| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+sehr
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
|
+ |
+0 - unbedeutend, keine Bindung. | +
| Ruhiges Wasser | +0 - unbedeutend, keine Bindung. | +
| Grundwasser | +0 - unbedeutend, keine Bindung. | +
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+253800 | +
| = | +
+ |
+
+ |
+1637 | +
| = | +
+ |
+
+ |
+1553 | +
| = | +
+ |
+
+ |
+1553 | +
| = | +
+ |
+
+ |
+253800 | +
| = | +
+ |
+
+ |
+2489 | +
| = | +
+ |
+
+ |
+2022 | +
| = | +
+ |
+
+ |
+253800 | +
| = | +
+ |
+
+ |
+1377 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +nicht anwendbar (Not Applicable) | +|
| Mittelland (MP) | +nicht anwendbar (Not Applicable) | +|
| Alpennordflanke (NA) | +verschollen, vermutlich in der Schweiz ausgestorben (Critically Endangered, Probably Extinct) | +|
|
+ |
+-- | +|
|
+ |
+verschollen, vermutlich in der Schweiz ausgestorben (Critically Endangered, Probably Extinct) | +|
| Westliche Zentralalpen (WA) | +
+stark
+ |
+B2ab(iii) | +
|
+ |
+
+3 - Mittlere nationale
+ |
+
|
+ |
+
+1 -
+ |
+
|
+ |
+1 - Gering | +
|
+ |
+
+0 -
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(02.03.2005) | +
|
+ |
+-- | + + +
|
+ |
+Z - Zielart | +weitere Informationen | +
|
+1.
+ |
+present present 12 | +spiny | +submargin | +
|---|---|---|---|
|
+2.
+ |
+present present 15 | +capitate | +caudal ridges | +
|
+3.
+ |
+absent present 14 | +capitate | +submargin | +
|
+4.
+ |
+present absent 12 | +capitate | +submargin | +
|
+5.
+ |
+present present 12 | +spiny | +submargin | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|
| Appendix | +Appendix | +
| Character | +
+ |
+
+ |
+
+ |
+
|---|---|---|---|
| th | +th | +th | +
| SPINES | +SOFT RAYS | +|||||
|---|---|---|---|---|---|---|
| VIII | +IX | +11 | +12 | +13 | +14 | +15 | +
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+116500 | +
| = | +
+ |
+
+ |
+116500 | +
| = | +
+ |
+
+ |
+1152 | +
| = | +
+ |
+
+ |
+116500 | +
|
+ |
+
+ |
+|||
|---|---|---|---|---|
| Standard length (mm) | +46.3 | +44.7 | +41.5 | +47.6 | +
|
+ |
+||||
| Head length | +25.3 | +23.7 | +25.0 | +24.6 | +
| Body depth at dorsal-fin origin | +16.2 | +14.1 | +16.5 | +13.7 | +
| Body width at dorsal-fin origin | +12.1 | +10.0 | +10.7 | +8.4 | +
| Predorsal length | +51.8 | +50.1 | +51.3 | +51.1 | +
| Postdorsal length | +36.5 | +36.9 | +35.2 | +37.5 | +
| Preanal length | +73.0 | +71.5 | +74.5 | +72.6 | +
| Prepelvic length | +50.0 | +51.3 | +50.9 | +52.9 | +
| Distance between pectoral- and pelvic-fin origins | +27.6 | +26.6 | +28.2 | +28.0 | +
| Distance between pelvic- and anal-fin origins | +22.8 | +21.7 | +23.8 | +20.2 | +
| Distance between vent and anal-fin origin | +2.4 | +1.9 | +2.2 | +2.2 | +
| Depth of caudal peduncle | +8.9 | +8.2 | +9.2 | +8.6 | +
| Length of caudal peduncle | +17.9 | +19.4 | +16.0 | +18.5 | +
| Dorsal-fin depth | +23.3 | +21.9 | +23.4 | +20.1 | +
| Anal-fin height | +18.7 | +17.7 | +19.6 | +18.1 | +
| Pectoral-fin length | +24.4 | +20.3 | +25.9 | +23.1 | +
| Pelvic-fin length | +19.3 | +17.1 | +18.8 | +17.2 | +
|
+ |
+||||
| Head depth at eye | +45 | +47 | +45 | +42 | +
| Snout length | +45 | +45 | +43 | +41 | +
| Eye diameter | +22 | +21 | +23 | +19 | +
| Postorbital distance | +36 | +42 | +39 | +40 | +
| Maximum head width | +57 | +60 | +53 | +53 | +
| Interorbital width | +24 | +26 | +29 | +23 | +
| Length of inner rostral barbel | +20 | +18 | +17 | +21 | +
| Length of outer rostral barbel | +28 | +29 | +37 | +33 | +
| Length of maxillary barbel | +27 | +36 | +37 | +31 | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+sehr feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +sauer (pH 3.5-6.5) | +Temperaturzahl T | +unter-montan und ober-kollin | +||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
|
+ |
+0.70 0.70 0.65 0.80 | +1.30 1.25 1.20 1.80 | +1.30 1.30 1.50 2.65 | +0.90 0.85 0.80 1.20 | +5.80 5.70 5.75 8.65 | +
|---|---|---|---|---|---|
| Female I 1.75 II 1.70 III 1.65 IV 2.35 | +0.80 0.80 0.75 0.90 | +1.40 1.25 1.25 1.90 | +1.30 1.30 1.60 2.80 | +0.90 0.90 0.85 1.25 | +6.15 5.95 6.10 9.20 | +
|
+ |
+1.55 1.50 1.40 1.65 | +3.00 2.75 2.55 3.50 | +2.95 2.80 3.05 5.00 | +1.80 1.65 1.50 2.10 | +12.75 12.10 11.65 16.45 | +
| Female I 2.85 II 2.80 III 2.60 IV 3.60 | +1.35 1.25 1.15 1.40 | +2.30 2.10 2.00 3.05 | +2.20 2.15 2.40 4.30 | +1.35 1.30 1.20 1.75 | +10.05 9.60 9.35 14.10 | +
|
+ |
+1.40 1.40 1.25 1.45 | +2.50 2.35 2.20 3.00 | +2.40 2.40 2.50 4.05 | +1.40 1.30 1.20 1.65 | +10.60 10.35 9.90 13.80 | +
| Female I 3.00 II 2.95 III 2.75 IV 3.60 | +1.40 1.40 1.25 1.50 | +2.50 2.30 2.10 3.00 | +2.30 2.30 2.50 4.20 | +1.40 1.35 1.20 1.70 | +10.60 10.30 9.80 14.00 | +
|
+ |
+0.80 0.80 | +1.35 1.20 | +1.40 1.30 | +1.00 0.90 | +6.20 5.80 continued next page | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
|
+ |
+0 - unbedeutend, keine Bindung. | +
| Ruhiges Wasser | +2 - Schwerpunktlebensraum | +
| Grundwasser | +0 - unbedeutend, keine Bindung. | +
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+433700 | +
| = | +
+ |
+
+ |
+1884 | +
| = | +
+ |
+
+ |
+1812 | +
| = | +
+ |
+
+ |
+1812 | +
| = | +
+ |
+
+ |
+433700 | +
| = | +
+ |
+
+ |
+2752 | +
| = | +
+ |
+
+ |
+2238 | +
| = | +
+ |
+
+ |
+433700 | +
| = | +
+ |
+
+ |
+1594 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +-- | +|
| Mittelland (MP) | +vom Aussterben bedroht (Critically Endangered) | +A3c | +
| Alpennordflanke (NA) | +regional beziehungsweise in der Schweiz ausgestorben (Regionally Extinct) | +|
|
+ |
+regional beziehungsweise in der Schweiz ausgestorben (Regionally Extinct) | +|
|
+ |
+-- | +|
| Westliche Zentralalpen (WA) | +vom Aussterben bedroht (Critically Endangered) | +A3c | +
|
+ |
+
+1 - Sehr hohe nationale
+ |
+
|
+ |
+2 - Klarer Massnahmebedarf | +
|
+ |
+2 - Mittel | +
|
+ |
+
+2 -
+ |
+
|
+ |
+Nein | +|
|
+ |
+
+ |
+(01.01.2010) | +
|
+ |
+-- | +|
|
+ |
+
+ |
+(23.01.2013) | +
| Character | +Holotype BMNH 1874.4.29.729 | +ZSI/WGRC/ IR.V/2710 | +ZSI/WGRC/ IR.V/2712 | +ZSI/WGRC/ IR.V/2711 | +ZSI/WGRC/ IR.V/2713 | +ZSI/WGRC/ IR.V/2714 | +
|---|---|---|---|---|---|---|
| Sex | +female | +male | +female | +male | +male | +female | +
| SVL | +42.4 | +40.5 | +42.7 | +38.4 | +39.8 | +47.0 | +
| AG | +18.1 | +17.3 | +18.8 | +15.4 | +17.1 | +21.8 | +
| TW | +5.6 | +6.9 | +8.7 | +6.9 | +7.8 | +8.5 | +
| ED | +1.6 | +2.4 | +2.5 | +1.7 | +2.4 | +2.7 | +
| EN | +2.9 | +3.2 | +3.4 | +3.2 | +3.4 | +3.7 | +
| ES | +5.0 | +4.8 | +4.6 | +4.4 | +5.1 | +5.0 | +
| ET | +4.0 | +3.6 | +3.8 | +3.1 | +3.8 | +4.0 | +
| IN | +1.4 | +1.3 | +1.2 | +1.2 | +1.5 | +1.6 | +
| TD | +0.4 | +0.8 | +0.5 | +0.7 | +0.8 | +0.9 | +
| HL | +10.1 | +10.5 | +11.1 | +9.3 | +11.1 | +11.3 | +
| HW | +7.5 | +7.0 | +7.4 | +6.6 | +7.2 | +7.3 | +
| HD | +5.1 | +4.3 | +4.3 | +3.9 | +4.4 | +4.5 | +
| IO | +4.1 | +2.9 | +3.5 | +4.1 | +3.2 | +3.1 | +
| UAL | +6.5 | +4.4 | +5.3 | +5.4 | +4.5 | +5.8 | +
| LAL | +6.1 | +6.2 | +4.3 | +3.3 | +6.4 | +6.7 | +
| PAL | +6.1 | +4.5 | +5.5 | +5.2 | +5.4 | +5.7 | +
| FL1 | +2.5 | +2.9 | +3.1 | +3.3 | +3.1 | +3.4 | +
| FL2 | +3.7 | +3.5 | +4.1 | +4.2 | +3.6 | +3.9 | +
| FL3 | +4.3 | +4.7 | +4.5 | +4.6 | +4.7 | +5.0 | +
| FL4 | +4.8 | +5.1 | +4.7 | +5.0 | +4.5 | +5.3 | +
| FL5 | +3.9 | +3.8 | +4.4 | +3.9 | +3.7 | +4.5 | +
| FEL | +8.6 | +7.2 | +7.8 | +7.3 | +6.9 | +7.5 | +
| TBL | +7.8 | +6.8 | +4.9 | +6.8 | +6.7 | +7.1 | +
| TOL1 | +2.9 | +2.2 | +3.5 | +3.9 | +2.5 | +2.5 | +
| TOL2 | +3.6 | +4.2 | +4.5 | +5.2 | +4.4 | +4.6 | +
| TOL3 | +4.9 | +4.9 | +5.5 | +5.8 | +5.3 | +5.3 | +
| TOL4 | +6.1 | +5.7 | +5.8 | +6.6 | +6.1 | +6.2 | +
| TOL5 | +4.9 | +5.0 | +4.9 | +5.2 | +5.1 | +5.3 | +
| TL | +32.6 | +53.0 | +52.6 | +42.8 | +- | +51.1 | +
| TBW | +5.6 | +4.6 | +4.8 | +4.5 | +4.7 | +4.9 | +
| Supralabials(R/L) | +8,8 | +7,8 | +7,7 | +7,7 | +7,7 | +8,8 | +
| Infralabials(R/L) | +8,7 | +7,7 | +7,7 | +7,7 | +7,7 | +7,7 | +
| LamIVmanus | +16 | +18 | +17 | +15 | +16 | +18 | +
| LamIVpes | +19 | +17 | +18 | +18 | +19 | +19 | +
| Femoral pores | +none | +5,5 | +none | +4,5 | +5,6 | +none | +
| Lοcality | +1 | +2 | +3 | +4 | +5 | +6 | +7 | +8 | +9 | +10 | +11 | +12 | +13 | +14 | +15 | +16 | +ref. | +|
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
+ |
+||||||||||||||||||
|
+ |
+Bοrneο | +
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|
|
+ |
+Bοrneο | +
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|
|
+ |
+Bοrneο | +40‾43 | +26‾32 | +- | +60‾68 | +4 | ++ | +0 | +2 | +6 | +5 | +6‾10 | +- | +0 | +0 | +0 | +0 | +A | +
|
+ |
+Philippines | +31 | +30 | +- | +- | +4 | ++ | ++ | +2 | +6 | +5 | +10 | +s | ++ | ++ | +0 | +0 | +B | +
|
+ |
+Philippines | +35 | +32 | +- | +- | +4 | ++ | +0 | +6 | +6 | +10 | +s | +0 | ++ | ++ | +0 | +C | +|
|
+ |
+Bοrneο | +48‾52 | +33‾41 | +- | +63‾73 | +4 | +0 | ++ | +2 | +- | +- | +10‾15 | +- | +0 | ++ | +0 | +0 | +A | +
|
+ |
+Sulawesi | +36 | +30 | +- | +- | +4 | +- | +0 | +- | +- | +- | +10 | +s | ++ | +0 | +0 | +0 | +D | +
|
+ |
+W Malaysia | +40.5‾44 | +31‾33 | +73‾74 | +61‾74 | +4 | ++ | ++ | +2 6 | +5 | +12‾13 | +k | +0 | +faint 0 | +0 | +A | +
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
+ |
+W Malaysia | +34‾44 | +31‾33 | +66‾67 | +62‾63 | +4 | ++ | ++ | +1,2 6 | +5‾6 | +12‾13 | +s | ++ | ++ 0 | +0 | +A | +
|
+ |
+W Malaysia | +38‾41 | +30‾32 | +68‾73 | +60‾70 | +4 | ++ | ++ | +2 6 | +5 | +11 | +k | ++ | +faint + | +0 | +A | +
|
+ |
+W Malaysia | +37.2 | +34‾37 | +60‾72 | +70‾72 | +4 | ++ | ++ | +2 6 | +5,6 | +11,12 | +k | ++ | ++ + | ++ | +A | +
|
+ |
+W Malaysia | +30 | +29 | +65 | +61 | +4 | ++ | ++ | +2 6 | +6 | +10 | +s | ++ | +0 0 | +0 | +A | +
|
+ |
+
+ |
+32 | +28 | +- | +- | +4 | +- | +0 | +- 5 | +- | +9 | +s | ++ | +0 0 | ++ | +E | +
|
+ |
+W Malaysia | +ϳuv | +29 | +58 | +52 | +4 | ++ | ++ | +1 6 | +5 | +9 | +k | ++ | ++ 0 | ++ | +A | +
|
+ |
+W Malaysia | +35.5 | +30‾35 | +68‾70 | +70,71 | +4 | ++ | ++ | +2 6 | +5 | +10,11 | +k | ++ | ++ + | ++ | +A | +
|
+ |
+56 | +30‾37 | +68‾80 | +62‾72 | +4 | ++ | +0,+ | +2 | +6 | +4‾5 | +9‾11 | +s | +0 | ++ | +0 | +0 | +A | +
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
+ |
+38 | +30 | +- | +- | +5 | +- | ++ | +- | +- | +- | +15 | +- | ++ | ++ | +0 | +0 | +F | +
|
+ |
+|||||||||||||||||
|
+ |
+35 | +24 | +- | +- | +4 | ++ | ++ | +- | +- | +- | +21‾23 | +s | ++ | ++ | +- | +0 | +A | +
|
+ |
+52‾60 | +32‾33 | +76‾80 | +74 | +4 | +0,+ | ++ | +2 | +7 | +7‾8 | +12‾13 | +k | ++ | +0,+ | +0,+ | +0 | +A | +
|
+ |
+39 | +28‾32 | +63‾75 | +- | +4 | ++ | +0,+ | +2 | +- | +- | +11‾15 | +- | ++ | ++ | +0 | +0 | +A | +
|
+ |
+32 | +22 | +- | +- | +4 | +- | ++ | +- | +- | +- | +10 | +s | ++ | ++ | +0 | ++ | +G | +
| Lοcality | +1 | +2 | +3 | +4 | +5 | +6 | +7 8 | +9 | +10 | +11 | +12 | +13 | +14 | +15 | +16 | +ref. | +|
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
+ |
+W Malaysia Java | +60‾65 20 | +32 31 | +72‾73 81 | +68 - | +4 4 | ++ + | +0 2 0 2 | +7 6 | +6 5 | +13‾17 14‾15 | +s s | ++ + | +0 0 | +0 0 | +0 0 | +H I | +
|
+ |
+
+ |
+42 | +27‾28 | +62‾64 | +60 | +5 | ++ | +0 2 | +5‾6 | +4‾5 | +8‾9 | +k | +0 | ++ | +0 | ++ | +J | +
|
+ |
+Bοrneο W Malaysia | +33 37 | +26 24 | +- 56 | +- - | +4 4 | +- + | +0 - 0 2 | +6 7 | +- 7 | +12 9 | +s - | +0 + | ++ + | +0 0 | +0 0 | +K A | +
|
+ |
+NE India | +35 | +24 | +- | +- | +4 | ++ | ++ - | +- | +- | +11 | +s | +- | ++ | +0 | ++ | +L | +
|
+ |
+Τhailand | +30 | +34 | +71 | +- | +4 | +- | ++ 2 | +6 | +5 | +12 | +- | +- | ++ | +0 | ++ | +M | +
|
+ |
+Τhailand | +28 | +30 | +- | +- | +4 | +- | +0 1 | +- | +- | +17 | +s | ++ | ++ | +0 | ++ | +N | +
|
+ |
+Bοrneο | +47 | +35‾36 | +79 | +84 | +7 | ++ | +0 1 | +6 | +6 | +18‾23 | +s | ++ | +0 | +0 | +0 | +A | +
|
+ |
+vietnam | +36 | +30 | +61 | +- | +4 | +- | ++ 2 | +7 | +6 | +16 | +- | ++ | ++ | +0 | +0 | +M | +
|
+ |
+Sumatra | +41 | +32 | +- | +- | +4 | +- | ++ - | +- | +- | +15 | +s | ++ | +0 | +0 | +0 | +Ο | +
|
+ |
+Java | +45 | +29 | +64 | +- | +5 | ++ | ++ 2 | +7 | +7 | +25 | +s | ++ | ++ | +0 | +0 | +A | +
|
+ |
+Java | +40‾45 | +32‾34 | +71 | +- | +5 | ++ | ++ 2 | +7 | +6 | +26‾27 | +- | ++ | +0 | +0 | +0 | +A | +
|
+ |
+W Malaysia | +50 | +28‾31 | +67‾74 | +- | +5 | ++ | +0,+ 2 | +7 | +6 | +22‾23 | +- | ++ | ++ | +0 | +0 | +A | +
|
+ |
+vietnam Bοrneο | +35.2 67 | +20 23‾34 | +53 - | +54 - | +4 4 | ++ + | +0 2 + - | +6 - | +5 - | +6 28‾29 | +s s | +- + | ++ + | +0 0 | ++ 0 | +A A | +
|
+ |
+Bοrneο | +48‾64 | +40‾42 | +76‾79 | +- | +5 | ++ | +0,+ 4 | +8‾9 | +7 | +16‾17 | +k | ++ | ++ | +0 | +0 | +A | +
|
+ |
+Bοrneο vietnam | +46 35 | +26 20 | +57 48 | +68 - | +4 4 | ++ 0 | +0 - 0 1 | +7 6 | +6 7 | +21‾24 10 | +s s | ++ + | ++ + | +0 0 | +0 + | +A P | +
|
+ |
+vietnam | +37.5‾48.8 | +32‾34 | +65‾72 | +61‾67 | +4 | ++ | ++ 2 | +7 | +6 | +15‾19 | +s | ++ | ++ | +0 | +0 | +Q | +
|
+ |
+vietnam | +36.5 | +18 | +50 | +- | +4 | ++ | +0 1 | +6 | +5 | +7‾8 | +- | +- | ++ | +0 | +0 | +R | +
| Species | +
+ |
+
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|---|---|
| Host | +
+ |
+
+Ciconia nigra, Ciconia
+ |
+Ciconia ciconia | +
+ |
+
+ |
+
+ |
+|
| Origin | +Ukraine | +Central Europe | +Ukraine | +India | +Belgian Congo | +India | +|
| Source | +Present study | +Dietz (1910) | +Iskova(1985) | +Mehra(1980) | +Skrjabin & Bashkirova (1956) | +
+ |
+|
| Parameter | +Range | +Mean | +Range | +Range | +Range | +Range | +Range | +
| Body length | +3,115–3,628 | +3,417 | +5,500–8,000 | +5,320–6,500 | +6,800 | +4,000–4,500 | +4,760 | +
| Body width in forebody | +1,182–1,709 | +1,452 | +1,870–2,290 | +1,280–1,360 | +– | +2,450–2,850 | +1,190 | +
| Body width in hindbody | +595–1,279 | +968 | +– | +800–900 | +– | +650–1,200 | +1,020 | +
| Collar length | +194–302 | +245 | +– | +200 | +187 | +530 | +200 | +
| Collar width | +405–479 | +448 | +– | +450–500 | +255 | +830–1,000 | +420 | +
| Number of spines | +27 | +27 | +27 | +25 | +27 | +26 | +|
| Angle spines length | +120–148 | +136 | +160–185 | +98–140 | +175–180 | +210–240 | +85 | +
| Angle spines width | +20–25 | +22 | +34 | +22–28 | +38 | +32 | +20 | +
| Lateral spines length | +90–115 | +102 | +– | +56–62 | +88–93 | +130–135 | +65 | +
| Lateral spines width | +13–19 | +15 | +– | +21 | +23 | +21–30 | +15 | +
| Dorsal spines length | +85–113 | +99 | +74–125 | +56–62 | +– | +130–135 | +65 | +
| Dorsal spines width | +13–18 | +15 | +– | +21 | +– | +21–30 | +15 | +
| Oral sucker length | +116–159 | +133 | +– | +120–130 | +– | +– | +– | +
| Oral sucker width | +131–182 | +155 | +130–150 | +140–160 | +136 | +380–400 | +136 | +
| Prepharynx length | +69–75 | +72 | +27 | +– | +0 | +– | +85 | +
| Pharynx length | +217–278 | +244 | +220 | +– | +204 | +– | +170 | +
| Pharynx width | +111–154 | +136 | +170 | +112–128 | +136 | +– | +136 | +
| Oesophagus length | +986–1,448 | +1,168 | +– | +– | +1,700 | +– | +1,730 | +
| Ventral sucker length | +502–690 | +610 | +– | +– | +– | +– | +– | +
| Ventral sucker width | +497–614 | +553 | +430–530 | +440–460 | +– | +560 | +425 | +
| Cirrus-sac length | +181 | +– | +– | +– | +120 | +– | +221 | +
| Cirrus-sac width | +146 | +– | +– | +– | +108 | +– | +135 | +
| Right testis length | +200–410 | +299 | +300 | +260–280 | +200 | +– | +370 | +
| Right testis width | +163–343 | +232 | +240 | +240 | +136 | +– | +220 | +
| Left testis length | +258–355 | +303 | +300 | +260–280 | +170 | +– | +220 | +
| Left testis width | +165–364 | +230 | +240 | +240 | +136 | +– | +150 | +
| Ovary length | +171–392 | +281 | +310 | +340 | +170 | +– | +220 | +
| Ovary width | +232–477 | +338 | +370 | +360 | +204 | +– | +200 | +
| Mehlis’ gland length | +222–410 | +305 | +– | +– | +– | +– | +
+ |
+
| Mehlis’ gland width | +280–572 | +382 | +– | +– | +– | +– | +– | +
| Egg-length | +87–111 | +99 | +88–92 | +88–96 | +93 | +– | +– | +
| Egg-width | +54–75 | +60 | +53–57 | +48 | +48 | +– | +– | +
| Species | +Ch. leucocephalus Vrat, 1947 | +Ch. odhneri Vrat, 1947 | +Ch. panduriformis Travassos, 1922 | +Ch. schulzi Gnedina, 1941 | +
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|---|---|
| Host | +
+ |
+
+Anastomus
+ |
+Ciconia maguari (Gmelin, 1789) | +Ciconia nigra | +
+Ephippiorhynchus
+ |
+Ciconia nigra | +
+ |
+
| Origin | +India | +India | +Brazil | +Russia | +India | +China | +India | +
| Source | +Mehra (1980) | +Mehra (1980) | +Skrjabin & Bashkirova(1956) | +Skrjabin & Bashkirova(1956) | +Skrjabin & Bashkirova(1956) | +Ku, Chiu, Li & Chu (1974) | +
+ |
+
| Parameter | +Range | +Range | +Range | +Range | +Range | +Range | +Range | +
| Body length | +8,040 | +5,491 | +6,000–10,000 | +4,820 | +4,410–8,800 | +4,850–6,600 | +6,760 | +
| Body width in forebody | +4,160 | +2,920 | +1,200–4,000 | +1,380 | +3,000–4,000 | +636–1,750 | +3,480 | +
| Body width in hindbody | +1,360 | +1,020 | +– | +– | +700–1,000 | +– | +– | +
| Collar length | +150 | +187 | +– | +270 | +– | +– | +170 | +
| Collar width | +750 | +560 | +870 | +440 | +590–670 | +412–448 | +660 | +
| Number of spines | +29 | +26 | +23–26 | +29 | +27 | +27 | +28 | +
| Angle spines length | +325 | +100 | +230 | +83–133 | +190–210 | +77–109 | +153 | +
| Angle spines width | +38 | +25 | +– | +– | +35–45 | +18–23 | +25 | +
| Lateral spines length | +– | +75 | +120 | +98 | +83–120 | +– | +100 | +
| Lateral spines width | +– | +15 | +– | +14 | +18–20 | +– | +18 | +
| Dorsal spines length | +– | +75 | +– | +– | +83–120 | +– | +100 | +
| Dorsal spines width | +– | +15 | +– | +– | +18–20 | +– | +18 | +
| Oral sucker length | +– | +– | +260 | +121 | +210–230 | +150–182 | +250 | +
| Oral sucker width | +290 | +102 | +220 | +103 | +300–330 | +136–173 | +170 | +
| Prepharynx length | +117 | +204 | +– | +– | +– | +72–77 | +85 | +
| Pharynx length | +250 | +255 | +290 | +220 | +– | +168–186 | +374 | +
| Pharynx width | +170 | +221 | +260 | +121 | +250 | +96–132 | +340 | +
| Oesophagus length | +4,500 | +380 | +3,000–4,000 | +– | +– | +– | +3,650 | +
| Ventral sucker length | +800 | +595 | +700 | +– | +506 | +466–591 | +540 | +
| Ventral sucker width | +600 | +544 | +– | +570–580 | +547 | +376–502 | +490 | +
| Cirrus-sac length | +204 | +– | +– | +311 | +170 | +223 | +510 | +
| Cirrus-sac width | +119 | +– | +– | +294 | +200 | +99 | +187 | +
| Right testis length | +220 | +510 | +207 | +420–460 | +295–394 | +420 | +|
| Right testis width | +500 | +340 | +340–430 | +276 | +290–330 | +161–287 | +290 | +
| Left testis length | +200 | +340 | +– | +207 | +300–330 | +269–403 | +340 | +
| Left testis width | +120 | +222 | +340–430 | +224 | +210–250 | +179–287 | +250 | +
| Ovary length | +– | +– | +– | +432 | +400–500 | +448–645 | +578 | +
| Ovary width | +600 | +510 | +500 | +449 | +500 | +358–518 | +470 | +
| Mehlis’ gland length | +935 | +830 | +– | +536 | +– | +305–466 | +850 | +
| Mehlis’ gland width | +395 | +440 | +– | +259 | +– | +537–645 | +320 | +
| Egg-length | +– | +110 | +120 | +103 | +67–109 | +70–108 | +80 | +
| Egg-width | +– | +55 | +63 | +69 | +42–65 | +59–73 | +
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+sehr warm-kollin (nur an
+ |
+||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| Leg | +
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+All | +Leg: body length | +
|---|---|---|---|---|---|---|---|
| I | +19 | +300 | +116 | +154 | +300 | +889 | +1.05 | +
| II | +19 | +232 | +97 | +131 | +259 | +738 | +0.87 | +
| III | +139 | +172 | +82 | +154 | +251 | +798 | +0.95 | +
| IV | +187 | +206 | +109 | +169 | +315 | +986 | +1.17 | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +basisch (pH 6.5->8.5) | +Temperaturzahl T | +
+subalpin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Fe | +ti | +ta1 | +ta2 | +ta3 | +ta4 | +ta5 | +LR | +BV | +SV | +
| p1 | +|||||||||
| p2 | +|||||||||
| p3 | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+109300 | +
| = | +
+ |
+
+ |
+294 | +
| = | +
+ |
+
+ |
+1108 | +
| = | +
+ |
+
+ |
+1108 | +
| = | +
+ |
+
+ |
+109300 | +
| = | +
+ |
+
+ |
+914 | +
| = | +
+ |
+
+ |
+794 | +
| = | +
+ |
+
+ |
+109300 | +
| = | +
+ |
+
+ |
+207 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+nicht
+ |
+|
| Mittelland (MP) | +
+nicht
+ |
+|
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+potenziell
+ |
+B2a | +
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
|
+ |
+L - Leitart | +weitere Informationen | +
| Slide nomber Measurements | +
+4290
+ |
+
+4290
+ |
+
+4293
+ |
+
+4290
+ |
+Difference between Max.-Min. values | +|||||
|---|---|---|---|---|---|---|---|---|---|---|
|
+ |
+pt | +
+ |
+pt | +
+ |
+pt | +
+ |
+pt | +|||
| 47.9 | +48.0 | +45.2 | +44.9 | +|||||||
| 60.5 | +58.6 | +58.5 | +58.0 | +|||||||
| 43.5 | +38.8 | +41.4 | +||||||||
| 33.9 | +32.8 | +35.4 | +||||||||
| 43.7 | +46.6 | +42.4 | +43.4 | +|||||||
| 36.0 | +37.8 | +33.3 | +35.4 | +|||||||
| 46.1 | +45.0 | +43.5 | +43.4 | +|||||||
| 36.7 | +36.6 | +34.7 | +34.9 | +|||||||
| 57.1 | +54.0 | +54.9 | +||||||||
| 40.2 | +38.0 | +39.1 | +||||||||
| Species Measurements | +
+ |
+
+ |
+||
|---|---|---|---|---|
| holotype * | +holotype ** | +|||
|
+ |
+pt | +
+ |
+pt | +|
| 49.3 | +36.7 | +|||
| 72.8 | +71.3 | +|||
| 40.5 | +35.2 | +|||
| 28.0 | +23.8 | +|||
| 49.1 | +||||
| 31.8 | +||||
| 48.0 | +||||
| 28.0 | +||||
| 53.0 | +52.5 | +|||
| 36.4 | +31.7 | +|||
| Holotype | +n | +Range | +Mean | +SD | +||
|---|---|---|---|---|---|---|
| 1 | +Standard length (mm) | +34.0 | +8 | +27.5–38.2 | +33.9 | +– | +
| Percentages of standard length | +||||||
| 2 | +Body depth | +11.0 | +8 | +11.6–16.0 | +13.1 | +1.9 | +
| 3 | +Predorsal length | +35.0 | +8 | +35.0–38.8 | +35.9 | +1.5 | +
| 4 | +Dorsal-fin base | +12.1 | +8 | +11.0–13.1 | +12.2 | +0.7 | +
| 5 | +Dorsal fin to adipose-fin | +16.1 | +8 | +13.1–18.5 | +16.4 | +1.7 | +
| 6 | +Preadipose length | +63.2 | +8 | +64.1–65.0 | +64.5 | +0.6 | +
| 7 | +Adipose-fin length | +20.6 | +8 | +21.1–23.6 | +22.2 | +1.0 | +
| 8 | +Adipose-fin depth | +4.1 | +8 | +3.4–4.0 | +3.8 | +0.2 | +
| 9 | +Prepectoral length | +20.8 | +8 | +16.5–23.4 | +20.4 | +2.6 | +
| 10 | +Prepelvic length | +39.7 | +8 | +40.0–42.7 | +41.1 | +1.1 | +
| 11 | +Preanal length | +63.9 | +8 | +62.8–67.2 | +65.2 | +1.4 | +
| 12 | +Pectoral fin length | +12.1 | +8 | +12.6–16.4 | +14.0 | +1.4 | +
| 13 | +Dorsal-fin length | +16.8 | +8 | +14.7–17.7 | +16.7 | +0.9 | +
| 14 | +Pelvic-fin length | +13.6 | +8 | +11.9–15.7 | +14.0 | +1.4 | +
| 15 | +Anal-fin length | +12.1 | +8 | +11.3–14.7 | +12.6 | +1.0 | +
| 16 | +Anal-fin base | +12.1 | +8 | +11.3–15.2 | +13.0 | +1.0 | +
| 17 | +Caudal-peduncle length | +21.6 | +8 | +20.4–22.7 | +21.6 | +0.8 | +
| 18 | +Caudal-peduncle depth | +9.5 | +8 | +6.9–8.6 | +8.0 | +0.8 | +
| 19 | +Upper caudal-fin lobe length | +18.1 | +8 | +19.4–21.8 | +20.4 | +1.2 | +
| 20 | +Lower caudal-fin lobe length | +18.5 | +7 | +18.8–20.8 | +19.3 | +0.8 | +
| 21 | +Maxillary-barbel length | +27.4 | +8 | +29.5–38.0 | +32.2 | +3.1 | +
| 22 | +Outer mental-barbel length | +18.9 | +8 | +18.3–24.3 | +19.9 | +2.1 | +
| 23 | +Inner mental-barbel length | +11.5 | +8 | +9.7–17.0 | +14.3 | +2.5 | +
| 24 | +Head length | +7.3 | +8 | +6.8–8.3 | +7.6 | +0.5 | +
| Percentages of head length | +||||||
| 25 | +Head width | +67.2 | +8 | +68.6–78.2 | +73.8 | +3.7 | +
| 26 | +Head depth | +49.0 | +8 | +46.9–53.3 | +49.6 | +2.1 | +
| 27 | +Snout length | +35.0 | +8 | +33.6–48.5 | +39.1 | +4.2 | +
| 28 | +Eye diameter | +16.8 | +8 | +13.5–16.0 | +15.3 | +1.2 | +
| 29 | +Fleshy interorbital | +29.6 | +8 | +20.4–30.7 | +24.6 | +3.6 | +
| 30 | +Mouth gape | +55.7 | +8 | +40.1–53.1 | +48.3 | +5.4 | +
|
+2.5.2 -
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+61300 | +
| = | +
+ |
+
+ |
+2101 | +
| = | +
+ |
+
+ |
+2094a | +
| = | +
+ |
+
+ |
+2094a | +
| = | +
+ |
+
+ |
+61300 | +
| = | +
+ |
+
+ |
+61300 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|
|
+ |
+19–22 | +18 (17–18) | +19 (17–23) | +
|
+ |
+66–74 | +56 (49–56) | +59 (59–66) | +
|
+ |
+Dorsal semiannuli 3– 19 irregularly bearing 8–9 wide central lobe | +Contiguous dorsal semiannuli 8–11 or 9–12 with 4 wide central lobe | +None | +
|
+ |
+Continuous on all dorsal semiannuli | +Not expressed | +Not expressed | +
|
+ |
+|||
|
+ |
+6 | +5 (4–5) | +4 (3–5) | +
|
+ |
+|||
|
+ |
+2 | +– | +2–3 | +
|
+ |
+4 | +6 | +4 | +
|
+ |
+17–21 | +19 (19–20) | +27 (22–27) | +
|
+ |
+11–13 | +11 (10–12) | +20 (14–22) | +
|
+ |
+49–59 | +22 (22–27) | +51 (41–51) | +
|
+ |
+13–17 | +9 (9–11) | +12 (7–12) | +
|
+ |
+21–26 | +18 (18–20) | +20 (17–20) | +
|
+ |
+4 | +2 (2–3) | +2 (2–3) | +
|
+ |
+15–18 | +7 (7–12) | +15 (14–16) | +
|
+ |
+21–22 | +17 (17–19) | +20 (17–20) | +
|
+ |
+12 | +8 (8–10) | +10 (10–13) | +
|
+ |
+15–18 | +32 (32–36) | +20 (16–20) | +
| Fe | +Ti | +Ta1 | +Ta2 | +Ta3 | +|
|---|---|---|---|---|---|
| p1 | +490–510, 505 | +255–290, 281 | +670–700, 690 | +450–470, 460 | +310–340, 327 | +
| p2 | +550–600, 570 | +420–450, 440 | +280–320, 307 | +150–160, 157 | +90–110, 100 | +
|
+ |
+570–620, 593 | +460–520, 495 | +420 | +220 | +180 | +
| Ta4 | +Ta5 | +LR | +BV | +
+ |
+|
|---|---|---|---|---|---|
| p1 | +230–250, 240 | +100–120, 113 | +2.31–2.41, 2.38 | +1.28–1.36, 1.31 | +1.14–1.19, 1.17 | +
| p2 | +50–70, 60 | +30–40, 35 | +0.67–0.72, 0.7 | +3.58–4.03, 3.74 | +3.15–3.46, 3.29 | +
|
+ |
+110 | +80 | +0.81 | +2.64 | +2.71 | +
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
| Features | +
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|
| Leaves | +Dimorphic | +dimorphic | +trimorphic | +homomorphic, obovate-ligulate | +
| Outer leaves | +Obovate | +spatulate to oblanceolate | +lanceolate | +n/a | +
| Inner leaves | +spatulate, petiole indistinct | +elliptic to suborbicular, petiole narrowly winged | +ovate-elliptic to suborbicular, petiole wingless | +n/a | +
| Bract | +Obovate | +lanceolate | +linear | +linear | +
| Calyx | +
+parted to 1/2 -
+ |
+parted to 1/2 | +parted less than 1/2 | +parted to less than 1/2 | +
| Corolla | +White | +pink | +pink | +Pink | +
| Capsule | +ellipsoidal | +ellipsoidal | +nearly spheroidal | +nearly spheroidal | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
| Pulau Tioman | +holotype 6284 | +M | +147.5 | +15.9 | +41.7 | +27.2 | +3.7 | +8.2 | +6.3 | +10.8 | +12.2 | +16.9 | +11.0 | +19.8 | +23.8 | +75.8 | +12.8 | +
| Pulau Tioman | +5399 | +F | +149.2 | +14.3 | +42.4 | +27.8 | +4.3 | +8.7 | +6.2 | +11.8 | +13.0 | +18.0 | +10.8 | +20.0 | +22.9 | +77.5 | +15.2 | +
| Pulau Tioman | +6283 | +F | +161.3 | +15.0 | +44.2 | +28.1 | +4.4 | +9.7 | +7.0 | +13.8 | +13.4 | +18.4 | +10.3 | +21.0 | +25.3 | +81.3 | +12.9 | +
| Pulau Tioman | +5152 | +F | +157.2 | +16.7 | +41.1 | +28.6 | +4.4 | +9.0 | +4.5 | +10.9 | +12.6 | +17.9 | +11.4 | +20.2 | +25.6 | +75.7 | +16.7 | +
| Pulau Tioman | +7263 | +F | +141.2 | +15.3 | +39.9 | +26.2 | +3.6 | +9.4 | +5.1 | +10.0 | +13.0 | +16.4 | +11.2 | +14.3 | +22.3 | +67.8 | +13.1 | +
| Pulau Tioman | +6890 | +M | +158.4 | +16.9 | +44.0 | +28.3 | +3.7 | +9.6 | +5.2 | +12.4 | +14.7 | +18.1 | +11.3 | +19.7 | +25.2 | +77.4 | +14.5 | +
| Pulau Tioman | +5151 | +M | +146.5 | +16.3 | +39.8 | +27.2 | +3.6 | +9.7 | +5.0 | +10.0 | +12.2 | +17.3 | +11.3 | +19.6 | +24.2 | +71.1 | +15.3 | +
| Pulau Tioman | +4681 | +M | +149.0 | +16.6 | +42.4 | +25.9 | +3.6 | +8.9 | +5.3 | +12.2 | +12.4 | +16.7 | +10.3 | +18.7 | +24.8 | +70.0 | +13.7 | +
| Pulau Tioman | +5390 | +F | +117.0 | +11.9 | +33.3 | +21.1 | +3.3 | +7.3 | +4.5 | +11.9 | +10.8 | +13.6 | +9.0 | +15.4 | +19.1 | +58.8 | +10.2 | +
| Pulau Tioman | +7299 | +F | +149.9 | +15.9 | +41.3 | +27.1 | +4.3 | +8.3 | +5.3 | +12.9 | +12.3 | +16.9 | +10.5 | +19.3 | +26.2 | +72.7 | +14.6 | +
| Pulau Tioman | +6260 | +F | +121.4 | +12.9 | +35.7 | +20.8 | +3.8 | +7.2 | +5.3 | +10.2 | +10.9 | +15.1 | +9.2 | +16.8 | +21.1 | +60.9 | +11.3 | +
| Pulau Tioman | +5849 | +M | +122.6 | +13.8 | +36.2 | +23.1 | +3.3 | +8.3 | +5.0 | +10.2 | +11.9 | +15.4 | +9.6 | +17.6 | +20.8 | +60.3 | +10.8 | +
| Pulau Tioman | +7264 | +F | +133.4 | +14.5 | +39.5 | +25.0 | +3.7 | +8.4 | +6.1 | +12.4 | +12.5 | +16.7 | +10.9 | +18.8 | +22.8 | +67.5 | +12.1 | +
| Pulau Tulai | +4694 | +F | +127.5 | +12.9 | +36.5 | +23.9 | +3.9 | +7.7 | +6.2 | +9.7 | +10.9 | +14.9 | +10.1 | +15.9 | +20.2 | +58.6 | +11.6 | +
| Pulau Tulai | +6282 | +M | +134.5 | +13.6 | +39.1 | +25.8 | +4.5 | +9.7 | +6.4 | +10.9 | +12.0 | +16.6 | +10.3 | +17.6 | +19.8 | +68.8 | +12.7 | +
| Pulau Tulai | +7026 | +F | +143.0 | +15.3 | +39.4 | +27.3 | +4.4 | +8.7 | +4.8 | +11.0 | +13.1 | +16.7 | +10.3 | +17.9 | +23.1 | +69.2 | +13.6 | +
| Pulau Tulai | +4697 | +F | +138.1 | +14.7 | +38.7 | +27.0 | +4.5 | +8.6 | +4.2 | +11.0 | +11.8 | +15.8 | +10.4 | +18.8 | +21.7 | +66.4 | +12.7 | +
| Pulau Tulai | +6278 | +F | +117.1 | +11.8 | +33.4 | +21.5 | +3.5 | +7.6 | +5.4 | +8.5 | +11.0 | +13.9 | +10.3 | +15.2 | +18.8 | +54.0 | +11.7 | +
| Pulau Tulai | +5063 | +F | +133.0 | +14.7 | +38.3 | +24.4 | +3.9 | +8.6 | +5.2 | +9.8 | +11.0 | +16.1 | +10.2 | +18.1 | +20.8 | +68.0 | +11.7 | +
| Pulau Tulai | +5062 | +M | +139.6 | +16.4 | +39.5 | +27.7 | +4.4 | +9.2 | +5.2 | +11.8 | +12.7 | +17.1 | +11.4 | +17.2 | +21.9 | +65.4 | +12.5 | +
| Pulau Tulai | +6265 | +F | +128.5 | +13.7 | +37.0 | +25.7 | +3.8 | +7.5 | +5.2 | +10.3 | +11.1 | +15.3 | +10.5 | +15.9 | +20.9 | +64.6 | +12.2 | +
| Pulau Tulai | +6277 | +F | +141.1 | +15.6 | +38.8 | +29.2 | +3.8 | +8.8 | +4.1 | +10.7 | +12.1 | +16.5 | +10.7 | +18.2 | +21.9 | +68.6 | +14.1 | +
| Pulau Tulai | +7025 | +M | +136.6 | +15.1 | +38.1 | +25.9 | +3.7 | +8.9 | +6.1 | +10.5 | +11.6 | +16.1 | +10.2 | +17.4 | +21.3 | +62.6 | +12.4 | +
| Pulau Tulai | +7257 | +F | +137.6 | +14.3 | +39.4 | +25.4 | +3.6 | +8.4 | +5.3 | +10.5 | +11.8 | +16.1 | +10.7 | +17.1 | +22.4 | +66.6 | +12.9 | +
| Pulau Tulai | +3891 | +F | +127.1 | +13.4 | +37.2 | +23.5 | +3.6 | +8.1 | +4.8 | +10.6 | +11.1 | +14.9 | +9.9 | +17.8 | +20.6 | +64.1 | +10.5 | +
| Pulau Tulai | +5059 | +F | +121.5 | +14.2 | +36.5 | +23.2 | +3.5 | +9.3 | +5.3 | +10.2 | +10.8 | +15.4 | +10.4 | +17.0 | +20.8 | +60.5 | +11.1 | +
| Pulau Tulai | +7024 | +M | +135.6 | +14.7 | +40.1 | +26.5 | +3.9 | +9.8 | +5.1 | +11.1 | +11.9 | +16.7 | +9.8 | +16.6 | +21.6 | +65.6 | +12.4 | +
| Pulau Tulai | +5061 | +F | +137.3 | +15.8 | +38.5 | +24.8 | +4.0 | +9.0 | +4.9 | +10.9 | +11.6 | +16.2 | +10.1 | +17.2 | +22.2 | +70.5 | +13.2 | +
| Pulau Tulai | +3990 | +M | +148.8 | +16.2 | +42.9 | +26.4 | +4.8 | +9.3 | +5.7 | +12.2 | +13.0 | +17.6 | +10.8 | +17.9 | +22.2 | +67.5 | +11.4 | +
| Pulau Tulai | +5060 | +M | +124.7 | +15.4 | +37.0 | +25.2 | +3.6 | +9.3 | +5.5 | +12.3 | +11.2 | +15.5 | +10.6 | +16.6 | +20.9 | +60.5 | +13.7 | +
| Pulau Tulai | +7251 | +F | +143.9 | +15.0 | +38.8 | +25.4 | +3.9 | +9.2 | +6.2 | +11.1 | +11.6 | +16.6 | +10.3 | +17.5 | +21.8 | +72.1 | +12.0 | +
| Endau Rompin | +7649 | +F | +143.5 | +15.9 | +40.2 | +27.1 | +3.7 | +8.6 | +5.7 | +11.4 | +12.1 | +16.6 | +11.0 | +16.9 | +21.7 | +72.0 | +11.8 | +
| Endau Rompin | +7694 | +F | +143.0 | +14.9 | +39.5 | +25.4 | +3.7 | +8.2 | +4.5 | +10.8 | +11.7 | +16.3 | +9.7 | +18.9 | +22.2 | +70.5 | +13.3 | +
| Endau Rompin | +7651 | +M | +116.4 | +11.9 | +34.3 | +21.2 | +3.4 | +7.2 | +4.1 | +8.9 | +11.2 | +13.5 | +9.7 | +15.3 | +19.1 | +58.4 | +10.5 | +
| Endau Rompin | +7702 | +F | +67.1 | +7.3 | +20.5 | +12.6 | +2.3 | +4.5 | +2.7 | +4.9 | +6.2 | +8.3 | +4.7 | +7.8 | +9.8 | +33.4 | +4.8 | +
| Endau Rompin | +7650 | +F | +61.2 | +6.1 | +19.5 | +11.8 | +2.3 | +4.9 | +2.4 | +4.4 | +5.1 | +7.4 | +5.1 | +8.0 | +9.6 | +32.0 | +4.6 | +
| Endau Rompin | +7648 | +M | +151.5 | +16.8 | +43.0 | +30.7 | +3.5 | +8.0 | +4.3 | +12.9 | +13.7 | +17.8 | +10.5 | +20.0 | +23.5 | +75.3 | +14.3 | +
| Gunung Lambak | +9959 | +M | +56.9 | +6.4 | +17.9 | +11.3 | +1.1 | +3.7 | +2.0 | +3.6 | +4.9 | +6.7 | +5.3 | +7.0 | +9.0 | +27.9 | +3.1 | +
| Gunung Ledang | +10585 | +F | +54.6 | +5.3 | +17.1 | +10.6 | +1.8 | +4.0 | +2.2 | +3.8 | +4.4 | +6.7 | +4.9 | +7.1 | +8.9 | +26.1 | +3.0 | +
|
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+
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+
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+
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+
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+
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+
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+
| Pulau Tioman | +holotype 6284 | +M | +12 | +10 | +18 | +6 | +110 | +17 | +10 | +25 | +15 | +19 | +8 | +4 | +
| Pulau Tioman | +5399 | +F | +14 | +12 | +21 | +5 | +104 | +18 | +10 | +22 | +17 | +21 | +0 | +4 | +
| Pulau Tioman | +6283 | +F | +14 | +11 | +18 | +6 | +95 | +18 | +10 | +26 | +16 | +20 | +0 | +4 | +
| Pulau Tioman | +5152 | +F | +12 | +10 | +18 | +5 | +91 | +17 | +10 | +21 | +16 | +21 | +0 | +4 | +
| Pulau Tioman | +7263 | +F | +15 | +13 | +18 | +6 | +90 | +16 | +9 | +22 | +18 | +22 | +0 | +2 | +
| Pulau Tioman | +6890 | +M | +14 | +12 | +19 | +7 | +98 | +19 | +11 | +27 | +17 | +21 | +7 | +4 | +
| Pulau Tioman | +5151 | +M | +14 | +12 | +21 | +6 | +109 | +19 | +10 | +23 | +19 | +24 | +8 | +4 | +
| Pulau Tioman | +4681 | +M | +15 | +12 | +20 | +5 | +91 | +20 | +10 | +21 | +16 | +20 | +7 | +6 | +
| Pulau Tioman | +5390 | +F | +13 | +11 | +17 | +7 | +106 | +21 | +11 | +26 | +17 | +21 | +0 | +4 | +
| Pulau Tioman | +7299 | +F | +14 | +11 | +18 | +6 | +105 | +19 | +10 | +27 | +15 | +19 | +0 | +2 | +
| Pulau Tioman | +6260 | +F | +14 | +11 | +21 | +7 | +108 | +21 | +11 | +32 | +17 | +21 | +0 | +4 | +
| Pulau Tioman | +5849 | +M | +13 | +11 | +22 | +7 | +91 | +21 | +11 | +22 | +16 | +21 | +6 | +4 | +
| Pulau Tioman | +7264 | +F | +12 | +9 | +20 | +5 | +93 | +17 | +11 | +26 | +16 | +21 | +0 | +4 | +
| Pulau Tulai | +4694 | +F | +13 | +11 | +19 | +8 | +104 | +17 | +9 | +27 | +19 | +23 | +0 | +2 | +
| Pulau Tulai | +6282 | +M | +13 | +11 | +18 | +6 | +101 | +19 | +10 | +24 | +16 | +20 | +10 | +4 | +
| Pulau Tulai | +7026 | +F | +12 | +10 | +21 | +6 | +98 | +18 | +10 | +25 | +17 | +21 | +0 | +2 | +
| Pulau Tulai | +4697 | +F | +12 | +10 | +17 | +6 | +98 | +18 | +11 | +27 | +16 | +20 | +0 | +4 | +
| Pulau Tulai | +6278 | +F | +13 | +11 | +18 | +6 | +102 | +17 | +10 | +24 | +17 | +21 | +0 | +2 | +
| Pulau Tulai | +5063 | +F | +13 | +11 | +17 | +6 | +99 | +17 | +10 | +25 | +16 | +20 | +0 | +4 | +
| Pulau Tulai | +5062 | +M | +13 | +11 | +20 | +5 | +106 | +18 | +10 | +27 | +15 | +19 | +6 | +2 | +
| Pulau Tulai | +6265 | +F | +13 | +11 | +21 | +7 | +109 | +19 | +10 | +28 | +17 | +21 | +0 | +2 | +
| Pulau Tulai | +6277 | +F | +14 | +12 | +19 | +6 | +106 | +17 | +10 | +26 | +17 | +22 | +0 | +2 | +
| Pulau Tulai | +7025 | +M | +14 | +12 | +23 | +6 | +96 | +17 | +10 | +24 | +19 | +22 | +9 | +2 | +
| Pulau Tulai | +7257 | +F | +13 | +11 | +20 | +7 | +95 | +18 | +9 | +23 | +16 | +20 | +0 | +2 | +
| Pulau Tulai | +3891 | +F | +13 | +11 | +20 | +5 | +102 | +18 | +10 | +26 | +18 | +22 | +0 | +2 | +
| Pulau Tulai | +5059 | +F | +14 | +12 | +22 | +6 | +104 | +18 | +10 | +25 | +16 | +21 | +0 | +2 | +
| Pulau Tulai | +7024 | +M | +14 | +12 | +21 | +6 | +100 | +20 | +10 | +24 | +17 | +21 | +8 | +4 | +
| Pulau Tulai | +5061 | +F | +14 | +11 | +19 | +6 | +98 | +19 | +10 | +27 | +17 | +21 | +0 | +2 | +
| Pulau Tulai | +3990 | +M | +13 | +10 | +22 | +6 | +104 | +21 | +10 | +23 | +17 | +20 | +9 | +3 | +
| Pulau Tulai | +5060 | +M | +13 | +10 | +15 | +7 | +92 | +18 | +10 | +26 | +19 | +23 | +9 | +4 | +
| Pulau Tulai | +7251 | +F | +13 | +11 | +21 | +5 | +101 | +19 | +11 | +29 | +19 | +22 | +0 | +4 | +
| Endau Rompin | +7649 | +F | +14 | +9 | +20 | +8 | +191 | +19 | +11 | +32 | +16 | +20 | +0 | +4 | +
| Endau Rompin | +7694 | +F | +13 | +11 | +18 | +6 | +102 | +18 | +10 | +29 | +15 | +19 | +0 | +4 | +
| Endau Rompin | +7651 | +M | +13 | +11 | +19 | +5 | +105 | +16 | +11 | +26 | +18 | +22 | +10 | +4 | +
| Endau Rompin | +7702 | +F | +10 | +10 | +13 | +4 | +89 | +19 | +9 | +23 | +16 | +20 | +0 | +0 | +
| Endau Rompin | +7650 | +F | +13 | +10 | +20 | +5 | +87 | +19 | +10 | +26 | +15 | +19 | +0 | +2 | +
| Endau Rompin | +7648 | +M | +13 | +10 | +21 | +7 | +95 | +19 | +11 | +30 | +14 | +18 | +11 | +6 | +
| Gunung Lambak | +9959 | +M | +13 | +10 | +18 | +6 | +84 | +18 | +10 | +24 | +16 | +20 | +10 | +0 | +
| Gunung Ledang | +10585 | +F | +14 | +11 | +24 | +5 | +89 | +18 | +10 | +23 | +16 | +21 | +0 | +2 | +
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|---|---|---|---|---|---|---|---|
| Color (dorsal view) | +black, shiny | +black, opaque | +black, moderately glossy | +black | +black | +black, shiny | +|
| BoL | +17.2–18.5 | +27.0–29.0 | +17.5–24.0 | +26.0–28.5 | +26.0 | +18.0–29.0 | +|
| BoW | +10.7–11.5 | +?? | +10.8–13.1 | +?? | +?? | +?? | +|
| Frons | +unarmed | +unarmed | +unarmed | +unarmed | +unarmed | +unarmed | +|
| Gen | +unexpanded | +unexpanded | +unexpanded | +unexpanded | +unexpanded | +unexpanded | +|
| HyC | +present | +present | +present | +present | +present | +present | +|
| Rusty setae covering HyC | +absent | +present (long and dense) | +absent | +present (long and dense) | +present present (long and dense) (long) | +||
| MeC | +absent | +absent | +absent | +absent | +absent | +absent | +|
| Elytral striae | +deep | +feeble | +feeble | +deep | +feeble | +deep | +|
| Punctures on elytral striae | +strong and dense | +impunctate | +weak | +weak | +strong and dense sparse | +||
| Interstriae shape | +convex | +flat | +flat | +weakly convex | +flat | +convex | +|
| Punctures on inter- striae | +absent | +absent | +absent | +absent | +absent | +present | +|
| Second interstria near not swollen base | +swollen | +swollen | +swollen | +swollen | +not swollen | +||
| MetaF | +densely punctate | +impunctate | +densely punctate | +sparsely punctate | +sparsely punctate sparsely punctate | +||
| MetaTibrush (male) | +absent | +absent | +absent | +absent | +absent | +absent | +|
| Distribution | +Central Vietnam: Thanh Hoa Province | +N+C Vietnam (Tuyen Quang, Thanh Hoa), Laos | +N. Vietnam: Vinh Phuc Prov., Cao Bang Prov. | +Northern Thai- land (Phukieo) | +Northern Thai- China (Yunnan) land (Chiang Mai) | +||
|
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+
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+||
| Color (dorsal view) | +opaque | +black, opaque | +black, opaque | +black, opaque | +black, Shiny | +||
| BoL | +21.1 | +21.8–26.0 | +21.0-26.0 | +27.0–30.0 | +22.0–28.0 | +||
| BoW | +12.0 | +11.7–13.8 | +14.5 | +?? | +max. 14.6 | +||
| Frons | +unarmed | +unarmed | +unarmed | +unarmed | +armed (minute horn) | +||
| Gen | +unexpanded | +unexpanded | +unexpanded | +unexpanded | +expanded | +||
| HyC | +present | +present | +present | +present | +present | +||
| Rusty setae covering HyC | +short and sparse | +long and dense | +long and dense | +absent | +?? | +||
| MeC | +absent | +absent | +absent | +absent | +absent | +||
| Elytral striae | +?? | +deep | +feeble | +feeble | +deep | +||
| Punctures on elytral striae | +strong and dense | +strong and dense | +strong and dense | +invisible | +strong and dense | +||
| Interstriae shape | +weakly convex | +flat | +flat | +flat | +flat | +||
| Punctures on inter- striae | +distinctly notched margin of interstriae | +distinctly notched slightly notched mar- margin of interstriae gin of interstriae | +absent | +fine, sparse and scattered | +|||
| Second interstria near not swollen base | +not swollen | +not swollen | +swollen | +not swollen | +|||
| MetaF | +densely punctate | +densely punctate | +densely punctate | +sparsely punctate | +densely punctate | +||
| MetaTibrush (male) | +absent | +absent | +absent | +absent | +absent | +||
| Distribution | +Myanmar | +Malaysia | +China (Yunnan), Malaysia, Myanmar, Thailand | +Taiwan | +Brunei (Kalimantan) | +||
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|
| Color (dorsal view) | +black | +black | +black, opaque | +black | +black, opaque | +
| BoL | +25.0 | +24.0 | +23.0–26.0 | +22.0–29.0 | +24.0 | +
| BoW | +?? | +?? | +?? | +?? | +14.0 | +
| Frons | +armed (minute horn) | +armed (minute horn) | +unarmed | +unarmed | +unarmed | +
| Gen | +expanded | +strongly expanded | +unexpanded | +unexpanded | +unexpanded | +
| HyC | +present | +present | +absent | +absent | +absent | +
| Rusty setae covering HyC | +?? | +present (long and dense) | +absent | +absent | +?? | +
| MeC | +absent | +absent | +present | +present | +present | +
| Elytral striae | +deep | +deep | +feeble | +feeble | +deep | +
| Punctures on elytral striae | +vague | +distinct | +irregular | +irregular | +distinct | +
| Interstriae shape | +flat | +weakly convex | +flat | +flat | +flat | +
| Punctures on interstriae | +fine, sparse and scattered | +sparse | +absent | +absent | +absent | +
| Second interstria near base | +not swollen | +not swollen | +not swollen | +not swollen | +not swollen | +
| MetaF | +?? | +weakly punctate | +sparsely punctate | +densely punctate | +impunctate | +
| MetaTibrush (male) | +absent | +absent | +absent | +absent | +absent | +
| Distribution | +Borneo, Java, Sumat- ra (Indonesia) | +Java, Sumatra (Indo- nesia) | +Laos, Thailand, Vietnam | +North Vietnam (Vinh Phuc, Lao Cai) | +
+Bangladesh,
+ |
+
|
+ |
+
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+
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+
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+
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+
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+
| Color (dorsal view) | +black, not shiny | +black, opaque | +black, opaque | +black, not shiny | +black, opaque | +
| BoL | +26.0–27.0 | +28.0–34.0 | +28.0–33.0 | +28.0–30.0 | +29.5 | +
| BoW | +?? | +17.0–21.0 | +?? | +17.0–18.0 | +16.3 | +
| Frons | +unarmed | +armed | +armed | +armed | +armed | +
| Gen | +unexpanded | +expanded | +expanded | +expanded | +expanded | +
| HyC | +absent | +absent | +absent | +absent | +absent | +
| Rusty setae covering HyC | +?? | +absent | +absent | +absent | +?? | +
| MeC | +present | +absent | +absent | +absent | +absent | +
| Elytral striae | +deep | +shallow | +shallow | +deep | +deep | +
| Punctures on elytral striae | +weak, notching interstriae | +weak | +weak | +weak | +weak, indistinct | +
| Interstriae shape | +flat | +flat | +flat | +slightly convex | +slightly convex | +
| Punctures on interstriae | +present (very small) | +absent | +absent | +absent | +absent | +
| Second interstria near base | +?? | +not swollen | +not swollen | +not swollen | +?? | +
| MetaF (ventral side) | +?? | +impunctate | +impunctate | +impunctate | +?? | +
| MetaTibrush (male) | +absent | +present | +present | +present | +present | +
| Distribution | +Thailand | +
+China,
+ |
+China, Taiwan | +
+Bhutan, Northeast
+ |
+Laos-Myanmar border | +
|
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|
| Color (dorsal view) | +black | +black | +black, shining | +
| BoL | +24.0–26.0 | +36.0–52.0 | +23.0–25.0 | +
| BoW | +?? | +?? | +?? | +
| Frons | +armed | +armed | +armed | +
| Gen | +unexpanded | +unexpanded | +unexpanded | +
| HyC | +absent | +absent | +absent | +
| Rusty setae covering HyC | +absent | +absent | +?? | +
| MeC | +absent | +absent | +absent | +
| Elytral striae | +deep | +deep | +deep | +
| Punctures on elytral striae | +invisible | +strong and close | +invisible | +
| Interstriae shape | +weakly convex | +weakly convex | +weakly convex | +
| Punctures on interstriae | +absent | +absent | +absent | +
| Second interstria near base | +not swollen | +not swollen | +not swollen | +
| MetaF (ventral side) | +impunctate | +impunctate | +?? | +
| MetaTibrush (male) | +present | +present | +present | +
| Distribution | +China (Yunnan), Laos, Myanmar, Thailand, Vietnam | +Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan, China | +Thailand | +
|
+ |
+
+ |
+||||
|---|---|---|---|---|---|
| Character | +Holotype | +Male (n = 2) | +Female (n = 4) | +Male (n = 3) | +Female (n = 1) | +
| BoL | +18.38 | +18.2 | +17.84 ± 0.59 | +18.72 ± 1.08 | +18.49 | +
| BoW | +11.32 | +11.19 | +10.97 ± 0.53 | +11.73 ± 0.71 | +11.79 | +
| HeadL | +3.67 | +3.82 | +3.77 ± 0.19 | +4.84 ± 0.46 | +5.21 | +
| HeadW | +7.44 | +7.43 | +7.17 ± 0.34 | +8.02 ± 0.41 | +8.17 | +
| PronL | +4.9 | +4.9 | +4.90 ± 0.26 | +4.93 ± 0.09 | +4.96 | +
| PronW | +10.08 | +9.89 | +9.57 ± 0.56 | +9.97 ± 0.54 | +9.98 | +
| ElyL | +11.4 | +11.24 | +10.97 ± 0.8 | +11.29 ± 0.74 | +11.61 | +
| MWoI123 | +2.51 | +2.51 | +2.48 ± 0.07 | +2.84 ± 0.2 | +2.86 | +
| DP10,15 | +1.03 | +1.04 | +0.98 ± 0.05 | +unclear | +unclear | +
| HoL | +unarmed | +unarmed | +unarmed | +unarmed | +unarmed | +
| PyL | +2.46 | +2.44 | +2.36 ± 0.13 | +2.33 ± 0.12 | +2.24 | +
| PyW | +4.5 | +4.42 | +4.28 ± 0.33 | +4.59 ± 0.41 | +4.66 | +
| ProTiL | +3.3 | +3.28 | +3.20 ± 0.12 | +3.41 ± 0.24 | +3.25 | +
| ProTiW | +2.35 | +2.35 | +2.28 ± 0.10 | +2.62 ± 0.23 | +2.68 | +
| ProTiSL | +1.21 | +1.22 | +1.23 ± 0.18 | +1.5 ± 0.09 | +1.48 | +
| MesoTiL | +3.34 | +3.45 | +3.54 ± 0.26 | +3.75 ± 0.21 | +3.72 | +
| MesoTiW | +1.33 | +1.32 | +1.28 ± 0.16 | +1.42 ± 0.1 | +1.49 | +
| 1st MesoTiSL | +2.09 | +2.02 | +1.88 ± 0.24 | +2.26 ± 0.2 | +2.36 | +
| 2nd MesoTiSL | +0.9 | +0.94 | +0.95 ± 0.14 | +1.17 ± 0.09 | +1.18 | +
| MetaTiL | +4.95 | +4.91 | +4.77 ± 0.21 | +4.89 ± 0.18 | +4.69 | +
| MetaTiW | +1.27 | +1.25 | +1.22 ± 0.13 | +1.41 ± 0.07 | +1.39 | +
| MetaTiSL | +1.55 | +1.5 | +1.49 ± 0.18 | +1.65 ± 0.06 | +1.71 | +
| MetaTaL | +3.72 | +3.71 | +3.68 ± 0.25 | +3.92 ± 0.11 | +4.08 | +
| MetaTa1L | +1.08 | +1.1 | +1.12 ± 0.07 | +1.24 ± 0.08 | +1.24 | +
| MetaTa1W | +0.68 | +0.68 | +0.67 ± 0.08 | +0.77 ± 0.07 | +0.74 | +
| MetaTa5W | +0.32 | +0.32 | +0.35 ± 0.03 | +0.39 ± 0.02 | +0.38 | +
| BoWeight | +0.67432 | +0.67 | +0.63 ± 0.04 | +0.86 ± 0.03 | +0.89676 | +
| DDC | +1.43 | +1.41 | +1.39 ± 0.11 | +1.45 ± 0.09 | +1.58 | +
| HyC | +present | +present | +present | +present | +present | +
| MeC | +absent | +absent | +absent | +absent | +absent | +
| Gen | +unexpanded | +unexpanded | +unexpanded | +unexpanded | +unexpanded | +
| MesoF | +densely punctured | +densely punctured | +densely punctured | +densely punctured | +densely punctured | +
| MetaF | +densely punctured | +densely punctured | +densely punctured | +densely punctured | +densely punctured | +
| MetaTibrush | +absent | +absent | +absent | +absent | +absent | +
| Characteristics | +Stage 25 (N=8) | +Stage 37 (N=5) | +Stage 40 (N=6) | +
|---|---|---|---|
| Total length | +24.26±0.50 | +50.66±1.21 | +53.28±0.26 | +
| Tail length | +16.83±0.36 | +34.32±1.29 | +35.88±0.20 | +
| Maximum tail height | +4.50±0.07 | +10.00±0.44 | +9.68±0.16 | +
| Tail muscle width | +1.96±0.07 | +5.14±0.18 | +5.03±0.14 | +
| Body length | +7.42±0.17 | +16.94±0.20 | +17.40±0.10 | +
| Tail muscle height | +1.89±0.07 | +5.42±0.23 | +5.38±0.09 | +
| Internarial distance | +1.55±0.05 | +2.94±0.07 | +2.75±0.12 | +
| Interorbital distance | +1.68±0.08 | +4.32±0.12 | +5.00±0.07 | +
| 1 | +Apical portion of flagellum cylindrical. Tooth of upper margin of mandible located subbasally. Subapical portion of clypeus with median tooth. Propodeum with costula and distinct median longitudinal carinae | +
+ |
+
| - | +Apical portion of flagellum distinctly flattened below. Tooth of upper margin of mandible located almost at middle. Clypeus without distinct tooth. Propodeum without costula, median longitudinal carinae vestigial | +
+ |
+
| 2 | +Tergite 3 and subsequent tergites black, basal margins brown | +
+ |
+
| - | +Tergite 2 and subsequent tergites entirely reddish brown | +
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+|
|
+ |
+|||||
|
+ |
+VI-CT | +Omnivore | +LC | +NE | +LC | +
|
+ |
+CT | +Omnivore | +LC | +NE | +LC | +
|
+ |
+|||||
|
+ |
+CT | +Insectivore | +VU | +VU | +CR | +
|
+ |
+CT | +Insectivore | +LC | +NE | +LC | +
|
+ |
+|||||
|
+ |
+CT | +Omnivore | +LC | +NE | +LC | +
|
+ |
+CT | +Omnivore | +LC | +NE | +LC | +
|
+ |
+|||||
|
+ |
+VI-CT | +Herbivore | +DD | +NE | +VU | +
|
+ |
+F-VI-CT | +Herbivore | +VU | +VU | +VU | +
|
+ |
+CT | +Herbivore | +LC | +NE | +LC | +
|
+ |
+|||||
|
+ |
+CT | +Omnivore | +LC | +LC | +VU | +
|
+ |
+|||||
|
+ |
+VI-CT | +Omnivore | +LC | +NE | +NE | +
|
+ |
+|||||
|
+ |
+VI-VO | +Omnivore | +NT | +NE | +DD | +
|
+ |
+|||||
|
+ |
+VI-VO | +Omnivore | +LC | +VU | +NT | +
|
+ |
+|||||
|
+ |
+F-VI-CT | +Omnivore | +NE | +NE | +NE | +
|
+ |
+F-VI-CT | +Omnivore | +LC | +NE | +LC | +
|
+ |
+F-VI | +Omnivore | +NT | +VU | +VU | +
|
+ |
+|||||
|
+ |
+F-CT | +Carnivore | +LC | +NE | +VU | +
|
+ |
+CT | +Carnivore | +VU | +VU | +VU | +
|
+ |
+CT | +Carnivore | +NT | +VU | +VU | +
|
+ |
+F-VI-CT | +Carnivore | +LC | +VU | +VU | +
|
+ |
+VI-CT | +Carnivore | +LC | +VU | +DD | +
|
+ |
+|||||
|
+ |
+CT | +Omnivore | +LC | +NE | +LC | +
|
+ |
+CT | +Omnivore | +LC | +NE | +NE | +
|
+ |
+|||||
|
+ |
+VI-CT | +Omnivore | +LC | +NE | +LC | +
|
+ |
+F-CT | +Omnivore | +LC | +NE | +LC | +
|
+ |
+|||||
|
+ |
+F-VI | +Herbivore | +LC | +NE | +NE | +
|
+ |
+|||||
|
+ |
+F-VI | +Herbivore | +LC | +NE | +LC | +
|
+ |
+|||||
|
+ |
+F-CT | +Frugivore | +LC | +NE | +EN | +
|
+ |
+|||||
|
+ |
+CT | +Frugivore | +DD | +NE | +LC | +
|
+ |
+|||||
|
+ |
+VI-CT | +Frugivore | +LC | +NE | +LC | +
|
+ |
+VI-CT | +Frugivore | +LC | +NE | +LC | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+39300 | +
| = | +
+ |
+
+ |
+39300 | +
| = | +
+ |
+
+ |
+1417 | +
| = | +
+ |
+
+ |
+39300 | +
| Male | +Female | +|||
|---|---|---|---|---|
| Character | +Range | +Average ± SE | +Range | +Average ± SE | +
| Body length, with wings | +2.51–2.52 | +2.52±0.01 | +2.56–2.58 | +2.57±0.02 | +
| Body length, no wings | +1.50–1.51 | +1.51±0.01 | +1.60–1.61 | +1.61±0.01 | +
| Forewing length | +2.20–2.21 | +2.20±0.01 | +2.26–2.27 | +2.26±0.01 | +
| Vertex length | +0.11–0.12 | +0.12±0.01 | +0.12–0.12 | +0.12±0.00 | +
| Vertex width, basal margin | +0.22–0.22 | +0.22±0.00 | +0.22–0.22 | +0.22±0.00 | +
| Vertex width, distal margin | +0.95–0.95 | +0.95±0.00 | +0.95–0.95 | +0.95±0.00 | +
| Pronotum length, midline | +0.10–0.10 | +0.10±0.00 | +0.10–0.10 | +0.10±0.00 | +
| Mesonotum length, midline | +0.41–0.41 | +0.41±0.00 | +0.41–0.42 | +0.42±0.01 | +
| Mesonotum width | +0.53–0.54 | +0.53±0.01 | +0.54–0.54 | +0.54±0.00 | +
| Frons width, dorsal margin | +0.95–0.95 | +0.95±0.00 | +0.95–0.95 | +0.95±0.00 | +
| Frons width, clypeal suture | +0.17–0.17 | +0.17±0.17 | +0.17–0.17 | +0.17±0.00 | +
| Frons width, widest | +0.17–0.17 | +0.17±0.17 | +0.17–0.17 | +0.17±0.00 | +
| Frons width, narrowest | +0.09–0.09 | +0.09±0.00 | +0.09–0.09 | +0.09±0.00 | +
| Frons length, midline | +0.34–0.34 | +0.34±0.00 | +0.35–0.35 | +0.35±0.00 | +
| Clypeus length | +0.23–0.23 | +0.23±0.23 | +0.23–0.23 | +0.23±0.23 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|
|
+ |
+1.48 | +0.32 | +1.55 | +1.57 | +0.92 | +
|
+ |
+1.34 | +0.31 | +1.27 | +1.34 | +0.77 | +
|
+ |
+1.05 | +0.27 | +0.84 | +1.01 | +0.55 | +
|
+ |
+1.41 | +0.29 | +1.23 | +1.40 | +0.71 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|
|
+ |
+1.43 | +0.33 | +1.44 | +1.41 | +0.90 | +
|
+ |
+1.27 | +0.32 | +1.16 | +1.18 | +0.76 | +
|
+ |
+1.02 | +0.28 | +0.80 | +0.90 | +0.55 | +
|
+ |
+1.37 | +0.28 | +1.18 | +1.26 | +0.71 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|
|
+ |
+1.19 | +0.28 | +1.25 | +1.26 | +0.82 | +
|
+ |
+1.05 | +0.27 | +1.02 | +1.07 | +0.68 | +
|
+ |
+0.80 | +0.22 | +0.65 | +0.76 | +0.48 | +
|
+ |
+1.04 | +0.24 | +0.98 | +1.09 | +0.63 | +
|
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
+ |
+
|---|---|---|---|---|---|
|
+ |
+1.05 | +0.26 | +0.99 | +0.97 | +0.67 | +
|
+ |
+0.91 | +0.24 | +0.81 | +0.81 | +0.57 | +
|
+ |
+0.70 | +0.22 | +0.53 | +0.60 | +0.39 | +
|
+ |
+0.95 | +0.22 | +0.78 | +0.83 | +0.55 | +
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Sex | +Type | +Country | +Label | +Latitude Longitude | +Code | +
|---|---|---|---|---|---|
| 9.164966, -79.837098 | +|||||
| 9.164966, -79.837098 | +|||||
| 9.164966, -79.837098 | +|||||
| 9.164966, -79.837098 | +|||||
| 9.164966, -79.837098 | +|||||
| 9.164966, -79.837098 | +|||||
| 9.164966, -79.837098 | +|||||
| 10.078445, -83.573754 | +|||||
| 9.989971, -83.763587 | +|||||
| 9.831438, -83.832067 | +|||||
| 8.490081, -82.620285 | +|||||
| 8.793367, -82.188833 | +|||||
| 10.974153, -84.343499 | +|||||
| 10.974309, -84.338318 | +|||||
| 10.972540, -84.338990 | +|||||
| 11.316590, -84.612565 | +|||||
| 9.164966, -79.837098 | +|||||
| 10.250000, -83.450000 | +|||||
| 10.250000, -83.450000 | +|||||
| 8.399580, -83.313750 | +|||||
| 9.164966, -79.837098 | +|||||
| 9.164966, -79.837098 | +
| 1 | +Malar suture complete and distinctly impressed; medio-posterior depression of mesoscutum present; second metasomal suture distinctly crenulate | +
+ |
+
| - | +Malar suture incomplete and obsolescent; medio-posterior depression of mesoscutum absent; second metasomal suture finely sculptured, without distinct crenulae | +
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+sehr feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +hell | +Salzzeichen | +1 | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +unter-montan und ober-kollin | +||
|
+ |
+
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+sehr feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +sauer (pH 3.5-6.5) | +Temperaturzahl T | +
+subalpin (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +frisch | +Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+kollin (
+ |
+||
|
+ |
+
+ |
+
+ |
+kontinental (sehr niedrige Luftfeuchtigkeit, sehr grosse Temperaturschwankungen, kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+116350 | +
| = | +
+ |
+
+ |
+156a | +
| 1 | +
+Pedipalp tibiotarsal claw present; 2 pedipalp tibiotarsal spine-like tubercles present (
+ |
+
+ |
+
| - | +
+Pedipalp tibiotarsal claw absent; 2 pedipalp tibiotarsal spine-like tubercles absent (
+ |
+
+ |
+
| - | +
+ |
+
+ |
+
|---|---|---|
| Body size | +Male size 4.56-4.80 mm long, 3.20-3.42 times as long as wide; | +Male size 3.32-3.40 mm long, 2.78-3.00 times as long as wide; | +
| Female size 4.8-5.34 mm long, 3.38-3.43 times as long as wide. | +Female size 3.44-3.58 mm long, 2.87-2.93 times as long as wide. | +|
| Frons | +Male frons almost flat, with shallower, irregularly placed punctures; circularly concave in median line. | +Male frons coarser, with deeper, irregularly placed punctures; linearly concave in median line. | +
| Female frons almost flat, without concave around median line. | +Female frons concave forming a big, circular impression around concave median line. | +|
| Elytra | +Male without lateral emargination at declivity base, semicircular lateral borders with serrated, lateral tubercles. | +Male with lateral emargination at declivity base, semicircular lateral borders rounded, without distinct serrated, lateral tubercles. | +
| KEINE ANGABE | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+405280 | +
| = | +
+ |
+
+ |
+2912 | +
|
+4.3.3 - Rostseggenhalde (
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+unter-alpin, supra-subalpin und ober-subalpin (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+194100 | +
| = | +
+ |
+
+ |
+1191 | +
| = | +
+ |
+
+ |
+677 | +
| = | +
+ |
+
+ |
+677 | +
| = | +
+ |
+
+ |
+194100 | +
| = | +
+ |
+
+ |
+194100 | +
| = | +
+ |
+
+ |
+924 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +-- | +|
| Mittelland (MP) | +-- | +|
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
|
+ |
+L - Leitart | +weitere Informationen | +
| Character | +
+ |
+
+ |
+
|---|---|---|
| Adaxial leaf blade | +bullate | +not bullate | +
| Abaxial leaf blade | +reticulate veinlets conspicuous | +veinlets inconspicuous | +
| Peduncle | +brown-purple | +green | +
| Corolla | +purple-red to purple with white spots on the face | +white, purple with purple or white striate on the face | +
| Corolla adaxial lip | +lobes triangular, apex acute | +lobes oblong, apex rounded | +
| Corolla abaxial lip | +lobe margin with 4-10 long teeth, apex acute | +lobe margin nearly entire, apex rounded | +
| Stamens | +adaxial 0.5-1.4 cm long, abaxial 0.8-1.8 cm long | +adaxial 2.0-2.6 cm long, abaxial 2.3-2.7 cm long | +
| Anthers | +thecae confluent at apex | +thecae not confluent | +
| Style | +glabrous | +glandular pubescent | +
| Characters | +
+ |
+
+ |
+
|---|---|---|
| Leaf epidermal cell | +adaxial and abaxial cuticular membranes smooth, anticlinal walls sinuate without knobs | +adaxial and abaxial cuticular membranes striate, anticlinal walls sinuate with knobs | +
| Leaf epidermal trichome | +both adaxial and abaxial membranes rugulate | +adaxial membranes smooth, abaxial membranes rugulate | +
| Stomata | +
+smaller, 34.8
+ |
+
+larger, 37.7
+ |
+
| Style epidermal cells | +quadrilateral or polygonal, cuticular membranes many granular derivatives | +long irregulate, cuticular membranes without granular derivative | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
| 4.3 - Gebirgs-Magerrasen | +
|
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +sehr hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +
+unter-alpin, supra-subalpin und ober-subalpin (
+ |
+||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+8400 | +
| = | +
+ |
+
+ |
+2762 | +
| = | +
+ |
+
+ |
+2942 | +
| = | +
+ |
+
+ |
+2942 | +
| = | +
+ |
+
+ |
+8400 | +
| = | +
+ |
+
+ |
+197 | +
| = | +
+ |
+
+ |
+185 | +
| = | +
+ |
+
+ |
+8400 | +
| = | +
+ |
+
+ |
+2304 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+ |
+|
| Mittelland (MP) | +-- | +|
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +
| - | +
+ |
+
+ |
+
|---|---|---|
| Habit | +erect | +erect to hanging | +
| Plant height | +5-9 cm long | +10-14 cm long | +
| Stem surface | +glabrous | +pubescent | +
| Leaf - relative size | +leaves not uniform on stem; central leaves relatively larger compared with basal and apical leaves | +leaves relatively uniform on stem | +
| Leaf margin | +glabrous | +ciliate | +
| Pedicel length | +1.5-4 cm long | +1-1.4 cm long | +
| Flower length | +0.8-1.5 cm long | +1.5-1.6 cm long | +
| Flower width during anthesis | +2-3 cm wide | +1.5-2 cm wide | +
| Sepal surface | +conspicuously pubescent | +sparsely pubescent at base only | +
| Sepal horn | +white to reddish horn at apex | +green to black horn at apex | +
| Petal coloration (=inner tepals) | +yellow with a red stripe and green tip. | +green suffused with yellow with three prominent red spots at the base | +
| Distribution | +Peru and Bolivia | +Ecuador (Tungurahua) | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+sehr feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +schattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter) | +||
| Taxon | +Sexes known | +Distribution | +Elevation (m.s.l.) | +Male | +Female | +|
|---|---|---|---|---|---|---|
| Antennal flagellum | +Spines of SV | +Antennal flagellum | +||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
|
+ |
+||||||
| guarnensis | +||||||
|
+ |
+||||||
|
+ |
+||||||
| + |
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +-- | +Lichtzahl L | +-- | +Salzzeichen | +-- | +
| Reaktionszahl R | +-- | +Temperaturzahl T | +-- | +||
|
+ |
+-- | +
+ |
+-- | +||
|
+ |
+
| 2 | +
|
+ |
+
| 3 | +
|
+ |
+
| 5 | +
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
| Species | +T3-R1 | +SMS | +
|---|---|---|
|
+ |
+||
|
+ |
+||
|
+ |
+||
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
|
+ |
+
|
+1.
+ |
+|
|
+2.
+ |
+
+ |
+
|
+2*.
+ |
+
+ |
+
|
+1*.
+ |
+|
|
+3. Stacheln gerade; Fruchtknoten zottig behaart;
+ |
+
+ |
+
|
+3*. Stacheln wenig gebogen; Fruchtknoten locker behaart bis kahl;
+ |
+
+ |
+
| 1 | +Ventrianal shield with three pairs of opisthogastric setae (JV1, JV2, ZV2) and three circum-anal setae; endopodal platelets II-III and III-IV connected, bearing metasternal setae (st4); genua II and III without ventral setae | +
+ |
+
| - | +Ventrianal shield absent or with at most two pairs of opisthogastric setae (JV2, ZV2) and three circum-anal setae; endopodal platelets II-III and III-IV separate or absent, setae st4 on soft integument; genua II and III each with at least one ventral seta | +
+ |
+
| 2 | +Dorsal setae simple, short, mostly smooth and pointed, needle-shaped; peritrematal shield reduced, not reaching the anterior tip of peritreme; peritrematal shields or peritremes not connected to dorsal shield; ambulacra of legs usually without claws | +
+ |
+
| - | +At least some of dorsal setae modified, expanded, thickened or flattened, often elongate and pilose, with acute or obtuse apex, lance-shaped or leaf-shaped; peritrematal shield developed along whole length of peritreme; anterior section of peritrematal shield wide and completely fused to dorsal shield; ambulacral claws in legs II-IV present or absent | +
+ |
+
| 3 | +Dorsal shield with 29 pairs of setae, dorsolateral soft integument besides the shield without setae; insemination ducts fused and entering sacculus foeminus by a common neck-like process of the sacculus before entering spermatheca | +
+ |
+
| - | +Dorsal shield with 18-25 pairs of setae, dorsolateral soft integument besides the shield with 3-20 pairs of setae; insemination ducts unfused and entering sacculus foeminus separately | +
+ |
+
| 4 | +Genu III with nine setae, including one posterolateral seta (2-2/1, 2/1-1) | +
+ |
+
| - | +Genu III with ten setae, including two posterolateral setae (2-2/1, 2/1-2) | +
+ |
+
| 5 | +Posterior venter with four pairs of setae on soft integument (JV1, JV5) and on ventrianal shield (JV2, ZV2), JV3 and JV4 absent; dorsal shield with 21 pairs of setae | +
+ |
+
| - | +Posterior venter with five or six pairs of setae (JV1-JV5, ZV2; JV4 sometimes absent); ventrianal shield (if present) with at most one pair of opisthogastric setae (JV3); dorsal shield with 22-30 pairs of setae | +
+ |
+
| 6 | +Dorsal shield ornamented with tubercles, or spines arranged in rows, bearing 22-28 pairs of setae; some dorsal shield setae (6-9 pairs) conspicuously thickened and greatly lengthened | +
+ |
+
| - | +Dorsal shield otherwise sculptured, never with tubercles or rows of spines in medial surface, bearing 28-30 pairs of setae; dorsal shield setae usually subequal or less differing in length | +
+ |
+
| 7 | +Sternal shield with three pairs of setae (st1-st3); genu IV with ten setae, including two ventral setae (2-2/1, 3/1-1); dorsal setae pinnate | +
+ |
+
| - | +Sternal shield with two pairs of setae (st1, st2); genu IV with at most nine setae, including one ventral seta (2-2/1, 3/0-1); dorsal setae usually otherwise formed | +
+ |
+
| 8 | +
+Some dorsal shield setae (6-7 pairs) inserted on distinct protuberances, these setae longer, thicker and more heavily pilose than those on flat surface; tibia IV with ten setae, including two posterolateral setae (2-2/1, 2/1-2); fixed digit of chelicera slender, straight, with three sharp teeth and widened bilobed tooth close to terminal hook (Plate
+ |
+
+ |
+
| - | +Dorsal shield setae not inserted on protuberances; tibia IV with nine setae, including one posterolateral seta (2-2/1, 2/1-1); fixed digit of chelicera more robust, at least slightly curved, without subterminal bilobed tooth | +
+ |
+
| 9 | +Cheliceral digits slender and terminally hooked: fixed digit edentate or with one weak tooth, having hyaline lobed appendage; corniculi slender and convergent, with undivided and pointed apex | +
+ |
+
| - | +Cheliceral digits robust and straight: fixed digit well dentate, normally with three teeth on proximal masticatory area (rarely with one robust tooth or two denticles), never with hyaline lobed appendage; corniculi stouter and parallel, with bi- or trifid apex | +
+ |
+
| 10 | +Tarsus I without ambulacrum and claws, bearing greatly lengthened sensory setae; posterior venter with anal shield bearing only three circum-anal setae | +
+ |
+
| - | +Tarsus I with well developed ambulacrum, claws present or absent; posterior venter with ventrianal shield bearing normally two pairs of opisthogastric setae (JV2, JV3) and three circum-anal setae | +
+ |
+
| 11 | +Fixed digit of chelicera with three proximal denticles; ambulacral claws on legs reduced or absent; epistome with bifurcate apex; dorsal setae relatively short, leaf-shaped, oblanceolate, spatulate or obovate, with rounded apex; anterior hypostomal setae (h1) thickened, curved and attenuate medially, subfalcate; anterior margin of coxa II produced into spiniform process | +
+ |
+
| - | +Fixed digit of chelicera with four proximal denticles; ambulacral claws on legs present; epistome with undivided apex; dorsal setae relatively longer, tubiform or leaf-shaped, with pointed apex; setae h1 thin or thickened, almost straight and regularly tapered; anterior margin of coxa II smooth | +
+ |
+
|
+ |
+
+ |
+
+ |
+
+ |
+|||||
| males | +females | +males | +females | +males | +females | +males | +females | +|
| (n = 5) | +(n = 5) | +(n = 5) | +(n = 5) | +(n = 10) | +(n = 10) | +(n = 1) | +(n = 1) | +|
| Standard length (mm) | +40.9-44.5 | +31.0-35.5 | +36.8-73.2 | +34.2-64.1 | +28.4-43.4 | +23.6-38.2 | +38.6 | +28.4 | +
| % of standard length | +||||||||
| Body depth | +38.5-40.2 | +38.4-40.9 | +35.8-42.7 | +37.6-44.0 | +37.8-39.6 | +38.0-41.4 | +37.4 | +38.1 | +
| Caudal peduncle depth | +16.5-17.1 | +15.1-17.0 | +14.4-16.3 | +13.5-17.2 | +14.1-16.0 | +13.0-14.6 | +14.6 | +14.9 | +
| Pre-dorsal length | +44.9-50.6 | +55.8-62.7 | +42.4-49.2 | +55.2-59.3 | +46.6-49.8 | +57.1-61.7 | +49.4 | +65.6 | +
| Pre-pelvic length | +44.2-46.6 | +49.1-53.8 | +40.2-42.4 | +47.0-51.5 | +41.0-45.3 | +49.6-51.4 | +42.5 | +49.2 | +
| Length of dorsal-fin base | +38.7-43.6 | +22.6-29.3 | +39.0-44.8 | +23.9-30.2 | +37.9-41.1 | +23.1-29.2 | +40.3 | +25.1 | +
| Length of anal-fin base | +39.9-45.3 | +27.3-29.9 | +41.2-43.7 | +27.2-33.7 | +39.9-42.8 | +25.1-27.4 | +40.2 | +27.8 | +
| Caudal-fin length | +33.1-35.8 | +33.4-37.0 | +34.6-35.4 | +32.6-37.0 | +34.6-40.4 | +33.0-37.1 | +36.1 | +35.3 | +
| Pectoral-fin length | +26.6-32.6 | +26.0-28.7 | +23.9-26.9 | +24.5-27.3 | +25.3-31.3 | +24.0-28.2 | +28.9 | +27.1 | +
| Pelvic-fin length | +8.6-10.9 | +10.8-12.0 | +9.1-9.5 | +10.1-11.9 | +9.7-11.6 | +10.1-12.8 | +10.5 | +12.4 | +
| Head length | +28.3-30.6 | +30.7-32.6 | +26.8-28.5 | +28.6-30.1 | +27.6-30.8 | +27.9-31.8 | +28.2 | +32.0 | +
| % of head length | +||||||||
| Head depth | +98.4-117.3 | +94.1-103.2 | +101.0-117.6 | +97.8-116.1 | +106.9-121.1 | +101.6-125.7 | +108.8 | +102.7 | +
| Head width | +74.7-78.4 | +65.9-69.7 | +69.2-78.0 | +69.6-82.5 | +65.0-70.2 | +62.5-72.2 | +69.4 | +67.0 | +
| Snout length | +12.5-14.7 | +11.9-13.5 | +15.7-16.3 | +13.5-16.1 | +13.2-16.6 | +13.2-15.4 | +15.9 | +14.7 | +
| Lower jaw length | +23.6-27.3 | +18.6-23.1 | +20.2-21.2 | +16.0-17.3 | +17.6-19.4 | +15.6-17.6 | +20.9 | +18.5 | +
| Eye diameter | +27.3-29.2 | +29.6-31.3 | +24.7-30.2 | +25.7-32.9 | +27.8-33.5 | +28.6-33.1 | +29.0 | +29.6 | +
|
+ |
+
+ |
+
+ |
+
+ |
+|||||
| males | +females | +males | +females | +males | +females | +males | +females | +|
| (n = 5) | +(n = 5) | +(n = 5) | +(n = 5) | +(n = 7) | +(n = 7) | +(n = 7) | +(n = 5) | +|
| Standard length (mm) | +40.8-44.9 | +32.8-35.3 | +32.7-41.1 | +32.1-35.9 | +30.0-43.1 | +24.7-31.6 | +33.6-46.2 | +31.9-36.4 | +
| % of standard length | +||||||||
| Body depth | +40.6-43.9 | +37.7-41.5 | +37.8-41.9 | +37.7-42.7 | +33.9-38.4 | +30.7-36.8 | +37.6-42.7 | +37.0-42.1 | +
| Caudal peduncle depth | +15.2-16.0 | +13.9-15.4 | +15.5-16.6 | +13.9-16.4 | +12.6-14.3 | +11.7-13.6 | +144-16.6 | +134-14.8 | +
| Pre-dorsal length | +47.3-49.4 | +56.5-61.6 | +42.5-47.3 | +58.7-61.7 | +44.6-45.3 | +56.1-60.5 | +41.8-44.8 | +56.7-59.7 | +
| Pre-pelvic length | +44.1-46.1 | +49.8-52.8 | +42.6-44.4 | +50.1-54.3 | +42.1-46.1 | +47.8-52.2 | +40.0-45.0 | +47.6-50.8 | +
| Length of dorsal-fin base | +38.4-43.6 | +23.8-26.2 | +39.6-42.7 | +23.0-25.8 | +39.6-42.3 | +22.2-26.4 | +41.4-45.3 | +23.1-25.9 | +
| Length of anal-fin base | +40.3-45.9 | +27.3-30.0 | +40.3-45.2 | +27.0-29.8 | +34.4-38.4 | +25.3-27.6 | +39.0-45.2 | +24.7-27.7 | +
| Caudal-fin length | +34.1-38.3 | +32.9-37.9 | +34.1 | +33.8 | +33.2-37.1 | +32.3-36.8 | +36.9-41.6 | +35.0-37.2 | +
| Pectoral-fin length | +28.2-31.0 | +26.8-27.7 | +26.3 | +25.5 | +24.2-29.7 | +22.7-26.8 | +27.0-29.0 | +25.2-28.3 | +
| Pelvic-fin length | +10.0-11.8 | +10.9-12.4 | +9.9 | +11.5 | +8.9-11.3 | +9.8-11.1 | +10.3-11.7 | +10.7-12.4 | +
| Head length | +29.4-30.2 | +30.2-32.7 | +28.3-29.5 | +30.7-32.2 | +27.6-31.4 | +28.1-31.8 | +26.8-29.6 | +29.6-31.7 | +
| % of head length | +||||||||
| Head depth | +108.5-119.0 | +98.2-101.4 | +97.5-108.1 | +93.2-100.3 | +97.1-108.7 | +88.6-100.1 | +105.8-115.5 | +97.6-101.5 | +
| Head width | +72.4-75.9 | +69.9-73.5 | +70.4-73.1 | +69.4-73.0 | +62.5-71.2 | +59.3-63.4 | +65.5-70.8 | +64.3-67.0 | +
| Snout length | +13.5-15.3 | +12.7-14.5 | +14.7 | +14.3 | +13.8-15.4 | +13.0-15.3 | +14.8-16.8 | +13.4-14.5 | +
| Lower jaw length | +28.0-32.9 | +28.0-29.5 | +21.2 | +17.6 | +17.5-20.0 | +17.8-19.8 | +17.9-21.0 | +15.8-18.9 | +
| Eye diameter | +24.9-29.9 | +27.9-30.0 | +27.2-29.4 | +28.8-32.4 | +27.6-30.7 | +29.6-32.8 | +29.1-34.1 | +30.7-32.1 | +
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+sehr feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +hell | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +unter-montan und ober-kollin | +||
|
+ |
+
+sehr
+ |
+
+ |
+subozeanisch (hohe Luftfeuchtigkeit, geringe Temperaturschwankungen, eher milde Winter) | +||
| ABLBNBNFNSONPEQCSKYT | +
| SaskatchewanDLCDLCLFCLFC | +
|
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +neutral bis basisch (pH 5.5-8.5) | +Temperaturzahl T | +warm-kollin | +||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
|
+ |
+
|
+6.2.3 - Waldmeister-Buchenwald (
+ |
+
|
+ |
+
|
+6.3.6 - Saurer Eichenmischwald (
+ |
+
| 6.3.7 - Kastanienwald | +
|
+6.4.4 - Kalkarmer
+ |
+
|
+ |
+
+ |
+
+ |
+|||
|---|---|---|---|---|---|
| Feuchtezahl F | +
+feucht; Feuchtigkeit stark wechselnd (mehr als
+ |
+Lichtzahl L | +halbschattig | +Salzzeichen | +-- | +
| Reaktionszahl R | +schwach sauer bis neutral (pH 4.5-7.5) | +Temperaturzahl T | +
+kollin (
+ |
+||
|
+ |
+
+ |
+
+ |
+
+subozeanisch bis subkontinental (mittlere Luftfeuchtigkeit,
+ |
+||
| Relation | +Nom | +Referenzwerke | +No | +
|---|---|---|---|
| = | +
+ |
+
+ |
+336100 | +
| = | +
+ |
+
+ |
+249 | +
| = | +
+ |
+
+ |
+331 | +
| = | +
+ |
+
+ |
+331 | +
| = | +
+ |
+
+ |
+336100 | +
| = | +
+ |
+
+ |
+834 | +
| = | +
+ |
+
+ |
+735 | +
| = | +
+ |
+
+ |
+336100 | +
| = | +
+ |
+
+ |
+143 | +
| Biogeografische Regionen | +Status | +Kriterien IUCN | +
|---|---|---|
| Jura (JU) | +
+nicht
+ |
+|
| Mittelland (MP) | +
+nicht
+ |
+|
| Alpennordflanke (NA) | +
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
|
+ |
+
+nicht
+ |
+|
| Westliche Zentralalpen (WA) | +
+nicht
+ |
+
|
+Keine nationale
+ |
+
| Kein internationaler, nationaler oder kantonaler Schutz | +