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<document id="24F7FDE0F9CC0F3DD8E6B44D71DDC5F4" ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e8013" ID-PMC="PMC4910507" ID-Pensoft-Pub="1314-2828--8013" ID-PubMed="27346954" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828--e8013" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="Checklist of British and Irish Hymenoptera - Chalcidoidea and Mymarommatoidea" checkinTime="1553126186093" checkinUser="pensoft" docAuthor="Dale-Skey, Natalie, Askew, Richard R., Noyes, John S., Livermore, Laurence &amp; Broad, Gavin R." docDate="2016" docId="F177384C6DF3257BD7620CEAB098D273" docLanguage="en" docName="BiodivDatJour 4: e8013" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e8013" docTitle="Melancistrus specularis Graham 1969" docType="treatment" docVersion="6" lastPageNumber="8013" masterDocId="FF97FFC8521CFFF7FFB6FF94962EFFD0" masterDocTitle="Checklist of British and Irish Hymenoptera - Chalcidoidea and Mymarommatoidea" masterLastPageNumber="8013" masterPageNumber="8013" pageNumber="8013" updateTime="1668123632585" updateUser="ExternalLinkService">
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<mods:title id="DA43C23E0F48244712338262F5CB482F">Checklist of British and Irish Hymenoptera - Chalcidoidea and Mymarommatoidea</mods:title>
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<mods:namePart id="E5FA25F0273CC9810C5C040F7E247F52">Dale-Skey, Natalie</mods:namePart>
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<taxonomicName id="8D18FBA56E5ACA05F2FA9B6FD736D4A7" ID-CoL="3Z68R" authority="Graham, 1969" authorityName="Graham" authorityYear="1969" class="Insecta" family="Pteromalidae" genus="Melancistrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melancistrus specularis" order="Hymenoptera" pageId="0" pageNumber="8013" phylum="Arthropoda" rank="species" species="specularis">Melancistrus specularis Graham, 1969</taxonomicName>
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<paragraph id="7378C839C782AAFA9D0E9470F010F936" pageId="0" pageNumber="8013">Distribution</paragraph>
<paragraph id="565750E4D0498235B1051386AF447907" pageId="0" pageNumber="8013">Scotland</paragraph>
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<taxonomicName id="BEDE41FF5B41FFE363BD03571B92FD8D" ID-CoL="7b5f7909-db8d-4b31-867e-5275e1f69064" box="[151,409,569,595]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="485" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B41FFE363BD03571B92FD8D" bold="true" box="[151,409,569,595]" italics="true" pageId="2" pageNumber="485">Batillipes brasiliensis</emphasis>
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<paragraph id="79613A7C5B41FFE363BD03321B67FDAA" blockId="2.[151,509,569,629]" box="[151,364,604,629]" pageId="2" pageNumber="485">
(
<figureCitation id="E1E526F95B41FFE363B503321B0FFDAA" box="[159,260,604,629]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[151,250,1850,1872]" captionStartId-1="4.[151,250,1691,1713]" captionTargetBox-0="[153,1436,195,1828]" captionTargetBox-1="[153,1432,195,1655]" captionTargetId-0="figure@3.[151,1436,193,1828]" captionTargetId-1="figure@4.[151,1436,193,1669]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." captionText-1="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri-0="https://zenodo.org/record/400155/files/figure.png" httpUri-1="https://zenodo.org/record/400156/files/figure.png" pageId="2" pageNumber="485">Figs 2–3</figureCitation>
; Table 1)
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B41FFE363BD03CA1987FC66" pageId="2" pageNumber="485" type="diagnosis">
<paragraph id="79613A7C5B41FFE363BD03CA1987FC66" blockId="2.[151,1437,676,2033]" pageId="2" pageNumber="485">
<emphasis id="4BAAE66E5B41FFE363BD03CA1B01FD63" bold="true" box="[151,266,676,701]" pageId="2" pageNumber="485">Diagnosis</emphasis>
. Small sized
<taxonomicName id="BEDE41FF5B41FFE3628C03CA1819FD62" box="[422,530,676,700]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="485" phylum="Tardigrada" rank="genus">
<emphasis id="4BAAE66E5B41FFE3628C03CA1819FD62" box="[422,530,676,700]" italics="true" pageId="2" pageNumber="485">Batillipes</emphasis>
</taxonomicName>
with tubular undivided primary clavae and papillar secondary clavae. Cephalic cirri with swollen distal tips. Sensorial spines on all legs. Toes 3 and 4 of legs IV with different lengths. Distinct head separated from the body by a neck constriction followed by well-developed lateral processes (“auricles”). Small lateral processes between legs I–III each with a small digit-shaped apex. Processes between legs III and IV fringed with digit-shaped expansions. The caudal apparatus consists of a wide ala-like cuticular expansion covered with detritus. A small semicircular cuticular projection is also present on the coxal region of each leg. Punctate cuticle comprised of small pillars. Rosette-like female gonopore surrounded by a four-valved cuticular structure. Males with circular gonopore with a cuticular crescent-shaped fold.
</paragraph>
</subSubSection>
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<paragraph id="79613A7C5B41FFE363ED02AA19E1FC03" blockId="2.[151,1437,676,2033]" box="[199,1002,964,989]" pageId="2" pageNumber="485">
<materialsCitation id="C9B630215B41FFE363ED02AA19E1FC03" ID-GBIF-Occurrence="1440156676" box="[199,1002,964,989]" country="Brazil" latitude="-9.8625" location="Gunga Beach" longLatPrecision="21" longitude="-35.904724" pageId="2" pageNumber="485" specimenCount="1" stateProvince="Alagoas" typeStatus="holotype">
<emphasis id="4BAAE66E5B41FFE363ED02AA1B6EFC03" bold="true" box="[199,357,964,989]" pageId="2" pageNumber="485">
<typeStatus id="A66584DE5B41FFE363ED02AA1B09FC03" box="[199,258,964,989]" pageId="2" pageNumber="485">Type</typeStatus>
locality.
</emphasis>
<location id="7C016CA75B41FFE3624602AA180DFC03" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE3624602AA180DFC03" box="[364,518,964,989]" country="Brazil" latitude="-9.8625" longLatPrecision="21" longitude="-35.904724" name="Gunga Beach" pageId="2" pageNumber="485" stateProvince="Alagoas">Gunga Beach</location>
,
<collectingRegion id="BB1AF49E5B41FFE3613802AA187BFC03" box="[530,624,964,989]" country="Brazil" name="Alagoas" pageId="2" pageNumber="485">Alagoas</collectingRegion>
,
<collectingCountry id="01C97AEC5B41FFE3615102AA18B4FC03" box="[635,703,964,989]" name="Brazil" pageId="2" pageNumber="485">Brazil</collectingCountry>
(
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,
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).
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<paragraph id="79613A7C5B41FFE363ED02891F32FB06" blockId="2.[151,1437,676,2033]" pageId="2" pageNumber="485">
<emphasis id="4BAAE66E5B41FFE363ED02891B71FBDE" bold="true" box="[199,378,999,1024]" pageId="2" pageNumber="485">Type material.</emphasis>
Holotype: adult, female (slide CVII-80) collected at Gunga Beach, mounted in glycerol. Allotype: adult, male (slide TARD/UFRPE 02-11) collected at Amor Beach, mounted in glycerol. Paratypes: 8 females collected at Sossego Beach, mounted in polyvinyl alcohol (slides TARD/UFRPE
<date id="0D601CBC5B41FFE367BE055E1F35FB96" box="[1172,1342,1071,1096]" pageId="2" pageNumber="485" value="2006-02-03" valueMax="2006-02-02" valueMin="2006-02-03">02-03 to 02-06</date>
and 02- 14); 2 females and one four-toed larva collected at Forte Orange Beach, mounted in polyvinyl alcohol (slides TARD/UFRPE 02-14); 5 specimens collected at Gunga Beach (3 females, 1 male and 1 juvenile), mounted in glycerol (slides TARD/UFRPE 02-08, 02-09 and CVII-81); and 14 specimens collected at Amor Beach (4 females, 4 males and 6 juveniles), mounted in glycerol (slides TARD/UFRPE
<date id="0D601CBC5B41FFE360B405AE1E4CFB06" box="[926,1095,1216,1240]" pageId="2" pageNumber="485" value="2013-02-10" valueMax="2013-02-02" valueMin="2013-02-10">02-10 to 02-13</date>
, 02-24 and CVII-82).
</paragraph>
<paragraph id="79613A7C5B41FFE363ED058A1B8DFAB6" blockId="2.[151,1437,676,2033]" pageId="2" pageNumber="485">
<materialsCitation id="C9B630215B41FFE363ED058A1B89FAB6" ID-GBIF-Occurrence="1440156678" country="Portugal" county="Department" location="Faculty" municipality="Biology" pageId="2" pageNumber="485" specimenCount="2" stateProvince="Pernambuco" typeStatus="holotype">
<emphasis id="4BAAE66E5B41FFE363ED058A1B80FB23" bold="true" box="[199,395,1252,1277]" pageId="2" pageNumber="485">
<typeStatus id="A66584DE5B41FFE363ED058A1B09FB23" box="[199,258,1252,1277]" pageId="2" pageNumber="485">Type</typeStatus>
repository.
</emphasis>
The
<typeStatus id="A66584DE5B41FFE362E3058B1BF2FB23" box="[457,505,1253,1277]" pageId="2" pageNumber="485">type</typeStatus>
material (slides CVII-80–82) is deposited in the collection of the
<collectingCounty id="900042F05B41FFE367DD058B1F70FB22" box="[1271,1403,1253,1276]" pageId="2" pageNumber="485">Department</collectingCounty>
of
<collectingMunicipality id="9905A0065B41FFE363BD04691AF9FAFE" box="[151,242,1287,1312]" pageId="2" pageNumber="485">Biology</collectingMunicipality>
,
<location id="7C016CA75B41FFE3622C04691B51FAFE" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE3622C04691B51FAFE" box="[262,346,1287,1312]" country="Portugal" county="Department" municipality="Biology" name="Faculty" pageId="2" pageNumber="485" stateProvince="Pernambuco">Faculty</location>
of
<location id="7C016CA75B41FFE362BB04691BFDFAFE" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE362BB04691BFDFAFE" box="[401,502,1287,1312]" country="Portugal" county="Department" municipality="Biology" name="Sciences" pageId="2" pageNumber="485" stateProvince="Pernambuco">Sciences</location>
,
<location id="7C016CA75B41FFE361200469188BFAFE" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE361200469188BFAFE" box="[522,640,1287,1312]" country="Portugal" county="Department" municipality="Biology" name="University" pageId="2" pageNumber="485" stateProvince="Pernambuco">University</location>
of
<location id="7C016CA75B41FFE3619D046618FCFAC1" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE3619D046618FCFAC1" box="[695,759,1288,1311]" country="Portugal" county="Department" municipality="Biology" name="Porto" pageId="2" pageNumber="485" stateProvince="Pernambuco">Porto</location>
,
<collectingCountry id="01C97AEC5B41FFE3602004691967FAFE" box="[778,876,1287,1312]" name="Portugal" pageId="2" pageNumber="485">Portugal</collectingCountry>
, and all the other slides in the collection of Tardigrades—UFRPE (
<location id="7C016CA75B41FFE3628D0442182EFA9B" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE3628D0442182EFA9B" box="[423,549,1324,1349]" country="Portugal" county="Department" municipality="Biology" name="Laboratory" pageId="2" pageNumber="485" stateProvince="Pernambuco">Laboratory</location>
of
<location id="7C016CA75B41FFE3614B044218D6FA9B" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE3614B044218D6FA9B" box="[609,733,1324,1349]" country="Portugal" county="Department" municipality="Biology" name="Meiofauna" pageId="2" pageNumber="485" stateProvince="Pernambuco">Meiofauna</location>
,
<location id="7C016CA75B41FFE361D90443197CFA9A" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE361D90443197CFA9A" box="[755,887,1325,1348]" country="Portugal" county="Department" municipality="Biology" name="Department" pageId="2" pageNumber="485" stateProvince="Pernambuco">Department</location>
of
<location id="7C016CA75B41FFE3609904421E05FA9B" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE3609904421E05FA9B" box="[947,1038,1324,1349]" country="Portugal" county="Department" municipality="Biology" name="Biology" pageId="2" pageNumber="485" stateProvince="Pernambuco">Biology</location>
, Universidade Federal Rural de
<collectingRegion id="BB1AF49E5B41FFE363BD04211B2DFAB6" box="[151,294,1359,1384]" country="Brazil" name="Pernambuco" pageId="2" pageNumber="485">Pernambuco</collectingRegion>
,
<collectingCountry id="01C97AEC5B41FFE3621B04211B71FAB6" box="[305,378,1359,1384]" name="Brazil" pageId="2" pageNumber="485">Brazil</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B41FFE363ED041A1887FA26" pageId="2" pageNumber="485" type="etymology">
<paragraph id="79613A7C5B41FFE363ED041A19C0FA53" blockId="2.[151,1437,676,2033]" box="[199,971,1396,1421]" pageId="2" pageNumber="485">
<emphasis id="4BAAE66E5B41FFE363ED041A1B45FA53" bold="true" box="[199,334,1396,1421]" pageId="2" pageNumber="485">Etymology.</emphasis>
The name
<taxonomicName id="BEDE41FF5B41FFE362E1041A1847FA52" box="[459,588,1396,1420]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="485" phylum="Tardigrada" rank="species" species="brasiliensis">
<emphasis id="4BAAE66E5B41FFE362E1041A1847FA52" box="[459,588,1396,1420]" italics="true" pageId="2" pageNumber="485">brasiliensis</emphasis>
</taxonomicName>
refers to the
<emphasis id="4BAAE66E5B41FFE361F5041A197BFA52" box="[735,880,1396,1420]" italics="true" pageId="2" pageNumber="485">locus typicus</emphasis>
,
<collectingCountry id="01C97AEC5B41FFE36055041A19CCFA53" box="[895,967,1396,1421]" name="Brazil" pageId="2" pageNumber="485">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="79613A7C5B41FFE363ED04F91887FA26" blockId="2.[151,1437,676,2033]" pageId="2" pageNumber="485">
<emphasis id="4BAAE66E5B41FFE363ED04F91B88FA6E" bold="true" box="[199,387,1431,1456]" pageId="2" pageNumber="485">Ecological note.</emphasis>
<taxonomicName id="BEDE41FF5B41FFE362A304F71822FA6E" box="[393,553,1432,1456]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="485" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B41FFE362A304F71822FA6E" box="[393,553,1432,1456]" italics="true" pageId="2" pageNumber="485">B. brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B41FFE3611A04F91882FA6E" bold="true" box="[560,649,1431,1456]" pageId="2" pageNumber="485">
<taxonomicNameLabel id="50995B155B41FFE3611A04F91882FA6E" box="[560,649,1431,1456]" pageId="2" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found in shallow sublittoral medium to coarse, gravels and quartz sands, in low energy (Sossego Beach) and estuarine beaches (Forte Orange Beach and Gunga Beach) and in reef pools of high energy beaches (Amor Beach).
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B41FFE463ED076A1F2BFBCE" lastPageId="5" lastPageNumber="488" pageId="2" pageNumber="485" type="description">
<paragraph id="79613A7C5B41FFE363ED076A1F3DF876" blockId="2.[151,1437,676,2033]" pageId="2" pageNumber="485">
<emphasis id="4BAAE66E5B41FFE363ED076A181BF9C3" bold="true" box="[199,528,1540,1565]" pageId="2" pageNumber="485">
Description of the
<typeStatus id="A66584DE5B41FFE36289076A1807F9C3" box="[419,524,1540,1565]" pageId="2" pageNumber="485" type="holotype">holotype</typeStatus>
.
</emphasis>
Female with a body length 124 µm (142 µm including the caudal apparatus) and 45.4 µm wide between the third and fourth pair of legs (
<figureCitation id="E1E526F95B41FFE3606E07491980F99E" box="[836,907,1575,1600]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
A, B). Trapezoid head with eleven cephalic appendages: Internal cirri inserted dorsally on the frontal margin of the head are 18.6 µm long, bearing cirrophores (about 2.2 µm long). External cirri 12.7 µm long, with indistinct cirrophores, inserted more ventrally, near the lateral cirri and primary clavae. The median cirrus, with cirrophore, is 15.5 µm long (
<figureCitation id="E1E526F95B41FFE3677407FA1EAAF972" box="[1118,1185,1684,1709]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
C). Lateral cirri (26.6 µm long) are located dorsally in relation to the unconstricted tubular primary clavae (12.9 µm long). These two appendages share a common pedestal. A van der Land’s organ is present at the base of the primary clava that exhibits a terminal hole (
<figureCitation id="E1E526F95B41FFE3629907911BFCF8C6" box="[435,503,1791,1816]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
D). An indentation is present in the frontal edge of the head between the external cephalic cirrus and the pedestal bearing the primary clava and lateral cirrus. In the frontal edge of the head papillar secondary clavae (
<figureCitation id="E1E526F95B41FFE3624106291BBBF8BE" box="[363,432,1863,1888]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
D) are also well visible (major diameter 2.9 µm). All cephalic cirri, including external cirri, and lateral cirri have swollen tips. Eye spots not recognized. Ovoid pharyngeal bulb 19.7 µm long and 17.7 µm wide. Ventral mouth opening in a protruded triangular cone. Placoids not visible after slide mounting.
</paragraph>
<paragraph id="79613A7C5B41FFE463ED06DB1F17FE9E" blockId="2.[151,1437,676,2033]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="488" pageId="2" pageNumber="485">
Four pairs of ventro-lateral body processes are present: a blunt lateral expansion, 4.1 µm long, between the head and the first pair of legs (
<figureCitation id="E1E526F95B41FFE362C706B61824F82E" box="[493,559,2008,2033]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
D). Lateral processes are also present between all leg pairs. Between the first two pair of legs these blunt processes (6.0, 4.6 µm between legs I–II and II–III respectively) exhibit a thin digitshaped apex (
<figureCitation id="E1E526F95B47FFE5620506191B7AF851" box="[303,369,1911,1936]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="4" pageNumber="487">Fig. 3</figureCitation>
A). Between legs III and IV (
<figureCitation id="E1E526F95B47FFE5619706191908F851" box="[701,771,1911,1936]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="4" pageNumber="487">Figs 3</figureCitation>
B, C) lateral processes are larger, 5.8 µm long and 10.8 µm wide. In these lateral processes the cuticle is punctated except for the membranous fringed external edge constituted by five digit-shaped expansions (
<figureCitation id="E1E526F95B47FFE561BB06D118DEF809" box="[657,725,1983,2008]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="4" pageNumber="487">Fig. 2</figureCitation>
A, B). The caudal apparatus is an ala-like structure, protruding 17.5 µm and covering almost all the posterior edge of the body. The precise shape of this structure cannot be described because it is covered by detritus (
<figureCitation id="E1E526F95B46FFE4618201D218FEFF0A" box="[680,757,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="5" pageNumber="488">Figs 2</figureCitation>
A, B, 3 C, D). Small semicircular cuticular projections, decreasing in length from the first to the third leg (3.5, 3.3 and 2.1 µm respectively), are also present on the coxal region of legs I–III. This structure is particularly evident in the first leg pair (
<figureCitation id="E1E526F95B46FFE460D4006A1E4DFEC2" box="[1022,1094,260,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="5" pageNumber="488">Figs 2</figureCitation>
D, 3 A) and difficult to see in legs III. Lateral processes between legs and cuticular projections on legs often covered by detritus also.
</paragraph>
<caption id="2DA16AF45B40FFE263BD06541832F875" httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="3" pageNumber="486" targetBox="[153,1436,195,1828]" targetPageId="3">
<paragraph id="79613A7C5B40FFE263BD06541832F875" blockId="3.[151,1436,1850,1963]" pageId="3" pageNumber="486">
<emphasis id="4BAAE66E5B40FFE263BD06541B1AF891" bold="true" box="[151,273,1850,1872]" pageId="3" pageNumber="486">FIGURE 2</emphasis>
.
<taxonomicName id="BEDE41FF5B40FFE2620806541809F88E" box="[290,514,1850,1872]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="486" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B40FFE2620806541809F88E" box="[290,514,1850,1872]" italics="true" pageId="3" pageNumber="486">Batillipes brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B40FFE2612706541869F891" bold="true" box="[525,610,1850,1871]" pageId="3" pageNumber="486">
<taxonomicNameLabel id="50995B155B40FFE2612706541869F891" box="[525,610,1850,1871]" pageId="3" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr1) and leg projection of the first leg (lpI). Scale bars: A–B = 50 µm; C–D = 10 µm.
</paragraph>
</caption>
<caption id="2DA16AF45B47FFE563BD07F51F07F8F5" httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="4" pageNumber="487" targetBox="[153,1432,195,1655]" targetPageId="4">
<paragraph id="79613A7C5B47FFE563BD07F51F07F8F5" blockId="4.[151,1436,1691,1835]" pageId="4" pageNumber="487">
<emphasis id="4BAAE66E5B47FFE563BD07F51B05F96E" bold="true" box="[151,270,1691,1713]" pageId="4" pageNumber="487">FIGURE 3</emphasis>
.
<taxonomicName id="BEDE41FF5B47FFE5623607F51BF0F96F" box="[284,507,1691,1713]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="4" pageNumber="487" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B47FFE5623607F51BF0F96F" box="[284,507,1691,1713]" italics="true" pageId="4" pageNumber="487">Batillipes brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B47FFE5612807F5185EF96E" bold="true" box="[514,597,1691,1712]" pageId="4" pageNumber="487">
<taxonomicNameLabel id="50995B155B47FFE5612807F5185EF96E" box="[514,597,1691,1712]" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Detail of the two first legs of a paratype showing first body projections (pr1; pr2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr4) of the holotype; D. Toes (t1–t6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm.
</paragraph>
</caption>
<paragraph id="79613A7C5B46FFE463ED0022188EFE2A" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
Sensorial spines present on all legs, increasing in length from leg I to leg IV (5.1; 8.2; 9.6, and 12.8 µm long on leg I, II, III and IV respectively). All leg sense organs with swollen tips. The sense organ on leg IV divided into a pronounced cirrophore (5.8 µm long) and a spine-like distal portion (7.0 µm long) with a basal van der Land’s organ (
<figureCitation id="E1E526F95B46FFE463C300D91B25FE0E" box="[233,302,439,464]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
B). Sharply pointed and short (13.1 µm long) cirri E with small cirrophores. Cirri E are dorsally placed between lateral processes and leg IV.
</paragraph>
<paragraph id="79613A7C5B46FFE463ED00911819FCE6" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
Telescopic legs with toes of different lengths. On the first three pair of legs (
<figureCitation id="E1E526F95B46FFE4671500911E8FFDC6" box="[1087,1156,511,536]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="5" pageNumber="488">Fig. 2</figureCitation>
B), toe 2 is the shortest (considering toe 1 the most cephalically), toes 3 and 5 are the longest, and toes 1, 3 and 4 are medium sized (in leg II toe 1 is 7.0 µm long, toe 2 is 3.2 µm, toe 3 is 13.0 µm long, toe 4 is 6.2 µm, toe 5 is 14.1 µm, toe 6 is 7.3 µm). On the fourth pair of legs toes (
<figureCitation id="E1E526F95B46FFE462FD03021810FD5A" box="[471,539,620,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
D) conform the pattern of the D group of species proposed by
<bibRefCitation id="1D4F478D5B46FFE467D903021B56FD76" author="Kristensen" pageId="5" pageNumber="488" refString="Kristensen, R. M. &amp; Mackness, B. S. (2000) First record of the marine Tardigrade genus Batillipes (Arthrotardigrada: Batillipedidae) from South Australia with a description of a new species. Records of the South Australian Museum, 33 (2), 73 - 87." type="journal article" year="2000">Kristensen and Mackness (2000)</bibRefCitation>
, with toes 3 and 4 of different lengths (6.4 and 10.4 µm long respectively for toes 3 and 4). In those legs toes 1 and 6 are similar in length (respectively 11.3 and 11.0 µm long); toes 2 and especially toes 5 are the longest (16.3 and 20.5 µm long respectively). Toes have the distal stalk enlarged distally (about 2.2 µm wide), ovoid suction discs (3.3 long x 2.8 µm wide measured on toes of leg IV) with conspicuous braces and well developed peripheral thickenings.
</paragraph>
<paragraph id="79613A7C5B46FFE463ED022A1E2AFC7B" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">Dorsal cuticle punctated, about 10 pillars/10 µm (each pillar about 1.4 µm high), without evident transverse folds. However, the large amount of detritus covering the dorsal region of the body can hide those lines. Transverse cuticular folds are visible ventrally where the cuticle punctation is more delicate.</paragraph>
<paragraph id="79613A7C5B46FFE463ED02C11F2BFBCE" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
Rosette-shaped gonopore (
<figureCitation id="E1E526F95B46FFE462D202C11836FC16" box="[504,573,943,968]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
C) separated from the anus by a weakly defined groove. The gonopore is surrounded by a four-valved cuticular structure. Two larger divergent valves are located posteriorly and two narrow valves are located anteriorly to the gonopore (
<figureCitation id="E1E526F95B46FFE461B3029918D7FBCE" box="[665,732,1015,1040]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
E). The anus is 10.9 µm distant from the gonopore.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B46FFE463ED05721F2AFA43" pageId="5" pageNumber="488" type="discussion">
<paragraph id="79613A7C5B46FFE463ED05721BFEFAEE" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
<emphasis id="4BAAE66E5B46FFE463ED05721B30FBEB" bold="true" box="[199,315,1052,1077]" pageId="5" pageNumber="488">Remarks.</emphasis>
Sexual dimorphism is not evident. Males are similar to females in both qualitative and metric characters (table 1), except for their circular gonopore with a cuticular crescent shaped fold (
<figureCitation id="E1E526F95B46FFE467FD05511F16FB86" box="[1239,1309,1087,1112]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
F) located nearer the anus (4.5 µm distant for the anus in the allotypic male). Juveniles, with six toes on each leg but without a visible gonopore, are also similar to adults. The four-toed larva differs from the other juveniles and the adults in having reduced lateral projections without digit-shaped apices; in particular the one located between legs III and IV which lacks the fringed edge. Moreover, the caudal apparatus of this larva is constituted by a conical process without any membranous ala. However, small projections on the coxal region of the three first legs, characteristic of the new species, are present.
</paragraph>
<paragraph id="79613A7C5B46FFE463ED04521F2AFA43" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
On the other hand, among adults, the development of the caudal apparatus (see
<figureCitation id="E1E526F95B46FFE4671404521E74FA8A" box="[1086,1151,1340,1365]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
B, C, F) and the shape of the fourth lateral projection (between legs III and IV), especially in the number (from 2 to 7) of digit-shaped membranous expansions, are very variable (
<figureCitation id="E1E526F95B46FFE461AE04EA18C0FA42" box="[644,715,1412,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Figs 3</figureCitation>
B, C). Variability in other projections is not relevant.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B46FFE763ED04C91D2CFD73" lastPageId="6" lastPageNumber="489" pageId="5" pageNumber="488" type="diagnosis">
<paragraph id="79613A7C5B46FFE463ED04C919C0F8DA" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
<emphasis id="4BAAE66E5B46FFE463ED04C91BDAFA1E" bold="true" box="[199,465,1447,1472]" pageId="5" pageNumber="488">Differential diagnosis.</emphasis>
The taxonomy of the genus
<taxonomicName id="BEDE41FF5B46FFE4600204C61998FA1E" box="[808,915,1448,1472]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="genus">
<emphasis id="4BAAE66E5B46FFE4600204C61998FA1E" box="[808,915,1448,1472]" italics="true" pageId="5" pageNumber="488">Batillipes</emphasis>
</taxonomicName>
is particular problematic due to the reduced number of taxonomic characters traditionally used to distinguish species. Intraspecific variability exhibited by some of those characters and deformations caused by slide mounting are also a source of taxonomic difficulties (
<emphasis id="4BAAE66E5B46FFE466AB049E1F9EF9D6" box="[1409,1429,1520,1544]" italics="true" pageId="5" pageNumber="488">cf</emphasis>
.
<bibRefCitation id="1D4F478D5B46FFE463BD077A1B34F9F3" author="McKirdy" box="[151,319,1556,1581]" pageId="5" pageNumber="488" refString="McKirdy, D. J. (1975) Batillipes (Heterotardigrada): Comparison of six species from Florida (USA) and a discussion of taxonomic characters within the genus. Memorie dell'Istituto Italiano di Idrobiologia, 32 (Supplement), 177223." type="journal article" year="1975">McKirdy 1975</bibRefCitation>
;
<bibRefCitation id="1D4F478D5B46FFE46266077A180EF9F2" author="Kristensen" box="[332,517,1556,1581]" pageId="5" pageNumber="488" refString="Kristensen, R. M. (1978) Notes on Marine Heterotardigrades. 1. Description of two Batillipes species, using the electron microscope. Zoologischer Anzeiger, 200, 1 - 17." type="journal article" year="1978">Kristensen 1978</bibRefCitation>
;
<bibRefCitation id="1D4F478D5B46FFE4613B077B1951F9F2" author="Morone" box="[529,858,1556,1581]" pageId="5" pageNumber="488" refString="Morone De Lucia, R. M., D'Addabo Gallo, M., Grimaldi de Zio, S. (1988) Descrizione di due nuove specie di Batillipedidae (Tardigrada: Heterotardigrada). Cahiers de Biologie Marine, 29, 361 - 373." type="journal article" year="1988">
Morone De Lucia
<emphasis id="4BAAE66E5B46FFE461CA077B191AF9F2" box="[736,785,1556,1580]" italics="true" pageId="5" pageNumber="488">et al</emphasis>
. 1988
</bibRefCitation>
; Villora Moreno and de
<bibRefCitation id="1D4F478D5B46FFE46741077A1F49F9F2" author="Zio" box="[1131,1346,1556,1581]" pageId="5" pageNumber="488" refString="Villora-Moreno, S. &amp; de Zio Grimaldi, S. (1993) Redescription and ecology of Batillipes phreaticus Renaud-Debyser, 1959 (Arthrotardigrada, Batillipedidae) in the gulf of Valencia (western Mediterranean). Cahiers de Biologie Marine, 34, 387 - 399." type="journal article" year="1993">Zio Grimaldi 1993</bibRefCitation>
). In this genus, one of the more reliable taxonomic characters is the relative dimension of toes. This indicator was introduced by
<bibRefCitation id="1D4F478D5B46FFE4621307321BEAF9AA" author="Pollock" box="[313,481,1628,1653]" pageId="5" pageNumber="488" refString="Pollock, L. W. (1970) Batillipes dicrocercus n. sp., Stygarctus granulatus n. sp. and other tardigrade from Woods Hole, Massachussets, U. S. A. Transactions of the American Microscopical Society, 89 (1), 38 - 52." type="journal article" year="1970">Pollock (1970)</bibRefCitation>
and modified by
<bibRefCitation id="1D4F478D5B46FFE4618207321E12F9AA" author="Kristensen" box="[680,1049,1628,1653]" pageId="5" pageNumber="488" refString="Kristensen, R. M. &amp; Mackness, B. S. (2000) First record of the marine Tardigrade genus Batillipes (Arthrotardigrada: Batillipedidae) from South Australia with a description of a new species. Records of the South Australian Museum, 33 (2), 73 - 87." type="journal article" year="2000">Kristensen and Mackness (2000)</bibRefCitation>
who proposed four patterns of toe lengths on the fourth foot, corresponding to four species groups.
<taxonomicName id="BEDE41FF5B46FFE4604707EE1E58F946" box="[877,1107,1664,1688]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="brasiliesis" status="sp. nov.">
<emphasis id="4BAAE66E5B46FFE4604707EE1E58F946" box="[877,1107,1664,1688]" italics="true" pageId="5" pageNumber="488">Batillipes brasiliesis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B46FFE4677007111EBFF946" bold="true" box="[1114,1204,1663,1688]" pageId="5" pageNumber="488">
<taxonomicNameLabel id="50995B155B46FFE4677007111EBFF946" box="[1114,1204,1663,1688]" pageId="5" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having toes 3 and 4 of different lengths and respectively different from toe 1 and toe 2, belongs to the D group of species. Only five species exhibit this toe pattern on leg IV:
<taxonomicName id="BEDE41FF5B46FFE4615A07A71AD8F8DA" authority="Morone De Lucia, D'Addabbo Gallo and Grimaldi de Zio, 1988" authorityName="Morone De Lucia, D'Addabbo Gallo and Grimaldi de Zio" authorityYear="1988" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B46FFE4615A07A718F0F93E" box="[624,763,1736,1760]" italics="true" pageId="5" pageNumber="488">B. africanus</emphasis>
Morone De Lucia, D’Addabbo Gallo and Grimaldi de
<bibRefCitation id="1D4F478D5B46FFE4664407A91AD8F8DA" author="Zio" pageId="5" pageNumber="488" refString="Morone De Lucia, R. M., D'Addabo Gallo, M., Grimaldi de Zio, S. (1988) Descrizione di due nuove specie di Batillipedidae (Tardigrada: Heterotardigrada). Cahiers de Biologie Marine, 29, 361 - 373." type="journal article" year="1988">Zio, 1988</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="BEDE41FF5B46FFE463F707831B48F8DA" box="[221,323,1772,1796]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B46FFE463F707831B48F8DA" box="[221,323,1772,1796]" italics="true" pageId="5" pageNumber="488">B. lesteri</emphasis>
</taxonomicName>
;
<taxonomicName id="BEDE41FF5B46FFE4627B07831BE9F8DA" box="[337,482,1772,1796]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="tubernatis">
<emphasis id="4BAAE66E5B46FFE4627B07831BE9F8DA" box="[337,482,1772,1796]" italics="true" pageId="5" pageNumber="488">B. tubernatis</emphasis>
</taxonomicName>
;
<taxonomicName id="BEDE41FF5B46FFE462DB078318F9F8DB" authority="Schulz, 1955" authorityName="Schulz" authorityYear="1955" box="[497,754,1772,1797]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="4BAAE66E5B46FFE462DB07831852F8DA" box="[497,601,1772,1796]" italics="true" pageId="5" pageNumber="488">B. similis</emphasis>
<bibRefCitation id="1D4F478D5B46FFE4614A078218F9F8DB" author="Schulz" box="[608,754,1772,1797]" pageId="5" pageNumber="488" refString="Schulz, E., (1955) Studien an marinen tardigraden. Kieler Meeresforschungen, 11, 74 - 79." type="journal article" year="1955">Schulz, 1955</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="BEDE41FF5B46FFE46003078319CFF8DA" box="[809,964,1772,1796]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="acuticauda">
<emphasis id="4BAAE66E5B46FFE46003078319CFF8DA" box="[809,964,1772,1796]" italics="true" pageId="5" pageNumber="488">B. acuticauda</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="79613A7C5B46FFE763ED06611E4EFDAC" blockId="5.[151,1437,151,2013]" lastBlockId="6.[119,1895,575,626]" lastPageId="6" lastPageNumber="489" pageId="5" pageNumber="488">
The new species, in having lateral projections, can be clearly distinguished from
<taxonomicName id="BEDE41FF5B46FFE46787067F1F35F8F6" box="[1197,1342,1808,1832]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B46FFE46787067F1F35F8F6" box="[1197,1342,1808,1832]" italics="true" pageId="5" pageNumber="488">B. africanus</emphasis>
</taxonomicName>
and
<emphasis id="4BAAE66E5B46FFE466AE067F1805F892" italics="true" pageId="5" pageNumber="488">
<taxonomicName id="BEDE41FF5B46FFE466AE067F1B07F892" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="tubernatis">B. tubernatis</taxonomicName>
.
<taxonomicName id="BEDE41FF5B46FFE46232065A1805F892" box="[280,526,1844,1868]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">Batillipes brasiliensis</taxonomicName>
</emphasis>
<emphasis id="4BAAE66E5B46FFE4613D065A1879F893" bold="true" box="[535,626,1844,1869]" pageId="5" pageNumber="488">
<taxonomicNameLabel id="50995B155B46FFE4613D065A1879F893" box="[535,626,1844,1869]" pageId="5" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="BEDE41FF5B46FFE46026065B197DF892" box="[780,886,1844,1868]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="4BAAE66E5B46FFE46026065B197DF892" box="[780,886,1844,1868]" italics="true" pageId="5" pageNumber="488">B. similis</emphasis>
</taxonomicName>
in the shape of lateral processes and the caudal appendage that, despite variability, are always conical in the last species. In addition, the primary clava of
<taxonomicName id="BEDE41FF5B46FFE466AE06371AEBF84A" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="4BAAE66E5B46FFE466AE06371AEBF84A" italics="true" pageId="5" pageNumber="488">B. similis</emphasis>
</taxonomicName>
, in having a swollen tip, is drumstick-like and not uniformly tubular as in the new species. On the other hand, the sense organ on leg IV is much longer, almost twice the length, in
<taxonomicName id="BEDE41FF5B46FFE460CF06CF1E46F866" box="[997,1101,1952,1976]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="4BAAE66E5B46FFE460CF06CF1E46F866" box="[997,1101,1952,1976]" italics="true" pageId="5" pageNumber="488">B. similis</emphasis>
</taxonomicName>
than in the new species; 18% of the body length in
<taxonomicName id="BEDE41FF5B46FFE4628006AB1819F802" box="[426,530,1988,2012]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="4BAAE66E5B46FFE4628006AB1819F802" box="[426,530,1988,2012]" italics="true" pageId="5" pageNumber="488">B. similis</emphasis>
</taxonomicName>
(mean values, according to Gallo d’Addabbo
<emphasis id="4BAAE66E5B46FFE4673506AB1E5BF802" box="[1055,1104,1988,2012]" italics="true" pageId="5" pageNumber="488">et al</emphasis>
. 1999) and 10% in the new
<emphasis id="4BAAE66E5B45FFE7635D03511ADCFD8B" bold="true" box="[119,215,575,597]" pageId="6" pageNumber="489">TABLE]</emphasis>
. Measurements (in µm) of selecteđ morphological structures of specimens of
<taxonomicName id="BEDE41FF5B45FFE7608303511E7AFD8B" box="[937,1137,575,597]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="489" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B45FFE7608303511E7AFD8B" box="[937,1137,575,597]" italics="true" pageId="6" pageNumber="489">Batillipes brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B45FFE7675203511EC8FD8B" bold="true" box="[1144,1219,575,597]" pageId="6" pageNumber="489">
<taxonomicNameLabel id="50995B155B45FFE7675203511EC8FD8B" box="[1144,1219,575,597]" pageId="6" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Holo—Holotype; SD—Stanđarđ đeviation; Range refers to the smallest largest measuređ specimen/structure; N—number of specimens/structures measuređ; Allo—Allotype).
</paragraph>
<paragraph id="79613A7C5B45FFE762F203F91D2CFD73" blockId="6.[119,1831,663,741]" box="[472,1831,663,685]" pageId="6" pageNumber="489">
<emphasis id="4BAAE66E5B45FFE762F203F91D2CFD73" bold="true" box="[472,1831,663,685]" pageId="6" pageNumber="489">FEMALES MALES JUVENILES LARVA</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B45FFE962F203DD1B66FC32" lastPageId="8" lastPageNumber="491" pageId="6" pageNumber="489" type="description">
<paragraph id="79613A7C5B45FFE762F203DD1D18FD17" blockId="6.[119,1831,663,741]" box="[472,1811,691,713]" pageId="6" pageNumber="489">
<emphasis id="4BAAE66E5B45FFE762F203DD1D18FD17" bold="true" box="[472,1811,691,713]" pageId="6" pageNumber="489">adults adults 6-toed 4-toed</emphasis>
</paragraph>
<paragraph id="79613A7C5B45FFE7635D03A11C59FD3B" blockId="6.[119,1831,663,741]" box="[119,1618,719,741]" pageId="6" pageNumber="489">
<emphasis id="4BAAE66E5B45FFE7635D03A11C59FD3B" bold="true" box="[119,1618,719,741]" pageId="6" pageNumber="489">STRUCTURES Holo Mean ± SD (Range); N Allo Mean ± SD (Range); N Mean ± SD (Range); N</emphasis>
</paragraph>
<paragraph id="79613A7C5B45FFE7635D03941CEBF9FD" pageId="6" pageNumber="489">
<table id="0BDEC8DC5B45001E635D03941D6DF9FD" box="[119,1894,762,1571]" gridcols="7" gridrows="20" pageId="6" pageNumber="489">
<tr id="C7EE383E5B45001E635D03941D6DFCCE" box="[119,1894,762,784]" gridrow="0" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D03941B5EFCCE" box="[119,341,762,784]" gridcol="0" gridrow="0" pageId="6" pageNumber="489">Bođy length</th>
<th id="843F51425B45001E62F20394180BFCCE" box="[472,512,762,784]" gridcol="1" gridrow="0" pageId="6" pageNumber="489">142</th>
<th id="843F51425B45001E61610394194AFCCE" box="[587,833,762,784]" gridcol="2" gridrow="0" pageId="6" pageNumber="489">130.3 ±19.3 (91­156); 16</th>
<th id="843F51425B45001E6086039419DFFCCE" box="[940,980,762,784]" gridcol="3" gridrow="0" pageId="6" pageNumber="489">159</th>
<th id="843F51425B45001E673503941F01FCCE" box="[1055,1290,762,784]" gridcol="4" gridrow="0" pageId="6" pageNumber="489">148.3 ±8.8 (142­159); 5</th>
<th id="843F51425B45001E665F03941C6BFCCE" box="[1397,1632,762,784]" gridcol="5" gridrow="0" pageId="6" pageNumber="489">140.1 ±7.8 (130­150); 5</th>
<th id="843F51425B45001E65FC03941D6DFCCE" box="[1750,1894,762,784]" gridcol="6" gridrow="0" pageId="6" pageNumber="489">92</th>
</tr>
<tr id="C7EE383E5B45001E635D024D1D6DFCE7" box="[119,1894,803,825]" gridrow="1" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D024D1B5EFCE7" box="[119,341,803,825]" gridcol="0" gridrow="1" pageId="6" pageNumber="489">Bođy without cauđal ala</th>
<td id="843F51425B45001E62F2024D180BFCE7" box="[472,512,803,825]" gridcol="1" gridrow="1" pageId="6" pageNumber="489">124</td>
<td id="843F51425B45001E6161024D194AFCE7" box="[587,833,803,825]" gridcol="2" gridrow="1" pageId="6" pageNumber="489">121.7 ±17.0 (87­151); 16</td>
<td id="843F51425B45001E6086024D19DFFCE7" box="[940,980,803,825]" gridcol="3" gridrow="1" pageId="6" pageNumber="489">143</td>
<td id="843F51425B45001E6735024D1F01FCE7" box="[1055,1290,803,825]" gridcol="4" gridrow="1" pageId="6" pageNumber="489">134.0 ±7.0 (127­144); 5</td>
<td id="843F51425B45001E665F024D1C6BFCE7" box="[1397,1632,803,825]" gridcol="5" gridrow="1" pageId="6" pageNumber="489">128.2 ±9.0 (116­137); 5</td>
<td id="843F51425B45001E65FC024D1D6DFCE7" box="[1750,1894,803,825]" gridcol="6" gridrow="1" pageId="6" pageNumber="489">87</td>
</tr>
<tr id="C7EE383E5B45001E635D02231D6DFCBD" box="[119,1894,845,867]" gridrow="2" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D02231B5EFCBD" box="[119,341,845,867]" gridcol="0" gridrow="2" pageId="6" pageNumber="489">Bođy wiđth</th>
<td id="843F51425B45001E62F20223180BFCBD" box="[472,512,845,867]" gridcol="1" gridrow="2" pageId="6" pageNumber="489">45.4</td>
<td id="843F51425B45001E61610223194AFCBD" box="[587,833,845,867]" gridcol="2" gridrow="2" pageId="6" pageNumber="489">46.9 ±8.7 (30.5­62.5); 17</td>
<td id="843F51425B45001E6086022319DFFCBD" box="[940,980,845,867]" gridcol="3" gridrow="2" pageId="6" pageNumber="489">40.0</td>
<td id="843F51425B45001E673502231F01FCBD" box="[1055,1290,845,867]" gridcol="4" gridrow="2" pageId="6" pageNumber="489">43.0 ±3.70 (37.6­46.3); 4</td>
<td id="843F51425B45001E665F02231C6BFCBD" box="[1397,1632,845,867]" gridcol="5" gridrow="2" pageId="6" pageNumber="489">48.1 ±7.78 (38.8­57.7); 6</td>
<td id="843F51425B45001E65FC02231D6DFCBD" box="[1750,1894,845,867]" gridcol="6" gridrow="2" pageId="6" pageNumber="489">25.9</td>
</tr>
<tr id="C7EE383E5B45001E635D02181D6DFC52" box="[119,1894,886,908]" gridrow="3" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D02181B5EFC52" box="[119,341,886,908]" gridcol="0" gridrow="3" pageId="6" pageNumber="489">Međian cirrus</th>
<td id="843F51425B45001E62F20218180BFC52" box="[472,512,886,908]" gridcol="1" gridrow="3" pageId="6" pageNumber="489">15.5</td>
<td id="843F51425B45001E61610218194AFC52" box="[587,833,886,908]" gridcol="2" gridrow="3" pageId="6" pageNumber="489">17.4 ±6.0 (11.2­28.8); 7</td>
<td id="843F51425B45001E6086021819DFFC52" box="[940,980,886,908]" gridcol="3" gridrow="3" pageId="6" pageNumber="489">14.0</td>
<td id="843F51425B45001E673502181F01FC52" box="[1055,1290,886,908]" gridcol="4" gridrow="3" pageId="6" pageNumber="489">- (12.4); 1</td>
<td id="843F51425B45001E665F02181C6BFC52" box="[1397,1632,886,908]" gridcol="5" gridrow="3" pageId="6" pageNumber="489">15.2 ±5.25 (10.4­20.8); 3</td>
<td id="843F51425B45001E65FC02181D6DFC52" box="[1750,1894,886,908]" gridcol="6" gridrow="3" pageId="6" pageNumber="489">13.0</td>
</tr>
<tr id="C7EE383E5B45001E635D02CE1D6DFC68" box="[119,1894,928,950]" gridrow="4" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D02CE1B5EFC68" box="[119,341,928,950]" gridcol="0" gridrow="4" pageId="6" pageNumber="489">Internal cirri</th>
<td id="843F51425B45001E62F202CE180BFC68" box="[472,512,928,950]" gridcol="1" gridrow="4" pageId="6" pageNumber="489">18.6</td>
<td id="843F51425B45001E616102CE194AFC68" box="[587,833,928,950]" gridcol="2" gridrow="4" pageId="6" pageNumber="489">18.0 ±2.19 (14.9­22.7); 15</td>
<td id="843F51425B45001E608602CE19DFFC68" box="[940,980,928,950]" gridcol="3" gridrow="4" pageId="6" pageNumber="489">18.5</td>
<td id="843F51425B45001E673502CE1F01FC68" box="[1055,1290,928,950]" gridcol="4" gridrow="4" pageId="6" pageNumber="489">18.1 ±3.61 (13.2­21.0); 5</td>
<td id="843F51425B45001E665F02CE1C6BFC68" box="[1397,1632,928,950]" gridcol="5" gridrow="4" pageId="6" pageNumber="489">15.3 ±3.27 (12.3­18.3); 4</td>
<td id="843F51425B45001E65FC02CE1D6DFC68" box="[1750,1894,928,950]" gridcol="6" gridrow="4" pageId="6" pageNumber="489">11.3</td>
</tr>
<tr id="C7EE383E5B45001E635D02A71D6DFC01" box="[119,1894,969,991]" gridrow="5" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D02A71B5EFC01" box="[119,341,969,991]" gridcol="0" gridrow="5" pageId="6" pageNumber="489">External cirri</th>
<td id="843F51425B45001E62F202A7180BFC01" box="[472,512,969,991]" gridcol="1" gridrow="5" pageId="6" pageNumber="489">12.7</td>
<td id="843F51425B45001E616102A7194AFC01" box="[587,833,969,991]" gridcol="2" gridrow="5" pageId="6" pageNumber="489">13.4 ±1.94 (11.0­17.9); 16</td>
<td id="843F51425B45001E608602A719DFFC01" box="[940,980,969,991]" gridcol="3" gridrow="5" pageId="6" pageNumber="489">8.7</td>
<td id="843F51425B45001E673502A71F01FC01" box="[1055,1290,969,991]" gridcol="4" gridrow="5" pageId="6" pageNumber="489">11.9 ±1.54 (9.9­14.0); 5</td>
<td id="843F51425B45001E665F02A71C6BFC01" box="[1397,1632,969,991]" gridcol="5" gridrow="5" pageId="6" pageNumber="489">11.6 ±1.78 (9.5­13.9); 6</td>
<td id="843F51425B45001E65FC02A71D6DFC01" box="[1750,1894,969,991]" gridcol="6" gridrow="5" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D029C1D6DFBD6" box="[119,1894,1010,1032]" gridrow="6" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D029C1B5EFBD6" box="[119,341,1010,1032]" gridcol="0" gridrow="6" pageId="6" pageNumber="489">Lateral cirri A</th>
<td id="843F51425B45001E62F2029C180BFBD6" box="[472,512,1010,1032]" gridcol="1" gridrow="6" pageId="6" pageNumber="489">26.6</td>
<td id="843F51425B45001E6161029C194AFBD6" box="[587,833,1010,1032]" gridcol="2" gridrow="6" pageId="6" pageNumber="489">27.7 ±2.15 (24.5­31.5); 16</td>
<td id="843F51425B45001E6086029C19DFFBD6" box="[940,980,1010,1032]" gridcol="3" gridrow="6" pageId="6" pageNumber="489">25.6</td>
<td id="843F51425B45001E6735029C1F01FBD6" box="[1055,1290,1010,1032]" gridcol="4" gridrow="6" pageId="6" pageNumber="489">27.6 ±2.84 (24.4­31.3); 4</td>
<td id="843F51425B45001E665F029C1C6BFBD6" box="[1397,1632,1010,1032]" gridcol="5" gridrow="6" pageId="6" pageNumber="489">22.2 ±6.77 (14.8­29.0); 4</td>
<td id="843F51425B45001E65FC029C1D6DFBD6" box="[1750,1894,1010,1032]" gridcol="6" gridrow="6" pageId="6" pageNumber="489">14.4</td>
</tr>
<tr id="C7EE383E5B45001E635D05721D6DFBEC" box="[119,1894,1052,1074]" gridrow="7" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D05721B5EFBEC" box="[119,341,1052,1074]" gridcol="0" gridrow="7" pageId="6" pageNumber="489">Primary clavae</th>
<td id="843F51425B45001E62F20572180BFBEC" box="[472,512,1052,1074]" gridcol="1" gridrow="7" pageId="6" pageNumber="489">12.9</td>
<td id="843F51425B45001E61610572194AFBEC" box="[587,833,1052,1074]" gridcol="2" gridrow="7" pageId="6" pageNumber="489">12.6 ±1.84 (9.2­15.8); 16</td>
<td id="843F51425B45001E6086057219DFFBEC" box="[940,980,1052,1074]" gridcol="3" gridrow="7" pageId="6" pageNumber="489">13.5</td>
<td id="843F51425B45001E673505721F01FBEC" box="[1055,1290,1052,1074]" gridcol="4" gridrow="7" pageId="6" pageNumber="489">13.1 ±1.0 (12.1­14.1); 4</td>
<td id="843F51425B45001E665F05721C6BFBEC" box="[1397,1632,1052,1074]" gridcol="5" gridrow="7" pageId="6" pageNumber="489">12.4 ±0.71 (11.2­13.0); 5</td>
<td id="843F51425B45001E65FC05721D6DFBEC" box="[1750,1894,1052,1074]" gridcol="6" gridrow="7" pageId="6" pageNumber="489">7.4</td>
</tr>
<tr id="C7EE383E5B45001E635D052B1D6DFB85" box="[119,1894,1093,1115]" gridrow="8" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D052B1B5EFB85" box="[119,341,1093,1115]" gridcol="0" gridrow="8" pageId="6" pageNumber="489">Primary clava / cirrus A</th>
<td id="843F51425B45001E62F2052B180BFB85" box="[472,512,1093,1115]" gridcol="1" gridrow="8" pageId="6" pageNumber="489">0.49</td>
<td id="843F51425B45001E6161052B194AFB85" box="[587,833,1093,1115]" gridcol="2" gridrow="8" pageId="6" pageNumber="489">0.46 ±0.06 (0.35­0.53); 16</td>
<td id="843F51425B45001E6086052B19DFFB85" box="[940,980,1093,1115]" gridcol="3" gridrow="8" pageId="6" pageNumber="489">0.53</td>
<td id="843F51425B45001E6735052B1F01FB85" box="[1055,1290,1093,1115]" gridcol="4" gridrow="8" pageId="6" pageNumber="489">0.48 ±0.06 (0.45­0.57); 4</td>
<td id="843F51425B45001E665F052B1C6BFB85" box="[1397,1632,1093,1115]" gridcol="5" gridrow="8" pageId="6" pageNumber="489">0.65 ±0.15 (0.47­0.76); 3</td>
<td id="843F51425B45001E65FC052B1D6DFB85" box="[1750,1894,1093,1115]" gridcol="6" gridrow="8" pageId="6" pageNumber="489">0.51</td>
</tr>
<tr id="C7EE383E5B45001E635D05011D6DFB5B" box="[119,1894,1135,1157]" gridrow="9" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D05011B5EFB5B" box="[119,341,1135,1157]" gridcol="0" gridrow="9" pageId="6" pageNumber="489">Seconđary clavae</th>
<td id="843F51425B45001E62F20501180BFB5B" box="[472,512,1135,1157]" gridcol="1" gridrow="9" pageId="6" pageNumber="489">2.9</td>
<td id="843F51425B45001E61610501194AFB5B" box="[587,833,1135,1157]" gridcol="2" gridrow="9" pageId="6" pageNumber="489">3.2±0.79 (1.2­4.7); 17</td>
<td id="843F51425B45001E6086050119DFFB5B" box="[940,980,1135,1157]" gridcol="3" gridrow="9" pageId="6" pageNumber="489">?</td>
<td id="843F51425B45001E673505011F01FB5B" box="[1055,1290,1135,1157]" gridcol="4" gridrow="9" pageId="6" pageNumber="489">3.3 ±0.63 (2.6­4.1); 4</td>
<td id="843F51425B45001E665F05011C6BFB5B" box="[1397,1632,1135,1157]" gridcol="5" gridrow="9" pageId="6" pageNumber="489">2.9±0.58 (1.8­3.4); 6</td>
<td id="843F51425B45001E65FC05011D6DFB5B" box="[1750,1894,1135,1157]" gridcol="6" gridrow="9" pageId="6" pageNumber="489">3.3</td>
</tr>
<tr id="C7EE383E5B45001E635D05F61D6DFB70" box="[119,1894,1176,1198]" gridrow="10" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D05F61B5EFB70" box="[119,341,1176,1198]" gridcol="0" gridrow="10" pageId="6" pageNumber="489">Cirri E</th>
<td id="843F51425B45001E62F205F6180BFB70" box="[472,512,1176,1198]" gridcol="1" gridrow="10" pageId="6" pageNumber="489">13.1</td>
<td id="843F51425B45001E616105F6194AFB70" box="[587,833,1176,1198]" gridcol="2" gridrow="10" pageId="6" pageNumber="489">9.8 ±3.02 (7.2­13.1); 3</td>
<td id="843F51425B45001E608605F619DFFB70" box="[940,980,1176,1198]" gridcol="3" gridrow="10" pageId="6" pageNumber="489">7.3</td>
<td id="843F51425B45001E673505F61F01FB70" box="[1055,1290,1176,1198]" gridcol="4" gridrow="10" pageId="6" pageNumber="489">- (8.0); 1</td>
<td id="843F51425B45001E665F05F61C6BFB70" box="[1397,1632,1176,1198]" gridcol="5" gridrow="10" pageId="6" pageNumber="489">9.0 ±2.84 (7.1­12.3); 3</td>
<td id="843F51425B45001E65FC05F61D6DFB70" box="[1750,1894,1176,1198]" gridcol="6" gridrow="10" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D05AC1D6DFB06" box="[119,1894,1218,1240]" gridrow="11" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D05AC1B5EFB06" box="[119,341,1218,1240]" gridcol="0" gridrow="11" pageId="6" pageNumber="489">Leg I spine</th>
<td id="843F51425B45001E62F205AC180BFB06" box="[472,512,1218,1240]" gridcol="1" gridrow="11" pageId="6" pageNumber="489">5.1</td>
<td id="843F51425B45001E616105AC194AFB06" box="[587,833,1218,1240]" gridcol="2" gridrow="11" pageId="6" pageNumber="489">5.6 ±0.87 (4.0­6.3); 7</td>
<td id="843F51425B45001E608605AC19DFFB06" box="[940,980,1218,1240]" gridcol="3" gridrow="11" pageId="6" pageNumber="489">5.6</td>
<td id="843F51425B45001E673505AC1F01FB06" box="[1055,1290,1218,1240]" gridcol="4" gridrow="11" pageId="6" pageNumber="489">5.4 ±1.10 (4.3­6.5); 3</td>
<td id="843F51425B45001E665F05AC1C6BFB06" box="[1397,1632,1218,1240]" gridcol="5" gridrow="11" pageId="6" pageNumber="489">6.3 ±0.66 (5.7­7.0); 3</td>
<td id="843F51425B45001E65FC05AC1D6DFB06" box="[1750,1894,1218,1240]" gridcol="6" gridrow="11" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D05851D6DFADF" box="[119,1894,1259,1281]" gridrow="12" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D05851B5EFADF" box="[119,341,1259,1281]" gridcol="0" gridrow="12" pageId="6" pageNumber="489">Leg II spine</th>
<td id="843F51425B45001E62F20585180BFADF" box="[472,512,1259,1281]" gridcol="1" gridrow="12" pageId="6" pageNumber="489">8.2</td>
<td id="843F51425B45001E61610585194AFADF" box="[587,833,1259,1281]" gridcol="2" gridrow="12" pageId="6" pageNumber="489">7.3 ±2.12 (4.6­10.0); 7</td>
<td id="843F51425B45001E6086058519DFFADF" box="[940,980,1259,1281]" gridcol="3" gridrow="12" pageId="6" pageNumber="489">6.3</td>
<td id="843F51425B45001E673505851F01FADF" box="[1055,1290,1259,1281]" gridcol="4" gridrow="12" pageId="6" pageNumber="489">- (10.5­10.6); 2</td>
<td id="843F51425B45001E665F05851C6BFADF" box="[1397,1632,1259,1281]" gridcol="5" gridrow="12" pageId="6" pageNumber="489">- (11.0); 1</td>
<td id="843F51425B45001E65FC05851D6DFADF" box="[1750,1894,1259,1281]" gridcol="6" gridrow="12" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D047B1D6DFAF5" box="[119,1894,1301,1323]" gridrow="13" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D047B1B5EFAF5" box="[119,341,1301,1323]" gridcol="0" gridrow="13" pageId="6" pageNumber="489">Leg III spine</th>
<td id="843F51425B45001E62F2047B180BFAF5" box="[472,512,1301,1323]" gridcol="1" gridrow="13" pageId="6" pageNumber="489">9.6</td>
<td id="843F51425B45001E6161047B194AFAF5" box="[587,833,1301,1323]" gridcol="2" gridrow="13" pageId="6" pageNumber="489">6.9 ±2.09 (4.6­9.6); 8</td>
<td id="843F51425B45001E6086047B19DFFAF5" box="[940,980,1301,1323]" gridcol="3" gridrow="13" pageId="6" pageNumber="489">8.2</td>
<td id="843F51425B45001E6735047B1F01FAF5" box="[1055,1290,1301,1323]" gridcol="4" gridrow="13" pageId="6" pageNumber="489">- (5.9­9.2); 2</td>
<td id="843F51425B45001E665F047B1C6BFAF5" box="[1397,1632,1301,1323]" gridcol="5" gridrow="13" pageId="6" pageNumber="489">- (8.5­8.7); 2</td>
<td id="843F51425B45001E65FC047B1D6DFAF5" box="[1750,1894,1301,1323]" gridcol="6" gridrow="13" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D04501D6DFA8A" box="[119,1894,1342,1364]" gridrow="14" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D04501B5EFA8A" box="[119,341,1342,1364]" gridcol="0" gridrow="14" pageId="6" pageNumber="489">Leg IV sense organ</th>
<td id="843F51425B45001E62F20450180BFA8A" box="[472,512,1342,1364]" gridcol="1" gridrow="14" pageId="6" pageNumber="489">12.8</td>
<td id="843F51425B45001E61610450194AFA8A" box="[587,833,1342,1364]" gridcol="2" gridrow="14" pageId="6" pageNumber="489">12.6 ±2.08 (9.3­16.0); 16</td>
<td id="843F51425B45001E6086045019DFFA8A" box="[940,980,1342,1364]" gridcol="3" gridrow="14" pageId="6" pageNumber="489">15.6</td>
<td id="843F51425B45001E673504501F01FA8A" box="[1055,1290,1342,1364]" gridcol="4" gridrow="14" pageId="6" pageNumber="489">13.7 ±1.47 (12.0­15.3); 4</td>
<td id="843F51425B45001E665F04501C6BFA8A" box="[1397,1632,1342,1364]" gridcol="5" gridrow="14" pageId="6" pageNumber="489">13.1 ±2.50 (9.6­15.7); 3</td>
<td id="843F51425B45001E65FC04501D6DFA8A" box="[1750,1894,1342,1364]" gridcol="6" gridrow="14" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D04091D6DFAA3" box="[119,1894,1383,1405]" gridrow="15" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D04091B5EFAA3" box="[119,341,1383,1405]" gridcol="0" gridrow="15" pageId="6" pageNumber="489">Leg IV base</th>
<td id="843F51425B45001E62F20409180BFAA3" box="[472,512,1383,1405]" gridcol="1" gridrow="15" pageId="6" pageNumber="489">5.8</td>
<td id="843F51425B45001E61610409194AFAA3" box="[587,833,1383,1405]" gridcol="2" gridrow="15" pageId="6" pageNumber="489">6.3 ±0.88 (5.1­7.6); 13</td>
<td id="843F51425B45001E6086040919DFFAA3" box="[940,980,1383,1405]" gridcol="3" gridrow="15" pageId="6" pageNumber="489">7.8</td>
<td id="843F51425B45001E673504091F01FAA3" box="[1055,1290,1383,1405]" gridcol="4" gridrow="15" pageId="6" pageNumber="489">6.2 ±1.15 (4.7­7.1); 4</td>
<td id="843F51425B45001E665F04091C6BFAA3" box="[1397,1632,1383,1405]" gridcol="5" gridrow="15" pageId="6" pageNumber="489">5.9 ±0.66 (5.1­6.7); 4</td>
<td id="843F51425B45001E65FC04091D6DFAA3" box="[1750,1894,1383,1405]" gridcol="6" gridrow="15" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D04FF1D6DFA79" box="[119,1894,1425,1447]" gridrow="16" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D04FF1B5EFA79" box="[119,341,1425,1447]" gridcol="0" gridrow="16" pageId="6" pageNumber="489">Leg IV spine</th>
<td id="843F51425B45001E62F204FF180BFA79" box="[472,512,1425,1447]" gridcol="1" gridrow="16" pageId="6" pageNumber="489">7.0</td>
<td id="843F51425B45001E616104FF194AFA79" box="[587,833,1425,1447]" gridcol="2" gridrow="16" pageId="6" pageNumber="489">7.2 ±1.61 (5.4­10.9); 11</td>
<td id="843F51425B45001E608604FF19DFFA79" box="[940,980,1425,1447]" gridcol="3" gridrow="16" pageId="6" pageNumber="489">7.8</td>
<td id="843F51425B45001E673504FF1F01FA79" box="[1055,1290,1425,1447]" gridcol="4" gridrow="16" pageId="6" pageNumber="489">6.4 ±1.28 (4.5­7.3); 4</td>
<td id="843F51425B45001E665F04FF1C6BFA79" box="[1397,1632,1425,1447]" gridcol="5" gridrow="16" pageId="6" pageNumber="489">7.6 ±1.70 (5.7­9.5); 5</td>
<td id="843F51425B45001E65FC04FF1D6DFA79" box="[1750,1894,1425,1447]" gridcol="6" gridrow="16" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D04D41D6DFA0E" box="[119,1894,1466,1488]" gridrow="17" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D04D41B5EFA0E" box="[119,341,1466,1488]" gridcol="0" gridrow="17" pageId="6" pageNumber="489">Leg I projection</th>
<td id="843F51425B45001E62F204D4180BFA0E" box="[472,512,1466,1488]" gridcol="1" gridrow="17" pageId="6" pageNumber="489">3.5</td>
<td id="843F51425B45001E616104D4194AFA0E" box="[587,833,1466,1488]" gridcol="2" gridrow="17" pageId="6" pageNumber="489">3.6 ±0.94 (2.0­5.0); 11</td>
<td id="843F51425B45001E608604D419DFFA0E" box="[940,980,1466,1488]" gridcol="3" gridrow="17" pageId="6" pageNumber="489">3.5</td>
<td id="843F51425B45001E673504D41F01FA0E" box="[1055,1290,1466,1488]" gridcol="4" gridrow="17" pageId="6" pageNumber="489">3.9 ±0.74 (3.1­4.8); 5</td>
<td id="843F51425B45001E665F04D41C6BFA0E" box="[1397,1632,1466,1488]" gridcol="5" gridrow="17" pageId="6" pageNumber="489">3.9 ±0.55 (3.3­4.5); 4</td>
<td id="843F51425B45001E65FC04D41D6DFA0E" box="[1750,1894,1466,1488]" gridcol="6" gridrow="17" pageId="6" pageNumber="489">2.2</td>
</tr>
<tr id="C7EE383E5B45001E635D048A1D6DFA24" box="[119,1894,1508,1530]" gridrow="18" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D048A1B5EFA24" box="[119,341,1508,1530]" gridcol="0" gridrow="18" pageId="6" pageNumber="489">Leg II projection</th>
<td id="843F51425B45001E62F2048A180BFA24" box="[472,512,1508,1530]" gridcol="1" gridrow="18" pageId="6" pageNumber="489">3.3</td>
<td id="843F51425B45001E6161048A194AFA24" box="[587,833,1508,1530]" gridcol="2" gridrow="18" pageId="6" pageNumber="489">3.4 ±0.43 (2.8­4.0); 7</td>
<td id="843F51425B45001E6086048A19DFFA24" box="[940,980,1508,1530]" gridcol="3" gridrow="18" pageId="6" pageNumber="489">5.1</td>
<td id="843F51425B45001E6735048A1F01FA24" box="[1055,1290,1508,1530]" gridcol="4" gridrow="18" pageId="6" pageNumber="489">3.5 ±1.56 (2.0­6.0); 5</td>
<td id="843F51425B45001E665F048A1C6BFA24" box="[1397,1632,1508,1530]" gridcol="5" gridrow="18" pageId="6" pageNumber="489">3.0 ±0.93 (1.8­3.7); 4</td>
<td id="843F51425B45001E65FC048A1D6DFA24" box="[1750,1894,1508,1530]" gridcol="6" gridrow="18" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D07631D6DF9FD" box="[119,1894,1549,1571]" gridrow="19" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D07631B5EF9FD" box="[119,341,1549,1571]" gridcol="0" gridrow="19" pageId="6" pageNumber="489">Leg III projection</th>
<td id="843F51425B45001E62F20763180BF9FD" box="[472,512,1549,1571]" gridcol="1" gridrow="19" pageId="6" pageNumber="489">2.1</td>
<td id="843F51425B45001E61610763194AF9FD" box="[587,833,1549,1571]" gridcol="2" gridrow="19" pageId="6" pageNumber="489">- (2.2); 1</td>
<td id="843F51425B45001E6086076319DFF9FD" box="[940,980,1549,1571]" gridcol="3" gridrow="19" pageId="6" pageNumber="489">2.4</td>
<td id="843F51425B45001E673507631F01F9FD" box="[1055,1290,1549,1571]" gridcol="4" gridrow="19" pageId="6" pageNumber="489">- (4.4); 2</td>
<td id="843F51425B45001E665F07631C6BF9FD" box="[1397,1632,1549,1571]" gridcol="5" gridrow="19" pageId="6" pageNumber="489">- (1.7­3.3); 2</td>
<td id="843F51425B45001E65FC07631D6DF9FD" box="[1750,1894,1549,1571]" gridcol="6" gridrow="19" pageId="6" pageNumber="489">?</td>
</tr>
</table>
</paragraph>
<paragraph id="79613A7C5B45FFE9656407561F96FF0A" blockId="6.[1614,1894,1592,1614]" lastBlockId="8.[151,1437,151,1005]" lastPageId="8" lastPageNumber="491" pageId="6" pageNumber="489">
<emphasis id="4BAAE66E5B45FFE7656407561D6DF990" box="[1614,1894,1592,1614]" italics="true" pageId="6" pageNumber="489">......continued on the next page</emphasis>
<emphasis id="4BAAE66E5B44FFE6635703CF1AEAFD69" bold="true" box="[125,225,673,695]" pageId="7" pageNumber="490">TABLE].</emphasis>
(Continueđ) species.
<taxonomicName id="BEDE41FF5B4BFFE963D001F61BE5FF6E" box="[250,494,152,176]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="acuticauda">
<emphasis id="4BAAE66E5B4BFFE963D001F61BE5FF6E" box="[250,494,152,176]" italics="true" pageId="8" pageNumber="491">Batillipes acuticauda</emphasis>
</taxonomicName>
, in contrary to the new species, has an acute spiky caudal appendage, cephalic appendages sharply pointed, and very different shaped lateral projections that are flattened between legs III and IV.
</paragraph>
<paragraph id="79613A7C5B44FFE66357026A1CEFF9EF" pageId="7" pageNumber="490">
<table id="0BDEC8DC5B44001E635703A21D21F9EF" box="[125,1834,716,1585]" gridcols="7" gridrows="20" pageId="7" pageNumber="490">
<tr id="C7EE383E5B44001E635703A21D21FCC4" box="[125,1834,716,794]" gridrow="0" pageId="7" pageNumber="490">
<th id="843F51425B44001E635703A21B6BFCC4" box="[125,352,716,794]" gridcol="0" gridrow="0" pageId="7" pageNumber="490">STRUCTURES</th>
<th id="843F51425B44001E62F703A2184DFCC4" box="[477,582,716,794]" gridcol="1" gridrow="0" pageId="7" pageNumber="490">FEMALES ađults Holo</th>
<th id="843F51425B44001E617B03A2194DFCC4" box="[593,838,716,794]" gridcol="2" gridrow="0" pageId="7" pageNumber="490">Mean ± SD (Range); N</th>
<th id="843F51425B44001E609B03A219F4FCC4" box="[945,1023,716,794]" gridcol="3" gridrow="0" pageId="7" pageNumber="490">MALES ađults Allo</th>
<th id="843F51425B44001E670E03A21F0FFCC4" box="[1060,1284,716,794]" gridcol="4" gridrow="0" pageId="7" pageNumber="490">Mean ± SD (Range); N</th>
<th id="843F51425B44001E665003A21C6EFCC4" box="[1402,1637,716,794]" gridcol="5" gridrow="0" pageId="7" pageNumber="490">JUVENILES 6-toeđ Mean ± SD (Range); N</th>
<th id="843F51425B44001E65F603A21D21FCC4" box="[1756,1834,716,794]" gridcol="6" gridrow="0" pageId="7" pageNumber="490">LARVA 4-toeđ</th>
</tr>
<tr id="C7EE383E5B44001E6357025E1D21FC98" box="[125,1834,816,838]" gridrow="1" pageId="7" pageNumber="490">
<th id="843F51425B44001E6357025E1B6BFC98" box="[125,352,816,838]" gridcol="0" gridrow="1" pageId="7" pageNumber="490">Bođy projection 1</th>
<td id="843F51425B44001E62F7025E184DFC98" box="[477,582,816,838]" gridcol="1" gridrow="1" pageId="7" pageNumber="490">4.1</td>
<td id="843F51425B44001E617B025E194DFC98" box="[593,838,816,838]" gridcol="2" gridrow="1" pageId="7" pageNumber="490">3.5 ±0.52 (2.8­4.1); 5</td>
<td id="843F51425B44001E609B025E19F4FC98" box="[945,1023,816,838]" gridcol="3" gridrow="1" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E025E1F0FFC98" box="[1060,1284,816,838]" gridcol="4" gridrow="1" pageId="7" pageNumber="490">- (3.2); 1</td>
<td id="843F51425B44001E6650025E1C6EFC98" box="[1402,1637,816,838]" gridcol="5" gridrow="1" pageId="7" pageNumber="490">- (2.5); 1</td>
<td id="843F51425B44001E65F6025E1D21FC98" box="[1756,1834,816,838]" gridcol="6" gridrow="1" pageId="7" pageNumber="490">1.8</td>
</tr>
<tr id="C7EE383E5B44001E635702371D21FCB1" box="[125,1834,857,879]" gridrow="2" pageId="7" pageNumber="490">
<th id="843F51425B44001E635702371B6BFCB1" box="[125,352,857,879]" gridcol="0" gridrow="2" pageId="7" pageNumber="490">Bođy projection 2</th>
<td id="843F51425B44001E62F70237184DFCB1" box="[477,582,857,879]" gridcol="1" gridrow="2" pageId="7" pageNumber="490">6.0</td>
<td id="843F51425B44001E617B0237194DFCB1" box="[593,838,857,879]" gridcol="2" gridrow="2" pageId="7" pageNumber="490">4.7 ±1.87 (2.7­6.6); 4</td>
<td id="843F51425B44001E609B023719F4FCB1" box="[945,1023,857,879]" gridcol="3" gridrow="2" pageId="7" pageNumber="490">1.9</td>
<td id="843F51425B44001E670E02371F0FFCB1" box="[1060,1284,857,879]" gridcol="4" gridrow="2" pageId="7" pageNumber="490">- (4.8); 1</td>
<td id="843F51425B44001E665002371C6EFCB1" box="[1402,1637,857,879]" gridcol="5" gridrow="2" pageId="7" pageNumber="490">- (3.0); 1</td>
<td id="843F51425B44001E65F602371D21FCB1" box="[1756,1834,857,879]" gridcol="6" gridrow="2" pageId="7" pageNumber="490">2.6</td>
</tr>
<tr id="C7EE383E5B44001E635702ED1D21FC47" box="[125,1834,899,921]" gridrow="3" pageId="7" pageNumber="490">
<th id="843F51425B44001E635702ED1B6BFC47" box="[125,352,899,921]" gridcol="0" gridrow="3" pageId="7" pageNumber="490">Bođy projection 3</th>
<td id="843F51425B44001E62F702ED184DFC47" box="[477,582,899,921]" gridcol="1" gridrow="3" pageId="7" pageNumber="490">4.6</td>
<td id="843F51425B44001E617B02ED194DFC47" box="[593,838,899,921]" gridcol="2" gridrow="3" pageId="7" pageNumber="490">5.1 ±1.51 (2.9­7.7); 10</td>
<td id="843F51425B44001E609B02ED19F4FC47" box="[945,1023,899,921]" gridcol="3" gridrow="3" pageId="7" pageNumber="490">5.6</td>
<td id="843F51425B44001E670E02ED1F0FFC47" box="[1060,1284,899,921]" gridcol="4" gridrow="3" pageId="7" pageNumber="490">5.6 ±0.58 (5.0­6.1); 3</td>
<td id="843F51425B44001E665002ED1C6EFC47" box="[1402,1637,899,921]" gridcol="5" gridrow="3" pageId="7" pageNumber="490">- (3.8); 1</td>
<td id="843F51425B44001E65F602ED1D21FC47" box="[1756,1834,899,921]" gridcol="6" gridrow="3" pageId="7" pageNumber="490">3.1</td>
</tr>
<tr id="C7EE383E5B44001E635702C21D21FC1C" box="[125,1834,940,962]" gridrow="4" pageId="7" pageNumber="490">
<th id="843F51425B44001E635702C21B6BFC1C" box="[125,352,940,962]" gridcol="0" gridrow="4" pageId="7" pageNumber="490">Bođy projection 4 wiđth</th>
<td id="843F51425B44001E62F702C2184DFC1C" box="[477,582,940,962]" gridcol="1" gridrow="4" pageId="7" pageNumber="490">10.8</td>
<td id="843F51425B44001E617B02C2194DFC1C" box="[593,838,940,962]" gridcol="2" gridrow="4" pageId="7" pageNumber="490">11.5 ±3.32 (5.5­18.4); 14</td>
<td id="843F51425B44001E609B02C219F4FC1C" box="[945,1023,940,962]" gridcol="3" gridrow="4" pageId="7" pageNumber="490">17.6</td>
<td id="843F51425B44001E670E02C21F0FFC1C" box="[1060,1284,940,962]" gridcol="4" gridrow="4" pageId="7" pageNumber="490">15.1 ±4.77 (7.5­19.8); 5</td>
<td id="843F51425B44001E665002C21C6EFC1C" box="[1402,1637,940,962]" gridcol="5" gridrow="4" pageId="7" pageNumber="490">11.0 ±4.71 (4.7­16.1); 6</td>
<td id="843F51425B44001E65F602C21D21FC1C" box="[1756,1834,940,962]" gridcol="6" gridrow="4" pageId="7" pageNumber="490">5.1</td>
</tr>
<tr id="C7EE383E5B44001E635702B81D21FC32" box="[125,1834,982,1004]" gridrow="5" pageId="7" pageNumber="490">
<th id="843F51425B44001E635702B81B6BFC32" box="[125,352,982,1004]" gridcol="0" gridrow="5" pageId="7" pageNumber="490">Bođy projection 4 length</th>
<td id="843F51425B44001E62F702B8184DFC32" box="[477,582,982,1004]" gridcol="1" gridrow="5" pageId="7" pageNumber="490">5.8</td>
<td id="843F51425B44001E617B02B8194DFC32" box="[593,838,982,1004]" gridcol="2" gridrow="5" pageId="7" pageNumber="490">9.2 ±3.29 (5.2­17.7); 14</td>
<td id="843F51425B44001E609B02B819F4FC32" box="[945,1023,982,1004]" gridcol="3" gridrow="5" pageId="7" pageNumber="490">10.9</td>
<td id="843F51425B44001E670E02B81F0FFC32" box="[1060,1284,982,1004]" gridcol="4" gridrow="5" pageId="7" pageNumber="490">11.1 ±2.84 (7.4­15.3); 5</td>
<td id="843F51425B44001E665002B81C6EFC32" box="[1402,1637,982,1004]" gridcol="5" gridrow="5" pageId="7" pageNumber="490">9.5 ±3.5 (3.6­14.0); 6</td>
<td id="843F51425B44001E65F602B81D21FC32" box="[1756,1834,982,1004]" gridcol="6" gridrow="5" pageId="7" pageNumber="490">5.3</td>
</tr>
<tr id="C7EE383E5B44001E635702911D21FBCB" box="[125,1834,1023,1045]" gridrow="6" pageId="7" pageNumber="490">
<th id="843F51425B44001E635702911B6BFBCB" box="[125,352,1023,1045]" gridcol="0" gridrow="6" pageId="7" pageNumber="490">Cauđal apparatus</th>
<td id="843F51425B44001E62F70291184DFBCB" box="[477,582,1023,1045]" gridcol="1" gridrow="6" pageId="7" pageNumber="490">17.5</td>
<td id="843F51425B44001E617B0291194DFBCB" box="[593,838,1023,1045]" gridcol="2" gridrow="6" pageId="7" pageNumber="490">10.1 ±4.78 (1.8­19.2); 15</td>
<td id="843F51425B44001E609B029119F4FBCB" box="[945,1023,1023,1045]" gridcol="3" gridrow="6" pageId="7" pageNumber="490">16.3</td>
<td id="843F51425B44001E670E02911F0FFBCB" box="[1060,1284,1023,1045]" gridcol="4" gridrow="6" pageId="7" pageNumber="490">13.2 ±3.14 (9.0­16.2); 5</td>
<td id="843F51425B44001E665002911C6EFBCB" box="[1402,1637,1023,1045]" gridcol="5" gridrow="6" pageId="7" pageNumber="490">12.7 ±4.20 (5.7­17.8); 6</td>
<td id="843F51425B44001E65F602911D21FBCB" box="[1756,1834,1023,1045]" gridcol="6" gridrow="6" pageId="7" pageNumber="490">4.8</td>
</tr>
<tr id="C7EE383E5B44001E635705471D21FBE1" box="[125,1834,1065,1087]" gridrow="7" pageId="7" pageNumber="490">
<th id="843F51425B44001E635705471B6BFBE1" box="[125,352,1065,1087]" gridcol="0" gridrow="7" pageId="7" pageNumber="490">Legs I­III: Toe 1</th>
<td id="843F51425B44001E62F70547184DFBE1" box="[477,582,1065,1087]" gridcol="1" gridrow="7" pageId="7" pageNumber="490">7.0</td>
<td id="843F51425B44001E617B0547194DFBE1" box="[593,838,1065,1087]" gridcol="2" gridrow="7" pageId="7" pageNumber="490">7.4 ±1.44 (5.3­10.1); 11</td>
<td id="843F51425B44001E609B054719F4FBE1" box="[945,1023,1065,1087]" gridcol="3" gridrow="7" pageId="7" pageNumber="490">6.3</td>
<td id="843F51425B44001E670E05471F0FFBE1" box="[1060,1284,1065,1087]" gridcol="4" gridrow="7" pageId="7" pageNumber="490">- (7.3); 1</td>
<td id="843F51425B44001E665005471C6EFBE1" box="[1402,1637,1065,1087]" gridcol="5" gridrow="7" pageId="7" pageNumber="490">- (5.1­10.5); 2</td>
<td id="843F51425B44001E65F605471D21FBE1" box="[1756,1834,1065,1087]" gridcol="6" gridrow="7" pageId="7" pageNumber="490">6.0</td>
</tr>
<tr id="C7EE383E5B44001E6357053C1D21FBB6" box="[125,1834,1106,1128]" gridrow="8" pageId="7" pageNumber="490">
<th id="843F51425B44001E6357053C1B6BFBB6" box="[125,352,1106,1128]" gridcol="0" gridrow="8" pageId="7" pageNumber="490">Toe 2</th>
<td id="843F51425B44001E62F7053C184DFBB6" box="[477,582,1106,1128]" gridcol="1" gridrow="8" pageId="7" pageNumber="490">3.2</td>
<td id="843F51425B44001E617B053C194DFBB6" box="[593,838,1106,1128]" gridcol="2" gridrow="8" pageId="7" pageNumber="490">4.2 ±0.94 (2.5­5.9); 11</td>
<td id="843F51425B44001E609B053C19F4FBB6" box="[945,1023,1106,1128]" gridcol="3" gridrow="8" pageId="7" pageNumber="490">2.4</td>
<td id="843F51425B44001E670E053C1F0FFBB6" box="[1060,1284,1106,1128]" gridcol="4" gridrow="8" pageId="7" pageNumber="490">- (3.8); 1</td>
<td id="843F51425B44001E6650053C1C6EFBB6" box="[1402,1637,1106,1128]" gridcol="5" gridrow="8" pageId="7" pageNumber="490">- (2.6­4.9); 2</td>
<td id="843F51425B44001E65F6053C1D21FBB6" box="[1756,1834,1106,1128]" gridcol="6" gridrow="8" pageId="7" pageNumber="490">-</td>
</tr>
<tr id="C7EE383E5B44001E635705121D21FB4C" box="[125,1834,1148,1170]" gridrow="9" pageId="7" pageNumber="490">
<th id="843F51425B44001E635705121B6BFB4C" box="[125,352,1148,1170]" gridcol="0" gridrow="9" pageId="7" pageNumber="490">Toe 3</th>
<td id="843F51425B44001E62F70512184DFB4C" box="[477,582,1148,1170]" gridcol="1" gridrow="9" pageId="7" pageNumber="490">13.0</td>
<td id="843F51425B44001E617B0512194DFB4C" box="[593,838,1148,1170]" gridcol="2" gridrow="9" pageId="7" pageNumber="490">11.8 ±2.35 (8.8­15.5); 11</td>
<td id="843F51425B44001E609B051219F4FB4C" box="[945,1023,1148,1170]" gridcol="3" gridrow="9" pageId="7" pageNumber="490">13.7</td>
<td id="843F51425B44001E670E05121F0FFB4C" box="[1060,1284,1148,1170]" gridcol="4" gridrow="9" pageId="7" pageNumber="490">- (11.1); 1</td>
<td id="843F51425B44001E665005121C6EFB4C" box="[1402,1637,1148,1170]" gridcol="5" gridrow="9" pageId="7" pageNumber="490">- (9.7­13.0); 2</td>
<td id="843F51425B44001E65F605121D21FB4C" box="[1756,1834,1148,1170]" gridcol="6" gridrow="9" pageId="7" pageNumber="490">7.4</td>
</tr>
<tr id="C7EE383E5B44001E635705CB1D21FB65" box="[125,1834,1189,1211]" gridrow="10" pageId="7" pageNumber="490">
<th id="843F51425B44001E635705CB1B6BFB65" box="[125,352,1189,1211]" gridcol="0" gridrow="10" pageId="7" pageNumber="490">Toe 4</th>
<td id="843F51425B44001E62F705CB184DFB65" box="[477,582,1189,1211]" gridcol="1" gridrow="10" pageId="7" pageNumber="490">6.2</td>
<td id="843F51425B44001E617B05CB194DFB65" box="[593,838,1189,1211]" gridcol="2" gridrow="10" pageId="7" pageNumber="490">6.7 ±1.47 (3.7­8.9); 11</td>
<td id="843F51425B44001E609B05CB19F4FB65" box="[945,1023,1189,1211]" gridcol="3" gridrow="10" pageId="7" pageNumber="490">6.4</td>
<td id="843F51425B44001E670E05CB1F0FFB65" box="[1060,1284,1189,1211]" gridcol="4" gridrow="10" pageId="7" pageNumber="490">- (6.0); 1</td>
<td id="843F51425B44001E665005CB1C6EFB65" box="[1402,1637,1189,1211]" gridcol="5" gridrow="10" pageId="7" pageNumber="490">- (5.2­7.1); 2</td>
<td id="843F51425B44001E65F605CB1D21FB65" box="[1756,1834,1189,1211]" gridcol="6" gridrow="10" pageId="7" pageNumber="490">7.9</td>
</tr>
<tr id="C7EE383E5B44001E635705A11D21FB3B" box="[125,1834,1231,1253]" gridrow="11" pageId="7" pageNumber="490">
<th id="843F51425B44001E635705A11B6BFB3B" box="[125,352,1231,1253]" gridcol="0" gridrow="11" pageId="7" pageNumber="490">Toe 5</th>
<td id="843F51425B44001E62F705A1184DFB3B" box="[477,582,1231,1253]" gridcol="1" gridrow="11" pageId="7" pageNumber="490">14.1</td>
<td id="843F51425B44001E617B05A1194DFB3B" box="[593,838,1231,1253]" gridcol="2" gridrow="11" pageId="7" pageNumber="490">13.1 ±2.05 (9.9­15.7); 10</td>
<td id="843F51425B44001E609B05A119F4FB3B" box="[945,1023,1231,1253]" gridcol="3" gridrow="11" pageId="7" pageNumber="490">16.0</td>
<td id="843F51425B44001E670E05A11F0FFB3B" box="[1060,1284,1231,1253]" gridcol="4" gridrow="11" pageId="7" pageNumber="490">- (15.5); 1</td>
<td id="843F51425B44001E665005A11C6EFB3B" box="[1402,1637,1231,1253]" gridcol="5" gridrow="11" pageId="7" pageNumber="490">- (11.0­14.1); 2</td>
<td id="843F51425B44001E65F605A11D21FB3B" box="[1756,1834,1231,1253]" gridcol="6" gridrow="11" pageId="7" pageNumber="490">-</td>
</tr>
<tr id="C7EE383E5B44001E635705961D21FAD0" box="[125,1834,1272,1294]" gridrow="12" pageId="7" pageNumber="490">
<th id="843F51425B44001E635705961B6BFAD0" box="[125,352,1272,1294]" gridcol="0" gridrow="12" pageId="7" pageNumber="490">Toe 6</th>
<td id="843F51425B44001E62F70596184DFAD0" box="[477,582,1272,1294]" gridcol="1" gridrow="12" pageId="7" pageNumber="490">7.3</td>
<td id="843F51425B44001E617B0596194DFAD0" box="[593,838,1272,1294]" gridcol="2" gridrow="12" pageId="7" pageNumber="490">8.2 ±1.51 (4.8­9.7); 10</td>
<td id="843F51425B44001E609B059619F4FAD0" box="[945,1023,1272,1294]" gridcol="3" gridrow="12" pageId="7" pageNumber="490">8.9</td>
<td id="843F51425B44001E670E05961F0FFAD0" box="[1060,1284,1272,1294]" gridcol="4" gridrow="12" pageId="7" pageNumber="490">- (9.4); 1</td>
<td id="843F51425B44001E665005961C6EFAD0" box="[1402,1637,1272,1294]" gridcol="5" gridrow="12" pageId="7" pageNumber="490">- (6.9­9.2); 2</td>
<td id="843F51425B44001E65F605961D21FAD0" box="[1756,1834,1272,1294]" gridcol="6" gridrow="12" pageId="7" pageNumber="490">13.5</td>
</tr>
<tr id="C7EE383E5B44001E6357044C1D21FAE6" box="[125,1834,1314,1336]" gridrow="13" pageId="7" pageNumber="490">
<th id="843F51425B44001E6357044C1B6BFAE6" box="[125,352,1314,1336]" gridcol="0" gridrow="13" pageId="7" pageNumber="490">Leg IV: Toe 1</th>
<td id="843F51425B44001E62F7044C184DFAE6" box="[477,582,1314,1336]" gridcol="1" gridrow="13" pageId="7" pageNumber="490">11.3</td>
<td id="843F51425B44001E617B044C194DFAE6" box="[593,838,1314,1336]" gridcol="2" gridrow="13" pageId="7" pageNumber="490">11.2 ±0.92 (8.8­12.5); 12</td>
<td id="843F51425B44001E609B044C19F4FAE6" box="[945,1023,1314,1336]" gridcol="3" gridrow="13" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E044C1F0FFAE6" box="[1060,1284,1314,1336]" gridcol="4" gridrow="13" pageId="7" pageNumber="490">- (9.6­11.8); 2</td>
<td id="843F51425B44001E6650044C1C6EFAE6" box="[1402,1637,1314,1336]" gridcol="5" gridrow="13" pageId="7" pageNumber="490">11.0 ±1.39 (9.3­12.7); 4</td>
<td id="843F51425B44001E65F6044C1D21FAE6" box="[1756,1834,1314,1336]" gridcol="6" gridrow="13" pageId="7" pageNumber="490">7.0</td>
</tr>
<tr id="C7EE383E5B44001E635704221D21FABC" box="[125,1834,1356,1378]" gridrow="14" pageId="7" pageNumber="490">
<th id="843F51425B44001E635704221B6BFABC" box="[125,352,1356,1378]" gridcol="0" gridrow="14" pageId="7" pageNumber="490">Toe 2</th>
<td id="843F51425B44001E62F70422184DFABC" box="[477,582,1356,1378]" gridcol="1" gridrow="14" pageId="7" pageNumber="490">16.3</td>
<td id="843F51425B44001E617B0422194DFABC" box="[593,838,1356,1378]" gridcol="2" gridrow="14" pageId="7" pageNumber="490">16.2 ±1.94 (12.6­18.9); 11</td>
<td id="843F51425B44001E609B042219F4FABC" box="[945,1023,1356,1378]" gridcol="3" gridrow="14" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E04221F0FFABC" box="[1060,1284,1356,1378]" gridcol="4" gridrow="14" pageId="7" pageNumber="490">- (14.8­17.2); 2</td>
<td id="843F51425B44001E665004221C6EFABC" box="[1402,1637,1356,1378]" gridcol="5" gridrow="14" pageId="7" pageNumber="490">14.7 ±2.07 (11.7­16.2); 4</td>
<td id="843F51425B44001E65F604221D21FABC" box="[1756,1834,1356,1378]" gridcol="6" gridrow="14" pageId="7" pageNumber="490">15.6</td>
</tr>
<tr id="C7EE383E5B44001E6357041B1D21FA55" box="[125,1834,1397,1419]" gridrow="15" pageId="7" pageNumber="490">
<th id="843F51425B44001E6357041B1B6BFA55" box="[125,352,1397,1419]" gridcol="0" gridrow="15" pageId="7" pageNumber="490">Toe 3</th>
<td id="843F51425B44001E62F7041B184DFA55" box="[477,582,1397,1419]" gridcol="1" gridrow="15" pageId="7" pageNumber="490">6.4</td>
<td id="843F51425B44001E617B041B194DFA55" box="[593,838,1397,1419]" gridcol="2" gridrow="15" pageId="7" pageNumber="490">6.5 ±1.10 (3.7­8.1); 12</td>
<td id="843F51425B44001E609B041B19F4FA55" box="[945,1023,1397,1419]" gridcol="3" gridrow="15" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E041B1F0FFA55" box="[1060,1284,1397,1419]" gridcol="4" gridrow="15" pageId="7" pageNumber="490">- (3.6­5.7); 2</td>
<td id="843F51425B44001E6650041B1C6EFA55" box="[1402,1637,1397,1419]" gridcol="5" gridrow="15" pageId="7" pageNumber="490">5.0 ±1.00 (4.0­6.0); 3</td>
<td id="843F51425B44001E65F6041B1D21FA55" box="[1756,1834,1397,1419]" gridcol="6" gridrow="15" pageId="7" pageNumber="490">-</td>
</tr>
<tr id="C7EE383E5B44001E635704F11D21FA6B" box="[125,1834,1439,1461]" gridrow="16" pageId="7" pageNumber="490">
<th id="843F51425B44001E635704F11B6BFA6B" box="[125,352,1439,1461]" gridcol="0" gridrow="16" pageId="7" pageNumber="490">Toe 4</th>
<td id="843F51425B44001E62F704F1184DFA6B" box="[477,582,1439,1461]" gridcol="1" gridrow="16" pageId="7" pageNumber="490">10.7</td>
<td id="843F51425B44001E617B04F1194DFA6B" box="[593,838,1439,1461]" gridcol="2" gridrow="16" pageId="7" pageNumber="490">10.6 ±0.92 (9.1­11.9); 12</td>
<td id="843F51425B44001E609B04F119F4FA6B" box="[945,1023,1439,1461]" gridcol="3" gridrow="16" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E04F11F0FFA6B" box="[1060,1284,1439,1461]" gridcol="4" gridrow="16" pageId="7" pageNumber="490">- (7.5­9.3); 2</td>
<td id="843F51425B44001E665004F11C6EFA6B" box="[1402,1637,1439,1461]" gridcol="5" gridrow="16" pageId="7" pageNumber="490">8.8 ±1.96 (7.4­10.2); 3</td>
<td id="843F51425B44001E65F604F11D21FA6B" box="[1756,1834,1439,1461]" gridcol="6" gridrow="16" pageId="7" pageNumber="490">-</td>
</tr>
<tr id="C7EE383E5B44001E635704A61D21FA00" box="[125,1834,1480,1502]" gridrow="17" pageId="7" pageNumber="490">
<th id="843F51425B44001E635704A61B6BFA00" box="[125,352,1480,1502]" gridcol="0" gridrow="17" pageId="7" pageNumber="490">Toe 5</th>
<td id="843F51425B44001E62F704A6184DFA00" box="[477,582,1480,1502]" gridcol="1" gridrow="17" pageId="7" pageNumber="490">20.5</td>
<td id="843F51425B44001E617B04A6194DFA00" box="[593,838,1480,1502]" gridcol="2" gridrow="17" pageId="7" pageNumber="490">19.3 ±2.31 (15.3­22.6); 11</td>
<td id="843F51425B44001E609B04A619F4FA00" box="[945,1023,1480,1502]" gridcol="3" gridrow="17" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E04A61F0FFA00" box="[1060,1284,1480,1502]" gridcol="4" gridrow="17" pageId="7" pageNumber="490">- (12.8­15.8); 2</td>
<td id="843F51425B44001E665004A61C6EFA00" box="[1402,1637,1480,1502]" gridcol="5" gridrow="17" pageId="7" pageNumber="490">17.9 ±2.73 (14.2­20.7); 4</td>
<td id="843F51425B44001E65F604A61D21FA00" box="[1756,1834,1480,1502]" gridcol="6" gridrow="17" pageId="7" pageNumber="490">16.2</td>
</tr>
<tr id="C7EE383E5B44001E6357049C1D21F9D6" box="[125,1834,1522,1544]" gridrow="18" pageId="7" pageNumber="490">
<th id="843F51425B44001E6357049C1B6BF9D6" box="[125,352,1522,1544]" gridcol="0" gridrow="18" pageId="7" pageNumber="490">Toe 6</th>
<td id="843F51425B44001E62F7049C184DF9D6" box="[477,582,1522,1544]" gridcol="1" gridrow="18" pageId="7" pageNumber="490">11.0</td>
<td id="843F51425B44001E617B049C194DF9D6" box="[593,838,1522,1544]" gridcol="2" gridrow="18" pageId="7" pageNumber="490">10.7 ±1.04 (9.0­12.1); 11</td>
<td id="843F51425B44001E609B049C19F4F9D6" box="[945,1023,1522,1544]" gridcol="3" gridrow="18" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E049C1F0FF9D6" box="[1060,1284,1522,1544]" gridcol="4" gridrow="18" pageId="7" pageNumber="490">- (8.5­10.8); 2</td>
<td id="843F51425B44001E6650049C1C6EF9D6" box="[1402,1637,1522,1544]" gridcol="5" gridrow="18" pageId="7" pageNumber="490">10.2 ±1.26 (8.5­11.5); 4</td>
<td id="843F51425B44001E65F6049C1D21F9D6" box="[1756,1834,1522,1544]" gridcol="6" gridrow="18" pageId="7" pageNumber="490">?</td>
</tr>
<tr id="C7EE383E5B44001E635707751D21F9EF" box="[125,1834,1563,1585]" gridrow="19" pageId="7" pageNumber="490">
<th id="843F51425B44001E635707751B6BF9EF" box="[125,352,1563,1585]" gridcol="0" gridrow="19" pageId="7" pageNumber="490">Gonopore - Anus</th>
<td id="843F51425B44001E62F70775184DF9EF" box="[477,582,1563,1585]" gridcol="1" gridrow="19" pageId="7" pageNumber="490">10.9</td>
<td id="843F51425B44001E617B0775194DF9EF" box="[593,838,1563,1585]" gridcol="2" gridrow="19" pageId="7" pageNumber="490">8.0 ±2.07 (4.7­11.0); 14</td>
<td id="843F51425B44001E609B077519F4F9EF" box="[945,1023,1563,1585]" gridcol="3" gridrow="19" pageId="7" pageNumber="490">4.5</td>
<td id="843F51425B44001E670E07751F0FF9EF" box="[1060,1284,1563,1585]" gridcol="4" gridrow="19" pageId="7" pageNumber="490">5.5 ±0.49 (5.0­6.1); 5</td>
<td id="843F51425B44001E665007751C6EF9EF" box="[1402,1637,1563,1585]" gridcol="5" gridrow="19" pageId="7" pageNumber="490">-</td>
<td id="843F51425B44001E65F607751D21F9EF" box="[1756,1834,1563,1585]" gridcol="6" gridrow="19" pageId="7" pageNumber="490">-</td>
</tr>
</table>
</paragraph>
<paragraph id="79613A7C5B4BFFE963ED018E1B66FC32" blockId="8.[151,1437,151,1005]" pageId="8" pageNumber="491">
<taxonomicName id="BEDE41FF5B4BFFE963ED018E1BB0FF26" box="[199,443,224,248]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B4BFFE963ED018E1BB0FF26" box="[199,443,224,248]" italics="true" pageId="8" pageNumber="491">Batillipes brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4BFFE962E801B11817FF26" bold="true" box="[450,540,223,248]" pageId="8" pageNumber="491">
<taxonomicNameLabel id="50995B155B4BFFE962E801B11817FF26" box="[450,540,223,248]" pageId="8" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having swollen external cephalic cirri and the caudal apparatus consisting of an ala-like cuticular expansion covered with detritus is most similar to
<taxonomicName id="BEDE41FF5B4BFFE96099006B1E12FEC2" box="[947,1049,260,284]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE96099006B1E12FEC2" box="[947,1049,260,284]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
. However,
<taxonomicName id="BEDE41FF5B4BFFE967B1006B1F30FEC2" box="[1179,1339,260,284]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B4BFFE967B1006B1F30FEC2" box="[1179,1339,260,284]" italics="true" pageId="8" pageNumber="491">B. brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4BFFE96668006A1F97FEC3" bold="true" box="[1346,1436,260,285]" pageId="8" pageNumber="491">
<taxonomicNameLabel id="50995B155B4BFFE96668006A1F97FEC3" box="[1346,1436,260,285]" pageId="8" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
exhibits different shaped lateral projections with digit-shaped apices and the one between leg III and IV distally fringed. In addition, external cephalic cirri are horn-shaped in
<taxonomicName id="BEDE41FF5B4BFFE96078002319BCFEBA" box="[850,951,332,356]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE96078002319BCFEBA" box="[850,951,332,356]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
and terminated by a swollen tip in the new species. The two species, that have comparable body lengths, differ in the length of cirri E (the longest cirrus E in the new species is 13.1 µm long while in
<taxonomicName id="BEDE41FF5B4BFFE9614100FB18DAFE72" box="[619,721,404,428]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE9614100FB18DAFE72" box="[619,721,404,428]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
the shortest cirrus E is 19.0 µm long) and leg sense organs that are shorter in the new species. In particular, the sense organ of leg IV is much shorter in the new species (10% of the body length) than in
<taxonomicName id="BEDE41FF5B4BFFE9629C00B31815FE2A" box="[438,542,476,500]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE9629C00B31815FE2A" box="[438,542,476,500]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
where it is comprised of between 20 and 26 % of the body length. Cuticular valved structures associated with the female gonopore are present in both species, but in
<taxonomicName id="BEDE41FF5B4BFFE96757036F1EE9FDC6" box="[1149,1250,512,536]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE96757036F1EE9FDC6" box="[1149,1250,512,536]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
only two valves are present posterior to the gonopore, while there are four in the new species, two anterior and two posterior. Moreover, the new species differs from
<taxonomicName id="BEDE41FF5B4BFFE96146032718DFFDBE" box="[620,724,584,608]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE96146032718DFFDBE" box="[620,724,584,608]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
(and from all
<taxonomicName id="BEDE41FF5B4BFFE960A9032619E4FDBE" box="[899,1007,584,608]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="genus">
<emphasis id="4BAAE66E5B4BFFE960A9032619E4FDBE" box="[899,1007,584,608]" italics="true" pageId="8" pageNumber="491">Batillipes</emphasis>
</taxonomicName>
species) by a peculiar character: the presence of small semicircular cuticular projections on the coxal region of legs I–III. This character, that cannot be confused with the coxal swelling of the leg articulation, is also present in specimens previously collected in
<collectingCountry id="01C97AEC5B4BFFE9667203E11F97FD76" box="[1368,1436,655,680]" name="Brazil" pageId="8" pageNumber="491">Brazil</collectingCountry>
and attributed to
<taxonomicName id="BEDE41FF5B4BFFE9627E03DB18B1FD12" authority="da Rocha et al. 2013" authorityName="da Rocha et al." authorityYear="2013" box="[340,698,692,717]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE9627E03DB1BB2FD12" box="[340,441,692,716]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
(da Rocha
<emphasis id="4BAAE66E5B4BFFE9611D03DB1864FD12" box="[567,623,692,716]" italics="true" pageId="8" pageNumber="491">et al.</emphasis>
2013)
</taxonomicName>
. The re-examination of those specimens, which also have external cephalic cirri with swollen tips, led to the conclusion that they should be attributed to the new species. In those specimens mounted in glycerin, eyes were observed which seem to confirm the presence of these structures in the new species. Eyes are soluble in some mounting media, as the glycerol. The re-examination also confirms the intraspecific variability of the caudal apparatus and lateral projections located between legs III and IV that, together with the large amount of adherent detritus were the reason of misidentification. With this correction,
<taxonomicName id="BEDE41FF5B4BFFE967D602071F97FC5E" box="[1276,1436,872,896]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B4BFFE967D602071F97FC5E" box="[1276,1436,872,896]" italics="true" pageId="8" pageNumber="491">B. brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4BFFE963BD02E21AF8FC7B" bold="true" box="[151,243,908,933]" pageId="8" pageNumber="491">
<taxonomicNameLabel id="50995B155B4BFFE963BD02E21AF8FC7B" box="[151,243,908,933]" pageId="8" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also recorded for Cupe Beach,
<collectingRegion id="BB1AF49E5B4BFFE961AA02E21907FC7B" box="[640,780,908,933]" country="Brazil" name="Pernambuco" pageId="8" pageNumber="491">Pernambuco</collectingRegion>
State (
<geoCoordinate id="1CEA5CBB5B4BFFE9607502E219A2FC7B" box="[863,937,908,933]" direction="south" orientation="latitude" pageId="8" pageNumber="491" precision="925" value="-8.416667">8°25'S</geoCoordinate>
,
<geoCoordinate id="1CEA5CBB5B4BFFE9609D02E21E17FC7B" box="[951,1052,908,933]" direction="west" orientation="longitude" pageId="8" pageNumber="491" precision="925" value="-34.916668">34°55'W</geoCoordinate>
), and for the continental shelf, in medium sands at
<quantity id="BE2697995B4BFFE9624A02C11B90FC19" box="[352,411,943,968]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="491" unit="m" value="35.0">35 m</quantity>
depth, of
<collectingRegion id="BB1AF49E5B4BFFE9613802DE1858FC16" box="[530,595,944,968]" country="Brazil" name="Ceara" pageId="8" pageNumber="491">Ceará</collectingRegion>
and
<collectingRegion id="BB1AF49E5B4BFFE961A402C1198BFC19" box="[654,896,943,968]" country="Brazil" name="Rio Grande do Norte" pageId="8" pageNumber="491">Rio Grande do Norte</collectingRegion>
States (4°20'–
<geoCoordinate id="1CEA5CBB5B4BFFE9670F02C11E65FC16" box="[1061,1134,943,968]" direction="south" orientation="latitude" pageId="8" pageNumber="491" precision="925" value="-4.8333335">4°50'S</geoCoordinate>
; 36°00–
<geoCoordinate id="1CEA5CBB5B4BFFE967E702C11F3DFC16" box="[1229,1334,943,968]" direction="west" orientation="longitude" pageId="8" pageNumber="491" precision="925" value="-37.25">37°15' W</geoCoordinate>
), (see da Rocha
<emphasis id="4BAAE66E5B4BFFE963CF02BB1B16FC32" box="[229,285,980,1004]" italics="true" pageId="8" pageNumber="491">et al.</emphasis>
2013).
</paragraph>
</subSubSection>
</treatment>
</document>

590
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View file

@ -0,0 +1,590 @@
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<mods:title id="D6907DB4689626A19F0B3406ED7C1454">Three new Batillipes species (Arthrotardigrada: Batillipedidae) from the Brazilian coast</mods:title>
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<treatment id="F1778B6A5B4BFFEC63BD05541977FC83" ID-DOI="http://doi.org/10.5281/zenodo.5617030" ID-GBIF-Taxon="127844695" ID-Zenodo-Dep="5617030" LSID="urn:lsid:plazi:treatment:F1778B6A5B4BFFEC63BD05541977FC83" httpUri="http://treatment.plazi.org/id/F1778B6A5B4BFFEC63BD05541977FC83" lastPageId="13" lastPageNumber="496" pageId="8" pageNumber="491">
<subSubSection id="31C469F75B4BFFE963BD05541B66FBA8" pageId="8" pageNumber="491" type="nomenclature">
<paragraph id="79613A7C5B4BFFE963BD05541BECFB8B" blockId="8.[151,487,1082,1142]" box="[151,487,1082,1109]" pageId="8" pageNumber="491">
<heading id="22298D105B4BFFE963BD05541BECFB8B" bold="true" box="[151,487,1082,1109]" fontSize="11" level="1" pageId="8" pageNumber="491" reason="1">
<emphasis id="4BAAE66E5B4BFFE963BD05541BECFB8B" bold="true" box="[151,487,1082,1109]" pageId="8" pageNumber="491">
<taxonomicName id="BEDE41FF5B4BFFE963BD05541B89FB8A" ID-CoL="9c4449fe-cb06-427e-8889-377b6ef16d40" box="[151,386,1082,1108]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4BFFE963BD05541B89FB8A" bold="true" box="[151,386,1082,1108]" italics="true" pageId="8" pageNumber="491">Batillipes dandarae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50995B155B4BFFE962A305551BECFB8B" box="[393,487,1083,1109]" pageId="8" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="79613A7C5B4BFFE963BD05331B66FBA8" blockId="8.[151,487,1082,1142]" box="[151,365,1117,1142]" pageId="8" pageNumber="491">
(
<figureCitation id="E1E526F95B4BFFE963B505331B0FFBA8" box="[159,260,1117,1142]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="9.[151,250,1733,1755]" captionStartId-1="11.[151,250,1908,1930]" captionTargetBox-0="[176,1418,183,1711]" captionTargetBox-1="[175,1418,414,1883]" captionTargetId-0="figure@9.[174,1418,181,1711]" captionTargetId-1="figure@11.[172,1418,411,1883]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionText-0="FIGURE 4. Batillipes dandarae sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view); C. Detailed drawing of the female gonopore (go) and anus (an); D. Dorsal view of the head showing the median cirrus (cm) and the eye (ey); E. Ventral view of the head showing the buccal cone (bc) and the external cephalic cirrus (ce). Scale bars: A – B = 50 µm; C – E = 10 µm." captionText-1="FIGURE 5. Batillipes dandarae sp. nov. A. Anterior portion of the body of the holotype showing cephalic appendages (ciinternal cirrus; cl—lateral cirrus); clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and blunt enlargement in the scapular region (sr); B. Leg I of a paratype showing the leg spine (pI) inserted on the posterior part of the leg and turning frontwards, mo—mouth; C. Detail of the female gonopore (go) and anus (an) of the holotype. Scale bars = 10 µm." httpUri-0="https://zenodo.org/record/400157/files/figure.png" httpUri-1="https://zenodo.org/record/400158/files/figure.png" pageId="8" pageNumber="491">Figs 4–5</figureCitation>
;
<tableCitation id="345C0FC75B4BFFE9623A05331B6FFBA8" box="[272,356,1117,1142]" captionStart="TABLE 2" captionStartId="10.[151,239,476,499]" captionTargetBox="[155,1408,590,2007]" captionTargetPageId="10" captionText="TABLE 2. Measurements (in µm) of selected morphological structures of specimens of Batillipes dandarae sp. nov. (Holo—Holotype; SD—Standard deviation; Range refers to the smallest and largest measured specimen / structure; Nnumber of specimens / structures measured; Allo—Allotype)." httpUri="http://table.plazi.org/id/2DA16AF45B49FFEB63BD00B21917FDED" pageId="8" pageNumber="491" tableUuid="2DA16AF45B49FFEB63BD00B21917FDED">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B4BFFE963BD05CB1BC3FA64" pageId="8" pageNumber="491" type="diagnosis">
<paragraph id="79613A7C5B4BFFE963BD05CB1BC3FA64" blockId="8.[151,1437,1189,2006]" pageId="8" pageNumber="491">
<emphasis id="4BAAE66E5B4BFFE963BD05CB1B01FB60" bold="true" box="[151,266,1189,1214]" pageId="8" pageNumber="491">Diagnosis</emphasis>
. Medium sized
<taxonomicName id="BEDE41FF5B4BFFE962EB05CB1827FB63" box="[449,556,1189,1213]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="genus">
<emphasis id="4BAAE66E5B4BFFE962EB05CB1827FB63" box="[449,556,1189,1213]" italics="true" pageId="8" pageNumber="491">Batillipes</emphasis>
</taxonomicName>
with cylindrical undivided primary clavae. Papillar secondary clavae present. Sensorial leg spines and cirri, including all cephalic cirri, lateral cirri and cirri E, with an optically dense dot near their blunt tips. Sensorial spine on leg I inserted on the posterior part of the leg and turning frontwards. Toes 3 and 4 of legs IV with different lengths. Lateral processes between legs absent. Pointed triangular caudal appendage very variable in shape and size among specimens. Finely punctated cuticle comprised of small pillars; dorsal punctations uniformly distributed. Rosette-like female gonopore separated from the anus by a groove. Males with circular gonopore with crescent-shaped cuticular fold. Anus surrounded by a peculiar cuticular structure constituted by six platelets.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B4BFFE963ED04AB1BE6F8FC" pageId="8" pageNumber="491" type="materials_examined">
<paragraph id="79613A7C5B4BFFE963ED04AB19E1FA00" blockId="8.[151,1437,1189,2006]" box="[199,1002,1477,1502]" pageId="8" pageNumber="491">
<materialsCitation id="C9B630215B4BFFE963ED04AB19E1FA00" ID-GBIF-Occurrence="1440156680" box="[199,1002,1477,1502]" country="Brazil" latitude="-9.8625" location="Gunga Beach" longLatPrecision="21" longitude="-35.904724" pageId="8" pageNumber="491" specimenCount="1" stateProvince="Alagoas" typeStatus="holotype">
<emphasis id="4BAAE66E5B4BFFE963ED04AB1B6EFA00" bold="true" box="[199,357,1477,1502]" pageId="8" pageNumber="491">
<typeStatus id="A66584DE5B4BFFE963ED04AB1B09FA00" box="[199,258,1477,1502]" pageId="8" pageNumber="491">Type</typeStatus>
locality.
</emphasis>
<location id="7C016CA75B4BFFE9624604A8180DFA00" LSID="urn:lsid:plazi:treatment:F1778B6A5B4BFFEC63BD05541977FC83:7C016CA75B4BFFE9624604A8180DFA00" box="[364,518,1477,1502]" country="Brazil" latitude="-9.8625" longLatPrecision="21" longitude="-35.904724" name="Gunga Beach" pageId="8" pageNumber="491" stateProvince="Alagoas">Gunga Beach</location>
,
<collectingRegion id="BB1AF49E5B4BFFE9613804AB187BFA00" box="[530,624,1477,1502]" country="Brazil" name="Alagoas" pageId="8" pageNumber="491">Alagoas</collectingRegion>
,
<collectingCountry id="01C97AEC5B4BFFE9615104AB18B4FA00" box="[635,703,1477,1502]" name="Brazil" pageId="8" pageNumber="491">Brazil</collectingCountry>
(
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,
<geoCoordinate id="1CEA5CBB5B4BFFE9606504AB19D5FA00" box="[847,990,1477,1502]" direction="west" orientation="longitude" pageId="8" pageNumber="491" precision="15" value="-35.904724">35°54'17&quot;W</geoCoordinate>
).
</materialsCitation>
</paragraph>
<paragraph id="79613A7C5B4BFFE963ED04871F38F94C" blockId="8.[151,1437,1189,2006]" pageId="8" pageNumber="491">
<emphasis id="4BAAE66E5B4BFFE963ED04871B71F9DC" bold="true" box="[199,378,1513,1538]" pageId="8" pageNumber="491">Type material.</emphasis>
Holotype: adult, female (slide CVII-81) collected at Gunga Beach, mounted in glycerol. Allotypic male (slide TARD/UFRPE 02-18), collected at Forte Orange Beach, mounted in polyvinyl alcohol. Paratypes: 23 specimens (6 females, 17 juveniles) collected at Forte Orange Beach, mounted in polyvinyl alcohol (slides TARD/UFRPE
<date id="0D601CBC5B4BFFE962B1073B1841F9B0" box="[411,586,1621,1646]" pageId="8" pageNumber="491" value="1921-02-15" valueMax="1921-02-02" valueMin="1921-02-15">02-15 to 02-21</date>
); 1 female collected in Ponta do Sal Beach, mounted in polyvinyl alcohol (slide CVII-83); 2 females collected in Patacho Beach, mounted in glycerol (slide TARD/UFRPE 02-22).
</paragraph>
<paragraph id="79613A7C5B4BFFE963ED07F31BE6F8FC" blockId="8.[151,1437,1189,2006]" pageId="8" pageNumber="491">
<materialsCitation id="C9B630215B4BFFE963ED07F31BE6F8FC" ID-GBIF-Occurrence="1440156684" country="Portugal" county="Department" location="Faculty" municipality="Biology" pageId="8" pageNumber="491" specimenCount="4" stateProvince="Pernambuco" typeStatus="holotype">
<emphasis id="4BAAE66E5B4BFFE963ED07F31B84F968" bold="true" box="[199,399,1693,1718]" pageId="8" pageNumber="491">
<typeStatus id="A66584DE5B4BFFE963ED07F31B09F968" box="[199,258,1693,1718]" pageId="8" pageNumber="491">Type</typeStatus>
repository.
</emphasis>
The
<typeStatus id="A66584DE5B4BFFE962FF07F0180EF968" box="[469,517,1694,1718]" pageId="8" pageNumber="491">type</typeStatus>
material (
<typeStatus id="A66584DE5B4BFFE961AE07F318E3F968" box="[644,744,1693,1718]" pageId="8" pageNumber="491" type="holotype">holotype</typeStatus>
slide CVII-81 and
<typeStatus id="A66584DE5B4BFFE960F007F01E31F968" box="[986,1082,1694,1718]" pageId="8" pageNumber="491" type="paratype">paratype</typeStatus>
CVII-83) is deposited in the collection of the
<collectingCounty id="900042F05B4BFFE9627F07AC1BD2F907" box="[341,473,1730,1753]" pageId="8" pageNumber="491">Department</collectingCounty>
of
<collectingMunicipality id="9905A0065B4BFFE962D507AF1851F904" box="[511,602,1729,1754]" pageId="8" pageNumber="491">Biology</collectingMunicipality>
,
<location id="7C016CA75B4BFFE9614C07AF18B1F904" LSID="urn:lsid:plazi:treatment:F1778B6A5B4BFFEC63BD05541977FC83:7C016CA75B4BFFE9614C07AF18B1F904" box="[614,698,1729,1754]" country="Portugal" county="Department" municipality="Biology" name="Faculty" pageId="8" pageNumber="491" stateProvince="Pernambuco">Faculty</location>
of
<location id="7C016CA75B4BFFE961CA07AF194EF904" LSID="urn:lsid:plazi:treatment:F1778B6A5B4BFFEC63BD05541977FC83:7C016CA75B4BFFE961CA07AF194EF904" box="[736,837,1729,1754]" country="Portugal" county="Department" municipality="Biology" name="Sciences" pageId="8" pageNumber="491" stateProvince="Pernambuco">Sciences</location>
,
<location id="7C016CA75B4BFFE9607B07AF19CCF904" LSID="urn:lsid:plazi:treatment:F1778B6A5B4BFFEC63BD05541977FC83:7C016CA75B4BFFE9607B07AF19CCF904" box="[849,967,1729,1754]" country="Portugal" county="Department" municipality="Biology" name="University" pageId="8" pageNumber="491" stateProvince="Pernambuco">University</location>
of
<location id="7C016CA75B4BFFE960C707AC1E27F907" LSID="urn:lsid:plazi:treatment:F1778B6A5B4BFFEC63BD05541977FC83:7C016CA75B4BFFE960C707AC1E27F907" box="[1005,1068,1730,1753]" country="Portugal" county="Department" municipality="Biology" name="Porto" pageId="8" pageNumber="491" stateProvince="Pernambuco">Porto</location>
,
<collectingCountry id="01C97AEC5B4BFFE9671207AF1E91F904" box="[1080,1178,1729,1754]" name="Portugal" pageId="8" pageNumber="491">Portugal</collectingCountry>
, and the other slides in the collection of Tardigrades—UFRPE (
<location id="7C016CA75B4BFFE96148078B18EBF920" LSID="urn:lsid:plazi:treatment:F1778B6A5B4BFFEC63BD05541977FC83:7C016CA75B4BFFE96148078B18EBF920" box="[610,736,1765,1790]" country="Portugal" county="Department" municipality="Biology" name="Laboratory" pageId="8" pageNumber="491" stateProvince="Pernambuco">Laboratory</location>
of
<location id="7C016CA75B4BFFE96020078B198DF920" LSID="urn:lsid:plazi:treatment:F1778B6A5B4BFFEC63BD05541977FC83:7C016CA75B4BFFE96020078B198DF920" box="[778,902,1765,1790]" country="Portugal" county="Department" municipality="Biology" name="Meiofauna" pageId="8" pageNumber="491" stateProvince="Pernambuco">Meiofauna</location>
,
<location id="7C016CA75B4BFFE960B907881E1CF923" LSID="urn:lsid:plazi:treatment:F1778B6A5B4BFFEC63BD05541977FC83:7C016CA75B4BFFE960B907881E1CF923" box="[915,1047,1766,1789]" country="Portugal" county="Department" municipality="Biology" name="Department" pageId="8" pageNumber="491" stateProvince="Pernambuco">Department</location>
of
<location id="7C016CA75B4BFFE9676B078B1E97F920" LSID="urn:lsid:plazi:treatment:F1778B6A5B4BFFEC63BD05541977FC83:7C016CA75B4BFFE9676B078B1E97F920" box="[1089,1180,1765,1790]" country="Portugal" county="Department" municipality="Biology" name="Biology" pageId="8" pageNumber="491" stateProvince="Pernambuco">Biology</location>
, Universidade Federal Rural de
<collectingRegion id="BB1AF49E5B4BFFE963D706671B87F8FC" box="[253,396,1801,1826]" country="Brazil" name="Pernambuco" pageId="8" pageNumber="491">Pernambuco</collectingRegion>
,
<collectingCountry id="01C97AEC5B4BFFE962BD06671BEAF8FC" box="[407,481,1801,1826]" name="Brazil" pageId="8" pageNumber="491">Brazil</collectingCountry>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B4BFFE963ED064319BCF808" pageId="8" pageNumber="491" type="etymology">
<paragraph id="79613A7C5B4BFFE963ED064318FFF8B4" blockId="8.[151,1437,1189,2006]" pageId="8" pageNumber="491">
<emphasis id="4BAAE66E5B4BFFE963ED06431B45F898" bold="true" box="[199,334,1837,1862]" pageId="8" pageNumber="491">Etymology.</emphasis>
The specific name,
<taxonomicName id="BEDE41FF5B4BFFE9610706431893F89B" box="[557,664,1837,1861]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="dandarae">
<emphasis id="4BAAE66E5B4BFFE9610706431893F89B" box="[557,664,1837,1861]" italics="true" pageId="8" pageNumber="491">dandarae</emphasis>
</taxonomicName>
, is in honor of “Dandara dos Palmares”, an ex-slave warrior, who led the fight against slavery in
<collectingCountry id="01C97AEC5B4BFFE962ED063F1800F8B4" box="[455,523,1873,1898]" name="Brazil" pageId="8" pageNumber="491">Brazil</collectingCountry>
in the XVII century.
</paragraph>
<paragraph id="79613A7C5B4BFFE963ED061B19BCF808" blockId="8.[151,1437,1189,2006]" pageId="8" pageNumber="491">
<emphasis id="4BAAE66E5B4BFFE963ED061B1B88F850" bold="true" box="[199,387,1909,1934]" pageId="8" pageNumber="491">Ecological note.</emphasis>
<taxonomicName id="BEDE41FF5B4BFFE962A10618181DF853" box="[395,534,1909,1933]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4BFFE962A10618181DF853" box="[395,534,1909,1933]" italics="true" pageId="8" pageNumber="491">B. dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4BFFE96137061B187CF850" bold="true" box="[541,631,1909,1934]" pageId="8" pageNumber="491">
<taxonomicNameLabel id="50995B155B4BFFE96137061B187CF850" box="[541,631,1909,1934]" pageId="8" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was mainly found in shallow sublittoral fine to medium calcareous and quartz sands in low energy and estuarine beaches as Gunga Beach and Forte Orange Beach and occasionally in high energy beaches (Ponta do Sal) and in reef pools as Patacho Beach.
</paragraph>
</subSubSection>
<caption id="2DA16AF45B4AFFE863BD07AB1F81F8C7" httpUri="https://zenodo.org/record/400157/files/figure.png" pageId="9" pageNumber="492" targetBox="[176,1418,183,1711]" targetPageId="9">
<paragraph id="79613A7C5B4AFFE863BD07AB1F81F8C7" blockId="9.[151,1436,1733,1817]" pageId="9" pageNumber="492">
<emphasis id="4BAAE66E5B4AFFE863BD07AB1B1BF904" bold="true" box="[151,272,1733,1755]" pageId="9" pageNumber="492">FIGURE 4</emphasis>
.
<taxonomicName id="BEDE41FF5B4AFFE8623507AB1BE0F905" box="[287,491,1733,1755]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="492" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4AFFE8623507AB1BE0F905" box="[287,491,1733,1755]" italics="true" pageId="9" pageNumber="492">Batillipes dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4AFFE862DE07A8184CF905" bold="true" box="[500,583,1734,1755]" pageId="9" pageNumber="492">
<taxonomicNameLabel id="50995B155B4AFFE862DE07A8184CF905" box="[500,583,1734,1755]" pageId="9" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Drawing (dorsal view); B. Habitus (dorsal view); C. Detailed drawing of the female gonopore (go) and anus (an); D. Dorsal view of the head showing the median cirrus (cm) and the eye (ey); E. Ventral view of the head showing the buccal cone (bc) and the external cephalic cirrus (ce). Scale bars: A–B = 50 µm; C–E = 10 µm.
</paragraph>
</caption>
<subSubSection id="31C469F75B4AFFED63ED062B1F62FC5E" lastPageId="12" lastPageNumber="495" pageId="9" pageNumber="492" type="description">
<paragraph id="79613A7C5B4AFFEB63ED062B1B55FE72" blockId="9.[151,1436,1861,2030]" lastBlockId="10.[151,1436,151,429]" lastPageId="10" lastPageNumber="493" pageId="9" pageNumber="492">
<emphasis id="4BAAE66E5B4AFFE863ED062B1BE2F880" bold="true" box="[199,489,1861,1886]" pageId="9" pageNumber="492">
Description of
<typeStatus id="A66584DE5B4AFFE86251062B1BEFF880" box="[379,484,1861,1886]" pageId="9" pageNumber="492" type="holotype">holotype</typeStatus>
.
</emphasis>
Female, 194 µm long (204 µm including the caudal apparatus) and 68.9 µm wide between the third and fourth pairs of legs (
<figureCitation id="E1E526F95B4AFFE861A5060718DEF85C" box="[655,725,1897,1922]" captionStart="FIGURE 4" captionStartId="9.[151,250,1733,1755]" captionTargetBox="[176,1418,183,1711]" captionTargetId="figure@9.[174,1418,181,1711]" captionTargetPageId="9" captionText="FIGURE 4. Batillipes dandarae sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view); C. Detailed drawing of the female gonopore (go) and anus (an); D. Dorsal view of the head showing the median cirrus (cm) and the eye (ey); E. Ventral view of the head showing the buccal cone (bc) and the external cephalic cirrus (ce). Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400157/files/figure.png" pageId="9" pageNumber="492">Fig. 4</figureCitation>
A, B). Eyes present (very faint and difficult to see, fig 4 D, probably dissolved by effect of the mounting media). Trapezoid head with eleven cephalic appendages (
<figureCitation id="E1E526F95B4AFFE8661B06E31F72F878" box="[1329,1401,1933,1958]" captionStart="FIGURE 4" captionStartId="9.[151,250,1733,1755]" captionTargetBox="[176,1418,183,1711]" captionTargetId="figure@9.[174,1418,181,1711]" captionTargetPageId="9" captionText="FIGURE 4. Batillipes dandarae sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view); C. Detailed drawing of the female gonopore (go) and anus (an); D. Dorsal view of the head showing the median cirrus (cm) and the eye (ey); E. Ventral view of the head showing the buccal cone (bc) and the external cephalic cirrus (ce). Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400157/files/figure.png" pageId="9" pageNumber="492">Figs 4</figureCitation>
D, E, 5 A): Internal cirri inserted dorsally on the frontal edge of the head are 23.2 µm long, bearing cirrophores (about 3.1 µm long). External cirri 16.1 µm long (
<figureCitation id="E1E526F95B4AFFE8615F06BB18BCF830" box="[629,695,2005,2030]" captionStart="FIGURE 4" captionStartId="9.[151,250,1733,1755]" captionTargetBox="[176,1418,183,1711]" captionTargetId="figure@9.[174,1418,181,1711]" captionTargetPageId="9" captionText="FIGURE 4. Batillipes dandarae sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view); C. Detailed drawing of the female gonopore (go) and anus (an); D. Dorsal view of the head showing the median cirrus (cm) and the eye (ey); E. Ventral view of the head showing the buccal cone (bc) and the external cephalic cirrus (ce). Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400157/files/figure.png" pageId="9" pageNumber="492">Fig. 4</figureCitation>
E), with indistinct cirrophores, inserted ventro-laterally, ventral to the pedestals bearing lateral cirri and primary clavae. The median cirrus, with cirrophore (about 3.4 µm long), is 24.8 µm long (
<figureCitation id="E1E526F95B49FFEB621601D21B73FF0A" box="[316,376,188,213]" captionStart="FIGURE 4" captionStartId="9.[151,250,1733,1755]" captionTargetBox="[176,1418,183,1711]" captionTargetId="figure@9.[174,1418,181,1711]" captionTargetPageId="9" captionText="FIGURE 4. Batillipes dandarae sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view); C. Detailed drawing of the female gonopore (go) and anus (an); D. Dorsal view of the head showing the median cirrus (cm) and the eye (ey); E. Ventral view of the head showing the buccal cone (bc) and the external cephalic cirrus (ce). Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400157/files/figure.png" pageId="10" pageNumber="493">Fig 4</figureCitation>
D). The lateral cirrus, 28.2 µm long, is located dorsally in relation to the unconstricted tubular primary clava which is 13.3 µm long. These two appendages share a common pedestal (
<figureCitation id="E1E526F95B49FFEB674701B11EA2FF26" box="[1133,1193,223,248]" captionStart="FIGURE 5" captionStartId="11.[151,250,1908,1930]" captionTargetBox="[175,1418,414,1883]" captionTargetId="figure@11.[172,1418,411,1883]" captionTargetPageId="11" captionText="FIGURE 5. Batillipes dandarae sp. nov. A. Anterior portion of the body of the holotype showing cephalic appendages (ciinternal cirrus; cl—lateral cirrus); clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and blunt enlargement in the scapular region (sr); B. Leg I of a paratype showing the leg spine (pI) inserted on the posterior part of the leg and turning frontwards, mo—mouth; C. Detail of the female gonopore (go) and anus (an) of the holotype. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400158/files/figure.png" pageId="10" pageNumber="493">Fig 5</figureCitation>
A). A van der Land’s organ is present at the base of the primary clava. All the cephalic cirri, including external cirri, and lateral cirri have an optically dense dot (black dot when observed under PHC) near their blunt tips. Indentation or notch in the frontal margin of the head between the external cirrus and the pedestal bearing the primary clava and lateral cirrus not observed. In the frontal edge of the head well developed (major diameter about 6.8 µm) secondary clavae are present (
<figureCitation id="E1E526F95B49FFEB63DD00FA1B39FE73" box="[247,306,404,429]" captionStart="FIGURE 5" captionStartId="11.[151,250,1908,1930]" captionTargetBox="[175,1418,414,1883]" captionTargetId="figure@11.[172,1418,411,1883]" captionTargetPageId="11" captionText="FIGURE 5. Batillipes dandarae sp. nov. A. Anterior portion of the body of the holotype showing cephalic appendages (ciinternal cirrus; cl—lateral cirrus); clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and blunt enlargement in the scapular region (sr); B. Leg I of a paratype showing the leg spine (pI) inserted on the posterior part of the leg and turning frontwards, mo—mouth; C. Detail of the female gonopore (go) and anus (an) of the holotype. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400158/files/figure.png" pageId="10" pageNumber="493">Fig 5</figureCitation>
A).
</paragraph>
<caption id="2DA16AF45B49FFEB63BD00B21917FDED" ID-Table-UUID="2DA16AF45B49FFEB63BD00B21917FDED" httpUri="http://table.plazi.org/id/2DA16AF45B49FFEB63BD00B21917FDED" pageId="10" pageNumber="493" targetBox="[155,1408,590,2007]" targetIsTable="true" targetPageId="10">
<paragraph id="79613A7C5B49FFEB63BD00B21917FDED" blockId="10.[151,1436,475,563]" pageId="10" pageNumber="493">
<emphasis id="4BAAE66E5B49FFEB63BD00B21B0DFE2D" bold="true" box="[151,262,476,499]" pageId="10" pageNumber="493">TABLE 2</emphasis>
. Measurements (in µm) of selected morphological structures of specimens of
<taxonomicName id="BEDE41FF5B49FFEB674F00B51F31FE2D" box="[1125,1338,475,499]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="10" pageNumber="493" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B49FFEB674F00B51F31FE2D" box="[1125,1338,475,499]" italics="true" pageId="10" pageNumber="493">Batillipes dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B49FFEB666E00B21F97FE2D" bold="true" box="[1348,1436,476,499]" pageId="10" pageNumber="493">
<taxonomicNameLabel id="50995B155B49FFEB666E00B21F97FE2D" box="[1348,1436,476,499]" pageId="10" pageNumber="493" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Holo—Holotype; SD—Standard deviation; Range refers to the smallest and largest measured specimen/structure; Nnumber of specimens/structures measured; Allo—Allotype).
</paragraph>
</caption>
<paragraph id="79613A7C5B49FFEB62C503201E4EF809" pageId="10" pageNumber="493">
<table id="0BDEC8DC5B49001E63B103201F8BF809" box="[155,1408,590,2007]" gridcols="4" gridrows="12" pageId="10" pageNumber="493">
<tr id="C7EE383E5B49001E63B103201F8BFD72" box="[155,1408,590,684]" gridrow="0" pageId="10" pageNumber="493" rowspan-0="1">
<th id="843F51425B49001E62C503201981FD72" box="[495,906,590,684]" gridcol="1" gridrow="0" pageId="10" pageNumber="493">FEMALES adults Holo Mean ± SD (Range); N</th>
<th id="843F51425B49001E60E103201E19FD72" box="[971,1042,590,684]" gridcol="2" gridrow="0" pageId="10" pageNumber="493">MALE adult Allo</th>
<th id="843F51425B49001E671703201F8BFD72" box="[1085,1408,590,684]" gridcol="3" gridrow="0" pageId="10" pageNumber="493">JUVENILES 6-toed Mean ± SD (Range); N</th>
</tr>
<tr id="C7EE383E5B49001E63B103AE1F8BFCF6" box="[155,1408,704,808]" gridrow="1" pageId="10" pageNumber="493">
<th id="843F51425B49001E63B103AE1BB5FCF6" box="[155,446,704,808]" gridcol="0" gridrow="1" pageId="10" pageNumber="493">STRUCTURES Body length (BL) BL without caudal apparatus</th>
<td id="843F51425B49001E62C503AE1981FCF6" box="[495,906,704,808]" gridcol="1" gridrow="1" pageId="10" pageNumber="493">204 176.5 ±36.1 (129.7–224.8); 8 194 165.4 ±34.6 (119–208); 10</td>
<td id="843F51425B49001E60E103AE1E19FCF6" box="[971,1042,704,808]" gridcol="2" gridrow="1" pageId="10" pageNumber="493">137.5 124.4</td>
<td id="843F51425B49001E671703AE1F8BFCF6" box="[1085,1408,704,808]" gridcol="3" gridrow="1" pageId="10" pageNumber="493">167.9 ±33.96 (127.4–219.1); 12 155.8 ±27.38 (122.2–200.1); 15</td>
</tr>
<tr id="C7EE383E5B49001E63B102521F8BFC7A" box="[155,1408,828,932]" gridrow="2" pageId="10" pageNumber="493">
<th id="843F51425B49001E63B102521BB5FC7A" box="[155,446,828,932]" gridcol="0" gridrow="2" pageId="10" pageNumber="493">Body width Median cirrus Internal cirri</th>
<td id="843F51425B49001E62C502521981FC7A" box="[495,906,828,932]" gridcol="1" gridrow="2" pageId="10" pageNumber="493">68.9 53.5 ±14.4 (33.6–73.9); 9 24.8 19.9 ±3.4 (13.5–24.8); 7 23.2 19.1 ±2.9 (14.3–23.2); 10</td>
<td id="843F51425B49001E60E102521E19FC7A" box="[971,1042,828,932]" gridcol="2" gridrow="2" pageId="10" pageNumber="493">56.5 15.1 15.3</td>
<td id="843F51425B49001E671702521F8BFC7A" box="[1085,1408,828,932]" gridcol="3" gridrow="2" pageId="10" pageNumber="493">53.2 ±10.95 (37.3–71.4); 9 19.4 ±4.29 (14.2–27.7); 9 18.2 ±2.51 (13.5–21.6); 11</td>
</tr>
<tr id="C7EE383E5B49001E63B102D61F8BFBFE" box="[155,1408,952,1056]" gridrow="3" pageId="10" pageNumber="493">
<th id="843F51425B49001E63B102D61BB5FBFE" box="[155,446,952,1056]" gridcol="0" gridrow="3" pageId="10" pageNumber="493">External cirri Lateral cirri A Primary clavae</th>
<td id="843F51425B49001E62C502D61981FBFE" box="[495,906,952,1056]" gridcol="1" gridrow="3" pageId="10" pageNumber="493">16.1 13.4 ±2.8 (9.7–17.3); 9 28.2 25.9 ±5.1 (18.2–33.8); 10 13.3 11.0 ±2.03 (6.7–14.4); 10</td>
<td id="843F51425B49001E60E102D61E19FBFE" box="[971,1042,952,1056]" gridcol="2" gridrow="3" pageId="10" pageNumber="493">8.6 19.6 8.7</td>
<td id="843F51425B49001E671702D61F8BFBFE" box="[1085,1408,952,1056]" gridcol="3" gridrow="3" pageId="10" pageNumber="493">14.3 ±3.36 (9.1–21.3); 10 23.3 ±2.80 (19.5–28.1); 11 11.1 ±1.78 (7.8–14.0); 11</td>
</tr>
<tr id="C7EE383E5B49001E63B1055A1F8BFB42" box="[155,1408,1076,1180]" gridrow="4" pageId="10" pageNumber="493">
<th id="843F51425B49001E63B1055A1BB5FB42" box="[155,446,1076,1180]" gridcol="0" gridrow="4" pageId="10" pageNumber="493">Primary clava / cirrus A Secondary clavae Cirri E</th>
<td id="843F51425B49001E62C5055A1981FB42" box="[495,906,1076,1180]" gridcol="1" gridrow="4" pageId="10" pageNumber="493">0.47 0.43 ±0.05 (0.35–0.51); 10 6.8 6.7 ±1.06 (5.2–8.4); 8 18.7 15.8 ±2.9 (11.1–18.7); 6</td>
<td id="843F51425B49001E60E1055A1E19FB42" box="[971,1042,1076,1180]" gridcol="2" gridrow="4" pageId="10" pageNumber="493">0.44 5.8 9.8</td>
<td id="843F51425B49001E6717055A1F8BFB42" box="[1085,1408,1076,1180]" gridcol="3" gridrow="4" pageId="10" pageNumber="493">0.47 ±0.04 (0.38–0.54); 11 5.5 ±1.20 (3.6–7.7); 11 13.4 ±4.93 (7.7–24.2); 9</td>
</tr>
<tr id="C7EE383E5B49001E63B105DE1F8BFAC6" box="[155,1408,1200,1304]" gridrow="5" pageId="10" pageNumber="493">
<th id="843F51425B49001E63B105DE1BB5FAC6" box="[155,446,1200,1304]" gridcol="0" gridrow="5" pageId="10" pageNumber="493">Leg I spine Leg II spine Leg III spine</th>
<td id="843F51425B49001E62C505DE1981FAC6" box="[495,906,1200,1304]" gridcol="1" gridrow="5" pageId="10" pageNumber="493">9.4 6.7 ±2.83 (3.4–10.8); 8 11.3 8.6 ±2.37 (4.1–11.3); 6 11.2 10.6 ±0.49 (10.0–11.2); 4</td>
<td id="843F51425B49001E60E105DE1E19FAC6" box="[971,1042,1200,1304]" gridcol="2" gridrow="5" pageId="10" pageNumber="493">???</td>
<td id="843F51425B49001E671705DE1F8BFAC6" box="[1085,1408,1200,1304]" gridcol="3" gridrow="5" pageId="10" pageNumber="493">7.5 ±3.21 (4.6–12); 4 6.0 ±1.66 (4.5–8.8); 5 7.0 ±0.86 (6.2–7.9); 4</td>
</tr>
<tr id="C7EE383E5B49001E63B104421F8BFA4A" box="[155,1408,1324,1428]" gridrow="6" pageId="10" pageNumber="493">
<th id="843F51425B49001E63B104421BB5FA4A" box="[155,446,1324,1428]" gridcol="0" gridrow="6" pageId="10" pageNumber="493">Leg IV sense organ Leg IV base Leg IV spine</th>
<td id="843F51425B49001E62C504421981FA4A" box="[495,906,1324,1428]" gridcol="1" gridrow="6" pageId="10" pageNumber="493">12.4 10.1 ±1.72 (7.1–12.7); 6 5.5 4.4 ±0.66 (3.6–5.5); 8 4.6 4.8 ±0.96 (3.6–6.7); 8</td>
<td id="843F51425B49001E60E104421E19FA4A" box="[971,1042,1324,1428]" gridcol="2" gridrow="6" pageId="10" pageNumber="493">8.6 3.4 5.2</td>
<td id="843F51425B49001E671704421F8BFA4A" box="[1085,1408,1324,1428]" gridcol="3" gridrow="6" pageId="10" pageNumber="493">10.3 ±2 (6.9–14.1); 11 4.2 ±0.97 (2.8–6); 8 5.0 ±1.01 (3.7–6.8); 8</td>
</tr>
<tr id="C7EE383E5B49001E63B104C61F8BF9CE" box="[155,1408,1448,1552]" gridrow="7" pageId="10" pageNumber="493">
<th id="843F51425B49001E63B104C61BB5F9CE" box="[155,446,1448,1552]" gridcol="0" gridrow="7" pageId="10" pageNumber="493">Caudal apparatus Legs I–III Toe 1 Toe 2</th>
<td id="843F51425B49001E62C504C61981F9CE" box="[495,906,1448,1552]" gridcol="1" gridrow="7" pageId="10" pageNumber="493">8.9 12.0 ±4.32 (6.2–19.3); 7 13.4 9.6 ±2.40 (6.3–13.4); 5 4.7 5.5 ±0.84 (4.5–6.7); 5</td>
<td id="843F51425B49001E60E104C61E19F9CE" box="[971,1042,1448,1552]" gridcol="2" gridrow="7" pageId="10" pageNumber="493">13.1 7.3 4.0</td>
<td id="843F51425B49001E671704C61F8BF9CE" box="[1085,1408,1448,1552]" gridcol="3" gridrow="7" pageId="10" pageNumber="493">11.9 ±5.59 (1.7–22.0); 12 8.3 ±1.61 (5.7–11.3); 9 4.3 ±1.10 (3.1–6.3); 9</td>
</tr>
<tr id="C7EE383E5B49001E63B1074A1F8BF952" box="[155,1408,1572,1676]" gridrow="8" pageId="10" pageNumber="493">
<th id="843F51425B49001E63B1074A1BB5F952" box="[155,446,1572,1676]" gridcol="0" gridrow="8" pageId="10" pageNumber="493">Toe 3 Toe 4 Toe 5</th>
<td id="843F51425B49001E62C5074A1981F952" box="[495,906,1572,1676]" gridcol="1" gridrow="8" pageId="10" pageNumber="493">16.2 12.6 ±3.20 (7.2–16.2); 5 10.2 9.7 ±1.23 (8.2–11.3); 5 17.9 15.9 ±2.89 (10.3–17.9); 5</td>
<td id="843F51425B49001E60E1074A1E19F952" box="[971,1042,1572,1676]" gridcol="2" gridrow="8" pageId="10" pageNumber="493">11.6 6.1 15.0</td>
<td id="843F51425B49001E6717074A1F8BF952" box="[1085,1408,1572,1676]" gridcol="3" gridrow="8" pageId="10" pageNumber="493">12.3 ±2.55 (8.1–16.9); 9 8.9 ±1.02 (7.2–10.6); 9 16.3 ±2.15 (14.3–20.0); 9</td>
</tr>
<tr id="C7EE383E5B49001E63B107CE1F8BF8D6" box="[155,1408,1696,1800]" gridrow="9" pageId="10" pageNumber="493">
<th id="843F51425B49001E63B107CE1BB5F8D6" box="[155,446,1696,1800]" gridcol="0" gridrow="9" pageId="10" pageNumber="493">Toe 6 Leg IV Toe 1 Toe 2</th>
<td id="843F51425B49001E62C507CE1981F8D6" box="[495,906,1696,1800]" gridcol="1" gridrow="9" pageId="10" pageNumber="493">12.7 11.3 ±1.53 (8.8–12.7); 5 14.4 13.0 ±1.96 (10.0–15.6); 6? 16.6 ±3.61 (12.5–22.5); 6</td>
<td id="843F51425B49001E60E107CE1E19F8D6" box="[971,1042,1696,1800]" gridcol="2" gridrow="9" pageId="10" pageNumber="493">9.0 11.6 16.0</td>
<td id="843F51425B49001E671707CE1F8BF8D6" box="[1085,1408,1696,1800]" gridcol="3" gridrow="9" pageId="10" pageNumber="493">9.6 ±1.59 (7.5–12.9); 9 11.7 ±1.48 (10.1–14.5); 8 15.8 ±2.38 (13.2–19.5); 8</td>
</tr>
<tr id="C7EE383E5B49001E63B106721F8BF85A" box="[155,1408,1820,1924]" gridrow="10" pageId="10" pageNumber="493">
<th id="843F51425B49001E63B106721BB5F85A" box="[155,446,1820,1924]" gridcol="0" gridrow="10" pageId="10" pageNumber="493">Toe 3 Toe 4 Toe 5</th>
<td id="843F51425B49001E62C506721981F85A" box="[495,906,1820,1924]" gridcol="1" gridrow="10" pageId="10" pageNumber="493">10.1 9.2 ±1.60 (6.8–12.3); 7 12.7 12.3 ±1.83 (10.0–16.0); 7? 17.6 ±2.40 (14.5–20.80; 6</td>
<td id="843F51425B49001E60E106721E19F85A" box="[971,1042,1820,1924]" gridcol="2" gridrow="10" pageId="10" pageNumber="493">7.2 10.2 16.4</td>
<td id="843F51425B49001E671706721F8BF85A" box="[1085,1408,1820,1924]" gridcol="3" gridrow="10" pageId="10" pageNumber="493">7.9 ±1.25 (6.2–9.8); 8 10.8 ±1.48 (9.0–13.6); 8 18.9 ±2.17 (16.2–21.8); 8</td>
</tr>
<tr id="C7EE383E5B49001E63B106F61F8BF809" box="[155,1408,1944,2007]" gridrow="11" pageId="10" pageNumber="493">
<th id="843F51425B49001E63B106F61BB5F809" box="[155,446,1944,2007]" gridcol="0" gridrow="11" pageId="10" pageNumber="493">Toe 6 Gonopore - Anus</th>
<td id="843F51425B49001E62C506F61981F809" box="[495,906,1944,2007]" gridcol="1" gridrow="11" pageId="10" pageNumber="493">? 11.8 ±1.97 (9.1–15.1); 6 7.3 7.6 ±1.34 (5.8–9.6); 6</td>
<td id="843F51425B49001E60E106F61E19F809" box="[971,1042,1944,2007]" gridcol="2" gridrow="11" pageId="10" pageNumber="493">11.8 6.5</td>
<td id="843F51425B49001E671706F61F8BF809" box="[1085,1408,1944,2007]" gridcol="3" gridrow="11" pageId="10" pageNumber="493">11.4 ±1.67 (9.6–14.6); 7 -</td>
</tr>
</table>
</paragraph>
<paragraph id="79613A7C5B48FFEA63ED01F91B93FF26" blockId="11.[151,1437,151,357]" pageId="11" pageNumber="494">
Ovoid pharyngeal bulb 19.5 µm long and 19.0 µm wide. Placoids not visible after slide mounting. Mouth opening located at the top of an ovoid buccal cone (12.1 µm long and 18.2 µm wide) about 10 µm distant from the frontal edge (
<figureCitation id="E1E526F95B48FFEA620701B11B64FF26" box="[301,367,223,248]" captionStart="FIGURE 4" captionStartId="9.[151,250,1733,1755]" captionTargetBox="[176,1418,183,1711]" captionTargetId="figure@9.[174,1418,181,1711]" captionTargetPageId="9" captionText="FIGURE 4. Batillipes dandarae sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view); C. Detailed drawing of the female gonopore (go) and anus (an); D. Dorsal view of the head showing the median cirrus (cm) and the eye (ey); E. Ventral view of the head showing the buccal cone (bc) and the external cephalic cirrus (ce). Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400157/files/figure.png" pageId="11" pageNumber="494">Fig. 4</figureCitation>
E).
</paragraph>
<paragraph id="79613A7C5B48FFEA63ED006A185EFEBB" blockId="11.[151,1437,151,357]" pageId="11" pageNumber="494">
A dorsal blunt enlargement is visible in the scapular region at the level of legs I
<figureCitation id="E1E526F95B48FFEA6775006A1EADFEC2" box="[1119,1190,260,285]" captionStart="FIGURE 4" captionStartId="9.[151,250,1733,1755]" captionTargetBox="[176,1418,183,1711]" captionTargetId="figure@9.[174,1418,181,1711]" captionTargetPageId="9" captionText="FIGURE 4. Batillipes dandarae sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view); C. Detailed drawing of the female gonopore (go) and anus (an); D. Dorsal view of the head showing the median cirrus (cm) and the eye (ey); E. Ventral view of the head showing the buccal cone (bc) and the external cephalic cirrus (ce). Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400157/files/figure.png" pageId="11" pageNumber="494">Figs 4</figureCitation>
A, B, 5 A). No other body lateral projections or cuticular lateral processes are present. The caudal apparatus is constituted by a cuticular conical-shaped projection, 8.9 µm long.
</paragraph>
<caption id="2DA16AF45B48FFEA63BD061A1F9CF838" httpUri="https://zenodo.org/record/400158/files/figure.png" pageId="11" pageNumber="494" targetBox="[175,1418,414,1883]" targetPageId="11">
<paragraph id="79613A7C5B48FFEA63BD061A1F9CF838" blockId="11.[151,1436,1908,2022]" pageId="11" pageNumber="494">
<emphasis id="4BAAE66E5B48FFEA63BD061A1B05F857" bold="true" box="[151,270,1908,1930]" pageId="11" pageNumber="494">FIGURE 5</emphasis>
.
<taxonomicName id="BEDE41FF5B48FFEA6231061A1BEFF854" box="[283,484,1908,1930]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="494" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B48FFEA6231061A1BEFF854" box="[283,484,1908,1930]" italics="true" pageId="11" pageNumber="494">Batillipes dandarae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50995B155B48FFEA62C6061A1836F857" box="[492,573,1908,1929]" pageId="11" pageNumber="494" rank="species">
<emphasis id="4BAAE66E5B48FFEA62C6061A1836F857" bold="true" box="[492,573,1908,1929]" pageId="11" pageNumber="494">sp. nov.</emphasis>
</taxonomicNameLabel>
A. Anterior portion of the body of the holotype showing cephalic appendages (ciinternal cirrus; cl—lateral cirrus); clavae (pc—primary clava; sc—secondary clava), first body projection (pr1) and blunt enlargement in the scapular region (sr); B. Leg I of a paratype showing the leg spine (pI) inserted on the posterior part of the leg and turning frontwards, mo—mouth; C. Detail of the female gonopore (go) and anus (an) of the holotype. Scale bars = 10 µm.
</paragraph>
</caption>
<paragraph id="79613A7C5B4FFFED63ED01F91876FE73" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="495">
Sensorial spines present on all legs. Sensorial spine on leg I is the shortest, about the same length on legs II and III, and longer on legs IV (9.4; 11.3; 11.2, and 12.7 µm long, respectively). Leg I sense organs are not subdivided into different parts. They are similar in shape to sense organs on legs II and III that are spine-like. However, leg I sense organs are inserted on the posterior part of the leg and revolved frontwards (
<figureCitation id="E1E526F95B4FFFED6702006A1E61FEC3" box="[1064,1130,260,285]" captionStart="FIGURE 5" captionStartId="11.[151,250,1908,1930]" captionTargetBox="[175,1418,414,1883]" captionTargetId="figure@11.[172,1418,411,1883]" captionTargetPageId="11" captionText="FIGURE 5. Batillipes dandarae sp. nov. A. Anterior portion of the body of the holotype showing cephalic appendages (ciinternal cirrus; cl—lateral cirrus); clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and blunt enlargement in the scapular region (sr); B. Leg I of a paratype showing the leg spine (pI) inserted on the posterior part of the leg and turning frontwards, mo—mouth; C. Detail of the female gonopore (go) and anus (an) of the holotype. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400158/files/figure.png" pageId="12" pageNumber="495">Fig. 5</figureCitation>
B). All the sense organs on legs I–III have an optically dense dot near their blunt tips. The sense organ on leg IV is divided into a cirrophore (1.3 µm long), a proximal portion (4.7 µm long) and a distal portion (6.4 µm long) with a basal van der Land’s organ and an optically dense dot near the tip. Dorsal cirri E are blunt and short (18.7 µm long) with no evident cirrophores and also with a dot near the tip.
</paragraph>
<paragraph id="79613A7C5B4FFFED63ED00D919FDFD13" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="495">
Telescopic legs without tibial papillae. Toes with distal stalks considerably enlarged (2.6 µm wide), terminated by small ovoid suction discs (3.7 µm long and 3.4 µm wide on leg IV) with conspicuous braces and well developed peripheral thickenings. In the first three pair of legs, toe 2 is the shortest, toes 3 and 5 are the longest and toes 1, 4 and 6 are medium sized (13.4, 4.7, 16.2, 10.2, 17.9 and 12.7 µm long, respectively for toes 1 to 6 of leg II). In the fourth pair of legs (
<figureCitation id="E1E526F95B4FFFED625A03291BBCFDBE" box="[368,439,583,608]" captionStart="FIGURE 4" captionStartId="9.[151,250,1733,1755]" captionTargetBox="[176,1418,183,1711]" captionTargetId="figure@9.[174,1418,181,1711]" captionTargetPageId="9" captionText="FIGURE 4. Batillipes dandarae sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view); C. Detailed drawing of the female gonopore (go) and anus (an); D. Dorsal view of the head showing the median cirrus (cm) and the eye (ey); E. Ventral view of the head showing the buccal cone (bc) and the external cephalic cirrus (ce). Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400157/files/figure.png" pageId="12" pageNumber="495">Figs 4</figureCitation>
A, B, 5 C) toes conform to the pattern of the D group of species proposed by
<bibRefCitation id="1D4F478D5B4FFFED660E03291B81FD5A" author="Kristensen" pageId="12" pageNumber="495" refString="Kristensen, R. M. &amp; Mackness, B. S. (2000) First record of the marine Tardigrade genus Batillipes (Arthrotardigrada: Batillipedidae) from South Australia with a description of a new species. Records of the South Australian Museum, 33 (2), 73 - 87." type="journal article" year="2000">Kristensen and Mackness (2000)</bibRefCitation>
, with toes 3 and 4 of different lengths (10.1 and 12.7 µm long respectively). In legs IV toes 1 and 6 are of intermediate size and similar in length; toes 2 and especially toes 5 are the longest (this character was observed in
<typeStatus id="A66584DE5B4FFFED620A03DB1B84FD13" box="[288,399,693,717]" pageId="12" pageNumber="495" type="paratype">paratypes</typeStatus>
; in the
<typeStatus id="A66584DE5B4FFFED62CA03DA1849FD13" box="[480,578,692,717]" pageId="12" pageNumber="495" type="holotype">holotype</typeStatus>
only toe 1, 14.4 µm, was measurable).
</paragraph>
<paragraph id="79613A7C5B4FFFED63ED03B918D3FCE6" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="495">
Dorsal cuticle uniformly and finely punctated, with about 15 pillars/10 µm, comprised of short pillars (
<emphasis id="4BAAE66E5B4FFFED664E03B71F8EFD2E" box="[1380,1413,729,752]" italics="true" pageId="12" pageNumber="495">ca.</emphasis>
1 µm high) with some delicate transverse lines (about eight) and without smooth regions. Less dense punctation on ventral cuticle which exhibit some transverse folds.
</paragraph>
<paragraph id="79613A7C5B4FFFED63ED022A1F62FC5E" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="495">
Gonopore rosette-shaped (
<figureCitation id="E1E526F95B4FFFED62DE022A1834FC82" box="[500,575,836,861]" captionStart="FIGURE 4" captionStartId="9.[151,250,1733,1755]" captionTargetBox="[176,1418,183,1711]" captionTargetId="figure@9.[174,1418,181,1711]" captionTargetPageId="9" captionText="FIGURE 4. Batillipes dandarae sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view); C. Detailed drawing of the female gonopore (go) and anus (an); D. Dorsal view of the head showing the median cirrus (cm) and the eye (ey); E. Ventral view of the head showing the buccal cone (bc) and the external cephalic cirrus (ce). Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400157/files/figure.png" pageId="12" pageNumber="495">Figs 4</figureCitation>
C, 5C) separated from the anus by a very small groove. The anus, 7.3 µm posterior from the gonopore, is surrounded by a peculiar cuticular round structure, constituted by six platelets.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B4FFFED63ED02E21F38FB86" pageId="12" pageNumber="495" type="discussion">
<paragraph id="79613A7C5B4FFFED63ED02E21F38FB86" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="495">
<emphasis id="4BAAE66E5B4FFFED63ED02E21B30FC7B" bold="true" box="[199,315,908,933]" pageId="12" pageNumber="495">Remarks.</emphasis>
Sexual dimorphism was not evident in the only male (138 µm long) found. This male is similar to females in both qualitative and metric characters (except for their circular gonopore with crescent shaped cuticular fold, distant 6.5 µm from the anus). Juveniles, with six toes on each leg but without a visible gonopore, were also similar to adults. Four-toed larvae were not found. Sense organ on legs I inserted on the posterior part of the leg and turning frontwards, similar to the one described for
<taxonomicName id="BEDE41FF5B4FFFED61CC05731E5DFBEA" authority="Kristensen, 1978" authorityName="Kristensen" authorityYear="1978" box="[742,1110,1052,1077]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="noerrevangi">
<emphasis id="4BAAE66E5B4FFFED61CC05731984FBEA" box="[742,911,1053,1076]" italics="true" pageId="12" pageNumber="495">B. noerrevangi</emphasis>
<bibRefCitation id="1D4F478D5B4FFFED60BD05721E5DFBEA" author="Kristensen" box="[919,1110,1052,1077]" pageId="12" pageNumber="495" refString="Kristensen, R. M. (1978) Notes on Marine Heterotardigrades. 1. Description of two Batillipes species, using the electron microscope. Zoologischer Anzeiger, 200, 1 - 17." type="journal article" year="1978">Kristensen, 1978</bibRefCitation>
</taxonomicName>
is consistently present in all the examined specimens. Shape and size of the caudal apparatus are strongly variable among individuals.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B4FFFED63ED050A18D8FAD2" pageId="12" pageNumber="495" type="diagnosis">
<paragraph id="79613A7C5B4FFFED63ED050A18D8FAD2" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="495">
<emphasis id="4BAAE66E5B4FFFED63ED050A1BDAFBA3" bold="true" box="[199,465,1124,1149]" pageId="12" pageNumber="495">Differential diagnosis.</emphasis>
<taxonomicName id="BEDE41FF5B4FFFED62F6050A18B6FBA2" box="[476,701,1124,1148]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4FFFED62F6050A18B6FBA2" box="[476,701,1124,1148]" italics="true" pageId="12" pageNumber="495">Batillipes dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4FFFED61E3050A192DFBA3" bold="true" box="[713,806,1124,1149]" pageId="12" pageNumber="495">
<taxonomicNameLabel id="50995B155B4FFFED61E3050A192DFBA3" box="[713,806,1124,1149]" pageId="12" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having medial toes 3 and 4 on leg IV of different lengths and respectively different from toe 1 and toe 2, belongs to the D group of species (
<bibRefCitation id="1D4F478D5B4FFFED675705E91ADFFB1A" author="Kristensen" pageId="12" pageNumber="495" refString="Kristensen, R. M. &amp; Mackness, B. S. (2000) First record of the marine Tardigrade genus Batillipes (Arthrotardigrada: Batillipedidae) from South Australia with a description of a new species. Records of the South Australian Museum, 33 (2), 73 - 87." type="journal article" year="2000">Kristensen and Mackness 2000</bibRefCitation>
). Only six known species, including
<taxonomicName id="BEDE41FF5B4FFFED615305C31911FB1A" box="[633,794,1196,1220]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B4FFFED615305C31911FB1A" box="[633,794,1196,1220]" italics="true" pageId="12" pageNumber="495">B. brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4FFFED600905C21975FB1B" bold="true" box="[803,894,1196,1221]" pageId="12" pageNumber="495">
<taxonomicNameLabel id="50995B155B4FFFED600905C21975FB1B" box="[803,894,1196,1221]" pageId="12" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
described above, exhibit this toe pattern on leg IV. However, similarly to
<taxonomicName id="BEDE41FF5B4FFFED629705BF1843FB36" box="[445,584,1232,1256]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4FFFED629705BF1843FB36" box="[445,584,1232,1256]" italics="true" pageId="12" pageNumber="495">B. dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4FFFED617A05A118A0FB36" bold="true" box="[592,683,1231,1256]" pageId="12" pageNumber="495">
<taxonomicNameLabel id="50995B155B4FFFED617A05A118A0FB36" box="[592,683,1231,1256]" pageId="12" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, only two other species included in this D group,
<taxonomicName id="BEDE41FF5B4FFFED67C805BF1F60FB36" box="[1250,1387,1232,1256]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B4FFFED67C805BF1F60FB36" box="[1250,1387,1232,1256]" italics="true" pageId="12" pageNumber="495">B. africanus</emphasis>
</taxonomicName>
and
<taxonomicName id="BEDE41FF5B4FFFED63BD059B1B23FAD2" box="[151,296,1268,1292]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="tubernatis">
<emphasis id="4BAAE66E5B4FFFED63BD059B1B23FAD2" box="[151,296,1268,1292]" italics="true" pageId="12" pageNumber="495">B. tubernatis</emphasis>
</taxonomicName>
, lack body cuticular lateral processes.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B4FFFEC63ED04771B55FE73" lastPageId="13" lastPageNumber="496" pageId="12" pageNumber="495" type="reference_group">
<paragraph id="79613A7C5B4FFFED63ED04771EACF9AA" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="495">
<taxonomicName id="BEDE41FF5B4FFFED63ED04771B53FAEE" ID-CoL="KYTC" box="[199,344,1304,1328]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="tubernatis">
<emphasis id="4BAAE66E5B4FFFED63ED04771B53FAEE" box="[199,344,1304,1328]" italics="true" pageId="12" pageNumber="495">B. tubernatis</emphasis>
</taxonomicName>
was originally described by
<bibRefCitation id="1D4F478D5B4FFFED61B40479194EFAEE" author="Pollock" box="[670,837,1303,1328]" pageId="12" pageNumber="495" refString="Pollock, L. W. (1971) On some British marine Tardigrada, including two new species of Batillipes. Journal of the. Marine Biological Association of the United Kingdom, 51, 93 - 103." type="journal article" year="1971">Pollock (1971)</bibRefCitation>
as having a rounded and swollen caudal contour and not a real caudal appendage. In the emended description (
<bibRefCitation id="1D4F478D5B4FFFED601B045219D5FA8B" author="McKirdy" box="[817,990,1340,1365]" pageId="12" pageNumber="495" refString="McKirdy, D. J. (1975) Batillipes (Heterotardigrada): Comparison of six species from Florida (USA) and a discussion of taxonomic characters within the genus. Memorie dell'Istituto Italiano di Idrobiologia, 32 (Supplement), 177223." type="journal article" year="1975">McKirdy 1975</bibRefCitation>
), some specimens of
<taxonomicName id="BEDE41FF5B4FFFED67F204531F67FA8A" ID-CoL="KYTC" box="[1240,1388,1340,1364]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="tubernatis">
<emphasis id="4BAAE66E5B4FFFED67F204531F67FA8A" box="[1240,1388,1340,1364]" italics="true" pageId="12" pageNumber="495">B. tubernatis</emphasis>
</taxonomicName>
can exhibit a “strong, single, swollen-based caudal spike inserting as a continuous extension of the body”, similar to the caudal apparatus of the new species. However, in
<taxonomicName id="BEDE41FF5B4FFFED61E404EA19A7FA42" ID-CoL="9c4449fe-cb06-427e-8889-377b6ef16d40" box="[718,940,1412,1436]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4FFFED61E404EA19A7FA42" box="[718,940,1412,1436]" italics="true" pageId="12" pageNumber="495">Batillipes dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4FFFED609E04EA1E04FA43" bold="true" box="[948,1039,1412,1437]" pageId="12" pageNumber="495">
<taxonomicNameLabel id="50995B155B4FFFED609E04EA1E04FA43" box="[948,1039,1412,1437]" pageId="12" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the caudal appendage is present in all the specimens, including young. Moreover, the new species differs from
<taxonomicName id="BEDE41FF5B4FFFED672B04C71E9EFA1E" ID-CoL="KYTC" box="[1025,1173,1448,1472]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="tubernatis">
<emphasis id="4BAAE66E5B4FFFED672B04C71E9EFA1E" box="[1025,1173,1448,1472]" italics="true" pageId="12" pageNumber="495">B. tubernatis</emphasis>
</taxonomicName>
in the aspect of dorsal punctations: they are uniformly distributed and finely punctate in the new species, while apunctate areas and different sized punctations occur in
<taxonomicName id="BEDE41FF5B4FFFED6115049F18DFF9D6" ID-CoL="KYTC" box="[575,724,1520,1544]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="tubernatis">
<emphasis id="4BAAE66E5B4FFFED6115049F18DFF9D6" box="[575,724,1520,1544]" italics="true" pageId="12" pageNumber="495">B. tubernatis</emphasis>
</taxonomicName>
. In
<taxonomicName id="BEDE41FF5B4FFFED6023049F1993F9D6" ID-CoL="9c4449fe-cb06-427e-8889-377b6ef16d40" box="[777,920,1520,1544]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4FFFED6023049F1993F9D6" box="[777,920,1520,1544]" italics="true" pageId="12" pageNumber="495">B. dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4FFFED608E04811E09F9D6" bold="true" box="[932,1026,1519,1544]" pageId="12" pageNumber="495">
<taxonomicNameLabel id="50995B155B4FFFED608E04811E09F9D6" box="[932,1026,1519,1544]" pageId="12" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and contrary to
<taxonomicName id="BEDE41FF5B4FFFED67F8049F1F6CF9D6" ID-CoL="KYTC" box="[1234,1383,1520,1544]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="tubernatis">
<emphasis id="4BAAE66E5B4FFFED67F8049F1F6CF9D6" box="[1234,1383,1520,1544]" italics="true" pageId="12" pageNumber="495">B. tubernatis</emphasis>
</taxonomicName>
, the secondary clavae are well developed, protruding from the frontal head border. In addition, according to
<bibRefCitation id="1D4F478D5B4FFFED661F077A1AE8F98E" author="McKirdy" pageId="12" pageNumber="495" refString="McKirdy, D. J. (1975) Batillipes (Heterotardigrada): Comparison of six species from Florida (USA) and a discussion of taxonomic characters within the genus. Memorie dell'Istituto Italiano di Idrobiologia, 32 (Supplement), 177223." type="journal article" year="1975">McKirdy (1975)</bibRefCitation>
, in
<taxonomicName id="BEDE41FF5B4FFFED622107571B97F98E" ID-CoL="KYTC" box="[267,412,1592,1616]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="tubernatis">
<emphasis id="4BAAE66E5B4FFFED622107571B97F98E" box="[267,412,1592,1616]" italics="true" pageId="12" pageNumber="495">B. tubernatis</emphasis>
</taxonomicName>
toe suction discs are rounded-quadrate to subcircular in shape with slightly indented frontal edge (see Fig. 12 B in
<bibRefCitation id="1D4F478D5B4FFFED62BE07321837F9AB" author="McKirdy" box="[404,572,1628,1653]" pageId="12" pageNumber="495" refString="McKirdy, D. J. (1975) Batillipes (Heterotardigrada): Comparison of six species from Florida (USA) and a discussion of taxonomic characters within the genus. Memorie dell'Istituto Italiano di Idrobiologia, 32 (Supplement), 177223." type="journal article" year="1975">McKirdy 1975</bibRefCitation>
), while they are ovoid in the new species (
<figureCitation id="E1E526F95B4FFFED673307321E50F9AB" box="[1049,1115,1628,1653]" captionStart="FIGURE 5" captionStartId="11.[151,250,1908,1930]" captionTargetBox="[175,1418,414,1883]" captionTargetId="figure@11.[172,1418,411,1883]" captionTargetPageId="11" captionText="FIGURE 5. Batillipes dandarae sp. nov. A. Anterior portion of the body of the holotype showing cephalic appendages (ciinternal cirrus; cl—lateral cirrus); clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and blunt enlargement in the scapular region (sr); B. Leg I of a paratype showing the leg spine (pI) inserted on the posterior part of the leg and turning frontwards, mo—mouth; C. Detail of the female gonopore (go) and anus (an) of the holotype. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400158/files/figure.png" pageId="12" pageNumber="495">Fig. 5</figureCitation>
A, C).
</paragraph>
<paragraph id="79613A7C5B4FFFEC63ED07EE1B55FE73" blockId="12.[151,1437,151,2013]" lastBlockId="13.[151,1436,151,861]" lastPageId="13" lastPageNumber="496" pageId="12" pageNumber="495">
<taxonomicName id="BEDE41FF5B4FFFED63ED07EE1BAFF946" ID-CoL="9c4449fe-cb06-427e-8889-377b6ef16d40" box="[199,420,1664,1688]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4FFFED63ED07EE1BAFF946" box="[199,420,1664,1688]" italics="true" pageId="12" pageNumber="495">Batillipes dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4FFFED62810711180FF946" bold="true" box="[427,516,1663,1688]" pageId="12" pageNumber="495">
<taxonomicNameLabel id="50995B155B4FFFED62810711180FF946" box="[427,516,1663,1688]" pageId="12" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="BEDE41FF5B4FFFED611307EF18C9F946" ID-CoL="KYSL" box="[569,706,1664,1688]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B4FFFED611307EF18C9F946" box="[569,706,1664,1688]" italics="true" pageId="12" pageNumber="495">B. africanus</emphasis>
</taxonomicName>
are very similar each other. In
<taxonomicName id="BEDE41FF5B4FFFED673007EF1EAFF946" ID-CoL="9c4449fe-cb06-427e-8889-377b6ef16d40" box="[1050,1188,1664,1688]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4FFFED673007EF1EAFF946" box="[1050,1188,1664,1688]" italics="true" pageId="12" pageNumber="495">B. dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4FFFED678107111F0FF946" bold="true" box="[1195,1284,1663,1688]" pageId="12" pageNumber="495">
<taxonomicNameLabel id="50995B155B4FFFED678107111F0FF946" box="[1195,1284,1663,1688]" pageId="12" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, compared to
<taxonomicName id="BEDE41FF5B4FFFED63BD07CB1B2BF962" ID-CoL="KYSL" box="[151,288,1700,1724]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B4FFFED63BD07CB1B2BF962" box="[151,288,1700,1724]" italics="true" pageId="12" pageNumber="495">B. africanus</emphasis>
</taxonomicName>
, pedestals bearing primary clavae and lateral cirri are less pronounced. Despite the variability in shape and size, the caudal apparatus is more consistently triangular shaped in
<taxonomicName id="BEDE41FF5B4FFFED60F007A71E63F93E" ID-CoL="9c4449fe-cb06-427e-8889-377b6ef16d40" box="[986,1128,1736,1760]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4FFFED60F007A71E63F93E" box="[986,1128,1736,1760]" italics="true" pageId="12" pageNumber="495">B. dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4FFFED675B07A91EC5F93E" bold="true" box="[1137,1230,1735,1760]" pageId="12" pageNumber="495">
<taxonomicNameLabel id="50995B155B4FFFED675B07A91EC5F93E" box="[1137,1230,1735,1760]" pageId="12" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and never in the shape of an elongated spine as sometimes occurs in
<taxonomicName id="BEDE41FF5B4FFFED61CA07831962F8DA" ID-CoL="KYSL" box="[736,873,1772,1796]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B4FFFED61CA07831962F8DA" box="[736,873,1772,1796]" italics="true" pageId="12" pageNumber="495">B. africanus</emphasis>
</taxonomicName>
. On the other hand, specimens of the new species without caudal apparatus were not found. Segmental folds are weakly marked in the new species, preventing the differentiation of body sections, contrary to
<taxonomicName id="BEDE41FF5B4FFFED61B9065B1915F892" ID-CoL="KYSL" box="[659,798,1844,1868]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B4FFFED61B9065B1915F892" box="[659,798,1844,1868]" italics="true" pageId="12" pageNumber="495">B. africanus</emphasis>
</taxonomicName>
where a cephalic, a caudal and three body sections are evident, including in first stage larvae. Another difference between the two species refers to tibial papillae that are present in
<taxonomicName id="BEDE41FF5B4FFFED623A06131B90F84A" ID-CoL="KYSL" box="[272,411,1916,1940]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B4FFFED623A06131B90F84A" box="[272,411,1916,1940]" italics="true" pageId="12" pageNumber="495">B. africanus</emphasis>
</taxonomicName>
only. In addition, the following characters, by their taxonomic relevance, deserve special attention.
<emphasis id="4BAAE66E5B4FFFED622C06CF1B05F866" box="[262,270,1953,1976]" italics="true" pageId="12" pageNumber="495">i</emphasis>
) Shape of the scapular region: in
<taxonomicName id="BEDE41FF5B4FFFED61AE06CF1904F866" ID-CoL="9c4449fe-cb06-427e-8889-377b6ef16d40" box="[644,783,1952,1976]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4FFFED61AE06CF1904F866" box="[644,783,1952,1976]" italics="true" pageId="12" pageNumber="495">B. dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4FFFED603E06CE1966F867" bold="true" box="[788,877,1952,1977]" pageId="12" pageNumber="495">
<taxonomicNameLabel id="50995B155B4FFFED603E06CE1966F867" box="[788,877,1952,1977]" pageId="12" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
a conspicuous dorsal blunt enlargement is present in the scapular region at the level of legs I, while
<taxonomicName id="BEDE41FF5B4FFFED61DF06AB1988F802" ID-CoL="KYSL" box="[757,899,1988,2012]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="495" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B4FFFED61DF06AB1988F802" box="[757,899,1988,2012]" italics="true" pageId="12" pageNumber="495">B. africanus</emphasis>
</taxonomicName>
lacks this enlargement and the body is more rectangular-shaped.
<emphasis id="4BAAE66E5B4EFFEC625001F61B82FF71" box="[378,393,152,175]" italics="true" pageId="13" pageNumber="496">ii</emphasis>
) Morphology of leg I sense organs, that are spine-like, inserted in the posterior region of the legs and revolved frontwards in the new species and not articulated, straight and parallel to the toes, as in
<taxonomicName id="BEDE41FF5B4EFFEC66AE01D31B0AFF26" ID-CoL="KYSL" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B4EFFEC66AE01D31B0AFF26" italics="true" pageId="13" pageNumber="496">B. africanus</emphasis>
</taxonomicName>
. In addition, in the new species sense organs on legs III are longer, about twice the length, than those of
<taxonomicName id="BEDE41FF5B4EFFEC63BD006B1B14FEC2" ID-CoL="KYSL" box="[151,287,260,284]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B4EFFEC63BD006B1B14FEC2" box="[151,287,260,284]" italics="true" pageId="13" pageNumber="496">B. africanus</emphasis>
</taxonomicName>
(range 10.0–11.2 µm in the new species; 5.0–6.0 µm in
<taxonomicName id="BEDE41FF5B4EFFEC60B9006B1E10FEC2" ID-CoL="KYSL" box="[915,1051,260,284]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B4EFFEC60B9006B1E10FEC2" box="[915,1051,260,284]" italics="true" pageId="13" pageNumber="496">B. africanus</emphasis>
</taxonomicName>
).
<emphasis id="4BAAE66E5B4EFFEC6718006B1E42FEC2" box="[1074,1097,261,284]" italics="true" pageId="13" pageNumber="496">iii</emphasis>
) Structure and morphology of the gonopore and the anus: in the new species the female gonopore has the typical rosette-like shape without additional structures or special cuticular areas and the anus is surrounded by a round structure constituted by six cuticular platelets. In
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<emphasis id="4BAAE66E5B4EFFEC62A1001F181FFE56" box="[395,532,368,392]" italics="true" pageId="13" pageNumber="496">B. africanus</emphasis>
</taxonomicName>
the female gonopore has two posterior apunctated areas and the anus is covered by three platelets.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B4EFFEC63ED00D91977FC83" pageId="13" pageNumber="496" type="description">
<paragraph id="79613A7C5B4EFFEC63ED00D91EADFDBE" blockId="13.[151,1436,151,861]" pageId="13" pageNumber="496">
Small dots near the tips of sensorial structures, appearing as black dots (optically dense) under phase contrast microscopy, present in
<taxonomicName id="BEDE41FF5B4EFFEC62B700B31823FE2A" box="[413,552,476,500]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4EFFEC62B700B31823FE2A" box="[413,552,476,500]" italics="true" pageId="13" pageNumber="496">B. dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4EFFEC611A00B21881FE2B" bold="true" box="[560,650,476,501]" pageId="13" pageNumber="496">
<taxonomicNameLabel id="50995B155B4EFFEC611A00B21881FE2B" box="[560,650,476,501]" pageId="13" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were not described for both
<taxonomicName id="BEDE41FF5B4EFFEC60FB00B31E68FE2A" box="[977,1123,476,500]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="tubernatis">
<emphasis id="4BAAE66E5B4EFFEC60FB00B31E68FE2A" box="[977,1123,476,500]" italics="true" pageId="13" pageNumber="496">B. tubernatis</emphasis>
</taxonomicName>
and
<emphasis id="4BAAE66E5B4EFFEC67B100B31F27FE2A" box="[1179,1324,476,500]" italics="true" pageId="13" pageNumber="496">
<taxonomicName id="BEDE41FF5B4EFFEC67B100B31F23FE2A" box="[1179,1320,476,500]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="africanus">B. africanus</taxonomicName>
.
</emphasis>
Optically dense regions below the tip of cephalic cirri were observed by Gallo D’Addabbo
<emphasis id="4BAAE66E5B4EFFEC6764036F1E81FDC6" box="[1102,1162,512,536]" italics="true" pageId="13" pageNumber="496">et al.</emphasis>
(2000), redescribing
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<emphasis id="4BAAE66E5B4EFFEC66AE036F1B12FDE2" italics="true" pageId="13" pageNumber="496">B. dicrocercus</emphasis>
</taxonomicName>
. Probably, the observation of these dots had been neglected in the description of other
<taxonomicName id="BEDE41FF5B4EFFEC661B034A1F97FDE2" box="[1329,1436,548,572]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="genus">
<emphasis id="4BAAE66E5B4EFFEC661B034A1F97FDE2" box="[1329,1436,548,572]" italics="true" pageId="13" pageNumber="496">Batillipes</emphasis>
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species; therefore this character should be carefully used in comparisons of different species.
</paragraph>
<paragraph id="79613A7C5B4EFFEC63ED03021977FC83" blockId="13.[151,1436,151,861]" pageId="13" pageNumber="496">
As previously mentioned, the structure of the revolved sense organ on leg I of
<taxonomicName id="BEDE41FF5B4EFFEC676903031EC4FD5A" box="[1091,1231,620,644]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="dandarae" status="sp. nov.">
<emphasis id="4BAAE66E5B4EFFEC676903031EC4FD5A" box="[1091,1231,620,644]" italics="true" pageId="13" pageNumber="496">B. dandarae</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4EFFEC67FF03021F3BFD5B" bold="true" box="[1237,1328,620,645]" pageId="13" pageNumber="496">
<taxonomicNameLabel id="50995B155B4EFFEC67FF03021F3BFD5B" box="[1237,1328,620,645]" pageId="13" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is shared with
<emphasis id="4BAAE66E5B4EFFEC63FD03FF1B86FD79" box="[215,397,656,680]" italics="true" pageId="13" pageNumber="496">
<taxonomicName id="BEDE41FF5B4EFFEC63FD03FF1B82FD79" box="[215,393,656,680]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="noerrevangi">B. noerrevangi</taxonomicName>
.
</emphasis>
The two species cannot be confused because they exhibit a different toe pattern (
<taxonomicName id="BEDE41FF5B4EFFEC66AE03FF1B14FD12" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="noerrevangi">
<emphasis id="4BAAE66E5B4EFFEC66AE03FF1B14FD12" italics="true" pageId="13" pageNumber="496">B. noerrevangi</emphasis>
</taxonomicName>
belongs to the A group of species proposed by
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, which have the middle toes of legs IV of equal length). Both species can be found in low saline waters. The new species occurring mainly in estuarine beaches, as it is the case of Forte Orange Beach that, according da Rocha
<emphasis id="4BAAE66E5B4EFFEC674603931E95FCCA" box="[1132,1182,764,788]" italics="true" pageId="13" pageNumber="496">et al</emphasis>
. (2004), has a salinity value of 5–29 PSU, while
<taxonomicName id="BEDE41FF5B4EFFEC62E9024F1867FCE9" box="[451,620,800,824]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="noerrevangi">
<emphasis id="4BAAE66E5B4EFFEC62E9024F1867FCE9" box="[451,620,800,824]" italics="true" pageId="13" pageNumber="496">B. noerrevangi</emphasis>
</taxonomicName>
can be found in low saline waters of the North Sea (4–20 PSU) and the Baltic Sea (3–11 PSU) (
<bibRefCitation id="1D4F478D5B4EFFEC6288022A1855FC82" author="Kristensen" box="[418,606,836,861]" pageId="13" pageNumber="496" refString="Kristensen, R. M. (1978) Notes on Marine Heterotardigrades. 1. Description of two Batillipes species, using the electron microscope. Zoologischer Anzeiger, 200, 1 - 17." type="journal article" year="1978">Kristensen 1978</bibRefCitation>
,
<bibRefCitation id="1D4F478D5B4EFFEC6143022A197BFC83" author="Zawierucha" box="[617,880,836,861]" pageId="13" pageNumber="496" refString="Zawierucha, K., Grzelakb, K., Kotwickib, L., Kaczmarekac, L. &amp; Kristensen, R. M. (2015) First observation of the marine tardigrades Batillipes mirus and Batillipes noerrevangi (Arthrotardigrada, Batillipedidae) from a strongly brackish part of the Polish Baltic Sea coast. Marine Biology Research, 11 (8), 859 - 868." type="journal article" year="2015">
Zawierucha
<emphasis id="4BAAE66E5B4EFFEC61DE022B1927FC82" box="[756,812,836,860]" italics="true" pageId="13" pageNumber="496">et al.</emphasis>
2015
</bibRefCitation>
).
</paragraph>
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<emphasis id="4BAAE66E5B4EFFEC63BD02C11B79FC17" bold="true" box="[151,370,943,969]" italics="true" pageId="13" pageNumber="496">Batillipes pennaki</emphasis>
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<emphasis id="4BAAE66E5B4EFFEC63BD02981B61FBD0" box="[151,362,1014,1038]" italics="true" pageId="13" pageNumber="496">Material examined</emphasis>
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<treatment id="F1778B6A5B4EFFF063BD05E81AD2FDC6" ID-DOI="http://doi.org/10.5281/zenodo.5617034" ID-GBIF-Taxon="127844693" ID-Zenodo-Dep="5617034" LSID="urn:lsid:plazi:treatment:F1778B6A5B4EFFF063BD05E81AD2FDC6" httpUri="http://treatment.plazi.org/id/F1778B6A5B4EFFF063BD05E81AD2FDC6" lastPageId="17" lastPageNumber="499" pageId="13" pageNumber="496">
<subSubSection id="31C469F75B4EFFEC63BD05E81B47FB1C" pageId="13" pageNumber="496" type="nomenclature">
<paragraph id="79613A7C5B4EFFEC63BD05E81813FB7E" blockId="13.[151,536,1158,1218]" box="[151,536,1158,1184]" pageId="13" pageNumber="496">
<heading id="22298D105B4EFFEC63BD05E81813FB7E" bold="true" box="[151,536,1158,1184]" fontSize="11" level="1" pageId="13" pageNumber="496" reason="1">
<emphasis id="4BAAE66E5B4EFFEC63BD05E81813FB7E" bold="true" box="[151,536,1158,1184]" pageId="13" pageNumber="496">
<taxonomicName id="BEDE41FF5B4EFFEC63BD05E81BB8FB7E" ID-CoL="f124ba8f-6286-4d2e-9892-031c6f409a89" box="[151,435,1158,1184]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="potiguarensis" status="sp. nov.">
<emphasis id="4BAAE66E5B4EFFEC63BD05E81BB8FB7E" bold="true" box="[151,435,1158,1184]" italics="true" pageId="13" pageNumber="496">Batillipes potiguarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50995B155B4EFFEC629005E81813FB7E" box="[442,536,1158,1184]" pageId="13" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="79613A7C5B4EFFEC63BD05C71B47FB1C" blockId="13.[151,536,1158,1218]" box="[151,332,1193,1218]" pageId="13" pageNumber="496">
(
<figureCitation id="E1E526F95B4EFFEC63B505C71AEFFB1C" box="[159,228,1193,1218]" captionStart="FIGURE 6" captionStartId="14.[151,250,1869,1891]" captionTargetBox="[154,1436,196,1848]" captionTargetId="figure@14.[151,1436,193,1848]" captionTargetPageId="14" captionText="FIGURE 6. Batillipes potiguarensis sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view of a paratype); C. Detail of clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg spine of the first leg (pI) of the holotype; D. Toes (t 1 – t 6) of the leg IV of a paratype; E—Gonopore (go) and anus (an) of the holotypic female. Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400159/files/figure.png" pageId="13" pageNumber="496">Fig. 6</figureCitation>
;
<tableCitation id="345C0FC75B4EFFEC63DA05C71B48FB1C" box="[240,323,1193,1218]" captionStart="TABLE 3" captionStartId="15.[151,239,152,175]" captionTargetBox="[162,1381,380,1848]" captionTargetPageId="15" captionText="TABLE 3. Measurements (in µm) of selected morphological structures of specimens of Batillipes potiguarensis sp. nov. (Holo—Holotype; SD—Standard deviation; Range refers to the smallest and largest measured specimen / structure; Nnumber of specimens / structures measured)." httpUri="http://table.plazi.org/id/2DA16AF45B4CFFEE63BD01F6186CFF31" pageId="13" pageNumber="496" tableUuid="2DA16AF45B4CFFEE63BD01F6186CFF31">Table 3</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B4EFFEC63BD059F189FF9D8" pageId="13" pageNumber="496" type="diagnosis">
<paragraph id="79613A7C5B4EFFEC63BD059F189FF9D8" blockId="13.[151,1437,1265,1974]" pageId="13" pageNumber="496">
<emphasis id="4BAAE66E5B4EFFEC63BD059F1B01FAD4" bold="true" box="[151,266,1265,1290]" pageId="13" pageNumber="496">Diagnosis</emphasis>
. Medium sized
<taxonomicName id="BEDE41FF5B4EFFEC62EA059F1820FAD7" box="[448,555,1265,1289]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="genus">
<emphasis id="4BAAE66E5B4EFFEC62EA059F1820FAD7" box="[448,555,1265,1289]" italics="true" pageId="13" pageNumber="496">Batillipes</emphasis>
</taxonomicName>
with tubular undivided primary clavae and well developed papillar secondary clavae. Cephalic cirri with swollen distal tips. Sensorial spines on all legs. Sensorial organ on leg IV very short. Legs terminated by long toes with spatula-like suction discs. Toes 3 and 4 of legs IV of equal lengths (pattern of the A group of species proposed by
<bibRefCitation id="1D4F478D5B4EFFEC612804331967FAA8" author="Kristensen" box="[514,876,1373,1398]" pageId="13" pageNumber="496" refString="Kristensen, R. M. &amp; Mackness, B. S. (2000) First record of the marine Tardigrade genus Batillipes (Arthrotardigrada: Batillipedidae) from South Australia with a description of a new species. Records of the South Australian Museum, 33 (2), 73 - 87." type="journal article" year="2000">Kristensen and Mackness, 2000</bibRefCitation>
). Distinct head separated from the body by a neck constriction. Scapular region well developed, protruding laterally at the level of the first pair of legs. Small lateral blunt processes between legs I–III. Well-developed blunt processes between legs III and IV. Prominent semicircular caudal projection. Cuticle finely punctated with transverse folds. Rosette-like female gonopore delimited by four punctate cuticular platelets.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B4EFFEC63ED077F1F8FF8F8" pageId="13" pageNumber="496" type="materials_examined">
<paragraph id="79613A7C5B4EFFEC63ED077F1E7CF9F4" blockId="13.[151,1437,1265,1974]" box="[199,1143,1553,1578]" pageId="13" pageNumber="496">
<materialsCitation id="C9B630215B4EFFEC63ED077F1E7CF9F4" ID-GBIF-Occurrence="1440156682" box="[199,1143,1553,1578]" country="Brazil" latitude="-6.2280555" location="Amor Beach" longLatPrecision="21" longitude="-35.04139" pageId="13" pageNumber="496" specimenCount="1" stateProvince="Rio Grande do Norte" typeStatus="holotype">
<emphasis id="4BAAE66E5B4EFFEC63ED077F1B6EF9F4" bold="true" box="[199,357,1553,1578]" pageId="13" pageNumber="496">
<typeStatus id="A66584DE5B4EFFEC63ED077F1B09F9F4" box="[199,258,1553,1578]" pageId="13" pageNumber="496">Type</typeStatus>
locality.
</emphasis>
<location id="7C016CA75B4EFFEC6246077C1BF6F9F4" LSID="urn:lsid:plazi:treatment:F1778B6A5B4EFFF063BD05E81AD2FDC6:7C016CA75B4EFFEC6246077C1BF6F9F4" box="[364,509,1553,1578]" country="Brazil" latitude="-6.2280555" longLatPrecision="21" longitude="-35.04139" name="Amor Beach" pageId="13" pageNumber="496" stateProvince="Rio Grande do Norte">Amor Beach</location>
,
<collectingRegion id="BB1AF49E5B4EFFEC6122077F18FCF9F7" box="[520,759,1553,1578]" country="Brazil" name="Rio Grande do Norte" pageId="13" pageNumber="496">
Rio
<collectingRegion id="BB1AF49E5B4EFFEC611D077F18FCF9F7" box="[567,759,1553,1578]" inferredBySuffix="true" pageId="13" pageNumber="496">Grande do Norte</collectingRegion>
</collectingRegion>
,
<collectingCountry id="01C97AEC5B4EFFEC6028077F1942F9F4" box="[770,841,1553,1578]" name="Brazil" pageId="13" pageNumber="496">Brazil</collectingCountry>
, (
<geoCoordinate id="1CEA5CBB5B4EFFEC6077077F19DAF9F4" box="[861,977,1553,1578]" direction="south" orientation="latitude" pageId="13" pageNumber="496" precision="15" value="-6.2280555">6°13'41&quot;S</geoCoordinate>
,
<geoCoordinate id="1CEA5CBB5B4EFFEC60F7077F1E60F9F4" box="[989,1131,1553,1578]" direction="west" orientation="longitude" pageId="13" pageNumber="496" precision="15" value="-35.04139">35°02'29&quot;W</geoCoordinate>
).
</materialsCitation>
</paragraph>
<paragraph id="79613A7C5B4EFFEC63ED075B18A5F964" blockId="13.[151,1437,1265,1974]" pageId="13" pageNumber="496">
<emphasis id="4BAAE66E5B4EFFEC63ED075B1B70F990" bold="true" box="[199,379,1589,1614]" pageId="13" pageNumber="496">Type material.</emphasis>
Holotype: adult, female (slide CVII-82) collected at Amor beach, mounted in glycerol. Paratypes mounted in glycerol: one female collected at Gunga beach (slide TARD/UFRPE 02-23); and 3 females collected at Amor beach (slides TARD/UFRPE 02-24 and CVII-82); 1 female and 1 juvenile collected at Francês Beach (slides TARD/UFRPE
<date id="0D601CBC5B4EFFEC62CE07CF18A9F964" box="[484,674,1697,1722]" pageId="13" pageNumber="496" value="1926-02-25" valueMax="1926-02-02" valueMin="1926-02-25">02-25 and 02-26</date>
).
</paragraph>
<paragraph id="79613A7C5B4EFFEC63ED07AB1F8FF8F8" blockId="13.[151,1437,1265,1974]" pageId="13" pageNumber="496">
<materialsCitation id="C9B630215B4EFFEC63ED07AB1F8BF8F8" ID-GBIF-Occurrence="1440156681" collectionCode="UFRPE" country="Portugal" county="Department" location="Faculty" municipality="Biology" pageId="13" pageNumber="496" specimenCount="2" stateProvince="Pernambuco" typeStatus="holotype">
<emphasis id="4BAAE66E5B4EFFEC63ED07AB1B85F900" bold="true" box="[199,398,1733,1758]" pageId="13" pageNumber="496">
<typeStatus id="A66584DE5B4EFFEC63ED07AB1B09F900" box="[199,258,1733,1758]" pageId="13" pageNumber="496">Type</typeStatus>
repository.
</emphasis>
The
<typeStatus id="A66584DE5B4EFFEC62FE07A8180FF900" box="[468,516,1734,1758]" pageId="13" pageNumber="496">type</typeStatus>
material (slide CVII-82) is deposited in the collection of the
<collectingCounty id="900042F05B4EFFEC67DE07A81F73F903" box="[1268,1400,1734,1757]" pageId="13" pageNumber="496">Department</collectingCounty>
of
<collectingMunicipality id="9905A0065B4EFFEC63BD07871AF9F8DC" box="[151,242,1769,1794]" pageId="13" pageNumber="496">Biology</collectingMunicipality>
,
<location id="7C016CA75B4EFFEC63D407871B59F8DC" LSID="urn:lsid:plazi:treatment:F1778B6A5B4EFFF063BD05E81AD2FDC6:7C016CA75B4EFFEC63D407871B59F8DC" box="[254,338,1769,1794]" country="Portugal" county="Department" municipality="Biology" name="Faculty" pageId="13" pageNumber="496" stateProvince="Pernambuco">Faculty</location>
of
<location id="7C016CA75B4EFFEC625207871BD6F8DC" LSID="urn:lsid:plazi:treatment:F1778B6A5B4EFFF063BD05E81AD2FDC6:7C016CA75B4EFFEC625207871BD6F8DC" box="[376,477,1769,1794]" country="Portugal" county="Department" municipality="Biology" name="Sciences" pageId="13" pageNumber="496" stateProvince="Pernambuco">Sciences</location>
,
<location id="7C016CA75B4EFFEC62C307871854F8DC" LSID="urn:lsid:plazi:treatment:F1778B6A5B4EFFF063BD05E81AD2FDC6:7C016CA75B4EFFEC62C307871854F8DC" box="[489,607,1769,1794]" country="Portugal" county="Department" municipality="Biology" name="University" pageId="13" pageNumber="496" stateProvince="Pernambuco">University</location>
of
<location id="7C016CA75B4EFFEC61AC078418CEF8DF" LSID="urn:lsid:plazi:treatment:F1778B6A5B4EFFF063BD05E81AD2FDC6:7C016CA75B4EFFEC61AC078418CEF8DF" box="[646,709,1770,1793]" country="Portugal" county="Department" municipality="Biology" name="Porto" pageId="13" pageNumber="496" stateProvince="Pernambuco">Porto</location>
,
<collectingCountry id="01C97AEC5B4EFFEC61FB07871938F8DC" box="[721,819,1769,1794]" name="Portugal" pageId="13" pageNumber="496">Portugal</collectingCountry>
, and the other slides in the collection of Tardigrades—
<collectionCode id="1FCFA2B95B4EFFEC63BD06601AE5F8FB" box="[151,238,1806,1829]" pageId="13" pageNumber="496">UFRPE</collectionCode>
(
<location id="7C016CA75B4EFFEC63D606631B71F8F8" LSID="urn:lsid:plazi:treatment:F1778B6A5B4EFFF063BD05E81AD2FDC6:7C016CA75B4EFFEC63D606631B71F8F8" box="[252,378,1805,1830]" country="Portugal" county="Department" municipality="Biology" name="Laboratory" pageId="13" pageNumber="496" stateProvince="Pernambuco">Laboratory</location>
of
<location id="7C016CA75B4EFFEC628A06631817F8F8" LSID="urn:lsid:plazi:treatment:F1778B6A5B4EFFF063BD05E81AD2FDC6:7C016CA75B4EFFEC628A06631817F8F8" box="[416,540,1805,1830]" country="Portugal" county="Department" municipality="Biology" name="Meiofauna" pageId="13" pageNumber="496" stateProvince="Pernambuco">Meiofauna</location>
,
<location id="7C016CA75B4EFFEC610D066018A0F8FB" LSID="urn:lsid:plazi:treatment:F1778B6A5B4EFFF063BD05E81AD2FDC6:7C016CA75B4EFFEC610D066018A0F8FB" box="[551,683,1806,1829]" country="Portugal" county="Department" municipality="Biology" name="Department" pageId="13" pageNumber="496" stateProvince="Pernambuco">Department</location>
of
<location id="7C016CA75B4EFFEC61FB06631927F8F8" LSID="urn:lsid:plazi:treatment:F1778B6A5B4EFFF063BD05E81AD2FDC6:7C016CA75B4EFFEC61FB06631927F8F8" box="[721,812,1805,1830]" country="Portugal" county="Department" municipality="Biology" name="Biology" pageId="13" pageNumber="496" stateProvince="Pernambuco">Biology</location>
, Universidade Federal Rural de
<collectingRegion id="BB1AF49E5B4EFFEC67BF06631F2FF8F8" box="[1173,1316,1805,1830]" country="Brazil" name="Pernambuco" pageId="13" pageNumber="496">Pernambuco</collectingRegion>
,
<collectingCountry id="01C97AEC5B4EFFEC660506631F73F8F8" box="[1327,1400,1805,1830]" name="Brazil" pageId="13" pageNumber="496">Brazil</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B4EFFEC63ED065F1F4EF868" pageId="13" pageNumber="496" type="etymology">
<paragraph id="79613A7C5B4EFFEC63ED065F1935F8B0" blockId="13.[151,1437,1265,1974]" pageId="13" pageNumber="496">
<emphasis id="4BAAE66E5B4EFFEC63ED065F1B45F894" bold="true" box="[199,334,1841,1866]" pageId="13" pageNumber="496">Etymology.</emphasis>
The name
<taxonomicName id="BEDE41FF5B4EFFEC62E6065C1868F897" box="[460,611,1842,1865]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="potiguarensis">
<emphasis id="4BAAE66E5B4EFFEC62E6065C1868F897" box="[460,611,1842,1865]" italics="true" pageId="13" pageNumber="496">potiguarensis</emphasis>
</taxonomicName>
refers the inhabitants of the State of
<collectingRegion id="BB1AF49E5B4EFFEC672E065F1EFAF897" box="[1028,1265,1841,1866]" country="Brazil" name="Rio Grande do Norte" pageId="13" pageNumber="496">Rio Grande do Norte</collectingRegion>
where the new species was found, primitively named the Potiguar Territory.
</paragraph>
<paragraph id="79613A7C5B4EFFEC63ED06171F4EF868" blockId="13.[151,1437,1265,1974]" pageId="13" pageNumber="496">
<emphasis id="4BAAE66E5B4EFFEC63ED06171B8CF84C" bold="true" box="[199,391,1913,1938]" pageId="13" pageNumber="496">Ecological note.</emphasis>
<taxonomicName id="BEDE41FF5B4EFFEC62BB06141840F84F" box="[401,587,1914,1937]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="13" pageNumber="496" phylum="Tardigrada" rank="species" species="potiguarensis" status="sp. nov.">
<emphasis id="4BAAE66E5B4EFFEC62BB06141840F84F" box="[401,587,1914,1937]" italics="true" pageId="13" pageNumber="496">B. potiguarensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4EFFEC617C061718BFF84C" bold="true" box="[598,692,1913,1938]" pageId="13" pageNumber="496">
<taxonomicNameLabel id="50995B155B4EFFEC617C061718BFF84C" box="[598,692,1913,1938]" pageId="13" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found in shallow sublittoral medium to coarse gravels and quartz sands, in reef pools of high energy beaches (Amor Beach) and low energy beaches (Francês Beach).
</paragraph>
</subSubSection>
<caption id="2DA16AF45B4DFFEF63BD06231B23F861" httpUri="https://zenodo.org/record/400159/files/figure.png" pageId="14" pageNumber="497" targetBox="[154,1436,196,1848]" targetPageId="14">
<paragraph id="79613A7C5B4DFFEF63BD06231B23F861" blockId="14.[151,1436,1869,1983]" pageId="14" pageNumber="497">
<emphasis id="4BAAE66E5B4DFFEF63BD06231B05F8BD" bold="true" box="[151,270,1869,1891]" pageId="14" pageNumber="497">FIGURE 6</emphasis>
.
<taxonomicName id="BEDE41FF5B4DFFEF623606231805F8BA" box="[284,526,1869,1892]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="14" pageNumber="497" phylum="Tardigrada" rank="species" species="potiguarensis" status="sp. nov.">
<emphasis id="4BAAE66E5B4DFFEF623606231805F8BA" box="[284,526,1869,1892]" italics="true" pageId="14" pageNumber="497">Batillipes potiguarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50995B155B4DFFEF613F0620186CF8BD" box="[533,615,1870,1891]" pageId="14" pageNumber="497" rank="species">
<emphasis id="4BAAE66E5B4DFFEF613F0620186AF8BD" bold="true" box="[533,609,1870,1891]" pageId="14" pageNumber="497">sp. nov</emphasis>
.
</taxonomicNameLabel>
A. Drawing (dorsal view); B. Habitus (dorsal view of a paratype); C. Detail of clavae (pc—primary clava; sc—secondary clava), first body projection (pr1) and leg spine of the first leg (pI) of the holotype; D. Toes (t1–t6) of the leg IV of a paratype; E—Gonopore (go) and anus (an) of the holotypic female. Scale bars: A–B = 50 µm; C–E = 10 µm.
</paragraph>
</caption>
<caption id="2DA16AF45B4CFFEE63BD01F6186CFF31" ID-Table-UUID="2DA16AF45B4CFFEE63BD01F6186CFF31" httpUri="http://table.plazi.org/id/2DA16AF45B4CFFEE63BD01F6186CFF31" pageId="15" pageNumber="498" targetBox="[162,1381,380,1848]" targetIsTable="true" targetPageId="15">
<paragraph id="79613A7C5B4CFFEE63BD01F6186CFF31" blockId="15.[151,1436,151,239]" pageId="15" pageNumber="498">
<emphasis id="4BAAE66E5B4CFFEE63BD01F61B02FF71" bold="true" box="[151,265,152,175]" pageId="15" pageNumber="498">TABLE 3.</emphasis>
Measurements (in µm) of selected morphological structures of specimens of
<taxonomicName id="BEDE41FF5B4CFFEE676901F91F4BFF71" box="[1091,1344,151,175]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="15" pageNumber="498" phylum="Tardigrada" rank="species" species="potiguarensis" status="sp. nov.">
<emphasis id="4BAAE66E5B4CFFEE676901F91F4BFF71" box="[1091,1344,151,175]" italics="true" pageId="15" pageNumber="498">Batillipes potiguarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50995B155B4CFFEE666D01F61F97FF71" box="[1351,1436,152,175]" pageId="15" pageNumber="498" rank="species">
<emphasis id="4BAAE66E5B4CFFEE666D01F61F97FF71" bold="true" box="[1351,1436,152,175]" pageId="15" pageNumber="498">sp. nov.</emphasis>
</taxonomicNameLabel>
(Holo—Holotype; SD—Standard deviation; Range refers to the smallest and largest measured specimen/structure; Nnumber of specimens/structures measured).
</paragraph>
</caption>
<subSubSection id="31C469F75B4CFFF1616300641E88FA8B" lastPageId="16" lastPageNumber="499" pageId="15" pageNumber="498" type="description">
<paragraph id="79613A7C5B4CFFEE616300641F26FEE1" blockId="15.[162,1381,266,360]" pageId="15" pageNumber="498">FEMALES JUVENILE adults 6-toed</paragraph>
<paragraph id="79613A7C5B4CFFEE6388003C19E2FEB6" blockId="15.[162,1381,266,360]" box="[162,1001,338,360]" pageId="15" pageNumber="498">STRUCTURES Holotype Mean ± SD (Range); N</paragraph>
<paragraph id="79613A7C5B4CFFEE638800121EFEF8E6" pageId="15" pageNumber="498">
<table id="0BDEC8DC5B4C001E638800121F6EF8E6" box="[162,1381,380,1848]" gridcols="4" gridrows="12" pageId="15" pageNumber="498">
<tr id="C7EE383E5B4C001E638800121F6EFE3A" box="[162,1381,380,484]" gridrow="0" pageId="15" pageNumber="498">
<th id="843F51425B4C001E638800121BCEFE3A" box="[162,453,380,484]" gridcol="0" gridrow="0" pageId="15" pageNumber="498">Body length (BL) BL without caudal apparatus Body width</th>
<th id="843F51425B4C001E61630012187EFE3A" box="[585,629,380,484]" gridcol="1" gridrow="0" pageId="15" pageNumber="498">187 171 62.1</th>
<th id="843F51425B4C001E61D700121E38FE3A" box="[765,1075,380,484]" gridcol="2" gridrow="0" pageId="15" pageNumber="498">184.5 ±18.04 (152.2–201.1); 6 168.8 ±21.55 (129.3–191.5); 6 64.5 ±11.26 (47.9–78.1); 6</th>
<th id="843F51425B4C001E67C700121F6EFE3A" box="[1261,1381,380,484]" gridcol="3" gridrow="0" pageId="15" pageNumber="498">183 175?</th>
</tr>
<tr id="C7EE383E5B4C001E638800961F6EFDBE" box="[162,1381,504,608]" gridrow="1" pageId="15" pageNumber="498">
<th id="843F51425B4C001E638800961BCEFDBE" box="[162,453,504,608]" gridcol="0" gridrow="1" pageId="15" pageNumber="498">Scapular region width Median cirrus Internal cirri</th>
<td id="843F51425B4C001E61630096187EFDBE" box="[585,629,504,608]" gridcol="1" gridrow="1" pageId="15" pageNumber="498">70.8 20.5 21.8</td>
<td id="843F51425B4C001E61D700961E38FDBE" box="[765,1075,504,608]" gridcol="2" gridrow="1" pageId="15" pageNumber="498">69.1 ±7.54 (58.0–80.2); 6 16.0 ±4.20 (10.4–20.5); 4 20.5 ±1.93 (17.0–22.5); 6</td>
<td id="843F51425B4C001E67C700961F6EFDBE" box="[1261,1381,504,608]" gridcol="3" gridrow="1" pageId="15" pageNumber="498">49.7 17.2 18.0</td>
</tr>
<tr id="C7EE383E5B4C001E6388031A1F6EFD02" box="[162,1381,628,732]" gridrow="2" pageId="15" pageNumber="498">
<th id="843F51425B4C001E6388031A1BCEFD02" box="[162,453,628,732]" gridcol="0" gridrow="2" pageId="15" pageNumber="498">External cirri Lateral cirri A Primary clavae</th>
<td id="843F51425B4C001E6163031A187EFD02" box="[585,629,628,732]" gridcol="1" gridrow="2" pageId="15" pageNumber="498">20.1 35.8 19.4</td>
<td id="843F51425B4C001E61D7031A1E38FD02" box="[765,1075,628,732]" gridcol="2" gridrow="2" pageId="15" pageNumber="498">16.0 ±3.26 (12.6–20.1); 5 28.8 ±4.27 (22.7–35.8); 6 16.5 ±2.26 (13.4–19.4); 6</td>
<td id="843F51425B4C001E67C7031A1F6EFD02" box="[1261,1381,628,732]" gridcol="3" gridrow="2" pageId="15" pageNumber="498">14.6 24.7 12.8</td>
</tr>
<tr id="C7EE383E5B4C001E6388039E1F6EFC86" box="[162,1381,752,856]" gridrow="3" pageId="15" pageNumber="498">
<th id="843F51425B4C001E6388039E1BCEFC86" box="[162,453,752,856]" gridcol="0" gridrow="3" pageId="15" pageNumber="498">Primary clava / cirrus A Secondary clavae Cirri E</th>
<td id="843F51425B4C001E6163039E187EFC86" box="[585,629,752,856]" gridcol="1" gridrow="3" pageId="15" pageNumber="498">0.54 5.6 20.1</td>
<td id="843F51425B4C001E61D7039E1E38FC86" box="[765,1075,752,856]" gridcol="2" gridrow="3" pageId="15" pageNumber="498">0.58 ±0.10 (0.47–0.75); 6 5.1 ±1.05 (3.7–6.2); 5 20.3 ±2.27 (17.0–22.8); 5</td>
<td id="843F51425B4C001E67C7039E1F6EFC86" box="[1261,1381,752,856]" gridcol="3" gridrow="3" pageId="15" pageNumber="498">0.52 6.2?</td>
</tr>
<tr id="C7EE383E5B4C001E638802021F6EFC0A" box="[162,1381,876,980]" gridrow="4" pageId="15" pageNumber="498">
<th id="843F51425B4C001E638802021BCEFC0A" box="[162,453,876,980]" gridcol="0" gridrow="4" pageId="15" pageNumber="498">Leg I spine Leg II spine Leg III spine</th>
<td id="843F51425B4C001E61630202187EFC0A" box="[585,629,876,980]" gridcol="1" gridrow="4" pageId="15" pageNumber="498">9.8 11.4 10.8</td>
<td id="843F51425B4C001E61D702021E38FC0A" box="[765,1075,876,980]" gridcol="2" gridrow="4" pageId="15" pageNumber="498">10.0 ±0.93 (8.3–10.9); 6 9.8 ±2.51 (7.3–13.5); 5 10.6 ±2.64 (7.2–13.1); 4</td>
<td id="843F51425B4C001E67C702021F6EFC0A" box="[1261,1381,876,980]" gridcol="3" gridrow="4" pageId="15" pageNumber="498">? 15.2 10.6</td>
</tr>
<tr id="C7EE383E5B4C001E638802861F6EFB8E" box="[162,1381,1000,1104]" gridrow="5" pageId="15" pageNumber="498">
<th id="843F51425B4C001E638802861BCEFB8E" box="[162,453,1000,1104]" gridcol="0" gridrow="5" pageId="15" pageNumber="498">Leg IV organ Leg IV base Leg IV spine</th>
<td id="843F51425B4C001E61630286187EFB8E" box="[585,629,1000,1104]" gridcol="1" gridrow="5" pageId="15" pageNumber="498">6.6 3.1 3.5</td>
<td id="843F51425B4C001E61D702861E38FB8E" box="[765,1075,1000,1104]" gridcol="2" gridrow="5" pageId="15" pageNumber="498">8.4 ±1.9 (6.6–11.1); 6 3.6 ±0.92 (3.0–5.4); 5 4.2 ±0.79 (3.5–5.4); 5</td>
<td id="843F51425B4C001E67C702861F6EFB8E" box="[1261,1381,1000,1104]" gridcol="3" gridrow="5" pageId="15" pageNumber="498">6.5??</td>
</tr>
<tr id="C7EE383E5B4C001E6388050A1F6EFB12" box="[162,1381,1124,1228]" gridrow="6" pageId="15" pageNumber="498">
<th id="843F51425B4C001E6388050A1BCEFB12" box="[162,453,1124,1228]" gridcol="0" gridrow="6" pageId="15" pageNumber="498">Body projection 1 Body projection 2 Body projection 3</th>
<td id="843F51425B4C001E6163050A187EFB12" box="[585,629,1124,1228]" gridcol="1" gridrow="6" pageId="15" pageNumber="498">6.8 4.6 5.8</td>
<td id="843F51425B4C001E61D7050A1E38FB12" box="[765,1075,1124,1228]" gridcol="2" gridrow="6" pageId="15" pageNumber="498">6.0 ±1.62 (3.2–7.8); 6 4.4 ±1.47 (2.5–6.1); 4 5.5 ±1.30 (3.8–7.4); 5</td>
<td id="843F51425B4C001E67C7050A1F6EFB12" box="[1261,1381,1124,1228]" gridcol="3" gridrow="6" pageId="15" pageNumber="498">3.9 3.5 5.1</td>
</tr>
<tr id="C7EE383E5B4C001E6388058E1F6EFA96" box="[162,1381,1248,1352]" gridrow="7" pageId="15" pageNumber="498">
<th id="843F51425B4C001E6388058E1BCEFA96" box="[162,453,1248,1352]" gridcol="0" gridrow="7" pageId="15" pageNumber="498">Body projection 4 Caudal apparatus Legs I–III Toe 1</th>
<td id="843F51425B4C001E6163058E187EFA96" box="[585,629,1248,1352]" gridcol="1" gridrow="7" pageId="15" pageNumber="498">9.1 15.2 10.3</td>
<td id="843F51425B4C001E61D7058E1E38FA96" box="[765,1075,1248,1352]" gridcol="2" gridrow="7" pageId="15" pageNumber="498">9.0 ±1.52 (6.6–10.8); 5 18.8 ±4.27 (14.0–23.3); 5 10.9 ±0.50 (10.3–11.5); 4</td>
<td id="843F51425B4C001E67C7058E1F6EFA96" box="[1261,1381,1248,1352]" gridcol="3" gridrow="7" pageId="15" pageNumber="498">? 8.0 9.1</td>
</tr>
<tr id="C7EE383E5B4C001E638804321F6EFA1A" box="[162,1381,1372,1476]" gridrow="8" pageId="15" pageNumber="498">
<th id="843F51425B4C001E638804321BCEFA1A" box="[162,453,1372,1476]" gridcol="0" gridrow="8" pageId="15" pageNumber="498">Toe 2 Toe 3 Toe 4</th>
<td id="843F51425B4C001E61630432187EFA1A" box="[585,629,1372,1476]" gridcol="1" gridrow="8" pageId="15" pageNumber="498">7.9 19.7 11.4</td>
<td id="843F51425B4C001E61D704321E38FA1A" box="[765,1075,1372,1476]" gridcol="2" gridrow="8" pageId="15" pageNumber="498">8.2 ±1.83 (6.5–10.8); 4 18.6 ±1.06 (17.6–19.7); 4 11.7 ±1.93 (9.5–14.2); 4</td>
<td id="843F51425B4C001E67C704321F6EFA1A" box="[1261,1381,1372,1476]" gridcol="3" gridrow="8" pageId="15" pageNumber="498">4.7 13.3 9</td>
</tr>
<tr id="C7EE383E5B4C001E638804B61F6EF99E" box="[162,1381,1496,1600]" gridrow="9" pageId="15" pageNumber="498">
<th id="843F51425B4C001E638804B61BCEF99E" box="[162,453,1496,1600]" gridcol="0" gridrow="9" pageId="15" pageNumber="498">Toe 5 Toe 6 Leg IV Toe 1</th>
<td id="843F51425B4C001E616304B6187EF99E" box="[585,629,1496,1600]" gridcol="1" gridrow="9" pageId="15" pageNumber="498">20.6 12.4 -</td>
<td id="843F51425B4C001E61D704B61E38F99E" box="[765,1075,1496,1600]" gridcol="2" gridrow="9" pageId="15" pageNumber="498">21.8 ±1.43 (20.6–23.4); 3 14.0 ±1.38 (12.4–15.0); 3 - (17.5–20.4); 2</td>
<td id="843F51425B4C001E67C704B61F6EF99E" box="[1261,1381,1496,1600]" gridcol="3" gridrow="9" pageId="15" pageNumber="498">14.7 9.8 12.3</td>
</tr>
<tr id="C7EE383E5B4C001E6388073A1F6EF962" box="[162,1381,1620,1724]" gridrow="10" pageId="15" pageNumber="498">
<th id="843F51425B4C001E6388073A1BCEF962" box="[162,453,1620,1724]" gridcol="0" gridrow="10" pageId="15" pageNumber="498">Toe 2 Toe 3 Toe 4</th>
<td id="843F51425B4C001E6163073A187EF962" box="[585,629,1620,1724]" gridcol="1" gridrow="10" pageId="15" pageNumber="498">- 15.5 15.7</td>
<td id="843F51425B4C001E61D7073A1E38F962" box="[765,1075,1620,1724]" gridcol="2" gridrow="10" pageId="15" pageNumber="498">- (25.6); 1 13.3 ±2.70 (10.3–15.5); 3 13.4 ±2.60 (10.6–15.7); 3</td>
<td id="843F51425B4C001E67C7073A1F6EF962" box="[1261,1381,1620,1724]" gridcol="3" gridrow="10" pageId="15" pageNumber="498">15.5 9.9 10.4</td>
</tr>
<tr id="C7EE383E5B4C001E638807BE1F6EF8E6" box="[162,1381,1744,1848]" gridrow="11" pageId="15" pageNumber="498">
<th id="843F51425B4C001E638807BE1BCEF8E6" box="[162,453,1744,1848]" gridcol="0" gridrow="11" pageId="15" pageNumber="498">Toe 5 Toe 6 Gonopore - Anus</th>
<td id="843F51425B4C001E616307BE187EF8E6" box="[585,629,1744,1848]" gridcol="1" gridrow="11" pageId="15" pageNumber="498">25.7 19.1 10.8</td>
<td id="843F51425B4C001E61D707BE1E38F8E6" box="[765,1075,1744,1848]" gridcol="2" gridrow="11" pageId="15" pageNumber="498">- (25.7–27.5); 2 18.8 ±1.01 (17.1–19.1); 3 9.0 ±2.21 (6.6–11.1); 5</td>
<td id="843F51425B4C001E67C707BE1F6EF8E6" box="[1261,1381,1744,1848]" gridcol="3" gridrow="11" pageId="15" pageNumber="498">18.2 11.0 -</td>
</tr>
</table>
</paragraph>
<paragraph id="79613A7C5B4CFFF163ED06131B8BFDC6" blockId="15.[151,1437,1917,2014]" lastBlockId="16.[151,1437,151,2013]" lastPageId="16" lastPageNumber="499" pageId="15" pageNumber="498">
<emphasis id="4BAAE66E5B4CFFEE63ED06131804F848" bold="true" box="[199,527,1917,1942]" pageId="15" pageNumber="498">
Description of the
<typeStatus id="A66584DE5B4CFFEE628B06131801F848" box="[417,522,1917,1942]" pageId="15" pageNumber="498" type="holotype">holotype</typeStatus>
.
</emphasis>
Female, 171 µm long (187 µm including the caudal apparatus) and 62.1 µm wide between the third and fourth pair of legs (
<figureCitation id="E1E526F95B4CFFEE61A806CF18C7F864" box="[642,716,1953,1978]" captionStart="FIGURE 6" captionStartId="14.[151,250,1869,1891]" captionTargetBox="[154,1436,196,1848]" captionTargetId="figure@14.[151,1436,193,1848]" captionTargetPageId="14" captionText="FIGURE 6. Batillipes potiguarensis sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view of a paratype); C. Detail of clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg spine of the first leg (pI) of the holotype; D. Toes (t 1 – t 6) of the leg IV of a paratype; E—Gonopore (go) and anus (an) of the holotypic female. Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400159/files/figure.png" pageId="15" pageNumber="498">Figs 6</figureCitation>
A, B). Typical trapezoid head separated from the body by an evident neck constriction. Head bearing eleven cephalic appendages. Internal cirri inserted dorsally on the frontal edge of the head are 21.8 µm long with cirrophores (about 3.8 µm long). External cirri 20.1 µm long, with indistinct cirrophores, inserted more ventrally, near the lateral cirri and primary clavae. The median cirrus, with cirrophore (2.6 µm), is 20.5 µm long. The lateral cirrus, 35.8 µm long, is located dorsally in relation to the unconstricted, but wrinkled, tubular primary clava, 19.4 µm long (
<figureCitation id="E1E526F95B53FFF1605D006A19B8FEC3" box="[887,947,260,285]" captionStart="FIGURE 6" captionStartId="14.[151,250,1869,1891]" captionTargetBox="[154,1436,196,1848]" captionTargetId="figure@14.[151,1436,193,1848]" captionTargetPageId="14" captionText="FIGURE 6. Batillipes potiguarensis sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view of a paratype); C. Detail of clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg spine of the first leg (pI) of the holotype; D. Toes (t 1 – t 6) of the leg IV of a paratype; E—Gonopore (go) and anus (an) of the holotypic female. Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400159/files/figure.png" pageId="16" pageNumber="499">Fig 6</figureCitation>
C). These two appendages share a common pedestal. A van der Land’s organ is present inside the base of the primary clava that has a terminal pore. Well developed and prominent papillar secondary clavae (diameter about 5.6 µm) are visible on the frontal edge of the head (
<figureCitation id="E1E526F95B53FFF163F700011B10FE56" box="[221,283,367,392]" captionStart="FIGURE 6" captionStartId="14.[151,250,1869,1891]" captionTargetBox="[154,1436,196,1848]" captionTargetId="figure@14.[151,1436,193,1848]" captionTargetPageId="14" captionText="FIGURE 6. Batillipes potiguarensis sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view of a paratype); C. Detail of clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg spine of the first leg (pI) of the holotype; D. Toes (t 1 – t 6) of the leg IV of a paratype; E—Gonopore (go) and anus (an) of the holotypic female. Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400159/files/figure.png" pageId="16" pageNumber="499">Fig 6</figureCitation>
C). All the cephalic cirri, including the external cirri and the lateral cirri have a optically dense dot (black dot when observed under PHC) near their swollen tips. There is no notch between internal and external cirrus nor an indentation between the external cephalic cirrus and the pedestal bearing the primary clava and lateral cirrus. Eye spots not observed. The ovoid pharyngeal bulb is 21.3 µm long and 20.7 µm wide. Placoids not visible after slide mounting.
</paragraph>
<paragraph id="79613A7C5B53FFF163ED034A1BBDFD2E" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="499">
A lateral blunt expansion (auricle), 6.8 µm long, is present between the head and the first pair of legs (
<figureCitation id="E1E526F95B53FFF1661D034A1F78FDE3" box="[1335,1395,548,573]" captionStart="FIGURE 6" captionStartId="14.[151,250,1869,1891]" captionTargetBox="[154,1436,196,1848]" captionTargetId="figure@14.[151,1436,193,1848]" captionTargetPageId="14" captionText="FIGURE 6. Batillipes potiguarensis sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view of a paratype); C. Detail of clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg spine of the first leg (pI) of the holotype; D. Toes (t 1 – t 6) of the leg IV of a paratype; E—Gonopore (go) and anus (an) of the holotypic female. Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400159/files/figure.png" pageId="16" pageNumber="499">Fig 6</figureCitation>
C). The scapular region is well developed (70.8 µm wide) protruding laterally at the level of the first pair of legs (
<figureCitation id="E1E526F95B53FFF1664503291AAEFD5B" captionStart="FIGURE 6" captionStartId="14.[151,250,1869,1891]" captionTargetBox="[154,1436,196,1848]" captionTargetId="figure@14.[151,1436,193,1848]" captionTargetPageId="14" captionText="FIGURE 6. Batillipes potiguarensis sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view of a paratype); C. Detail of clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg spine of the first leg (pI) of the holotype; D. Toes (t 1 – t 6) of the leg IV of a paratype; E—Gonopore (go) and anus (an) of the holotypic female. Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400159/files/figure.png" pageId="16" pageNumber="499">Fig. 6</figureCitation>
A, B). At the level of the second pair of legs a weak body protrusion is also visible. Ventrolateral blunt processes (
<figureCitation id="E1E526F95B53FFF163B503E11AE9FD76" box="[159,226,655,680]" captionStart="FIGURE 6" captionStartId="14.[151,250,1869,1891]" captionTargetBox="[154,1436,196,1848]" captionTargetId="figure@14.[151,1436,193,1848]" captionTargetPageId="14" captionText="FIGURE 6. Batillipes potiguarensis sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view of a paratype); C. Detail of clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg spine of the first leg (pI) of the holotype; D. Toes (t 1 – t 6) of the leg IV of a paratype; E—Gonopore (go) and anus (an) of the holotypic female. Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400159/files/figure.png" pageId="16" pageNumber="499">Fig. 6</figureCitation>
A, B) are present between all leg pairs, particularly developed between legs III and IV (4.6, 5.8 and 9.1 µm between legs I–II, II–III and III–IV respectively). The caudal region has a conspicuous semicircle-shaped protrusion, 15.2 µm long.
</paragraph>
<paragraph id="79613A7C5B53FFF163ED03921B05FC7B" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="499">
Sharply pointed (20.1 µm long) cirri E with small cirrophores. Sensorial organs present on all legs (9.8; 11.4; 10.8, and 6.6 on leg I, II, III and IV respectively). Leg sense organs of legs I and IV divided into a basal and a distal portion with a dot near their swollen tips. The short sense organ on leg IV (
<figureCitation id="E1E526F95B53FFF160CF022A1E2AFC83" box="[997,1057,836,861]" captionStart="FIGURE 6" captionStartId="14.[151,250,1869,1891]" captionTargetBox="[154,1436,196,1848]" captionTargetId="figure@14.[151,1436,193,1848]" captionTargetPageId="14" captionText="FIGURE 6. Batillipes potiguarensis sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view of a paratype); C. Detail of clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg spine of the first leg (pI) of the holotype; D. Toes (t 1 – t 6) of the leg IV of a paratype; E—Gonopore (go) and anus (an) of the holotypic female. Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400159/files/figure.png" pageId="16" pageNumber="499">Fig 6</figureCitation>
A, B) has a van der Land’s organ separating the two portions (basal portion 3.1 µm long; distal portion 3.5 µm long). Sense organs on legs II and III are spines.
</paragraph>
<paragraph id="79613A7C5B53FFF163ED02C11EE5FB1B" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="499">
Telescopic legs with long toes. Toes with the distal stalk with a distal enlargement (3.0 µm wide), spatula-like suction discs (5.3 µm long and 4.7 µm wide on leg IV) with straight frontal edge, short and thin braces and slightly thickened lateral edges. In the first three pair of legs, toe 2 is the shortest, toes 3 and 5 are the longest and toes 1, 4 and 6 are medium sized (in leg I 10.3, 7.9, 19.7, 11.4, 20.6 and 12.4 µm long for toes 1 to 6 respectively). In the fourth pair of legs (
<figureCitation id="E1E526F95B53FFF1625B05511BB8FB86" box="[369,435,1087,1112]" captionStart="FIGURE 6" captionStartId="14.[151,250,1869,1891]" captionTargetBox="[154,1436,196,1848]" captionTargetId="figure@14.[151,1436,193,1848]" captionTargetPageId="14" captionText="FIGURE 6. Batillipes potiguarensis sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view of a paratype); C. Detail of clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg spine of the first leg (pI) of the holotype; D. Toes (t 1 – t 6) of the leg IV of a paratype; E—Gonopore (go) and anus (an) of the holotypic female. Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400159/files/figure.png" pageId="16" pageNumber="499">Fig. 6</figureCitation>
D) the toes conform to the pattern of the A group of species proposed by
<bibRefCitation id="1D4F478D5B53FFF167DF05511B50FBA2" author="Kristensen" pageId="16" pageNumber="499" refString="Kristensen, R. M. &amp; Mackness, B. S. (2000) First record of the marine Tardigrade genus Batillipes (Arthrotardigrada: Batillipedidae) from South Australia with a description of a new species. Records of the South Australian Museum, 33 (2), 73 - 87." type="journal article" year="2000">Kristensen and Mackness (2000)</bibRefCitation>
, with medial toes 3 and 4 of equal lengths (15.5 and 15.7 µm long respectively). Toes 5 and 6 of legs IV are respectively 25.7 and 19.1 µm long and toes 1 and 2 were not measurable (in
<typeStatus id="A66584DE5B53FFF1675105E71EECFB7F" box="[1147,1255,1161,1185]" pageId="16" pageNumber="499" type="paratype">paratypes</typeStatus>
toes 2 and 5 are the longest and similar each other; toes 1 and 6, also similar each other, are of intermediate length).
</paragraph>
<paragraph id="79613A7C5B53FFF163ED05A11897FAD3" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="499">Dorsal cuticle punctated (about 12 pillars/10 µm; each pillar with about 1 µm high) with about eight transverse folds. Ventral transverse folds are also visible.</paragraph>
<paragraph id="79613A7C5B53FFF163ED04791E88FA8B" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="499">
Rosette-like gonopore delimited by four punctate cuticular platelets, two anterior and two posterior forming a groove in direction to the anus (
<figureCitation id="E1E526F95B53FFF162D604521835FA8B" box="[508,574,1340,1365]" captionStart="FIGURE 6" captionStartId="14.[151,250,1869,1891]" captionTargetBox="[154,1436,196,1848]" captionTargetId="figure@14.[151,1436,193,1848]" captionTargetPageId="14" captionText="FIGURE 6. Batillipes potiguarensis sp. nov. A. Drawing (dorsal view); B. Habitus (dorsal view of a paratype); C. Detail of clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg spine of the first leg (pI) of the holotype; D. Toes (t 1 – t 6) of the leg IV of a paratype; E—Gonopore (go) and anus (an) of the holotypic female. Scale bars: A – B = 50 µm; C – E = 10 µm." httpUri="https://zenodo.org/record/400159/files/figure.png" pageId="16" pageNumber="499">Fig. 6</figureCitation>
E). The anus is 10.8 µm distant from the gonopore.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B53FFF163ED043118DAFA1E" pageId="16" pageNumber="499" type="discussion">
<paragraph id="79613A7C5B53FFF163ED043118DAFA1E" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="499">
<emphasis id="4BAAE66E5B53FFF163ED04311B30FAA6" bold="true" box="[199,315,1375,1400]" pageId="16" pageNumber="499">Remarks.</emphasis>
Males and four-toed larvae were not found. Morphometric variations in lateral processes and caudal projection of adult specimens were not observed. The only juvenile observed, with six toes on each leg but without a visible gonopore, is similar to adult females also.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B53FFF063ED04A21AD2FDC6" lastPageId="17" lastPageNumber="500" pageId="16" pageNumber="499" type="diagnosis">
<paragraph id="79613A7C5B53FFF163ED04A218ABF962" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="499">
<emphasis id="4BAAE66E5B53FFF163ED04A21BD9FA3B" bold="true" box="[199,466,1484,1509]" pageId="16" pageNumber="499">Differential diagnosis.</emphasis>
<taxonomicName id="BEDE41FF5B53FFF162F704A218E0FA3A" box="[477,747,1484,1508]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="potiguarensis" status="sp. nov.">
<emphasis id="4BAAE66E5B53FFF162F704A218E0FA3A" box="[477,747,1484,1508]" italics="true" pageId="16" pageNumber="499">Batillipes potiguarensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B53FFF161DC04A2195FFA3B" bold="true" box="[758,852,1484,1509]" pageId="16" pageNumber="499">
<taxonomicNameLabel id="50995B155B53FFF161DC04A2195FFA3B" box="[758,852,1484,1509]" pageId="16" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having toes 3 and 4 of the hind legs of equal lengths and respectively different from toe 1 and toe 2, belongs to the A group of species (
<bibRefCitation id="1D4F478D5B53FFF1675704811ADFF9F2" author="Kristensen" pageId="16" pageNumber="499" refString="Kristensen, R. M. &amp; Mackness, B. S. (2000) First record of the marine Tardigrade genus Batillipes (Arthrotardigrada: Batillipedidae) from South Australia with a description of a new species. Records of the South Australian Museum, 33 (2), 73 - 87." type="journal article" year="2000">Kristensen and Mackness 2000</bibRefCitation>
). There are 20 known species included in this group. However, similarly to the new species, only two species, both with limited geographic distribution in the Pacific Ocean, have simultaneously a blunt or semicircular caudal protrusion and blunt lateral processes between legs III and IV:
<taxonomicName id="BEDE41FF5B53FFF160A507331F93F9AA" authority="Rho, Min and Chang, 1999" authorityName="Rho, Min and Chang" authorityYear="1999" box="[911,1432,1628,1653]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="rotundiculus">
<emphasis id="4BAAE66E5B53FFF160A507331E4AF9AA" box="[911,1089,1628,1652]" italics="true" pageId="16" pageNumber="499">B. rotundiculus</emphasis>
<bibRefCitation id="1D4F478D5B53FFF1676707321F93F9AA" author="Rho" box="[1101,1432,1628,1653]" pageId="16" pageNumber="499" refString="Rho, H. S., Min, B. H. &amp; Chang, C. Y. (1999) Taxonomic study of Marine Tardigrades from Korea I. Genus Batillipes (Heterotardigrada: Batillipedidae). The Korean Journal of Systematic Zoology, 15 (1), 107 - 118." type="journal article" year="1999">Rho, Min and Chang, 1999</bibRefCitation>
</taxonomicName>
, described from the Korean coasts (Sea of
<collectingCountry id="01C97AEC5B53FFF1615A07EE18BAF946" box="[624,689,1664,1688]" name="Japan" pageId="16" pageNumber="499">Japan</collectingCountry>
) and
<taxonomicName id="BEDE41FF5B53FFF161DB07EF1AD9F962" authority="Jorgensen, Boesgaard, Mobjerg and Kristensen, 2014" authorityName="Jorgensen, Boesgaard, Mobjerg and Kristensen" authorityYear="2014" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="solitarius">
<emphasis id="4BAAE66E5B53FFF161DB07EF1970F946" box="[753,891,1664,1688]" italics="true" pageId="16" pageNumber="499">B. solitarius</emphasis>
<bibRefCitation id="1D4F478D5B53FFF160A907EF1AD9F962" author="Jorgensen" pageId="16" pageNumber="499" refString="Jorgensen, A., Boesgaard, T. M., Mobjerg, N. &amp; Kristensen, R. M. (2014) The tardigrade fauna of Australian marine caves: with descriptions of nine new species of Arthrotardigrada. Zootaxa, 3802 (4), 401 - 443." type="journal article" year="2014">Jørgensen, Boesgaard, Møbjerg and Kristensen, 2014</bibRefCitation>
</taxonomicName>
, from the Australian coast (Tasman Sea).
</paragraph>
<paragraph id="79613A7C5B53FFF163ED07A71E80F8AE" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="499">
<taxonomicName id="BEDE41FF5B53FFF163ED07A71B8AF901" box="[199,385,1736,1760]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="potiguarensis" status="sp. nov.">
<emphasis id="4BAAE66E5B53FFF163ED07A71B8AF901" box="[199,385,1736,1760]" italics="true" pageId="16" pageNumber="499">B. potiguarensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B53FFF162A707A91BE0F93E" bold="true" box="[397,491,1735,1760]" pageId="16" pageNumber="499">
<taxonomicNameLabel id="50995B155B53FFF162A707A91BE0F93E" box="[397,491,1735,1760]" pageId="16" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="BEDE41FF5B53FFF161A607A71911F93E" box="[652,794,1736,1760]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="solitarius">
<emphasis id="4BAAE66E5B53FFF161A607A71911F93E" box="[652,794,1736,1760]" italics="true" pageId="16" pageNumber="499">B. solitarius</emphasis>
</taxonomicName>
in having a much more protruded semicircular caudal projection; smaller and different shaped secondary clavae (prominent papilla, less than 6.2 µm wide in the new species, concave and elongate lens-shaped, 13.0 µm wide, in
<taxonomicName id="BEDE41FF5B53FFF1606B067F19C1F8F6" box="[833,970,1808,1832]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="solitarius">
<emphasis id="4BAAE66E5B53FFF1606B067F19C1F8F6" box="[833,970,1808,1832]" italics="true" pageId="16" pageNumber="499">B. solitarius</emphasis>
</taxonomicName>
) and lateral projection between legs I and II (blunt in the new species; sharp pointed in
<taxonomicName id="BEDE41FF5B53FFF16188065B1926F892" box="[674,813,1844,1868]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="solitarius">
<emphasis id="4BAAE66E5B53FFF16188065B1926F892" box="[674,813,1844,1868]" italics="true" pageId="16" pageNumber="499">B. solitarius</emphasis>
</taxonomicName>
). Moreover, cephalic cirri in
<taxonomicName id="BEDE41FF5B53FFF16755065B1F3CF892" box="[1151,1335,1845,1868]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="potiguarensis" status="sp. nov.">
<emphasis id="4BAAE66E5B53FFF16755065B1F3CF892" box="[1151,1335,1845,1868]" italics="true" pageId="16" pageNumber="499">B. potiguarensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B53FFF1666B065A1F97F893" bold="true" box="[1345,1436,1844,1869]" pageId="16" pageNumber="499">
<taxonomicNameLabel id="50995B155B53FFF1666B065A1F97F893" box="[1345,1436,1844,1869]" pageId="16" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have a plain tip, while in
<taxonomicName id="BEDE41FF5B53FFF1629906371837F8AE" box="[435,572,1880,1904]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="solitarius">
<emphasis id="4BAAE66E5B53FFF1629906371837F8AE" box="[435,572,1880,1904]" italics="true" pageId="16" pageNumber="499">B. solitarius</emphasis>
</taxonomicName>
median, internal and lateral cirri have terminal tufts.
</paragraph>
<paragraph id="79613A7C5B53FFF063ED06121AD2FDC6" blockId="16.[151,1437,151,2013]" lastBlockId="17.[151,1437,151,536]" lastPageId="17" lastPageNumber="500" pageId="16" pageNumber="499">
The new species is very similar to
<emphasis id="4BAAE66E5B53FFF16163061318F6F84A" box="[585,765,1916,1940]" italics="true" pageId="16" pageNumber="499">
<taxonomicName id="BEDE41FF5B53FFF16163061318F3F84A" box="[585,760,1916,1940]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="rotundiculus">B. rotundiculus</taxonomicName>
.
</emphasis>
These two species share a similar caudal protrusion, a welldeveloped scapular region at the level of the first pair of legs, and similar blunt-shaped ventrolateral processes. However
<taxonomicName id="BEDE41FF5B53FFF1622D06AA1819F802" box="[263,530,1988,2012]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="potiguarensis" status="sp. nov.">
<emphasis id="4BAAE66E5B53FFF1622D06AA1819F802" box="[263,530,1988,2012]" italics="true" pageId="16" pageNumber="499">Batillipes potiguarensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B53FFF1613606AA1873F803" bold="true" box="[540,632,1988,2013]" pageId="16" pageNumber="499">
<taxonomicNameLabel id="50995B155B53FFF1613606AA1873F803" box="[540,632,1988,2013]" pageId="16" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="BEDE41FF5B53FFF1609906AB1E6AF802" box="[947,1121,1988,2012]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="16" pageNumber="499" phylum="Tardigrada" rank="species" species="rotundiculus">
<emphasis id="4BAAE66E5B53FFF1609906AB1E6AF802" box="[947,1121,1988,2012]" italics="true" pageId="16" pageNumber="499">B. rotundiculus</emphasis>
</taxonomicName>
in having swollen cephalic cirri (sharply pointed in
<taxonomicName id="BEDE41FF5B52FFF062EA01F71879FF6E" box="[448,626,152,176]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="17" pageNumber="500" phylum="Tardigrada" rank="species" species="rotundiculus">
<emphasis id="4BAAE66E5B52FFF062EA01F71879FF6E" box="[448,626,152,176]" italics="true" pageId="17" pageNumber="500">B. rotundiculus</emphasis>
</taxonomicName>
) and an evident and protruded secondary clavae (not present in
<taxonomicName id="BEDE41FF5B52FFF066AE01F71B28FF0A" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="17" pageNumber="500" phylum="Tardigrada" rank="species" species="rotundiculus">
<emphasis id="4BAAE66E5B52FFF066AE01F71B28FF0A" italics="true" pageId="17" pageNumber="500">B. rotundiculus</emphasis>
</taxonomicName>
). In addition, the two species also differ in the relative length of leg sense organs: contrary to
<taxonomicName id="BEDE41FF5B52FFF066AE01D31B28FF26" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="17" pageNumber="500" phylum="Tardigrada" rank="species" species="rotundiculus">
<emphasis id="4BAAE66E5B52FFF066AE01D31B28FF26" italics="true" pageId="17" pageNumber="500">B. rotundiculus</emphasis>
</taxonomicName>
, in the new species sense organs on leg IV are particularly shorter and shorter than sense organs on leg I (in the
<typeStatus id="A66584DE5B52FFF063DF006A1B5CFEC3" box="[245,343,260,285]" pageId="17" pageNumber="500" type="holotype">holotype</typeStatus>
of
<taxonomicName id="BEDE41FF5B52FFF06256006B1823FEC2" box="[380,552,260,284]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="17" pageNumber="500" phylum="Tardigrada" rank="species" species="rotundiculus">
<emphasis id="4BAAE66E5B52FFF06256006B1823FEC2" box="[380,552,260,284]" italics="true" pageId="17" pageNumber="500">B. rotundiculus</emphasis>
</taxonomicName>
leg spine I is 8 µm and sense organ on leg IV is 9.2 µm; in
<taxonomicName id="BEDE41FF5B52FFF06794006B1F78FEC2" box="[1214,1395,261,284]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="17" pageNumber="500" phylum="Tardigrada" rank="species" species="potiguarensis" status="sp. nov.">
<emphasis id="4BAAE66E5B52FFF06794006B1F78FEC2" box="[1214,1395,261,284]" italics="true" pageId="17" pageNumber="500">B. potiguarensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B52FFF06650006A1AC3FE9E" bold="true" pageId="17" pageNumber="500">
<taxonomicNameLabel id="50995B155B52FFF06650006A1AC3FE9E" pageId="17" pageNumber="500" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
leg spine I is 9.8 µm and sense organ IV is 6.6 µm, specimens respectively 195 and 187 µm long, including caudal apparatus). Moreover, the leg IV sensory organs of
<taxonomicName id="BEDE41FF5B52FFF06065002319F4FEBA" box="[847,1023,332,356]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="17" pageNumber="500" phylum="Tardigrada" rank="species" species="rotundiculus">
<emphasis id="4BAAE66E5B52FFF06065002319F4FEBA" box="[847,1023,332,356]" italics="true" pageId="17" pageNumber="500">B. rotundiculus</emphasis>
</taxonomicName>
are spike-shaped while in the new species they are divided into two portions, the distal portion with swollen tips. Another important differentiating character between the two species is the relative size of medial (toes 3 and 4) and lateral toes (1 and 6) on legs IV. In
<taxonomicName id="BEDE41FF5B52FFF0639F00D71B60FE11" box="[181,363,440,464]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="17" pageNumber="500" phylum="Tardigrada" rank="species" species="potiguarensis" status="sp. nov.">
<emphasis id="4BAAE66E5B52FFF0639F00D71B60FE11" box="[181,363,440,464]" italics="true" pageId="17" pageNumber="500">B. potiguarensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B52FFF0625900D91BC7FE0E" bold="true" box="[371,460,439,464]" pageId="17" pageNumber="500">
<taxonomicNameLabel id="50995B155B52FFF0625900D91BC7FE0E" box="[371,460,439,464]" pageId="17" pageNumber="500" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
toes 3 and 4 of hind legs are shorter than toes 1 and 6 while in
<taxonomicName id="BEDE41FF5B52FFF067BA00D71F37FE0E" box="[1168,1340,440,464]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="17" pageNumber="500" phylum="Tardigrada" rank="species" species="rotundiculus">
<emphasis id="4BAAE66E5B52FFF067BA00D71F37FE0E" box="[1168,1340,440,464]" italics="true" pageId="17" pageNumber="500">B. rotundiculus</emphasis>
</taxonomicName>
they are similar or longer (in the
<typeStatus id="A66584DE5B52FFF0629F00B2181CFE2B" box="[437,535,476,501]" pageId="17" pageNumber="500" type="holotype">holotype</typeStatus>
lateral toes are 16.6 and 17.1 µm long and medial toes are 17.5 and 17.1 µm long).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="1BA47EF9490D7D721ABB143049F5E270">Three new Batillipes species (Arthrotardigrada: Batillipedidae) from the Brazilian coast</mods:title>
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One specimen in a dirty preparation and not identified to species level, collected in medium to coarse, gravels and quartz sands, in a low energy beach, Sossego Beach (
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<emphasis id="4BAAE66E5B52FFF063BD05F31B72FB69" bold="true" box="[151,377,1181,1207]" italics="true" pageId="17" pageNumber="500">Orzeliscus belopus</emphasis>
du
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Four specimens collected in medium to coarse, gravels and quartz sands, in low energy beaches, one specimen from Forte Orange Beach (
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<emphasis id="4BAAC442FFE0FF86FE39FB51FD78948C" bold="true" box="[439,671,1179,1203]" italics="true" pageId="3" pageNumber="638">Amphisternus alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50997939FFE0FF86FD28FB56FD1C948C" box="[678,763,1180,1203]" pageId="3" pageNumber="638" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="31C44BDBFFE0FF86FD9CFB70FD7894F2" box="[530,671,1210,1229]" pageId="3" pageNumber="638" type="description">
<paragraph id="79611850FFE0FF86FD9CFB70FD7894F2" blockId="3.[113,1089,1210,1351]" box="[530,671,1210,1229]" pageId="3" pageNumber="638">
(
<figureCitation id="E1E504D5FFE0FF86FD96FB70FD8394F2" box="[536,612,1210,1229]" captionStart="Figs 1–8" captionStartId="4.[151,188,1506,1525]" captionTargetBox="[176,1112,224,1445]" captionTargetId="figure-10@4.[151,1126,224,1456]" captionTargetPageId="4" captionText="Figs 1–8. Habitus. 1–5 – Amphisternus alberti sp. nov.; 6–8 – Gerstaeckerus rufiplagiatus sp. nov. 1, 6 – dorsal habitus; 2 – lateral habitus; 3, 8 – pronotum; 4, 7 – antenna; 5 – habitus, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5299675" httpUri="https://zenodo.org/record/5299675/files/figure.png" pageId="3" pageNumber="638">Figs 1–5</figureCitation>
,
<figureCitation id="E1E504D5FFE0FF86FDE0FB70FD7E94F2" box="[622,665,1210,1229]" captionStart="Figs 9–14" captionStartId="5.[113,150,1528,1547]" captionTargetBox="[195,1035,433,1509]" captionTargetId="figure-10@5.[144,1056,430,1512]" captionTargetPageId="5" captionText="Figs 9–14. Genitalia. 9–13 – Amphisternus alberti sp. nov.; 14 – Gerstaeckerus rufiplagiatus, sp. nov. 9 – aedeagus, dorsal view; 10 – aedeagus, ventral view; 11 – abdominal segment VIII, male, ventral view; 12 – male genital segment, ventral view; 13, 14 – female genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5299677" httpUri="https://zenodo.org/record/5299677/files/figure.png" pageId="3" pageNumber="638">9–13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="31C44BDBFFE0FF86FFFFFB2EFEFD9578" pageId="3" pageNumber="638" type="materials_examined">
<paragraph id="79611850FFE0FF86FFFFFB2EFEFD9578" blockId="3.[113,1089,1210,1351]" pageId="3" pageNumber="638">
<emphasis id="4BAAC442FFE0FF86FFFFFB2EFF1C94C8" bold="true" box="[113,251,1252,1271]" pageId="3" pageNumber="638">Type material.</emphasis>
<materialsCitation id="C9B6120DFFE0FF86FE8DFB2EFBA794C8" box="[259,1088,1246,1271]" collectingDate="2011-02" collectorName="I. Lumawig" country="Philippines" location="Zamboanga del Norte" pageId="3" pageNumber="638" specimenCount="1" stateProvince="Zamboanga del Norte" typeStatus="holotype">
<typeStatus id="A665A6F2FFE0FF86FE8DFB2EFEB094CA" box="[259,343,1252,1271]" pageId="3" pageNumber="638" type="holotype">HOLOTYPE</typeStatus>
: ♁, ‘
<collectingCountry id="01C958C0FFE0FF86FE08FB2EFDEE94C8" box="[390,521,1252,1271]" name="Philippines" pageId="3" pageNumber="638">PHILIPPINES</collectingCountry>
/ MINDANAO/
<collectingRegion id="BB1AD6B2FFE0FF86FD0EFB2EFCA594C8" box="[640,834,1252,1271]" country="Philippines" name="Zamboanga del Norte" pageId="3" pageNumber="638">Zamboanga del Norte</collectingRegion>
/
<date id="0D603E90FFE0FF86FCDEFB2EFC7194C8" box="[848,918,1252,1271]" pageId="3" pageNumber="638" value="2011-02">
<collectingDate id="1D24C778FFE0FF86FCDEFB2EFC7194C8" box="[848,918,1252,1271]" pageId="3" pageNumber="638" value="2011-02">II. 2011</collectingDate>
</date>
/ leg.
<collectorName id="D42B7D86FFE0FF86FC43FB2EFBD494C8" box="[973,1075,1252,1271]" pageId="3" pageNumber="638">I. Lumawig</collectorName>
//
</materialsCitation>
<materialsCitation id="C9B6120DFFE0FF86FFFFFB35FD0C952D" box="[113,747,1279,1298]" collectionCode="MZPW" country="Philippines" pageId="3" pageNumber="638" specimenCount="1" stateProvince="Zamboanga del Norte" typeStatus="holotype">
<typeStatus id="A665A6F2FFE0FF86FFFFFB35FF25952D" box="[113,194,1279,1298]" pageId="3" pageNumber="638" type="holotype">Holotype</typeStatus>
<taxonomicName id="BEDE63D3FFE0FF86FF44FB35FE63952D" authority="Tomaszewska, 2014" authorityName="Tomaszewska" authorityYear="2014" box="[202,388,1279,1298]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
<emphasis id="4BAAC442FFE0FF86FF44FB35FE63952D" box="[202,388,1279,1298]" italics="true" pageId="3" pageNumber="638">Amphisternus alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50997939FFE0FF86FE02FB35FE33952D" box="[396,468,1279,1298]" pageId="3" pageNumber="638" rank="species">sp. nov.</taxonomicNameLabel>
, Tomaszewska 2014’ (
<collectionCode id="1FCF8095FFE0FF86FD11FB35FD02952D" box="[671,741,1279,1298]" country="Poland" httpUri="http://grbio.org/cool/ynr4-t34e" name="Polish Academy of Science, Museum of the Institute of Zoology" pageId="3" pageNumber="638">MZPW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C9B6120DFFE0FF86FD78FB35FF249513" collectionCode="MZPW" country="Philippines" pageId="3" pageNumber="638" specimenCount="1" specimenCount-female="1" stateProvince="Zamboanga del Norte" typeStatus="paratype">
<typeStatus id="A665A6F2FFE0FF86FD78FB35FCA8952F" box="[758,847,1279,1298]" pageId="3" pageNumber="638" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="6FD8D3D9FFE0FF86FCD3FB31FC8A952D" box="[861,877,1275,1298]" pageId="3" pageNumber="638" type="female">♀</specimenCount>
, same data as holotype (
<collectionCode id="1FCF8095FFE0FF86FFF9FAD3FF5A9513" box="[119,189,1305,1324]" country="Poland" httpUri="http://grbio.org/cool/ynr4-t34e" name="Polish Academy of Science, Museum of the Institute of Zoology" pageId="3" pageNumber="638">MZPW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9B6120DFFE0FF86FF40FADCFE169513" box="[206,497,1302,1325]" collectingDate="2010-12" collectionCode="AABI" country="Philippines" pageId="3" pageNumber="638" specimenCount="1" specimenCount-female="1" stateProvince="Zamboanga del Norte" typeStatus="paratype">
<specimenCount id="6FD8D3D9FFE0FF86FF40FADCFF399512" box="[206,222,1302,1325]" pageId="3" pageNumber="638" type="female">♀</specimenCount>
, same but ‘
<date id="0D603E90FFE0FF86FECAFAD3FE469513" box="[324,417,1305,1324]" pageId="3" pageNumber="638" value="2010-12">
<collectingDate id="1D24C778FFE0FF86FECAFAD3FE469513" box="[324,417,1305,1324]" pageId="3" pageNumber="638" value="2010-12">XII. 2010</collectingDate>
</date>
’ (
<collectionCode id="1FCF8095FFE0FF86FE3FFAD3FE0C9513" box="[433,491,1305,1324]" pageId="3" pageNumber="638">AABI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9B6120DFFE0FF86FE70FADCFEF19578" collectingDate="2013-10" collectionCode="AABI" collectorName="I. Lumawig'" country="Philippines" location="Misamis Oriental" pageId="3" pageNumber="638" specimenCount="1" specimenCount-female="1" stateProvince="Misamis Oriental" typeStatus="paratype">
<specimenCount id="6FD8D3D9FFE0FF86FE70FADCFDEB9512" box="[510,524,1302,1325]" pageId="3" pageNumber="638" type="female">♀</specimenCount>
, ‘
<collectingCountry id="01C958C0FFE0FF86FD95FAD3FD469513" box="[539,673,1305,1324]" name="Philippines" pageId="3" pageNumber="638">PHILIPPINES</collectingCountry>
, MINDANAO,
<collectingRegion id="BB1AD6B2FFE0FF86FC9DFAD3FC489513" box="[787,943,1305,1324]" country="Philippines" name="Misamis Oriental" pageId="3" pageNumber="638">Misamis Oriental</collectingRegion>
,
<date id="0D603E90FFE0FF86FC37FAD3FBE49513" box="[953,1027,1305,1324]" pageId="3" pageNumber="638" value="2013-10">
<collectingDate id="1D24C778FFE0FF86FC37FAD3FBE49513" box="[953,1027,1305,1324]" pageId="3" pageNumber="638" value="2013-10">X. 2013</collectingDate>
</date>
, leg.
<collectorName id="D42B7D86FFE0FF86FBBAFAD3FF2D9578" pageId="3" pageNumber="638">I. Lumawig’</collectorName>
(
<collectionCode id="1FCF8095FFE0FF86FF58FAFEFEF79578" box="[214,272,1332,1351]" pageId="3" pageNumber="638">AABI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="31C44BDBFFE0FF83FFFFFAAAFED7914C" lastPageId="6" lastPageNumber="641" pageId="3" pageNumber="638" type="description">
<paragraph id="79611850FFE0FF83FFFFFAAAFE56910C" blockId="3.[113,1089,1376,1559]" lastBlockId="6.[151,1127,220,563]" lastPageId="6" lastPageNumber="641" pageId="3" pageNumber="638">
<emphasis id="4BAAC442FFE0FF86FFFFFAAAFEE49548" bold="true" box="[113,259,1376,1399]" pageId="3" pageNumber="638">Description.</emphasis>
Length
<quantity id="BE26B5B5FFE0FF86FEE9FAAAFE139548" box="[359,500,1376,1399]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" metricValueMax="7.8" metricValueMin="6.8" pageId="3" pageNumber="638" unit="mm" value="7.3" valueMax="7.8" valueMin="6.8">6.8–7.8 mm</quantity>
. Body 1.70–1.83 times as long as wide; pronotum 0.50–0.60 times as long as wide; elytra 1.15–1.25 times as long as wide; 2.75–2.90 times longer than pronotum, 1.23–1.32 times wider than pronotum. Body (
<figureCitation id="E1E504D5FFE0FF86FCD1FA6AFC3D9588" box="[863,986,1440,1463]" captionStart="Figs 1–8" captionStartId="4.[151,188,1506,1525]" captionTargetBox="[176,1112,224,1445]" captionTargetId="figure-10@4.[151,1126,224,1456]" captionTargetPageId="4" captionText="Figs 1–8. Habitus. 1–5 – Amphisternus alberti sp. nov.; 6–8 – Gerstaeckerus rufiplagiatus sp. nov. 1, 6 – dorsal habitus; 2 – lateral habitus; 3, 8 – pronotum; 4, 7 – antenna; 5 – habitus, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5299675" httpUri="https://zenodo.org/record/5299675/files/figure.png" pageId="3" pageNumber="638">Figs 1, 2, 5</figureCitation>
) short-oval, convex, opaque; colour black with only elytral tubercles red-orange. Antenna (
<figureCitation id="E1E504D5FFE0FF86FB9AFA0AFF9895C8" captionStart="Figs 1–8" captionStartId="4.[151,188,1506,1525]" captionTargetBox="[176,1112,224,1445]" captionTargetId="figure-10@4.[151,1126,224,1456]" captionTargetPageId="4" captionText="Figs 1–8. Habitus. 1–5 – Amphisternus alberti sp. nov.; 6–8 – Gerstaeckerus rufiplagiatus sp. nov. 1, 6 – dorsal habitus; 2 – lateral habitus; 3, 8 – pronotum; 4, 7 – antenna; 5 – habitus, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5299675" httpUri="https://zenodo.org/record/5299675/files/figure.png" pageId="3" pageNumber="638">Fig. 4</figureCitation>
) with antennomere 3 about 1.3 times longer than antennomere 4 or 5. Pronotum (
<figureCitation id="E1E504D5FFE0FF86FC79FA2AFBD095C8" box="[1015,1079,1504,1527]" captionStart="Figs 1–8" captionStartId="4.[151,188,1506,1525]" captionTargetBox="[176,1112,224,1445]" captionTargetId="figure-10@4.[151,1126,224,1456]" captionTargetPageId="4" captionText="Figs 1–8. Habitus. 1–5 – Amphisternus alberti sp. nov.; 6–8 – Gerstaeckerus rufiplagiatus sp. nov. 1, 6 – dorsal habitus; 2 – lateral habitus; 3, 8 – pronotum; 4, 7 – antenna; 5 – habitus, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5299675" httpUri="https://zenodo.org/record/5299675/files/figure.png" pageId="3" pageNumber="638">Fig. 3</figureCitation>
) impunctate, weakly reticulated; pronotal disk with two pairs of weak convexities/tubercles; anterior angles blunt, posterior angles weakly acute. Elytra weakly wider than base of pronotum; shoulders obliquely widened and carinate; humeral carina with prominent red-orange tubercle; each elytron with basal moderately high, round-oval, red-orange tubercle, a pair of similar tubercles in apical third and a large, carinate black tubercle, somewhat rectangular in shape (in lateral view) a little in front of mid length (
<figureCitation id="E1E504D5FFE6FF80FC62FE96FBD5914C" box="[1004,1074,348,371]" captionStart="Figs 1–8" captionStartId="4.[151,188,1506,1525]" captionTargetBox="[176,1112,224,1445]" captionTargetId="figure-10@4.[151,1126,224,1456]" captionTargetPageId="4" captionText="Figs 1–8. Habitus. 1–5 – Amphisternus alberti sp. nov.; 6–8 – Gerstaeckerus rufiplagiatus sp. nov. 1, 6 – dorsal habitus; 2 – lateral habitus; 3, 8 – pronotum; 4, 7 – antenna; 5 – habitus, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5299675" httpUri="https://zenodo.org/record/5299675/files/figure.png" pageId="5" pageNumber="640">Fig. 2</figureCitation>
); elytral punctures coarse and almost touching one another. Hind wings absent. Male fore tibia with small tooth in apical third and male fore trochanters with small triangular tubercle on outer edge. Aedeagus (
<figureCitation id="E1E504D5FFE5FF83FE78FF36FD85912C" box="[502,610,252,275]" captionStart="Figs 9–14" captionStartId="5.[113,150,1528,1547]" captionTargetBox="[195,1035,433,1509]" captionTargetId="figure-10@5.[144,1056,430,1512]" captionTargetPageId="5" captionText="Figs 9–14. Genitalia. 9–13 – Amphisternus alberti sp. nov.; 14 – Gerstaeckerus rufiplagiatus, sp. nov. 9 – aedeagus, dorsal view; 10 – aedeagus, ventral view; 11 – abdominal segment VIII, male, ventral view; 12 – male genital segment, ventral view; 13, 14 – female genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5299677" httpUri="https://zenodo.org/record/5299677/files/figure.png" pageId="6" pageNumber="641">Figs 9, 10</figureCitation>
) short and stout; median lobe with inner margin denticulate in mid length.
</paragraph>
<caption id="2DA148D8FFE7FF81FF19FA28FC9E962F" ID-DOI="http://doi.org/10.5281/zenodo.5299675" ID-Zenodo-Dep="5299675" httpUri="https://zenodo.org/record/5299675/files/figure.png" pageId="4" pageNumber="639" startId="4.[151,188,1506,1525]" targetBox="[176,1112,224,1445]" targetPageId="4">
<paragraph id="79611850FFE7FF81FF19FA28FC9E962F" blockId="4.[151,1126,1506,1552]" pageId="4" pageNumber="639">
Figs 1–8. Habitus. 1–5 –
<taxonomicName id="BEDE63D3FFE7FF81FEFBFA28FDCD95CA" authority="Tomaszewska, 2014" authorityName="Tomaszewska" authorityYear="2014" box="[373,554,1506,1525]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="639" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
<emphasis id="4BAAC442FFE7FF81FEFBFA28FDCD95CA" box="[373,554,1506,1525]" italics="true" pageId="4" pageNumber="639">Amphisternus alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50997939FFE7FF81FDBFFA28FD9F95CA" box="[561,632,1506,1525]" pageId="4" pageNumber="639" rank="species">sp. nov.</taxonomicNameLabel>
; 6–8 –
<taxonomicName id="BEDE63D3FFE7FF81FD37FA28FC4F95CA" authority="Tomaszewska, 2014" authorityName="Tomaszewska" authorityYear="2014" box="[697,936,1506,1525]" class="Insecta" family="Endomychidae" genus="Gerstaeckerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="639" phylum="Arthropoda" rank="species" species="rufiplagiatus" status="sp. nov.">
<emphasis id="4BAAC442FFE7FF81FD37FA28FC4F95CA" box="[697,936,1506,1525]" italics="true" pageId="4" pageNumber="639">Gerstaeckerus rufiplagiatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50997939FFE7FF81FC21FA28FC1495CA" box="[943,1011,1506,1525]" pageId="4" pageNumber="639" rank="species">sp. nov.</taxonomicNameLabel>
1, 6 – dorsal habitus; 2 – lateral habitus; 3, 8 – pronotum; 4, 7 – antenna; 5 – habitus, anterior view.
</paragraph>
</caption>
<caption id="2DA148D8FFE6FF80FFFFFA32FD8D967E" ID-DOI="http://doi.org/10.5281/zenodo.5299677" ID-Zenodo-Dep="5299677" httpUri="https://zenodo.org/record/5299677/files/figure.png" pageId="5" pageNumber="640" startId="5.[113,150,1528,1547]" targetBox="[195,1035,433,1509]" targetPageId="5">
<paragraph id="79611850FFE6FF80FFFFFA32FD8D967E" blockId="5.[113,1089,1528,1601]" pageId="5" pageNumber="640">
Figs 9–14. Genitalia. 9–13 –
<taxonomicName id="BEDE63D3FFE6FF80FEE8FA32FDFE9634" authority="Tomaszewska, 2014" authorityName="Tomaszewska" authorityYear="2014" box="[358,537,1528,1547]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="640" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
<emphasis id="4BAAC442FFE6FF80FEE8FA32FDFE9634" box="[358,537,1528,1547]" italics="true" pageId="5" pageNumber="640">Amphisternus alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50997939FFE6FF80FD93FA32FD859634" box="[541,610,1528,1547]" pageId="5" pageNumber="640" rank="species">sp. nov.</taxonomicNameLabel>
; 14 –
<taxonomicName id="BEDE63D3FFE6FF80FD1CFA32FC989634" authority="Tomaszewska, 2014" authorityName="Tomaszewska" authorityYear="2014" box="[658,895,1528,1547]" class="Insecta" family="Endomychidae" genus="Gerstaeckerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="640" phylum="Arthropoda" rank="species" species="rufiplagiatus" status="sp. nov.">
<emphasis id="4BAAC442FFE6FF80FD1CFA32FC989634" box="[658,895,1528,1547]" italics="true" pageId="5" pageNumber="640">Gerstaeckerus rufiplagiatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="50997939FFE6FF80FC07FA32FC2C9634" box="[905,971,1528,1547]" pageId="5" pageNumber="640" rank="species">sp. nov.</taxonomicNameLabel>
9 – aedeagus, dorsal view; 10 – aedeagus, ventral view; 11 – abdominal segment VIII, male, ventral view; 12 – male genital segment, ventral view; 13, 14 – female genitalia, ventral view.
</paragraph>
</caption>
<paragraph id="79611850FFE5FF83FF3BFEF6FED7914C" blockId="6.[151,1127,220,563]" pageId="6" pageNumber="641">
Female genitalia (
<figureCitation id="E1E504D5FFE5FF83FEFAFEF6FE24916C" box="[372,451,316,339]" captionStart="Figs 9–14" captionStartId="5.[113,150,1528,1547]" captionTargetBox="[195,1035,433,1509]" captionTargetId="figure-10@5.[144,1056,430,1512]" captionTargetPageId="5" captionText="Figs 9–14. Genitalia. 9–13 – Amphisternus alberti sp. nov.; 14 – Gerstaeckerus rufiplagiatus, sp. nov. 9 – aedeagus, dorsal view; 10 – aedeagus, ventral view; 11 – abdominal segment VIII, male, ventral view; 12 – male genital segment, ventral view; 13, 14 – female genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5299677" httpUri="https://zenodo.org/record/5299677/files/figure.png" pageId="6" pageNumber="641">Fig. 13</figureCitation>
). Ovipositor with styli distinct, subterminal; spermatheca large, elongate-oval.
</paragraph>
</subSubSection>
<subSubSection id="31C44BDBFFE5FF83FF19FEB6FBD191EC" pageId="6" pageNumber="641" type="diagnosis">
<paragraph id="79611850FFE5FF83FF19FEB6FBD191EC" blockId="6.[151,1127,220,563]" pageId="6" pageNumber="641">
<emphasis id="4BAAC442FFE5FF83FF19FEB6FE7691AC" bold="true" box="[151,401,380,403]" pageId="6" pageNumber="641">Differential diagnosis.</emphasis>
This species is most similar to
<taxonomicName id="BEDE63D3FFE5FF83FD55FEB6FCDF91AC" authorityName="Strohecker" authorityYear="1964" box="[731,824,380,403]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="641" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="4BAAC442FFE5FF83FD55FEB6FCDF91AC" box="[731,824,380,403]" italics="true" pageId="6" pageNumber="641">A. nanus</emphasis>
</taxonomicName>
, but can be distinguished by larger body, which is deeply black and opaque, humeral carina less oblique and with large rounded, red-orange tubercle, and discal tubercle larger and rectangular in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="31C44BDBFFE5FF83FF19FE16FDC7922C" pageId="6" pageNumber="641" type="etymology">
<paragraph id="79611850FFE5FF83FF19FE16FDC7922C" blockId="6.[151,1127,220,563]" pageId="6" pageNumber="641">
<emphasis id="4BAAC442FFE5FF83FF19FE16FEFF91CC" bold="true" box="[151,280,476,499]" pageId="6" pageNumber="641">Etymology.</emphasis>
The new species is dedicated to Mr. Albert Allen, who kindly sent me the specimens of this new species for study.
</paragraph>
</subSubSection>
<subSubSection id="31C44BDBFFE5FF83FF19FDD6FDD3920C" box="[151,564,540,563]" pageId="6" pageNumber="641" type="distribution">
<paragraph id="79611850FFE5FF83FF19FDD6FDD3920C" blockId="6.[151,1127,220,563]" box="[151,564,540,563]" pageId="6" pageNumber="641">
<emphasis id="4BAAC442FFE5FF83FF19FDD6FECD920C" bold="true" box="[151,298,540,563]" pageId="6" pageNumber="641">Distribution.</emphasis>
<collectingCountry id="01C958C0FFE5FF83FEBEFDD6FE4F920C" box="[304,424,540,563]" name="Philippines" pageId="6" pageNumber="641">Philippines</collectingCountry>
(Mindanao).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="5C9B301E4F2647CF5F2AC1F9928304A7">New species of the Amphisternus-group of Lycoperdininae from Philippines (Coleoptera: Endomychidae)</mods:title>
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Key to the species of
<taxonomicName id="BEDE63D3FFE1FF87FD27FC4AFCA493A7" authorityName=": TOMASZEWSKA" authorityYear="2005" box="[681,835,896,920]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="637" phylum="Arthropoda" rank="genus">
<emphasis id="4BAAC442FFE1FF87FD27FC4AFCA493A7" bold="true" box="[681,835,896,920]" italics="true" pageId="2" pageNumber="637">Amphisternus</emphasis>
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(based on
<bibRefCitation id="1D4F65A1FFE1FF87FE67FC55FD6B938D" author="STROHECKER H. F." box="[489,652,927,946]" pageId="2" pageNumber="637" pagination="319 - 357" refId="ref4179" refString="STROHECKER H. F. 1964: A synopsis of the Amphisternini (Coleoptera: Endomychidae). Pacific Insects 6: 319 - 357." type="journal article" year="1964">STROHECKER (1964)</bibRefCitation>
and
<bibRefCitation id="1D4F65A1FFE1FF87FD3BFC55FC8A938D" author="WANG J. - L. &amp; REN G. - D." box="[693,877,927,946]" pageId="2" pageNumber="637" pagination="459 - 461" refId="ref4582" refString="WANG J. - L. &amp; REN G. - D. 2007: A new record of genus and new species of Endomychidae (Coleoptera, Cucujoidea) from China. Acta Zootaxonomica Sinica 32: 459 - 461." type="journal article" year="2007">WANG &amp; REN (2007))</bibRefCitation>
</paragraph>
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<paragraph id="79611850FFE1FF87FF19FC01FB81943D" blockId="2.[151,1127,971,1570]" pageId="2" pageNumber="637">
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1 Dorsum covered with short, dense pubescence; body length
<quantity id="BE26B5B5FFE1FF87FCB1FC01FC7393DD" box="[831,916,971,994]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="2" pageNumber="637" unit="mm" value="8.7">8.7 mm</quantity>
;
<collectingCountry id="01C958C0FFE1FF87FC11FC01FC0693DD" box="[927,993,971,994]" name="China" pageId="2" pageNumber="637">China</collectingCountry>
,
<collectingRegion id="BB1AD6B2FFE1FF87FC65FC01FBA793DD" box="[1003,1088,971,994]" country="China" name="Yunnan" pageId="2" pageNumber="637">Yunnan</collectingRegion>
. .... ....................................................................................
<taxonomicName id="BEDE63D3FFE1FF87FD74FC20FB81943D" authority="Chang &amp; Ren, 2013" authorityName="Chang &amp; Ren" authorityYear="2013" box="[762,1126,1002,1026]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="637" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="4BAAC442FFE1FF87FD74FC20FC6D943D" bold="true" box="[762,906,1002,1026]" italics="true" pageId="2" pageNumber="637">A. pubescens</emphasis>
<bibRefCitation id="1D4F65A1FFE1FF87FC1FFC21FB81943D" author="CHANG L. - X. &amp; REN G. - D." box="[913,1126,1003,1026]" pageId="2" pageNumber="637" pagination="95 - 100" refId="ref3839" refString="CHANG L. - X. &amp; REN G. - D. 2013: Humerus medogicus, new genus and new species from Xizang, China (Coleoptera: Endomychidae). Annales Zoologici 63: 95 - 100." type="journal article" year="2013">Chang &amp; Ren, 2013</bibRefCitation>
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<keyLead id="C22FBB65FFE1FF87FF19FBC1FB81941D" box="[151,1126,1035,1058]" pageId="2" pageNumber="637">– Dorsum without pubescence. ......................................................................................... 2</keyLead>
</paragraph>
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<keyLead id="C22FBB65FFE1FF87FF19FBE1FB81947D" box="[151,1126,1067,1090]" pageId="2" pageNumber="637">2 Elytral umbo without spine. ........................................................................................... 3</keyLead>
</paragraph>
<paragraph id="79611850FFE1FF87FF19FB81FB81945D" blockId="2.[151,1127,971,1570]" box="[151,1126,1099,1122]" pageId="2" pageNumber="637">
<keyLead id="C22FBB65FFE1FF87FF19FB81FB81945D" box="[151,1126,1099,1122]" pageId="2" pageNumber="637">– Elytral umbo with cylindrical or conical spine. ........................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="C22A00F5FFE1FF87FF19FBA1FB81949D" pageId="2" pageNumber="637">
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<keyLead id="C22FBB65FFE1FF87FF19FBA1FB8194BD" box="[151,1126,1131,1154]" pageId="2" pageNumber="637">3 Wingless, elytral humeri strongly oblique. .................................................................... 4</keyLead>
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<paragraph id="79611850FFE1FF87FF19FB41FB81949D" blockId="2.[151,1127,971,1570]" box="[151,1126,1163,1186]" pageId="2" pageNumber="637">
<keyLead id="C22FBB65FFE1FF87FF19FB41FB81949D" box="[151,1126,1163,1186]" pageId="2" pageNumber="637">– Winged, humeri subrectangular. ..................................................................................... 7</keyLead>
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<paragraph id="79611850FFE1FF87FF19FB61FB8194DD" blockId="2.[151,1127,971,1570]" pageId="2" pageNumber="637">
<keyLead id="C22FBB65FFE1FF87FF19FB61FB8194DD" pageId="2" pageNumber="637">
4 Elytra wholly black, very coarsely punctured; body length
<quantity id="BE26B5B5FFE1FF87FCBEFB61FC6294FD" box="[816,901,1195,1218]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="637" unit="mm" value="7.5">7.5 mm</quantity>
;
<collectingCountry id="01C958C0FFE1FF87FC01FB61FC0B94FD" box="[911,1004,1195,1218]" name="Vietnam" pageId="2" pageNumber="637">Vietnam</collectingCountry>
,
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. ...... ...................................................................................................
<taxonomicName id="BEDE63D3FFE1FF87FCD2FB00FB8194DD" authority="Arrow, 1928" authorityName="Arrow" authorityYear="1928" box="[860,1126,1226,1250]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="637" phylum="Arthropoda" rank="species" species="sordidus">
<emphasis id="4BAAC442FFE1FF87FCD2FB00FC3F94DD" bold="true" box="[860,984,1226,1250]" italics="true" pageId="2" pageNumber="637">A. sordidus</emphasis>
Arrow, 1928
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<keyLead id="C22FBB65FFE1FF87FF19FB21FB81953D" box="[151,1126,1259,1282]" pageId="2" pageNumber="637">– Elytra with red or yellow areas. ..................................................................................... 5</keyLead>
</paragraph>
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<keyLead id="C22FBB65FFE1FF87FF19FAC1FB81951D" box="[151,1126,1291,1314]" pageId="2" pageNumber="637">5 Elytra with 2 preapical spots. ......................................................................................... 6</keyLead>
</paragraph>
<paragraph id="79611850FFE1FF87FF19FAE1FB81955D" blockId="2.[151,1127,971,1570]" pageId="2" pageNumber="637">
<keyLead id="C22FBB65FFE1FF87FF19FAE1FB81955D" pageId="2" pageNumber="637">
– Elytra with single pre-apical spot; body length 6.5–7.0 mm;
<collectingRegion id="BB1AD6B2FFE1FF87FCCFFAE1FC93957D" box="[833,884,1323,1346]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="637">Java</collectingRegion>
. .................................. ............................................................................................
<taxonomicName id="BEDE63D3FFE1FF87FCA0FA80FB81955D" authority="Gorham, 1897" authorityName="Gorham" authorityYear="1897" box="[814,1126,1354,1378]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="637" phylum="Arthropoda" rank="species" species="verrucosus">
<emphasis id="4BAAC442FFE1FF87FCA0FA80FC22955E" bold="true" box="[814,965,1354,1378]" italics="true" pageId="2" pageNumber="637">A. verrucosus</emphasis>
Gorham, 1897
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="C22A00F5FFE1FF87FF19FAA1FB81961D" pageId="2" pageNumber="637">
<paragraph id="79611850FFE1FF87FF19FAA1FB8195FD" blockId="2.[151,1127,971,1570]" pageId="2" pageNumber="637">
<keyLead id="C22FBB65FFE1FF87FF19FAA1FB8195FD" pageId="2" pageNumber="637">
6 Elytral humeri dark; discal prominence smaller, rounded; preapical spots transversely- -oval, sometimes connected in form of transverse band; body length
<quantity id="BE26B5B5FFE1FF87FC10FA41FC13959D" box="[926,1012,1419,1442]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="2" pageNumber="637" unit="mm" value="6.3">6.3 mm</quantity>
;
<collectingCountry id="01C958C0FFE1FF87FC71FA41FB80959D" box="[1023,1127,1419,1442]" name="Indonesia" pageId="2" pageNumber="637">Indonesia</collectingCountry>
(
<collectingRegion id="BB1AD6B2FFE1FF87FF4AFA61FEC495FD" box="[196,291,1451,1474]" country="Indonesia" name="Sumatera Utara" pageId="2" pageNumber="637">Sumatra</collectingRegion>
). ..............................................................................
<taxonomicName id="BEDE63D3FFE1FF87FCC9FA60FB8195FD" authority="Strohecker, 1964" authorityName="Strohecker" authorityYear="1964" box="[839,1126,1450,1474]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="637" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="4BAAC442FFE1FF87FCC9FA60FC4D95FE" bold="true" box="[839,938,1450,1474]" italics="true" pageId="2" pageNumber="637">A. nanus</emphasis>
<bibRefCitation id="1D4F65A1FFE1FF87FC3FFA61FB8195FD" author="STROHECKER H. F." box="[945,1126,1451,1474]" pageId="2" pageNumber="637" pagination="319 - 357" refId="ref4179" refString="STROHECKER H. F. 1964: A synopsis of the Amphisternini (Coleoptera: Endomychidae). Pacific Insects 6: 319 - 357." type="journal article" year="1964">Strohecker, 1964</bibRefCitation>
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</keyLead>
</paragraph>
<paragraph id="79611850FFE1FF87FF19FA01FB81961D" blockId="2.[151,1127,971,1570]" pageId="2" pageNumber="637">
<keyLead id="C22FBB65FFE1FF87FF19FA01FB81961D" pageId="2" pageNumber="637">
– Elytral humeri with red-orange spot; discal prominence large, rectangular; preapical spots elongate-oval; body length
<quantity id="BE26B5B5FFE1FF87FD91FA21FD44963D" box="[543,675,1515,1538]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" metricValueMax="7.8" metricValueMin="6.8" pageId="2" pageNumber="637" unit="mm" value="7.3" valueMax="7.8" valueMin="6.8">6.8–7.8 mm</quantity>
;
<collectingCountry id="01C958C0FFE1FF87FD23FA21FCC2963D" box="[685,805,1515,1538]" name="Philippines" pageId="2" pageNumber="637">Philippines</collectingCountry>
(Mindanao). ......................... ..............................................................................................................
<emphasis id="4BAAC442FFE1FF87FC2AF9C0FB81961D" bold="true" box="[932,1126,1546,1570]" pageId="2" pageNumber="637">
<taxonomicName id="BEDE63D3FFE1FF87FC2AF9C0FBED961D" authority="Tomaszewska, 2014" authorityName="Tomaszewska" authorityYear="2014" box="[932,1034,1546,1570]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="637" phylum="Arthropoda" rank="species" species="alberti" status="sp. nov.">
<emphasis id="4BAAC442FFE1FF87FC2AF9C0FBED961D" bold="true" box="[932,1034,1546,1570]" italics="true" pageId="2" pageNumber="637">A. alberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50997939FFE1FF87FB9FF9C1FB81961D" box="[1041,1126,1547,1570]" pageId="2" pageNumber="637" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<keyStep id="C22A00F5FFE0FF86FFFFFF16FBA7912C" pageId="3" pageNumber="638">
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<keyLead id="C22FBB65FFE0FF86FFFFFF16FBA790CC" box="[113,1088,220,243]" pageId="3" pageNumber="638">7 Umbonal carina without tubercle. .................................................................................. 8</keyLead>
</paragraph>
<paragraph id="79611850FFE0FF86FFFFFF36FBA7912C" blockId="3.[113,1089,220,1139]" box="[113,1088,252,275]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFF36FBA7912C" box="[113,1088,252,275]" pageId="3" pageNumber="638">– Umbonal carina with tubercle. ..................................................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="C22A00F5FFE0FF86FFFFFED6FBA7914C" pageId="3" pageNumber="638">
<paragraph id="79611850FFE0FF86FFFFFED6FBA7910C" blockId="3.[113,1089,220,1139]" box="[113,1088,284,307]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFED6FBA7910C" box="[113,1088,284,307]" pageId="3" pageNumber="638">8 Elytra slightly longer than their combined width. .......................................................... 9</keyLead>
</paragraph>
<paragraph id="79611850FFE0FF86FFFFFEF6FBA7914C" blockId="3.[113,1089,220,1139]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFEF6FBA7914C" pageId="3" pageNumber="638">
– Elytra 1.5 times as long as wide; body length 9.5–10.0 mm;
<collectingCountry id="01C958C0FFE0FF86FC94FEF6FC98916C" box="[794,895,316,339]" name="Malaysia" pageId="3" pageNumber="638">Malaysia</collectingCountry>
. ........................... ..............................................................................................
<taxonomicName id="BEDE63D3FFE0FF86FC96FE91FBA7914C" authority="Strohecker, 1957" authorityName="Strohecker" authorityYear="1957" box="[792,1088,347,371]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="opacus">
<emphasis id="4BAAC442FFE0FF86FC96FE91FC63914C" bold="true" box="[792,900,347,371]" italics="true" pageId="3" pageNumber="638">A. opacus</emphasis>
Strohecker, 1957
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9 Elytra abruptly rounded to apex; body length 8.0–
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;
<collectingRegion id="BB1AD6B2FFE0FF86FC82FEB6FCD891AC" box="[780,831,380,403]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="638">Java</collectingRegion>
. .................................... ..........................................................................................
<taxonomicName id="BEDE63D3FFE0FF86FD73FE51FBA7918C" authority="Germar, 1843" authorityName="Germar" authorityYear="1843" box="[765,1088,411,435]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="4BAAC442FFE0FF86FD73FE51FC40918C" bold="true" box="[765,935,411,435]" italics="true" pageId="3" pageNumber="638">A. tuberculatus</emphasis>
<bibRefCitation id="1D4F65A1FFE0FF86FC23FE56FBA7918C" author="GERMAR E. F." box="[941,1088,412,435]" pageId="3" pageNumber="638" pagination="84 - 86" refId="ref3880" refString="GERMAR E. F. 1843: Eumorphus. Pp. 84 - 86. In: ERSCH J. S. &amp; GRUBER J. G. (eds): Algemeine Encyclopaedie der Wissenschaften und Kuenste, Vol. 39. Eugen - Ezzelino, Leipzig, 492 pp." type="book chapter" year="1843">Germar, 1843</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="79611850FFE0FF86FFFFFE76FBA791CC" blockId="3.[113,1089,220,1139]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFE76FBA791CC" pageId="3" pageNumber="638">
– Elytra distinctly tapering behind middle; body length 7.0 mm;
<collectingCountry id="01C958C0FFE0FF86FCBDFE76FC7A91EC" box="[819,925,444,467]" name="Indonesia" pageId="3" pageNumber="638">Indonesia</collectingCountry>
,
<collectingCountry id="01C958C0FFE0FF86FC26FE76FBEA91EC" box="[936,1037,444,467]" name="Malaysia" pageId="3" pageNumber="638">Malaysia</collectingCountry>
. ..... ..................................................................................................
<taxonomicName id="BEDE63D3FFE0FF86FCA3FE11FBA791CC" authority="Gorham, 1901" authorityName="Gorham" authorityYear="1901" box="[813,1088,475,499]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="eruptus">
<emphasis id="4BAAC442FFE0FF86FCA3FE11FC7891CC" bold="true" box="[813,927,475,499]" italics="true" pageId="3" pageNumber="638">A. eruptus</emphasis>
Gorham, 1901
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C22A00F5FFE0FF86FFFFFE36FBA7926C" pageId="3" pageNumber="638">
<paragraph id="79611850FFE0FF86FFFFFE36FBA7922C" blockId="3.[113,1089,220,1139]" box="[113,1088,508,531]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFE36FBA7922C" box="[113,1088,508,531]" pageId="3" pageNumber="638">10. Elytra with a discal subcarinate tubercle. .................................................................... 11</keyLead>
</paragraph>
<paragraph id="79611850FFE0FF86FFFFFDD6FBA7926C" blockId="3.[113,1089,220,1139]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFDD6FBA7926C" pageId="3" pageNumber="638">
– Elytra with a high discal prominence; body length
<quantity id="BE26B5B5FFE0FF86FD3FFDD6FCE2920C" box="[689,773,540,563]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="638" unit="mm" value="7.5">7.5 mm</quantity>
;
<collectingCountry id="01C958C0FFE0FF86FC9EFDD6FC9D920C" box="[784,890,540,563]" name="Indonesia" pageId="3" pageNumber="638">Indonesia</collectingCountry>
,
<collectingCountry id="01C958C0FFE0FF86FC0AFDD6FC01920C" box="[900,998,540,563]" name="Malaysia" pageId="3" pageNumber="638">Malaysia</collectingCountry>
(
<collectingCountry id="01C958C0FFE0FF86FC7AFDD6FF55926C" name="Malaysia" pageId="3" pageNumber="638">Malacca</collectingCountry>
). ......................................................................................
<taxonomicName id="BEDE63D3FFE0FF86FC80FDF1FBA7926C" authority="Gorham, 1901" authorityName="Gorham" authorityYear="1901" box="[782,1088,571,595]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="vomeratus">
<emphasis id="4BAAC442FFE0FF86FC80FDF1FC78926C" bold="true" box="[782,927,571,595]" italics="true" pageId="3" pageNumber="638">A. vomeratus</emphasis>
Gorham, 1901
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C22A00F5FFE0FF86FFFFFD96FBA792EC" pageId="3" pageNumber="638">
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<keyLead id="C22FBB65FFE0FF86FFFFFD96FBA792AC" pageId="3" pageNumber="638">
11 Scape of antenna and the outer half of each femur blood-red; body length 9.0 mm;
<collectingCountry id="01C958C0FFE0FF86FB9EFD96FF5C92AC" name="Myanmar" pageId="3" pageNumber="638">Burma</collectingCountry>
. ..................................................................................
<taxonomicName id="BEDE63D3FFE0FF86FD7DFDB1FBA792AC" authority="Gerstaecker, 1857" authorityName="Gerstaecker" authorityYear="1857" box="[755,1088,635,659]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="corallifer">
<emphasis id="4BAAC442FFE0FF86FD7DFDB1FC9F92AC" bold="true" box="[755,888,635,659]" italics="true" pageId="3" pageNumber="638">A. corallifer</emphasis>
<bibRefCitation id="1D4F65A1FFE0FF86FCF1FDB6FBA792AC" author="GERSTAECKER A." box="[895,1088,636,659]" pageId="3" pageNumber="638" pagination="211 - 243" refId="ref3932" refString="GERSTAECKER A. 1857: Versuch einer systematischen Auseinandersetzung der Gattungen Eumorphus Web. und Endomychus Payk. Archiv fur Naturgeschichte 23 (1): 211 - 243." type="journal article" year="1857">Gerstaecker, 1857</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="79611850FFE0FF86FFFFFD56FBA792EC" blockId="3.[113,1089,220,1139]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFD56FBA792EC" pageId="3" pageNumber="638">
– Scape of antennae and femora black; body length
<quantity id="BE26B5B5FFE0FF86FD22FD56FCAD928C" box="[684,842,668,691]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.13" metricValueMin="1.0699999999999998" pageId="3" pageNumber="638" unit="mm" value="11.0" valueMax="11.3" valueMin="10.7">10.7–11.3 mm</quantity>
;
<collectingCountry id="01C958C0FFE0FF86FCDAFD56FC70928C" box="[852,919,668,691]" name="China" pageId="3" pageNumber="638">China</collectingCountry>
. ....................... ....................................................................................
<taxonomicName id="BEDE63D3FFE0FF86FD5DFD71FBA792EC" authority="Wang &amp; Ren, 2007" authorityName="Wang &amp; Ren" authorityYear="2007" box="[723,1088,699,723]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="rufituberus">
<emphasis id="4BAAC442FFE0FF86FD5DFD71FC8A92EC" bold="true" box="[723,877,699,723]" italics="true" pageId="3" pageNumber="638">A. rufituberus</emphasis>
<bibRefCitation id="1D4F65A1FFE0FF86FCFDFD76FBA792EC" author="WANG J. - L. &amp; REN G. - D." box="[883,1088,700,723]" pageId="3" pageNumber="638" pagination="459 - 461" refId="ref4582" refString="WANG J. - L. &amp; REN G. - D. 2007: A new record of genus and new species of Endomychidae (Coleoptera, Cucujoidea) from China. Acta Zootaxonomica Sinica 32: 459 - 461." type="journal article" year="2007">Wang &amp; Ren, 2007</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C22A00F5FFE0FF86FFFFFD16FBA7930C" pageId="3" pageNumber="638">
<paragraph id="79611850FFE0FF86FFFFFD16FBA7932C" blockId="3.[113,1089,220,1139]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFD16FBA7932C" pageId="3" pageNumber="638">
12 Elytra much prolonged at apex; body length
<quantity id="BE26B5B5FFE0FF86FDFDFD16FD2F92CC" box="[627,712,732,755]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="3" pageNumber="638" unit="mm" value="7.4">7.4 mm</quantity>
;
<collectingCountry id="01C958C0FFE0FF86FD5DFD16FCDF92CC" box="[723,824,732,755]" name="Malaysia" pageId="3" pageNumber="638">Malaysia</collectingCountry>
. ..................................... ...........................................................................................
<taxonomicName id="BEDE63D3FFE0FF86FC8CFD31FBA7932C" authority="Strohecker, 1964" authorityName="Strohecker" authorityYear="1964" box="[770,1088,763,787]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="caudatus">
<emphasis id="4BAAC442FFE0FF86FC8CFD31FC63932C" bold="true" box="[770,900,763,787]" italics="true" pageId="3" pageNumber="638">A. caudatus</emphasis>
<bibRefCitation id="1D4F65A1FFE0FF86FC05FD36FBA7932C" author="STROHECKER H. F." box="[907,1088,764,787]" pageId="3" pageNumber="638" pagination="319 - 357" refId="ref4179" refString="STROHECKER H. F. 1964: A synopsis of the Amphisternini (Coleoptera: Endomychidae). Pacific Insects 6: 319 - 357." type="journal article" year="1964">Strohecker, 1964</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="79611850FFE0FF86FFFFFCD6FBA7930C" blockId="3.[113,1089,220,1139]" box="[113,1088,796,819]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFCD6FBA7930C" box="[113,1088,796,819]" pageId="3" pageNumber="638">– Elytra rather abruptly rounded to apex. ........................................................................ 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="C22A00F5FFE0FF86FFFFFCF6FBA793AC" pageId="3" pageNumber="638">
<paragraph id="79611850FFE0FF86FFFFFCF6FBA7934C" blockId="3.[113,1089,220,1139]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFCF6FBA7934C" pageId="3" pageNumber="638">
13 Discal prominence of elytron truncate; body length 7.0 mm;
<collectingCountry id="01C958C0FFE0FF86FC90FCF6FC64936C" box="[798,899,828,851]" name="Malaysia" pageId="3" pageNumber="638">Malaysia</collectingCountry>
. .......................... ....................................................................................................
<taxonomicName id="BEDE63D3FFE0FF86FCB0FC91FBA7934C" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[830,1088,859,883]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="grandjeani">
<emphasis id="4BAAC442FFE0FF86FCB0FC91FC32934C" bold="true" box="[830,981,859,883]" italics="true" pageId="3" pageNumber="638">A. grandjeani</emphasis>
Pic, 1930
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="79611850FFE0FF86FFFFFCB6FBA793AC" blockId="3.[113,1089,220,1139]" box="[113,1088,892,915]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFCB6FBA793AC" box="[113,1088,892,915]" pageId="3" pageNumber="638">– Discal prominence of elytron rounded at apex. ............................................................ 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="C22A00F5FFE0FF86FFFFFC56FBA793CC" pageId="3" pageNumber="638">
<paragraph id="79611850FFE0FF86FFFFFC56FBA793EC" blockId="3.[113,1089,220,1139]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFC56FBA793EC" pageId="3" pageNumber="638">
14 Elytron with broad post-median pale bar; body length 8.0 mm;
<collectingCountry id="01C958C0FFE0FF86FCB4FC56FC7B938C" box="[826,924,924,947]" name="Malaysia" pageId="3" pageNumber="638">Malaysia</collectingCountry>
(
<collectingCountry id="01C958C0FFE0FF86FC24FC56FBEC938C" box="[938,1035,924,947]" name="Malaysia" pageId="3" pageNumber="638">Malacca</collectingCountry>
). ..... .................................................................................................
<taxonomicName id="BEDE63D3FFE0FF86FCBCFC71FBA793EC" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[818,1088,955,979]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="malaccanus">
<emphasis id="4BAAC442FFE0FF86FCBCFC71FC3293EC" bold="true" box="[818,981,955,979]" italics="true" pageId="3" pageNumber="638">A. malaccanus</emphasis>
Pic, 1930
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="79611850FFE0FF86FFFFFC16FBA793CC" blockId="3.[113,1089,220,1139]" box="[113,1088,988,1011]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFC16FBA793CC" box="[113,1088,988,1011]" pageId="3" pageNumber="638">
– Elytron with 2 small spots behind middle; body length
<quantity id="BE26B5B5FFE0FF86FD6DFC16FCDF93CC" box="[739,824,988,1011]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="638" unit="mm" value="7.5">7.5 mm</quantity>
. ................................. 15
</keyLead>
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<paragraph id="79611850FFE0FF86FFFFFC36FBA7940C" blockId="3.[113,1089,220,1139]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFC36FBA7940C" pageId="3" pageNumber="638">
15 At least hind elytral pustules red; discal elytral prominence at least as high as long; ......
<collectingCountry id="01C958C0FFE0FF86FF19FBD6FF1E940C" box="[151,249,1052,1075]" name="Malaysia" pageId="3" pageNumber="638">Malaysia</collectingCountry>
(
<collectingCountry id="01C958C0FFE0FF86FE89FBD6FE8E940C" box="[263,361,1052,1075]" name="Malaysia" pageId="3" pageNumber="638">Sarawak</collectingCountry>
). ..................................................
<taxonomicName id="BEDE63D3FFE0FF86FD5BFBD1FBA7940C" authority="Gerstaecker, 1857" authorityName="Gerstaecker" authorityYear="1857" box="[725,1088,1051,1075]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="4BAAC442FFE0FF86FD5BFBD1FC9E940C" bold="true" box="[725,889,1051,1075]" italics="true" pageId="3" pageNumber="638">A. mucronatus</emphasis>
<bibRefCitation id="1D4F65A1FFE0FF86FCF1FBD6FBA7940C" author="GERSTAECKER A." box="[895,1088,1052,1075]" pageId="3" pageNumber="638" pagination="211 - 243" refId="ref3932" refString="GERSTAECKER A. 1857: Versuch einer systematischen Auseinandersetzung der Gattungen Eumorphus Web. und Endomychus Payk. Archiv fur Naturgeschichte 23 (1): 211 - 243." type="journal article" year="1857">Gerstaecker, 1857</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="79611850FFE0FF86FFFFFBF6FBA7944C" blockId="3.[113,1089,220,1139]" pageId="3" pageNumber="638">
<keyLead id="C22FBB65FFE0FF86FFFFFBF6FBA7944C" pageId="3" pageNumber="638">
– Elytral pustules dark; discal elytral prominence lower than long,
<collectingCountry id="01C958C0FFE0FF86FCC5FBF6FC4A946C" box="[843,941,1084,1107]" name="Malaysia" pageId="3" pageNumber="638">Malaysia</collectingCountry>
(
<collectingRegion id="BB1AD6B2FFE0FF86FC35FBF6FBE5946C" box="[955,1026,1084,1107]" country="Malaysia" name="Sabah" pageId="3" pageNumber="638">Sabah</collectingRegion>
). ...... ............................................................................................
<taxonomicName id="BEDE63D3FFE0FF86FC87FB91FBA7944C" authority="Strohecker, 1968" authorityName="Strohecker" authorityYear="1968" box="[777,1088,1115,1139]" class="Insecta" family="Endomychidae" genus="Amphisternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="638" phylum="Arthropoda" rank="species" species="lugubris">
<emphasis id="4BAAC442FFE0FF86FC87FB91FC63944C" bold="true" box="[777,900,1115,1139]" italics="true" pageId="3" pageNumber="638">A. lugubris</emphasis>
<bibRefCitation id="1D4F65A1FFE0FF86FC05FB96FBA7944C" author="STROHECKER H. F." box="[907,1088,1116,1139]" pageId="3" pageNumber="638" pagination="113 - 114" refId="ref4205" refString="STROHECKER H. F. 1968: Addenda to the Amphisternini (Coleoptera: Endomychidae). Pacific Insects 10: 113 - 114." type="journal article" year="1968">Strohecker, 1968</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

133
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<paragraph id="79611850FFEBFF8DFE37FF11FCA290CC" blockId="8.[441,837,219,243]" box="[441,837,219,243]" pageId="8" pageNumber="643">
<heading id="2229AF3CFFEBFF8DFE37FF11FCA290CC" bold="true" box="[441,837,219,243]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="643" reason="3">
<emphasis id="4BAAC442FFEBFF8DFE37FF11FCA290CC" bold="true" box="[441,837,219,243]" pageId="8" pageNumber="643">
<taxonomicName id="BEDE63D3FFEBFF8DFE37FF11FD0E90CC" authority="Tomaszewska, 2014" authorityName="Tomaszewska" authorityYear="2014" box="[441,745,219,243]" class="Insecta" family="Endomychidae" genus="Gerstaeckerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="643" phylum="Arthropoda" rank="species" species="rufiplagiatus" status="sp. nov.">
<emphasis id="4BAAC442FFEBFF8DFE37FF11FD0E90CC" bold="true" box="[441,745,219,243]" italics="true" pageId="8" pageNumber="643">Gerstaeckerus rufiplagiatus</emphasis>
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<subSubSection id="31C44BDBFFEBFF8DFDCDFF30FD5C9132" box="[579,699,250,269]" pageId="8" pageNumber="643" type="description">
<paragraph id="79611850FFEBFF8DFDCDFF30FD5C9132" blockId="8.[151,1127,250,337]" box="[579,699,250,269]" pageId="8" pageNumber="643">
(
<figureCitation id="E1E504D5FFEBFF8DFDC7FF30FD719132" box="[585,662,250,269]" captionStart="Figs 1–8" captionStartId="4.[151,188,1506,1525]" captionTargetBox="[176,1112,224,1445]" captionTargetId="figure-10@4.[151,1126,224,1456]" captionTargetPageId="4" captionText="Figs 1–8. Habitus. 1–5 – Amphisternus alberti sp. nov.; 6–8 – Gerstaeckerus rufiplagiatus sp. nov. 1, 6 – dorsal habitus; 2 – lateral habitus; 3, 8 – pronotum; 4, 7 – antenna; 5 – habitus, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5299675" httpUri="https://zenodo.org/record/5299675/files/figure.png" pageId="8" pageNumber="643">Figs 6–8</figureCitation>
,
<figureCitation id="E1E504D5FFEBFF8DFD11FF30FD529132" box="[671,693,250,269]" captionStart="Figs 9–14" captionStartId="5.[113,150,1528,1547]" captionTargetBox="[195,1035,433,1509]" captionTargetId="figure-10@5.[144,1056,430,1512]" captionTargetPageId="5" captionText="Figs 9–14. Genitalia. 9–13 – Amphisternus alberti sp. nov.; 14 – Gerstaeckerus rufiplagiatus, sp. nov. 9 – aedeagus, dorsal view; 10 – aedeagus, ventral view; 11 – abdominal segment VIII, male, ventral view; 12 – male genital segment, ventral view; 13, 14 – female genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5299677" httpUri="https://zenodo.org/record/5299677/files/figure.png" pageId="8" pageNumber="643">14</figureCitation>
)
</paragraph>
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<subSubSection id="31C44BDBFFEBFF8DFF19FEEEFCD5916E" pageId="8" pageNumber="643" type="materials_examined">
<paragraph id="79611850FFEBFF8DFF19FEEEFCD5916E" blockId="8.[151,1127,250,337]" pageId="8" pageNumber="643">
<emphasis id="4BAAC442FFEBFF8DFF19FEEEFEFA9108" bold="true" box="[151,285,292,311]" pageId="8" pageNumber="643">Type material.</emphasis>
<materialsCitation id="C9B6120DFFEBFF8DFEAAFEEEFB809108" box="[292,1127,288,311]" collectingDate="2013-10" collectorName="I. Lumawig" country="Philippines" location="Calahinan" municipality="Mindanao" pageId="8" pageNumber="643" specimenCount="1" specimenCount-female="1" stateProvince="Bukidnon" typeStatus="holotype">
<typeStatus id="A665A6F2FFEBFF8DFEAAFEEEFE90910A" box="[292,375,292,311]" pageId="8" pageNumber="643" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="6FD8D3D9FFEBFF8DFE0AFEEAFE739108" box="[388,404,288,311]" pageId="8" pageNumber="643" type="female">♀</specimenCount>
, ‘
<collectingCountry id="01C958C0FFEBFF8DFE2DFEEEFDC29108" box="[419,549,292,311]" name="Philippines" pageId="8" pageNumber="643">PHILIPPINES</collectingCountry>
/
<collectingMunicipality id="9905822AFFEBFF8DFDB7FEEEFD689108" box="[569,655,292,311]" pageId="8" pageNumber="643">Mindanao</collectingMunicipality>
/
<collectingRegion id="BB1AD6B2FFEBFF8DFD12FEEEFD179108" box="[668,752,292,311]" country="Philippines" name="Bukidnon" pageId="8" pageNumber="643">Bukidnon</collectingRegion>
/
<location id="7C014E8BFFEBFF8DFC8AFEEEFCBC9108" LSID="urn:lsid:plazi:treatment:F177A946FFEBFF8DFE37FF11FDD3937E:7C014E8BFFEBFF8DFC8AFEEEFCBC9108" box="[772,859,292,311]" country="Philippines" municipality="Mindanao" name="Calahinan" pageId="8" pageNumber="643" stateProvince="Bukidnon">Calahinan</location>
/
<date id="0D603E90FFEBFF8DFCE0FEEEFC529108" box="[878,949,292,311]" pageId="8" pageNumber="643" value="2013-10">
<collectingDate id="1D24C778FFEBFF8DFCE0FEEEFC529108" box="[878,949,292,311]" pageId="8" pageNumber="643" value="2013-10">X. 2013</collectingDate>
</date>
/ leg.
<collectorName id="D42B7D86FFEBFF8DFC61FEEEFBB39108" box="[1007,1108,292,311]" pageId="8" pageNumber="643">I. Lumawig</collectorName>
//
</materialsCitation>
<materialsCitation id="C9B6120DFFEBFF8DFF19FEF4FCC9916E" box="[151,814,318,337]" collectionCode="MZPW" country="Philippines" pageId="8" pageNumber="643" specimenCount="1" stateProvince="Bukidnon" typeStatus="holotype">
<typeStatus id="A665A6F2FFEBFF8DFF19FEF4FF01916E" box="[151,230,318,337]" pageId="8" pageNumber="643" type="holotype">Holotype</typeStatus>
<taxonomicName id="BEDE63D3FFEBFF8DFF62FEF4FE3E916E" authority="Tomaszewska, 2014" authorityName="Tomaszewska" authorityYear="2014" box="[236,473,318,337]" class="Insecta" family="Endomychidae" genus="Gerstaeckerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="643" phylum="Arthropoda" rank="species" species="rufiplagiatus" status="sp. nov.">
<emphasis id="4BAAC442FFEBFF8DFF62FEF4FE3E916E" box="[236,473,318,337]" italics="true" pageId="8" pageNumber="643">Gerstaeckerus rufiplagiatus</emphasis>
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<taxonomicNameLabel id="50997939FFEBFF8DFE51FEF4FDC3916E" box="[479,548,318,337]" pageId="8" pageNumber="643" rank="species">sp. nov.</taxonomicNameLabel>
, Tomaszewska 2014’ (
<collectionCode id="1FCF8095FFEBFF8DFD6DFEF4FCCF916E" box="[739,808,318,337]" country="Poland" httpUri="http://grbio.org/cool/ynr4-t34e" name="Polish Academy of Science, Museum of the Institute of Zoology" pageId="8" pageNumber="643">MZPW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="31C44BDBFFEBFF8DFF19FEA0FEB992FE" pageId="8" pageNumber="643" type="description">
<paragraph id="79611850FFEBFF8DFF19FEA0FBFB929E" blockId="8.[151,1127,362,833]" pageId="8" pageNumber="643">
<emphasis id="4BAAC442FFEBFF8DFF19FEA0FEC591BE" bold="true" box="[151,290,362,385]" pageId="8" pageNumber="643">Description.</emphasis>
Length 9.00 mm. Body 1.88 times as long as wide; pronotum 0.55 times as long as wide; elytra 1.27 times as long as wide; 3.40 times longer than pronotum, 1.48 times wider than pronotum. Body black, with only each elytron covered with elongate, reddish fascia (
<figureCitation id="E1E504D5FFEBFF8DFF6DFE00FEC391DE" box="[227,292,458,481]" captionStart="Figs 1–8" captionStartId="4.[151,188,1506,1525]" captionTargetBox="[176,1112,224,1445]" captionTargetId="figure-10@4.[151,1126,224,1456]" captionTargetPageId="4" captionText="Figs 1–8. Habitus. 1–5 – Amphisternus alberti sp. nov.; 6–8 – Gerstaeckerus rufiplagiatus sp. nov. 1, 6 – dorsal habitus; 2 – lateral habitus; 3, 8 – pronotum; 4, 7 – antenna; 5 – habitus, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5299675" httpUri="https://zenodo.org/record/5299675/files/figure.png" pageId="8" pageNumber="643">Fig. 6</figureCitation>
); shiny; moderately convex; glabrous. Antenna (
<figureCitation id="E1E504D5FFEBFF8DFCA8FE00FC8291DE" box="[806,869,458,481]" captionStart="Figs 1–8" captionStartId="4.[151,188,1506,1525]" captionTargetBox="[176,1112,224,1445]" captionTargetId="figure-10@4.[151,1126,224,1456]" captionTargetPageId="4" captionText="Figs 1–8. Habitus. 1–5 – Amphisternus alberti sp. nov.; 6–8 – Gerstaeckerus rufiplagiatus sp. nov. 1, 6 – dorsal habitus; 2 – lateral habitus; 3, 8 – pronotum; 4, 7 – antenna; 5 – habitus, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5299675" httpUri="https://zenodo.org/record/5299675/files/figure.png" pageId="8" pageNumber="643">Fig. 7</figureCitation>
) about 0.5 times as long as body; club flattened and comparatively narrow. Pronotum (
<figureCitation id="E1E504D5FFEBFF8DFCB5FE20FC9B923E" box="[827,892,490,513]" captionStart="Figs 1–8" captionStartId="4.[151,188,1506,1525]" captionTargetBox="[176,1112,224,1445]" captionTargetId="figure-10@4.[151,1126,224,1456]" captionTargetPageId="4" captionText="Figs 1–8. Habitus. 1–5 – Amphisternus alberti sp. nov.; 6–8 – Gerstaeckerus rufiplagiatus sp. nov. 1, 6 – dorsal habitus; 2 – lateral habitus; 3, 8 – pronotum; 4, 7 – antenna; 5 – habitus, anterior view." figureDoi="http://doi.org/10.5281/zenodo.5299675" httpUri="https://zenodo.org/record/5299675/files/figure.png" pageId="8" pageNumber="643">Fig. 8</figureCitation>
) widest near anterior third; lateral margins rounded along anterior half then constricted in basal third; posterior angles about right-angled, anterior angles subacute; punctures fine, 2.0–3.0 diameters apart. Prosternal process about 0.5 times as wide as procoxal diameter, excised at apex. Elytra with lateral margins weakly rounded toward apex; widest in front of mid length; with punctures as large as the pronotal ones, 1.5–2.5 diameters apart. Female genitalia as in
<figureCitation id="E1E504D5FFEBFF8DFC47FD40FBFF929E" box="[969,1048,650,673]" captionStart="Figs 9–14" captionStartId="5.[113,150,1528,1547]" captionTargetBox="[195,1035,433,1509]" captionTargetId="figure-10@5.[144,1056,430,1512]" captionTargetPageId="5" captionText="Figs 9–14. Genitalia. 9–13 – Amphisternus alberti sp. nov.; 14 – Gerstaeckerus rufiplagiatus, sp. nov. 9 – aedeagus, dorsal view; 10 – aedeagus, ventral view; 11 – abdominal segment VIII, male, ventral view; 12 – male genital segment, ventral view; 13, 14 – female genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5299677" httpUri="https://zenodo.org/record/5299677/files/figure.png" pageId="8" pageNumber="643">Fig. 14</figureCitation>
.
</paragraph>
<paragraph id="79611850FFEBFF8DFF3BFD60FEB992FE" blockId="8.[151,1127,362,833]" box="[181,350,682,705]" pageId="8" pageNumber="643">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="31C44BDBFFEBFF8DFF19FD00FD8D933E" pageId="8" pageNumber="643" type="diagnosis">
<paragraph id="79611850FFEBFF8DFF19FD00FD8D933E" blockId="8.[151,1127,362,833]" pageId="8" pageNumber="643">
<emphasis id="4BAAC442FFEBFF8DFF19FD00FE7692DE" bold="true" box="[151,401,714,737]" pageId="8" pageNumber="643">Differential diagnosis.</emphasis>
This species is easily distinguished from all its congeners by having elongate, red-orange fascia on each elytron.
</paragraph>
</subSubSection>
<subSubSection id="31C44BDBFFEBFF8DFF19FCC0FBD3931E" box="[151,1076,778,801]" pageId="8" pageNumber="643" type="etymology">
<paragraph id="79611850FFEBFF8DFF19FCC0FBD3931E" blockId="8.[151,1127,362,833]" box="[151,1076,778,801]" pageId="8" pageNumber="643">
<emphasis id="4BAAC442FFEBFF8DFF19FCC0FEFF931E" bold="true" box="[151,280,778,801]" pageId="8" pageNumber="643">Etymology.</emphasis>
The specific epithet refers to the elongate red-orange fasciae on the elytra.
</paragraph>
</subSubSection>
<subSubSection id="31C44BDBFFEBFF8DFF19FCE0FDD3937E" box="[151,564,810,833]" pageId="8" pageNumber="643" type="distribution">
<paragraph id="79611850FFEBFF8DFF19FCE0FDD3937E" blockId="8.[151,1127,362,833]" box="[151,564,810,833]" pageId="8" pageNumber="643">
<emphasis id="4BAAC442FFEBFF8DFF19FCE0FECD937E" bold="true" box="[151,298,810,833]" pageId="8" pageNumber="643">Distribution.</emphasis>
<collectingCountry id="01C958C0FFEBFF8DFEBEFCE0FE4F937E" box="[304,424,810,833]" name="Philippines" pageId="8" pageNumber="643">Philippines</collectingCountry>
(Mindanao).
</paragraph>
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</treatment>
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140
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<taxonomicName id="C481FEEF486A60F7D2B6D1480B54DEB7" ID-CoL="4R3T" authority="Haldeman, 1849" authorityName="Haldeman" authorityYear="1849" class="Insecta" family="Chrysomelidae" genus="Griburius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Griburius" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="genus">Griburius Haldeman, 1849</taxonomicName>
Fig. 24
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<taxonomicName id="578C7027096AC2BDC0912DDF2ECE648D" class="Insecta" family="Chrysomelidae" genus="Griburius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Griburius" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="genus">Griburius</taxonomicName>
Haldeman, 1849: 245.
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<taxonomicName id="A4B3F34AE4FBFE03B1680DE49E83F35F" class="Insecta" equal="=" family="Chrysomelidae" genus="Scolochrus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scolochrus" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="genus">= Scolochrus</taxonomicName>
Suffrian, 1852.
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<paragraph id="609F0AFCF6DB16ED965ADF35348D39F6" pageId="34" pageNumber="45">Type species.</paragraph>
<paragraph id="D29973633633F0AD6E66CB950EB213BF" pageId="34" pageNumber="45">
<taxonomicName id="B394757AFF015308B1E2A72E0547F490" class="Insecta" family="Chrysomelidae" genus="Griburius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Griburius scutellaris" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="species" species="scutellaris">Griburius scutellaris</taxonomicName>
<bibRefCitation id="96315EE95C6FD2D45CEECC52718D1C23" author="Haldeman, SS" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="58" pageNumber="69" pagination="169 - 171" title="Cryptocephalinarum Boreali-Americae diagnoses, cum speciebus novis museilecontiani." volume="4" year="1849">Haldeman 1849</bibRefCitation>
: 245 (=
<taxonomicName id="065616B2B5C7B41407F9B9A3CB4C8133" class="Insecta" family="Chrysomelidae" genus="Cryptocephalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptocephalus scutellaris" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="species" species="scutellaris">Cryptocephalus scutellaris</taxonomicName>
Fabricius, 1801), by monotypy.
</paragraph>
<caption id="C57DC5B567C71E518556562104F34C98" pageId="34" pageNumber="45">
<paragraph id="095373163A317254826BDFE791A10C74" pageId="34" pageNumber="45">
Figure 24.
<taxonomicName id="99AC9579DDA9AB47B2003746BE7675B8" class="Insecta" family="Chrysomelidae" genus="Griburius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Griburius cultus" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="species" species="cultus">Griburius cultus</taxonomicName>
(Suffrian) (4), left: habitus (dorsal view), right: habitus (lateral view).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="3170C0A6C7567424C26444A26F94FFFA" pageId="34" pageNumber="45" type="diagnosis">
<paragraph id="2E1AE1E5CA2CFD42842E5E05B23D3886" pageId="34" pageNumber="45">Diagnosis.</paragraph>
<paragraph id="6ECD7CB91C186DDF72CF13D565C100EA" pageId="34" pageNumber="45">
Recently,
<bibRefCitation id="ED98A7BD227766CD3E52352CD5F11365" author="Chamorro, ML" journalOrPublisher="Caucasian Entomological Bulletin" pageId="56" pageNumber="67" pagination="201 - 206" title="On the identity of Mastacanthus Suffrian, 1852 and Sternoglosus Suffrian, 1866 and key to world genera of Pachybrachina (Chrysomelidae: Cryptocephalinae: Cryptocephalini)." volume="9" year="2013">Chamorro (2013)</bibRefCitation>
provided the following characters to identify the species of this genus: posterior margin of intercoxal prosternal process rounded; lateral margin of elytra deeply excised exposing abdomen caudally, elytra length approximately 2
<normalizedToken id="7B3E3D0457853A5F6F54636FB24108AF" originalValue="×">x</normalizedToken>
or less length of pronotum. However, the limits of
<taxonomicName id="E84AC773384618D4A4917C9407B58EE9" class="Insecta" family="Chrysomelidae" genus="Griburius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Griburius" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="genus">Griburius</taxonomicName>
and
<taxonomicName id="EA56DA0CC2A9E8DD9EDB7D3CC03B03B4" genus="Metallactus" lsidName="Metallactus" pageId="34" pageNumber="45" rank="genus">Metallactus</taxonomicName>
are confused and require revision.
</paragraph>
</subSubSection>
<subSubSection id="9E75DDE3FE50B8ED5B5DCAD7061AFE24" pageId="34" pageNumber="45" type="distribution">
<paragraph id="9F760D1689115C89CD8082D93469BEB5" pageId="34" pageNumber="45">Distribution.</paragraph>
<paragraph id="4F4D8BE6FDC4BE8F7A9A3AB9038983F6" pageId="34" pageNumber="45">Nearctic and Neotropical.</paragraph>
</subSubSection>
<subSubSection id="AAD22A3EB15513A6C14900FEA14E8358" pageId="34" pageNumber="45" type="argentinian species checklist">
<paragraph id="1DBF24EC573C666CC936A523DE9D2C8C" pageId="34" pageNumber="45">Argentinian species checklist.</paragraph>
<paragraph id="FD87E943892C1A196C23484E8782F85E" pageId="34" pageNumber="45">
<taxonomicName id="A6F2431ED9CA99C7D64FC0E622C0D4AA" class="Insecta" family="Chrysomelidae" genus="Griburius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Griburius bilineolatus" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="species" species="bilineolatus">Griburius bilineolatus</taxonomicName>
(Suffrian, 1866) (BAS, COR).
</paragraph>
<paragraph id="E9D39FBC1533EC9446F0CA67E4E2A243" pageId="34" pageNumber="45">
<taxonomicName id="471F0D7FF8F6372A31A035576E17E724" class="Insecta" family="Chrysomelidae" genus="Griburius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Griburius conspurcatus" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="species" species="conspurcatus">Griburius conspurcatus</taxonomicName>
(Suffrian, 1866) (BAS).
</paragraph>
<paragraph id="8722850A50E0A5ACE9CECBBF3CF8776D" pageId="34" pageNumber="45">
<taxonomicName id="C486320569A1C578B5D392DBF12B735A" class="Insecta" family="Chrysomelidae" genus="Griburius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Griburius cultus" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="species" species="cultus">Griburius cultus</taxonomicName>
(Suffrian, 1866) (BAS).
</paragraph>
<paragraph id="B6BF661996BC8B584FF6D95BAC4511B1" pageId="34" pageNumber="45">
<taxonomicName id="24F04D1DBC7F85C3700C0DBC113E601E" class="Insecta" family="Chrysomelidae" genus="Griburius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Griburius fastidiosus" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="species" species="fastidiosus">Griburius fastidiosus</taxonomicName>
(Suffrian, 1866) (BAS).
</paragraph>
<paragraph id="7AC25478BE5AA8184E2E5EE8499F01F1" pageId="34" pageNumber="45">
<taxonomicName id="5180ACE95AE3ABB5CB92B7FD6B736B62" class="Insecta" family="Chrysomelidae" genus="Griburius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Griburius octoguttatus" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="species" species="octoguttatus">Griburius octoguttatus</taxonomicName>
Burmeister, 1877 (ERS).
</paragraph>
<paragraph id="984AB296A29A5B2F3D0EFB01346F2860" pageId="34" pageNumber="45">
<taxonomicName id="C8F1ED7C1CF87DCE4C0B659BF3151275" class="Insecta" family="Chrysomelidae" genus="Griburius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Griburius persimilis" order="Coleoptera" pageId="34" pageNumber="45" phylum="Arthropoda" rank="species" species="persimilis">Griburius persimilis</taxonomicName>
Burmeister, 1877 (BAS).
</paragraph>
</subSubSection>
</treatment>
</document>

121
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<mods:title>Taxonomic study of Baeosega and its allies, with description of a new species of Nipponosega (Hymenoptera, Chrysididae, Amiseginae)</mods:title>
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<mods:namePart>Mita, Toshiharu</mods:namePart>
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<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Kyushu University, Motooka 744, Nishi-ku, Fukuoka 819 - 0395, Japan</mods:affiliation>
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<treatment ID-GBIF-Taxon="182473856" LSID="urn:lsid:plazi:treatment:F177DA38780052F2910DBBE4865EAB2E" httpUri="http://treatment.plazi.org/id/F177DA38780052F2910DBBE4865EAB2E" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="F177DA38-7800-52F2-910D-BBE4865EAB2E" authority="Xu, He &amp; Terayama, 2003" authorityName="Xu, He &amp; Terayama" authorityYear="2003" class="Insecta" family="Chrysididae" genus="Nipponosega" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nipponosega kurzenkoi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kurzenkoi">Nipponosega kurzenkoi Xu, He &amp; Terayama, 2003</taxonomicName>
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<taxonomicName authorityName="Xu, He &amp; Terayama" authorityYear="2003" class="Insecta" family="Chrysididae" genus="Nipponosega" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nipponosega kurzenkoi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kurzenkoi">Nipponosega kurzenkoi</taxonomicName>
Xu, He &amp; Terayama, 2003: 195, holotype ♀ by original designation. Type locality: Mt. Jiulong, Suichang, Zhejiang Province, China (Type no. 944347, no type depository information, but likely Zhejiang University, Hangzhou). Not examined.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Xu, He &amp; Terayama" authorityYear="2003" class="Insecta" family="Chrysididae" genus="Nipponosega" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nipponosega kurzenkoi" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kurzenkoi">
<emphasis italics="true" pageId="0" pageNumber="1">Nipponosega kurzenkoi</emphasis>
</taxonomicName>
is closely related to
<taxonomicName lsidName="N. yamanei" pageId="0" pageNumber="1" rank="species" species="yamanei">
<emphasis italics="true" pageId="0" pageNumber="1">N. yamanei</emphasis>
</taxonomicName>
from Japan. It can be distinguished from
<taxonomicName lsidName="N. yamanei" pageId="0" pageNumber="1" rank="species" species="yamanei">
<emphasis italics="true" pageId="0" pageNumber="1">N. yamanei</emphasis>
</taxonomicName>
by the fully testaceous mesopleuron, whereas it is blackish in
<taxonomicName lsidName="N. yamanei" pageId="0" pageNumber="1" rank="species" species="yamanei">
<emphasis italics="true" pageId="0" pageNumber="1">N. yamanei</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. General habitus of Amiseginae A Nipponosega yamanei Kurzenko &amp; Lelej, holotype ♀ B ditto, ♂ C Okinawasega eguchii Terayama ♀ D ditto, holotype ♂. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1041.66267.figure2" httpUri="https://binary.pensoft.net/fig/550152" pageId="0" pageNumber="1">2A</figureCitation>
). Also, the frons is wider than in
<taxonomicName lsidName="N. yamanei" pageId="0" pageNumber="1" rank="species" species="yamanei">
<emphasis italics="true" pageId="0" pageNumber="1">N. yamanei</emphasis>
</taxonomicName>
. The maximum interocular distance (measured at the lower end of the face) is 1.2
<normalizedToken originalValue="×">x</normalizedToken>
longer than the width of frons (1.5-2.0 in
<taxonomicName lsidName="N. yamanei" pageId="0" pageNumber="1" rank="species" species="yamanei">
<emphasis italics="true" pageId="0" pageNumber="1">N. yamanei</emphasis>
</taxonomicName>
). Other diagnostic characters are as follows: body length 3.0 mm; head black, smooth with scattered punctures; lateral ocelli well separated from the inner margin of eye; mesosoma largely testaceous with mesoscutum and mesoscutellum black; pronotum smooth with scattered punctures; legs yellow; metasoma dark brown. Male is unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">China (Zhejiang).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Although the body color could be variable to some extent, no specimen of
<taxonomicName lsidName="N. yamanei" pageId="0" pageNumber="1" rank="species" species="yamanei">
<emphasis italics="true" pageId="0" pageNumber="1">N. yamanei</emphasis>
</taxonomicName>
with completely testaceous mesopleuron has been found; a closely related species showing different distribution. The morphology of
<taxonomicName lsidName="N. kurzenkoi" pageId="0" pageNumber="1" rank="species" species="kurzenkoi">
<emphasis italics="true" pageId="0" pageNumber="1">N. kurzenkoi</emphasis>
</taxonomicName>
should be evaluated in more detail to discuss their identification. However, diagnostic characters shown above do not overlap with those of
<taxonomicName lsidName="N. yamanei" pageId="0" pageNumber="1" rank="species" species="yamanei">
<emphasis italics="true" pageId="0" pageNumber="1">N. yamanei</emphasis>
</taxonomicName>
. In the original description of
<taxonomicName lsidName="N. kurzenkoi" pageId="0" pageNumber="1" rank="species" species="kurzenkoi">
<emphasis italics="true" pageId="0" pageNumber="1">N. kurzenkoi</emphasis>
</taxonomicName>
<bibRefCitation author="Xu, Z-F" journalOrPublisher="Biogeography" pageId="0" pageNumber="1" refId="B17" refString="Xu, Z-F, He, J-H, Terayama, M, 2003. The genus Nipponosega Kurzenko et Lelej, 1994 firstly recorded from China, with a new species description (Hymenoptera, Chrysididae, Amiseginae) Bulletin de L'Institut Royal des Sciences Naturelles de Belgique, Entomologie 73: 195-196." title="The genus Nipponosega Kurzenko et Lelej, 1994 firstly recorded from China, with a new species description (Hymenoptera, Chrysididae, Amiseginae) Bulletin de L'Institut Royal des Sciences Naturelles de Belgique, Entomologie 73: 195 - 196." year="2003">Xu et al., 2003</bibRefCitation>
, the evaluation of POD is different from the POD (= POL herein) used in
<bibRefCitation author="Nagase, H" journalOrPublisher="Series A, Zoology" pageId="0" pageNumber="1" pagination="103 - 107" refId="B14" refString="Nagase, H, 1995. A new species of Nipponosega (Hymenoptera, Chrysididae, Amiseginae) from central Japan. Bulletin of the National Science Museum. Series A, Zoology 21: 103 - 107" title="A new species of Nipponosega (Hymenoptera, Chrysididae, Amiseginae) from central Japan. Bulletin of the National Science Museum." volume="21" year="1995">Nagase (1995)</bibRefCitation>
. The POD
<emphasis italics="true" pageId="0" pageNumber="1">sensu</emphasis>
<bibRefCitation author="Xu, Z-F" journalOrPublisher="Biogeography" pageId="0" pageNumber="1" refId="B17" refString="Xu, Z-F, He, J-H, Terayama, M, 2003. The genus Nipponosega Kurzenko et Lelej, 1994 firstly recorded from China, with a new species description (Hymenoptera, Chrysididae, Amiseginae) Bulletin de L'Institut Royal des Sciences Naturelles de Belgique, Entomologie 73: 195-196." title="The genus Nipponosega Kurzenko et Lelej, 1994 firstly recorded from China, with a new species description (Hymenoptera, Chrysididae, Amiseginae) Bulletin de L'Institut Royal des Sciences Naturelles de Belgique, Entomologie 73: 195 - 196." year="2003">Xu et al. (2003)</bibRefCitation>
is correspond to OOL herein.
</paragraph>
</subSubSection>
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</document>