A3656A42DAC6" pageId="0" pageNumber="1">The species was collected at 175 m in elevation, in tropical dry rainforest. Nest was located under a stone.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ehazoara" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ehazoara">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ehazoara</emphasis>
<subSubSection id="FC6EAD1745AE943B72BB4406A8FF4E02" pageId="0" pageNumber="1" type="comments">
<paragraph id="9C54AD585C309C090B86A53441ED7818" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="DA8BE091F0E95C0D040E4F4C168055A2" pageId="0" pageNumber="1">
<taxonomicName id="09F2BBCC015860E4151EDF7A7E4A0D60" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ehazoara" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ehazoara">
<emphasis id="237A618714A5ED5BA82760A9865BA276" italics="true" pageId="0" pageNumber="1">Pheidole ehazoara</emphasis>
</taxonomicName>
sp. nov. is most similar to
<taxonomicName lsidName="P. fitarata" pageId="0" pageNumber="1" rank="species" species="fitarata">
<emphasis italics="true" pageId="0" pageNumber="1">P. fitarata</emphasis>
<taxonomicName id="5BA3EC27B3D009342EDD9A34B5A77B85" lsidName="P. fitarata" pageId="0" pageNumber="1" rank="species" species="fitarata">
<emphasis id="04EAD88BB4FBE5166D1A471D91270095" italics="true" pageId="0" pageNumber="1">P. fitarata</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
<emphasis id="423ADFC9E874C9130E9A790BDF9B43A2" bold="true" pageId="0" pageNumber="1">
<emphasis id="DAB263A1FA81EEC027B8A3D9270CBEE0" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
It differs from
<taxonomicName lsidName="P. fitarata" pageId="0" pageNumber="1" rank="species" species="fitarata">
<emphasis italics="true" pageId="0" pageNumber="1">P. fitarata</emphasis>
<taxonomicName id="D9B16D5264BC0225AC5ABC05FED75016" lsidName="P. fitarata" pageId="0" pageNumber="1" rank="species" species="fitarata">
<emphasis id="0D79D40CD6A142067C89F01C254FF11B" italics="true" pageId="0" pageNumber="1">P. fitarata</emphasis>
</taxonomicName>
sp. nov. in dark yellow body colouration, frons with rugae connected with rugae on the occipital lobes, absence of foveolae on head, shorter propodeal spines with wider base, and dentate outer hypostomal teeth directed outward.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
<emphasis id="4E34B6FBFDDA2491973C391666232443" bold="true" pageId="0" pageNumber="1">
<emphasis id="3E658CC5EE530CD0307DFB516561FF9B" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
It differs from
<taxonomicName lsidName="P. fitarata" pageId="0" pageNumber="1" rank="species" species="fitarata">
<emphasis italics="true" pageId="0" pageNumber="1">P. fitarata</emphasis>
<taxonomicName id="AD5D794E69931BD513B29D794AC5A142" lsidName="P. fitarata" pageId="0" pageNumber="1" rank="species" species="fitarata">
<emphasis id="CFADD20566FBFC34DB9279221682C49D" italics="true" pageId="0" pageNumber="1">P. fitarata</emphasis>
</taxonomicName>
sp. nov. in presence of the additional longitudinal rugae on frons and malar area, longer postpetiole, and never smooth genae.
</paragraph>