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<mods:title id="80676413701685A8F506EE0940F2D432">Ameisen des Herrn Prof. v. Ihering aus Brasilien (Sao Paulo usw.) nebst einigen anderen aus Südamerika und Afrika (Hym.).</mods:title>
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<taxonomicNameLabel id="F646EA09CE2077C62C84B7F36EF33C2B" rank="species">n. sp.</taxonomicNameLabel>
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[[worker]]. L. 4,7-5 mm. Kiefer mit 5 starken
<normalizedToken id="E4BE6B3C4615A17463238A32ACEAF5E1" originalValue="Zähnen">Zaehnen</normalizedToken>
,
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, zerstreut, grob punktiert, mit einigen stumpfen Streifen und Spuren einer feinen Runzelung. Kopf gerundet rechteckig, um 1/4
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als breit, hinten etwas
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als vorn, aber mit deutlichem, schwach konvexem Hinterrand, ohne erhabenen Gelenkrand, mit konvexen Seiten (also wie bei Foreli und
<taxonomicName id="DC2C27D7E029688F88D613096723BA34" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:30561" class="Insecta" family="Formicidae" genus="Megalomyrmex" kingdom="Animalia" lsidName-HNS="Megalomyrmex modestus Emery" order="Hymenoptera" pageNumber="304" phylum="Arthropoda" rank="species" species="modestus">modestus Em.</taxonomicName>
geformt). Clypeus mit konvexem Vorderrand, hinten recht schmal zwischen den Stirnleisten; diese mit einem kleinen, relativ horizontalen Vorderlappen. Augen nur wenig vor der Mitte,
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, konvex (klein bei
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). Der
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<normalizedToken id="D4ABCC836C1D09A5320CD5BC0D9B0EE2" originalValue="überragt">ueberragt</normalizedToken>
den Hinterrand um mehr als 1/3 seiner
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.
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und sechstes
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<normalizedToken id="38CFBEEE1545FF253DCC9C9858189B5E" originalValue="ungefähr">ungefaehr</normalizedToken>
so dick als
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; die anderen etwas
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. Keule, scharf dreigliedrig. Neuntes
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so lang wie das zehnte und doppelt so lang wie das achte. Erstes
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fast doppelt so lang wie das zweite. Thorax wie bei
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und
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, mit
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, unbewehrter
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des Epinotums; aber die
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ist tiefer und ziemlich schmal, und die
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des Epinotums steiler,
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bogig in die
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<normalizedToken id="5772AF66309D5529E5C58847C7B0C69C" originalValue="übergehend">uebergehend</normalizedToken>
. Unten, am Episternum, 2 kleine erhabene Lamellen. Stielchen genau wie bei
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. Beide Knoten unten ohne
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(der erste mit einem Zahn bei
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).
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Ganz glatt und sehr
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, mit zerstreuten haartragenden Punkten. Abstehende Behaarung gelblich, spitz, ziemlich kurz,
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verteilt, an den Schienen und am
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etwas schief. Anliegende Behaarung fast null.
<normalizedToken id="B5A9DC3C4BBA3568F1D207BC041431BD" originalValue="Gleichmäßig">Gleichmaessig</normalizedToken>
gelblich rot.
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<subSubSection id="36D49940EA7466555E1BC8C8F0223B6B" type="materials_examined">
<paragraph id="C803CD0412F882CA8A17AEFB6F59F221" pageNumber="305">
Bahnhof Alto da Serra (v. Ihering); Park
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bei Alto da Serra (
<normalizedToken id="CBAA2C7AEAAD8D2C660BA31A062038CA" originalValue="Lüderwaldt">Luederwaldt</normalizedToken>
), Est. Sao Paulo, Brasilien.
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<subSubSection id="12E5583CC5050C645F2DA02BAD0EDE51" type="discussion">
<paragraph id="6A262679351312A7BC581D0609AA4D70" pageNumber="305">
Dem
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am
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, aber dunkler und lebhafter
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,
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, und durch die Augen und die
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besonders scharf zu unterscheiden, auch durch den
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Hinterteil des Clypeus.
</paragraph>
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</treatment>
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<taxonomicName id="7BB14D101D40FFB59071E582FB155C6D" box="[1032,1217,1515,1539]" class="Magnoliopsida" family="Hamamelidaceae" genus="Bucklandia" kingdom="Plantae" order="Saxifragales" pageId="12" pageNumber="37" phylum="Tracheophyta" rank="species" species="undefined-A">Bucklandia sp. A</taxonomicName>
.
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</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D40FFB5978DE665FB0D5C4A" box="[1012,1241,1548,1572]" pageId="12" pageNumber="37" type="description">
<paragraph id="BC0E36931D40FFB5978DE665FB0D5C4A" blockId="12.[831,1422,1349,1572]" box="[1012,1241,1548,1572]" pageId="12" pageNumber="37">
<figureCitation id="248A2A161D40FFB5978DE665FBE05C4A" box="[1012,1076,1548,1572]" captionStart="Fig" captionStartId="13.[166,201,1948,1969]" captionTargetBox="[166,1443,149,1939]" captionTargetId="figure-19@13.[166,1443,149,1939]" captionTargetPageId="13" captionText="Fig. 6 Buckladia sp. A. - inventory no. 27719 (field no. 1107). a-f: Cross section - gymnosperm structure with tracheids rectangular with rounded corners, thin curled walls in the early-wood, thick walled in the late-wood; idioblasts between tracheids; d-e: Numerous secondary rays; g-i: Tangential section - unpitted tracheids; uniseriate rays." figureDoi="http://doi.org/10.5281/zenodo.10520863" httpUri="https://zenodo.org/record/10520863/files/figure.png" pageId="12" pageNumber="37">Fig. 6</figureCitation>
, a-i;
<figureCitation id="248A2A161D40FFB59013E665FB7E5C4A" box="[1130,1194,1548,1572]" captionStart="Fig" captionStartId="14.[167,202,1919,1940]" captionTargetBox="[167,1421,179,1910]" captionTargetId="figure-38@14.[167,1421,179,1910]" captionTargetPageId="14" captionText="Fig. 7 Buckladia sp. A - inventory no. 27719 (field no. 1107). a-i: Radial section - d-g: - scalariform thickenings of Zamia type on tracheids, with reticular aspect; g-i: - araucarioid cross-fields with alternate in 2-3 superposed rows." figureDoi="http://doi.org/10.5281/zenodo.10520865" httpUri="https://zenodo.org/record/10520865/files/figure.png" pageId="12" pageNumber="37">Fig. 7</figureCitation>
, a-i.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D40FFB59749E626FCAF5D43" pageId="12" pageNumber="37" type="materials_examined">
<paragraph id="BC0E36931D40FFB59749E626FC475C09" blockId="12.[816,1436,1615,1837]" box="[816,915,1615,1639]" pageId="12" pageNumber="37">
<emphasis id="8EC5EA811D40FFB59749E626FC475C09" bold="true" box="[816,915,1615,1639]" pageId="12" pageNumber="37">Material</emphasis>
</paragraph>
<paragraph id="BC0E36931D40FFB59749E606FCAF5D43" blockId="12.[816,1436,1615,1837]" pageId="12" pageNumber="37">
<materialsCitation id="0CD93CCE1D40FFB59749E606FCA25D43" location="National Museum of Geology" municipality="Holbav area" pageId="12" pageNumber="37" specimenCount="1" stateProvince="Bucuresti">
The studied material is represented by a piece of silicified wood collected from
<collectingMunicipality id="5C6AACE91D40FFB5906CE6F9FB4D5CC6" box="[1045,1177,1680,1704]" pageId="12" pageNumber="37">Holbav area</collectingMunicipality>
, on Maiului brook. This specimen has the field-number 1107, in &quot;Grădinaru Collection&quot;, and is stored now at the
<location id="B96E60481D40FFB590CFE6BBFC585D62" LSID="urn:lsid:plazi:treatment:341887851D40FFA99071E582FBF55AB9:B96E60481D40FFB590CFE6BBFC585D62" municipality="Holbav area" name="National Museum of Geology" pageId="12" pageNumber="37" stateProvince="Bucuresti">National Museum of Geology</location>
in
<collectingRegion id="7E75F8711D40FFB597B1E69DFBE15D62" box="[968,1077,1780,1804]" country="Romania" name="Bucuresti" pageId="12" pageNumber="37">Bucharest</collectingRegion>
, under the inventory number 27719
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D40FFB69749E73EFF2B5CC6" lastPageId="15" lastPageNumber="40" pageId="12" pageNumber="37" type="description">
<paragraph id="BC0E36931D40FFB59749E73EFB955D01" blockId="12.[816,1437,1879,2068]" box="[816,1089,1879,1903]" pageId="12" pageNumber="37">
<emphasis id="8EC5EA811D40FFB59749E73EFB955D01" bold="true" box="[816,1089,1879,1903]" pageId="12" pageNumber="37">Microscopic description</emphasis>
</paragraph>
<paragraph id="BC0E36931D40FFB69749E710FDFE5B12" blockId="12.[816,1437,1879,2068]" lastBlockId="15.[151,771,158,1208]" lastPageId="15" lastPageNumber="40" pageId="12" pageNumber="37">
In cross-section, the structure of the wood is of gymnospermous
<typeStatus id="630A88311D40FFB597E6E7F3FC045DDC" box="[927,976,1946,1970]" pageId="12" pageNumber="37">type</typeStatus>
, and some details (such as the presence of the typical idioblasts and axial mucilaginous canals) suggest a cycadeoid polyxylic stem structure, which is also supported by the presence of scalariform thickenings of
<taxonomicName id="7BB14D101D43FFB694EEE0F6FF0F5AD8" authorityName="Kutorga" authorityYear="1842" box="[151,219,159,182]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D43FFB694EEE0F6FF0F5AD8" box="[151,219,159,182]" italics="true" pageId="15" pageNumber="40">Zamia</emphasis>
</taxonomicName>
<typeStatus id="630A88311D43FFB6949EE0F7FEC15AD8" box="[231,277,158,182]" pageId="15" pageNumber="40">type</typeStatus>
on the tracheids in vertical view (
<bibRefCitation id="D8204B621D43FFB696DAE0F7FF045AB9" author="Greguss" pageId="15" pageNumber="40" pagination="1 - 260" refId="ref14798" refString="Greguss, 1968. Xylotomy of the living Cycads, with a description of their leaves and epidermis. 1 - 260 pages, Akademiai Kiado." type="book chapter" year="1968">Greguss, 1968</bibRefCitation>
), so, the xylotomic description will be adequate:
<emphasis id="8EC5EA811D43FFB694EEE089FF175A96" box="[151,195,224,248]" italics="true" pageId="15" pageNumber="40">Pith</emphasis>
unfortunately, is absent in our fragmentary samples which does not retain the axial part of the stem, so we did not have the possibility to describe its component elements (as parenchyma cells, idioblasts, medullary vascular bundles and mucilaginous canals).
</paragraph>
<caption id="E8CE661B1D41FFB494DFE7F5FEB45264" ID-DOI="http://doi.org/10.5281/zenodo.10520863" ID-Zenodo-Dep="10520863" httpUri="https://zenodo.org/record/10520863/files/figure.png" pageId="13" pageNumber="38" startId="13.[166,201,1948,1969]" targetBox="[166,1443,149,1939]" targetPageId="13" targetType="figure">
<paragraph id="BC0E36931D41FFB494DFE7F5FD2F5DDF" blockId="13.[166,1444,1947,2058]" box="[166,763,1947,1969]" pageId="13" pageNumber="38">
<emphasis id="8EC5EA811D41FFB494DFE7F5FF0B5DDF" bold="true" box="[166,223,1948,1969]" pageId="13" pageNumber="38">Fig. 6</emphasis>
<emphasis id="8EC5EA811D41FFB4949FE7F2FE505DDF" box="[230,388,1947,1969]" italics="true" pageId="13" pageNumber="38">Buckladia sp. A.</emphasis>
- inventory no. 27719 (field no. 1107).
</paragraph>
<paragraph id="BC0E36931D41FFB494DFE7D0FEB45264" blockId="13.[166,1444,1947,2058]" pageId="13" pageNumber="38">
<emphasis id="8EC5EA811D41FFB494DFE7D0FF1D5DA0" bold="true" box="[166,201,1977,1998]" pageId="13" pageNumber="38">a-f:</emphasis>
Cross section - gymnosperm structure with tracheids rectangular with rounded corners, thin curled walls in the early-wood, thick walled in the late-wood; idioblasts between tracheids;
<emphasis id="8EC5EA811D41FFB49690E7BEFCDD5D82" bold="true" box="[745,777,2007,2028]" pageId="13" pageNumber="38">d-e</emphasis>
: Numerous secondary rays;
<emphasis id="8EC5EA811D41FFB49067E7BEFB945D82" bold="true" box="[1054,1088,2007,2028]" pageId="13" pageNumber="38">g-i:</emphasis>
Tangential section - unpitted tracheids; uniseriate rays.
</paragraph>
</caption>
<caption id="E8CE661B1D42FFB794DEE716FDC15DBE" ID-DOI="http://doi.org/10.5281/zenodo.10520865" ID-Zenodo-Dep="10520865" httpUri="https://zenodo.org/record/10520865/files/figure.png" pageId="14" pageNumber="39" startId="14.[167,202,1919,1940]" targetBox="[167,1421,179,1910]" targetPageId="14" targetType="figure">
<paragraph id="BC0E36931D42FFB794DEE716FD225DFB" blockId="14.[167,1421,1919,2001]" box="[167,758,1919,1941]" pageId="14" pageNumber="39">
<emphasis id="8EC5EA811D42FFB794DEE716FF345DFA" bold="true" box="[167,224,1919,1940]" pageId="14" pageNumber="39">Fig. 7</emphasis>
<emphasis id="8EC5EA811D42FFB7949FE716FEAA5DFB" box="[230,382,1919,1941]" italics="true" pageId="14" pageNumber="39">Buckladia sp. A</emphasis>
- inventory no. 27719 (field no. 1107).
</paragraph>
<paragraph id="BC0E36931D42FFB794DEE7F4FDC15DBE" blockId="14.[167,1421,1919,2001]" pageId="14" pageNumber="39">
<emphasis id="8EC5EA811D42FFB794DEE7F4FF1E5DDC" bold="true" box="[167,202,1949,1970]" pageId="14" pageNumber="39">a-i:</emphasis>
Radial section -
<emphasis id="8EC5EA811D42FFB79517E7F4FE4C5DDC" bold="true" box="[366,408,1949,1970]" pageId="14" pageNumber="39">d-g:</emphasis>
- scalariform thickenings of
<taxonomicName id="7BB14D101D42FFB796D6E7F4FD385DDC" authorityName="Kutorga" authorityYear="1842" box="[687,748,1949,1970]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" kingdom="Plantae" order="Cycadales" pageId="14" pageNumber="39" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D42FFB796D6E7F4FD385DDC" box="[687,748,1949,1970]" italics="true" pageId="14" pageNumber="39">Zamia</emphasis>
</taxonomicName>
type on tracheids, with reticular aspect;
<emphasis id="8EC5EA811D42FFB7900CE7F4FB4C5DDC" bold="true" box="[1141,1176,1949,1970]" pageId="14" pageNumber="39">g-i:</emphasis>
- araucarioid cross-fields with alternate in 2-3 superposed rows.
</paragraph>
</caption>
<paragraph id="BC0E36931D43FFB694EEE1ECFE9C5ED6" blockId="15.[151,771,158,1208]" pageId="15" pageNumber="40">
<emphasis id="8EC5EA811D43FFB694EEE1ECFE1C5BF3" box="[151,456,389,413]" italics="true" pageId="15" pageNumber="40">Cylindrical vascular system</emphasis>
in cross section, is represented by some alternative growth rings of secondary xylem and phloem (not as annual rings) and numerous thin secondary rays, so giving a polyxylic structure (
<figureCitation id="248A2A161D43FFB696AEE180FF66584C" captionStart="Fig" captionStartId="13.[166,201,1948,1969]" captionTargetBox="[166,1443,149,1939]" captionTargetId="figure-19@13.[166,1443,149,1939]" captionTargetPageId="13" captionText="Fig. 6 Buckladia sp. A. - inventory no. 27719 (field no. 1107). a-f: Cross section - gymnosperm structure with tracheids rectangular with rounded corners, thin curled walls in the early-wood, thick walled in the late-wood; idioblasts between tracheids; d-e: Numerous secondary rays; g-i: Tangential section - unpitted tracheids; uniseriate rays." figureDoi="http://doi.org/10.5281/zenodo.10520863" httpUri="https://zenodo.org/record/10520863/files/figure.png" pageId="15" pageNumber="40">Fig. 6a</figureCitation>
), crossed by wide multiseriate primary medullary rays which connect the axial pith with the cortex (but this aspect does not appear in our sections). The cambial cells are recognized with difficulty, since they are very similar to the neighboring cells. The xylem has rectangular or square tracheids, in gradual transition from early- to the late-wood, with r/tg diameter of 25-65 / 10-15-30 μm, with relatively thick walls, of 4-8 μm the double wall, (
<figureCitation id="248A2A161D43FFB694E6E37AFF0B5945" box="[159,223,787,811]" captionStart="Fig" captionStartId="13.[166,201,1948,1969]" captionTargetBox="[166,1443,149,1939]" captionTargetId="figure-19@13.[166,1443,149,1939]" captionTargetPageId="13" captionText="Fig. 6 Buckladia sp. A. - inventory no. 27719 (field no. 1107). a-f: Cross section - gymnosperm structure with tracheids rectangular with rounded corners, thin curled walls in the early-wood, thick walled in the late-wood; idioblasts between tracheids; d-e: Numerous secondary rays; g-i: Tangential section - unpitted tracheids; uniseriate rays." figureDoi="http://doi.org/10.5281/zenodo.10520863" httpUri="https://zenodo.org/record/10520863/files/figure.png" pageId="15" pageNumber="40">Fig. 6</figureCitation>
a-f). Their walls are often corrugated, by compression. On the radial walls, the tracheids show uni- to biseriate bordered pits, rounded to hexagonal, contiguous and flattened, sometimes alternate, when biseriate (
<figureCitation id="248A2A161D43FFB696D3E31FFD2A59E1" box="[682,766,886,911]" captionStart="Fig" captionStartId="14.[167,202,1919,1940]" captionTargetBox="[167,1421,179,1910]" captionTargetId="figure-38@14.[167,1421,179,1910]" captionTargetPageId="14" captionText="Fig. 7 Buckladia sp. A - inventory no. 27719 (field no. 1107). a-i: Radial section - d-g: - scalariform thickenings of Zamia type on tracheids, with reticular aspect; g-i: - araucarioid cross-fields with alternate in 2-3 superposed rows." figureDoi="http://doi.org/10.5281/zenodo.10520865" httpUri="https://zenodo.org/record/10520865/files/figure.png" pageId="15" pageNumber="40">
Fig.
<quantity id="7B499B761D43FFB6969BE31EFD2A59E1" box="[738,766,887,911]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="15" pageNumber="40" unit="g" value="7.0">7g</quantity>
</figureCitation>
, arrow). The pits have diameters of 9-12 /8-15 μm, with round to elliptical apertures. Very obvious are the scalariform thickenings of
<taxonomicName id="7BB14D101D43FFB6950BE3B3FE62599F" authorityName="Kutorga" authorityYear="1842" box="[370,438,986,1009]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D43FFB6950BE3B3FE62599F" box="[370,438,986,1009]" italics="true" pageId="15" pageNumber="40">Zamia</emphasis>
</taxonomicName>
<typeStatus id="630A88311D43FFB695C4E3B3FE39599C" box="[445,493,986,1010]" pageId="15" pageNumber="40">type</typeStatus>
, which are often connected by inclined bars, giving a reticular appearance (
<figureCitation id="248A2A161D43FFB696AEE393FF705E5D" captionStart="Fig" captionStartId="14.[167,202,1919,1940]" captionTargetBox="[167,1421,179,1910]" captionTargetId="figure-38@14.[167,1421,179,1910]" captionTargetPageId="14" captionText="Fig. 7 Buckladia sp. A - inventory no. 27719 (field no. 1107). a-i: Radial section - d-g: - scalariform thickenings of Zamia type on tracheids, with reticular aspect; g-i: - araucarioid cross-fields with alternate in 2-3 superposed rows." figureDoi="http://doi.org/10.5281/zenodo.10520865" httpUri="https://zenodo.org/record/10520865/files/figure.png" pageId="15" pageNumber="40">Fig. 7</figureCitation>
a-e). No perforations were observed. The tangential walls are not pitted (
<figureCitation id="248A2A161D43FFB695FAE455FE1C5E3A" box="[387,456,1084,1108]" captionStart="Fig" captionStartId="13.[166,201,1948,1969]" captionTargetBox="[166,1443,149,1939]" captionTargetId="figure-19@13.[166,1443,149,1939]" captionTargetPageId="13" captionText="Fig. 6 Buckladia sp. A. - inventory no. 27719 (field no. 1107). a-f: Cross section - gymnosperm structure with tracheids rectangular with rounded corners, thin curled walls in the early-wood, thick walled in the late-wood; idioblasts between tracheids; d-e: Numerous secondary rays; g-i: Tangential section - unpitted tracheids; uniseriate rays." figureDoi="http://doi.org/10.5281/zenodo.10520863" httpUri="https://zenodo.org/record/10520863/files/figure.png" pageId="15" pageNumber="40">Fig. 6</figureCitation>
g-i). The phloem has parenchymal cells, (unclear if they are sieve cells), thick-walled fibres, secretory idioblasts, and sclerenchymal cells (
<figureCitation id="248A2A161D43FFB694AFE4C9FEC25ED6" box="[214,278,1184,1208]" captionStart="Fig" captionStartId="13.[166,201,1948,1969]" captionTargetBox="[166,1443,149,1939]" captionTargetId="figure-19@13.[166,1443,149,1939]" captionTargetPageId="13" captionText="Fig. 6 Buckladia sp. A. - inventory no. 27719 (field no. 1107). a-f: Cross section - gymnosperm structure with tracheids rectangular with rounded corners, thin curled walls in the early-wood, thick walled in the late-wood; idioblasts between tracheids; d-e: Numerous secondary rays; g-i: Tangential section - unpitted tracheids; uniseriate rays." figureDoi="http://doi.org/10.5281/zenodo.10520863" httpUri="https://zenodo.org/record/10520863/files/figure.png" pageId="15" pageNumber="40">Fig. 6</figureCitation>
d-e).
</paragraph>
<paragraph id="BC0E36931D43FFB694EEE48AFE7F5C2B" blockId="15.[151,771,1250,1704]" pageId="15" pageNumber="40">
<emphasis id="8EC5EA811D43FFB694EEE48AFF1F5E94" box="[151,203,1251,1274]" italics="true" pageId="15" pageNumber="40">Rays</emphasis>
in transversal and tangential sections, the primary rays were not observed in our fragmentary material, but numerous secondary rays appear, giving a manoxylic aspect to the structure. They are uniseriate, sometimes 2(- 3)-seriate, of low to mid tall, and are composed of parenchyma cells and few secretory idioblasts, so they are heterogeneous. The ray cells are rounded to rectangular in outline, the idioblasts are similar or are slightly elongate and, in radial view, appear all procumbent. In cross-fields they have 2-6(9) oculipores usually in two superposed rows arranged (
<figureCitation id="248A2A161D43FFB69547E644FEA95C2B" box="[318,381,1581,1605]" captionStart="Fig" captionStartId="14.[167,202,1919,1940]" captionTargetBox="[167,1421,179,1910]" captionTargetId="figure-38@14.[167,1421,179,1910]" captionTargetPageId="14" captionText="Fig. 7 Buckladia sp. A - inventory no. 27719 (field no. 1107). a-i: Radial section - d-g: - scalariform thickenings of Zamia type on tracheids, with reticular aspect; g-i: - araucarioid cross-fields with alternate in 2-3 superposed rows." figureDoi="http://doi.org/10.5281/zenodo.10520865" httpUri="https://zenodo.org/record/10520865/files/figure.png" pageId="15" pageNumber="40">Fig. 7</figureCitation>
d-i).
</paragraph>
<paragraph id="BC0E36931D43FFB694EEE626FF2B5CC6" blockId="15.[151,771,1250,1704]" pageId="15" pageNumber="40">
<emphasis id="8EC5EA811D43FFB694EEE626FF0B5C08" box="[151,223,1615,1638]" italics="true" pageId="15" pageNumber="40">Cortex</emphasis>
is not present in our sample, so parenchymal cells, idioblasts, mucilage canals and leaf traces, were not observed.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D43FFA994EEE6BAFBF55AB9" lastPageId="16" lastPageNumber="41" pageId="15" pageNumber="40" type="discussion">
<paragraph id="BC0E36931D43FFB694EEE6BAFE775C85" blockId="15.[151,771,1747,2069]" box="[151,419,1747,1771]" pageId="15" pageNumber="40">
<emphasis id="8EC5EA811D43FFB694EEE6BAFE775C85" bold="true" box="[151,419,1747,1771]" pageId="15" pageNumber="40">Afinities and discussion</emphasis>
</paragraph>
<paragraph id="BC0E36931D43FFB694EEE69DFE6D5DDF" blockId="15.[151,771,1747,2069]" pageId="15" pageNumber="40">
Evaluating the general image of the wood structure in cross section, we noticed some details that indicated a possible cycadeoid stem-structure of polyxylic
<typeStatus id="630A88311D43FFB696D1E75EFD0D5D21" box="[680,729,1847,1871]" pageId="15" pageNumber="40">type</typeStatus>
, by the presence of idioblasts and mucilaginous canals. Thus, we searched for more xylotomical details in order to identify the unknown structure.
</paragraph>
<paragraph id="BC0E36931D43FFB694EEE7D3FB475F33" blockId="15.[151,771,1747,2069]" lastBlockId="15.[816,1437,158,2035]" pageId="15" pageNumber="40">
As is known, the Cycadeoidophyta is a Mesozoic extinct taxon, which include two families: Cycadeoidaceae Buck. (or
<taxonomicName id="7BB14D101D43FFB694C6E795FE74527A" authority="Engl." authorityName="Engl." box="[191,416,2044,2068]" class="Cycadopsida" family="Bennettitaceae" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="family">Bennettitaceae Engl.</taxonomicName>
) with the genera
<taxonomicName id="7BB14D101D43FFB69601E795FCA45AD8" authority="Buck." authorityName="Buckland" authorityYear="1828" box="[632,880,158,2068]" class="Cycadopsida" family="Cycadeoideaceae" genus="Cycadeoidea" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D43FFB69601E795FCD7527A" box="[632,771,2044,2068]" italics="true" pageId="15" pageNumber="40">Cycadeoidea</emphasis>
Buck.
</taxonomicName>
(or
<taxonomicName id="7BB14D101D43FFB697DDE0F6FB5A5AD8" authority="Caruthers" authorityName="Caruthers" box="[932,1166,158,182]" class="Cycadopsida" family="Cycadeoideaceae" genus="Bennettites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D43FFB697DDE0F6FBCE5AD8" box="[932,1050,159,182]" italics="true" pageId="15" pageNumber="40">Bennettites</emphasis>
Caruthers
</taxonomicName>
) and
<taxonomicName id="7BB14D101D43FFB690ABE0F7FCB55AB9" authority="Wieland" authorityName="Wieland" class="Cycadopsida" family="Cycadeoideaceae" genus="Monanthesia" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D43FFB690ABE0F7FA895AD8" box="[1234,1373,158,182]" italics="true" pageId="15" pageNumber="40">Monanthesia</emphasis>
Wieland
</taxonomicName>
; they have a barrel-shaped trunk, like a large pineapple, with a crown of leaves; the second family,
<taxonomicName id="7BB14D101D43FFB69145E089FB485B77" authority="(Carruthers) Nathorst" authorityName="Nathorst" authorityYear="1913" baseAuthorityName="Carruthers" baseAuthorityYear="1870" class="Cycadopsida" family="Williamsoniaceae" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="family">Williamsoniaceae (Carruthers) Nathorst</taxonomicName>
- with
<taxonomicName id="7BB14D101D43FFB69176E168FC4C5B54" authority="Caruthers" authorityName="Caruthers" class="Insecta" family="Corduliidae" genus="Williamsonia" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="8EC5EA811D43FFB69176E168FA495B77" box="[1295,1437,257,281]" italics="true" pageId="15" pageNumber="40">Williamsonia</emphasis>
Caruthers
</taxonomicName>
as
<typeStatus id="630A88311D43FFB697BDE14AFC265B55" box="[964,1010,291,315]" pageId="15" pageNumber="40">type</typeStatus>
genus, a
<taxonomicName id="7BB14D101D43FFB6901DE14BFB065B54" box="[1124,1234,290,314]" form="with" pageId="15" pageNumber="40" rank="form">form with</taxonomicName>
a columnar trunk with frond like leaves at branch tips (see
<bibRefCitation id="D8204B621D43FFB6916AE12AFC085B12" author="Stockey, R. &amp; Rothwell, G. W." pageId="15" pageNumber="40" pagination="251 - 262" refId="ref17276" refString="Stockey, R. and Rothwell, G. W., 2003. Anatomically Preserved Williamsonia (Williamsoniaceae): Evidence for Bennettitalean Reproduction in the Late Cretaceous of Western North America. International Journal of Plant Sciences 164 (2): 251 - 262. https: // doi. org / 10.1086 / 346166" type="journal article" year="2003">Stockey and Rothwell, 2003</bibRefCitation>
). This family includes a lot of described genera of plant-parts, variably named, which send sometimes to a current Cycadalean genus, as follows:
<emphasis id="8EC5EA811D43FFB69116E1CEFB3F58EB" italics="true" pageId="15" pageNumber="40">
<taxonomicName id="7BB14D101D43FFB69116E1CEFC625B8E" authorityName="H.H.Thomas" authorityYear="1915" class="Cycadopsida" family="Williamsoniellaceae" genus="Williamsoniella" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Williamsoniella</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB697ADE1A1FBAC5B8E" authorityName="A.G.Nathorst" authorityYear="1909" box="[980,1144,456,480]" class="Cycadopsida" family="Williamsoniaceae" genus="Nilssoniopteris" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Nilssoniopteris</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB690EFE1A1FA945B8E" authorityName="A.I.Turutanova-Ketova" authorityYear="1930" box="[1174,1344,456,480]" class="Cycadopsida" family="Williamsoniaceae" genus="Bennettitanthus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Bennettitanthus</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB69124E1A1FC78586F" authorityName="Nathorst" authorityYear="1902" class="Cycadopsida" family="Williamsoniaceae" genus="Cycadocephalus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Cycadocephalus</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB697A8E180FB91586F" authorityName="J.M.Anderson &amp; H.M.Anderson" authorityYear="2003" box="[977,1093,489,513]" class="Cycadopsida" family="Williamsoniaceae" genus="Fredlindia" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Fredlindia</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB69010E180FB2F586F" authorityName="T.Delevoryas &amp; R.C.Hope" authorityYear="1976" box="[1129,1275,489,513]" class="Cycadopsida" family="Williamsoniaceae" genus="Ischnophyton" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Ischnophyton</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB69166E180FA4C586F" authorityName="Daugherty" authorityYear="1941" box="[1311,1432,489,513]" class="Cycadopsida" family="Williamsoniaceae" genus="Lyssoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Lyssoxylon</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB69749E263FC72584C" authorityName="Hollick" authorityYear="1893" box="[816,934,522,546]" class="Cycadopsida" family="Williamsoniaceae" genus="Serenopsis" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Serenopsis</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB697C1E263FBFF584C" authorityName="C.F.W.Braun" authorityYear="1847" box="[952,1067,522,546]" class="Cycadopsida" family="Williamsoniaceae" genus="Weltrichia" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Weltrichia</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB69045E262FB47584C" authorityName="A.T.Brongniart" authorityYear="1828" box="[1084,1171,523,546]" class="Cycadopsida" family="Williamsoniaceae" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB690DDE263FA97584C" authorityName="W.P.Schimper" authorityYear="1870" box="[1188,1347,522,546]" class="Cycadopsida" genus="Anomozamites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Anomozamites</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB6912DE263FCBF582D" authorityName="C.F.W.Braun" authorityYear="1843" class="Cycadopsida" family="Williamsoniaceae" genus="Otozamites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Otozamites</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB69701E242FB78582C" authority=", Ceratozamites" authorityName="Ceratozamites" box="[888,1196,555,579]" class="Pinopsida" family="Podocarpaceae" genus="Podozamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Podozamites, Ceratozamites</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB690C0E245FA8D582D" authorityName="Z.Czier" authorityYear="1996" box="[1209,1369,556,579]" class="Cycadopsida" genus="Banatozamites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Banatozamites</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB6911CE245FB3F58EB" authority=", Wielandella, Crossozamia, Dioonites, Eostangeria, Palaeocycas, Pseudoctenis, Ticoa" authorityName="Wielandella, Crossozamia, Dioonites, Eostangeria, Palaeocycas, Pseudoctenis, Ticoa" family="Cichlidae" genus="Pterophyllum" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="40" phylum="Chordata" rank="genus">Pterophyllum, Wielandella, Crossozamia, Dioonites, Eostangeria, Palaeocycas, Pseudoctenis, Ticoa</taxonomicName>
</emphasis>
, and many others (see
<bibRefCitation id="D8204B621D43FFB697F5E2E7FB8D58C8" author="Taylor, T. N. &amp; Taylor, E. &amp; Krings, M." box="[908,1113,654,678]" pageId="15" pageNumber="40" refId="ref17564" refString="Taylor, T. N., Taylor, E. and Krings, M., 2009. Paleobotany. The Biology and Evolution of Fossil Plants. Edition: 2 nd, 1252 p. Publisher: Academic Press / Elsevier. https: // doi. org / 10.2307 / 1223352" type="book" year="2009">Taylor et al., 2009</bibRefCitation>
;
<bibRefCitation id="D8204B621D43FFB6901CE2E7FA9E58C8" author="Martinez, L. &amp; Artabe, A. &amp; Bodnar, J." box="[1125,1354,654,678]" pageId="15" pageNumber="40" pagination="436 - 458" refId="ref15741" refString="Martinez, L., Artabe, A. and Bodnar, J. 2012. A new Cycad stem from the Cretaceous in Argentina and its phylogenetic relationships with other Cycadales. Botanical Journal of the Linnean Society, 170: 436 - 458." type="journal article" year="2012">Martínez et al., 2012</bibRefCitation>
,
<bibRefCitation id="D8204B621D43FFB6912EE2E7FA4458C8" author="Martinez, L. C. A. &amp; Iglesias, A. &amp; Artabe A. E. &amp; Varela A. N. &amp; Apesteguia, S." box="[1367,1424,654,678]" pageId="15" pageNumber="40" pagination="81 - 94" refId="ref15787" refString="Martinez, L. C. A., Iglesias, A., Artabe A. E., Varela A. N. and Apesteguia, S., 2017. A new Encephalarteae trunk (Cycadales) from the Cretaceous of Patagonia (Mata Amarilla Formation, Austral Basin), Argentina. Cretaceous Research, 72: 81 - 94 http: // dx. doi. org / 10.1016 / j." type="journal article" year="2017">2017</bibRefCitation>
). Numerous fossil remains of
<taxonomicName id="7BB14D101D43FFB6901EE2C6FB2658A9" box="[1127,1266,687,711]" order="Bennettitales" pageId="15" pageNumber="43" rank="order">Bennettitales</taxonomicName>
were described from
<collectingCountry id="C4A676031D43FFB6970CE2B8FC015887" box="[885,981,721,745]" name="Romania" pageId="15" pageNumber="40">Romania</collectingCountry>
. Thus, in the Holbav-Cristian-Vulcan-Codlea area
<bibRefCitation id="D8204B621D43FFB697C1E29BFBA75964" author="Givulescu, R." box="[952,1139,754,778]" pageId="15" pageNumber="40" pagination="17 - 21" refId="ref14639" refString="Givulescu, R., 1991 a. Zamites vachrameevii Doludenko 1969 in dem Unteren Lias, Rumanien. Acta Palaeobotanica, 31 (1 - 2): 17 - 21." type="journal article" year="1991">Givulescu (1991)</bibRefCitation>
cited an Early Jurassic flora (or “Liasic flora”) with the presence of
<emphasis id="8EC5EA811D43FFB6917DE37AFC725922" italics="true" pageId="15" pageNumber="40">
<taxonomicName id="7BB14D101D43FFB6917DE37AFA8F5944" authorityName="A.T.Brongniart" authorityYear="1828" box="[1284,1371,787,810]" class="Cycadopsida" family="Williamsoniaceae" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB69110E37AFC725922" authorityName="A.G.Nathorst" authorityYear="1909" class="Cycadopsida" family="Williamsoniaceae" genus="Nilssoniopteris" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">Nilssoniopteris</taxonomicName>
</emphasis>
and
<taxonomicName id="7BB14D101D43FFB69793E35DFBAF5922" box="[1002,1147,820,844]" family="Cichlidae" genus="Pterophyllum" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="40" phylum="Chordata" rank="genus">
<emphasis id="8EC5EA811D43FFB69793E35DFBAF5922" box="[1002,1147,820,844]" italics="true" pageId="15" pageNumber="40">Pterophyllum</emphasis>
</taxonomicName>
(from
<taxonomicName id="7BB14D101D43FFB690ADE35DFAB55922" box="[1236,1377,820,844]" order="Bennettitales" pageId="15" pageNumber="43" rank="order">Bennettitales</taxonomicName>
) and
<taxonomicName id="7BB14D101D43FFB69749E33CFC6A5903" authorityName="O.Heer" authorityYear="1876" box="[816,958,853,877]" class="Pinopsida" family="Czekanowskiaceae" genus="Phoenicopsis" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D43FFB69749E33CFC6A5903" box="[816,958,853,877]" italics="true" pageId="15" pageNumber="40">Phoenicopsis</emphasis>
</taxonomicName>
(
<taxonomicName id="7BB14D101D43FFB697AEE33CFBBD5903" box="[983,1129,853,877]" class="Ginkgoopsida" family="Ginkgoaceae" kingdom="Plantae" order="Ginkgoales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="family">Ginkgoaceae</taxonomicName>
). In recent decades,
<bibRefCitation id="D8204B621D43FFB69112E33FFBEB59E0" author="Dragastan, O. N. &amp; Popa, M. E." pageId="15" pageNumber="40" pagination="51 - 60" refId="ref14175" refString="Dragastan, O. N. and Popa, M. E., 1997. Early Jurassic Phytostratigraphy of the Holbav Formation, Getic Nappe, Brasov County. Revue Roumaine de Geologie, 41: 51 - 60." type="journal article" year="1997">Dragastan and Popa (1997)</bibRefCitation>
,
<bibRefCitation id="D8204B621D43FFB69029E31EFB0559E0" author="Popa, M. E." box="[1104,1233,886,910]" pageId="15" pageNumber="40" pagination="81 - 87" refId="ref16468" refString="Popa, M. E., 1997. Corystospermal pteridosperms in the Liassic continental deposits of Romania. Acta Palaeontologica Romaniae, 1: 81 - 87." type="journal article" year="1997">Popa (1997</bibRefCitation>
a,
<bibRefCitation id="D8204B621D43FFB69096E31FFAF359E0" author="Popa, M. E." box="[1263,1319,886,910]" pageId="15" pageNumber="40" pagination="177 - 184" refId="ref16497" refString="Popa, M. E., 1998. The Liassic continental flora of Romania: Systematics, Stratigraphy and Paleoecology. Acta Botanica Horti Bucurestensis: 177 - 184." type="book chapter" year="1998">1998</bibRefCitation>
,
<bibRefCitation id="D8204B621D43FFB69141E31FFAAA59E0" author="Popa, M. E." box="[1336,1406,886,910]" pageId="15" pageNumber="40" pagination="377 - 386" refId="ref16528" refString="Popa, M. E., 2000 a. Aspects of Romanian Early Jurassic palaeobotany and palynology. Part III. Phytostratigraphy of the Getic Nappe. Acta Palaeontologica Romaniae, 2: 377 - 386." type="journal article" year="2000">2000a</bibRefCitation>
,b), and Popa and van Konijnenburg-van Cittert (1999, 2006) contributed more to the Early Jurassic flora in the Holbav area, describing
<taxonomicName id="7BB14D101D43FFB69038E3B0FAE1599F" box="[1089,1333,985,1009]" class="Testudines" family="Trionychidae" genus="Nilssonia" isUncertain="true" kingdom="Animalia" order="Cryptodira" pageId="15" pageNumber="40" phylum="Chordata" rank="species" species="undulata">
<emphasis id="8EC5EA811D43FFB69038E3B0FB72599F" box="[1089,1190,985,1009]" italics="true" pageId="15" pageNumber="40">Nilssonia</emphasis>
cf.
<emphasis id="8EC5EA811D43FFB690AEE3B0FAE1599F" box="[1239,1333,985,1009]" italics="true" pageId="15" pageNumber="40">undulata</emphasis>
</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB6913EE3B3FCAA5E7C" class="Cycadopsida" genus="Paracycas" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="8EC5EA811D43FFB6913EE3B3FC805E7C" italics="true" pageId="15" pageNumber="40">Paracycas</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB697F5E392FB855E7C" authorityName="Fontaine" authorityYear="1896" box="[908,1105,1018,1042]" class="Cycadopsida" genus="Ctenis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="species" species="grandifolia">
<emphasis id="8EC5EA811D43FFB697F5E392FB855E7C" box="[908,1105,1018,1042]" italics="true" pageId="15" pageNumber="40">Ctenis grandifolia</emphasis>
</taxonomicName>
(
<taxonomicName id="7BB14D101D43FFB6901BE393FB075E7C" box="[1122,1235,1018,1042]" class="Cycadopsida" kingdom="Plantae" order="Cycadales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="order">Cycadales</taxonomicName>
) and
<emphasis id="8EC5EA811D43FFB6916AE393FADE5E5D" italics="true" pageId="15" pageNumber="40">
<taxonomicName id="7BB14D101D43FFB6916AE393FC535E5D" class="Polypodiopsida" family="Hymenophyllaceae" genus="Ptilophyllum" kingdom="Plantae" order="Hymenophyllales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="species" species="rigidum">Ptilophyllum rigidum</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB697E2E472FB535E5D" box="[923,1159,1051,1075]" family="Cichlidae" genus="Pterophyllum" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="40" phylum="Chordata" rank="species" species="magoti">Pterophyllum magoti</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB690E2E475FADE5E5D" box="[1179,1290,1052,1075]" family="Cichlidae" genus="Pterophyllum" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="40" phylum="Chordata" rank="species" species="neagui">P. neagui</taxonomicName>
</emphasis>
(as
<taxonomicName id="7BB14D101D43FFB69131E475FCA55E3A" order="Bennettitales" pageId="15" pageNumber="43" rank="order">Bennettitales</taxonomicName>
) and beside them -
<taxonomicName id="7BB14D101D43FFB6902BE454FAD55E3A" box="[1106,1281,1085,1108]" class="Polypodiopsida" family="Pteridaceae" genus="Taeniopteris" kingdom="Plantae" order="Polypodiales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="8EC5EA811D43FFB6902BE454FB0D5E3A" box="[1106,1241,1085,1108]" italics="true" pageId="15" pageNumber="40">Taeniopteris</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="7BB14D101D43FFB6916BE455FA455E3A" authorityName="Gorozhankin" authorityYear="1904" box="[1298,1425,1084,1108]" class="Ginkgoopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ginkgoales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="order">Ginkgoales</taxonomicName>
),
<taxonomicName id="7BB14D101D43FFB69749E434FB215E1B" authority="(Czekanowkiales)" baseAuthorityName="Czekanowkiales" box="[816,1269,1117,1141]" class="Pinopsida" family="Czekanowskiaceae" genus="Phoenicopsis" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="species" species="potoniei">
<emphasis id="8EC5EA811D43FFB69749E434FBF25E1B" box="[816,1062,1117,1141]" italics="true" pageId="15" pageNumber="40">Phoenicopsis potoniei</emphasis>
(Czekanowkiales)
</taxonomicName>
and
<taxonomicName id="7BB14D101D43FFB69142E437FC9B5EF8" class="Insecta" family="Geinitziidae" genus="Geinitzia" kingdom="Animalia" order="Grylloblattodea" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8EC5EA811D43FFB69142E437FA485E1B" box="[1339,1436,1118,1141]" italics="true" pageId="15" pageNumber="40">Geinitzia</emphasis>
sp.
</taxonomicName>
(Conifers). Also, from the Cristian area were described
<taxonomicName id="7BB14D101D43FFB697FEE4C9FB575ED6" box="[903,1155,1184,1208]" class="Magnoliopsida" family="Brassicaceae" genus="Pachypteris" kingdom="Plantae" order="Brassicales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="species" species="gradinarui">
<emphasis id="8EC5EA811D43FFB697FEE4C9FB575ED6" box="[903,1155,1184,1208]" italics="true" pageId="15" pageNumber="40">Pachypteris grădinarui</emphasis>
</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB690EAE4C9FA415ED6" box="[1171,1429,1184,1208]" class="Cycadopsida" family="Williamsoniaceae" genus="Otozamites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="species" species="mandelslohi">
<emphasis id="8EC5EA811D43FFB690EAE4C9FA415ED6" box="[1171,1429,1184,1208]" italics="true" pageId="15" pageNumber="40">Otozamites mandelslohi</emphasis>
</taxonomicName>
,
<taxonomicName id="7BB14D101D43FFB69749E4ABFC1F5EB7" box="[816,971,1217,1241]" class="Cycadopsida" family="Williamsoniaceae" genus="Otozamites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="species" species="molinianus">
<emphasis id="8EC5EA811D43FFB69749E4ABFC1F5EB7" box="[816,971,1217,1241]" italics="true" pageId="15" pageNumber="40">O. molinianus</emphasis>
</taxonomicName>
and a
<taxonomicName id="7BB14D101D43FFB69064E4A8FB395EB7" box="[1053,1261,1217,1241]" class="Pinopsida" family="Czekanowskiaceae" genus="Desmiophyllum" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="8EC5EA811D43FFB69064E4A8FB105EB7" box="[1053,1220,1217,1241]" italics="true" pageId="15" pageNumber="40">Desmiophyllum</emphasis>
sp.
</taxonomicName>
which belongs, probably, to Czekanowskiales. Also,
<bibRefCitation id="D8204B621D43FFB690BDE48AFCAC5F75" author="Dragastan, O. N. &amp; Popa, M. E." pageId="15" pageNumber="40" pagination="51 - 60" refId="ref14175" refString="Dragastan, O. N. and Popa, M. E., 1997. Early Jurassic Phytostratigraphy of the Holbav Formation, Getic Nappe, Brasov County. Revue Roumaine de Geologie, 41: 51 - 60." type="journal article" year="1997">Dragastan and Popa (1997)</bibRefCitation>
and
<bibRefCitation id="D8204B621D43FFB697C2E56AFB8C5F75" author="Czier, Z." box="[955,1112,1283,1307]" pageId="15" pageNumber="40" pagination="538 - 573" refId="ref14045" refString="Czier, Z., 2016 b. Probleme de stratigrafie cu privire la depozitele continentale ale Jurasicului inferior din Romania. In: Moisa, G., Goman, I., Sipos, S. (coord.), Omagiu Chiriac Aurel. Fragmentarium, studii interdisciplinare: 538 - 573, Oradea (Editura Muzeului Țarii Crisurilor)." type="book chapter" year="2016">Czier (2016b)</bibRefCitation>
advanced some phytostratigraphic schemes based on the Early Jurassic foliar material identified in the Holbav area.
</paragraph>
<paragraph id="BC0E36931D43FFB69749E50EFC005CA4" blockId="15.[816,1437,158,2035]" pageId="15" pageNumber="40">
However, there are few genera described in the world from the study of fossil stem remains, often with local importance and with names related to the original fossiliferous site or people (see list in
<bibRefCitation id="D8204B621D43FFB690D2E5A3FA4C5F8C" author="Martinez, L. &amp; Artabe, A. &amp; Bodnar, J." box="[1195,1432,1482,1506]" pageId="15" pageNumber="40" pagination="436 - 458" refId="ref15741" refString="Martinez, L., Artabe, A. and Bodnar, J. 2012. A new Cycad stem from the Cretaceous in Argentina and its phylogenetic relationships with other Cycadales. Botanical Journal of the Linnean Society, 170: 436 - 458." type="journal article" year="2012">Martínez et al., 2012</bibRefCitation>
,
<bibRefCitation id="D8204B621D43FFB69749E582FCBD5C6D" author="Martinez, L. C. A. &amp; Iglesias, A. &amp; Artabe A. E. &amp; Varela A. N. &amp; Apesteguia, S." box="[816,873,1515,1539]" pageId="15" pageNumber="40" pagination="81 - 94" refId="ref15787" refString="Martinez, L. C. A., Iglesias, A., Artabe A. E., Varela A. N. and Apesteguia, S., 2017. A new Encephalarteae trunk (Cycadales) from the Cretaceous of Patagonia (Mata Amarilla Formation, Austral Basin), Argentina. Cretaceous Research, 72: 81 - 94 http: // dx. doi. org / 10.1016 / j." type="journal article" year="2017">2017</bibRefCitation>
;
<bibRefCitation id="D8204B621D43FFB69701E582FB9D5C6D" author="Zhang W. &amp; Yang X. - J. &amp; Fu X. - P. &amp; Zheng S. - L. &amp; Wang Y. - D." box="[888,1097,1515,1539]" pageId="15" pageNumber="40" pagination="50 - 60" refId="ref17946" refString="Zhang W., Yang X. - J., Fu X. - P., Zheng S. - L. and Wang Y. - D., 2012. A polyxylic cycad trunk from the Middle Jurassic of western Liaoning, China, and its evolutionary implications. Review of Palaeobotany and Palynology, 183: 50 - 60." type="journal article" year="2012">Zhang et al., 2012</bibRefCitation>
). For example,
<bibRefCitation id="D8204B621D43FFB69080E582FCAE5C4A" author="Martinez, L. C. A. &amp; Iglesias, A. &amp; Artabe A. E. &amp; Varela A. N. &amp; Apesteguia, S." pageId="15" pageNumber="40" pagination="81 - 94" refId="ref15787" refString="Martinez, L. C. A., Iglesias, A., Artabe A. E., Varela A. N. and Apesteguia, S., 2017. A new Encephalarteae trunk (Cycadales) from the Cretaceous of Patagonia (Mata Amarilla Formation, Austral Basin), Argentina. Cretaceous Research, 72: 81 - 94 http: // dx. doi. org / 10.1016 / j." type="journal article" year="2017">Martinez et al. (2017)</bibRefCitation>
, have described a well-preserved Jurassic bennettitalean specimen from Patagonia, as
<emphasis id="8EC5EA811D43FFB690A2E647FA495C2B" box="[1243,1437,1581,1605]" italics="true" pageId="15" pageNumber="40">Zamuneria amyla</emphasis>
Martínez, Iglesias and Artabe, and made a carefully documented comparison with the xylotomy of the extant cycadalean genera and also, with all Mesozoic Bennettitalean genera.
</paragraph>
<paragraph id="BC0E36931D43FFB69749E6BBFBA65D9D" blockId="15.[816,1437,158,2035]" pageId="15" pageNumber="40">
The studied specimens show a cycadeoid polyxylic structure with scalariform thickenings on tracheids which indicate similarities with stem of
<taxonomicName id="7BB14D101D43FFB690F9E77CFB2D5D43" authorityName="Brongniart, Zamuneria Martinez, Iglesias and Artabe" authorityYear="1828" box="[1152,1273,1813,1837]" class="Magnoliopsida" family="Hamamelidaceae" genus="Bucklandia" kingdom="Plantae" order="Saxifragales" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D43FFB690F9E77CFB2D5D43" box="[1152,1273,1813,1837]" italics="true" pageId="15" pageNumber="40">Bucklandia</emphasis>
</taxonomicName>
<typeStatus id="630A88311D43FFB69178E77CFAFB5D43" box="[1281,1327,1813,1837]" pageId="15" pageNumber="40">type</typeStatus>
(see
<bibRefCitation id="D8204B621D43FFB69749E75EFC625D20" author="Bosse, M. N." box="[816,950,1846,1870]" pageId="15" pageNumber="40" pagination="41 - 50" refId="ref13235" refString="Bosse, M. N. 1953. Bucklandia sahnii sp. nov. from the Jurassic of the Rajmahal Hills, Bihar. Palaeobotanist, 2: 41 - 50." type="journal article" year="1953">Bosse, 1953</bibRefCitation>
;
<bibRefCitation id="D8204B621D43FFB697B8E75FFB8C5D20" author="Sharma, B. D." box="[961,1112,1846,1870]" pageId="15" pageNumber="40" pagination="35 - 46" refId="ref17086" refString="Sharma, B. D. 1967. Investigations on the Jurassic flora of Rajmahal Hills, India. 4. On a new species of Indi- an Bucklandia, B. guptai, with remarks on B. sahnii of Bose. Ameghiniana, 4: 35 - 46." type="journal article" year="1967">Sharma, 1967</bibRefCitation>
,
<bibRefCitation id="D8204B621D43FFB6901BE75FFB4E5D20" author="Sharma, B. D." box="[1122,1178,1846,1870]" pageId="15" pageNumber="40" pagination="303 - 308" refId="ref17137" refString="Sharma, B. D. 1969. Bucklandia dichotoma sp. nov. from the Middle Jurassic of Rajmahal Hills, India. Ameghiniana, 6: 303 - 308." type="journal article" year="1969">1969</bibRefCitation>
,
<bibRefCitation id="D8204B621D43FFB690DCE75FFB095D20" author="Sharma, B. D." box="[1189,1245,1846,1870]" pageId="15" pageNumber="40" pagination="43 - 49" refId="ref17169" refString="Sharma, B. D., 1973. Anatomy of petrified rachises collected from the Jurassic of Amarjola in the Rajmahal Hills, India. The Proceedings of the Linnean Society of NSW, 98: 43 - 49." type="journal article" year="1973">1973</bibRefCitation>
,
<bibRefCitation id="D8204B621D43FFB6909EE75FFACB5D20" author="Sharma, B. D." box="[1255,1311,1846,1870]" pageId="15" pageNumber="40" pagination="260 - 265" refId="ref17210" refString="Sharma, B. D. 1991. Indian Williamsoniaceae - an overview. Paleobotanist, 4: 260 - 265" type="journal article" year="1991">1991</bibRefCitation>
;
<bibRefCitation id="D8204B621D43FFB69152E75FFC1A5D01" author="Saiki, K. &amp; Yoshida, Y." pageId="15" pageNumber="40" pagination="326 - 332" refId="ref16888" refString="Saiki, K. and Yoshida, Y., 1999. A New Bennettitalean Trunk with Unilacunar Five-Trace Nodal Structure from the Upper Cretaceous of Hokkaido, Japan. American Journal of Botany, 86 (3): 326 - 332. https: // doi. org / 10.2307 / 2656754; https: // www. jstor. org / stable / 2656754" type="journal article" year="1999">Saiki and Yoshida, 1999</bibRefCitation>
) or even with
<taxonomicName id="7BB14D101D43FFB69017E73EFB265D01" authorityName="M.N.Bose &amp; S.C.D.Sah" authorityYear="1955" box="[1134,1266,1879,1903]" class="Cycadopsida" genus="Sahnioxylon" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="15" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D43FFB69017E73EFB265D01" box="[1134,1266,1879,1903]" italics="true" pageId="15" pageNumber="40">Sahnioxylon</emphasis>
</taxonomicName>
(see in Philippe et al. 1999) - the latter being considered “nomen ilegitimum” (see
<collectionCode id="DAA0AE561D43FFB69793E7F3FBF85DDF" box="[1002,1068,1946,1969]" pageId="15" pageNumber="40">IFPNI</collectionCode>
- www.fossilplants.info/genus.htm?id=
<uuid id="C8170C461D43FFB690B8E7D3FBB65D9D" pageId="15" pageNumber="40">D46FC51A-7AEB-421E-AF52-1A3F90515E43</uuid>
).
</paragraph>
<paragraph id="BC0E36931D5CFFA994EEE0F7FDF55BF3" blockId="16.[151,772,158,2035]" pageId="16" pageNumber="41">Because the structure of our specimen indicates a columnar polyxylic stem, the sample does not preserve the pith from the central part of the stem and also the cortical zone, both of parenchymal tissue, which were likely easily destroyed during the early stages of fossilization. We cannot see typical parenchymal cells, idioblasts, medullary vascular bundles, mucilaginous canals, and transfusion cells or leaf bases, in the cortex.</paragraph>
<paragraph id="BC0E36931D5CFFA994EEE1CEFF285886" blockId="16.[151,772,158,2035]" pageId="16" pageNumber="41">
However, the polyxylic cycadeoid structure with idioblasts and scalariform thickenings on tracheids, that resemble those in the current
<taxonomicName id="7BB14D101D5CFFA9959FE183FDFE586F" authorityName="Kutorga" authorityYear="1842" box="[486,554,490,513]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA9959FE183FDFE586F" box="[486,554,490,513]" italics="true" pageId="16" pageNumber="41">Zamia</emphasis>
</taxonomicName>
,
<taxonomicName id="7BB14D101D5CFFA99639E183FD1D586F" box="[576,713,490,513]" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA99639E183FD1D586F" box="[576,713,490,513]" italics="true" pageId="16" pageNumber="41">Ceratozamia</emphasis>
</taxonomicName>
(see
<bibRefCitation id="D8204B621D5CFFA994EEE263FEE7584C" author="Greguss" box="[151,307,522,546]" pageId="16" pageNumber="41" pagination="1 - 260" refId="ref14798" refString="Greguss, 1968. Xylotomy of the living Cycads, with a description of their leaves and epidermis. 1 - 260 pages, Akademiai Kiado." type="book chapter" year="1968">Greguss, 1968</bibRefCitation>
) and are partially similar to the structure of the Jurassic genera
<taxonomicName id="7BB14D101D5CFFA99507E242FE0F580A" authority="Brongniart, Zamuneria Martinez, Iglesias and Artabe" authorityName="Brongniart, Zamuneria Martinez, Iglesias and Artabe" authorityYear="1828" class="Magnoliopsida" family="Hamamelidaceae" genus="Bucklandia" kingdom="Plantae" order="Saxifragales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA99507E242FE23582D" box="[382,503,555,579]" italics="true" pageId="16" pageNumber="41">Bucklandia</emphasis>
Brongniart,
<emphasis id="8EC5EA811D5CFFA996F7E245FCD7582D" box="[654,771,556,579]" italics="true" pageId="16" pageNumber="41">Zamuneria</emphasis>
Martínez, Iglesias and Artabe
</taxonomicName>
and
<taxonomicName id="7BB14D101D5CFFA9966FE225FD1A580A" authority="Sharma" authorityName="Sharma" box="[534,718,588,612]" class="Cycadopsida" genus="Lioxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA9966FE225FDA7580A" box="[534,627,588,612]" italics="true" pageId="16" pageNumber="41">Lioxylon</emphasis>
Sharma
</taxonomicName>
(see
<bibRefCitation id="D8204B621D5CFFA994EEE204FE5358EB" author="Martinez, L. C. A. &amp; Iglesias, A. &amp; Artabe A. E. &amp; Varela A. N. &amp; Apesteguia, S." box="[151,391,621,645]" pageId="16" pageNumber="41" pagination="81 - 94" refId="ref15787" refString="Martinez, L. C. A., Iglesias, A., Artabe A. E., Varela A. N. and Apesteguia, S., 2017. A new Encephalarteae trunk (Cycadales) from the Cretaceous of Patagonia (Mata Amarilla Formation, Austral Basin), Argentina. Cretaceous Research, 72: 81 - 94 http: // dx. doi. org / 10.1016 / j." type="journal article" year="2017">Martínez et al., 2017</bibRefCitation>
), which are also characterized by polyxylic stem with centripetal and centrifugal vascular systems and by the presence of numerous idioblasts in the structure.
</paragraph>
<paragraph id="BC0E36931D5CFFA994EEE29BFD5B5E94" blockId="16.[151,772,158,2035]" pageId="16" pageNumber="41">
In the studied Mesozoic areas of Romanian Carpathians, many fossil macro-remains have been described and attributed especially to taxa from
<taxonomicName id="7BB14D101D5CFFA99586E35CFD875922" authorityName="A.T.Brongniart" authorityYear="1828" box="[511,595,821,844]" class="Cycadopsida" family="Williamsoniaceae" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA99586E35CFD875922" box="[511,595,821,844]" italics="true" pageId="16" pageNumber="41">Zamites</emphasis>
</taxonomicName>
group:
<emphasis id="8EC5EA811D5CFFA996D6E35DFF2859E0" italics="true" pageId="16" pageNumber="41">
<taxonomicName id="7BB14D101D5CFFA996D6E35DFF3D5903" authorityName="W.P.Schimper" authorityYear="1870" class="Cycadopsida" genus="Anomozamites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">Anomozamites</taxonomicName>
,
<taxonomicName id="7BB14D101D5CFFA9948EE33CFEA55902" authorityName="C.F.W.Braun" authorityYear="1843" box="[247,369,853,876]" class="Cycadopsida" family="Williamsoniaceae" genus="Otozamites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">Otozamites</taxonomicName>
,
<taxonomicName id="7BB14D101D5CFFA99506E33CFD675902" authority=", Ceratozamites" authorityName="Ceratozamites" box="[383,691,853,877]" class="Pinopsida" family="Podocarpaceae" genus="Podozamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">Podozamites, Ceratozamites</taxonomicName>
,
<taxonomicName id="7BB14D101D5CFFA996BBE33FFF2859E0" authorityName="Z.Czier" authorityYear="1996" class="Cycadopsida" genus="Banatozamites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">Banatozamites</taxonomicName>
</emphasis>
(see
<bibRefCitation id="D8204B621D5CFFA9953FE31FFDD559E0" author="Givulescu, R." box="[326,513,886,910]" pageId="16" pageNumber="41" pagination="17 - 21" refId="ref14639" refString="Givulescu, R., 1991 a. Zamites vachrameevii Doludenko 1969 in dem Unteren Lias, Rumanien. Acta Palaeobotanica, 31 (1 - 2): 17 - 21." type="journal article" year="1991">Givulescu, 1991</bibRefCitation>
;
<bibRefCitation id="D8204B621D5CFFA9966CE31EFF0459C1" author="Dragastan, O. N. &amp; Popa, M. E." pageId="16" pageNumber="41" pagination="51 - 60" refId="ref14175" refString="Dragastan, O. N. and Popa, M. E., 1997. Early Jurassic Phytostratigraphy of the Holbav Formation, Getic Nappe, Brasov County. Revue Roumaine de Geologie, 41: 51 - 60." type="journal article" year="1997">Dragastan and Popa, 1997</bibRefCitation>
;
<bibRefCitation id="D8204B621D5CFFA994A5E3F1FE8C59C1" author="Popa, M. E." box="[220,344,919,943]" pageId="16" pageNumber="41" pagination="81 - 87" refId="ref16468" refString="Popa, M. E., 1997. Corystospermal pteridosperms in the Liassic continental deposits of Romania. Acta Palaeontologica Romaniae, 1: 81 - 87." type="journal article" year="1997">Popa, 1997</bibRefCitation>
,
<bibRefCitation id="D8204B621D5CFFA9951CE3FEFE4959C1" author="Popa, M. E." box="[357,413,919,943]" pageId="16" pageNumber="41" pagination="177 - 184" refId="ref16497" refString="Popa, M. E., 1998. The Liassic continental flora of Romania: Systematics, Stratigraphy and Paleoecology. Acta Botanica Horti Bucurestensis: 177 - 184." type="book chapter" year="1998">1998</bibRefCitation>
,
<bibRefCitation id="D8204B621D5CFFA995D0E3FEFE3A59C1" author="Popa, M. E." box="[425,494,919,943]" pageId="16" pageNumber="41" refId="ref16566" refString="Popa, M. E., 2000 b. Early Jurassic land flora of the Getic Nappe. PhD thesis, University of Bucharest, 258 pp, Bucharest." type="book" year="2000">2000b</bibRefCitation>
; Popa and van Konijnenburg-van Cittert, 1999, 2006;
<bibRefCitation id="D8204B621D5CFFA99592E3D1FDBB59BE" author="Czier, Z." box="[491,623,952,976]" pageId="16" pageNumber="41" pagination="43 - 58" refId="ref13784" refString="Czier, Z., 1999. Biostratigraphy of the Lower Jurassic from Romania, based on the macroflora fossil record. Nymphaea, Folia naturae Bihariae, 27: 43 - 58." type="journal article" year="1999">Czier, 1999</bibRefCitation>
,
<bibRefCitation id="D8204B621D5CFFA99606E3D1FD6359BE" author="Czier, Z." box="[639,695,952,976]" pageId="16" pageNumber="41" pagination="27 - 46" refId="ref13849" refString="Czier, Z., 2001. Rezumatul tezei de doctorat &quot; Flora liasica din Crisana si Banat &quot;. (Universitatea Eotvos Lorand - Budapesta, 1994). Nymphaea, Folia naturae Bihariae, 28: 27 - 46." type="journal article" year="2001">2001</bibRefCitation>
,
<bibRefCitation id="D8204B621D5CFFA996BEE3D1FD2A59BE" author="Czier, Z." box="[711,766,952,976]" pageId="16" pageNumber="41" pagination="5 - 26" refId="ref13923" refString="Czier, Z., 2008. Banatozamites remotus Czier sp. nov. (Cycadeoidales) from the Lower Jurassic of Anina, Romania. Nymphaea, Folia naturae Bihariae, 35: 5 - 26." type="journal article" year="2008">2008</bibRefCitation>
,
<bibRefCitation id="D8204B621D5CFFA994EEE3B0FF1B599F" author="Czier, Z." box="[151,207,985,1009]" pageId="16" pageNumber="41" pagination="5 - 26" refId="ref13961" refString="Czier, Z., 2009. Banatozamites calvus Czier sp. nov. and Bucklandia aninaensis Czier sp. nov. from the Lower Jurassic of Anina, Romania. Nymphaea, Folia naturae Bihariae, 36: 5 - 26." type="journal article" year="2009">2009</bibRefCitation>
,
<bibRefCitation id="D8204B621D5CFFA9949AE3B0FEFD599F" author="Czier, Z." box="[227,297,985,1009]" pageId="16" pageNumber="41" pagination="133 - 184" refId="ref14004" refString="Czier, Z., 2016 a. The Bennettitalen genus Pterophyllum in the fossil flora of Romania, and the global phytostratigraphic - paleophytogeographic context. Vestnik TvGU, Seriya Biologiya i Ekologiya, 3: 133 - 184." type="journal article" year="2016">2016a</bibRefCitation>
,c, 2018), and such taxa have also been commonly described in Europe (see
<bibRefCitation id="D8204B621D5CFFA99637E393FF055E5D" author="Barbacka, M. &amp; Bodor, E. &amp; Jarzynka, A. &amp; Kustatscher, E. &amp; Pacyna, G. &amp; Popa, M. &amp; Scanu, G. G. &amp; Thevenard, F. &amp; Ziaja, J." pageId="16" pageNumber="41" pagination="173 - 195" refId="ref12863" refString="Barbacka, M., Bodor, E., Jarzynka, A., Kustatscher, E., Pacyna, G., Popa, M. E, Scanu, G. G., Thevenard, F. and Ziaja, J., 2014. European Jurassic floras: statistics and palaeoenvironmental proxies. Acta Palaeobotanica, 54: 173 - 195." type="journal article" year="2014">Barbacka et al., 2014</bibRefCitation>
). Thus, from the Lower Jurassic of Anina, Banat region,
<collectingCountry id="C4A676031D5CFFA99492E455FE9A5E3A" box="[235,334,1084,1108]" name="Romania" pageId="16" pageNumber="41">Romania</collectingCountry>
,
<bibRefCitation id="D8204B621D5CFFA99523E455FE375E3A" author="Czier, Z." box="[346,483,1084,1108]" pageId="16" pageNumber="41" pagination="5 - 26" refId="ref13961" refString="Czier, Z., 2009. Banatozamites calvus Czier sp. nov. and Bucklandia aninaensis Czier sp. nov. from the Lower Jurassic of Anina, Romania. Nymphaea, Folia naturae Bihariae, 36: 5 - 26." type="journal article" year="2009">Czier (2009)</bibRefCitation>
described two Bennettitalean species:
<taxonomicName id="7BB14D101D5CFFA9955EE437FD805E1B" authority="Czier" authorityName="Czier" box="[295,596,1117,1141]" class="Cycadopsida" genus="Banatozamites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="species" species="calvus">
<emphasis id="8EC5EA811D5CFFA9955EE437FDC45E1B" box="[295,528,1117,1141]" italics="true" pageId="16" pageNumber="41">Banatozamites calvus</emphasis>
Czier
</taxonomicName>
, by the study of leaf impresions and
<taxonomicName id="7BB14D101D5CFFA9950EE416FD655EF9" authority="Czier" authorityName="Czier" box="[375,689,1151,1175]" class="Magnoliopsida" family="Hamamelidaceae" genus="Bucklandia" kingdom="Plantae" order="Saxifragales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="species" species="aninaensis">
<emphasis id="8EC5EA811D5CFFA9950EE416FDBF5EF9" box="[375,619,1151,1175]" italics="true" pageId="16" pageNumber="41">Bucklandia aninaensis</emphasis>
Czier
</taxonomicName>
, by the study of a small stem with petioles, interpreted to belong to the same plant; they have fine longitudinal ridges, and rows of rectangular to rounded-elongated cells.
</paragraph>
<paragraph id="BC0E36931D5CFFA994EEE56AFD7D5C6D" blockId="16.[151,772,158,2035]" pageId="16" pageNumber="41">
The specimen studied here has details that resemble a
<taxonomicName id="7BB14D101D5CFFA994EEE54CFF0F5F52" authorityName="Kutorga" authorityYear="1842" box="[151,219,1317,1340]" class="Cycadopsida" family="Zamiaceae" genus="Zamia" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA994EEE54CFF0F5F52" box="[151,219,1317,1340]" italics="true" pageId="16" pageNumber="41">Zamia</emphasis>
</taxonomicName>
-
<typeStatus id="630A88311D5CFFA9949DE54CFEC65F53" box="[228,274,1317,1341]" pageId="16" pageNumber="41">type</typeStatus>
structure (see above), especially due to presence of scalariform thickenings on tracheids, even with reticular appearance (see
<bibRefCitation id="D8204B621D5CFFA995D3E50FFD9D5F10" author="Greguss" box="[426,585,1382,1406]" pageId="16" pageNumber="41" pagination="1 - 260" refId="ref14798" refString="Greguss, 1968. Xylotomy of the living Cycads, with a description of their leaves and epidermis. 1 - 260 pages, Akademiai Kiado." type="book chapter" year="1968">Greguss, 1968</bibRefCitation>
). Also due to the presence of idioblasts and mucillaginous canals, we assign it to
<taxonomicName id="7BB14D101D5CFFA9957CE5C0FE145FAF" box="[261,448,1449,1473]" class="Magnoliopsida" family="Hamamelidaceae" genus="Bucklandia" kingdom="Plantae" order="Saxifragales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="species" species="undefined-A">
<emphasis id="8EC5EA811D5CFFA9957CE5C0FE145FAF" box="[261,448,1449,1473]" italics="true" pageId="16" pageNumber="41">Bucklandia sp. A</emphasis>
</taxonomicName>
, probably representing wood of an extinct
<taxonomicName id="7BB14D101D5CFFA9954AE5A3FE735F8C" box="[307,423,1482,1506]" form="from" pageId="16" pageNumber="41" rank="form">form from</taxonomicName>
the
<taxonomicName id="7BB14D101D5CFFA99599E5A2FDE05F8C" authorityName="A.T.Brongniart" authorityYear="1828" box="[480,564,1483,1506]" class="Cycadopsida" family="Williamsoniaceae" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA99599E5A2FDE05F8C" box="[480,564,1483,1506]" italics="true" pageId="16" pageNumber="41">Zamites</emphasis>
</taxonomicName>
group, commonly present in the Romanian Carpathians (see above).
</paragraph>
<paragraph id="BC0E36931D5CFFA994EEE665FE615D43" blockId="16.[151,772,158,2035]" pageId="16" pageNumber="41">
Regarding the genus
<taxonomicName id="7BB14D101D5CFFA995FFE665FE2B5C4A" authorityName="Brongniart, Zamuneria Martinez, Iglesias and Artabe" authorityYear="1828" box="[390,511,1548,1572]" class="Magnoliopsida" family="Hamamelidaceae" genus="Bucklandia" kingdom="Plantae" order="Saxifragales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA995FFE665FE2B5C4A" box="[390,511,1548,1572]" italics="true" pageId="16" pageNumber="41">Bucklandia</emphasis>
</taxonomicName>
, erected by
<bibRefCitation id="D8204B621D5CFFA996E9E665FF355C2B" author="Brongniart, A." pageId="16" pageNumber="41" refId="ref13310" refString="Brongniart, A., 1828. Prodrome d'une histoire des vegetaux fossils, pp. 1 - 223; Paris." type="book" year="1828">Brongniart (1828)</bibRefCitation>
, we consider it as a still valid genus, because it was proposed by
<bibRefCitation id="D8204B621D5CFFA99523E627FE2A5C08" author="Doweld, A. B." box="[346,510,1614,1638]" pageId="16" pageNumber="41" pagination="1131 - 1132" refId="ref14216" refString="Doweld, A. B., 2012. (2099) Proposal to conserve the name Bucklandia Brongn. against Bucklandia Sternb. and Conites (fossil Gymnospermae, Cycadeoideopsida). Taxon, 61 (5): 1131 - 1132" type="journal article" year="2012">Doweld (2012)</bibRefCitation>
to be conserved. However,
<bibRefCitation id="D8204B621D5CFFA994ADE606FDD25CE9" author="Herbst, R. &amp; Crisafulli, A." box="[212,518,1647,1671]" pageId="16" pageNumber="41" pagination="78 - 102" refId="ref14905" refString="Herbst, R. and Crisafulli, A., 2016. La flora triasica del grupo El Tranquilo, provincia de Santa Cruz (Patagonia): lenos fosiles. Ameghiniana, 48 (3): 78 - 102." type="journal article" year="2016">Herbst and Crisafulli (2016)</bibRefCitation>
ignoring this proposal, described a new genus,
<emphasis id="8EC5EA811D5CFFA995D8E6F9FE3B5CC6" box="[417,495,1680,1704]" italics="true" pageId="16" pageNumber="41">Buckya</emphasis>
Herbst and Crisafulli, to replace the old genus
<taxonomicName id="7BB14D101D5CFFA995DCE6D8FD705CA7" authority="Brongniart" authorityName="Brongniart, Zamuneria Martinez, Iglesias and Artabe" authorityYear="1828" box="[421,676,1713,1737]" class="Magnoliopsida" family="Hamamelidaceae" genus="Bucklandia" kingdom="Plantae" order="Saxifragales" pageId="16" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA995DCE6D8FDCA5CA7" box="[421,542,1713,1737]" italics="true" pageId="16" pageNumber="41">Bucklandia</emphasis>
Brongniart
</taxonomicName>
, recommending transfer of all the species previously described to the new proposed genus, as nov. comb. (see
<bibRefCitation id="D8204B621D5CFFA996F7E69DFE955D43" author="Herbst, R. &amp; Crisafulli, A." pageId="16" pageNumber="41" pagination="78 - 102" refId="ref14905" refString="Herbst, R. and Crisafulli, A., 2016. La flora triasica del grupo El Tranquilo, provincia de Santa Cruz (Patagonia): lenos fosiles. Ameghiniana, 48 (3): 78 - 102." type="journal article" year="2016">Herbst and Crisafulli, 2016</bibRefCitation>
, p. 97-98).
</paragraph>
<paragraph id="BC0E36931D5CFFA994EEE75EFBF55AB9" blockId="16.[151,772,158,2035]" lastBlockId="16.[816,1437,158,215]" pageId="16" pageNumber="41">
But, we observed that there is another possible problem of this name,
<emphasis id="8EC5EA811D5CFFA9954DE73EFE565D01" box="[308,386,1879,1903]" italics="true" pageId="16" pageNumber="41">Buckya</emphasis>
, which could become illegitimate, since there are two quasi-homonyms:
<taxonomicName id="7BB14D101D5CFFA99627E711FF1B5DDF" authority="Cobos, 1956" authorityName="Cobos" authorityYear="1956" class="Bryopsida" family="Pylaisiaceae" genus="Buckia" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Hypnales" pageId="16" pageNumber="41" phylum="Bryophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA99627E711FD7C5DFE" box="[606,680,1912,1936]" italics="true" pageId="16" pageNumber="41">Buckia</emphasis>
<bibRefCitation id="D8204B621D5CFFA996C0E711FF1B5DDF" author="Cobos, A. S." pageId="16" pageNumber="41" pagination="1 - 16" refId="ref13508" refString="Cobos, A. S., 1956. Nuevos generos y especies de Eucnemidae (Insecta, Coleoptera) de la fauna neotropical. Bulletin del'Institut Royal des Sciences Naturelles de Belgique, 32 (no. 41): 1 - 16." type="journal article" year="1956">Cobos, 1956</bibRefCitation>
</taxonomicName>
, which is the name of a current insect from the Order
<taxonomicName id="7BB14D101D5CFFA994BBE7D3FEE35DBC" box="[194,311,1978,2002]" class="Bryopsida" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Coleoptera" pageId="16" pageNumber="41" phylum="Bryophyta" rank="order">Coleoptera</taxonomicName>
<collectionCode id="DAA0AE561D5CFFA99546E7D2FE805DBC" box="[319,340,1979,2002]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="16" pageNumber="41" type="Herbarium">L</collectionCode>
., and
<taxonomicName id="7BB14D101D5CFFA995EDE7D3FD2A5DBC" authority="Rios, Gallego and Guerra" authorityName="Rios, Gallego and Guerra" box="[404,766,1978,2002]" class="Bryopsida" family="Pylaisiaceae" genus="Buckia" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Hypnales" pageId="16" pageNumber="41" phylum="Bryophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA995EDE7D3FE0A5DBC" box="[404,478,1978,2002]" italics="true" pageId="16" pageNumber="41">Buckia</emphasis>
Rios, Gallego and Guerra
</taxonomicName>
, (see in GBIF-online), but is also used to describe a moss, a homotypic synonym for
<taxonomicName id="7BB14D101D5CFFA99025E0F6FADD5AD8" authority="Hedw." authorityName="Hedw." box="[1116,1289,158,182]" class="Bryopsida" family="Hypnaceae" genus="Hypnum" kingdom="Plantae" order="Hypnales" pageId="16" pageNumber="41" phylum="Bryophyta" rank="genus">
<emphasis id="8EC5EA811D5CFFA99025E0F6FB625AD8" box="[1116,1206,159,182]" italics="true" pageId="16" pageNumber="41">Hypnum</emphasis>
Hedw.
</taxonomicName>
(Order
<taxonomicName id="7BB14D101D5CFFA9911FE0F7FBC15AB9" authority="Buck. and Vitt." authorityName="Buck. and Vitt." class="Bryopsida" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Hypnales" pageId="16" pageNumber="41" phylum="Bryophyta" rank="order">Hypnales Buck. and Vitt.</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,190 @@
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<paragraph id="BC0E36931D44FFB1972BE35DFB695903" blockId="8.[850,1402,820,910]" pageId="8" pageNumber="33">
<taxonomicName id="7BB14D101D44FFB1972BE35DFB695903" authority="Iamandei, Iamandei and Gradinaru, 2018" authorityName="Iamandei, Iamandei and Gradinaru" authorityYear="2018" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="33" phylum="Tracheophyta" rank="species" species="cristianicum">
<emphasis id="8EC5EA811D44FFB1972BE35DFBA65922" box="[850,1138,820,844]" italics="true" pageId="8" pageNumber="33">Brachyoxylon cristianicum</emphasis>
<bibRefCitation id="D8204B621D44FFB19000E35DFB695903" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." pageId="8" pageNumber="33" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei, Iamandei and Grădinaru, 2018</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D44FFB19056E31FFB4A59E0" box="[1071,1182,886,910]" pageId="8" pageNumber="33" type="description">
<paragraph id="BC0E36931D44FFB19056E31FFB4A59E0" blockId="8.[850,1402,820,910]" box="[1071,1182,886,910]" pageId="8" pageNumber="33">
<figureCitation id="248A2A161D44FFB19056E31FFBBB59E0" box="[1071,1135,886,910]" captionStart="Fig" captionStartId="9.[178,213,1935,1956]" captionTargetBox="[178,1433,149,1926]" captionTargetId="figure-19@9.[178,1433,149,1926]" captionTargetPageId="9" captionText="Fig. 4 Brachyoxylon cristianicum Iamandei, Iamandei and Grădinaru, 2018 - inventory no. 27690(field no. 1113b). a-c: Cross section - thick-walled tracheids, polygonal, with rounded corners and curled walls; dispersed parenchyma cells; linear rays; d-f: Tangential section - uniseriate rays with polygonal ray-cells and biseriate storeys; unpitted tracheids; parenchyma with end-walls smooth or nodular and resin content inside; g-i: Radial section - radial pitting on tracheids of mixed type; cross-fields with 2-3 cupressoid or araucarioid in a single row or, when more numerous, in 2-3 superposed rows." figureDoi="http://doi.org/10.5281/zenodo.10520859" httpUri="https://zenodo.org/record/10520859/files/figure.png" pageId="8" pageNumber="33">Fig. 4</figureCitation>
, a-i.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D44FFB19749E3D1FA8E5F10" pageId="8" pageNumber="33" type="materials_examined">
<paragraph id="BC0E36931D44FFB19749E3D1FC4759BE" blockId="8.[816,1436,952,1406]" box="[816,915,952,976]" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB19749E3D1FC4759BE" bold="true" box="[816,915,952,976]" pageId="8" pageNumber="33">Material</emphasis>
</paragraph>
<paragraph id="BC0E36931D44FFB19749E3B0FA8E5F10" blockId="8.[816,1436,952,1406]" pageId="8" pageNumber="33">
<materialsCitation id="0CD93CCE1D44FFB19749E3B0FA825F10" location="National Museum of Geology" pageId="8" pageNumber="33" specimenCount="1" stateProvince="Bucuresti">
The studied material is represented by 13 samples of silicified wood collected from the Holbav locality, on Maiului brook. All the studied samples are fragments of trunk or thick-branch fragments, with centimetric to decimetric size, dark color and, by hand lens or even naked eye, the regular fibrous structure without vessels is visible, evidence of a conifer wood. The specimens, with the following field numbers: 1071, 1072, 1073, 1075, 1076, 1080, 1081, 1101, 1103, 1108, 1110, 1112, 1113b belong to “Grădinaru Collection” and are stored at the
<location id="B96E60481D44FFB19138E56AFBCC5F52" LSID="urn:lsid:plazi:treatment:341887851D44FFB3972BE35DFB3A5B35:B96E60481D44FFB19138E56AFBCC5F52" name="National Museum of Geology" pageId="8" pageNumber="33" stateProvince="Bucuresti">National Museum of Geology</location>
, in
<collectingRegion id="7E75F8711D44FFB1903EE54DFB605F52" box="[1095,1204,1316,1340]" country="Romania" name="Bucuresti" pageId="8" pageNumber="33">Bucharest</collectingRegion>
, under the inventory numbers 27678, 276789, 27680, 27681, 27682, 27684, 27685, 27686, 27717, 27687, 27688, 27689, 27690
</materialsCitation>
.
</paragraph>
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<subSubSection id="F4AB65181D44FFB39749E5C0FE395EF9" lastPageId="10" lastPageNumber="35" pageId="8" pageNumber="33" type="description">
<paragraph id="BC0E36931D44FFB19749E5C0FB955FAF" blockId="8.[816,1437,1449,2068]" box="[816,1089,1449,1473]" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB19749E5C0FB955FAF" bold="true" box="[816,1089,1449,1473]" pageId="8" pageNumber="33">Microscopic description</emphasis>
</paragraph>
<paragraph id="BC0E36931D44FFB19749E5A3FAD55C2B" blockId="8.[816,1437,1449,2068]" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB19749E5A3FC115F8C" box="[816,965,1482,1506]" italics="true" pageId="8" pageNumber="33">Growth rings</emphasis>
in cross-section show a tracheidoxylic structure without resin canals, usually with abrupt transition from early- to late-wood, and with rather distinct growth-ring boundary. Resin canals absent.
</paragraph>
<paragraph id="BC0E36931D44FFB39749E627FE8A5B77" blockId="8.[816,1437,1449,2068]" lastBlockId="10.[151,772,158,1175]" lastPageId="10" lastPageNumber="35" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB19749E627FC4F5C08" box="[816,923,1614,1638]" italics="true" pageId="8" pageNumber="33">Tracheids</emphasis>
with polygonal (quadrangular) cross-section with rounded corners, unequal in size, diameters of 20- 35-(45-65) μm and with thick walls, of 5-8(9) μm the double wall, usually curled by compression. The late-wood is represented by 2-3 rows of thick-walled and radially flattened cells (radial diameter of 5-10 μm). Intercellular spaces are present. The interradial bundles have 1-10 radial regular rows of tracheids. Density is of 992- 1518 tracheids per mm
<superScript id="4BC49BDB1D44FFB19053E73CFBE75D0D" attach="right" box="[1066,1075,1877,1891]" fontSize="6" pageId="8" pageNumber="33">2</superScript>
. Tangential pitting is absent. Radial pitting is usually uniseriate or biseriate of mixed
<typeStatus id="630A88311D44FFB19749E7F3FCB55DDC" box="[816,865,1946,1970]" pageId="8" pageNumber="33">type</typeStatus>
. When biseriate, the pits appear alternate or opposite and contiguous on short rows often continued by uniseriate portions and do not occupy all the length of the wall. The pits have round to hexagonal borders, diameters of 18-22 μm, and are variably flattened when contiguous (d/D=0.65-0.88), bearing round or tilted elliptic apertures of 7.5-10 / 2-3 μm. Sometimes crassulae or helical thickenings can appear.
</paragraph>
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<paragraph id="BC0E36931D45FFB094CBE7E6FAD05DCA" blockId="9.[178,1434,1934,2075]" box="[178,1284,1934,1956]" pageId="9" pageNumber="34">
<emphasis id="8EC5EA811D45FFB094CBE7E6FF3F5DCA" bold="true" box="[178,235,1935,1956]" pageId="9" pageNumber="34">Fig. 4</emphasis>
<taxonomicName id="7BB14D101D45FFB0948BE7E7FC515DCA" authority="Iamandei, Iamandei and Gradinaru, 2018" authorityName="Iamandei, Iamandei and Gradinaru" authorityYear="2018" box="[242,901,1934,1956]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="34" phylum="Tracheophyta" rank="species" species="cristianicum">
<emphasis id="8EC5EA811D45FFB0948BE7E7FE215DCA" box="[242,501,1934,1956]" italics="true" pageId="9" pageNumber="34">Brachyoxylon cristianicum</emphasis>
<bibRefCitation id="D8204B621D45FFB09582E7E7FC515DCA" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." box="[507,901,1934,1956]" pageId="9" pageNumber="34" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei, Iamandei and Grădinaru, 2018</bibRefCitation>
</taxonomicName>
- inventory no. 27690(field no. 1113b).
</paragraph>
<paragraph id="BC0E36931D45FFB094CBE7C5FBA35275" blockId="9.[178,1434,1934,2075]" pageId="9" pageNumber="34">
<emphasis id="8EC5EA811D45FFB094CBE7C5FF055DAF" bold="true" box="[178,209,1964,1985]" pageId="9" pageNumber="34">a-c</emphasis>
: Cross section - thick-walled tracheids, polygonal, with rounded corners and curled walls; dispersed parenchyma cells; linear rays;
<emphasis id="8EC5EA811D45FFB09491E7A3FED15DB1" bold="true" box="[232,261,1994,2015]" pageId="9" pageNumber="34">d-f</emphasis>
: Tangential section - uniseriate rays with polygonal ray-cells and biseriate storeys; unpitted tracheids; parenchyma with end-walls smooth or nodular and resin content inside;
<emphasis id="8EC5EA811D45FFB096B0E781FD305D93" bold="true" box="[713,740,2024,2045]" pageId="9" pageNumber="34">g-i</emphasis>
: Radial section - radial pitting on tracheids of mixed type; cross-fields with 2-3 cupressoid or araucarioid in a single row or, when more numerous, in 2-3 superposed rows.
</paragraph>
</caption>
<paragraph id="BC0E36931D46FFB394EEE14BFE825B8E" blockId="10.[151,772,158,1175]" pageId="10" pageNumber="35">
<emphasis id="8EC5EA811D46FFB394EEE14BFEB65B54" box="[151,354,290,314]" italics="true" pageId="10" pageNumber="35">Axial parenchyma</emphasis>
in cross-section appear few, dispersed, often difficult to identify among the tracheids. Vertically, sometimes they can show simple pits. The horizontal walls are thin and smooth, or slightly nodular. Sometimes, a resin content is present inside the cells, as plugs or granules.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE183FE745E3A" blockId="10.[151,772,158,1175]" pageId="10" pageNumber="35">
<emphasis id="8EC5EA811D46FFB394EEE183FF1F586F" box="[151,203,490,513]" italics="true" pageId="10" pageNumber="35">Rays</emphasis>
in cross-section appear usually uniseriate and linear. In tangential section, the rays show 1-20(-25) cells in height (i.e. 30-570(-650) μm). The taller uniseriate rays have biseriations, which give sometimes a fusiform aspect. The ray-cells have polygonal rounded to oval section, marginal spaces and sometimes have dark content. The ray density is of 5-11 rays per horizontal millimeter. The rays appear homogeneous, with cells all procumbent, of 20-30 μm tall, relatively thick walled, of 4-5(7) μm the double wall. Within the marginal rows, but not only, the ray cells are taller, of 30-40 μm, and have slightly corrugated outer walls. The cross-fields are of araucarioid
<typeStatus id="630A88311D46FFB394EEE31EFF1C59E1" box="[151,200,887,911]" pageId="10" pageNumber="35">type</typeStatus>
, have 1-6(-8) pits, hexagonal or slightly rounded to oval, of cupressoid
<typeStatus id="630A88311D46FFB39514E3F1FE4959DE" box="[365,413,920,944]" pageId="10" pageNumber="35">type</typeStatus>
, tending to podocarpoid or araucarioid. They have 15-19/8-12 μm in diameters, and show circular or short elliptic tilted apertures of 10-14/5-7 μm. The pits are arranged in 1-2(3) rows per cross field, within ray body fields, or in the marginal fields and are alternate to slightly irregular.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE437FE785E1B" blockId="10.[151,772,158,1175]" box="[151,428,1117,1141]" pageId="10" pageNumber="35">
<emphasis id="8EC5EA811D46FFB394EEE437FECB5E1B" box="[151,287,1117,1141]" italics="true" pageId="10" pageNumber="35">Resin canals</emphasis>
are absent.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE416FE395EF9" blockId="10.[151,772,158,1175]" box="[151,493,1151,1175]" pageId="10" pageNumber="35">
<emphasis id="8EC5EA811D46FFB394EEE416FE8B5EF9" box="[151,351,1151,1175]" italics="true" pageId="10" pageNumber="35">Mineral inclusions</emphasis>
are absent.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D46FFB394EEE4A8FB3A5B35" pageId="10" pageNumber="35" type="discussion">
<paragraph id="BC0E36931D46FFB394EEE4A8FE785EB7" blockId="10.[151,771,1217,2035]" box="[151,428,1217,1241]" pageId="10" pageNumber="35">
<emphasis id="8EC5EA811D46FFB394EEE4A8FE785EB7" bold="true" box="[151,428,1217,1241]" pageId="10" pageNumber="35">Affinities and discussion</emphasis>
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE48AFE1C5F8C" blockId="10.[151,771,1217,2035]" pageId="10" pageNumber="35">
From the microscopic study of all the standard oriented thin sections, we selected
<specimenCount id="AAB7FD1A1D46FFB395BBE56AFD8C5F75" box="[450,600,1283,1307]" count="13" pageId="10" pageNumber="35" type="generic">13 specimens</specimenCount>
having similar generic characters, represented by tracheidoxylic structure, radial pitting of mixed
<typeStatus id="630A88311D46FFB395ADE52FFDD65F33" box="[468,514,1350,1373]" pageId="10" pageNumber="35">type</typeStatus>
on tracheids and araucarioid cross-fields, which are consistent with the structure of
<taxonomicName id="7BB14D101D46FFB3948EE5EEFE5F5FF1" box="[247,395,1415,1439]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D46FFB3948EE5EEFE5F5FF1" box="[247,395,1415,1439]" italics="true" pageId="10" pageNumber="35">Brachyoxylon</emphasis>
</taxonomicName>
wood-type (see
<bibRefCitation id="D8204B621D46FFB3962EE5EEFD2A5FF1" author="Philippe, M." box="[599,766,1415,1439]" pageId="10" pageNumber="35" pagination="74 - 80" refId="ref16155" refString="Philippe, M., 1993. Nomenclature generique des tracheidoxyles mesozoiques a champs araucarioides. Taxon, 42: 74 - 80." type="journal article" year="1993">Philippe, 1993</bibRefCitation>
; Philipe and Bamford, 2008), but a slightly different from the above-described species.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE582FE755CC6" blockId="10.[151,771,1217,2035]" pageId="10" pageNumber="35">
We do not repeat the discussion from above on the genus creation and about the previously described species, but, consulting the comparative Table 1, from
<bibRefCitation id="D8204B621D46FFB39618E644FF355C08" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." pageId="10" pageNumber="35" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei et al. (2018)</bibRefCitation>
, we observed beside the generic characters some other specific details that are different of those of the above-described species.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE6D8FD575D43" blockId="10.[151,771,1217,2035]" pageId="10" pageNumber="35">
Thus, we observed that the radial tracheidal pitting of mixed
<typeStatus id="630A88311D46FFB39498E6BAFEDB5C85" box="[225,271,1747,1771]" pageId="10" pageNumber="35">type</typeStatus>
are commonly discontinuous, the axial parenchyma is present, as dispersed cells with slightly nodular horizontal walls and with resin content inside.
</paragraph>
<paragraph id="BC0E36931D46FFB394EEE75FFC635AB9" blockId="10.[151,771,1217,2035]" lastBlockId="10.[816,1436,158,645]" pageId="10" pageNumber="35">
Other features are: the relatively tall uniseriate rays with biseriations, up to fusiform aspect, homogeneous and with cross-fields with 1-6(-8) hexagonal to rounded cupressoid oculipores, alternately arranged (araucarioid) to slightly irregular. All these details almost identical to those of the species
<taxonomicName id="7BB14D101D46FFB395F1E7B2FD7A5D9D" box="[392,686,2011,2035]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="35" phylum="Tracheophyta" rank="species" species="cristianicum">
<emphasis id="8EC5EA811D46FFB395F1E7B2FD7A5D9D" box="[392,686,2011,2035]" italics="true" pageId="10" pageNumber="35">Brachyoxylon cristianicum</emphasis>
</taxonomicName>
, which was previously described from the same area (
<bibRefCitation id="D8204B621D46FFB3914EE0F7FC785AB9" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." pageId="10" pageNumber="35" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei et al., 2018</bibRefCitation>
).
</paragraph>
<paragraph id="BC0E36931D46FFB39749E088FB3A5B35" blockId="10.[816,1436,158,645]" pageId="10" pageNumber="35">
From these characteristics, we attribute all the 13 here studied specimens coming from Holbav and Cristian to the above specified form:
<taxonomicName id="7BB14D101D46FFB3900BE14BFB3F5B35" authority="Iamandei, Iamandei and Gradinaru, 2018" authorityName="Iamandei, Iamandei and Gradinaru" authorityYear="2018" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="35" phylum="Tracheophyta" rank="species" species="cristianicum">
<emphasis id="8EC5EA811D46FFB3900BE14BFA485B54" box="[1138,1436,290,314]" italics="true" pageId="10" pageNumber="35">Brachyoxylon cristianicum</emphasis>
<bibRefCitation id="D8204B621D46FFB39749E12AFB3F5B35" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." box="[816,1259,323,347]" pageId="10" pageNumber="35" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei, Iamandei and Grădinaru, 2018</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="7BB14D101D48FFBD948FE2E7FD9D58C8" authority="Iamandei &amp; Iamandei &amp; Grădinaru, 2022" authorityName="Iamandei &amp; Iamandei &amp; Grădinaru" authorityYear="2022" box="[246,585,654,678]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="species" species="holbavicum" status="sp. nov.">
<emphasis id="8EC5EA811D48FFBD948FE2E7FD9D58C8" box="[246,585,654,678]" italics="true" pageId="4" pageNumber="29">Protojuniperoxylon holbavicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95F657FA1D48FFBD9629E2E6FD7158C8" box="[592,677,655,678]" pageId="4" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="248A2A161D48FFBD95EFE2C6FE0258A9" box="[406,470,687,711]" captionStart="Fig" captionStartId="5.[169,204,1925,1946]" captionTargetBox="[169,1412,154,1916]" captionTargetId="figure-19@5.[169,1412,154,1916]" captionTargetPageId="5" captionText="Fig. 2 Protojuniperoxylon holbavicum sp. nov. (inventory no. 27706). a-c: Cross section - tracheids polygonal with rounded corners, and linear rays; d-f: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells, with lateral empty spaces; unpitted tracheids; g-j: Radial section uniseriate radial pitting on tracheids as hexagonal pits; araucarioid cross-fields with vertical pairs of pits or slightly irregular; end-wall with juniperoid thickenings (see arrow)." figureDoi="http://doi.org/10.5281/zenodo.10520855" httpUri="https://zenodo.org/record/10520855/files/figure.png" pageId="4" pageNumber="29">Fig. 2</figureCitation>
, a-j.
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<emphasis id="8EC5EA811D48FFBD94EEE29BFF2E5964" bold="true" box="[151,250,754,778]" pageId="4" pageNumber="29">Material</emphasis>
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<materialsCitation id="0CD93CCE1D48FFBD94EEE37AFEF95ED6" collectionCode="NMG" location="Holbav" municipality="Maiului" pageId="4" pageNumber="29" specimenCode="27704, 27705, 27706, 27707, 27708, 27709, 27710" specimenCount="7">
The studied material is represented by
<specimenCount id="AAB7FD1A1D48FFBD964EE37AFD905944" box="[567,580,787,810]" pageId="4" pageNumber="29">7</specimenCount>
samples of silicified wood, collected from the
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area, on
<collectingMunicipality id="5C6AACE91D48FFBD96D6E35DFCD75922" box="[687,771,820,844]" pageId="4" pageNumber="29">Maiului</collectingMunicipality>
brook. All studied samples are fragments of trunk or thick branches, of decimetric size, dark to black color. Viewing through a hand lens, or even by naked eye, it is possible to see the regular fibrous structure without vessels, which indicate a conifer wood. The specimens attributed to this species have very similar details and have the following field numbers within the &quot;Grădinaru Collection&quot;: 1078, 1083, 1085, 1087, 1091, 1095, 1110 and are stored at the
<collectionCode id="DAA0AE561D48FFBD94C7E434FDD45E1B" box="[190,512,1117,1141]" pageId="4" pageNumber="29">National Museum of Geology</collectionCode>
, in Bucharest, under the inventory numbers:
<specimenCode id="EC179EE81D48FFBD950BE416FE6C5EF9" box="[370,440,1151,1175]" pageId="4" pageNumber="29">27704</specimenCode>
,
<specimenCode id="EC179EE81D48FFBD95BAE416FDDC5EF9" box="[451,520,1151,1175]" pageId="4" pageNumber="29">27705</specimenCode>
,
<specimenCode id="EC179EE81D48FFBD966CE416FD8F5EF9" box="[533,603,1151,1175]" pageId="4" pageNumber="29">27706</specimenCode>
,
<specimenCode id="EC179EE81D48FFBD961EE416FD785EF9" box="[615,684,1151,1175]" pageId="4" pageNumber="29">27707</specimenCode>
,
<specimenCode id="EC179EE81D48FFBD96C0E416FD2A5EF9" box="[697,766,1151,1175]" pageId="4" pageNumber="29">27708</specimenCode>
,
<specimenCode id="EC179EE81D48FFBD94EEE4C9FF085ED6" box="[151,220,1184,1208]" pageId="4" pageNumber="29">27709</specimenCode>
,
<specimenCode id="EC179EE81D48FFBD949EE4C9FEF95ED6" box="[231,301,1184,1208]" pageId="4" pageNumber="29">27710</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D48FFBD94EEE48BFB655EB7" pageId="4" pageNumber="29" type="description">
<paragraph id="BC0E36931D48FFBD94EEE48BFE7C5E94" blockId="4.[151,772,1250,2068]" box="[151,424,1250,1274]" pageId="4" pageNumber="29">
<emphasis id="8EC5EA811D48FFBD94EEE48BFE7C5E94" bold="true" box="[151,424,1250,1274]" pageId="4" pageNumber="29">Microscopic description</emphasis>
</paragraph>
<paragraph id="BC0E36931D48FFBD94EEE56AFEDE5FAF" blockId="4.[151,772,1250,2068]" pageId="4" pageNumber="29">
<emphasis id="8EC5EA811D48FFBD94EEE56AFEFF5F75" box="[151,299,1283,1307]" italics="true" pageId="4" pageNumber="29">Growth rings</emphasis>
well developed in the studied samples, representing the secondary xylem which is tracheidoxylic, showing a fairly gradual transition but sometimes abrupt, from early- to late-wood, and has distinct growth ring boundaries. Normal or traumatic axial resin canals are absent.
</paragraph>
<paragraph id="BC0E36931D48FFBD94EEE5A3FBE65A96" blockId="4.[151,772,1250,2068]" lastBlockId="4.[816,1437,163,1241]" pageId="4" pageNumber="29">
<emphasis id="8EC5EA811D48FFBD94EEE5A3FED65F8C" box="[151,258,1482,1506]" italics="true" pageId="4" pageNumber="29">Tracheids</emphasis>
in cross section appear quasi-polygonal, usually quadrangular, and with moderately thick walls in the early-wood. They have radial / tangential diameters of 25-60 / 25-55 μm in the early- wood and wall thickness of 3-7 μm double wall. When gradual transition is present, the tracheids gradually diminish in section and usually the tracheidal wall in the late-wood becomes slightly thicker, of 7-9 μm in the final 3-8 rows of smaller sized and radially flattened cells, with r/tg diameters of 10-25 / 20-25 μm, marking the growth- ring boundary. Between two successive rays, 2-12 radial rows of tracheids can be counted. The density ranges 750-1216 tracheids / mm
<superScript id="4BC49BDB1D48FFBD968DE75DFD285D2C" attach="left" box="[756,764,1844,1858]" fontSize="6" pageId="4" pageNumber="29">2</superScript>
. Longitudinally, the tracheids have, on the radial walls, uniseriate pits, often with biseriate portions, often contiguous, or biseriate of mixed
<typeStatus id="630A88311D48FFBD95BAE7F3FE275DDC" box="[451,499,1946,1970]" pageId="4" pageNumber="29">type</typeStatus>
, sometimes separated by crassulae. The pits are of abietinean
<typeStatus id="630A88311D48FFBD965AE7D2FD805DBD" box="[547,596,1979,2003]" pageId="4" pageNumber="29">type</typeStatus>
, have 14-21 μm in diameter and round apertures of 4-5 μm. Details on torus not observed. On the tangential walls, pits are usually absent. Organic deposits - absent. Average tracheids length difficult to measure. Helical thickenings usually absent, or rarely appear.
</paragraph>
<paragraph id="BC0E36931D48FFBD9749E168FC075BD1" blockId="4.[816,1437,163,1241]" pageId="4" pageNumber="29">
<emphasis id="8EC5EA811D48FFBD9749E168FC2D5B77" box="[816,1017,257,281]" italics="true" pageId="4" pageNumber="29">Axial parenchyma</emphasis>
in the cross sections is absent, or rarely present as dispersed thin-walled cells with dark content. In longitudinal view, the parenchyma appears as vertical rows of rectangular cells, with the horizontal wall thin and smooth or feebly nodular and with some dark remains inside.
</paragraph>
<paragraph id="BC0E36931D48FFBD9749E1A0FAFD5EF9" blockId="4.[816,1437,163,1241]" pageId="4" pageNumber="29">
<emphasis id="8EC5EA811D48FFBD9749E1A0FCB05B8E" box="[816,868,457,480]" italics="true" pageId="4" pageNumber="29">Rays</emphasis>
usually thin in cross section, appear in tangential section as uniseriate, but sometimes with some biseriate storeys, or even as 2-3-seriate rays. The ray height varies from 232 to up to 56 cells (average height - 16 cells). The ray cells appear ovoid-rectangular, with 2639 μm in height and 2330 μm in width. Lateral triangular spaces are present. Their density is 6-9 rays on tangential milimeter. In radial section, the rays appear heterogeneous, because they have thicker walled cells, as ray tracheids, beside the thinner-walled parenchymal body cells, and are all procumbent. The ray cells have 8-20 μm tall, the marginals are slightly taller, of 25-30 μm. The horizontal walls of the ray cells are smooth and usually pitted, and the tangential end-walls are relatively thick, of 1.5-2 μm the simple wall and are nodular, as typical juniperoid thickenings. The ray tracheids are located in the ray-body or marginally, and have smooth or slightly rough thick walls. The indentures are missing or difficult to observe. Spiral thickenings are usually absent. The cross-fields are typical cupressoid, with 1-2-4(-6) cupressoid oculipores of 8-10 μm in diameter, usually in vertical pairs arranged, and with slit-like inclined apertures, of 2-5 μm.
</paragraph>
<paragraph id="BC0E36931D48FFBD9749E4C9FB3C5ED6" blockId="4.[816,1437,163,1241]" box="[816,1256,1184,1208]" pageId="4" pageNumber="29">
<emphasis id="8EC5EA811D48FFBD9749E4C9FB8E5ED6" box="[816,1114,1184,1208]" italics="true" pageId="4" pageNumber="29">Radial or axial resin canals</emphasis>
are absent.
</paragraph>
<paragraph id="BC0E36931D48FFBD9749E4A8FB655EB7" blockId="4.[816,1437,163,1241]" box="[816,1201,1217,1241]" pageId="4" pageNumber="29">
<emphasis id="8EC5EA811D48FFBD9749E4A8FC2D5EB7" box="[816,1017,1217,1241]" italics="true" pageId="4" pageNumber="29">Mineral inclusions</emphasis>
are not present
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D48FFBF9749E56DFA475B35" lastPageId="6" lastPageNumber="31" pageId="4" pageNumber="29" type="discussion">
<paragraph id="BC0E36931D48FFBD9749E56DFB925F72" blockId="4.[816,1437,1284,2069]" box="[816,1094,1284,1308]" pageId="4" pageNumber="29">
<emphasis id="8EC5EA811D48FFBD9749E56DFB925F72" bold="true" box="[816,1094,1284,1308]" pageId="4" pageNumber="29">Affinities and discussion</emphasis>
</paragraph>
<paragraph id="BC0E36931D48FFBD9749E54DFB465CE9" blockId="4.[816,1437,1284,2069]" pageId="4" pageNumber="29">
The cupressaceous xylotomy of the
<specimenCount id="AAB7FD1A1D48FFBD90C6E54CFA915F52" box="[1215,1349,1316,1340]" count="7" pageId="4" pageNumber="29" type="generic">7 specimens</specimenCount>
studied here, assigned to the genus
<taxonomicName id="7BB14D101D48FFBD901AE52CFA4C5F33" authority="Eckhold" authorityName="Eckhold" authorityYear="1922" box="[1123,1432,1349,1373]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D48FFBD901AE52CFAE05F33" box="[1123,1332,1349,1373]" italics="true" pageId="4" pageNumber="29">Protojuniperoxylon</emphasis>
Eckhold
</taxonomicName>
, are definitely pycnoxylic and homoxylic (or tracheidoxylic sensu
<bibRefCitation id="D8204B621D48FFBD97B9E5EEFB865FF1" author="Creber, G. T." box="[960,1106,1415,1439]" pageId="4" pageNumber="29" pagination="655 - 661" refId="ref13670" refString="Creber, G. T., 1972. Gymnospermous wood from the Kimmeridgian of East Sutherland and from the Sandringham Sands of Norfolk. Palaeontology, 15 (4): 655 - 661." type="journal article" year="1972">Creber, 1972</bibRefCitation>
), having radial pitting on tracheids of mixed
<typeStatus id="630A88311D48FFBD9797E5C0FBCB5FAF" box="[1006,1055,1449,1473]" pageId="4" pageNumber="29">type</typeStatus>
, no resin canals, rare parenchyma but with nodular horizontal walls, rays exclusively uniseriate and cross-fields with 1-2(4-6) cupressoid pits with slit-like inclined apertures, arranged in 1-2 vertical pairs, or irregularly to slightly alternately and with nodular tangential end-walls of ray cells as typical juniperoid nodules (i.e. juniperoid thickenings).
</paragraph>
<paragraph id="BC0E36931D48FFBD9749E6F8FC0C5DFE" blockId="4.[816,1437,1284,2069]" pageId="4" pageNumber="29">
We have discussed above the current status of “
<taxonomicName id="7BB14D101D48FFBD9133E6F9FC545CA7" class="Pinopsida" family="Protopinaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="family">Protopinaceae</taxonomicName>
family” and of “Cheirlepidiaceae family” and, lastly, the proposal to be accepted as valid - the family name “
<taxonomicName id="7BB14D101D48FFBD9704E69DFBF85D62" authorityName="Harris" authorityYear="1979" box="[893,1068,1780,1804]" family="Hirmeriellaceae" pageId="4" pageNumber="29" rank="family">Hirmeriellaceae</taxonomicName>
” (
<bibRefCitation id="D8204B621D48FFBD903FE69DFB075D62" author="Harris, T. M." box="[1094,1235,1780,1804]" pageId="4" pageNumber="29" refId="ref14877" refString="Harris, T. M., 1979. The Yorkshire Jurassic Flora. 5. Coniferales. British Museum Natural History, London, 167 pp." type="book" year="1979">Harris, 1979</bibRefCitation>
), which is considered correct and valid to be used for the Mesozoic gymnosperms characterized by pollen of
<taxonomicName id="7BB14D101D48FFBD90A2E75FFA8B5D20" authorityName="H.D.Pflug" authorityYear="1953" box="[1243,1375,1846,1870]" class="Pinopsida" family="Cheirolepidiaceae" genus="Classopollis" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D48FFBD90A2E75FFA8B5D20" box="[1243,1375,1846,1870]" italics="true" pageId="4" pageNumber="29">Classopollis</emphasis>
</taxonomicName>
-
<typeStatus id="630A88311D48FFBD9111E75EFA4C5D21" box="[1384,1432,1847,1871]" pageId="4" pageNumber="29">type</typeStatus>
, and was proposed for taxonomic conservation (see
<bibRefCitation id="D8204B621D48FFBD9749E711FC185DFE" author="Doweld, A. B." box="[816,972,1912,1936]" pageId="4" pageNumber="29" pagination="1092 - 1098" refId="ref14259" refString="Doweld, A. B., 2020. The controversial nomenclature of the fossil plant names Cheirolepis, Cheirolepidium and Hirmeriella (Cheirolepidaceae / Cheirolepidiaceae / Hirmeriellaceae). 69 (5): 1092 - 1098." type="journal article" year="2020">Doweld, 2020</bibRefCitation>
).
</paragraph>
<paragraph id="BC0E36931D48FFBF9749E7F0FE715B54" blockId="4.[816,1437,1284,2069]" lastBlockId="6.[151,771,158,2068]" lastPageId="6" lastPageNumber="31" pageId="4" pageNumber="29">
The genus name
<taxonomicName id="7BB14D101D48FFBD978BE7F0FB165DDF" authorityName="Eckhold" authorityYear="1922" box="[1010,1218,1945,1969]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D48FFBD978BE7F0FB165DDF" box="[1010,1218,1945,1969]" italics="true" pageId="4" pageNumber="29">Protojuniperoxylon</emphasis>
</taxonomicName>
was established by Eckhold in his PhD Thesis (
<bibRefCitation id="D8204B621D48FFBD9019E7D3FAD45DBC" author="Eckhold, W." box="[1120,1280,1978,2002]" pageId="4" pageNumber="29" refId="ref14298" refString="Eckhold, W., 1921. Die Hoftupfel bei rezenten und fossilen Koniferen. [PhD Thesis] Breslau Schlesische Fridrich-Wilhelms Universitat (Univ. of Wroclaw Poland)" type="book" year="1921">Eckhold, 1921</bibRefCitation>
,
<bibRefCitation id="D8204B621D48FFBD9175E7D3FA905DBC" author="Eckhold, W." box="[1292,1348,1978,2002]" pageId="4" pageNumber="29" pagination="472 - 505" refId="ref14328" refString="Eckhold, W., 1922. Die Hoftupfel bei rezenten und fossilen Coniferen. Jahrbuch Konigliche Preussische Geologische Landesansalt, 42: 472 - 505." type="journal article" year="1922">1922</bibRefCitation>
, p.491), to designate Mesozoic cupressaceous wood of
<taxonomicName id="7BB14D101D48FFBD9152E7B5FA405D9D" box="[1323,1428,2012,2035]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D48FFBD9152E7B5FA405D9D" box="[1323,1428,2012,2035]" italics="true" pageId="4" pageNumber="29">Juniperus</emphasis>
</taxonomicName>
-
<typeStatus id="630A88311D48FFBD9749E794FCB5527B" box="[816,865,2045,2069]" pageId="4" pageNumber="29">type</typeStatus>
, described like this: “Annual rings more or less marked, tracheid pits in various transitional arrangements, Abietineentüpfelung absent, but Juniperustüpfelung present, no resin canals, cupressoid pits in cross-field, axial parenchyma occasionally present” (
<bibRefCitation id="D8204B621D4AFFBF96DBE168FF195B54" author="Eckhold, W." pageId="6" pageNumber="31" refId="ref14298" refString="Eckhold, W., 1921. Die Hoftupfel bei rezenten und fossilen Koniferen. [PhD Thesis] Breslau Schlesische Fridrich-Wilhelms Universitat (Univ. of Wroclaw Poland)" type="book" year="1921">Eckhold, 1921</bibRefCitation>
fide
<bibRefCitation id="D8204B621D4AFFBF957DE14BFE4D5B54" author="Bodnar, J." box="[260,409,290,314]" pageId="6" pageNumber="31" pagination="1480 - 1481" refId="ref13105" refString="Bodnar, J., 2017. (2575) Proposal to conserve the name Protocupressinoxylon with a conserved type against Protobrachyoxylon (fossil plants). Taxon (Journal of IAPT), 66 (6): 1480 - 1481. https: // doi. org / 10.12705 / 666.25" type="journal article" year="2017">Bodnar, 2017</bibRefCitation>
).
</paragraph>
<caption id="E8CE661B1D49FFBC94D0E7ECFEB5527F" ID-DOI="http://doi.org/10.5281/zenodo.10520855" ID-Zenodo-Dep="10520855" httpUri="https://zenodo.org/record/10520855/files/figure.png" pageId="5" pageNumber="30" startId="5.[169,204,1925,1946]" targetBox="[169,1412,154,1916]" targetPageId="5" targetType="figure">
<paragraph id="BC0E36931D49FFBC94D0E7ECFC9E5DF4" blockId="5.[169,1429,1924,2065]" box="[169,842,1924,1946]" pageId="5" pageNumber="30">
<emphasis id="8EC5EA811D49FFBC94D0E7ECFF365DF4" bold="true" box="[169,226,1925,1946]" pageId="5" pageNumber="30">Fig. 2</emphasis>
<taxonomicName id="7BB14D101D49FFBC9491E7EDFDCE5DF4" authority="Iamandei &amp; Iamandei &amp; Grădinaru, 2022" authorityName="Iamandei &amp; Iamandei &amp; Grădinaru" authorityYear="2022" box="[232,538,1924,1946]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="30" phylum="Tracheophyta" rank="species" species="holbavicum" status="sp. nov.">
<emphasis id="8EC5EA811D49FFBC9491E7EDFDCE5DF4" box="[232,538,1924,1946]" italics="true" pageId="5" pageNumber="30">Protojuniperoxylon holbavicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95F657FA1D49FFBC9659E7ECFDB85DF4" box="[544,620,1925,1946]" pageId="5" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
(inventory no. 27706).
</paragraph>
<paragraph id="BC0E36931D49FFBC94D0E7CAFEB5527F" blockId="5.[169,1429,1924,2065]" pageId="5" pageNumber="30">
<emphasis id="8EC5EA811D49FFBC94D0E7CAFF1C5DD6" bold="true" box="[169,200,1955,1976]" pageId="5" pageNumber="30">a-c</emphasis>
: Cross section - tracheids polygonal with rounded corners, and linear rays;
<emphasis id="8EC5EA811D49FFBC97DDE7CAFC155DD6" bold="true" box="[932,961,1955,1976]" pageId="5" pageNumber="30">d-f</emphasis>
: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells, with lateral empty spaces; unpitted tracheids;
<emphasis id="8EC5EA811D49FFBC97B6E7A9FC3F5DBB" bold="true" box="[975,1003,1984,2005]" pageId="5" pageNumber="30">g-j</emphasis>
: Radial section uniseriate radial pitting on tracheids as hexagonal pits; araucarioid cross-fields with vertical pairs of pits or slightly irregular; end-wall with juniperoid thickenings (see arrow).
</paragraph>
</caption>
<paragraph id="BC0E36931D4AFFBF94EEE12DFE735B12" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">In fact, few species were described for this genus, sometimes revised, as follows:</paragraph>
<paragraph id="BC0E36931D4AFFBF94EEE1ECFDBC580A" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
<taxonomicName id="7BB14D101D4AFFBF94EEE1ECFCD75BF3" authority="(Stopes) Eckhold, 1921" authorityName="Eckhold" authorityYear="1921" baseAuthorityName="Stopes" baseAuthorityYear="1915" box="[151,771,389,413]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="maidstonense">
<emphasis id="8EC5EA811D4AFFBF94EEE1ECFE2B5BF3" box="[151,511,389,413]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon maidstonense</emphasis>
(Stopes)
<bibRefCitation id="D8204B621D4AFFBF961EE1ECFCD75BF3" author="Eckhold, W." box="[615,771,389,413]" pageId="6" pageNumber="31" refId="ref14298" refString="Eckhold, W., 1921. Die Hoftupfel bei rezenten und fossilen Koniferen. [PhD Thesis] Breslau Schlesische Fridrich-Wilhelms Universitat (Univ. of Wroclaw Poland)" type="book" year="1921">Eckhold, 1921</bibRefCitation>
</taxonomicName>
was taken as type-species, even if, initially, this wood
<typeStatus id="630A88311D4AFFBF94EEE1A1FF115B8E" box="[151,197,456,480]" pageId="6" pageNumber="31">type</typeStatus>
was described from the Cretaceous of Maidstone (
<collectingCountry id="C4A676031D4AFFBF94E7E180FEE2586F" box="[158,310,489,513]" name="United Kingdom" pageId="6" pageNumber="31">Great Britain</collectingCountry>
), as a species of
<taxonomicName id="7BB14D101D4AFFBF967CE180FD55586F" authorityName="G.Kraus" authorityYear="1870" box="[517,641,489,513]" class="Pinopsida" family="Pinaceae" genus="Cedroxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF967CE180FD55586F" box="[517,641,489,513]" italics="true" pageId="6" pageNumber="31">Cedroxylon</emphasis>
</taxonomicName>
, by
<bibRefCitation id="D8204B621D4AFFBF96C5E180FF35584C" author="Stopes, M." pageId="6" pageNumber="31" refId="ref17231" refString="Stopes, M., 1915. Catalogue of the Mesozoic plants in the British Museum - The Cretaceous flora, Part 2, Lower Greensand (Aptian) plants of Britain: London, British Museum (Natural History), 360 p." type="book" year="1915">Stopes (1915)</bibRefCitation>
. It has uniseriate radial pitting on tracheids, few axial parenchyma with nodular end wall, and cupressoid cross-fields (see Bodnar and Artabe, 2007).
</paragraph>
<paragraph id="BC0E36931D4AFFBF94EEE204FDEA5887" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
<taxonomicName id="7BB14D101D4AFFBF94EEE204FED858C8" authority="(Seward and Bancroft) Eckhold, 1921" authorityName="Eckhold" authorityYear="1921" baseAuthorityName="Seward and Bancroft" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="hornei">
<emphasis id="8EC5EA811D4AFFBF94EEE204FE6E58EB" box="[151,442,621,645]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon hornei</emphasis>
(Seward and Bancroft)
<bibRefCitation id="D8204B621D4AFFBF96A8E204FED858C8" author="Eckhold, W." pageId="6" pageNumber="31" refId="ref14298" refString="Eckhold, W., 1921. Die Hoftupfel bei rezenten und fossilen Koniferen. [PhD Thesis] Breslau Schlesische Fridrich-Wilhelms Universitat (Univ. of Wroclaw Poland)" type="book" year="1921">Eckhold, 1921</bibRefCitation>
</taxonomicName>
was initially described as Jurassic species of
<taxonomicName id="7BB14D101D4AFFBF94EEE2C6FEC758A9" authorityName="G.Kraus" authorityYear="1870" box="[151,275,687,711]" class="Pinopsida" family="Pinaceae" genus="Cedroxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF94EEE2C6FEC758A9" box="[151,275,687,711]" italics="true" pageId="6" pageNumber="31">Cedroxylon</emphasis>
</taxonomicName>
and later, of
<taxonomicName id="7BB14D101D4AFFBF95DFE2C6FD2B58A9" authority="(Vogellehner, 1968)" baseAuthorityName="Vogellehner" baseAuthorityYear="1968" box="[422,767,687,711]" class="Pinopsida" genus="Scotoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF95DFE2C6FDCF58A9" box="[422,539,687,711]" italics="true" pageId="6" pageNumber="31">Scotoxylon</emphasis>
(
<bibRefCitation id="D8204B621D4AFFBF9652E2C6FD2258A9" author="Vogellehner, D." box="[555,758,687,711]" pageId="6" pageNumber="31" pagination="125 - 162" refId="ref17771" refString="Vogellehner, D., 1968. Zur Anatomie und Phylogenie de Mesozoischer Gymnospermenholzer, 7: Prodromus zu einer Monographie de Protopinaceae. II. Die Protopinoiden Holzer des Jura. Palaeontographica Abt. B, Stuttgart, 124 (4 - 6): 125 - 162." type="journal article" year="1968">Vogellehner, 1968</bibRefCitation>
)
</taxonomicName>
, so, it could be doubtful for comparison.
</paragraph>
<paragraph id="BC0E36931D4AFFBF94EEE29BFDAF5903" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
However, Zhang et al. (2000) described a
<taxonomicName id="7BB14D101D4AFFBF96F7E29BFE3B5945" authority="Zhang and Zengh" authorityName="Zhang and Zengh" class="Pinopsida" genus="Scotoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="yanqingense">
<emphasis id="8EC5EA811D4AFFBF96F7E29BFEC85944" italics="true" pageId="6" pageNumber="31">Scotoxylon yanqingense</emphasis>
Zhang and Zengh
</taxonomicName>
, a Late Jurassic Chinese species, as representing secondary wood of
<taxonomicName id="7BB14D101D4AFFBF9609E35CFF0A5903" authorityName="Eckhold" authorityYear="1922" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF9609E35CFF0A5903" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon</emphasis>
</taxonomicName>
-
<typeStatus id="630A88311D4AFFBF949EE33FFEC35900" box="[231,279,854,878]" pageId="6" pageNumber="31">type</typeStatus>
, but devoid of axial parenchyma.
</paragraph>
<paragraph id="BC0E36931D4AFFBF94EEE31FFEB35E3A" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
<taxonomicName id="7BB14D101D4AFFBF94EEE31FFE7C59C1" authority="(Bonetti 1966) Bodnar and Artabe, 2007" authorityName="Bodnar and Artabe" authorityYear="2007" baseAuthorityName="Bonetti" baseAuthorityYear="1966" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="ischigualastensis">
<emphasis id="8EC5EA811D4AFFBF94EEE31FFDEC59E0" box="[151,568,886,910]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon ischigualastensis</emphasis>
(
<bibRefCitation id="D8204B621D4AFFBF9623E31FFD2F59E0" author="Bonetti, M. I. R." box="[602,763,886,910]" pageId="6" pageNumber="31" pagination="211 - 218" refId="ref13201" refString="Bonetti, M. I. R., 1966. Protojuniperoxylon ischigualastensis sp. nov. del Triasico de Ischigualasto (San Juan). Ameghiniana, 4: 211 - 218." type="journal article" year="1966">Bonetti 1966</bibRefCitation>
) Bodnar and Artabe, 2007
</taxonomicName>
is a Triassic wood from
<collectingCountry id="C4A676031D4AFFBF96CEE3FEFF1659BE" name="Argentina" pageId="6" pageNumber="31">Argentina</collectingCountry>
, which has mixed radial pitting on tracheids, cupressoid cross fields, juniperoid thickenings on the end walls of ray parenchymal cells and nodules on the horizontal walls axial parenchyma, which are typical for
<taxonomicName id="7BB14D101D4AFFBF9698E475FF335E3A" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF9698E475FF335E3A" italics="true" pageId="6" pageNumber="31">Juniperus</emphasis>
</taxonomicName>
wood-type.
</paragraph>
<paragraph id="BC0E36931D4AFFBF94EEE434FEEF5F52" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
<taxonomicName id="7BB14D101D4AFFBF94EEE434FDB95E1B" authority="Selling, 1951" authorityName="Selling" authorityYear="1951" box="[151,621,1117,1141]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="arcticum">
<emphasis id="8EC5EA811D4AFFBF94EEE434FE055E1B" box="[151,465,1117,1141]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon arcticum</emphasis>
<bibRefCitation id="D8204B621D4AFFBF95A5E434FDB95E1B" author="Selling, O. H." box="[476,621,1117,1141]" pageId="6" pageNumber="31" pagination="538 - 539" refId="ref17060" refString="Selling, O. H., 1951. On Protojuniperoxylon arcticum. Journal of Paleontology, 25 (4): 538 - 539." type="journal article" year="1951">Selling, 1951</bibRefCitation>
</taxonomicName>
described on some Mesozoic root-wood remains from Hope Island (
<collectingCountry id="C4A676031D4AFFBF94E7E4C9FED55ED6" box="[158,257,1184,1208]" name="Svalbard and Jan Mayen" pageId="6" pageNumber="31">Svalbard</collectingCountry>
,
<collectingCountry id="C4A676031D4AFFBF956BE4C9FEBD5ED6" box="[274,361,1184,1208]" name="Norway" pageId="6" pageNumber="31">Norway</collectingCountry>
) is poorly identified, for it is most probably a bennettitalean taxon (
<taxonomicName id="7BB14D101D4AFFBF9582E4A8FD2C5EB7" authority="Buckland" authorityName="Buckland" authorityYear="1828" box="[507,760,1217,1241]" class="Cycadopsida" family="Cycadeoideaceae" genus="Cycadeoidea" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF9582E4A8FD525EB7" box="[507,646,1217,1241]" italics="true" pageId="6" pageNumber="31">Cycadeoidea</emphasis>
Buckland
</taxonomicName>
), showing interactions with chytrid fungi and bacteria (see
<bibRefCitation id="D8204B621D4AFFBF94EEE56AFE075F75" author="Strullu-Derrien, C. &amp; McLoughlin, S. &amp; Philippe, M. &amp; Mork, A. &amp; Strullu, D. G." box="[151,467,1283,1307]" pageId="6" pageNumber="31" pagination="45 - 58" refId="ref17334" refString="Strullu-Derrien, C., McLoughlin, S., Philippe, M., Mork, A. and Strullu, D. G., 2012. Arthropod interactions with bennettitalean roots in a Triassic permineralized peat from Hopen, Svalbard Archipelago (Arctic). Palaeogeography Palaeoclimatology Palaeoecology 348 - 349: 45 - 58. https: // doi. org / 10.1016 / j. palaeo. 2012.06.006" type="journal article" year="2012">Strullu-Derrien et al., 2012</bibRefCitation>
; McLoughlin and Strullu-Derrien. 2016).
</paragraph>
<paragraph id="BC0E36931D4AFFBF94EEE52CFD0B5C08" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
<bibRefCitation id="D8204B621D4AFFBF94EEE52CFE4E5F33" author="Ruiz, D. P. &amp; Bodnar, J." box="[151,410,1349,1373]" pageId="6" pageNumber="31" pagination="481 - 488" refId="ref16844" refString="Ruiz, D. P. and Bodnar, J., 2019. The oldest record of Juniperoxylon, a cupressaceous fossil wood from the Middle Triassic of Argentina. Acta Palaeontologica Polonica, 64 (3): 481 - 488." type="journal article" year="2019">Ruiz and Bodnar (2019)</bibRefCitation>
revised a Mid Triassic species of
<taxonomicName id="7BB14D101D4AFFBF94EEE50FFE9E5F10" authorityName="Göppert" authorityYear="1850" box="[151,330,1382,1406]" class="Pinopsida" family="Cupressaceae" genus="Cupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF94EEE50FFE9E5F10" box="[151,330,1382,1406]" italics="true" pageId="6" pageNumber="31">Cupressinoxylon</emphasis>
</taxonomicName>
of Bodnar et al. (2015) as
<taxonomicName id="7BB14D101D4AFFBF9612E50FFDA45FF1" authority="(Bodnar et al.) Ruiz and Bodnar" authorityName="Ruiz and Bodnar" baseAuthorityName="Bodnar" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="zamunerae">
<emphasis id="8EC5EA811D4AFFBF9612E50FFED85FF1" italics="true" pageId="6" pageNumber="31">Juniperoxylon zamunerae</emphasis>
(Bodnar et al.) Ruiz and Bodnar
</taxonomicName>
, and made an extensive discussion on the wood structure of
<taxonomicName id="7BB14D101D4AFFBF96E8E5C3FD2E5FAF" box="[657,762,1450,1473]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF96E8E5C3FD2E5FAF" box="[657,762,1450,1473]" italics="true" pageId="6" pageNumber="31">Juniperus</emphasis>
</taxonomicName>
-
<typeStatus id="630A88311D4AFFBF94EEE5A3FF115F8C" box="[151,197,1482,1506]" pageId="6" pageNumber="31">type</typeStatus>
including in a table the described species of
<taxonomicName id="7BB14D101D4AFFBF96D0E5A2FEC25C6D" authorityName="Eckhold" authorityYear="1922" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF96D0E5A2FEC25C6D" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon</emphasis>
</taxonomicName>
together with Mesozoic and Cenozoic species of
<taxonomicName id="7BB14D101D4AFFBF9497E665FE525C4A" authorityName="C.Houlbert" authorityYear="1910" box="[238,390,1548,1572]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF9497E665FE525C4A" box="[238,390,1548,1572]" italics="true" pageId="6" pageNumber="31">Juniperoxylon</emphasis>
</taxonomicName>
, all of them characterized by the presence of nodular ray parenchyma end-walls (i.e. Juniperustüpfelung), details also observed in our material.
</paragraph>
<paragraph id="BC0E36931D4AFFBF94EEE606FD0A5D65" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
Also, very useful is the discussion of
<bibRefCitation id="D8204B621D4AFFBF963EE606FD2A5CE9" author="Akkemik, U." box="[583,766,1647,1671]" pageId="6" pageNumber="31" pagination="15 - 26" refId="ref12580" refString="Akkemik, U., 2021. A new species of Juniperoxylon from the Early Miocene of Northwestern Turkey. Acta Palaeontologica Romaniae, 17 (1): 15 - 26. https: // doi. org / 10.35463 / j. apr. 2021.01.02" type="journal article" year="2021">Akkemik (2021)</bibRefCitation>
, who described and identified an Early Miocene species of
<taxonomicName id="7BB14D101D4AFFBF94EEE6D8FEE45CA7" authorityName="C.Houlbert" authorityYear="1910" box="[151,304,1713,1737]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF94EEE6D8FEE45CA7" box="[151,304,1713,1737]" italics="true" pageId="6" pageNumber="31">Juniperoxylon</emphasis>
</taxonomicName>
from
<collectingCountry id="C4A676031D4AFFBF9501E6D8FE135CA7" box="[376,455,1713,1737]" name="Turkey" pageId="6" pageNumber="31">Turkey</collectingCountry>
, as having typical xylotomy of
<taxonomicName id="7BB14D101D4AFFBF94C0E6BAFEF65C84" box="[185,290,1747,1770]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF94C0E6BAFEF65C84" box="[185,290,1747,1770]" italics="true" pageId="6" pageNumber="31">Juniperus</emphasis>
</taxonomicName>
<typeStatus id="630A88311D4AFFBF9554E6BAFE895C85" box="[301,349,1747,1771]" pageId="6" pageNumber="31">type</typeStatus>
, proposing even an identification key that includes also some species of
<taxonomicName id="7BB14D101D4AFFBF967EE69DFD0C5D62" authorityName="Eckhold" authorityYear="1922" box="[519,728,1780,1804]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF967EE69DFD0C5D62" box="[519,728,1780,1804]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="BC0E36931D4AFFBF94EEE77CFA475B35" blockId="6.[151,771,158,2068]" lastBlockId="6.[816,1436,158,347]" pageId="6" pageNumber="31">
Our material displays the typical xylotomical features of
<taxonomicName id="7BB14D101D4AFFBF94EEE75EFED45D20" box="[151,256,1847,1870]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4AFFBF94EEE75EFED45D20" box="[151,256,1847,1870]" italics="true" pageId="6" pageNumber="31">Juniperus</emphasis>
</taxonomicName>
type as cupressaceous wood with mixed 1-2 seriate radial pitting on tracheids, rare or absent parenchyma, and usually uniseriate rays with cupressoid cross-fields in vertical pairs and juniperoid nodules on the tangential walls of the ray cells. Evaluating these details comparatively with those included in Table 1 from
<bibRefCitation id="D8204B621D4AFFBF96A9E7B2FEB9527A" author="Ruiz, D. P. &amp; Bodnar, J." pageId="6" pageNumber="31" pagination="481 - 488" refId="ref16844" refString="Ruiz, D. P. and Bodnar, J., 2019. The oldest record of Juniperoxylon, a cupressaceous fossil wood from the Middle Triassic of Argentina. Acta Palaeontologica Polonica, 64 (3): 481 - 488." type="journal article" year="2019">Ruiz and Bodnar (2019)</bibRefCitation>
, and using the identification key of
<bibRefCitation id="D8204B621D4AFFBF9749E0F7FC335AD8" author="Akkemik, U." box="[816,999,158,182]" pageId="6" pageNumber="31" pagination="15 - 26" refId="ref12580" refString="Akkemik, U., 2021. A new species of Juniperoxylon from the Early Miocene of Northwestern Turkey. Acta Palaeontologica Romaniae, 17 (1): 15 - 26. https: // doi. org / 10.35463 / j. apr. 2021.01.02" type="journal article" year="2021">Akkemik (2021)</bibRefCitation>
, we observe that our specimens are not identical to any described species. Thus, we describe a new species named here
<taxonomicName id="7BB14D101D4AFFBF903FE089FA485A96" authority="Iamandei &amp; Iamandei &amp; Grădinaru, 2022" authorityName="Iamandei &amp; Iamandei &amp; Grădinaru" authorityYear="2022" box="[1094,1436,224,248]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="holbavicum" status="sp. nov.">
<emphasis id="8EC5EA811D4AFFBF903FE089FA485A96" box="[1094,1436,224,248]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon holbavicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95F657FA1D4AFFBF9749E16BFC5F5B77" box="[816,907,258,281]" pageId="6" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
The designated
<typeStatus id="630A88311D4AFFBF9035E168FB7D5B77" box="[1100,1193,257,281]" pageId="6" pageNumber="31" type="holotype">holotype</typeStatus>
is the sample 27706 (inventory number), kept in “Grădinaru Collection”, hosted by National Geological Museum, from
<collectingRegion id="7E75F8711D4AFFBF9158E12AFA5B5B35" box="[1313,1423,323,347]" country="Romania" name="Bucuresti" pageId="6" pageNumber="31">Bucharest</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D4AFFBF9749E1EFFAE058A9" pageId="6" pageNumber="31" type="diagnosis">
<paragraph id="BC0E36931D4AFFBF9749E1EFFBDB5BD1" blockId="6.[816,1437,390,711]" pageId="6" pageNumber="31">
<emphasis id="8EC5EA811D4AFFBF9749E1EFFA485BF0" bold="true" box="[816,1436,390,414]" pageId="6" pageNumber="31">
The diagnosis of the species
<taxonomicName id="7BB14D101D4AFFBF90BCE1EFFA485BF0" authority="Iamandei &amp; Iamandei &amp; Grădinaru, 2022" authorityName="Iamandei &amp; Iamandei &amp; Grădinaru" authorityYear="2022" box="[1221,1436,390,414]" pageId="6" pageNumber="31" status="sp. nov.">
<emphasis id="8EC5EA811D4AFFBF90BCE1EFFA485BF0" bold="true" box="[1221,1436,390,414]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon</emphasis>
</taxonomicName>
</emphasis>
<emphasis id="8EC5EA811D4AFFBF9749E1CEFBDB5BD1" bold="true" box="[816,1039,423,447]" pageId="6" pageNumber="31">
<emphasis id="8EC5EA811D4AFFBF9749E1CEFC655BD1" bold="true" box="[816,945,423,447]" italics="true" pageId="6" pageNumber="31">holbavicum</emphasis>
<taxonomicNameLabel id="95F657FA1D4AFFBF97C1E1CEFBDB5BD1" box="[952,1039,423,447]" pageId="6" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="BC0E36931D4AFFBF9749E1A1FAE058A9" blockId="6.[816,1437,390,711]" pageId="6" pageNumber="31">
Secondary tracheidoxylic wood, with thick-walled tracheids with uniseriate pitting, or even biseriate of mixed
<typeStatus id="630A88311D4AFFBF9749E262FCB5584D" box="[816,865,523,547]" pageId="6" pageNumber="31">type</typeStatus>
, parenchyma absent or few and with nodular horizontal walls, heterogeneous uniseriate rays with juniperoid nodules on tangential walls or ray cells, cross-fields with 1-2(4-6) small cupressoid pits per field, as 1-2 vertical pairs pits and when more, slightly irregular or alternate, and having slit-like inclined apertures.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:classification id="0DC538CDD6D32B648FB9570CB5999867">journal article</mods:classification>
<mods:identifier id="1D2D65377AEB48B5BB938C71EBA999CA" type="DOI">10.35463/j.apr.2022.01.04</mods:identifier>
<mods:identifier id="F8A11A95F34C529D31B67EAD58ECC795" type="ISSN">1842-371x</mods:identifier>
<mods:identifier id="50108E5A19F4A32EFA073C5CF86C4C2D" type="Zenodo-Dep">10520851</mods:identifier>
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<subSubSection id="F4AB65181D4AFFBF972EE37AFB695922" pageId="6" pageNumber="31" type="nomenclature">
<paragraph id="BC0E36931D4AFFBF972EE37AFB695922" blockId="6.[855,1397,787,877]" pageId="6" pageNumber="31">
<taxonomicName id="7BB14D101D4AFFBF972EE37AFB695922" authority="Iamandei, Iamandei and Gradinaru, 2018" authorityName="Iamandei, Iamandei and Gradinaru" authorityYear="2018" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="holbavicum">
<emphasis id="8EC5EA811D4AFFBF972EE37AFBBA5945" box="[855,1134,787,811]" italics="true" pageId="6" pageNumber="31">Brachyoxylon holbavicum</emphasis>
<bibRefCitation id="D8204B621D4AFFBF900CE37AFB695922" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." pageId="6" pageNumber="31" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei, Iamandei and Grădinaru, 2018</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D4AFFBF9056E33CFB4A5903" box="[1071,1182,853,877]" pageId="6" pageNumber="31" type="description">
<paragraph id="BC0E36931D4AFFBF9056E33CFB4A5903" blockId="6.[855,1397,787,877]" box="[1071,1182,853,877]" pageId="6" pageNumber="31">
<figureCitation id="248A2A161D4AFFBF9056E33CFBBB5903" box="[1071,1135,853,877]" captionStart="Fig" captionStartId="7.[181,216,1943,1964]" captionTargetBox="[181,1432,154,1935]" captionTargetId="figure-19@7.[181,1432,154,1935]" captionTargetPageId="7" captionText="Fig. 3 Brachyoxylon holbavicum Iamandei, Iamandei and Grădinaru, 2018. (inventory no. 27711). a-c: Cross section - tracheids polygonal with rounded corners, and linear rays; d-f: Tangential section - pitting on tracheids - absent; rays uniseriate; g-i: Radial section radial pitting of mixed type on tracheids, pits hexagonal and, when biseriate, alternate; cross-fields with 2-3(4) cupressoid to araucarioid in 1 row or, when more numerous, in 2-3 superposed rows." figureDoi="http://doi.org/10.5281/zenodo.10520857" httpUri="https://zenodo.org/record/10520857/files/figure.png" pageId="6" pageNumber="31">Fig. 3</figureCitation>
, a-i.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D4AFFBF9749E3FEFA8F5F75" pageId="6" pageNumber="31" type="materials_examined">
<paragraph id="BC0E36931D4AFFBF9749E3FEFC4759C1" blockId="6.[816,1437,919,1307]" box="[816,915,919,943]" pageId="6" pageNumber="31">
<emphasis id="8EC5EA811D4AFFBF9749E3FEFC4759C1" bold="true" box="[816,915,919,943]" pageId="6" pageNumber="31">Material</emphasis>
</paragraph>
<paragraph id="BC0E36931D4AFFBF9749E3D1FA8F5F75" blockId="6.[816,1437,919,1307]" pageId="6" pageNumber="31">
<materialsCitation id="0CD93CCE1D4AFFBF9749E3D1FA835F75" location="National Museum of Geology" pageId="6" pageNumber="31" specimenCount="1" stateProvince="Bucuresti">
The studied material is represented by 7 samples of silicified wood collected from the Holbav locality, on Maiului brook. All the studied samples are fragments of trunk or thick branches, with decimetric size and dark to black colored. By hand lens or even by naked eye, the regular fibrous structure without vessels suggests a conifer wood. The specimens, with the following field numbers: 1089, 1090, 1093, 1094, 1096, 1102, 1105, belong to &quot;Grădinaru Collection&quot;, and are stored at the
<location id="B96E60481D4AFFBF90A2E4A8FC7B5E94" LSID="urn:lsid:plazi:treatment:341887851D4AFFB1972EE37AFBAB5964:B96E60481D4AFFBF90A2E4A8FC7B5E94" name="National Museum of Geology" pageId="6" pageNumber="31" stateProvince="Bucuresti">National Museum of Geology</location>
, in
<collectingRegion id="7E75F8711D4AFFBF97A5E48BFB9D5E94" box="[988,1097,1250,1274]" country="Romania" name="Bucuresti" pageId="6" pageNumber="31">Bucharest</collectingRegion>
, under the inventory numbers: 27711, 27712, 27713, 27714, 27715, 27716, 27718
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D4AFFB19749E52FFDCB59E0" lastPageId="8" lastPageNumber="33" pageId="6" pageNumber="31" type="description">
<paragraph id="BC0E36931D4AFFBF9749E52FFB955F30" blockId="6.[816,1437,1350,2035]" box="[816,1089,1350,1374]" pageId="6" pageNumber="31">
<emphasis id="8EC5EA811D4AFFBF9749E52FFB955F30" bold="true" box="[816,1089,1350,1374]" pageId="6" pageNumber="31">Microscopic description</emphasis>
</paragraph>
<paragraph id="BC0E36931D4AFFBF9749E50FFC615FAF" blockId="6.[816,1437,1350,2035]" pageId="6" pageNumber="31">
<emphasis id="8EC5EA811D4AFFBF9749E50FFC6B5F10" box="[816,959,1382,1406]" italics="true" pageId="6" pageNumber="31">Growth rings</emphasis>
well developed, with tracheidoxylic structure, abrupt or gradual transition, distinct boundaries, no resin canals.
</paragraph>
<paragraph id="BC0E36931D4AFFBF9749E5A3FA585DBC" blockId="6.[816,1437,1350,2035]" pageId="6" pageNumber="31">
<emphasis id="8EC5EA811D4AFFBF9749E5A3FC4F5F8C" box="[816,923,1482,1506]" italics="true" pageId="6" pageNumber="31">Tracheids</emphasis>
with polygonal cross section, usually quadrangular with rather thick and slightly corrugated walls in the early-wood, diameters of 20-40(60) μm and wall thickness of 5-7(8) μm (double wall), quite similar in the late-wood. Between two successive rays, 1-12 radial regular rows tracheids appear and intercellular spaces are present. Density is 836-1520 tracheids per square mm. On tangential walls the pitting is usually absent. The radial pits are of mixed
<typeStatus id="630A88311D4AFFBF9051E6BAFB8D5C85" box="[1064,1113,1747,1771]" pageId="6" pageNumber="31">type</typeStatus>
, either uniseriate, with round pits, spaced or contiguous and variably flattened (flattening index: d/D=0.79-0.88), or biseriate, with round to oval pits, of 19-24 μm the diameter, with round to oblique-elliptic apertures of 7-10/3-5 μm, in opposite or alternate arrangement, sometimes contiguous, combined with short uniseriate portions. There are no crassulae or helical thickenings, but sometimes striations are present.
</paragraph>
<paragraph id="BC0E36931D4AFFBF9749E7B2FBA65D9D" blockId="6.[816,1437,1350,2035]" box="[816,1138,2011,2035]" pageId="6" pageNumber="31">
<emphasis id="8EC5EA811D4AFFBF9749E7B2FC275D9D" box="[816,1011,2011,2035]" italics="true" pageId="6" pageNumber="31">Axial parenchyma</emphasis>
is absent.
</paragraph>
<caption id="E8CE661B1D4BFFBE94CCE7FEFB1F5268" ID-DOI="http://doi.org/10.5281/zenodo.10520857" ID-Zenodo-Dep="10520857" httpUri="https://zenodo.org/record/10520857/files/figure.png" pageId="7" pageNumber="32" startId="7.[181,216,1943,1964]" targetBox="[181,1432,154,1935]" targetPageId="7" targetType="figure">
<paragraph id="BC0E36931D4BFFBE94CCE7FEFBB15DC3" blockId="7.[181,1433,1943,2054]" box="[181,1125,1943,1965]" pageId="7" pageNumber="32">
<emphasis id="8EC5EA811D4BFFBE94CCE7FEFF3A5DC2" bold="true" box="[181,238,1943,1964]" pageId="7" pageNumber="32">Fig. 3</emphasis>
<taxonomicName id="7BB14D101D4BFFBE948CE7FEFC575DC3" authority="Iamandei, Iamandei and Gradinaru, 2018" authorityName="Iamandei, Iamandei and Gradinaru" authorityYear="2018" box="[245,899,1943,1965]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="32" phylum="Tracheophyta" rank="species" species="holbavicum">
<emphasis id="8EC5EA811D4BFFBE948CE7FEFE245DC3" box="[245,496,1943,1965]" italics="true" pageId="7" pageNumber="32">Brachyoxylon holbavicum</emphasis>
<bibRefCitation id="D8204B621D4BFFBE958FE7FEFC575DC3" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." box="[502,899,1943,1965]" pageId="7" pageNumber="32" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei, Iamandei and Grădinaru, 2018</bibRefCitation>
</taxonomicName>
. (inventory no. 27711).
</paragraph>
<paragraph id="BC0E36931D4BFFBE94CCE7DCFB1F5268" blockId="7.[181,1433,1943,2054]" pageId="7" pageNumber="32">
<emphasis id="8EC5EA811D4BFFBE94CCE7DCFF005DA4" bold="true" box="[181,212,1973,1994]" pageId="7" pageNumber="32">a-c</emphasis>
: Cross section - tracheids polygonal with rounded corners, and linear rays;
<emphasis id="8EC5EA811D4BFFBE97CFE7DCFC075DA4" bold="true" box="[950,979,1973,1994]" pageId="7" pageNumber="32">d-f</emphasis>
: Tangential section - pitting on tracheids - absent; rays uniseriate;
<emphasis id="8EC5EA811D4BFFBE95FAE7BAFE4A5D86" bold="true" box="[387,414,2003,2024]" pageId="7" pageNumber="32">g-i</emphasis>
: Radial section radial pitting of mixed type on tracheids, pits hexagonal and, when biseriate, alternate; cross-fields with 2-3(4) cupressoid to araucarioid in 1 row or, when more numerous, in 2-3 superposed rows.
</paragraph>
</caption>
<paragraph id="BC0E36931D44FFB194EEE0F6FD8A5922" blockId="8.[151,771,158,910]" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB194EEE0F6FF1F5AD8" box="[151,203,159,182]" italics="true" pageId="8" pageNumber="33">Rays</emphasis>
in cross-section appear thin, with linear trajectory, from rectangular cells radially elongated, with smooth unpitted horizontal walls. Tangentially they appear exclusively uniseriate, with 2-17(-25) cells in height, sometimes more. And, sometimes, the taller rays have 1-2 short biseriate storeys of the same thickness as a single cell. Lateral empty spaces commonly appear. Ray-density is of 7-12 rays per tangential horizontal millimeter. The rays are homogeneous, with parenchymal cells all procumbent, of 20-24 μm in height, in the marginal rows taller, of 28-35(-50) μm. The horizontal walls are moderately thick walled, 4-6 μm for the double wall (sometimes thicker), and the outer wall of the marginals is slightly corrugated. Tangential wall is relatively thick. Indentures are not visible. The cross-fields have 1-6 cupressoid pits, tending to podocarpoid, usually hexagonal-rounded to oval, with diameters of 9-15(18) / 6-8(10) μm and with tilted elliptic apertures of 5-8 / 3-5 μm. In normal fields the pits arrangement is 1-3(-4) pits in horizontal row and, in the taller rays, 4-6(-9) pits in 1-2(3) horizontal rows, alternately or slightly irregularly arranged.
</paragraph>
<paragraph id="BC0E36931D44FFB194EEE33CFD9A5903" blockId="8.[151,771,158,910]" box="[151,590,853,877]" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB194EEE33CFE155903" box="[151,449,853,877]" italics="true" pageId="8" pageNumber="33">Radial or axial resin canals</emphasis>
are absent.
</paragraph>
<paragraph id="BC0E36931D44FFB194EEE31FFDCB59E0" blockId="8.[151,771,158,910]" box="[151,543,886,910]" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB194EEE31FFE8B59E0" box="[151,351,886,910]" italics="true" pageId="8" pageNumber="33">Mineral inclusions</emphasis>
are not present.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D44FFB194EEE3D1FBAB5964" pageId="8" pageNumber="33" type="discussion">
<paragraph id="BC0E36931D44FFB194EEE3D1FE7859BE" blockId="8.[151,772,952,2068]" box="[151,428,952,976]" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB194EEE3D1FE7859BE" bold="true" box="[151,428,952,976]" pageId="8" pageNumber="33">Affinities and discussion</emphasis>
</paragraph>
<paragraph id="BC0E36931D44FFB194EEE3B0FE6E5ED6" blockId="8.[151,772,952,2068]" pageId="8" pageNumber="33">
All the 7 samples of fossil wood studied here present the next xylotomy: typical tracheidoxylic structure with thick-walled tracheids, having radial pitting of mixed
<typeStatus id="630A88311D44FFB194EEE454FF1C5E3B" box="[151,200,1085,1109]" pageId="8" pageNumber="33">type</typeStatus>
, with rays mostly uniseriate showing cupressoid cross-fields (tending to araucarioid). All these xylotomical details are consistent with those of the genus
<taxonomicName id="7BB14D101D44FFB196D7E416FE6E5ED6" authority="Hollick and Jeffrey." authorityName="Hollick and Jeffrey." class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="33" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D44FFB196D7E416FF0A5ED6" italics="true" pageId="8" pageNumber="33">Brachyoxylon</emphasis>
Hollick and Jeffrey.
</taxonomicName>
</paragraph>
<paragraph id="BC0E36931D44FFB194EEE4A8FF085F8C" blockId="8.[151,772,952,2068]" pageId="8" pageNumber="33">
This genus was created by
<bibRefCitation id="D8204B621D44FFB195A8E4A8FD2A5EB7" author="Hollick, A. &amp; Jeffrey, E. C." box="[465,766,1217,1241]" pageId="8" pageNumber="33" pagination="1 - 137" refId="ref15032" refString="Hollick, A. and Jeffrey, E. C., 1909. Studies of coniferous Cretaceous remains from Kreischerville (New-York). Memoirs of the New York Botanical Garden, 3: 1 - 137." type="journal article" year="1909">Hollick and Jeffrey (1909)</bibRefCitation>
, having as
<typeStatus id="630A88311D44FFB19570E48AFEE35E95" box="[265,311,1251,1275]" pageId="8" pageNumber="33">type</typeStatus>
species
<taxonomicName id="7BB14D101D44FFB195EEE48BFF375F72" authority="Hollick et Jeffrey" authorityName="Hollick et Jeffrey" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="33" phylum="Tracheophyta" rank="species" species="notabile">
<emphasis id="8EC5EA811D44FFB195EEE48BFD585E94" box="[407,652,1250,1274]" italics="true" pageId="8" pageNumber="33">Brachyoxylon notabile</emphasis>
Hollick et Jeffrey
</taxonomicName>
, with a very elliptic diagnosis: “tracheidoxyl devoid of normal secretory ducts, with radial pitting of mixed
<typeStatus id="630A88311D44FFB1949AE52FFEC05F33" box="[227,276,1350,1373]" pageId="8" pageNumber="33">type</typeStatus>
, araucarioid cross-fields and other ray walls, integer” (see
<bibRefCitation id="D8204B621D44FFB1955EE50FFE115F10" author="Philippe, M." box="[295,453,1382,1406]" pageId="8" pageNumber="33" pagination="74 - 80" refId="ref16155" refString="Philippe, M., 1993. Nomenclature generique des tracheidoxyles mesozoiques a champs araucarioides. Taxon, 42: 74 - 80." type="journal article" year="1993">Philippe, 1993</bibRefCitation>
; Philipe and Bamford, 2008). The occasionally presence of traumatic canals is considered as a result of injury, or of freezing (see Phillipe, 1995).
</paragraph>
<paragraph id="BC0E36931D44FFB194EEE582FE5C5CC6" blockId="8.[151,772,952,2068]" pageId="8" pageNumber="33">
In order to identify the specimens studied here at species level, we evaluated other fossil forms previously described. We noted that some species are no longer considered to be
<taxonomicName id="7BB14D101D44FFB195C1E627FD7E5C08" box="[440,682,1614,1638]" pageId="8" pageNumber="33">
<emphasis id="8EC5EA811D44FFB195C1E627FD985C08" box="[440,588,1614,1638]" italics="true" pageId="8" pageNumber="33">Brachyoxylon</emphasis>
species
</taxonomicName>
, or are doubtful, so they should be excluded from discussion (see
<bibRefCitation id="D8204B621D44FFB194EEE6F9FEA85CC6" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." box="[151,380,1680,1704]" pageId="8" pageNumber="33" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei et al., 2018</bibRefCitation>
).
</paragraph>
<paragraph id="BC0E36931D44FFB194EEE6D8FE1E5DBC" blockId="8.[151,772,952,2068]" pageId="8" pageNumber="33">
There are many forms of
<taxonomicName id="7BB14D101D44FFB195CCE6D8FD9D5CA7" box="[437,585,1713,1737]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="33" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D44FFB195CCE6D8FD9D5CA7" box="[437,585,1713,1737]" italics="true" pageId="8" pageNumber="33">Brachyoxylon</emphasis>
</taxonomicName>
described showing a large spread of this Mesozoic genus on the planet and we have quoted and discussed, in detail, some of them (in
<bibRefCitation id="D8204B621D44FFB1957CE77CFE2A5D43" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." box="[261,510,1813,1837]" pageId="8" pageNumber="33" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei et al., 2018</bibRefCitation>
). They were described from
<collectingCountry id="C4A676031D44FFB194ADE75EFEC55D20" box="[212,273,1847,1870]" name="Japan" pageId="8" pageNumber="33">Japan</collectingCountry>
(
<bibRefCitation id="D8204B621D44FFB19559E75FFE3C5D20" author="Iijima, A. &amp; Kakuwa, Y. &amp; Matsuda, H." box="[288,488,1846,1870]" pageId="8" pageNumber="33" refId="ref15297" refString="Iijima, A., Kakuwa, Y. and Matsuda, H., 1989. Silicified Wood from the Adoyama Chert, Kuzuh, Central Honshu, and Its Bearing on Compaction and Depositional Environment of Radiolarian Bedded Chert. In: Hein J. R., Obradovic J. (eds.), Siliceous Deposits of the Tethys and Pacific Regions. Springer, New York." type="book" year="1989">Iijima et al., 1989</bibRefCitation>
), from
<collectingCountry id="C4A676031D44FFB19640E75FFD5F5D20" box="[569,651,1846,1870]" name="Mexico" pageId="8" pageNumber="33">Mexico</collectingCountry>
(
<bibRefCitation id="D8204B621D44FFB196E2E75FFECB5D01" author="Cevallos-Ferriz, S. R. S." pageId="8" pageNumber="33" pagination="111 - 137" refId="ref13402" refString="Cevallos-Ferriz, S. R. S., 1992. Tres Maderas de Gimnospermas Cretacicas del Norte de Mexico. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico, Serie Botanica, 63 (2): 111 - 137." type="journal article" year="1992">Cevallos-Ferriz, 1992</bibRefCitation>
), from
<collectingCountry id="C4A676031D44FFB19508E73EFE725D01" box="[369,422,1879,1903]" name="United States of America" pageId="8" pageNumber="33">USA</collectingCountry>
(
<bibRefCitation id="D8204B621D44FFB195CFE73EFD585D01" author="Hickey, L. J. &amp; Hu, S. &amp; Skinner, B. J." box="[438,652,1879,1903]" pageId="8" pageNumber="33" pagination="608 - 631" refId="ref14947" refString="Hickey, L. J., Hu, S. and Skinner, B. J., 2011. A New genus of silicified conifer wood from the Late Triassic of Connecticut. American Journal of Science, 311 (7): 608 - 631." type="journal article" year="2011">Hickey et al., 2011</bibRefCitation>
), from
<collectingCountry id="C4A676031D44FFB196A4E73EFF325DFE" name="Argentina" pageId="8" pageNumber="33">Argentina</collectingCountry>
(
<bibRefCitation id="D8204B621D44FFB1948FE711FE245DFE" author="Vera, E. I. &amp; Cesari, S. N." box="[246,496,1912,1936]" pageId="8" pageNumber="33" pagination="617 - 625" refId="ref17664" refString="Vera, E. I. and Cesari, S. N., 2012. Fossil woods (Coniferales) from the Baquero Group (Aptian), Santa Cruz Province, Argentina. Anais da Academia Brasileira de Ciencias (Annals of the Brazilian Academy of Sciences), 84 (3): 617 - 625." type="journal article" year="2012">Vera and Césari, 2012</bibRefCitation>
;
<bibRefCitation id="D8204B621D44FFB19584E711FD075DFE" author="Bodnar, J. &amp; Escapa, I. &amp; Cuneo, N. R. &amp; Gnaedinger, S." box="[509,723,1912,1936]" pageId="8" pageNumber="33" pagination="227 - 239" refId="ref13053" refString="Bodnar, J., Escapa, I., Cuneo, N. R., Gnaedinger, S., 2013. First record of conifer wood from the Canadon Asfalto Formation (Early-Middle Jurassic), Chubut Province, Argentina. Ameghiniana, 50: 227 - 239." type="journal article" year="2013">Bodnar et al., 2013</bibRefCitation>
), all of them having various specific xylotomical details, partially similar to our material.
</paragraph>
<paragraph id="BC0E36931D44FFB194EEE7B5FBE25BD1" blockId="8.[151,772,952,2068]" lastBlockId="8.[816,1436,158,778]" pageId="8" pageNumber="33">
But the European species
<taxonomicName id="7BB14D101D44FFB195AEE7B2FD045D9D" box="[471,720,2011,2035]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="33" phylum="Tracheophyta" rank="species" species="saurinii">
<emphasis id="8EC5EA811D44FFB195AEE7B2FD045D9D" box="[471,720,2011,2035]" italics="true" pageId="8" pageNumber="33">Brachyoxylon saurinii</emphasis>
</taxonomicName>
described by
<bibRefCitation id="D8204B621D44FFB1956BE794FDE4527A" author="Boureau, E. &amp; Serra, C." box="[274,560,2044,2068]" pageId="8" pageNumber="33" pagination="373 - 393" refId="ref13267" refString="Boureau, E. and Serra, C., 1961. Sur les structures normales et traumatiques d'un bois mesozoique du Cambodge, le Brachyoxylon saurinii, n. sp. Revue generale de botanique, 68: 373 - 393." type="journal article" year="1961">Boureau and Serra (1961)</bibRefCitation>
,
<taxonomicName id="7BB14D101D44FFB19646E794FD67527A" box="[575,691,2045,2068]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="33" phylum="Tracheophyta" rank="species" species="avramii">
<emphasis id="8EC5EA811D44FFB19646E794FD67527A" box="[575,691,2045,2068]" italics="true" pageId="8" pageNumber="33">B. avramii</emphasis>
</taxonomicName>
and
<taxonomicName id="7BB14D101D44FFB19694E794FC6A5AD8" box="[749,958,158,2068]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="33" phylum="Tracheophyta" rank="species" species="dobrogiacum">
<emphasis id="8EC5EA811D44FFB19694E794FC6A5AD8" box="[749,958,158,2068]" italics="true" pageId="8" pageNumber="33">B. dobrogiacum</emphasis>
</taxonomicName>
described by
<bibRefCitation id="D8204B621D44FFB190FAE0F7FCAE5AB9" author="Iamandei, E. &amp; Iamandei, S." pageId="8" pageNumber="33" pagination="231 - 247" refId="ref15205" refString="Iamandei, E. and Iamandei, S., 2005. Early Cretaceous protopinaceous fossil wood from South Dobrogea, Romania. Acta Palaeontologica Romaniae, 5: 231 - 247." type="journal article" year="2005">Iamandei and Iamandei (2005)</bibRefCitation>
,
<taxonomicName id="7BB14D101D44FFB197FEE0A9FC3F5AB9" authorityName="Philippe, Suteethorn &amp; Buffetaut" authorityYear="2011" box="[903,1003,192,215]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="33" phylum="Tracheophyta" rank="species" species="serrae">
<emphasis id="8EC5EA811D44FFB197FEE0A9FC3F5AB9" box="[903,1003,192,215]" italics="true" pageId="8" pageNumber="33">B. serrae</emphasis>
</taxonomicName>
described by Phillipe et al. (2011) and, especially
<emphasis id="8EC5EA811D44FFB197DDE088FB935A96" box="[932,1095,224,248]" italics="true" pageId="8" pageNumber="33">
<taxonomicName id="7BB14D101D44FFB197DDE088FBEB5A96" box="[932,1087,224,248]" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="33" phylum="Tracheophyta" rank="species" species="holbavicum">B. holbavicum</taxonomicName>
,
</emphasis>
described by us from the same area (
<bibRefCitation id="D8204B621D44FFB19712E168FB9A5B77" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." box="[875,1102,257,281]" pageId="8" pageNumber="33" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei et al., 2018</bibRefCitation>
) are closer to our studied specimens. And, related to this, the debate on Mesozoic palaeoenvironment of large European forests made by
<bibRefCitation id="D8204B621D44FFB19112E12AFC0F5B12" author="Garcia, J. - P. &amp; Philippe, M. &amp; Gaumet, F." pageId="8" pageNumber="33" pagination="199 - 214" refId="ref14545" refString="Garcia, J. - P., Philippe, M. and Gaumet, F., 1998. Fossil wood in Middle-Upper Jurassic marine sedimentary cycles of France: relations with climate, sea-level dynamics, and carbonate-platform environments. Palaeogeography, Palaeoclimatology, Palaeoecology, 141 (3 - 4): 199 - 214." type="journal article" year="1998">Garcia et al. (1998)</bibRefCitation>
is very interesting and, also, the complex biogeographic analysis of Gondwanian Mesozoic forests of
<bibRefCitation id="D8204B621D44FFB19734E1CEFBE65BD1" author="Philippe, M. &amp; Bamford, M. &amp; McLoughlin, S. &amp; Alves, L. S. R. &amp; Falcon Lang, H. J. &amp; Gnaedinger, S. &amp; Ottone, E. G. &amp; Pole, M. &amp; Rajanikanthi, A. &amp; Shoemaker, R. E. &amp; Torres, T. &amp; Zamuner, A." box="[845,1074,423,447]" pageId="8" pageNumber="33" pagination="141 - 173" refId="ref16274" refString="Philippe, M., Bamford, M., McLoughlin, S., Alves, L. S. R., Falcon Lang, H. J., Gnaedinger, S., Ottone, E. G., Pole, M., Rajanikanthi, A., Shoemaker, R. E., Torres, T. and Zamuner, A., 2004. Biogeographic analysis of Jurassic - Early Cretaceous wood assemblages from Gondwana. Review of Palaeobotany and Palynology, 129: 141 - 173." type="journal article" year="2004">Philippe et al. (2004)</bibRefCitation>
.
</paragraph>
<paragraph id="BC0E36931D44FFB19749E1A1FBAB5964" blockId="8.[816,1436,158,778]" pageId="8" pageNumber="33">
In summary, taking into account the comparative table (see Table 1, in
<bibRefCitation id="D8204B621D44FFB1979EE180FB05586F" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." box="[999,1233,489,513]" pageId="8" pageNumber="33" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei et al., 2018</bibRefCitation>
) and the synthetic description of the specimens studied here (mixed radial pitting on tracheids, axial parenchyma absent, uniseriate rays medium-tall, cross-fields with cupressoid pits, tending to podocarpoid, in quassi-araucarioid arrangement), we consider our samples to perfectly fit to the diagnosis of
<taxonomicName id="7BB14D101D44FFB1972EE2C6FC4B5887" authority="Iamandei, Iamandei and Gradinaru" authorityName="Iamandei, Iamandei and Gradinaru" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="33" phylum="Tracheophyta" rank="species" species="holbavicum">
<emphasis id="8EC5EA811D44FFB1972EE2C6FBAC58A9" box="[855,1144,687,711]" italics="true" pageId="8" pageNumber="33">Brachyoxylon holbavicum</emphasis>
Iamandei, Iamandei and Grădinaru
</taxonomicName>
, a species previously identified in the same area (see
<bibRefCitation id="D8204B621D44FFB197ECE29BFBA75964" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." box="[917,1139,754,778]" pageId="8" pageNumber="33" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei et al. 2018</bibRefCitation>
).
</paragraph>
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</document>

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<taxonomicName id="7BB14D101D4DFFB894DBE455FE2A5E1B" authority="Iamandei and Iamandei, 2000" authorityName="Iamandei and Iamandei" authorityYear="2000" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="26" phylum="Tracheophyta" rank="species" species="dragastanii">
<emphasis id="8EC5EA811D4DFFB894DBE455FDDA5E3A" box="[162,526,1084,1108]" italics="true" pageId="1" pageNumber="26">Protocupressinoxylon dragastanii</emphasis>
<bibRefCitation id="D8204B621D4DFFB8966CE455FE2A5E1B" author="Iamandei E. &amp; Iamandei S." pageId="1" pageNumber="26" pagination="191 - 199" refId="ref15071" refString="Iamandei E. and Iamandei S., 2000. Bois fossiles de Coniferes dans l'Aptien de Dobroudja de Sud, Romania. Acta Palaeontologica Romaniae, 2 (1999): 191 - 199." type="journal article" year="2000">Iamandei and Iamandei, 2000</bibRefCitation>
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<figureCitation id="248A2A161D4DFFB895EFE416FE005EF9" box="[406,468,1151,1175]" captionStart="Fig" captionStartId="2.[183,218,1925,1946]" captionTargetBox="[183,1404,154,1916]" captionTargetId="figure-30@2.[183,1404,154,1916]" captionTargetPageId="2" captionText="Fig. 1 Protocupressinoxylon dragastanii Iamandei and Iamandei, 2000 (inventory no. 27699). a-c: Cross section - tracheids polygonal with rounded corners, linear rays and diffuse parenchyma; d-f: Tangential section - uniseriate rays with polygonal ray-cells, sometimes with biseriate storeys; g-i: Radial section - radial pitting on tracheids 1-2 - seriate, of mixed-type; when biseriate, pits are alternate (g, i); cross-fields with 1-2(4) cupressoid pits, when more - are slightly irregular or alternate, in two superposed rows (g, h)." figureDoi="http://doi.org/10.5281/zenodo.10520853" httpUri="https://zenodo.org/record/10520853/files/figure.png" pageId="1" pageNumber="26">Fig. 1</figureCitation>
, a-i.
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<subSubSection id="F4AB65181D4DFFB894EEE4A8FDFA5CC6" pageId="1" pageNumber="26" type="materials_examined">
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<emphasis id="8EC5EA811D4DFFB894EEE4A8FF2E5EB7" bold="true" box="[151,250,1217,1241]" pageId="1" pageNumber="26">Material</emphasis>
</paragraph>
<paragraph id="BC0E36931D4DFFB894EEE48BFDFA5CC6" blockId="1.[151,771,1217,1704]" pageId="1" pageNumber="26">
The studied material is represented by 14 samples of silicified wood collected from the Holbav area. All studied samples are fragments of trunk or thick-branches, of decimetric size, dark to black color and by magnifying glass or even by naked eye, the regular fibrous structure without vessels suggesting a conifer wood. The described specimens attributed to this taxon have very similar details and have the following field-numbers within the &quot;Grădinaru Collection&quot;: 1074, 1077, 1082, 1084, 1086, 1088, 1092, 1097, 1098, 1099, 1100, 1104, 1106, 1109 and are stored at the National Museum of Geology, in
<collectingRegion id="7E75F8711D4DFFB894EEE627FED05C08" box="[151,260,1614,1638]" country="Romania" name="Bucuresti" pageId="1" pageNumber="26">Bucharest</collectingRegion>
, under the inventory numbers: 27691, 27692, 27693, 27694, 27695, 27696, 27697, 27698, 27699, 27700, 27701, 27702, 277203, 27720.
</paragraph>
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<subSubSection id="F4AB65181D4DFFB894EEE6BAFAD85DDF" pageId="1" pageNumber="26" type="description">
<paragraph id="BC0E36931D4DFFB894EEE6BAFE7C5C85" blockId="1.[151,771,1747,2068]" box="[151,424,1747,1771]" pageId="1" pageNumber="26">
<emphasis id="8EC5EA811D4DFFB894EEE6BAFE7C5C85" bold="true" box="[151,424,1747,1771]" pageId="1" pageNumber="26">Microscopic description</emphasis>
</paragraph>
<paragraph id="BC0E36931D4DFFB894EEE69DFEF85DBC" blockId="1.[151,771,1747,2068]" pageId="1" pageNumber="26">
<emphasis id="8EC5EA811D4DFFB894EEE69DFEFD5D62" box="[151,297,1780,1804]" italics="true" pageId="1" pageNumber="26">Growth rings</emphasis>
in the studied samples represent secondary xylem, which is tracheidoxylic. Thus, in cross-section they appear more or less well developed, sometimes wide with more than 60 cells, showing a fairly abrupt or gradual transition from early- to late-wood and distinct boundaries of the growth-rings. Normal or traumatic axial resin canals absent.
</paragraph>
<paragraph id="BC0E36931D4DFFB894EEE7B2FC3A5E7C" blockId="1.[151,771,1747,2068]" lastBlockId="1.[816,1437,158,1969]" pageId="1" pageNumber="26">
<emphasis id="8EC5EA811D4DFFB894EEE7B2FED65D9D" box="[151,258,2011,2035]" italics="true" pageId="1" pageNumber="26">Tracheids</emphasis>
in cross-section appear large and with relatively thin walls in the early-wood, having quasi-polygonal cross-section (4-5 sides), and are sometimes deformed by compression. They have radial/tangential diameters of 30-60(70) / 25-60(70) μm and the wall thickness of 4-7 μm double wall. Where gradual size transition is present, a well-developed transitional wood is visible, with tracheids gradually diminishing their section and tracheidal wall becoming progressively thicker, until the final 3-5 rows of the late-wood, which consist of smaller radially flattened cells (with r/tg diameters of 20- 25 / 10-20 μm) and thicker walled, of 8-10 μm across the double-wall, marking the growth-ring boundary. Between two successive rays, 1-9 radial rows of tracheids can be counted. Sometimes, radial rows of smaller-sized tracheids are intermingled. The density is between 750-1216 tracheids / mm
<superScript id="4BC49BDB1D4DFFB897ACE202FC0A5817" attach="right" box="[981,990,619,633]" fontSize="6" pageId="1" pageNumber="26">2</superScript>
. In longitudinal view, tracheids have on the radial walls 1-2-seriate pits, sometimes contiguous, and of mixed
<typeStatus id="630A88311D4DFFB897BCE2D9FC2158A6" box="[965,1013,688,712]" pageId="1" pageNumber="26">type</typeStatus>
, because when are biseriate the pits are often alternately arranged. Sometimes crassulae are present. The pits are of abietinean
<typeStatus id="630A88311D4DFFB890E9E29BFB155964" box="[1168,1217,754,778]" pageId="1" pageNumber="26">type</typeStatus>
, have 10-12 μm diameter and round to oval apertures, of 5-6 μm. No details about torus. On the tangential walls, pits are usually absent, or rarely present, as poorly preserved, uniseriate, small round pits, of 5-6 μm in diameter, with small round to oval aperture. Inside the tracheids, organic deposits are sometimes present as dark remains. Average tracheid length is difficult to measure. Helical thickenings on tracheids are absent.
</paragraph>
<paragraph id="BC0E36931D4DFFB89749E472FAB35ED6" blockId="1.[816,1437,158,1969]" pageId="1" pageNumber="26">
<emphasis id="8EC5EA811D4DFFB89749E472FC205E5D" box="[816,1012,1051,1075]" italics="true" pageId="1" pageNumber="26">Axial parenchyma</emphasis>
in the cross sections is usually absent or rarely present, as dispersed thin-walled cells with dark content. In longitudinal view it appears as rows of rectangular cells, with the horizontal wall thin and smooth, rarely feebly nodular and with some dark remains inside.
</paragraph>
<paragraph id="BC0E36931D4DFFB89749E4ABFB8A5D01" blockId="1.[816,1437,158,1969]" pageId="1" pageNumber="26">
<emphasis id="8EC5EA811D4DFFB89749E4ABFCB05EB7" box="[816,868,1218,1241]" italics="true" pageId="1" pageNumber="26">Rays</emphasis>
usually appear thin in cross section while in tangential sections appear exclusively uniseriate, sometimes with some biseriate storeys, having up to 15 cells in height, sometimes more. Their density is 6-9 rays per tangential millimeter. In radial view the rays show homocellular aspect, with parenchymal cells all procumbent, 18-20 μm tall, marginals slightly taller, of 25-30 μm. The horizontal ray-cell walls are smooth or slightly rough and usually pitted, and are relatively thin (2-3.5 μm double wall); the tangential end-walls are relatively thick, of 1.5- 2 μm the simple wall, smooth or slightly nodular, but usually poorly preserved. The indentures are missing or difficult to notice. Spiral thickenings were not observed, most probably are absent. The cross-field pitting is of cupressoid
<typeStatus id="630A88311D4DFFB897D2E6F8FC085CC7" box="[939,988,1681,1705]" pageId="1" pageNumber="26">type</typeStatus>
, showing 1-2(4-5) pits, in 1-2 rows horizontally arranged, and sometimes irregular or even to slightly alternate aspect. The pits are oculipores of cupressoid
<typeStatus id="630A88311D4DFFB897E8E69DFC155D62" box="[913,961,1780,1804]" pageId="1" pageNumber="26">type</typeStatus>
, 8-10 μm in diameter, with slit- like or lenslike apertures of 2-5 μm. Sometimes cross- field pitting is difficult to observe, due to poor preservation or to the presence of organic content.
</paragraph>
<paragraph id="BC0E36931D4DFFB89749E711FB3C5DFE" blockId="1.[816,1437,158,1969]" box="[816,1256,1912,1936]" pageId="1" pageNumber="26">
<emphasis id="8EC5EA811D4DFFB89749E711FB8F5DFE" box="[816,1115,1912,1936]" italics="true" pageId="1" pageNumber="26">Radial or axial resin canals</emphasis>
are absent.
</paragraph>
<paragraph id="BC0E36931D4DFFB89749E7F0FAD85DDF" blockId="1.[816,1437,158,1969]" box="[816,1292,1945,1969]" pageId="1" pageNumber="26">
<emphasis id="8EC5EA811D4DFFB89749E7F0FC2D5DDF" box="[816,1017,1945,1969]" italics="true" pageId="1" pageNumber="26">Mineral inclusions</emphasis>
usually are not present.
</paragraph>
</subSubSection>
<caption id="E8CE661B1D4EFFBB94CEE7ECFD79527F" ID-DOI="http://doi.org/10.5281/zenodo.10520853" ID-Zenodo-Dep="10520853" httpUri="https://zenodo.org/record/10520853/files/figure.png" pageId="2" pageNumber="27" startId="2.[183,218,1925,1946]" targetBox="[183,1404,154,1916]" targetPageId="2" targetType="figure">
<paragraph id="BC0E36931D4EFFBB94CEE7ECFB945DF4" blockId="2.[183,1405,1924,2065]" box="[183,1088,1924,1946]" pageId="2" pageNumber="27">
<emphasis id="8EC5EA811D4EFFBB94CEE7ECFF255DF4" bold="true" box="[183,241,1925,1946]" pageId="2" pageNumber="27">Fig. 1</emphasis>
<taxonomicName id="7BB14D101D4EFFBB948EE7EDFCB65DF4" authority="Iamandei and Iamandei, 2000" authorityName="Iamandei and Iamandei" authorityYear="2000" box="[247,866,1924,1946]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="27" phylum="Tracheophyta" rank="species" species="dragastanii">
<emphasis id="8EC5EA811D4EFFBB948EE7EDFDEA5DF4" box="[247,574,1924,1946]" italics="true" pageId="2" pageNumber="27">Protocupressinoxylon dragastanii</emphasis>
<bibRefCitation id="D8204B621D4EFFBB963CE7EDFCB65DF4" author="Iamandei E. &amp; Iamandei S." box="[581,866,1924,1946]" pageId="2" pageNumber="27" pagination="191 - 199" refId="ref15071" refString="Iamandei E. and Iamandei S., 2000. Bois fossiles de Coniferes dans l'Aptien de Dobroudja de Sud, Romania. Acta Palaeontologica Romaniae, 2 (1999): 191 - 199." type="journal article" year="2000">Iamandei and Iamandei, 2000</bibRefCitation>
</taxonomicName>
(inventory no. 27699).
</paragraph>
<paragraph id="BC0E36931D4EFFBB94CEE7CAFD79527F" blockId="2.[183,1405,1924,2065]" pageId="2" pageNumber="27">
<emphasis id="8EC5EA811D4EFFBB94CEE7CAFF0A5DD6" bold="true" box="[183,222,1955,1976]" pageId="2" pageNumber="27">a-c:</emphasis>
Cross section - tracheids polygonal with rounded corners, linear rays and diffuse parenchyma;
<emphasis id="8EC5EA811D4EFFBB90F4E7CAFB665DD6" bold="true" box="[1165,1202,1955,1976]" pageId="2" pageNumber="27">d-f:</emphasis>
Tangential section - uniseriate rays with polygonal ray-cells, sometimes with biseriate storeys;
<emphasis id="8EC5EA811D4EFFBB97F0E7A9FC785DBB" bold="true" box="[905,940,1984,2005]" pageId="2" pageNumber="27">g-i:</emphasis>
Radial section - radial pitting on tracheids 1-2 - seriate, of mixed-type; when biseriate, pits are alternate (g, i); cross-fields with 1-2(4) cupressoid pits, when more - are slightly irregular or alternate, in two superposed rows (g, h).
</paragraph>
</caption>
<subSubSection id="F4AB65181D4FFFBD94EEE0F7FE61582D" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28" type="discussion">
<paragraph id="BC0E36931D4FFFBA94EEE0F7FE785AD8" blockId="3.[151,772,158,2068]" box="[151,428,158,182]" pageId="3" pageNumber="28">
<emphasis id="8EC5EA811D4FFFBA94EEE0F7FE785AD8" bold="true" box="[151,428,158,182]" pageId="3" pageNumber="28">Affinities and discussion</emphasis>
</paragraph>
<paragraph id="BC0E36931D4FFFBA94EEE0D6FF3158A9" blockId="3.[151,772,158,2068]" pageId="3" pageNumber="28">
The xylotomy of the here studied
<specimenCount id="AAB7FD1A1D4FFFBA966FE0D6FD7E5AB9" box="[534,682,191,215]" count="14" pageId="3" pageNumber="28" type="generic">14 specimens</specimenCount>
is definitely pycnoxylic and homoxylic (or tracheidoxylic
<emphasis id="8EC5EA811D4FFFBA94EEE16BFF065B77" box="[151,210,258,281]" italics="true" pageId="3" pageNumber="28">sensu</emphasis>
<bibRefCitation id="D8204B621D4FFFBA94A5E168FEB95B77" author="Creber, G. T." box="[220,365,257,281]" pageId="3" pageNumber="28" pagination="655 - 661" refId="ref13670" refString="Creber, G. T., 1972. Gymnospermous wood from the Kimmeridgian of East Sutherland and from the Sandringham Sands of Norfolk. Palaeontology, 15 (4): 655 - 661." type="journal article" year="1972">Creber, 1972</bibRefCitation>
), having tracheids with radial pitting of mixed
<typeStatus id="630A88311D4FFFBA957CE14AFEE15B55" box="[261,309,291,315]" pageId="3" pageNumber="28">type</typeStatus>
, no resin canals, little or no parenchyma, exclusively uniseriate rays and cross-fields with 1-2(4-5) cupressoid pits with slit-like inclined apertures, arranged in one or two horizontal rows, which are slightly irregularly to even alternately arranged; all these characters are evidence of a typical cupressaceous structure. Such details as: radial pitting on tracheids of mixed
<typeStatus id="630A88311D4FFFBA96E6E183FD1B586C" box="[671,719,490,514]" pageId="3" pageNumber="28">type</typeStatus>
, cupressoid pits in cross-fields, to fairly araucarioid arrangement, axial parenchyma showing few and radial pitting of tracheids of mixed
<typeStatus id="630A88311D4FFFBA959DE224FDC6580B" box="[484,530,589,613]" pageId="3" pageNumber="28">type</typeStatus>
in a Mesozoic wood structure indicate a wood-type from “
<taxonomicName id="7BB14D101D4FFFBA9639E204FCD758EB" box="[576,771,621,645]" class="Pinopsida" family="Cheirolepidiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="family">Cheirolepidiaceae</taxonomicName>
family”, as defined by
<bibRefCitation id="D8204B621D4FFFBA95D1E2E7FD5758C8" author="Alvin, K. L. &amp; Fraser, C. J. &amp; Spicer, R. A." box="[424,643,654,678]" pageId="3" pageNumber="28" pagination="759 - 778" refId="ref12656" refString="Alvin, K. L., Fraser, C. J. and Spicer, R. A., 1981. Anatomy and paleoecology of Pseudofrenelopsis and associated conifers in the English Wealden. Palaeontology, 24: 759 - 778." type="journal article" year="1981">Alvin et al. (1981)</bibRefCitation>
and
<bibRefCitation id="D8204B621D4FFFBA96BFE2E7FF3558A9" author="Alvin, K. L." pageId="3" pageNumber="28" pagination="71 - 98" refId="ref12627" refString="Alvin, K. L., 1982. Cheirolepidiaceae: Biology, structure and paleoecology. Review of Palaeobotany and Palynology, 37: 71 - 98." type="journal article" year="1982">Alvin (1982)</bibRefCitation>
.
</paragraph>
<paragraph id="BC0E36931D4FFFBA94EEE2B8FECD5E1B" blockId="3.[151,772,158,2068]" pageId="3" pageNumber="28">
The synthetic description of all the studied specimens was compared with the diagnoses of Mesozoic genera with cupressoid oculipores in the cross-fields as described by
<bibRefCitation id="D8204B621D4FFFBA94C7E35DFEB05922" author="Phillips, E. W. J." box="[190,356,820,844]" pageId="3" pageNumber="28" pagination="259 - 320" refId="ref16431" refString="Phillips, E. W. J., 1941. The identification of coniferous woods by their microscopic structure. Journal of the Linnean Society of London, Botany, 52: 259 - 320." type="journal article" year="1941">Phillips (1941)</bibRefCitation>
,
<bibRefCitation id="D8204B621D4FFFBA950CE35DFDCA5922" author="Krausel, R." box="[373,542,820,844]" pageId="3" pageNumber="28" pagination="83 - 203" refId="ref15485" refString="Krausel, R., 1949. Die fossilen Koniferen-Holzer (Unter Ausschluss von Araucarioxylon Kraus). II: Kritische Untersuchungenzur Diagnostik lebender und fossiler Koniferen-Holzer. Palaeontographica, Abt. B, 89: 83 - 203." type="journal article" year="1949">Kräusel (1949)</bibRefCitation>
,
<bibRefCitation id="D8204B621D4FFFBA9657E35DFD2A5922" author="Vogellehner, D." box="[558,766,820,844]" pageId="3" pageNumber="28" pagination="30 - 51" refId="ref17724" refString="Vogellehner, D., 1967. Zur Anatomie und Phylogenie de Mesozoischer Gymnospermenholzer, 7: Prodromus zueiner Monographie de Protopinaceae. I. Die Protopinoiden Holzer des Trias. Palaeontographica Abt. B, 121 (1 - 3): 30 - 51." type="journal article" year="1967">Vogellehner (1967</bibRefCitation>
,
<bibRefCitation id="D8204B621D4FFFBA94EEE33CFF045903" author="Vogellehner, D." box="[151,208,853,877]" pageId="3" pageNumber="28" pagination="125 - 162" refId="ref17771" refString="Vogellehner, D., 1968. Zur Anatomie und Phylogenie de Mesozoischer Gymnospermenholzer, 7: Prodromus zu einer Monographie de Protopinaceae. II. Die Protopinoiden Holzer des Jura. Palaeontographica Abt. B, Stuttgart, 124 (4 - 6): 125 - 162." type="journal article" year="1968">1968</bibRefCitation>
), Müller Stoll and Schultze-Motel (1989),
<bibRefCitation id="D8204B621D4FFFBA96D3E33CFF0C59E0" author="Philippe, M." pageId="3" pageNumber="28" pagination="45 - 103" refId="ref16179" refString="Philippe, M., 1995. Bois fossils du Jurassique de Franche-Comte (NE-France). Palaeontographica Abt. B, 236: 45 - 103." type="journal article" year="1995">Philippe (1995</bibRefCitation>
,
<bibRefCitation id="D8204B621D4FFFBA949DE31FFEC959E0" author="Philippe, M." box="[228,285,886,910]" pageId="3" pageNumber="28" pagination="319 - 326" refId="ref16207" refString="Philippe, M., 2002. Reappraisal of five genera designed for fossil coniferous woods by early American wood anatomists. IAWA Journal, 23: 319 - 326." type="journal article" year="2002">2002</bibRefCitation>
),
<bibRefCitation id="D8204B621D4FFFBA9548E31FFE0659E0" author="Bodnar, J." box="[305,466,886,910]" pageId="3" pageNumber="28" pagination="1480 - 1481" refId="ref13105" refString="Bodnar, J., 2017. (2575) Proposal to conserve the name Protocupressinoxylon with a conserved type against Protobrachyoxylon (fossil plants). Taxon (Journal of IAPT), 66 (6): 1480 - 1481. https: // doi. org / 10.12705 / 666.25" type="journal article" year="2017">Bodnar (2017)</bibRefCitation>
, and it appears that our material is mostly consistent, at generic level, with
<taxonomicName id="7BB14D101D4FFFBA96DCE3F1FEFF59BE" authorityName="Eckhold" authorityYear="1923" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4FFFBA96DCE3F1FEFF59BE" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon</emphasis>
</taxonomicName>
wood-type. In fact, the Mesozoic conifers were grouped, a long time ago, based on anatomical features, in the &quot;
<taxonomicName id="7BB14D101D4FFFBA9552E393FE115E7C" box="[299,453,1018,1042]" class="Pinopsida" family="Protopinaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="family">Protopinaceae</taxonomicName>
family&quot; (
<bibRefCitation id="D8204B621D4FFFBA9649E393FD1D5E7C" author="Krausel, R." box="[560,713,1018,1042]" pageId="3" pageNumber="28" pagination="305 - 311" refId="ref15456" refString="Krausel, R., 1917. Die Bedeutung der Anatomielebender und fossiler Holzer fur die Phylogenie der Koniferen. Naturwissenschaften Wochenschrift, 16: 305 - 311." type="journal article" year="1917">Krausel, 1917</bibRefCitation>
This family name was noted by
<bibRefCitation id="D8204B621D4FFFBA95B9E472FD785E5D" author="Bamford, M. K. &amp; Philippe, M. &amp; Thevenard, F." box="[448,684,1051,1075]" pageId="3" pageNumber="28" pagination="25 - 30" refId="ref12825" refString="Bamford, M. K., Philippe, M. and Thevenard, F., 2016. Long overdue extinction of the Protopinaceae. Review of Palaeobotany and Palynology, 234: 25 - 30." type="journal article" year="2016">Bamford et al. (2016)</bibRefCitation>
as having a dubious taxonomic status, and they recommended its abandon.
</paragraph>
<paragraph id="BC0E36931D4FFFBA94EEE416FDEC5F8C" blockId="3.[151,772,158,2068]" pageId="3" pageNumber="28">
However, the name
<taxonomicName id="7BB14D101D4FFFBA9516E416FDE65EF9" box="[367,562,1151,1175]" class="Pinopsida" family="Cheirolepidiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="family">Cheirolepidiaceae</taxonomicName>
, attributed to Turutanova-Ketova (see Fossil works online), is not a valid name, as remarked by
<bibRefCitation id="D8204B621D4FFFBA95F3E4A8FDFB5EB7" author="Doweld, A. B." box="[394,559,1217,1241]" pageId="3" pageNumber="28" pagination="1092 - 1098" refId="ref14259" refString="Doweld, A. B., 2020. The controversial nomenclature of the fossil plant names Cheirolepis, Cheirolepidium and Hirmeriella (Cheirolepidaceae / Cheirolepidiaceae / Hirmeriellaceae). 69 (5): 1092 - 1098." type="journal article" year="2020">Doweld (2020)</bibRefCitation>
, who considered an alternative name as being more correct for the Mesozoic gymnosperms characterized by pollen of
<taxonomicName id="7BB14D101D4FFFBA960EE56AFD2F5F75" authorityName="H.D.Pflug" authorityYear="1953" box="[631,763,1283,1307]" class="Pinopsida" family="Cheirolepidiaceae" genus="Classopollis" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4FFFBA960EE56AFD2F5F75" box="[631,763,1283,1307]" italics="true" pageId="3" pageNumber="28">Classopollis</emphasis>
</taxonomicName>
-
<typeStatus id="630A88311D4FFFBA94EEE54CFF115F53" box="[151,197,1317,1341]" pageId="3" pageNumber="28">type</typeStatus>
. Thus, he applied the valid family name
<taxonomicName id="7BB14D101D4FFFBA96EBE54DFEBD5F33" authority="Harris, 1979" authorityName="Harris" authorityYear="1979" family="Hirmeriellaceae" pageId="3" pageNumber="28" rank="family">
Hirmeriellaceae
<bibRefCitation id="D8204B621D4FFFBA949BE52CFEBD5F33" author="Harris, T. M." box="[226,361,1349,1373]" pageId="3" pageNumber="28" refId="ref14877" refString="Harris, T. M., 1979. The Yorkshire Jurassic Flora. 5. Coniferales. British Museum Natural History, London, 167 pp." type="book" year="1979">Harris, 1979</bibRefCitation>
</taxonomicName>
- and proposed this family name “for conservation against
<taxonomicName id="7BB14D101D4FFFBA9506E50FFD905F10" box="[383,580,1382,1406]" class="Pinopsida" family="Cheirolepidiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="family">Cheirolepidiaceae</taxonomicName>
”, the latter being considered as &quot;inadmissible, being based on the illegitimate later homonym
<taxonomicName id="7BB14D101D4FFFBA95EDE5C0FD5C5FAF" authority="Schimper" authorityName="Schimper" box="[404,648,1449,1473]" class="Magnoliopsida" family="Asteraceae" genus="Cheirolepis" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4FFFBA95EDE5C0FDC45FAF" box="[404,528,1449,1473]" italics="true" pageId="3" pageNumber="28">Cheirolepis</emphasis>
Schimper
</taxonomicName>
, and thus, itself illegitimate” (see
<bibRefCitation id="D8204B621D4FFFBA95E9E5A3FDF85F8C" author="Doweld, A. B." box="[400,556,1482,1506]" pageId="3" pageNumber="28" pagination="1092 - 1098" refId="ref14259" refString="Doweld, A. B., 2020. The controversial nomenclature of the fossil plant names Cheirolepis, Cheirolepidium and Hirmeriella (Cheirolepidaceae / Cheirolepidiaceae / Hirmeriellaceae). 69 (5): 1092 - 1098." type="journal article" year="2020">Doweld, 2020</bibRefCitation>
).
</paragraph>
<paragraph id="BC0E36931D4FFFBA94EEE582FEF05D62" blockId="3.[151,772,158,2068]" pageId="3" pageNumber="28">
This genus name,
<taxonomicName id="7BB14D101D4FFFBA9519E582FD9E5C6D" authorityName="Eckhold" authorityYear="1923" box="[352,586,1515,1539]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4FFFBA9519E582FD9E5C6D" box="[352,586,1515,1539]" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon</emphasis>
</taxonomicName>
, was established by
<bibRefCitation id="D8204B621D4FFFBA94C5E665FEB45C4A" author="Eckhold, W." box="[188,352,1548,1572]" pageId="3" pageNumber="28" pagination="472 - 505" refId="ref14328" refString="Eckhold, W., 1922. Die Hoftupfel bei rezenten und fossilen Coniferen. Jahrbuch Konigliche Preussische Geologische Landesansalt, 42: 472 - 505." type="journal article" year="1922">Eckhold (1922</bibRefCitation>
, pp. 490-491), to designate Mesozoic fossil conifer wood-types described like this: “wood with annual rings more or less distinct, tracheid pitting in various transitional arrangements, both horizontal and end walls of ray parenchyma cells smooth, resin canals absent, axial parenchyma occasionally present, cross-field pits cupressoid”, but not specifying their number and arrangement.
</paragraph>
<paragraph id="BC0E36931D4FFFBA94EEE77CFCA25B8E" blockId="3.[151,772,158,2068]" lastBlockId="3.[816,1437,158,2068]" pageId="3" pageNumber="28">
Initially,
<bibRefCitation id="D8204B621D4FFFBA9486E77CFE7F5D43" author="Eckhold, W." box="[255,427,1813,1837]" pageId="3" pageNumber="28" pagination="472 - 505" refId="ref14328" refString="Eckhold, W., 1922. Die Hoftupfel bei rezenten und fossilen Coniferen. Jahrbuch Konigliche Preussische Geologische Landesansalt, 42: 472 - 505." type="journal article" year="1922">Eckhold (1922)</bibRefCitation>
included seven species in the protologue:
<emphasis id="8EC5EA811D4FFFBA9563E75FFF255DFE" italics="true" pageId="3" pageNumber="28">
<taxonomicName id="7BB14D101D4FFFBA9563E75FFD4C5D20" authorityName="Eckhold" authorityYear="1922" baseAuthorityName="Holden" baseAuthorityYear="1913" box="[282,664,1846,1870]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="cupressoides">Protocupressinoxylon cupressoides</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA96DDE75EFF0C5D01" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="jurassicum">P. jurassicum</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA9491E731FE4E5D01" box="[232,410,1880,1903]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="potomacense">P. potomacense</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA95D3E731FD9D5D01" box="[426,585,1879,1903]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="eboracense">P. eboracense</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA9620E731FD0F5D01" box="[601,731,1879,1903]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="koettlitzi">P. koettlitzi</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA9692E731FF255DFE" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="vectense">P. vectense</taxonomicName>
</emphasis>
and
<taxonomicName id="7BB14D101D4FFFBA955EE710FE135DFE" box="[295,455,1913,1936]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="mesozoicum">
<emphasis id="8EC5EA811D4FFFBA955EE710FE135DFE" box="[295,455,1913,1936]" italics="true" pageId="3" pageNumber="28">P. mesozoicum</emphasis>
</taxonomicName>
, but without designation of a clear generotype. Only later,
<bibRefCitation id="D8204B621D4FFFBA95AEE7F0FD5C5DDF" author="Andrews, H. N., Jr." box="[471,648,1945,1969]" pageId="3" pageNumber="28" pagination="1 - 262" refId="ref12700" refString="Andrews, H. N., Jr. 1955. Index of generic names of fossil plants, 1820 - 1950. [U. S.] Geological Survey Bulletin, 1013: 1 - 262." type="journal article" year="1955">Andrews (1955)</bibRefCitation>
designated
<taxonomicName id="7BB14D101D4FFFBA94EEE7D3FF195D9D" authority="(Holden) Eckhold, 1922" authorityName="Eckhold" authorityYear="1922" baseAuthorityName="Holden" baseAuthorityYear="1913" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="cupressoides">
<emphasis id="8EC5EA811D4FFFBA94EEE7D3FDC95DBC" box="[151,541,1978,2002]" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon cupressoides</emphasis>
(Holden)
<bibRefCitation id="D8204B621D4FFFBA96DDE7D3FF195D9D" author="Eckhold, W." pageId="3" pageNumber="28" pagination="472 - 505" refId="ref14328" refString="Eckhold, W., 1922. Die Hoftupfel bei rezenten und fossilen Coniferen. Jahrbuch Konigliche Preussische Geologische Landesansalt, 42: 472 - 505." type="journal article" year="1922">Eckhold, 1922</bibRefCitation>
</taxonomicName>
(not 1923, see Jogmans and Dijkstra, 1974, p. 794) - as the species-type for that generic name. However, the species
<taxonomicName id="7BB14D101D4FFFBA97E8E0F7FADE5AD8" box="[913,1290,158,182]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="eboracense">
<emphasis id="8EC5EA811D4FFFBA97E8E0F7FADE5AD8" box="[913,1290,158,182]" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon eboracense</emphasis>
</taxonomicName>
which was initially assigned as type-species of the
<taxonomicName id="7BB14D101D4FFFBA9091E0D6FC865A96" authorityName="R.Holden" authorityYear="1913" class="Pinopsida" family="Araucariaceae" genus="Protobrachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4FFFBA9091E0D6FC865A96" italics="true" pageId="3" pageNumber="28">Protobrachyoxylon</emphasis>
</taxonomicName>
genus (
<bibRefCitation id="D8204B621D4FFFBA97C3E089FB8D5A96" author="Holden, R." box="[954,1113,224,248]" pageId="3" pageNumber="28" pagination="33 - 549" refId="ref14996" refString="Holden, R., 1913. Contributions to the Anatomy of Mesozoic Conifers. 1. Jurassic coniferous woods from Yorkshire. Annals of Botany, 27 (5): 33 - 549." type="journal article" year="1913">Holden, 1913</bibRefCitation>
) was included by
<bibRefCitation id="D8204B621D4FFFBA913AE089FCAC5B77" author="Eckhold, W." pageId="3" pageNumber="28" refId="ref14298" refString="Eckhold, W., 1921. Die Hoftupfel bei rezenten und fossilen Koniferen. [PhD Thesis] Breslau Schlesische Fridrich-Wilhelms Universitat (Univ. of Wroclaw Poland)" type="book" year="1921">Eckhold (1921)</bibRefCitation>
along with other species in the protologue of
<taxonomicName id="7BB14D101D4FFFBA9115E16BFC265B54" authorityName="Eckhold" authorityYear="1923" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4FFFBA9115E16BFC265B54" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon</emphasis>
</taxonomicName>
genus. But the original material seems to be lost, and the analysis of Holdens description and figuration suggest either identity with
<taxonomicName id="7BB14D101D4FFFBA90B5E10CFBD05BF3" authority="Eckhold, 1921" authorityName="Eckhold" authorityYear="1921" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4FFFBA90B5E10CFC865BF3" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon</emphasis>
<bibRefCitation id="D8204B621D4FFFBA9718E1ECFBD05BF3" author="Eckhold, W." box="[865,1028,389,413]" pageId="3" pageNumber="28" refId="ref14298" refString="Eckhold, W., 1921. Die Hoftupfel bei rezenten und fossilen Koniferen. [PhD Thesis] Breslau Schlesische Fridrich-Wilhelms Universitat (Univ. of Wroclaw Poland)" type="book" year="1921">Eckhold, 1921</bibRefCitation>
</taxonomicName>
or with
<taxonomicName id="7BB14D101D4FFFBA900CE1ECFC145BD1" authority="Hollick and Jeffrey, 1909" authorityName="Hollick and Jeffrey" authorityYear="1909" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4FFFBA900CE1ECFADD5BF3" box="[1141,1289,389,413]" italics="true" pageId="3" pageNumber="28">Brachyoxylon</emphasis>
<bibRefCitation id="D8204B621D4FFFBA9161E1ECFC145BD1" author="Hollick, A. &amp; Jeffrey, E. C." pageId="3" pageNumber="28" pagination="1 - 137" refId="ref15032" refString="Hollick, A. and Jeffrey, E. C., 1909. Studies of coniferous Cretaceous remains from Kreischerville (New-York). Memoirs of the New York Botanical Garden, 3: 1 - 137." type="journal article" year="1909">Hollick and Jeffrey, 1909</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="D8204B621D4FFFBA9784E1CEFAFB5BD1" author="Bamford, M. K. &amp; Philippe, M." box="[1021,1327,423,447]" pageId="3" pageNumber="28" pagination="287 - 297" refId="ref12781" refString="Bamford, M. K. and Philippe, M., 2001. Jurassic - Early Cretaceous Gondwanan homoxylous wood: a nomenclatural revision of the genera with taxonomic notes. Review of Palaeobotany and Palynology, 113: 287 - 297." type="journal article" year="2001">Bamford and Philippe 2001</bibRefCitation>
,
<bibRefCitation id="D8204B621D4FFFBA9145E1CEFCBE5B8E" author="Philippe, M." pageId="3" pageNumber="28" pagination="319 - 326" refId="ref16207" refString="Philippe, M., 2002. Reappraisal of five genera designed for fossil coniferous woods by early American wood anatomists. IAWA Journal, 23: 319 - 326." type="journal article" year="2002">Philippe, 2002</bibRefCitation>
).
</paragraph>
<paragraph id="BC0E36931D4FFFBA9749E180FACF5964" blockId="3.[816,1437,158,2068]" pageId="3" pageNumber="28">
Recently,
<bibRefCitation id="D8204B621D4FFFBA97D9E180FB94586F" author="Bodnar, J." box="[928,1088,489,513]" pageId="3" pageNumber="28" pagination="1480 - 1481" refId="ref13105" refString="Bodnar, J., 2017. (2575) Proposal to conserve the name Protocupressinoxylon with a conserved type against Protobrachyoxylon (fossil plants). Taxon (Journal of IAPT), 66 (6): 1480 - 1481. https: // doi. org / 10.12705 / 666.25" type="journal article" year="2017">Bodnar (2017)</bibRefCitation>
proposed the genus name
<taxonomicName id="7BB14D101D4FFFBA9115E183FC26584C" authorityName="Eckhold" authorityYear="1921" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4FFFBA9115E183FC26584C" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon</emphasis>
</taxonomicName>
to be conserved, together with a proposed generotype
<taxonomicName id="7BB14D101D4FFFBA907EE245FA8E582D" authority="Roggeveen, 1932" authorityName="Roggeveen" authorityYear="1932" box="[1031,1370,555,580]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="malayense">
<emphasis id="8EC5EA811D4FFFBA907EE245FB40582D" box="[1031,1172,555,579]" italics="true" pageId="3" pageNumber="28">P. malayense</emphasis>
<bibRefCitation id="D8204B621D4FFFBA90E2E245FA8E582D" author="Roggeveen, P. M." box="[1179,1370,555,580]" pageId="3" pageNumber="28" pagination="580 - 584" refId="ref16743" refString="Roggeveen, P. M., 1932. Mesozoisches Koniferenholz (Protocupressinoxylon malayense n. s.) von der Insel Soegi im Riouw-Archipel. Niederlandisch Ost Indien. Proceedings of the Section of Science, Koninklijke (Nederlandse) Akademie van Wetenschappente Amsterdam, 35: 580 - 584." type="journal article" year="1932">Roggeveen, 1932</bibRefCitation>
</taxonomicName>
. Also, she proposed the necessity of a systematic revision of several described taxa, since abnormal details were described for some species of
<bibRefCitation id="D8204B621D4FFFBA9000E2E7FAF758C8" author="Krausel, R." box="[1145,1315,654,678]" pageId="3" pageNumber="28" pagination="83 - 203" refId="ref15485" refString="Krausel, R., 1949. Die fossilen Koniferen-Holzer (Unter Ausschluss von Araucarioxylon Kraus). II: Kritische Untersuchungenzur Diagnostik lebender und fossiler Koniferen-Holzer. Palaeontographica, Abt. B, 89: 83 - 203." type="journal article" year="1949">Kräusel (1949)</bibRefCitation>
, like resin ducts in
<taxonomicName id="7BB14D101D4FFFBA97F5E2C6FAD358A9" box="[908,1287,687,711]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="potomacense">
<emphasis id="8EC5EA811D4FFFBA97F5E2C6FAD358A9" box="[908,1287,687,711]" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon potomacense</emphasis>
</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA916FE2D9FAAC58A9" box="[1302,1400,687,711]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="solmsi">
<emphasis id="8EC5EA811D4FFFBA916FE2D9FAAC58A9" box="[1302,1400,687,711]" italics="true" pageId="3" pageNumber="28">P. solmsi</emphasis>
</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA91FFE2D9FC5E5887" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="vectense">
<emphasis id="8EC5EA811D4FFFBA91FFE2D9FC5E5887" italics="true" pageId="3" pageNumber="28">P. vectense</emphasis>
</taxonomicName>
,
<emphasis id="8EC5EA811D4FFFBA97E2E2BBFB9F5887" box="[923,1099,721,745]" italics="true" pageId="3" pageNumber="28">
<taxonomicName id="7BB14D101D4FFFBA97E2E2BBFB935887" box="[923,1095,721,745]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="luccombense">P. luccombense</taxonomicName>
,
</emphasis>
or multiseriate rays, like in
<taxonomicName id="7BB14D101D4FFFBA91FFE2BBFC6F5964" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="weidlingense">
<emphasis id="8EC5EA811D4FFFBA91FFE2BBFC6F5964" italics="true" pageId="3" pageNumber="28">P. weidlingense</emphasis>
</taxonomicName>
(in
<bibRefCitation id="D8204B621D4FFFBA979EE29BFBAB5964" author="Krausel, R." box="[999,1151,754,778]" pageId="3" pageNumber="28" pagination="83 - 203" refId="ref15485" refString="Krausel, R., 1949. Die fossilen Koniferen-Holzer (Unter Ausschluss von Araucarioxylon Kraus). II: Kritische Untersuchungenzur Diagnostik lebender und fossiler Koniferen-Holzer. Palaeontographica, Abt. B, 89: 83 - 203." type="journal article" year="1949">Kräusel, 1949</bibRefCitation>
, pp. 183-185).
</paragraph>
<paragraph id="BC0E36931D4FFFBA9749E37AFC3459BE" blockId="3.[816,1437,158,2068]" pageId="3" pageNumber="28">
For this genus many species of fossil wood from all over the world ranging from Permian to Cretaceous have been described, especially after
<bibRefCitation id="D8204B621D4FFFBA9037E33CFB245903" author="Krausel, R." box="[1102,1264,853,877]" pageId="3" pageNumber="28" pagination="83 - 203" refId="ref15485" refString="Krausel, R., 1949. Die fossilen Koniferen-Holzer (Unter Ausschluss von Araucarioxylon Kraus). II: Kritische Untersuchungenzur Diagnostik lebender und fossiler Koniferen-Holzer. Palaeontographica, Abt. B, 89: 83 - 203." type="journal article" year="1949">Kräusel (1949)</bibRefCitation>
, who admited it as valid genus, and he even has described two species:
<taxonomicName id="7BB14D101D4FFFBA91FCE31EFBAA59C1" authority="(Torrey) Krausel" authorityName="Krausel" baseAuthorityName="Torrey" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="dockumense">
<emphasis id="8EC5EA811D4FFFBA91FCE31EFC6759C1" italics="true" pageId="3" pageNumber="28">P. dockumense</emphasis>
(Torrey) Kräusel
</taxonomicName>
and
<taxonomicName id="7BB14D101D4FFFBA90BBE3F1FC3459BE" authority="(Sahni) Krausel." authorityName="Krausel." baseAuthorityName="Sahni" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="coromandelinum">
<emphasis id="8EC5EA811D4FFFBA90BBE3F1FA4959C1" box="[1218,1437,919,943]" italics="true" pageId="3" pageNumber="28">P. coromandelinum</emphasis>
(Sahni) Kräusel.
</taxonomicName>
</paragraph>
<paragraph id="BC0E36931D4FFFBA9749E3B3FC085EB7" blockId="3.[816,1437,158,2068]" pageId="3" pageNumber="28">
Later, new European species were described, as for example:
<taxonomicName id="7BB14D101D4FFFBA97F5E393FCB25E5D" authority="Schultze-Motel 1960" authorityName="Schultze-Motel" authorityYear="1960" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="catenatum">
<emphasis id="8EC5EA811D4FFFBA97F5E393FB3A5E7C" box="[908,1262,1018,1042]" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon catenatum</emphasis>
<bibRefCitation id="D8204B621D4FFFBA9081E393FCB25E5D" author="Schultze-Motel, J." pageId="3" pageNumber="28" pagination="1 - 15" refId="ref17004" refString="Schultze-Motel, J., 1960. Gymnospermen Holzer aus den Oberen Kreide von Quedlinburg. Palaontologische Zeitschrift, 34: 1 - 15." type="journal article" year="1960">Schultze-Motel 1960</bibRefCitation>
</taxonomicName>
(see also
<bibRefCitation id="D8204B621D4FFFBA97A9E472FAC45E5D" author="Iamandei, S. &amp; Iamandei, E. &amp; Tibuleac, P." box="[976,1296,1051,1075]" pageId="3" pageNumber="28" pagination="211 - 218" refId="ref15107" refString="Iamandei, S., Iamandei, E. and Tibuleac, P., 2001. Fossil Wood from Coal-layer &quot; B &quot; of Volhinian Formation in Leucusesti - Falticeni area. Analele Stiintifice ale Universitatii &quot; Al. I. Cuza &quot; (s. n.), Geologie, 47: 211 - 218." type="journal article" year="2001">Iamandei and Iamandei, 2001</bibRefCitation>
),
<taxonomicName id="7BB14D101D4FFFBA915AE475FBC85E3A" authority="Schultze-Motel 1960" authorityName="Schultze-Motel" authorityYear="1960" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="liasinum">
<emphasis id="8EC5EA811D4FFFBA915AE475FA485E5D" box="[1315,1436,1051,1075]" italics="true" pageId="3" pageNumber="28">P. liasinum</emphasis>
<bibRefCitation id="D8204B621D4FFFBA9749E455FBC85E3A" author="Schultze-Motel, J." box="[816,1052,1084,1108]" pageId="3" pageNumber="28" pagination="1 - 15" refId="ref17004" refString="Schultze-Motel, J., 1960. Gymnospermen Holzer aus den Oberen Kreide von Quedlinburg. Palaontologische Zeitschrift, 34: 1 - 15." type="journal article" year="1960">Schultze-Motel 1960</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA9056E454FC0B5E1B" authority="(Charrier) Vogellehner 1968" authorityName="Vogellehner" authorityYear="1968" baseAuthorityName="Charrier" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="leonardianum">
<emphasis id="8EC5EA811D4FFFBA9056E454FB3F5E3A" box="[1071,1259,1084,1108]" italics="true" pageId="3" pageNumber="28">P. leonardianum</emphasis>
(Charrier)
<bibRefCitation id="D8204B621D4FFFBA910AE454FC0B5E1B" author="Vogellehner, D." pageId="3" pageNumber="28" pagination="125 - 162" refId="ref17771" refString="Vogellehner, D., 1968. Zur Anatomie und Phylogenie de Mesozoischer Gymnospermenholzer, 7: Prodromus zu einer Monographie de Protopinaceae. II. Die Protopinoiden Holzer des Jura. Palaeontographica Abt. B, Stuttgart, 124 (4 - 6): 125 - 162." type="journal article" year="1968">Vogellehner 1968</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA978FE437FABB5E1B" authority="Vogellehner 1968" authorityName="Vogellehner" authorityYear="1968" box="[1014,1391,1117,1141]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="rhaeticum">
<emphasis id="8EC5EA811D4FFFBA978FE437FB585E1B" box="[1014,1164,1117,1141]" italics="true" pageId="3" pageNumber="28">P. rhaeticum</emphasis>
<bibRefCitation id="D8204B621D4FFFBA90D9E434FABB5E1B" author="Vogellehner, D." box="[1184,1391,1117,1141]" pageId="3" pageNumber="28" pagination="125 - 162" refId="ref17771" refString="Vogellehner, D., 1968. Zur Anatomie und Phylogenie de Mesozoischer Gymnospermenholzer, 7: Prodromus zu einer Monographie de Protopinaceae. II. Die Protopinoiden Holzer des Jura. Palaeontographica Abt. B, Stuttgart, 124 (4 - 6): 125 - 162." type="journal article" year="1968">Vogellehner 1968</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA91FFE437FB445EF9" authority="Shilkina and Blokhina" authorityName="Shilkina and Blokhina" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="munense">
<emphasis id="8EC5EA811D4FFFBA91FFE437FC595EF9" italics="true" pageId="3" pageNumber="28">P. munense</emphasis>
Shilkina and Blokhina
</taxonomicName>
(in
<bibRefCitation id="D8204B621D4FFFBA90BAE416FABB5EF9" author="Blokhina, N. I." box="[1219,1391,1151,1175]" pageId="3" pageNumber="28" pagination="47 - 54" refId="ref13001" refString="Blokhina, N. I., 1975. New species of conifers from the Jurassic of North Siberia (Fossil woods). Proceedings of Institute of Biology and Pedology, Vladivostok, n. sr., 27 (130): 47 - 54. [in Russian]" type="journal article" year="1975">Blokhina, 1975</bibRefCitation>
),
<taxonomicName id="7BB14D101D4FFFBA91FFE4E9FB8E5ED6" authority="Francis 1983" authorityName="Francis" authorityYear="1983" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="purbeckensis">
<emphasis id="8EC5EA811D4FFFBA91FFE4E9FC6F5ED6" italics="true" pageId="3" pageNumber="28">P. purbeckensis</emphasis>
<bibRefCitation id="D8204B621D4FFFBA97B1E4C9FB8E5ED6" author="Francis, J. E." box="[968,1114,1184,1208]" pageId="3" pageNumber="28" pagination="277 - 294" refId="ref14517" refString="Francis, J. E., 1983. The dominant conifer of the Jurassic Purbeck Formation, England. Palaeontology, 26: 277 - 294." type="journal article" year="1983">Francis 1983</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA9013E4C8FAA65ED6" authority="Barale" authorityName="Barale" box="[1130,1394,1184,1208]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="carentanensis">
<emphasis id="8EC5EA811D4FFFBA9013E4C8FAF55ED6" box="[1130,1313,1185,1208]" italics="true" pageId="3" pageNumber="28">P. carentanensis</emphasis>
Barale
</taxonomicName>
(in
<bibRefCitation id="D8204B621D4FFFBA9749E4A8FC045EB7" author="Philippe, M." box="[816,976,1217,1241]" pageId="3" pageNumber="28" pagination="45 - 103" refId="ref16179" refString="Philippe, M., 1995. Bois fossils du Jurassique de Franche-Comte (NE-France). Palaeontographica Abt. B, 236: 45 - 103." type="journal article" year="1995">Philippe, 1995</bibRefCitation>
).
</paragraph>
<paragraph id="BC0E36931D4FFFBA9749E48BFC4E5F8C" blockId="3.[816,1437,158,2068]" pageId="3" pageNumber="28">
Also,
<bibRefCitation id="D8204B621D4FFFBA9700E48BFB025E94" author="Iamandei E. &amp; Iamandei S." box="[889,1238,1250,1274]" pageId="3" pageNumber="28" pagination="191 - 199" refId="ref15071" refString="Iamandei E. and Iamandei S., 2000. Bois fossiles de Coniferes dans l'Aptien de Dobroudja de Sud, Romania. Acta Palaeontologica Romaniae, 2 (1999): 191 - 199." type="journal article" year="2000">Iamandei and Iamandei (2000</bibRefCitation>
,
<bibRefCitation id="D8204B621D4FFFBA9093E48BFAF55E94" author="Iamandei, E. &amp; Iamandei, S." box="[1258,1313,1250,1274]" pageId="3" pageNumber="28" pagination="231 - 247" refId="ref15205" refString="Iamandei, E. and Iamandei, S., 2005. Early Cretaceous protopinaceous fossil wood from South Dobrogea, Romania. Acta Palaeontologica Romaniae, 5: 231 - 247." type="journal article" year="2005">2005</bibRefCitation>
) have described an Early Cretaceous species from Dobrogea,
<collectingCountry id="C4A676031D4FFFBA910DE56DFCA15F52" name="Romania" pageId="3" pageNumber="28">Romania</collectingCountry>
:
<taxonomicName id="7BB14D101D4FFFBA97F2E54DFC425F33" authority="Iamandei et Iamandei" authorityName="Iamandei et Iamandei" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="dragastanii">
<emphasis id="8EC5EA811D4FFFBA97F2E54DFAD65F52" box="[907,1282,1316,1340]" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon dragastanii</emphasis>
Iamandei et Iamandei
</taxonomicName>
, which has radial pitting uni- to biseriate, opposite or alternate, no parenchyma, rays of mid tall, uniseriate or partially biseriate and cross fields of cupressoid
<typeStatus id="630A88311D4FFFBA9749E5C0FCB55FAF" box="[816,865,1449,1473]" pageId="3" pageNumber="28">type</typeStatus>
, sometimes with an araucarioid tendency (i.e. alternate pits).
</paragraph>
<paragraph id="BC0E36931D4FFFBA9749E582FB775D20" blockId="3.[816,1437,158,2068]" pageId="3" pageNumber="28">
More recently, Correa et al. (2018) studying an abundant population of wood remains (58 permineralized trunks) from
<collectingCountry id="C4A676031D4FFFBA9715E644FC0D5C2B" box="[876,985,1581,1605]" name="Argentina" pageId="3" pageNumber="28">Argentina</collectingCountry>
, described a new species as
<taxonomicName id="7BB14D101D4FFFBA9169E647FBF75C08" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="carrizalense">
<emphasis id="8EC5EA811D4FFFBA9169E647FBF75C08" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon carrizalense</emphasis>
</taxonomicName>
, and made a comparative table of 32 species. They observed that only 24 from them fit the generic diagnosis, but eight species (i.e.
<emphasis id="8EC5EA811D4FFFBA9087E6F8FA4F5C84" italics="true" pageId="3" pageNumber="28">
<taxonomicName id="7BB14D101D4FFFBA9087E6F8FC235CA7" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="alternans">Protocupressinoxylon alternans</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA907EE6DBFB145CA7" box="[1031,1216,1714,1737]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="cenomanicum">P. cenomanicum</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA90B6E6DBFAA25CA7" box="[1231,1398,1714,1737]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="mesozoicum">P. mesozoicum</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA91FCE6DBFC685C84" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="luccombense">P. luccombense</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA97B2E6BAFBB05C84" box="[971,1124,1747,1770]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="jurassicum">P. jurassicum</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA9008E6BAFB355C84" box="[1137,1249,1746,1770]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="koetlizi">P. koetlizi</taxonomicName>
,
<taxonomicName id="7BB14D101D4FFFBA9096E6BAFA4F5C84" box="[1263,1435,1747,1770]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="potomacense">P. potomacense</taxonomicName>
</emphasis>
and
<taxonomicName id="7BB14D101D4FFFBA9727E69CFC025D62" box="[862,982,1781,1804]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="species" species="vectense">
<emphasis id="8EC5EA811D4FFFBA9727E69CFC025D62" box="[862,982,1781,1804]" italics="true" pageId="3" pageNumber="28">P. vectense</emphasis>
</taxonomicName>
) present cross-fields pits that are different from the cupressoid
<typeStatus id="630A88311D4FFFBA906DE77CFB925D43" box="[1044,1094,1813,1837]" pageId="3" pageNumber="28">type</typeStatus>
: either not bordered, or typical taxodioid (see Correa et al., 2018).
</paragraph>
<paragraph id="BC0E36931D4FFFBD9749E73EFD405BD1" blockId="3.[816,1437,158,2068]" lastBlockId="4.[151,771,163,579]" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28">
Since the specific details inside the genus
<taxonomicName id="7BB14D101D4FFFBA9087E731FC515DFE" authorityName="Eckhold" authorityYear="1921" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="28" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D4FFFBA9087E731FC515DFE" italics="true" pageId="3" pageNumber="28">Protocupressinoxylon</emphasis>
</taxonomicName>
are very similar, it is very difficult to discriminate between them, but through comparative study of their descriptions, and using also the comparative table from Correa et al. (2018), we tried to identify our material, which present these clear xylotomical features: tracheidoxylic secondary wood, having radial pitting 1-2 seriate, of mixed
<typeStatus id="630A88311D48FFBD9524E0A9FE5F5AB6" box="[349,395,192,216]" pageId="4" pageNumber="29">type</typeStatus>
on tracheids, few parenchyma or absent and uniseriate rays with cross-fields with 1-2(4-5) cupressoid pits per field, spaced arranged in 1-2 horizontal rows when more, to irregular, or to slightly alternate. These xylotomical details are very similar with those described by
<bibRefCitation id="D8204B621D48FFBD956DE10DFD8B5B12" author="Iamandei E. &amp; Iamandei S." box="[276,607,356,380]" pageId="4" pageNumber="29" pagination="191 - 199" refId="ref15071" refString="Iamandei E. and Iamandei S., 2000. Bois fossiles de Coniferes dans l'Aptien de Dobroudja de Sud, Romania. Acta Palaeontologica Romaniae, 2 (1999): 191 - 199." type="journal article" year="2000">Iamandei and Iamandei (2000</bibRefCitation>
and
<bibRefCitation id="D8204B621D48FFBD96E2E10DFD065B12" author="Iamandei, E. &amp; Iamandei, S." box="[667,722,356,380]" pageId="4" pageNumber="29" pagination="231 - 247" refId="ref15205" refString="Iamandei, E. and Iamandei, S., 2005. Early Cretaceous protopinaceous fossil wood from South Dobrogea, Romania. Acta Palaeontologica Romaniae, 5: 231 - 247." type="journal article" year="2005">2005</bibRefCitation>
) for the species
<taxonomicName id="7BB14D101D48FFBD9559E1ECFD475BF3" box="[288,659,389,413]" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="species" species="dragastanii">
<emphasis id="8EC5EA811D48FFBD9559E1ECFD475BF3" box="[288,659,389,413]" italics="true" pageId="4" pageNumber="29">Protocupressinoxylon dragastanii</emphasis>
</taxonomicName>
from the Early Cretaceous of South Dobrogea,
<collectingCountry id="C4A676031D48FFBD9654E1CEFD455BD1" box="[557,657,423,447]" name="Romania" pageId="4" pageNumber="29">Romania</collectingCountry>
.
</paragraph>
<paragraph id="BC0E36931D48FFBD94EEE1A1FE61582D" blockId="4.[151,771,163,579]" pageId="4" pageNumber="29">
Thus, based on the above discussion, we attribute all the 14 studied specimens to the species
<taxonomicName id="7BB14D101D48FFBD9660E180FDAF584C" authority="Iamandei and Iamandei, 2000" authorityName="Iamandei and Iamandei" authorityYear="2000" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="species" species="dragastanii">
<emphasis id="8EC5EA811D48FFBD9660E180FEC6584C" italics="true" pageId="4" pageNumber="29">Protocupressinoxylon dragastanii</emphasis>
<bibRefCitation id="D8204B621D48FFBD955BE263FDAF584C" author="Iamandei E. &amp; Iamandei S." box="[290,635,522,546]" pageId="4" pageNumber="29" pagination="191 - 199" refId="ref15071" refString="Iamandei E. and Iamandei S., 2000. Bois fossiles de Coniferes dans l'Aptien de Dobroudja de Sud, Romania. Acta Palaeontologica Romaniae, 2 (1999): 191 - 199." type="journal article" year="2000">Iamandei and Iamandei, 2000</bibRefCitation>
</taxonomicName>
, as another Mesozoic European taxon.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,273 @@
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<emphasis id="8EC5EA811D5CFFA99075E168FB155B77" box="[1036,1217,257,281]" italics="true" pageId="16" pageNumber="41">Bucklandia sp. B</emphasis>
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<figureCitation id="248A2A161D5CFFA99056E14BFBBB5B54" box="[1071,1135,290,314]" captionStart="Fig" captionStartId="17.[166,201,1917,1938]" captionTargetBox="[166,1424,151,1907]" captionTargetId="figure-19@17.[166,1424,151,1907]" captionTargetPageId="17" captionText="Fig. 8 Bucklandia sp. B - inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i). a-f: Cross section b: - tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem; c: - mucilaginous canals lined by secretory epithelial cells; d-e: - phloem cells with idioblasts bearing mucillages; f: - or crystals (f); g: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids; h-i: Radial section mucilaginous canal (h); araucarian radial pitting on tracheids with uniseriate hexagonal pits (i), cross-fields with 2-3 superposed rows of hexagonal pits (h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="16" pageNumber="41">Fig. 8</figureCitation>
, a-i.
</paragraph>
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<emphasis id="8EC5EA811D5CFFA99749E10CFC475B13" bold="true" box="[816,915,357,381]" pageId="16" pageNumber="41">Material</emphasis>
</paragraph>
<paragraph id="BC0E36931D5CFFA99749E1ECFB77580A" blockId="16.[816,1437,357,612]" pageId="16" pageNumber="41">
<materialsCitation id="0CD93CCE1D5CFFA99749E1ECFB4B580A" location="National Museum of Geology" pageId="16" pageNumber="41" specimenCount="1" stateProvince="Bucuresti">
The studied material is represented by three small (centimetric) samples of silicified wood, collected from around Holbav locality area, on Maiului brook. These samples have the field-numbers: 1037, 1038 and 1079, in &quot;Grădinaru Collection&quot; and are stored now at the
<location id="B96E60481D5CFFA99173E263FC2A582D" LSID="urn:lsid:plazi:treatment:341887851D5CFFAB9075E168FD4C5F33:B96E60481D5CFFA99173E263FC2A582D" name="National Museum of Geology" pageId="16" pageNumber="41" stateProvince="Bucuresti">National Museum of Geology</location>
, in
<collectingRegion id="7E75F8711D5CFFA9904EE242FB70582D" box="[1079,1188,555,579]" country="Romania" name="Bucuresti" pageId="16" pageNumber="41">Bucharest</collectingRegion>
, under the inventory numbers 27676, 27677 and 27683
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D5CFFAB9749E2E6FD785A96" lastPageId="18" lastPageNumber="43" pageId="16" pageNumber="41" type="description">
<paragraph id="BC0E36931D5CFFA99749E2E6FB9558C9" blockId="16.[816,1437,655,2068]" box="[816,1089,655,679]" pageId="16" pageNumber="41">
<emphasis id="8EC5EA811D5CFFA99749E2E6FB9558C9" bold="true" box="[816,1089,655,679]" pageId="16" pageNumber="41">Microscopic description</emphasis>
</paragraph>
<paragraph id="BC0E36931D5CFFA99749E2C6FB545903" blockId="16.[816,1437,655,2068]" pageId="16" pageNumber="41">
Cross-sections of the studied specimens show a wood structure of gymnospermous
<typeStatus id="630A88311D5CFFA99000E2B8FB735887" box="[1145,1191,721,745]" pageId="16" pageNumber="41">type</typeStatus>
(
<figureCitation id="248A2A161D5CFFA990C5E2B8FADA5887" box="[1212,1294,721,745]" captionStart="Fig" captionStartId="17.[166,201,1917,1938]" captionTargetBox="[166,1424,151,1907]" captionTargetId="figure-19@17.[166,1424,151,1907]" captionTargetPageId="17" captionText="Fig. 8 Bucklandia sp. B - inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i). a-f: Cross section b: - tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem; c: - mucilaginous canals lined by secretory epithelial cells; d-e: - phloem cells with idioblasts bearing mucillages; f: - or crystals (f); g: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids; h-i: Radial section mucilaginous canal (h); araucarian radial pitting on tracheids with uniseriate hexagonal pits (i), cross-fields with 2-3 superposed rows of hexagonal pits (h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="16" pageNumber="41">Fig. 8a</figureCitation>
), with some details indicative of a cycadeoid stem structure, due to the presence of idioblasts and mucilaginous canals (
<figureCitation id="248A2A161D5CFFA9913EE37AFC945922" captionStart="Fig" captionStartId="17.[166,201,1917,1938]" captionTargetBox="[166,1424,151,1907]" captionTargetId="figure-19@17.[166,1424,151,1907]" captionTargetPageId="17" captionText="Fig. 8 Bucklandia sp. B - inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i). a-f: Cross section b: - tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem; c: - mucilaginous canals lined by secretory epithelial cells; d-e: - phloem cells with idioblasts bearing mucillages; f: - or crystals (f); g: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids; h-i: Radial section mucilaginous canal (h); araucarian radial pitting on tracheids with uniseriate hexagonal pits (i), cross-fields with 2-3 superposed rows of hexagonal pits (h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="16" pageNumber="41">Fig. 8c, h</figureCitation>
); the xylotomical description is appropriate, in order to identify the unknown structure.
</paragraph>
<paragraph id="BC0E36931D5CFFA99749E31FFAB959C1" blockId="16.[816,1437,655,2068]" pageId="16" pageNumber="41">
<emphasis id="8EC5EA811D5CFFA99749E31FFC5D59E0" box="[816,905,886,910]" italics="true" pageId="16" pageNumber="41">The pith</emphasis>
is not present in our sections because the axial part of the stem is absent in our fragmentary samples.
</paragraph>
<paragraph id="BC0E36931D5CFFA99749E3D1FB535CE9" blockId="16.[816,1437,655,2068]" pageId="16" pageNumber="41">
<emphasis id="8EC5EA811D5CFFA99749E3D1FB4759BE" box="[816,1171,952,976]" italics="true" pageId="16" pageNumber="41">The cylindrical vascular system</emphasis>
in cross section, appears to consist of some alternate growth rings of secondary xylem and phloem, which indicate a polyxylic structure, traversed by numerous thin secondary rays (
<figureCitation id="248A2A161D5CFFA99138E472FA5B5E5D" box="[1345,1423,1051,1075]" captionStart="Fig" captionStartId="17.[166,201,1917,1938]" captionTargetBox="[166,1424,151,1907]" captionTargetId="figure-19@17.[166,1424,151,1907]" captionTargetPageId="17" captionText="Fig. 8 Bucklandia sp. B - inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i). a-f: Cross section b: - tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem; c: - mucilaginous canals lined by secretory epithelial cells; d-e: - phloem cells with idioblasts bearing mucillages; f: - or crystals (f); g: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids; h-i: Radial section mucilaginous canal (h); araucarian radial pitting on tracheids with uniseriate hexagonal pits (i), cross-fields with 2-3 superposed rows of hexagonal pits (h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="16" pageNumber="41">Fig. 8a</figureCitation>
). The wide multiseriate primary medullary rays appear only locally, in contact with the xylem of growth ring (see
<figureCitation id="248A2A161D5CFFA9971AE416FC675EF9" box="[867,947,1151,1175]" captionStart="Fig" captionStartId="17.[166,201,1917,1938]" captionTargetBox="[166,1424,151,1907]" captionTargetId="figure-19@17.[166,1424,151,1907]" captionTargetPageId="17" captionText="Fig. 8 Bucklandia sp. B - inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i). a-f: Cross section b: - tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem; c: - mucilaginous canals lined by secretory epithelial cells; d-e: - phloem cells with idioblasts bearing mucillages; f: - or crystals (f); g: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids; h-i: Radial section mucilaginous canal (h); araucarian radial pitting on tracheids with uniseriate hexagonal pits (i), cross-fields with 2-3 superposed rows of hexagonal pits (h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="16" pageNumber="41">Fig. 8b</figureCitation>
). Through the xylem and phloem cells, obvious idioblasts appear, carrying mucilages or crystals often grouped as druses (
<figureCitation id="248A2A161D5CFFA9907BE4A8FB955EB7" box="[1026,1089,1217,1241]" captionStart="Fig" captionStartId="17.[166,201,1917,1938]" captionTargetBox="[166,1424,151,1907]" captionTargetId="figure-19@17.[166,1424,151,1907]" captionTargetPageId="17" captionText="Fig. 8 Bucklandia sp. B - inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i). a-f: Cross section b: - tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem; c: - mucilaginous canals lined by secretory epithelial cells; d-e: - phloem cells with idioblasts bearing mucillages; f: - or crystals (f); g: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids; h-i: Radial section mucilaginous canal (h); araucarian radial pitting on tracheids with uniseriate hexagonal pits (i), cross-fields with 2-3 superposed rows of hexagonal pits (h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="16" pageNumber="41">Fig. 8</figureCitation>
d-f). The cambium cells are difficult to recognize. The xylem has rectangular to square tracheids (ra/tg diameters of 18-50/20-45 μm), with rounded corners, with often slightly wavy walls, 3-6 μm as a double wall. On the radial walls, the tracheids show uni- to biseriate bordered pits, rounded or hexagonal, of 6-8 μm, with round to elliptic apertures and are contiguous and flattened (
<figureCitation id="248A2A161D5CFFA99786E5C0FB9D5FAF" box="[1023,1097,1449,1473]" captionStart="Fig" captionStartId="17.[166,201,1917,1938]" captionTargetBox="[166,1424,151,1907]" captionTargetId="figure-19@17.[166,1424,151,1907]" captionTargetPageId="17" captionText="Fig. 8 Bucklandia sp. B - inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i). a-f: Cross section b: - tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem; c: - mucilaginous canals lined by secretory epithelial cells; d-e: - phloem cells with idioblasts bearing mucillages; f: - or crystals (f); g: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids; h-i: Radial section mucilaginous canal (h); araucarian radial pitting on tracheids with uniseriate hexagonal pits (i), cross-fields with 2-3 superposed rows of hexagonal pits (h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="16" pageNumber="41">Fig. 8i</figureCitation>
), usually alternate, where they are biseriate. Perforations were not seen. The tangential walls are not pitted. The phloem appears as parenchymal cells, with secretory idioblasts and sclerenchymal cells often with large crystals, or druses (
<figureCitation id="248A2A161D5CFFA990A8E644FAC15C2B" box="[1233,1301,1581,1605]" captionStart="Fig" captionStartId="17.[166,201,1917,1938]" captionTargetBox="[166,1424,151,1907]" captionTargetId="figure-19@17.[166,1424,151,1907]" captionTargetPageId="17" captionText="Fig. 8 Bucklandia sp. B - inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i). a-f: Cross section b: - tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem; c: - mucilaginous canals lined by secretory epithelial cells; d-e: - phloem cells with idioblasts bearing mucillages; f: - or crystals (f); g: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids; h-i: Radial section mucilaginous canal (h); araucarian radial pitting on tracheids with uniseriate hexagonal pits (i), cross-fields with 2-3 superposed rows of hexagonal pits (h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="16" pageNumber="41">Fig. 8</figureCitation>
d-e). At the border with the pith area, mucilaginous canals appear, lined by rows of epithelial cells.
</paragraph>
<paragraph id="BC0E36931D5CFFA99749E6F9FAE3527A" blockId="16.[816,1437,655,2068]" pageId="16" pageNumber="41">
<emphasis id="8EC5EA811D5CFFA99749E6F9FC5A5CC6" box="[816,910,1680,1704]" italics="true" pageId="16" pageNumber="41">The rays</emphasis>
in cross- and tangential sections the secondary rays appear numerous and thin, giving a manoxylic appearance to the structure. The primary rays were not observed; but they do appear in cross-section, in contact with xylem (see
<figureCitation id="248A2A161D5CFFA9979FE77CFBE25D43" box="[998,1078,1813,1837]" captionStart="Fig" captionStartId="17.[166,201,1917,1938]" captionTargetBox="[166,1424,151,1907]" captionTargetId="figure-19@17.[166,1424,151,1907]" captionTargetPageId="17" captionText="Fig. 8 Bucklandia sp. B - inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i). a-f: Cross section b: - tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem; c: - mucilaginous canals lined by secretory epithelial cells; d-e: - phloem cells with idioblasts bearing mucillages; f: - or crystals (f); g: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids; h-i: Radial section mucilaginous canal (h); araucarian radial pitting on tracheids with uniseriate hexagonal pits (i), cross-fields with 2-3 superposed rows of hexagonal pits (h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="16" pageNumber="41">Fig. 8b</figureCitation>
). The secondary rays are usually uniseriate, very numerous and are composed of parenchymal cells, and few secretory idioblasts, so that the rays are heterogeneous (
<figureCitation id="248A2A161D5CFFA9907BE711FB9E5DFE" box="[1026,1098,1912,1936]" captionStart="Fig" captionStartId="17.[166,201,1917,1938]" captionTargetBox="[166,1424,151,1907]" captionTargetId="figure-19@17.[166,1424,151,1907]" captionTargetPageId="17" captionText="Fig. 8 Bucklandia sp. B - inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i). a-f: Cross section b: - tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem; c: - mucilaginous canals lined by secretory epithelial cells; d-e: - phloem cells with idioblasts bearing mucillages; f: - or crystals (f); g: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids; h-i: Radial section mucilaginous canal (h); araucarian radial pitting on tracheids with uniseriate hexagonal pits (i), cross-fields with 2-3 superposed rows of hexagonal pits (h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="16" pageNumber="41">
Fig.
<quantity id="7B499B761D5CFFA99049E711FB9E5DFE" box="[1072,1098,1912,1936]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="16" pageNumber="41" unit="g" value="8.0">8g</quantity>
</figureCitation>
). The ray cells have rounded to rectangular outlines, the idioblasts are similar, or slightly elongate and, in radial view, are all procumbent. The cross-fields show 2-6 oculipores, in one or two superposed rows and are slightly alternately arranged.
</paragraph>
<caption id="E8CE661B1D5DFFA894DFE714FE575249" ID-DOI="http://doi.org/10.5281/zenodo.10520867" ID-Zenodo-Dep="10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="17" pageNumber="42" startId="17.[166,201,1917,1938]" targetBox="[166,1424,151,1907]" targetPageId="17" targetType="figure">
<paragraph id="BC0E36931D5DFFA894DFE714FC305DFC" blockId="17.[166,1425,1916,2087]" box="[166,996,1916,1938]" pageId="17" pageNumber="42">
<emphasis id="8EC5EA811D5DFFA894DFE714FF0B5DFC" bold="true" box="[166,223,1917,1938]" pageId="17" pageNumber="42">Fig. 8</emphasis>
<taxonomicName id="7BB14D101D5DFFA8949FE715FE5D5DFC" box="[230,393,1916,1938]" class="Magnoliopsida" family="Hamamelidaceae" genus="Bucklandia" kingdom="Plantae" order="Saxifragales" pageId="17" pageNumber="42" phylum="Tracheophyta" rank="species" species="undefined-B">
<emphasis id="8EC5EA811D5DFFA8949FE715FE5D5DFC" box="[230,393,1916,1938]" italics="true" pageId="17" pageNumber="42">Bucklandia sp. B</emphasis>
</taxonomicName>
- inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i).
</paragraph>
<paragraph id="BC0E36931D5DFFA894DFE7F2FE575249" blockId="17.[166,1425,1916,2087]" pageId="17" pageNumber="42">
<emphasis id="8EC5EA811D5DFFA894DFE7F2FF165DDE" bold="true" box="[166,194,1947,1968]" pageId="17" pageNumber="42">a-f</emphasis>
: Cross section
<emphasis id="8EC5EA811D5DFFA89515E7F2FE555DDE" bold="true" box="[364,385,1947,1968]" pageId="17" pageNumber="42">b:</emphasis>
- tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem;
<emphasis id="8EC5EA811D5DFFA89149E7F2FA855DDE" bold="true" box="[1328,1361,1947,1968]" pageId="17" pageNumber="42">c: -</emphasis>
mucilaginous canals lined by secretory epithelial cells;
<emphasis id="8EC5EA811D5DFFA896E2E7D1FD075DA3" bold="true" box="[667,723,1976,1997]" pageId="17" pageNumber="42">d-e: -</emphasis>
phloem cells with idioblasts bearing mucillages;
<emphasis id="8EC5EA811D5DFFA890B9E7D1FB345DA3" bold="true" box="[1216,1248,1976,1997]" pageId="17" pageNumber="42">f: -</emphasis>
or crystals (
<emphasis id="8EC5EA811D5DFFA89126E7D1FAB35DA3" bold="true" box="[1375,1383,1976,1997]" pageId="17" pageNumber="42">f</emphasis>
);
<emphasis id="8EC5EA811D5DFFA89107E7D1FA5E5DA3" bold="true" box="[1406,1418,1976,1997]" pageId="17" pageNumber="42">g</emphasis>
: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids;
<emphasis id="8EC5EA811D5DFFA890F2E7BFFB735D85" bold="true" box="[1163,1191,2006,2027]" pageId="17" pageNumber="42">h-i</emphasis>
: Radial section mucilaginous canal (
<emphasis id="8EC5EA811D5DFFA89538E79DFE9A5267" bold="true" box="[321,334,2036,2057]" pageId="17" pageNumber="42">h</emphasis>
); araucarian radial pitting on tracheids with uniseriate hexagonal pits (
<emphasis id="8EC5EA811D5DFFA8907AE79DFBDE5267" bold="true" box="[1027,1034,2036,2057]" pageId="17" pageNumber="42">i</emphasis>
), cross-fields with 2-3 superposed rows of hexagonal pits (
<emphasis id="8EC5EA811D5DFFA89520E87BFEA15249" bold="true" box="[345,373,2066,2087]" pageId="17" pageNumber="42">h-i</emphasis>
).
</paragraph>
</caption>
<paragraph id="BC0E36931D5EFFAB94EEE0F7FD785A96" blockId="18.[151,772,158,1373]" pageId="18" pageNumber="43">
<emphasis id="8EC5EA811D5EFFAB94EEE0F7FECA5AD8" box="[151,286,158,182]" italics="true" pageId="18" pageNumber="43">Axial canals</emphasis>
- appear at the border with the pith area, as large mucilaginous canals, with diameters of 360-930 μm, lined by secretory epithelial cells (
<figureCitation id="248A2A161D5EFFAB964FE089FD4A5A96" box="[566,670,224,248]" captionStart="Fig" captionStartId="17.[166,201,1917,1938]" captionTargetBox="[166,1424,151,1907]" captionTargetId="figure-19@17.[166,1424,151,1907]" captionTargetPageId="17" captionText="Fig. 8 Bucklandia sp. B - inventory no. 27676 and 27677 (field no. 1037 a-f, 1038g-i). a-f: Cross section b: - tracheids polygonal with rounded corners, and linear rays; primary ray in contact with xylem; c: - mucilaginous canals lined by secretory epithelial cells; d-e: - phloem cells with idioblasts bearing mucillages; f: - or crystals (f); g: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells; unpitted tracheids; h-i: Radial section mucilaginous canal (h); araucarian radial pitting on tracheids with uniseriate hexagonal pits (i), cross-fields with 2-3 superposed rows of hexagonal pits (h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520867" httpUri="https://zenodo.org/record/10520867/files/figure.png" pageId="18" pageNumber="43">Fig. 8c, h</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F4AB65181D5EFFAB94EEE16BFD4C5F33" pageId="18" pageNumber="43" type="discussion">
<paragraph id="BC0E36931D5EFFAB94EEE16BFE775B74" blockId="18.[151,772,158,1373]" box="[151,419,258,282]" pageId="18" pageNumber="43">
<emphasis id="8EC5EA811D5EFFAB94EEE16BFE775B74" bold="true" box="[151,419,258,282]" pageId="18" pageNumber="43">Afinities and discussion</emphasis>
</paragraph>
<paragraph id="BC0E36931D5EFFAB94EEE14BFE95586F" blockId="18.[151,772,158,1373]" pageId="18" pageNumber="43">
Having a cycadeoid stem structure, indicated by the presence of idioblasts with druses and of mucilaginous canals, we searched also for other xylotomical details, but the absence of scalariform thickenings on tracheids makes it different from the above-described form, and indicated a resemblance to a
<typeStatus id="630A88311D5EFFAB951DE1A1FE455B8E" box="[356,401,456,480]" pageId="18" pageNumber="43">type</typeStatus>
close to the current
<taxonomicName id="7BB14D101D5EFFAB96F3E1A1FD1F5BB1" box="[650,715,456,479]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5EFFAB96F3E1A1FD1F5BB1" box="[650,715,456,479]" italics="true" pageId="18" pageNumber="43">Cycas</emphasis>
</taxonomicName>
(see
<bibRefCitation id="D8204B621D5EFFAB94EEE180FEE1586F" author="Greguss" box="[151,309,489,513]" pageId="18" pageNumber="43" pagination="1 - 260" refId="ref14798" refString="Greguss, 1968. Xylotomy of the living Cycads, with a description of their leaves and epidermis. 1 - 260 pages, Akademiai Kiado." type="book chapter" year="1968">Greguss, 1968</bibRefCitation>
).
</paragraph>
<paragraph id="BC0E36931D5EFFAB94EEE262FD1C58EB" blockId="18.[151,772,158,1373]" pageId="18" pageNumber="43">
In the Early Jurassic flora from Holbav, species of
<taxonomicName id="7BB14D101D5EFFAB96B6E263FF05582D" class="Testudines" family="Trionychidae" genus="Nilssonia" kingdom="Animalia" order="Cryptodira" pageId="18" pageNumber="43" phylum="Chordata" rank="genus">
<emphasis id="8EC5EA811D5EFFAB96B6E263FF05582D" italics="true" pageId="18" pageNumber="43">Nilssonia</emphasis>
</taxonomicName>
,
<taxonomicName id="7BB14D101D5EFFAB9493E245FE8F582D" authorityName="T.M.Harris" authorityYear="1964" box="[234,347,556,579]" class="Cycadopsida" genus="Paracycas" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5EFFAB9493E245FE8F582D" box="[234,347,556,579]" italics="true" pageId="18" pageNumber="43">Paracycas</emphasis>
</taxonomicName>
,
<emphasis id="8EC5EA811D5EFFAB950AE242FD8D582D" box="[371,601,555,579]" italics="true" pageId="18" pageNumber="43">
<taxonomicName id="7BB14D101D5EFFAB950AE242FE6D582C" authorityName="Lindley &amp; W.Hutton" authorityYear="1834" box="[371,441,555,578]" class="Cycadopsida" genus="Ctenis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="genus">Ctenis</taxonomicName>
,
<taxonomicName id="7BB14D101D5EFFAB95B6E242FD8D582D" authorityName="Guérin-Ménéville" authorityYear="1845" box="[463,601,555,579]" class="Polypodiopsida" family="Hymenophyllaceae" genus="Ptilophyllum" kingdom="Plantae" order="Hymenophyllales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="genus">Ptilophyllum</taxonomicName>
</emphasis>
,
<taxonomicName id="7BB14D101D5EFFAB960BE242FCD7582D" box="[626,771,555,579]" family="Cichlidae" genus="Pterophyllum" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="43" phylum="Chordata" rank="genus">
<emphasis id="8EC5EA811D5EFFAB960BE242FCD7582D" box="[626,771,555,579]" italics="true" pageId="18" pageNumber="43">Pterophyllum</emphasis>
</taxonomicName>
appear, in addition to forms of
<emphasis id="8EC5EA811D5EFFAB958EE224FD80580A" box="[503,596,589,612]" italics="true" pageId="18" pageNumber="43">Zamites-</emphasis>
group (see
<bibRefCitation id="D8204B621D5EFFAB96ABE224FE5E58EB" author="Dragastan, O. N. &amp; Popa, M. E." pageId="18" pageNumber="43" pagination="51 - 60" refId="ref14175" refString="Dragastan, O. N. and Popa, M. E., 1997. Early Jurassic Phytostratigraphy of the Holbav Formation, Getic Nappe, Brasov County. Revue Roumaine de Geologie, 41: 51 - 60." type="journal article" year="1997">Dragastan and Popa, 1997</bibRefCitation>
;
<bibRefCitation id="D8204B621D5EFFAB95ECE207FDC458EB" author="Popa, M. E." box="[405,528,621,645]" pageId="18" pageNumber="43" pagination="81 - 87" refId="ref16468" refString="Popa, M. E., 1997. Corystospermal pteridosperms in the Liassic continental deposits of Romania. Acta Palaeontologica Romaniae, 1: 81 - 87." type="journal article" year="1997">Popa, 1997</bibRefCitation>
,
<bibRefCitation id="D8204B621D5EFFAB9663E204FD8658EB" author="Popa, M. E." box="[538,594,621,645]" pageId="18" pageNumber="43" pagination="177 - 184" refId="ref16497" refString="Popa, M. E., 1998. The Liassic continental flora of Romania: Systematics, Stratigraphy and Paleoecology. Acta Botanica Horti Bucurestensis: 177 - 184." type="book chapter" year="1998">1998</bibRefCitation>
,
<bibRefCitation id="D8204B621D5EFFAB9624E204FD7558EB" author="Popa, M. E." box="[605,673,621,645]" pageId="18" pageNumber="43" pagination="377 - 386" refId="ref16528" refString="Popa, M. E., 2000 a. Aspects of Romanian Early Jurassic palaeobotany and palynology. Part III. Phytostratigraphy of the Getic Nappe. Acta Palaeontologica Romaniae, 2: 377 - 386." type="journal article" year="2000">2000a</bibRefCitation>
, b).
</paragraph>
<paragraph id="BC0E36931D5EFFAB94EEE2E7FD4159E0" blockId="18.[151,772,158,1373]" pageId="18" pageNumber="43">
Comparing to wood description of
<emphasis id="8EC5EA811D5EFFAB966AE2E6FD0458C8" box="[531,720,654,678]" italics="true" pageId="18" pageNumber="43">Zamuneria amyla</emphasis>
(see
<bibRefCitation id="D8204B621D5EFFAB94EEE2C6FE5458A9" author="Martinez, L. C. A. &amp; Iglesias, A. &amp; Artabe A. E. &amp; Varela A. N. &amp; Apesteguia, S." box="[151,384,687,711]" pageId="18" pageNumber="43" pagination="81 - 94" refId="ref15787" refString="Martinez, L. C. A., Iglesias, A., Artabe A. E., Varela A. N. and Apesteguia, S., 2017. A new Encephalarteae trunk (Cycadales) from the Cretaceous of Patagonia (Mata Amarilla Formation, Austral Basin), Argentina. Cretaceous Research, 72: 81 - 94 http: // dx. doi. org / 10.1016 / j." type="journal article" year="2017">Martinez et al., 2017</bibRefCitation>
) and, also, their identification key, we observed that our studied specimens indicate a columnar polyxylic stem, with several growth rings in the cylindrical vascular system, and with idioblasts and mucilaginous canals, similar to the genus
<taxonomicName id="7BB14D101D5EFFAB9640E35DFF1B5903" authority="Bosse, 1953" authorityName="Bosse" authorityYear="1953" class="Magnoliopsida" family="Hamamelidaceae" genus="Bucklandia" kingdom="Plantae" order="Saxifragales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5EFFAB9640E35DFD665922" box="[569,690,820,844]" italics="true" pageId="18" pageNumber="43">Bucklandia</emphasis>
<bibRefCitation id="D8204B621D5EFFAB96C4E35CFF1B5903" author="Bosse, M. N." pageId="18" pageNumber="43" pagination="41 - 50" refId="ref13235" refString="Bosse, M. N. 1953. Bucklandia sahnii sp. nov. from the Jurassic of the Rajmahal Hills, Bihar. Palaeobotanist, 2: 41 - 50." type="journal article" year="1953">Bosse, 1953</bibRefCitation>
</taxonomicName>
, which is a stem of
<taxonomicName id="7BB14D101D5EFFAB95C6E33CFD175903" authority="Caruthers" authorityName="Caruthers" box="[447,707,853,877]" class="Insecta" family="Corduliidae" genus="Williamsonia" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="8EC5EA811D5EFFAB95C6E33CFD995903" box="[447,589,853,877]" italics="true" pageId="18" pageNumber="43">Williamsonia</emphasis>
Caruthers
</taxonomicName>
, as it was described by
<bibRefCitation id="D8204B621D5EFFAB952FE31FFD4659E0" author="Stockey, R. &amp; Rothwell, G. W." box="[342,658,886,910]" pageId="18" pageNumber="43" pagination="251 - 262" refId="ref17276" refString="Stockey, R. and Rothwell, G. W., 2003. Anatomically Preserved Williamsonia (Williamsoniaceae): Evidence for Bennettitalean Reproduction in the Late Cretaceous of Western North America. International Journal of Plant Sciences 164 (2): 251 - 262. https: // doi. org / 10.1086 / 346166" type="journal article" year="2003">Stockey and Rothwell (2003)</bibRefCitation>
.
</paragraph>
<paragraph id="BC0E36931D5EFFAB94EEE3FEFDF35E5D" blockId="18.[151,772,158,1373]" pageId="18" pageNumber="43">
Our specimens have a structure similar to a Jurassic stem described by
<bibRefCitation id="D8204B621D5EFFAB954DE3D1FDC159BE" author="Zhang W. &amp; Yang X. - J. &amp; Fu X. - P. &amp; Zheng S. - L. &amp; Wang Y. - D." box="[308,533,952,976]" pageId="18" pageNumber="43" pagination="50 - 60" refId="ref17946" refString="Zhang W., Yang X. - J., Fu X. - P., Zheng S. - L. and Wang Y. - D., 2012. A polyxylic cycad trunk from the Middle Jurassic of western Liaoning, China, and its evolutionary implications. Review of Palaeobotany and Palynology, 183: 50 - 60." type="journal article" year="2012">Zhang et al. (2012)</bibRefCitation>
as
<taxonomicName id="7BB14D101D5EFFAB9631E3D1FD2859BE" authorityName="W.Zhang &amp; X.-J.Yang" authorityYear="2012" box="[584,764,952,976]" class="Cycadopsida" genus="Sinocycadoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5EFFAB9631E3D1FD2859BE" box="[584,764,952,976]" italics="true" pageId="18" pageNumber="43">Sinocycadoxylon</emphasis>
</taxonomicName>
, which is also characterized by a polyxylic vascular cylinder with centrifugal rings of secondary xylem and araucarioid radial pitting on the tracheids.
</paragraph>
<paragraph id="BC0E36931D5EFFAB94EEE455FF035EB7" blockId="18.[151,772,158,1373]" pageId="18" pageNumber="43">
Also,
<bibRefCitation id="D8204B621D5EFFAB94A0E455FE1F5E3A" author="Iamandei E. &amp; Iamandei S. &amp; Grinea, D." box="[217,459,1084,1108]" pageId="18" pageNumber="43" pagination="43 - 53" refId="ref15166" refString="Iamandei E., Iamandei S. and Grinea, D., 2003, Cycadoxylon sp., a Fossil Wood from Upper Jurassic of Haghimas. Revue roumaine de Geologie, 47, 43 - 53." type="journal article" year="2003">Iamandei et al. (2003)</bibRefCitation>
studied a fragmentary fossil wood from the Upper Jurassic of Hăghimaş Mts., Eastern Carpathians, which is identified as
<taxonomicName id="7BB14D101D5EFFAB963BE416FD2A5EF8" box="[578,766,1151,1175]" class="Cycadopsida" family="Nilssoniaceae" genus="Cycadoxylon" kingdom="Plantae" order="Cycadales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="8EC5EA811D5EFFAB963BE416FD195EF9" box="[578,717,1151,1175]" italics="true" pageId="18" pageNumber="43">Cycadoxylon</emphasis>
sp.
</taxonomicName>
, which has a xylotomy very similar to our studied specimens.
</paragraph>
<paragraph id="BC0E36931D5EFFAB94EEE48BFD4C5F33" blockId="18.[151,772,158,1373]" pageId="18" pageNumber="43">
Thus, the studied specimens show details that indicate a columnar form of
<taxonomicName id="7BB14D101D5EFFAB951DE56DFE715F75" box="[356,421,1284,1307]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="8EC5EA811D5EFFAB951DE56DFE715F75" box="[356,421,1284,1307]" italics="true" pageId="18" pageNumber="43">Cycas</emphasis>
</taxonomicName>
-
<typeStatus id="630A88311D5EFFAB95D7E56DFE085F72" box="[430,476,1284,1308]" pageId="18" pageNumber="43">type</typeStatus>
(see
<bibRefCitation id="D8204B621D5EFFAB9663E56AFD6F5F75" author="Greguss" box="[538,699,1283,1307]" pageId="18" pageNumber="43" pagination="1 - 260" refId="ref14798" refString="Greguss, 1968. Xylotomy of the living Cycads, with a description of their leaves and epidermis. 1 - 260 pages, Akademiai Kiado." type="book chapter" year="1968">Greguss, 1968</bibRefCitation>
), as is discussed above, so we assign them to
<taxonomicName id="7BB14D101D5EFFAB963AE54DFD285F52" box="[579,764,1316,1340]" class="Magnoliopsida" family="Hamamelidaceae" genus="Bucklandia" kingdom="Plantae" order="Saxifragales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="species" species="undefined-B">
<emphasis id="8EC5EA811D5EFFAB963AE54DFD285F52" box="[579,764,1316,1340]" italics="true" pageId="18" pageNumber="43">Bucklandia sp. B</emphasis>
</taxonomicName>
, clearly different from the
<taxonomicName id="7BB14D101D5EFFAB95D3E52CFD9E5F33" box="[426,586,1349,1373]" form="described" pageId="18" pageNumber="43" rank="form">form described</taxonomicName>
above.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,103 @@
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Order
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-
<taxonomicName id="7BB14D101D5EFFAB973BE1ECFC475BD1" authority="Iamandei and Iamandei, 2000" authorityName="Iamandei and Iamandei" authorityYear="2000" class="Pinopsida" family="Araucariaceae" genus="Protocupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="species" species="dragastanii">
<emphasis id="8EC5EA811D5EFFAB973BE1ECFB7B5BF3" box="[834,1199,389,413]" italics="true" pageId="18" pageNumber="43">Protocupressinoxylon dragastanii</emphasis>
<bibRefCitation id="D8204B621D5EFFAB90CEE1ECFC475BD1" author="Iamandei E. &amp; Iamandei S." pageId="18" pageNumber="43" pagination="191 - 199" refId="ref15071" refString="Iamandei E. and Iamandei S., 2000. Bois fossiles de Coniferes dans l'Aptien de Dobroudja de Sud, Romania. Acta Palaeontologica Romaniae, 2 (1999): 191 - 199." type="journal article" year="2000">Iamandei and Iamandei, 2000</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="BC0E36931D5EFFAB9749E1A1FC2A586F" blockId="18.[816,1436,389,811]" pageId="18" pageNumber="43">
-
<taxonomicName id="7BB14D101D5EFFAB9739E1A1FC4B586F" authority="Iamandei, Iamandei and Gradinaru" authorityName="Iamandei, Iamandei and Gradinaru" authorityYear="2022" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="species" species="holbavicum" status="sp. nov.">
<emphasis id="8EC5EA811D5EFFAB9739E1A1FB405B8E" box="[832,1172,456,480]" italics="true" pageId="18" pageNumber="43">Protojuniperoxylon holbavicum</emphasis>
Iamandei, Iamandei and Grădinaru
</taxonomicName>
,
<taxonomicNameLabel id="95F657FA1D5EFFAB97D3E183FC2A586F" box="[938,1022,490,513]" pageId="18" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
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-
<taxonomicName id="7BB14D101D5EFFAB9735E263FC0B582D" authority="Iamandei, Iamandei and Gradinaru, 2018" authorityName="Iamandei, Iamandei and Gradinaru" authorityYear="2018" class="Pinopsida" family="Araucariaceae" genus="Brachyoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="18" pageNumber="43" phylum="Tracheophyta" rank="species" species="holbavicum">
<emphasis id="8EC5EA811D5EFFAB9735E263FBA4584C" box="[844,1136,522,546]" italics="true" pageId="18" pageNumber="43">Brachyoxylon holbavicum</emphasis>
<bibRefCitation id="D8204B621D5EFFAB90FAE263FC0B582D" author="Iamandei, S. &amp; Iamandei, E. &amp; Gradinaru, E." pageId="18" pageNumber="43" pagination="3 - 34" refId="ref15238" refString="Iamandei, S., Iamandei, E. and Gradinaru, E., 2018. Contributions to the study of the Early Jurassic Petrified Forest of Holbav and Cristian areas (Brasov region, South Carpathians, Romania). 1 st part. Acta Palaeontologica Romaniae, 14 (2): 3 - 34." type="journal article" year="2018">Iamandei, Iamandei and Grădinaru, 2018</bibRefCitation>
</taxonomicName>
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<paragraph id="BC0E36931D5EFFAB9749E224FC0B58EB" blockId="18.[816,1436,389,811]" pageId="18" pageNumber="43">
-
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<emphasis id="8EC5EA811D5EFFAB9732E225FBA1580A" box="[843,1141,588,612]" italics="true" pageId="18" pageNumber="43">Brachyoxylon cristianicum</emphasis>
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