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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.88.807" ID-GBIF-Dataset="1858994c-378e-4e6e-bec9-ee565e3b3be1" ID-Pensoft-Pub="1313-2970-88-1" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-88-1" ModsDocOrigin="ZooKeys 88" ModsDocTitle="Family-group names in Coleoptera (Insecta)" checkinTime="1451250740194" checkinUser="pensoft" docAuthor="Bouchard, Patrice, Bousquet, Yves, Davies, Anthony E., Alonso-Zarazaga, Miguel A., Lawrence, John F., Lyal, Chris H. C., Newton, Alfred F., Reid, Chris A. M., Schmitt, Michael, Ślipinski, S. Adam &amp; Smith, Andrew B. T." docDate="2011" docId="34442A7609D48BE40D2CB9F83A0C587F" docLanguage="en" docName="ZooKeys 88: 1-972" docOrigin="ZooKeys 88" docSource="http://dx.doi.org/10.3897/zookeys.88.807" docTitle="Trachykelina Holynski 1988" docType="treatment" docVersion="6" lastPageNumber="140" masterDocId="FFC1FFDA4E1CE6158E11FFCD0115FF8E" masterDocTitle="Family-group names in Coleoptera (Insecta)" masterLastPageNumber="972" masterPageNumber="1" pageNumber="140" updateTime="1661732154594" updateUser="admin">
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<mods:namePart>Bouchard, Patrice</mods:namePart>
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<mods:namePart>Davies, Anthony E.</mods:namePart>
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<mods:namePart>Alonso-Zarazaga, Miguel A.</mods:namePart>
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<mods:namePart>Lawrence, John F.</mods:namePart>
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<mods:namePart>Lyal, Chris H. C.</mods:namePart>
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<mods:namePart>Newton, Alfred F.</mods:namePart>
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<mods:namePart>Reid, Chris A. M.</mods:namePart>
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<mods:namePart>Schmitt, Michael</mods:namePart>
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<mods:namePart>Ślipinski, S. Adam</mods:namePart>
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<mods:namePart>Smith, Andrew B. T.</mods:namePart>
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Subtribe
<taxonomicName authority="Holynski, 1988" authorityName="Holynski" authorityYear="1988" lsidName="Trachykelina" pageId="139" pageNumber="140" rank="tribe" tribe="Trachykelina">
Trachykelina
<normalizedToken originalValue="Holyński">Holynski</normalizedToken>
, 1988
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<paragraph pageId="139" pageNumber="140">
<taxonomicName genus="Trachykelina" lsidName="Trachykelina" pageId="139" pageNumber="140" rank="genus">Trachykelina</taxonomicName>
<normalizedToken originalValue="Holyński">Holynski</normalizedToken>
, 1988: 51, in key [stem: Trachykel-]. Type genus:
<taxonomicName class="Insecta" family="Buprestidae" genus="Trachykele" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trachykele" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="genus">Trachykele</taxonomicName>
Marseul, 1865.
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<emphasis id="8E99EAB6FFC1FFAA5FBCFCBEFD52FC35" bold="true" box="[444,713,884,909]" italics="true" pageId="10" pageNumber="2335">Marcusenius altisambesi</emphasis>
<bibRefCitation id="D87C4B55FFC1FFAA5CD1FCBDFC31FC35" author="Kramer B &amp; Skelton PH &amp; Van der Bank FH &amp; Wink M" box="[721,938,885,909]" pageId="10" pageNumber="2335" pagination="647 - 708" refId="ref16697" refString="Kramer B, Skelton PH, Van der Bank FH, Wink M. 2007. Allopatric differentiation in the Marcusenius macrolepidotus species complex in southern and eastern Africa: the resurrection of M. pongolensis and M. angolensis, and the description of two new species (Mormyridae, Teleostei). J Nat Hist. 41: 647 - 708." type="journal article" year="2007">Kramer et al., 2007</bibRefCitation>
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(
<figureCitation id="24D62A21FFC1FFAA5C69FC5EFD66FC15" box="[617,765,917,941]" captionStart="Figure 4" captionStartId="11.[143,210,1506,1528]" captionTargetBox="[168,1081,161,1473]" captionTargetId="figure-6@11.[168,1081,161,1473]" captionTargetPageId="11" captionText="Figure 4. Photographs of bulldog fish from various origins. (A) Marcusenius macrolepidotus (Peters, 1852), SAIAB 60847, coll. R. Bills 1 August 1999, Lower Zambezi; (B) Marcusenius angolensis (Boulenger, 1905), BMNH 1905.5.29.64 (holotype); (C) Marcusenius altisambesi, coll. F. H. van der Bank and B. Kramer, 21 August 1999, Upper Zambezi, Kalimbeza, live specimen of 16.5 cm SL photographed 3 July 2003 in aquarium; (D) M. altisambesi, coll. F.H. van der Bank and B. Kramer, 11/12 August 2004, Okavango River, live fish of SL 13 cm photographed 20 April 2006 in aquarium; (E) Marcusenius multisquamatus sp. nov., coll. B. Kramer and Ernst Swartz, coll. and photographed on 19 August 2006, Cunene River, below Ruacana Falls, specimen RUAC01, SL 15.4 cm." figureDoi="http://doi.org/10.5281/zenodo.5197598" httpUri="https://zenodo.org/record/5197598/files/figure.png" pageId="10" pageNumber="2335">Figure 4C, D</figureCitation>
)
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<taxonomicName id="7BED4D27FFC1FFAA5ED3FC1EFD47FC54" ID-CoL="6KNGJ" authority="Pappenheim, 1907" authorityName="Pappenheim" authorityYear="1907" box="[211,732,980,1005]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="10" pageNumber="2335" phylum="Chordata" rank="species" species="okavangensis">
<emphasis id="8E99EAB6FFC1FFAA5ED3FC1EFD9AFC55" box="[211,513,981,1005]" italics="true" pageId="10" pageNumber="2335">Gnathonemus okavangensis</emphasis>
<bibRefCitation id="D87C4B55FFC1FFAA5C09FC1FFD47FC54" author="Pappenheim P" box="[521,732,980,1004]" pageId="10" pageNumber="2335" pagination="339 - 367" refId="ref17154" refString="Pappenheim P. 1907. Zur Systematik und Variationsstatistik der Mormyriden hauptsachlich aus den deutsch-afrikanischen Schutzgebieten. Nach dem Material des Koniglichen Zoologischen Museums zu Berlin. Mitteilungen aus dem Zoologischen Museum in Berlin. 3: 339 - 367." type="journal article" year="1907">Pappenheim, 1907</bibRefCitation>
</taxonomicName>
. “Appeared as a form of
<taxonomicName id="7BED4D27FFC1FFAA5DFDFC1EFE72FBB4" ID-CoL="3GLTR" authority="Peters" authorityName="Peters" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="10" pageNumber="2335" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFC1FFAA5DFDFC1EFE02FBB5" italics="true" pageId="10" pageNumber="2335">Gnathonemus macrolepidotus</emphasis>
Peters
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from the
<collectingRegion id="7E29F846FFC1FFAA5C65FC3FFD43FBB4" box="[613,728,1012,1036]" country="Namibia" name="Kavango West" pageId="10" pageNumber="2335">Okavango</collectingRegion>
R., Damaraland, Africa; regarded as infrasubspecific and not available” (Eschmeyer 2013). “Nomen dubium” according to
<bibRefCitation id="D87C4B55FFC1FFAA5F11FBFFFE38FBF4" author="Gosse J-P" box="[273,419,1076,1100]" pageId="10" pageNumber="2335" pagination="63 - 124" refId="ref16269" refString="Gosse J-P. 1984. Mormyriformes. In: Daget J, Gosse J-P, Thys van den Audenaerde DFE, editors. Check-list of the freshwater fishes of Africa. Bondy (France) / Tervuren. Belgium: ORSTOM / MRAC; p. 63 - 124." type="book chapter" year="1984">Gosse (1984)</bibRefCitation>
and
<bibRefCitation id="D87C4B55FFC1FFAA5FDBFBFFFDE8FBF4" author="Seegers L" box="[475,627,1076,1100]" pageId="10" pageNumber="2335" pagination="1 - 407" refId="ref17257" refString="Seegers L. 1996. The fishes of the Lake Rukwa drainage. Annales Sciences Zoologiques (Tervuren). 278: 1 - 407." type="journal article" year="1996">Seegers (1996</bibRefCitation>
, p. 73).
</paragraph>
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<taxonomicName id="7BED4D27FFC1FFAA5ED3FB9FFB94FBD4" ID-CoL="3GLTR" authority=": Gilchrist and Thompson 1913, pp. 330 - 331" authorityName="Gilchrist and Thompson" authorityPageNumber="330 - 331" authorityYear="1913" box="[211,1039,1108,1132]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="10" pageNumber="2335" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFC1FFAA5ED3FB9FFD8DFBD4" box="[211,534,1108,1132]" italics="true" pageId="10" pageNumber="2335">Gnathonemus macrolepidotus</emphasis>
:
<bibRefCitation id="D87C4B55FFC1FFAA5C24FB9FFCEDFBD4" author="Gilchrist JDF &amp; Thompson WW" box="[548,886,1108,1132]" pageId="10" pageNumber="2335" pagination="321 - 463" refId="ref16195" refString="Gilchrist JDF, Thompson WW. 1913. The freshwater fishes of South Africa. Annals of the South African Museum (Cape Town, etc). XI (V): 321 - 463." type="journal article" year="1913">Gilchrist and Thompson 1913</bibRefCitation>
, pp. 330–331
</taxonomicName>
.
</paragraph>
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<taxonomicName id="7BED4D27FFC1FFAA5ED3FBBEFD57FB34" ID-CoL="3Y375" authority="Kramer et al. (2007)" authorityName="Kramer" authorityYear="2007" box="[211,716,1140,1164]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="10" pageNumber="2335" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFC1FFAA5ED3FBBEFE7BFB34" box="[211,480,1140,1164]" italics="true" pageId="10" pageNumber="2335">Marcusenius altisambesi</emphasis>
<bibRefCitation id="D87C4B55FFC1FFAA5FE7FBBEFD57FB34" author="Kramer B &amp; Skelton PH &amp; Van der Bank FH &amp; Wink M" box="[487,716,1140,1164]" pageId="10" pageNumber="2335" pagination="647 - 708" refId="ref16697" refString="Kramer B, Skelton PH, Van der Bank FH, Wink M. 2007. Allopatric differentiation in the Marcusenius macrolepidotus species complex in southern and eastern Africa: the resurrection of M. pongolensis and M. angolensis, and the description of two new species (Mormyridae, Teleostei). J Nat Hist. 41: 647 - 708." type="journal article" year="2007">Kramer et al. (2007)</bibRefCitation>
</taxonomicName>
, pp. 681–684.
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<emphasis id="8E99EAB6FFC1FFAA5ED3FB1CFEE5FB56" box="[211,382,1239,1262]" italics="true" pageId="10" pageNumber="2335">Type specimens</emphasis>
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</paragraph>
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<materialsCitation id="0C853CF9FFC1FFAA5ED3FB35FEE7FA8E" collectionCode="SAIAB" country="Namibia" location="Lisikili on Upper" municipality="Strip" pageId="10" pageNumber="2335" specimenCode="SAIAB 79135" specimenCount="1" stateProvince="Zambezi" typeStatus="holotype">
<typeStatus id="63568806FFC1FFAA5ED3FB35FED9FAAE" box="[211,322,1278,1302]" pageId="10" pageNumber="2335" type="holotype">Holotype</typeStatus>
:
<specimenCode id="EC4B9EDFFFC1FFAA5F4CFB35FE73FAAE" box="[332,488,1278,1302]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="10" pageNumber="2335" type="Other type of research institution/biorepository">SAIAB 79135</specimenCode>
(specimen L39isi),
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:
<collectingRegion id="7E29F846FFC1FFAA5D32FB35FC1DFAAE" box="[818,902,1278,1302]" country="Namibia" name="Zambezi" pageId="10" pageNumber="2335">Caprivi</collectingRegion>
<collectingMunicipality id="5C36ACDEFFC1FFAA5D8CFB35FC5CFAAE" box="[908,967,1278,1302]" pageId="10" pageNumber="2335">Strip</collectingMunicipality>
:
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<collectingRegion id="7E29F846FFC1FFAA5ED3FAD5FEE7FA8E" box="[211,380,1310,1334]" country="Namibia" name="Zambezi" pageId="10" pageNumber="2335">Zambezi River</collectingRegion>
</materialsCitation>
.
<materialsCitation id="0C853CF9FFC1FFAA5F86FAD4FD5DFA8E" box="[390,710,1310,1335]" collectionCode="SAIAB" country="Namibia" pageId="10" pageNumber="2335" specimenCode="SAIAB 79136" specimenCount="1" stateProvince="Zambezi" typeStatus="paratype">
<typeStatus id="63568806FFC1FFAA5F86FAD4FE63FA8F" box="[390,504,1311,1335]" pageId="10" pageNumber="2335" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="EC4B9EDFFFC1FFAA5C02FAD5FD05FA8E" box="[514,670,1310,1334]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="10" pageNumber="2335" type="Other type of research institution/biorepository">SAIAB 79136</specimenCode>
(6)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC1FFAA5CD2FAD5FC0EFA8E" box="[722,917,1310,1334]" collectionCode="SAIAB" country="Namibia" pageId="10" pageNumber="2335" specimenCode="SAIAB 79137" specimenCount="1" stateProvince="Zambezi" typeStatus="paratype">
<specimenCode id="EC4B9EDFFFC1FFAA5CD2FAD5FCF5FA8E" box="[722,878,1310,1334]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="10" pageNumber="2335" type="Other type of research institution/biorepository">SAIAB 79137</specimenCode>
(3)
</materialsCitation>
,
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35086 (5)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC1FFAA5A59FAD5FEA5FAEE" accessionNumber="ZSM 35085" collectionCode="ZSM" country="Namibia" pageId="10" pageNumber="2335" specimenCount="1" stateProvince="Zambezi" typeStatus="paratype">
<collectionCode id="DAFCAE61FFC1FFAA5A59FAD5FB0FFA8E" box="[1113,1172,1310,1334]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="10" pageNumber="2335" type="Museum">ZSM</collectionCode>
35085 (2)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC1FFAA5F49FAF5FE6FFAEE" accessionNumber="ZSM 35097" box="[329,500,1342,1366]" collectionCode="ZSM" country="Namibia" pageId="10" pageNumber="2335" specimenCount="1" stateProvince="Zambezi" typeStatus="paratype">
<collectionCode id="DAFCAE61FFC1FFAA5F49FAF5FE1FFAEE" box="[329,388,1342,1366]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="10" pageNumber="2335" type="Museum">ZSM</collectionCode>
35097 (1)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC1FFAA5C00FAF5FD37FAEE" accessionNumber="ZSM 35082" box="[512,684,1342,1366]" collectionCode="ZSM" country="Namibia" pageId="10" pageNumber="2335" specimenCount="1" stateProvince="Zambezi" typeStatus="paratype">
<collectionCode id="DAFCAE61FFC1FFAA5C00FAF5FDA0FAEE" box="[512,571,1342,1366]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="10" pageNumber="2335" type="Museum">ZSM</collectionCode>
35082 (2)
</materialsCitation>
; all examined.
</paragraph>
<paragraph id="BC5236A4FFC1FFAA5EFEFA48FD46FA63" blockId="10.[254,1172,1411,1690]" pageId="10" pageNumber="2335">
<emphasis id="8E99EAB6FFC1FFAA5EFEFA48FE90FA23" bold="true" box="[254,267,1411,1435]" pageId="10" pageNumber="2335">–</emphasis>
<materialsCitation id="0C853CF9FFC1FFAA5F1CFA4FFD42FA63" country="Namibia" county="Aquarium" location="Upper" municipality="Kalimbeza" pageId="10" pageNumber="2335" specimenCount="1" stateProvince="Zambezi">
Non-types, examined. One hundred and
<specimenCount id="AAEBFD2DFFC1FFAA5D11FA48FC59FA23" box="[785,962,1411,1435]" pageId="10" pageNumber="2335" type="generic">four specimens</specimenCount>
from the
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<collectingRegion id="7E29F846FFC1FFAA5F1CFA68FE4AFA03" box="[284,465,1443,1467]" country="Namibia" name="Zambezi" pageId="10" pageNumber="2335">Zambezi River</collectingRegion>
<location id="B932607FFFC1FFAA5FE5FA68FDA2FA03" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC1FFAA5FE5FA68FDA2FA03" box="[485,569,1443,1467]" country="Namibia" county="Aquarium" municipality="Kalimbeza" name="System" pageId="10" pageNumber="2335" stateProvince="Zambezi">System</location>
,
<location id="B932607FFFC1FFAA5C52FA6FFD1FFA03" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC1FFAA5C52FA6FFD1FFA03" box="[594,644,1444,1467]" country="Namibia" county="Aquarium" municipality="Kalimbeza" name="East" pageId="10" pageNumber="2335" stateProvince="Zambezi">East</location>
<collectingRegion id="7E29F846FFC1FFAA5C98FA68FD74FA03" box="[664,751,1443,1467]" country="Namibia" name="Zambezi" pageId="10" pageNumber="2335">Caprivi</collectingRegion>
,
<collectingCountry id="C4FA7634FFC1FFAA5D08FA68FCF5FA03" box="[776,878,1443,1467]" name="Namibia" pageId="10" pageNumber="2335">Namibia</collectingCountry>
, some specimens from
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presently alive in
<collectingCounty id="55334E28FFC1FFAA5C60FA08FD42FA63" box="[608,729,1475,1499]" pageId="10" pageNumber="2335">Aquarium</collectingCounty>
</materialsCitation>
:
</paragraph>
<paragraph id="BC5236A4FFC1FFAA5EFEFA28FE42F982" blockId="10.[254,1172,1411,1690]" pageId="10" pageNumber="2335">
<emphasis id="8E99EAB6FFC1FFAA5EFEFA28FE90FA43" bold="true" box="[254,267,1507,1531]" pageId="10" pageNumber="2335">–</emphasis>
<materialsCitation id="0C853CF9FFC1FFAA5F1CFA28FE4EF982" accessionNumber="SMF 28264" collectingDate="1994-03-05" collectingDateMax="1994-03-07" collectingDateMin="1994-03-05" collectionCode="SMF" collectorName="F. H. van der Bank &amp; B. Kramer" latitude="-17.55" location="Lisikili" longLatPrecision="1278" longitude="24.483334" pageId="10" pageNumber="2335" specimenCount="22" stateProvince="Zambezi">
<collectionCode id="DAFCAE61FFC1FFAA5F1CFA28FECCFA43" box="[284,343,1507,1531]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="10" pageNumber="2335">SMF</collectionCode>
28264 (
<specimenCount id="AAEBFD2DFFC1FFAA5FC1FA28FDF9FA43" box="[449,610,1507,1531]" pageId="10" pageNumber="2335" type="generic">22 specimens</specimenCount>
), from the
<collectingRegion id="7E29F846FFC1FFAA5CF7FA28FC33FA43" box="[759,936,1507,1531]" country="Namibia" name="Zambezi" pageId="10" pageNumber="2335">Zambezi River</collectingRegion>
,
<location id="B932607FFFC1FFAA5DBCFA28FB90FA43" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC1FFAA5DBCFA28FB90FA43" box="[956,1035,1507,1531]" latitude="-17.55" longLatPrecision="1278" longitude="24.483334" name="Lisikili" pageId="10" pageNumber="2335" stateProvince="Zambezi">Lisikili</location>
backwater,
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17
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33
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S
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,
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24
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29
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E
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(
<typeStatus id="63568806FFC1FFAA5C04F9C9FDADF9A2" box="[516,566,1538,1562]" pageId="10" pageNumber="2335">type</typeStatus>
locality), coll.:
<collectorName id="11185372FFC1FFAA5CF3F9C8FC4AF9A2" box="[755,977,1538,1562]" pageId="10" pageNumber="2335">F.H. van der Bank</collectorName>
and
<collectorName id="11185372FFC1FFAA5A13F9C8FB0BF9A2" box="[1043,1168,1539,1562]" pageId="10" pageNumber="2335">B. Kramer</collectorName>
,
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<collectingDate id="D817E98CFFC1FFAA5F1CF9E9FE4EF982" box="[284,469,1570,1594]" pageId="10" pageNumber="2335" value="1994-03-05" valueMax="1994-03-07" valueMin="1994-03-05">5–7 March 1994</collectingDate>
</date>
</materialsCitation>
,
</paragraph>
<paragraph id="BC5236A4FFC1FFAA5EFEF989FDCDF922" blockId="10.[254,1172,1411,1690]" pageId="10" pageNumber="2335">
<emphasis id="8E99EAB6FFC1FFAA5EFEF989FE90F9E2" bold="true" box="[254,267,1602,1626]" pageId="10" pageNumber="2335">–</emphasis>
<materialsCitation id="0C853CF9FFC1FFAA5F1CF989FDD5F9E2" accessionNumber="SMF 28264" box="[284,590,1602,1626]" collectionCode="SMF" pageId="10" pageNumber="2335" specimenCount="45">
<collectionCode id="DAFCAE61FFC1FFAA5F1CF989FECCF9E2" box="[284,343,1602,1626]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="10" pageNumber="2335">SMF</collectionCode>
28264 (
<specimenCount id="AAEBFD2DFFC1FFAA5FAEF989FDDDF9E2" box="[430,582,1602,1626]" pageId="10" pageNumber="2335" type="generic">45 specimens</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC1FFAA5C59F989FC98F9E2" accessionNumber="ZSM 35084" box="[601,771,1602,1626]" collectionCode="ZSM" pageId="10" pageNumber="2335" specimenCount="1">
<collectionCode id="DAFCAE61FFC1FFAA5C59F989FD0FF9E2" box="[601,660,1602,1626]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="10" pageNumber="2335" type="Museum">ZSM</collectionCode>
35084 (1)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC1FFAA5D0FF989FDD4F922" collectingDate="1994-03-09" collectingDateMax="1994-03-15" collectingDateMin="1994-03-09" collectionCode="ZSM" collectorName="B. Kramer" county="Mudumu National Park" latitude="-18.1" location="Kwando River" longLatPrecision="1276" longitude="23.383333" municipality="Nakatwa" pageId="10" pageNumber="2335" specimenCount="1">
from the
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,
<collectingMunicipality id="5C36ACDEFFC1FFAA5A25F989FB14F9E2" box="[1061,1167,1602,1626]" pageId="10" pageNumber="2335">Nakatwa</collectingMunicipality>
,
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18
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06
<superScript id="4B989BECFFC1FFAA5F5CF995FEFAF9D7" attach="left" box="[348,353,1630,1647]" fontSize="7" pageId="10" pageNumber="2335">′</superScript>
S
</geoCoordinate>
,
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23
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23
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E
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, in
<collectingCounty id="55334E28FFC1FFAA5C1AF9A9FCA8F9C2" box="[538,819,1634,1658]" pageId="10" pageNumber="2335">Mudumu National Park</collectingCounty>
, coll.:
<collectorName id="11185372FFC1FFAA5D81F9A8FC67F9C2" box="[897,1020,1635,1658]" pageId="10" pageNumber="2335">B. Kramer</collectorName>
,
<date id="C8531064FFC1FFAA5A09F9A9FECFF922" pageId="10" pageNumber="2335" value="1994-03-09" valueMax="1994-03-15" valueMin="1994-03-09">
<collectingDate id="D817E98CFFC1FFAA5A09F9A9FECFF922" pageId="10" pageNumber="2335" value="1994-03-09" valueMax="1994-03-15" valueMin="1994-03-09">9–15 March 1994</collectingDate>
</date>
, locality 6 on
<figureCitation id="24D62A21FFC1FFAA5FF1F949FDD4F922" box="[497,591,1666,1690]" captionStart="Figure 1" captionStartId="4.[211,278,932,954]" captionTargetBox="[213,1170,167,899]" captionTargetId="figure-5@4.[213,1170,167,899]" captionTargetPageId="4" captionText="Figure 1. Map of sampling locations in southern Africa. (A) Type locality for Marcusenius macrolepidotus (Peters, 1852), Tete on the Lower Zambezi River; (B) origin of M. macrolepidotus, SAIAB 60847, from Marromeu, Lower Zambezi delta (Figure 4A); (C) origin of M. macrolepidotus from the Buzi System, same location as SAIAB 67369, presently alive in aquarium; (D) likely type region for Marcusenius angolensis (Boulenger, 1905), the Lower Quanza; (E) Marcusenius altisambesi Kramer et al., 2007, type locality on the Upper Zambezi River; (F) M. altisambesi from the Kwando River; (G) M. altisambesi from the Okavango delta; (H, I) Marcusenius multisquamatus sp. nov., stretching from below the Ruacana Falls to above the Epupa Falls (I), the type locality; (J) specimens from the Cunene River Mouth." figureDoi="http://doi.org/10.5281/zenodo.5197592" httpUri="https://zenodo.org/record/5197592/files/figure.png" pageId="10" pageNumber="2335">Figure 1</figureCitation>
</materialsCitation>
,
</paragraph>
<caption id="E892662CFFC0FFAB5E8FFA29FD8BF95A" ID-DOI="http://doi.org/10.5281/zenodo.5197598" ID-Zenodo-Dep="5197598" httpUri="https://zenodo.org/record/5197598/files/figure.png" pageId="11" pageNumber="2336" startId="11.[143,210,1506,1528]" targetBox="[168,1081,161,1473]" targetPageId="11">
<paragraph id="BC5236A4FFC0FFAB5E8FFA29FD8BF95A" blockId="11.[143,1104,1506,1762]" pageId="11" pageNumber="2336">
Figure 4. Photographs of bulldog fish from various origins. (A)
<taxonomicName id="7BED4D27FFC0FFAB5D33FA28FE86F9AE" authority="(Peters, 1852)" baseAuthorityName="Peters" baseAuthorityYear="1852" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="11" pageNumber="2336" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFC0FFAB5D33FA28FBD4FA41" box="[819,1103,1507,1529]" italics="true" pageId="11" pageNumber="2336">Marcusenius macrolepidotus</emphasis>
(
<bibRefCitation id="D87C4B55FFC0FFAB5E96F9CBFE8EF9AE" author="Peters WCH" box="[150,277,1536,1558]" pageId="11" pageNumber="2336" pagination="273 - 276" refId="ref17195" refString="Peters WCH. 1852. Einige neue Saugethiere und Flussfische aus Mossambique. Monatsber Konigl Preuss Akad Wiss Berlin. 1852: 273 - 276." type="journal article" year="1852">Peters, 1852</bibRefCitation>
)
</taxonomicName>
, SAIAB 60847, coll. R. Bills 1 August 1999, Lower Zambezi; (B)
<taxonomicName id="7BED4D27FFC0FFAB5DD2F9CAFE37F98B" authority="(Boulenger, 1905)" baseAuthorityName="Boulenger" baseAuthorityYear="1905" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="11" pageNumber="2336" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="8E99EAB6FFC0FFAB5DD2F9CAFF6FF98B" italics="true" pageId="11" pageNumber="2336">Marcusenius angolensis</emphasis>
(
<bibRefCitation id="D87C4B55FFC0FFAB5F00F9D6FE3EF98B" author="Boulenger GA" box="[256,421,1565,1587]" pageId="11" pageNumber="2336" pagination="457 - 459" refId="ref15927" refString="Boulenger GA. 1905. Descriptions of four new freshwater fishes discovered by Dr. W. J. Ansorge in Angola. Ann Mag Nat Hist (Ser. 7). 15: 457 - 459." type="journal article" year="1905">Boulenger, 1905</bibRefCitation>
)
</taxonomicName>
, BMNH 1905.5.29.64 (holotype); (C)
<taxonomicName id="7BED4D27FFC0FFAB5D27F9D5FB82F98B" box="[807,1049,1565,1587]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="11" pageNumber="2336" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFC0FFAB5D27F9D5FB82F98B" box="[807,1049,1565,1587]" italics="true" pageId="11" pageNumber="2336">Marcusenius altisambesi</emphasis>
</taxonomicName>
, coll. F. H. van der Bank and B. Kramer, 21 August 1999, Upper Zambezi, Kalimbeza, live specimen of 16.5 cm SL photographed 3 July 2003 in aquarium; (D)
<taxonomicName id="7BED4D27FFC0FFAB5D46F993FC46F9D6" box="[838,989,1624,1646]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="11" pageNumber="2336" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFC0FFAB5D46F993FC46F9D6" box="[838,989,1624,1646]" italics="true" pageId="11" pageNumber="2336">M. altisambesi</emphasis>
</taxonomicName>
, coll. F.H. van der Bank and B. Kramer, 11/12 August 2004, Okavango River, live fish of SL 13 cm photographed 20 April 2006 in aquarium; (E)
<taxonomicName id="7BED4D27FFC0FFAB5C38F958FCCCF910" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[568,855,1682,1704]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="11" pageNumber="2336" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFC0FFAB5C38F958FCCCF910" box="[568,855,1682,1704]" italics="true" pageId="11" pageNumber="2336">Marcusenius multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFC0FFAB5D5CF958FC30F910" box="[860,939,1683,1704]" pageId="11" pageNumber="2336" rank="species">sp. nov.</taxonomicNameLabel>
, coll. B. Kramer and Ernst Swartz, coll. and photographed on 19 August 2006, Cunene River, below Ruacana Falls, specimen RUAC01, SL 15.4 cm.
</paragraph>
</caption>
<paragraph id="BC5236A4FFC7FFAC5EFEFF6CFCFFFF47" blockId="12.[254,1172,164,988]" pageId="12" pageNumber="2337">
<emphasis id="8E99EAB6FFC7FFAC5EFEFF6CFE90FF07" bold="true" box="[254,267,167,191]" pageId="12" pageNumber="2337">–</emphasis>
<materialsCitation id="0C853CF9FFC7FFAC5F1CFF6CFCFBFF47" accessionNumber="SMF 28264" collectingDate="1993-09-09" collectingDateMax="1993-09-10" collectingDateMin="1993-09-09" collectionCode="SMF" collectorName="F. H. van der Bank &amp; B. Kramer" county="Mamili National Park" latitude="-18.45" location="Kwando River" longLatPrecision="1275" longitude="23.7" municipality="Nkasa Island" pageId="12" pageNumber="2337" specimenCount="1">
<collectionCode id="DAFCAE61FFC7FFAC5F1CFF6CFECCFF07" box="[284,343,167,191]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="2337">SMF</collectionCode>
28264 (
<specimenCount id="AAEBFD2DFFC7FFAC5FBAFF6CFDFCFF07" box="[442,615,167,191]" pageId="12" pageNumber="2337" type="generic">two specimens</specimenCount>
), from
<location id="B932607FFFC7FFAC5CC1FF6CFCF7FF07" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC7FFAC5CC1FF6CFCF7FF07" box="[705,876,167,191]" county="Mamili National Park" latitude="-18.45" longLatPrecision="1275" longitude="23.7" municipality="Nkasa Island" name="Kwando River" pageId="12" pageNumber="2337">Kwando River</location>
,
<collectingMunicipality id="5C36ACDEFFC7FFAC5D7BFF6CFB8CFF07" box="[891,1047,167,191]" pageId="12" pageNumber="2337">Nkasa Island</collectingMunicipality>
(
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18
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27
<superScript id="4B989BECFFC7FFAC5A6CFF6FFBEAFF0D" attach="left" box="[1132,1137,164,181]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC7FFAC5F1CFF0CFEE6FF67" box="[284,381,195,223]" degrees="23" direction="east" minutes="42" orientation="longitude" pageId="12" pageNumber="2337" precision="925" value="23.7">
23
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42
<superScript id="4B989BECFFC7FFAC5F5CFF08FEFAFF6C" attach="left" box="[348,353,195,212]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
E
</geoCoordinate>
) in
<collectingCounty id="55334E28FFC7FFAC5FACFF0CFD36FF67" box="[428,685,199,223]" pageId="12" pageNumber="2337">Mamili National Park</collectingCounty>
, close to locality 6 on
<figureCitation id="24D62A21FFC7FFAC5DB1FF0CFB8AFF67" box="[945,1041,199,223]" captionStart="Figure 1" captionStartId="4.[211,278,932,954]" captionTargetBox="[213,1170,167,899]" captionTargetId="figure-5@4.[213,1170,167,899]" captionTargetPageId="4" captionText="Figure 1. Map of sampling locations in southern Africa. (A) Type locality for Marcusenius macrolepidotus (Peters, 1852), Tete on the Lower Zambezi River; (B) origin of M. macrolepidotus, SAIAB 60847, from Marromeu, Lower Zambezi delta (Figure 4A); (C) origin of M. macrolepidotus from the Buzi System, same location as SAIAB 67369, presently alive in aquarium; (D) likely type region for Marcusenius angolensis (Boulenger, 1905), the Lower Quanza; (E) Marcusenius altisambesi Kramer et al., 2007, type locality on the Upper Zambezi River; (F) M. altisambesi from the Kwando River; (G) M. altisambesi from the Okavango delta; (H, I) Marcusenius multisquamatus sp. nov., stretching from below the Ruacana Falls to above the Epupa Falls (I), the type locality; (J) specimens from the Cunene River Mouth." figureDoi="http://doi.org/10.5281/zenodo.5197592" httpUri="https://zenodo.org/record/5197592/files/figure.png" pageId="12" pageNumber="2337">Figure 1</figureCitation>
, coll.:
<collectorName id="11185372FFC7FFAC5A61FF03FE28FF47" pageId="12" pageNumber="2337">F.H. van der Bank</collectorName>
and
<collectorName id="11185372FFC7FFAC5FEBFF23FDF8FF47" box="[491,611,232,255]" pageId="12" pageNumber="2337">B. Kramer</collectorName>
,
<date id="C8531064FFC7FFAC5C6EFF2CFCFBFF47" box="[622,864,231,255]" pageId="12" pageNumber="2337" value="1993-09-09" valueMax="1993-09-10" valueMin="1993-09-09">
<collectingDate id="D817E98CFFC7FFAC5C6EFF2CFCFBFF47" box="[622,864,231,255]" pageId="12" pageNumber="2337" value="1993-09-09" valueMax="1993-09-10" valueMin="1993-09-09">9–10 September 1993</collectingDate>
</date>
</materialsCitation>
,
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5EFEFECCFCB4FE66" blockId="12.[254,1172,164,988]" pageId="12" pageNumber="2337">
<emphasis id="8E99EAB6FFC7FFAC5EFEFECCFE90FEA7" bold="true" box="[254,267,263,287]" pageId="12" pageNumber="2337">–</emphasis>
<specimenCount id="AAEBFD2DFFC7FFAC5F1CFECCFE2FFEA7" box="[284,436,263,287]" pageId="12" pageNumber="2337" type="generic">31 specimens</specimenCount>
, about
<quantity id="7B159B41FFC7FFAC5C0CFECCFDCFFEA7" box="[524,596,263,287]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="2337" unit="m" value="500.0">500 m</quantity>
from opposite Kalimbeza fishing camp, at downstream tip of small island between Lisikili side channel and main channel, coll.: F.H. van der Bank and B. Kramer, caught
<date id="C8531064FFC7FFAC5D10FE8CFC5CFEE7" box="[784,967,327,351]" pageId="12" pageNumber="2337" value="1999-08-21">21 August 1999</date>
, water conductivity and temperature, 84 µS cm, 22
<superScript id="4B989BECFFC7FFAC5CBDFEA8FD5DFECC" attach="right" box="[701,710,355,372]" fontSize="7" pageId="12" pageNumber="2337">◦</superScript>
C, size range
<quantity id="7B159B41FFC7FFAC5D5FFEACFC7DFEC7" box="[863,998,359,383]" metricMagnitude="-1" metricUnit="m" metricValue="1.025" metricValueMax="1.33" metricValueMin="0.72" pageId="12" pageNumber="2337" unit="cm" value="10.25" valueMax="13.3" valueMin="7.2">7.2–13.3 cm</quantity>
SL, arrival live in Regensburg
<date id="C8531064FFC7FFAC5FC8FE4CFD0DFE27" box="[456,662,391,415]" pageId="12" pageNumber="2337" value="1999-09-02">2 September 1999</date>
,
<collectionCode id="DAFCAE61FFC7FFAC5CA4FE4CFD7BFE27" box="[676,736,391,415]" pageId="12" pageNumber="2337">EOD</collectionCode>
recording
<date id="C8531064FFC7FFAC5D5DFE4CFECAFE06" pageId="12" pageNumber="2337" value="1999-09-28" valueMax="1999-10-07" valueMin="1999-09-28">28 September to 7 October 1999</date>
at 100 µS
<superScript id="4B989BECFFC7FFAC5FC7FE6CFD9BFE0B" attach="left" box="[455,512,418,446]" fontSize="7" pageId="12" pageNumber="2337">cm−1</superScript>
water conductivity and 21
<superScript id="4B989BECFFC7FFAC5D2EFE68FCACFE0C" attach="right" box="[814,823,419,436]" fontSize="7" pageId="12" pageNumber="2337">◦</superScript>
C (
<collectionCode id="DAFCAE61FFC7FFAC5D59FE6DFC0DFE06" box="[857,918,422,446]" pageId="12" pageNumber="2337">EOD</collectionCode>
recording in
<collectingCountry id="C4FA7634FFC7FFAC5A29FE6DFB08FE06" box="[1065,1171,422,446]" name="Germany" pageId="12" pageNumber="2337">Germany</collectingCountry>
for quicker transport in Africa), presently alive,
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5C73FEA8FD10FECC" blockId="12.[254,1172,164,988]" box="[627,651,355,372]" pageId="12" pageNumber="2337">
<superScript id="4B989BECFFC7FFAC5C73FEA8FD10FECC" attach="left" box="[627,651,355,372]" fontSize="7" pageId="12" pageNumber="2337">−1</superScript>
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5EFEFE2DFBD3FDA6" blockId="12.[254,1172,164,988]" pageId="12" pageNumber="2337">
<emphasis id="8E99EAB6FFC7FFAC5EFEFE2DFE90FE46" bold="true" box="[254,267,486,510]" pageId="12" pageNumber="2337">–</emphasis>
<materialsCitation id="0C853CF9FFC7FFAC5F1CFE2DFBDFFDA6" accessionNumber="ZSM 35083" collectingDate="2001-01-24" collectionCode="ZSM" collectorName="F. H. van der Bank &amp; B. Kramer" latitude="-17.790749" location="Kwando River" longLatPrecision="1" longitude="23.344444" municipality="Kongola Bridge" pageId="12" pageNumber="2337" specimenCount="1">
<collectionCode id="DAFCAE61FFC7FFAC5F1CFE2DFECCFE46" box="[284,343,486,510]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="12" pageNumber="2337" type="Museum">ZSM</collectionCode>
35083 (1), from
<location id="B932607FFFC7FFAC5C46FE2DFD63FE46" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC7FFAC5C46FE2DFD63FE46" box="[582,760,486,510]" latitude="-17.790749" longLatPrecision="1" longitude="23.344444" municipality="Kongola Bridge" name="Kwando River" pageId="12" pageNumber="2337">Kwando River</location>
,
<collectingMunicipality id="5C36ACDEFFC7FFAC5D10FE2DFC48FE46" box="[784,979,486,510]" pageId="12" pageNumber="2337">Kongola Bridge</collectingMunicipality>
,
<geoCoordinate id="D9D95063FFC7FFAC5DEAFE2DFB14FE46" box="[1002,1167,482,510]" degrees="17" direction="south" minutes="47" orientation="latitude" pageId="12" pageNumber="2337" precision="1" seconds="26.7" value="-17.790749">
17
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47
<superScript id="4B989BECFFC7FFAC5A2AFE29FBB4FE4B" attach="none" box="[1066,1071,482,499]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
26.7
<superScript id="4B989BECFFC7FFAC5A5FFE29FBF2FE4B" attach="left" box="[1119,1129,482,499]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC7FFAC5F1CFDCDFE23FDA6" box="[284,440,514,542]" degrees="23" direction="east" minutes="20" orientation="longitude" pageId="12" pageNumber="2337" precision="1" seconds="40.0" value="23.344444">
23
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20
<superScript id="4B989BECFFC7FFAC5F5CFDC9FEFAFDAB" attach="none" box="[348,353,514,531]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
40.0
<superScript id="4B989BECFFC7FFAC5F91FDC9FE00FDAB" attach="left" box="[401,411,514,531]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
E
</geoCoordinate>
,
<date id="C8531064FFC7FFAC5FC3FDCDFDE7FDA6" box="[451,636,518,543]" pageId="12" pageNumber="2337" value="2001-01-24">
<collectingDate id="D817E98CFFC7FFAC5FC3FDCDFDE7FDA6" box="[451,636,518,543]" pageId="12" pageNumber="2337" value="2001-01-24">24 January 2001</collectingDate>
</date>
, coll.:
<collectorName id="11185372FFC7FFAC5CC4FDCCFC0FFDA6" box="[708,916,518,542]" pageId="12" pageNumber="2337">F.H. van der Bank</collectorName>
and
<collectorName id="11185372FFC7FFAC5DCCFDCCFBDFFDA6" box="[972,1092,519,542]" pageId="12" pageNumber="2337">B. Kramer</collectorName>
</materialsCitation>
,
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5EFEFDEDFDDCFDC6" blockId="12.[254,1172,164,988]" pageId="12" pageNumber="2337">
<emphasis id="8E99EAB6FFC7FFAC5EFEFDEDFE90FD86" bold="true" box="[254,267,550,574]" pageId="12" pageNumber="2337">–</emphasis>
<materialsCitation id="0C853CF9FFC7FFAC5F1CFDECFDD8FDC6" country="Botswana" location="Okavango River" pageId="12" pageNumber="2337" specimenCount="73" specimenCount-male="10">
Non-types (
<specimenCount id="AAEBFD2DFFC7FFAC5FA8FDEDFDD9FD86" box="[424,578,550,574]" pageId="12" pageNumber="2337" type="generic">63 specimens</specimenCount>
)from the
<location id="B932607FFFC7FFAC5CBFFDEDFCE5FD86" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC7FFAC5CBFFDEDFCE5FD86" box="[703,894,550,574]" country="Botswana" name="Okavango River" pageId="12" pageNumber="2337">Okavango River</location>
,
<collectingCountry id="C4FA7634FFC7FFAC5D8FFDECFB9BFD86" box="[911,1024,551,574]" name="Botswana" pageId="12" pageNumber="2337">Botswana</collectingCountry>
, totalling at least
<specimenCount id="AAEBFD2DFFC7FFAC5F5BFD8DFE58FDE6" box="[347,451,582,606]" pageId="12" pageNumber="2337" type="male">10 males</specimenCount>
, male size range
<quantity id="7B159B41FFC7FFAC5C93FD8DFCB3FDE6" box="[659,808,582,606]" metricMagnitude="-1" metricUnit="m" metricValue="1.455" metricValueMax="1.81" metricValueMin="1.1" pageId="12" pageNumber="2337" unit="mm" value="145.5" valueMax="181.0" valueMin="110.0">110–181 mm</quantity>
SL, juvenile/female size range
<quantity id="7B159B41FFC7FFAC5F1CFDADFE04FDC6" box="[284,415,614,638]" metricMagnitude="-1" metricUnit="m" metricValue="1.115" metricValueMax="1.69" metricValueMin="0.54" pageId="12" pageNumber="2337" unit="mm" value="111.5" valueMax="169.0" valueMin="54.0">54–169 mm</quantity>
SL, examined
</materialsCitation>
:
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5EFEFD4DFD64FD45" blockId="12.[254,1172,164,988]" pageId="12" pageNumber="2337">
<emphasis id="8E99EAB6FFC7FFAC5EFEFD4DFE90FD26" bold="true" box="[254,267,646,670]" pageId="12" pageNumber="2337">–</emphasis>
<materialsCitation id="0C853CF9FFC7FFAC5F1CFD4DFE75FD26" box="[284,494,646,670]" collectionCode="SAIAB" pageId="12" pageNumber="2337" specimenCode="SAIAB 79140" specimenCount="1">
<specimenCode id="EC4B9EDFFFC7FFAC5F1CFD4DFE24FD26" box="[284,447,646,670]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="12" pageNumber="2337" type="Other type of research institution/biorepository">SAIAB 79140</specimenCode>
(9)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC7FFAC5C01FD4DFD22FD26" accessionNumber="ZSM 35079" box="[513,697,646,670]" collectionCode="ZSM" pageId="12" pageNumber="2337" specimenCount="1">
<collectionCode id="DAFCAE61FFC7FFAC5C01FD4DFDA7FD26" box="[513,572,646,670]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="12" pageNumber="2337" type="Museum">ZSM</collectionCode>
35079 (1)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC7FFAC5CCCFD4DFC1EFD26" accessionNumber="ZSM 35080" box="[716,901,646,670]" collectionCode="ZSM" pageId="12" pageNumber="2337" specimenCount="1">
<collectionCode id="DAFCAE61FFC7FFAC5CCCFD4DFC9CFD26" box="[716,775,646,670]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="12" pageNumber="2337" type="Museum">ZSM</collectionCode>
35080 (3)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC7FFAC5D98FD4DFBD4FD26" accessionNumber="ZSM 35081" box="[920,1103,646,670]" collectionCode="ZSM" pageId="12" pageNumber="2337" specimenCount="1">
<collectionCode id="DAFCAE61FFC7FFAC5D98FD4DFC48FD26" box="[920,979,646,670]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="12" pageNumber="2337" type="Museum">ZSM</collectionCode>
35081 (6)
</materialsCitation>
<materialsCitation id="0C853CF9FFC7FFAC5A5DFD4DFD63FD45" collectingDate="2001-01-20" collectingDateMax="2001-01-22" collectingDateMin="2001-01-20" collectorName="F. H. van der Bank &amp; J. Engelbrecht &amp; B. Kramer" county="River" latitude="-19.125" location="Etsha" longLatPrecision="874" longitude="22.366667" municipality="Makwena Lodge" pageId="12" pageNumber="2337" specimenCount="1" stateProvince="Kavango West">
from the
<collectingRegion id="7E29F846FFC7FFAC5F4DFD6EFE5BFD05" box="[333,448,677,701]" country="Namibia" name="Kavango West" pageId="12" pageNumber="2337">Okavango</collectingRegion>
<collectingCounty id="55334E28FFC7FFAC5FCDFD6EFD97FD05" box="[461,524,677,701]" pageId="12" pageNumber="2337">River</collectingCounty>
,
<collectingMunicipality id="5C36ACDEFFC7FFAC5C1EFD6EFD7EFD05" box="[542,741,677,701]" pageId="12" pageNumber="2337">Makwena Lodge</collectingMunicipality>
, near the township of
<location id="B932607FFFC7FFAC5A00FD6EFBDAFD05" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC7FFAC5A00FD6EFBDAFD05" box="[1024,1089,677,701]" county="River" latitude="-19.125" longLatPrecision="874" longitude="22.366667" municipality="Makwena Lodge" name="Etsha" pageId="12" pageNumber="2337" stateProvince="Kavango West">Etsha</location>
no. 6,
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19
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07
<superScript id="4B989BECFFC7FFAC5F5CFD09FEFAFD6B" attach="none" box="[348,353,706,723]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
30
<superScript id="4B989BECFFC7FFAC5F7DFD09FE1CFD6B" attach="left" box="[381,391,706,723]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC7FFAC5FADFD0EFD94FD65" box="[429,527,706,733]" degrees="22" direction="east" minutes="22" orientation="longitude" pageId="12" pageNumber="2337" precision="925" value="22.366667">
22
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22
<superScript id="4B989BECFFC7FFAC5FECFD09FE6AFD6B" attach="left" box="[492,497,706,723]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
E
</geoCoordinate>
, coll.:
<collectorName id="11185372FFC7FFAC5C5AFD0DFCABFD65" box="[602,816,709,733]" pageId="12" pageNumber="2337">F.H. van der Bank</collectorName>
,
<collectorName id="11185372FFC7FFAC5D3DFD0DFC47FD65" box="[829,988,709,733]" pageId="12" pageNumber="2337">J. Engelbrecht</collectorName>
and
<collectorName id="11185372FFC7FFAC5A16FD0DFB0BFD65" box="[1046,1168,710,733]" pageId="12" pageNumber="2337">B. Kramer</collectorName>
,
<date id="C8531064FFC7FFAC5F1CFD2EFE66FD45" box="[284,509,741,766]" pageId="12" pageNumber="2337" value="2001-01-20" valueMax="2001-01-22" valueMin="2001-01-20">
<collectingDate id="D817E98CFFC7FFAC5F1CFD2EFE66FD45" box="[284,509,741,766]" pageId="12" pageNumber="2337" value="2001-01-20" valueMax="2001-01-22" valueMin="2001-01-20">20–22 January 2001</collectingDate>
</date>
, locality 7 on
<figureCitation id="24D62A21FFC7FFAC5C9AFD2EFD63FD45" box="[666,760,741,765]" captionStart="Figure 1" captionStartId="4.[211,278,932,954]" captionTargetBox="[213,1170,167,899]" captionTargetId="figure-5@4.[213,1170,167,899]" captionTargetPageId="4" captionText="Figure 1. Map of sampling locations in southern Africa. (A) Type locality for Marcusenius macrolepidotus (Peters, 1852), Tete on the Lower Zambezi River; (B) origin of M. macrolepidotus, SAIAB 60847, from Marromeu, Lower Zambezi delta (Figure 4A); (C) origin of M. macrolepidotus from the Buzi System, same location as SAIAB 67369, presently alive in aquarium; (D) likely type region for Marcusenius angolensis (Boulenger, 1905), the Lower Quanza; (E) Marcusenius altisambesi Kramer et al., 2007, type locality on the Upper Zambezi River; (F) M. altisambesi from the Kwando River; (G) M. altisambesi from the Okavango delta; (H, I) Marcusenius multisquamatus sp. nov., stretching from below the Ruacana Falls to above the Epupa Falls (I), the type locality; (J) specimens from the Cunene River Mouth." figureDoi="http://doi.org/10.5281/zenodo.5197592" httpUri="https://zenodo.org/record/5197592/files/figure.png" pageId="12" pageNumber="2337">Figure 1</figureCitation>
</materialsCitation>
,
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5EFEFCCEFE21FCC5" blockId="12.[254,1172,164,988]" pageId="12" pageNumber="2337">
<emphasis id="8E99EAB6FFC7FFAC5EFEFCCEFE90FCA5" bold="true" box="[254,267,773,797]" pageId="12" pageNumber="2337">–</emphasis>
<materialsCitation id="0C853CF9FFC7FFAC5F1CFCCEFDB6FCA5" box="[284,557,773,797]" collectionCode="ZSM" pageId="12" pageNumber="2337" specimenCode="SAIAB 79143" specimenCount="1">
<specimenCode id="EC4B9EDFFFC7FFAC5F1CFCCEFE21FCA5" box="[284,442,773,797]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="12" pageNumber="2337" type="Other type of research institution/biorepository">SAIAB 79143</specimenCode>
(6),
<collectionCode id="DAFCAE61FFC7FFAC5FF2FCCEFDB6FCA5" box="[498,557,773,797]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="12" pageNumber="2337" type="Museum">ZSM</collectionCode>
</materialsCitation>
35096 (5),
<materialsCitation id="0C853CF9FFC7FFAC5CAFFCCEFE29FCC5" collectingDate="2004-08-10" collectingDateMax="2004-08-12" collectingDateMin="2004-08-10" collectionCode="ZSM" collectorName="F. H. van der Bank &amp; B. Kramer" latitude="-18.962946" location="Guma Lagoon" longLatPrecision="1" longitude="22.373695" municipality="River" pageId="12" pageNumber="2337" specimenCount="24" stateProvince="Kavango West">
and
<specimenCount id="AAEBFD2DFFC7FFAC5CE2FCCEFCE3FCA5" box="[738,888,773,797]" pageId="12" pageNumber="2337" type="generic">24 specimens</specimenCount>
presently alive in aquarium, from the
<collectingRegion id="7E29F846FFC7FFAC5FBEFCEEFDAAFC85" box="[446,561,805,829]" country="Namibia" name="Kavango West" pageId="12" pageNumber="2337">Okavango</collectingRegion>
<collectingMunicipality id="5C36ACDEFFC7FFAC5C39FCEEFDECFC85" box="[569,631,805,829]" pageId="12" pageNumber="2337">River</collectingMunicipality>
at
<location id="B932607FFFC7FFAC5C9AFCEEFCDFFC86" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC7FFAC5C9AFCEEFCDFFC86" box="[666,836,805,830]" latitude="-18.962946" longLatPrecision="1" longitude="22.373695" municipality="River" name="Guma Lagoon" pageId="12" pageNumber="2337" stateProvince="Kavango West">Guma Lagoon</location>
,
<geoCoordinate id="D9D95063FFC7FFAC5D4FFCEEFC7CFC85" box="[847,999,801,829]" degrees="18" direction="south" minutes="57" orientation="latitude" pageId="12" pageNumber="2337" precision="1" seconds="46.6" value="-18.962946">
18
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57
<superScript id="4B989BECFFC7FFAC5D8FFCEAFC0FFC8A" attach="none" box="[911,916,801,818]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
46.6
<superScript id="4B989BECFFC7FFAC5DC4FCEAFC55FC8A" attach="left" box="[964,974,801,818]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC7FFAC5DF3FCEEFB15FC85" box="[1011,1166,801,829]" degrees="22" direction="east" minutes="22" orientation="longitude" pageId="12" pageNumber="2337" precision="1" seconds="25.3" value="22.373695">
22
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22
<superScript id="4B989BECFFC7FFAC5A32FCEAFBACFC8A" attach="left" box="[1074,1079,801,818]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
25.3
<superScript id="4B989BECFFC7FFAC5A68FCEAFBE9FC8A" attach="left" box="[1128,1138,801,818]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
E
</geoCoordinate>
, coll.:
<collectorName id="11185372FFC7FFAC5F5CFC8DFDA9FCE5" box="[348,562,837,861]" pageId="12" pageNumber="2337">F.H. van der Bank</collectorName>
and
<collectorName id="11185372FFC7FFAC5C6FFC8DFD72FCE5" box="[623,745,838,861]" pageId="12" pageNumber="2337">B. Kramer</collectorName>
,
<date id="C8531064FFC7FFAC5CF6FC8EFC48FCE5" box="[758,979,837,861]" pageId="12" pageNumber="2337" value="2004-08-10" valueMax="2004-08-12" valueMin="2004-08-10">
<collectingDate id="D817E98CFFC7FFAC5CF6FC8EFC48FCE5" box="[758,979,837,861]" pageId="12" pageNumber="2337" value="2004-08-10" valueMax="2004-08-12" valueMin="2004-08-10">10–12 August 2004</collectingDate>
</date>
, close to locality 7 on
<figureCitation id="24D62A21FFC7FFAC5F54FCAEFE29FCC5" box="[340,434,869,893]" captionStart="Figure 1" captionStartId="4.[211,278,932,954]" captionTargetBox="[213,1170,167,899]" captionTargetId="figure-5@4.[213,1170,167,899]" captionTargetPageId="4" captionText="Figure 1. Map of sampling locations in southern Africa. (A) Type locality for Marcusenius macrolepidotus (Peters, 1852), Tete on the Lower Zambezi River; (B) origin of M. macrolepidotus, SAIAB 60847, from Marromeu, Lower Zambezi delta (Figure 4A); (C) origin of M. macrolepidotus from the Buzi System, same location as SAIAB 67369, presently alive in aquarium; (D) likely type region for Marcusenius angolensis (Boulenger, 1905), the Lower Quanza; (E) Marcusenius altisambesi Kramer et al., 2007, type locality on the Upper Zambezi River; (F) M. altisambesi from the Kwando River; (G) M. altisambesi from the Okavango delta; (H, I) Marcusenius multisquamatus sp. nov., stretching from below the Ruacana Falls to above the Epupa Falls (I), the type locality; (J) specimens from the Cunene River Mouth." figureDoi="http://doi.org/10.5281/zenodo.5197592" httpUri="https://zenodo.org/record/5197592/files/figure.png" pageId="12" pageNumber="2337">Figure 1</figureCitation>
</materialsCitation>
,
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5EFEFC4EFCB3FC64" blockId="12.[254,1172,164,988]" pageId="12" pageNumber="2337">
<emphasis id="8E99EAB6FFC7FFAC5EFEFC4EFE90FC25" bold="true" box="[254,267,901,925]" pageId="12" pageNumber="2337">–</emphasis>
<materialsCitation id="0C853CF9FFC7FFAC5F1CFC4EFDA0FC25" box="[284,571,901,925]" collectionCode="ZSM" pageId="12" pageNumber="2337" specimenCode="SAIAB 79141" specimenCount="1">
<specimenCode id="EC4B9EDFFFC7FFAC5F1CFC4EFE24FC25" box="[284,447,901,925]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="12" pageNumber="2337" type="Other type of research institution/biorepository">SAIAB 79141</specimenCode>
(1),
<collectionCode id="DAFCAE61FFC7FFAC5C00FC4EFDA0FC25" box="[512,571,901,925]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="12" pageNumber="2337" type="Museum">ZSM</collectionCode>
</materialsCitation>
35095 (1), and
<specimenCount id="AAEBFD2DFFC7FFAC5D02FC4DFC26FC25" box="[770,957,901,925]" pageId="12" pageNumber="2337" type="generic">seven specimens</specimenCount>
presently alive in aquarium, details as in preceding paragraph, except for locality at Makwena,
<geoCoordinate id="D9D95063FFC7FFAC5F1CFC0FFE5AFC64" box="[284,449,961,988]" degrees="19" direction="south" minutes="03" orientation="latitude" pageId="12" pageNumber="2337" precision="1" seconds="13.85" value="-19.053846">
19
<superScript id="4B989BECFFC7FFAC5F37FC0AFEDBFC6A" attach="none" box="[311,320,961,978]" fontSize="7" pageId="12" pageNumber="2337">◦</superScript>
03
<superScript id="4B989BECFFC7FFAC5F5CFC0AFEFAFC6A" attach="none" box="[348,353,961,978]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
13.85
<superScript id="4B989BECFFC7FFAC5F9FFC0AFE32FC6A" attach="left" box="[415,425,961,978]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC7FFAC5FCCFC0FFDFCFC64" box="[460,615,961,988]" degrees="22" direction="east" minutes="22" orientation="longitude" pageId="12" pageNumber="2337" precision="1" seconds="42.6" value="22.3785">
22
<superScript id="4B989BECFFC7FFAC5FE7FC0AFE6BFC6A" attach="none" box="[487,496,961,978]" fontSize="7" pageId="12" pageNumber="2337">◦</superScript>
22
<superScript id="4B989BECFFC7FFAC5C0CFC0AFD8AFC6A" attach="none" box="[524,529,961,978]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
42.6
<superScript id="4B989BECFFC7FFAC5C41FC0AFDD0FC6A" attach="left" box="[577,587,961,978]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
E
</geoCoordinate>
,
<date id="C8531064FFC7FFAC5C73FC0FFCBFFC64" box="[627,804,964,988]" pageId="12" pageNumber="2337" value="2004-08-12">12 August 2004</date>
.
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5ED3FBCFFCC7FB84" blockId="12.[211,1172,1028,1084]" pageId="12" pageNumber="2337">
<emphasis id="8E99EAB6FFC7FFAC5ED3FBCFFDB0FBA4" box="[211,555,1028,1052]" italics="true" pageId="12" pageNumber="2337">Samples examined for genetics.</emphasis>
DNA samples are stored at Institute of Pharmacy and Molecular Biotechnology, Heidelberg University (
<collectionCode id="DAFCAE61FFC7FFAC5D04FBEFFCD4FB84" box="[772,847,1060,1084]" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="12" pageNumber="2337">IPMB</collectionCode>
).
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5EFEFBAFFBA2FB24" blockId="12.[254,1172,1120,1307]" pageId="12" pageNumber="2337">
<emphasis id="8E99EAB6FFC7FFAC5EFEFBAFFE90FBC4" bold="true" box="[254,267,1124,1148]" pageId="12" pageNumber="2337">–</emphasis>
<materialsCitation id="0C853CF9FFC7FFAC5F1CFBAEFBAEFB24" collectingDate="1999-08-21" collectionCode="IPMB" collectorName="F. H. van der Bank &amp; B. Kramer" country="Namibia" latitude="-17.540916" location="Kalimbeza" longLatPrecision="1" longitude="24.523945" municipality="Upper" pageId="12" pageNumber="2337" specimenCode="IPMB 44903-44905" specimenCount="1" stateProvince="Zambezi">
<specimenCode id="EC4B9EDFFFC7FFAC5F1CFBAEFD9DFBC4" box="[284,518,1124,1148]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="12" pageNumber="2337">IPMB 44903–44905</specimenCode>
,
<collectingCountry id="C4FA7634FFC7FFAC5C18FBAFFDE4FBC4" box="[536,639,1124,1148]" name="Namibia" pageId="12" pageNumber="2337">Namibia</collectingCountry>
:
<collectingMunicipality id="5C36ACDEFFC7FFAC5C91FBAEFD41FBC4" box="[657,730,1125,1148]" pageId="12" pageNumber="2337">Upper</collectingMunicipality>
<collectingRegion id="7E29F846FFC7FFAC5CE9FBAFFCD4FBC4" box="[745,847,1124,1148]" country="Namibia" name="Zambezi" pageId="12" pageNumber="2337">Zambezi</collectingRegion>
:
<location id="B932607FFFC7FFAC5D60FBAFFC46FBC4" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC7FFAC5D60FBAFFC46FBC4" box="[864,989,1124,1148]" country="Namibia" latitude="-17.540916" longLatPrecision="1" longitude="24.523945" municipality="Upper" name="Kalimbeza" pageId="12" pageNumber="2337" stateProvince="Zambezi">Kalimbeza</location>
,
<geoCoordinate id="D9D95063FFC7FFAC5DF0FBAFFB14FBC4" box="[1008,1167,1120,1148]" degrees="17" direction="south" minutes="32" orientation="latitude" pageId="12" pageNumber="2337" precision="1" seconds="27.3" value="-17.540916">
17
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32
<superScript id="4B989BECFFC7FFAC5A30FBABFBAEFBC9" attach="none" box="[1072,1077,1120,1137]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
27.3
<superScript id="4B989BECFFC7FFAC5A65FBABFBF4FBC9" attach="left" box="[1125,1135,1120,1137]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC7FFAC5F1CFB4FFE23FB23" box="[284,440,1152,1180]" degrees="24" direction="east" minutes="31" orientation="longitude" pageId="12" pageNumber="2337" precision="1" seconds="26.2" value="24.523945">
24
<superScript id="4B989BECFFC7FFAC5F37FB4BFEDBFB29" attach="none" box="[311,320,1152,1169]" fontSize="7" pageId="12" pageNumber="2337">◦</superScript>
31
<superScript id="4B989BECFFC7FFAC5F5CFB4BFEFAFB29" attach="none" box="[348,353,1152,1169]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
26.2
<superScript id="4B989BECFFC7FFAC5F91FB4BFE00FB29" attach="left" box="[401,411,1152,1169]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
E
</geoCoordinate>
, coll.
<collectorName id="11185372FFC7FFAC5FFAFB4FFD52FB24" box="[506,713,1156,1180]" pageId="12" pageNumber="2337">F.H. van der Bank</collectorName>
and
<collectorName id="11185372FFC7FFAC5D01FB4FFCE2FB23" box="[769,889,1156,1179]" pageId="12" pageNumber="2337">B. Kramer</collectorName>
,
<date id="C8531064FFC7FFAC5D84FB4FFBAEFB24" box="[900,1077,1156,1180]" pageId="12" pageNumber="2337" value="1999-08-21">
<collectingDate id="D817E98CFFC7FFAC5D84FB4FFBAEFB24" box="[900,1077,1156,1180]" pageId="12" pageNumber="2337" value="1999-08-21">21 August 1999</collectingDate>
</date>
</materialsCitation>
;
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5EFEFB6FFDABFAA3" blockId="12.[254,1172,1120,1307]" pageId="12" pageNumber="2337">
<emphasis id="8E99EAB6FFC7FFAC5EFEFB6FFE90FB04" bold="true" box="[254,267,1188,1212]" pageId="12" pageNumber="2337">–</emphasis>
<materialsCitation id="0C853CF9FFC7FFAC5F1CFB6FFBDAFB63" collectingDate="2004-08-10" collectionCode="IPMB" collectorName="F. H. van der Bank &amp; B. Kramer" country="Botswana" latitude="-18.962946" location="Okavango" longLatPrecision="1" longitude="22.373695" municipality="Guma Lagoon" pageId="12" pageNumber="2337" specimenCode="IPMB 44638-44640" specimenCount="1">
<specimenCode id="EC4B9EDFFFC7FFAC5F1CFB6FFD9EFB04" box="[284,517,1188,1212]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="12" pageNumber="2337">IPMB 44638–44640</specimenCode>
<collectingCountry id="C4FA7634FFC7FFAC5C16FB6FFD13FB03" box="[534,648,1188,1211]" name="Botswana" pageId="12" pageNumber="2337">Botswana</collectingCountry>
:
<location id="B932607FFFC7FFAC5C9CFB6FFC88FB04" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC7FFAC5C9CFB6FFC88FB04" box="[668,787,1188,1212]" country="Botswana" latitude="-18.962946" longLatPrecision="1" longitude="22.373695" municipality="Guma Lagoon" name="Okavango" pageId="12" pageNumber="2337">Okavango</location>
:
<collectingMunicipality id="5C36ACDEFFC7FFAC5D26FB6FFC42FB04" box="[806,985,1188,1212]" pageId="12" pageNumber="2337">Guma Lagoon</collectingMunicipality>
,
<geoCoordinate id="D9D95063FFC7FFAC5DEEFB6FFB14FB04" box="[1006,1167,1184,1212]" degrees="18" direction="south" minutes="57" orientation="latitude" pageId="12" pageNumber="2337" precision="1" seconds="46.6" value="-18.962946">
18
<superScript id="4B989BECFFC7FFAC5A08FB6BFB8AFB09" attach="left" box="[1032,1041,1184,1201]" fontSize="7" pageId="12" pageNumber="2337">◦</superScript>
57
<superScript id="4B989BECFFC7FFAC5A2DFB6BFBA9FB09" attach="left" box="[1069,1074,1184,1201]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
46.6
<superScript id="4B989BECFFC7FFAC5A63FB6BFBF6FB09" attach="left" box="[1123,1133,1184,1201]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC7FFAC5F1CFB08FE22FB63" box="[284,441,1216,1243]" degrees="22" direction="east" minutes="22" orientation="longitude" pageId="12" pageNumber="2337" precision="1" seconds="25.3" value="22.373695">
22
<superScript id="4B989BECFFC7FFAC5F37FB0BFEDBFB69" attach="none" box="[311,320,1216,1233]" fontSize="7" pageId="12" pageNumber="2337">◦</superScript>
22
<superScript id="4B989BECFFC7FFAC5F5CFB0BFEFAFB69" attach="none" box="[348,353,1216,1233]" fontSize="7" pageId="12" pageNumber="2337">′</superScript>
25.3
<superScript id="4B989BECFFC7FFAC5F91FB0BFE00FB69" attach="left" box="[401,411,1216,1233]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
E
</geoCoordinate>
, coll.
<collectorName id="11185372FFC7FFAC5FFDFB0FFD54FB63" box="[509,719,1219,1243]" pageId="12" pageNumber="2337">F.H. van der Bank</collectorName>
and
<collectorName id="11185372FFC7FFAC5D0AFB0FFC18FB63" box="[778,899,1220,1243]" pageId="12" pageNumber="2337">B. Kramer</collectorName>
,
<date id="C8531064FFC7FFAC5D8EFB08FBDAFB63" box="[910,1089,1219,1243]" pageId="12" pageNumber="2337" value="2004-08-10">
<collectingDate id="D817E98CFFC7FFAC5D8EFB08FBDAFB63" box="[910,1089,1219,1243]" pageId="12" pageNumber="2337" value="2004-08-10">10 August 2004</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC7FFAC5A4DFB0FFDB7FAA3" accessionNumber="KC202230, KC202237" collectingDate="2004-08-10" collectingDateMax="2004-08-12" collectingDateMin="2004-08-10" collectionCode="KC" country="Botswana" pageId="12" pageNumber="2337" specimenCode="IPMB 44641, 44642" specimenCount="1">
<specimenCode id="EC4B9EDFFFC7FFAC5A4DFB0FFEF9FB43" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="12" pageNumber="2337">IPMB 44641</specimenCode>
,
<specimenCode id="EC4B9EDFFFC7FFAC5F6EFB28FE2FFB43" box="[366,436,1251,1275]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="12" pageNumber="2337">44642</specimenCode>
, as before, but
<date id="C8531064FFC7FFAC5C5FFB28FCA1FB43" box="[607,826,1251,1275]" pageId="12" pageNumber="2337" value="2004-08-10" valueMax="2004-08-12" valueMin="2004-08-10">
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; GenBank accession numbers: (
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202230
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202237
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)
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5ED3FA92FEF9FAC8" blockId="12.[211,1172,1368,1719]" box="[211,354,1368,1392]" pageId="12" pageNumber="2337">
<heading id="E71A81C8FFC7FFAC5ED3FA92FEF9FAC8" box="[211,354,1368,1392]" fontSize="10" level="3" pageId="12" pageNumber="2337" reason="5">
<emphasis id="8E99EAB6FFC7FFAC5ED3FA92FEF9FAC8" box="[211,354,1368,1392]" italics="true" pageId="12" pageNumber="2337">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFC7FFAC5ED3FA4BFE81FA4F" blockId="12.[211,1172,1368,1719]" pageId="12" pageNumber="2337">
<materialsCitation id="0C853CF9FFC7FFAC5ED3FA4BFE8CFA4F" country="Namibia" county="Katima Mulilo" latitude="-17.540916" location="Upper" longLatPrecision="1" longitude="24.523945" municipality="Kalambesa" pageId="12" pageNumber="2337" specimenCount="1" stateProvince="Zambezi">
<location id="B932607FFFC7FFAC5ED3FA4BFE87FA2F" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC7FFAC5ED3FA4BFE87FA2F" box="[211,284,1408,1431]" country="Namibia" county="Katima Mulilo" latitude="-17.540916" longLatPrecision="1" longitude="24.523945" municipality="Kalambesa" name="Upper" pageId="12" pageNumber="2337" stateProvince="Zambezi">Upper</location>
<collectingRegion id="7E29F846FFC7FFAC5F25FA4BFE54FA20" box="[293,463,1408,1432]" country="Namibia" name="Zambezi" pageId="12" pageNumber="2337">Zambezi River</collectingRegion>
in
<location id="B932607FFFC7FFAC5FF7FA4BFDB2FA2F" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC7FFAC5FF7FA4BFDB2FA2F" box="[503,553,1408,1431]" country="Namibia" county="Katima Mulilo" latitude="-17.540916" longLatPrecision="1" longitude="24.523945" municipality="Kalambesa" name="East" pageId="12" pageNumber="2337" stateProvince="Zambezi">East</location>
<collectingRegion id="7E29F846FFC7FFAC5C32FA4BFD1DFA20" box="[562,646,1408,1432]" country="Namibia" name="Zambezi" pageId="12" pageNumber="2337">Caprivi</collectingRegion>
(
<collectingCountry id="C4FA7634FFC7FFAC5C97FA4BFD64FA20" box="[663,767,1408,1432]" name="Namibia" pageId="12" pageNumber="2337">Namibia</collectingCountry>
); specifically
<location id="B932607FFFC7FFAC5D98FA4BFC7AFA2F" LSID="urn:lsid:plazi:treatment:344487B2FFC1FFAD5FBCFCBEFE7AFEA7:B932607FFFC7FFAC5D98FA4BFC7AFA2F" box="[920,993,1408,1431]" country="Namibia" county="Katima Mulilo" latitude="-17.540916" longLatPrecision="1" longitude="24.523945" municipality="Kalambesa" name="Upper" pageId="12" pageNumber="2337" stateProvince="Zambezi">Upper</location>
<collectingRegion id="7E29F846FFC7FFAC5DEAFA4BFB08FA20" box="[1002,1171,1408,1432]" country="Namibia" name="Zambezi" pageId="12" pageNumber="2337">Zambezi River</collectingRegion>
<locationDeviation id="ADE02F86FFC7FFAC5ED3FA54FDA6FA0F" box="[211,573,1439,1463]" firstLocation="Lisikili" pageId="12" pageNumber="2337" secondLocation="Kalimbeza">between Lisikili and Kalimbeza</locationDeviation>
(or
<collectingMunicipality id="5C36ACDEFFC7FFAC5C70FA54FD69FA0F" box="[624,754,1439,1463]" pageId="12" pageNumber="2337">Kalambesa</collectingMunicipality>
,
<geoCoordinate id="D9D95063FFC7FFAC5CFFFA54FCC5FA0F" box="[767,862,1436,1463]" degrees="17" direction="south" minutes="33" orientation="latitude" pageId="12" pageNumber="2337" precision="925" value="-17.55">
17
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S
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,
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24
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29
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E
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to
<geoCoordinate id="D9D95063FFC7FFAC5DF5FA54FB14FA0F" box="[1013,1167,1436,1463]" degrees="17" direction="south" minutes="32" orientation="latitude" pageId="12" pageNumber="2337" precision="1" seconds="27.3" value="-17.540916">
17
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32
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27.3
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S
</geoCoordinate>
,
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24
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31
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26.2
<superScript id="4B989BECFFC7FFAC5F48FA70FEC9FA74" attach="left" box="[328,338,1467,1484]" fontSize="7" pageId="12" pageNumber="2337">′′</superScript>
E
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;
<quantity id="7B159B41FFC7FFAC5F82FA74FE6EFA6F" box="[386,501,1471,1495]" metricMagnitude="4" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="12" pageNumber="2337" unit="km" value="24.0" valueMax="26.0" valueMin="22.0">22–26 km</quantity>
straight line downstream from
<collectingCounty id="55334E28FFC7FFAC5D6AFA74FB81FA6F" box="[874,1050,1471,1495]" pageId="12" pageNumber="2337">Katima Mulilo</collectingCounty>
;
<figureCitation id="24D62A21FFC7FFAC5A2AFA74FB17FA6F" box="[1066,1164,1471,1495]" captionStart="Figure 1" captionStartId="4.[211,278,932,954]" captionTargetBox="[213,1170,167,899]" captionTargetId="figure-5@4.[213,1170,167,899]" captionTargetPageId="4" captionText="Figure 1. Map of sampling locations in southern Africa. (A) Type locality for Marcusenius macrolepidotus (Peters, 1852), Tete on the Lower Zambezi River; (B) origin of M. macrolepidotus, SAIAB 60847, from Marromeu, Lower Zambezi delta (Figure 4A); (C) origin of M. macrolepidotus from the Buzi System, same location as SAIAB 67369, presently alive in aquarium; (D) likely type region for Marcusenius angolensis (Boulenger, 1905), the Lower Quanza; (E) Marcusenius altisambesi Kramer et al., 2007, type locality on the Upper Zambezi River; (F) M. altisambesi from the Kwando River; (G) M. altisambesi from the Okavango delta; (H, I) Marcusenius multisquamatus sp. nov., stretching from below the Ruacana Falls to above the Epupa Falls (I), the type locality; (J) specimens from the Cunene River Mouth." figureDoi="http://doi.org/10.5281/zenodo.5197592" httpUri="https://zenodo.org/record/5197592/files/figure.png" pageId="12" pageNumber="2337">Figure 1</figureCitation>
, no. 5)
</materialsCitation>
.
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<paragraph id="BC5236A4FFC7FFAD5EFEFA34FE7AFEA7" blockId="12.[211,1172,1368,1719]" lastBlockId="13.[143,1103,167,287]" lastPageId="13" lastPageNumber="2338" pageId="12" pageNumber="2337">
The first record of
<taxonomicName id="7BED4D27FFC7FFAC5FCBFA34FD14F9AF" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[459,655,1535,1559]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="12" pageNumber="2337" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFC7FFAC5FCBFA34FD14F9AF" box="[459,655,1535,1559]" italics="true" pageId="12" pageNumber="2337">G. macrolepidotus</emphasis>
</taxonomicName>
for the Upper
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is that of
<bibRefCitation id="D87C4B55FFC7FFAC5A00FA34FE0FF98F" author="Gilchrist JDF &amp; Thompson WW" pageId="12" pageNumber="2337" pagination="465 - 579" refId="ref16232" refString="Gilchrist JDF, Thompson WW. 1917. The freshwater fishes of South Africa. Annals of the South African Museum (Cape Town, etc). XI (VI): 465 - 579." type="journal article" year="1917">Gilchrist and Thompson (1917</bibRefCitation>
, p. 562), specifying Lialui, Barotseland as origin. For a description, the authors refer to
<bibRefCitation id="D87C4B55FFC7FFAC5FBEF9F4FCBAF9EF" author="Gilchrist JDF &amp; Thompson WW" box="[446,801,1599,1623]" pageId="12" pageNumber="2337" pagination="321 - 463" refId="ref16195" refString="Gilchrist JDF, Thompson WW. 1913. The freshwater fishes of South Africa. Annals of the South African Museum (Cape Town, etc). XI (V): 321 - 463." type="journal article" year="1913">Gilchrist and Thompson (1913</bibRefCitation>
, p. 330), a description of South African specimens that
<bibRefCitation id="D87C4B55FFC7FFAC5FE0F9ABFD53F9CF" author="Kramer B &amp; Skelton PH &amp; Van der Bank FH &amp; Wink M" box="[480,712,1631,1655]" pageId="12" pageNumber="2337" pagination="647 - 708" refId="ref16697" refString="Kramer B, Skelton PH, Van der Bank FH, Wink M. 2007. Allopatric differentiation in the Marcusenius macrolepidotus species complex in southern and eastern Africa: the resurrection of M. pongolensis and M. angolensis, and the description of two new species (Mormyridae, Teleostei). J Nat Hist. 41: 647 - 708." type="journal article" year="2007">Kramer et al. (2007)</bibRefCitation>
have referred to
<taxonomicName id="7BED4D27FFC7FFAC5D89F9ABFE8EF92F" authority="(Fowler, 1934)" baseAuthorityName="Fowler" baseAuthorityYear="1934" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="12" pageNumber="2337" phylum="Chordata" rank="species" species="pongolensis">
<emphasis id="8E99EAB6FFC7FFAC5D89F9ABFBABF9CF" box="[905,1072,1631,1655]" italics="true" pageId="12" pageNumber="2337">M. pongolensis</emphasis>
(Fowler, 1934)
</taxonomicName>
. The presence of
<taxonomicName id="7BED4D27FFC7FFAC5FD4F9B4FD03F92F" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[468,664,1663,1687]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="12" pageNumber="2337" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFC7FFAC5FD4F9B4FD03F92F" box="[468,664,1663,1687]" italics="true" pageId="12" pageNumber="2337">G. macrolepidotus</emphasis>
</taxonomicName>
in the Upper
<collectingRegion id="7E29F846FFC7FFAC5D32F9B4FC0FF92F" box="[818,916,1663,1687]" country="Namibia" name="Zambezi" pageId="12" pageNumber="2337">Zambezi</collectingRegion>
System was confirmed by
<bibRefCitation id="D87C4B55FFC7FFAC5EF4F955FEE3F90E" author="Jubb RA" box="[244,376,1694,1718]" pageId="12" pageNumber="2337" pagination="177 - 189" refId="ref16442" refString="Jubb RA. 1958. A preliminary report on the collections of freshwater fishes made by the Bernard Carp expeditions to the Caprivi Strip, 1949, the lower Sabi river, 1950, and to Barotseland, 1952. Occasional Papers of the National Museums of Southern Rhodesia. 3: 177 - 189." type="journal article" year="1958">Jubb (1958)</bibRefCitation>
. Another possible synonym would be
<taxonomicName id="7BED4D27FFC7FFAC5D1CF954FC52F90F" authorityName="Pappenheim" authorityYear="1907" box="[796,969,1695,1719]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="12" pageNumber="2337" phylum="Chordata" rank="species" species="okavangensis">
<emphasis id="8E99EAB6FFC7FFAC5D1CF954FC52F90F" box="[796,969,1695,1719]" italics="true" pageId="12" pageNumber="2337">G. okavangensis</emphasis>
</taxonomicName>
if it were available (this name should be dropped from a list of synonyms, as suggested by
<bibRefCitation id="D87C4B55FFC6FFAD5DB7FF63FF5FFF67" author="Kramer B &amp; Skelton PH &amp; Van der Bank FH &amp; Wink M" pageId="13" pageNumber="2338" pagination="647 - 708" refId="ref16697" refString="Kramer B, Skelton PH, Van der Bank FH, Wink M. 2007. Allopatric differentiation in the Marcusenius macrolepidotus species complex in southern and eastern Africa: the resurrection of M. pongolensis and M. angolensis, and the description of two new species (Mormyridae, Teleostei). J Nat Hist. 41: 647 - 708." type="journal article" year="2007">Kramer et al. 2007</bibRefCitation>
). Upper
<collectingRegion id="7E29F846FFC6FFAD5F32FF0CFE0FFF67" box="[306,404,199,223]" country="Namibia" name="Zambezi" pageId="13" pageNumber="2338">Zambezi</collectingRegion>
and
<collectingRegion id="7E29F846FFC6FFAD5FD2FF0CFDDEFF67" box="[466,581,199,223]" country="Namibia" name="Kavango West" pageId="13" pageNumber="2338">Okavango</collectingRegion>
specimens were recognized as representing a new species,
<taxonomicName id="7BED4D27FFC6FFAD5F1DFF23FE5BFF47" box="[285,448,231,255]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="13" pageNumber="2338" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFC6FFAD5F1DFF23FE5BFF47" box="[285,448,231,255]" italics="true" pageId="13" pageNumber="2338">M. altisambesi</emphasis>
</taxonomicName>
, that is well differentiated from
<taxonomicName id="7BED4D27FFC6FFAD5D27FF23FF56FEA7" authority="(Peters, 1852)" baseAuthorityName="Peters" baseAuthorityYear="1852" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="13" pageNumber="2338" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFC6FFAD5D27FF23FC6EFF47" box="[807,1013,231,255]" italics="true" pageId="13" pageNumber="2338">M. macrolepidotus</emphasis>
(
<bibRefCitation id="D87C4B55FFC6FFAD5A04FF2CFF5EFEA7" author="Peters WCH" pageId="13" pageNumber="2338" pagination="273 - 276" refId="ref17195" refString="Peters WCH. 1852. Einige neue Saugethiere und Flussfische aus Mossambique. Monatsber Konigl Preuss Akad Wiss Berlin. 1852: 273 - 276." type="journal article" year="1852">Peters, 1852</bibRefCitation>
)
</taxonomicName>
by
<bibRefCitation id="D87C4B55FFC6FFAD5EF6FEC3FE47FEA7" author="Kramer B &amp; Skelton PH &amp; Van der Bank FH &amp; Wink M" box="[246,476,263,287]" pageId="13" pageNumber="2338" pagination="647 - 708" refId="ref16697" refString="Kramer B, Skelton PH, Van der Bank FH, Wink M. 2007. Allopatric differentiation in the Marcusenius macrolepidotus species complex in southern and eastern Africa: the resurrection of M. pongolensis and M. angolensis, and the description of two new species (Mormyridae, Teleostei). J Nat Hist. 41: 647 - 708." type="journal article" year="2007">Kramer et al. (2007)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

924
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<mods:detail id="BD53D5EC9F5D64D3630DBC5E70A45C6A" type="issue">
<mods:number id="1B1EAE4629C01913DB38A56796FA3F5B">35 - 36</mods:number>
</mods:detail>
<mods:extent id="0507FBB589EA568D7FBEF21FF85AAFF3" unit="page">
<mods:start id="B518CAD1B0BD97F978C7B71AC1BDF5A1">2327</mods:start>
<mods:end id="EAB89D5F9620AFA8551EA93F8681A969">2362</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="9E559CFB3018247F976A89E600604B04">
<mods:url id="A1D66410F2149A0974CB6E229DFE9812">http://dx.doi.org/10.1080/00222933.2013.798699</mods:url>
</mods:location>
<mods:classification id="43FBBB67429280C6F1A3A19CD5D861B0">journal article</mods:classification>
<mods:identifier id="03DF70DE3E721BF4B45ACFA4708630C8" type="DOI">10.1080/00222933.2013.798699</mods:identifier>
<mods:identifier id="0133B9E7F294827A395E0F53A289E5DC" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="1CF66AFBED9AA0B68EB969EEDB23A543" type="Zenodo-Dep">5197590</mods:identifier>
<mods:identifier id="ADB5AB1BACE1694760F274D6A2F3CCF3" type="ZooBank">8FE68494-9ED9-428E-B181-E814D25493F2</mods:identifier>
</mods:mods>
<treatment id="344487B2FFC5FFB25FE6FC8EFB0BFCA6" LSID="urn:lsid:zoobank.org:act:CE3474C9-C821-420C-9548-0D8F917BFCB9" httpUri="http://treatment.plazi.org/id/344487B2FFC5FFB25FE6FC8EFB0BFCA6" lastPageId="18" lastPageNumber="2343" pageId="14" pageNumber="2339">
<subSubSection id="F4F7652FFFC5FFAE5FE6FC8EFC1AFCE5" box="[486,897,836,861]" pageId="14" pageNumber="2339" type="nomenclature">
<paragraph id="BC5236A4FFC5FFAE5FE6FC8EFC1AFCE5" blockId="14.[486,897,836,893]" box="[486,897,836,861]" pageId="14" pageNumber="2339">
<heading id="E71A81C8FFC5FFAE5FE6FC8EFC1AFCE5" box="[486,897,836,861]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="2339" reason="2">
<taxonomicName id="7BED4D27FFC5FFAE5FE6FC8EFCBDFCE5" ID-CoL="3Y39B" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[486,806,836,861]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="14" pageNumber="2339" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFC5FFAE5FE6FC8EFCBDFCE5" bold="true" box="[486,806,836,861]" italics="true" pageId="14" pageNumber="2339">Marcusenius multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFC5FFAE5D2DFC8DFC1AFCE5" box="[813,897,838,861]" pageId="14" pageNumber="2339" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFC5FFAE5C72FCAEFD6FFCC5" box="[626,756,869,893]" pageId="14" pageNumber="2339" type="description">
<paragraph id="BC5236A4FFC5FFAE5C72FCAEFD6FFCC5" blockId="14.[486,897,836,893]" box="[626,756,869,893]" pageId="14" pageNumber="2339">
(
<figureCitation id="24D62A21FFC5FFAE5C7BFCAEFD70FCC5" box="[635,747,869,893]" captionStart="Figure 4" captionStartId="11.[143,210,1506,1528]" captionTargetBox="[168,1081,161,1473]" captionTargetId="figure-6@11.[168,1081,161,1473]" captionTargetPageId="11" captionText="Figure 4. Photographs of bulldog fish from various origins. (A) Marcusenius macrolepidotus (Peters, 1852), SAIAB 60847, coll. R. Bills 1 August 1999, Lower Zambezi; (B) Marcusenius angolensis (Boulenger, 1905), BMNH 1905.5.29.64 (holotype); (C) Marcusenius altisambesi, coll. F. H. van der Bank and B. Kramer, 21 August 1999, Upper Zambezi, Kalimbeza, live specimen of 16.5 cm SL photographed 3 July 2003 in aquarium; (D) M. altisambesi, coll. F.H. van der Bank and B. Kramer, 11/12 August 2004, Okavango River, live fish of SL 13 cm photographed 20 April 2006 in aquarium; (E) Marcusenius multisquamatus sp. nov., coll. B. Kramer and Ernst Swartz, coll. and photographed on 19 August 2006, Cunene River, below Ruacana Falls, specimen RUAC01, SL 15.4 cm." figureDoi="http://doi.org/10.5281/zenodo.5197598" httpUri="https://zenodo.org/record/5197598/files/figure.png" pageId="14" pageNumber="2339">Figure 4E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFC5FFAF5ED3FC09FF6EFD7A" lastPageId="15" lastPageNumber="2340" pageId="14" pageNumber="2339" type="materials_examined">
<paragraph id="BC5236A4FFC5FFAE5ED3FC09FEE5FC61" blockId="14.[211,1172,962,1726]" box="[211,382,962,985]" pageId="14" pageNumber="2339">
<heading id="E71A81C8FFC5FFAE5ED3FC09FEE5FC61" box="[211,382,962,985]" fontSize="10" level="3" pageId="14" pageNumber="2339" reason="5">
<emphasis id="8E99EAB6FFC5FFAE5ED3FC09FEE5FC61" box="[211,382,962,985]" italics="true" pageId="14" pageNumber="2339">Type specimens</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFC5FFAE5ED3FC22FDDDFA98" blockId="14.[211,1172,962,1726]" pageId="14" pageNumber="2339">
<materialsCitation id="0C853CF9FFC5FFAE5ED3FC22FB0BFBD9" collectingDate="2006-08-15" collectionCode="SAIAB, KUNE" collectorName="E. Swartz &amp; Swartz, B &amp; L. da Costa" country="Namibia" county="Falls" elevation="300" latitude="-17.001944" location="Cunene River" longLatPrecision="21" longitude="13.2491665" municipality="Hot Springs Campsite" pageId="14" pageNumber="2339" specimenCode="SAIAB 78781, KUNE24" specimenCount="1" typeStatus="holotype">
<typeStatus id="63568806FFC5FFAE5ED3FC22FED9FBB9" box="[211,322,1001,1025]" pageId="14" pageNumber="2339" type="holotype">Holotype</typeStatus>
:
<specimenCode id="EC4B9EDFFFC5FFAE5F4CFC22FE73FBB9" box="[332,488,1001,1025]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="14" pageNumber="2339" type="Other type of research institution/biorepository">SAIAB 78781</specimenCode>
(field no.
<specimenCode id="EC4B9EDFFFC5FFAE5C58FC22FD51FBB9" box="[600,714,1001,1025]" collectionCode="KUNE" country="China" lsid="urn:lsid:biocol.org:col:13450" name="Kunming Institute of Ecology, Academia Sinica" pageId="14" pageNumber="2339" type="Herbarium">KUNE24</specimenCode>
), live SL
<quantity id="7B159B41FFC5FFAE5D31FC22FC11FBB9" box="[817,906,1001,1025]" metricMagnitude="-1" metricUnit="m" metricValue="2.09" pageId="14" pageNumber="2339" unit="cm" value="20.9">20.9 cm</quantity>
, fixed SL
<quantity id="7B159B41FFC5FFAE5DF9FC22FBC9FBB9" box="[1017,1106,1001,1025]" metricMagnitude="-1" metricUnit="m" metricValue="2.02" pageId="14" pageNumber="2339" unit="cm" value="20.2">20.2 cm</quantity>
, fixed TL
<quantity id="7B159B41FFC5FFAE5EFEFBC2FEC2FB99" box="[254,345,1033,1057]" metricMagnitude="-1" metricUnit="m" metricValue="2.29" pageId="14" pageNumber="2339" unit="cm" value="22.9">22.9 cm</quantity>
, male,
<collectingCountry id="C4FA7634FFC5FFAE5FA8FBC2FD94FB99" box="[424,527,1033,1057]" name="Namibia" pageId="14" pageNumber="2339">Namibia</collectingCountry>
:
<location id="B932607FFFC5FFAE5C1BFBC2FD21FB99" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC5FFAE5C1BFBC2FD21FB99" box="[539,698,1033,1057]" country="Namibia" county="Falls" latitude="-17.001944" longLatPrecision="21" longitude="13.2491665" municipality="Hot Springs Campsite" name="Cunene River" pageId="14" pageNumber="2339">Cunene River</location>
:
<location id="B932607FFFC5FFAE5CC5FBC1FCCBFB99" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC5FFAE5CC5FBC1FCCBFB99" box="[709,848,1033,1057]" country="Namibia" county="Falls" latitude="-17.001944" longLatPrecision="21" longitude="13.2491665" municipality="Hot Springs Campsite" name="Epupa Falls" pageId="14" pageNumber="2339">Epupa Falls</location>
,
<collectingMunicipality id="5C36ACDEFFC5FFAE5D5BFBC1FBC2FB99" box="[859,1113,1033,1057]" pageId="14" pageNumber="2339">Hot Springs Campsite</collectingMunicipality>
, estimated
<quantity id="7B159B41FFC5FFAE5F1FFBE2FEFFFBF9" box="[287,356,1065,1089]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="2339" unit="m" value="300.0">
<elevation id="37C0D197FFC5FFAE5F1FFBE2FEFFFBF9" box="[287,356,1065,1089]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="2339" unit="m" value="300.0">300 m</elevation>
</quantity>
upstream from the
<collectingCounty id="55334E28FFC5FFAE5C40FBE2FDE3FBF9" box="[576,632,1065,1089]" pageId="14" pageNumber="2339">Falls</collectingCounty>
,
<geoCoordinate id="D9D95063FFC5FFAE5C83FBE2FC9CFBF9" box="[643,775,1061,1089]" degrees="17" direction="south" minutes="00" orientation="latitude" pageId="14" pageNumber="2339" precision="15" seconds="07" value="-17.001944">
17
<superScript id="4B989BECFFC5FFAE5C9DFBEEFD3DFB8E" attach="left" box="[669,678,1061,1078]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
00
<superScript id="4B989BECFFC5FFAE5CC2FBEEFD5CFB8E" attach="left" box="[706,711,1061,1078]" fontSize="7" pageId="14" pageNumber="2339">′</superScript>
07
<superScript id="4B989BECFFC5FFAE5CE3FBEEFD75FB8E" attach="left" box="[739,750,1061,1078]" fontSize="7" pageId="14" pageNumber="2339">′′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC5FFAE5D12FBE2FC01FBF9" box="[786,922,1061,1089]" degrees="13" direction="east" minutes="14" orientation="longitude" pageId="14" pageNumber="2339" precision="15" seconds="57" value="13.2491665">
13
<superScript id="4B989BECFFC5FFAE5D2CFBEEFCAEFB8E" attach="left" box="[812,821,1061,1078]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
14
<superScript id="4B989BECFFC5FFAE5D51FBEEFCCDFB8E" attach="left" box="[849,854,1061,1078]" fontSize="7" pageId="14" pageNumber="2339">′</superScript>
57
<superScript id="4B989BECFFC5FFAE5D72FBEEFCE6FB8E" attach="left" box="[882,893,1061,1078]" fontSize="7" pageId="14" pageNumber="2339">′′</superScript>
E
</geoCoordinate>
, about
<quantity id="7B159B41FFC5FFAE5DECFBE2FBAAFBF9" box="[1004,1073,1065,1089]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="2339" unit="m" value="600.0">
<elevation id="37C0D197FFC5FFAE5DECFBE2FBAAFBF9" box="[1004,1073,1065,1089]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="2339" unit="m" value="600.0">600 m</elevation>
</quantity>
altitude,
<date id="C8531064FFC5FFAE5ED3FB82FE1DFBD9" box="[211,390,1097,1121]" pageId="14" pageNumber="2339" value="2006-08-15">
<collectingDate id="D817E98CFFC5FFAE5ED3FB82FE1DFBD9" box="[211,390,1097,1121]" pageId="14" pageNumber="2339" value="2006-08-15">15 August 2006</collectingDate>
</date>
, coll.
<collectorName id="11185372FFC5FFAE5FC9FB81FDA1FBD9" box="[457,570,1097,1121]" pageId="14" pageNumber="2339">E. Swartz</collectorName>
, B. Kramer and
<collectorName id="11185372FFC5FFAE5CF8FB81FCE4FBD9" box="[760,895,1097,1121]" pageId="14" pageNumber="2339">L. da Costa</collectorName>
at ≤
<quantity id="7B159B41FFC5FFAE5DBFFB82FC64FBD9" box="[959,1023,1097,1121]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="2339" unit="m" value="1.5">
<elevation id="37C0D197FFC5FFAE5DBFFB82FC64FBD9" box="[959,1023,1097,1121]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="2339" unit="m" value="1.5">1.5 m</elevation>
</quantity>
water depth
</materialsCitation>
.
<materialsCitation id="0C853CF9FFC5FFAE5ED3FBA2FD8AFB39" box="[211,529,1129,1153]" collectionCode="SAIAB" country="Namibia" pageId="14" pageNumber="2339" specimenCode="SAIAB 78780" specimenCount="1" typeStatus="paratype">
<typeStatus id="63568806FFC5FFAE5ED3FBA2FEDEFB39" box="[211,325,1129,1153]" pageId="14" pageNumber="2339" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="EC4B9EDFFFC5FFAE5F4EFBA2FE72FB39" box="[334,489,1129,1153]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="14" pageNumber="2339" type="Other type of research institution/biorepository">SAIAB 78780</specimenCode>
(2)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC5FFAE5C1BFBA2FD2FFB39" box="[539,692,1129,1153]" collectionCode="SAIAB" country="Namibia" pageId="14" pageNumber="2339" specimenCode="SAIAB78792" specimenCount="1" typeStatus="paratype">
<specimenCode id="EC4B9EDFFFC5FFAE5C1BFBA2FD2FFB39" box="[539,692,1129,1153]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="14" pageNumber="2339" type="Other type of research institution/biorepository">SAIAB78792</specimenCode>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC5FFAE5CBEFBA2FCF3FB39" accessionNumber="ZSM 38526" box="[702,872,1129,1153]" collectionCode="ZSM" country="Namibia" pageId="14" pageNumber="2339" specimenCount="1" typeStatus="paratype">
<collectionCode id="DAFCAE61FFC5FFAE5CBEFBA2FD62FB39" box="[702,761,1129,1153]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
38526 (2)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC5FFAE5D72FBA2FB80FB39" accessionNumber="ZSM 38527" box="[882,1051,1129,1153]" collectionCode="ZSM" country="Namibia" pageId="14" pageNumber="2339" specimenCount="1" typeStatus="paratype">
<collectionCode id="DAFCAE61FFC5FFAE5D72FBA2FC36FB39" box="[882,941,1129,1153]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
38527 (2)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC5FFAE5A26FBA2FDD9FA98" collectingDate="2006-08-12" collectingDateMax="2006-08-17" collectingDateMin="2006-08-12" collectorName="E. Swartz &amp; B. Kramer &amp; L. da Costa" country="Namibia" county="Cunene River" elevation="300" latitude="-17.001944" location="Cunene River" longLatPrecision="21" longitude="13.2491665" municipality="Falls" pageId="14" pageNumber="2339" specimenCount="1" typeStatus="paratype">
size range
<quantity id="7B159B41FFC5FFAE5ED3FB43FEF2FB18" box="[211,361,1160,1184]" metricMagnitude="-1" metricUnit="m" metricValue="1.515" metricValueMax="2.02" metricValueMin="1.01" pageId="14" pageNumber="2339" unit="cm" value="15.149999999999999" valueMax="20.2" valueMin="10.1">10.1–20.2 cm</quantity>
SL,
<collectingCountry id="C4FA7634FFC5FFAE5FA1FB43FD93FB18" box="[417,520,1160,1184]" name="Namibia" pageId="14" pageNumber="2339">Namibia</collectingCountry>
:
<location id="B932607FFFC5FFAE5C14FB43FD2EFB18" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC5FFAE5C14FB43FD2EFB18" box="[532,693,1160,1184]" country="Namibia" county="Cunene River" latitude="-17.001944" longLatPrecision="21" longitude="13.2491665" municipality="Falls" name="Cunene River" pageId="14" pageNumber="2339">Cunene River</location>
:
<location id="B932607FFFC5FFAE5CC0FB42FCD7FB18" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC5FFAE5CC0FB42FCD7FB18" box="[704,844,1160,1184]" country="Namibia" county="Cunene River" latitude="-17.001944" longLatPrecision="21" longitude="13.2491665" municipality="Falls" name="Epupa Falls" pageId="14" pageNumber="2339">Epupa Falls</location>
, Hot Springs Campsite, estimated
<quantity id="7B159B41FFC5FFAE5F1FFB63FEFFFB78" box="[287,356,1192,1216]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="2339" unit="m" value="300.0">
<elevation id="37C0D197FFC5FFAE5F1FFB63FEFFFB78" box="[287,356,1192,1216]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="2339" unit="m" value="300.0">300 m</elevation>
</quantity>
upstream from the
<collectingMunicipality id="5C36ACDEFFC5FFAE5C40FB63FDE3FB78" box="[576,632,1192,1216]" pageId="14" pageNumber="2339">Falls</collectingMunicipality>
,
<geoCoordinate id="D9D95063FFC5FFAE5C83FB63FC9CFB78" box="[643,775,1188,1216]" degrees="17" direction="south" minutes="00" orientation="latitude" pageId="14" pageNumber="2339" precision="15" seconds="07" value="-17.001944">
17
<superScript id="4B989BECFFC5FFAE5C9DFB6FFD3DFB0D" attach="left" box="[669,678,1188,1205]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
00
<superScript id="4B989BECFFC5FFAE5CC2FB6FFD5CFB0D" attach="left" box="[706,711,1188,1205]" fontSize="7" pageId="14" pageNumber="2339">′</superScript>
07
<superScript id="4B989BECFFC5FFAE5CE3FB6FFD75FB0D" attach="left" box="[739,750,1188,1205]" fontSize="7" pageId="14" pageNumber="2339">′′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC5FFAE5D12FB63FC01FB78" box="[786,922,1188,1216]" degrees="13" direction="east" minutes="14" orientation="longitude" pageId="14" pageNumber="2339" precision="15" seconds="57" value="13.2491665">
13
<superScript id="4B989BECFFC5FFAE5D2CFB6FFCAEFB0D" attach="left" box="[812,821,1188,1205]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
14
<superScript id="4B989BECFFC5FFAE5D51FB6FFCCDFB0D" attach="left" box="[849,854,1188,1205]" fontSize="7" pageId="14" pageNumber="2339">′</superScript>
57
<superScript id="4B989BECFFC5FFAE5D72FB6FFCE6FB0D" attach="left" box="[882,893,1188,1205]" fontSize="7" pageId="14" pageNumber="2339">′′</superScript>
E
</geoCoordinate>
, about
<quantity id="7B159B41FFC5FFAE5DECFB63FBAAFB78" box="[1004,1073,1192,1216]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="2339" unit="m" value="600.0">
<elevation id="37C0D197FFC5FFAE5DECFB63FBAAFB78" box="[1004,1073,1192,1216]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="2339" unit="m" value="600.0">600 m</elevation>
</quantity>
altitude, coll.
<collectorName id="11185372FFC5FFAE5F0CFB02FEE5FB58" box="[268,382,1224,1248]" pageId="14" pageNumber="2339">E. Swartz</collectorName>
,
<collectorName id="11185372FFC5FFAE5F8CFB02FD9CFB58" box="[396,519,1225,1248]" pageId="14" pageNumber="2339">B. Kramer</collectorName>
, and
<collectorName id="11185372FFC5FFAE5C47FB02FD4BFB58" box="[583,720,1224,1248]" pageId="14" pageNumber="2339">L. da Costa</collectorName>
at ≤
<quantity id="7B159B41FFC5FFAE5D14FB03FCCEFB58" box="[788,853,1224,1248]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="2339" unit="m" value="1.5">
<elevation id="37C0D197FFC5FFAE5D14FB03FCCEFB58" box="[788,853,1224,1248]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="2339" unit="m" value="1.5">1.5 m</elevation>
</quantity>
water depth,
<collectingCounty id="55334E28FFC5FFAE5DF6FB03FB08FB58" box="[1014,1171,1224,1248]" pageId="14" pageNumber="2339">Cunene River</collectingCounty>
water at Hot Springs: Saturday,
<date id="C8531064FFC5FFAE5C42FB23FD6EFAB8" box="[578,757,1256,1280]" pageId="14" pageNumber="2339" value="2006-08-12">
<collectingDate id="D817E98CFFC5FFAE5C42FB23FD6EFAB8" box="[578,757,1256,1280]" pageId="14" pageNumber="2339" value="2006-08-12">12 August 2006</collectingDate>
</date>
, 12.50 h: 19.9
<superScript id="4B989BECFFC5FFAE5D9BFB2FFC3FFB4D" attach="right" box="[923,932,1252,1269]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
C, 48 µS
<superScript id="4B989BECFFC5FFAE5A13FB22FBD7FB4D" attach="left" box="[1043,1100,1252,1280]" fontSize="7" pageId="14" pageNumber="2339">cm−1</superScript>
, from 11 Augure 2006–
<date id="C8531064FFC5FFAE5F93FAC3FDD9FA98" box="[403,578,1288,1312]" pageId="14" pageNumber="2339" value="2006-08-17">
<collectingDate id="D817E98CFFC5FFAE5F93FAC3FDD9FA98" box="[403,578,1288,1312]" pageId="14" pageNumber="2339" value="2006-08-17">17 August 2006</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFC5FFAE5EFEFAE2FBE5FA67" blockId="14.[211,1172,962,1726]" pageId="14" pageNumber="2339">
<emphasis id="8E99EAB6FFC5FFAE5EFEFAE2FEF4FAF8" box="[254,367,1321,1344]" italics="true" pageId="14" pageNumber="2339">Non-types</emphasis>
.
<materialsCitation id="0C853CF9FFC5FFAE5F7DFAE3FDD8FAF8" box="[381,579,1320,1344]" collectionCode="SAIAB" pageId="14" pageNumber="2339" specimenCode="SAIAB 78785" specimenCount="1">
<specimenCode id="EC4B9EDFFFC5FFAE5F7DFAE3FD81FAF8" box="[381,538,1320,1344]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="14" pageNumber="2339" type="Other type of research institution/biorepository">SAIAB 78785</specimenCode>
(2)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC5FFAE5C4EFAE3FC8FFAF8" box="[590,788,1320,1344]" collectionCode="SAIAB" pageId="14" pageNumber="2339" specimenCode="SAIAB 78789" specimenCount="1">
<specimenCode id="EC4B9EDFFFC5FFAE5C4EFAE3FD70FAF8" box="[590,747,1320,1344]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="14" pageNumber="2339" type="Other type of research institution/biorepository">SAIAB 78789</specimenCode>
(2)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC5FFAE5D20FAE3FC30FAF8" accessionNumber="ZSM 38528" box="[800,939,1320,1344]" collectionCode="ZSM" pageId="14" pageNumber="2339" specimenCount="1">
<collectionCode id="DAFCAE61FFC5FFAE5D20FAE3FCC0FAF8" box="[800,859,1320,1344]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
38528,
</materialsCitation>
<materialsCitation id="0C853CF9FFC5FFAE5DB2FAE3FBC5FAF8" accessionNumber="ZSM 38529" box="[946,1118,1320,1344]" collectionCode="ZSM" pageId="14" pageNumber="2339" specimenCount="1">
<collectionCode id="DAFCAE61FFC5FFAE5DB2FAE3FC76FAF8" box="[946,1005,1320,1344]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
38529 (2)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC5FFAE5A6AFAE3FBE1FA67" collectingDate="2004-08-19" collectingDateMax="2006-08-23" collectingDateMin="2004-08-19" collectorName="E. Swartz &amp; B. Kramer" elevation="800" latitude="-17.406666" location="Falls" longLatPrecision="21" longitude="14.216944" municipality="Ruacana Falls" pageId="14" pageNumber="2339" specimenCount="1" stateProvince="Cunene">
size range
<quantity id="7B159B41FFC5FFAE5F1CFA83FE2FFAD8" box="[284,436,1352,1376]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="1.54" metricValueMin="1.16" pageId="14" pageNumber="2339" unit="cm" value="13.5" valueMax="15.4" valueMin="11.6">11.6–15.4 cm</quantity>
SL (live),
<collectingMunicipality id="5C36ACDEFFC5FFAE5C31FA82FD4CFAD8" box="[561,727,1352,1376]" pageId="14" pageNumber="2339">Ruacana Falls</collectingMunicipality>
, Hippo Pool Campsite, just below the
<location id="B932607FFFC5FFAE5ED3FAA3FE90FA38" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC5FFAE5ED3FAA3FE90FA38" box="[211,267,1384,1408]" latitude="-17.406666" longLatPrecision="21" longitude="14.216944" municipality="Ruacana Falls" name="Falls" pageId="14" pageNumber="2339" stateProvince="Cunene">Falls</location>
,
<geoCoordinate id="D9D95063FFC5FFAE5F17FAA3FE07FA38" box="[279,412,1380,1408]" degrees="17" direction="south" minutes="24" orientation="latitude" pageId="14" pageNumber="2339" precision="15" seconds="24" value="-17.406666">
17
<superScript id="4B989BECFFC5FFAE5F31FAAFFEA1FACD" attach="left" box="[305,314,1380,1397]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
24
<superScript id="4B989BECFFC5FFAE5F56FAAFFEC0FACD" attach="left" box="[342,347,1380,1397]" fontSize="7" pageId="14" pageNumber="2339">′</superScript>
24
<superScript id="4B989BECFFC5FFAE5F77FAAFFE19FACD" attach="left" box="[375,386,1380,1397]" fontSize="7" pageId="14" pageNumber="2339">′′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC5FFAE5FA8FAA3FDA9FAC7" box="[424,562,1380,1408]" degrees="14" direction="east" minutes="13" orientation="longitude" pageId="14" pageNumber="2339" precision="15" seconds="01" value="14.216944">
14
<superScript id="4B989BECFFC5FFAE5FC2FAAFFE50FACD" attach="left" box="[450,459,1380,1397]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
13’01” E
</geoCoordinate>
, about
<quantity id="7B159B41FFC5FFAE5C87FAA3FD56FAC7" box="[647,717,1384,1408]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="2339" unit="m" value="800.0">
<elevation id="37C0D197FFC5FFAE5C87FAA3FD56FAC7" box="[647,717,1384,1408]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="2339" unit="m" value="800.0">800 m</elevation>
</quantity>
altitude, coll.
<collectorName id="11185372FFC5FFAE5D6FFAA3FC47FA38" box="[879,988,1384,1408]" pageId="14" pageNumber="2339">E. Swartz</collectorName>
and
<collectorName id="11185372FFC5FFAE5A16FAA3FB0BFAC7" box="[1046,1168,1384,1407]" pageId="14" pageNumber="2339">B. Kramer</collectorName>
, at ≤
<quantity id="7B159B41FFC5FFAE5F09FA4CFEDCFA27" box="[265,327,1415,1439]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="2339" unit="m" value="1.5">
<elevation id="37C0D197FFC5FFAE5F09FA4CFEDCFA27" box="[265,327,1415,1439]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="2339" unit="m" value="1.5">1.5 m</elevation>
</quantity>
water depth,
<collectingRegion id="7E29F846FFC5FFAE5FDFFA4CFDAEFA27" box="[479,565,1415,1439]" country="Angola" name="Cunene" pageId="14" pageNumber="2339">Cunene</collectingRegion>
River water at Hippo Pool: Saturday,
<date id="C8531064FFC5FFAE5DDFFA4CFB14FA27" box="[991,1167,1415,1439]" pageId="14" pageNumber="2339" value="2004-08-19">
<collectingDate id="D817E98CFFC5FFAE5DDFFA4CFB14FA27" box="[991,1167,1415,1439]" pageId="14" pageNumber="2339" value="2004-08-19">19 August 2004</collectingDate>
</date>
, 10.00 h: 21.1
<superScript id="4B989BECFFC5FFAE5F74FA6FFEE6FA0D" attach="right" box="[372,381,1444,1461]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
C, 45.8 µS
<superScript id="4B989BECFFC5FFAE5C08FA63FDDAFA0C" attach="left" box="[520,577,1443,1471]" fontSize="7" pageId="14" pageNumber="2339">cm−1</superScript>
;
<date id="C8531064FFC5FFAE5C55FA6CFC96FA07" box="[597,781,1447,1471]" pageId="14" pageNumber="2339" value="2006-08-20">
<collectingDate id="D817E98CFFC5FFAE5C55FA6CFC96FA07" box="[597,781,1447,1471]" pageId="14" pageNumber="2339" value="2006-08-20">20 August 2006</collectingDate>
</date>
, 10.00 h, 19.8
<superScript id="4B989BECFFC5FFAE5DBEFA6FFC5CFA0D" attach="right" box="[958,967,1444,1461]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
C, 45.4 µS
<superScript id="4B989BECFFC5FFAE5A51FA63FB11FA0C" attach="left" box="[1105,1162,1443,1471]" fontSize="7" pageId="14" pageNumber="2339">cm−1</superScript>
; 21 August, 10.18 h, 19.4
<superScript id="4B989BECFFC5FFAE5FE8FA08FE6AFA6C" attach="right" box="[488,497,1475,1492]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
C, 44.2 µS
<superScript id="4B989BECFFC5FFAE5C70FA03FD32FA6C" attach="left" box="[624,681,1475,1503]" fontSize="7" pageId="14" pageNumber="2339">cm−1</superScript>
; from
<date id="C8531064FFC5FFAE5CF6FA0CFC38FA67" box="[758,931,1479,1503]" pageId="14" pageNumber="2339" value="2006-08-18">
<collectingDate id="D817E98CFFC5FFAE5CF6FA0CFC38FA67" box="[758,931,1479,1503]" pageId="14" pageNumber="2339" value="2006-08-18">18 August 2006</collectingDate>
</date>
to
<date id="C8531064FFC5FFAE5DC8FA0CFBE1FA67" box="[968,1146,1479,1503]" pageId="14" pageNumber="2339" value="2006-08-23">
<collectingDate id="D817E98CFFC5FFAE5DC8FA0CFBE1FA67" box="[968,1146,1479,1503]" pageId="14" pageNumber="2339" value="2006-08-23">23 August 2006</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFC5FFAE5EFEFA2CFD30F906" blockId="14.[211,1172,962,1726]" pageId="14" pageNumber="2339">
<materialsCitation id="0C853CF9FFC5FFAE5EFEFA2CFDA8FA47" accessionNumber="ZSM 41761" box="[254,563,1511,1535]" collectionCode="ZSM" pageId="14" pageNumber="2339" specimenCount="1">
<collectionCode id="DAFCAE61FFC5FFAE5EFEFA2CFEA2FA47" box="[254,313,1511,1535]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
41761 (11), specimens
</materialsCitation>
<materialsCitation id="0C853CF9FFC5FFAE5C39FA2CFDC3FA47" box="[569,600,1511,1535]" collectionCode="R" pageId="14" pageNumber="2339" specimenCode="R1" specimenCount="1">
<specimenCode id="EC4B9EDFFFC5FFAE5C39FA2CFDC3FA47" box="[569,600,1511,1535]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="2339">R1</specimenCode>
</materialsCitation>
–
<materialsCitation id="0C853CF9FFC5FFAE5C69FA2CFB9AF9A7" collectingDate="2009-12-15" collectionCode="R" collectorName="F. H. van der Bank" elevation="2" latitude="-17.2601" location="River" longLatPrecision="1" longitude="11.764867" pageId="14" pageNumber="2339" specimenCode="R11" specimenCount="1" stateProvince="Cunene">
<specimenCode id="EC4B9EDFFFC5FFAE5C69FA2CFD01FA47" box="[617,666,1511,1535]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="2339">R11</specimenCode>
, from the
<collectingRegion id="7E29F846FFC5FFAE5D0AFA2CFCFBFA47" box="[778,864,1511,1535]" country="Angola" name="Cunene" pageId="14" pageNumber="2339">Cunene</collectingRegion>
<location id="B932607FFFC5FFAE5D67FA2CFC3EFA47" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC5FFAE5D67FA2CFC3EFA47" box="[871,933,1511,1535]" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" name="River" pageId="14" pageNumber="2339" stateProvince="Cunene">River</location>
mouth,
<geoCoordinate id="D9D95063FFC5FFAE5A03FA2CFB14FA47" box="[1027,1167,1507,1535]" degrees="17" direction="south" minutes="15.606" orientation="latitude" pageId="14" pageNumber="2339" precision="1" value="-17.2601">
17
<superScript id="4B989BECFFC5FFAE5A1EFA28FBBCFA4C" attach="none" box="[1054,1063,1507,1524]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
15.606
<superScript id="4B989BECFFC5FFAE5A72FA28FBECFA4C" attach="left" box="[1138,1143,1507,1524]" fontSize="7" pageId="14" pageNumber="2339">′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC5FFAE5ED3F9CCFEFEF9A7" box="[211,357,1539,1567]" degrees="11" direction="east" minutes="45.892" orientation="longitude" pageId="14" pageNumber="2339" precision="1" value="11.764867">
11
<superScript id="4B989BECFFC5FFAE5EEEF9C8FF6CF9AC" attach="none" box="[238,247,1539,1556]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
45.892
<superScript id="4B989BECFFC5FFAE5F42F9C8FEDCF9AC" attach="left" box="[322,327,1539,1556]" fontSize="7" pageId="14" pageNumber="2339">′</superScript>
E
</geoCoordinate>
, altitude
<quantity id="7B159B41FFC5FFAE5FD1F9CCFD9BF9A7" box="[465,512,1543,1567]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="2339" unit="m" value="2.0">
<elevation id="37C0D197FFC5FFAE5FD1F9CCFD9BF9A7" box="[465,512,1543,1567]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="2339" unit="m" value="2.0">2 m</elevation>
</quantity>
,
<date id="C8531064FFC5FFAE5C0CF9CCFD78F9A7" box="[524,739,1543,1567]" pageId="14" pageNumber="2339" value="2009-12-15">
<collectingDate id="D817E98CFFC5FFAE5C0CF9CCFD78F9A7" box="[524,739,1543,1567]" pageId="14" pageNumber="2339" value="2009-12-15">15 December 2009</collectingDate>
</date>
, coll.
<collectorName id="11185372FFC5FFAE5D28F9C3FB9AF9A7" box="[808,1025,1543,1567]" pageId="14" pageNumber="2339">F.H. van der Bank</collectorName>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC5FFAE5A0EF9CCFB0BF9F8" accessionNumber="ZSM 41762" collectingDate="2010-11-08" collectionCode="ZSM" collectorName="S. Voges" latitude="-17.272083" longLatPrecision="1" longitude="11.7862835" pageId="14" pageNumber="2339" specimenCount="1" stateProvince="Cunene">
<collectionCode id="DAFCAE61FFC5FFAE5A0EF9CCFBD2F9A7" box="[1038,1097,1543,1567]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
41762 (2), specimens 49 and 49,
<geoCoordinate id="D9D95063FFC5FFAE5FF3F9ECFDE4F987" box="[499,639,1571,1599]" degrees="17" direction="south" minutes="16.325" orientation="latitude" pageId="14" pageNumber="2339" precision="1" value="-17.272083">
17
<superScript id="4B989BECFFC5FFAE5C0DF9E8FD8DF98C" attach="left" box="[525,534,1571,1588]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
16.325
<superScript id="4B989BECFFC5FFAE5C62F9E8FDFCF98C" attach="left" box="[610,615,1571,1588]" fontSize="7" pageId="14" pageNumber="2339">′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC5FFAE5C8AF9ECFC81F987" box="[650,794,1571,1599]" degrees="11" direction="east" minutes="47.177" orientation="longitude" pageId="14" pageNumber="2339" precision="1" value="11.7862835">
11
<superScript id="4B989BECFFC5FFAE5CA4F9E8FD36F98C" attach="left" box="[676,685,1571,1588]" fontSize="7" pageId="14" pageNumber="2339">◦</superScript>
47.177
<superScript id="4B989BECFFC5FFAE5CF9F9E8FD65F98C" attach="left" box="[761,766,1571,1588]" fontSize="7" pageId="14" pageNumber="2339">′</superScript>
E
</geoCoordinate>
,
<date id="C8531064FFC5FFAE5D25F9ECFC75F987" box="[805,1006,1575,1599]" pageId="14" pageNumber="2339" value="2010-11-08">
<collectingDate id="D817E98CFFC5FFAE5D25F9ECFC75F987" box="[805,1006,1575,1599]" pageId="14" pageNumber="2339" value="2010-11-08">8 November 2010</collectingDate>
</date>
, coll.
<collectorName id="11185372FFC5FFAE5A2FF9ECFB0BF9F8" box="[1071,1168,1575,1600]" pageId="14" pageNumber="2339">S. Voges</collectorName>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC5FFAE5ED3F98CFC65F9D8" accessionNumber="ZSM 41765" box="[211,1022,1607,1632]" collectingDate="2011-01-17" collectionCode="ZSM" collectorName="S. Voges" pageId="14" pageNumber="2339" specimenCount="1" stateProvince="Cunene">
<collectionCode id="DAFCAE61FFC5FFAE5ED3F98CFE95F9E7" box="[211,270,1607,1631]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
41765, specimen C113, same place,
<date id="C8531064FFC5FFAE5CA3F98CFCC7F9E7" box="[675,860,1607,1632]" pageId="14" pageNumber="2339" value="2011-01-17">
<collectingDate id="D817E98CFFC5FFAE5CA3F98CFCC7F9E7" box="[675,860,1607,1632]" pageId="14" pageNumber="2339" value="2011-01-17">17 January 2011</collectingDate>
</date>
, coll.
<collectorName id="11185372FFC5FFAE5D9DF98CFC65F9D8" box="[925,1022,1607,1632]" pageId="14" pageNumber="2339">S. Voges</collectorName>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC5FFAE5A08F98CFCD4F938" accessionNumber="ZSM 41763" collectingDate="2011-07-13" collectionCode="ZSM" collectorName="S. Voges" pageId="14" pageNumber="2339" specimenCount="1" stateProvince="Cunene">
<collectionCode id="DAFCAE61FFC5FFAE5A08F98CFBD8F9E7" box="[1032,1091,1607,1631]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
41763, specimen Ü7, same place,
<date id="C8531064FFC5FFAE5C0AF9ACFD3BF9C7" box="[522,672,1639,1663]" pageId="14" pageNumber="2339" value="2011-07-13">
<collectingDate id="D817E98CFFC5FFAE5C0AF9ACFD3BF9C7" box="[522,672,1639,1663]" pageId="14" pageNumber="2339" value="2011-07-13">13 July 2011</collectingDate>
</date>
, coll.
<collectorName id="11185372FFC5FFAE5CEAF9ACFCD4F938" box="[746,847,1639,1664]" pageId="14" pageNumber="2339">S. Voges</collectorName>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC5FFAE5D5DF9ACFC35F927" accessionNumber="ZSM 41764" collectingDate="2011-11-22" collectionCode="ZSM" collectorName="S. Voges. The" pageId="14" pageNumber="2339" specimenCount="1" stateProvince="Cunene">
<collectionCode id="DAFCAE61FFC5FFAE5D5DF9ACFC03F9C7" box="[861,920,1639,1663]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
41764 (9), specimens Ä110–Ä118, same place,
<date id="C8531064FFC5FFAE5FF4F94CFD54F927" box="[500,719,1671,1695]" pageId="14" pageNumber="2339" value="2011-11-22">
<collectingDate id="D817E98CFFC5FFAE5FF4F94CFD54F927" box="[500,719,1671,1695]" pageId="14" pageNumber="2339" value="2011-11-22">22 November 2011</collectingDate>
</date>
, coll.
<collectorName id="11185372FFC5FFAE5D14F94CFC35F927" box="[788,942,1671,1695]" pageId="14" pageNumber="2339">S. Voges. The</collectorName>
</materialsCitation>
specimens from the
<collectingRegion id="7E29F846FFC5FFAE5ED3F96DFEB2F906" box="[211,297,1702,1726]" country="Angola" name="Cunene" pageId="14" pageNumber="2339">Cunene</collectingRegion>
mouth were not studied for
<collectionCode id="DAFCAE61FFC5FFAE5C69F96DFD3CF906" box="[617,679,1702,1726]" pageId="14" pageNumber="2339">EOD</collectionCode>
.
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5EBAFF6CFD90FEA7" blockId="15.[143,1104,167,542]" pageId="15" pageNumber="2340">
<materialsCitation id="0C853CF9FFC4FFAF5EBAFF6CFD9CFEA7" collectingDate="2009-12-15" collectionCode="IPMB" collectorName="F. H. van der Bank" country="Namibia" county="River Mouth" latitude="-17.2601" location="Samples" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" pageId="15" pageNumber="2340" specimenCode="IPMB 57459-57469" specimenCount="1">
<location id="B932607FFFC4FFAF5EBAFF6CFE83FF07" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5EBAFF6CFE83FF07" box="[186,280,167,191]" country="Namibia" county="River Mouth" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" name="Samples" pageId="15" pageNumber="2340">Samples</location>
examined for genetics. DNA samples are stored at the
<location id="B932607FFFC4FFAF5DCAFF6CFE99FF67" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5DCAFF6CFE99FF67" country="Namibia" county="River Mouth" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" name="Institute of Pharmacy" pageId="15" pageNumber="2340">Institute of Pharmacy</location>
and
<location id="B932607FFFC4FFAF5F4CFF0CFDEEFF67" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5F4CFF0CFDEEFF67" box="[332,629,199,223]" country="Namibia" county="River Mouth" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" name="Molecular Biotechnology" pageId="15" pageNumber="2340">Molecular Biotechnology</location>
,
<location id="B932607FFFC4FFAF5C89FF0CFC11FF67" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5C89FF0CFC11FF67" box="[649,906,199,223]" country="Namibia" county="River Mouth" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" name="Heidelberg University" pageId="15" pageNumber="2340">Heidelberg University</location>
(
<collectionCode id="DAFCAE61FFC4FFAF5DA2FF0CFC76FF67" box="[930,1005,199,223]" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB</collectionCode>
).
<specimenCode id="EC4B9EDFFFC4FFAF5A09FF03FEBEFF47" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 57459–57469</specimenCode>
,
<collectingCountry id="C4FA7634FFC4FFAF5F31FF2CFE03FF47" box="[305,408,231,255]" name="Namibia" pageId="15" pageNumber="2340">Namibia</collectingCountry>
:
<location id="B932607FFFC4FFAF5FA4FF2CFD07FF47" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5FA4FF2CFD07FF47" box="[420,668,231,255]" country="Namibia" county="River Mouth" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" name="Cunene River Mouth" pageId="15" pageNumber="2340">
<collectingMunicipality id="5C36ACDEFFC4FFAF5FA4FF2CFDDAFF47" box="[420,577,231,255]" pageId="15" pageNumber="2340">
Cunene
<collectingCounty id="55334E28FFC4FFAF5C03FF2CFDDAFF47" box="[515,577,231,255]" pageId="15" pageNumber="2340">River</collectingCounty>
</collectingMunicipality>
Mouth
</location>
,
<geoCoordinate id="D9D95063FFC4FFAF5CA9FF2CFCACFF47" box="[681,823,227,255]" degrees="17" direction="south" minutes="15.606" orientation="latitude" pageId="15" pageNumber="2340" precision="1" value="-17.2601">
17
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15.606’ S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC4FFAF5D44FF2CFC4CFF47" box="[836,983,227,255]" degrees="11" direction="east" minutes="45.892" orientation="longitude" pageId="15" pageNumber="2340" precision="1" value="11.764867">
11
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45.892’ E
</geoCoordinate>
, coll.
<collectorName id="11185372FFC4FFAF5A1DFF23FEB2FEA7" pageId="15" pageNumber="2340">F.H. van der Bank</collectorName>
,
<date id="C8531064FFC4FFAF5F34FECCFD9CFEA7" box="[308,519,263,287]" pageId="15" pageNumber="2340" value="2009-12-15">
<collectingDate id="D817E98CFFC4FFAF5F34FECCFD9CFEA7" box="[308,519,263,287]" pageId="15" pageNumber="2340" value="2009-12-15">15 December 2009</collectingDate>
</date>
</materialsCitation>
;
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5EBAFEE3FE58FE27" blockId="15.[143,1104,167,542]" pageId="15" pageNumber="2340">
<materialsCitation id="0C853CF9FFC4FFAF5EBAFEE3FC38FEE7" collectingDate="2006-08-14" collectionCode="IPMB" collectorName="E. Swartz &amp; B. Kramer" country="Namibia" latitude="-17.001944" location="Cunene River" longLatPrecision="21" longitude="13.2491665" municipality="Epupa Falls" pageId="15" pageNumber="2340" specimenCode="IPMB 43971-43974" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5EBAFEE3FE31FE87" box="[186,426,295,319]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 43971–43974</specimenCode>
,
<collectingCountry id="C4FA7634FFC4FFAF5FC3FEECFDB1FE87" box="[451,554,295,319]" name="Namibia" pageId="15" pageNumber="2340">Namibia</collectingCountry>
:
<location id="B932607FFFC4FFAF5C42FEECFD74FE87" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5C42FEECFD74FE87" box="[578,751,295,319]" country="Namibia" latitude="-17.001944" longLatPrecision="21" longitude="13.2491665" municipality="Epupa Falls" name="Cunene River" pageId="15" pageNumber="2340">Cunene River</location>
:
<collectingMunicipality id="5C36ACDEFFC4FFAF5D07FEE3FC05FE87" box="[775,926,295,319]" pageId="15" pageNumber="2340">Epupa Falls</collectingMunicipality>
,
<geoCoordinate id="D9D95063FFC4FFAF5DB7FEECFBD0FE87" box="[951,1099,291,319]" degrees="17" direction="south" minutes="00" orientation="latitude" pageId="15" pageNumber="2340" precision="15" seconds="07" value="-17.001944">
17
<superScript id="4B989BECFFC4FFAF5DD2FEE8FC40FE8C" attach="none" box="[978,987,291,308]" fontSize="7" pageId="15" pageNumber="2340">◦</superScript>
00’07” S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC4FFAF5E8FFE8CFEBFFEE7" box="[143,292,323,351]" degrees="13" direction="east" minutes="14" orientation="longitude" pageId="15" pageNumber="2340" precision="15" seconds="57" value="13.2491665">
13
<superScript id="4B989BECFFC4FFAF5EAAFE88FF28FEEC" attach="none" box="[170,179,323,340]" fontSize="7" pageId="15" pageNumber="2340">◦</superScript>
14’57” E
</geoCoordinate>
, coll.
<collectorName id="11185372FFC4FFAF5F7DFE83FE6FFEE7" box="[381,500,327,351]" pageId="15" pageNumber="2340">E. Swartz</collectorName>
and
<collectorName id="11185372FFC4FFAF5C43FE83FD5DFEE7" box="[579,710,328,351]" pageId="15" pageNumber="2340">B. Kramer</collectorName>
,
<date id="C8531064FFC4FFAF5CDCFE8CFC38FEE7" box="[732,931,327,351]" pageId="15" pageNumber="2340" value="2006-08-14">
<collectingDate id="D817E98CFFC4FFAF5CDCFE8CFC38FEE7" box="[732,931,327,351]" pageId="15" pageNumber="2340" value="2006-08-14">14 August 2006</collectingDate>
</date>
</materialsCitation>
,
<geoCoordinate id="D9D95063FFC4FFAF5DBAFE8CFBD0FEE7" box="[954,1099,323,351]" degrees="17" direction="south" minutes="00" orientation="latitude" pageId="15" pageNumber="2340" precision="15" seconds="07" value="-17.001944">
17
<superScript id="4B989BECFFC4FFAF5DD4FE88FC46FEEC" attach="left" box="[980,989,323,340]" fontSize="7" pageId="15" pageNumber="2340">◦</superScript>
00’07” S
</geoCoordinate>
, E
<geoCoordinate id="D9D95063FFC4FFAF5EA0FEACFEA2FEC7" box="[160,313,355,383]" degrees="013" direction="east" minutes="14" orientation="longitude" pageId="15" pageNumber="2340" precision="15" seconds="57" value="13.2491665">
013
<superScript id="4B989BECFFC4FFAF5EC9FEA8FF49FECC" attach="none" box="[201,210,355,372]" fontSize="7" pageId="15" pageNumber="2340">◦</superScript>
14’57” E
</geoCoordinate>
;
<materialsCitation id="0C853CF9FFC4FFAF5F46FEA3FC10FEC7" box="[326,907,359,383]" collectingDate="2006-08-15" collectionCode="IPMB" country="Namibia" pageId="15" pageNumber="2340" specimenCode="IPMB 43975-43978" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5F46FEA3FDB1FEC7" box="[326,554,359,383]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 43975–43978</specimenCode>
, as before, but
<date id="C8531064FFC4FFAF5CD8FEACFC10FEC7" box="[728,907,359,383]" pageId="15" pageNumber="2340" value="2006-08-15">
<collectingDate id="D817E98CFFC4FFAF5CD8FEACFC10FEC7" box="[728,907,359,383]" pageId="15" pageNumber="2340" value="2006-08-15">15 August 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC4FFAF5D98FEA3FE24FE27" collectingDate="2006-08-17" collectionCode="IPMB" country="Namibia" pageId="15" pageNumber="2340" specimenCode="IPMB 43993" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5D98FEA3FBB7FEC7" box="[920,1068,359,383]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 43993</specimenCode>
, as before, but
<date id="C8531064FFC4FFAF5F0EFE4CFE24FE27" box="[270,447,391,415]" pageId="15" pageNumber="2340" value="2006-08-17">
<collectingDate id="D817E98CFFC4FFAF5F0EFE4CFE24FE27" box="[270,447,391,415]" pageId="15" pageNumber="2340" value="2006-08-17">17 August 2006</collectingDate>
</date>
</materialsCitation>
;
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5EBAFE6CFC8AFDA6" blockId="15.[143,1104,167,542]" pageId="15" pageNumber="2340">
<materialsCitation id="0C853CF9FFC4FFAF5EBAFE6CFD2AFE66" collectingDate="2006-08-19" collectionCode="IPMB" collectorName="E. Swartz &amp; B. Kramer" country="Namibia" latitude="-17.406666" location="Cunene River" longLatPrecision="21" longitude="14.216944" municipality="Ruacana Falls" pageId="15" pageNumber="2340" specimenCode="IPMB 43980" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5EBAFE6CFED6FE06" box="[186,333,422,446]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 43980</specimenCode>
,
<collectingCountry id="C4FA7634FFC4FFAF5F58FE6DFE24FE06" box="[344,447,422,446]" name="Namibia" pageId="15" pageNumber="2340">Namibia</collectingCountry>
:
<location id="B932607FFFC4FFAF5FCAFE6DFDF2FE06" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5FCAFE6DFDF2FE06" box="[458,617,422,446]" country="Namibia" latitude="-17.406666" longLatPrecision="21" longitude="14.216944" municipality="Ruacana Falls" name="Cunene River" pageId="15" pageNumber="2340">Cunene River</location>
:
<collectingMunicipality id="5C36ACDEFFC4FFAF5C74FE6CFC8DFE06" box="[628,790,422,446]" pageId="15" pageNumber="2340">Ruacana Falls</collectingMunicipality>
,
<geoCoordinate id="D9D95063FFC4FFAF5D21FE6DFC3CFE06" box="[801,935,419,446]" degrees="17" direction="south" minutes="24" orientation="latitude" pageId="15" pageNumber="2340" precision="15" seconds="24" value="-17.406666">
17
<superScript id="4B989BECFFC4FFAF5D3CFE68FCDEFE0C" attach="none" box="[828,837,419,436]" fontSize="7" pageId="15" pageNumber="2340">◦</superScript>
24’24” S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC4FFAF5DB3FE6DFBD1FE06" box="[947,1098,419,446]" degrees="014" direction="east" minutes="13" orientation="longitude" pageId="15" pageNumber="2340" precision="15" seconds="01" value="14.216944">
014
<superScript id="4B989BECFFC4FFAF5DDBFE68FC7FFE0C" attach="none" box="[987,996,419,436]" fontSize="7" pageId="15" pageNumber="2340">◦</superScript>
13’01” E
</geoCoordinate>
, coll.
<collectorName id="11185372FFC4FFAF5EC8FE0CFEACFE66" box="[200,311,454,478]" pageId="15" pageNumber="2340">E. Swartz</collectorName>
and
<collectorName id="11185372FFC4FFAF5F74FE0CFE75FE66" box="[372,494,455,478]" pageId="15" pageNumber="2340">B. Kramer</collectorName>
,
<date id="C8531064FFC4FFAF5FFCFE0DFD2AFE66" box="[508,689,454,478]" pageId="15" pageNumber="2340" value="2006-08-19">
<collectingDate id="D817E98CFFC4FFAF5FFCFE0DFD2AFE66" box="[508,689,454,478]" pageId="15" pageNumber="2340" value="2006-08-19">19 August 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC4FFAF5CBFFE0CFED3FE46" collectingDate="2006-08-21" collectionCode="IPMB" country="Namibia" pageId="15" pageNumber="2340" specimenCode="IPMB 43986, 43988" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5CBFFE0CFCCFFE66" box="[703,852,454,478]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 43986</specimenCode>
,
<specimenCode id="EC4B9EDFFFC4FFAF5D61FE0DFC3CFE66" box="[865,935,454,478]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">43988</specimenCode>
, as before, but
<date id="C8531064FFC4FFAF5E8FFE2DFED3FE46" box="[143,328,486,510]" pageId="15" pageNumber="2340" value="2006-08-21">
<collectingDate id="D817E98CFFC4FFAF5E8FFE2DFED3FE46" box="[143,328,486,510]" pageId="15" pageNumber="2340" value="2006-08-21">21 August 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC4FFAF5F57FE2CFCFBFE46" box="[343,864,486,510]" collectingDate="2006-08-22" collectionCode="IPMB" country="Namibia" pageId="15" pageNumber="2340" specimenCode="IPMB 43990" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5F57FE2CFE75FE46" box="[343,494,486,510]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 43990</specimenCode>
, as before, but
<date id="C8531064FFC4FFAF5CA7FE2DFCFBFE46" box="[679,864,486,510]" pageId="15" pageNumber="2340" value="2006-08-22">
<collectingDate id="D817E98CFFC4FFAF5CA7FE2DFCFBFE46" box="[679,864,486,510]" pageId="15" pageNumber="2340" value="2006-08-22">22 August 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC4FFAF5D6FFE2DFC96FDA6" accessionNumber="KC202227, KC202230, KC202238, KC202258" collectionCode="KC" country="Namibia" location="GenBank" pageId="15" pageNumber="2340" specimenCount="1">
<location id="B932607FFFC4FFAF5D6FFE2DFC40FE46" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5D6FFE2DFC40FE46" box="[879,987,486,510]" country="Namibia" name="GenBank" pageId="15" pageNumber="2340">GenBank</location>
accession numbers: (
<accessionNumber id="A3BEAB47FFC4FFAF5F05FDCDFEE5FDA6" box="[261,382,518,542]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC202227" pageId="15" pageNumber="2340">
<collectionCode id="DAFCAE61FFC4FFAF5F05FDCDFEB0FDA6" box="[261,299,518,542]" pageId="15" pageNumber="2340">KC</collectionCode>
202227
</accessionNumber>
-
<accessionNumber id="A3BEAB47FFC4FFAF5F86FDCDFD9BFDA6" box="[390,512,518,542]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC202230" pageId="15" pageNumber="2340">KC202230</accessionNumber>
;
<accessionNumber id="A3BEAB47FFC4FFAF5C0BFDCDFD1EFDA6" box="[523,645,518,542]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC202238" pageId="15" pageNumber="2340">KC202238</accessionNumber>
-
<accessionNumber id="A3BEAB47FFC4FFAF5C8CFDCDFC9DFDA6" box="[652,774,518,542]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC202258" pageId="15" pageNumber="2340">
<collectionCode id="DAFCAE61FFC4FFAF5C8CFDCDFD29FDA6" box="[652,690,518,542]" pageId="15" pageNumber="2340">KC</collectionCode>
202258
</accessionNumber>
)
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5E8FFDAFFE85FDC3" blockId="15.[143,1103,611,706]" box="[143,286,611,635]" pageId="15" pageNumber="2340">
<heading id="E71A81C8FFC4FFAF5E8FFDAFFE85FDC3" box="[143,286,611,635]" fontSize="10" level="3" pageId="15" pageNumber="2340" reason="5">
<emphasis id="8E99EAB6FFC4FFAF5E8FFDAFFE85FDC3" box="[143,286,611,635]" italics="true" pageId="15" pageNumber="2340">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5E8FFD41FF6EFD7A" blockId="15.[143,1103,611,706]" pageId="15" pageNumber="2340">
<collectingRegion id="7E29F846FFC4FFAF5E8FFD41FF7EFD1A" box="[143,229,650,674]" country="Angola" name="Cunene" pageId="15" pageNumber="2340">Cunene</collectingRegion>
River just above the Epupa Falls (Angolan/Namibian border, locality no. 9 on
<figureCitation id="24D62A21FFC4FFAF5E8FFD61FF76FD7A" box="[143,237,682,706]" captionStart="Figure 1" captionStartId="4.[211,278,932,954]" captionTargetBox="[213,1170,167,899]" captionTargetId="figure-5@4.[213,1170,167,899]" captionTargetPageId="4" captionText="Figure 1. Map of sampling locations in southern Africa. (A) Type locality for Marcusenius macrolepidotus (Peters, 1852), Tete on the Lower Zambezi River; (B) origin of M. macrolepidotus, SAIAB 60847, from Marromeu, Lower Zambezi delta (Figure 4A); (C) origin of M. macrolepidotus from the Buzi System, same location as SAIAB 67369, presently alive in aquarium; (D) likely type region for Marcusenius angolensis (Boulenger, 1905), the Lower Quanza; (E) Marcusenius altisambesi Kramer et al., 2007, type locality on the Upper Zambezi River; (F) M. altisambesi from the Kwando River; (G) M. altisambesi from the Okavango delta; (H, I) Marcusenius multisquamatus sp. nov., stretching from below the Ruacana Falls to above the Epupa Falls (I), the type locality; (J) specimens from the Cunene River Mouth." figureDoi="http://doi.org/10.5281/zenodo.5197592" httpUri="https://zenodo.org/record/5197592/files/figure.png" pageId="15" pageNumber="2340">Figure 1</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFC4FFAF5E8FFCCBFC89FBA6" pageId="15" pageNumber="2340" type="diagnosis">
<paragraph id="BC5236A4FFC4FFAF5E8FFCCBFF60FCAF" blockId="15.[143,1104,768,1054]" box="[143,251,768,791]" pageId="15" pageNumber="2340">
<heading id="E71A81C8FFC4FFAF5E8FFCCBFF60FCAF" box="[143,251,768,791]" fontSize="10" level="3" pageId="15" pageNumber="2340" reason="5">
<emphasis id="8E99EAB6FFC4FFAF5E8FFCCBFF60FCAF" box="[143,251,768,791]" italics="true" pageId="15" pageNumber="2340">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5E8FFCECFC89FBA6" blockId="15.[143,1104,768,1054]" pageId="15" pageNumber="2340">Body moderately long, prominent mobile and forward-extending mental lobe on lower jaw, median fins set well back with dorsal fin shorter than and originating behind anal fin, depth of caudal peduncle 38% (34–43%) of its length, 24 (23–25) dorsal fin rays, 30 (28–31) anal fin rays, 59 (56–64) scales in lateral series, 13 (12–16) scales around caudal peduncle, HL (head length) 20% (19–21%) of SL, BD (body depth) 29% (27–32%) of SL, LD (dorsal fin length) 19.4% (18.1–22.3%) of SL, LSo (length of snout) 48% (45–50%) HL, LA (anal fin length) 24.2% (22.7–25.3%) of SL, CPL (length of caudal peduncle) 18.3% (16.3–19.9%) of SL. (See also Remarks.)</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFC4FFB05E8FFB90FD72FA47" lastPageId="16" lastPageNumber="2341" pageId="15" pageNumber="2340" type="description">
<paragraph id="BC5236A4FFC4FFAF5E8FFB90FE95FBCA" blockId="15.[143,1104,1115,1689]" box="[143,270,1115,1138]" pageId="15" pageNumber="2340">
<heading id="E71A81C8FFC4FFAF5E8FFB90FE95FBCA" box="[143,270,1115,1138]" fontSize="10" level="3" pageId="15" pageNumber="2340" reason="5">
<emphasis id="8E99EAB6FFC4FFAF5E8FFB90FE95FBCA" box="[143,270,1115,1138]" italics="true" pageId="15" pageNumber="2340">Description</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFC4FFB05E8FFB49FD6FFA32" blockId="15.[143,1104,1115,1689]" lastBlockId="16.[211,1172,1362,1418]" lastPageId="16" lastPageNumber="2341" pageId="15" pageNumber="2340">
Head with terminal mouth well in front of eye, mental lobe on lower jaw protruding beyond upper jaw. Head and body dorsolaterally compressed. Dorsal fin situated about two-thirds of standard length from snout, obliquely oriented, anteriorly higher and posteriorly lower, distal margin sometimes only slightly crescent-shaped with anterior two or three rays longer than posterior rays, number of rays 23 (
<emphasis id="8E99EAB6FFC4FFAF5D88FAC8FC0EFAA2" box="[904,917,1283,1306]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 5), 24 (
<emphasis id="8E99EAB6FFC4FFAF5A04FAC8FB8AFAA2" box="[1028,1041,1283,1306]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 4), 25 (
<emphasis id="8E99EAB6FFC4FFAF5EBAFAE8FF5CFA82" box="[186,199,1315,1338]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 6); anal fin opposite dorsal fin with distinctly more anterior origin, obliquely oriented, anteriorly lower and posteriorly higher, anterior rays longer than posterior ones, especially in males where they also appear stronger and often darkened, distal margin crescent-shaped (in males only posterior to rounded, elongated anterior part of fin), number of rays 28 (
<emphasis id="8E99EAB6FFC4FFAF5FB9FA69FE5DFA01" box="[441,454,1442,1465]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 1), 29 (
<emphasis id="8E99EAB6FFC4FFAF5C34FA69FDDAFA01" box="[564,577,1442,1465]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 4), 30 (
<emphasis id="8E99EAB6FFC4FFAF5CAFFA69FD27FA01" box="[687,700,1442,1465]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 7), 31 (
<emphasis id="8E99EAB6FFC4FFAF5D2BFA69FCA3FA01" box="[811,824,1442,1465]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 3). Scales cycloid with reticulate striae, scales extending anteriorly to operculum and pectoral fins (beyond pelvic fins). Scales on caudal peduncle circumference, 12 (
<emphasis id="8E99EAB6FFC4FFAF5D0FFA29FC87FA41" box="[783,796,1506,1529]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 5), 13 (
<emphasis id="8E99EAB6FFC4FFAF5D8AFA29FC0CFA41" box="[906,919,1506,1529]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 4), 14 (
<emphasis id="8E99EAB6FFC4FFAF5A04FA29FB8AFA41" box="[1028,1041,1506,1529]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 5), 16 (
<emphasis id="8E99EAB6FFC4FFAF5EB9F9C9FF5DF9A1" box="[185,198,1538,1561]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 1) Caudal peduncle relatively deep, subcylindrical entire length, usually 18.3% (16.3–19.9%) in SL (
<tableCitation id="F16F031FFFC4FFAF5F73F9EAFE47F981" box="[371,476,1569,1593]" captionStart="Table 1" captionStartId="21.[58,113,594,616]" captionText="Table 1. Comparison of anatomical characters in three Marcusenius species from different southern African origins (Cunene MS = multisquamatus sp. nov., L Zambezi ML = macrolepidotus, Okavango AS and U Zambezi AS = altisambesi). Multivariate analysis of variance (MANOVA). p values in the body of the table not shown when&gt;0.05." pageId="15" pageNumber="2340">Table A1</tableCitation>
, in Appendix A). Electric organ discharge biphasic with weak pre-potential (
<figureCitation id="24D62A21FFC4FFAF5F7BF98AFE45F9E1" box="[379,478,1601,1625]" captionStart="Figure 5" captionStartId="16.[211,278,1076,1098]" captionTargetBox="[213,1170,168,1043]" captionTargetId="figure-5@16.[213,1170,168,1043]" captionTargetPageId="16" captionText="Figure 5. Electric organ discharges (EODs) recorded from fish from various origins. (A) Five specimens of Marcusenius macrolepidotus originating from the Buzi River. (B) Marcusenius altisambesi originating from the the Upper Zambezi System. Note P and N phases of exaggerated duration in male individual. (C) M. altisambesi originating from the Okavango delta. (D) Marcusenius multisquamatus sp. nov. originating from the Cunene River. M, male, F, female, J, juvenile individuals below size where sexual maturity is possible. Size given is SL. Note that in Buzi individuals an initial miniature potential is lacking which is present in the other forms of bulldog fish shown here (see EODs with clipped peaks, amplified × 3)." figureDoi="http://doi.org/10.5281/zenodo.5197600" httpUri="https://zenodo.org/record/5197600/files/figure.png" pageId="15" pageNumber="2340">Figure 5</figureCitation>
). Males approaching sexual maturity develop a kink in the base of the anal fin (e.g.
<figureCitation id="24D62A21FFC4FFAF5C08F9ABFDE6F9C0" box="[520,637,1632,1656]" captionStart="Figure 4" captionStartId="11.[143,210,1506,1528]" captionTargetBox="[168,1081,161,1473]" captionTargetId="figure-6@11.[168,1081,161,1473]" captionTargetPageId="11" captionText="Figure 4. Photographs of bulldog fish from various origins. (A) Marcusenius macrolepidotus (Peters, 1852), SAIAB 60847, coll. R. Bills 1 August 1999, Lower Zambezi; (B) Marcusenius angolensis (Boulenger, 1905), BMNH 1905.5.29.64 (holotype); (C) Marcusenius altisambesi, coll. F. H. van der Bank and B. Kramer, 21 August 1999, Upper Zambezi, Kalimbeza, live specimen of 16.5 cm SL photographed 3 July 2003 in aquarium; (D) M. altisambesi, coll. F.H. van der Bank and B. Kramer, 11/12 August 2004, Okavango River, live fish of SL 13 cm photographed 20 April 2006 in aquarium; (E) Marcusenius multisquamatus sp. nov., coll. B. Kramer and Ernst Swartz, coll. and photographed on 19 August 2006, Cunene River, below Ruacana Falls, specimen RUAC01, SL 15.4 cm." figureDoi="http://doi.org/10.5281/zenodo.5197598" httpUri="https://zenodo.org/record/5197598/files/figure.png" pageId="15" pageNumber="2340">Figure 4C</figureCitation>
) that is absent in juveniles and females where the anal fin base is straight. Colour in life: brownish grey with many distinct dark-brown blotches, except on head and belly, purple hue depending on the angle of light incidence, paired fins light and transparent.
</paragraph>
<caption id="E892662CFFDBFFB05ED3FBFFFC02FAAE" ID-DOI="http://doi.org/10.5281/zenodo.5197600" ID-Zenodo-Dep="5197600" httpUri="https://zenodo.org/record/5197600/files/figure.png" pageId="16" pageNumber="2341" startId="16.[211,278,1076,1098]" targetBox="[213,1170,168,1043]" targetPageId="16">
<paragraph id="BC5236A4FFDBFFB05ED3FBFFFC02FAAE" blockId="16.[211,1172,1076,1302]" pageId="16" pageNumber="2341">
Figure 5. Electric organ discharges (EODs) recorded from fish from various origins. (A) Five specimens of
<taxonomicName id="7BED4D27FFDBFFB05F64FB99FD19FBDF" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[356,642,1105,1127]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="16" pageNumber="2341" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFDBFFB05F64FB99FD19FBDF" box="[356,642,1105,1127]" italics="true" pageId="16" pageNumber="2341">Marcusenius macrolepidotus</emphasis>
</taxonomicName>
originating from the Buzi River. (B)
<taxonomicName id="7BED4D27FFDBFFB05A16FB99FED9FB3C" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="16" pageNumber="2341" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFDBFFB05A16FB99FED9FB3C" italics="true" pageId="16" pageNumber="2341">Marcusenius altisambesi</emphasis>
</taxonomicName>
originating from the the Upper Zambezi System. Note
<emphasis id="8E99EAB6FFDBFFB05D7AFBA4FC11FB3C" box="[890,906,1135,1156]" italics="true" pageId="16" pageNumber="2341">P</emphasis>
and
<emphasis id="8E99EAB6FFDBFFB05DBEFBA4FC4BFB3C" box="[958,976,1135,1156]" italics="true" pageId="16" pageNumber="2341">N</emphasis>
phases of exaggerated duration in male individual. (C)
<taxonomicName id="7BED4D27FFDBFFB05C52FB47FD7DFB1A" box="[594,742,1164,1186]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="16" pageNumber="2341" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFDBFFB05C52FB47FD7DFB1A" box="[594,742,1164,1186]" italics="true" pageId="16" pageNumber="2341">M. altisambesi</emphasis>
</taxonomicName>
originating from the Okavango delta. (D)
<taxonomicName id="7BED4D27FFDBFFB05ED3FB61FE68FB07" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[211,499,1193,1215]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="16" pageNumber="2341" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDBFFB05ED3FB61FE68FB07" box="[211,499,1193,1215]" italics="true" pageId="16" pageNumber="2341">Marcusenius multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDBFFB05FFAFB62FDDCFB06" box="[506,583,1193,1214]" pageId="16" pageNumber="2341" rank="species">sp. nov.</taxonomicNameLabel>
originating from the Cunene River. M, male, F, female, J, juvenile individuals below size where sexual maturity is possible. Size given is SL. Note that in Buzi individuals an initial miniature potential is lacking which is present in the other forms of bulldog fish shown here (see EODs with clipped peaks, amplified × 3).
</paragraph>
</caption>
<paragraph id="BC5236A4FFDBFFB05ED3FA74FE5DFA6F" blockId="16.[211,745,1471,1535]" box="[211,454,1471,1495]" pageId="16" pageNumber="2341">
<heading id="E71A81C8FFDBFFB05ED3FA74FE5DFA6F" box="[211,454,1471,1495]" fontSize="10" level="3" pageId="16" pageNumber="2341" reason="5">
<emphasis id="8E99EAB6FFDBFFB05ED3FA74FE5DFA6F" box="[211,454,1471,1495]" italics="true" pageId="16" pageNumber="2341">Colour in preservation</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDBFFB05ED3FA2CFD72FA47" blockId="16.[211,745,1471,1535]" box="[211,745,1511,1535]" pageId="16" pageNumber="2341">Medium brown, with darker, irregular blotches.</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFDBFFB15ED3F9FFFCC4FD86" lastPageId="17" lastPageNumber="2342" pageId="16" pageNumber="2341" type="biology_ecology">
<paragraph id="BC5236A4FFDBFFB05ED3F9FFFEB7F9F4" blockId="16.[211,1172,1588,1716]" box="[211,300,1588,1612]" pageId="16" pageNumber="2341">
<heading id="E71A81C8FFDBFFB05ED3F9FFFEB7F9F4" box="[211,300,1588,1612]" fontSize="10" level="3" pageId="16" pageNumber="2341" reason="5">
<emphasis id="8E99EAB6FFDBFFB05ED3F9FFFEB7F9F4" box="[211,300,1588,1612]" italics="true" pageId="16" pageNumber="2341">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDBFFB15ED3F997FCC4FD86" blockId="16.[211,1172,1588,1716]" lastBlockId="17.[143,1104,167,574]" lastPageId="17" lastPageNumber="2342" pageId="16" pageNumber="2341">
The
<collectingRegion id="7E29F846FFDBFFB05F06F997FEC7F9CC" box="[262,348,1628,1652]" country="Angola" name="Cunene" pageId="16" pageNumber="2341">Cunene</collectingRegion>
is a major, perennial and independent river that arises from the Angolan central highlands of
<collectingRegion id="7E29F846FFDBFFB05FC0F9B0FE7EF92B" box="[448,485,1659,1683]" country="Angola" name="Bie" pageId="16" pageNumber="2341">Bié</collectingRegion>
and flows southward towards the Namibian border, shortly before it turns west and breaches the coastal mountain ranges (Zebra and Baynes Mountains) to drain into the Atlantic. In the section between Ruacana Falls and Epupa Falls, water level was regulated by a hydroelectric company (NamPower) at Ruacana Falls. The Ruacana Falls were bare rock and completely dry, apparently because the water dammed above Ruacana Falls (Calueque Dam) was all fed into the hydroelectric power turbines. When the water level below the dam was kept high, fishing with gill nets and other methods generally proved unproductive. The Epupa Falls consist of a main fall with many lesser falls beside this over a wide front, and mormyrid EODs were demonstrated with an electro-acoustic, custom-built “fish detector” also below the Falls although the fish were not caught. Although we were warned of a high incidence of crocodiles we saw only a few, and no hippopotami. River borders were covered mainly by dense semi-aquatic shrubs at Epupa Falls, and dense reed beds also with shrubs at Ruacana Falls. Palm trees (Makalani palms) were common at Epupa, much less so at Ruacana where dicotyledonous trees dominated.
</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFDAFFB15E8FFDB8FC08FCDF" pageId="17" pageNumber="2342" type="distribution">
<paragraph id="BC5236A4FFDAFFB15E8FFDB8FE8FFD33" blockId="17.[143,1103,627,723]" box="[143,276,627,651]" pageId="17" pageNumber="2342">
<heading id="E71A81C8FFDAFFB15E8FFDB8FE8FFD33" box="[143,276,627,651]" fontSize="10" level="3" pageId="17" pageNumber="2342" reason="5">
<emphasis id="8E99EAB6FFDAFFB15E8FFDB8FE8FFD33" box="[143,276,627,651]" italics="true" pageId="17" pageNumber="2342">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDAFFB15E8FFD51FB9BFD6A" blockId="17.[143,1103,627,723]" pageId="17" pageNumber="2342">
<materialsCitation id="0C853CF9FFDAFFB15E8FFD51FC67FD6A" location="This river" municipality="Ruacana Falls" pageId="17" pageNumber="2342" specimenCount="1" stateProvince="Cunene">
At present known only from the lower
<collectingRegion id="7E29F846FFDAFFB15C4BFD51FD3AFD0A" box="[587,673,666,690]" country="Angola" name="Cunene" pageId="17" pageNumber="2342">Cunene</collectingRegion>
River, from just below
<collectingMunicipality id="5C36ACDEFFDAFFB15DAEFD50FBD4FD0A" box="[942,1103,666,690]" pageId="17" pageNumber="2342">Ruacana Falls</collectingMunicipality>
to the
<collectingRegion id="7E29F846FFDAFFB15ED7FD71FEB6FD6A" box="[215,301,698,722]" country="Angola" name="Cunene" pageId="17" pageNumber="2342">Cunene</collectingRegion>
mouth.
<location id="B932607FFFDAFFB15F8DFD71FE62FD6A" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFDAFFB15F8DFD71FE62FD6A" box="[397,505,698,722]" municipality="Ruacana Falls" name="This river" pageId="17" pageNumber="2342" stateProvince="Cunene">This river</location>
section forms the Angolan/Namibian border
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFDAFFB15E8FFCCCFEB8FCA7" blockId="17.[143,1103,775,871]" box="[143,291,775,799]" pageId="17" pageNumber="2342">
<heading id="E71A81C8FFDAFFB15E8FFCCCFEB8FCA7" box="[143,291,775,799]" fontSize="10" level="3" pageId="17" pageNumber="2342" reason="5">
<emphasis id="8E99EAB6FFDAFFB15E8FFCCCFEB8FCA7" box="[143,291,775,799]" italics="true" pageId="17" pageNumber="2342">Relationships</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDAFFB15E8FFCE4FC08FCDF" blockId="17.[143,1103,775,871]" pageId="17" pageNumber="2342">
Closest relationships are assumed with
<taxonomicName id="7BED4D27FFDAFFB15C40FCFBFD79FCFF" box="[576,738,815,839]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFDAFFB15C40FCFBFD79FCFF" box="[576,738,815,839]" italics="true" pageId="17" pageNumber="2342">M. altisambesi</emphasis>
</taxonomicName>
to the east of
<taxonomicName id="7BED4D27FFDAFFB15D7BFCFBFBD4FCFF" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[891,1103,815,839]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15D7BFCFBFBD4FCFF" box="[891,1103,815,839]" italics="true" pageId="17" pageNumber="2342">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDAFFB15E8FFC84FF7FFCDE" box="[143,228,847,870]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
on the basis of morphological similarity, EODs and genetics.
</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFDAFFB15E8FFC50FC18FB83" pageId="17" pageNumber="2342" type="etymology">
<paragraph id="BC5236A4FFDAFFB15E8FFC50FE9CFC0B" blockId="17.[143,1104,923,1083]" box="[143,263,923,947]" pageId="17" pageNumber="2342">
<heading id="E71A81C8FFDAFFB15E8FFC50FE9CFC0B" box="[143,263,923,947]" fontSize="10" level="3" pageId="17" pageNumber="2342" reason="5">
<emphasis id="8E99EAB6FFDAFFB15E8FFC50FE9CFC0B" box="[143,263,923,947]" italics="true" pageId="17" pageNumber="2342">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDAFFB15E8FFC0FFC18FB83" blockId="17.[143,1104,923,1083]" pageId="17" pageNumber="2342">
<taxonomicName id="7BED4D27FFDAFFB15E8FFC0FFE54FC63" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[143,463,963,987]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15E8FFC0FFE54FC63" box="[143,463,963,987]" italics="true" pageId="17" pageNumber="2342">Marcusenius multisquamatus</emphasis>
</taxonomicName>
<materialsCitation id="0C853CF9FFDAFFB15FD6FC0FFCE4FB83" collectionCode="BMNH" location="Mossamedes" municipality="southern African" pageId="17" pageNumber="2342" specimenCount="1" stateProvince="Cunene">
<taxonomicNameLabel id="95AA57CDFFDAFFB15FD6FC0FFDB0FC63" box="[470,555,964,987]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
refers to the highest number of lateral line scales among the different forms of
<collectingMunicipality id="5C36ACDEFFDAFFB15FFAFC28FD5AFC43" box="[506,705,995,1019]" pageId="17" pageNumber="2342">southern African</collectingMunicipality>
bulldog fish (excluding the three
<location id="B932607FFFDAFFB15E8FFBC8FEBBFBA3" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFDAFFB15E8FFBC8FEBBFBA3" box="[143,288,1027,1051]" municipality="southern African" name="Mossamedes" pageId="17" pageNumber="2342" stateProvince="Cunene">Mossamedes</location>
/
<collectingRegion id="7E29F846FFDAFFB15F2CFBC8FE19FBA3" box="[300,386,1027,1051]" country="Angola" name="Cunene" pageId="17" pageNumber="2342">Cunene</collectingRegion>
specimens (
<collectionCode id="DAFCAE61FFDAFFB15C12FBC8FDF0FBA3" box="[530,619,1027,1051]" pageId="17" pageNumber="2342">BMNH</collectionCode>
1907.6.29.231–233) from any location on the
<collectingRegion id="7E29F846FFDAFFB15EB9FBE8FE94FB83" box="[185,271,1059,1083]" country="Angola" name="Cunene" pageId="17" pageNumber="2342">Cunene</collectingRegion>
up to
<quantity id="7B159B41FFDAFFB15F58FBE8FE37FB83" box="[344,428,1059,1083]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="2342" unit="km" value="300.0">300 km</quantity>
north of the Angolan/Namibian border)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFDAFFB25E8FFBA4FB0BFCA6" lastPageId="18" lastPageNumber="2343" pageId="17" pageNumber="2342" type="discussion">
<paragraph id="BC5236A4FFDAFFB15E8FFBA4FF6AFB3F" blockId="17.[143,1104,1135,1709]" box="[143,241,1135,1159]" pageId="17" pageNumber="2342">
<heading id="E71A81C8FFDAFFB15E8FFBA4FF6AFB3F" box="[143,241,1135,1159]" fontSize="10" level="3" pageId="17" pageNumber="2342" reason="5">
<emphasis id="8E99EAB6FFDAFFB15E8FFBA4FF6AFB3F" box="[143,241,1135,1159]" italics="true" pageId="17" pageNumber="2342">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDAFFB15E8FFB5CFDEFFAD6" blockId="17.[143,1104,1135,1709]" pageId="17" pageNumber="2342">
Compared with the
<taxonomicName id="7BED4D27FFDAFFB15F6DFB53FD9EFB17" authorityName="Boulenger" authorityYear="1905" box="[365,517,1175,1199]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="8E99EAB6FFDAFFB15F6DFB53FD9EFB17" box="[365,517,1175,1199]" italics="true" pageId="17" pageNumber="2342">M. angolensis</emphasis>
</taxonomicName>
<typeStatus id="63568806FFDAFFB15C0AFB5CFDF4FB17" box="[522,623,1175,1199]" pageId="17" pageNumber="2342" type="holotype">holotype</typeStatus>
,
<taxonomicName id="7BED4D27FFDAFFB15C7AFB53FCD5FB17" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[634,846,1175,1199]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15C7AFB53FCD5FB17" box="[634,846,1175,1199]" italics="true" pageId="17" pageNumber="2342">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDAFFB15D54FB53FC33FB17" box="[852,936,1176,1199]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
specimens had lower counts in nD, no. of dorsal fin rays (maximum,
<quantity id="7B159B41FFDAFFB15D0CFB7CFCDCFB77" box="[780,839,1207,1231]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" pageId="17" pageNumber="2342" unit="in" value="25.0">25 in</quantity>
the latter versus
<quantity id="7B159B41FFDAFFB15A15FB7CFBD4FB77" box="[1045,1103,1207,1231]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="17" pageNumber="2342" unit="in" value="26.0">26 in</quantity>
the former, that is, no overlap) and nA, number of anal fin rays (maximum, 31 versus 33), shorter LA, anal fin length (maximum, 0.253 versus 0.258 of SL) and PDL, predorsal length (maximum, 0.665 versus 0.674 of SL), smaller ratio HL/Na, head length/separation of nares (maximum, 15.52 versus 15.96), but a greater BD, body depth (minimum 0.271 versus 0.266 of SL).
</paragraph>
<paragraph id="BC5236A4FFDAFFB25EBAFABDFD63FE27" blockId="17.[143,1104,1135,1709]" lastBlockId="18.[211,1172,167,798]" lastPageId="18" lastPageNumber="2343" pageId="17" pageNumber="2342">
When compared with the other
<taxonomicName id="7BED4D27FFDAFFB15C4CFABCFCAEFA36" box="[588,821,1398,1422]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="8E99EAB6FFDAFFB15C4CFABCFD4CFA36" box="[588,727,1399,1422]" italics="true" pageId="17" pageNumber="2342">Marcusenius</emphasis>
species
</taxonomicName>
within the
<collectingRegion id="7E29F846FFDAFFB15DCFFABDFBDAFA36" box="[975,1089,1398,1422]" country="Namibia" name="Kavango West" pageId="17" pageNumber="2342">Okavango</collectingRegion>
– Kwando–
<collectingRegion id="7E29F846FFDAFFB15EFBFA5DFEC6FA16" box="[251,349,1430,1454]" country="Namibia" name="Zambezi" pageId="17" pageNumber="2342">Zambezi</collectingRegion>
System,
<taxonomicName id="7BED4D27FFDAFFB15FC8FA5CFD04FA16" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[456,671,1430,1454]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15FC8FA5CFD04FA16" box="[456,671,1430,1454]" italics="true" pageId="17" pageNumber="2342">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDAFFB15CA9FA5CFC9BFA16" box="[681,768,1431,1454]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by a specific morphology and EOD in multivariate analysis, specific bands in genomic ISSR fingerprinting, and as a monophyletic taxon in mitochondrial DNA (mtDNA) cytochrome
<emphasis id="8E99EAB6FFDAFFB15A42FA1DFBD4FA56" box="[1090,1103,1494,1518]" italics="true" pageId="17" pageNumber="2342">b</emphasis>
analysis. To identify a specimen in hand it is best to rely on several characters in combination to exclude mistakes due to outliers. The 90th percentile of the distribution of HL (measured as HL/SL, head length to standard length) of
<taxonomicName id="7BED4D27FFDAFFB15D77F9FCFBCBF9F6" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[887,1104,1590,1614]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15D77F9FCFBCBF9F6" box="[887,1104,1590,1614]" italics="true" pageId="17" pageNumber="2342">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDAFFB15E8FF99DFF7FF9D5" box="[143,228,1622,1645]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
specimens, is shorter than the 10th percentile for
<taxonomicName id="7BED4D27FFDAFFB15D13F99CFC79F9D6" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[787,994,1622,1646]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFDAFFB15D13F99CFC79F9D6" box="[787,994,1622,1646]" italics="true" pageId="17" pageNumber="2342">M. macrolepidotus</emphasis>
</taxonomicName>
(together with
<taxonomicName id="7BED4D27FFDAFFB15ECBF9BDFEFDF936" authorityName="Boulenger" authorityYear="1905" box="[203,358,1654,1678]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="8E99EAB6FFDAFFB15ECBF9BDFEFDF936" box="[203,358,1654,1678]" italics="true" pageId="17" pageNumber="2342">M. angolensis</emphasis>
</taxonomicName>
, shortest HL of all). The BD/SL ratio (body depth to standard length) of
<taxonomicName id="7BED4D27FFDAFFB15F0CF95DFE7DF915" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[268,486,1685,1709]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15F0CF95DFE7DF915" box="[268,486,1685,1709]" italics="true" pageId="17" pageNumber="2342">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDAFFB15FF2F95DFDD0F915" box="[498,587,1686,1709]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
overlaps with that of
<taxonomicName id="7BED4D27FFDAFFB15D56F95DFBB3F915" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[854,1064,1685,1709]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFDAFFB15D56F95DFBB3F915" box="[854,1064,1685,1709]" italics="true" pageId="17" pageNumber="2342">M. macrolepidotus</emphasis>
</taxonomicName>
by less than one quartile; the same holds true for the distributions of nA (no. of anal fin rays), nD (no. of dorsal fin rays), LD/SL (ratio of dorsal fin length to standard length) in which the means or medians are greater for
<taxonomicName id="7BED4D27FFD9FFB25D52FF23FBB1FF47" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[850,1066,231,255]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25D52FF23FBB1FF47" box="[850,1066,231,255]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25A35FF23FB14FF47" box="[1077,1167,232,255]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
, and SPc, number of scales around caudal peduncle in which the median for
<taxonomicName id="7BED4D27FFD9FFB25A38FEC3FECCFE87" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25A38FEC3FECCFE87" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25F5FFEE3FE2FFE87" box="[351,436,296,319]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
is smaller. The EOD of
<taxonomicName id="7BED4D27FFD9FFB25CC6FEE3FC07FE87" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[710,924,295,319]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25CC6FEE3FC07FE87" box="[710,924,295,319]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25DA4FEE3FC62FE87" box="[932,1017,296,319]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
has a leading head-negativity of miniature amplitude that is usually not present in the
<taxonomicName id="7BED4D27FFD9FFB25DF8FE83FEC1FEC7" authority="EOD." authorityName="EOD." class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25DF8FE83FE95FEC7" italics="true" pageId="18" pageNumber="2343">M. macrolepidotus</emphasis>
EOD.
</taxonomicName>
<taxonomicName id="7BED4D27FFD9FFB25F63FEA3FD3EFEC7" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[355,677,359,383]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25F63FEA3FD3EFEC7" box="[355,677,359,383]" italics="true" pageId="18" pageNumber="2343">Marcusenius multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25CAFFEA3FC9DFEC7" box="[687,774,360,383]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="7BED4D27FFD9FFB25D43FEA3FB8FFEC7" box="[835,1044,359,383]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25D43FEA3FB8FFEC7" box="[835,1044,359,383]" italics="true" pageId="18" pageNumber="2343">M. macrolepidotus</emphasis>
</taxonomicName>
are clearly differentiated in ISSR bands 2, 6 and 8 (
<tableCitation id="F16F031FFFD9FFB25C97FE4CFD7CFE27" box="[663,743,391,415]" captionStart="Table 4" captionStartId="30.[213,268,169,191]" captionText="Table 4. Inter-simple-sequence-repeat profile of Marcusenius altisambesi, Marcusenius multisquamatus sp. nov. and Marcusenius macrolepidotus (selection of informative bands, numbers 1 to 13); x = band is present; formatting [bold/italic/bold-italic/clean]: highlighting bands characteristic of species." pageId="18" pageNumber="2343">Table 4</tableCitation>
).
</paragraph>
<paragraph id="BC5236A4FFD9FFB25EFEFE6DFB0BFCA6" blockId="18.[211,1172,167,798]" pageId="18" pageNumber="2343">
There is less than a 10% overlap of distributions between the greater LSo/HL (length of snout to head length) of
<taxonomicName id="7BED4D27FFD9FFB25C6AFE0CFCDAFE67" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[618,833,455,479]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25C6AFE0CFCDAFE67" box="[618,833,455,479]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25D4AFE0CFC3AFE66" box="[842,929,455,478]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
specimens compared with that of
<taxonomicName id="7BED4D27FFD9FFB25F63FE2CFD93FE47" box="[355,520,487,511]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFD9FFB25F63FE2CFD93FE47" box="[355,520,487,511]" italics="true" pageId="18" pageNumber="2343">M. altisambesi</emphasis>
</taxonomicName>
from the Upper
<collectingRegion id="7E29F846FFD9FFB25CCDFE2DFCA9FE46" box="[717,818,486,510]" country="Namibia" name="Zambezi" pageId="18" pageNumber="2343">Zambezi</collectingRegion>
, and less than 25% overlap for
<taxonomicName id="7BED4D27FFD9FFB25ED3FDCCFEE2FDA6" box="[211,377,518,542]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFD9FFB25ED3FDCCFEE2FDA6" box="[211,377,518,542]" italics="true" pageId="18" pageNumber="2343">M. altisambesi</emphasis>
</taxonomicName>
from the
<collectingRegion id="7E29F846FFD9FFB25FF0FDCDFDFDFDA6" box="[496,614,518,542]" country="Namibia" name="Kavango West" pageId="18" pageNumber="2343">Okavango</collectingRegion>
. Also, there is less than one quartile overlap for LA/SL (anal fin length to SL), LSc/HL (length of snout to head length) and CL/HL (chin length to head length) of
<taxonomicName id="7BED4D27FFD9FFB25C28FD8CFD51FDE6" box="[552,714,582,606]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFD9FFB25C28FD8CFD51FDE6" box="[552,714,582,606]" italics="true" pageId="18" pageNumber="2343">M. altisambesi</emphasis>
</taxonomicName>
from the Upper
<collectingRegion id="7E29F846FFD9FFB25D84FD8DFC7DFDE6" box="[900,998,582,606]" country="Namibia" name="Zambezi" pageId="18" pageNumber="2343">Zambezi</collectingRegion>
compared with
<taxonomicName id="7BED4D27FFD9FFB25ED3FDACFE30FDC6" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[211,427,614,638]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25ED3FDACFE30FDC6" box="[211,427,614,638]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25FB4FDACFD91FDC6" box="[436,522,615,638]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
and less than one quartile overlap for lower SLS (no. of lateral line scales) of
<taxonomicName id="7BED4D27FFD9FFB25FBDFD4CFDFBFD26" box="[445,608,646,670]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFD9FFB25FBDFD4CFDFBFD26" box="[445,608,646,670]" italics="true" pageId="18" pageNumber="2343">M. altisambesi</emphasis>
</taxonomicName>
from the
<collectingRegion id="7E29F846FFD9FFB25CCEFD4DFCDAFD26" box="[718,833,646,670]" country="Namibia" name="Kavango West" pageId="18" pageNumber="2343">Okavango</collectingRegion>
compared with
<taxonomicName id="7BED4D27FFD9FFB25DF7FD4CFE8EFD06" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25DF7FD4CFE8EFD06" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25F1BFD6DFEEBFD06" box="[283,368,678,702]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
<collectingRegion id="7E29F846FFD9FFB25F76FD6DFE72FD06" box="[374,489,678,702]" country="Namibia" name="Kavango West" pageId="18" pageNumber="2343">Okavango</collectingRegion>
bulldog fish are distinguished by an SPc (no. of scales around caudal peduncle) of exclusively 12 (median 12, same median for Upper
<collectingRegion id="7E29F846FFD9FFB25DF6FD0EFBC3FD65" box="[1014,1112,709,733]" country="Namibia" name="Zambezi" pageId="18" pageNumber="2343">Zambezi</collectingRegion>
bulldog fish) whereas SPc ranges from
<quantity id="7B159B41FFD9FFB25C5EFD2EFD26FD45" box="[606,701,741,765]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="4.064" metricValueMin="3.048" pageId="18" pageNumber="2343" unit="in" value="14.0" valueMax="16.0" valueMin="12.0">12–16 in</quantity>
<taxonomicName id="7BED4D27FFD9FFB25CC5FD2DFC01FD46" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[709,922,742,766]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25CC5FD2DFC01FD46" box="[709,922,742,766]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25DA2FD2DFC6CFD45" box="[930,1015,742,765]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
(median, 13). ISSR band 2 is specific for
<taxonomicName id="7BED4D27FFD9FFB25C02FCCDFD4CFCA6" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[514,727,774,798]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25C02FCCDFD4CFCA6" box="[514,727,774,798]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25CDEFCCDFCA9FCA5" box="[734,818,774,797]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
and band 6 for
<emphasis id="8E99EAB6FFD9FFB25DE5FCCDFB0BFCA6" box="[997,1168,774,798]" italics="true" pageId="18" pageNumber="2343">
<taxonomicName id="7BED4D27FFD9FFB25DE5FCCDFB17FCA6" box="[997,1164,774,798]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="altisambesi">M. altisambesi</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="1B08775F9D60A2DD8962A5DC1F83A19D">East-west differentiation in the Marcusenius macrolepidotus species complex in Southern Africa: the description of a new species for the lower Cunene River, Namibia (Teleostei: Mormyridae)</mods:title>
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<mods:namePart id="D4C4987E301EC942AC66EF6E0DF3E3EE">Kramer, Bernd</mods:namePart>
<mods:affiliation id="F972FA11BB078701D42FF8F16139EC1C">University of Regensburg, Zoological Institute, Universitätsstrasse 31, D- 93040 Regensburg, Germany;</mods:affiliation>
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<mods:namePart id="99A1E14C183C5782D4787D1782F01ABE">Wink, Michael</mods:namePart>
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<mods:title id="E9B0C9ACCFA9FE7DF49079F51ECB87BF">Journal of Natural History</mods:title>
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<mods:date id="C7DCBABB53BFED890EC69F599F96F7EB">2013</mods:date>
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<mods:title id="3C45FACFDEF794C61785261681DACE88">J. Nat. Hist.</mods:title>
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<mods:number id="F6A1A009228903AD26C782732D821BD1">2013-07-18</mods:number>
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<heading id="E71A81C8FFD9FFB25FC8FC9EFC05FCD5" box="[456,926,853,878]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="2343" reason="2">
<taxonomicName id="7BED4D27FFD9FFB25FC8FC9EFC05FCD5" ID-CoL="3Y38T" authority="(Peters, 1852)" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[456,926,853,878]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25FC8FC9EFC9AFCD6" bold="true" box="[456,769,853,878]" italics="true" pageId="18" pageNumber="2343">Marcusenius macrolepidotus</emphasis>
(
<bibRefCitation id="D87C4B55FFD9FFB25D0FFC9EFC0CFCD5" author="Peters WCH" box="[783,919,853,877]" pageId="18" pageNumber="2343" pagination="273 - 276" refId="ref17195" refString="Peters WCH. 1852. Einige neue Saugethiere und Flussfische aus Mossambique. Monatsber Konigl Preuss Akad Wiss Berlin. 1852: 273 - 276." type="journal article" year="1852">Peters, 1852</bibRefCitation>
)
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<paragraph id="BC5236A4FFD9FFB25C71FCBEFD6DFC35" blockId="18.[456,926,853,909]" box="[625,758,885,909]" pageId="18" pageNumber="2343">
(
<figureCitation id="24D62A21FFD9FFB25C7AFCBEFD77FC35" box="[634,748,885,909]" captionStart="Figure 4" captionStartId="11.[143,210,1506,1528]" captionTargetBox="[168,1081,161,1473]" captionTargetId="figure-6@11.[168,1081,161,1473]" captionTargetPageId="11" captionText="Figure 4. Photographs of bulldog fish from various origins. (A) Marcusenius macrolepidotus (Peters, 1852), SAIAB 60847, coll. R. Bills 1 August 1999, Lower Zambezi; (B) Marcusenius angolensis (Boulenger, 1905), BMNH 1905.5.29.64 (holotype); (C) Marcusenius altisambesi, coll. F. H. van der Bank and B. Kramer, 21 August 1999, Upper Zambezi, Kalimbeza, live specimen of 16.5 cm SL photographed 3 July 2003 in aquarium; (D) M. altisambesi, coll. F.H. van der Bank and B. Kramer, 11/12 August 2004, Okavango River, live fish of SL 13 cm photographed 20 April 2006 in aquarium; (E) Marcusenius multisquamatus sp. nov., coll. B. Kramer and Ernst Swartz, coll. and photographed on 19 August 2006, Cunene River, below Ruacana Falls, specimen RUAC01, SL 15.4 cm." figureDoi="http://doi.org/10.5281/zenodo.5197598" httpUri="https://zenodo.org/record/5197598/files/figure.png" pageId="18" pageNumber="2343">Figure 4A</figureCitation>
)
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<paragraph id="BC5236A4FFD9FFB25ED3FC7EFD86FC75" blockId="18.[211,1172,949,1355]" box="[211,541,949,973]" pageId="18" pageNumber="2343">Nominal species in bold face.</paragraph>
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<paragraph id="BC5236A4FFD9FFB25ED3FC1EFD73FC54" blockId="18.[211,1172,949,1355]" box="[211,744,980,1005]" pageId="18" pageNumber="2343">
<taxonomicName id="7BED4D27FFD9FFB25ED3FC1EFD7FFC54" ID-CoL="6RRC3" authority="Peters, 1852 a, p. 275" authorityName="Peters" authorityPageNumber="275" authorityYear="1852" box="[211,740,980,1005]" class="Actinopterygii" family="Mormyridae" genus="Mormyrus" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25ED3FC1EFE6DFC55" bold="true" box="[211,502,980,1005]" italics="true" pageId="18" pageNumber="2343">Mormyrus macrolepidotus</emphasis>
<bibRefCitation id="D87C4B55FFD9FFB25FFDFC1EFD1DFC54" author="Peters WCH" box="[509,646,980,1004]" pageId="18" pageNumber="2343" pagination="273 - 276" refId="ref17195" refString="Peters WCH. 1852. Einige neue Saugethiere und Flussfische aus Mossambique. Monatsber Konigl Preuss Akad Wiss Berlin. 1852: 273 - 276." type="journal article" year="1852">Peters, 1852</bibRefCitation>
a, p. 275
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.
</paragraph>
<paragraph id="BC5236A4FFD9FFB25ED3FC3EFC88FBB4" blockId="18.[211,1172,949,1355]" box="[211,787,1012,1037]" pageId="18" pageNumber="2343">
<treatmentCitationGroup id="9CFD118AFFD9FFB25ED3FC3EFC88FBB4" box="[211,787,1012,1037]" pageId="18" pageNumber="2343">
<treatmentCitation id="3D4C10B5FFD9FFB25ED3FC3EFC94FBB4" author="Marcusen J" box="[211,783,1012,1037]" page="142" pageId="18" pageNumber="2343" year="1864">
<taxonomicName id="7BED4D27FFD9FFB25ED3FC3EFC94FBB4" ID-CoL="6S3CK" authority=": Marcusen 1864, p. 142" authorityName="Marcusen" authorityPageNumber="142" authorityYear="1864" box="[211,783,1012,1037]" class="Actinopterygii" family="Mormyridae" genus="Mormyrops" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25ED3FC3EFD9AFBB5" box="[211,513,1013,1037]" italics="true" pageId="18" pageNumber="2343">Mormyrops macrolepidotus</emphasis>
:
<bibRefCitation id="D87C4B55FFD9FFB25C0FFC3EFD25FBB4" author="Marcusen J" box="[527,702,1012,1036]" pageId="18" pageNumber="2343" pagination="1 - 162" refId="ref17002" refString="Marcusen J. 1864. Die Familie der Mormyren. Eine anatomisch-zoologische Abhandlung. Mem Acad Sci St Petersburg. 7: 1 - 162." type="journal article" year="1864">Marcusen 1864</bibRefCitation>
, p. 142
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.
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<paragraph id="BC5236A4FFD9FFB25ED3FBDEFCB7FB94" blockId="18.[211,1172,949,1355]" box="[211,812,1044,1069]" pageId="18" pageNumber="2343">
<treatmentCitationGroup id="9CFD118AFFD9FFB25ED3FBDEFCB7FB94" box="[211,812,1044,1069]" pageId="18" pageNumber="2343">
<treatmentCitation id="3D4C10B5FFD9FFB25ED3FBDEFCB2FB94" author="Boulenger GA" box="[211,809,1044,1069]" page="804" pageId="18" pageNumber="2343" year="1898">
<taxonomicName id="7BED4D27FFD9FFB25ED3FBDEFCB2FB94" ID-CoL="3GLTR" authority=": Boulenger 1898, p. 804" authorityName=": Boulenger" authorityPageNumber="804" authorityYear="1898" box="[211,809,1044,1069]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25ED3FBDEFD8DFB95" box="[211,534,1045,1069]" italics="true" pageId="18" pageNumber="2343">Gnathonemus macrolepidotus</emphasis>
:
<bibRefCitation id="D87C4B55FFD9FFB25C25FBDFFD4CFB94" author="Boulenger GA" box="[549,727,1044,1068]" pageId="18" pageNumber="2343" pagination="775 - 821" refId="ref15896" refString="Boulenger GA. 1898. A revision of the genera and species of the fishes of the family Mormyridae. Proc Zool Soc Lond. 1898: 775 - 821." type="journal article" year="1898">Boulenger 1898</bibRefCitation>
, p. 804
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.
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</paragraph>
<paragraph id="BC5236A4FFD9FFB25ED3FBFFFB9BFBF4" blockId="18.[211,1172,949,1355]" box="[211,1024,1076,1100]" pageId="18" pageNumber="2343">
<treatmentCitationGroup id="9CFD118AFFD9FFB25ED3FBFFFB9BFBF4" box="[211,1024,1076,1100]" pageId="18" pageNumber="2343">
<treatmentCitation id="3D4C10B5FFD9FFB25ED3FBFFFC67FBF4" author="Poll M &amp; Gosse J-P" box="[211,1020,1076,1100]" page="93" pageId="18" pageNumber="2343" year="1963">
<taxonomicName id="7BED4D27FFD9FFB25ED3FBFFFC67FBF4" ID-CoL="3GLTR" authority="macrolepidotus: Poll and Gosse 1963, p. 93" authorityName="Poll and Gosse" authorityPageNumber="93" authorityYear="1963" box="[211,1020,1076,1100]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="subSpecies" species="macrolepidotus" subSpecies="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25ED3FBFFFD58FBF4" box="[211,707,1076,1100]" italics="true" pageId="18" pageNumber="2343">Gnathonemus macrolepidotus macrolepidotus</emphasis>
:
<bibRefCitation id="D87C4B55FFD9FFB25CD2FBFFFC22FBF4" author="Poll M &amp; Gosse J-P" box="[722,953,1076,1100]" pageId="18" pageNumber="2343" pagination="43 - 111" refId="ref17221" refString="Poll M, Gosse J-P. 1963. Contribution a l'etude systematique de la faune ichthyologique du Congo Central. Annales Musee Royal de l'Afrique Centrale, Sciences Zoologiques. 116: 43 - 111." type="journal article" year="1963">Poll and Gosse 1963</bibRefCitation>
, p. 93
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.
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</paragraph>
<paragraph id="BC5236A4FFD9FFB25ED3FB9EFC88FBD4" blockId="18.[211,1172,949,1355]" box="[211,787,1108,1132]" pageId="18" pageNumber="2343">
<treatmentCitationGroup id="9CFD118AFFD9FFB25ED3FB9EFC88FBD4" box="[211,787,1108,1132]" pageId="18" pageNumber="2343">
<treatmentCitation id="3D4C10B5FFD9FFB25ED3FB9EFC95FBD4" author="Taverne L" box="[211,782,1108,1132]" page="103" pageId="18" pageNumber="2343" year="1971">
<taxonomicName id="7BED4D27FFD9FFB25ED3FB9EFC95FBD4" ID-CoL="3Y38T" authority=": Taverne 1971 a, p. 103" authorityName=": Taverne" authorityPageNumber="103" authorityYear="1971" box="[211,782,1108,1132]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25ED3FB9EFD90FBD4" box="[211,523,1108,1132]" italics="true" pageId="18" pageNumber="2343">Marcusenius macrolepidotus</emphasis>
:
<bibRefCitation id="D87C4B55FFD9FFB25C1AFB9EFD25FBD4" author="Taverne L" box="[538,702,1108,1132]" pageId="18" pageNumber="2343" pagination="99 - 110" refId="ref17631" refString="Taverne L. 1971 a. Note sur la systematique des poissons Mormyriformes. Le probleme des genres Gnathonemus Gill, Marcusenius Gill, Hippopotamyrus Pappenheim, Cyphomyrus Myers et les nouveaux genres Pollimyrus et Brienomyrus. Revue de Zoologie et de Botanique Africaines. 84 (1 - 2): 99 - 110." type="journal article" year="1971">Taverne 1971a</bibRefCitation>
, p. 103
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.
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</paragraph>
<paragraph id="BC5236A4FFD9FFB25ED3FBBFFC7BFB34" blockId="18.[211,1172,949,1355]" box="[211,992,1140,1164]" pageId="18" pageNumber="2343">
<treatmentCitationGroup id="9CFD118AFFD9FFB25ED3FBBFFC7BFB34" box="[211,992,1140,1164]" pageId="18" pageNumber="2343">
<treatmentCitation id="3D4C10B5FFD9FFB25ED3FBBFFCBAFB34" author="Pappenheim P" box="[211,801,1140,1164]" page="354" pageId="18" pageNumber="2343" year="1907">
<taxonomicName id="7BED4D27FFD9FFB25ED3FBBFFCBAFB34" ID-CoL="6KNGJ" authority="Pappenheim 1907, p. 354" authorityName="Pappenheim" authorityPageNumber="354" authorityYear="1907" box="[211,801,1140,1164]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="okavangensis">
<emphasis id="8E99EAB6FFD9FFB25ED3FBBFFD9BFB34" box="[211,512,1140,1164]" italics="true" pageId="18" pageNumber="2343">Gnathonemus okavangensis</emphasis>
<bibRefCitation id="D87C4B55FFD9FFB25C07FBBFFD4AFB34" author="Pappenheim P" box="[519,721,1140,1164]" pageId="18" pageNumber="2343" pagination="339 - 367" refId="ref17154" refString="Pappenheim P. 1907. Zur Systematik und Variationsstatistik der Mormyriden hauptsachlich aus den deutsch-afrikanischen Schutzgebieten. Nach dem Material des Koniglichen Zoologischen Museums zu Berlin. Mitteilungen aus dem Zoologischen Museum in Berlin. 3: 339 - 367." type="journal article" year="1907">Pappenheim 1907</bibRefCitation>
, p. 354
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, nomen dubium.
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</paragraph>
<paragraph id="BC5236A4FFD9FFB25ED3FB58FCCCFB74" blockId="18.[211,1172,949,1355]" pageId="18" pageNumber="2343">
<treatmentCitationGroup id="9CFD118AFFD9FFB25ED3FB58FCCCFB74" pageId="18" pageNumber="2343">
<treatmentCitation id="3D4C10B5FFD9FFB25ED3FB58FCAAFB14" author="Boulenger GA" box="[211,817,1171,1196]" page="163" pageId="18" pageNumber="2343" year="1915">
<taxonomicName id="7BED4D27FFD9FFB25ED3FB58FCAAFB14" ID-CoL="3GLTV" authority="Boulenger, 1915, p. 163" authorityName="Boulenger" authorityPageNumber="163" authorityYear="1915" box="[211,817,1171,1196]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="moeruensis">
<emphasis id="8E99EAB6FFD9FFB25ED3FB58FE6FFB14" bold="true" box="[211,500,1171,1196]" italics="true" pageId="18" pageNumber="2343">Gnathonemus moeruensis</emphasis>
<bibRefCitation id="D87C4B55FFD9FFB25C07FB5FFD52FB14" author="Boulenger GA" box="[519,713,1172,1196]" pageId="18" pageNumber="2343" pagination="162 - 171" refId="ref16011" refString="Boulenger GA. 1915. Diagnoses de poissons nouveaux. II. Mormyrides, Kneriides, Characinides, Cyprinides, Silurides. Revue de Zoologie Africaine. 4 (2): 162 - 171." type="journal article" year="1915">Boulenger, 1915</bibRefCitation>
, p. 163
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;
<treatmentCitation id="3D4C10B5FFD9FFB25D47FB5FFBDAFB14" author="Jackson PBN" box="[839,1089,1172,1196]" page="32" pageId="18" pageNumber="2343" year="1961">
<bibRefCitation id="D87C4B55FFD9FFB25D47FB5FFC7CFB14" author="Jackson PBN" box="[839,999,1172,1196]" pageId="18" pageNumber="2343" refId="ref16419" refString="Jackson PBN. 1961. The fishes of Northern Rhodesia: a check-list of indigenous species. Lusaka: Govt. Printer." type="book" year="1961">Jackson 1961</bibRefCitation>
, p. 32
</treatmentCitation>
(most probably a subspecies of
<taxonomicName id="7BED4D27FFD9FFB25C0BFB7FFCD5FB74" ID-CoL="3GLTR" authorityName=": Boulenger" authorityYear="1898" box="[523,846,1204,1228]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25C0BFB7FFCD5FB74" box="[523,846,1204,1228]" italics="true" pageId="18" pageNumber="2343">Gnathonemus macrolepidotus</emphasis>
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)
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<paragraph id="BC5236A4FFD9FFB25ED3FB1EFD7DFB54" blockId="18.[211,1172,949,1355]" box="[211,742,1236,1260]" pageId="18" pageNumber="2343">
<treatmentCitationGroup id="9CFD118AFFD9FFB25ED3FB1EFD7DFB54" box="[211,742,1236,1260]" pageId="18" pageNumber="2343">
<treatmentCitation id="3D4C10B5FFD9FFB25ED3FB1EFD79FB54" author="Taverne L" box="[211,738,1236,1260]" page="102" pageId="18" pageNumber="2343" year="1971">
<taxonomicName id="7BED4D27FFD9FFB25ED3FB1EFD79FB54" ID-CoL="3Y396" authority=": Taverne 1971 a, p. 102" authorityName="Taverne" authorityPageNumber="102" authorityYear="1971" box="[211,738,1236,1260]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="moeruensis">
<emphasis id="8E99EAB6FFD9FFB25ED3FB1EFE45FB54" box="[211,478,1237,1260]" italics="true" pageId="18" pageNumber="2343">Marcusenius moeruensis</emphasis>
:
<bibRefCitation id="D87C4B55FFD9FFB25FEDFB1FFD0AFB54" author="Taverne L" box="[493,657,1236,1260]" pageId="18" pageNumber="2343" pagination="99 - 110" refId="ref17631" refString="Taverne L. 1971 a. Note sur la systematique des poissons Mormyriformes. Le probleme des genres Gnathonemus Gill, Marcusenius Gill, Hippopotamyrus Pappenheim, Cyphomyrus Myers et les nouveaux genres Pollimyrus et Brienomyrus. Revue de Zoologie et de Botanique Africaines. 84 (1 - 2): 99 - 110." type="journal article" year="1971">Taverne 1971a</bibRefCitation>
, p. 102
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC5236A4FFD9FFB25ED3FB38FE77FAF3" blockId="18.[211,1172,949,1355]" pageId="18" pageNumber="2343">
<treatmentCitationGroup id="9CFD118AFFD9FFB25ED3FB38FE77FAF3" pageId="18" pageNumber="2343">
<treatmentCitation id="3D4C10B5FFD9FFB25ED3FB38FD6CFAB3" author="Steindachner F" box="[211,759,1267,1292]" page="536" pageId="18" pageNumber="2343" year="1914">
<taxonomicName id="7BED4D27FFD9FFB25ED3FB38FD6CFAB3" ID-CoL="3GLTB" authority="Steindachner, 1914, p. 536" authorityName="Steindachner" authorityPageNumber="536" authorityYear="1914" box="[211,759,1267,1292]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="graeverti">
<emphasis id="8E99EAB6FFD9FFB25ED3FB38FE4BFAB4" bold="true" box="[211,464,1267,1292]" italics="true" pageId="18" pageNumber="2343">Gnathonemus graeverti</emphasis>
<bibRefCitation id="D87C4B55FFD9FFB25FD6FB38FD37FAB3" author="Steindachner F" box="[470,684,1267,1291]" pageId="18" pageNumber="2343" pagination="536 - 538" refId="ref17455" refString="Steindachner F. 1914. Bericht uber die ichthyologischen Aufsammlungen der Bruder Adolf und Albin Horn wahrend einer im Sommer 1913 ausgefuhrten Reise nach Deutsch-Ostafrika. Anzeiger der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse. 51: 536 - 538." type="journal article" year="1914">Steindachner, 1914</bibRefCitation>
, p. 536
</taxonomicName>
</treatmentCitation>
(in full:
<bibRefCitation id="D87C4B55FFD9FFB25D53FB38FBBAFAB3" author="Steindachner F" box="[851,1057,1267,1291]" pageId="18" pageNumber="2343" pagination="59 - 86" refId="ref17497" refString="Steindachner F. 1916. Bericht uber die ichthyologischen Aufsammlungen der Bruder Adolf und Albin Horn wahrend einer im Sommer 1913 ausgefuhrten Reise nach Deutsch-Ostafrika. Denkschriften der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse. 92: 59 - 86." type="journal article" year="1916">Steindachner 1916</bibRefCitation>
);
<treatmentCitation id="3D4C10B5FFD9FFB25A36FB38FEFDFA93" author="Matthes H" page="4" pageId="18" pageNumber="2343" year="1967">
<bibRefCitation id="D87C4B55FFD9FFB25A36FB38FEB3FA93" author="Matthes H" pageId="18" pageNumber="2343" refId="ref17028" refString="Matthes H. 1967. The fishes and fisheries of the Ruaha river basin, Tanzania (systematic, ecology, zoogeography, fisheries). East African Freshwater Fisheries Research Organization Occasional Paper [9]." type="book" year="1967">Matthes 1967</bibRefCitation>
, p. 4
</treatmentCitation>
(synonymy with
<taxonomicName id="7BED4D27FFD9FFB25C34FADFFCE0FA94" ID-CoL="3GLTR" authorityName=": Boulenger" authorityYear="1898" box="[564,891,1300,1324]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25C34FADFFCE0FA94" box="[564,891,1300,1324]" italics="true" pageId="18" pageNumber="2343">Gnathonemus macrolepidotus</emphasis>
</taxonomicName>
).
<treatmentCitation id="3D4C10B5FFD9FFB25D96FADFFE73FAF3" author="Taverne L" page="103" pageId="18" pageNumber="2343" year="1971">
<taxonomicName id="7BED4D27FFD9FFB25D96FADFFE73FAF3" ID-CoL="3Y386" authority=": Taverne 1971 a, p. 103" authorityName="Taverne" authorityPageNumber="103" authorityYear="1971" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="graeverti">
<emphasis id="8E99EAB6FFD9FFB25D96FADFFB17FA93" box="[918,1164,1300,1323]" italics="true" pageId="18" pageNumber="2343">Marcusenius graeverti</emphasis>
:
<bibRefCitation id="D87C4B55FFD9FFB25EF3FAFFFE0FFAF3" author="Taverne L" box="[243,404,1331,1355]" pageId="18" pageNumber="2343" pagination="99 - 110" refId="ref17631" refString="Taverne L. 1971 a. Note sur la systematique des poissons Mormyriformes. Le probleme des genres Gnathonemus Gill, Marcusenius Gill, Hippopotamyrus Pappenheim, Cyphomyrus Myers et les nouveaux genres Pollimyrus et Brienomyrus. Revue de Zoologie et de Botanique Africaines. 84 (1 - 2): 99 - 110." type="journal article" year="1971">Taverne 1971a</bibRefCitation>
, p. 103
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFD9FFB35ED3FA5AFE9EFB48" lastPageId="19" lastPageNumber="2344" pageId="18" pageNumber="2343" type="materials_examined">
<paragraph id="BC5236A4FFD9FFB25ED3FA5AFEE5FA10" blockId="18.[211,1171,1425,1584]" box="[211,382,1425,1448]" pageId="18" pageNumber="2343">
<heading id="E71A81C8FFD9FFB25ED3FA5AFEE5FA10" box="[211,382,1425,1448]" fontSize="10" level="3" pageId="18" pageNumber="2343" reason="5">
<emphasis id="8E99EAB6FFD9FFB25ED3FA5AFEE5FA10" box="[211,382,1425,1448]" italics="true" pageId="18" pageNumber="2343">
<typeStatus id="63568806FFD9FFB25ED3FA5AFE92FA10" box="[211,265,1425,1448]" pageId="18" pageNumber="2343">Type</typeStatus>
specimens
</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFD9FFB25ED3FA73FD0EF9B7" blockId="18.[211,1171,1425,1584]" pageId="18" pageNumber="2343">
<materialsCitation id="0C853CF9FFD9FFB25ED3FA73FE34FA57" collectingDate="1993-04-13" collectionCode="ZMB" collectorName="L. Seegers" country="Mozambique" location="Lower Zambezi" pageId="18" pageNumber="2343" specimenCode="ZMB 3678" specimenCount="2" stateProvince="Tete" typeStatus="lectotype">
<specimenCode id="EC4B9EDFFFD9FFB25ED3FA73FED0FA77" box="[211,331,1463,1487]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="18" pageNumber="2343" type="Museum">ZMB 3678</specimenCode>
(
<typeStatus id="63568806FFD9FFB25F58FA7CFE5BFA77" box="[344,448,1463,1487]" pageId="18" pageNumber="2343" type="lectotype">lectotype</typeStatus>
<collectorName id="11185372FFD9FFB25FC6FA73FDACFA77" box="[454,567,1463,1487]" pageId="18" pageNumber="2343">L. Seegers</collectorName>
<date id="C8531064FFD9FFB25C3CFA7CFD2DFA77" box="[572,694,1463,1487]" pageId="18" pageNumber="2343" value="1993-04-13">
<collectingDate id="D817E98CFFD9FFB25C3CFA7CFD2DFA77" box="[572,694,1463,1487]" pageId="18" pageNumber="2343" value="1993-04-13">13.04.1993</collectingDate>
</date>
) from
<collectingRegion id="7E29F846FFD9FFB25CFFFA73FCAAFA77" box="[767,817,1464,1487]" country="Mozambique" name="Tete" pageId="18" pageNumber="2343">Tete</collectingRegion>
,
<location id="B932607FFFD9FFB25D3BFA73FC76FA77" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD9FFB25D3BFA73FC76FA77" box="[827,1005,1463,1487]" country="Mozambique" name="Lower Zambezi" pageId="18" pageNumber="2343" stateProvince="Tete">Lower Zambezi</location>
,
<collectingCountry id="C4FA7634FFD9FFB25DF7FA7CFB14FA77" box="[1015,1167,1463,1487]" name="Mozambique" pageId="18" pageNumber="2343">Mozambique</collectingCountry>
, with
<typeStatus id="63568806FFD9FFB25F0BFA1CFE34FA57" box="[267,431,1495,1519]" pageId="18" pageNumber="2343" type="paralectotype">paralectotypes</typeStatus>
</materialsCitation>
<materialsCitation id="0C853CF9FFD9FFB25FB5FA13FD0AF9B7" collectionCode="ZMB" country="Mozambique" pageId="18" pageNumber="2343" specimenCode="ZMB 6730" specimenCount="3" stateProvince="Tete" typeStatus="holotype">
<specimenCode id="EC4B9EDFFFD9FFB25FB5FA13FDB6FA57" box="[437,557,1495,1519]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="18" pageNumber="2343" type="Museum">ZMB 6730</specimenCode>
(
<specimenCount id="AAEBFD2DFFD9FFB25C3BFA1CFD4CFA57" box="[571,727,1495,1519]" pageId="18" pageNumber="2343" type="generic">one specimen</specimenCount>
), 3677 (
<specimenCount id="AAEBFD2DFFD9FFB25D2CFA1CFC5CFA57" box="[812,967,1495,1519]" pageId="18" pageNumber="2343" type="generic">one specimen</specimenCount>
), and 32043 (
<specimenCount id="AAEBFD2DFFD9FFB25A59FA1CFED3F9B7" pageId="18" pageNumber="2343" type="generic">three specimens</specimenCount>
;
<bibRefCitation id="D87C4B55FFD9FFB25F53FA3CFE77F9B7" author="Seegers L" box="[339,492,1527,1551]" pageId="18" pageNumber="2343" pagination="1 - 407" refId="ref17257" refString="Seegers L. 1996. The fishes of the Lake Rukwa drainage. Annales Sciences Zoologiques (Tervuren). 278: 1 - 407." type="journal article" year="1996">Seegers, 1996</bibRefCitation>
), all examined
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFD9FFB25EFEF9D3FE6EF997" blockId="18.[211,1171,1425,1584]" box="[254,501,1559,1584]" pageId="18" pageNumber="2343">Non-types, examined:</paragraph>
<paragraph id="BC5236A4FFD9FFB25EFEF99CFBEFF916" blockId="18.[254,1172,1623,1711]" pageId="18" pageNumber="2343">
<emphasis id="8E99EAB6FFD9FFB25EFEF99CFE90F9D7" bold="true" box="[254,267,1623,1647]" pageId="18" pageNumber="2343">–</emphasis>
<materialsCitation id="0C853CF9FFD9FFB25F1CF993FBF6F916" collectingDate="1999-08-01" collectionCode="R" county="Lower" latitude="-18.564999" location="Bills" longLatPrecision="21" longitude="35.66278" municipality="System" pageId="18" pageNumber="2343" specimenCode="SAIAB 60847" specimenCount="81" stateProvince="Zambezi">
<collectingCounty id="55334E28FFD9FFB25F1CF993FEFFF9D7" box="[284,356,1624,1647]" pageId="18" pageNumber="2343">Lower</collectingCounty>
<collectingRegion id="7E29F846FFD9FFB25F6CF99CFE55F9D7" box="[364,462,1623,1647]" country="Namibia" name="Zambezi" pageId="18" pageNumber="2343">Zambezi</collectingRegion>
<collectingMunicipality id="5C36ACDEFFD9FFB25FD6F99CFDB1F9D7" box="[470,554,1623,1647]" pageId="18" pageNumber="2343">System</collectingMunicipality>
:
<specimenCode id="EC4B9EDFFFD9FFB25C35F99CFD4CF9D7" box="[565,727,1623,1647]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="18" pageNumber="2343" type="Other type of research institution/biorepository">SAIAB 60847</specimenCode>
, coll:
<location id="B932607FFFD9FFB25D19F99CFCD0F9D7" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD9FFB25D19F99CFCD0F9D7" box="[793,843,1623,1647]" county="Lower" latitude="-18.564999" longLatPrecision="21" longitude="35.66278" municipality="System" name="Bills" pageId="18" pageNumber="2343" stateProvince="Zambezi">Bills</location>
<collectionCode id="DAFCAE61FFD9FFB25D53F993FCF7F9D7" box="[851,876,1624,1647]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="2343">R</collectionCode>
.,
<date id="C8531064FFD9FFB25D7DF99CFBB9F9D7" box="[893,1058,1623,1647]" pageId="18" pageNumber="2343" value="1999-08-01">
<collectingDate id="D817E98CFFD9FFB25D7DF99CFBB9F9D7" box="[893,1058,1623,1647]" pageId="18" pageNumber="2343" value="1999-08-01">1 August 1999</collectingDate>
</date>
, tributary of the
<location id="B932607FFFD9FFB25F64F9BCFE37F936" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD9FFB25F64F9BCFE37F936" box="[356,428,1655,1678]" county="Lower" latitude="-18.564999" longLatPrecision="21" longitude="35.66278" municipality="System" name="Lower" pageId="18" pageNumber="2343" stateProvince="Zambezi">Lower</location>
<collectingRegion id="7E29F846FFD9FFB25FB2F9BCFD8FF937" box="[434,532,1655,1679]" country="Namibia" name="Zambezi" pageId="18" pageNumber="2343">Zambezi</collectingRegion>
in the delta region, southeast of
<location id="B932607FFFD9FFB25D82F9BCFC64F936" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD9FFB25D82F9BCFC64F936" box="[898,1023,1655,1678]" county="Lower" latitude="-18.564999" longLatPrecision="21" longitude="35.66278" municipality="System" name="Marromeu" pageId="18" pageNumber="2343" stateProvince="Zambezi">Marromeu</location>
,
<geoCoordinate id="D9D95063FFD9FFB25A09F9BCFB14F937" box="[1033,1167,1651,1679]" degrees="18" direction="south" minutes="33" orientation="latitude" pageId="18" pageNumber="2343" precision="15" seconds="54" value="-18.564999">
18
<superScript id="4B989BECFFD9FFB25A24F9B8FBB6F93C" attach="none" box="[1060,1069,1651,1668]" fontSize="7" pageId="18" pageNumber="2343">◦</superScript>
33’54” S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFD9FFB25F1CF95DFE3DF916" box="[284,422,1683,1710]" degrees="35" direction="east" minutes="39" orientation="longitude" pageId="18" pageNumber="2343" precision="15" seconds="46" value="35.66278">
35
<superScript id="4B989BECFFD9FFB25F37F958FEDBF91C" attach="none" box="[311,320,1683,1700]" fontSize="7" pageId="18" pageNumber="2343">◦</superScript>
39’46” E
</geoCoordinate>
,
<specimenCount id="AAEBFD2DFFD9FFB25FB1F95DFDDCF916" box="[433,583,1686,1710]" pageId="18" pageNumber="2343" type="generic">81 specimens</specimenCount>
, size range
<quantity id="7B159B41FFD9FFB25CC6F95DFCD3F916" box="[710,840,1686,1710]" metricMagnitude="-2" metricUnit="m" metricValue="7.8" metricValueMax="10.4" metricValueMin="5.2" pageId="18" pageNumber="2343" unit="mm" value="78.0" valueMax="104.0" valueMin="52.0">52–104 mm</quantity>
SL, locality 2 on
<figureCitation id="24D62A21FFD9FFB25A0FF95DFBF6F916" box="[1039,1133,1686,1710]" captionStart="Figure 1" captionStartId="4.[211,278,932,954]" captionTargetBox="[213,1170,167,899]" captionTargetId="figure-5@4.[213,1170,167,899]" captionTargetPageId="4" captionText="Figure 1. Map of sampling locations in southern Africa. (A) Type locality for Marcusenius macrolepidotus (Peters, 1852), Tete on the Lower Zambezi River; (B) origin of M. macrolepidotus, SAIAB 60847, from Marromeu, Lower Zambezi delta (Figure 4A); (C) origin of M. macrolepidotus from the Buzi System, same location as SAIAB 67369, presently alive in aquarium; (D) likely type region for Marcusenius angolensis (Boulenger, 1905), the Lower Quanza; (E) Marcusenius altisambesi Kramer et al., 2007, type locality on the Upper Zambezi River; (F) M. altisambesi from the Kwando River; (G) M. altisambesi from the Okavango delta; (H, I) Marcusenius multisquamatus sp. nov., stretching from below the Ruacana Falls to above the Epupa Falls (I), the type locality; (J) specimens from the Cunene River Mouth." figureDoi="http://doi.org/10.5281/zenodo.5197592" httpUri="https://zenodo.org/record/5197592/files/figure.png" pageId="18" pageNumber="2343">Figure 1</figureCitation>
</materialsCitation>
,
</paragraph>
<paragraph id="BC5236A4FFD8FFB35EBAFF6CFBABFEA7" blockId="19.[186,1104,167,670]" pageId="19" pageNumber="2344">
<emphasis id="8E99EAB6FFD8FFB35EBAFF6CFF5CFF07" bold="true" box="[186,199,167,191]" pageId="19" pageNumber="2344">–</emphasis>
<materialsCitation id="0C853CF9FFD8FFB35ED8FF6CFBB2FEA7" collectingDate="2002-09-29" collectionCode="R" county="Bills" latitude="-19.966667" location="Dombe Nyangapwa Stream" longLatPrecision="21" longitude="33.414448" municipality="Lucite River" pageId="19" pageNumber="2344" specimenCode="SAIAB 67369" specimenCount="25" stateProvince="Manica">
Buzi System:
<specimenCode id="EC4B9EDFFFD8FFB35F71FF6CFD89FF07" box="[369,530,167,191]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="19" pageNumber="2344" type="Other type of research institution/biorepository">SAIAB 67369</specimenCode>
, coll:
<collectingCounty id="55334E28FFD8FFB35C53FF6CFD1EFF07" box="[595,645,167,191]" pageId="19" pageNumber="2344">Bills</collectingCounty>
<collectionCode id="DAFCAE61FFD8FFB35C8CFF63FD3EFF07" box="[652,677,168,191]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="2344">R</collectionCode>
.,
<date id="C8531064FFD8FFB35CB6FF6CFC15FF07" box="[694,910,167,191]" pageId="19" pageNumber="2344" value="2002-09-29">
<collectingDate id="D817E98CFFD8FFB35CB6FF6CFC15FF07" box="[694,910,167,191]" pageId="19" pageNumber="2344" value="2002-09-29">29 September 2002</collectingDate>
</date>
, first tributary of the
<collectingMunicipality id="5C36ACDEFFD8FFB35F01FF0CFE17FF67" box="[257,396,199,223]" pageId="19" pageNumber="2344">Lucite River</collectingMunicipality>
leaving
<location id="B932607FFFD8FFB35FE6FF0CFC84FF67" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35FE6FF0CFC84FF67" box="[486,799,199,224]" county="Bills" latitude="-19.966667" longLatPrecision="21" longitude="33.414448" municipality="Lucite River" name="Dombe Nyangapwa Stream" pageId="19" pageNumber="2344" stateProvince="Manica">Dombe Nyangapwa Stream</location>
, which is part of the middle
<location id="B932607FFFD8FFB35ED8FF2CFE36FF47" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35ED8FF2CFE36FF47" box="[216,429,231,255]" county="Bills" latitude="-19.966667" longLatPrecision="21" longitude="33.414448" municipality="Lucite River" name="Buzi River System" pageId="19" pageNumber="2344" stateProvince="Manica">
<location id="B932607FFFD8FFB35ED8FF2CFECAFF47" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35ED8FF2CFECAFF47" box="[216,337,231,255]" county="Bills" latitude="-19.966667" longLatPrecision="21" longitude="33.414448" municipality="Lucite River" name="Buzi River" pageId="19" pageNumber="2344" stateProvince="Manica">
Buzi
<location id="B932607FFFD8FFB35F13FF2CFECAFF47" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35F13FF2CFECAFF47" box="[275,337,231,255]" county="Bills" latitude="-19.966667" longLatPrecision="21" longitude="33.414448" municipality="Lucite River" name="River System" pageId="19" pageNumber="2344" stateProvince="Manica">River</location>
</location>
System
</location>
, on the road east of
<location id="B932607FFFD8FFB35C95FF2CFD70FF47" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35C95FF2CFD70FF47" box="[661,747,231,255]" county="Bills" latitude="-19.966667" longLatPrecision="21" longitude="33.414448" municipality="Lucite River" name="Dombe" pageId="19" pageNumber="2344" stateProvince="Manica">Dombe</location>
,
<collectingRegion id="7E29F846FFD8FFB35CF7FF2CFC22FF47" box="[759,953,231,255]" country="Mozambique" name="Manica" pageId="19" pageNumber="2344">Manica Province</collectingRegion>
,
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19
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58’00” S
</geoCoordinate>
,
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33
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24’52” E
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,
<specimenCount id="AAEBFD2DFFD8FFB35F6DFECCFD98FEA7" box="[365,515,263,287]" pageId="19" pageNumber="2344" type="generic">25 specimens</specimenCount>
, size range
<quantity id="7B159B41FFD8FFB35C82FECCFC9FFEA7" box="[642,772,263,287]" metricMagnitude="-1" metricUnit="m" metricValue="1.065" metricValueMax="1.43" metricValueMin="0.7" pageId="19" pageNumber="2344" unit="mm" value="106.5" valueMax="143.0" valueMin="70.0">70–143 mm</quantity>
SL, locality 3 on
<figureCitation id="24D62A21FFD8FFB35DCBFECCFBB2FEA7" box="[971,1065,263,287]" captionStart="Figure 1" captionStartId="4.[211,278,932,954]" captionTargetBox="[213,1170,167,899]" captionTargetId="figure-5@4.[213,1170,167,899]" captionTargetPageId="4" captionText="Figure 1. Map of sampling locations in southern Africa. (A) Type locality for Marcusenius macrolepidotus (Peters, 1852), Tete on the Lower Zambezi River; (B) origin of M. macrolepidotus, SAIAB 60847, from Marromeu, Lower Zambezi delta (Figure 4A); (C) origin of M. macrolepidotus from the Buzi System, same location as SAIAB 67369, presently alive in aquarium; (D) likely type region for Marcusenius angolensis (Boulenger, 1905), the Lower Quanza; (E) Marcusenius altisambesi Kramer et al., 2007, type locality on the Upper Zambezi River; (F) M. altisambesi from the Kwando River; (G) M. altisambesi from the Okavango delta; (H, I) Marcusenius multisquamatus sp. nov., stretching from below the Ruacana Falls to above the Epupa Falls (I), the type locality; (J) specimens from the Cunene River Mouth." figureDoi="http://doi.org/10.5281/zenodo.5197592" httpUri="https://zenodo.org/record/5197592/files/figure.png" pageId="19" pageNumber="2344">Figure 1</figureCitation>
</materialsCitation>
,
</paragraph>
<paragraph id="BC5236A4FFD8FFB35EBAFEECFCE9FEC7" blockId="19.[186,1104,167,670]" pageId="19" pageNumber="2344">
<emphasis id="8E99EAB6FFD8FFB35EBAFEECFF5CFE87" bold="true" box="[186,199,295,319]" pageId="19" pageNumber="2344">–</emphasis>
<materialsCitation id="0C853CF9FFD8FFB35ED8FEECFCF6FEC7" collectingDate="2002-10-08" collectionCode="R, EOD" collectorName="Five &amp; Buzi System &amp; Bills" country="Germany" location="Regensburg" pageId="19" pageNumber="2344" specimenCount="1">
<collectorName id="11185372FFD8FFB35ED8FEECFE92FE87" box="[216,265,295,319]" pageId="19" pageNumber="2344">Five</collectorName>
live specimens from the
<collectorName id="11185372FFD8FFB35C25FEECFD2EFE87" box="[549,693,295,319]" pageId="19" pageNumber="2344">Buzi System</collectorName>
, same location as SAIAB 67369,
<collectionCode id="DAFCAE61FFD8FFB35A34FEE3FBD7FE87" box="[1076,1100,296,319]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="2344">R</collectionCode>
.
<collectorName id="11185372FFD8FFB35ED8FE8CFE91FEE7" box="[216,266,327,351]" pageId="19" pageNumber="2344">Bills</collectorName>
coll., arrived
<date id="C8531064FFD8FFB35FABFE8CFDC0FEE7" box="[427,603,327,351]" pageId="19" pageNumber="2344" value="2002-10-08">
<collectingDate id="D817E98CFFD8FFB35FABFE8CFDC0FEE7" box="[427,603,327,351]" pageId="19" pageNumber="2344" value="2002-10-08">
8 October
<quantity id="7B159B41FFD8FFB35C26FE8CFDC0FEE7" box="[550,603,327,351]" metricMagnitude="1" metricUnit="m" metricValue="5.08508" pageId="19" pageNumber="2344" unit="in" value="2002.0">2002</quantity>
</collectingDate>
</date>
in
<location id="B932607FFFD8FFB35C83FE8CFC91FEE7" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35C83FE8CFC91FEE7" box="[643,778,327,351]" country="Germany" name="Regensburg" pageId="19" pageNumber="2344">Regensburg</location>
,
<collectingCountry id="C4FA7634FFD8FFB35D17FE8CFC19FEE7" box="[791,898,327,351]" name="Germany" pageId="19" pageNumber="2344">Germany</collectingCountry>
, studied for
<collectionCode id="DAFCAE61FFD8FFB35A13FE8CFBD4FEE7" box="[1043,1103,327,351]" pageId="19" pageNumber="2344">EOD</collectionCode>
and meristic characters, size range
<quantity id="7B159B41FFD8FFB35C5EFEACFD48FEC7" box="[606,723,359,383]" metricMagnitude="-2" metricUnit="m" metricValue="8.35" metricValueMax="8.8" metricValueMin="7.9" pageId="19" pageNumber="2344" unit="mm" value="83.5" valueMax="88.0" valueMin="79.0">79–88 mm</quantity>
SL, still alive
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFD8FFB35EBAFE4CFD5CFD26" blockId="19.[186,1104,167,670]" pageId="19" pageNumber="2344">
<emphasis id="8E99EAB6FFD8FFB35EBAFE4CFF5CFE27" bold="true" box="[186,199,391,415]" pageId="19" pageNumber="2344">–</emphasis>
<materialsCitation id="0C853CF9FFD8FFB35ED8FE4CFCD5FE46" collectionCode="IPMB, R" country="Mozambique" county="Nyangapwa Stream" latitude="-19.966667" location="Samples" longLatPrecision="21" longitude="33.414448" municipality="Lucite River" pageId="19" pageNumber="2344" specimenCode="IPMB 22682, 22684, 22685, SAIAB 67369" specimenCount="1">
<location id="B932607FFFD8FFB35ED8FE4CFEADFE27" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35ED8FE4CFEADFE27" box="[216,310,391,415]" country="Mozambique" county="Nyangapwa Stream" latitude="-19.966667" longLatPrecision="21" longitude="33.414448" municipality="Lucite River" name="Samples" pageId="19" pageNumber="2344">Samples</location>
examined for genetics. DNA samples are stored at Institute of Pharmacy and
<location id="B932607FFFD8FFB35F81FE6DFD04FE06" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35F81FE6DFD04FE06" box="[385,671,422,446]" country="Mozambique" county="Nyangapwa Stream" latitude="-19.966667" longLatPrecision="21" longitude="33.414448" municipality="Lucite River" name="Molecular Biotechnology" pageId="19" pageNumber="2344">Molecular Biotechnology</location>
,
<location id="B932607FFFD8FFB35CA9FE6DFC04FE06" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35CA9FE6DFC04FE06" box="[681,927,422,446]" country="Mozambique" county="Nyangapwa Stream" latitude="-19.966667" longLatPrecision="21" longitude="33.414448" municipality="Lucite River" name="Heidelberg University" pageId="19" pageNumber="2344">Heidelberg University</location>
(
<collectionCode id="DAFCAE61FFD8FFB35DADFE6DFC63FE06" box="[941,1016,422,446]" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="19" pageNumber="2344">IPMB</collectionCode>
).
<specimenCode id="EC4B9EDFFFD8FFB35A09FE6CFE85FE66" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="19" pageNumber="2344">IPMB 22682</specimenCode>
,
<specimenCode id="EC4B9EDFFFD8FFB35F29FE0DFEF4FE66" box="[297,367,454,478]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="19" pageNumber="2344">22684</specimenCode>
,
<specimenCode id="EC4B9EDFFFD8FFB35F79FE0DFE20FE66" box="[377,443,454,478]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="19" pageNumber="2344">22685</specimenCode>
(
<specimenCode id="EC4B9EDFFFD8FFB35FCAFE0DFDF6FE66" box="[458,621,454,478]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="19" pageNumber="2344" type="Other type of research institution/biorepository">SAIAB 67369</specimenCode>
),
<collectingCountry id="C4FA7634FFD8FFB35C7FFE0DFC82FE66" box="[639,793,454,478]" name="Mozambique" pageId="19" pageNumber="2344">Mozambique</collectingCountry>
:
<location id="B932607FFFD8FFB35D23FE0DFC2AFE66" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35D23FE0DFC2AFE66" box="[803,945,454,478]" country="Mozambique" county="Nyangapwa Stream" latitude="-19.966667" longLatPrecision="21" longitude="33.414448" municipality="Lucite River" name="Buzi System" pageId="19" pageNumber="2344">Buzi System</location>
:
<collectingMunicipality id="5C36ACDEFFD8FFB35DBCFE0DFBD7FE66" box="[956,1100,454,478]" pageId="19" pageNumber="2344">Lucite River</collectingMunicipality>
:
<collectingCounty id="55334E28FFD8FFB35ED8FE2CFE27FE46" box="[216,444,486,511]" pageId="19" pageNumber="2344">Nyangapwa Stream</collectingCounty>
,
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19
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58
<superScript id="4B989BECFFD8FFB35C0AFE29FD94FE4B" attach="left" box="[522,527,482,499]" fontSize="7" pageId="19" pageNumber="2344">′</superScript>
00” S
</geoCoordinate>
,
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33
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24
<superScript id="4B989BECFFD8FFB35CA1FE29FD3DFE4B" attach="none" box="[673,678,482,499]" fontSize="7" pageId="19" pageNumber="2344">′</superScript>
52” E
</geoCoordinate>
, coll.
<collectionCode id="DAFCAE61FFD8FFB35D36FE2CFCD5FE46" box="[822,846,487,510]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="2344">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="0C853CF9FFD8FFB35D5CFE2DFCD6FD86" collectingDate="2002-09-29" collectionCode="R" collectorName="Bills &amp; S. Chimela &amp; A. Chivindzi" country="Mozambique" latitude="-18.283333" location="Lower Zambezi" longLatPrecision="1275" longitude="35.95" municipality="Marromeu" pageId="19" pageNumber="2344" specimenCode="IPMB 22686, SAIAB 61592" specimenCount="1">
<collectorName id="11185372FFD8FFB35D5CFE2DFC14FE46" box="[860,911,486,510]" pageId="19" pageNumber="2344">Bills</collectorName>
,
<collectorName id="11185372FFD8FFB35D9DFE2DFB80FE46" box="[925,1051,486,510]" pageId="19" pageNumber="2344">S. Chimela</collectorName>
and
<collectorName id="11185372FFD8FFB35ED8FDCDFEF0FDA6" box="[216,363,518,542]" pageId="19" pageNumber="2344">A. Chivindzi</collectorName>
,
<date id="C8531064FFD8FFB35F77FDCDFDCBFDA6" box="[375,592,518,542]" pageId="19" pageNumber="2344" value="2002-09-29">
<collectingDate id="D817E98CFFD8FFB35F77FDCDFDCBFDA6" box="[375,592,518,542]" pageId="19" pageNumber="2344" value="2002-09-29">29 September 2002</collectingDate>
</date>
;
<specimenCode id="EC4B9EDFFFD8FFB35C5CFDCCFD77FDA6" box="[604,748,518,542]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="19" pageNumber="2344">IPMB 22686</specimenCode>
(
<specimenCode id="EC4B9EDFFFD8FFB35CFBFDCDFC04FDA6" box="[763,927,518,542]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="19" pageNumber="2344" type="Other type of research institution/biorepository">SAIAB 61592</specimenCode>
),
<collectingCountry id="C4FA7634FFD8FFB35DB2FDCDFBD7FDA6" box="[946,1100,518,542]" name="Mozambique" pageId="19" pageNumber="2344">Mozambique</collectingCountry>
:
<location id="B932607FFFD8FFB35ED8FDECFE14FD86" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35ED8FDECFE14FD86" box="[216,399,550,574]" country="Mozambique" latitude="-18.283333" longLatPrecision="1275" longitude="35.95" municipality="Marromeu" name="Lower Zambezi" pageId="19" pageNumber="2344">Lower Zambezi</location>
:
<collectingMunicipality id="5C36ACDEFFD8FFB35F9AFDECFD8CFD86" box="[410,535,551,574]" pageId="19" pageNumber="2344">Marromeu</collectingMunicipality>
,
<geoCoordinate id="D9D95063FFD8FFB35C23FDEDFD1AFD86" box="[547,641,546,574]" degrees="18" direction="south" minutes="17" orientation="latitude" pageId="19" pageNumber="2344" precision="925" value="-18.283333">
18
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17
<superScript id="4B989BECFFD8FFB35C63FDE9FDF3FD8B" attach="left" box="[611,616,546,563]" fontSize="7" pageId="19" pageNumber="2344">′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFD8FFB35C8EFDEDFD6BFD86" box="[654,752,546,574]" degrees="35" direction="east" minutes="57" orientation="longitude" pageId="19" pageNumber="2344" precision="925" value="35.95">
35
<superScript id="4B989BECFFD8FFB35CA8FDE9FD2AFD8B" attach="left" box="[680,689,546,563]" fontSize="7" pageId="19" pageNumber="2344">◦</superScript>
57
<superScript id="4B989BECFFD8FFB35CCDFDE9FD49FD8B" attach="left" box="[717,722,546,563]" fontSize="7" pageId="19" pageNumber="2344">′</superScript>
E
</geoCoordinate>
, coll.
<collectionCode id="DAFCAE61FFD8FFB35D35FDECFCD6FD86" box="[821,845,551,574]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="2344">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="0C853CF9FFD8FFB35D58FDEDFD21FDC6" collectingDate="1999-08-11" collectionCode="R" collectorName="Bills" country="Mozambique" latitude="-18.316668" location="Lower Zambezi" longLatPrecision="1275" longitude="35.916668" municipality="Lagoon" pageId="19" pageNumber="2344" specimenCode="IPMB 22689, SAIAB 61603" specimenCount="1">
<collectorName id="11185372FFD8FFB35D58FDEDFC10FD86" box="[856,907,550,574]" pageId="19" pageNumber="2344">Bills</collectorName>
,
<date id="C8531064FFD8FFB35D98FDEDFBD0FD86" box="[920,1099,550,574]" pageId="19" pageNumber="2344" value="1999-08-11">
<collectingDate id="D817E98CFFD8FFB35D98FDEDFBD0FD86" box="[920,1099,550,574]" pageId="19" pageNumber="2344" value="1999-08-11">11 August 1999</collectingDate>
</date>
;
<specimenCode id="EC4B9EDFFFD8FFB35ED8FD8CFEF1FDE6" box="[216,362,582,606]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="19" pageNumber="2344">IPMB 22689</specimenCode>
(
<specimenCode id="EC4B9EDFFFD8FFB35F7BFD8DFDBAFDE6" box="[379,545,582,606]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="19" pageNumber="2344" type="Other type of research institution/biorepository">SAIAB 61603</specimenCode>
),
<collectingCountry id="C4FA7634FFD8FFB35C35FD8DFD54FDE6" box="[565,719,582,606]" name="Mozambique" pageId="19" pageNumber="2344">Mozambique</collectingCountry>
:
<location id="B932607FFFD8FFB35CDDFD8CFC0FFDE6" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35CDDFD8CFC0FFDE6" box="[733,916,582,606]" country="Mozambique" latitude="-18.316668" longLatPrecision="1275" longitude="35.916668" municipality="Lagoon" name="Lower Zambezi" pageId="19" pageNumber="2344">Lower Zambezi</location>
:
<collectingMunicipality id="5C36ACDEFFD8FFB35DA1FD8CFC62FDE7" box="[929,1017,583,607]" pageId="19" pageNumber="2344">Lagoon</collectingMunicipality>
behind sugar fields,
<geoCoordinate id="D9D95063FFD8FFB35F72FDADFE4DFDC6" box="[370,470,610,638]" degrees="18" direction="south" minutes="19" orientation="latitude" pageId="19" pageNumber="2344" precision="925" value="-18.316668">
18
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19
<superScript id="4B989BECFFD8FFB35FB2FDA9FE2CFDCB" attach="left" box="[434,439,610,627]" fontSize="7" pageId="19" pageNumber="2344">′</superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFD8FFB35FE9FDADFDCAFDC6" box="[489,593,610,638]" degrees="35" direction="east" minutes="55" orientation="longitude" pageId="19" pageNumber="2344" precision="925" value="35.916668">
35
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55
<superScript id="4B989BECFFD8FFB35C29FDA9FDB5FDCB" attach="left" box="[553,558,610,627]" fontSize="7" pageId="19" pageNumber="2344">′</superScript>
E
</geoCoordinate>
, coll.
<collectionCode id="DAFCAE61FFD8FFB35CA2FDACFD21FDC6" box="[674,698,615,638]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="2344">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="0C853CF9FFD8FFB35CCCFDADFD59FD26" accessionNumber="KC202222, KC202226" collectingDate="1999-08-13" collectionCode="KC" collectorName="Bills" country="Mozambique" location="GenBank" pageId="19" pageNumber="2344" specimenCount="1">
<collectorName id="11185372FFD8FFB35CCCFDADFD64FDC6" box="[716,767,614,638]" pageId="19" pageNumber="2344">Bills</collectorName>
,
<date id="C8531064FFD8FFB35D11FDADFC4BFDC6" box="[785,976,614,638]" pageId="19" pageNumber="2344" value="1999-08-13">
<collectingDate id="D817E98CFFD8FFB35D11FDADFC4BFDC6" box="[785,976,614,638]" pageId="19" pageNumber="2344" value="1999-08-13">13 August 1999</collectingDate>
</date>
;
<location id="B932607FFFD8FFB35DE3FDADFBD4FDC6" LSID="urn:lsid:plazi:treatment:344487B2FFD9FFB35FC8FC9EFE9EFB48:B932607FFFD8FFB35DE3FDADFBD4FDC6" box="[995,1103,614,638]" country="Mozambique" name="GenBank" pageId="19" pageNumber="2344">GenBank</location>
accession numbers: (
<accessionNumber id="A3BEAB47FFD8FFB35FBEFD4DFDA1FD26" box="[446,570,646,670]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC202222" pageId="19" pageNumber="2344">
<collectionCode id="DAFCAE61FFD8FFB35FBEFD4DFE7FFD26" box="[446,484,646,670]" pageId="19" pageNumber="2344">KC</collectionCode>
202222
</accessionNumber>
-
<accessionNumber id="A3BEAB47FFD8FFB35C40FD4DFD20FD26" box="[576,699,646,670]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC202226" pageId="19" pageNumber="2344">
<collectionCode id="DAFCAE61FFD8FFB35C40FD4DFDFDFD26" box="[576,614,646,670]" pageId="19" pageNumber="2344">KC</collectionCode>
202226
</accessionNumber>
)
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFD8FFB35E8FFD18FE85FD52" blockId="19.[143,1104,722,1264]" box="[143,286,722,746]" pageId="19" pageNumber="2344">
<heading id="E71A81C8FFD8FFB35E8FFD18FE85FD52" box="[143,286,722,746]" fontSize="10" level="3" pageId="19" pageNumber="2344" reason="5">
<emphasis id="8E99EAB6FFD8FFB35E8FFD18FE85FD52" box="[143,286,722,746]" italics="true" pageId="19" pageNumber="2344">
<typeStatus id="63568806FFD8FFB35E8FFD18FF5EFD52" box="[143,197,723,746]" pageId="19" pageNumber="2344">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFD8FFB35E8FFD31FF46FCEA" blockId="19.[143,1104,722,1264]" pageId="19" pageNumber="2344">
<materialsCitation id="0C853CF9FFD8FFB35E8FFD31FF43FCEA" country="Mozambique" county="Lower Zambezi" location="Licuare River" municipality="Lower Zambezi" pageId="19" pageNumber="2344" specimenCount="1" stateProvince="Tete" typeStatus="holotype">
<collectingCountry id="C4FA7634FFD8FFB35E8FFD31FEB2FCAA" box="[143,297,762,786]" name="Mozambique" pageId="19" pageNumber="2344">Mozambique</collectingCountry>
:
<collectingCounty id="55334E28FFD8FFB35F37FD30FE74FCAA" box="[311,495,762,786]" pageId="19" pageNumber="2344">Lower Zambezi</collectingCounty>
:
<collectingRegion id="7E29F846FFD8FFB35FFCFD30FDB7FCAA" box="[508,556,763,786]" country="Mozambique" name="Tete" pageId="19" pageNumber="2344">Tete</collectingRegion>
(no. 1,
<figureCitation id="24D62A21FFD8FFB35C8AFD31FD70FCAA" box="[650,747,762,786]" captionStart="Figure 1" captionStartId="4.[211,278,932,954]" captionTargetBox="[213,1170,167,899]" captionTargetId="figure-5@4.[213,1170,167,899]" captionTargetPageId="4" captionText="Figure 1. Map of sampling locations in southern Africa. (A) Type locality for Marcusenius macrolepidotus (Peters, 1852), Tete on the Lower Zambezi River; (B) origin of M. macrolepidotus, SAIAB 60847, from Marromeu, Lower Zambezi delta (Figure 4A); (C) origin of M. macrolepidotus from the Buzi System, same location as SAIAB 67369, presently alive in aquarium; (D) likely type region for Marcusenius angolensis (Boulenger, 1905), the Lower Quanza; (E) Marcusenius altisambesi Kramer et al., 2007, type locality on the Upper Zambezi River; (F) M. altisambesi from the Kwando River; (G) M. altisambesi from the Okavango delta; (H, I) Marcusenius multisquamatus sp. nov., stretching from below the Ruacana Falls to above the Epupa Falls (I), the type locality; (J) specimens from the Cunene River Mouth." figureDoi="http://doi.org/10.5281/zenodo.5197592" httpUri="https://zenodo.org/record/5197592/files/figure.png" pageId="19" pageNumber="2344">Figure 1</figureCitation>
), and tributaries to the
<collectingMunicipality id="5C36ACDEFFD8FFB35A07FD30FF6FFC8A" pageId="19" pageNumber="2344">Lower Zambezi</collectingMunicipality>
, also
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(Zambezi delta region). For more detail,
<bibRefCitation id="D87C4B55FFD8FFB35DB5FCD0FF43FCEA" author="Kramer B &amp; Skelton PH &amp; Van der Bank FH &amp; Wink M" pageId="19" pageNumber="2344" pagination="647 - 708" refId="ref16697" refString="Kramer B, Skelton PH, Van der Bank FH, Wink M. 2007. Allopatric differentiation in the Marcusenius macrolepidotus species complex in southern and eastern Africa: the resurrection of M. pongolensis and M. angolensis, and the description of two new species (Mormyridae, Teleostei). J Nat Hist. 41: 647 - 708." type="journal article" year="2007">Kramer et al. (2007)</bibRefCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFD8FFB35EBAFC91FE9EFB48" blockId="19.[143,1104,722,1264]" pageId="19" pageNumber="2344">
Since
<bibRefCitation id="D87C4B55FFD8FFB35EFCFC91FE59FCCA" author="Taverne L" box="[252,450,858,882]" pageId="19" pageNumber="2344" refId="ref17687" refString="Taverne L. 1971 b. Osteologie des genres Marcusenius Gill, Hippopotamyrus Pappenheim, Cyphomyrus Myers, Pollimyrus Taverne et Brienomyrus Taverne (Pisces Mormyriformes). Annales Musee Royal de l'Afrique Centrale, Sciences Zoologiques. 188: x - 144 + 3 plates." type="book" year="1971">Taverne’s (1971b)</bibRefCitation>
major revision of several genera of the
<taxonomicName id="7BED4D27FFD8FFB35D7BFC91FB97FCCA" box="[891,1036,858,882]" class="Actinopterygii" family="Mormyridae" kingdom="Animalia" order="Osteoglossiformes" pageId="19" pageNumber="2344" phylum="Chordata" rank="family">Mormyridae</taxonomicName>
, there are no members of
<taxonomicName id="7BED4D27FFD8FFB35F68FCB1FE65FC2A" authorityName="Worthington" authorityYear="1929" box="[360,510,890,914]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="19" pageNumber="2344" phylum="Chordata" rank="genus">
<emphasis id="8E99EAB6FFD8FFB35F68FCB1FE65FC2A" box="[360,510,890,914]" italics="true" pageId="19" pageNumber="2344">Gnathonemus</emphasis>
</taxonomicName>
in southern Africa. Two subspecies of
<taxonomicName id="7BED4D27FFD8FFB35DB3FCB0FF51FC0A" authorityName=": Boulenger" authorityYear="1898" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="19" pageNumber="2344" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD8FFB35DB3FCB0FF51FC0A" italics="true" pageId="19" pageNumber="2344">M. macrolepidotus</emphasis>
</taxonomicName>
were recognized (
<bibRefCitation id="D87C4B55FFD8FFB35F8FFC52FDEBFC09" author="Poll M &amp; Gosse J-P" box="[399,624,921,945]" pageId="19" pageNumber="2344" pagination="43 - 111" refId="ref17221" refString="Poll M, Gosse J-P. 1963. Contribution a l'etude systematique de la faune ichthyologique du Congo Central. Annales Musee Royal de l'Afrique Centrale, Sciences Zoologiques. 116: 43 - 111." type="journal article" year="1963">Poll and Gosse 1963</bibRefCitation>
):
<emphasis id="8E99EAB6FFD8FFB35C86FC50FCEFFC0A" box="[646,884,922,946]" italics="true" pageId="19" pageNumber="2344">
M.
<taxonomicName id="7BED4D27FFD8FFB35CADFC50FCEFFC0A" authorityName=": Boulenger" authorityYear="1898" box="[685,884,922,946]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="19" pageNumber="2362" phylum="Chordata" rank="species" species="macrolepidotus">m. macrolepidotus</taxonomicName>
</emphasis>
and
<emphasis id="8E99EAB6FFD8FFB35DAAFC50FF30FC69" italics="true" pageId="19" pageNumber="2344">
M.
<taxonomicName id="7BED4D27FFD8FFB35DD1FC50FF30FC69" authorityName="Boulenger" authorityYear="1905" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="19" pageNumber="2357" phylum="Chordata" rank="species" species="angolensis">m. angolensis</taxonomicName>
</emphasis>
, listed by
<bibRefCitation id="D87C4B55FFD8FFB35F1CFC72FE2BFC69" author="Gosse J-P" box="[284,432,953,977]" pageId="19" pageNumber="2344" pagination="63 - 124" refId="ref16269" refString="Gosse J-P. 1984. Mormyriformes. In: Daget J, Gosse J-P, Thys van den Audenaerde DFE, editors. Check-list of the freshwater fishes of Africa. Bondy (France) / Tervuren. Belgium: ORSTOM / MRAC; p. 63 - 124." type="book chapter" year="1984">Gosse (1984)</bibRefCitation>
. Based on morphology,
<bibRefCitation id="D87C4B55FFD8FFB35CBFFC71FC38FC69" author="Kramer B &amp; Skelton PH &amp; Van der Bank FH &amp; Wink M" box="[703,931,953,977]" pageId="19" pageNumber="2344" pagination="647 - 708" refId="ref16697" refString="Kramer B, Skelton PH, Van der Bank FH, Wink M. 2007. Allopatric differentiation in the Marcusenius macrolepidotus species complex in southern and eastern Africa: the resurrection of M. pongolensis and M. angolensis, and the description of two new species (Mormyridae, Teleostei). J Nat Hist. 41: 647 - 708." type="journal article" year="2007">Kramer et al. (2007)</bibRefCitation>
resurrected the latter subspecies as a valid species, and in consequence dropped the former subspecies. The origin of
<taxonomicName id="7BED4D27FFD8FFB35F26FC32FDEBFBA9" authority="Boulenger 1915" authorityName="Boulenger" authorityYear="1915" box="[294,624,1017,1041]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="19" pageNumber="2344" phylum="Chordata" rank="species" species="moeruensis">
<emphasis id="8E99EAB6FFD8FFB35F26FC32FE27FBA9" box="[294,444,1017,1041]" italics="true" pageId="19" pageNumber="2344">G. moeruensis</emphasis>
<bibRefCitation id="D87C4B55FFD8FFB35FC2FC32FDEBFBA9" author="Boulenger GA" box="[450,624,1017,1041]" pageId="19" pageNumber="2344" pagination="162 - 171" refId="ref16011" refString="Boulenger GA. 1915. Diagnoses de poissons nouveaux. II. Mormyrides, Kneriides, Characinides, Cyprinides, Silurides. Revue de Zoologie Africaine. 4 (2): 162 - 171." type="journal article" year="1915">Boulenger 1915</bibRefCitation>
</taxonomicName>
, a single specimen from Lake Mweru in the
<collectingCountry id="C4FA7634FFD8FFB35E8FFBD2FE76FB89" box="[143,493,1049,1073]" name="Democratic Republic of the Congo" pageId="19" pageNumber="2344">Democratic Republic of Congo</collectingCountry>
(
<collectingCountry id="C4FA7634FFD8FFB35FFCFBD2FDA1FB89" box="[508,570,1049,1073]" name="Democratic Republic of the Congo" pageId="19" pageNumber="2344">Zaïre</collectingCountry>
)/
<collectingCountry id="C4FA7634FFD8FFB35C4EFBD2FD32FB89" box="[590,681,1049,1073]" name="Zambia" pageId="19" pageNumber="2344">Zambia</collectingCountry>
, west of Lake Tanganyika, forms part of the Lualaba/
<collectingCountry id="C4FA7634FFD8FFB35F47FBF2FE08FBE9" box="[327,403,1081,1105]" name="Democratic Republic of the Congo" pageId="19" pageNumber="2344">Congo</collectingCountry>
drainage system, and from morphology the synonymization appears questionable to
<bibRefCitation id="D87C4B55FFD8FFB35FA8FB91FD0DFBC9" author="Kramer B &amp; Skelton PH &amp; Van der Bank FH &amp; Wink M" box="[424,662,1113,1137]" pageId="19" pageNumber="2344" pagination="647 - 708" refId="ref16697" refString="Kramer B, Skelton PH, Van der Bank FH, Wink M. 2007. Allopatric differentiation in the Marcusenius macrolepidotus species complex in southern and eastern Africa: the resurrection of M. pongolensis and M. angolensis, and the description of two new species (Mormyridae, Teleostei). J Nat Hist. 41: 647 - 708." type="journal article" year="2007">Kramer et al. (2007)</bibRefCitation>
. Also synonymized with
<taxonomicName id="7BED4D27FFD8FFB35DB9FB92FF51FB29" authorityName=": Boulenger" authorityYear="1898" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="19" pageNumber="2344" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD8FFB35DB9FB92FF51FB29" italics="true" pageId="19" pageNumber="2344">G. macrolepidotus</emphasis>
</taxonomicName>
was
<taxonomicName id="7BED4D27FFD8FFB35F05FBB2FDFBFB29" authority="Steindachner 1914" authorityName="Steindachner" authorityYear="1914" box="[261,608,1145,1169]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="19" pageNumber="2344" phylum="Chordata" rank="species" species="graeverti">
<emphasis id="8E99EAB6FFD8FFB35F05FBB2FE1EFB29" box="[261,389,1145,1169]" italics="true" pageId="19" pageNumber="2344">G. graeverti</emphasis>
<bibRefCitation id="D87C4B55FFD8FFB35F8FFBB2FDFBFB29" author="Steindachner F" box="[399,608,1145,1169]" pageId="19" pageNumber="2344" pagination="536 - 538" refId="ref17455" refString="Steindachner F. 1914. Bericht uber die ichthyologischen Aufsammlungen der Bruder Adolf und Albin Horn wahrend einer im Sommer 1913 ausgefuhrten Reise nach Deutsch-Ostafrika. Anzeiger der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse. 51: 536 - 538." type="journal article" year="1914">Steindachner 1914</bibRefCitation>
</taxonomicName>
whose origin is the Ruaha, an east-flowing river south of
<collectingRegion id="7E29F846FFD8FFB35F30FB52FE42FB09" box="[304,473,1177,1201]" country="Tanzania" name="Dar es Salaam" pageId="19" pageNumber="2344">Dar es Salaam</collectingRegion>
,
<collectingCountry id="C4FA7634FFD8FFB35FE4FB52FDD4FB09" box="[484,591,1177,1201]" name="Tanzania" pageId="19" pageNumber="2344">Tanzania</collectingCountry>
; the unique
<typeStatus id="63568806FFD8FFB35CDBFB52FCA4FB09" box="[731,831,1177,1201]" pageId="19" pageNumber="2344" type="holotype">holotype</typeStatus>
is lost; the synonymization is questionable (
<bibRefCitation id="D87C4B55FFD8FFB35F7FFB73FDC1FB68" author="Kramer B &amp; Skelton PH &amp; Van der Bank FH &amp; Wink M" box="[383,602,1208,1232]" pageId="19" pageNumber="2344" pagination="647 - 708" refId="ref16697" refString="Kramer B, Skelton PH, Van der Bank FH, Wink M. 2007. Allopatric differentiation in the Marcusenius macrolepidotus species complex in southern and eastern Africa: the resurrection of M. pongolensis and M. angolensis, and the description of two new species (Mormyridae, Teleostei). J Nat Hist. 41: 647 - 708." type="journal article" year="2007">Kramer et al. 2007</bibRefCitation>
).
<taxonomicName id="7BED4D27FFD8FFB35C74FB72FC63FB68" authority="Pappenheim 1907" authorityName="Pappenheim" authorityYear="1907" box="[628,1016,1208,1233]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="19" pageNumber="2344" phylum="Chordata" rank="species" species="okavangensis">
<emphasis id="8E99EAB6FFD8FFB35C74FB72FCBEFB69" box="[628,805,1209,1233]" italics="true" pageId="19" pageNumber="2344">G. okavangensis</emphasis>
<bibRefCitation id="D87C4B55FFD8FFB35D2EFB73FC63FB68" author="Pappenheim P" box="[814,1016,1208,1232]" pageId="19" pageNumber="2344" pagination="339 - 367" refId="ref17154" refString="Pappenheim P. 1907. Zur Systematik und Variationsstatistik der Mormyriden hauptsachlich aus den deutsch-afrikanischen Schutzgebieten. Nach dem Material des Koniglichen Zoologischen Museums zu Berlin. Mitteilungen aus dem Zoologischen Museum in Berlin. 3: 339 - 367." type="journal article" year="1907">Pappenheim 1907</bibRefCitation>
</taxonomicName>
is “not available”.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:name id="6D57D7889F9FE904340898CCF8D4CC66" type="personal">
<mods:role id="1B28E798189A27C3B45CACE1BEF0D5CA">
<mods:roleTerm id="08897D8025E7786AD4E2706AD16C937F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="00E4E67BCA8934E70A885C9A3E62D735">Eichmeier, Ales</mods:namePart>
<mods:nameIdentifier id="388A474656E81D3985A64B043C29496C" type="ORCID">0000-0001-7659-8852</mods:nameIdentifier>
<mods:affiliation id="5195707171C6B887FFD31265E83768F0">Mendeleum-Institute of Genetics, Faculty of Horticulture, Mendel University in Brno, Valtická 334, 691 44 Lednice, Czech Republic. milan. spetik @ mendelu. cz; https: // orcid. org / 0000 - 0001 - 7659 - 8852</mods:affiliation>
<mods:nameIdentifier id="3683A9E1D5E1AD1E7C218B84BC967416" type="email">milan.spetik@mendelu.cz</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="767F7E81651034C723E97A8757DA0DB3">text</mods:typeOfResource>
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<mods:titleInfo id="C9DBF49FB1BE36A61CC666695B25D7D5">
<mods:title id="4A9AB80C7EE5AD5FBA742EEE3B62925D">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="F160C5AFB7DB374263F6FBF8F30995B7">
<mods:date id="F55113037A9394535A873339A22B116F">2022</mods:date>
<mods:detail id="33FA5DD82FF19D950D34BC94163E31E5" type="pubDate">
<mods:number id="46D5898BB1695842864BF1505716B1AB">2022-08-25</mods:number>
</mods:detail>
<mods:detail id="DB94903393B0E5A4B771709EBEF79BE9" type="volume">
<mods:number id="CB753BCD2193B23C62B2F52487E1E320">559</mods:number>
</mods:detail>
<mods:detail id="F41A84C922413222093B44910C9BCED4" type="issue">
<mods:number id="E2B90490DEA08BC8FA95B771982549DD">3</mods:number>
</mods:detail>
<mods:extent id="F88E63E3F971CD87598CA4B1827ECA7B" unit="page">
<mods:start id="CDEB0E18966EAD9EAE18319F85FCE466">238</mods:start>
<mods:end id="3F4CBBB4FC945F9D65FAA450ACA19469">246</mods:end>
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<mods:classification id="BE4D97D2584DBD4777E45BAEF11DFC5E">journal article</mods:classification>
<mods:identifier id="366BDD54822FDC7E374DFDA6B377957E" type="CLB-Dataset">128622</mods:identifier>
<mods:identifier id="13EBC9DEB7702EB7495941336D84D666" type="DOI">10.11646/phytotaxa.559.3.2</mods:identifier>
<mods:identifier id="C149AE0A016872079577F73AA7ACE94B" type="GBIF-Dataset">066c00f0-d6bf-4344-8829-7b709fffe392</mods:identifier>
<mods:identifier id="3E19B362A19A0ED9FE8F9A760A9C5630" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="4833A9F73837F8C0ABF0BBE98EF07420" type="Zenodo-Dep">7021706</mods:identifier>
</mods:mods>
<treatment id="344487E9FFBE7364A8C13FDAFEB3FDF8" ID-DOI="http://doi.org/10.5281/zenodo.7021694" ID-GBIF-Taxon="199818190" ID-Zenodo-Dep="7021694" LSID="urn:lsid:plazi:treatment:344487E9FFBE7364A8C13FDAFEB3FDF8" httpUri="http://treatment.plazi.org/id/344487E9FFBE7364A8C13FDAFEB3FDF8" lastPageId="5" lastPageNumber="243" pageId="4" pageNumber="242">
<subSubSection id="F4F76574FFBE7365A8C13FDAFBD2F92F" box="[136,1117,1756,1783]" pageId="4" pageNumber="242" type="nomenclature">
<paragraph id="BC5236FFFFBE7365A8C13FDAFBD2F92F" blockId="4.[136,1117,1756,1783]" box="[136,1117,1756,1783]" pageId="4" pageNumber="242">
<heading id="E71A8193FFBE7365A8C13FDAFBD2F92F" box="[136,1117,1756,1783]" fontSize="11" level="2" pageId="4" pageNumber="242" reason="3">
<taxonomicName id="7BED4D7CFFBE7365A8C13FDAFC2FF92F" ID-CoL="B2QS7" authority="Spetik, Eichmeier, Gramaje &amp; Berraf-Tebbal" authorityName="Spetik, Eichmeier, Gramaje &amp; Berraf-Tebbal" authorityYear="2022" box="[136,928,1756,1783]" class="Eurotiomycetes" family="Aspergillaceae" genus="Paecilomyces" kingdom="Fungi" order="Eurotiales" pageId="4" pageNumber="242" phylum="Ascomycota" rank="species" species="clematidis" status="sp. nov.">
<emphasis id="8E99EAEDFFBE7365A8C13FDAFE2FF92E" bold="true" box="[136,416,1756,1783]" italics="true" pageId="4" pageNumber="242">Paecilomyces clematidis</emphasis>
Spetik, Eichmeier, Gramaje &amp; Berraf-Tebbal
</taxonomicName>
,
<taxonomicNameLabel id="95AA5796FFBE7365ABE33FDBFB8EF92E" box="[938,1025,1757,1783]" pageId="4" pageNumber="242" rank="species">
<emphasis id="8E99EAEDFFBE7365ABE33FDBFC76F92F" box="[938,1017,1757,1782]" italics="true" pageId="4" pageNumber="242">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="24D62A7AFFBE7365AC583FDAFBDBF92F" box="[1041,1108,1756,1782]" captionStart="FIGURE 2" captionStartId="5.[136,229,1995,2017]" captionTargetBox="[230,1365,815,1953]" captionTargetId="figure-278@5.[215,1372,802,1967]" captionTargetPageId="5" captionText="FIGURE 2. Paecilomyces clematidis (ex-type CBS 148466). A, F–G. Chlamydospores. C. Conidiophores with phialides. B, D. Phialides forming conidia. E. Conidia. H. Colony on PDA. I. Colony on MEA. J. Colony on OA. Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.7021710" httpUri="https://zenodo.org/record/7021710/files/figure.png" pageId="4" pageNumber="242">
<emphasis id="8E99EAEDFFBE7365AC583FDAFBDBF92F" bold="true" box="[1041,1108,1756,1782]" pageId="4" pageNumber="242">Fig. 2</emphasis>
</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F4F76574FFBE7365A8C13E06FE64F8C2" box="[136,491,1792,1819]" pageId="4" pageNumber="242" type="multiple">
<paragraph id="BC5236FFFFBE7365A8C13E06FE64F8C2" blockId="4.[136,1451,1792,1890]" box="[136,491,1792,1819]" pageId="4" pageNumber="242">Mycobank number: MB843540</paragraph>
</subSubSection>
<subSubSection id="F4F76574FFBE7365A8C13E2EFA17F8BB" pageId="4" pageNumber="242" type="materials_examined">
<paragraph id="BC5236FFFFBE7365A8C13E2EFA17F8BB" blockId="4.[136,1451,1792,1890]" pageId="4" pageNumber="242">
Type:
<materialsCitation id="0C853CA2FFBE7365A88D3E2EFA1AF8BB" ID-GBIF-Occurrence="3897120302" collectingDate="2021-05" collectionCode="BRNU" collectorName="M. Spetik" country="Czech Republic" latitude="48.792583" location="Lednice" longLatPrecision="1" longitude="16.798805" municipality="Breclav" pageId="4" pageNumber="242" specimenCode="BRNU 677844" specimenCount="1" typeStatus="holotype">
<collectingCountry id="C4FA766FFFBE7365A88D3E2EFE00F8E4" box="[196,399,1832,1853]" name="Czech Republic" pageId="4" pageNumber="242">CZECH REPUBLIC</collectingCountry>
,
<collectingMunicipality id="5C36AC85FFBE7365A9DE3E2EFE6AF8E7" box="[407,485,1832,1854]" pageId="4" pageNumber="242">Breclav</collectingMunicipality>
:
<location id="B9326024FFBE7365A9A73E2EFDB1F8E7" LSID="urn:lsid:plazi:treatment:344487E9FFBE7364A8C13FDAFEB3FDF8:B9326024FFBE7365A9A73E2EFDB1F8E7" box="[494,574,1832,1854]" country="Czech Republic" latitude="48.792583" longLatPrecision="1" longitude="16.798805" municipality="Breclav" name="Lednice" pageId="4" pageNumber="242">Lednice</location>
, university garden (
<geoCoordinate id="D9D95038FFBE7365AAB33E2EFC0BF8E4" box="[762,900,1832,1853]" degrees="48" direction="north" minutes="47" orientation="latitude" pageId="4" pageNumber="242" precision="1" seconds="33.3" value="48.792583">48°47’33.3”N</geoCoordinate>
,
<geoCoordinate id="D9D95038FFBE7365ABC73E2EFB99F8E7" box="[910,1046,1832,1854]" degrees="16" direction="east" minutes="47" orientation="longitude" pageId="4" pageNumber="242" precision="1" seconds="55.7" value="16.798805">16°47’55.7”E</geoCoordinate>
), isolated from the root of
<taxonomicName id="7BED4D7CFFBE7365AD5B3E2EFAE8F8E4" authority="Snow" authorityName="Snow" box="[1298,1383,1832,1853]" class="Magnoliopsida" family="Ranunculaceae" genus="Clematis" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="242" phylum="Tracheophyta" rank="genus">
<emphasis id="8E99EAEDFFBE7365AD5B3E2EFAE8F8E4" box="[1298,1383,1832,1853]" italics="true" pageId="4" pageNumber="242">Clematis</emphasis>
</taxonomicName>
‘Snow Queen’ (
<taxonomicName id="7BED4D7CFFBE7365A9593E4AFE2BF8B8" box="[272,420,1868,1889]" class="Magnoliopsida" family="Ranunculaceae" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="242" phylum="Tracheophyta" rank="family">
<emphasis id="8E99EAEDFFBE7365A9593E4AFE2BF8B8" box="[272,420,1868,1889]" italics="true" pageId="4" pageNumber="242">Ranunculaceae</emphasis>
</taxonomicName>
),
<date id="C853103FFFBE7365A9F13E4AFD93F8B8" box="[440,540,1868,1889]" pageId="4" pageNumber="242" value="2021-05">
<collectingDate id="D817E9D7FFBE7365A9F13E4AFD93F8B8" box="[440,540,1868,1889]" pageId="4" pageNumber="242" value="2021-05">May 2021</collectingDate>
</date>
,
<collectorName id="11185329FFBE7365AA6F3E4AFD08F8BB" box="[550,647,1868,1890]" pageId="4" pageNumber="242">M. Spetik</collectorName>
,
<typeStatus id="6356885DFFBE7365AAD83E4AFD61F8BB" box="[657,750,1868,1890]" pageId="4" pageNumber="242" type="holotype">Holotype</typeStatus>
:
<specimenCode id="EC4B9E84FFBE7365AABE3E4AFC05F8BB" box="[759,906,1868,1890]" pageId="4" pageNumber="242">BRNU 677844</specimenCode>
, ex-type living culture CBS 148466 = MEND-F-0560
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F4F76574FFBE7364A8C13E96FAFEFE49" lastPageId="5" lastPageNumber="243" pageId="4" pageNumber="242" type="description">
<paragraph id="BC5236FFFFBE7364A8C13E96FAFEFE49" blockId="4.[136,1452,1936,2071]" lastBlockId="5.[136,1452,158,545]" lastPageId="5" lastPageNumber="243" pageId="4" pageNumber="242">
<emphasis id="8E99EAEDFFBE7365A8C13E96FEB8F873" bold="true" box="[136,311,1936,1962]" pageId="4" pageNumber="242">Sexual morph:</emphasis>
not observed.
<emphasis id="8E99EAEDFFBE7365A9903E96FD18F873" bold="true" box="[473,663,1936,1962]" pageId="4" pageNumber="242">Asexual morph:</emphasis>
<emphasis id="8E99EAEDFFBE7365AAD43E96FCDFF873" box="[669,848,1936,1962]" italics="true" pageId="4" pageNumber="242">Hyphomycetous</emphasis>
.
<emphasis id="8E99EAEDFFBE7365AB143E96FB8DF873" box="[861,1026,1936,1962]" italics="true" pageId="4" pageNumber="242">Conidiophores</emphasis>
hyaline, septate, irregularly branched.
<emphasis id="8E99EAEDFFBE7365A8C13EB2FF7DF817" box="[136,242,1972,1998]" italics="true" pageId="4" pageNumber="242">Phialides</emphasis>
with a cylindrical basal portion tapering to a thin long neck, solitary directly on vegetative hyphae or in groups of two–four per metula.
<emphasis id="8E99EAEDFFBE7365A9BE3EDEFDDEF82B" box="[503,593,2008,2034]" italics="true" pageId="4" pageNumber="242">Conidia</emphasis>
(2.5–)3.3–3.6(–4.2) × (1.7–)2.2–2.5(–3.2) µm, mean ± SD 3.4 ± 0.4 × 2.4 ± 0.4 µm hyaline, ellipsoidal with flattened base, pyriform, produced in chains.
<emphasis id="8E99EAEDFFBE7365AC6D3EFAFB50F7CF" box="[1060,1247,2044,2070]" italics="true" pageId="4" pageNumber="242">Chlamydospores</emphasis>
usually produced laterally or terminally on short side branches of superficial or immersed hyphae, thick-walled, initially smooth later becoming rough and warted, subglobose to globose, dark green, 4.5–6.2 µm diam.
<emphasis id="8E99EAEDFFBF7364AC7139C4FACCFF05" box="[1080,1347,194,220]" italics="true" pageId="5" pageNumber="243">Culture characteristics.</emphasis>
On PDA after seven days: white to olivaceous, floccose, aerial mycelium, reverse olivaceous, almost no growth at 18 °C;&gt;
<quantity id="7B159B1AFFBF7364A8C1380CFF57FEFC" box="[136,216,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="243" unit="mm" value="90.0">90 mm</quantity>
diam at 25°, 30° and 35 °C;
<quantity id="7B159B1AFFBF7364AA6F380CFDF8FEFC" box="[550,631,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="243" unit="mm" value="42.0">42 mm</quantity>
diam at 40 °C; no growth at 45 °C. On MEA after seven days: white to olivaceous, floccose, aerial mycelium, reverse olivaceous,&gt;
<quantity id="7B159B1AFFBF7364AB7C3828FC0BFE90" box="[821,900,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="243" unit="mm" value="90.0">90 mm</quantity>
diam at 25° and 30 °C. On OA after seven days: white to olivaceous, floccose, aerial mycelium, reverse olivaceous,&gt;
<quantity id="7B159B1AFFBF7364ABEA3854FC7BFEB4" box="[931,1012,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="243" unit="mm" value="90.0">90 mm</quantity>
diam at 25° and 30 °C. On CYA after seven days: white to olivaceous, floccose, aerial mycelium, reverse olivaceous,&gt;
<quantity id="7B159B1AFFBF7364AC553870FBE4FE48" box="[1052,1131,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="243" unit="mm" value="90.0">90 mm</quantity>
diam at 25° and 30 °C.
</paragraph>
</subSubSection>
<subSubSection id="F4F76574FFBF7364A8F4389CFD63FE6D" box="[189,748,410,437]" pageId="5" pageNumber="243" type="etymology">
<paragraph id="BC5236FFFFBF7364A8F4389CFD63FE6D" blockId="5.[136,1452,158,545]" box="[189,748,410,437]" pageId="5" pageNumber="243">
<emphasis id="8E99EAEDFFBF7364A8F4389CFEC7FE6D" bold="true" box="[189,328,410,436]" pageId="5" pageNumber="243">Etymology:</emphasis>
—refers to the host genus (
<taxonomicName id="7BED4D7CFFBF7364AA31389CFD53FE6D" box="[632,732,410,436]" class="Magnoliopsida" family="Ranunculaceae" genus="Clematis" kingdom="Plantae" order="Ranunculales" pageId="5" pageNumber="243" phylum="Tracheophyta" rank="genus">
<emphasis id="8E99EAEDFFBF7364AA31389CFD53FE6D" box="[632,732,410,436]" italics="true" pageId="5" pageNumber="243">Clematis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="F4F76574FFBF7364A8F438B8FEB3FDF8" pageId="5" pageNumber="243" type="materials_examined">
<paragraph id="BC5236FFFFBF7364A8F438B8FEB3FDF8" blockId="5.[136,1452,158,545]" pageId="5" pageNumber="243">
<emphasis id="8E99EAEDFFBF7364A8F438B8FDCAFE01" bold="true" box="[189,581,446,472]" pageId="5" pageNumber="243">Additional specimens examined:</emphasis>
<materialsCitation id="0C853CA2FFBF7364AA0738B8FEB7FDF8" ID-GBIF-Occurrence="3897120301" collectingDate="2021-05" collectorName="M. Spetik" country="Czech Republic" latitude="48.79261" location="Lednice" longLatPrecision="1" longitude="16.798805" municipality="Breclav" pageId="5" pageNumber="243" specimenCount="1">
<emphasis id="8E99EAEDFFBF7364AA0738B8FCC3FE01" bold="true" box="[590,844,446,472]" pageId="5" pageNumber="243">
<collectingCountry id="C4FA766FFFBF7364AA0738B8FCC3FE01" box="[590,844,446,472]" name="Czech Republic" pageId="5" pageNumber="243">CZECH REPUBLIC</collectingCountry>
</emphasis>
,
<collectingMunicipality id="5C36AC85FFBF7364AB1538B8FC38FE01" box="[860,951,446,472]" pageId="5" pageNumber="243">Breclav</collectingMunicipality>
:
<location id="B9326024FFBF7364AB8D38B8FBAEFE01" LSID="urn:lsid:plazi:treatment:344487E9FFBE7364A8C13FDAFEB3FDF8:B9326024FFBF7364AB8D38B8FBAEFE01" box="[964,1057,446,472]" country="Czech Republic" latitude="48.79261" longLatPrecision="1" longitude="16.798805" municipality="Breclav" name="Lednice" pageId="5" pageNumber="243">Lednice</location>
, university garden (
<geoCoordinate id="D9D95038FFBF7364AD4C38B8FA29FE01" box="[1285,1446,446,472]" degrees="48" direction="north" minutes="47" orientation="latitude" pageId="5" pageNumber="243" precision="1" seconds="33.4" value="48.79261">48°47’33.4”N</geoCoordinate>
,
<geoCoordinate id="D9D95038FFBF7364A8C138E4FEA9FE24" box="[136,294,482,509]" degrees="16" direction="east" minutes="47" orientation="longitude" pageId="5" pageNumber="243" precision="1" seconds="55.7" value="16.798805">16°47’55.7”E</geoCoordinate>
), isolated from the root of
<emphasis id="8E99EAEDFFBF7364AA0138E4FD35FE24" box="[584,698,482,509]" italics="true" pageId="5" pageNumber="243">
<taxonomicName id="7BED4D7CFFBF7364AA0138E4FD23FE25" authority="Snow" authorityName="Snow" box="[584,684,482,508]" class="Magnoliopsida" family="Ranunculaceae" genus="Clematis" kingdom="Plantae" order="Ranunculales" pageId="5" pageNumber="243" phylum="Tracheophyta" rank="genus">Clematis</taxonomicName>
‘
</emphasis>
Snow Queen’ (
<taxonomicName id="7BED4D7CFFBF7364AB1738E4FB84FE25" box="[862,1035,482,508]" class="Magnoliopsida" family="Ranunculaceae" kingdom="Plantae" order="Ranunculales" pageId="5" pageNumber="243" phylum="Tracheophyta" rank="family">
<emphasis id="8E99EAEDFFBF7364AB1738E4FB84FE25" box="[862,1035,482,508]" italics="true" pageId="5" pageNumber="243">Ranunculaceae</emphasis>
</taxonomicName>
),
<date id="C853103FFFBF7364AC6838E5FB1BFE25" box="[1057,1172,482,509]" pageId="5" pageNumber="243" value="2021-05">
<collectingDate id="D817E9D7FFBF7364AC6838E5FB1BFE25" box="[1057,1172,482,509]" pageId="5" pageNumber="243" value="2021-05">May 2021</collectingDate>
</date>
,
<collectorName id="11185329FFBF7364ACD738E5FA82FE25" box="[1182,1293,482,509]" pageId="5" pageNumber="243">M. Spetik</collectorName>
, living culture MEND-F-0561
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

81
data/34/44/F9/3444F95557EC563B9B8D439F962229CD.xml Normal file
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@ -0,0 +1,81 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1184.112437" ID-Pensoft-Pub="1313-2970-1184-81" ID-Pensoft-UUID="8C1CA9892B8F565DA5BE8D16D85FAC48" ID-ZooBank="654C20B58303463E86B8ACA39BA34EA6" ModsDocID="1313-2970-1184-81" checkinTime="1699992646542" checkinUser="pensoft" docAuthor="Packova, Gabriela &amp; Kundrata, Robin" docDate="2023" docId="3444F95557EC563B9B8D439F962229CD" docLanguage="en" docName="ZooKeys 1184: 81-89" docOrigin="ZooKeys 1184" docPubDate="2023-11-14" docSource="http://dx.doi.org/10.3897/zookeys.1184.112437" docTitle="Rhagophthalmus Motschulsky 1854" docType="treatment" docVersion="1" id="8C1CA9892B8F565DA5BE8D16D85FAC48" lastPageNumber="81" masterDocId="8C1CA9892B8F565DA5BE8D16D85FAC48" masterDocTitle="A new species of Rhagophthalmus Motschulsky, 1854 (Coleoptera, Rhagophthalmidae) from Laos represents the smallest known member of the genus" masterLastPageNumber="89" masterPageNumber="81" pageNumber="81" updateTime="1699992646542" updateUser="pensoft">
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<mods:titleInfo id="A14E869321BB2A70B0B5E370952DCD55">
<mods:title id="AF0C3A7F8ED8F700A79AB446D1C22B39">A new species of Rhagophthalmus Motschulsky, 1854 (Coleoptera, Rhagophthalmidae) from Laos represents the smallest known member of the genus</mods:title>
</mods:titleInfo>
<mods:name id="48D0613D0408748327B1247D32DED289" type="personal">
<mods:role id="053B561EA8C0348B789B5DDD1A82706D">
<mods:roleTerm id="CF814E03595B0C979A3715ABA38449D8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7835352F40F199A1AD19778A7263AE00">Packova, Gabriela</mods:namePart>
<mods:nameIdentifier id="7A4F544C64AF126B70F38F9F1CD7E07F" type="ORCID">https://orcid.org/0000-0001-7949-619X</mods:nameIdentifier>
<mods:affiliation id="FE29FCEC61FBB108844C0563F40389D0">Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 77146 Olomouc, Czech Republic</mods:affiliation>
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<mods:namePart id="DB31605C0AF37BDFAD66E2C8F800AC4A">Kundrata, Robin</mods:namePart>
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<mods:affiliation id="B21633FFA679B248FF1013E25E8CDFDC">Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 77146 Olomouc, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="E747D4CE199B38174886B41CDCC5DAE9" type="email">robin.kundrata@upol.cz</mods:nameIdentifier>
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<mods:date id="DBC66D133BEDF9EE45CF793C23C54850">2023</mods:date>
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<mods:number id="2364825CF99AAF22BE9431C3B1EFA3A4">2023-11-14</mods:number>
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<mods:url id="3F3952FC66D425D9F91A21CDFEA5546D">http://dx.doi.org/10.3897/zookeys.1184.112437</mods:url>
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<treatment id="3444F95557EC563B9B8D439F962229CD" LSID="urn:lsid:plazi:treatment:3444F95557EC563B9B8D439F962229CD" httpUri="http://treatment.plazi.org/id/3444F95557EC563B9B8D439F962229CD" lastPageNumber="81" pageId="0" pageNumber="81">
<subSubSection id="69D5B7D31EFF09B5D862E70C78A4DE6D" pageId="0" pageNumber="81" type="nomenclature">
<paragraph id="ECDA9600A96CD028B68E0C765E2B2A58" pageId="0" pageNumber="81">
Genus
<taxonomicName id="C8CE7A354E369B7A4A36FE86A32E242B" ID-CoL="78BL" ID-ENA="71224" LSID="3444F955-57EC-563B-9B8D-439F962229CD" authority="Motschulsky, 1854" authorityName="Motschulsky" authorityYear="1854" class="Insecta" family="Rhagophthalmidae" genus="Rhagophthalmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhagophthalmus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">Rhagophthalmus Motschulsky, 1854</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C843D3C0FC696FA09298E166DBDF11CA" pageId="0" pageNumber="81" type="reference_group">
<paragraph id="D78A216B6D21AFCD1E2AF3868696FA8F" pageId="0" pageNumber="81">
<taxonomicName id="437616DA60B0911B285A95F01A85C4D1" authorityName="Motschulsky" authorityYear="1854" class="Insecta" family="Rhagophthalmidae" genus="Rhagophthalmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhagophthalmus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">Rhagophthalmus</taxonomicName>
Motschulsky, 1854: 45. Type species:
<taxonomicName id="AF96293A53E442BA0E796BE677032CBF" authorityName="Motschulsky" authorityYear="1854" class="Insecta" family="Rhagophthalmidae" genus="Rhagophthalmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhagophthalmus scutellatus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="scutellatus">Rhagophthalmus scutellatus</taxonomicName>
Motschulsky, 1854.
</paragraph>
<paragraph id="3E7F78FB9C87A1C58A66B4D43ED0015D" pageId="0" pageNumber="81">
<taxonomicName id="37C6429718EF0D54BE9FEAACC359CA6A" authorityName="Pascoe" authorityYear="1862" class="Insecta" family="Rhagophthalmidae" genus="Ochotyra" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ochotyra" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">Ochotyra</taxonomicName>
=
<taxonomicName id="7313577456B0A704B550145B2A1B4DDB" authorityName="Pascoe" authorityYear="1862" class="Insecta" family="Rhagophthalmidae" genus="Ochotyra" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ochotyra" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">Ochotyra</taxonomicName>
Pascoe, 1862: 323. Type species:
<taxonomicName id="2605EAAE3DF88E358B081130CFB0434B" class="Insecta" family="Rhagophthalmidae" genus="Ochotyra" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ochotyra semiusta" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="semiusta">Ochotyra semiusta</taxonomicName>
Pascoe, 1862: 323 (Wittmer in
<bibRefCitation id="005E3834455146B724C71253BA1CB4C7" author="Wittmer, W" journalOrPublisher="Japanese Journal of Entomology" pageId="0" pageNumber="81" pagination="341 - 355" refId="B19" refString="Wittmer, W, Ohba, N, 1994. Neue Rhagophthalmidae (Coleoptera) aus China und benachbarten Laendern. Japanese Journal of Entomology 62: 341 - 355" title="Neue Rhagophthalmidae (Coleoptera) aus China und benachbarten Laendern." volume="62" year="1994">Wittmer and Ohba 1994</bibRefCitation>
: 342).*
</paragraph>
</subSubSection>
<subSubSection id="83EB90507B490E65949957956881E429" pageId="0" pageNumber="81" type="diagnosis">
<paragraph id="DFC8EEDBABD1065F6152521221A4B6DB" pageId="0" pageNumber="81">Diagnosis.</paragraph>
<paragraph id="E396A1C214405EA6249BDBAEFE9D6144" pageId="0" pageNumber="81">This genus can be currently characterized by deeply emarginate eyes, medium-sized mandibles, and relatively short antennae. Its limits and relationships with morphologically similar genera need to be critically revised.</paragraph>
</subSubSection>
</treatment>
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