diff --git a/data/FE/38/E4/FE38E4545826FFD444FD1E0CFBF0F97F.xml b/data/FE/38/E4/FE38E4545826FFD444FD1E0CFBF0F97F.xml index 706137e7777..8d4e8c0d688 100644 --- a/data/FE/38/E4/FE38E4545826FFD444FD1E0CFBF0F97F.xml +++ b/data/FE/38/E4/FE38E4545826FFD444FD1E0CFBF0F97F.xml @@ -1,59 +1,59 @@ - - - -On the taxonomy of the genus Lethe Hübner (Lepidoptera, Papilionoidea, Nymphalidae, Satyrinae) with description of a new species + + + +On the taxonomy of the genus Lethe Hübner (Lepidoptera, Papilionoidea, Nymphalidae, Satyrinae) with description of a new species - - -Author + + +Author -Huang, Si-Yao -Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauer Allee 127, Bonn 53113, Germany +Huang, Si-Yao +Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauer Allee 127, Bonn 53113, Germany - - -Author + + +Author -Lang, Song-Yun -Chongqing Museum of Natural History, Beibei, 400700, Chongqing, China +Lang, Song-Yun +Chongqing Museum of Natural History, Beibei, 400700, Chongqing, China - - -Author + + +Author -Müller, Günter C. -University of Sciences, Techniques and Technology of Bamako, BP 1805, Bamako, Mali +Müller, Günter C. +University of Sciences, Techniques and Technology of Bamako, BP 1805, Bamako, Mali -text - - -Zootaxa +text + + +Zootaxa - -2024 - -2024-11-13 + +2024 + +2024-11-13 - -5538 + +5538 - -1 + +1 - -59 -73 + +59 +73 - -https://doi.org/10.11646/zootaxa.5538.1.6 + +https://doi.org/10.11646/zootaxa.5538.1.6 -journal article -10.11646/zootaxa.5538.1.6 -1175-5326 -14611457 -80FE2B01-EC06-416C-90C2-712EE95CE2C3 +journal article +10.11646/zootaxa.5538.1.6 +1175-5326 +14611457 +80FE2B01-EC06-416C-90C2-712EE95CE2C3 @@ -122,11 +122,13 @@ Lang, 2016: 342 Material examined: - + Holotype -(fig. 9): male, altitude +(fig. 9): +male +, altitude 3000 m @@ -135,48 +137,45 @@ Lang, 2016: 342 14. VIII. 2014 , -Mt. Luoji +Mt. Luoji , Puge County , -Sichuan Province -, +Sichuan +Province, China , leg. Song-yun Lang -, genitalia preparation -No. +, genitalia preparation No. SATY0246 -, androconia preparation -No. -ANDR0046 -( +, androconia preparation No. ANDR0046 ( CQMNH -); - -holotype - +); + + +holotype of Lethe armandina yanbiana Lang, 2016 (figs. 11, 37): male, -Gesala +Gesala , -Yanbian County +Yanbian County , Sichuan Province , China , leg. Song-yun Lang -, genitalia preparation -No. +, genitalia preparation No. SATY0413 ( CSYL -); +); + + SICHUAN : @@ -187,33 +186,28 @@ Lang, 2016 3000 m , -Gesala -, -Yanbian County -, -Sichuan Province +Gesala , +Yanbian County +, Sichuan Province, China , leg. Song-yun Lang -, genitalia preparation -No. +, genitalia preparation No. SATY0419 ( CSYL -); +); + + YUNNAN : 2 males , -Li-kiang -( -Lijiang -), -Provinz Nord-Yuennan -( +Li-kiang (Lijiang) +, Provinz Nord-Yuennan ( Yunnan ), @@ -222,8 +216,10 @@ Lang, 2016 , H. Höne ( -ZFMK -); +ZFMK +); + + 4 males , 1 female @@ -232,7 +228,7 @@ Lang, 2016 30.7.1935 ( -ZFMK +ZFMK ). diff --git a/data/FE/38/E4/FE38E454582DFFDD44FD1B8DFC01FCD7.xml b/data/FE/38/E4/FE38E454582DFFDD44FD1B8DFC01FCD7.xml new file mode 100644 index 00000000000..81ee5c655aa --- /dev/null +++ b/data/FE/38/E4/FE38E454582DFFDD44FD1B8DFC01FCD7.xml @@ -0,0 +1,604 @@ + + + +On the taxonomy of the genus Lethe Hübner (Lepidoptera, Papilionoidea, Nymphalidae, Satyrinae) with description of a new species + + + +Author + +Huang, Si-Yao +Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauer Allee 127, Bonn 53113, Germany + + + +Author + +Lang, Song-Yun +Chongqing Museum of Natural History, Beibei, 400700, Chongqing, China + + + +Author + +Müller, Günter C. +University of Sciences, Techniques and Technology of Bamako, BP 1805, Bamako, Mali + +text + + +Zootaxa + + +2024 + +2024-11-13 + + +5538 + + +1 + + +59 +73 + + + + +https://doi.org/10.11646/zootaxa.5538.1.6 + +journal article +10.11646/zootaxa.5538.1.6 +1175-5326 +14611457 +80FE2B01-EC06-416C-90C2-712EE95CE2C3 + + + + + + + +Lethe lyncus +de Nicéville, 1897 + + + + + + + +Figs 21–27 +, +41–44 +, +54 +, +60 + + + + + + + +Lethe lyncus +de Nicéville, 1897: 544 + + +, pl. 1, fig. 8; + +Tytler 1914: 222 + +; H. + +Huang 1999: 130 + +; + +Lang 2016b: 216 + +, fig. 4; + +Lang 2017: 49 + +. + + + + + +Lethe tristigmata lyncus + +: + +Fruhstorfer 1911: 313 + +. + + + + + +Lethe ocellata lyncus + +: + +Evans 1923: 532 + +; + +Talbot 1947: 181 + +. + + + + + + +Lethe wui +H. +Huang, 1999: 129 + + +; figs 1a, 2A, 3A; + +Lang 2016b: 216 + +, figs 5, 6, 25, 28b; + +Lang 2017: 49 + +. +syn. nov. + + + + + +FIGURES 37–40. +Male genitalia of + +Lethe jianqingi + +and + +L. armandina + +sspp. Depository of the genitalia preparation: 37 & 40 in CQMNH; 38 & 39 in CSYL. a=genitalia capsule with left valva removed; b=lateral view of the phallus; c=dorsal view of the distal end of left valva + + + + +Material examined: + + +INDIA +: + +1 male +, +Sikkim +, (18)86, +Möll(er) +. (leg.), genitalia & androconia preparation No. +MfNLe001 +( +MfN +) + +; + +1male +, +Sikkim +,Gantok( +Gangtok +), + +VIII.1891 + +, +G.C.Dudgeon +leg.,1905–259, +NHMUK015107301 +( +NHMUK +) + +; + + +BHUTAN +: + +1 male +, +Bhutan +, +Bela la +, 10000 (ft), 1. 7. (19)22, +Bailey +(leg.), +NHMUK015107302 +( +NHMUK +) + +; + + +CHINA + +: + +holotype + +of + +Lethe wui +H. +Huang, 1999 + +(figs. 20, 35): male, handwritten blue label “ +Tibet +: +Metok +( +Motuo County +) +Lage +, + +2700 m + +. + +1995-VIII.24 + +H. Huang +(leg.)”; white handwritten label “ +ī̎-ŝö +(Hanmi-Lage) + +95-8-24 + +”; red handwritten label “HOLOTYPE + +Lethe wui +Huang + +” ( +CQEC +) + +; + +2 males +, altitude + +2800 m + +, + +24. VII. 2015 + +, +Yunnan +, +Gongshan +, +East slope of Mts. Gaoligong +, leg. +Song-yun Lang +, genitalia preparation No. +SATY0359 + +, + +androconia preparation No. +ANDR0069 +( +CSYL +) + +. + + + + +FIGURES 41–46. +Male genitalia of + +Lethe +spp. + +Depository of the genitalia preparation: 41 & 42 in MfN; 43 & 46 in CSYL; 44 in CQEC; 45 in ZFMK. a=genitalia capsule with left valva removed; b=lateral view of the phallus; c=dorsal view of the distal end of left valva. Scale bar for all figures except Fig. 44. + + + + +FIGURES 47–56. +Dorsal view of the uncus and tegumen of the male genitalia of + +Lethe +spp. + +Depository of the genitalia preparation: 47 in CGM; 48, 50, 52, 54b & 56 in CSYL; 49, 51 & 53 in CQMNH; 54a in MfN; 55 in ZFMK. + + + + +Diagnosis. + +Lethe lyncus + +can be distinguished from + +L. tristigmata +Elwes, 1887 + +(figs. 28, 29, 45, 55, 61) and + +L. ocellata +( +Poujade, 1885 +) + +(figs. 30, 46, 56, 63) by the combination of the following characters: 1) The subapical spots on the forewing underside are tightly conjoined from space 6 to 8 as in + +L. ocellata + +, while in + +L. tristigmata + +the subapical series of spots are widely separated from each other and situated from space 3 to space 7. 2) The pale margin next to the dark postdiscal band on the forewing underside is well-developed, while in + +L. tristigmata + +, it is less developed and in + +L. ocellata + +it is totally absent. 3) On the forewing underside, the area between the two dark cell bars of the discal cell is lighter and brighter than the remaining parts, similar to that in + +L. tristigmata + +, while in + +L. ocellata + +the whole discal cell is uniform in coloration. 4) In male genitalia, the uncus in lateral view is much wider than that in + +L. tristigmata + +and + +L. ocellata + +. 5) The uncus in dorsal view is widened in the medial section as in + +L. ocellata + +, while in + +L. tristigmata + +the uncus is generally even in width from the medial to distal section. 6) The gnathos are thicker than those of + +L. ocellata + +and sinuated, similar to those in + +L. tristigmata + +. + + + +FIGURES 57–63. +Androconia of + +Lethe +spp. + +Depository of the androconia preparation: 57 in CGM; 58, 59, 60b, 62 in CSYL; 60a in MfN; 61 in ZFMK; 63 in CQMNH; + + + + +Lethe lyncus + +also somewhat resembles + +L. armandina + +but can be distinguished by lacking the purplish tint from basal to postmedial zone on the underside of hindwing, the shorter androconia and the broader basal section of uncus in the male genitalia in lateral view. + + + + +Discussion. +Since the original description, the status of this rare taxon has been uncertain for a long time. +Fruhstorfer (1911) +treated this taxon as a subspecies of + +L. tristigmata + +. Later both +Evans (1923) +and +Talbot (1947) +considered it as a subspecies of + +L. ocellata +, + +and subsequently H. +Huang (1999) +regarded it as a synonym of + +L. tristigmata + +. Recently, the senior author examined the specimens of this species held in MfN and NHMUK, and the study of the male genitalia as well as the androconia led to the conclusion that + +L. lyncus + +is a bona species different from both + +L. tristigmata + +and + +L. ocellata + +, and the differences are summarized above. + + +The descriptions of the male brand in this species differ from each other in several works. Although, in the original description, +de Nicéville (1897) +mentioned that the male brand was “barely traceable”, +Evans (1923) +and +Talbot (1947) +both stated that + +L. lyncus + +bore “broken brand from mid v1 to base v4” or “interrupted male brand”, respectively. During the course of the current study, we examined +three males +of this species together with the +syntype +photos published in + +Sheela +et al. +(2019) + +( +Fig. 21 +), and we found that the male brand of this species varied from vestigial ( +Fig. 22 +) to relatively prominent ( +Fig. 24 +) with intermediate individuals connecting both forms ( +Fig. 23 +). Therefore, it is probable that the different descriptions of the male brand in historical literatures are caused by the different individuals examined by the authors. + + + +Lethe wui +H. +Huang, 1999 + +is sunk as a synonym of + +L. lyncus + +in the present study. Since the original description, + +L. wui + +was only known from a few individuals captured in +Xizang +and +Yunnan +in +China +as well as +Kachin State +in northern +Myanmar +( +Koiwaya & Shizuya 2011 +), and the relationship between it and + +L. lyncus + +remains unclear for the lack of available material of the latter species. +Lang (2016b +, +2017 +) indicated that + +L. wui + +was probably only a subspecies or synonym of + +L. lyncus + +due to the similarity in wing pattern. After examining the specimens of + +L. lyncus + +, it turned out that the male genitalia as well as the androconia of + +L. wui + +did not show significant difference from those of + +L. lyncus + +. The male brand in + +L. wui + +is also variable ( +Figs 25–27 +), and, taking the specimens of + +L. lyncus + +examined in the present study into account, all the individuals together form a complete chain of variation in male brand from vestigial to well-developed, same as the situation of + +L. jianqingi + +and + +L. armandina yanbiana + +. Therefore, + +L. wui + +should be synonymized with + +L. lyncus + +( +syn. nov. +). + + + + +Distribution. +northeastern +India +( +Sikkim +, +Manipur +), western +Bhutan +, northern +Myanmar +( +Kachin state +), southwestern +China +(southeastern +Xizang +, western +Yunnan +) ( +Fig. 55 +). + + + + \ No newline at end of file