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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="07DDAEE963EF306D8B93DEF24ED8D889" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole mavesatra Salata &amp; Fisher 2020, sp. nov." docType="treatment" docUuid="EFA92733-E4F6-4351-9E36-91E266A59FD7" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:title id="D8DBD1A8152F4EFBC36FB3A3B7BEC1AA">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<treatment ID-GBIF-Taxon="161776995" LSID="urn:lsid:zoobank.org:act:EFA92733-E4F6-4351-9E36-91E266A59FD7" httpUri="http://treatment.plazi.org/id/07DDAEE963EF306D8B93DEF24ED8D889" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/EFA92733-E4F6-4351-9E36-91E266A59FD7" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavesatra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavesatra" status="sp. nov.">Pheidole mavesatra</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">Figs 69A-F</figureCitation>
<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85J</figureCitation>
<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87N</figureCitation>
<treatment id="07DDAEE963EF306D8B93DEF24ED8D889" ID-GBIF-Taxon="161776995" LSID="urn:lsid:zoobank.org:act:EFA92733-E4F6-4351-9E36-91E266A59FD7" httpUri="http://treatment.plazi.org/id/07DDAEE963EF306D8B93DEF24ED8D889" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="5DA25581FB1693B544DB8BE082996D73" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="DE02D977D96D1FA33275AFA387F104CB" pageId="0" pageNumber="1">
<taxonomicName id="CC8D964DFD336F04AC0A065F4447C2DB" ID-CoL="CMLGJ" LSID="http://zoobank.org/EFA92733-E4F6-4351-9E36-91E266A59FD7" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavesatra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavesatra" status="sp. nov.">Pheidole mavesatra</taxonomicName>
<taxonomicNameLabel id="85BCFA1E8A9466824C3E945DB9B49F89" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="D7EFA5F3BCAD359A6D70260130BFDE8C" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">Figs 69A-F</figureCitation>
<figureCitation id="2C0EE0A7836CA5299B552AF2F0DF5D8C" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85J</figureCitation>
<figureCitation id="61541EEBA23BB6CD0CF7EAB90D3E0359" captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87N</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
<subSubSection id="DE66BBE762947ACAC18EF1B20B51A83F" pageId="0" pageNumber="1" type="type material">
<paragraph id="3433525EFA16CCF946753F3557D0A94E" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="01C743F2E948A2F643F1C8B967A3F8FD" pageId="0" pageNumber="1">
<emphasis id="01E23C070523501B34B54BF8F7109F3B" bold="true" pageId="0" pageNumber="1">
<emphasis id="B35DFED93FD978E9785FE15C1F990456" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Madagascar. •1 major worker; Toamasina; Tampolo, 39.4 km SSE Maroantsetra;
<geoCoordinate degrees="15.70978" direction="south" orientation="latitude" precision="1" value="-15.70978">-15.70978</geoCoordinate>
<geoCoordinate id="F3EEEA598D1B2A1979D092F54686536D" degrees="15.70978" direction="south" orientation="latitude" precision="1" value="-15.70978">-15.70978</geoCoordinate>
,
<geoCoordinate degrees="49.96965" direction="east" orientation="longitude" precision="1" value="49.96965">49.96965</geoCoordinate>
<geoCoordinate id="8E45F48CF39DEC3D6CE83D376BFD6212" degrees="49.96965" direction="east" orientation="longitude" precision="1" value="49.96965">49.96965</geoCoordinate>
; alt. 218 m; 30 Aug 2007; B.L. Fisher et al. leg.; BLF17995, CASENT0134807 (CASC).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
<emphasis id="4B5514D6C2D3C3475ADB65D862EF794C" bold="true" pageId="0" pageNumber="1">
<emphasis id="F3557D22116EA377F7D95AB27107E9C5" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
.
</emphasis>
Madagascar. •1w.; same data as for holotype; CASENT0923177 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
<subSubSection id="74EDCA74B10F18E38859B23D5D4E56EC" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="36C844578B56AD592E56A4BA4E4D40C3" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="B4D0059BBBA44B9BC80FD385B85A1538" pageId="0" pageNumber="1">
<emphasis id="875B94B199EA987DD9A672A01DC8DD8F" bold="true" pageId="0" pageNumber="1">
<emphasis id="76AD82A2B8C0864DCC4841E40AEC1E67" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
Head in full-face view rectangular, anterior of eyes relatively straight, posterior of eyes slightly convex; sides of the head with sparse, relatively long, suberect pilosity; scrobe surface shiny, foveolate with sparse, thin, longitudinal rugae; closely spaced, low, triangular, with rounded apex directed inward; outer hypostomal teeth thinner and higher than outer hypostomal teeth, dentate, and with relatively narrow base; gaster finely shagreened; body brown to dark brown.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
<emphasis id="FE380678C76023808941805166A3C0FA" bold="true" pageId="0" pageNumber="1">
<emphasis id="FB1E9E47FA630ADC13C30B4BC6705575" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
Head foveolate, genae with reduced sculpture; promesonotum low, long, relatively flat, with steep posterior declivity; mesosoma foveolate; katepisternum and sometimes mesosoma smooth; propodeal spines moderately long, triangular, narrow; body yellow.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
<subSubSection id="61CDFA475865A89DA83D8AD993665F75" pageId="0" pageNumber="1" type="description">
<paragraph id="BEC8D9A009FABD48148AE31B7363A70F" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="F6ED0BD796872431705022979868B9FF" pageId="0" pageNumber="1">
<emphasis id="33C77D04C9E9E8DE195BCC144DF9E1C9" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
<emphasis id="28C20E3FD91123D5DBD2AA1D5AE6D4F2" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 1): HL: 1.3; HW: 1.2; SL: 0.53; EL: 0.14; WL: 0.92; PSL: 0.21; MTL: 0.52; PNW: 0.62; PTW: 0.15; PPW: 0.47; CI: 92.6; SI: 43.8; PSLI: 15.8; PPI: 32.5; PNI: 51.5; MTI: 43.1.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
<emphasis id="E656FEE33DECEC797A86516E93E152AE" bold="true" pageId="0" pageNumber="1">
<emphasis id="4074D6A6AD41EB075FC316CD49BBA991" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
In full-face view rectangular, anterior of eyes relatively straight, posterior of eyes slightly convex (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69B</figureCitation>
<figureCitation id="65C513927B66BAD71B5B84301436E324" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69B</figureCitation>
). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming shallow transverse depression between frons and occipital lobes; inner hypostomal teeth visible. Sides of the head with sparse, relatively long, suberect pilosity; whole head with dense, moderately short, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, foveolate with sparse, thin, longitudinal rugae. Occipital lobes shiny, with indistinct foveolae and distinct, sparse, thick, irregular rugae, foveolae fading posteriorly; frons with dense, thick, and longitudinal rugae, interspaces superficially foveolate; genae shiny, with distinct and thin rugulae; malar area with thin, dense, longitudinal rugae. Centre of clypeus smooth and shiny, lateral sides with longitudinal rugae; median notch present, moderately wide, and shallow; median longitudinal carina indistinct; lateral longitudinal carinae present. Scape, when laid back, reaching the midlength of head; pilosity suberect to erect (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69B, D</figureCitation>
<figureCitation id="A5A05A6ABEE65A6C76DB4820D8C1EF7C" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69B, D</figureCitation>
). Inner hypostomal teeth distinct, closely spaced, low, triangular, with rounded apex directed inward; outer hypostomal teeth thinner and higher than outer hypostomal teeth, dentate, and with relatively narrow base (Fig.
<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85J</figureCitation>
<figureCitation id="8AAA8B0A828C5C04551831B4D28CB523" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85J</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
<emphasis id="608FE5BD625F66A346940AAACFCF5AF2" bold="true" pageId="0" pageNumber="1">
<emphasis id="F35961A5017EDCD7B8580863CE267ABD" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum relatively low and angular, posterior mesonotum steep, with indistinct tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, triangular, with rounded apex and very wide base; humeral area laterally well produced (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D</figureCitation>
<figureCitation id="D72F6FB3E1022BABE7451BCF9E642CDE" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D</figureCitation>
). Surface shiny and foveolate, foveolae fading on the dorsal surface of promesonotum; promesonotum and lateral surfaces of propodeum with additional thick, sparse, and irregular rugae; katepisternum smooth. Pilosity sparse, long and erect (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D, F</figureCitation>
<figureCitation id="CFA3A8C1423859854E0A9F9F3E4B0600" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="FDCF50C7E8E6E9E4182FC32824ED31C8" bold="true" pageId="0" pageNumber="1">
<emphasis id="54D72C5CC471104540EA97EBD4C5CE28" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny and with fine foveolae; peduncle moderately long, with indistinct horizontal lobes on its basal part; node relatively high, triangular, with rounded apex, in rear view node slightly convex; pilosity long and erect (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D, F</figureCitation>
<figureCitation id="74A9E0D0C10D799868A5D6A2A4B58C8D" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
<emphasis id="7304FAAD4457A90C187F483ED7816724" bold="true" pageId="0" pageNumber="1">
<emphasis id="4B20C16D22F802251F1F3E4BC777E9F2" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Shiny, finely shagreened; in dorsal view sides with moderately long, acute, and horn-like projections; pilosity long and erect (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D, F</figureCitation>
<figureCitation id="A48132A12F361F59EE9DAD8E22522874" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="61D0D9E5DA9901AF67694CDDEF93FA07" bold="true" pageId="0" pageNumber="1">
<emphasis id="F0BEBE59C6FBFB16486A918D61D88054" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny and finely shagreened; pilosity moderately dense, long, and erect (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D, F</figureCitation>
<figureCitation id="A45A497B8DB0F27190C181ED41E94C64" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
<emphasis id="7B51DE9775A3C5927BE71DA0188EB041" bold="true" pageId="0" pageNumber="1">
<emphasis id="93FBD5324B861A9B650B8AE11A58EC2E" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, brown to dark brown; lower part of frons and malar area brighter that the rest of head (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D, F</figureCitation>
<figureCitation id="3849AC1D1185E785CAFE4EFE3EF9EFD2" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69D, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1" start="Figure 69" startId="F69">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 69.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavesatra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavesatra">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mavesatra</emphasis>
<caption id="9ECDF404138B5D349D1401108FFE32DE" doi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1" start="Figure 69" startId="F69">
<paragraph id="DBDA960EC60B53C3E1F99788DBDC5900" pageId="0" pageNumber="1">
<emphasis id="E82CECBE5563D6F2DD4457734D5AB99C" bold="true" pageId="0" pageNumber="1">Figure 69.</emphasis>
<taxonomicName id="B90CFFC026C45A3C5CE034FDF4066572" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavesatra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavesatra">
<emphasis id="72F214070045ED9E802082FFD2570FFF" italics="true" pageId="0" pageNumber="1">Pheidole mavesatra</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<emphasis id="5B2DB7CA091CF90074D9AD88EDEA91EC" bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<emphasis id="2802F7CBFF383F5F77789265833FB107" bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<emphasis id="8A9A2C09B6E457794A3DFA0510FAF4BE" bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0923177) and full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<emphasis id="8C642C7AE3DFC32E75B86407CE768C81" bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<emphasis id="A4EAE1A23920EB9E3ECA27A7C7F56CFA" bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<emphasis id="9A05523BE34851496B5EECD14880ED8A" bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0134807).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
<paragraph id="471D8E15B22867C0AE9511C2A4BB59E9" pageId="0" pageNumber="1">
<emphasis id="D15F1C8B087A825D8D5A60238DC47B04" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
<emphasis id="F6E33841AB3BACBBF89A0C02F9C6A372" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 1): HL: 0.51; HW: 0.48; SL: 0.51; EL: 0.11; WL: 0.6; PSL: 0.09; MTL: 0.37; PNW: 0.31; PTW: 0.05; PPW: 0.11; CI: 93.9; SI: 105.6; PSLI: 18.1; PPI: 46.5; PNI: 65.1; MTI: 78.0.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
<emphasis id="1724ED11943D26EE8E7C361E1DE83D1B" bold="true" pageId="0" pageNumber="1">
<emphasis id="4B562054FDD282B2706C1265EBA12EE7" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Occipital margin straight or indistinctly concave; occipital carina absent (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69A</figureCitation>
<figureCitation id="656FA6EBF7C00C01DFC4629501BBB298" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69A</figureCitation>
). Pilosity moderately sparse, moderately long, suberect to erect. Head foveolate; genae with reduced sculpture. Clypeus with fine and sometimes reduced foveolae; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity erect (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69A, C</figureCitation>
<figureCitation id="F4F2CFB9F7D13CE25F6B2C69077B5C1F" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69A, C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
<emphasis id="80CB347302A57951ECBB71B8F6BB250B" bold="true" pageId="0" pageNumber="1">
<emphasis id="FDDB35F13BAC358AFE7D3873EFE947EE" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum low, long, relatively flat, with steep posterior declivity; promesonotal groove absent; metanotal groove indistinct; propodeal spines moderately long, triangular, narrow, and with acute apex (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C</figureCitation>
<figureCitation id="29F24472E56AF6295CD05F6FD14498C8" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C</figureCitation>
). Sculpture foveolate; katepisternum and sometimes mesosoma smooth. Pilosity moderately sparse, long, and erect (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C, E</figureCitation>
<figureCitation id="DB5EF7671D5E4EE1914C945DF0C1D0E7" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="855203D1E93B21BEA2B43E680665F5B7" bold="true" pageId="0" pageNumber="1">
<emphasis id="29BC53357D4EE6270C0758498B209B7C" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Peduncle short and thin with ventral face slightly convex; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C, E</figureCitation>
<figureCitation id="B15DD8B5FE2BAC2D62C203E18E5145F9" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
<emphasis id="2B3444DB6218E5C4ECA918994A949268" bold="true" pageId="0" pageNumber="1">
<emphasis id="C490641280C6BD0379640D1B907E275F" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Moderately short, low and convex; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C, E</figureCitation>
<figureCitation id="6355C6F0D238E38130C164247466A074" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
<emphasis id="AC4C4EB6D8145462AA7C33888A60CCBD" bold="true" pageId="0" pageNumber="1">
<emphasis id="E7925BC0AE9777CD90C0739D6208A08E" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
With sparse, erect pilosity (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C, E</figureCitation>
<figureCitation id="A56BA6BECCF873AE42DA84ADE1CA2C02" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
<emphasis id="3E1C40E68EFD3B2169CBBF2A97B0C134" bold="true" pageId="0" pageNumber="1">
<emphasis id="4796D28CBBF51ADB34AAF2B413FA3636" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, yellow (Fig.
<figureCitation captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C, E</figureCitation>
<figureCitation id="E3FDF2CDE16E166DFF106F38E806D831" captionStart="Figure 69" captionStartId="F69" captionText="Figure 69. Pheidole mavesatra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923177) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134807)." figureDoi="10.3897/zookeys.905.39592.figure69" httpUri="https://binary.pensoft.net/fig/373116" pageId="0" pageNumber="1">69C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Malagasy for heavy, in reference to massive head of major workers.</paragraph>
<subSubSection id="5C6C5E260257674377EF92B6F23B3639" pageId="0" pageNumber="1" type="etymology">
<paragraph id="D379E538CD5ED3A0FB9FDD76360963B2" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="FB5C280F47C0611BB7FB3DB59D39B368" pageId="0" pageNumber="1">Malagasy for heavy, in reference to massive head of major workers.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was collected at 218 m in elevation, in disturbed rainforest. Nests were located under moss.</paragraph>
<subSubSection id="A3B395A99BD51B435DCC9B12A60C35AB" pageId="0" pageNumber="1" type="biology_ecology">
<paragraph id="A351B2918519447696F3A94F2FF2750C" pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph id="1A945CCDA5704C213507B1204ED80695" pageId="0" pageNumber="1">The species was collected at 218 m in elevation, in disturbed rainforest. Nests were located under moss.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavesatra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavesatra">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mavesatra</emphasis>
<subSubSection id="4B1C9E7DE9E852D8104E4D224B4A839A" pageId="0" pageNumber="1" type="comments">
<paragraph id="49F9A3D40746DA10DB4E1037BBF66C77" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="A19F946101087A68C361A9E0F34644D3" pageId="0" pageNumber="1">
<taxonomicName id="F6C42129B5C75FAD2EE7C7246EEA0FAB" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavesatra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavesatra">
<emphasis id="912662BE9FBA20C4930A6448D21BF34F" italics="true" pageId="0" pageNumber="1">Pheidole mavesatra</emphasis>
</taxonomicName>
sp. nov. is most similar to
<taxonomicName lsidName="P. goavana" pageId="0" pageNumber="1" rank="species" species="goavana">
<emphasis italics="true" pageId="0" pageNumber="1">P. goavana</emphasis>
<taxonomicName id="1CC6E52410FF19D80B9F5DC265EFBA6C" lsidName="P. goavana" pageId="0" pageNumber="1" rank="species" species="goavana">
<emphasis id="B5514EBDE05E9428C5E1E0211BAF0537" italics="true" pageId="0" pageNumber="1">P. goavana</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
<emphasis id="D06A1C818B33DDE216E9A0F8379005A4" bold="true" pageId="0" pageNumber="1">
<emphasis id="CD2333E4B9D7F9F417165A2965C2E5FE" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavesatra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavesatra">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mavesatra</emphasis>
<taxonomicName id="52C3F152E50C800DDF16A9888B552AA4" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavesatra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavesatra">
<emphasis id="F84C0C657CE085891A93A4F367B8A1E2" italics="true" pageId="0" pageNumber="1">Pheidole mavesatra</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName lsidName="P. goavana" pageId="0" pageNumber="1" rank="species" species="goavana">
<emphasis italics="true" pageId="0" pageNumber="1">P. goavana</emphasis>
<taxonomicName id="D15742BA4F15459398AAC3ADF89DA96A" lsidName="P. goavana" pageId="0" pageNumber="1" rank="species" species="goavana">
<emphasis id="AD9C42146DB069DC5643894A443FE116" italics="true" pageId="0" pageNumber="1">P. goavana</emphasis>
</taxonomicName>
sp. nov. in brown to dark brown body colouration, smooth katepisternum, and promesonotum and lateral surfaces of propodeum with additional thick, sparse, and irregular rugae.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
<emphasis id="DBADB27DF60C0648B14E8ACD0EB42DCB" bold="true" pageId="0" pageNumber="1">
<emphasis id="6D81D6FF58D59EB459E91BF77063DA20" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavesatra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavesatra">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mavesatra</emphasis>
<taxonomicName id="50A43D6A68FFCDA3460EC2246AD574C6" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mavesatra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mavesatra">
<emphasis id="30268991DDE7A3428EA4CBFBD80A4778" italics="true" pageId="0" pageNumber="1">Pheidole mavesatra</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName lsidName="P. goavana" pageId="0" pageNumber="1" rank="species" species="goavana">
<emphasis italics="true" pageId="0" pageNumber="1">P. goavana</emphasis>
<taxonomicName id="7D903B35FBDCD533B73DE38D23B4BE39" lsidName="P. goavana" pageId="0" pageNumber="1" rank="species" species="goavana">
<emphasis id="5761DF72BB0607FE54DECF68C460A6D0" italics="true" pageId="0" pageNumber="1">P. goavana</emphasis>
</taxonomicName>
sp. nov. in promesonotum low, long, relatively flat, and narrow propodeal spines.
</paragraph>