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<document ENCODING="UTF8" ID-DOI="http://doi.org/10.5281/zenodo.6172619" ID-GBIF-Dataset="9998aa6b-9348-45f8-82a1-55a07ac10bad" ID-GBIF-Taxon="100105345" ID-HNS-Pub="26064" ID-ISSN="175-5326" ID-Zenodo-Dep="6172619" ModsDocAuthor="Hita Garcia, F. & B. L. Fisher" ModsDocDate="2012" ModsDocID="26064" ModsDocOrigin="http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf" ModsDocTitle="The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups." checkinTime="1373743554799" checkinUser="donat" docAuthor="Hita Garcia, F. & B. L. Fisher" docDate="2012" docId="08860F754A634F237BBB4702887E39D5" docLanguage="en" docName="26064.xml" docOrigin="Zootaxa 3592" docSource="http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf" docTitle="Tetramorium elf Garcia & Fisher, 2012, sp. n." docType="treatment" docVersion="11" lastPageNumber="37" masterDocId="FFBCFFC2FFB3FFACFE12FFF966101E6A" masterDocTitle="The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups." masterLastPageNumber="85" masterPageNumber="1" pageNumber="34" updateTime="1645306871304" updateUser="ExternalLinkService">
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<mods:title>The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups.</mods:title>
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<mods:title id="8815C89FCF9A1E46AB91A88ED50D4CD5">The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups.</mods:title>
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<mods:namePart>Hita Garcia, F.</mods:namePart>
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<mods:namePart>B. L. Fisher</mods:namePart>
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<mods:namePart id="A1A881DFFF82BF7BEF59C07E8E9E78F1">B. L. Fisher</mods:namePart>
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<mods:date>2012</mods:date>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172619" ID-GBIF-Taxon="100105345" ID-Zenodo-Dep="6172619" LSID="urn:lsid:plazi:treatment:08860F754A634F237BBB4702887E39D5" httpUri="http://treatment.plazi.org/id/08860F754A634F237BBB4702887E39D5" lastPageNumber="37" pageNumber="34">
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<subSubSection type="nomenclature">
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<paragraph box="[118,543,1292,1312]" pageId="33" pageNumber="34" type="mainText">
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<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="elf">
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<treatment id="08860F754A634F237BBB4702887E39D5" ID-DOI="http://doi.org/10.5281/zenodo.6172619" ID-GBIF-Taxon="100105345" ID-Zenodo-Dep="6172619" LSID="urn:lsid:plazi:treatment:08860F754A634F237BBB4702887E39D5" httpUri="http://treatment.plazi.org/id/08860F754A634F237BBB4702887E39D5" lastPageNumber="37" pageNumber="34" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="3BF619A4BFE5024D1F20D813A2B50768" type="nomenclature">
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<paragraph id="CB1B174D5C57E09EA0CE702E5AB5AA81" box="[118,543,1292,1312]" pageId="33" pageNumber="34" type="mainText">
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<taxonomicName id="4C08E43641149C15DDD9116453526D35" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="elf">
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Tetramorium elf
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<authority>Hita</authority>
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<authority id="93E13F64E34F880BF24D07D71876A3A7">Hita</authority>
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</taxonomicName>
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Garcia & Fisher
|
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<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
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<taxonomicNameLabel id="56924C7CC7F10CE830B37CF0CF50150D" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection type="description">
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<paragraph box="[118,476,1319,1338]" pageId="33" pageNumber="34" type="mainText">(Figs. 12, 56, 58, 62, 63, 87, 88, 89, 141)</paragraph>
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<subSubSection id="2CFA9268BE6B3D61CEAF8767B6E73671" type="description">
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<paragraph id="AA2FC580E118F2D3454B663E5BC24BA8" box="[118,476,1319,1338]" pageId="33" pageNumber="34" type="mainText">(Figs. 12, 56, 58, 62, 63, 87, 88, 89, 141)</paragraph>
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||||
</subSubSection>
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||||
<subSubSection type="materials_examined">
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||||
<paragraph box="[118,1122,1370,1586]" pageId="33" pageNumber="34" type="mainText">Holotype worker, MADAGASCAR, Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, sifted litter (leaf mold, rotten wood), collection code BLF08872, 15.-18.XI.2003 (B.L. Fisher et al.) (CASC: CASENT0045788). Paratypes, one worker with same data as holotype (CASC: CASENT0045787); one worker with same data as holotype except sampled from yellow pan trap and collection code BLF08873 (CASC: CASENT0048893); and one worker from Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, ground foragers, collection code BLF09077, 17.XI.2003 (B.L. Fisher) (CASC: CASENT0487782).</paragraph>
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||||
<subSubSection id="6312376B6F536FB6710258C5A0786F27" type="materials_examined">
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||||
<paragraph id="2F1A9401D03D1C5C61A380CDEF8732A4" box="[118,1122,1370,1586]" pageId="33" pageNumber="34" type="mainText">Holotype worker, MADAGASCAR, Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, sifted litter (leaf mold, rotten wood), collection code BLF08872, 15.-18.XI.2003 (B.L. Fisher et al.) (CASC: CASENT0045788). Paratypes, one worker with same data as holotype (CASC: CASENT0045787); one worker with same data as holotype except sampled from yellow pan trap and collection code BLF08873 (CASC: CASENT0048893); and one worker from Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, ground foragers, collection code BLF09077, 17.XI.2003 (B.L. Fisher) (CASC: CASENT0487782).</paragraph>
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</subSubSection>
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||||
<subSubSection type="multiple">
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||||
<caption box="[118,1122,1544,1589]" pageId="34" pageNumber="35" type="caption">
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||||
<paragraph box="[118,1122,1544,1589]" pageId="34" pageNumber="35" type="caption">
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||||
<pageBreakToken pageId="34" pageNumber="35" start="start">FIGURES</pageBreakToken>
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||||
<subSubSection id="D0DB06835571D17550857E10EE3EEE18" type="multiple">
|
||||
<caption id="C1937F408BE641C0CC2E064A8193D200" box="[118,1122,1544,1589]" pageId="34" pageNumber="35" type="caption">
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||||
<paragraph id="AB6167DD93744E84A4754DA1C34739AB" box="[118,1122,1544,1589]" pageId="34" pageNumber="35" type="caption">
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<pageBreakToken id="C3160E86E4BF7FE6973CE9B91F3DE391" pageId="34" pageNumber="35" start="start">FIGURES</pageBreakToken>
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87-89.
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="elf">T. elf,</taxonomicName>
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<taxonomicName id="182B4266173CD3D3DF023ACC738183EE" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="elf">T. elf,</taxonomicName>
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holotype (CASENT0045788). 87. Body in profile. 88. Body in dorsal view. 89. Head in full-face view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection type="diagnosis">
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<paragraph box="[118,208,118,137]" pageId="35" pageNumber="36" type="mainText">
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<pageBreakToken pageId="35" pageNumber="36" start="start">Diagnosis</pageBreakToken>
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||||
<subSubSection id="3B5A5E54B6DB01F37FAB913C05102B63" type="diagnosis">
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||||
<paragraph id="86740686507EDE0C122BCAF7EE04B190" box="[118,208,118,137]" pageId="35" pageNumber="36" type="mainText">
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||||
<pageBreakToken id="4EED36C9BED32C0B9A04D8CDB07DB239" pageId="35" pageNumber="36" start="start">Diagnosis</pageBreakToken>
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</paragraph>
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<paragraph box="[118,1121,146,246]" pageId="35" pageNumber="36" type="mainText">
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">Tetramorium elf</taxonomicName>
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<paragraph id="E57D438DD0563C275D76A23F3167D56D" box="[118,1121,146,246]" pageId="35" pageNumber="36" type="mainText">
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||||
<taxonomicName id="9185167678FDC734D0A48B688E080C91" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">Tetramorium elf</taxonomicName>
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is easily separable from the remainder of the species group due to the following combination of characters: propodeal spines extremely long (PSLI 59-64); petiolar node around 1.2 times higher than long (LPeI 80-83); posterodorsal corner of petiole not strongly protruding posteriorly; mandibles unsculptured, smooth, and shiny; body of yellowish colour.
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</paragraph>
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</subSubSection>
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<subSubSection type="description">
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<paragraph box="[118,226,281,299]" pageId="35" pageNumber="36" type="mainText">Description</paragraph>
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<paragraph box="[118,1121,308,435]" pageId="35" pageNumber="36" type="mainText">HL 0.97-1.13 (1.07); HW 0.92-1.06 (1.02); SL 0.76-0.84 (0.81); EL 0.16-0.19 (0.18); PH 0.51-0.65 (0.55); PW 0.68-0.76 (0.73); WL 1.29-1.41 (1.36); PSL 0.59-0.70 (0.66); PTL 0.33-0.37 (0.35); PTH 0.40-0.45 (0.42); PTW 0.29-0.35 (0.32); PPL 0.33-0.38 (0.37); PPH 0.39-0.46 (0.42); PPW 0.35-0.41 (0.38); CI 92-96 (95); SI 78-83 (80); OI 17-18 (18); DMI 53-54 (54); LMI 38-46 (40); PSLI 59-64 (62); PeNI 43-45 (44); LPeI 80-83 (82); DPeI 88-96 (92); PpNI 51-54 (52); LPpI 84-93 (88); DPpI 100-108 (104); PPI 117-122 (119) (ten measured).</paragraph>
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<paragraph box="[118,1122,443,1058]" pageId="35" pageNumber="36" type="mainText">Head longer than wider (CI 95-96). Anterior clypeal margin medially impressed. Posterior head margin only weakly concave. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short, not reaching posterior head margin (SI 78-83). Eyes small (OI 17-18). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-46). Propodeal spines extremely long, spinose, and acute (PSLI 59-64); propodeal lobes short, triangular, and blunt. Petiolar node in profile rectangular nodiform with comparatively rounded angles, around 1.2 times higher than long (LPeI 80-83), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum straight to weakly convex; node in dorsal view around 1.1 times longer than wide (DPeI 88-96). Postpetiole in profile subglobular, approximately 1.1 to 1.2 times higher than long (LPpI 84-93); in dorsal view as wide as long to weakly wider than long (DPpI 100-108). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view approximately 1.2 times wider than petiolar node (PPI 117-122). Mandibles unsculptured, smooth, and shining; clypeus with longitudinally rugulose, usually with three rugulae, median rugula better developed than lateral rugulae; cephalic dorsum between frontal carinae with eight to ten longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae never with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose without cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae with weak, superficial punctate or rugulose sculpture only. Waist segments with weak to moderate rugulose sculpture, especially weakly developed laterally. Ground sculpture everywhere on body faint to absent. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. First gastral tergite without appressed pubescence. Anterior edges of antennal scapes with subdecumbent to erect standing hairs. Body of uniform yellowish colour.</paragraph>
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<subSubSection id="12FB479296612624E1932857AA350D0F" type="description">
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<paragraph id="EDA58673CA2E66531CB8106548A8233B" box="[118,226,281,299]" pageId="35" pageNumber="36" type="mainText">Description</paragraph>
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<paragraph id="030FB7890C9236E3AF1B1CFDAB7F802D" box="[118,1121,308,435]" pageId="35" pageNumber="36" type="mainText">HL 0.97-1.13 (1.07); HW 0.92-1.06 (1.02); SL 0.76-0.84 (0.81); EL 0.16-0.19 (0.18); PH 0.51-0.65 (0.55); PW 0.68-0.76 (0.73); WL 1.29-1.41 (1.36); PSL 0.59-0.70 (0.66); PTL 0.33-0.37 (0.35); PTH 0.40-0.45 (0.42); PTW 0.29-0.35 (0.32); PPL 0.33-0.38 (0.37); PPH 0.39-0.46 (0.42); PPW 0.35-0.41 (0.38); CI 92-96 (95); SI 78-83 (80); OI 17-18 (18); DMI 53-54 (54); LMI 38-46 (40); PSLI 59-64 (62); PeNI 43-45 (44); LPeI 80-83 (82); DPeI 88-96 (92); PpNI 51-54 (52); LPpI 84-93 (88); DPpI 100-108 (104); PPI 117-122 (119) (ten measured).</paragraph>
|
||||
<paragraph id="06748E29B8245526248A1F4B88188E83" box="[118,1122,443,1058]" pageId="35" pageNumber="36" type="mainText">Head longer than wider (CI 95-96). Anterior clypeal margin medially impressed. Posterior head margin only weakly concave. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short, not reaching posterior head margin (SI 78-83). Eyes small (OI 17-18). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-46). Propodeal spines extremely long, spinose, and acute (PSLI 59-64); propodeal lobes short, triangular, and blunt. Petiolar node in profile rectangular nodiform with comparatively rounded angles, around 1.2 times higher than long (LPeI 80-83), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum straight to weakly convex; node in dorsal view around 1.1 times longer than wide (DPeI 88-96). Postpetiole in profile subglobular, approximately 1.1 to 1.2 times higher than long (LPpI 84-93); in dorsal view as wide as long to weakly wider than long (DPpI 100-108). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view approximately 1.2 times wider than petiolar node (PPI 117-122). Mandibles unsculptured, smooth, and shining; clypeus with longitudinally rugulose, usually with three rugulae, median rugula better developed than lateral rugulae; cephalic dorsum between frontal carinae with eight to ten longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae never with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose without cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae with weak, superficial punctate or rugulose sculpture only. Waist segments with weak to moderate rugulose sculpture, especially weakly developed laterally. Ground sculpture everywhere on body faint to absent. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. First gastral tergite without appressed pubescence. Anterior edges of antennal scapes with subdecumbent to erect standing hairs. Body of uniform yellowish colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,1093,1111]" pageId="35" pageNumber="36" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1122,1121,1221]" pageId="35" pageNumber="36" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">Tetramorium elf</taxonomicName>
|
||||
<subSubSection id="8583C54A6E15536F940B06341201CB45" type="discussion">
|
||||
<paragraph id="1C3F20FF4329E12B185FC3333462067D" box="[118,170,1093,1111]" pageId="35" pageNumber="36" type="mainText">Notes</paragraph>
|
||||
<paragraph id="D3EDAC888D2B3AB029E21C5196C1F402" box="[118,1122,1121,1221]" pageId="35" pageNumber="36" type="mainText">
|
||||
<taxonomicName id="79BC9C2F8ACAD4FF563B51701D936F2C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">Tetramorium elf</taxonomicName>
|
||||
was collected from rainforests in Makirovana, Marojejy, Antalaha, and Ambalagoavy at elevations from 625 to 900 m. Despite being known from four localities, the available material consists of just 12 specimens, making
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
<taxonomicName id="C39ABD6569BB72A0B7541B6F19CED848" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
a relatively rarely sampled species. The microhabitat of these ants is not clear since the specimens were collected from the ground, leaf litter, and malaise or pan traps.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,1229,1465]" pageId="35" pageNumber="36" type="mainText">
|
||||
<paragraph id="BB02301FD6B387A06F8B0AE1BA9F26B1" box="[118,1122,1229,1465]" pageId="35" pageNumber="36" type="mainText">
|
||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="D365019CB054E6548441036464044178" rank="species">new species</taxonomicNameLabel>
|
||||
is highly conspicuous and not easily confused with another member of the complex. The small eyes (OI 17-18), extremely long propodeal spines (PSLI 59-64), smooth and shiny mandibles, and the distinct yellowish colouration are quite unique in the complex and group. Nevertheless,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
<taxonomicName id="0544C8B856A28744D668FB50FAE2AF31" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
seems to be morphologically near to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="9A60C5B15613E77D57FDAABF6AA55B10" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum.</taxonomicName>
|
||||
<taxonomicName id="3C5FDADDCEE01C9443F0E4AB63557A6E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum.</taxonomicName>
|
||||
All three share the small eyes (OI 16-19), the enormously developed propodeal spines (PSLI 46-64), usually higher than 50), and relatively small and blunted propodeal lobes. The conspicuous yellowish body colour, longer antennal scapes (SI 78-83), and a lower petiolar node (LPeI 80-83), however, separate
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
<taxonomicName id="4D90801356C417C4B3CB2DD4C1E683E0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
from the very dark brown to black
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="FEF168D0F31B4C15523B40018456C007" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
(SI 67-74; LPeI 64-74). In addition, the likewise very darkly coloured
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="0EDA7FEBE66719D6C828BC4C851E4512" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
has strongly sculptured mandibles, which contrasts with the unsculptured mandibles of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">T. elf.</taxonomicName>
|
||||
<taxonomicName id="C4F6F48673BB63E7F4F48CF80DCD1F3C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="elf">T. elf.</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,1499,1518]" pageId="35" pageNumber="36" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,562,1527,1573]" pageId="35" pageNumber="36" type="mainText">
|
||||
<subSubSection id="37C323A32921974CA0E1E42949E8D676" type="etymology">
|
||||
<paragraph id="6A82DF9ECCB0B47A883038EF62AFC9D8" box="[118,219,1499,1518]" pageId="35" pageNumber="36" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="6060BFE77F9D0093383D8AC04C6847E6" box="[118,562,1527,1573]" pageId="35" pageNumber="36" type="mainText">
|
||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="A6CF98B44311CA630CC711D7DF87C40A" rank="species">new species</taxonomicNameLabel>
|
||||
is named after the "supernatural epithet is a noun in apposition, and thus invariant.
|
||||
</paragraph>
|
||||
<paragraph box="[570,1008,1527,1546]" pageId="35" pageNumber="36" type="mainText">beings" from Old Norse and Old English myths.</paragraph>
|
||||
<paragraph box="[1016,1120,1527,1546]" pageId="35" pageNumber="36" type="mainText">The species</paragraph>
|
||||
<paragraph id="14C4673BC12778C12C925E6371DFB287" box="[570,1008,1527,1546]" pageId="35" pageNumber="36" type="mainText">beings" from Old Norse and Old English myths.</paragraph>
|
||||
<paragraph id="575A49D634AECBC3D7B3175F80C7D991" box="[1016,1120,1527,1546]" pageId="35" pageNumber="36" type="mainText">The species</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,118,137]" pageId="36" pageNumber="37" type="mainText">
|
||||
<pageBreakToken pageId="36" pageNumber="37" start="start">Material</pageBreakToken>
|
||||
<subSubSection id="F0761119C13CAB20977E1B9AD066DD99" type="materials_examined">
|
||||
<paragraph id="20C5E9A94076A0416D2288C892877433" box="[118,295,118,137]" pageId="36" pageNumber="37" type="mainText">
|
||||
<pageBreakToken id="7D36A97E50B542C14991B9177BFFB97E" pageId="36" pageNumber="37" start="start">Material</pageBreakToken>
|
||||
examined
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,146,362]" pageId="36" pageNumber="37" type="mainText">MADAGASCAR: Antsiranana, 4 km SW of Antalaha, 31.I.1990 (G.D. Alpert); Antsiranana, Makirovana forest, 14.16666 S, 49.95 E, 715 m, rainforest, 30.IV.2011 (B.L. Fisher et al.); Antsiranana, Makirovana forest, 14.16506 S, 49.9477 E, 900 m, rainforest, 30.IV.-2.V.2011 (B.L. Fisher et al.); Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, 15.-18.XI.2003 (B.L. Fisher et al.); Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, 17.XI.2003 (B.L. Fisher); Fianarantsoa, Foret d'Ambalagoavy Nord, Ikongo, Ambatombe, 21.8275 S, 47.33889 E, 625 m, 1.XII.2000 (R. Harin'Hala & M.E. Irwin).</paragraph>
|
||||
<paragraph id="7BEEE26E87236DD0C02EA755756861F1" box="[118,1122,146,362]" pageId="36" pageNumber="37" type="mainText">MADAGASCAR: Antsiranana, 4 km SW of Antalaha, 31.I.1990 (G.D. Alpert); Antsiranana, Makirovana forest, 14.16666 S, 49.95 E, 715 m, rainforest, 30.IV.2011 (B.L. Fisher et al.); Antsiranana, Makirovana forest, 14.16506 S, 49.9477 E, 900 m, rainforest, 30.IV.-2.V.2011 (B.L. Fisher et al.); Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, 15.-18.XI.2003 (B.L. Fisher et al.); Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, 17.XI.2003 (B.L. Fisher); Fianarantsoa, Foret d'Ambalagoavy Nord, Ikongo, Ambatombe, 21.8275 S, 47.33889 E, 625 m, 1.XII.2000 (R. Harin'Hala & M.E. Irwin).</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,97 +1,97 @@
|
|||
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|
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|
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<taxonomicName LSID="http://species-id.net/wikiDolichosoma_foveicolle" authority="(Kirby, 1837) **" class="Insecta" family="Melyridae" genus="Dolichosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolichosoma foveicolle" order="Coleoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="foveicolle">Dolichosoma foveicolle (Kirby, 1837)**</taxonomicName>
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|
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|
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|
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<taxonomicName id="3B20B33BF261743B8ACBB21D21A7FA43" LSID="http://species-id.net/wikiDolichosoma_foveicolle" authority="(Kirby, 1837) **" genus="Dolichosoma" lsidName="Dolichosoma foveicolle" pageId="9" pageNumber="150" rank="species" species="foveicolle">Dolichosoma foveicolle (Kirby, 1837)**</taxonomicName>
|
||||
Map 10
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="10" lastPageNumber="151" pageId="9" pageNumber="150" type="material examined">
|
||||
<paragraph pageId="9" pageNumber="150">Material examined.</paragraph>
|
||||
<paragraph lastPageId="10" lastPageNumber="151" pageId="9" pageNumber="150">
|
||||
<subSubSection id="4C30667356BFB94139E011AB174DEBF6" lastPageId="10" lastPageNumber="151" pageId="9" pageNumber="150" type="material examined">
|
||||
<paragraph id="EA4E4912ED40228D88A2D4EBBEE1D7EB" pageId="9" pageNumber="150">Material examined.</paragraph>
|
||||
<paragraph id="01824657C084475C0F972EF41C7A8320" lastPageId="10" lastPageNumber="151" pageId="9" pageNumber="150">
|
||||
New Brunswick, Albert Co., Shepody N.W.A.,
|
||||
<normalizedToken originalValue="Mary’s">Mary's</normalizedToken>
|
||||
<normalizedToken id="26C3B2AF3A3A67CCB1693ED1D9D6E2A2" originalValue="Mary’s">Mary's</normalizedToken>
|
||||
Point Section,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.732">45.7320°N</geoCoordinate>
|
||||
<geoCoordinate id="B7206395D4C5A529537699B972072F6D" direction="north" orientation="latitude" precision="5" value="45.732">45.7320°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-64.6765">64.6765°W</geoCoordinate>
|
||||
<geoCoordinate id="D255996056FB834E443D869B9A814DC8" direction="west" orientation="longitude" precision="5" value="-64.6765">64.6765°W</geoCoordinate>
|
||||
, 16.VI.2004, R. P. Webster, margin of salt marsh near for
|
||||
<pageBreakToken pageId="10" pageNumber="151" start="start">est</pageBreakToken>
|
||||
<pageBreakToken id="84E6B454E6535CD0E78E270D0717EBE8" pageId="10" pageNumber="151" start="start">est</pageBreakToken>
|
||||
, sweeping (1, RWC). Gloucester Co., near Acadian Historical Village,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="47.7873">47.7873°N</geoCoordinate>
|
||||
<geoCoordinate id="96509A5F177A8B9EF1F3AE63D625BB0A" direction="north" orientation="latitude" precision="5" value="47.7873">47.7873°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-65.0756">65.0756°W</geoCoordinate>
|
||||
<geoCoordinate id="478D78B0A678202D7FC7396873574C73" direction="west" orientation="longitude" precision="5" value="-65.0756">65.0756°W</geoCoordinate>
|
||||
, 29.VI.2006, R. P. Webster, inland margin of salt marsh, sweeping vegetation (6, NBM, RWC). Sunbury Co., Burton, near Sunpoke Lake,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.7662">45.7662°N</geoCoordinate>
|
||||
<geoCoordinate id="11F8B2730E11422E29B2BE0C109C9F07" direction="north" orientation="latitude" precision="5" value="45.7662">45.7662°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.5526">66.5526°W</geoCoordinate>
|
||||
<geoCoordinate id="1884BE76FC64312D8C746E43EAB23C47" direction="west" orientation="longitude" precision="5" value="-66.5526">66.5526°W</geoCoordinate>
|
||||
, 20.VI.2007, R. P. Webster, seasonally flooded marsh, sweeping vegetation (1, RWC). York Co., Mazerolle Settlement,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.8765">45.8765°N</geoCoordinate>
|
||||
<geoCoordinate id="E6C44DECFAF07C9219FE08BB2A64A7C2" direction="north" orientation="latitude" precision="5" value="45.8765">45.8765°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.826">66.8260°W</geoCoordinate>
|
||||
<geoCoordinate id="A96AB0386819C47BBF509A94870D327E" direction="west" orientation="longitude" precision="5" value="-66.826">66.8260°W</geoCoordinate>
|
||||
, 8.VI.2008, R. P. Webster, beaver meadow, sweeping vegetation along brook margin (2, RWC).
|
||||
</paragraph>
|
||||
<caption pageId="10" pageNumber="151">
|
||||
<paragraph pageId="10" pageNumber="151">
|
||||
<caption id="DB30DAEB6CB0B44CA137E45456158122" pageId="10" pageNumber="151">
|
||||
<paragraph id="293FB22A84CCFD4A4877B98DC18B2482" pageId="10" pageNumber="151">
|
||||
Map 10. Collection localities in New Brunswick, Canada of
|
||||
<taxonomicName class="Insecta" family="Melyridae" genus="Dolichosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolichosoma foveicolle" order="Coleoptera" pageId="10" pageNumber="151" phylum="Arthropoda" rank="species" species="foveicolle">Dolichosoma foveicolle</taxonomicName>
|
||||
<taxonomicName id="4561AFA8F9A1025ADA622EAD10BE9662" genus="Dolichosoma" lsidName="Dolichosoma foveicolle" pageId="10" pageNumber="151" rank="species" species="foveicolle">Dolichosoma foveicolle</taxonomicName>
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="10" pageNumber="151" type="collection and habitat data">
|
||||
<paragraph pageId="10" pageNumber="151">Collection and habitat data.</paragraph>
|
||||
<paragraph pageId="10" pageNumber="151">Adults were found in June on the inner margin of salt marshes, seasonally flooded (freshwater) marshes, and in a beaver meadow, and were collected by sweeping the marsh vegetation.</paragraph>
|
||||
<subSubSection id="0FC1ADAE88ADF50B25E147569C95A6E6" pageId="10" pageNumber="151" type="collection and habitat data">
|
||||
<paragraph id="3C0E90B5D4752100FB17EE9F24F7F28D" pageId="10" pageNumber="151">Collection and habitat data.</paragraph>
|
||||
<paragraph id="4E836D5366DBED2F223F1B6B508C1E77" pageId="10" pageNumber="151">Adults were found in June on the inner margin of salt marshes, seasonally flooded (freshwater) marshes, and in a beaver meadow, and were collected by sweeping the marsh vegetation.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="10" pageNumber="151" type="distribution">
|
||||
<paragraph pageId="10" pageNumber="151">Distribution in Canada and Alaska.</paragraph>
|
||||
<paragraph pageId="10" pageNumber="151">
|
||||
<subSubSection id="2DE24E43239BEA77120A035FEF3DF885" pageId="10" pageNumber="151" type="distribution">
|
||||
<paragraph id="6F67C77FA004EC832A66296014661849" pageId="10" pageNumber="151">Distribution in Canada and Alaska.</paragraph>
|
||||
<paragraph id="C1ADC36A14E06A0878BE54B05DB5DDC9" pageId="10" pageNumber="151">
|
||||
BC, AB, MB, ON, QC, NB, (
|
||||
<bibRefCitation pageId="10" pageNumber="151">Bright 1991</bibRefCitation>
|
||||
<bibRefCitation id="1B873719DF96A1534AADBD95FBC7EB1D" pageId="10" pageNumber="151">Bright 1991</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,204 +1,203 @@
|
|||
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|
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electrum
|
||||
<authority>Bolton</authority>
|
||||
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|
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,
|
||||
</taxonomicName>
|
||||
1979
|
||||
</paragraph>
|
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</subSubSection>
|
||||
<subSubSection type="description">
|
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<paragraph box="[118,410,146,165]" pageId="30" pageNumber="31" type="mainText">(Figs. 31, 53, 55, 84, 85, 86, 141)</paragraph>
|
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<paragraph id="1F7A9D300717ED5BB6908EB458BB01CB" box="[118,410,146,165]" pageId="30" pageNumber="31" type="mainText">(Figs. 31, 53, 55, 84, 85, 86, 141)</paragraph>
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|
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<subSubSection type="reference_group">
|
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|
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|
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<taxonomicName id="33B80D63F1ECAC307376934FF9173D8F" authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="electrum">
|
||||
Tetramorium electrum
|
||||
<authority>Bolton</authority>
|
||||
<authority id="89376E2AB6250DD3ADAD725698908211">Bolton</authority>
|
||||
,
|
||||
</taxonomicName>
|
||||
1979:144. Holotype worker, MADAGASCAR, Rte d'Anosibe, km 33, forest humus and litter, AB 44, 4.-12.IV.1975 (A. Peyrieras) (MCZ: CASENT0280589) [examined]. Paratypes, 11 workers with same data as holotype, and one worker from vic. Andasibe (= Perinet) 950-980 m, 2.-6.II.1977 (W.L. & D.E. Brown) (BMNH: CASENT0102350; MCZ: CASENT0280850; MCZ_paratype_32378) [examined].
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,325,344]" pageId="30" pageNumber="31" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,354,457]" pageId="30" pageNumber="31" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
<subSubSection id="2775ADEA66B3E585D67CFBE648E9D524" type="diagnosis">
|
||||
<paragraph id="2FF8D1AAE17D3F7FE7EA4239B1480749" box="[118,208,325,344]" pageId="30" pageNumber="31" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="F7459EDB4431959414D56961A5930D0C" box="[118,1121,354,457]" pageId="30" pageNumber="31" type="mainText">
|
||||
<taxonomicName id="164510D890A15ABFF2CBFE2B28B86FDD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
is easily recognisable within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="37467A242C8361E65CDE5931ADDB6C76" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group in the Malagasy region due to the following combination of characters: propodeal spines very long to extremely long (PSLI 46-52); petiolar node around 1.3 to 1.6 times higher than long (LPeI 64-74); posterodorsal corner of petiole not strongly protruding posteriorly; body dark brown to black in colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,494,513]" pageId="30" pageNumber="31" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,523,654]" pageId="30" pageNumber="31" type="mainText">HL 0.89-1.20 (1.07); HW 0.87-1.25 (1.08); SL 0.65-0.84 (0.77); EL 0.16-0.22 (0.19); PH 0.49-0.63 (0.55); PW 0.63-0.82 (0.75); WL 1.14-1.59 (1.34); PSL 0.44-0.60 (0.53); PTL 0.28-0.35 (0.31); PTH 0.38-0.47 (0.44); PTW 0.29-0.37 (0.33); PPL 0.32-0.40 (0.36); PPH 0.38-0.48 (0.43); PPW 0.37-0.44 (0.40); CI 98-104 (100); SI 67-74 (71); OI 16-19 (17); DMI 52-58 (56); LMI 39-44 (41); PSLI 46-52 (50); PeNI 41-46 (44); LPeI 64-74 (71); DPeI 100-114 (105); PpNI 50-59 (53); LPpI 77-89 (84); DPpI 106-116 (110); PPI 115-128 (122) (20 measured).</paragraph>
|
||||
<paragraph box="[118,1122,663,1442]" pageId="30" pageNumber="31" type="mainText">Head approximately as long as wide (CI 98-104). Anterior clypeal margin medially impressed. Posterior head margin weakly to moderately concave. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes very short, not reaching posterior head margin (SI 67-74). Eyes small (OI 16-19). Mesosomal outline in profile weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 39-44). Propodeal spines very long, spinose, and acute (PSLI 46-52); propodeal lobes strongly reduced, generally vestigial, sometimes very short, broadly triangular and blunt. Petiolar node in profile rectangular nodiform with moderately to sharply defined margins, around 1.3 to 1.6 times higher than long (LPeI 64-74), anterior and posterior faces often not parallel narrowing towards dorsum, anterodorsal and posterodorsal margins generally at approximately same height, dorsum straight to weakly convex, sometimes anterodorsal margin slightly higher than posterodorsal with dorsum tapering weakly backwards posteriorly; node in dorsal view as wide as long to 1.2 times wider than long (DPeI 100-114). Postpetiole in profile subglobular, approximately 1.1 to 1.3 times higher than long (LPpI 77-89); in dorsal view around 1.1 times wider than long (DPpI 106-116). Postpetiole in profile usually appearing a bit more voluminous than petiolar node, in dorsal view approximately 1.1 to 1.3 times wider than petiolar node (PPI 115-128). Mandibles ranging from completely unsculptured, smooth, and shining to finely striate, often partly striate and partly smooth; clypeus with three to six longitudinal rugulae, rugulae often broken and irregularly arranged; cephalic dorsum between frontal carinae with 8 to 12 longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae only very rarely with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose with few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose, often rugae on lateral mesosoma more irregularly arranged than on the dorsum. Forecoxae usually unsculptured, smooth, and shining, sometimes with superficial sculpture, rarely weakly reticulate-punctate or reticulate-rugulose. Waist segments with reticulate-rugose/rugulose to longitudinally rugose/rugulose sculpture, often weaker laterally. Ground sculpture everywhere on body faint to absent. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. First gastral tergite without appressed pubescence. Anterior edges of antennal scapes with decumbent to erect standing hairs. Body very dark brown to black in colour, sometimes of lighter brown.</paragraph>
|
||||
<subSubSection id="EE96297D480B9F3D50C3DAF5FFCFF5F3" type="description">
|
||||
<paragraph id="DB2235FD7EC32136EF2212878A3AFFAC" box="[118,226,494,513]" pageId="30" pageNumber="31" type="mainText">Description</paragraph>
|
||||
<paragraph id="58D22118D33339E8FA314A7AF79261F1" box="[118,1121,523,654]" pageId="30" pageNumber="31" type="mainText">HL 0.89-1.20 (1.07); HW 0.87-1.25 (1.08); SL 0.65-0.84 (0.77); EL 0.16-0.22 (0.19); PH 0.49-0.63 (0.55); PW 0.63-0.82 (0.75); WL 1.14-1.59 (1.34); PSL 0.44-0.60 (0.53); PTL 0.28-0.35 (0.31); PTH 0.38-0.47 (0.44); PTW 0.29-0.37 (0.33); PPL 0.32-0.40 (0.36); PPH 0.38-0.48 (0.43); PPW 0.37-0.44 (0.40); CI 98-104 (100); SI 67-74 (71); OI 16-19 (17); DMI 52-58 (56); LMI 39-44 (41); PSLI 46-52 (50); PeNI 41-46 (44); LPeI 64-74 (71); DPeI 100-114 (105); PpNI 50-59 (53); LPpI 77-89 (84); DPpI 106-116 (110); PPI 115-128 (122) (20 measured).</paragraph>
|
||||
<paragraph id="AA2E120626B0DAAFDA2B8DF269A32362" box="[118,1122,663,1442]" pageId="30" pageNumber="31" type="mainText">Head approximately as long as wide (CI 98-104). Anterior clypeal margin medially impressed. Posterior head margin weakly to moderately concave. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes very short, not reaching posterior head margin (SI 67-74). Eyes small (OI 16-19). Mesosomal outline in profile weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 39-44). Propodeal spines very long, spinose, and acute (PSLI 46-52); propodeal lobes strongly reduced, generally vestigial, sometimes very short, broadly triangular and blunt. Petiolar node in profile rectangular nodiform with moderately to sharply defined margins, around 1.3 to 1.6 times higher than long (LPeI 64-74), anterior and posterior faces often not parallel narrowing towards dorsum, anterodorsal and posterodorsal margins generally at approximately same height, dorsum straight to weakly convex, sometimes anterodorsal margin slightly higher than posterodorsal with dorsum tapering weakly backwards posteriorly; node in dorsal view as wide as long to 1.2 times wider than long (DPeI 100-114). Postpetiole in profile subglobular, approximately 1.1 to 1.3 times higher than long (LPpI 77-89); in dorsal view around 1.1 times wider than long (DPpI 106-116). Postpetiole in profile usually appearing a bit more voluminous than petiolar node, in dorsal view approximately 1.1 to 1.3 times wider than petiolar node (PPI 115-128). Mandibles ranging from completely unsculptured, smooth, and shining to finely striate, often partly striate and partly smooth; clypeus with three to six longitudinal rugulae, rugulae often broken and irregularly arranged; cephalic dorsum between frontal carinae with 8 to 12 longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae only very rarely with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose with few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose, often rugae on lateral mesosoma more irregularly arranged than on the dorsum. Forecoxae usually unsculptured, smooth, and shining, sometimes with superficial sculpture, rarely weakly reticulate-punctate or reticulate-rugulose. Waist segments with reticulate-rugose/rugulose to longitudinally rugose/rugulose sculpture, often weaker laterally. Ground sculpture everywhere on body faint to absent. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. First gastral tergite without appressed pubescence. Anterior edges of antennal scapes with decumbent to erect standing hairs. Body very dark brown to black in colour, sometimes of lighter brown.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,1479,1497]" pageId="30" pageNumber="31" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1121,1507,1582]" pageId="30" pageNumber="31" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
<subSubSection id="F42758FB2A4BCCD05C261ECA0143B977" type="discussion">
|
||||
<paragraph id="52A01025573646D3F2E668CBA6C2C770" box="[118,170,1479,1497]" pageId="30" pageNumber="31" type="mainText">Notes</paragraph>
|
||||
<paragraph id="BA380B08FC82BE58425D91FBE29EA4F6" box="[118,1121,1507,1582]" pageId="30" pageNumber="31" type="mainText">
|
||||
<taxonomicName id="CCF4437B89E271356CD4B0D897A878BD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
is one of the most common and conspicuous genus members encountered in the eastern rainforests of Madagascar. Its distribution ranges from Andohahela in the southeast to Marojejy in the northeast. It is found at elevations from 25 to 1080 m, and appears to prefer the leaf litter stratum.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,118,417]" pageId="32" pageNumber="33" type="mainText">
|
||||
<pageBreakToken pageId="32" pageNumber="33" start="start">Within</pageBreakToken>
|
||||
<paragraph id="87F634CDE16F35EACE7090828C2AFAE1" box="[118,1122,118,417]" pageId="32" pageNumber="33" type="mainText">
|
||||
<pageBreakToken id="BB1B5CC34C1EB4DB3D2A4A5C8DC98E6E" pageId="32" pageNumber="33" start="start">Within</pageBreakToken>
|
||||
the species complex
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="F1D90854B885BB60884CDC369A199D88" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
can be grouped together with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
<taxonomicName id="3CCC21A35A594E4A5E6FA725D52FD2FE" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="D444A321516A565A78CADA7741207ADA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
on the basis of shared morphology.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="elf">Tetramorium elf</taxonomicName>
|
||||
<taxonomicName id="D603E426F13813BABF11ACE2BE8450D3" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="elf">Tetramorium elf</taxonomicName>
|
||||
is not easily confused with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum,</taxonomicName>
|
||||
<taxonomicName id="0D158F6413A0EFDD2CF868C826447C93" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum,</taxonomicName>
|
||||
however, since the first is yellowish in colour with longer antennal scapes (SI 78-83) and petiolar node (DPeI 88-96), whereas
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="A4AB2B5B84C141822E8D9618B54D5D23" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
is very dark brown to black in colour and the scapes and petiolar node are shorter (SI 67-74; DPeI 100-114). The head of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
<taxonomicName id="C72BD7285A1D1F549431A255677BFC9D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
(CI 92-96) is also narrower than that of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="29D6D3C1935AD0A292921A149B67C530" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
(CI 98-104). More challenging is the separation of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="B9AA105C771F1A0EC2C8D50455B0EF8F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="6483153329BF4B84CD4FEAFD4056E253" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
since they are superficially very similar. Much like
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="32" pageNumber="33" rank="species" species="elf">
|
||||
<taxonomicName id="D66C7C335AF2E01F5AD131CE05565227" authorityName="," genus="Tetamorium" pageId="32" pageNumber="33" rank="species" species="elf">
|
||||
T. elf
|
||||
<authority>,</authority>
|
||||
<authority id="82255C0119C09CF6A33E5C72A4B3B0E0">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="F7425272CF2215E5B898CB2AA36CED4B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
also has longer antennal scapes (SI 81-84), a longer petiolar node (DPeI 87-97), and a less broad head (CI 93-96) compared to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum.</taxonomicName>
|
||||
<taxonomicName id="0B0B9569C5801306F67EABD96AA03BE9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum.</taxonomicName>
|
||||
Another difference is the petiolar node, which is lower and more square in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="687844E820D9BE4C95C11DFB3B613F3E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
(LPeI 77-86) but higher and narrows towards the dorsum in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="DC3C73253590F0D20C3772BB5654C76A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
(LPeI 64-74). Furthermore,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="D8A656AC8A8EAAF056F446D27AD43F4F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
is not likely to be misidentified with the remaining species of the complex due to the combination of very long to extremely long propodeal spines (PSLI 46-52), low and inconspicuous propodeal lobes, and comparatively high petiolar node.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,426,668]" pageId="32" pageNumber="33" type="mainText">
|
||||
<paragraph id="C9E93D44A69ECF169C5B38932A8DA5BE" box="[118,1122,426,668]" pageId="32" pageNumber="33" type="mainText">
|
||||
The general similarity between
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="F717D26F7C8B99609AA96427AF2F2F13" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="0F531AC0D94195ADC72AE900C6893DBC" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
offers the slight possibility that they may belong to the same species; however, this does not seem likely given the material examined in this study.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
<taxonomicName id="EE99165B8C236B3E9393A3F148F73DAB" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
is very common and we were able to examine several hundred specimens. Despite the variation mentioned below, the species seems to be relatively consistent throughout its range in eastern Madagascar.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">Tetramorium isoelectrum</taxonomicName>
|
||||
<taxonomicName id="090B5609F8B9E52FC0D38C5510B8F7FF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="isoelectrum">Tetramorium isoelectrum</taxonomicName>
|
||||
is less common and only found in northeastern Madagascar, mostly north of the distribution range of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum.</taxonomicName>
|
||||
<taxonomicName id="EA2202FFA899CEE6AD8656CB6FC7467E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum.</taxonomicName>
|
||||
However, both species overlap in their distribution ranges and they are found in sympatry in Marojejy. There, both species retain their species-specific characteristics, and can be distinguished using the diagnostic notes provided in this revision. This piece of biogeographic evidence, in combination with the significant morphometric differences between the two species, provides strong support for their heterospecificity.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,677,891]" pageId="32" pageNumber="33" type="mainText">
|
||||
<paragraph id="F595D953DEF8BDE138F72B823648592E" box="[118,1122,677,891]" pageId="32" pageNumber="33" type="mainText">
|
||||
It must be noted that some morphological variation exists within
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum.</taxonomicName>
|
||||
<taxonomicName id="6F335260580D0B944162FD67710A881F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="electrum">T. electrum.</taxonomicName>
|
||||
This fact is not surprising considering its large distribution range in eastern Madagascar. The mandibles are usually finely striate, but some populations have completely unsculptured mandibles while only partly sculptured mandibles are found in other series. The shape of the petiolar node is also somewhat variable. The node is usually rectangular nodiform but comparatively high. In many specimens the anterodorsal margin is slightly higher than the posterodorsal, which causes the dorsum to taper backwards, even if weakly (as in the holotype CASENT0280589). In other specimens the node narrows distinctly towards the dorsum but with the anterior and posterior faces almost mirror-inverted (like in the paratype specimens CASENT0102350 and CASENT0280850).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,928,946]" pageId="32" pageNumber="33" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1128,956,1580]" pageId="32" pageNumber="33" type="mainText">
|
||||
<subSubSection id="ED3006702432916C47D963D90574A2BB" type="materials_examined">
|
||||
<paragraph id="93F4200F95DAF810018C9EA852A77C3F" box="[118,295,928,946]" pageId="32" pageNumber="33" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="F2AF51A364DC44EC4FCDED8B32032A9B" box="[118,1128,956,1580]" pageId="32" pageNumber="33" type="mainText">
|
||||
MADAGASCAR: Antsiranana, Marojejy National Park, Sambava district, 5 km W of Manantenina village, 1st Camp site (Mantella), 14.4382 S, 49.774 E, 487 m, low altitude rainforest, 28.IV.-7.V.2005 (Rin'Ha & Mike); Antsiranana, Parc National de Marojejy, Manantenina River, 28.0 km 38° NE Andapa, 8.2 km 333° NNW Manantenina, 14.4367 S, 49.775 E, 450 m, rainforest, 12.-15.XI.2003 (B.L. Fisher et al.); Antsiranana, Masoala Peninsula, Ambavoany Forest, 15° 12' 28.7'' S, 50° 17' 20'' E, 30 m, primary rainforest, 26.IV.1996 (G.D. Alpert et al.); Fianarantsoa, 45 km S. Ambalavao, 22.2167 S, 47.0167 E, 785 m, rainforest, 24.-25.IX.1993 (B.L. Fisher); Fianarantsoa,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="09841060147B15C5277F8B21FFB7F262" originalValue="Rés">Res</normalizedToken>
|
||||
. Andringitra, 43 km S Ambalavao, 22.2333 S, 47 E, 825 m, rainforest, 4.-5.X.1993 (B.L. Fisher); Fianarantsoa, Parc National Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango, 23.8352 S, 46.9637 E, 940 m, rainforest, 14.XI.2006 (B.L. Fisher et al.); Fianarantsoa, 9.0 km NE Ivohibe, 22.4267 S, 46.9383 E, 900 m, rainforest, 12.-17.XI.1997 (B.L. Fisher); Fianarantsoa,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="27BC60433BD4256E867B983B8B2E21B4" originalValue="Réserve">Reserve</normalizedToken>
|
||||
Speciale Manombo 24.5 km 228° Farafangana, 23.0158 S, 47.719 E, 30 m, rainforest, 20.IV.2006 (B.L. Fisher et al.); Fianarantsoa, Ranomafana Nat. Park, Ambohila, 21° 10' 9'' S, 47° 33' 7'' E, 700 m, montane rainforest, 23.VII.1992 (A. Kingman); Fianarantsoa, Ranomafana Nat. Park, Miaranony forest, 700 m, montane rainforest, 26.X.1992 (E. Raferiarison); Fianarantsoa, P.N. Ranomafana, Tolongoina-Ampasimpotsy, 21.4799 S, 47.5571 E, 577 m, rainforest, 13.III.-1.IV.2003 (V. Clark); Fianarantsoa,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken originalValue="Classée">Classee</normalizedToken>
|
||||
<normalizedToken id="21509068502D9E22F7BE84A99E7B8699" originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="83D5FD3535579177EB910967C4026506" originalValue="Classée">Classee</normalizedToken>
|
||||
Vatovavy, 7.6 km 122
|
||||
<normalizedToken originalValue="º">°</normalizedToken>
|
||||
<normalizedToken id="8C5E42115D7B5D56D1500F63579200E9" originalValue="º">°</normalizedToken>
|
||||
Kianjavato, 21.4 S, 47.94 E, 175 m, rainforest, 6.-8.VI.2005 (B.L. Fisher et al.); Fianarantsoa,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="B8C5CC0AD302242B861D2D0DA60351B1" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Vevembe, 66.6 km 293° Farafangana, 22.791 S, 47.1818 E, 600 m, rainforest, transition to montane forest, 23.IV.2006 (B.L. Fisher et al.); Mahajanga,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="BC35F141F356C5552EF00C05F92A3E0F" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="C0B9A8F32764230B8E63E18E9D2A0454" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Marotandrano, Marotandrano 48.3 km S Mandritsara, 16.2832 S, 48.8144 E, 865 m, transition humid forest, 6.-8.XII.2007 (B.L. Fisher et al.); Toamasina, Montagne d'Akirindro 7.6 km 341° NNW Ambinanitelo, 15.2883 S, 49.5483 E, 600 m, rainforest, 17.-21.III.2003 (B.L. Fisher, C. Griswold et al.); Toamasina, 6.3 km S Ambanizana, Andranobe, 15.6813 S, 49.958 E, 25 m, rainforest, 15.XI.1993 (B.L. Fisher); Toamasina, 6.9 km NE Ambanizana, Ambohitsitondroina, 15.5667 S, 50 E, 825 m, rainforest, 2.-8.XII.1993 (B.L. Fisher); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="8F6A73A19666250E50F1E6BAA698E93B" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="D48982BDB0E657C96A945EA3C8BDF675" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato river, 16.7727 S, 49.2655 E, 450 m, rainforest, 20.-22.II.2010 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="3F5B1B7567F1956FA8D5D7FC658D24A3" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="0ACD21A87F4C3B674DFC189F506CF742" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato River, 16.7691 S, 49.267 E, 475 m, rainforest,
|
||||
</paragraph>
|
||||
<paragraph box="[118,1127,118,1237]" pageId="33" pageNumber="34" type="mainText">
|
||||
<pageBreakToken pageId="33" pageNumber="34" start="start">21</pageBreakToken>
|
||||
<paragraph id="AD6ACE69F5E6D21A9477035CB2CE6205" box="[118,1127,118,1237]" pageId="33" pageNumber="34" type="mainText">
|
||||
<pageBreakToken id="9E21249E0F0405456DA95E4AB27049A6" pageId="33" pageNumber="34" start="start">21</pageBreakToken>
|
||||
.II.2010 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="AC09FDEE2067B411D7563B777B01480D" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="FA8B97B03CE9F469A90A8679E5BE62E0" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato River, 16.7633 S, 49.2669 E, 520 m, rainforest, 22.-24.II.2010 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="7ADFAF452639B5B1EC48E28B80DA9FE2" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="11CCB916D2F1A90CE0B8E1E8B4C757CB" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato river, 16.7702 S, 49.2664 E, 470 m, rainforest, 23.II.2010 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="A3870F56A5A17B08DED65F3784782930" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="844730E67F3ADC28D34A2327CE315376" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato River, 16.7674 S, 49.2681 E, 500 m, rainforest, 23.II.2010 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="341BA44A9E9425B76C3A23EAC3369051" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="27B77BFB89A3EAC0109E0E16927C4001" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato River, 16.8175 S, 49.295 E, 360 m, rainforest, 25.-27.II.2010 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="A6061B67BFA1273E1BC7F1906DFCAE08" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="F54CE55D6453BD0E9A791709D8B2F675" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato River, 16.8121 S, 49.2922 E, 460 m, rainforest, 26.II.2010 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="F4AE477D3E48A70342DBD62A2834D448" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="2E0ED5507DABAF6580080EB7DA0824B5" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato River, 16.8174 S, 49.2925 E, 400 m, rainforest, 26.II.2010 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="6F9A873E27C2492824B641188F458577" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="F5D819E742F2688DB1F2A5116A458042" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato River, 16.8056 S, 49.2951 E, 480 m, rainforest, 27.II.2010 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="9EBA6EA0F7F5D0F149C3E502C08949A9" originalValue="Forêt">Foret</normalizedToken>
|
||||
Ambatovy, 14.3 km 57° Moramanga, 18.8508 S, 48.32 E, 1075 m, montane rainforest, 18.XII.2004 (B.L. Fisher); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.8496 S, 48.2947 E, 1010 m, montane rainforest, 3.-6.III.2007 (B.L. Fisher et al.); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.8394 S, 48.3084 E, 1080 m, montane rainforest, 4.-8.III.2007 (B.L. Fisher et al.); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.8477 S, 48.2957 E, 1000 m, montane rainforest, 5.-8.III.2007 (B.L. Fisher et al.); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.8581 S, 48.2849 E, 1040 m, montane rainforest, 5.-8.III.2007 (B.L. Fisher et al.); Toamasina, Amparihibe, 15° 2' S, 49° 34' E, II.-III.2003 (K.A. Jackson & D. Carpenter); Toamasina, Analamay, 18.8062 S, 48.3371 E, 1068 m, montane rainforest, 21.III.2004 (Malagasy ant team); Toamasina, vic. Andasibe (= Perinet), 950-980 m, 2.-6.II.1977 (W.L. & D.E. Brown); Toamasina, F.C. Andriantantely, 18.695 S, 48.8133 E, 530 m, rainforest, 4.-10.XII.1998 (H.J. Ratsirarson); Toamasina, Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo, 15.1883 S, 49.615 E, 470 m, rainforest, 8.-12.III.2003 (B.L. Fisher, C. Griswold et al.); Toamasina, Rte d'Anosibe, km 33, 4.-12.IV.1975 (A. Peyrieras); Toamasina, Betampona, Ambodiriana, 14.V.1993 (P. Rabeson); Toamasina, Reserve Betampona, Camp Vohitsivalana, 37.1 km 338° Toamasina, 17.8867 S, 49.2025 E, 520 m, rainforest, 1.-3.XII.2005 (B.L. Fisher et al.); Toamasina, F.C. Didy, 18.1983 S, 48.5783 E, 960 m, rainforest, 16.-23.XII.1998 (H.J. Ratsirarson); Toamasina, Mahavelona (Foulpointe), 17.6667 S, 49.5 E, sandy forest & Pandanus marsh, 11.XI.-2.XII.1993 (A. Pauly); Toamasina, Parc National Mananara-Nord, 7.1 km 261° Antanambe, 16.455 S, 49.7875 E, 225 m, rainforest, 14.-16.XI.2005 (B.L. Fisher et al.); Toamasina, P.N. Mantadia, 18.7917 S, 48.4267 E, 895 m, rainforest, 25.XI.-1.XII.1998 (H.J. Ratsirarson); Toamasina, 19 km ESE Maroantsetra, 350 m, rainforest, 22.IV.1989 (P.S. Ward); Toamasina, F.C. Sandranantitra, 18.0483 S, 49.0917 E, 450 m, 18.-24.I.1999 (H.J. Ratsirarson); Toamasina, Torotorofotsy, 18.8708 S, 48.3474 E, 1070 m, montane rainforest, marsh edge, 24.-29.III.2004 (Malagasy ant team); Toamasina, Parc National de Zahamena, Besaky River, 17.7524 S, 48.8532 E, 760 m, rainforest, 22.II.2009 (B.L. Fisher et al.); Toamasina, Parc National de Zahamena, Onibe River, 17.7591 S, 48.8547 E, 780 m, rainforest, 21.-23.II.2009 (B.L. Fisher et al.); Toamasina, Parc National de Zahamena, Sahavorondrano River, 17.7526 S, 48.8573 E, 765 m, rainforest, 23.II.2009 (B.L. Fisher et al.); Toamasina, Parc National de Zahamena, Tetezambatana Forest, near junction of Nosivola and Manakambahiny Rivers, 17.743 S, 48.7294 E, 860 m, rainforest, 18.-19.II.2009 (B.L. Fisher et al.); Toliara,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="79C26D8B2E0E632CC3050BE7D9DDEEE6" originalValue="Rés">Res</normalizedToken>
|
||||
. Andohahela, 10 km NW Enakara, 24.5667 S, 46.8167 E, 430 m, rainforest, 22.XI.1992 (B.L. Fisher); Toliara,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="C642D1A1B5534E4BD9E73EBEE82F1A11" originalValue="Rés">Res</normalizedToken>
|
||||
. Andohahela, 11 km NW Enakara, 24.5667 S, 46.8333 E, 800 m, rainforest, 17.XI.1992 (B.L. Fisher); Toliara, Res. Andohahela, 6 km SSW Eminiminy, 24° 44' S, 46° 48' E, 330 m, wet forest, 4.II.1993 (G.D. Alpert et al.); Toliara, Res. Andohahela, 6 km SSW Eminiminy, 24° 44' S, 46° 48' E, 330 m, rainforest, 4.II.1993 (P.S. Ward); Toliara, Parc National Andohahela, Col de Tanatana, 33.3 km NW Tolagnaro, 24.7585 S, 46.8537 E, 275 m, rainforest,22.-24.XI.2006 (B.L. Fisher et al.); Toliara,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="F3DD9DE158084B1CC952526C538EDFD4" originalValue="Forêt">Foret</normalizedToken>
|
||||
Ivohibe 55.0 km N Tolagnaro, 24.569 S, 47.204 E, 200 m, rainforest, 2.-4.XII.2006 (B.L. Fisher et al.); Toliara,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="44CE71B81DBF44C3ADC8CB584863F504" originalValue="Forêt">Foret</normalizedToken>
|
||||
Ivohibe 55.6 km N Tolagnaro, 24.5617 S, 47.2002 E, 650 m, rainforest, 4.XII.2006 (B.L. Fisher et al.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,138 +1,137 @@
|
|||
<document ENCODING="UTF8" ID-DOI="http://doi.org/10.5281/zenodo.6172603" ID-GBIF-Dataset="9998aa6b-9348-45f8-82a1-55a07ac10bad" ID-GBIF-Taxon="100105326" ID-HNS-Pub="26064" ID-ISSN="175-5326" ID-Zenodo-Dep="6172603" ModsDocAuthor="Hita Garcia, F. & B. L. Fisher" ModsDocDate="2012" ModsDocID="26064" ModsDocOrigin="http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf" ModsDocTitle="The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups." checkinTime="1373743554799" checkinUser="donat" docAuthor="Hita Garcia, F. & B. L. Fisher" docDate="2012" docId="0FFEC2B9C7FA68A4F47159BE2EFD3FD9" docLanguage="en" docName="26064.xml" docOrigin="Zootaxa 3592" docSource="http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf" docTitle="Tetramorium ankarana Garcia & Fisher, 2012, sp. n." docType="treatment" docVersion="11" lastPageNumber="11" masterDocId="FFBCFFC2FFB3FFACFE12FFF966101E6A" masterDocTitle="The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups." masterLastPageNumber="85" masterPageNumber="1" pageNumber="9" updateTime="1645306871304" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo>
|
||||
<mods:title>The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups.</mods:title>
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<mods:title id="2F586C9E77D2497C4A4865CDF42A3B7D">The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups.</mods:title>
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<mods:namePart>Hita Garcia, F.</mods:namePart>
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<mods:namePart>B. L. Fisher</mods:namePart>
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<mods:namePart id="4D4930FB69153FC38D194DE2ABD5D559">B. L. Fisher</mods:namePart>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172603" ID-GBIF-Taxon="100105326" ID-Zenodo-Dep="6172603" LSID="urn:lsid:plazi:treatment:0FFEC2B9C7FA68A4F47159BE2EFD3FD9" httpUri="http://treatment.plazi.org/id/0FFEC2B9C7FA68A4F47159BE2EFD3FD9" lastPageNumber="11" pageNumber="9">
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<subSubSection type="nomenclature">
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<paragraph box="[118,610,118,138]" pageId="8" pageNumber="9" type="mainText">
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<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">
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<pageBreakToken pageId="8" pageNumber="9" start="start">Tetramorium</pageBreakToken>
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<treatment id="0FFEC2B9C7FA68A4F47159BE2EFD3FD9" ID-DOI="http://doi.org/10.5281/zenodo.6172603" ID-GBIF-Taxon="100105326" ID-Zenodo-Dep="6172603" LSID="urn:lsid:plazi:treatment:0FFEC2B9C7FA68A4F47159BE2EFD3FD9" httpUri="http://treatment.plazi.org/id/0FFEC2B9C7FA68A4F47159BE2EFD3FD9" lastPageNumber="11" pageNumber="9" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="F0F47444977D9AF43CDB1F64C73E2DD5" type="nomenclature">
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<paragraph id="06C31246C4399D3B93FEA29CFCC553A3" box="[118,610,118,138]" pageId="8" pageNumber="9" type="mainText">
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<taxonomicName id="AD5FF7A9BEADE33B4491E75F29855679" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">
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<pageBreakToken id="E3F63AE1706032485FAEE1AF45E73C03" pageId="8" pageNumber="9" start="start">Tetramorium</pageBreakToken>
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ankarana
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<authority>Hita</authority>
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<authority id="C781D16F6EB04BDA2871490DEF305F67">Hita</authority>
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</taxonomicName>
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Garcia & Fisher
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<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
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<taxonomicNameLabel id="0C772CA17B5131C2CAAA55920A56CE0F" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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<subSubSection type="description">
|
||||
<paragraph box="[118,322,146,165]" pageId="8" pageNumber="9" type="mainText">(Figs. 1, 2, 5, 6, 7, 141)</paragraph>
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<subSubSection id="206F407D93BF661AE6E9638A4872F075" type="description">
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<paragraph id="438D94EEB5E02EF1D599EEE6192654A3" box="[118,322,146,165]" pageId="8" pageNumber="9" type="mainText">(Figs. 1, 2, 5, 6, 7, 141)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1122,202,361]" pageId="8" pageNumber="9" type="mainText">
|
||||
<subSubSection id="27D1D120EAEA45E90ACBE87C00344F3A" type="materials_examined">
|
||||
<paragraph id="94D53F60394E4208709AE1399DE5BC4F" box="[118,1122,202,361]" pageId="8" pageNumber="9" type="mainText">
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Holotype worker, MADAGASCAR, Antsiranana,
|
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="724F214E15E6531D0547BF09A078EF89" originalValue="Réserve">Reserve</normalizedToken>
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Ankarana, 7 km SE Matsaborimanga, 12° 54' S, 49° 07' E, 150 m, rainforest, ground foragers, collection code PSW11043, 30.XI.1990 (P.S. Ward) (CASC: CASENT0247543). Paratypes, one worker with same data as holotype (BMNH: CASENT0247311); and two workers from Antsiranana,
|
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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<normalizedToken id="FD13FDC148C74DBA6B3FF499DE9EC902" originalValue="Réserve">Reserve</normalizedToken>
|
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<normalizedToken id="A762D2C83693AD19B04667469A1F5C76" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
de l'Ankarana, 22.9 km 224° SW Anivorano Nord, 12.90889 S, 49.10983 E, 80 m, tropical dry forest, on low vegetation, collection code BLF03007, 10.-16.II.2001 (B.L. Fisher, C. Griswold et al.) (CASC: CASENT0404423; CASENT0427943).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,398,417]" pageId="8" pageNumber="9" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,427,474]" pageId="8" pageNumber="9" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">Tetramorium ankarana</taxonomicName>
|
||||
<subSubSection id="45C277BDD34A2299764888DD52D28572" type="diagnosis">
|
||||
<paragraph id="F731263A2A6F0CDDBFBD87E9869D5985" box="[118,208,398,417]" pageId="8" pageNumber="9" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="ECC72999DEBD8967FD9ACCE7D10343E1" box="[118,1121,427,474]" pageId="8" pageNumber="9" type="mainText">
|
||||
<taxonomicName id="DA870D989D664D1A13C9DE815E0AF135" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">Tetramorium ankarana</taxonomicName>
|
||||
can be easily distinguished from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
<taxonomicName id="BFCC3A1F055473A09672D78D85467238" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
by the following character combination: much smaller eyes (OI 20); relatively longer antennal scapes (SI 101-104); and long propodeal spines (PSLI 35-38).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,511,529]" pageId="8" pageNumber="9" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,539,671]" pageId="8" pageNumber="9" type="mainText">HL 0.97-1.02 (0.99); HW 0.85-0.89 (0.86); SL 0.88-0.90 (0.88); EL 0.17-0.18 (0.17); PH 0.47-0.50 (0.49); PW 0.66-0.72 (0.69); WL 1.21-1.31 (1.26); PSL 0.34-0.37 (0.36); PTL 0.33-0.36 (0.35); PTH 0.34-0.36 (0.35); PTW 0.26-0.29 (0.28); PPL 0.28-0.31 (0.29); PPH 0.34-0.37 (0.35); PPW 0.33-0.37 (0.35); CI 87-88 (87); SI 101-104 (102); OI 20 (20); DMI 54-56 (55); LMI 37-40 (39); PSLI 35-38 (36); PeNI 39-42 (41); LPeI 97-100 (99); DPeI 79-84 (81); PpNI 50-52 (51); LPpI 81-85 (83); DPpI 116-125 (120); PPI 122-128 (125) (four measured).</paragraph>
|
||||
<paragraph box="[118,1122,680,1233]" pageId="8" pageNumber="9" type="mainText">Head much longer than wide (CI 87-88); posterior head margin very weakly concave. Anterior clypeal margin medially impressed, often weakly so. Frontal carinae strongly developed, approaching corners of posterior head margin. Antennal scrobes well-developed, moderately deep, narrow, and without defined posterior margin; ventral margin moderately defined. Antennal scapes long, reaching posterior head margin (SI 101-104). Eyes small (OI 20). Mesosomal outline in profile flat to weakly convex, very weakly marginate from lateral to dorsal mesosoma, sides rounding smoothly onto the dorsum; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-40). Propodeal spines long, spinose and acute (PSLI 35-38); propodeal lobes short, triangular, and rounded. Petiolar node in profile clublike with fairly rounded margins, approximately as long as high (LPeI 97-100), anterodorsal margin situated lower than posterodorsal margin, dorsum noticeably convex; node in dorsal view approximately 1.2 to 1.3 times longer than wide (DPeI 79-84). Postpetiole in profile globular, approximately 1.2 times higher than long (LPpI 81-85); in dorsal view around 1.1 to 1.3 times wider than long (DPpI 116-125). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view 1.2 to 1.3 times wider than petiolar node (PPI 122-128). Mandibles strongly striate; clypeus longitudinally rugose/rugulose, with three to four rugae/rugulae, median ruga always present and distinct, remaining rugae weaker; cephalic dorsum between frontal carinae reticulate-rugose to longitudinally rugose, posteriorly more reticulate-rugose and anteriorly more longitudinally rugose; lateral and ventral head mostly reticulate-rugose. Mesosoma dorsally mainly reticulate-rugose, laterally reticulate-rugose to longitudinally rugose. Forecoxae usually with weak to moderately developed longitudinal rugae/rugulae. Waist segments mainly longitudinally rugulose, less reticulate-rugulose. First gastral tergite unsculptured, smooth, and shining. Whole body covered with numerous, very long, fine standing hairs. Body of uniform dark reddish brown colour.</paragraph>
|
||||
<subSubSection id="46CBC3F7A7639801E7C8928A4C74A897" type="description">
|
||||
<paragraph id="556FECF96E01775E62A48F108E1549D4" box="[118,226,511,529]" pageId="8" pageNumber="9" type="mainText">Description</paragraph>
|
||||
<paragraph id="2F860AB6A10F2FAE109C76E0B39D38B8" box="[118,1121,539,671]" pageId="8" pageNumber="9" type="mainText">HL 0.97-1.02 (0.99); HW 0.85-0.89 (0.86); SL 0.88-0.90 (0.88); EL 0.17-0.18 (0.17); PH 0.47-0.50 (0.49); PW 0.66-0.72 (0.69); WL 1.21-1.31 (1.26); PSL 0.34-0.37 (0.36); PTL 0.33-0.36 (0.35); PTH 0.34-0.36 (0.35); PTW 0.26-0.29 (0.28); PPL 0.28-0.31 (0.29); PPH 0.34-0.37 (0.35); PPW 0.33-0.37 (0.35); CI 87-88 (87); SI 101-104 (102); OI 20 (20); DMI 54-56 (55); LMI 37-40 (39); PSLI 35-38 (36); PeNI 39-42 (41); LPeI 97-100 (99); DPeI 79-84 (81); PpNI 50-52 (51); LPpI 81-85 (83); DPpI 116-125 (120); PPI 122-128 (125) (four measured).</paragraph>
|
||||
<paragraph id="B64E5A7FFB740ADB4CDB1F84235DD7A0" box="[118,1122,680,1233]" pageId="8" pageNumber="9" type="mainText">Head much longer than wide (CI 87-88); posterior head margin very weakly concave. Anterior clypeal margin medially impressed, often weakly so. Frontal carinae strongly developed, approaching corners of posterior head margin. Antennal scrobes well-developed, moderately deep, narrow, and without defined posterior margin; ventral margin moderately defined. Antennal scapes long, reaching posterior head margin (SI 101-104). Eyes small (OI 20). Mesosomal outline in profile flat to weakly convex, very weakly marginate from lateral to dorsal mesosoma, sides rounding smoothly onto the dorsum; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-40). Propodeal spines long, spinose and acute (PSLI 35-38); propodeal lobes short, triangular, and rounded. Petiolar node in profile clublike with fairly rounded margins, approximately as long as high (LPeI 97-100), anterodorsal margin situated lower than posterodorsal margin, dorsum noticeably convex; node in dorsal view approximately 1.2 to 1.3 times longer than wide (DPeI 79-84). Postpetiole in profile globular, approximately 1.2 times higher than long (LPpI 81-85); in dorsal view around 1.1 to 1.3 times wider than long (DPpI 116-125). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view 1.2 to 1.3 times wider than petiolar node (PPI 122-128). Mandibles strongly striate; clypeus longitudinally rugose/rugulose, with three to four rugae/rugulae, median ruga always present and distinct, remaining rugae weaker; cephalic dorsum between frontal carinae reticulate-rugose to longitudinally rugose, posteriorly more reticulate-rugose and anteriorly more longitudinally rugose; lateral and ventral head mostly reticulate-rugose. Mesosoma dorsally mainly reticulate-rugose, laterally reticulate-rugose to longitudinally rugose. Forecoxae usually with weak to moderately developed longitudinal rugae/rugulae. Waist segments mainly longitudinally rugulose, less reticulate-rugulose. First gastral tergite unsculptured, smooth, and shining. Whole body covered with numerous, very long, fine standing hairs. Body of uniform dark reddish brown colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,1270,1289]" pageId="8" pageNumber="9" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1120,1299,1346]" pageId="8" pageNumber="9" type="mainText">
|
||||
<subSubSection id="90B2B16013896925C5968084E8560EF5" type="discussion">
|
||||
<paragraph id="F85A6A8DAC3FB4D8AB14CAAC00AC3511" box="[118,170,1270,1289]" pageId="8" pageNumber="9" type="mainText">Notes</paragraph>
|
||||
<paragraph id="B74C473F8A5370547751A840F7D186A4" box="[118,1120,1299,1346]" pageId="8" pageNumber="9" type="mainText">
|
||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="66EB70D11EBD30EEAB3D8200199941A6" rank="species">new species</taxonomicNameLabel>
|
||||
is currently only known from the type locality Ankarana where it was collected in tropical dry forest and rainforest at elevations of 80 to 150 m.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,1355,1543]" pageId="8" pageNumber="9" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">Tetramorium ankarana</taxonomicName>
|
||||
<paragraph id="1CA61761D158656F6C2134E6DF50F56B" box="[118,1122,1355,1543]" pageId="8" pageNumber="9" type="mainText">
|
||||
<taxonomicName id="A42611E0A7069962FC297379C4125A9C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">Tetramorium ankarana</taxonomicName>
|
||||
is the second species known from the species group, and it is easily distinguishable from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri.</taxonomicName>
|
||||
<taxonomicName id="AE12964D68D004832C3A673CE90491FA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri.</taxonomicName>
|
||||
The latter species has much larger eyes (OI 24-26), relatively shorter antennal scapes (SI 89-99), and extremely long propodeal spines (PSLI 49-68), whereas T.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
<taxonomicName id="3CC2D98317100191310C950BE4672513" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
possesses significantly smaller eyes (OI 20), slightly longer antennal scapes (SI 101-104), and much shorter propodeal spines (PSLI 35-38). Furthermore, T.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
<taxonomicName id="84B5564584A45117BF3F4AF7F8E29C77" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
is also much darker in colour than most of the material of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri,</taxonomicName>
|
||||
<taxonomicName id="C344D384F42DBB2650CD9B0CBB8D4C40" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri,</taxonomicName>
|
||||
which ranges from whitish-yellowish to brown. However, some series were almost the same colour as T.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
<taxonomicName id="698AE3CD679A4F127EC3F244BA98459F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
, and we do not consider colouration a good diagnostic character in this case.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="multiple">
|
||||
<caption box="[118,1122,1539,1584]" pageId="9" pageNumber="10" type="caption">
|
||||
<paragraph box="[118,1122,1539,1584]" pageId="9" pageNumber="10" type="caption">
|
||||
<pageBreakToken pageId="9" pageNumber="10" start="start">FIGURES</pageBreakToken>
|
||||
<subSubSection id="51DACB931B57A6CA44044C3E6BE1A170" type="multiple">
|
||||
<caption id="B245AB944AE5C7BC7FE3EF2F5D591AFA" box="[118,1122,1539,1584]" pageId="9" pageNumber="10" type="caption">
|
||||
<paragraph id="E8C5C2D051F57284C754CDEED5577E6D" box="[118,1122,1539,1584]" pageId="9" pageNumber="10" type="caption">
|
||||
<pageBreakToken id="6A95281A7E8747DFAA1603D45B6EE4BE" pageId="9" pageNumber="10" start="start">FIGURES</pageBreakToken>
|
||||
5-7. T.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
<taxonomicName id="7BA7B2BF9E50312C6846185D3CAE12D5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
, holotype (CASENT0247543). 5. Body in profile. 6. Body in dorsal view. 7. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,118,136]" pageId="10" pageNumber="11" type="mainText">
|
||||
<pageBreakToken pageId="10" pageNumber="11" start="start">Etymology</pageBreakToken>
|
||||
<subSubSection id="6F8293833BFE1B298ABF0E4D36182723" type="etymology">
|
||||
<paragraph id="C86F75C35ADB113853405952969B9913" box="[118,219,118,136]" pageId="10" pageNumber="11" type="mainText">
|
||||
<pageBreakToken id="CFD902FEFA84E11C0C75924FA37E7446" pageId="10" pageNumber="11" start="start">Etymology</pageBreakToken>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1120,147,222]" pageId="10" pageNumber="11" type="mainText">
|
||||
<paragraph id="F9B86FC8E338E3C7E97D220396E22CAB" box="[118,1120,147,222]" pageId="10" pageNumber="11" type="mainText">
|
||||
The name of the
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="BA9C2CB66BEEA4A215270DFAC2E93B30" rank="species">new species</taxonomicNameLabel>
|
||||
refers to the type locality, the
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="8D1354D483FB31DB14A3C59D97CDD2A6" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="5835AA953A615E06BF07B23239087F95" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
de l'Ankarana. The reserve is of high importance for the conservation of animal biodiversity, and the dedication of this
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="223F82EE91AF62B8D2934E40C0E9A623" rank="species">new species</taxonomicNameLabel>
|
||||
accounts for its importance. The species epithet is a noun in apposition, and thus invariant.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,259,277]" pageId="10" pageNumber="11" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,287,362]" pageId="10" pageNumber="11" type="mainText">
|
||||
<subSubSection id="EB0BBA579177863EB506D044EC71B699" type="materials_examined">
|
||||
<paragraph id="BE6AF44FF9CFE03891917ACC4686DF51" box="[118,295,259,277]" pageId="10" pageNumber="11" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="6762C93436A7C0B037D816EF84D83665" box="[118,1122,287,362]" pageId="10" pageNumber="11" type="mainText">
|
||||
MADAGASCAR: Antsiranana,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="618D5B3647EF59B3BF1D7502992ABDA7" originalValue="Réserve">Reserve</normalizedToken>
|
||||
Ankarana, 7 km SE Matsaborimanga, 12° 54' S, 49° 07' E, 150 m, rainforest, 30.XI.1990 (P.S. Ward); Antsiranana,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="A282748F6CE4E2A0B26822C7A0D2443A" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="58B25CD8F5AE352C853673EF4450DE14" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
de l'Ankarana, 22.9 km 224° SW Anivorano Nord, 12.90889 S, 49.10983 E, 80 m, tropical dry forest, 10.-16.II.2001 (B.L. Fisher, C. Griswold et al.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,63 +1,62 @@
|
|||
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<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="adamsi">
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|
||||
Tetramorium adamsi
|
||||
<authority>Hita</authority>
|
||||
<authority id="45C02D747674CA6D3589736F74961441">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="85A73E5F3B1D75132B2EAD68C8F210D8" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,410,707,726]" pageId="65" pageNumber="66" type="mainText">(Figs. 14, 17, 123, 124, 125, 142)</paragraph>
|
||||
<subSubSection id="8F55F1C81360281405128DBFF93FBCA4" type="description">
|
||||
<paragraph id="0D5BAA25D86E3C70C5A43335A26DDE2C" box="[118,410,707,726]" pageId="65" pageNumber="66" type="mainText">(Figs. 14, 17, 123, 124, 125, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1123,763,951]" pageId="65" pageNumber="66" type="mainText">Holotype worker, MADAGASCAR, Antsiranana, R.S. Manongarivo 17.3 km 218° SW Antanambao, 14.02167 S, 48.41833 E, 1580 m, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF01970, 27.X.1998 (B.L. Fisher) (CASC: CASENT0247345). Paratypes, 21 workers with same data as holotype (BMNH: CASENT0247299; CASC: CASENT0189393; CASENT0218051; CASENT0247290; CASENT0247291; CASENT0247292; CASENT0247293; CASENT0247294; CASENT0247300; CASENT0247304; CASENT0247305; CASENT0247306; CASENT0247307; CASENT0247308; CASENT0247341; CASENT0247342; CASENT0247343; CASENT0247344; MCZ: CASENT0247301; MHNG: CASENT0247297; NHMB: CASENT0247303).</paragraph>
|
||||
<subSubSection id="A7DB66944DE05680D7E871E0D82F76B0" type="materials_examined">
|
||||
<paragraph id="EA5E2B221E14421EDC118CEAE5E71E1B" box="[118,1123,763,951]" pageId="65" pageNumber="66" type="mainText">Holotype worker, MADAGASCAR, Antsiranana, R.S. Manongarivo 17.3 km 218° SW Antanambao, 14.02167 S, 48.41833 E, 1580 m, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF01970, 27.X.1998 (B.L. Fisher) (CASC: CASENT0247345). Paratypes, 21 workers with same data as holotype (BMNH: CASENT0247299; CASC: CASENT0189393; CASENT0218051; CASENT0247290; CASENT0247291; CASENT0247292; CASENT0247293; CASENT0247294; CASENT0247300; CASENT0247304; CASENT0247305; CASENT0247306; CASENT0247307; CASENT0247308; CASENT0247341; CASENT0247342; CASENT0247343; CASENT0247344; MCZ: CASENT0247301; MHNG: CASENT0247297; NHMB: CASENT0247303).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,988,1007]" pageId="65" pageNumber="66" type="mainText">Diagnosis</paragraph>
|
||||
<subSubSection id="DCACAFA7F4C9FA30D8BD79A5B2EFF6A6" type="diagnosis">
|
||||
<paragraph id="B07B01A58C2206E0883FBDC4C9EB5B61" box="[118,208,988,1007]" pageId="65" pageNumber="66" type="mainText">Diagnosis</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,91 +1,90 @@
|
|||
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">Tetramorium kelleri</taxonomicName>
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<taxonomicName id="88EF62F91FB0C3FC9A7CA5A521E29AF2" ID-CoL="55PP2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">Tetramorium kelleri</taxonomicName>
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species group
|
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<paragraph box="[118,208,679,697]" pageId="7" pageNumber="8" type="mainText">Diagnosis</paragraph>
|
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<paragraph box="[118,1127,707,923]" pageId="7" pageNumber="8" type="mainText">Eleven-segmented antennae; anterior clypeal margin medially impressed; frontal carinae well-developed and usually running to posterior head margin; anterior face of mesosoma weakly developed; margination between lateral and dorsal mesosoma very weakly developed, sides round smoothly onto dorsum; propodeal spines always long to extremely long, and spinose; propodeal lobes short, triangular, and blunted or acute; petiolar node clublike, anterodorsal margin situated lower and much less pronounced than posterodorsal margin, in profile approximately as long as high, in dorsal view distinctly longer than wide; postpetiole globular to subglobular; mandibles strongly sculptured; sculpture on head, mesosoma, and waist segments distinct and predominantly reticulate-rugose; gaster unsculptured, smooth, and shiny; whole body covered with numerous, very long, fine, standing hairs; sting appendage spatulate.</paragraph>
|
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<subSubSection id="522A31FB2FD3556532B0B4E64340A074" type="diagnosis">
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<paragraph id="1971E9C6F3E8749FA502FE5CFA52C190" box="[118,208,679,697]" pageId="7" pageNumber="8" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="A1CC39EF5615473A2031B2BEEC11C205" box="[118,1127,707,923]" pageId="7" pageNumber="8" type="mainText">Eleven-segmented antennae; anterior clypeal margin medially impressed; frontal carinae well-developed and usually running to posterior head margin; anterior face of mesosoma weakly developed; margination between lateral and dorsal mesosoma very weakly developed, sides round smoothly onto dorsum; propodeal spines always long to extremely long, and spinose; propodeal lobes short, triangular, and blunted or acute; petiolar node clublike, anterodorsal margin situated lower and much less pronounced than posterodorsal margin, in profile approximately as long as high, in dorsal view distinctly longer than wide; postpetiole globular to subglobular; mandibles strongly sculptured; sculpture on head, mesosoma, and waist segments distinct and predominantly reticulate-rugose; gaster unsculptured, smooth, and shiny; whole body covered with numerous, very long, fine, standing hairs; sting appendage spatulate.</paragraph>
|
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|
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<subSubSection type="discussion">
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<paragraph box="[118,219,960,978]" pageId="7" pageNumber="8" type="mainText">Comments</paragraph>
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<paragraph box="[118,1121,988,1064]" pageId="7" pageNumber="8" type="mainText">
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<paragraph id="FD7558FD4B6C73AC243F067C428CDD2B" box="[118,219,960,978]" pageId="7" pageNumber="8" type="mainText">Comments</paragraph>
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<paragraph id="E8298947A3DBE421C8F42BD7180E692B" box="[118,1121,988,1064]" pageId="7" pageNumber="8" type="mainText">
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The recently (Hita Garcia & Fisher, 2012) proposed
|
||||
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<taxonomicName id="BD21FDDD1194A1D5133FC8C4753C4704" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
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||||
species group contains
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
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<taxonomicName id="7634AA34019810462491795FFEC7AD7D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
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and the
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
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<taxonomicNameLabel id="2768C1928D6D25C10328D5646E388B0E" rank="species">new species</taxonomicNameLabel>
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T.
|
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
<taxonomicName id="9864FA40611133F8C1A09F3F204CAA3D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
. It is mainly distributed in the north and west of Madagascar, as well as on Nosy Be and Mayotte, and both species prefer forested habitats.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,1073,1148]" pageId="7" pageNumber="8" type="mainText">
|
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<paragraph id="E8F5CA1901108B18F0D876E138B4F4B3" box="[118,1122,1073,1148]" pageId="7" pageNumber="8" type="mainText">
|
||||
Other
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
<taxonomicName id="590A9981241A50ECC14CE6E045127413" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
species from a different species group are unlikely to be confused with T.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
<taxonomicName id="8CEB4AD99FDD5E41679CD5F03E10FBA4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
or
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri.</taxonomicName>
|
||||
<taxonomicName id="05A671E5F4505DE9C5D0B239C85C40D9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri.</taxonomicName>
|
||||
The possession of 11-segmented antennae and the very conspicuous petiolar node shape clearly distinguish them from all other Malagasy species groups.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="key">
|
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<paragraph box="[118,643,1213,1233]" pageId="7" pageNumber="8" type="mainText">
|
||||
<subSubSection id="B3FA4E84F39FBDDF1D2A6F4B4315D6BF" type="key">
|
||||
<paragraph id="09869BF3B54C35411205344ED9BF3B54" box="[118,643,1213,1233]" pageId="7" pageNumber="8" type="mainText">
|
||||
Key to species of the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
<taxonomicName id="6A37334F10A5A063BB1CB6B34981C9C6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
species group (workers)
|
||||
</paragraph>
|
||||
<paragraph box="[118,1123,1268,1308]" pageId="7" pageNumber="8" type="mainText">
|
||||
<paragraph id="9A099432790EC850057D176E8597B5AE" box="[118,1123,1268,1308]" pageId="7" pageNumber="8" type="mainText">
|
||||
1. Species with smaller eyes (OI 20); relatively longer antennal scapes (SI 101-104); and long propodeal spines (PSLI 35-38) (Figs. 1, 2).................................................................................. T.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
<taxonomicName id="7D2714B4964A9597209211F89B81C089" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1123,1314,1354]" pageId="7" pageNumber="8" type="mainText">
|
||||
<paragraph id="10B1CB1859DF633F6CF8ABAD1CD9075B" box="[175,1123,1314,1354]" pageId="7" pageNumber="8" type="mainText">
|
||||
Species with larger eyes (OI 24-26); relatively shorter antennal scapes (SI 89-99); and extremely long propodeal spines (PSLI 49-68) (Figs. 3, 4)...............................................................................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
<taxonomicName id="2CDB08C93EA91B9993E2BF1CDC6283FF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
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</treatment>
|
||||
|
|
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|||
|
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@ -1,275 +1,274 @@
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<mods:title id="B4AB5CC28700C1ACA491A91FE8BAA7AA">The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups.</mods:title>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172615" ID-GBIF-Taxon="100105324" ID-Zenodo-Dep="6172615" LSID="urn:lsid:plazi:treatment:285A85A6897E3ECC849B5BDE4B9770E4" httpUri="http://treatment.plazi.org/id/285A85A6897E3ECC849B5BDE4B9770E4" lastPageNumber="29" pageNumber="26">
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<paragraph box="[118,434,118,138]" pageId="25" pageNumber="26" type="heading">
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<taxonomicName authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="andrei">
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<pageBreakToken pageId="25" pageNumber="26" start="start">Tetramorium</pageBreakToken>
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<treatment id="285A85A6897E3ECC849B5BDE4B9770E4" ID-DOI="http://doi.org/10.5281/zenodo.6172615" ID-GBIF-Taxon="100105324" ID-Zenodo-Dep="6172615" LSID="urn:lsid:plazi:treatment:285A85A6897E3ECC849B5BDE4B9770E4" httpUri="http://treatment.plazi.org/id/285A85A6897E3ECC849B5BDE4B9770E4" lastPageNumber="29" pageNumber="26" scope_family="Formicidae" scope_order="Hymenoptera">
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<taxonomicName id="CD5D1DFCF056C539D9A132187E4A24D7" ID-CoL="55PHN" authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="andrei">
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<pageBreakToken id="A21B300FDC3DF2426759855170364113" pageId="25" pageNumber="26" start="start">Tetramorium</pageBreakToken>
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andrei
|
||||
<authority>Forel</authority>
|
||||
<authority id="559B454A933DFEEBDEF1C759E89B66C2">Forel</authority>
|
||||
,
|
||||
</taxonomicName>
|
||||
1892a
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,509,146,165]" pageId="25" pageNumber="26" type="heading">(Figs. 13, 54, 67, 70, 73, 74, 81, 82, 83, 141)</paragraph>
|
||||
<subSubSection id="4DA8415FF6DF15A387F0D4991112567F" type="description">
|
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<paragraph id="E03C92C1906327ECC77193B14B54D1B3" box="[118,509,146,165]" pageId="25" pageNumber="26" type="heading">(Figs. 13, 54, 67, 70, 73, 74, 81, 82, 83, 141)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="reference_group">
|
||||
<paragraph box="[118,1122,202,291]" pageId="25" pageNumber="26" type="mainText">
|
||||
<taxonomicName authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="andrei" subGenus="Xiphomyrmex">
|
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<subSubSection id="8981632BDF038C923C13F038CCE72D08" type="reference_group">
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<paragraph id="D6F16F134B44A88A3E6E57D1A36DA8F6" box="[118,1122,202,291]" pageId="25" pageNumber="26" type="mainText">
|
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<taxonomicName id="6ABF76104514DBCEEEEB6C3BC7864B91" authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="andrei" subGenus="Xiphomyrmex">
|
||||
Tetramorium (Xiphomyrmex) andrei
|
||||
<authority>Forel</authority>
|
||||
<authority id="482F2A710FC9A55C3CECB9583ABF6FA2">Forel</authority>
|
||||
,
|
||||
</taxonomicName>
|
||||
1892a:263. Lectotype worker [designated here], MADAGASCAR, Toamasina, province de Bezanozano,
|
||||
<normalizedToken originalValue="Nosibé">Nosibe</normalizedToken>
|
||||
<normalizedToken id="D1DF4E8537B198E4EBB4F3AAAA3A93D3" originalValue="Nosibé">Nosibe</normalizedToken>
|
||||
Village de
|
||||
<normalizedToken originalValue="I’Imerina">I'Imerina</normalizedToken>
|
||||
<normalizedToken id="1BB6B578F55B8BEF392DAEA25F50739E" originalValue="I’Imerina">I'Imerina</normalizedToken>
|
||||
(M. Sikora) (MHNG: CASENT0101821) [examined]. Paralectotypes, two workers with same data as holotype (MHNG: CASENT0101281; CASENT0101282) [examined]. [Combination in
|
||||
<taxonomicName authority="Forel, 1887" authorityName="Forel" authorityYear="1887" class="Insecta" family="Formicidae" genus="Xiphomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subGenus" subGenus="Xiphomyrmex">Xiphomyrmex</taxonomicName>
|
||||
<taxonomicName id="601856E37ED9D828FCDBE8C04A501A63" genus="Xiphomyrmex" pageId="25" pageNumber="26" rank="subGenus" subGenus="Xiphomyrmex">Xiphomyrmex</taxonomicName>
|
||||
by Wheeler, W.M. 1922:1031; in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
<taxonomicName id="06294D22A33823916D6443D28C01FA7A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
by Bolton, 1979:143].
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,297,339]" pageId="25" pageNumber="26" type="mainText">
|
||||
<taxonomicName authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="andrei" subGenus="Xyphomyrmex" subSpecies="robustior">
|
||||
<paragraph id="690C58AA59F84069C03D336178F56F1F" box="[118,1122,297,339]" pageId="25" pageNumber="26" type="mainText">
|
||||
<taxonomicName id="E6B304A90038D029DF13DEA919197B88" authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="andrei" subGenus="Xyphomyrmex" subSpecies="robustior">
|
||||
Tetramorium (Xyphomyrmex) andrei st. robustior
|
||||
<authority>Forel</authority>
|
||||
<authority id="145B0899ABE32A4D69128A36456F360D">Forel</authority>
|
||||
,
|
||||
</taxonomicName>
|
||||
1892b:521. Syntype workers, Foret d'Andrangoloaca (M. Sikora) (MHNG: CASENT0101279, CASENT0101280) [examined]. [Raised to species by Bolton, 1979:147].
|
||||
<taxonomicNameLabel>Syn. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="F2D44435B4F22341CEE6E2EB5D49AA6A">Syn. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[155,1122,345,531]" pageId="25" pageNumber="26" type="mainText">
|
||||
<subSubSection id="638C9BF8B0A4F3E21932443982CCC90A" type="discussion">
|
||||
<paragraph id="A89A2F8C64D147D0961A99582A7837ED" box="[155,1122,345,531]" pageId="25" pageNumber="26" type="mainText">
|
||||
[Note: there is a bit of confusion concerning the type locality of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
<taxonomicName id="D4253AB834D6F3830EBCE304D3BCF27E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
The type locality information on the labels from MHNG is clearly "
|
||||
<normalizedToken originalValue="Nosibé">Nosibe</normalizedToken>
|
||||
<normalizedToken id="D7005EE21F16F7B02A2935695DF153A5" originalValue="Nosibé">Nosibe</normalizedToken>
|
||||
Village del
|
||||
<normalizedToken originalValue="I’Imerina">I'Imerina</normalizedToken>
|
||||
<normalizedToken id="CE5014D11F2E0EB36A80CC0669C39ACE" originalValue="I’Imerina">I'Imerina</normalizedToken>
|
||||
(Sikora)", whereas the description by Forel gives the type locality as "Environs de la ville
|
||||
<normalizedToken originalValue="d'Anosibé">d'Anosibe</normalizedToken>
|
||||
<normalizedToken id="C8DBC47D2643080F2DD089683CFCF547" originalValue="d'Anosibé">d'Anosibe</normalizedToken>
|
||||
(province de Bezanozano), a trois
|
||||
<normalizedToken originalValue="journées">journees</normalizedToken>
|
||||
<normalizedToken id="7BA045A28DF169244D712AF5DE8E7E0A" originalValue="journées">journees</normalizedToken>
|
||||
a l'est-sud-est d'Antananarivo (M. Sikora)." There are several places with the names
|
||||
<normalizedToken originalValue="Nosibé">Nosibe</normalizedToken>
|
||||
<normalizedToken id="67426E89FDE23F45CE3CC7532F5DCB0A" originalValue="Nosibé">Nosibe</normalizedToken>
|
||||
or
|
||||
<normalizedToken originalValue="Anosibé">Anosibe</normalizedToken>
|
||||
<normalizedToken id="ED8B1B171FF6D92F26AD4DD519A820C4" originalValue="Anosibé">Anosibe</normalizedToken>
|
||||
in Madagascar, and additional information is necessary to ascertain which was meant by Forel. Since he mentioned that the locality was three
|
||||
<normalizedToken originalValue="days’">days'</normalizedToken>
|
||||
<normalizedToken id="4F2FC42AD3F74046B3B339F88069AB23" originalValue="days’">days'</normalizedToken>
|
||||
march ESE of Antananarivo in the province of Bezanozano we can hypothesize that it might be close to present-day Anosibe an 'Ala in Toamasina, which is ESE of Antananarivo approximately 30 km south of Moramanga and in the traditional area of the Bezanozano people. Some uncertainty remains, but we think that this locality data is the closest to Forel's (1891) description.]
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,565,584]" pageId="25" pageNumber="26" type="heading">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,593,781]" pageId="25" pageNumber="26" type="mainText">
|
||||
<subSubSection id="4AC8484922E9DD5C66D70A20CE11DEA4" type="diagnosis">
|
||||
<paragraph id="0458050CAD1927A1CC43A0C9FBADF8F0" box="[118,208,565,584]" pageId="25" pageNumber="26" type="heading">Diagnosis</paragraph>
|
||||
<paragraph id="50AC5BCBCCE0BCCE9B1CF3372CFBD272" box="[118,1121,593,781]" pageId="25" pageNumber="26" type="mainText">
|
||||
Within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="FC4E388033359AEAF03C2F67D6DF76B8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
species complex
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="4D36BB359B58D56F9B5747CE74A6C12D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
is clearly identifiable based on the following character combination: eyes short to moderately sized (OI 19-25); propodeal spines long to very long (PSLI 32-45, usually below 40); in profile petiolar node with well-developed anterodorsal and posterodorsal angles, usually sharply defined, often one or both angles weakly rounded; posterodorsal corner of petiole not strongly protruding posteriorly; node in dorsal view ranging from as long as wide to 1.2 times longer than wide (DPeI 82-98); postpetiole in dorsal view as wide as long to approximately 1.4 times wider than long (DPpI 103-136); body colour ranging from uniform reddish-brown to very dark brown.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,818,837]" pageId="25" pageNumber="26" type="heading">Description</paragraph>
|
||||
<paragraph box="[118,1121,847,978]" pageId="25" pageNumber="26" type="mainText">HL 0.77-1.11 (0.95); HW 0.73-1.06 (0.91); SL 0.60-0.86 (0.70); EL 0.15-0.23 (0.20); PH 0.37-0.55 (0.47); PW 0.53-0.75 (0.67); WL 0.96-1.40 (1.20); PSL 0.27-0.46 (0.36); PTL 0.27-0.36 (0.32); PTH 0.29-0.45 (0.37); PTW 0.23-0.33 (0.28); PPL 0.26-0.37 (0.32); PPH 0.29-0.45 (0.37); PPW 0.30-0.44 (0.37); CI 91-100 (96); SI 68-86 (77); OI 19-25 (22); DMI 50-60 (56); LMI 37-43 (39); PSLI 32-45 (38); PeNI 36-48 (42); LPeI 76-97 (86); DPeI 82-98 (90); PpNI 48-63 (55); LPpI 75-98 (88); DPpI 103-136 (116); PPI 117-149 (131) (65 measured).</paragraph>
|
||||
<paragraph box="[118,1122,987,1569][118,1122,118,362]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="mainText">
|
||||
<subSubSection id="DFEBC3853C8E52C9001F142C91D778FD" type="description">
|
||||
<paragraph id="BC2BC5DBA25BB5C2CFC4387CF82B9558" box="[118,226,818,837]" pageId="25" pageNumber="26" type="heading">Description</paragraph>
|
||||
<paragraph id="B72666237D2D6C4217CC91888B3AF0E1" box="[118,1121,847,978]" pageId="25" pageNumber="26" type="mainText">HL 0.77-1.11 (0.95); HW 0.73-1.06 (0.91); SL 0.60-0.86 (0.70); EL 0.15-0.23 (0.20); PH 0.37-0.55 (0.47); PW 0.53-0.75 (0.67); WL 0.96-1.40 (1.20); PSL 0.27-0.46 (0.36); PTL 0.27-0.36 (0.32); PTH 0.29-0.45 (0.37); PTW 0.23-0.33 (0.28); PPL 0.26-0.37 (0.32); PPH 0.29-0.45 (0.37); PPW 0.30-0.44 (0.37); CI 91-100 (96); SI 68-86 (77); OI 19-25 (22); DMI 50-60 (56); LMI 37-43 (39); PSLI 32-45 (38); PeNI 36-48 (42); LPeI 76-97 (86); DPeI 82-98 (90); PpNI 48-63 (55); LPpI 75-98 (88); DPpI 103-136 (116); PPI 117-149 (131) (65 measured).</paragraph>
|
||||
<paragraph id="DE2C33DF61A5B91925FD3240A68358E9" box="[118,1122,987,1569][118,1122,118,362]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="mainText">
|
||||
Head distinctly longer than wide to as long as wide (CI 91-100). Posterior head margin weakly concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short to moderately long, not reaching posterior head margin (SI 68-86). Eyes short to moderately sized (OI 19-25). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-43). Propodeal spines long, rarely very long, spinose, and acute (PSLI 32-45, usually below 40); propodeal lobes variable, short and weakly developed to well-developed and moderately long, triangular to elongate-triangular, and rounded and blunt to acute. Petiolar node in profile rectangular nodiform with usually well-defined angles, node ranging from as high as long to 1.3 times higher than long (LPeI 76-97), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum straight to weakly convex, sometimes anterodorsal or posterodorsal margins slightly higher than other; node in dorsal view ranging from as long as wide to 1.2 times longer than wide (DPeI 82-98). Postpetiole in profile globular to subglobular, approximately as high as long to 1.3 times higher than long (LPpI 75-98); in dorsal view between as wide as long to approximately 1.4 wider than long (DPpI 103-136). Generally, postpetiole in profile approximately as voluminous as petiolar node, in dorsal view postpetiole approximately 1.2 to 1.5 times wider than petiolar node (PPI 117-149). Mandibles usually distinctly longitudinally rugose, in some localities mandibles weakly sculptured to completely unsculptured, smooth, and shining; clypeus usually longitudinally rugose/rugulose, almost always with one distinct median ruga/rugula, often two well-developed rugae/rugulae laterally, sometimes only median ruga/rugula distinct and remaining sculpture irregularly arranged; cephalic dorsum between frontal carinae with 7 to 12 longitudinal rugae running from posterior head
|
||||
<pageBreakToken pageId="26" pageNumber="27" start="start">margin</pageBreakToken>
|
||||
<pageBreakToken id="6FE0F75BAD56DCFA399A2A78E9CF67F8" pageId="26" pageNumber="27" start="start">margin</pageBreakToken>
|
||||
to posterior clypeus, rugae often broken, rarely with cross-meshes; lateral and ventral head mainly reticulate-rugose and to lesser extent longitudinally rugose. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae usually completely unsculptured, smooth, and shiny, sometimes with partial superficial sculpture. Waist segments longitudinally to irregularly rugose. Generally ground sculpture faint to absent everywhere on body, rarely head with weak to moderate reticulate-punctate ground sculpture. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with subdecumbent to erect standing hairs. Body colour ranging from uniform reddish-brown (most commonly encountered) to very dark brown, often appendages of much lighter colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,399,418]" pageId="26" pageNumber="27" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1122,428,559]" pageId="26" pageNumber="27" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">Tetramorium andrei</taxonomicName>
|
||||
<subSubSection id="4A3FDD45DD4DB7D1DBFDBFDA8D7720C4" type="discussion">
|
||||
<paragraph id="1B75DAFCA3A9163ED3618B633C50D408" box="[118,170,399,418]" pageId="26" pageNumber="27" type="mainText">Notes</paragraph>
|
||||
<paragraph id="6E479D0F9288C07392B0B4E0B0DE9CDC" box="[118,1122,428,559]" pageId="26" pageNumber="27" type="mainText">
|
||||
<taxonomicName id="C8C68154228BE166C3C53C910B00CB10" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">Tetramorium andrei</taxonomicName>
|
||||
is a widespread and common species found in most of the rainforests and montane rainforests of eastern and northern Madagascar. The southernmost locality is Andohahela in the southeast, and from there it ranges in an almost continuous band north to Montagne Francais in the northern tip of Madagascar. Fairly isolated from the main distribution,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="8518282ADE8CD520428DF6CF95229443" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
is also encountered in Ambohijanahary and further north in Manongarivo and Ampasindava. The species was sampled at elevations from 10 to 1625 m, and lives in leaf litter.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,568,784]" pageId="26" pageNumber="27" type="mainText">
|
||||
<paragraph id="697AA3FA45F338D0DC13752330939410" box="[118,1122,568,784]" pageId="26" pageNumber="27" type="mainText">
|
||||
It must be pointed out that
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="EED45BA10EA6FFEA7367722A24589AC6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
displays an extraordinary level of intraspecific variation, and we are confident that the material included under this name consists of three to four
|
||||
<normalizedToken originalValue="“good”">"good"</normalizedToken>
|
||||
<normalizedToken id="51F44DBBFC95D0B59DFAC63BBD79F018" originalValue="“good”">"good"</normalizedToken>
|
||||
species that we are unfortunately unable to delimit in this study. The morphometric range outlined above, based on 65 measured specimens, shows a remarkable variability not found in another Afrotropical or Malagasy
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
<taxonomicName id="51FB213F8D63C86995E550886F0FB9F2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
. There are a number of characters that are variably developed, often within the same locality, which do not allow a consistent species delimitation. The most important characters are head shape (CI 91-100), antennal scape length (SI 68-86), the development of the propodeal lobes and the petiolar node, and the sculpture on the mandibles. There seem to be three to four relatively discrete groups that can be recognised by morphometrics and few important characters.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,793,1234]" pageId="26" pageNumber="27" type="mainText">
|
||||
<paragraph id="44EDF6C0AD3FDBAA16A09E2BD2454BFE" box="[118,1122,793,1234]" pageId="26" pageNumber="27" type="mainText">
|
||||
The first group is found in the southeast from Andohahela north to Andringitra, and is mainly characterised by relatively short antennal scapes (SI 68-74), short, rounded propodeal lobes, and a petiolar node without very sharply defined antero- and posterodorsal margins. The second group, which contains the type material of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
<taxonomicName id="4CC4E3A5E4417A9D742E709144E28582" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
has significantly longer antennal scapes (SI 80-86), short but acute propodeal lobes, and a petiolar node with very well-developed and sharply defined antero- and posterodorsal margins. This group is mainly found in central-eastern Madagascar from Vevembe and Vatovavy north to Mandrisy. The third group contains the types of the junior synonym
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="robustior">T. robustior,</taxonomicName>
|
||||
<taxonomicName id="79F36E1ECAE4C457C80F9C9BB203A633" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="robustior">T. robustior,</taxonomicName>
|
||||
and is found from central-eastern to north-eastern Madagascar. It is characterised by comparatively longer antennal scapes (SI 80-85), longer, elongate-triangular, and very acute propodeal lobes, and a petiolar node with the posterodorsal margin weakly higher and more angular than the anterodorsal margin. The head in this group shows the widest shape with a CI approaching 100. The last group seems restricted to northern parts of Madagascar and is encountered north of Manongarivo in the West and Marotandrano in the east. In this group the mandibles are often unsculptured or almost unsculptured, and generally very smooth and shining. Also, the antennal scapes are relatively short (SI 70-77), the propodeal lobes are short, triangular, and often blunted, and the petiolar node has less strongly marginate antero- and posterodorsal margins. Sometimes, especially in Manongarivo, the anterodorsal margin of the petiole is situated slightly higher, although this is not consistent in that series.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,1243,1516]" pageId="26" pageNumber="27" type="mainText">
|
||||
<paragraph id="10A2BE4B73FE37B065EB77FF5E1035B0" box="[118,1121,1243,1516]" pageId="26" pageNumber="27" type="mainText">
|
||||
Despite encountering these four groups within the extensive material of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="AEFE56FDCC1B6178A1DD7C966A4C761F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
examined during this study, we are reluctant to describe these as different species at the moment, even though some or all likely deserve species status. We observed that several of the characters mentioned above show a gradual overlap from one group to another, and it was not possible to draw significant lines between these groups recognisable for non-taxonomists to describe them as distinct species. Therefore, even though we examined several hundred specimens, we postpone a solution to this species delimitation problem within
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
<taxonomicName id="AB2B8AA67D6D7EFD50A7A0C0CC554E89" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
We are still confident that there are several more or less cryptic species involved, and for future studies we recommend measuring significantly more specimens than the 65 measured here. Morphometrics appears to be of high importance to find species limits, and might be a useful means to solve this problem. Molecular data, if possible from multiple genes, may also be necessary to support morphological and morphometric diagnostics.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,1525,1572][118,1122,118,559]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="mainText">
|
||||
<paragraph id="BDC73D83F93833B5F65E858DCD5CC70E" box="[118,1121,1525,1572][118,1122,118,559]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="mainText">
|
||||
Within the species complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="DE31022FD49CBFC8017D0503EE217A9F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
is not characterised and distinguishable by specialised characters, but instead by their lack.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="A8AAF3B6286D37E2BE71B66CAA6BF6B8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
(OI 19-25, usually above 20) differs strikingly from the two species with very small
|
||||
<pageBreakToken pageId="27" pageNumber="28" start="start">eyes</pageBreakToken>
|
||||
<pageBreakToken id="E07B7FF1189AF56365E7EC9F64D0B26F" pageId="27" pageNumber="28" start="start">eyes</pageBreakToken>
|
||||
:
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="4439D58ECC6524CD15A5FC03B36B35E6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
(OI 15-16) and
|
||||
<taxonomicName genus="Tetamorium" pageId="26" pageNumber="27" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="8CD77D13983AAFC37C9EA302BA366762" genus="Tetamorium" pageId="26" pageNumber="27" rank="species" species="nify">T. nify</taxonomicName>
|
||||
(OI 16-17). Furthermore,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="176012C99883850A19A5D6FF010E08CF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
can be easily distinguished from
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="DF60184578634A1A223665D3D02B7EB1" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="electrum">
|
||||
T. electrum
|
||||
<authority>,</authority>
|
||||
<authority id="94C5F06EF741514EA06CF43C8EFA1526">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="elf">
|
||||
<taxonomicName genus="Tetamorium" pageId="26" pageNumber="27" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="44C6F79C682547FC64A3ED48349F6E6A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="elf">
|
||||
<taxonomicName id="996FE21F33140CC147BF288C2C379C8B" genus="Tetamorium" pageId="26" pageNumber="27" rank="genus">T.</taxonomicName>
|
||||
elf,
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="6F8636EAF3E4204FFA945B13A6B17A91" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
based on their smaller eyes (16-19), very long to extremely long propodeal spines (PSLI 46-64), and different body colours.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
<taxonomicName id="213FA79F293E2DF95EBD4D00FD4F22C7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="98FFA4CF12B73CA6356EF1DCFCE64956" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
are very dark brown to black and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
<taxonomicName id="2274374E65DDC4FB769FD40427257AD3" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
is of whitish to yellowish colour, and even though
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="8BD1ABFF400F0191EF8F34070E0D1FBD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
shows a wide variation in colour, it is almost never yellowish as
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
<taxonomicName id="95AC8E22BA9FE1A88DD760BA58F00619" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
or as darkly coloured as
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="94234493F9C1787D6B06C3A2A0FBBEED" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum.</taxonomicName>
|
||||
<taxonomicName id="A806BD7FA6A43746A9B5D62EA235BECC" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum.</taxonomicName>
|
||||
In addition, T.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
<taxonomicName id="B94B560FE1E8A53B8C1B0090578DF996" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
with its very distinctive petiolar node with the posterodorsal corner strongly protruding posteriorly cannot be misidentified with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
<taxonomicName id="D9B1B684B549BF7FE6657473A36014F4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
which does not have a node shape like that.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ala">Tetramorium ala,</taxonomicName>
|
||||
<taxonomicName id="D873DCF878544B29FAFCCC6FEA0AF335" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ala">Tetramorium ala,</taxonomicName>
|
||||
however, can be confused with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
<taxonomicName id="842D20BE1426BC52A502FCCA18D6C185" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
They are both morphologically fairly similar, and in profile view almost indistinguishable. Nevertheless, in dorsal view they can be clearly separated from each other since the petiolar node of T. ala is much longer and thinner (DPeI 72-76) than that of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="F518FB93849BCF8EF8F0A8D95623F998" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
(DPeI 82-98). The last species of the complex,
|
||||
<taxonomicName genus="Tetamorium" pageId="26" pageNumber="27" rank="species" species="voasary">T. voasary,</taxonomicName>
|
||||
<taxonomicName id="50AB903AC7C61591963F1B80EF177659" genus="Tetamorium" pageId="26" pageNumber="27" rank="species" species="voasary">T. voasary,</taxonomicName>
|
||||
is morphologically also comparatively close to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
<taxonomicName id="D3A35F2F16B7441CEC5837D0C5784C49" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
Their separation is based on the shape differences of both waist segments. In
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="94F3EE2327A9BEE8F92D1686E05177B5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
the petiolar node in lateral view usually has very well defined, almost sharp, anterodorsal and posterodorsal margins, although they are often moderately rounded, and the postpetiole in dorsal view is between 1.0 to 1.4 times wider than long (DPpI 103-136). In contrast, the petiolar node of
|
||||
<taxonomicName genus="Tetamorium" pageId="26" pageNumber="27" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
<taxonomicName id="964205C8277EAF3C9AD8BCBDC2B497C0" genus="Tetamorium" pageId="26" pageNumber="27" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
has anterodorsal and posterodorsal margins that are fairly rounded, and a relatively longer postpetiole which is slightly longer than wide to slightly wider than long (DPpI 95-101).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,591,610]" pageId="27" pageNumber="28" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1124,619,1588]" pageId="27" pageNumber="28" type="mainText">
|
||||
<subSubSection id="5AB59A32E8B6D4C93D543B2CB97E446E" type="materials_examined">
|
||||
<paragraph id="5F1C910D1FC42A8EB3A969E15DE0900A" box="[118,295,591,610]" pageId="27" pageNumber="28" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="0C2B5CD2C49AB9E2E5108DCBBA70AD50" box="[118,1124,619,1588]" pageId="27" pageNumber="28" type="mainText">
|
||||
MADAGASCAR: Antananarivo, 3 km 41° NE Andranomay, 11.5 km 147° SSE Anjozorobe, 18.4733 S, 47.96 E, 1300 m, montane rainforest, 5.-13.XII.2000 (B.L. Fisher, C. Griswold et al.); Antsiranana, Parc Nat. Mont. d'Ambre, 1000-1100 m, wet forest, 12.II.1977 (W.L. & D.E. Brown); Antsiranana, Parc National Montagne d'Ambre [1st campsite], 12.5144 S, 49.1814 E, 960 m, rainforest, 21.-26.I.2001 (M.E. Irwin, E. Schlinger & R. Harin'Hala); Antsiranana, Parc National Montagne d'Ambre [Petit Lac road], 12.5203 S, 49.1792 E, 1125 m, rainforest, 29.I.-11.II.2001 (R. Harin'Hala); Antsiranana, Parc National Montagne d'Ambre, 12.2 km 211° SSW Joffreville, 12.5964 S, 49.1595 E, 1300 m, montane rainforest, 2.-7.II.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana, Parc National Montagne d'Ambre, 3.6 km 235° SW Joffreville, 12.5344 S, 49.1795 E, 925 m, 20.-26.I.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana, Parc National Montagne d'Ambre, Lac Maudit, 12.585 S, 49.1515 E, 1250 m, montane rainforest, 14.XI.2007 (B.L. Fisher et al.); Antsiranana, Parc National Montagne d'Ambre, Ambre Grand Lac, 12.5966 S, 49.1593 E, 1350 m, 13.XI.2007 (B.L. Fisher et al.); Antsiranana, Parc National Montagne d'Ambre, Antomboka, 12.5127 S, 49.1781 E, 970 m, 17.XI.2007 (B.L. Fisher et al.); Antsiranana, Parc National Montagne d'Ambre, Pic Bades, 12.5186 S, 49.1862 E, 900 m, montane rainforest, 20.XI.2007 (B.L. Fisher et al.); Antsiranana, Parc National Montagne d'Ambre, 12.5139 S, 49.1778 E, 984 m, montane rainforest, 28.II.2011 (B.L. Fisher et al.); Antsiranana, Parc National Montagne d'Ambre, 12.5178 S, 49.1796 E, 1000 m, 4.-7.III.2011 (B.L. Fisher et al.); Antsiranana, Parc National Montagne d'Ambre, 12.5286 S, 49.1772 E, 1100 m, 12.III.2011 (B.L. Fisher et al.); Antsiranana, Ampasindava,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="B9B485B14F18F09578AA62DFD35D7403" originalValue="Forêt">Foret</normalizedToken>
|
||||
d'Ambilanivy, 3.9 km 181° S Ambaliha, 13.7986 S, 48.1617 E, 600 m, 4.-9.III.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana, 11.0 km WSW Befingotra,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="21CC5A390AA5EAF749E4D9575AD0D03F" originalValue="Rés">Res</normalizedToken>
|
||||
. Anjanaharibe-Sud, 14.75 S, 49.45 E, 1550 m, montane rainforest, 18.XI.1994 (B.L. Fisher); Antsiranana, Betaolana Forest, along Bekona River, 14.53 S, 49.4404 E, 880 m, rainforest, 4.III.2009 (B.L. Fisher et al.); Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="880FF2B12D963A250737E16C19AD8951" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Binara, 9.4 km 235° SW Daraina, 13.2633 S, 49.6 E, 1100 m, montane rainforest, 5.XII.2003 (B.L. Fisher); Antsiranana, Makirovana forest, 14.1667 S, 49.95 E, 715 m, rainforest, 1.-2.V.2011 (B.L. Fisher et al.); Antsiranana, R.S. Manongarivo, 12.8 km 228° SW Antanambao, 13.9767 S, 48.4233 E, 780 m, rainforest, 11.X.1998 (B.L. Fisher); Antsiranana, R.S. Manongarivo, 14.5 km 220° SW Antanambao, 13.9983 S, 48.4283 E, 1175 m, montane rainforest, 20.X.1998 (B.L. Fisher); Antsiranana, R.S. Manongarivo, 10.8 km 229° SW Antanambao, 13.9617 S, 48.4333 E, 400 m, rainforest, 8.XI.1998 (B.L. Fisher); Antsiranana, RNI Marojejy, 10.5 km NW Manantenina, 14.4333 S, 49.75 E, 1625 m, 6.-12.XI.1996 (E.L. Quinter); Antsiranana, Parc National de Marojejy, Antranohofa, 26.6 km 31° NNE Andapa, 10.7 km 318° NW Manantenina, 14.4433 S, 49.7433 E, 1325 m, montane rainforest, 19.III.2003 (B.L. Fisher); Antsiranana, Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, 15.-18.XI.2003 (B.L. Fisher et al.); Parc National de Marojejy, 25.7 km 32° NNE Andapa, 10.3 km 314° NW Manantenina, 14.445 S, 49.7417 E, 1575 m, montane rainforest, 21.XI.2003 (B.L. Fisher); Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, 14.XII.2005 (B.L. Fisher et al.); Fianarantsoa, Foret d'Ambalagoavy Nord, Ikongo, Ambatombe, 21.8275 S, 47.3389 E, 625 m, 1.XII.2000 (R. Harin'Hala & M.E. Irwin); Fianarantsoa, 45km S. Ambalavao, 22.2167 S, 47.0167 E, 785 m, rainforest,
|
||||
</paragraph>
|
||||
<paragraph box="[118,1124,118,1589]" pageId="28" pageNumber="29" type="mainText">
|
||||
<pageBreakToken pageId="28" pageNumber="29" start="start">24</pageBreakToken>
|
||||
<paragraph id="31CC2888BFA81BE3ABAAF4F07323329D" box="[118,1124,118,1589]" pageId="28" pageNumber="29" type="mainText">
|
||||
<pageBreakToken id="8EB5AAF37069502A9BEC10E566FDB29D" pageId="28" pageNumber="29" start="start">24</pageBreakToken>
|
||||
.-25.IX.1993 (B.L. Fisher); Fianarantsoa, 43 km S Ambalavao,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="DCC7EECD821477CAC8C3C460B94167C1" originalValue="Rés">Res</normalizedToken>
|
||||
. Andringitra, 22.2333 S, 47 E, 825 m, rainforest, 5.X.1993 (B.L. Fisher); Fianarantsoa, R.S. Ivohibe, 7.5 km ENE Ivohibe, 22.47 S, 46.96 E, 900 m, rainforest, 7.-12.X.1997 (B.L. Fisher); Fianarantsoa,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="27DBF1328F948675F0E9F5C98CF791BB" originalValue="Réserve">Reserve</normalizedToken>
|
||||
Speciale Manombo 24.5 km 228° Farafangana, 23.0158 S, 47.719 E, 30 m, rainforest, 20.-22.IV.2006 (B.L. Fisher et al.); Fianarantsoa,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken originalValue="Classée">Classee</normalizedToken>
|
||||
<normalizedToken id="EDE9947CD9CA2CE2C540DD9E896AFA98" originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="F1AA85168EE9806C745C4141C828F553" originalValue="Classée">Classee</normalizedToken>
|
||||
Vatovavy, 7.6 km 122
|
||||
<normalizedToken originalValue="º">°</normalizedToken>
|
||||
<normalizedToken id="18F7ABD4B564AC1BB2B6716858F01BD7" originalValue="º">°</normalizedToken>
|
||||
Kianjavato, 21.4 S, 47.94 E, 175 m, rainforest, 6.-8.VI.2005 (B.L. Fisher et al.); Fianarantsoa,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="ED8C174FE318727FD0E0EFD4990A3C9A" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Vevembe, 66.6 km 293° Farafangana, 22.791 S, 47.1818 E, 600 m, rainforest, transition to montane forest, 23.-24.IV.2006 (B.L. Fisher et al.); Mahajanga,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="FDB232FDF212643AA99C0498F349FF0E" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="F5442C9C3D5B490E20A1D9791E7969B2" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Marotandrano, Marotandrano 48.3 km S Mandritsara, 16.2832 S, 48.8144 E, 865 m, transitional humid forest, 6.-8.XII.2007 (B.L. Fisher et al.); Toamasina, Montagne d'Akirindro 7.6 km 341° NNW Ambinanitelo, 15.2883 S, 49.5483 E, 600 m, 17.-21.III.2003 (B.L. Fisher, C. Griswold et al.); Toamasina, 6.3 km S Ambanizana, Andranobe, 15.6813 S, 49.958 E, 25 m, rainforest, 13.-23.XI.1993 (B.L. Fisher); Toamasina, 6.9 km NE Ambanizana, Ambohitsitondroina, 15.5667 S, 50 E, 825 m, rainforest, 2.XII.1993 (B.L. Fisher); Toamasina, Ambanizana, Parc National Masoala, 15.5722 S, 50.0069 E, 1020 m, montane rainforest, 2.-6.III.2003 (D. Andriamalala, D. Silva et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="B96DD1255BE94E48B9E4B46CABC705FB" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="7256FD1E8FBFA58AAF96BA502F8F5289" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato River, 16.7727 S, 49.2655 E, 450 m, rainforest, 20.-22.II.2010 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="A3659B9D45C30763794E200EFE654792" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="8C6B182EEC0A9C201B53436A3B83ECA6" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato River, 16.7633 S, 49.2669 E, 520 m, rainforest, 22.-24.II.2010 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="2CD390A4EDA6CE20FCBF03008787C508" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="D26548E86A414D6FDBFA18FEEDC8DE23" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato River, 16.8175 S, 49.295 E, 360 m, rainforest, 25.-27.II.2010 (B.L. Fisher et al.); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.8496 S, 48.2947 E, 1010 m, montane rainforest, 3.-6.III.2007 (B.L. Fisher et al.); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.8477 S, 48.2957 E, 1000 m, montane rainforest, 5.-8.III.2007 (B.L. Fisher et al.); Toamasina, Amparihibe, 15° 2' S, 49° 34' E, II.-III.2003 (K.A. Jackson & D. Carpenter); Toamasina, Analamay, 18.8062 S, 48.3371 E, 1068 m, montane rainforest, 21.III.2004 (Malagasy ant team); Toamasina, Station
|
||||
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
|
||||
<normalizedToken id="1F0DAEB96B5455663E89753A18D22928" originalValue="forestière">forestiere</normalizedToken>
|
||||
Analamazaotra, Analamazaotra 1.3 km S Andasibe, 18.3847 S, 48.4127 E, 980 m, montane rainforest, 11.-13.XII.2007 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="8D484A066A556A986EC76361507D2FBB" originalValue="Forêt">Foret</normalizedToken>
|
||||
d'Analava Mandrisy, 5.9 km 195
|
||||
<normalizedToken originalValue="º">°</normalizedToken>
|
||||
<normalizedToken id="1A7E54885AE6490D61A11791285DB5BA" originalValue="º">°</normalizedToken>
|
||||
Antanambe, 16.4857 S, 49.847 E, 10 m, littoral rainforest, 13.XI.2005 (B.L. Fisher et al.); Toamasina, 6 km ESE Andasibe (= Perinet), 18.95 S, 48.4667 E, 900 m, 17.XI.1990 (P.S. Ward); Toamasina, F.C. Andriantantely, 18.695 S, 48.8133 E, 530 m, 4.-10.XII.1998 (H.J. Ratsirarson); Toamasina, Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo, 15.1883 S, 49.615 E, 470 m, rainforest, 8.-12.III.2003 (B.L. Fisher, C. Griswold et al.); Toamasina, Reserve Betampona, Camp Rendrirendry 34.1 km 332° Toamasina, 17.924 S, 49.1997 E, 390 m, rainforest, 28.XI.2005 (B.L. Fisher et al.); Toamasina, Reserve Betampona, Camp Vohitsivalana, 37.1 km 338° Toamasina, 17.8867 S, 49.2025 E, 520 m, rainforest, 1.-3.XII.2005 (B.L. Fisher et al.); Toamasina, F.C. Didy, 18.1983 S, 48.5783 E, 960 m, rainforest, 16.-23.XII.1998 (H.J. Ratsirarson); Toamasina, Parc National Mananara-Nord, 7.1 km 261° Antanambe, 16.455 S, 49.7875 E, 225 m, rainforest, 14.XI.2005 (B.L. Fisher et al.); Toamasina, P.N. Mantadia, 18.7917 S, 48.4267 E, 895 m, rainforest, 28.XI.-1.XII.1998 (H.J. Ratsirarson); Toamasina,
|
||||
<normalizedToken originalValue="Nosibé">Nosibe</normalizedToken>
|
||||
<normalizedToken id="E1857C5E680C9ACC4E1EE4CD6E3C4E11" originalValue="Nosibé">Nosibe</normalizedToken>
|
||||
Village de
|
||||
<normalizedToken originalValue="I’Imerina">I'Imerina</normalizedToken>
|
||||
<normalizedToken id="FB9965E898915542DE82C1AD2204D5E8" originalValue="I’Imerina">I'Imerina</normalizedToken>
|
||||
, province de Bezanozano (M. Sikora); Toamasina, Nosy Mangabe, 15° 30' S, 49° 46' E, 300 m, rainforest, 18.IV.1989 (P.S. Ward); Toamasina, Tamatave, Ampasimbe, 450 m, rainforest (J.M. Betsch); Toamasina, S.F. Tampolo, 10 km NNE Fenoarivo Atn., 17.2825 S, 49.43 E, 10 m, littoral rainforest, 10.IV.1997 (B.L. Fisher); Toamasina, Parcelle K9 Tampolo, 17.175 S, 49.268 E, 10 m, littoral forest, 19.IV.2004 (Malagasy ant team); Toamasina, Torotorofotsy, 18.8708 S, 48.3474 E, 1070 m, montane rainforest, marsh edge, 24.III.2004 (Malagasy ant team); Toamasina, Parc National de Zahamena, Tetezambatana forest, near junction of Nosivola and Manakambahiny Rivers, 17.743 S, 48.7294 E, 860 m, rainforest, 18.-19.II.2009 (B.L. Fisher et al.); Toamasina, Parc National de Zahamena, 17.7336 S, 48.7262 E, 950 m, rainforest, 19.II.2009 (B.L. Fisher et al.); Toamasina, Parc National de Zahamena, Besaky River, 17.7524 S, 48.8532 E, 760 m, rainforest, 22.II.2009 (B.L. Fisher et al.); Toamasina, Parc National de Zahamena, Onibe River, 17.7591 S, 48.8547 E, 780 m, rainforest, 21.-23.II.2009 (B.L. Fisher et al.); Toliara,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="BB8CC6A6B96021B852B95CDE7D8A4A13" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="58C3024C01FC6A9EA1FD826D57953FEF" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
d'Ambohijanahary,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="52A6F68DA18398572D54DE165ED79100" originalValue="Forêt">Foret</normalizedToken>
|
||||
d'Ankazotsihitafototra, 35.2 km 312° NW Ambaravaranala, 18.26 S, 45.4183 E, 1100 m, montane rainforest, 13.-17.I.2003 (B.L. Fisher, C. Griswold et al.); Toliara,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="A09C35DA975E8A58AB6EC69E359543B1" originalValue="Rés">Res</normalizedToken>
|
||||
. Andohahela, 11 km NW Enakara, 24.5667 S, 46.8333 E, 800 m, rainforest, 16.XI.1992 (B.L. Fisher); Toliara,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="639B70510454F226F0BA8016F8201B9F" originalValue="Rés">Res</normalizedToken>
|
||||
. Andohahela, 10 km NW Enakara, 24.5667 S, 46.8167 E, 430 m, rainforest, 22.XI.1992 (B.L. Fisher); Toliara,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="F946921F030A5EEA7DF97F66E6D9CBFF" originalValue="Rés">Res</normalizedToken>
|
||||
. Andohahela, 6 km SSW Eminiminy, 24° 44' S, 46° 48' E, 330 m, rainforest, 4.II.1993 (P.S. Ward); Toliara, Parc National d'Andohahela, Manampanihy River, 5.4 km 113° ESE Mahamavo, 36.7 km 343° NNW Tolagnaro, 24.7639 S, 46.7668 E, 650 m, rainforest, 24.I.2002 (B.L. Fisher et al.); Toliara, Parc National Andohahela, Col de Tanatana, 33.3 km NW Tolagnaro, 24.7585 S, 46.8537 E, 275 m, rainforest, 22.-24.XI.2006 (B.L. Fisher et al.); Toliara,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="AF79EAA013971685E9325EAAE189849E" originalValue="Forêt">Foret</normalizedToken>
|
||||
Ivohibe 55.0 km N Tolagnaro, 24.569 S, 47.204 E, 200 m, rainforest, 2.-4.XII.2006 (B.L. Fisher et al.); Toliara, Grand Lavasoa, 25.9 km W Tolagnaro, 25.0877 S, 46.749 E, 450 m, rainforest, 30.XI.-2.XII.2006 (B.L. Fisher et al.); Toliara, 2.7 km WNW 302° Ste. Luce, 24.7717 S, 47.1717 E, 20 m, littoral rainforest, 9.-11.XII.1998 (B.L. Fisher).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,83 +1,82 @@
|
|||
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172629" ID-GBIF-Taxon="100105360" ID-Zenodo-Dep="6172629" LSID="urn:lsid:plazi:treatment:298B167E41AB351168CE8CC53525B0B0" httpUri="http://treatment.plazi.org/id/298B167E41AB351168CE8CC53525B0B0" lastPageNumber="47" pageNumber="47">
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<subSubSection type="nomenclature">
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<paragraph box="[118,441,906,926]" pageId="46" pageNumber="47" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">Tetramorium jedi</taxonomicName>
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<treatment id="298B167E41AB351168CE8CC53525B0B0" ID-DOI="http://doi.org/10.5281/zenodo.6172629" ID-GBIF-Taxon="100105360" ID-Zenodo-Dep="6172629" LSID="urn:lsid:plazi:treatment:298B167E41AB351168CE8CC53525B0B0" httpUri="http://treatment.plazi.org/id/298B167E41AB351168CE8CC53525B0B0" lastPageNumber="47" pageNumber="47" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="29CA9B40CBAD8A71AB6F82877CA30306" type="nomenclature">
|
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<paragraph id="25A7FC6AC1B95D5C15ABD29DB707EF13" box="[118,441,906,926]" pageId="46" pageNumber="47" type="mainText">
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<taxonomicName id="A175A069B742F9ED7C7241E88F6236E8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">Tetramorium jedi</taxonomicName>
|
||||
species complex
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,1122,961,1036]" pageId="46" pageNumber="47" type="mainText">
|
||||
<subSubSection id="C339BDC2F50D5A7F42E5FBB1366E4FE7" type="discussion">
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<paragraph id="DB203EC4928B6A505C3F55C17A4AFAE4" box="[118,1122,961,1036]" pageId="46" pageNumber="47" type="mainText">
|
||||
The
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="3AFC808521FC259426E3CB2FE853A2C4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
complex is relatively small with just the three species
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="avaratra">
|
||||
<taxonomicName id="35A1811135F797CA36944AE6AFFDD61B" authorityName="," genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="avaratra">
|
||||
T. avaratra
|
||||
<authority>,</authority>
|
||||
<authority id="5D91A8DC8C7FC534E47793FF97CCB023">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">T. jedi,</taxonomicName>
|
||||
<taxonomicName id="A83D93CD1B788F22CA35E7C639F4AB58" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">T. jedi,</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon.</taxonomicName>
|
||||
<taxonomicName id="184B5E2B5CCE1763C2D6EB5899B5FC78" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon.</taxonomicName>
|
||||
It is characterized by the absence of sculpture on the forecoxae and the presence of reticulate-punctate sculpture on the first gastral tergite.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,1046,1205]" pageId="46" pageNumber="47" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">Tetramorium jedi</taxonomicName>
|
||||
<paragraph id="DCE356562632B3F9E19A5B5C99760527" box="[118,1122,1046,1205]" pageId="46" pageNumber="47" type="mainText">
|
||||
<taxonomicName id="1E5539D63173DCC9F0B133021E886FFB" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">Tetramorium jedi</taxonomicName>
|
||||
does not seem to be morphologically close to the other two species of the complex, and they were grouped together on the basis of the reticulate-punctate sculpture on the first gastral tergite present in all three, even though the development of the sculpture from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="246EE0EB66A2E5C653E5DDD5C0C09F1F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="BFF830E4524DBA27C8E22B606DCC067A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="93D29FB2729E29C511840835DF867FED" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
is fairly different. The latter two, as mentioned above, may not be closely associated with any other
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="4D3E56EACF62D50EFBDCD6E04167C992" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group members.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">Tetramorium jedi,</taxonomicName>
|
||||
<taxonomicName id="5D21361EDBDD6C284511C3AA275BFFE7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">Tetramorium jedi,</taxonomicName>
|
||||
however, might be distantly related to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="11F356E94339D58F9EBAE2DAC3BF1C2A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
and allies since the main separating character is the conspicuous sculpture on the first gastral tergite present in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="7399105F4FF9C3F93AA4AB3DEC57BF56" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
but absent in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="4599893D85FAC89B8A620BD955986759" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
complex.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,189 +1,188 @@
|
|||
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<mods:title>The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups.</mods:title>
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<mods:namePart>B. L. Fisher</mods:namePart>
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<mods:namePart id="05F0BCDD56FE3AF5259EDCF025064247">B. L. Fisher</mods:namePart>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172661" ID-GBIF-Taxon="100105387" ID-Zenodo-Dep="6172661" LSID="urn:lsid:plazi:treatment:2BF1F9E82CB2C39898CB85AFC70E4E6B" httpUri="http://treatment.plazi.org/id/2BF1F9E82CB2C39898CB85AFC70E4E6B" lastPageNumber="79" pageNumber="76">
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<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="sabatra">
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<treatment id="2BF1F9E82CB2C39898CB85AFC70E4E6B" ID-DOI="http://doi.org/10.5281/zenodo.6172661" ID-GBIF-Taxon="100105387" ID-Zenodo-Dep="6172661" LSID="urn:lsid:plazi:treatment:2BF1F9E82CB2C39898CB85AFC70E4E6B" httpUri="http://treatment.plazi.org/id/2BF1F9E82CB2C39898CB85AFC70E4E6B" lastPageNumber="79" pageNumber="76" scope_family="Formicidae" scope_order="Hymenoptera">
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<taxonomicName id="415E07F827096A2CAA1BCCA02EC634E1" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="sabatra">
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Tetramorium sabatra
|
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<authority>Hita</authority>
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<authority id="3D760D1ABDD84D81192539D337698EEA">Hita</authority>
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Garcia & Fisher
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<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
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<taxonomicNameLabel id="787E4DB0874A31DC128056397BC0AF5F" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection type="description">
|
||||
<paragraph box="[118,508,961,980]" pageId="75" pageNumber="76" type="mainText">(Figs. 11, 15, 24, 25, 26, 135, 136, 137, 142)</paragraph>
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<subSubSection id="0D3688FA06AD6ED9EA9E8AB4F8AB862E" type="description">
|
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<paragraph id="7B75E997D028DD679FEB671EFC0A8487" box="[118,508,961,980]" pageId="75" pageNumber="76" type="mainText">(Figs. 11, 15, 24, 25, 26, 135, 136, 137, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1122,1017,1233]" pageId="75" pageNumber="76" type="mainText">
|
||||
<subSubSection id="FC1219903872CFAD3101C09647574259" type="materials_examined">
|
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<paragraph id="D33B0AA4FF263AB77C54D8846FB6AC4C" box="[118,1122,1017,1233]" pageId="75" pageNumber="76" type="mainText">
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||||
Holotype worker, MADAGASCAR, Toliara,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="C6E2C8D6C00700CCCE54578F38E16560" originalValue="Rés">Res</normalizedToken>
|
||||
. Andohahela, 11 km NW Enakara, 24.56667 S, 46.83333 E, 800 m, montane rainforest, pitfall trap, collection code BLF00490, 16.XI.1992 (B.L. Fisher) (CASC: CASENT0189241). Paratypes, three workers with same data as holotype (CASC: CASENT0218056; CASENT0218057; CASENT0270780); one worker from Toliara,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="8344A0076715AAB123D4214162915596" originalValue="Rés">Res</normalizedToken>
|
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. Andohahela, 11 km NW Enakara, 24.56667 S, 46.83333 E, 800 m, montane rainforest, from sifted litter, collection code BLF00492, 17.XI.1992 (B.L. Fisher) (CASC: CASENT0189240); and one worker from Toliara,
|
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<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="7233614C9C4C60112C7090F0904D38D2" originalValue="Rés">Res</normalizedToken>
|
||||
. Andohahela, 10 km NW Enakara, 24.56667 S, 46.81667 E, 430 m, rainforest, sifted litter, collection code BLF00522, 22.XI.1992 (B.L. Fisher) (CASC: CASENT0189239).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,1265,1284]" pageId="75" pageNumber="76" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,1293,1453]" pageId="75" pageNumber="76" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="sabatra">Tetramorium sabatra</taxonomicName>
|
||||
<subSubSection id="03F2AE8FE22A06DB5D5E452495B3EA93" type="diagnosis">
|
||||
<paragraph id="2486A868055D0F54A7AACC0A9BD5465B" box="[118,208,1265,1284]" pageId="75" pageNumber="76" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="14B8567257F10EC0B01CA97C842A7D24" box="[118,1121,1293,1453]" pageId="75" pageNumber="76" type="mainText">
|
||||
<taxonomicName id="F21D579F50334629F88140D21D8F2129" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="sabatra">Tetramorium sabatra</taxonomicName>
|
||||
is easily distinguished from the other group members by the following character combination: antennal scapes comparatively short (SI 73-80); extremely long and massively constructed propodeal spines (PSLI 48-72); anterodorsal and posterodorsal margins of petiolar node situated at about same height; mesosoma with one or two pairs of standing hairs, restricted to dorsal pronotum; hairs on leading edge of antennal scapes usually strongly appressed; first gastral tergite with few standing hairs and very sparse, short pubescence; very dark brown to black colouration.
|
||||
</paragraph>
|
||||
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|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,1485,1503]" pageId="75" pageNumber="76" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1124,1513,1589][118,1124,118,166]" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="mainText">
|
||||
<subSubSection id="2B7F7A020F4556B6056101C587D9EBB0" type="description">
|
||||
<paragraph id="D69A8962CC66BBAD07602FA74ABBB212" box="[118,226,1485,1503]" pageId="75" pageNumber="76" type="mainText">Description</paragraph>
|
||||
<paragraph id="CD0688590E4B17303B9EA2E45A13C326" box="[118,1124,1513,1589][118,1124,118,166]" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="mainText">
|
||||
HL 1.00-1.12 (1.04); HW 1.02-1.13 (1.05); SL 0.76-0.90 (0.82); EL 0.20-0.23 (0.21); PH 0.48-0.54 (0.50); PW 0.71-0.79 (0.75); WL 1.27-1.41 (1.33); PSL 0.50-0.81 (0.60); PTL 0.34-0.37 (0.36); PTH 0.39-0.44 (0.41); PTW 0.27-0.31 (0.28); PPL 0.30-0.34 (0.32); PPH 0.40-0.46 (0.42); PPW 0.37-0.44 (0.39); CI 100-103 (101); SI 73-80
|
||||
<pageBreakToken pageId="76" pageNumber="77" start="start">(</pageBreakToken>
|
||||
<pageBreakToken id="2AACFC334C3CF2C99596626B195D5124" pageId="76" pageNumber="77" start="start">(</pageBreakToken>
|
||||
77); OI 18-21 (20); DMI 55-58 (56); LMI 37-40 (38); PSLI 48-72 (57); PeNI 36-39 (38); LPeI 82-90 (87); DPeI 75-86 (80); PpNI 50-56 (52); LPpI 71-77 (75); DPpI 119-129 (125); PPI 134-143 (138) (ten measured).
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,175,869]" pageId="76" pageNumber="77" type="mainText">Head as long as wide to weakly wider than long (CI 100-103); posterior head margin strongly concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes well-developed, moderately deep, but narrow, and without defined posterior and ventral margins. Antennal scapes short to moderate, not reaching posterior head margin (SI 73-80). Eyes small to moderate (OI 18-21). Mesosomal outline in profile flat, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-40). Propodeal spines massively developed with very broad base, extremely long, and acute (PSLI 48-72); propodeal lobes short and blunt. Petiolar node in profile rectangular nodiform, approximately 1.1 to 1.2 times higher than long (LPeI 82-90), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum slightly convex; node in dorsal view approximately 1.1 to 1.3 times longer than wide (DPeI 75-86). Postpetiole in profile subglobular, weakly anteroposteriorly compressed, approximately 1.3 to 1.4 times higher than long (LPpI 71-77); in dorsal view around 1.2 to 1.3 times wider than long (DPpI 119-129). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view approximately 1.3 to 1.5 times wider than petiolar node (PPI 134-143). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose, with three to five rugae; cephalic dorsum between frontal carinae with 9 to 12 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted or with cross-meshes; lateral and ventral head longitudinally rugose, rarely with cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae with very distinct and pronounced longitudinal rugae. Waist segments longitudinally rugose, rugae on waist segments weaker than on head and mesosoma, especially dorsally. Gaster completely unsculptured, smooth, and shining. Ground sculpture generally faint to absent everywhere on body. Head with abundant standing hairs; hairs on mesosoma restricted to dorsal pronotum, usually one or two pairs of hairs present, rarely three pairs; waist segments and first gastral tergite with few to numerous standing hairs; first gastral tergite with very sparse, short, and appressed pubescence. Anterior edges of antennal scapes usually with appressed hairs (decumbent in one specimen). Body a uniform very dark brown to black colour.</paragraph>
|
||||
<paragraph id="8A3C494E786ECC78D00F73682B5E591B" box="[118,1122,175,869]" pageId="76" pageNumber="77" type="mainText">Head as long as wide to weakly wider than long (CI 100-103); posterior head margin strongly concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes well-developed, moderately deep, but narrow, and without defined posterior and ventral margins. Antennal scapes short to moderate, not reaching posterior head margin (SI 73-80). Eyes small to moderate (OI 18-21). Mesosomal outline in profile flat, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-40). Propodeal spines massively developed with very broad base, extremely long, and acute (PSLI 48-72); propodeal lobes short and blunt. Petiolar node in profile rectangular nodiform, approximately 1.1 to 1.2 times higher than long (LPeI 82-90), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum slightly convex; node in dorsal view approximately 1.1 to 1.3 times longer than wide (DPeI 75-86). Postpetiole in profile subglobular, weakly anteroposteriorly compressed, approximately 1.3 to 1.4 times higher than long (LPpI 71-77); in dorsal view around 1.2 to 1.3 times wider than long (DPpI 119-129). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view approximately 1.3 to 1.5 times wider than petiolar node (PPI 134-143). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose, with three to five rugae; cephalic dorsum between frontal carinae with 9 to 12 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted or with cross-meshes; lateral and ventral head longitudinally rugose, rarely with cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae with very distinct and pronounced longitudinal rugae. Waist segments longitudinally rugose, rugae on waist segments weaker than on head and mesosoma, especially dorsally. Gaster completely unsculptured, smooth, and shining. Ground sculpture generally faint to absent everywhere on body. Head with abundant standing hairs; hairs on mesosoma restricted to dorsal pronotum, usually one or two pairs of hairs present, rarely three pairs; waist segments and first gastral tergite with few to numerous standing hairs; first gastral tergite with very sparse, short, and appressed pubescence. Anterior edges of antennal scapes usually with appressed hairs (decumbent in one specimen). Body a uniform very dark brown to black colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,906,924]" pageId="76" pageNumber="77" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1125,934,1178]" pageId="76" pageNumber="77" type="mainText">
|
||||
<subSubSection id="0B8447AE4D67301AF5CA89FA26C4ACEE" type="discussion">
|
||||
<paragraph id="8A25B53254C5729B8F05E9D65D62EF50" box="[118,170,906,924]" pageId="76" pageNumber="77" type="mainText">Notes</paragraph>
|
||||
<paragraph id="CE8CDADC8F8BEA6E7E6C087911FEBD0F" box="[118,1125,934,1178]" pageId="76" pageNumber="77" type="mainText">
|
||||
Despite the comparatively small number of known specimens (15 in total),
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="2E656B300C3184AAE790AB12EB3D12D1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
seems to be widely distributed in the rainforests and montane rainforests of eastern Madagascar, as well as in Analavelona in the southwest. The southernmost locality is the type locality, Andohahela, and the northernmost locality is Montagne d'Akirindro. Inbetween it is only known from few more localities. Furthermore,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="05EBB66375E2DE0CAD39D7770877A81D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
seems to inhabit forests at elevations of 430 to 1300 m, and was mainly collected from the ground. The scarcity of material and wide distribution range suggests that this species is fairly rare, uncommon, or just sampled inappropriately. Four out of the fifteen specimens were collected while visiting the flowers of
|
||||
<taxonomicName authorityName="Fischer & Rahelivololona" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" order="Ericales" pageId="76" pageNumber="77" phylum="Tracheophyta" rank="species" species="mandrakae">
|
||||
<taxonomicName id="495A35007F4CA6C26DB462175C2F3460" authorityName="Fischer & Rahelivololona" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" order="Ericales" pageId="76" pageNumber="77" phylum="Tracheophyta" rank="species" species="mandrakae">
|
||||
Impatiens mandrakae
|
||||
<authority>Fischer & Rahelivololona</authority>
|
||||
<authority id="0B00775EFD2F4D4F9B6032B7454C2AF3">Fischer & Rahelivololona</authority>
|
||||
</taxonomicName>
|
||||
(
|
||||
<taxonomicName family="Balsaminaceae" genus="Balsaminaceae" pageId="76" pageNumber="77" rank="family">Balsaminaceae</taxonomicName>
|
||||
<taxonomicName id="F058C2E2CAC22ACAF8EE7BED670A84F4" class="Magnoliopsida" family="Balsaminaceae" higherTaxonomySource="CoL" kingdom="Plantae" order="Ericales" pageId="76" pageNumber="77" phylum="Tracheophyta" rank="family">Balsaminaceae</taxonomicName>
|
||||
) at Mandraka. This suggests that
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="741229F09E700A3982463CBE92C46EA5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
lives in the vegetation and is therefore rarely sampled from the ground. This could be true for
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="D64DBF4498FBEAD7732BF02A228227D7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">T. smaug,</taxonomicName>
|
||||
<taxonomicName id="DDF09378E85A8FA750D0E5AD506F4462" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">T. smaug,</taxonomicName>
|
||||
too, and might explain the rarity of these three species.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1124,1187,1572]" pageId="76" pageNumber="77" type="mainText">
|
||||
<paragraph id="D6EBD159A79BE54A84750544BA768584" box="[118,1124,1187,1572]" pageId="76" pageNumber="77" type="mainText">
|
||||
Inside the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="3BC1E98A2B510837785524B0E0BEA06F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="2E3A4D72901BD0EDC05FA3ACB4DC8651" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
is unlikely to be misidentified with
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="adamsi">
|
||||
<taxonomicName id="1B562F6105E2F5F7DEA70B60040FB4C4" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="adamsi">
|
||||
T. adamsi
|
||||
<authority>,</authority>
|
||||
<authority id="9E27A7505BF62C0715CA9540CC4270EA">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy,</taxonomicName>
|
||||
<taxonomicName id="7F6E57BB5255F0994D8FADB8EB6FBA6A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy,</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul.</taxonomicName>
|
||||
<taxonomicName id="87564321CED9CC26C96DE179F8B5A567" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul.</taxonomicName>
|
||||
The latter three have numerous standing hairs on the first gastral tergite, and are much hairier than
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
<taxonomicName id="8074E7C668764ED170013A333BF784BA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
Furthermore,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
<taxonomicName id="D9447D4CCE674C67F841B42F119EA298" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
has a petiolar node shape with the posterodorsal margin situated higher than the anterodorsal, while both margins are at the same height in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="nazgul">Tetramorium nazgul</taxonomicName>
|
||||
<taxonomicName id="C2504B1EDC8B4F37E9E080A986D7571D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
<taxonomicName id="06F7AAB26F37BE148B8D45D556030A7E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="nazgul">Tetramorium nazgul</taxonomicName>
|
||||
also has much longer antennal scapes (SI 89-93), and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
<taxonomicName id="4D1A1AFE8F6538A0525C8CA794E3A9EF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
is of orange to light brown body colour. Nevertheless,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="12735E18832D3C5C82DDEDC667A5EC92" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
appears to be morphologically most closely associated with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="179C490E27A7E0C3EAA843AF7643E929" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="EACB7BA589E0F6D70E6F6A031F7AD655" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
since they share the same morphometric range and have a very similar gestalt. They are very darkly coloured species with massively developed propodeal spines and reduced hairiness. However,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="B21C0B3EB676A77E2027C0567045E199" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
can be well separated from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="573E6046CF02F2483607F26976305510" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
due to the absence of standing hairs on the first gastral tergite in the latter, whereas a few to several standing hairs are always present in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
<taxonomicName id="1F4DB032CA7D72027DB4206EA0894D34" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
In addition, the latter species has only very sparse and inconspicuous pubescence on the first gastral tergite while
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="720B4D8BC54AADCDC841B7A38673CC4D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
has moderately dense and distinct pubescence.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
<taxonomicName id="F3054C404529D041A4ECBFE7B899D44E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
mainly differs from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="4B4DABA0B3925DB8EF933767BAA708BA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
in the number of hairs on the mesosomal dorsum. The latter has just one or two pairs on the pronotal dorsum, whereas
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="71869B5435BD67986D7863343E9C98DF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
has 7 to 14 pairs throughout the whole mesosomal dorsum. Also, in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="E7AAF48C9789D3430C84784186AA5CA7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
the hairs on the leading edges of the antennal scapes are usually appressed but subdecumbent to suberect in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
<taxonomicName id="F7B602D08079BFE3E4E2C480716E0733" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
</paragraph>
|
||||
<caption box="[118,1122,1534,1579]" pageId="77" pageNumber="78" type="caption">
|
||||
<paragraph box="[118,1122,1534,1579]" pageId="77" pageNumber="78" type="caption">
|
||||
<pageBreakToken pageId="77" pageNumber="78" start="start">FIGURES</pageBreakToken>
|
||||
<caption id="F98814581BE75D53384DE1F2CA7936B0" box="[118,1122,1534,1579]" pageId="77" pageNumber="78" type="caption">
|
||||
<paragraph id="2756BD7578C62C4C002CC8FA53BB99B5" box="[118,1122,1534,1579]" pageId="77" pageNumber="78" type="caption">
|
||||
<pageBreakToken id="1CF2AB1A45D87EC1947F8215EEFD504D" pageId="77" pageNumber="78" start="start">FIGURES</pageBreakToken>
|
||||
135-137.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
<taxonomicName id="7BEBFB2F92F174BB63C553FA8CB65504" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
holotype (CASENT0189241). 135. Body in profile. 136. Body in dorsal view. 137. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,1122,118,250]" pageId="78" pageNumber="79" type="mainText">
|
||||
<pageBreakToken pageId="78" pageNumber="79" start="start">As</pageBreakToken>
|
||||
<paragraph id="5570CA66FC299F399742DDC03CC0C3F9" box="[118,1122,118,250]" pageId="78" pageNumber="79" type="mainText">
|
||||
<pageBreakToken id="5616DE4D68D9A1E426BD59983E6B871B" pageId="78" pageNumber="79" start="start">As</pageBreakToken>
|
||||
noted in the above descriptions,
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
<taxonomicName id="F4B2E07F6FA80C737EB4F9056B52E94E" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
T. latreillei
|
||||
<authority>,</authority>
|
||||
<authority id="EDF8F4AC9C80308DCDF4CC60B0C15AE8">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
<taxonomicName id="A070B9A9114CFEB94FBB80D9BA3AFE20" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="8592CAFDA717E1D4E411165B5EAEC4A3" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
could all belong to one species with considerable variation in patterns of pilosity/pubescence. However, as previously discussed, we are of the opinion that these characters are generally of high diagnostic value at the species level, which leads us to treat the three as distinct species. The available material is limited, however, and more material could demonstrate that our current species delimitations are incorrect.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,287,305]" pageId="78" pageNumber="79" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,615,315,334]" pageId="78" pageNumber="79" type="mainText">The species epithet is an arbitrary combination of letters.</paragraph>
|
||||
<subSubSection id="00541747C8853F364D0BB193E7EC2DE0" type="etymology">
|
||||
<paragraph id="9292CB6B4F7C8D9E4344C428CF96D5AF" box="[118,219,287,305]" pageId="78" pageNumber="79" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="FA5B724846F0EE8F8F13B4F8D4021FA1" box="[118,615,315,334]" pageId="78" pageNumber="79" type="mainText">The species epithet is an arbitrary combination of letters.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,371,390]" pageId="78" pageNumber="79" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,399,644]" pageId="78" pageNumber="79" type="mainText">
|
||||
<subSubSection id="FBC334D8244F0A324D58E80C1E1376AD" type="materials_examined">
|
||||
<paragraph id="4EEC9227C14BE1F84CC8C47D79BE8EAD" box="[118,295,371,390]" pageId="78" pageNumber="79" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="69D84417B0E5416037E0AC269B29043B" box="[118,1122,399,644]" pageId="78" pageNumber="79" type="mainText">
|
||||
MADAGASCAR: Antananarivo, Mandraka, 18° 54' 46" S, 47° 53' 32" E, ca. 1200 m, 23.I.2006 (A. Erpenbach); Fianarantsoa, Parc National Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango, 23.83517 S, 46.96367 E, 940 m, 15.XI.2006 (B.L. Fisher et al.); Toamasina, F.C. Didy, 18.19833 S, 48.57833 E, 960 m, 16.-23.XII.1998 (H.J. Ratsirarson); Toamasina, Montagne d'Akirindro, 7.6 km 341° NNW Ambinanitelo, 15.28833 S, 49.54833 E, 600 m, rainforest, 17.-21.III.2003 (B.L. Fisher, C. Griswold et al.); Toliara,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken originalValue="Classée">Classee</normalizedToken>
|
||||
<normalizedToken id="E0BBFB55D7065830D7996F9145396F14" originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="32EA7582BA8BF921703B944D843117B9" originalValue="Classée">Classee</normalizedToken>
|
||||
d'Analavelona, 33.2 km 344° NNW Mahaboboka, 22.64333 S, 44.17167 E, 1300 m, 22.-26.II.2003 (B.L. Fisher, C. Griswold et al.); Toliara,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="CF0AF96393C9B2ADB5BD6226A52C3D86" originalValue="Rés">Res</normalizedToken>
|
||||
. Andohahela, 11 km NW Enakara, 24.56667 S, 46.83333 E, 800 m, montane rainforest, 16.-17.XI.1992 (B.L. Fisher); Toliara,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="108435BD6A2ECFE9886C9449D043187E" originalValue="Rés">Res</normalizedToken>
|
||||
. Andohahela, 10 km NW Enakara, 24.56667 S, 46.81667 E, 430 m, rainforest, 22.XI.1992 (B.L. Fisher).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,382 +1,381 @@
|
|||
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|
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">
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|
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<taxonomicName id="849B9EA52FC22388ADDFEFCB1B077142" ID-CoL="7C4VH" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">
|
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<pageBreakToken id="EFE90BFCA84BE0479E03402957FBC386" pageId="13" pageNumber="14" start="start">Tetramorium</pageBreakToken>
|
||||
tortuosum
|
||||
</taxonomicName>
|
||||
species group
|
||||
</paragraph>
|
||||
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|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,173,192]" pageId="13" pageNumber="14" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,202,502]" pageId="13" pageNumber="14" type="mainText">Eleven-segmented antennae; anterior clypeal margin medially impressed; frontal carinae well-developed and usually running to posterior head margin; anterior face of mesosoma weakly developed; margination between lateral and dorsal mesosoma generally well-developed; propodeal spines always long to extremely long, spinose, and acute (PSLI 28-72); propodeal lobes usually well-developed, triangular to elongate-triangular, generally short and acute, rarely strongly reduced to almost absent; petiolar node rectangular nodiform, anterodorsal and posterodorsal margins usually well-defined, anterior and posterior faces often parallel, node longer than wide in most species, broader than long in few species; postpetiole usually globular to subglobular; mandibles strongly sculptured in most species; head and mesosoma with distinct and predominantly longitudinally rugose sculpture; waist segments with distinctly rugose, rarely rugulose, sculpture; gaster unsculptured, smooth and shiny in many species, but sculpture present on the first gastral tergite in several species; in most species all dorsal surfaces of head, mesosoma, waist segments and gaster with abundant, long, standing hairs; sting appendage spatulate.</paragraph>
|
||||
<subSubSection id="871AAF4FC552E8A02D93C93143D882A2" type="diagnosis">
|
||||
<paragraph id="8CE2DFE25A62C746804AF5C9756713B7" box="[118,208,173,192]" pageId="13" pageNumber="14" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="B9BB797F9A09FD0FB7ADAABFD1D2E509" box="[118,1121,202,502]" pageId="13" pageNumber="14" type="mainText">Eleven-segmented antennae; anterior clypeal margin medially impressed; frontal carinae well-developed and usually running to posterior head margin; anterior face of mesosoma weakly developed; margination between lateral and dorsal mesosoma generally well-developed; propodeal spines always long to extremely long, spinose, and acute (PSLI 28-72); propodeal lobes usually well-developed, triangular to elongate-triangular, generally short and acute, rarely strongly reduced to almost absent; petiolar node rectangular nodiform, anterodorsal and posterodorsal margins usually well-defined, anterior and posterior faces often parallel, node longer than wide in most species, broader than long in few species; postpetiole usually globular to subglobular; mandibles strongly sculptured in most species; head and mesosoma with distinct and predominantly longitudinally rugose sculpture; waist segments with distinctly rugose, rarely rugulose, sculpture; gaster unsculptured, smooth and shiny in many species, but sculpture present on the first gastral tergite in several species; in most species all dorsal surfaces of head, mesosoma, waist segments and gaster with abundant, long, standing hairs; sting appendage spatulate.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,219,539,558]" pageId="13" pageNumber="14" type="mainText">Comments</paragraph>
|
||||
<paragraph box="[118,1123,567,1121]" pageId="13" pageNumber="14" type="mainText">
|
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|
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<paragraph id="1F8A2F402C540CB9411B10E76CCB6E9F" box="[118,219,539,558]" pageId="13" pageNumber="14" type="mainText">Comments</paragraph>
|
||||
<paragraph id="AAED0E4A673910F5C520389386312781" box="[118,1123,567,1121]" pageId="13" pageNumber="14" type="mainText">
|
||||
Within the Malagasy region, the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="E81E5C41CA5E6B582459F369DCA86637" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group is easily recognisable within the groups with 11-segmented antennae. The distinction from the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
<taxonomicName id="39D5296E7974B633D7A39355AFB93E7D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
group was explained in detail above. The
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="plesiarum">T. plesiarum</taxonomicName>
|
||||
<taxonomicName id="A8C90AF507389E68D56C1FE56F968A24" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="plesiarum">T. plesiarum</taxonomicName>
|
||||
group is characterised by its distinct and conspicuous antennal scrobes with well-defined margins all-around. By contrast, the scrobes are usually developed, but less conspicuous and without well-defined posterior and ventral margins in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="C263EA89F6FE2D10A793C7BAA311F50A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group. Due to the presence of distinct sculpture on both waist segments, the group cannot be mistaken for the
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bessonii">
|
||||
<taxonomicName id="08E56086F33F02AB444136F76F04D920" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bessonii">
|
||||
T. bessonii
|
||||
<authority>,</authority>
|
||||
<authority id="31C14204932D345EB42208BD6D17D789">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="marginatum">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="marginatum">
|
||||
<taxonomicName id="D90D0296FC58A21DD79DA869C9AD3CF0" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="marginatum">
|
||||
<taxonomicName id="69B767D130CD3EE638D8333E692BDC23" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="marginatum">
|
||||
T. marginatum
|
||||
<authority>,</authority>
|
||||
<authority id="1C15C99F8F5DA6BD86E5AC27E4842E7C">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="naganum">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="naganum">
|
||||
<taxonomicName id="91D06F5044512D7A1E188F595421F3DB" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="naganum">
|
||||
<taxonomicName id="CAAB5BBA71D935AAC6038C7F5FF92FA1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="naganum">
|
||||
T. naganum
|
||||
<authority>,</authority>
|
||||
<authority id="FC6561F744C7C9956483AF34E1AA94BB">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="schaufussii">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="schaufussii">
|
||||
<taxonomicName id="A72520B808F6B0506D7C0F7AAE0CFC92" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="schaufussii">
|
||||
<taxonomicName id="6C025168423C8A7F61E863C1CD36FA87" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="schaufussii">
|
||||
T. schaufussii
|
||||
<authority>,</authority>
|
||||
<authority id="6D3ED9B554837FB7E56AA6080F973F72">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="severini">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="severini">
|
||||
<taxonomicName id="0504BFC5844EE999CE00A49B32637622" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="severini">
|
||||
<taxonomicName id="9836FEFCFC09497400F77F450DD3BB41" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="severini">
|
||||
T. severini
|
||||
<authority>,</authority>
|
||||
<authority id="BBA81F7D3C6C7638CED397B46FB1E11C">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName genus="Tetamorium" pageId="13" pageNumber="14" rank="species" species="tsingy">T. tsingy,</taxonomicName>
|
||||
<taxonomicName id="9CD4948812B25C749D59FD2907E06E7F" genus="Tetamorium" pageId="13" pageNumber="14" rank="species" species="tsingy">T. tsingy,</taxonomicName>
|
||||
or
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="weitzeckeri">T. weitzeckeri</taxonomicName>
|
||||
<taxonomicName id="518BD56C9100B43A76F3E98E2003F4C5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="weitzeckeri">T. weitzeckeri</taxonomicName>
|
||||
groups, nor parts of the
|
||||
<taxonomicName genus="Tetamorium" pageId="13" pageNumber="14" rank="species" species="bonibony">T. bonibony</taxonomicName>
|
||||
<taxonomicName id="2DAA30AED3E000211AEC280CCFF55E69" genus="Tetamorium" pageId="13" pageNumber="14" rank="species" species="bonibony">T. bonibony</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
<taxonomicName id="804E05BB2FE97E17FA18A18AB7BD832C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
groups. The species of the
|
||||
<taxonomicName genus="Tetamorium" pageId="13" pageNumber="14" rank="species" species="bonibony">T. bonibony</taxonomicName>
|
||||
<taxonomicName id="728510E82AF0C7B6AD1901061E4E0EE4" genus="Tetamorium" pageId="13" pageNumber="14" rank="species" species="bonibony">T. bonibony</taxonomicName>
|
||||
group with sculptured waist segments have reticulate-rugose sculpture on the posterior head and anterior mesosoma which is not present in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="D94178BD44B45A5D0051656AD2394F8E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group. Most species of the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
<taxonomicName id="6435E7CBF6FF76772C249533D2FAA2E1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
group with sculptured waist segments usually have a weakly sculptured petiolar node which distinguishes them from the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="A0C4B064257761A55F55BAB457C27FE2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group species.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dysalum">Tetramorium dysalum</taxonomicName>
|
||||
<taxonomicName id="5AF8AF54C445217EC70E8A048F87A6D1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dysalum">Tetramorium dysalum</taxonomicName>
|
||||
has more sculpture, and could be misidentified with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="4BA3CA3770D074CBD2655306AE945A40" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
or
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="AE16D6579406CC0A09FC4CE63D105FA1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
on the basis of petiolar node shape, but the latter species have sculptured mandibles and sculpture on the first gastral tergite, whereas
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
<taxonomicName id="04C11BBA146E0CDFD6136E3F9D9F216D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
possesses smooth mandibles and lacks sculpture on the first gastral tergite. Moreover, the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ranarum">T. ranarum</taxonomicName>
|
||||
<taxonomicName id="E4A8F2777C9092ED4A023EFA31AC93BD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ranarum">T. ranarum</taxonomicName>
|
||||
group, despite sharing the rectangular nodiform shape of the petiolar node in some species, differs in several other important aspects, making it easily separable from the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="3AD913595D01CA9DBCB301A3E64F7488" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group. The sculpture on posterior head and mesosoma is usually reticulate-rugose in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ranarum">T. ranarum</taxonomicName>
|
||||
<taxonomicName id="A9F1AC0609A0E0C11D2A6BD0811DE953" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ranarum">T. ranarum</taxonomicName>
|
||||
group but longitudinally rugose in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="3E223BA86AA9C493217EC3C100604878" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group. Also, the propodeal spines are also much longer in the latter group than in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ranarum">T. ranarum</taxonomicName>
|
||||
<taxonomicName id="7E1648D1472243FA19861D5ABEA4975F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ranarum">T. ranarum</taxonomicName>
|
||||
group. In addition, the frontal carinae and antennal scrobes are often much weaker in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ranarum">T. ranarum</taxonomicName>
|
||||
<taxonomicName id="16C0CC45ADDFA4AE0BDE845D42C3EA6D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ranarum">T. ranarum</taxonomicName>
|
||||
group, and several species have one or even both waist segments unsculptured, whereas all Malagasy
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="7759AA21F9AE829017CB8B54222F6C94" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group species have both waist segments noticeably sculptured.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,1130,1290]" pageId="13" pageNumber="14" type="mainText">
|
||||
<paragraph id="57062E4C1955515A7A0DB9DF8EED11B8" box="[118,1122,1130,1290]" pageId="13" pageNumber="14" type="mainText">
|
||||
Most Malagasy
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="50C149473A4279862C270A47A8487A6D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group members appear to form a natural grouping except for
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="6388CC64F107B2C48BE8CE99AEBA2128" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon.</taxonomicName>
|
||||
<taxonomicName id="B9B6617A311F67F8E6DBC600351EAFC4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon.</taxonomicName>
|
||||
In these two the petiolar node has a shape with the anterodorsal margin situated higher than the posterodorsal and the dorsum tapers distinctly backwards posteriorly, which, as mentioned above, is also seen in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
<taxonomicName id="7BB180C0AFB554E6204903BE892D8BE9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
group. The true affinities of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="509C4E5BF2762FE8C74C9C1318BC7DC7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="208C11B1881E2256D687CED81375146B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
are unclear at the moment, and might be revealed by the use of molecular data in future studies. At present, however, we place them in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="469EC7287F974B7ADA88300F81CA14C4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group since they seem to fit best in this group until more data becomes available.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,1299,1402]" pageId="13" pageNumber="14" type="mainText">
|
||||
<paragraph id="CFC3C133A14791CD4C38F71978C7D0FB" box="[118,1121,1299,1402]" pageId="13" pageNumber="14" type="mainText">
|
||||
Our revision has revealed a remarkable 22 species within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="CEFD80202FB1210F9FFAA503394DF48F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group, which makes it the most species-rich
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
<taxonomicName id="A133FD5A284353F488BFB3BC2BB99576" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
species group in the Malagasy region. In order to facilitate the work with this comparatively high number of species, we have split the group into four species complexes on the basis of few important and conspicuous morphological key characters.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="key">
|
||||
<paragraph box="[118,680,1467,1487]" pageId="13" pageNumber="14" type="mainText">
|
||||
<subSubSection id="705E2A9346E3C8B2488A489735756DE2" type="key">
|
||||
<paragraph id="81C140E75CF4E4F6C67CEC9205C09385" box="[118,680,1467,1487]" pageId="13" pageNumber="14" type="mainText">
|
||||
Key to species of the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="BE4F8D39FD663D41630995452CC98AF6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
species group (workers)
|
||||
</paragraph>
|
||||
<paragraph box="[118,1123,1523,1539]" pageId="13" pageNumber="14" type="mainText">
|
||||
<paragraph id="A838619C363CA44AB08463C7319A9220" box="[118,1123,1523,1539]" pageId="13" pageNumber="14" type="mainText">
|
||||
1. In profile forecoxae completely covered with very distinct, strong, longitudinal rugae (
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="841E478CED11A40D06379473AB6EDBA5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
species complex) (Fig. 11).... 2
|
||||
</paragraph>
|
||||
<paragraph box="[175,1122,1546,1585]" pageId="13" pageNumber="14" type="mainText">In profile forecoxae without very distinct, strong, longitudinal rugae; usually unsculptured, smooth and shining, but often with traces of rugulae or punctate sculpture (Figs. 12, 13)..........................................................9</paragraph>
|
||||
<paragraph box="[118,1120,501,541]" pageId="14" pageNumber="15" type="mainText">
|
||||
<pageBreakToken pageId="14" pageNumber="15" start="start">2</pageBreakToken>
|
||||
<paragraph id="37A384DD762ACF525A8DBBC9260577BC" box="[175,1122,1546,1585]" pageId="13" pageNumber="14" type="mainText">In profile forecoxae without very distinct, strong, longitudinal rugae; usually unsculptured, smooth and shining, but often with traces of rugulae or punctate sculpture (Figs. 12, 13)..........................................................9</paragraph>
|
||||
<paragraph id="291757544778B668A168CEDB93484626" box="[118,1120,501,541]" pageId="14" pageNumber="15" type="mainText">
|
||||
<pageBreakToken id="E4692FCFC69206AFBB4065B61D0E870B" pageId="14" pageNumber="15" start="start">2</pageBreakToken>
|
||||
. Propodeal spines long to extremely long (PSLI 34-50, generally below 45); comparatively hairy species with numerous long standing hairs on the first gastral tergite (Fig. 14)............................................................ 3
|
||||
</paragraph>
|
||||
<paragraph box="[175,1123,547,587]" pageId="14" pageNumber="15" type="mainText">Propodeal spines always extremely long (PSLI 48-72; generally above 50); less hairy species with no or few scattered, long standing hairs (Figs. 15, 16)............................................................................. 7</paragraph>
|
||||
<paragraph box="[118,1121,834,851]" pageId="14" pageNumber="15" type="mainText">
|
||||
<paragraph id="486F3716F765AEA14E9AAE422C56F9B7" box="[175,1123,547,587]" pageId="14" pageNumber="15" type="mainText">Propodeal spines always extremely long (PSLI 48-72; generally above 50); less hairy species with no or few scattered, long standing hairs (Figs. 15, 16)............................................................................. 7</paragraph>
|
||||
<paragraph id="92BD1EF52AA4F45487B49CD0DFBF9FDC" box="[118,1121,834,851]" pageId="14" pageNumber="15" type="mainText">
|
||||
3. Petiolar node with posterodorsal margin situated higher than anterodorsal, dorsum convex (Fig. 17).............
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
<taxonomicName id="C03AE9C6B91986886952F2B52AD22350" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[176,1122,856,872]" pageId="14" pageNumber="15" type="mainText">Petiolar node with anterodorsal and posterodorsal margins at about the same height, dorsum flat to weakly convex (Fig. 18)...4</paragraph>
|
||||
<paragraph box="[118,1120,1256,1295]" pageId="14" pageNumber="15" type="mainText">
|
||||
<paragraph id="27BE8FD2B6F88E5163731A9C63C3C346" box="[176,1122,856,872]" pageId="14" pageNumber="15" type="mainText">Petiolar node with anterodorsal and posterodorsal margins at about the same height, dorsum flat to weakly convex (Fig. 18)...4</paragraph>
|
||||
<paragraph id="73696E778E7D0C6DC728C60D1C829E72" box="[118,1120,1256,1295]" pageId="14" pageNumber="15" type="mainText">
|
||||
4. Antennal scapes comparatively long (SI 89-92); petiolar node higher (LPeI 81-88); body colour dark brown to black (Figs. 19, 20)..................................................................................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nazgul">Tetramorium nazgul</taxonomicName>
|
||||
<taxonomicName id="2722BA665DA12BD5FD8F1EAC613DB938" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nazgul">Tetramorium nazgul</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1121,1300,1338]" pageId="14" pageNumber="15" type="mainText">
|
||||
<paragraph id="B8FE6A979A81FB36428EEA95825F1342" box="[175,1121,1300,1338]" pageId="14" pageNumber="15" type="mainText">
|
||||
Antennal scapes shorter than above (SI 79-85); petiolar node lower (LPeI 89-97); body colour orange to light brown (Figs. 21, 22)................................................................................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="marojejy">Tetramorium marojejy</taxonomicName>
|
||||
<taxonomicName id="9FDF103440B720CCB0657ED7F2C8EC8F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="marojejy">Tetramorium marojejy</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,118,158]" pageId="15" pageNumber="16" type="mainText">
|
||||
<pageBreakToken pageId="15" pageNumber="16" start="start">7</pageBreakToken>
|
||||
<paragraph id="704B8184C1FBA8A2E4F2C19070585414" box="[118,1122,118,158]" pageId="15" pageNumber="16" type="mainText">
|
||||
<pageBreakToken id="041D6BB9E22312971C2C8AFEB391C5BE" pageId="15" pageNumber="16" start="start">7</pageBreakToken>
|
||||
. First gastral tergite without any standing hairs, only with moderately dense appressed pubescence (Fig. 23)..................................................................................................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="latreillei">Tetramorium latreillei</taxonomicName>
|
||||
<taxonomicName id="3B57875C659F4C685B068FC75F70EBF2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="latreillei">Tetramorium latreillei</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1121,164,181]" pageId="15" pageNumber="16" type="mainText">First gastral tergite with several standing hairs and scarce pubescence (Fig. 24)..................................... 8</paragraph>
|
||||
<caption box="[118,1115,621,639]" pageId="15" pageNumber="16" type="caption">
|
||||
<paragraph box="[118,1115,621,639]" pageId="15" pageNumber="16" type="caption">
|
||||
<paragraph id="CB231C2F775759FF955FBA77ED74D069" box="[175,1121,164,181]" pageId="15" pageNumber="16" type="mainText">First gastral tergite with several standing hairs and scarce pubescence (Fig. 24)..................................... 8</paragraph>
|
||||
<caption id="DFB7C8D49E046006454F338AF683FA9A" box="[118,1115,621,639]" pageId="15" pageNumber="16" type="caption">
|
||||
<paragraph id="01E9A1031C72A927318D6DC609018206" box="[118,1115,621,639]" pageId="15" pageNumber="16" type="caption">
|
||||
FIGURES 23 & 24. 23.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="6EF681508A83F15692B68C4ED108B1D2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
(CASENT0101291) gaster in profile. 24.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="1D8277C34FCFBBD69DA3AE8EC3F3BFC7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
(CASENT0189240) gaster in profile.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,1124,673,713]" pageId="15" pageNumber="16" type="mainText">
|
||||
<paragraph id="9FE4D80CC176BD837A540A1E91F8996B" box="[118,1124,673,713]" pageId="15" pageNumber="16" type="mainText">
|
||||
8. Leading edges of antennal scapes with appressed hairs; mesosoma with just one to two pairs of standing hairs on the pronotal dorsum (Figs. 25, 26)..................................................................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sabatra">Tetramorium sabatra</taxonomicName>
|
||||
<taxonomicName id="92F3016910AD4E348218E999B0FDE225" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sabatra">Tetramorium sabatra</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1122,719,759]" pageId="15" pageNumber="16" type="mainText">
|
||||
<paragraph id="732505F5E9D57A52C986E5FB8925007D" box="[175,1122,719,759]" pageId="15" pageNumber="16" type="mainText">
|
||||
Leading edges of antennal scapes with subdecumbent to suberect hairs; mesosoma with 7 to 14 pairs of standing hairs throughout the whole mesosomal dorsum (Figs. 27, 28)...............................................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
<taxonomicName id="DECCBF39F8437509B0FE381C17C4F633" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
</paragraph>
|
||||
<caption box="[118,1122,983,1049]" pageId="15" pageNumber="16" type="caption">
|
||||
<paragraph box="[118,1122,983,1049]" pageId="15" pageNumber="16" type="caption">
|
||||
<caption id="F99B03182CA0385B7423B2995BA32248" box="[118,1122,983,1049]" pageId="15" pageNumber="16" type="caption">
|
||||
<paragraph id="BC8293E6D10C65083D3CB2F6BA5B5D9C" box="[118,1122,983,1049]" pageId="15" pageNumber="16" type="caption">
|
||||
FIGURES 25-28. 25.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="153B1907975D73AF9EE84941B7E4AAF8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
(CASENT0048680) antennal scape in frontal view. 26.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="35E64BAF71B06C4B1F89F4243887B918" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
(CASENT0189241) mesosoma in profile. 27.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="13D7A6CA1E6A7F271FEFEB3A723367A0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
(CASENT0121244) antennal scape in frontal view. 28.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="17E35B3157FB1EE9CB43CE19BFABB52C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
(CASENT0121244) mesosoma in profile.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,1121,1083,1100]" pageId="15" pageNumber="16" type="mainText">9. First gastral tergite with either reticulate-punctate or costulate sculpture (Figs. 29, 30)..............................10</paragraph>
|
||||
<paragraph box="[175,1122,1106,1447]" pageId="15" pageNumber="16" type="mainText">
|
||||
<paragraph id="094DEE7CFBBB6F9F902BD66AC8134276" box="[118,1121,1083,1100]" pageId="15" pageNumber="16" type="mainText">9. First gastral tergite with either reticulate-punctate or costulate sculpture (Figs. 29, 30)..............................10</paragraph>
|
||||
<paragraph id="9F5A283CBE8FBDE3A9D221B873E1BE75" box="[175,1122,1106,1447]" pageId="15" pageNumber="16" type="mainText">
|
||||
First gastral tergite unsculptured, smooth, and shining (
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="43137E1434AD564A4D4992F94C84D860" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
species complex) (Fig. 31)..........................16 first
|
||||
</paragraph>
|
||||
<caption box="[118,1085,1429,1471]" pageId="15" pageNumber="16" type="caption">
|
||||
<paragraph box="[118,1085,1429,1471]" pageId="15" pageNumber="16" type="caption">
|
||||
<caption id="0008C0DF0CAB44A44C0356C36F755498" box="[118,1085,1429,1471]" pageId="15" pageNumber="16" type="caption">
|
||||
<paragraph id="B823477D6D3BA27FBCBDEE8DE636BFAA" box="[118,1085,1429,1471]" pageId="15" pageNumber="16" type="caption">
|
||||
FIGURES 29-31. 29.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="0D4BC3196B13796656722C1D337410BA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
(CASENT0046064) first gastral tergite in dorsal view. 30.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
<taxonomicName id="F790163C7585788AF30D7058E8182516" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
(CASENT0247161) gastral tergite in dorsal view. 31.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="9D3B95D05740ABE5C68CF67736487800" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
(CASENT0280850) first gastral tergite in dorsal view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,141,1506,1522]" pageId="15" pageNumber="16" type="mainText">10.</paragraph>
|
||||
<paragraph box="[175,1119,1506,1545]" pageId="15" pageNumber="16" type="mainText">
|
||||
<paragraph id="0764338371A250E6F8995DBC1709407C" box="[118,141,1506,1522]" pageId="15" pageNumber="16" type="mainText">10.</paragraph>
|
||||
<paragraph id="116A7E332EB09DA9FE80483D82F02031" box="[175,1119,1506,1545]" pageId="15" pageNumber="16" type="mainText">
|
||||
First gastral tergite with basigastral costulae (
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="12275445AA32C917A19DB8270D7A60FA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
species complex) (Fig. 32)................................... 11 First gastral tergite differently sculptured (
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="94A389974FFB22A187B85169BB75C998" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
species complex) (Fig. 33)......................................14
|
||||
</paragraph>
|
||||
<caption box="[118,1122,526,568]" pageId="16" pageNumber="17" type="caption">
|
||||
<paragraph box="[118,1122,526,568]" pageId="16" pageNumber="17" type="caption">
|
||||
<pageBreakToken pageId="16" pageNumber="17" start="start">FIGURES</pageBreakToken>
|
||||
<caption id="3EFAE40BE8C25E7159AF5A01A18A15BB" box="[118,1122,526,568]" pageId="16" pageNumber="17" type="caption">
|
||||
<paragraph id="A7B1325F936CA483E3F1D68C008849DA" box="[118,1122,526,568]" pageId="16" pageNumber="17" type="caption">
|
||||
<pageBreakToken id="2E6FA2709A95E861662C32D0EBB91727" pageId="16" pageNumber="17" start="start">FIGURES</pageBreakToken>
|
||||
32 & 33. 32.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
<taxonomicName id="564CD7495731E85A120A8DB78D1E39E5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
(CASENT0247161) first gastral tergite in dorsal view. 33.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="73116C5AB606D3A019005A60763A2C55" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
(CASENT0046064) first gastral tergite in dorsal view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,1124,602,642]" pageId="16" pageNumber="17" type="mainText">11. In profile view, petiolar node with anterodorsal and posterodorsal margins at about the same height, and the dorsum flat to weakly convex (Fig. 34)...............................................................................12</paragraph>
|
||||
<paragraph box="[175,1122,648,688]" pageId="16" pageNumber="17" type="mainText">In profile view, petiolar node with the posterodorsal margin situated higher than the anterodorsal margin, and the dorsum convex (Fig. 35)........................................................................................13</paragraph>
|
||||
<caption box="[118,1122,1056,1098]" pageId="16" pageNumber="17" type="caption">
|
||||
<paragraph box="[118,1122,1056,1098]" pageId="16" pageNumber="17" type="caption">
|
||||
<paragraph id="451C6A3A012407A8189D2862AB708246" box="[118,1124,602,642]" pageId="16" pageNumber="17" type="mainText">11. In profile view, petiolar node with anterodorsal and posterodorsal margins at about the same height, and the dorsum flat to weakly convex (Fig. 34)...............................................................................12</paragraph>
|
||||
<paragraph id="D2746AEC70866E2645D3548EAED7D557" box="[175,1122,648,688]" pageId="16" pageNumber="17" type="mainText">In profile view, petiolar node with the posterodorsal margin situated higher than the anterodorsal margin, and the dorsum convex (Fig. 35)........................................................................................13</paragraph>
|
||||
<caption id="5B531A12B823682A8269247008463A11" box="[118,1122,1056,1098]" pageId="16" pageNumber="17" type="caption">
|
||||
<paragraph id="65B9FCED0F18C6CEC6829BAB24452007" box="[118,1122,1056,1098]" pageId="16" pageNumber="17" type="caption">
|
||||
FIGURES 34 & 35. 34.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
<taxonomicName id="FE4279BA6EE7EA88F75BD3A2FABF70E4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
(CASENT0045755) petiolar node in profile. 35.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="3B4681794D8A7ABCE85EF39644701B7A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
(CASENT0043554) petiolar node in profile.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,1122,1133,1172]" pageId="16" pageNumber="17" type="mainText">
|
||||
<paragraph id="3EC07358C17E66C25AD5091BF1FDAD63" box="[118,1122,1133,1172]" pageId="16" pageNumber="17" type="mainText">
|
||||
12. Eyes comparatively moderate to large (OI 22-24); propodeal spines comparatively long (PSLI 41-45) (Figs. 36, 37).............................................................................................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aherni">Tetramorium aherni</taxonomicName>
|
||||
<taxonomicName id="88012DA4A7FFB3A0C6AE1ED2671877BD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aherni">Tetramorium aherni</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1122,1179,1218]" pageId="16" pageNumber="17" type="mainText">
|
||||
<paragraph id="04E3FE001019AE56961908C1A8739EF5" box="[175,1122,1179,1218]" pageId="16" pageNumber="17" type="mainText">
|
||||
Eyes comparatively small (OI 16-17); propodeal spines comparatively short (PSLI 27-29) (Figs. 38, 39)..................................................................................................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="singletonae">Tetramorium singletonae</taxonomicName>
|
||||
<taxonomicName id="EA4ACD13413AD97E2314848E55FA242D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="singletonae">Tetramorium singletonae</taxonomicName>
|
||||
</paragraph>
|
||||
<caption box="[118,1123,1416,1457]" pageId="16" pageNumber="17" type="caption">
|
||||
<paragraph box="[118,1123,1416,1457]" pageId="16" pageNumber="17" type="caption">
|
||||
<caption id="4FD910345993E2D4B08D4F91EF2EEEF7" box="[118,1123,1416,1457]" pageId="16" pageNumber="17" type="caption">
|
||||
<paragraph id="0D5B4BB93C8B04A73CAD20E6942BC1D1" box="[118,1123,1416,1457]" pageId="16" pageNumber="17" type="caption">
|
||||
FIGURES 36-39. 36.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
<taxonomicName id="3D4DE9AAB8D28816A3366B497D0E84E9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
(CASENT0045755) head in full-face view. 37.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
<taxonomicName id="027D9CFAFF9BB1F7C3A1E5D8AD8B9C89" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
(CASENT0045755) mesosoma in profile. 38.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
<taxonomicName id="DD212B49EC88E58D02308EEFA11C04AB" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
(CASENT0247161) head in full-face view. 39.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
<taxonomicName id="5D4ABE59E578AA60511FDBDA58F52FA4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
(CASENT0247161) mesosoma in profile.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,1121,1492,1531]" pageId="16" pageNumber="17" type="mainText">
|
||||
<paragraph id="0D16E5CC46667AEC64DD2D6F794E1EFA" box="[118,1121,1492,1531]" pageId="16" pageNumber="17" type="mainText">
|
||||
13. Head distinctly longer than wide (CI 92-95); propodeal spines extremely long but comparatively shorter than below (PSLI 38-48); postpetiole relatively higher (LPpI 68-72) and broader (DPpI 119-129) (Figs. 40, 41)..........
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="noeli">Tetramorium noeli</taxonomicName>
|
||||
<taxonomicName id="477C3CFD09BFD508D6C67C6FC6B68666" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="noeli">Tetramorium noeli</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1123,1538,1577]" pageId="16" pageNumber="17" type="mainText">
|
||||
<paragraph id="E9A674B0666C8C97A538289E3C3903A6" box="[175,1123,1538,1577]" pageId="16" pageNumber="17" type="mainText">
|
||||
Head weakly longer than wide (CI 97-98); propodeal spines extremely long but comparatively longer than above (PSLI 60-65); postpetiole relatively lower (LPpI 77-81) and narrower (DPpI 104-112) (Figs. 42, 43)....
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ambanizana">Tetramorium ambanizana</taxonomicName>
|
||||
<taxonomicName id="EB7BC6260C0D98E9717F5BA9319DC99A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ambanizana">Tetramorium ambanizana</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1123,334,373]" pageId="17" pageNumber="18" type="mainText">
|
||||
<pageBreakToken pageId="17" pageNumber="18" start="start">14</pageBreakToken>
|
||||
<paragraph id="301009B7D794D679919BA364E620D76C" box="[118,1123,334,373]" pageId="17" pageNumber="18" type="mainText">
|
||||
<pageBreakToken id="DE220C09390908C4F4D0BB63E76386AC" pageId="17" pageNumber="18" start="start">14</pageBreakToken>
|
||||
. Petiolar node in dorsal view distinctly longer than wide (DPeI 79-85), in profile anterodorsal and posterodorsal margins at about same height; first gastral tergite completely covered with distinct reticulate-punctate sculpture (Figs. 44, 45)....
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="8336488F1709F67B4340D57D398C985F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1122,378,438]" pageId="17" pageNumber="18" type="mainText">Petiolar node in dorsal view distinctly wider than long (DPeI111-137); in profile anterodorsal margin situated higher than posterodorsal margin and dorsum tapering backwards; first gastral tergite never completely covered with distinct reticulate-punctate sculpture, generally with superficial reticulate-punctate sculpture restricted to the basal first half of the tergite (Figs. 46, 47).........15</paragraph>
|
||||
<paragraph box="[118,1122,780,819]" pageId="17" pageNumber="18" type="mainText">
|
||||
<paragraph id="2E2841FF0ABCB3FD0CF59C2EC1EDDD18" box="[175,1122,378,438]" pageId="17" pageNumber="18" type="mainText">Petiolar node in dorsal view distinctly wider than long (DPeI111-137); in profile anterodorsal margin situated higher than posterodorsal margin and dorsum tapering backwards; first gastral tergite never completely covered with distinct reticulate-punctate sculpture, generally with superficial reticulate-punctate sculpture restricted to the basal first half of the tergite (Figs. 46, 47).........15</paragraph>
|
||||
<paragraph id="C443F4A593DBA72700F7001E04B6D3FB" box="[118,1122,780,819]" pageId="17" pageNumber="18" type="mainText">
|
||||
15. Propodeal spines very long (PSLI 37-44); petiolar node relatively lower (LPeI 63-73) and narrower (DPeI 111-118); petiolar dorsum strongly rugose (Fig. 48)........................................................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
<taxonomicName id="642DE310A0BA9E20BEE930821BBF9389" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
ple ganon
|
||||
</paragraph>
|
||||
<paragraph box="[175,1124,824,862]" pageId="17" pageNumber="18" type="mainText">
|
||||
<paragraph id="57EC3C78F0BEC94F49550ADA9D630B96" box="[175,1124,824,862]" pageId="17" pageNumber="18" type="mainText">
|
||||
Propodeal spines long, but shorter than above (PSLI 27-37, usually below 34); petiolar node relatively higher (LPeI 54-66) and broader (DPeI 126-137); petiolar dorsum weakly rugose (Fig. 49)..........................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra</taxonomicName>
|
||||
<taxonomicName id="BD8EA1A88100D597C74DA0A979FD63AF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1120,1186,1203]" pageId="17" pageNumber="18" type="mainText">
|
||||
<paragraph id="53F716ECB2AA5247CE22188C0949EB5B" box="[118,1120,1186,1203]" pageId="17" pageNumber="18" type="mainText">
|
||||
16. Posterodorsal corner of petiolar node strongly protruding posteriorly (Fig. 50)..................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="andohahela">Tetramorium andohahela</taxonomicName>
|
||||
<taxonomicName id="2275DF27A8AC2281745ECA4A4AAF9370" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="andohahela">Tetramorium andohahela</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1121,1208,1224]" pageId="17" pageNumber="18" type="mainText">Posterodorsal corner of petiolar node not protruding posteriorly (Fig. 51).........................................17</paragraph>
|
||||
<paragraph box="[118,1122,118,158]" pageId="18" pageNumber="19" type="mainText">
|
||||
<pageBreakToken pageId="18" pageNumber="19" start="start">17</pageBreakToken>
|
||||
<paragraph id="0458EF013913893A676BA5FDF408BECB" box="[175,1121,1208,1224]" pageId="17" pageNumber="18" type="mainText">Posterodorsal corner of petiolar node not protruding posteriorly (Fig. 51).........................................17</paragraph>
|
||||
<paragraph id="E0DF0C87135DBAB737957CA5F2C22CA4" box="[118,1122,118,158]" pageId="18" pageNumber="19" type="mainText">
|
||||
<pageBreakToken id="D107D5394731E25379E5192FCB6B7E7E" pageId="18" pageNumber="19" start="start">17</pageBreakToken>
|
||||
. Eyes very small (OI 15-16); propodeal spines long (PSLI 28-33); anterodorsal margin of petiolar node situated slightly higher than posterodorsal margin; body orange to light reddish brown in colour (Fig. 52)..................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
<taxonomicName id="1DD810E00457B55D701CC951D85CFF07" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1121,164,181]" pageId="18" pageNumber="19" type="mainText">Character combination never as above; if eyes small, then propodeal spines very long to extremely long (Figs. 53, 54)....18</paragraph>
|
||||
<paragraph box="[118,1122,458,497]" pageId="18" pageNumber="19" type="mainText">
|
||||
<paragraph id="2F0C19A8F0AFDD8A2ACCBAAED01D21EC" box="[175,1121,164,181]" pageId="18" pageNumber="19" type="mainText">Character combination never as above; if eyes small, then propodeal spines very long to extremely long (Figs. 53, 54)....18</paragraph>
|
||||
<paragraph id="1237D737EAA74DE677485520117EE187" box="[118,1122,458,497]" pageId="18" pageNumber="19" type="mainText">
|
||||
18. Petiolar node in profile comparatively high, between 1.3 to 1.6 times higher than long (LPeI 64-74); propodeal lobes strongly reduced and inconspicuous (Fig. 55)......................................................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
<taxonomicName id="508E20B8466E93349CDA635E06618245" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1122,504,543]" pageId="18" pageNumber="19" type="mainText">Petiolar node in profile less high than above, between 1.1 times longer than high to 1.3 times higher than long (LPeI 76-105); propodeal lobes variably developed, usually conspicuous (Figs. 56, 57)..........................................19</paragraph>
|
||||
<paragraph box="[118,1120,821,837]" pageId="18" pageNumber="19" type="mainText">19. Propodeal spines extremely long (PSLI 58-64) (Figs. 58, 59)..................................................20</paragraph>
|
||||
<paragraph box="[175,1121,844,860]" pageId="18" pageNumber="19" type="mainText">Propodeal spines long to very long, but always significantly much shorter than above (PSLI 29-45) (Figs. 60, 61)........21</paragraph>
|
||||
<paragraph box="[118,1120,1192,1208]" pageId="18" pageNumber="19" type="mainText">
|
||||
<paragraph id="E3B01A9AA7CD4AB00EF3E36C4F086366" box="[175,1122,504,543]" pageId="18" pageNumber="19" type="mainText">Petiolar node in profile less high than above, between 1.1 times longer than high to 1.3 times higher than long (LPeI 76-105); propodeal lobes variably developed, usually conspicuous (Figs. 56, 57)..........................................19</paragraph>
|
||||
<paragraph id="52059B110AFC023ED9611DDC5F323D41" box="[118,1120,821,837]" pageId="18" pageNumber="19" type="mainText">19. Propodeal spines extremely long (PSLI 58-64) (Figs. 58, 59)..................................................20</paragraph>
|
||||
<paragraph id="22245D7986BEC86E846D54515B949502" box="[175,1121,844,860]" pageId="18" pageNumber="19" type="mainText">Propodeal spines long to very long, but always significantly much shorter than above (PSLI 29-45) (Figs. 60, 61)........21</paragraph>
|
||||
<paragraph id="21655E3FB6D5D21ADBC1711231B64D2A" box="[118,1120,1192,1208]" pageId="18" pageNumber="19" type="mainText">
|
||||
20. Mandibles unsculptured, smooth, and shining; body colour yellow (Figs. 62, 63).......................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elf">Tetramorium elf</taxonomicName>
|
||||
<taxonomicName id="4D9542CFBC33FF6537C253E3E9393C14" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elf">Tetramorium elf</taxonomicName>
|
||||
</paragraph>
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<paragraph box="[176,1121,1215,1231]" pageId="18" pageNumber="19" type="mainText">
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<paragraph id="8B0473F205A6A16CDF788EB2C08B3D36" box="[176,1121,1215,1231]" pageId="18" pageNumber="19" type="mainText">
|
||||
Mandibles noticeably longitudinally sculptured; body colour very dark brown to black (Fig. 64, 65)...
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="isoelectrum">Tetramorium isoelectrum</taxonomicName>
|
||||
<taxonomicName id="6ADE026156A5E21F4948CE76AB0B37E1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="isoelectrum">Tetramorium isoelectrum</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,1518,1535]" pageId="18" pageNumber="19" type="mainText">
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||||
<paragraph id="CD3EF153CE1B570F1F81E96B4DBEF7AE" box="[118,1122,1518,1535]" pageId="18" pageNumber="19" type="mainText">
|
||||
21. Petiolar node in dorsal view relatively long, always more than 1.3 times longer than wide (DPeI 72-76) (Fig. 66)..
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ala">Tetramorium ala</taxonomicName>
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<taxonomicName id="DCE748B8345E12FA171E9A137F88A054" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ala">Tetramorium ala</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1123,1541,1580]" pageId="18" pageNumber="19" type="mainText">Petiolar node in dorsal view relatively wider, always between 1.0 to approximately 1.2 times longer than wide (DPeI 82-98) (Figs. 67, 68)........................................................................................22</paragraph>
|
||||
<caption box="[118,1122,353,394]" pageId="19" pageNumber="20" type="caption">
|
||||
<paragraph box="[118,1122,353,394]" pageId="19" pageNumber="20" type="caption">
|
||||
<pageBreakToken pageId="19" pageNumber="20" start="start">FIGURES</pageBreakToken>
|
||||
<paragraph id="8EA046BCABE22341B3B5108B905E48D1" box="[175,1123,1541,1580]" pageId="18" pageNumber="19" type="mainText">Petiolar node in dorsal view relatively wider, always between 1.0 to approximately 1.2 times longer than wide (DPeI 82-98) (Figs. 67, 68)........................................................................................22</paragraph>
|
||||
<caption id="76A44498E06FEB87549134F2C1D4A895" box="[118,1122,353,394]" pageId="19" pageNumber="20" type="caption">
|
||||
<paragraph id="75C4DC09F4B6B2691035670185915B1F" box="[118,1122,353,394]" pageId="19" pageNumber="20" type="caption">
|
||||
<pageBreakToken id="E994B677596C40FC800C43DD2D06B6B4" pageId="19" pageNumber="20" start="start">FIGURES</pageBreakToken>
|
||||
66-68. 66. T. ala (CASENT0038473) petiolar node in dorsal view. 67.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="8354E6353C2A99BA3AF19A0B3E2B0593" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
(CASENT0102395) petiolar node in dorsal view. 68.
|
||||
<taxonomicName genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="C0048A3C73CEF04732474007ED0BF820" genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="nify">T. nify</taxonomicName>
|
||||
(CASENT0163155) petiolar node in dorsal view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,1122,429,445]" pageId="19" pageNumber="20" type="mainText">
|
||||
<paragraph id="E6BD26E367C4DBC018B7538D338BF497" box="[118,1122,429,445]" pageId="19" pageNumber="20" type="mainText">
|
||||
22. Eyes relatively small (OI 16-17); propodeal spines relatively short (PSLI 29-30) (Fig. 69)..............
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nify">Tetramorium nify</taxonomicName>
|
||||
<taxonomicName id="8696E84C4D514159BBE9B0DE8D3591F6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nify">Tetramorium nify</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1121,452,468]" pageId="19" pageNumber="20" type="mainText">Eyes always much larger (OI 19-25); propodeal spines relatively longer (PSLI 32-45) (Figs. 70, 71)..................23</paragraph>
|
||||
<caption box="[118,1122,708,750]" pageId="19" pageNumber="20" type="caption">
|
||||
<paragraph box="[118,1122,708,750]" pageId="19" pageNumber="20" type="caption">
|
||||
<paragraph id="D5CA847418D70F315CFB92BF714FD077" box="[175,1121,452,468]" pageId="19" pageNumber="20" type="mainText">Eyes always much larger (OI 19-25); propodeal spines relatively longer (PSLI 32-45) (Figs. 70, 71)..................23</paragraph>
|
||||
<caption id="9197F8ECC9B82E8A17659A2BBD1083F1" box="[118,1122,708,750]" pageId="19" pageNumber="20" type="caption">
|
||||
<paragraph id="744A37D6ED1C16FD9536E575C219638E" box="[118,1122,708,750]" pageId="19" pageNumber="20" type="caption">
|
||||
FIGURES 69-71. 69.
|
||||
<taxonomicName genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="F86D973D44C4859142C7E79B7C73DF47" genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="nify">T. nify</taxonomicName>
|
||||
(CASENT0163155) body in profile. 70.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="53922605F30688FE9EFFD990E55048ED" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
(CASENT0102395) body in profile. 71.
|
||||
<taxonomicName genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
<taxonomicName id="E32C389B536FC139D5096B1156AA0EC6" genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
(CASENT0247162) body in profile.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,1121,785,824]" pageId="19" pageNumber="20" type="mainText">
|
||||
<paragraph id="99C6BB0D607188FFB73BD5B198269F1B" box="[118,1121,785,824]" pageId="19" pageNumber="20" type="mainText">
|
||||
23. Petiolar node with weakly defined and fairly rounded anterodorsal and posterodorsal margins; postpetiole relatively longer, longer than wide to slightly wider than long (DPpI 95-101) (Fig. 72).............................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="voasary">Tetramorium voasary</taxonomicName>
|
||||
<taxonomicName id="0D1D8BA1CFD8317A6795E86ED6887970" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="voasary">Tetramorium voasary</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[175,1122,831,870]" pageId="19" pageNumber="20" type="mainText">
|
||||
<paragraph id="739A1F66BF86552A754AB5FA172F6692" box="[175,1122,831,870]" pageId="19" pageNumber="20" type="mainText">
|
||||
Petiolar node with moderately to sharply defined anterodorsal and posterodorsal margins; postpetiole relatively wider, between 1.0 to 1.4 times wider than long (DPpI 103-136) (Fig. 73, 74)...................................
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="andrei">Tetramorium andrei</taxonomicName>
|
||||
<taxonomicName id="BFA4AEBC796759F22FA789A194D98AA0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="andrei">Tetramorium andrei</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
|
|
|
|||
|
|
@ -1,114 +1,113 @@
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Tetramorium andohahela
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<authority>Hita</authority>
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Garcia & Fisher
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<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
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<taxonomicNameLabel id="8BBCE40FEF0A5D9EB0435A944E1D2B5C" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph box="[118,345,840,859]" pageId="22" pageNumber="23" type="mainText">(Figs. 50, 78, 79, 80, 141)</paragraph>
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<paragraph id="84926C3A1B76A95D833A15F6B95195E0" box="[118,345,840,859]" pageId="22" pageNumber="23" type="mainText">(Figs. 50, 78, 79, 80, 141)</paragraph>
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<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1122,897,1141]" pageId="22" pageNumber="23" type="mainText">Holotype worker, MADAGASCAR, Toliara, Parc National d'Andohahela, Col du Sedro, 3.8 km 113° ESE Mahamavo, 37.6 km 341° NNW Tolagnaro, 24.76389 S, 46.75167 E, 900 m, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF05010, 21.-25.I.2002 (B.L. Fisher, C. Griswold, et al.) (CASC: CASENT0454449). Paratypes, 25 workers with same data as holotype (BMNH: CASENT0484441; CASC: CASENT0454555; CASENT0484356; CASENT0484390; CASENT0484395; CASENT0484407; CASENT0484417; CASENT0484430; CASENT0484433; CASENT0484434; CASENT0484435; CASENT0484472; CASENT0484482; CASENT0484487; CASENT0484488; CASENT0484489; CASENT0484494; CASENT0484499; CASENT0484508; CASENT0484520; CASENT0484524; MCZ: CASENT0484469; MHNG: CASENT0484510; NHMB: CASENT0484555).</paragraph>
|
||||
<subSubSection id="8D7F29AE98611D5852DF68B0658A44DA" type="materials_examined">
|
||||
<paragraph id="D2A66E6378082D8DB3301951E2A8D1AE" box="[118,1122,897,1141]" pageId="22" pageNumber="23" type="mainText">Holotype worker, MADAGASCAR, Toliara, Parc National d'Andohahela, Col du Sedro, 3.8 km 113° ESE Mahamavo, 37.6 km 341° NNW Tolagnaro, 24.76389 S, 46.75167 E, 900 m, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF05010, 21.-25.I.2002 (B.L. Fisher, C. Griswold, et al.) (CASC: CASENT0454449). Paratypes, 25 workers with same data as holotype (BMNH: CASENT0484441; CASC: CASENT0454555; CASENT0484356; CASENT0484390; CASENT0484395; CASENT0484407; CASENT0484417; CASENT0484430; CASENT0484433; CASENT0484434; CASENT0484435; CASENT0484472; CASENT0484482; CASENT0484487; CASENT0484488; CASENT0484489; CASENT0484494; CASENT0484499; CASENT0484508; CASENT0484520; CASENT0484524; MCZ: CASENT0484469; MHNG: CASENT0484510; NHMB: CASENT0484555).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,1178,1196]" pageId="22" pageNumber="23" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,1206,1253]" pageId="22" pageNumber="23" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andohahela">Tetramorium andohahela</taxonomicName>
|
||||
<subSubSection id="C9DBF0073207D738BFBEA272FF4CD283" type="diagnosis">
|
||||
<paragraph id="D7C07AC42DA73113758CC5E205BBFD0F" box="[118,208,1178,1196]" pageId="22" pageNumber="23" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="3BA0A5899E1AF507A12E23762B60D31D" box="[118,1121,1206,1253]" pageId="22" pageNumber="23" type="mainText">
|
||||
<taxonomicName id="CD746327910876C1A983A6F22B97A697" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andohahela">Tetramorium andohahela</taxonomicName>
|
||||
can be straightforwardly identified within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="786339533A032B356A20FD9CB02DB000" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
species complex due to the very characteristic petiolar node shape with the posterodorsal corner strongly protruding posteriorly.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,1290,1309]" pageId="22" pageNumber="23" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,1318,1478]" pageId="22" pageNumber="23" type="mainText">HL 0.94-1.10 (1.02); HW 0.95-1.15 (1.05); SL 0.75-0.90 (0.83); EL 0.20-0.25 (0.22); PH 0.49-0.60 (0.53); PW 0.71-0.85 (0.78); WL 1.28-1.53 (1.40); PSL 0.44-0.58 (0.50); PTL 0.36-0.46 (0.42); PTH 0.37-0.48 (0.43); PTW 0.28-0.35 (0.32); PPL 0.32-0.36 (0.34); PPH 0.37-0.48 (0.43); PPW 0.40-0.48 (0.44); CI 101-106 (103); SI 76-82 (79); OI 20-22 (21); DMI 54-58 (56); LMI 37-40 (38); PSLI 45-53 (49); PeNI 38-43 (41); LPeI 93-105 (99); DPeI 73-80 (76); PpNI 55-59 (56); LPpI 76-88 (80); DPpI 121-137 (129); PPI 133-145 (139) (14 measured).</paragraph>
|
||||
<paragraph box="[118,1121,1487,1591][118,1121,118,672]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="mainText">
|
||||
<subSubSection id="5C3041246DE7AD6F62D3A1C1625677C0" type="description">
|
||||
<paragraph id="30AC26F80ABF186E6AFE10C9BA68815E" box="[118,226,1290,1309]" pageId="22" pageNumber="23" type="mainText">Description</paragraph>
|
||||
<paragraph id="86F815EF9E4028826EA7CB39992B27A4" box="[118,1121,1318,1478]" pageId="22" pageNumber="23" type="mainText">HL 0.94-1.10 (1.02); HW 0.95-1.15 (1.05); SL 0.75-0.90 (0.83); EL 0.20-0.25 (0.22); PH 0.49-0.60 (0.53); PW 0.71-0.85 (0.78); WL 1.28-1.53 (1.40); PSL 0.44-0.58 (0.50); PTL 0.36-0.46 (0.42); PTH 0.37-0.48 (0.43); PTW 0.28-0.35 (0.32); PPL 0.32-0.36 (0.34); PPH 0.37-0.48 (0.43); PPW 0.40-0.48 (0.44); CI 101-106 (103); SI 76-82 (79); OI 20-22 (21); DMI 54-58 (56); LMI 37-40 (38); PSLI 45-53 (49); PeNI 38-43 (41); LPeI 93-105 (99); DPeI 73-80 (76); PpNI 55-59 (56); LPpI 76-88 (80); DPpI 121-137 (129); PPI 133-145 (139) (14 measured).</paragraph>
|
||||
<paragraph id="1F9D964BD07AFAEBF7233613FD912FB0" box="[118,1121,1487,1591][118,1121,118,672]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="mainText">
|
||||
Head weakly to distinctly wider than long (CI 101-106); posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes short to moderate, not reaching posterior head margin (SI 76-82). Eyes of
|
||||
<pageBreakToken pageId="23" pageNumber="24" start="start">moderate</pageBreakToken>
|
||||
<pageBreakToken id="26F859A37D0990743EE3748F1C533E4A" pageId="23" pageNumber="24" start="start">moderate</pageBreakToken>
|
||||
size (OI 20-22). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-40). Propodeal spines very long to extremely long, spinose and acute (PSLI 45-53); propodeal lobes well-developed, elongate-triangular, and acute. Petiolar node in profile rectangular nodiform with well-defined angles, weakly higher than long to weakly higher than long (LPeI 93-105), anterior and posterior faces almost parallel, anterior face much lower than posterior face, posterodorsal margin situated much higher than anterodorsal, posterior face weakly concave, posterodorsal corner strongly protruding posteriorly; node in dorsal view between 1.2 to 1.4 times longer than wide (DPeI 73-80). Postpetiole in profile subglobular, approximately 1.1 to 1.3 times higher than long (LPpI 76-88); in dorsal view around 1.2 to 1.4 times wider than long (DPpI 121-137). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view approximately 1.3 to 1.5 times wider than petiolar node (PPI 133-145). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose/rugulose, with four to seven rugae/rugulae, median ruga/rugula often developed and conspicuous, remaining rugae/rugulae often short or irregularly arranged; cephalic dorsum between frontal carinae with eight to ten longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted or with cross-meshes; lateral and ventral head mainly reticulate-rugose, less longitudinally rugose. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae usually unsculptured, smooth, and shining, rarely with superficial, weak sculpture. Waist segments mainly longitudinally rugose. Gaster completely unsculptured, smooth, and shining. Ground sculpture generally faint to absent everywhere on body. Body with numerous long, fine, standing hairs. Anterior edges of antennal scapes with suberect to erect hairs. Head, mesosoma, waist segments, and gaster brown to dark brown, mandibles, antennae, and legs of much lighter colour, usually yellow.
|
||||
</paragraph>
|
||||
<paragraph box="[118,170,704,722]" pageId="23" pageNumber="24" type="mainText">Notes</paragraph>
|
||||
<paragraph id="748B65D42F55C420B5DB73411A4E13D4" box="[118,170,704,722]" pageId="23" pageNumber="24" type="mainText">Notes</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="reference_group">
|
||||
<paragraph box="[118,1122,732,807]" pageId="23" pageNumber="24" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="andohahela">Tetramorium andohahela</taxonomicName>
|
||||
<subSubSection id="20536F8EB60D0633B9E9E67357F298AE" type="reference_group">
|
||||
<paragraph id="48519FDB09F8AAC4DC2C4717BB6375B6" box="[118,1122,732,807]" pageId="23" pageNumber="24" type="mainText">
|
||||
<taxonomicName id="149A4B8B2EE8CDAED201C90951CEFF2B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="andohahela">Tetramorium andohahela</taxonomicName>
|
||||
is only found in a strip of localities in southern-eastern Madagascar that ranges from the southernmost locality, Andohahela, north to Ranomafana. All localities are rainforests or montane rainforests located at elevations of 800 to 1250 m. Also, the species was mostly collected from leaf litter.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,1122,816,948]" pageId="23" pageNumber="24" type="mainText">
|
||||
<subSubSection id="6BCE33D3502CB7E690FC9061DF3D8571" type="discussion">
|
||||
<paragraph id="898971852ED088E93EFAE35E53102347" box="[118,1122,816,948]" pageId="23" pageNumber="24" type="mainText">
|
||||
The petiolar node is somewhat variable in its length since it ranges from longer than wide to wider than long (LPeI 93-105). This is not of much diagnostic importance, however, since the node always retains its characteristic shape with the posterodorsal corner protruding strongly posteriorly. This extraordinary node shape is absent in the remainder of the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="145C7300872045CC92BD468BCF483D49" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
complex and the whole
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="D9116823DA96763BC9D8604A92E71A64" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group in Madagascar, making T.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
<taxonomicName id="FC61B41819DE9A05791D0EDBC98CC356" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
straightforwardly recognisable.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,980,998]" pageId="23" pageNumber="24" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,1120,1008,1083]" pageId="23" pageNumber="24" type="mainText">
|
||||
<subSubSection id="CAB7BADE3653AFD80494234CABB664B0" type="etymology">
|
||||
<paragraph id="D9A0DA0B545356F704EB5884DEC829CF" box="[118,219,980,998]" pageId="23" pageNumber="24" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="D904BF61192D3150D8896B52092205FE" box="[118,1120,1008,1083]" pageId="23" pageNumber="24" type="mainText">
|
||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="8FBF95D10FBC4543F6584ECF5FBA3877" rank="species">new species</taxonomicNameLabel>
|
||||
is named after the type locality, the Andohahela National Park, which harbours an extraordinary variety of landscapes and habitats, and is of special importance for the conservation of biodiversity in Madagascar. The species epithet is a noun in apposition, and thus invariant.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,1120,1139]" pageId="23" pageNumber="24" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,1148,1590]" pageId="23" pageNumber="24" type="mainText">
|
||||
<subSubSection id="F8474094AE1E393E5F959B7A174397C3" type="materials_examined">
|
||||
<paragraph id="1B30DDD6F3D46DBD68E270E07623D286" box="[118,295,1120,1139]" pageId="23" pageNumber="24" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="4D02541A7B5F241F83432093B9B25B5A" box="[118,1122,1148,1590]" pageId="23" pageNumber="24" type="mainText">
|
||||
MADAGASCAR: Fianarantsoa, 2 km W Andrambovato, along river Tatamaly, 21.51167 S, 47.41 E, 1075 m, montane rainforest, 3.-5.VI.2005 (B.L. Fisher et al.); Fianarantsoa,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="7F761BA0E5457734C02444804F4D6C2A" originalValue="Rés">Res</normalizedToken>
|
||||
. Andringitra, 43 km S Ambalavao, 22.23333 S, 47 E, 825 m, rainforest, 4.X.1993 (B.L. Fisher); Fianarantsoa, Parc National Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango,23.83517 S, 46.96367 E, 940 m, rainforest, 13.-15.XI.2006 (B.L. Fisher et al.); Fianarantsoa, Parc National Befotaka-Midongy, Papango 28.5 km S Midongy-Sud, Mount Papango, 23.84083 S, 46.9575 E, 1250 m, montane rainforest, 17.XI.2006 (B.L. Fisher et al.); Fianarantsoa, R.S. Ivohibe, 7.5 km ENE Ivohibe, 22.47 S, 46.96 E, 900 m, rainforest, 7.-12.X.1997 (B.L. Fisher); Fianarantsoa, 9.0 km NE Ivohibe, 22.42667 S, 46.93833 E, 900 m, rainforest, 12.-17.X.1997 (B.L. Fisher); Fianarantsoa, Parc Nationale Ranomafana, Talatakely, 21.24833 S, 47.42667 E, in guava forest, 9.-26.IV.1998 (C. Griswold et al.); Fianarantsoa, Parc National de Ranomafana, Vatoharanana River, 4.1 km 231° SW Ranomafana, 21.29 S, 47.43333 E, 1100 m, montane rainforest, 27.-31.III.2003 (B.L. Fisher, C. Griswold, et al.); Toliara,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="364C69F5E98D363A2F245ADF0DD0EC49" originalValue="Rés">Res</normalizedToken>
|
||||
. Andohahela, 11 km NW Enakara, 24.56667 S, 46.83333 E, 800 m, rainforest, 16.XI.1992 (B.L. Fisher); Toliara,13 km NW Enakara,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="B955031D1A1F8D4A7B3122F587E82A33" originalValue="Rés">Res</normalizedToken>
|
||||
. Andohahela, 24.55 S, 46.8 E, 1250 m, montane rainforest, 30.XI.1992 (B.L. Fisher); Toliara, Parc National d'Andohahela, Col du Sedro, 3.8 km 113° ESE Mahamavo, 37.6 km 341° NNW Tolagnaro, 24.76389 S, 46.75167 E, 900 m, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF05010, 21.-25.I.2002 (B.L. Fisher, C. Griswold, et al.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,98 +1,97 @@
|
|||
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172649" ID-GBIF-Taxon="100105393" ID-Zenodo-Dep="6172649" LSID="urn:lsid:plazi:treatment:575E61C31E5BDCFE52C8E7D9514737CA" httpUri="http://treatment.plazi.org/id/575E61C31E5BDCFE52C8E7D9514737CA" lastPageNumber="66" pageNumber="66">
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
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<taxonomicName id="860D4CADF4CD5FF9E90EDBABE0698AAC" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
species complex
|
||||
</paragraph>
|
||||
</subSubSection>
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<subSubSection type="discussion">
|
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<paragraph box="[118,1122,427,530]" pageId="65" pageNumber="66" type="mainText">
|
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<subSubSection id="1D567B9EC1397E1A02BBBB1D8EB1FF37" type="discussion">
|
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|
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This complex includes the species
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="adamsi">
|
||||
<taxonomicName id="46C6537E1DC0C435AE7706776138B1F1" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="adamsi">
|
||||
T. adamsi
|
||||
<authority>,</authority>
|
||||
<authority id="27106899C36AA9C68281E19D0E1DC624">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
<taxonomicName id="CFA420F77FC39479A9775EB05B2B443A" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
<taxonomicName id="3A6484155CAAE8AF669172B2C0958FA9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
T. latreillei
|
||||
<authority>,</authority>
|
||||
<authority id="D50C827FCFC8C3F80330127F070DB1AF">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="marojejy">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="marojejy">
|
||||
<taxonomicName id="D5B3583B2691CE7287C114DA156AA2CF" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="marojejy">
|
||||
<taxonomicName id="7FA5A7D1C2700729135E0BD1263ABBB3" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="marojejy">
|
||||
T. marojejy
|
||||
<authority>,</authority>
|
||||
<authority id="5DD84D2E4E56F4BA72BD03F8DFBD7988">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="65" pageNumber="66" rank="species" species="nazgul">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="nazgul">
|
||||
<taxonomicName id="77AF3D4D1AAF518957940D63A32842BB" authorityName="," genus="Tetamorium" pageId="65" pageNumber="66" rank="species" species="nazgul">
|
||||
<taxonomicName id="A8C0B3EDA2D568DB30529ABBABA68EEE" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="nazgul">
|
||||
T. nazgul
|
||||
<authority>,</authority>
|
||||
<authority id="95A1216BAC3E33E595B0E9D77480DA27">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
<taxonomicName id="36471FF31F258B3AF35EC063BD56367E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
<taxonomicName id="5A6370E282EFD36839E7615F647D587D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
They are all characterized by the presence of very strongly developed longitudinal rugae on the forecoxae, and the absence of sculpture on the first gastral tergite (except in some specimens of T. manongarivo, which have costulate sculpture on the basal half of the first gastral tergite).
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,539,615]" pageId="65" pageNumber="66" type="mainText">
|
||||
<paragraph id="AE7045E37827415B5659085B2118D09A" box="[118,1122,539,615]" pageId="65" pageNumber="66" type="mainText">
|
||||
The species within this group can be further divided into those with massive propodeal spines and reduced hairiness (
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
<taxonomicName id="099C4B9DEAE67FB1CF393B2FD260F489" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
T. latreillei
|
||||
<authority>,</authority>
|
||||
<authority id="D63BEDF711F5CB96D5C513F3F0E22D87">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
<taxonomicName id="094982E877D77C7D6DC96C82578ED8E3" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="smaug">T. smaug),</taxonomicName>
|
||||
<taxonomicName id="A05D048803E0265D684F0E4D8B4AE665" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="smaug">T. smaug),</taxonomicName>
|
||||
while the remaining three species are much hairier and have less massive propodeal spines (
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="adamsi">
|
||||
<taxonomicName id="55FDCE7011EA0C42B90C47CD2C791716" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="adamsi">
|
||||
T. adamsi
|
||||
<authority>,</authority>
|
||||
<authority id="EB9D202E7D257E03D31A511284CCDBB2">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy,</taxonomicName>
|
||||
<taxonomicName id="F89A055147063E987F271151693408BA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy,</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul).</taxonomicName>
|
||||
<taxonomicName id="42C0F7A2A2AFA49C381197539AF91CFD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul).</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
|
|
|
|||
|
|
@ -1,168 +1,167 @@
|
|||
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<mods:title id="825CE701A91B25B1627B08596E574C11">Zootaxa</mods:title>
|
||||
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|
||||
<mods:part>
|
||||
<mods:date>2012</mods:date>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>3592</mods:number>
|
||||
<mods:part id="A822706A62F883F1F9C8646498B8E9AC">
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||||
<mods:date id="8885F1F2EDD45104E4DD0171F22EF11B">2012</mods:date>
|
||||
<mods:detail id="F300D47062716148B971D10FEAD25A54" type="volume">
|
||||
<mods:number id="483CC95921F56E057EFC5D7CC85C8C3B">3592</mods:number>
|
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|
||||
<mods:extent unit="page">
|
||||
<mods:start>1</mods:start>
|
||||
<mods:end>85</mods:end>
|
||||
<mods:extent id="C29ABD68714B87EF11BE0216A21D50B3" unit="page">
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||||
<mods:start id="7DF9F49DF43095EF188A4195D6C3815F">1</mods:start>
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||||
<mods:end id="6A3C6EA7A0FB058EB68A163DDB976462">85</mods:end>
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<mods:url>http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
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<mods:location id="8939DECD1D67785F6B40C849D0137AD8">
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<mods:url id="761CF6DD73E8684EC4744F23F40D54CF">http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
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||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="HNS-Pub">26064</mods:identifier>
|
||||
<mods:identifier type="ISSN">175-5326</mods:identifier>
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||||
<mods:identifier type="ZooBank">A2D9C9ED-C0BA-4B5F-A330-C9AB7D625704</mods:identifier>
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<mods:classification id="396D80C5B85972BC9ED48243F9C9D8F9">journal article</mods:classification>
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<mods:identifier id="DA33C6D5820C38B3B36C66B730B45069" type="HNS-Pub">26064</mods:identifier>
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<mods:identifier id="F1FA16CF188088BD194EB6BCB0A07109" type="ISSN">175-5326</mods:identifier>
|
||||
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|
||||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172663" ID-GBIF-Taxon="100105391" ID-Zenodo-Dep="6172663" LSID="urn:lsid:plazi:treatment:5F39CF90488FDD707A2D5F5193D6F50A" httpUri="http://treatment.plazi.org/id/5F39CF90488FDD707A2D5F5193D6F50A" lastPageNumber="80" pageNumber="79">
|
||||
<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,581,709,729]" pageId="78" pageNumber="79" type="mainText">
|
||||
<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="smaug">
|
||||
<treatment id="5F39CF90488FDD707A2D5F5193D6F50A" ID-DOI="http://doi.org/10.5281/zenodo.6172663" ID-GBIF-Taxon="100105391" ID-Zenodo-Dep="6172663" LSID="urn:lsid:plazi:treatment:5F39CF90488FDD707A2D5F5193D6F50A" httpUri="http://treatment.plazi.org/id/5F39CF90488FDD707A2D5F5193D6F50A" lastPageNumber="80" pageNumber="79" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="4060B158BB7BE91DB691F0CC2EC8D1BD" type="nomenclature">
|
||||
<paragraph id="6F840C30E15262B990E0167E3CDBB373" box="[118,581,709,729]" pageId="78" pageNumber="79" type="mainText">
|
||||
<taxonomicName id="A89DBF2364951D5CF0AB82E05097B133" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="smaug">
|
||||
Tetramorium smaug
|
||||
<authority>Hita</authority>
|
||||
<authority id="13584BA97038114E944398ECD08A9F7E">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="B3FA882D578AF08CBEDC91FCB28BD02B" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,410,736,755]" pageId="78" pageNumber="79" type="mainText">(Figs. 27, 28, 138, 139, 140, 142)</paragraph>
|
||||
<subSubSection id="8ED8ACB7DF15C6A60CD1998C68BB3A08" type="description">
|
||||
<paragraph id="0020A3107DA17E8C461D64DCD34E43A8" box="[118,410,736,755]" pageId="78" pageNumber="79" type="mainText">(Figs. 27, 28, 138, 139, 140, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1122,792,868]" pageId="78" pageNumber="79" type="mainText">Holotype worker, MADAGASCAR, Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.83937 S, 48.30842 E, 1080 m, montane rainforest, pitfall trap, collection code BLF16917, 4.-7.V.2007 (B.L. Fisher et al.) (CASC: CASENT0121244). Paratype, one worker with same data as holotype (CASC: CASENT0124788).</paragraph>
|
||||
<subSubSection id="F36662D2AEFA98B80D94BD4FE973900B" type="materials_examined">
|
||||
<paragraph id="C0817F0B4F3E20577381A42CAD559760" box="[118,1122,792,868]" pageId="78" pageNumber="79" type="mainText">Holotype worker, MADAGASCAR, Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.83937 S, 48.30842 E, 1080 m, montane rainforest, pitfall trap, collection code BLF16917, 4.-7.V.2007 (B.L. Fisher et al.) (CASC: CASENT0121244). Paratype, one worker with same data as holotype (CASC: CASENT0124788).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,899,918]" pageId="78" pageNumber="79" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1122,928,1087]" pageId="78" pageNumber="79" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
<subSubSection id="101F461C7411AD93102E098715B233EE" type="diagnosis">
|
||||
<paragraph id="CB8FA8F201EB1E12DE21397F43850F90" box="[118,208,899,918]" pageId="78" pageNumber="79" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="737C862D44823CCB4F59BABC4E7F3758" box="[118,1122,928,1087]" pageId="78" pageNumber="79" type="mainText">
|
||||
<taxonomicName id="DD5F2C02EB09E7C3D73366D1192BA769" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
can be easily discriminated from the remainder of the species complex by the following character set: antennal scapes comparatively short (SI 77-81); extremely long and massively constructed propodeal spines (PSLI 57-63); anterodorsal and posterodorsal margins of petiolar node situated at about same height; mesosoma with 7 to 14 pairs of standing hairs; hairs on leading edge of antennal scapes subdecumbent to suberect; first gastral tergite with several scattered standing hairs and very sparse, short, appressed pubescence; very dark brown to black colouration.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,1119,1138]" pageId="78" pageNumber="79" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1124,1148,1279]" pageId="78" pageNumber="79" type="mainText">HL 0.99-1.04 (1.02); HW 0.99-1.06 (1.02); SL 0.80-0.85 (0.81); EL 0.19-0.23 (0.21); PH 0.50-0.54 (0. 52); PW 0.70-0.77 (0.74); WL 1.30-1.38 (1.34); PSL 0.57-0.66 (0.60); PTL 0.30-0.38 (0.34); PTH 0.36-0.43 (0.40); PTW 0.25-0.32 (0. 29); PPL 0.30-0.32 (0.31); PPH 0.38-0.42 (0.41); PPW 0.36-0.41 (0.38); CI 100-102 (101); SI 77-81 (79); OI 19-22 (20); DMI 54-58 (56); LMI 38-40 (39); PSLI 57-63 (59); PeNI 35-41 (38); LPeI 77-91 (86); DPeI 79-88 (83); PpNI 48-55 (52); LPpI 73-79 (76); DPpI 120-132 (125); PPI 128-144 (135) (six measured).</paragraph>
|
||||
<paragraph box="[118,1121,1288,1589][118,1122,118,475]" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79" type="mainText">
|
||||
<subSubSection id="73F33AF36D2E1762B820B32D1CBE7C16" type="description">
|
||||
<paragraph id="BCAD9ABDFB1A08CF03C1B09F156C2695" box="[118,226,1119,1138]" pageId="78" pageNumber="79" type="mainText">Description</paragraph>
|
||||
<paragraph id="C9FD9514BEBF3BED72641B38FBF60B09" box="[118,1124,1148,1279]" pageId="78" pageNumber="79" type="mainText">HL 0.99-1.04 (1.02); HW 0.99-1.06 (1.02); SL 0.80-0.85 (0.81); EL 0.19-0.23 (0.21); PH 0.50-0.54 (0. 52); PW 0.70-0.77 (0.74); WL 1.30-1.38 (1.34); PSL 0.57-0.66 (0.60); PTL 0.30-0.38 (0.34); PTH 0.36-0.43 (0.40); PTW 0.25-0.32 (0. 29); PPL 0.30-0.32 (0.31); PPH 0.38-0.42 (0.41); PPW 0.36-0.41 (0.38); CI 100-102 (101); SI 77-81 (79); OI 19-22 (20); DMI 54-58 (56); LMI 38-40 (39); PSLI 57-63 (59); PeNI 35-41 (38); LPeI 77-91 (86); DPeI 79-88 (83); PpNI 48-55 (52); LPpI 73-79 (76); DPpI 120-132 (125); PPI 128-144 (135) (six measured).</paragraph>
|
||||
<paragraph id="C8B20CD13174D7648EB298B486E4D5F5" box="[118,1121,1288,1589][118,1122,118,475]" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79" type="mainText">
|
||||
Head as long as wide to weakly wider than long (CI 100-102); posterior head margin strongly concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes well-developed, moderately deep, but narrow and without defined posterior and ventral margins. Antennal scapes of moderate length, not reaching posterior head margin (SI 77-81). Eyes small to moderate (OI 19-22). Mesosomal outline in profile flat, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-40). Propodeal spines massively constructed with very broad base, extremely long, and acute (PSLI 57-63); propodeal lobes short and rounded. Petiolar node in profile rectangular nodiform, approximately 1.1 to 1.3 times higher than long (LPeI 77-91), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum slightly convex; node in dorsal view approximately 1.1 to 1.3 times longer than wide (DPeI 79-88). Postpetiole in profile subglobular, weakly anteroposteriorly
|
||||
<pageBreakToken pageId="79" pageNumber="80" start="start">compressed</pageBreakToken>
|
||||
<pageBreakToken id="91529AF33B7F6110F1F569F626BD1AFB" pageId="79" pageNumber="80" start="start">compressed</pageBreakToken>
|
||||
, approximately 1.2 to 1.4 times higher than long (LPpI 73-79); in dorsal view around 1.2 to 1.3 times wider than long (DPpI 120-132). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view approximately 1.3 to 1.5 times wider than petiolar node (PPI 128-144). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose, with three to five rugae; cephalic dorsum between frontal carinae with 9 to 12 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted or with cross-meshes; lateral and ventral head longitudinally rugose, rarely with cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae with very distinct and pronounced longitudinal rugae. Waist segments longitudinally rugose, rugae on waist segments weaker than on head and mesosoma, especially dorsally. Gaster completely unsculptured, smooth, and shining. Ground sculpture generally faint to absent everywhere on body. Head with abundant standing hairs; mesosoma with 7 to 14 pairs of hairs; waist segments and first gastral tergite with few to several scattered, standing hairs; first gastral tergite with very sparse, short, and appressed pubescence. Anterior edges of antennal scapes with subdecumbent to suberect hairs. Body a uniform very dark brown to black colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,512,530]" pageId="79" pageNumber="80" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1122,540,756]" pageId="79" pageNumber="80" type="mainText">
|
||||
<subSubSection id="B1B6752FAB309D69A4CA330AB4DD11E4" type="discussion">
|
||||
<paragraph id="59538C7E6D96E2A44B0495117E4A63C3" box="[118,170,512,530]" pageId="79" pageNumber="80" type="mainText">Notes</paragraph>
|
||||
<paragraph id="FCBC301249073A05F92CCA7E294E048B" box="[118,1122,540,756]" pageId="79" pageNumber="80" type="mainText">
|
||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="F34FB9EC14870B83EB133B78B9FB9C9C" rank="species">new species</taxonomicNameLabel>
|
||||
is currently only known from Ambatovy, Montagne d'Ambre, and Ivohibe. All three localities are montane rainforests at altitudes of 900 to 1300 m. However, these sites are geographically widely separated, one being located in the southeast, one in the east, and one in the northernmost tip of the island. This represents a fairly disjunctive distribution. Furthermore, the species is only known from six specimens, which could mean that it is very rare and uncommon, as seems to be the case with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="47AB7BB85767545C50FF47CF03F5B30C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
<taxonomicName id="1409BCE5C8706FE3CABEDCB9E7D20624" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
The scant available material was collected from pitfall traps, hand collecting, or rotten logs, which suggests that
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="07E57DF03DDAD8D25168A9380572DC49" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
is rarely encountered on the ground. As already stated in the description of
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="79" pageNumber="80" rank="species" species="sabatra">
|
||||
<taxonomicName id="6AD8F6CEFD480D0C76CE9835F5B6FC96" authorityName="," genus="Tetamorium" pageId="79" pageNumber="80" rank="species" species="sabatra">
|
||||
T. sabatra
|
||||
<authority>,</authority>
|
||||
<authority id="D3E4A7B6CB48A95403BD9597CF191EC3">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="1FF204A36DDAEB619685069B45D1C899" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
might live in the vegetation, which could explain why it was only seldom sampled.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,765,925]" pageId="79" pageNumber="80" type="mainText">
|
||||
<paragraph id="11E64798A0EEDBCC912C213AAD5B32A9" box="[118,1122,765,925]" pageId="79" pageNumber="80" type="mainText">
|
||||
Within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="1D587AB3C2B0B45B74419D14CDAA30F6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
species complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="4A0C33CF12B7F107070093EAD5C2D5ED" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
cannot be mistaken for
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
<taxonomicName id="D496F412437E6DC97D8C456A2030923C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
since the latter has a petiolar node with the posterodorsal margin situated higher than the anterodorsal margin. Also, it is unlikely to be confused with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
<taxonomicName id="7089E5CB12FB7435BDF89E4324421D80" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
or
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
<taxonomicName id="A3A704E630A7311E25017DA7C2A7759C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
because they are both much hairier than
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
<taxonomicName id="B375C38CB40230F72BAF036C9DECD3E2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
In addition,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
<taxonomicName id="CC9D1B7CE07001277EFEABA76A276620" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
possesses the longest antennal scapes of the complex (SI 89-93) while the scapes of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="C2A95777EEA132C13E78523913908BA9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
are much shorter (SI 77-81). Moreover,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
<taxonomicName id="E1ECD2BF2E60F490068EA5E7017CCE2B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
differs also in body colour, which is orange to pale brown, whereas the body colour of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="D45FF9B13A6E4606B17F1654088CD0C6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
is very dark brown to black.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,934,1206]" pageId="79" pageNumber="80" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
<paragraph id="5E0ECB0C45C84FB8B8C9CDA22F6C3E36" box="[118,1122,934,1206]" pageId="79" pageNumber="80" type="mainText">
|
||||
<taxonomicName id="100D6017BC5FD1AC645897D61547735D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
is morphologically most closely associated with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="E2979F3E76C73BED690FDC2E76D7318B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
<taxonomicName id="3661681A76148A2EB0D8C1E4908BB7F8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
As mentioned above, the three species share the character combination of massive, extremely long propodeal spines, reduced hairiness, and very dark brown to back colouration. This character combination makes them unlikely to be confused with another species of the complex.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
<taxonomicName id="8A420BF5F6AD4CC8C80E7CDEFAF13242" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
is also easily distinguished from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="0C49FF33732F13D36F5A46FA17D78AE5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="latreillei">Tetramorium latreillei</taxonomicName>
|
||||
<taxonomicName id="B05092877750395E0244463634CA37F7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra.</taxonomicName>
|
||||
<taxonomicName id="CEA1C43CCBF1E4655FDCBD43C10077C8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="latreillei">Tetramorium latreillei</taxonomicName>
|
||||
has no standing hairs on the waist segments or the first gastral tergite, and displays moderately dense appressed pubescence on the first gastral tergite. This separates it clearly form
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="817688A2C2E003805F5E945406D5ECF3" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
<taxonomicName id="8C21541C57531136B833A9EC0B9283FB" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
The latter two can also be easily distinguished from each other.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
<taxonomicName id="0B1F9B0CCF38CE580002EB86A83E10D2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">Tetramorium smaug</taxonomicName>
|
||||
has 7 to 14 pairs of hairs throughout the mesosomal dorsum, whereas only one or two pairs are present in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
<taxonomicName id="CC113A62320C8E7CCBACEDB602255095" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
and these are restricted to the pronotal dorsum. In addition, the hairs on the leading edge of the antennal scapes are generally strongly appressed in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="0BC6753205FD0EEF10AD91F56C44ED10" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
while they are subdecumbent to suberect in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
<taxonomicName id="6626667DFB3721958CAD4B3EECC7DBDF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,1215,1319]" pageId="79" pageNumber="80" type="mainText">
|
||||
<paragraph id="A3EFE31C8657297A78114A843102B16C" box="[118,1122,1215,1319]" pageId="79" pageNumber="80" type="mainText">
|
||||
As discussed in the descriptions of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="4955660611A5C41840585D454540EAE0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
<taxonomicName id="AC1DCD50DE786BCAAD1EB326F2E99DCB" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
there is a possibility that
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="E666A3BF87C72CBE064C50684DF91C04" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
is conspecific with one or even both of them. The lack of material in all three species makes species delimitations difficult, although we think that the differences in pilosity/pubescence are sufficient on the basis of the material examined in this study. At present, we treat all three as distinct species until more material becomes available.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,1351,1369]" pageId="79" pageNumber="80" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,1122,1379,1426]" pageId="79" pageNumber="80" type="mainText">
|
||||
<subSubSection id="F523298B516BE52EC97C6888C268EDE1" type="etymology">
|
||||
<paragraph id="755C0CA563E75A62AD7BC3167748BC82" box="[118,219,1351,1369]" pageId="79" pageNumber="80" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="A6608F93CC2DBF929665DBFD4C4097D7" box="[118,1122,1379,1426]" pageId="79" pageNumber="80" type="mainText">
|
||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="A31C22403F720873CFE423549E821FA9" rank="species">new species</taxonomicNameLabel>
|
||||
was named after
|
||||
<normalizedToken originalValue="“Smaug”">"Smaug"</normalizedToken>
|
||||
<normalizedToken id="DF51FEC379D54ACBD3AA4A04934517D0" originalValue="“Smaug”">"Smaug"</normalizedToken>
|
||||
, the villain dragon from the fantasy novel "The Hobbit" written by J.R.R. Tolkien. The species epithet is an arbitrary combination of letters.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,1458,1476]" pageId="79" pageNumber="80" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1123,1486,1590]" pageId="79" pageNumber="80" type="mainText">MADAGASCAR: Antsiranana, Parc National Montagne d'Ambre, 12.2 km 211° SSW Joffreville, 12.59639 S, 49.1595 E, 1300 m, montane rainforest, 2.-7.II.2001 (B.L. Fisher, C. Griswold et al.); Fianarantsoa, R.S. Ivohibe, 7.5 km ENE Ivohibe, 22.47 S, 46.96 E, 900 m, montane rainforest, 7.-12.X.1997 (B.L. Fisher); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.83937 S, 48.30842 E, 1080 m, montane rainforest, 4.-7.V.2007 (B.L. Fisher et al.).</paragraph>
|
||||
<subSubSection id="107643025479380C7A572DA1D99FD6A3" type="materials_examined">
|
||||
<paragraph id="92855AB9293C95A1212337E18CE7CF5E" box="[118,295,1458,1476]" pageId="79" pageNumber="80" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="D21E8D191F5F69ED60A724CC095039CC" box="[118,1123,1486,1590]" pageId="79" pageNumber="80" type="mainText">MADAGASCAR: Antsiranana, Parc National Montagne d'Ambre, 12.2 km 211° SSW Joffreville, 12.59639 S, 49.1595 E, 1300 m, montane rainforest, 2.-7.II.2001 (B.L. Fisher, C. Griswold et al.); Fianarantsoa, R.S. Ivohibe, 7.5 km ENE Ivohibe, 22.47 S, 46.96 E, 900 m, montane rainforest, 7.-12.X.1997 (B.L. Fisher); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.83937 S, 48.30842 E, 1080 m, montane rainforest, 4.-7.V.2007 (B.L. Fisher et al.).</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,216 +1,215 @@
|
|||
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<mods:date id="BA282928D48F826ECB1380B4EE5853B3">2012</mods:date>
|
||||
<mods:detail id="DE976B675F07AD50FE27879385CF3FA5" type="volume">
|
||||
<mods:number id="ACE1479DA44B713020EA23F6722B7B0C">3592</mods:number>
|
||||
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|
||||
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|
||||
<mods:start>1</mods:start>
|
||||
<mods:end>85</mods:end>
|
||||
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<mods:start id="837EACA5AB9921E8E2EC634A7DAF0EC4">1</mods:start>
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<mods:end id="AAB01534AA232B2EA6F9ED8A4F3257A5">85</mods:end>
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||||
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|
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|
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|
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<mods:location>
|
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<mods:url>http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
|
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<mods:location id="2644CE8038B3BDDE933542A98AD5A377">
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<mods:url id="FB814FB0A9C7430ECD7ED177E00DCA12">http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="HNS-Pub">26064</mods:identifier>
|
||||
<mods:identifier type="ISSN">175-5326</mods:identifier>
|
||||
<mods:identifier type="ZooBank">A2D9C9ED-C0BA-4B5F-A330-C9AB7D625704</mods:identifier>
|
||||
<mods:classification id="58694355B65408E24C7EF2404E72F609">journal article</mods:classification>
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<mods:identifier id="E3E5E2D18FE57B286648A8D371415C47" type="HNS-Pub">26064</mods:identifier>
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<mods:identifier id="1358B07CFF278FC2F6353B02F9BF570C" type="ISSN">175-5326</mods:identifier>
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||||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172631" ID-GBIF-Taxon="100105331" ID-Zenodo-Dep="6172631" LSID="urn:lsid:plazi:treatment:641AEFF92ABFEDDC681F0186327D41FD" httpUri="http://treatment.plazi.org/id/641AEFF92ABFEDDC681F0186327D41FD" lastPageNumber="50" pageNumber="47">
|
||||
<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,598,1270,1290]" pageId="46" pageNumber="47" type="mainText">
|
||||
<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
<treatment id="641AEFF92ABFEDDC681F0186327D41FD" ID-DOI="http://doi.org/10.5281/zenodo.6172631" ID-GBIF-Taxon="100105331" ID-Zenodo-Dep="6172631" LSID="urn:lsid:plazi:treatment:641AEFF92ABFEDDC681F0186327D41FD" httpUri="http://treatment.plazi.org/id/641AEFF92ABFEDDC681F0186327D41FD" lastPageNumber="50" pageNumber="47" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="D49E08CA1A9F2EAF2B57F2A09F95522E" type="nomenclature">
|
||||
<paragraph id="E54E049F174DCB4BA708B95A6594EC73" box="[118,598,1270,1290]" pageId="46" pageNumber="47" type="mainText">
|
||||
<taxonomicName id="46F2CDEC5026DA886907845EB6B19118" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="avaratra">
|
||||
Tetramorium avaratra
|
||||
<authority>Hita</authority>
|
||||
<authority id="F971F93365D0169CA4C5384D058026CD">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="6CDEB0798ADD2A6DC0E1E96D57F10B69" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,444,1298,1317]" pageId="46" pageNumber="47" type="mainText">(Figs. 46, 47, 49, 102, 103, 104, 141)</paragraph>
|
||||
<subSubSection id="A7D5DD10273EDC3C35D3F91C595006D1" type="description">
|
||||
<paragraph id="96ACE9A52BE3C61735DDEF8521A226D8" box="[118,444,1298,1317]" pageId="46" pageNumber="47" type="mainText">(Figs. 46, 47, 49, 102, 103, 104, 141)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1125,1354,1542]" pageId="46" pageNumber="47" type="mainText">
|
||||
<subSubSection id="BB6E770723C2506729D4A5FB76AF92E6" type="materials_examined">
|
||||
<paragraph id="FB38C8224F10D02FBE2B7551C345C64D" box="[118,1125,1354,1542]" pageId="46" pageNumber="47" type="mainText">
|
||||
Holotype worker, MADAGASCAR, Antsiranana,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="1586F86A0956169F928B891EEF85AC8F" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="733242D00233675A87C2ED7BD33A8E6E" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
de l'Ankarana, 22.9 km 224° SW Anivorano Nord, 12.90889 S, 49.10983 E, 80 m, tropical dry forest, sifted litter (leaf mold, rotten wood), collection code BLF02858, 10.-16.II.2001 (B.L. Fisher, C. Griswold et al.) (CASC: CASENT0445167). Paratypes, 15 workers with same data as holotype (BMNH: CASENT0443679; CASC: CASENT0439463; CASENT0443689; CASENT0445147; CASENT0445161; CASENT0445174; CASENT0448424; CASENT0448440; CASENT0448557; CASENT0448579; CASENT0448643; CASENT0448665; MCZ: CASENT0443659; MHNG: CASENT0448417; NHMB: CASENT0448568).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="multiple">
|
||||
<caption box="[118,1122,1537,1582]" pageId="47" pageNumber="48" type="caption">
|
||||
<paragraph box="[118,1122,1537,1582]" pageId="47" pageNumber="48" type="caption">
|
||||
<pageBreakToken pageId="47" pageNumber="48" start="start">FIGURES</pageBreakToken>
|
||||
<subSubSection id="10A6B12A52CF2646EBF17320BA12CB91" type="multiple">
|
||||
<caption id="8CA6F9020C817B4985EF15DC4D71CD65" box="[118,1122,1537,1582]" pageId="47" pageNumber="48" type="caption">
|
||||
<paragraph id="9BA33D998822BBC829C056184451F3F5" box="[118,1122,1537,1582]" pageId="47" pageNumber="48" type="caption">
|
||||
<pageBreakToken id="262CD26C4385792D4550C2D241CD74C7" pageId="47" pageNumber="48" start="start">FIGURES</pageBreakToken>
|
||||
102-104.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra,</taxonomicName>
|
||||
<taxonomicName id="B00D0DF77FF167170873D7F3C58F4822" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra,</taxonomicName>
|
||||
holotype (CASENT0445167). 102. Body in profile. 103. Body in dorsal view. 104. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,118,137]" pageId="48" pageNumber="49" type="mainText">
|
||||
<pageBreakToken pageId="48" pageNumber="49" start="start">Diagnosis</pageBreakToken>
|
||||
<subSubSection id="27C3C79C6FB07473E6E670DFBD57A843" type="diagnosis">
|
||||
<paragraph id="40E499099D5B661788B15003DC88FB39" box="[118,208,118,137]" pageId="48" pageNumber="49" type="mainText">
|
||||
<pageBreakToken id="7F607DCD7BB7EDF073A78BFE7ACCC37D" pageId="48" pageNumber="49" start="start">Diagnosis</pageBreakToken>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,147,250]" pageId="48" pageNumber="49" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra</taxonomicName>
|
||||
<paragraph id="4B5389C4981FBA67CB98824A7F8E67B3" box="[118,1121,147,250]" pageId="48" pageNumber="49" type="mainText">
|
||||
<taxonomicName id="E2822B4C499BF07F08CFF8AF84980F2B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra</taxonomicName>
|
||||
is easily recognisable within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="0F723BA37DEFB8CE3C39AA6C314997A0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
complex due to the following character combination: propodeal spines long (PSLI 27-34 without the Nosy Be specimens, and PSLI 27-37 with the Nosy Be material); petiolar node in dorsal view between 1.2 to 1.4 times wider than long (DPeI 126-137); dorsum of petiolar node only weakly rugose; base of first gastral tergite with superficial, fine, reticulate-punctate sculpture.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,287,305]" pageId="48" pageNumber="49" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,315,447]" pageId="48" pageNumber="49" type="mainText">HL 0.81-0.94 (0.88); HW 0.78-0.94 (0.87); SL 0.56-0.65 (0.60); EL 0.15-0.18 (0.17); PH 0.41-0.48 (0.45); PW 0.59-0.71 (0.65); WL 1.02-1.20 (1.12); PSL 0.24-0.33 (0.28); PTL 0.19-0.24 (0.22); PTH 0.33-0.40 (0.37); PTW 0.26-0.31 (0.29); PPL 0.23-0.30 (0.27); PPH 0.29-0.37 (0.34); PPW 0.30-0.38 (0.35); CI 97-101 (99); SI 66-72 (69); OI 18-20 (19); DMI 56-60 (57); LMI 38-41 (40); PSLI 27-37 (32); PeNI 43-46 (44); LPeI 54-66 (60); DPeI 126-137 (130); PpNI 50-56 (54); LPpI 74-85 (79); DPpI 124-137 (130); PPI 116-125 (120) (15 measured).</paragraph>
|
||||
<paragraph box="[118,1121,456,1178]" pageId="48" pageNumber="49" type="mainText">Head approximately as long as wide (CI 97-101); posterior head margin moderately concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, moderately deep, and broad, without defined ventral margins. Antennal scapes very short, not reaching posterior head margin (SI 66-72). Eyes short (OI 18-20). Mesosomal outline in profile flat to weakly convex, strongly marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-41). Propodeal spines long, spinose and acute (PSLI 27-37); propodeal lobes short, triangular, and moderately acute. Petiolar node in profile rectangular nodiform, approximately 1.5 to 1.8 times higher than long (LPeI 54-66), anterior and posterior faces approximately parallel, anterodorsal margin situated higher than posterodorsal, dorsum noticeably tapering backwards posteriorly; node in dorsal view approximately 1.2 to 1.4 times wider than long (DPeI 126-137). Postpetiole in profile rounded, approximately 1.2 to 1.4 times higher than long (LPpI 74-85); in dorsal view around 1.2 to 1.4 times wider than long (DPpI 124-137). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view 1.1 to 1.3 times wider than petiolar node (PPI 116-125). Mandibles finely to strongly striate; clypeus longitudinally rugose, with four to eight rugae, median ruga always present, well-developed and distinct, remaining rugae variably developed, usually weaker and sometimes irregularly arranged; cephalic dorsum between frontal carinae with 11 to 13 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted and none with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose with very few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose, lateral mesosoma sometimes weaker sculptured than dorsum. Forecoxae generally unsculptured, smooth, and shining, at most with superficial sculpture. Ground sculpture on head and mesosoma generally faint to absent. Waist segments weakly to moderately rugose, sculpture on petiolar node weaker than on postpetiole; both waist segments with conspicuous reticulate-punctate ground sculpture. First gastral tergite with superficial, fine reticulate-punctate sculpture, generally restricted to basal third of the tergite, in several specimens sculpture fairly reduced, but always present. All dorsal surfaces of body with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with decumbent to suberect hairs. Body of uniform brown to dark brown colour, appendages often of lighter colour.</paragraph>
|
||||
<subSubSection id="3406189FE563AF4C048D2274614D81AE" type="description">
|
||||
<paragraph id="11DDF250C45155B6106BA31609D0FAC5" box="[118,226,287,305]" pageId="48" pageNumber="49" type="mainText">Description</paragraph>
|
||||
<paragraph id="E1DB48B923AEDBE2484379D48A425B7F" box="[118,1121,315,447]" pageId="48" pageNumber="49" type="mainText">HL 0.81-0.94 (0.88); HW 0.78-0.94 (0.87); SL 0.56-0.65 (0.60); EL 0.15-0.18 (0.17); PH 0.41-0.48 (0.45); PW 0.59-0.71 (0.65); WL 1.02-1.20 (1.12); PSL 0.24-0.33 (0.28); PTL 0.19-0.24 (0.22); PTH 0.33-0.40 (0.37); PTW 0.26-0.31 (0.29); PPL 0.23-0.30 (0.27); PPH 0.29-0.37 (0.34); PPW 0.30-0.38 (0.35); CI 97-101 (99); SI 66-72 (69); OI 18-20 (19); DMI 56-60 (57); LMI 38-41 (40); PSLI 27-37 (32); PeNI 43-46 (44); LPeI 54-66 (60); DPeI 126-137 (130); PpNI 50-56 (54); LPpI 74-85 (79); DPpI 124-137 (130); PPI 116-125 (120) (15 measured).</paragraph>
|
||||
<paragraph id="02132D95E95703B1D7ABB4552A8AAB7F" box="[118,1121,456,1178]" pageId="48" pageNumber="49" type="mainText">Head approximately as long as wide (CI 97-101); posterior head margin moderately concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, moderately deep, and broad, without defined ventral margins. Antennal scapes very short, not reaching posterior head margin (SI 66-72). Eyes short (OI 18-20). Mesosomal outline in profile flat to weakly convex, strongly marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-41). Propodeal spines long, spinose and acute (PSLI 27-37); propodeal lobes short, triangular, and moderately acute. Petiolar node in profile rectangular nodiform, approximately 1.5 to 1.8 times higher than long (LPeI 54-66), anterior and posterior faces approximately parallel, anterodorsal margin situated higher than posterodorsal, dorsum noticeably tapering backwards posteriorly; node in dorsal view approximately 1.2 to 1.4 times wider than long (DPeI 126-137). Postpetiole in profile rounded, approximately 1.2 to 1.4 times higher than long (LPpI 74-85); in dorsal view around 1.2 to 1.4 times wider than long (DPpI 124-137). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view 1.1 to 1.3 times wider than petiolar node (PPI 116-125). Mandibles finely to strongly striate; clypeus longitudinally rugose, with four to eight rugae, median ruga always present, well-developed and distinct, remaining rugae variably developed, usually weaker and sometimes irregularly arranged; cephalic dorsum between frontal carinae with 11 to 13 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted and none with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose with very few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose, lateral mesosoma sometimes weaker sculptured than dorsum. Forecoxae generally unsculptured, smooth, and shining, at most with superficial sculpture. Ground sculpture on head and mesosoma generally faint to absent. Waist segments weakly to moderately rugose, sculpture on petiolar node weaker than on postpetiole; both waist segments with conspicuous reticulate-punctate ground sculpture. First gastral tergite with superficial, fine reticulate-punctate sculpture, generally restricted to basal third of the tergite, in several specimens sculpture fairly reduced, but always present. All dorsal surfaces of body with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with decumbent to suberect hairs. Body of uniform brown to dark brown colour, appendages often of lighter colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,1215,1233]" pageId="48" pageNumber="49" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1122,1243,1403]" pageId="48" pageNumber="49" type="mainText">
|
||||
<subSubSection id="BA15ABA5CEBC51C2065AA707C105EED4" type="discussion">
|
||||
<paragraph id="715A50D805E5087CC51988E2C23AEB5A" box="[118,170,1215,1233]" pageId="48" pageNumber="49" type="mainText">Notes</paragraph>
|
||||
<paragraph id="F8DA112CC1615C1136C503C1F975B12E" box="[118,1122,1243,1403]" pageId="48" pageNumber="49" type="mainText">
|
||||
This
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="7CDE68FFE97A5256EC58306D3F56C34E" rank="species">new species</taxonomicNameLabel>
|
||||
is restricted in its distribution to several localities in the northern tip of Madagascar and Nosy Be. Interestingly, there is no material known from the area between the population on Nosy Be and the other localities from Andavakoera, Ankarana, Bekaraoka, Ampondrabe, and Analamerana.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra</taxonomicName>
|
||||
<taxonomicName id="B845D92713F3DE022AE9A09A89A267D8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra</taxonomicName>
|
||||
appears to have comparatively flexible habitat requirements, having been found in rainforests, tropical dry forests, and on tsingy. Also,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="A350693B44B815823AB0DFD0E038B449" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
appears to be a ground-active species sampled mainly from leaf litter, and seems restricted to low elevations of 30 to 425 m.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,1412,1572][118,1120,118,194]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra</taxonomicName>
|
||||
<paragraph id="97323E5C1054F399C17BE42EF2BD0D35" box="[118,1122,1412,1572][118,1120,118,194]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="mainText">
|
||||
<taxonomicName id="0ED16215F8DB863BA3752D2A35527B3E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra</taxonomicName>
|
||||
shows variation in two important morphological characters. First, the propodeal spines of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="741636FA6FDFAAE4C1AE985638C0C03B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
are usually long, but relatively short for the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="BF25E491DD1D2E41859F4D872B03FB48" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group, with a PSLI of 27-34 with a mean value of 31. The few specimens known from the island of Nosy Be, however, have significantly longer spines (PSLI 37-39). This is the only difference observed, and we consider it to be geographic variation since, as mentioned above, the Nosy Be population is fairly widely separated from the other
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="0E1C035A5D311397C7CF913A51AB470C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
populations. The second interesting character, which is variable, is the sculpture on the first gastral tergite. Generally, the reticulate-punctate
|
||||
<pageBreakToken pageId="49" pageNumber="50" start="start">ground</pageBreakToken>
|
||||
<pageBreakToken id="2A606872A094C189268747957EDFE5FA" pageId="49" pageNumber="50" start="start">ground</pageBreakToken>
|
||||
sculpture is well-developed on the basal third and fairly conspicuous. Nevertheless, in some specimens from Ankarana and Andavakoera, this character is often weakly developed, but still always present and visible.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1123,203,644]" pageId="49" pageNumber="50" type="mainText">
|
||||
<paragraph id="2202166BD88C7781BC7B4DB6338C0C88" box="[118,1123,203,644]" pageId="49" pageNumber="50" type="mainText">
|
||||
Within the complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="7244A1E6AB385AE7C59E8C8617557D3E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
cannot be mistaken with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="jedi">T. jedi.</taxonomicName>
|
||||
<taxonomicName id="1DF5C48AB248E8F4B4AC93D2EF754F5C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="jedi">T. jedi.</taxonomicName>
|
||||
The latter species has much more pronounced sculpture on the whole first gastral tergite, which is densely and strongly reticulate-punctate, while the sculpture in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="93642E4C0479F97B1CCE9567C18F85F2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
is only superficial, much less developed, and restricted to the basal third of the first gastral tergite. In addition, both differ in the shape of the petiolar node since it is distinctly longer than wide in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="EF86805A468737A5E032E64D953845E1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
(DPeI 79-85), whereas it is wider than long in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="F4509C1F22DE6139DED8FC32DD14FDED" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
(DPeI 126-137). The third species in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="C719EC3E07DBDC5C801DCE2F5763625C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon,</taxonomicName>
|
||||
<taxonomicName id="E7DCAC7A3EA2A087A8B968109AE9169F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon,</taxonomicName>
|
||||
is morphologically much closer to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra.</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">Tetramorium pleganon</taxonomicName>
|
||||
<taxonomicName id="706E1445C90580B2BBA996C92B91B84F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra.</taxonomicName>
|
||||
<taxonomicName id="52AC55D4AB001700892340A913A0DE6C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">Tetramorium pleganon</taxonomicName>
|
||||
differs from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="213C4342A5B6430124C99B8B013D62E2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
in several aspects, although some are not obvious at first glance. The propodeal spines are generally much longer in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="F65856C97A33D79617F7926B0D53532F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
(PSLI 37-44) than in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="8C3FD57AA17B2085E096920951CCD7EA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
(PSLI 27-34 without the Nosy Be specimens, PSLI 27-37 with the Nosy Be material). Additionally, the petiolar node dorsum is wider and higher in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="3943EA3E30FDE3AF54D773CB2AE630DC" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
(DPeI 126-137; LPeI 54-66) than in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="2927A09CDF3E690863B32580CB3DEA4B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
(DPeI 111-118; LPeI 63-73. Apart from these differences, both species differ also in the development of sculpture on the waist segments and the first gastral tergite, which is generally less well-developed in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="DA9151868F66B06FE9509CD5687EB859" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
than in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon.</taxonomicName>
|
||||
<taxonomicName id="3CA4CAFD79E66DCA4E2866E9AB37C685" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon.</taxonomicName>
|
||||
This is especially visible on the dorsum of the petiolar node, which is always strongly rugose in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="3E5C31072B461395B414B4572B21259D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
but much less rugose in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra,</taxonomicName>
|
||||
<taxonomicName id="3242CE6D338F5AA58DC42193419ADF8C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra,</taxonomicName>
|
||||
where it is partly smooth. The distribution ranges of both species overlap. Indeed,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="4CE0B8B0B86AF7A30B6D603DD6A4BE1E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="7A05852C4655D505EBE733E398156F2A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
are found in sympatry in Ankarana while retaining their species-characteristics, providing additional evidence for their heterospecificity.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1123,653,953]" pageId="49" pageNumber="50" type="mainText">
|
||||
<paragraph id="E83CEE0CD701522B02B140D02B1F7683" box="[118,1123,653,953]" pageId="49" pageNumber="50" type="mainText">
|
||||
As already mentioned above, it is possible that
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="167723E13FF530912333643875805991" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="048ED1925150DE9B6F0D9CA46BB4B0A3" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
are not members of the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="613CD16666F841FA6EBEC148B664192B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group. The petiolar nodes of both are wider than long in dorsal view (DPeI 111-137) while all other
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="02DAA253A245C20EA6FC4211C636DBA2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group species have petiolar node shapes that range from distinctly longer than wide to weakly wider than long (DPeI 72-114, usually below 100). Also, the general morphology of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="1677F33B910DF4227636F21D92C00425" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon,</taxonomicName>
|
||||
<taxonomicName id="6DF5686869343A736CD8381B214D168E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon,</taxonomicName>
|
||||
in particular the shape of the petiolar node and strong margination, is very close to some species from the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
<taxonomicName id="E1DDB2BB724C21CCB544F43DB347A008" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
species group, especially
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum.</taxonomicName>
|
||||
<taxonomicName id="4C6A809729B36A07B91C3A574A5EC5A7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum.</taxonomicName>
|
||||
However, no member of the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
<taxonomicName id="347B131356E981D500519CBB90C0ADBF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum</taxonomicName>
|
||||
group has a sculptured first gastral tergite, which clearly distinguishes
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="64AA930C942B7A8816BB0095F9E7B414" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="F946C69740A517C70E10523839FD96F5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
from that group despite the strong general similarity with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum.</taxonomicName>
|
||||
<taxonomicName id="2A6C7FB50EB58AB13717BEB83CA25FAB" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum.</taxonomicName>
|
||||
The latter species also has completely unsculptured mandibles, whereas the mandibles of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="3C116D4EA7285C275C44CE0FEAE5F4E7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="4DABBC4C163457C31CE30B1E2901B7B9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
are always sculptured, even though sometimes only finely so. At present, we consider these two species best placed within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="F08B5E0C7F6FA5306F9118FA24D1FE97" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group, but cannot discount that they might be more closely related to another species group or represent a lineage parallel to other Malagasy species groups.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,990,1008]" pageId="49" pageNumber="50" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,615,1018,1037]" pageId="49" pageNumber="50" type="mainText">The species epithet is an arbitrary combination of letters.</paragraph>
|
||||
<subSubSection id="F7E2972096D9A371F595D6215D10B972" type="etymology">
|
||||
<paragraph id="091F4FF0EE75910549CAC04E3FDE0E75" box="[118,219,990,1008]" pageId="49" pageNumber="50" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="A067419EE0DD447C24398D793CF42B04" box="[118,615,1018,1037]" pageId="49" pageNumber="50" type="mainText">The species epithet is an arbitrary combination of letters.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,1074,1093]" pageId="49" pageNumber="50" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,1103,1375]" pageId="49" pageNumber="50" type="mainText">
|
||||
<subSubSection id="67F7076EDF2C4F93414617B31841E3B5" type="materials_examined">
|
||||
<paragraph id="B32AD9A7FB9C5A238722C9043B809104" box="[118,295,1074,1093]" pageId="49" pageNumber="50" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="980D23A27784E731DB5244417AEBA85D" box="[118,1122,1103,1375]" pageId="49" pageNumber="50" type="mainText">
|
||||
MADAGASCAR: Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="411BCC71A26EBAF58207F57EE5AF4E35" originalValue="Forêt">Foret</normalizedToken>
|
||||
d'Ampondrabe, 26.3km 10° NNE Daraina, 12.97 S, 49.7 E, 175 m, tropical dry forest, 10.XII.2003 (B.L. Fisher); Antsiranana,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="7F37CA015F7EC41AE351B39A998DC8E9" originalValue="Rés">Res</normalizedToken>
|
||||
. Analamerana, 28.4 km 99° Anivorano-Nord, 12.74667 S, 49.49483 E, 60 m, tropical dry forest, 5.XII.2004 (B.L. Fisher); Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="CE550725AD7DE01BD92CAE9F432C760A" originalValue="Forêt">Foret</normalizedToken>
|
||||
d' Andavakoera, 21.4 km 75° ENE Ambilobe, 13.11833 S, 49.23 E, 425 m, rainforest 15.XII.2003 (B.L. Fisher); Antsiranana,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="C89ED98367D101A4672A9BB3EB110E9F" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="2259F3AE1509B20770ABF65C82CBD919" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
de l'Ankarana, 22.9 km 224° SW Anivorano Nord, 12.90889 S, 49.10983 E, 80 m, tropical dry forest, 10.-16.II.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="9DA047BEFE04729CFA81693ABF629262" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="61BE2BAA39B1E0477D8F781FC9A8396E" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
de l'Ankarana, 13.6 km 192° SSW Anivorano Nord, 12.86361 S, 49.22583 E, 210 m, tropical dry forest, 16.-21.II.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="0B5B23164778620555CAD763C8EC5494" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Bekaraoka, 6.8 km 60° ENE Daraina, 13.16667 S, 49.71 E, 150 m, tropical dry forest, 7.XII.2003 (B.L. Fisher); Antsiranana, Nosy Be,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="E78490FC9D9CBF0072060862A3C5F459" originalValue="Réserve">Reserve</normalizedToken>
|
||||
Naturelle
|
||||
<normalizedToken originalValue="Intégrale">Integrale</normalizedToken>
|
||||
<normalizedToken id="FE5F04E3986568366BCFBD7DE688410C" originalValue="Intégrale">Integrale</normalizedToken>
|
||||
de Lokobe, 6.3 km 112° ESE Hellville, 13.41933 S, 48.33117 E, 30 m, rainforest, 19.-24.III.2001 (B.L. Fisher, C. Griswold et al.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,115 +1,115 @@
|
|||
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<treatment ID-GBIF-Taxon="152033759" LSID="urn:lsid:plazi:treatment:7A7DFC885587389DC2D626B76F4EBA38" httpUri="http://treatment.plazi.org/id/7A7DFC885587389DC2D626B76F4EBA38" lastPageNumber="146" pageId="5" pageNumber="146">
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<subSubSection pageId="5" pageNumber="146" type="nomenclature">
|
||||
<paragraph pageId="5" pageNumber="146">
|
||||
<taxonomicName LSID="http://species-id.net/wiki/Calitys_scabra" authority="Thunberg, 1784" authorityName="Thunberg" authorityYear="1784" class="Insecta" family="Trogossitidae" genus="Calitys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Calitys scabra" order="Coleoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="scabra">Calitys scabra (Thunberg, 1784)</taxonomicName>
|
||||
<treatment id="7A7DFC885587389DC2D626B76F4EBA38" ID-GBIF-Taxon="152033759" LSID="urn:lsid:plazi:treatment:7A7DFC885587389DC2D626B76F4EBA38" httpUri="http://treatment.plazi.org/id/7A7DFC885587389DC2D626B76F4EBA38" lastPageNumber="146" pageId="5" pageNumber="146" scope_family="Trogossitidae" scope_order="Coleoptera" scope_subFamily="Trogossitinae" scope_tribe="Calityini">
|
||||
<subSubSection id="50EEAD688276147AA3DDFE3A1270757D" pageId="5" pageNumber="146" type="nomenclature">
|
||||
<paragraph id="29F5E7FB1DE042BE516C07145699D69C" pageId="5" pageNumber="146">
|
||||
<taxonomicName id="7F5A6D8682E2160947EEAB2C884AC256" LSID="http://species-id.net/wiki/Calitys_scabra" authority="Thunberg, 1784" authorityName="Thunberg" authorityYear="1784" genus="Calitys" lsidName="Calitys scabra" pageId="5" pageNumber="146" rank="species" species="scabra">Calitys scabra (Thunberg, 1784)</taxonomicName>
|
||||
Map 4
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="5" pageNumber="146" type="material examined">
|
||||
<paragraph pageId="5" pageNumber="146">Material examined.</paragraph>
|
||||
<paragraph pageId="5" pageNumber="146">
|
||||
<subSubSection id="2C278BDE3610283FED5904E17C79CA3E" pageId="5" pageNumber="146" type="material examined">
|
||||
<paragraph id="C39F9F539182BC377E4DBF917C90AF79" pageId="5" pageNumber="146">Material examined.</paragraph>
|
||||
<paragraph id="21FB5A63D9612DF74C904D507DF00723" pageId="5" pageNumber="146">
|
||||
Additional New Brunswick records. Queens Co., Cranberry Lake P.N.A,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="46.1125">46.1125°N</geoCoordinate>
|
||||
<geoCoordinate id="CD56A7A5552086DF0629D9F0485FA6BC" direction="north" orientation="latitude" precision="5" value="46.1125">46.1125°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-65.6075">65.6075°W</geoCoordinate>
|
||||
<geoCoordinate id="D3AFDB3DE1B94DD0EBC541E1E7EFA6C5" direction="west" orientation="longitude" precision="5" value="-65.6075">65.6075°W</geoCoordinate>
|
||||
, 21-27.V.2009, R. Webster & M.-A.
|
||||
<normalizedToken originalValue="Giguère">Giguere</normalizedToken>
|
||||
<normalizedToken id="BAA43AC6AD00C982357A7DC590EE3C37" originalValue="Giguère">Giguere</normalizedToken>
|
||||
, old red oak forest, Lindgren funnel trap (1, RWC); same locality and habitat data, 25.
|
||||
<normalizedToken originalValue="V–">V-</normalizedToken>
|
||||
<normalizedToken id="6D51A08A0F56B57B39B61D73ED7DB4A3" originalValue="V–">V-</normalizedToken>
|
||||
7.VI.2011, 29.
|
||||
<normalizedToken originalValue="VI–">VI-</normalizedToken>
|
||||
<normalizedToken id="10D97071B282E89BB725DAA7726DF8FC" originalValue="VI–">VI-</normalizedToken>
|
||||
7.VII.2011, M. Roy & V. Webster, Lindgren funnel traps (2, NBM, RWC). Sunbury Co., Acadia Research Forest, 11.VI.2008, Brawn/Harrison (2, AFC); same locality but
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.9866">45.9866°N</geoCoordinate>
|
||||
<geoCoordinate id="59F40C87ABB981728A34B05D54B1BC99" direction="north" orientation="latitude" precision="5" value="45.9866">45.9866°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.3841">66.3841°W</geoCoordinate>
|
||||
<geoCoordinate id="9945623DA910C9263D028C371F5F961A" direction="west" orientation="longitude" precision="5" value="-66.3841">66.3841°W</geoCoordinate>
|
||||
, 13-19.V.2009, 19-25.V.2009, 25.V-2.VI.2009, R. Webster & M.-A.
|
||||
<normalizedToken originalValue="Giguère">Giguere</normalizedToken>
|
||||
<normalizedToken id="D69783A91C2EBC8FF0F7204AF8318439" originalValue="Giguère">Giguere</normalizedToken>
|
||||
, mature (110-year-old) red spruce forest with scattered red maple and balsam fir, Lindgren funnel traps (3, AFC, RWC). York Co., 14 km WSW of Tracy, S of Rt. 645,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.6741">45.6741°N</geoCoordinate>
|
||||
<geoCoordinate id="BA58563B20563D57A9D085E31E75BB12" direction="north" orientation="latitude" precision="5" value="45.6741">45.6741°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.8661">66.8661°W</geoCoordinate>
|
||||
<geoCoordinate id="BED0BE96902AD2A461005876E88304E6" direction="west" orientation="longitude" precision="5" value="-66.8661">66.8661°W</geoCoordinate>
|
||||
, 10-26.V.2010, R. Webster & C. MacKay, old mixed forest with red and white spruce, red and white pine, balsam fir, eastern white cedar, red maple, and
|
||||
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="5" pageNumber="146" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
|
||||
<taxonomicName id="46CB34FDE0DA12434EBF1B7BE4D2BEB2" class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="5" pageNumber="146" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
|
||||
sp., Lindgren funnel trap (1, RWC); 15 km W of Tracy off Rt. 645,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.6848">45.6848°N</geoCoordinate>
|
||||
<geoCoordinate id="AA8199D861BD256548D61C46CC36AD1E" direction="north" orientation="latitude" precision="5" value="45.6848">45.6848°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.8821">66.8821°W</geoCoordinate>
|
||||
<geoCoordinate id="D1B28B3C9B9362795813B81B1B154FC4" direction="west" orientation="longitude" precision="5" value="-66.8821">66.8821°W</geoCoordinate>
|
||||
, 30.V-8.VI.2011, 8-20.VI.2011, M. Roy & V. Webster, old red pine forest, Lindgren funnel traps (6, AFC, NBM, RWC).
|
||||
</paragraph>
|
||||
<caption pageId="5" pageNumber="146">
|
||||
<paragraph pageId="5" pageNumber="146">
|
||||
<caption id="6D6F038FBA3BB97300D459A7DE893FD1" pageId="5" pageNumber="146">
|
||||
<paragraph id="A065CC3F58D4FCB47BC3FA39F044F7D4" pageId="5" pageNumber="146">
|
||||
Map 4. Collection localities in New Brunswick, Canada of
|
||||
<taxonomicName class="Insecta" family="Trogossitidae" genus="Calitys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Calitys scabra" order="Coleoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="scabra">Calitys scabra</taxonomicName>
|
||||
<taxonomicName id="FF2958C5683AAC7E5F3DE0C5DDDCE7DE" genus="Calitys" lsidName="Calitys scabra" pageId="5" pageNumber="146" rank="species" species="scabra">Calitys scabra</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="5" pageNumber="146" type="collection and habitat data">
|
||||
<paragraph pageId="5" pageNumber="146">Collection and habitat data.</paragraph>
|
||||
<paragraph pageId="5" pageNumber="146">
|
||||
<taxonomicName class="Insecta" family="Trogossitidae" genus="Calitys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Calitys scabra" order="Coleoptera" pageId="5" pageNumber="146" phylum="Arthropoda" rank="species" species="scabra">Calitys scabra</taxonomicName>
|
||||
<subSubSection id="EB2E9F4B7C35BBF4DD73F0B21BEA0C5C" pageId="5" pageNumber="146" type="collection and habitat data">
|
||||
<paragraph id="46E9E3083AD1033DBB2C59C736FB6DF3" pageId="5" pageNumber="146">Collection and habitat data.</paragraph>
|
||||
<paragraph id="985A1210DFEA9A5E1FAB93E74013CEE1" pageId="5" pageNumber="146">
|
||||
<taxonomicName id="17847230F7E7E06D5AF9F856A2E8BB3D" genus="Calitys" lsidName="Calitys scabra" pageId="5" pageNumber="146" rank="species" species="scabra">Calitys scabra</taxonomicName>
|
||||
was captured in Lindgren funnel traps deployed in an old red oak forest, a mature red spruce stand, an old-growth red pine forest, and an old mixed forest. This species was reported from under bark of dead pine and from
|
||||
<taxonomicName class="Agaricomycetes" family="Fomitopsidaceae" genus="Fomitopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Fomitopsis pinicola" order="Polyporales" pageId="5" pageNumber="146" phylum="Basidiomycota" rank="species" species="pinicola">Fomitopsis pinicola</taxonomicName>
|
||||
<taxonomicName id="8DD889E986B0EC0E27B5859F3ACB58FC" class="Agaricomycetes" family="Fomitopsidaceae" genus="Fomitopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Fomitopsis pinicola" order="Polyporales" pageId="5" pageNumber="146" phylum="Basidiomycota" rank="species" species="pinicola">Fomitopsis pinicola</taxonomicName>
|
||||
(Fr.) Kar. (
|
||||
<bibRefCitation author="Barron, JR" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="11" pageNumber="152" pagination="1 - 143" title="A revision of the Trogossitidae of America north of Mexico." url="10.4039/entm10375fv" volume="75" year="1971">Barron 1971</bibRefCitation>
|
||||
<bibRefCitation id="2B9A72790FD4624E2D4A9B04BB4994D9" author="Barron, JR" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="11" pageNumber="152" pagination="1 - 143" title="A revision of the Trogossitidae of America north of Mexico." url="10.4039/entm10375fv" volume="75" year="1971">Barron 1971</bibRefCitation>
|
||||
). Adults were collected during May, June, and July.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="5" pageNumber="146" type="distribution">
|
||||
<paragraph pageId="5" pageNumber="146">Distribution in Canada and Alaska.</paragraph>
|
||||
<paragraph pageId="5" pageNumber="146">
|
||||
<subSubSection id="B836FFF534E1F0CCC7A1B88AC1808903" pageId="5" pageNumber="146" type="distribution">
|
||||
<paragraph id="C2866C81DEF4E8375942908E23283AAB" pageId="5" pageNumber="146">Distribution in Canada and Alaska.</paragraph>
|
||||
<paragraph id="B6137390287AB426ACEEF7FC504F0B60" pageId="5" pageNumber="146">
|
||||
AK, NT, BC, AB, MB, ON, QC, NB, NS (
|
||||
<bibRefCitation pageId="5" pageNumber="146">Bousquet 1991</bibRefCitation>
|
||||
<bibRefCitation id="FA430A417C292D362BA7820FDCC87B3C" pageId="5" pageNumber="146">Bousquet 1991</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka 2011</bibRefCitation>
|
||||
<bibRefCitation id="808121CD9E26028DEC33D4AA864F18EF" author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka 2011</bibRefCitation>
|
||||
).
|
||||
<bibRefCitation author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka (2011)</bibRefCitation>
|
||||
<bibRefCitation id="665721079468B7A5E10D02414C65BC90" author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka (2011)</bibRefCitation>
|
||||
reported this Holarctic species for the first time from New Brunswick based on a specimen collected by W. McIntosh in Saint John, ca. 1900. The above records are the first recent records of this species for New Brunswick.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,70 +1,69 @@
|
|||
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<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,454,1018,1038]" pageId="55" pageNumber="56" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="noeli">Tetramorium noeli</taxonomicName>
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<treatment id="833881B6157525F6D0931E692B1E9CB1" ID-DOI="http://doi.org/10.5281/zenodo.6172637" ID-GBIF-Taxon="100105383" ID-Zenodo-Dep="6172637" LSID="urn:lsid:plazi:treatment:833881B6157525F6D0931E692B1E9CB1" httpUri="http://treatment.plazi.org/id/833881B6157525F6D0931E692B1E9CB1" lastPageNumber="56" pageNumber="56" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="63CA70425AA58330CC8B1A1FD1FC2CFE" type="nomenclature">
|
||||
<paragraph id="471E33C298B6EAAEE4EFE29E7F55561C" box="[118,454,1018,1038]" pageId="55" pageNumber="56" type="mainText">
|
||||
<taxonomicName id="89F941A951DF56DE8FCD25203773FA17" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="noeli">Tetramorium noeli</taxonomicName>
|
||||
species complex
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,1122,1074,1149]" pageId="55" pageNumber="56" type="mainText">
|
||||
<subSubSection id="C0E5A7365C3C0A6DEC8597D39C39CEA9" type="discussion">
|
||||
<paragraph id="CC5EBA79634516E0E5FA47E807BB443C" box="[118,1122,1074,1149]" pageId="55" pageNumber="56" type="mainText">
|
||||
This species complex contains the species
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="55" pageNumber="56" rank="species" species="ambanizana">
|
||||
<taxonomicName id="EE335F8A4AFAA21FA382C63F18AAABE0" authorityName="," genus="Tetamorium" pageId="55" pageNumber="56" rank="species" species="ambanizana">
|
||||
T. ambanizana
|
||||
<authority>,</authority>
|
||||
<authority id="FB7AE7C248F4F1BF826A5D62D1D34ABB">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="55" pageNumber="56" rank="species" species="aherni">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="aherni">
|
||||
<taxonomicName id="C5ABC156B1ABD54866DB7FE1F4B878A5" authorityName="," genus="Tetamorium" pageId="55" pageNumber="56" rank="species" species="aherni">
|
||||
<taxonomicName id="8774EE11160155F2A950E184558ECBBF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="aherni">
|
||||
T. aherni
|
||||
<authority>,</authority>
|
||||
<authority id="575E17A6E2F3E12D2126779C2EDC76D6">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae,</taxonomicName>
|
||||
<taxonomicName id="063690219393C27AE920458400DD835D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae,</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="noeli">T. noeli.</taxonomicName>
|
||||
<taxonomicName id="F057CCF8DC6B941531646432097858CD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="noeli">T. noeli.</taxonomicName>
|
||||
The characters that define the complex are the lack of sculpture on the forecoxae and the presence of basigastral costulae on the first gastral tergite.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,1158,1205]" pageId="55" pageNumber="56" type="mainText">The four species of this complex appear to be morphologically close to each other, and it is likely that they represent a natural group with shared common ancestry.</paragraph>
|
||||
<paragraph id="2922F86EAB3AF619E8C9BB92A5C2F541" box="[118,1121,1158,1205]" pageId="55" pageNumber="56" type="mainText">The four species of this complex appear to be morphologically close to each other, and it is likely that they represent a natural group with shared common ancestry.</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,166 +1,165 @@
|
|||
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<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="nify">
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<taxonomicName id="80913A7CD4106330721F14E5F3123E9C" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="nify">
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Tetramorium nify
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<authority>Hita</authority>
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<authority id="7438DF459407A45000B5C652F7ECC526">Hita</authority>
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Garcia & Fisher
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<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
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<taxonomicNameLabel id="A8AB4CFCE8C9488B296DA69DA3EDE460" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection type="description">
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<paragraph box="[118,366,792,811]" pageId="41" pageNumber="42" type="mainText">(Figs. 61, 68, 69, 96, 97, 98)</paragraph>
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|
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<paragraph id="4C4709ACD4A5B70BF54112F282DA0662" box="[118,366,792,811]" pageId="41" pageNumber="42" type="mainText">(Figs. 61, 68, 69, 96, 97, 98)</paragraph>
|
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</subSubSection>
|
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<subSubSection type="materials_examined">
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<paragraph box="[118,1121,849,952]" pageId="41" pageNumber="42" type="mainText">
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<subSubSection id="918E07A5BE9E3FB3DFB3BC96192DC675" type="materials_examined">
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Holotype worker, MADAGASCAR, Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
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<normalizedToken id="D3EE8FAE89DEE4F0F3E5415EEDAFFEAF" originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken id="9E835AA8802C8A983CFA6226BDB66445" originalValue="Spéciale">Speciale</normalizedToken>
|
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Ambatovaky, Sandrangato River, 16.81753 S, 49.29498 E, 360 m, rainforest, ex rotten log, collection code BLF24814, 25.II.2010 (B.L. Fisher et al.) (CASC: CASENT0163155). Paratype, one worker with same data as holotype (CASC: CASENT0163345).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,989,1008]" pageId="41" pageNumber="42" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1120,1017,1121]" pageId="41" pageNumber="42" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="nify">Tetramorium nify</taxonomicName>
|
||||
<subSubSection id="2607E10A31E957037C2C95DD6731CBDF" type="diagnosis">
|
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<paragraph id="194C8B5E67754485FF9ECF7B296C775D" box="[118,208,989,1008]" pageId="41" pageNumber="42" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="BFB6AE75109E154096F91534DCFEBDDD" box="[118,1120,1017,1121]" pageId="41" pageNumber="42" type="mainText">
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<taxonomicName id="32229C9D5472D6CEE022BE2E69658B57" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="nify">Tetramorium nify</taxonomicName>
|
||||
can be clearly distinguished from the other species of the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="B9843D751244B8E380C0956008D3D04F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
complex by the following character combination: small eyes (OI 16-17); long propodeal spines (PSLI 29-30); posterodorsal corner of petiolar node not strongly protruding posteriorly; body very dark brown to black in colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,1158,1176]" pageId="41" pageNumber="42" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1117,1186,1346]" pageId="41" pageNumber="42" type="mainText">HL 0.83-0.95 (0.87); HW 0.82-0.93 (0.86); SL 0.60-0.66 (0.63); EL 0.14-0.15 (0.14); PH 0.41-0.51 (0.44); PW 0.61-0.72 (0.65); WL 1.02-1.21 (1.08); PSL 0.25-0.28 (0.26); PTL 0.30-0.37 (0.33); PTH 0.32-0.38 (0.35); PTW 0.26-0.31 (0.28); PPL 0.29-0.33 (0.31); PPH 0.32-0.37 (0.34); PPW 0.33-0.38 (0.35); CI 98-100 (99); SI 69-74 (73); OI 16-17 (17); DMI 59-63 (60); LMI 39-42 (41); PSLI 29-30 (30); PeNI 43-44 (43); LPeI 91-100 (96); DPeI 82-88 (84); PpNI 52-54 (53); LPpI 88-91 (90); DPpI 112-116 (113); PPI 121-125 (124) (five measured).</paragraph>
|
||||
<paragraph box="[118,1119,1355,1575][118,1120,118,540]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="mainText">
|
||||
<subSubSection id="4F6A9F5C729F77059E42F3B2BCD7B6B6" type="description">
|
||||
<paragraph id="A0B3E044753A4BFBA502E89D0789AD59" box="[118,226,1158,1176]" pageId="41" pageNumber="42" type="mainText">Description</paragraph>
|
||||
<paragraph id="5967EA522F33ACE4F1CFC3211E9FA8AB" box="[118,1117,1186,1346]" pageId="41" pageNumber="42" type="mainText">HL 0.83-0.95 (0.87); HW 0.82-0.93 (0.86); SL 0.60-0.66 (0.63); EL 0.14-0.15 (0.14); PH 0.41-0.51 (0.44); PW 0.61-0.72 (0.65); WL 1.02-1.21 (1.08); PSL 0.25-0.28 (0.26); PTL 0.30-0.37 (0.33); PTH 0.32-0.38 (0.35); PTW 0.26-0.31 (0.28); PPL 0.29-0.33 (0.31); PPH 0.32-0.37 (0.34); PPW 0.33-0.38 (0.35); CI 98-100 (99); SI 69-74 (73); OI 16-17 (17); DMI 59-63 (60); LMI 39-42 (41); PSLI 29-30 (30); PeNI 43-44 (43); LPeI 91-100 (96); DPeI 82-88 (84); PpNI 52-54 (53); LPpI 88-91 (90); DPpI 112-116 (113); PPI 121-125 (124) (five measured).</paragraph>
|
||||
<paragraph id="AFF4C08DEA4DC58953C28EA78B09F36F" box="[118,1119,1355,1575][118,1120,118,540]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="mainText">
|
||||
Head weakly longer than wider to as long as wide (CI 98-100). Posterior had margin weakly concave. Anterior clypeal margin weakly medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes moderately developed, but shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short, not reaching posterior head margin (SI 69-74). Eyes very small (OI 16-17). Mesosomal outline in profile flat, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 39-42). Propodeal spines with very broad base, up-curved, elongate-triangular, and moderately long (PSLI 29-30); propodeal lobes well-developed, triangular and acute. Petiolar node in profile
|
||||
<pageBreakToken pageId="42" pageNumber="43" start="start">rectangular</pageBreakToken>
|
||||
<pageBreakToken id="342696BE427BC2F7E53696554099D224" pageId="42" pageNumber="43" start="start">rectangular</pageBreakToken>
|
||||
nodiform with well-defined angles, around 1.0 to 1.1 times higher than long (LPeI 91-100), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum straight; node in dorsal view approximately 1.2 times longer than wide (DPeI 82-88). Postpetiole in profile globular, approximately 1.1 times higher than long (LPpI 88-91); in dorsal view around 1.1 to 1.2 times wider than long (112-116). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view approximately 1.2 times wider than petiolar node (PPI 121-125). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose, with three to five rugae; cephalic dorsum between frontal carinae with 8 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, few rugae interrupted or with cross-meshes; lateral and ventral head longitudinally rugose, rarely with cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured. Waist segments strongly irregularly longitudinally rugose. Gaster completely unsculptured, smooth and shining. Ground sculpture generally faint to absent everywhere on body. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. First gastral tergite without appressed pubescence. Anterior edges of antennal scapes with erect, standing hairs. Body uniform dark brown to black in colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,578,596]" pageId="42" pageNumber="43" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1044,607,626]" pageId="42" pageNumber="43" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="nify">Tetramorium nify</taxonomicName>
|
||||
<subSubSection id="14819AFACAAE8E3432E8E940C0D832C5" type="discussion">
|
||||
<paragraph id="30260A9EC8889A15A7B8CA07D869CFBA" box="[118,170,578,596]" pageId="42" pageNumber="43" type="mainText">Notes</paragraph>
|
||||
<paragraph id="F80DB78DA978624EEB690ADB4EBF2EA7" box="[118,1044,607,626]" pageId="42" pageNumber="43" type="mainText">
|
||||
<taxonomicName id="23F8E57FD09BB484E4C7CDD44C9F0218" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="nify">Tetramorium nify</taxonomicName>
|
||||
is only known from five specimens from Ambatovaky, Befingotra, and Isle Saint Marie.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,636,971]" pageId="42" pageNumber="43" type="mainText">
|
||||
<paragraph id="94ABFCFE61678839A48E937F45F9CA35" box="[118,1122,636,971]" pageId="42" pageNumber="43" type="mainText">
|
||||
Within the species complex,
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="4958EBE4FBFDEA2976F605DA6735E1D3" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
is another species with fairly small eyes (OI 16-17). The other species with comparatively small eyes are
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="electrum">T. elf,</taxonomicName>
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="07B01F44E2F277D13AC582A3A50B5501" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="electrum">T. elf,</taxonomicName>
|
||||
<taxonomicName id="BEE6E13908AA5B9F8B3820B2AE707BF2" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="8FE0A9FB814B1A525B2867B55FD53CB9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="electrum">
|
||||
T. electrum
|
||||
<authority>,</authority>
|
||||
<authority id="3365C1ECBFAA2E1FCE580A8D1BDCDBA5">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
<taxonomicName id="75F16D1E42C81B3B99912683625C9F8A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="isectum">T. isectum.</taxonomicName>
|
||||
<taxonomicName id="313989D6F26B38F2072A554E7BD1C43C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="isectum">T. isectum.</taxonomicName>
|
||||
The first three have very long to extremely long propodeal pines (PSLI 46-64), short and blunted propodeal lobes, and differently shaped petiolar nodes.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
<taxonomicName id="5986F9CB301EFBAF2C619C33B8DC551F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
has even smaller eyes (OI 15-16) than
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify,</taxonomicName>
|
||||
<taxonomicName id="4D46A6962C2A33B0622D975A38CF9CAA" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify,</taxonomicName>
|
||||
a petiolar node with the anterodorsal margin situated higher than the posterodorsal and the dorsum weakly tapering backwards, and bright orange body colour, whereas
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="0D1313706E10E6E2741D766721F9BB34" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
is very dark brown to black in colour and has anterodorsal and posterodorsal margins of the petiolar node at about the same height. Disregarding eye size,
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="CF6DCEF09E637CA88DCC2C177106433B" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
also cannot be confused with T.
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="ala">
|
||||
<taxonomicName id="AE327FD421C2AF62B5DE6528A939B3B5" authorityName="," genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="ala">
|
||||
ala
|
||||
<authority>,</authority>
|
||||
<authority id="E94CFCC13BC2E8F406B7DC72D0D3BD52">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
<taxonomicName id="709DC791F2ADBA20FCD6572C608663A9" genus="Tetamorium" pageId="42" pageNumber="43" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="ED826B44C21CCAEE1593D0455F5715DC" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
, or
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="voasary">T. voasary.</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andohahela">Tetramorium andohahela</taxonomicName>
|
||||
<taxonomicName id="657E01A4D768AA01189093AD744B4B5F" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="voasary">T. voasary.</taxonomicName>
|
||||
<taxonomicName id="DC138A9FB3B49825CE49B89C1A41FB52" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andohahela">Tetramorium andohahela</taxonomicName>
|
||||
has the posterodorsal corner of the petiolar node strongly protruding posteriorly, T. ala has the posterior corners of the head weakly angular and marginate, and
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
<taxonomicName id="CE8C080B4D629276446EF2CC3DF561AC" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
has a petiolar node with the anterodorsal and posterodorsal angles fairly rounded. The defining characters of the first two are absent in
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify,</taxonomicName>
|
||||
<taxonomicName id="2524D47D66F5D952DA0EF5F0509E365B" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify,</taxonomicName>
|
||||
and the petiole of the latter has a node with well-defined anterodorsal and posterodorsal angles, which separates it from
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="voasary">T. voasary.</taxonomicName>
|
||||
<taxonomicName id="FAB1EE2769383CC024D04E34195D4538" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="voasary">T. voasary.</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,981,1230]" pageId="42" pageNumber="43" type="mainText">
|
||||
<paragraph id="AAC6F56C0EFD36123249042756E7FCEA" box="[118,1122,981,1230]" pageId="42" pageNumber="43" type="mainText">
|
||||
The last species of the complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
<taxonomicName id="B317CB101BE2F76222557B99E16F2562" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
is morphologically very close to
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify,</taxonomicName>
|
||||
<taxonomicName id="0EC6CC8EFC045025D7B76EDF7A4A36DD" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify,</taxonomicName>
|
||||
and we have treated the material as conspecific for a while during this revision. The only good diagnostic character that divides them is eye size, which is much larger in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="EA6608A0B185CD96471F2381A690C841" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
(OI 19-25, usually above 20). However, we observed both in sympatry in Ambatovaky and Befingotra and specimens of
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="2308E2CC163BA7F22E69D7AA25F9FACB" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
are easily separable from sympatric
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="C45FBDAF2E914E19CB7949F8B6A01A3D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
due to their fairly small eyes and a petiolar node with sharper defined antero- and posterodorsal margins. The mesosoma of
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="A7A0A7430D6E155C90CC4048AE94834B" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
is also a bit shorter (DMI 59-63) than the one of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="FEAA059BB5C3498F33701B8DD12F9445" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
(DMI 50-60). As noted in its description,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="0186BF3DF3786944BAEE8A7DA46814BE" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
is a remarkably variable species, which very likely includes several cryptic species. However, especially on the basis of the co-occurrence without intermediate forms, we treat
|
||||
<taxonomicName genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="6690A8DE617AC6B4774246B734E8B888" genus="Tetamorium" pageId="42" pageNumber="43" rank="species" species="nify">T. nify</taxonomicName>
|
||||
as its own species distinct from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
<taxonomicName id="8FD765F4160F8AB20B4D8E30AB4F170F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,1268,1286]" pageId="42" pageNumber="43" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,615,1297,1316]" pageId="42" pageNumber="43" type="mainText">The species epithet is an arbitrary combination of letters.</paragraph>
|
||||
<subSubSection id="FFAFE4F65E0D6DE2849AC38328043449" type="etymology">
|
||||
<paragraph id="BDE411660183E7BFEAD80D94A5A5E019" box="[118,219,1268,1286]" pageId="42" pageNumber="43" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="A44BD1F2316D1AE46415831A879EE191" box="[118,615,1297,1316]" pageId="42" pageNumber="43" type="mainText">The species epithet is an arbitrary combination of letters.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,1354,1373]" pageId="42" pageNumber="43" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,1383,1517]" pageId="42" pageNumber="43" type="mainText">
|
||||
<subSubSection id="F4125CB8EF096F67FACEB46D40F3204D" type="materials_examined">
|
||||
<paragraph id="13091632FEA495768B73A86DE782F057" box="[118,295,1354,1373]" pageId="42" pageNumber="43" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="4B00AB0ECDD53EE1D4E52924DC424809" box="[118,1122,1383,1517]" pageId="42" pageNumber="43" type="mainText">
|
||||
MADAGASCAR: Antsiranana,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="2952C5437CB551D8739967A3B228E1D1" originalValue="Rés">Res</normalizedToken>
|
||||
. Anjanaharibe-Sud, 6.5 km SSW Befingotra, 14.75 S, 49.5 E, 875 m, rainforest, 18.-22.X.1994 (B.L. Fisher); Toamasina, Ile Sainte Marie,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="559127C1F32904C9EBE9893B0E464799" originalValue="Forêt">Foret</normalizedToken>
|
||||
Kalalao, 9.9 km 34° Ambodifotatra, 16.9225 S, 49.88733 E, 100 m, rainforest, 24.-27.XI.2005 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="DB63FC1F106DA7E5EDF626D29925A4A6" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="3DD28A6228F9D2F6ED9128EB55E30F88" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato River, 16.81753 S, 49.29498 E, 360 m, rainforest, 25.II.2010 (B.L. Fisher et al.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,128 +1,127 @@
|
|||
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|
||||
<mods:title id="CC986067933504B05CC6BC2487EBC8B2">Zootaxa</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:part>
|
||||
<mods:date>2012</mods:date>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>3592</mods:number>
|
||||
<mods:part id="6651A9349CBDD8CA067177228A22E62E">
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<mods:date id="42D6566EF7EF428D43F36125B23CFB10">2012</mods:date>
|
||||
<mods:detail id="7C91D8AE59F797AC2974DB2E076DAA30" type="volume">
|
||||
<mods:number id="FEAE815674EF173DFCBE00044D1247CE">3592</mods:number>
|
||||
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|
||||
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|
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<mods:start>1</mods:start>
|
||||
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|
||||
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<mods:start id="4474657367CAFD59372C2466AF43B07B">1</mods:start>
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<mods:end id="78E4BC2A8984F125E8D86E3604380EA9">85</mods:end>
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<mods:location id="64D535D79239251CCDDD85A4268139B8">
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<mods:url id="8DC8C79BB50550D112FA0C788CEF48F8">http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="HNS-Pub">26064</mods:identifier>
|
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<mods:identifier type="ISSN">175-5326</mods:identifier>
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<mods:identifier type="ZooBank">A2D9C9ED-C0BA-4B5F-A330-C9AB7D625704</mods:identifier>
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<mods:classification id="1F838E107254F5F151E9B6AFF502FD9B">journal article</mods:classification>
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<mods:identifier id="ECC930F6556CBF73A02388AEA3449EE4" type="HNS-Pub">26064</mods:identifier>
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<mods:identifier id="517851E362824FA4878DD8375E2ED3ED" type="ISSN">175-5326</mods:identifier>
|
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</mods:mods>
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||||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172657" ID-GBIF-Taxon="100105373" ID-Zenodo-Dep="6172657" LSID="urn:lsid:plazi:treatment:90D3E00B8BE86588B3C7DE12E6A4C44B" httpUri="http://treatment.plazi.org/id/90D3E00B8BE86588B3C7DE12E6A4C44B" lastPageNumber="72" pageNumber="71">
|
||||
<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,601,906,926]" pageId="70" pageNumber="71" type="mainText">
|
||||
<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="marojejy">
|
||||
<treatment id="90D3E00B8BE86588B3C7DE12E6A4C44B" ID-DOI="http://doi.org/10.5281/zenodo.6172657" ID-GBIF-Taxon="100105373" ID-Zenodo-Dep="6172657" LSID="urn:lsid:plazi:treatment:90D3E00B8BE86588B3C7DE12E6A4C44B" httpUri="http://treatment.plazi.org/id/90D3E00B8BE86588B3C7DE12E6A4C44B" lastPageNumber="72" pageNumber="71" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="222531F904D1D473F4BEA04935F91988" type="nomenclature">
|
||||
<paragraph id="09C231EC9C6021B26E509157E6E063E2" box="[118,601,906,926]" pageId="70" pageNumber="71" type="mainText">
|
||||
<taxonomicName id="7E97517516FDCA4F8FC5B1F5E770F2D5" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="marojejy">
|
||||
Tetramorium marojejy
|
||||
<authority>Hita</authority>
|
||||
<authority id="C862E0F18942BD56AF5B10B18BF544F8">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="223747A3CE76EC26EB1DAE9F171ABB3B" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,443,933,952]" pageId="70" pageNumber="71" type="mainText">(Figs. 18, 21, 22, 129, 130, 131, 142)</paragraph>
|
||||
<subSubSection id="A256A801FA7E2EED1E3C04DA1B99C390" type="description">
|
||||
<paragraph id="E0EB6EA506AFBB2CE52D4AF9DE406A1D" box="[118,443,933,952]" pageId="70" pageNumber="71" type="mainText">(Figs. 18, 21, 22, 129, 130, 131, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1121,989,1121]" pageId="70" pageNumber="71" type="mainText">Holotype worker, MADAGASCAR, Antsiranana, Parc National de Marojejy, Antranohofa, 26.6 km 31° NNE Andapa, 10.7 km 318° NW Manantenina, 14.44333 S, 49.74333 E, 1325 m, montane rainforest, canopy moss and leaf litter, collection code BLF09192, 19.XI.2003 (B.L. Fisher) (CASC: CASENT0247334). Paratypes, 8 workers with same data as holotype (CASC: CASENT0247327; CASENT0247328; CASENT0247329; CASENT0247330; CASENT0247333; CASENT0499782; CASENT0499783; CASENT0499784).</paragraph>
|
||||
<subSubSection id="C0F79EA08AFCDE8733E5EAD47B938D98" type="materials_examined">
|
||||
<paragraph id="263CEDCB9CA7456A5FD3BB2C674DA9D8" box="[118,1121,989,1121]" pageId="70" pageNumber="71" type="mainText">Holotype worker, MADAGASCAR, Antsiranana, Parc National de Marojejy, Antranohofa, 26.6 km 31° NNE Andapa, 10.7 km 318° NW Manantenina, 14.44333 S, 49.74333 E, 1325 m, montane rainforest, canopy moss and leaf litter, collection code BLF09192, 19.XI.2003 (B.L. Fisher) (CASC: CASENT0247334). Paratypes, 8 workers with same data as holotype (CASC: CASENT0247327; CASENT0247328; CASENT0247329; CASENT0247330; CASENT0247333; CASENT0499782; CASENT0499783; CASENT0499784).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,1158,1176]" pageId="70" pageNumber="71" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1122,1186,1290]" pageId="70" pageNumber="71" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="marojejy">Tetramorium marojejy</taxonomicName>
|
||||
<subSubSection id="84428417AEC22B26EB702CB39DC80645" type="diagnosis">
|
||||
<paragraph id="67636C0B101B8EAC05EFBD3405B59589" box="[118,208,1158,1176]" pageId="70" pageNumber="71" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="D06C8AE6CA2F9FF51E52FC8997ABF012" box="[118,1122,1186,1290]" pageId="70" pageNumber="71" type="mainText">
|
||||
<taxonomicName id="33A2E658E7E88F80492911994D94FBFE" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="marojejy">Tetramorium marojejy</taxonomicName>
|
||||
can be easily distinguished from the remainder of the species complex by the following character combination: antennal scapes short to moderate (SI 79-85); propodeal spines long to very long (PSLI 34-37); anterodorsal and posterodorsal margins of petiolar node situated at about same height; first gastral tergite with numerous standing hairs; uniform orange to pale brown body colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,1327,1345]" pageId="70" pageNumber="71" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1117,1355,1515]" pageId="70" pageNumber="71" type="mainText">HL 1.07-1.14 (1.11); HW 1.00-1.07 (1.05); SL 0.82-0.88 (0.85); EL 0.23-0.25 (0.24); PH 0.50-0.59 (0.54); PW 0.72-0.78 (0.75); WL 1.36-1.49 (1.42); PSL 0.36-0.41 (0.40); PTL 0.35-0.37 (0.36); PTH 0.37-0.41 (0.39); PTW 0.27-0.31 (0.29); PPL 0.35-0.37 (0.36); PPH 0.37-0.41 (0.39); PPW 0.36-0.39 (0.37); CI 91-96 (94); SI 79-85 (81); OI 22-24 (23); DMI 52-54 (53); LMI 37-39 (38); PSLI 34-37 (36); PeNI 37-39 (39); LPeI 89-97 (93); DPeI 77-84 (80); PpNI 49-51 (50); LPpI 88-95 (91); DPpI 101-109 (105); PPI 125-135 (129) (12 measured).</paragraph>
|
||||
<paragraph box="[118,1120,1524,1571][118,1120,118,700]" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="mainText">
|
||||
<subSubSection id="549EDD49C5A956B75C489B1215C61328" type="description">
|
||||
<paragraph id="5D878D481ACBF5DBED926C1BBDEA618A" box="[118,226,1327,1345]" pageId="70" pageNumber="71" type="mainText">Description</paragraph>
|
||||
<paragraph id="FFA448AC2110EF7EB6C0E8BD5AC8DB40" box="[118,1117,1355,1515]" pageId="70" pageNumber="71" type="mainText">HL 1.07-1.14 (1.11); HW 1.00-1.07 (1.05); SL 0.82-0.88 (0.85); EL 0.23-0.25 (0.24); PH 0.50-0.59 (0.54); PW 0.72-0.78 (0.75); WL 1.36-1.49 (1.42); PSL 0.36-0.41 (0.40); PTL 0.35-0.37 (0.36); PTH 0.37-0.41 (0.39); PTW 0.27-0.31 (0.29); PPL 0.35-0.37 (0.36); PPH 0.37-0.41 (0.39); PPW 0.36-0.39 (0.37); CI 91-96 (94); SI 79-85 (81); OI 22-24 (23); DMI 52-54 (53); LMI 37-39 (38); PSLI 34-37 (36); PeNI 37-39 (39); LPeI 89-97 (93); DPeI 77-84 (80); PpNI 49-51 (50); LPpI 88-95 (91); DPpI 101-109 (105); PPI 125-135 (129) (12 measured).</paragraph>
|
||||
<paragraph id="51ECBD51AD6E722D7BC267680A79822F" box="[118,1120,1524,1571][118,1120,118,700]" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="mainText">
|
||||
Head longer than wide (CI 91-96); posterior head margin distinctly concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior
|
||||
<pageBreakToken pageId="71" pageNumber="72" start="start">head</pageBreakToken>
|
||||
<pageBreakToken id="6CF3F6C7E15DB44F0562C8B79813DD23" pageId="71" pageNumber="72" start="start">head</pageBreakToken>
|
||||
margin. Antennal scrobes developed, but very shallow and narrow, without defined posterior and ventral margins. Antennal scapes of moderate length, not reaching posterior head margin (SI 79-85). Eyes of moderate size (OI 22-24). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-39). Propodeal spines long, spinose and acute (PSLI 34-37); propodeal lobes well-developed, triangular to elongate-triangular, and acute. Petiolar node in profile rectangular nodiform, approximately 1.0 to 1.1 times higher than long (LPeI 89-97), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins situated at about same height, dorsum flat to weakly convex; node in dorsal view approximately 1.2 to 1.3 times longer than wide (DPeI 77-84). Postpetiole in profile globular, approximately 1.1 times higher than long (LPpI 88-95); in dorsal view around 1.0 to 1.1 times wider than long (DPpI 101-109). Postpetiole in profile approximately as high as petiolar node and generally appearing slightly less voluminous, in dorsal view approximately 1.2 to 1.4 times wider than petiolar node (PPI 125-135). Mandibles striate; clypeus longitudinally rugose, with four to six rugae, median ruga always present and distinct, remaining rugae variably developed; cephalic dorsum between frontal carinae with 9 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted, rarely with cross-meshes; lateral and ventral head longitudinally rugose to reticulate-rugose. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae with well-developed and conspicuous longitudinal rugae. Waist segments strongly longitudinally rugose. Gaster unsculptured, smooth, and shining. Ground sculpture generally faint to absent everywhere on body. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with suberect to erect hairs. Body of uniform orange to pale brown colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,737,755]" pageId="71" pageNumber="72" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1122,765,841]" pageId="71" pageNumber="72" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">Tetramorium marojejy</taxonomicName>
|
||||
<subSubSection id="A5692C064ED928D62FDD70D7AA0DD639" type="discussion">
|
||||
<paragraph id="D1BE63F9AE625792458FDB6A44C6CFCB" box="[118,170,737,755]" pageId="71" pageNumber="72" type="mainText">Notes</paragraph>
|
||||
<paragraph id="6DA0CBA593B7237C4961BC49F9B56614" box="[118,1122,765,841]" pageId="71" pageNumber="72" type="mainText">
|
||||
<taxonomicName id="BA2E85A5793B46A8FB6BA1D5F37288D6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">Tetramorium marojejy</taxonomicName>
|
||||
is currently only known from two localities: the type locality Marojejy and Anjanaharibe-Sud. Both localities are montane rainforests located in north-eastern Madagascar at altitudes of 1200 to 1325 m. In addition, the
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="E986A6CB203E641EA6FA83190326EC5A" rank="species">new species</taxonomicNameLabel>
|
||||
was mainly collected from leaf litter.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,850,1206]" pageId="71" pageNumber="72" type="mainText">
|
||||
<paragraph id="4F80E73D13FD5C106A92492CD8FB6244" box="[118,1122,850,1206]" pageId="71" pageNumber="72" type="mainText">
|
||||
Inside the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="D59ED038B6863E95CEB3EEE4BD69E7F9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
complex the orange to pale brownish
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
<taxonomicName id="094C39E86ACC3AC8A67AAE2D076C0262" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
is the only species without a dark brown to blackish body colouration, and thus is easil y recognisable. However, not considering body colour, the species differs from
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
<taxonomicName id="31B4802F8C74215A3F12326F7D34A770" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
T. latreillei
|
||||
<authority>,</authority>
|
||||
<authority id="2B7BC91345EEB222647EA4396DACE049">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
<taxonomicName id="0684121386B8132F506E9AE363538F86" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="6EE6B201D6B5F102C686F29B2C68D198" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
in the development of the propodeal spines and pilosity. The latter three species all have massive and extremely long propodeal spines (PSLI 48-72) and have either no, few, or several scattered standing hairs on the first gastral tergite, whereas
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
<taxonomicName id="2B82AE6A0751C7B61BECBEB075CF0F76" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
has much less extremely developed propodeal spines (PSLI 34-37) and numerous standing hairs on the first gastral tergite.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">Tetramorium marojejy</taxonomicName>
|
||||
<taxonomicName id="3CA8B9970F85C97A205DA531886AA203" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">Tetramorium marojejy</taxonomicName>
|
||||
also cannot be confused with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
<taxonomicName id="EAC9E008E3B1F4B71E3D8BF59C2804E1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
since the latter species possesses a petiolar node with the posterodorsal margin situated higher than the anterodorsal while both margins are at about the same height in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy.</taxonomicName>
|
||||
<taxonomicName id="8500997843E9F7A3D1E410E3AC1744CD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy.</taxonomicName>
|
||||
The remaining species,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul,</taxonomicName>
|
||||
<taxonomicName id="247738E5A8FCCB9A938872E29CEC0B59" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul,</taxonomicName>
|
||||
is dark brown to black in colour and therefore is also unlikely to be misidentified with the orange to pale brown
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy.</taxonomicName>
|
||||
<taxonomicName id="7C5E689F9C23FB390DA89361210742A3" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy.</taxonomicName>
|
||||
The latter has shorter antennal scapes (SI 79-85), a comparatively lower petiolar node (LPeI 89-97), and a lower and narrower postpetiole (LPpI 88-95; DPpI 101-109) than
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
<taxonomicName id="9A90D766571AC35F577ACF214238F906" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
(SI 89-92; LPeI 81-88; LPpI 78-85; DPpI 113-122).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,1243,1261]" pageId="71" pageNumber="72" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,1122,1272,1347]" pageId="71" pageNumber="72" type="mainText">
|
||||
<subSubSection id="2C3719B6AE6E2731AC9511FD0BEE3D93" type="etymology">
|
||||
<paragraph id="31129124CD9918AD6C4E11548DA3939E" box="[118,219,1243,1261]" pageId="71" pageNumber="72" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="DCDB46A898F4315CC2AFE3AEED6B8CBE" box="[118,1122,1272,1347]" pageId="71" pageNumber="72" type="mainText">
|
||||
This
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="A7AA49AB48159E5C59F9BC4AF50288F2" rank="species">new species</taxonomicNameLabel>
|
||||
is named after the type locality, the Park National de Marojejy, which is one of the most important places for the conservation of biodiversity in Madagascar due to the extraordinarily high diversity of landscapes, habitats, fauna and flora. The species epithet is a noun in apposition, and thus invariant.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,1384,1402]" pageId="71" pageNumber="72" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,1412,1516]" pageId="71" pageNumber="72" type="mainText">
|
||||
<subSubSection id="D9326277A489DD2F80552BBB825B4057" type="materials_examined">
|
||||
<paragraph id="AAED29074E3AC17B14849F49FEC5245D" box="[118,295,1384,1402]" pageId="71" pageNumber="72" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="0E9BF51CC36D075B9C7D08FDFA912416" box="[118,1122,1412,1516]" pageId="71" pageNumber="72" type="mainText">
|
||||
MADAGASCAR: Antsiranana, 9.2 km WSW Befingotra,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="D31E7D5541B6921B22AF8294036E7DA2" originalValue="Rés">Res</normalizedToken>
|
||||
. Anjanaharibe-Sud, 14.75 S, 49.46667 E, 1200-1260 m, montane rainforest, 9.-11.XI.1994 (B.L. Fisher); Antsiranana, Parc National de Marojejy, Antranohofa, 26.6 km 31° NNE Andapa, 10.7 km 318° NW Manantenina, 14.44333 S, 49.74333 E, 1325 m, montane rainforest, 19.XI.2003 (B.L. Fisher).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,137 +1,136 @@
|
|||
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<subSubSection type="nomenclature">
|
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|
||||
<taxonomicName authorityName="voasary Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
|
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<pageBreakToken pageId="44" pageNumber="45" start="start">Tetramorium</pageBreakToken>
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<authority>voasary Hita</authority>
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|
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|
||||
<taxonomicName id="F3FE931467BC0646FC34AE3BD013022C" ID-CoL="63V54" authorityName="voasary Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
|
||||
<pageBreakToken id="834243536631A742285C7D8764906C2B" pageId="44" pageNumber="45" start="start">Tetramorium</pageBreakToken>
|
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<authority id="B33DE3A86AEC73F2DD8CADF63C8B4247">voasary Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="A28D048C279A2BEB332CC3BB2FADADAE" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,432,146,165]" pageId="44" pageNumber="45" type="mainText">(Figs. 60, 71, 72, 99, 100, 101, 141)</paragraph>
|
||||
<subSubSection id="0D96651A6DD918B45A69091C37969D38" type="description">
|
||||
<paragraph id="D9F837694F2BA059FEBF6188072B7E8D" box="[118,432,146,165]" pageId="44" pageNumber="45" type="mainText">(Figs. 60, 71, 72, 99, 100, 101, 141)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1121,202,361]" pageId="44" pageNumber="45" type="mainText">Holotype worker, MADAGASCAR, Toamasina, Montagne d'Anjanaharibe, 19.5 km 27° NNE Ambinanitelo, 15.17833 S, 49.635 E, 1100 m, montane rainforest, canopy moss and leaf litter, collection code BLF08213, 12.-16.III.2003 (B.L. Fisher, C. Griswold et al.) (CASC: CASENT0247162). Paratypes, 11 workers with same data as holotype (CASC: CASENT0497903; CASENT0497904; CASENT0497905; CASENT0497906); and one worker with same data as holotype except sampled from beating low vegetation and collection code BLF08151 (CASC: CASENT0489080).</paragraph>
|
||||
<subSubSection id="F2A157D14B437E7E52EB66F328981D25" type="materials_examined">
|
||||
<paragraph id="A71ADC2072FFA43D81A74C4B800AAC24" box="[118,1121,202,361]" pageId="44" pageNumber="45" type="mainText">Holotype worker, MADAGASCAR, Toamasina, Montagne d'Anjanaharibe, 19.5 km 27° NNE Ambinanitelo, 15.17833 S, 49.635 E, 1100 m, montane rainforest, canopy moss and leaf litter, collection code BLF08213, 12.-16.III.2003 (B.L. Fisher, C. Griswold et al.) (CASC: CASENT0247162). Paratypes, 11 workers with same data as holotype (CASC: CASENT0497903; CASENT0497904; CASENT0497905; CASENT0497906); and one worker with same data as holotype except sampled from beating low vegetation and collection code BLF08151 (CASC: CASENT0489080).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,398,417]" pageId="44" pageNumber="45" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,427,558]" pageId="44" pageNumber="45" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="voasary">Tetramorium voasary</taxonomicName>
|
||||
<subSubSection id="C6C80EA35EF82C87A2BAD7627D06728B" type="diagnosis">
|
||||
<paragraph id="3EC4CC677CE8AD23CC79BCD2E82BCF34" box="[118,208,398,417]" pageId="44" pageNumber="45" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="B0A661765F5481AF435A946E9B00E507" box="[118,1121,427,558]" pageId="44" pageNumber="45" type="mainText">
|
||||
<taxonomicName id="82519209A1AB51A8F25A9B5ACDD561F0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="voasary">Tetramorium voasary</taxonomicName>
|
||||
can be clearly distinguished from the remainder of the species complex by the following character combination: eyes moderately sized (OI 21-24); propodeal spines long to very long (PSLI 35-39); petiolar node rectangular nodiform but with relatively rounded anterodorsal and posterodorsal angles; posterodorsal corner of petiolar node not strongly protruding posteriorly; postpetiole in dorsal view usually slightly longer than wide, rarely as wide as long or longer than wide (DPpI 95-101); body uniform bright orange in colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,590,609]" pageId="44" pageNumber="45" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,618,750]" pageId="44" pageNumber="45" type="mainText">HL 0.94-1.23 (1.07); HW 0.87-1.15 (0.98); SL 0.70-0.99 (0.81); EL 0.18-0.27 (0.23); PH 0.49-0.59 (0.53); PW 0.69-0.80 (0.73); WL 1.20-1.49 (1.36); PSL 0.36-0.46 (0.39); PTL 0.32-0.36 (0.35); PTH 0.37-0.42 (0.40); PTW 0.29-0.34 (0.31); PPL 0.34-0.42 (0.38); PPH 0.36-0.43 (0.39); PPW 0.34-0.41 (0.38); CI 88-93 (92); SI 80-89 (82); OI 21-24 (23); DMI 52-58 (54); LMI 38-42 (39); PSLI 35-39 (37); PeNI 41-47 (43); LPeI 85-93 (89); DPeI 85-94 (90); PpNI 48-55 (52); LPpI 90-104 (97); DPpI 95-101 (98); PPI 113-125 (120) (ten measured).</paragraph>
|
||||
<paragraph box="[118,1124,759,1369]" pageId="44" pageNumber="45" type="mainText">Head distinctly longer than wider (CI 88-93). Posterior head margin weakly concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively moderately long, not reaching posterior head margin (SI 80-89). Eyes small to moderately sized (OI 21-24). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-42). Propodeal spines long to very long, spinose, and acute (PSLI 35-39); propodeal lobes short, triangular, and rounded, rarely acute. Petiolar node in profile rectangular nodiform with fairly rounded margins, around 1.1 to 1.2 times higher than long (LPeI 85-93), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum weakly to moderately convex; node in dorsal view around 1.1 to 1.2 times longer than wide (DPeI 85-94). Postpetiole in profile globular, ranging from weakly longer than high to 1.1 times higher than long (LPpI 90-104); in dorsal view ranging from weakly longer than wide to feebly wider than long (DPpI 95-101). Postpetiole in profile appearing approximately as voluminous as petiolar node, in dorsal view approximately 1.1 to 1.3 times wider than petiolar node (PPI 113-125). Mandibles distinctly longitudinally rugose, sometimes weakly so; sculpture on clypeus variable, often longitudinally rugulose with three to five rugulae, sometimes more irregularly rugulose; cephalic dorsum between frontal carinae with seven to ten longitudinal rugae, rugae often broken or with cross-meshes; lateral and ventral head mostly reticulate-rugose. Mesosoma laterally and dorsally mainly longitudinally rugose. Forecoxae usually completely unsculptured, smooth, and shiny, sometimes with partial superficial sculpture. Waist segments rugulose, usually longitudinally so. Generally ground sculpture everywhere on body faint to absent. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with suberect to erect standing hairs. Body of bright orange to light orange brown colour.</paragraph>
|
||||
<subSubSection id="755E93BC54D1AF591EE5CB916F0EE5FA" type="description">
|
||||
<paragraph id="9E86D43FE392FF8972C031E5799C8833" box="[118,226,590,609]" pageId="44" pageNumber="45" type="mainText">Description</paragraph>
|
||||
<paragraph id="4A53856E3C526E58FD12573762BCDF9C" box="[118,1121,618,750]" pageId="44" pageNumber="45" type="mainText">HL 0.94-1.23 (1.07); HW 0.87-1.15 (0.98); SL 0.70-0.99 (0.81); EL 0.18-0.27 (0.23); PH 0.49-0.59 (0.53); PW 0.69-0.80 (0.73); WL 1.20-1.49 (1.36); PSL 0.36-0.46 (0.39); PTL 0.32-0.36 (0.35); PTH 0.37-0.42 (0.40); PTW 0.29-0.34 (0.31); PPL 0.34-0.42 (0.38); PPH 0.36-0.43 (0.39); PPW 0.34-0.41 (0.38); CI 88-93 (92); SI 80-89 (82); OI 21-24 (23); DMI 52-58 (54); LMI 38-42 (39); PSLI 35-39 (37); PeNI 41-47 (43); LPeI 85-93 (89); DPeI 85-94 (90); PpNI 48-55 (52); LPpI 90-104 (97); DPpI 95-101 (98); PPI 113-125 (120) (ten measured).</paragraph>
|
||||
<paragraph id="04BA428FB8D71863A754EA35B3FC27BE" box="[118,1124,759,1369]" pageId="44" pageNumber="45" type="mainText">Head distinctly longer than wider (CI 88-93). Posterior head margin weakly concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively moderately long, not reaching posterior head margin (SI 80-89). Eyes small to moderately sized (OI 21-24). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-42). Propodeal spines long to very long, spinose, and acute (PSLI 35-39); propodeal lobes short, triangular, and rounded, rarely acute. Petiolar node in profile rectangular nodiform with fairly rounded margins, around 1.1 to 1.2 times higher than long (LPeI 85-93), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum weakly to moderately convex; node in dorsal view around 1.1 to 1.2 times longer than wide (DPeI 85-94). Postpetiole in profile globular, ranging from weakly longer than high to 1.1 times higher than long (LPpI 90-104); in dorsal view ranging from weakly longer than wide to feebly wider than long (DPpI 95-101). Postpetiole in profile appearing approximately as voluminous as petiolar node, in dorsal view approximately 1.1 to 1.3 times wider than petiolar node (PPI 113-125). Mandibles distinctly longitudinally rugose, sometimes weakly so; sculpture on clypeus variable, often longitudinally rugulose with three to five rugulae, sometimes more irregularly rugulose; cephalic dorsum between frontal carinae with seven to ten longitudinal rugae, rugae often broken or with cross-meshes; lateral and ventral head mostly reticulate-rugose. Mesosoma laterally and dorsally mainly longitudinally rugose. Forecoxae usually completely unsculptured, smooth, and shiny, sometimes with partial superficial sculpture. Waist segments rugulose, usually longitudinally so. Generally ground sculpture everywhere on body faint to absent. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with suberect to erect standing hairs. Body of bright orange to light orange brown colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,1401,1419]" pageId="44" pageNumber="45" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1122,1429,1589]" pageId="44" pageNumber="45" type="mainText">
|
||||
<subSubSection id="13E583E5EE638AFA8DDCE2F6196AE06E" type="discussion">
|
||||
<paragraph id="56FB1C228FF83D4907C089C530D8CA44" box="[118,170,1401,1419]" pageId="44" pageNumber="45" type="mainText">Notes</paragraph>
|
||||
<paragraph id="31280B77E2D8C4448816C67D35ED5073" box="[118,1122,1429,1589]" pageId="44" pageNumber="45" type="mainText">
|
||||
This
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="9D8B8D60E8035DFE870B11068EAD3293" rank="species">new species</taxonomicNameLabel>
|
||||
is distributed in the rainforests and montane rainforests of eastern Madagascar from BefotakaMidongo in the south to Makirovana in the northeast. The distribution range is comparatively large, but localities where
|
||||
<taxonomicName genus="Tetamorium" pageId="44" pageNumber="45" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
<taxonomicName id="4ED319B965450E36247106DDBD3E3207" genus="Tetamorium" pageId="44" pageNumber="45" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
was encountered are often widely separated. Despite being known from approximately ten localities,
|
||||
<taxonomicName genus="Tetamorium" pageId="44" pageNumber="45" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
<taxonomicName id="799947C2E79C9919A292E4A63F3949CD" genus="Tetamorium" pageId="44" pageNumber="45" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
was collected relatively rarely with less than 25 specimens in total. One explanation might be that the species lives and forages in vegetation; it was mostly collected from lower vegetation, and only rarely from the ground or leaf litter.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,118,306]" pageId="46" pageNumber="47" type="mainText">
|
||||
<pageBreakToken pageId="46" pageNumber="47" start="start">Due</pageBreakToken>
|
||||
<paragraph id="E51461C780BEFC1147E4999410609BC0" box="[118,1122,118,306]" pageId="46" pageNumber="47" type="mainText">
|
||||
<pageBreakToken id="00F2B0DC6A36255BDA0DEDD91A8F38A1" pageId="46" pageNumber="47" start="start">Due</pageBreakToken>
|
||||
to its well-developed eyes (OI 21-24),
|
||||
<taxonomicName genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
<taxonomicName id="2E769F42FBF1AB96AE19C08576CE8208" genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
is unlikely to be mistaken for
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="21876731BAB286B8CF09D54A42E53D6C" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="electrum">
|
||||
T. electrum
|
||||
<authority>,</authority>
|
||||
<authority id="12244B105B0B33C43B3B6ABC2A53D233">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="isectum">
|
||||
<taxonomicName genus="Tetamorium" pageId="46" pageNumber="47" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="1F0D205BC0C5B371025F67F4C600D62E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="isectum">
|
||||
<taxonomicName id="807D030F5140D76088A5D6AF23AA7002" genus="Tetamorium" pageId="46" pageNumber="47" rank="genus">T.</taxonomicName>
|
||||
elf,
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="isectum">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="isectum">
|
||||
<taxonomicName id="E3A37037D38DD168F43F8C88A355B32B" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="isectum">
|
||||
<taxonomicName id="F4F1979A27E376AB494ABFD96BD11AF4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="isectum">
|
||||
T. isectum
|
||||
<authority>,</authority>
|
||||
<authority id="90380B736157714ADEBDAFC9B38EA256">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
<taxonomicName id="8A0D6B819C4FDD63FD21B16F2A669AE9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
or
|
||||
<taxonomicName genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="A67042E03E13B82434FAF777F5690500" genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="nify">T. nify</taxonomicName>
|
||||
since they all have much smaller eyes (OI 15-19). The remaining three species, T.
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="ala">
|
||||
<taxonomicName id="A45963B79BF1B7EC47C0B5B39440A43A" authorityName="," genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="ala">
|
||||
ala
|
||||
<authority>,</authority>
|
||||
<authority id="16E63B5401D7329BDB42C531C8C6F57A">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName genus="Tetamorium" pageId="46" pageNumber="47" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
<taxonomicName id="E3676EB048813A8E361C7EAAD6062F79" genus="Tetamorium" pageId="46" pageNumber="47" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="DB69D8D29A20728F259941013A63A08A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
, and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
<taxonomicName id="4A598120A4069DC84021D65C5B32F6C0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
all have a petiolar node with well-defined antero- and posterodorsal margins while the node of
|
||||
<taxonomicName genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
<taxonomicName id="9CF61603B883336E3D09168CCB6C2FD9" genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
has fairly rounded margins. This character is shared with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
<taxonomicName id="99259B6D4283B701A3418BFD63112BA7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="elf">T. elf</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
<taxonomicName id="8FEA281B88CB24A51285336BD87FA22D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
although they are not likely to be confused with
|
||||
<taxonomicName genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="voasary">T. voasary.</taxonomicName>
|
||||
<taxonomicName id="64F35C206CE447CF4D6AE79F16DA1191" genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="voasary">T. voasary.</taxonomicName>
|
||||
Apart from the small eyes mentioned above, the first two also have extremely long propodeal spines (PSLI 58-64) that contrast with the shorter spines of
|
||||
<taxonomicName genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
<taxonomicName id="26847EAE22EAC3D632F31AD8F267F4A3" genus="Tetamorium" pageId="46" pageNumber="47" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
(PSLI 35-39).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,343,361]" pageId="46" pageNumber="47" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,615,372,391]" pageId="46" pageNumber="47" type="mainText">The species epithet is an arbitrary combination of letters.</paragraph>
|
||||
<subSubSection id="6AB007B7BED16AE64D25F096276E8C13" type="etymology">
|
||||
<paragraph id="5D1BA291BC49FEED221014481B1D3DE2" box="[118,219,343,361]" pageId="46" pageNumber="47" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="7F83646FDC0BCA67FFDC77B63DACE27C" box="[118,615,372,391]" pageId="46" pageNumber="47" type="mainText">The species epithet is an arbitrary combination of letters.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,428,446]" pageId="46" pageNumber="47" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,456,841]" pageId="46" pageNumber="47" type="mainText">
|
||||
<subSubSection id="431F0F83ED0AB60D933A1EE74419E868" type="materials_examined">
|
||||
<paragraph id="DAFBD4D8F60CB6CA0A4000F2B0CAA0EA" box="[118,295,428,446]" pageId="46" pageNumber="47" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="B44E0BD5478361BFBE2DA666E2875FD9" box="[118,1122,456,841]" pageId="46" pageNumber="47" type="mainText">
|
||||
MADAGASCAR: Antsiranana, Makirovana forest, 14.1707 S, 49.9541 E, 415 m, rainforest, 28.IV.2011 (B.L. Fisher et al.); Fianarantsoa, Foret d'Ambalagoavy Nord, Ikongo, Ambatombe, 21.8275 S, 47.3389 E, 625 m, 1.XII.2000 (R. Harin'Hala & M.E. Irwin); Fianarantsoa, Parc National Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango,23.83517 S, 46.96367 E, 940 m, rainforest, 13.-15.XI.2006 (B.L. Fisher et al.); Fianarantsoa,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="EB719BA09A1EA862BCA18816FBADB34A" originalValue="Réserve">Reserve</normalizedToken>
|
||||
Speciale Manombo 24.5 km 228° Farafangana, 23.0158 S, 47.719 E, 30 m, rainforest, 22.IV.2006 (B.L. Fisher et al.); Fianarantsoa, Ranomafana, 21.25 S, 47.3667 E, in forest along riverbank, 1.III.1994 (A. Pauly); Fianarantsoa, Ranomafana National Park, Talatakely, 30.X.-20.XI.1998 (V.F. Lee & K.J. Ribardo); Fianarantsoa, Parc National de Ranomafana, Vatoharanana River, 4.1 km 231° SW Ranomafana, 21.29 S, 47.4333 E, 1100 m, montane rainforest, 27.-31.III.2003 (B.L. Fisher, C. Griswold et al.); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.8496 S, 48.2947 E, 1010 m, 3.-6.III.2007 (B.L. Fisher et al.); Toamasina, Amparihibe, 15° 2' S, 49° 34' E, II.-III.2003 (K.A. Jackson & D. Carpenter); Toamasina, Montagne d'Anjanaharibe, 19.5 km 27° NNE Ambinanitelo, 15.1783 S, 49.635 E, 1100 m, montane rainforest, 12.-16.III.2003 (B.L. Fisher, C. Griswold et al.); Toamasina, Torotorofotsy, 18.8708 S, 48.3474 E, 1070 m, montane rainforest, marsh edge, 24.III.2004 (Malagasy ant team).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,128 +1,127 @@
|
|||
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|
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|
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:name id="10EF0A5B21B0577F14E78E2BF542CF23" type="personal">
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|
||||
</mods:role>
|
||||
<mods:namePart>B. L. Fisher</mods:namePart>
|
||||
<mods:namePart id="62AB7C162E8B06D99C8550A60A00B705">B. L. Fisher</mods:namePart>
|
||||
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|
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<mods:title>Zootaxa</mods:title>
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<mods:title id="C9097FD7556282542A985BB63E18EFF7">Zootaxa</mods:title>
|
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|
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<mods:part>
|
||||
<mods:date>2012</mods:date>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>3592</mods:number>
|
||||
<mods:part id="3E8E181A626038413532B9CD18BEBB6D">
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||||
<mods:date id="D7AE993F3075AF9FEF0812F16572F79F">2012</mods:date>
|
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<mods:detail id="16F895A95156064B8977BF7983A3A984" type="volume">
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||||
<mods:number id="96F2C194539D1C5026DAB7C0A034D848">3592</mods:number>
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</mods:detail>
|
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<mods:extent unit="page">
|
||||
<mods:start>1</mods:start>
|
||||
<mods:end>85</mods:end>
|
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<mods:start id="E5F2F0C8B241A2329A9A38BE28DBB743">1</mods:start>
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<mods:end id="4D127DC1BFC928D9F679A5173BCF940B">85</mods:end>
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<mods:url>http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
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<mods:url id="2860F2730E4E55DA03EEC2CF8A49E4EE">http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="HNS-Pub">26064</mods:identifier>
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<mods:identifier type="ISSN">175-5326</mods:identifier>
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<mods:identifier type="ZooBank">A2D9C9ED-C0BA-4B5F-A330-C9AB7D625704</mods:identifier>
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<mods:identifier id="999D5151A042913A6572727C248AB5C0" type="HNS-Pub">26064</mods:identifier>
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<mods:identifier id="58363AA9318E64384EE603287B194FB9" type="ISSN">175-5326</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172611" ID-GBIF-Taxon="100105316" ID-Zenodo-Dep="6172611" LSID="urn:lsid:plazi:treatment:9C1995A5694DF9E09C9B887EF7DD35CD" httpUri="http://treatment.plazi.org/id/9C1995A5694DF9E09C9B887EF7DD35CD" lastPageNumber="23" pageNumber="21">
|
||||
<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,548,118,138]" pageId="20" pageNumber="21" type="mainText">
|
||||
<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ala">
|
||||
<pageBreakToken pageId="20" pageNumber="21" start="start">Tetramorium</pageBreakToken>
|
||||
<treatment id="9C1995A5694DF9E09C9B887EF7DD35CD" ID-DOI="http://doi.org/10.5281/zenodo.6172611" ID-GBIF-Taxon="100105316" ID-Zenodo-Dep="6172611" LSID="urn:lsid:plazi:treatment:9C1995A5694DF9E09C9B887EF7DD35CD" httpUri="http://treatment.plazi.org/id/9C1995A5694DF9E09C9B887EF7DD35CD" lastPageNumber="23" pageNumber="21" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="DBCF203C23D3DC8055EBD56D4BA93FBB" type="nomenclature">
|
||||
<paragraph id="9C932ACA63FE480CD87B70FD71343A49" box="[118,548,118,138]" pageId="20" pageNumber="21" type="mainText">
|
||||
<taxonomicName id="AAE9DF858A7A0891AA3BBE206294E078" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ala">
|
||||
<pageBreakToken id="B09BC15EC4F58021ED10D9C4010CFC6B" pageId="20" pageNumber="21" start="start">Tetramorium</pageBreakToken>
|
||||
ala
|
||||
<authority>Hita</authority>
|
||||
<authority id="8301A18765C56F3BF9DC7FCB1CB09B5B">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="E7DEFFF2026AA0A20BD9EDAB76478DE8" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,410,146,165]" pageId="20" pageNumber="21" type="mainText">(Figs. 51, 57, 66, 75, 76, 77, 141)</paragraph>
|
||||
<subSubSection id="D7818444FDA9DAB5A9D0C2C73858A493" type="description">
|
||||
<paragraph id="881FEDC7190E7AFCB371FF7B45EBB2B0" box="[118,410,146,165]" pageId="20" pageNumber="21" type="mainText">(Figs. 51, 57, 66, 75, 76, 77, 141)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1124,202,446]" pageId="20" pageNumber="21" type="mainText">Holotype worker, MADAGASCAR, Toamasina, Montagne d'Anjanaharibe, 19.5 km 27° NNE Ambinanitelo, 15.17833 S, 49.635 E, 1100 m, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF08150, 12.-16.III.2003 (B.L. Fisher, C. Griswold et al.) (CASC: CASENT0038473). Paratypes, 13 workers with same data as holotype (BMNH: CASENT0038387; CASC: CASENT0038371; CASENT0038374; CASENT0038389; CASENT0038410; CASENT0038414; CASENT0038453; CASENT0038459; CASENT0038464; CASENT0038467; MCZ: CASENT0038404; MHNG: CASENT0038423; NHMB: CASENT0038373); 1 worker with same data as holotype except sampled from ground and collection code BLF08230 (CASC: CASENT0497817); 9 workers with same data as holotype except sampled from root mat at ground layer and collection code BLF08185 (CASC: CASENT0498013; CASENT0498014; CASENT0498015).</paragraph>
|
||||
<subSubSection id="657663164E60027327286A1B5605FCB7" type="materials_examined">
|
||||
<paragraph id="E22CC626F109CF8185858F84D09B706E" box="[118,1124,202,446]" pageId="20" pageNumber="21" type="mainText">Holotype worker, MADAGASCAR, Toamasina, Montagne d'Anjanaharibe, 19.5 km 27° NNE Ambinanitelo, 15.17833 S, 49.635 E, 1100 m, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF08150, 12.-16.III.2003 (B.L. Fisher, C. Griswold et al.) (CASC: CASENT0038473). Paratypes, 13 workers with same data as holotype (BMNH: CASENT0038387; CASC: CASENT0038371; CASENT0038374; CASENT0038389; CASENT0038410; CASENT0038414; CASENT0038453; CASENT0038459; CASENT0038464; CASENT0038467; MCZ: CASENT0038404; MHNG: CASENT0038423; NHMB: CASENT0038373); 1 worker with same data as holotype except sampled from ground and collection code BLF08230 (CASC: CASENT0497817); 9 workers with same data as holotype except sampled from root mat at ground layer and collection code BLF08185 (CASC: CASENT0498013; CASENT0498014; CASENT0498015).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,483,501]" pageId="20" pageNumber="21" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,511,643]" pageId="20" pageNumber="21" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ala">Tetramorium ala</taxonomicName>
|
||||
<subSubSection id="5A7D579893730E49E4C7F524AA5D7100" type="diagnosis">
|
||||
<paragraph id="C9F034CAC6A955480C583A110731974B" box="[118,208,483,501]" pageId="20" pageNumber="21" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="CC2D9F90865BB1E220A1ED17E1654AA7" box="[118,1121,511,643]" pageId="20" pageNumber="21" type="mainText">
|
||||
<taxonomicName id="F6D44F80856CF03CE24FA5FA737532AD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ala">Tetramorium ala</taxonomicName>
|
||||
can be distinguished from the other
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="950406D4A4973A50B51BD90E948C75AA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
species complex members by the following combination of characters: eyes usually moderately sized (OI 20-21); propodeal spines long to very long (PSLI 38-42); petiolar node in dorsal view relatively slender and long, around 1.3 to 1.4 times longer than wide (DPeI 72-76); posterodorsal corner of petiolar node not strongly protruding posteriorly; body bright orange to light brown in colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,680,698]" pageId="20" pageNumber="21" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1117,708,868]" pageId="20" pageNumber="21" type="mainText">HL 0.88-1.01 (0.94); HW 0.83-0.96 (0.88); SL 0.69-0.76 (0.72); EL 0.17-0.19 (0.18); PH 0.43-0.49 (0.46); PW 0.63-0.74 (0.69); WL 1.16-1.29 (1.22); PSL 0.34-0.41 (0.37); PTL 0.29-0.34 (0.31); PTH 0.30-0.35 (0.32); PTW 0.21-0.26 (0.23); PPL 0.27-0.33 (0.30); PPH 0.30-0.36 (0.32); PPW 0.30-0.36 (0.32); CI 93-95 (94); SI 79-85 (81); OI 20-21 (20); DMI 54-58 (57); LMI 37-39 (38); PSLI 38-42 (40); PeNI 31-34 (33); LPeI 93-98 (95); DPeI 72-76 (75); PpNI 44-48 (46); LPpI 87-95 (92); DPpI 102-112 (107); PPI 135-143 (139) (12 measured).</paragraph>
|
||||
<paragraph box="[118,1122,877,1543]" pageId="20" pageNumber="21" type="mainText">Head distinctly longer than wide (CI 93-95). Posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, strongly diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, but shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short to moderately long, not reaching posterior head margin (SI 79-85). Eyes short to moderately sized (OI 20-21). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-39). Propodeal spines long to very long, spinose, and acute (PSLI 38-42); propodeal lobes well-developed and comparatively long, elongate-triangular, and acute. Petiolar node in profile rectangular nodiform with sharply defined angles, between 1.0 to 1.1 times higher than long (LPeI 93-98), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum straight; node in dorsal view around 1.3 to 1.4 times longer than wide (DPeI 72-76). Postpetiole in profile globular to subglobular, approximately 1.1 times higher than long (LPpI 87-95); in dorsal view 1.0 to 1.1 times wider than long (DPpI 102-112). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view approximately 1.3 to 1.4 times wider than petiolar node (PPI 135-143). Mandibles distinctly longitudinally rugose; clypeus always with a very well-developed and pronounced longitudinal median ruga and few much weaker, and often broken lateral rugae/rugulae; cephalic dorsum between frontal carinae with seven to ten longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae only very rarely with cross-meshes; lateral and ventral head usually longitudinally rugose, sometimes partly reticulate-rugose. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae usually completely unsculptured, smooth, and shiny, sometimes with partial superficial sculpture. Waist segments longitudinally rugose. Ground sculpture generally faint to absent everywhere on body. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with suberect to erect standing hairs. Body colour uniformly bright orange to orange-brown.</paragraph>
|
||||
<caption box="[118,1122,1543,1589]" pageId="21" pageNumber="22" type="caption">
|
||||
<paragraph box="[118,1122,1543,1589]" pageId="21" pageNumber="22" type="caption">
|
||||
<pageBreakToken pageId="21" pageNumber="22" start="start">FIGURES</pageBreakToken>
|
||||
<subSubSection id="9504481D29777586BD6819F952502F1F" type="description">
|
||||
<paragraph id="4CFCA9D740926B9F2670B953AD7DEDF7" box="[118,226,680,698]" pageId="20" pageNumber="21" type="mainText">Description</paragraph>
|
||||
<paragraph id="14FBA48AFCF52FF9D274FC5887FD1912" box="[118,1117,708,868]" pageId="20" pageNumber="21" type="mainText">HL 0.88-1.01 (0.94); HW 0.83-0.96 (0.88); SL 0.69-0.76 (0.72); EL 0.17-0.19 (0.18); PH 0.43-0.49 (0.46); PW 0.63-0.74 (0.69); WL 1.16-1.29 (1.22); PSL 0.34-0.41 (0.37); PTL 0.29-0.34 (0.31); PTH 0.30-0.35 (0.32); PTW 0.21-0.26 (0.23); PPL 0.27-0.33 (0.30); PPH 0.30-0.36 (0.32); PPW 0.30-0.36 (0.32); CI 93-95 (94); SI 79-85 (81); OI 20-21 (20); DMI 54-58 (57); LMI 37-39 (38); PSLI 38-42 (40); PeNI 31-34 (33); LPeI 93-98 (95); DPeI 72-76 (75); PpNI 44-48 (46); LPpI 87-95 (92); DPpI 102-112 (107); PPI 135-143 (139) (12 measured).</paragraph>
|
||||
<paragraph id="B4139B7E1090D2FD8E095C5C4B82120D" box="[118,1122,877,1543]" pageId="20" pageNumber="21" type="mainText">Head distinctly longer than wide (CI 93-95). Posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, strongly diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, but shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short to moderately long, not reaching posterior head margin (SI 79-85). Eyes short to moderately sized (OI 20-21). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-39). Propodeal spines long to very long, spinose, and acute (PSLI 38-42); propodeal lobes well-developed and comparatively long, elongate-triangular, and acute. Petiolar node in profile rectangular nodiform with sharply defined angles, between 1.0 to 1.1 times higher than long (LPeI 93-98), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum straight; node in dorsal view around 1.3 to 1.4 times longer than wide (DPeI 72-76). Postpetiole in profile globular to subglobular, approximately 1.1 times higher than long (LPpI 87-95); in dorsal view 1.0 to 1.1 times wider than long (DPpI 102-112). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view approximately 1.3 to 1.4 times wider than petiolar node (PPI 135-143). Mandibles distinctly longitudinally rugose; clypeus always with a very well-developed and pronounced longitudinal median ruga and few much weaker, and often broken lateral rugae/rugulae; cephalic dorsum between frontal carinae with seven to ten longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae only very rarely with cross-meshes; lateral and ventral head usually longitudinally rugose, sometimes partly reticulate-rugose. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae usually completely unsculptured, smooth, and shiny, sometimes with partial superficial sculpture. Waist segments longitudinally rugose. Ground sculpture generally faint to absent everywhere on body. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with suberect to erect standing hairs. Body colour uniformly bright orange to orange-brown.</paragraph>
|
||||
<caption id="C1430C0EB1220826EE968D4683A81EAC" box="[118,1122,1543,1589]" pageId="21" pageNumber="22" type="caption">
|
||||
<paragraph id="CDA3AD24D45348DF545A29C563BC86AF" box="[118,1122,1543,1589]" pageId="21" pageNumber="22" type="caption">
|
||||
<pageBreakToken id="D07634845AE194CBB351DBDFBD2D4FC0" pageId="21" pageNumber="22" start="start">FIGURES</pageBreakToken>
|
||||
75-77. T. ala, holotype (CASENT0038473). 75. Body in profile. 76. Body in dorsal view. 77. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,170,118,136]" pageId="22" pageNumber="23" type="mainText">
|
||||
<pageBreakToken pageId="22" pageNumber="23" start="start">Notes</pageBreakToken>
|
||||
<paragraph id="65FBB2106B9C449D91BA9C219E37A89E" box="[118,170,118,136]" pageId="22" pageNumber="23" type="mainText">
|
||||
<pageBreakToken id="86D3EFB511D785284A57C481F4543A8B" pageId="22" pageNumber="23" start="start">Notes</pageBreakToken>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1120,147,194]" pageId="22" pageNumber="23" type="mainText">
|
||||
<paragraph id="63C58255FF0A983DB8949C2747F468F5" box="[118,1120,147,194]" pageId="22" pageNumber="23" type="mainText">
|
||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="28A027108606240256234302CBAC009C" rank="species">new species</taxonomicNameLabel>
|
||||
is currently only known to occur in the type locality Montagne d'Anjanaharibe where it was sampled in montane rainforest at 1100 m elevation. Most specimens were found in leaf litter.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,1122,203,559]" pageId="22" pageNumber="23" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ala">Tetramorium ala</taxonomicName>
|
||||
<subSubSection id="566F19EB02BFFCB9433400E15D2FA487" type="discussion">
|
||||
<paragraph id="1DD23CE1A254F139F4D560D5DB3C3AAB" box="[118,1122,203,559]" pageId="22" pageNumber="23" type="mainText">
|
||||
<taxonomicName id="254AC2D3A4BCBDF5223ED582F464B9FD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ala">Tetramorium ala</taxonomicName>
|
||||
has the longest petiolar node in dorsal view within the species complex, which is around 1.3 to 1.4 times longer than wide (DPeI 72-76), whereas all other species (except T.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
<taxonomicName id="6608AC8A2979DFE854342392028C29A7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
) have much shorter and wider nodes (DPeI 82-114).
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andohahela">Tetramorium andohahela</taxonomicName>
|
||||
<taxonomicName id="320BCDFF834F2647D84819827BE40267" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andohahela">Tetramorium andohahela</taxonomicName>
|
||||
also has a relatively long and slender node in dorsal view (DPeI 73-79), but the posterodorsal corner of the petiolar node distinctly protrudes posteriorly in this species, a character absent in T. ala. Not considering the node shape in dorsal view, T. ala is not easily confused with
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="90C91EF96CF7433BEA678E44D5E60213" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="electrum">
|
||||
T. electrum
|
||||
<authority>,</authority>
|
||||
<authority id="74FDD7DA69A6AA07841754CEC5798A05">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="elf">
|
||||
<taxonomicName genus="Tetamorium" pageId="22" pageNumber="23" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="94AD696FA30D4CFFF6C4A39C6F63897D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="elf">
|
||||
<taxonomicName id="556562D1E07995B9DE64ECA9CBEAE827" genus="Tetamorium" pageId="22" pageNumber="23" rank="genus">T.</taxonomicName>
|
||||
elf,
|
||||
</taxonomicName>
|
||||
or
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="123AD7BB72F7105139137A4DA16EBB48" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
since the latter species have significantly smaller eyes (OI 16-19), much longer propodeal spines (PSLI 46-64), and much shorter propodeal lobes. The other two species with very small eyes,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="300F358ED20C963224AC77F12BF4ABDB" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
(OI 15 -16) and
|
||||
<taxonomicName genus="Tetamorium" pageId="22" pageNumber="23" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="103FE4B069DE59519C66E14BCC0A1868" genus="Tetamorium" pageId="22" pageNumber="23" rank="species" species="nify">T. nify</taxonomicName>
|
||||
(OI 16-17), are morphologically not very close to T. ala with its larger eyes (OI 20-21). Due to its fairly rounded anterodorsal and posterodorsal margins of the petiolar node,
|
||||
<taxonomicName genus="Tetamorium" pageId="22" pageNumber="23" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
<taxonomicName id="1DF2C414231456757FD74909DD324782" genus="Tetamorium" pageId="22" pageNumber="23" rank="species" species="voasary">T. voasary</taxonomicName>
|
||||
can be easily separated from T. ala, which has very sharply defined margins. The last and most variable species of the complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
<taxonomicName id="27A8FFE54D9292FD457FD4CF4559C29D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
could be confused with T. ala since they share most morphological characters, especially in profile view. However, as mentioned above, the different node shape in dorsal view distinguishes them clearly (DPeI 72-76 in T. ala versus DPeI 82-98 in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andrei">T. andrei).</taxonomicName>
|
||||
<taxonomicName id="4ECE9B187C99FA70A4D69D1350E45AB9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="andrei">T. andrei).</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,592,610]" pageId="22" pageNumber="23" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,615,621,640]" pageId="22" pageNumber="23" type="mainText">The species epithet is an arbitrary combination of letters.</paragraph>
|
||||
<subSubSection id="E2D127D81C21B33C20860BE54F969198" type="etymology">
|
||||
<paragraph id="EF6F57D588ECA778D77A2676184E9670" box="[118,219,592,610]" pageId="22" pageNumber="23" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="B0FED841E35ADDF16AFBFC7305D5CC50" box="[118,615,621,640]" pageId="22" pageNumber="23" type="mainText">The species epithet is an arbitrary combination of letters.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,672,691]" pageId="22" pageNumber="23" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,921,701,748]" pageId="22" pageNumber="23" type="mainText">MADAGASCAR: Toamasina, Montagne d'Anjanaharibe, 19.5 km 27° NNE Ambinanitelo, 1100 m, montane rainforest, 12.-16.III.2003 (B.L. Fisher, C. Griswold et al.).</paragraph>
|
||||
<paragraph box="[926,1120,701,719]" pageId="22" pageNumber="23" type="mainText">15.17833 S, 49.635 E,</paragraph>
|
||||
<subSubSection id="F04F9549AA90AD7C56894E3C40CAC3E6" type="materials_examined">
|
||||
<paragraph id="89A7F737C07124F2A6D26C22DE73188A" box="[118,295,672,691]" pageId="22" pageNumber="23" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="A2F448D73847ED12E1EEE6EBE831C768" box="[118,921,701,748]" pageId="22" pageNumber="23" type="mainText">MADAGASCAR: Toamasina, Montagne d'Anjanaharibe, 19.5 km 27° NNE Ambinanitelo, 1100 m, montane rainforest, 12.-16.III.2003 (B.L. Fisher, C. Griswold et al.).</paragraph>
|
||||
<paragraph id="5C058A88101820D6FD6C9EFC10BDAD0B" box="[926,1120,701,719]" pageId="22" pageNumber="23" type="mainText">15.17833 S, 49.635 E,</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,49 +1,50 @@
|
|||
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Key to Fijian
|
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<taxonomicName id="D0974D48562BFFD3E94EFF5AFE53FF21" ID-CoL="77f81588-9b0d-4ab2-b82e-7265a66ef220" box="[323,472,222,248]" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="6" rank="order">Trichoptera</taxonomicName>
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families (adults) and
|
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<taxonomicName id="D0974D48562BFFD3EAEBFF5AFC20FF21" box="[742,939,222,248]" class="Insecta" family="Philopotamidae" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Philopotamidae</taxonomicName>
|
||||
species (males)
|
||||
|
|
|
|||
|
|
@ -1,128 +1,127 @@
|
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<mods:title id="A914214A5008EC45DCA197AE0448C4AD">The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups.</mods:title>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172605" ID-GBIF-Taxon="100105363" ID-Zenodo-Dep="6172605" LSID="urn:lsid:plazi:treatment:A59B39DF7906619A055C17313AF3AE89" httpUri="http://treatment.plazi.org/id/A59B39DF7906619A055C17313AF3AE89" lastPageNumber="12" pageNumber="11">
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<taxonomicName authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kelleri">
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<taxonomicName id="C7B9CFAE44D82F06DD109F6C9BADA81B" ID-CoL="55PP2" authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kelleri">
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Tetramorium kelleri
|
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<authority>Forel</authority>
|
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<authority id="E0A3C4E8D1FE2800ABB5388AAB232965">Forel</authority>
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,
|
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</taxonomicName>
|
||||
1887
|
||||
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|
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|
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|
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<paragraph box="[118,333,455,474]" pageId="10" pageNumber="11" type="mainText">(Figs. 3, 4, 8, 9, 10, 141)</paragraph>
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<paragraph id="6601EEA11AC057D8B730B992B57E6E99" box="[118,333,455,474]" pageId="10" pageNumber="11" type="mainText">(Figs. 3, 4, 8, 9, 10, 141)</paragraph>
|
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</subSubSection>
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<subSubSection type="materials_examined">
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<paragraph box="[118,1122,511,576]" pageId="10" pageNumber="11" type="mainText">
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<taxonomicName authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kelleri" subGenus="Xiphomyrmex">
|
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<subSubSection id="794C20328C061A83DBBFFCDA1CAB2BEA" type="materials_examined">
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<paragraph id="8A7E308C5BB9FA86C82D35091BF9AC08" box="[118,1122,511,576]" pageId="10" pageNumber="11" type="mainText">
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<taxonomicName id="BFB25C0AB7C4C47EFCC78CA0D4C7B756" authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kelleri" subGenus="Xiphomyrmex">
|
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Tetramorium (Xiphomyrmex) kelleri
|
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<authority>Forel</authority>
|
||||
<authority id="78BD5D37530BDD274D1E88EECE54BA21">Forel</authority>
|
||||
,
|
||||
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|
||||
1887:385. Syntype workers, MADAGASCAR, Toamasina, Ivondro, pr. Tamatave (C. Keller). (BMNH: CASENT0102339; NHMB: CASENT0101138; MCZ: CASENT0247309; CASENT0247310; MHNG: CASENT0101293; CASENT0101294; CASENT0101938; USNM) [all examined, except material from USNM].
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,611,629]" pageId="10" pageNumber="11" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,639,686]" pageId="10" pageNumber="11" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kelleri">Tetramorium kelleri</taxonomicName>
|
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<subSubSection id="FDE482FF2D62FC53D4A362DB35DB84AC" type="diagnosis">
|
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<paragraph id="9B9CF45D92FFDCCBA652B242A6EFA111" box="[118,208,611,629]" pageId="10" pageNumber="11" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="229F911342DC89D9070838334837A9DC" box="[118,1121,639,686]" pageId="10" pageNumber="11" type="mainText">
|
||||
<taxonomicName id="07486A392E507070B48FE2880F4FFB4B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kelleri">Tetramorium kelleri</taxonomicName>
|
||||
differs from T.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
<taxonomicName id="6BEDD32CB4CB3C9C42C65E6CDD232130" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
by the character combination of: large eyes (OI 24-26); relatively shorter antennal scapes (SI 89-99); and extremely long propodeal spines (PSLI 49-68).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,723,742]" pageId="10" pageNumber="11" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,752,883]" pageId="10" pageNumber="11" type="mainText">HL 0.98-1.11 (1.03); HW 0.85-0.97 (0.90); SL 0.79-0.90 (0.85); EL 0.21-0.24 (0.22); PH 0.45-0.58 (0.52); PW 0.66-0.80 (0.73); WL 1.29-1.48 (1.37); PSL 0.50-0.73 (0.59); PTL 0.36-0.44 (0.40); PTH 0.37-0.44 (0.40); PTW 0.29-0.36 (0.33); PPL 0.32-0.37 (0.34); PPH 0.38-0.44 (0.40); PPW 0.37-0.43 (0.40); CI 84-89 (87); SI 89-99 (94); OI 24-26 (25); DMI 50-59 (53); LMI 35-40 (38); PSLI 49-68 (57); PeNI 41-50 (46); LPeI 95-102 (99); DPeI 79-86 (83); PpNI 49-59 (55); LPpI 81-90 (86); DPpI 109-124 (117); PPI 114-126 (121) (25 measured).</paragraph>
|
||||
<paragraph box="[118,1121,892,1474]" pageId="10" pageNumber="11" type="mainText">Head much longer than wide (CI 84-89); posterior head margin very weakly concave. Anterior clypeal margin medially impressed, often weakly so. Frontal carinae strongly developed, approaching corners of posterior head margin. Antennal scrobes well-developed, moderately deep, narrow, and without defined posterior margin; ventral margin moderately defined. Antennal scapes moderately long to long, reaching posterior head margin (SI 89-99). Eyes large (OI 24-26). Mesosomal outline in profile flat to weakly convex, very weakly marginate from lateral to dorsal mesosoma, sides rounding smoothly onto the dorsum; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 35-40). Propodeal spines extremely long, spinose and acute (PSLI 49-68); propodeal lobes short, triangular, and rounded, sometimes weakly acute. Petiolar node in profile clublike with fairly rounded margins, ranging from weakly longer than high to weakly higher than long (LPeI 95-102), anterodorsal margin situated lower than posterodorsal margin, dorsum noticeably convex; node in dorsal view approximately 1.2 to 1.3 times longer than wide (DPeI 79-86). Postpetiole in profile globular to subglobular, approximately 1.1 to 1.2 times higher than long (LPpI 81-90); in dorsal view around 1.1 to 1.3 times wider than long (DPpI 109-124). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view 1.1 to 1.3 times wider than petiolar node (PPI 114-126). Mandibles strongly striate; clypeus longitudinally rugose/rugulose, median ruga always present and distinct, remaining rugae/rugulae weaker and variably developed; cephalic dorsum between frontal carinae reticulate-rugose to longitudinally rugose, posteriorly more reticulate-rugose and anteriorly more longitudinally rugose; lateral and ventral head mostly reticulate-rugose. Mesosoma dorsally mainly reticulate-rugose, laterally reticulate-rugose to longitudinally rugose. Forecoxae usually with weak to moderately developed longitudinal rugae/rugulae, sometimes reduced. Waist segments mainly longitudinally rugulose, less reticulate-rugulose. First gastral tergite unsculptured, smooth, and shining. Whole body covered with numerous, very long, fine standing hairs. Body of uniform whitish-yellowish to brown, mostly yellowish to orange-brown.</paragraph>
|
||||
<subSubSection id="A03B6F4C2F9C11B7970AE02F36BA263C" type="description">
|
||||
<paragraph id="D6EEE3D4C57C6D503BBE85BF2830FC9C" box="[118,226,723,742]" pageId="10" pageNumber="11" type="mainText">Description</paragraph>
|
||||
<paragraph id="0A247B507B41D3046572BBCE5C22BBED" box="[118,1121,752,883]" pageId="10" pageNumber="11" type="mainText">HL 0.98-1.11 (1.03); HW 0.85-0.97 (0.90); SL 0.79-0.90 (0.85); EL 0.21-0.24 (0.22); PH 0.45-0.58 (0.52); PW 0.66-0.80 (0.73); WL 1.29-1.48 (1.37); PSL 0.50-0.73 (0.59); PTL 0.36-0.44 (0.40); PTH 0.37-0.44 (0.40); PTW 0.29-0.36 (0.33); PPL 0.32-0.37 (0.34); PPH 0.38-0.44 (0.40); PPW 0.37-0.43 (0.40); CI 84-89 (87); SI 89-99 (94); OI 24-26 (25); DMI 50-59 (53); LMI 35-40 (38); PSLI 49-68 (57); PeNI 41-50 (46); LPeI 95-102 (99); DPeI 79-86 (83); PpNI 49-59 (55); LPpI 81-90 (86); DPpI 109-124 (117); PPI 114-126 (121) (25 measured).</paragraph>
|
||||
<paragraph id="B5EBAACF6A3EBE2FB36A2DBDA09BB960" box="[118,1121,892,1474]" pageId="10" pageNumber="11" type="mainText">Head much longer than wide (CI 84-89); posterior head margin very weakly concave. Anterior clypeal margin medially impressed, often weakly so. Frontal carinae strongly developed, approaching corners of posterior head margin. Antennal scrobes well-developed, moderately deep, narrow, and without defined posterior margin; ventral margin moderately defined. Antennal scapes moderately long to long, reaching posterior head margin (SI 89-99). Eyes large (OI 24-26). Mesosomal outline in profile flat to weakly convex, very weakly marginate from lateral to dorsal mesosoma, sides rounding smoothly onto the dorsum; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 35-40). Propodeal spines extremely long, spinose and acute (PSLI 49-68); propodeal lobes short, triangular, and rounded, sometimes weakly acute. Petiolar node in profile clublike with fairly rounded margins, ranging from weakly longer than high to weakly higher than long (LPeI 95-102), anterodorsal margin situated lower than posterodorsal margin, dorsum noticeably convex; node in dorsal view approximately 1.2 to 1.3 times longer than wide (DPeI 79-86). Postpetiole in profile globular to subglobular, approximately 1.1 to 1.2 times higher than long (LPpI 81-90); in dorsal view around 1.1 to 1.3 times wider than long (DPpI 109-124). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view 1.1 to 1.3 times wider than petiolar node (PPI 114-126). Mandibles strongly striate; clypeus longitudinally rugose/rugulose, median ruga always present and distinct, remaining rugae/rugulae weaker and variably developed; cephalic dorsum between frontal carinae reticulate-rugose to longitudinally rugose, posteriorly more reticulate-rugose and anteriorly more longitudinally rugose; lateral and ventral head mostly reticulate-rugose. Mesosoma dorsally mainly reticulate-rugose, laterally reticulate-rugose to longitudinally rugose. Forecoxae usually with weak to moderately developed longitudinal rugae/rugulae, sometimes reduced. Waist segments mainly longitudinally rugulose, less reticulate-rugulose. First gastral tergite unsculptured, smooth, and shining. Whole body covered with numerous, very long, fine standing hairs. Body of uniform whitish-yellowish to brown, mostly yellowish to orange-brown.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,1122,1539,1586][118,1122,118,334]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kelleri">Tetramorium kelleri</taxonomicName>
|
||||
<subSubSection id="0BFE73AC6D63CEB672195CBB5767BBE2" type="discussion">
|
||||
<paragraph id="6FDE03A39A43DC4EA18053C179E8988C" box="[118,1122,1539,1586][118,1122,118,334]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="mainText">
|
||||
<taxonomicName id="D5FC405515DA9EC5CCC81D562FF6C4DE" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kelleri">Tetramorium kelleri</taxonomicName>
|
||||
is mainly encountered in northern and western Madagascar, and on several islands including Nosy Be, Nosy Mangabe, and Nosy Ngontsy. Interestingly, most localities are in relatively close proximity to the
|
||||
<pageBreakToken pageId="11" pageNumber="12" start="start">ocean</pageBreakToken>
|
||||
<pageBreakToken id="DEBD55085F3B15550FB5C76064F1520F" pageId="11" pageNumber="12" start="start">ocean</pageBreakToken>
|
||||
since no material was collected more than 50 km inland. One aspect that deserves attention, however, is the type locality, the Ivondro River, which is relatively far from the currently known distribution range mentioned above. The area around the Ivondro River in eastern Madagascar was intensively sampled by the Malagasy ant inventory (see Fisher, 2005), but no additional
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
<taxonomicName id="8912812F75C4B4E99626FCF6FA03C84E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
material was found. Indeed, not a single modern specimen is known from eastern Madagascar south of Nosy Mangabe. One possible explanation could be that the species was present in the area over 120 years ago, but did not survive until the present day. This is surprising, though, since the species is comparatively flexible in its habitat requirements. It was collected from rainforest, littoral rainforest, tropical dry forest, and secondary forest at elevations from 5 to 780 m, although mostly at the lower range.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,343,531]" pageId="11" pageNumber="12" type="mainText">
|
||||
<paragraph id="2760EEB8741FA9E9BD3308D89487702A" box="[118,1122,343,531]" pageId="11" pageNumber="12" type="mainText">
|
||||
Despite being a very common species with a relatively broad distribution range, which includes several islands,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
<taxonomicName id="C436A7B70351C24BF4FC5426FA098652" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
remains remarkably invariable. Some minor variation in colouration is observed ranging from whitish-yellowish to brown, although we do not think it significant for species diagnostics. In general, as already stated by Bolton (1979), it is a highly conspicuous and easily recognisable Malagasy ant species. The only other species it could be confused with is the second species of the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
<taxonomicName id="623E5EF922252966B7ACC02BB8C2A54E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
group, T.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
<taxonomicName id="07515DFD8E9ACDCCB821E8A037F3FA3D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ankarana">ankarana</taxonomicName>
|
||||
. The latter species, however, has much smaller eyes (OI 20) and propodeal spines (PSLI 35-38), and slightly longer antennal scapes (SI 101-104) than
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
<taxonomicName id="186A516EB450388639E8732EF75721E1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kelleri">T. kelleri</taxonomicName>
|
||||
(OI 24-26; SI 89-99; PSLI 49-68).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,568,587]" pageId="11" pageNumber="12" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,596,1516]" pageId="11" pageNumber="12" type="mainText">
|
||||
<subSubSection id="9990B2DA4E91D1F59F887A5D9804CCE1" type="materials_examined">
|
||||
<paragraph id="2C0E4E188593258C1EC803E39DB4AB52" box="[118,295,568,587]" pageId="11" pageNumber="12" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="D5642F4329A33852164CE0C8B7026927" box="[118,1122,596,1516]" pageId="11" pageNumber="12" type="mainText">
|
||||
MADAGASCAR: Antsiranana, Ambondrobe, 41.1 km 175° Vohemar, 13.7153 S, 50.1017 E, 10 m, littoral rainforest, 30.XI.-1.XII.2004 (B.L. Fisher); Antsiranana,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="D4F3ACEFD4618FC136656047B5FB3228" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="E5B55392DC2BDED984BBFB7C1FEA43B8" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
d'Ambre, 3.5 km 235° SW Sakaramy, 12.4689 S, 49.2422 E, 325 m, tropical dry forest, 26.-31.I.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana, Ampahana, 18 km N Antalaha, 14° 46' S, 50° 13' E, 1 m, lowland forest, 5.II.1991 (G.D. Alpert); Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="CD0585306C0827E631022C5D050B1AD5" originalValue="Forêt">Foret</normalizedToken>
|
||||
d' Andavakoera, 21.4 km 75° ENE Ambilobe, 4.6 km 356° N Betsiaka, 13.1183 S, 49.23 E, 425 m, rainforest, 15.XII.2003 (B.L. Fisher); Antsiranana, 5 km SW Antalaha, 14° 56' 17" S, 50° 15' 42" E, 50 m, secondary forest, 10.II.1991 (G.D. Alpert); Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="AFBA1CF3CA3100B1B6B7C666795C0BF7" originalValue="Forêt">Foret</normalizedToken>
|
||||
d' Antsahabe, 11.4 km 275° W Daraina, 13.2117 S, 49.5567 E, 550 m, tropical dry forest, 12.XII.2003-16.XI.2004 (B.L. Fisher); Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="A5F04182E85578061EA0E08E4A4C8FF8" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Binara, 7.5 km 230° SW Daraina, 13.255 S, 49.6167 E, 375 m, tropical dry forest, 1.-2.XII.2003 (B.L. Fisher); Antsiranana, 6 km N Cap Est, 5 m, 20.I.1991 (G.D. Alpert); Antsiranana, R.S. Manongarivo, 10.8 km 229° SW Antanambao, 13.9767 S, 48.4233 E, 780 m, rainforest, 11.-17.X.1998 (B.L. Fisher); Antsiranana, R.S. Manongarivo, 10.8 km 229° SW Antanambao, 13.9617 S, 48.4333 E, 400 m, rainforest, 8.-13.XI.1998 (B.L. Fisher); Antsiranana, 2 km NE Marofinaritra, 15° 3' S, 50° 9' E, 50 m, lowland forest, 8.II.1991 (G.D. Alpert); Antsiranana, Nosy Be, 4 km ENE Andoany, (=Helleville), 13° 25' S, 48° 18' E, 200 m, rainforest, 2.V.1989 (P.S. Ward); Antsiranana, Nosy Be,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="7F5622C50E1B5335B4734E6A4863F616" originalValue="Réserve">Reserve</normalizedToken>
|
||||
Naturelle
|
||||
<normalizedToken originalValue="Intégrale">Integrale</normalizedToken>
|
||||
<normalizedToken id="C67A258F133C6318B61C6FBC4865828E" originalValue="Intégrale">Integrale</normalizedToken>
|
||||
de Lokobe, 6.3 km 112° ESE Hellville, 13.4193 S, 48.3312 E, 30 m, rainforest, 19.-24.III.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana, Nosy Ngontsy, 55 km S Antalaha, 15° 15' 51.9" S, 50° 29' 21.5" E, 1m, secondary rainforest, 21.I.1991 (G.D. Alpert); Antsiranana, Sakaramy, 12.4411 S, 49.232 E, 260 m, tropical dry forest, 11.-12.V.2011 (B.L. Fisher et al.); Mahajanga,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="B977DD5B192A47FDF0E26B6A4B837F9D" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="06876F89EBBE846D427E5E239E7B5DFB" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
de Bemarivo, 23.8 km 223° SW Besalampy, 16.925 S, 44.3683 E, 30 m, tropical dry forest, 19.-23.XI.2002 (B.L. Fisher, C. Griswold et al.); Mahajanga, Parc National de Namoroka, 16.9 km 317° NW Vilanandro, 16.4067 S, 45.31 E, 100 m, tropical dry forest, 12.-16.XI.2002 (B.L. Fisher, C. Griswold et al.); Toamasina, Ivondro, pr. Tamatave (C. Keller); Toamasina, Nosy Mangabe, 7.43 km S Maroantsetra, 15.4973 S, 49.7622 E, 5 m, littoral rainforest edge, 26.VII.2007 (B.L. Fisher et al.); MAYOTTE: Mt. Benara, 12.8758 S, 45.1567 E, 425 m, 30.XI.-2.XII.2007 (B.L. Fisher et al.); Mt. Chongui, 12.8 S, 45.1 E, 360 m, forest near fallen tree, 15.II.-3.III.1999 (R. Jocque & G. DeSmet); Mt. Chongui, 12.9578 S, 45.134 E, 470 m, rainforest, 28.XI.2007 (B.L. Fisher et al.); Mt. Chongui, 12.959 S, 45.1341 E, 380 m, rainforest, 28.-30.XI.2007 (B.L. Fisher et al.); Mt. Combani, 12.8 S, 45.1333 E, 470 m, forest litter, 22.-24.II.1999 (R. Jocque & G. DeSmet); Mt. Combani, 12.8063 S, 45.1531 E, 370 m, rainforest, 25.XI.-4.XII.2007 (B.L. Fisher et al.); Mt. Combani, 12.8049 S, 45.1527 E, 460 m, rainforest, 29.XI.2007 (B.L. Fisher et al.); Coconi, DAF Campus, 12.8333 S, 45.1333 E, 15.I.1999 (R. Jocque & G. DeSmet); Dapani, 12.9628 S, 45.1504 E, 135 m, rainforest, 2.-4.XII.2007 (B.L. Fisher et al.); Reserve
|
||||
<normalizedToken originalValue="Forestiére">Forestiere</normalizedToken>
|
||||
<taxonomicName pageId="11" pageNumber="12" rank="tribe" tribe="Majimbini">Majimbini</taxonomicName>
|
||||
<normalizedToken id="476FA19414F53D5B242B39C4204944A7" originalValue="Forestiére">Forestiere</normalizedToken>
|
||||
<taxonomicName id="830E19DD9C538A4A4780F59E80593F5A" pageId="11" pageNumber="12" rank="tribe" tribe="Majimbini">Majimbini</taxonomicName>
|
||||
, 12.768 S, 45.1861 E, 525 m, rainforest, 3.XII.2007 (B.L. Fisher et al.); Reserve
|
||||
<normalizedToken originalValue="Forestiére">Forestiere</normalizedToken>
|
||||
<taxonomicName pageId="11" pageNumber="12" rank="tribe" tribe="Majimbini">Majimbini</taxonomicName>
|
||||
<normalizedToken id="B9C4379DAA40CD671AACD8BCAE00274F" originalValue="Forestiére">Forestiere</normalizedToken>
|
||||
<taxonomicName id="10993E56FA5C9D1CF45A40F93B8A0689" pageId="11" pageNumber="12" rank="tribe" tribe="Majimbini">Majimbini</taxonomicName>
|
||||
, 12.7689 S, 45.1902 E, 350 m, rainforest, 3.XII.2007 (B.L. Fisher et al.); Tsingoni, 12.7833 S, 45.1 E, litter of shrubs on mangrove edge, 27.II.-4.III.1999 (R. Jocque & G. DeSmet).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,151 +1,151 @@
|
|||
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<taxonomicName LSID="http://species-id.net/wiki/Grynocharis_quadrilineata" authority="Melsheimer, 1844" authorityName="Melsheimer" authorityYear="1844" class="Insecta" family="Trogossitidae" genus="Grynocharis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Grynocharis quadrilineata" order="Coleoptera" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="quadrilineata">Grynocharis quadrilineata (Melsheimer, 1844)</taxonomicName>
|
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<treatment id="B3C8B92ECFA42F9AB4F2A2927828E5FA" ID-GBIF-Taxon="152033755" LSID="urn:lsid:plazi:treatment:B3C8B92ECFA42F9AB4F2A2927828E5FA" httpUri="http://treatment.plazi.org/id/B3C8B92ECFA42F9AB4F2A2927828E5FA" lastPageId="3" lastPageNumber="144" pageId="2" pageNumber="143" scope_family="Trogossitidae" scope_order="Coleoptera" scope_subFamily="Peltinae" scope_tribe="Lophocaterini">
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<taxonomicName id="CD9BD0D1DDDCFAF22E9C54DFE910B509" LSID="http://species-id.net/wiki/Grynocharis_quadrilineata" authority="Melsheimer, 1844" authorityName="Melsheimer" authorityYear="1844" genus="Grynocharis" lsidName="Grynocharis quadrilineata" pageId="2" pageNumber="143" rank="species" species="quadrilineata">Grynocharis quadrilineata (Melsheimer, 1844)</taxonomicName>
|
||||
Map 1
|
||||
</paragraph>
|
||||
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|
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|
||||
<paragraph pageId="2" pageNumber="143">Material examined.</paragraph>
|
||||
<paragraph lastPageId="3" lastPageNumber="144" pageId="2" pageNumber="143">
|
||||
<subSubSection id="98C85F497CFA1D3905469F38253C4AE4" lastPageId="3" lastPageNumber="144" pageId="2" pageNumber="143" type="material examined">
|
||||
<paragraph id="6CE2261ED61D0E3D323E6AB9DD252565" pageId="2" pageNumber="143">Material examined.</paragraph>
|
||||
<paragraph id="BC9CEA2C1A7F6A7092C3297789FD5131" lastPageId="3" lastPageNumber="144" pageId="2" pageNumber="143">
|
||||
New Brunswick, Carleton Co., Meduxnekeag Valley Nature Preserve,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="46.1907">46.1907°N</geoCoordinate>
|
||||
<geoCoordinate id="8598DBF547C12D6B2D3C80A156C1A434" direction="north" orientation="latitude" precision="5" value="46.1907">46.1907°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-67.674">67.6740°W</geoCoordinate>
|
||||
<geoCoordinate id="36162905C1AE21CE2848A3ADC4F79BC0" direction="west" orientation="longitude" precision="5" value="-67.674">67.6740°W</geoCoordinate>
|
||||
, 20.VI.2006, R. P. Webster, mature mixed forest, in
|
||||
<taxonomicName class="Agaricomycetes" family="Pleurotaceae" genus="Pleurotus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Pleurotus" order="Agaricales" pageId="2" pageNumber="143" phylum="Basidiomycota" rank="genus">Pleurotus</taxonomicName>
|
||||
<taxonomicName id="7FF47947C73177305070223206C4C608" class="Agaricomycetes" family="Pleurotaceae" genus="Pleurotus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Pleurotus" order="Agaricales" pageId="2" pageNumber="143" phylum="Basidiomycota" rank="genus">Pleurotus</taxonomicName>
|
||||
sp. on dead standing
|
||||
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="2" pageNumber="143" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
|
||||
<taxonomicName id="26F0A9059C4595398E14B7B8639A44F3" class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="2" pageNumber="143" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
|
||||
sp. (1, NBM); same locality and forest type but 7.VI.2007, R. P. Webster, under bark of standing dead beech (4, RWC); Jackson Falls, Bell Forest,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="46.22">46.2200°N</geoCoordinate>
|
||||
<geoCoordinate id="8BB6A71B9E4C365DBC1202A8CCACE7B4" direction="north" orientation="latitude" precision="5" value="46.22">46.2200°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-67.7231">67.7231°W</geoCoordinate>
|
||||
<geoCoordinate id="907A9B9A31D891C9E21E08EB2D30D439" direction="west" orientation="longitude" precision="5" value="-67.7231">67.7231°W</geoCoordinate>
|
||||
, 5-12.VII.2008, R. P. Webster, mature hardwood forest, Lindgren funnel traps (2, AFC, RWC); same locality and habitat data but 23-28.IV.2009, 28.IV-9.V.2009, 14-20.V.2009, 20-26.V.2009, 16-21.VI.2009, R. Webster & M.-A.
|
||||
<normalizedToken originalValue="Giguère">Giguere</normalizedToken>
|
||||
<normalizedToken id="FC9274E920BB2AF65655B472A69920F3" originalValue="Giguère">Giguere</normalizedToken>
|
||||
, Lindgren funnel traps (7, AFC, RWC). Queens Co., near Queenstown,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.6904">45.6904°N</geoCoordinate>
|
||||
<geoCoordinate id="4092B57CFF58FE988DBDEDA8A5149E2C" direction="north" orientation="latitude" precision="5" value="45.6904">45.6904°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.1455">66.1455°W</geoCoordinate>
|
||||
<geoCoordinate id="CE7D2279DF3C1AD83888F9F54CC5B859" direction="west" orientation="longitude" precision="5" value="-66.1455">66.1455°W</geoCoordinate>
|
||||
, 13.V.2008, R. P. Webster, old growth hardwood forest, under
|
||||
<pageBreakToken pageId="3" pageNumber="144" start="start">bark</pageBreakToken>
|
||||
<pageBreakToken id="C6156EEB4508B7A2212A6CD1701E9051" pageId="3" pageNumber="144" start="start">bark</pageBreakToken>
|
||||
of standing dead sugar maple (1, RWC); Cranberry Lake P.N.A. (Protected Natural Area),
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="46.1125">46.1125°N</geoCoordinate>
|
||||
<geoCoordinate id="6916793E532F1A25BB4DB27710DD198E" direction="north" orientation="latitude" precision="5" value="46.1125">46.1125°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-65.6075">65.6075°W</geoCoordinate>
|
||||
<geoCoordinate id="E89CE5BEBC2757037867F194A129707F" direction="west" orientation="longitude" precision="5" value="-65.6075">65.6075°W</geoCoordinate>
|
||||
, 24.IV-5.V.2009, 5-12.V.2009, 12-21.V.2009, R. Webster & M.-A.
|
||||
<normalizedToken originalValue="Giguère">Giguere</normalizedToken>
|
||||
<normalizedToken id="40F5506EB5798A61780DE9292168BD84" originalValue="Giguère">Giguere</normalizedToken>
|
||||
, old red oak forest, Lindgren funnel traps (4, AFC, RWC); same locality and habitat data but, 21.V.2009, R. Webster & M.-A.
|
||||
<normalizedToken originalValue="Giguère">Giguere</normalizedToken>
|
||||
<normalizedToken id="F6382FB03D7B3CCD37A0AE28E8264D1E" originalValue="Giguère">Giguere</normalizedToken>
|
||||
, under bark of red oak (1, AFC); same locality data and forest type, 13-25.V.2011, 25.V-7.VI.2011, M. Roy & V. Webster, Lindgren funnel traps (5, AFC, NBM). Restigouche Co., Jacquet River Gorge P.N.A.,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="47.804">47.804°N</geoCoordinate>
|
||||
<geoCoordinate id="07AD449167A230901C9B3F4169605A7F" direction="north" orientation="latitude" precision="55" value="47.804">47.804°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-65.993">65.993°W</geoCoordinate>
|
||||
<geoCoordinate id="0D2F1A6EA915F1847B57F361688EFA0D" direction="west" orientation="longitude" precision="55" value="-65.993">65.993°W</geoCoordinate>
|
||||
, 13-23.VI.2009, G. J. McBriarty (2, NBM); Dionne Brook P.N.A.,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="47.903">47.9030°N</geoCoordinate>
|
||||
<geoCoordinate id="0C3494B07499BD884B72EFBBFC368A33" direction="north" orientation="latitude" precision="5" value="47.903">47.9030°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-68.3503">68.3503°W</geoCoordinate>
|
||||
<geoCoordinate id="1ED0FFF52FBDC1E2C958FE7FF426CBBE" direction="west" orientation="longitude" precision="5" value="-68.3503">68.3503°W</geoCoordinate>
|
||||
, 30.V-15.VI.2011, M. Roy & V. Webster, old-growth northern hardwood forest, Lindgren funnel traps (4, AFC, NBM). Sunbury Co., Acadia Research Forest,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.9866">45.9866°N</geoCoordinate>
|
||||
<geoCoordinate id="40DE61D2AD9C267383B3226584006946" direction="north" orientation="latitude" precision="5" value="45.9866">45.9866°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.3841">66.3841°W</geoCoordinate>
|
||||
<geoCoordinate id="AFC322988B187C0BD77B6558E775AF06" direction="west" orientation="longitude" precision="5" value="-66.3841">66.3841°W</geoCoordinate>
|
||||
, 19-25.V.2009, 2-9.VI.2009, 9-16.VI.2009, 24-30.VI.2009, R. Webster & M.-A.
|
||||
<normalizedToken originalValue="Giguère">Giguere</normalizedToken>
|
||||
<normalizedToken id="DE5AB7B136723FE9D677E8CB909B2FCB" originalValue="Giguère">Giguere</normalizedToken>
|
||||
, mature (110-year-old) red spruce forest with scattered red maple and balsam fir, Lindgren funnel traps (4, AFC). York Co., 15 km W of Tracy off Rt. 645,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="45.6848">45.6848°N</geoCoordinate>
|
||||
<geoCoordinate id="F41DF42D6059A50EE086F2BECE353396" direction="north" orientation="latitude" precision="5" value="45.6848">45.6848°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.8821">66.8821°W</geoCoordinate>
|
||||
<geoCoordinate id="FA6762F87C6F7570E3878A50E4954E67" direction="west" orientation="longitude" precision="5" value="-66.8821">66.8821°W</geoCoordinate>
|
||||
, 19-25.V.2009, 21-28.VI.2009, R. Webster & M.-A.
|
||||
<normalizedToken originalValue="Giguère">Giguere</normalizedToken>
|
||||
<normalizedToken id="2AFF067E19D6677DF82CD7B2EC5A0C2B" originalValue="Giguère">Giguere</normalizedToken>
|
||||
, old red pine forest, Lindgren funnel traps (2, AFC); same locality and habitat data, 10-26.V.2010, 4-16.VI.2010, R. Webster & C. MacKay, Lindgren funnel traps (1, AFC, RWC); same locality and habitat data, 8-20.VI.2011, M. Roy & V. Webster, Lindgren funnel trap (1, NBM).
|
||||
</paragraph>
|
||||
<caption pageId="3" pageNumber="144">
|
||||
<paragraph pageId="3" pageNumber="144">
|
||||
<caption id="DC733002A378B7DF390BD4DBDA42AB91" pageId="3" pageNumber="144">
|
||||
<paragraph id="C923F2663988571F783278F276EA413A" pageId="3" pageNumber="144">
|
||||
Map 1. Collection localities in New Brunswick, Canada of
|
||||
<taxonomicName class="Insecta" family="Trogossitidae" genus="Grynocharis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Grynocharis quadrilineata" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="quadrilineata">Grynocharis quadrilineata</taxonomicName>
|
||||
<taxonomicName id="5C49E61216031BA6955DD7BA1D84103D" genus="Grynocharis" lsidName="Grynocharis quadrilineata" pageId="3" pageNumber="144" rank="species" species="quadrilineata">Grynocharis quadrilineata</taxonomicName>
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="3" pageNumber="144" type="collection and habitat data">
|
||||
<paragraph pageId="3" pageNumber="144">Collection and habitat data.</paragraph>
|
||||
<paragraph pageId="3" pageNumber="144">
|
||||
<subSubSection id="66615EDDC32AD9D1A61E7A95E1196783" pageId="3" pageNumber="144" type="collection and habitat data">
|
||||
<paragraph id="B8E88A090E6642767F995087E3DD4EE8" pageId="3" pageNumber="144">Collection and habitat data.</paragraph>
|
||||
<paragraph id="0B935C994D41CB11728B3FFCC6A49F6E" pageId="3" pageNumber="144">
|
||||
In New Brunswick,
|
||||
<taxonomicName class="Insecta" family="Trogossitidae" genus="Grynocharis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Grynocharis quadrilineata" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="quadrilineata">Grynocharis quadrilineata</taxonomicName>
|
||||
<taxonomicName id="D0740BC50C8D2D3A60CCF1201504C945" genus="Grynocharis" lsidName="Grynocharis quadrilineata" pageId="3" pageNumber="144" rank="species" species="quadrilineata">Grynocharis quadrilineata</taxonomicName>
|
||||
was found in mature and old hardwood forests with sugar maple (
|
||||
<taxonomicName class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acer saccharum" order="Sapindales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="saccharum">Acer saccharum</taxonomicName>
|
||||
<taxonomicName id="9C12BC741EC6FA7671411DCA432D649C" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acer saccharum" order="Sapindales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="saccharum">Acer saccharum</taxonomicName>
|
||||
Marsh.) and American beech (
|
||||
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus grandifolia" order="Fagales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="grandifolia">Fagus grandifolia</taxonomicName>
|
||||
<taxonomicName id="067756D73E9F0159E743F3FC1276EA55" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus grandifolia" order="Fagales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="grandifolia">Fagus grandifolia</taxonomicName>
|
||||
Ehrh.), an old-growth northern hardwood forest with sugar maple and yellow birch (
|
||||
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Betula alleghaniensis" order="Fagales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="alleghaniensis">Betula alleghaniensis</taxonomicName>
|
||||
<taxonomicName id="2A2FAB13BACD80FEE27D57F7719D3182" class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Betula alleghaniensis" order="Fagales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="alleghaniensis">Betula alleghaniensis</taxonomicName>
|
||||
Britt.), an old red oak (
|
||||
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus rubra" order="Fagales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="rubra">Quercus rubra</taxonomicName>
|
||||
<taxonomicName id="264253FDE00267CF0C3912E32DBA8EBA" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus rubra" order="Fagales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="rubra">Quercus rubra</taxonomicName>
|
||||
L.) forest, a mature (110-year-old) red spruce (
|
||||
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea rubens" order="Pinales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="rubens">Picea rubens</taxonomicName>
|
||||
<taxonomicName id="481804E916151AA777243B0A08ECB245" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea rubens" order="Pinales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="rubens">Picea rubens</taxonomicName>
|
||||
Sarg.) forest, and an old (180-year-old) red pine (
|
||||
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus resinosa" order="Pinales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="resinosa">Pinus resinosa</taxonomicName>
|
||||
<taxonomicName id="C25B2F98E56A8145368262AA0BAEA215" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus resinosa" order="Pinales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="species" species="resinosa">Pinus resinosa</taxonomicName>
|
||||
Ait.) forest. Adults were captured in Lindgren funnel traps in most of these forest types. Specimens with microhabitat data were collected from
|
||||
<taxonomicName class="Agaricomycetes" family="Pleurotaceae" genus="Pleurotus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Pleurotus" order="Agaricales" pageId="3" pageNumber="144" phylum="Basidiomycota" rank="genus">Pleurotus</taxonomicName>
|
||||
<taxonomicName id="15C62F285989550E71E15A5525F06C98" class="Agaricomycetes" family="Pleurotaceae" genus="Pleurotus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Pleurotus" order="Agaricales" pageId="3" pageNumber="144" phylum="Basidiomycota" rank="genus">Pleurotus</taxonomicName>
|
||||
mushrooms on a dead, standing poplar (
|
||||
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
|
||||
<taxonomicName id="90E9652B75E2EF2FFA3093ED74D1715E" class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
|
||||
sp.), from under bark of a dead, standing American beech, from under bark of a standing, dead sugar maple, and from under bark of a red oak. This species has been reported from under bark of a dead poplar (
|
||||
<bibRefCitation author="Barron, JR" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="11" pageNumber="152" pagination="1 - 143" title="A revision of the Trogossitidae of America north of Mexico." url="10.4039/entm10375fv" volume="75" year="1971">Barron 1971</bibRefCitation>
|
||||
<bibRefCitation id="8BE6F5C0B1D21305A559732CC2235C90" author="Barron, JR" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="11" pageNumber="152" pagination="1 - 143" title="A revision of the Trogossitidae of America north of Mexico." url="10.4039/entm10375fv" volume="75" year="1971">Barron 1971</bibRefCitation>
|
||||
). Adults were collected during April, May, June, and July in New Brunswick.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="3" pageNumber="144" type="distribution">
|
||||
<paragraph pageId="3" pageNumber="144">Distribution in Canada and Alaska.</paragraph>
|
||||
<paragraph pageId="3" pageNumber="144">
|
||||
<subSubSection id="45190FD2B0D271E5F54418C7128D323E" pageId="3" pageNumber="144" type="distribution">
|
||||
<paragraph id="83C1EE58ABDEE52ECF52752057700680" pageId="3" pageNumber="144">Distribution in Canada and Alaska.</paragraph>
|
||||
<paragraph id="658EFFBD410E9BB94950AF8F997D3F6A" pageId="3" pageNumber="144">
|
||||
MB, ON, QC, NB, NS (
|
||||
<bibRefCitation pageId="3" pageNumber="144">Bousquet 1991</bibRefCitation>
|
||||
<bibRefCitation id="FC2BE0489E68B795D0D82F3B30FA7DD6" pageId="3" pageNumber="144">Bousquet 1991</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka 2011</bibRefCitation>
|
||||
<bibRefCitation id="9FC314AA3CDE19183228D28EE0BF89FE" author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka 2011</bibRefCitation>
|
||||
). This species was first reported from Nova Scotia and the Maritime provinces by
|
||||
<bibRefCitation author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka (2011)</bibRefCitation>
|
||||
<bibRefCitation id="1B67B192A086544A436F2236E8E8E0F0" author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka (2011)</bibRefCitation>
|
||||
based on one specimen and was considered regionally rare. This species appears to be widespread in New Brunswick and was commonly detected using Lindgren funnel traps.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,100 +1,99 @@
|
|||
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="adamsi">Tetramorium adamsi</taxonomicName>
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<taxonomicName id="925211325503DC0173018DA3BD34BC88" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="adamsi">Tetramorium adamsi</taxonomicName>
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can be straightforwardly recognised within the
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
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<taxonomicName id="490A4CA5466710C3D445D32E52355712" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
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species complex since it is the only species in which the posterodorsal margin of the petiolar node is situated higher than the anterodorsal margin.
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<paragraph box="[118,226,1129,1147]" pageId="65" pageNumber="66" type="mainText">Description</paragraph>
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<paragraph box="[118,1117,1157,1317]" pageId="65" pageNumber="66" type="mainText">HL 0.94-1.06 (1.01); HW 0.89-1.03 (0.96); SL 0.69-0.82 (0.76); EL 0.17-0.21 (0.19); PH 0.43-0.55 (0.50); PW 0.63-0.78 (0.72); WL 1.13-1.45 (1.30); PSL 0.34-0.51 (0.44); PTL 0.31-0.39 (0.36); PTH 0.34-0.42 (0.38); PTW 0.25-0.31 (0.28); PPL 0.29-0.32 (0.31); PPH 0.34-0.42 (0.38); PPW 0.34-0.39 (0.36); CI 93-97 (95); SI 73-84 (79); OI 18-22 (20); DMI 51-59 (56); LMI 34-40 (38); PSLI 36-50 (43); PeNI 36-41 (39); LPeI 88-97 (93); DPeI 76-84 (79); PpNI 47-54 (50); LPpI 75-86 (81); DPpI 113-126 (117); PPI 116-138 (127) (18 measured).</paragraph>
|
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|
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<paragraph id="8EC47523C63D7889C95261F04F1EF874" box="[118,226,1129,1147]" pageId="65" pageNumber="66" type="mainText">Description</paragraph>
|
||||
<paragraph id="2B98E78E5464C58C9CBD0ABF06209986" box="[118,1117,1157,1317]" pageId="65" pageNumber="66" type="mainText">HL 0.94-1.06 (1.01); HW 0.89-1.03 (0.96); SL 0.69-0.82 (0.76); EL 0.17-0.21 (0.19); PH 0.43-0.55 (0.50); PW 0.63-0.78 (0.72); WL 1.13-1.45 (1.30); PSL 0.34-0.51 (0.44); PTL 0.31-0.39 (0.36); PTH 0.34-0.42 (0.38); PTW 0.25-0.31 (0.28); PPL 0.29-0.32 (0.31); PPH 0.34-0.42 (0.38); PPW 0.34-0.39 (0.36); CI 93-97 (95); SI 73-84 (79); OI 18-22 (20); DMI 51-59 (56); LMI 34-40 (38); PSLI 36-50 (43); PeNI 36-41 (39); LPeI 88-97 (93); DPeI 76-84 (79); PpNI 47-54 (50); LPpI 75-86 (81); DPpI 113-126 (117); PPI 116-138 (127) (18 measured).</paragraph>
|
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||||
Head longer than wide (CI 93-97); posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, and narrow, without defined posterior and ventral margins. Antennal scapes short to moderate in length, not reaching posterior head margin (SI 73-84). Eyes short to moderately sized (OI 18-22). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 34-40). Propodeal spines very long to extremely long, spinose and acute (PSLI 36-50); propodeal lobes well-developed, triangular to elongate-triangular, and usually acute. Petiolar node in profile rectangular nodiform, approximately 1.0 to 1.1 times higher than long (LPeI 81-88), anterior
|
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<pageBreakToken pageId="66" pageNumber="67" start="start">and</pageBreakToken>
|
||||
<pageBreakToken id="73A8133B52567FCDD1B5EA5EA9700A6A" pageId="66" pageNumber="67" start="start">and</pageBreakToken>
|
||||
posterior faces approximately parallel, posterodorsal margin situated higher than anterodorsal margin, dorsum convex; node in dorsal view approximately 1.2 to 1.3 times longer than wide (DPeI 76-84). Postpetiole in profile subglobular, approximately 1.2 to 1.3 times higher than long (LPpI 75-86); in dorsal view around 1.1 to 1.3 times wider than long (DPpI 113-126). Postpetiole in profile generally appearing less voluminous than petiolar node, in dorsal view between 1.1 to 1.4 time s wider than petiolar node (PPI 116-138). Mandibles striate; clypeus longitudinally rugose/rugulose, with four to eight rugae/rugulae, median ruga usually present and distinct, but rarely more conspicuous than other rugae/rugulae; cephalic dorsum between frontal carinae with eight to 13 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted and with cross-meshes; lateral and ventral head mostly reticulate-rugose. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae with well-developed and conspicuous longitudinal rugae. Waist segments strongly rugose. Gaster mostly unsculptured, smooth, and shining, several specimens from higher elevations with dist inct costulae on the basal half of first gastral tergite. Ground sculpture generally faint to absent everywhere on body, better developed at higher elevations, but still weak. All dorsal surfaces with abundant, long, and fine st anding hairs. Anterior edges of antennal scapes with suberect to erect hairs. Body generally uniformly dark brown to black in colour, sometimes appendages of slightly lighter colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,596,615]" pageId="66" pageNumber="67" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1120,625,700]" pageId="66" pageNumber="67" type="mainText">
|
||||
<subSubSection id="016FBF00C2E3FE04E635B509D685B7AB" type="discussion">
|
||||
<paragraph id="89CC45BE79AF4ED6E8CCF274060DF2D4" box="[118,170,596,615]" pageId="66" pageNumber="67" type="mainText">Notes</paragraph>
|
||||
<paragraph id="27E940B839D1AD3AAB17A5F7158F3FC2" box="[118,1120,625,700]" pageId="66" pageNumber="67" type="mainText">
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||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="5B35F4E997EB692E5A1552D36EAAB565" rank="species">new species</taxonomicNameLabel>
|
||||
is currently only known from Manongarivo and Bemanevika, where it was collected from montane forest leaf litter at elevations from 1175 to 1860 m. Almost all of the material was sampled in Manongarivo while just one specimen from Bemanevika was available.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1122,709,1094]" pageId="66" pageNumber="67" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="adamsi">Tetramorium adamsi</taxonomicName>
|
||||
<subSubSection id="D7DC45A436547E8A7E0A096E84CA5DFF" type="materials_examined">
|
||||
<paragraph id="1E8D37E6081F1A0F6799DE2FF7F6947D" box="[118,1122,709,1094]" pageId="66" pageNumber="67" type="mainText">
|
||||
<taxonomicName id="BDE1A1123DBB10B721EE93D6254196A1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="adamsi">Tetramorium adamsi</taxonomicName>
|
||||
shows a remarkable variation in Manongarivo. The examined material was collected from three sub-localities located at different altitudes separated by few kilometres from each other. The lowest was situated at 1175 m, the middle at 1580, and the highest at 1860 m. The specimens from 1175 m had longer antennal scapes (SI 81-84), longer propodeal spines (PSLI 47-50), and a narrower postpetiole in relation to the petiolar node (PPI 116-125) than specimens from the higher elevations (SI 73-78; PSLI 36-41; PPI 129-138). These differences were fairly constant between the series, and we have considered separating the material into two species. However, the only arguments for separation are morphometric divergences, and apart from this there is a great amount of morphological similarity. For these reasons, we have decided against a split. In addition, we found that on half of the workers from the highest elevation at 1860 m, the basal half of the first gastral tergite is distinctly costulate while it is unsculptured and smooth in the remaining specimens. Sculpture on the first gastral tergite is generally of high diagnostic importance in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="B97D079263282CDF08BDE99B14AF8B45" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group, and could have been a reason to split the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
<taxonomicName id="7009B069EC0F3DE374A39E70FB6188D6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
material. Nevertheless, no other character could separate the sculptured specimens from those collected at the two higher sub-localities. This has convinced us to keep all the material as one variable, but well-defined species.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,1103,1178]" pageId="66" pageNumber="67" type="mainText">
|
||||
<paragraph id="6484143644A0C27741BFF1F5D936A491" box="[118,1121,1103,1178]" pageId="66" pageNumber="67" type="mainText">
|
||||
Within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="C376DD4C900EEBFA312E877A94271872" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
species complex, it is fairly easy to discriminate
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
<taxonomicName id="FF0F6DBDA91DC37ABCE5CCA69957831F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
from the other species on the basis of the petiolar node shape. The node has the posterodorsal margin situated higher than the anterodorsal margin in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi,</taxonomicName>
|
||||
<taxonomicName id="9FAB77905CD040EB28E8A735D3A856E9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi,</taxonomicName>
|
||||
whereas both margins are at about the same height in the rest of the complex.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="66" pageNumber="67" type="etymology">
|
||||
<paragraph box="[118,219,1215,1233]" pageId="66" pageNumber="67" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,950,1243,1291]" pageId="66" pageNumber="67" type="mainText">
|
||||
<subSubSection id="0B6383E8409657BB8C862FBF6277441B" pageId="66" pageNumber="67" type="etymology">
|
||||
<paragraph id="90C4E0142ACB253D2258346F023AF735" box="[118,219,1215,1233]" pageId="66" pageNumber="67" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="1A2FFBE4FC295472F1351A8545FD3299" box="[118,950,1243,1291]" pageId="66" pageNumber="67" type="mainText">
|
||||
The name of the
|
||||
<taxonomicNameLabel pageId="66" pageNumber="67" rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="B5BBAD28F4D2ABF14260E90E03DC2F05" pageId="66" pageNumber="67" rank="species">new species</taxonomicNameLabel>
|
||||
is a patronym in honour of the British writer Douglas Adams best known for "The Hitchhiker's Guide to the Galaxy" series. (1952-2001.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,1328,1346]" pageId="66" pageNumber="67" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,1356,1516]" pageId="66" pageNumber="67" type="mainText">MADAGASCAR: Antsiranana, R.S. Manongarivo, 14.5 km 220° SW Antanambao, 13.99833 S, 48.42833 E, 1175 m, montane rainforest, 19.-25.X.1998 (B.L. Fisher); Mahajanga, Bemanevika, Souspref. de Bealanana, forest, 20. X. 1975 (A. Peyrieras); Antsiranana, R.S. Manongarivo 17.3 km 218° SW Antanambao, 14.02167 S, 48.41833 E, 1580 m, montane rainforest, 27.X.1998 (B.L. Fisher); Antsiranana, R.S. Manongarivo, 20.4 km 219° SW Antanambao, 14.04667 S, 48.40167 E, 1860 m, montane rainforest, 3.XI.1998 (B.L. Fisher); Mahajanga, Bemanevika, Soupref. de Bealanana, 20.X.1975 (A. Peyrieras).</paragraph>
|
||||
<subSubSection id="BBFF42E32EDA922B409096C804536735" type="materials_examined">
|
||||
<paragraph id="87FDFE88B9CB4A06593D2C802E0B7041" box="[118,295,1328,1346]" pageId="66" pageNumber="67" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="94223F12284DF6FAB66450849B89E208" box="[118,1122,1356,1516]" pageId="66" pageNumber="67" type="mainText">MADAGASCAR: Antsiranana, R.S. Manongarivo, 14.5 km 220° SW Antanambao, 13.99833 S, 48.42833 E, 1175 m, montane rainforest, 19.-25.X.1998 (B.L. Fisher); Mahajanga, Bemanevika, Souspref. de Bealanana, forest, 20. X. 1975 (A. Peyrieras); Antsiranana, R.S. Manongarivo 17.3 km 218° SW Antanambao, 14.02167 S, 48.41833 E, 1580 m, montane rainforest, 27.X.1998 (B.L. Fisher); Antsiranana, R.S. Manongarivo, 20.4 km 219° SW Antanambao, 14.04667 S, 48.40167 E, 1860 m, montane rainforest, 3.XI.1998 (B.L. Fisher); Mahajanga, Bemanevika, Soupref. de Bealanana, 20.X.1975 (A. Peyrieras).</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,102 +1,101 @@
|
|||
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|
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="andrei">Tetramorium andrei</taxonomicName>
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|
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|
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<taxonomicName id="BFF1D38F803B3BE23482DB0EB5329526" ID-CoL="55PHN" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="andrei">Tetramorium andrei</taxonomicName>
|
||||
species complex
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,1122,1248,1520]" pageId="19" pageNumber="20" type="mainText">
|
||||
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|
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|
||||
This species complex is the largest within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="931B508368CFF6FBE8693F73193A5951" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group with the nine species T.
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="ala">
|
||||
<taxonomicName id="E5CF3032301C19E71EED18CA40BF1B50" authorityName="," genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="ala">
|
||||
ala
|
||||
<authority>,</authority>
|
||||
<authority id="C92082AB8948188DEC5C38283BDEC23D">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName genus="Tetamorium" pageId="19" pageNumber="20" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName authorityName="," genus="Tetramoirum" pageId="19" pageNumber="20" rank="species" species="andohahela">
|
||||
<taxonomicName id="B148F6BAEC0CECC9D65F215A1C13F62B" genus="Tetamorium" pageId="19" pageNumber="20" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="1199749B7C8543AFACA29014D0A68F2F" authorityName="," genus="Tetramoirum" pageId="19" pageNumber="20" rank="species" species="andohahela">
|
||||
andohahela
|
||||
<authority>,</authority>
|
||||
<authority id="EBA21504CFB400FDFF8058F3394079CB">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="andrei">
|
||||
<taxonomicName genus="Tetamorium" pageId="19" pageNumber="20" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="78BB4C0430ABAC3D64BDBA50BB832F6C" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="andrei">
|
||||
<taxonomicName id="BB891D8E8DA3A6A77BADF2719DBE95CA" genus="Tetamorium" pageId="19" pageNumber="20" rank="genus">T.</taxonomicName>
|
||||
andrei
|
||||
<authority>,</authority>
|
||||
<authority id="8E45A436621243F8C34D5F088A23CFDA">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="isectum">
|
||||
<taxonomicName genus="Tetamorium" pageId="19" pageNumber="20" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="6E4485B35702BB2ADAB65282C26F533E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="isectum">
|
||||
<taxonomicName id="49A50FE4E8C582E894BB6EB35818C427" genus="Tetamorium" pageId="19" pageNumber="20" rank="genus">T.</taxonomicName>
|
||||
elf,
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="isectum">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="isectum">
|
||||
<taxonomicName id="54D4C1858FBE03C6DCCBAFADCC03B93C" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="isectum">
|
||||
<taxonomicName id="34B0546631A968B25FE1DED8544B6B3F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="isectum">
|
||||
T. isectum
|
||||
<authority>,</authority>
|
||||
<authority id="E7C1C064D5F10D98D95E9B5DE06FB2EE">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
<taxonomicName id="0E52FAA88764A553181BAECD41AEC58F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="01A5ECB0E30E67750E53952C1F15A3B1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="8ABDC923F5289D6861BDE586F3B9C8F5" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="F7A91C6C21EC55DFE26D92F1C018B1E4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="electrum">
|
||||
T. electrum
|
||||
<authority>,</authority>
|
||||
<authority id="CD3E0590C13F66189C9CECD590B97023">,</authority>
|
||||
</taxonomicName>
|
||||
</taxonomicName>
|
||||
<taxonomicName genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="nify">T. nify,</taxonomicName>
|
||||
<taxonomicName id="FE3256059EF24275FC49B48512CB1029" genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="nify">T. nify,</taxonomicName>
|
||||
and
|
||||
<taxonomicName genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="voasary">T. voasary.</taxonomicName>
|
||||
<taxonomicName id="79A2660F9B83474AEC289A81838D42A6" genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="voasary">T. voasary.</taxonomicName>
|
||||
The forecoxae are usually completely or mostly unsculptured, smooth, and shining, although sculpture is sometimes present. What sculpture is present is mostly superficial and generally does not consist of strongly longitudinally arranged rugae. Instead, it is reticulate-punctate with few superimposed rugulae or traces of rugulae. In a few species (e.g.
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="elf">
|
||||
<taxonomicName id="490456CF51E327E5F2E0E3F0BB8491EA" authorityName="," genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="elf">
|
||||
T. elf
|
||||
<authority>,</authority>
|
||||
<authority id="E02C741C9049FD6B4178D67FF62F2EEB">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="voasary">T. voasary)</taxonomicName>
|
||||
<taxonomicName id="709A6038CBDD17BACB494BBB436FAAB3" genus="Tetamorium" pageId="19" pageNumber="20" rank="species" species="voasary">T. voasary)</taxonomicName>
|
||||
the forecoxae are partly rugulose, but the sculpture is comparatively weak, and never covers the whole coxa as in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="F26B928EFEF96A2240DB9433A423774B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
species complex. The sculpture may appear longitudinal at first glance, but this is due to sections of linearly arranged reticulate-punctate sculpture, not true rugae/rugulae (e.g.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum).</taxonomicName>
|
||||
<taxonomicName id="E7A91FD9495F645E2EA5BF7C5C323846" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum).</taxonomicName>
|
||||
The first gastral tergite is always completely devoid of any sculpture, and fairly smooth and shining.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,132 +1,131 @@
|
|||
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<mods:namePart>Hita Garcia, F.</mods:namePart>
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<mods:namePart>B. L. Fisher</mods:namePart>
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<mods:namePart id="DD1907B1CFB0936A9639152E152ADB57">B. L. Fisher</mods:namePart>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172641" ID-GBIF-Taxon="100105318" ID-Zenodo-Dep="6172641" LSID="urn:lsid:plazi:treatment:C890408D62CC153A4BEACFD982AAE259" httpUri="http://treatment.plazi.org/id/C890408D62CC153A4BEACFD982AAE259" lastPageNumber="61" pageNumber="59">
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<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,634,343,363]" pageId="58" pageNumber="59" type="mainText">
|
||||
<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ambanizana">
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<treatment id="C890408D62CC153A4BEACFD982AAE259" ID-DOI="http://doi.org/10.5281/zenodo.6172641" ID-GBIF-Taxon="100105318" ID-Zenodo-Dep="6172641" LSID="urn:lsid:plazi:treatment:C890408D62CC153A4BEACFD982AAE259" httpUri="http://treatment.plazi.org/id/C890408D62CC153A4BEACFD982AAE259" lastPageNumber="61" pageNumber="59" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="0F0AE03034F7AC6459A158064FB9FE36" type="nomenclature">
|
||||
<paragraph id="B2B82AF8C180B80C80D13CB916F50D6E" box="[118,634,343,363]" pageId="58" pageNumber="59" type="mainText">
|
||||
<taxonomicName id="4F88946810EC677490C6AED01245200F" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ambanizana">
|
||||
Tetramorium ambanizana
|
||||
<authority>Hita</authority>
|
||||
<authority id="D2BFAC5E7EE4B68318F1DF9B42FF10B1">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="A5C53A221292FE0DC997C1AFEBD0F63A" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,408,371,390]" pageId="58" pageNumber="59" type="mainText">(Figs. 42, 43, 114, 115, 116, 142)</paragraph>
|
||||
<subSubSection id="EA6D90DB6CDBDC10C6696E2A44519F82" type="description">
|
||||
<paragraph id="FBA24135DF05FE2112FACE7CD7BE9153" box="[118,408,371,390]" pageId="58" pageNumber="59" type="mainText">(Figs. 42, 43, 114, 115, 116, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1122,425,525]" pageId="58" pageNumber="59" type="mainText">Holotype worker, MADAGASCAR, Toamasina, 5.3 km SSE Ambanizana, Andranobe, 15.66667 S, 49.96667 E, 425 m, rainforest, leaf litter (sifted mold, rotten wood), collection code BLF00926, 21.XI.2003 (B.L. Fisher) (CASC: CASENT0189238). Paratypes, four workers with same data as holotype (CASC: CASENT0189214, CASENT0218048, CASENT0218050, CASENT0270779).</paragraph>
|
||||
<subSubSection id="70C2B5FD5AD7E885FA37E959C4439C6E" type="materials_examined">
|
||||
<paragraph id="037D8D18579FE76F30484788104FA4AE" box="[118,1122,425,525]" pageId="58" pageNumber="59" type="mainText">Holotype worker, MADAGASCAR, Toamasina, 5.3 km SSE Ambanizana, Andranobe, 15.66667 S, 49.96667 E, 425 m, rainforest, leaf litter (sifted mold, rotten wood), collection code BLF00926, 21.XI.2003 (B.L. Fisher) (CASC: CASENT0189238). Paratypes, four workers with same data as holotype (CASC: CASENT0189214, CASENT0218048, CASENT0218050, CASENT0270779).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,560,578]" pageId="58" pageNumber="59" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,587,715]" pageId="58" pageNumber="59" type="mainText">
|
||||
<subSubSection id="E71C79C1694148D4A00992AFAE057454" type="diagnosis">
|
||||
<paragraph id="1540BCF30758C4E15B6757D3BE62F07F" box="[118,208,560,578]" pageId="58" pageNumber="59" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="968CBA013FDD5C535FE697AABD2E6CDF" box="[118,1121,587,715]" pageId="58" pageNumber="59" type="mainText">
|
||||
Within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="C69B461973509FF0A2239A3598BC0385" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
species complex
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
<taxonomicName id="A5706BDFC825903585A6BAD61237FD2F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
can be recognised by the following character combination: head weakly longer than wide (CI 97-98); propodeal spines extremely long (PSLI 60-65); petiolar node rectangular nodiform with the posterodorsal margin situated much higher than the anterodorsal margin, and the dorsum convex; postpetiole in profile around 1.2 to 1.3 times higher than long (LPpI 77-81), and in dorsal view approximately 1.1 times wider than long (DPpI 104-112).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,749,768]" pageId="58" pageNumber="59" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,777,904]" pageId="58" pageNumber="59" type="mainText">HL 0.85-0.95 (0.92); HW 0.82-0.93 (0.90); SL 0.68-0.76 (0.73); EL 0.20-0.21 (0.20); PH 0.43-0.48 (0.46); PW 0.61-0.69 (0.66); WL 1.08-1.23 (1.17); PSL 0.55-0.58 (0.57); PTL 0.31-0.35 (0.33); PTH 0.38-0.43 (0.40); PTW 0.26-0.30 (0.28); PPL 0.33-0.35 (0.34); PPH 0.41-0.46 (0.43); PPW 0.36-0.38 (0.37); CI 97-98 (98); SI 78-83 (81); OI 21-24 (22); DMI 56-58 (57); LMI 38-41 (39); PSLI 60-65 (62); PeNI 40-43 (42); LPeI 79-83 (81); DPeI 82-86 (84); PpNI 54-58 (56); LPpI 77-81 (79); DPpI 104-112 (108); PPI 127-138 (134) (five measured).</paragraph>
|
||||
<paragraph box="[118,1122,912,1581]" pageId="58" pageNumber="59" type="mainText">Head weakly longer than wide (CI 97-98); posterior head margin moderately concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, moderately deep, but narrow and without defined posterior and ventral margins. Antennal scapes short to moderate, not reaching posterior head margin (SI 78-83). Eyes small to moderate (OI 21-24). Mesosomal outline in profile weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-41). Propodeal spines extremely long, spinose, and acute (PSLI 60-65); propodeal lobes well-developed, triangular to elongate-triangular, and acute. Petiolar node in profile rectangular nodiform but comparatively high, approximately 1.2 to 1.3 times higher than long (LPeI 79-83), anterior and posterior faces approximately parallel, posterodorsal margin situated higher than anterodorsal, dorsum convex; node in dorsal view approximately 1.2 times longer than wide (DPeI 82-86). Postpetiole in profile subglobular and anteroposteriorly compressed, approximately 1.2 to 1.3 times higher than long (LPpI 77-81); in dorsal view around 1.1 times wider than long (DPpI 104-112). Postpetiole in profile higher and less voluminous than petiolar node, in dorsal view approximately 1.3 to 1.4 times wider than petiolar node (PPI 127-138). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugulose, with one to six rugulae, median rugula always present, remaining rugulae variably developed; cephalic dorsum between frontal carinae with six to nine longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted and none with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose, very rarely with cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured, smooth, and shining, with weak, superficial ground sculpture. Waist segments strongly and very regularly longitudinally rugose, rugae absent from anterior face, but running unbroken from one side to other through posterior face. Base of first gastral tergite distinctly costulate, remainder of gaster unsculptured, smooth and shining. Ground sculpture generally faint to absent everywhere on body. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs; first gastral tergite without appressed pubescence. Anterior edges of antennal scapes suberect to erect hairs. Body uniform dark brown to black in colour.</paragraph>
|
||||
<subSubSection id="67243CB0C20767041CE52BC70B247F05" type="description">
|
||||
<paragraph id="73B49A3BF71A42EF9EDE6CA367D63513" box="[118,226,749,768]" pageId="58" pageNumber="59" type="mainText">Description</paragraph>
|
||||
<paragraph id="EC2F09DAC96995DD2B1AB9FFB3A50FD3" box="[118,1121,777,904]" pageId="58" pageNumber="59" type="mainText">HL 0.85-0.95 (0.92); HW 0.82-0.93 (0.90); SL 0.68-0.76 (0.73); EL 0.20-0.21 (0.20); PH 0.43-0.48 (0.46); PW 0.61-0.69 (0.66); WL 1.08-1.23 (1.17); PSL 0.55-0.58 (0.57); PTL 0.31-0.35 (0.33); PTH 0.38-0.43 (0.40); PTW 0.26-0.30 (0.28); PPL 0.33-0.35 (0.34); PPH 0.41-0.46 (0.43); PPW 0.36-0.38 (0.37); CI 97-98 (98); SI 78-83 (81); OI 21-24 (22); DMI 56-58 (57); LMI 38-41 (39); PSLI 60-65 (62); PeNI 40-43 (42); LPeI 79-83 (81); DPeI 82-86 (84); PpNI 54-58 (56); LPpI 77-81 (79); DPpI 104-112 (108); PPI 127-138 (134) (five measured).</paragraph>
|
||||
<paragraph id="6F747C7E15F2D1DEF1A617A12BD50D5A" box="[118,1122,912,1581]" pageId="58" pageNumber="59" type="mainText">Head weakly longer than wide (CI 97-98); posterior head margin moderately concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, moderately deep, but narrow and without defined posterior and ventral margins. Antennal scapes short to moderate, not reaching posterior head margin (SI 78-83). Eyes small to moderate (OI 21-24). Mesosomal outline in profile weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-41). Propodeal spines extremely long, spinose, and acute (PSLI 60-65); propodeal lobes well-developed, triangular to elongate-triangular, and acute. Petiolar node in profile rectangular nodiform but comparatively high, approximately 1.2 to 1.3 times higher than long (LPeI 79-83), anterior and posterior faces approximately parallel, posterodorsal margin situated higher than anterodorsal, dorsum convex; node in dorsal view approximately 1.2 times longer than wide (DPeI 82-86). Postpetiole in profile subglobular and anteroposteriorly compressed, approximately 1.2 to 1.3 times higher than long (LPpI 77-81); in dorsal view around 1.1 times wider than long (DPpI 104-112). Postpetiole in profile higher and less voluminous than petiolar node, in dorsal view approximately 1.3 to 1.4 times wider than petiolar node (PPI 127-138). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugulose, with one to six rugulae, median rugula always present, remaining rugulae variably developed; cephalic dorsum between frontal carinae with six to nine longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted and none with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose, very rarely with cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured, smooth, and shining, with weak, superficial ground sculpture. Waist segments strongly and very regularly longitudinally rugose, rugae absent from anterior face, but running unbroken from one side to other through posterior face. Base of first gastral tergite distinctly costulate, remainder of gaster unsculptured, smooth and shining. Ground sculpture generally faint to absent everywhere on body. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs; first gastral tergite without appressed pubescence. Anterior edges of antennal scapes suberect to erect hairs. Body uniform dark brown to black in colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,118,136]" pageId="60" pageNumber="61" type="mainText">
|
||||
<pageBreakToken pageId="60" pageNumber="61" start="start">Notes</pageBreakToken>
|
||||
<subSubSection id="E822E0BBC3B00B9DD8596484CAE6D64C" type="discussion">
|
||||
<paragraph id="06FC8C736EE7890CEEE0DFACE75ED4C7" box="[118,170,118,136]" pageId="60" pageNumber="61" type="mainText">
|
||||
<pageBreakToken id="8F0525C63C3CFF3961E4F1E7AD0E95B2" pageId="60" pageNumber="61" start="start">Notes</pageBreakToken>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,146,192]" pageId="60" pageNumber="61" type="mainText">
|
||||
<paragraph id="72441D061CBA61AD9AE062C3B1E12536" box="[118,1121,146,192]" pageId="60" pageNumber="61" type="mainText">
|
||||
At present,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
<taxonomicName id="168D3A3C07803AEA1E49DF58F13F24A2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
is only known from the type series from Ambanizana where it was collected from rainforest at an elevation of 425 m. The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="E08C29052E313F5DA44BF37107FABEB0" rank="species">new species</taxonomicNameLabel>
|
||||
appears to inhabit leaf litter.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,201,577]" pageId="60" pageNumber="61" type="mainText">
|
||||
<paragraph id="2B0088D0BA53C3346FDB8DD440F0A0F3" box="[118,1122,201,577]" pageId="60" pageNumber="61" type="mainText">
|
||||
Within the species complex its petiolar node shape distinguishes it from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
<taxonomicName id="C11949E69A97D7BB6D50EE435B3D06E8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
<taxonomicName id="D88DE31CEB23DCBD38F741852445CDAC" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
since the latter two have a node with the anterodorsal and posterodorsal margins at about the same height. The node of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
<taxonomicName id="6DDEC4B5D5063B267EA0AB657E38613C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
has a posterodorsal margin situated much higher than the anterodorsal margin. This node development is shared with the remaining species,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli,</taxonomicName>
|
||||
<taxonomicName id="537374D2430AD0B32C9724A7630CB14C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli,</taxonomicName>
|
||||
which is generally fairly similar to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana.</taxonomicName>
|
||||
<taxonomicName id="ADD10F444C7FB21FF6CF74E7D6DF78F4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana.</taxonomicName>
|
||||
Both are described as new and distinct species here, but it is possible that they are conspecific. Their general appearance is very much alike, and differentiating between the two requires measuring a few key characters.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">Tetramorium ambanizana</taxonomicName>
|
||||
<taxonomicName id="758C093DDE05BD4FC6EC797D3803F39B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">Tetramorium ambanizana</taxonomicName>
|
||||
is only known from the type locality, whereas
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="D32A6744A747D352ED74308D1AD17FA7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
is found in several localities further north. This could mean that the first might be a geographic variation of the latter. However, we consider the following morphometric differences sufficient to consider them different species until more material, especially from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana,</taxonomicName>
|
||||
<taxonomicName id="07A21F3AF034BC6EC72F4CD5DE06A4BC" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana,</taxonomicName>
|
||||
becomes available. The head of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="944413644B0505283BB4772F306CD9EF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
is always distinctly longer than wide (CI 92-95) while it is only weakly longer than wide in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
<taxonomicName id="CCBA569A426AE42B9A097CFFA2346FEA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
(CI 97-98). In addition, both species have extremely long propodeal spines, but the spines of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
<taxonomicName id="6B9DE352A96DE1B4E4C2CEF6AA6D1643" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
are still much longer (PSLI 60-65) than the spines of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="0E0BD83F9A57EBEF1DEC0622912C2130" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
(PSLI 38-48). The third separating character is the shape of the postpetiole, which is relatively higher and broader in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="099B78B1BB4DA9A44FFC0DFB6AC1E773" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
(LPpI 68-72; DPpI 119-129) than in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
<taxonomicName id="F928B58E6E7F68384BE75B49D6A323E5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
(LPpI 77-81; DPpI 104-112).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,613,631]" pageId="60" pageNumber="61" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,1090,641,660]" pageId="60" pageNumber="61" type="mainText">
|
||||
<subSubSection id="153BD5E8C2A1A5F0EF89C8CF73093A57" type="etymology">
|
||||
<paragraph id="981C433EA2113938CE38A53D65F2F7E8" box="[118,219,613,631]" pageId="60" pageNumber="61" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="9676F58774164BBC97909C1635F303A2" box="[118,1090,641,660]" pageId="60" pageNumber="61" type="mainText">
|
||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="F9B9C715392D53BF4F4F2896C2F3244D" rank="species">new species</taxonomicNameLabel>
|
||||
is named after the type locality. The species epithet is a noun in apposition, and thus invariant.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,696,714]" pageId="60" pageNumber="61" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,624,723,770]" pageId="60" pageNumber="61" type="mainText">MADAGASCAR: Toamasina, 5.3 km SSE Ambanizana, 21.XI.2003 (B.L. Fisher).</paragraph>
|
||||
<paragraph box="[630,733,723,742]" pageId="60" pageNumber="61" type="mainText">Andranobe,</paragraph>
|
||||
<paragraph box="[739,1121,723,742]" pageId="60" pageNumber="61" type="mainText">15.66667 S, 49.96667 E, 425 m, rainforest,</paragraph>
|
||||
<subSubSection id="0F26FB870D4A2D8B905BB3AFA3F0AF46" type="materials_examined">
|
||||
<paragraph id="11497837D309AB2D239D4590C1C2E3E8" box="[118,295,696,714]" pageId="60" pageNumber="61" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="45E4767B041DE6BCBFCBEBA12182AE59" box="[118,624,723,770]" pageId="60" pageNumber="61" type="mainText">MADAGASCAR: Toamasina, 5.3 km SSE Ambanizana, 21.XI.2003 (B.L. Fisher).</paragraph>
|
||||
<paragraph id="9F890AB9E0992812373B618EEBE9694B" box="[630,733,723,742]" pageId="60" pageNumber="61" type="mainText">Andranobe,</paragraph>
|
||||
<paragraph id="9DFEAFC6FA5487FC9693D0ADE69610D0" box="[739,1121,723,742]" pageId="60" pageNumber="61" type="mainText">15.66667 S, 49.96667 E, 425 m, rainforest,</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,144 +1,143 @@
|
|||
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|
||||
<mods:namePart>B. L. Fisher</mods:namePart>
|
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<mods:namePart id="96DEA0C7283554DAAB5640C954C64041">B. L. Fisher</mods:namePart>
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<mods:title>Zootaxa</mods:title>
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<mods:title id="BED3394B8FD50147ACDEBD318B285DCA">Zootaxa</mods:title>
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<mods:part>
|
||||
<mods:date>2012</mods:date>
|
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<mods:detail type="volume">
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||||
<mods:number>3592</mods:number>
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<mods:part id="A272EA9AEA22F5307EF030BCA3C8CC69">
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<mods:date id="D7FA60E6FC461DBAAD895E20F792BBE9">2012</mods:date>
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<mods:detail id="60D57DF1BB3001B04CE7A88FE446DC05" type="volume">
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||||
<mods:number id="B19C1BF1C294ADBB4A125F319E6B8131">3592</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>85</mods:end>
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<mods:start id="C8128158CCB05827846CDEE0AB48DE6C">1</mods:start>
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<mods:end id="C9C91A2124DD944936899E5B9873A43C">85</mods:end>
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<mods:url>http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
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<mods:url id="CEE5201EAF85029B405203CD15D8569D">http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="HNS-Pub">26064</mods:identifier>
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<mods:identifier type="ISSN">175-5326</mods:identifier>
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<mods:identifier type="ZooBank">A2D9C9ED-C0BA-4B5F-A330-C9AB7D625704</mods:identifier>
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<mods:classification id="5AB1F4E367DE07807E270006018EA4BF">journal article</mods:classification>
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<mods:identifier id="66E9D2F84CBAECB645C3E1A08786C50C" type="HNS-Pub">26064</mods:identifier>
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<mods:identifier id="FB81E0D445DF17FA1FDDCBF28CE735A9" type="ISSN">175-5326</mods:identifier>
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||||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172643" ID-GBIF-Taxon="100105381" ID-Zenodo-Dep="6172643" LSID="urn:lsid:plazi:treatment:CB6142FA482F247AA968CDB6B73CD0D0" httpUri="http://treatment.plazi.org/id/CB6142FA482F247AA968CDB6B73CD0D0" lastPageNumber="62" pageNumber="61">
|
||||
<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,565,833,880]" pageId="60" pageNumber="61" type="mainText">
|
||||
<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">
|
||||
<treatment id="CB6142FA482F247AA968CDB6B73CD0D0" ID-DOI="http://doi.org/10.5281/zenodo.6172643" ID-GBIF-Taxon="100105381" ID-Zenodo-Dep="6172643" LSID="urn:lsid:plazi:treatment:CB6142FA482F247AA968CDB6B73CD0D0" httpUri="http://treatment.plazi.org/id/CB6142FA482F247AA968CDB6B73CD0D0" lastPageNumber="62" pageNumber="61" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="766CD3DBFBB76323272EECB19F99DE1E" type="nomenclature">
|
||||
<paragraph id="50C985D43868F834CFB52692BE100B8A" box="[118,565,833,880]" pageId="60" pageNumber="61" type="mainText">
|
||||
<taxonomicName id="5C682C0E94EB3122C0B471C5BF4970FB" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">
|
||||
Tetramorium noeli
|
||||
<authority>Hita</authority>
|
||||
<authority id="384D70F19F3902856DF9C54CAAB989BF">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="6839C5F146D4382D3484B9D542C00B43" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,565,833,880]" pageId="60" pageNumber="61" type="mainText">(Figs. 35, 40, 41, 117, 118, 119, 142)</paragraph>
|
||||
<subSubSection id="98AEEFF52791084C4C77C63F3116FE59" type="description">
|
||||
<paragraph id="D63BA1C1D8CE81E4373433B82D38E4E3" box="[118,565,833,880]" pageId="60" pageNumber="61" type="mainText">(Figs. 35, 40, 41, 117, 118, 119, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1122,917,1076]" pageId="60" pageNumber="61" type="mainText">
|
||||
<subSubSection id="BEF32621FF9FF3EC18190D2D13F7884E" type="materials_examined">
|
||||
<paragraph id="F526B0970CADA4A65173AA7B5A1826E5" box="[118,1122,917,1076]" pageId="60" pageNumber="61" type="mainText">
|
||||
Holotype worker, MADAGASCAR, Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="F959B6591DECD85A64C6F556050D5EC8" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Binara, 9.1 km 233° SW Daraina, 13.26333 S, 49.60333 E, 800 m, rainforest, sifted litter, (leaf mold, rotten wood), collection code BLF09656, 3.XII.2003 (B.L. Fisher) (CASC: CASENT0043554). Paratypes, nine workers with same data as holotype (BMNH: CASENT0043563; CASC: CASENT0043546; CASENT0043547; CASENT0043549; CASENT0043550; CASENT0043559; CASENT0043571; CASENT0043577; CASENT0043688); and one worker with same data as holotype but collected ex rotten log the 20.XI.2004 and collection code BLF10901 (MCZ: CASENT0053894).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,1113,1132]" pageId="60" pageNumber="61" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,1142,1273]" pageId="60" pageNumber="61" type="mainText">
|
||||
<subSubSection id="09CE338FEB7507A5AF194644D0920A8B" type="diagnosis">
|
||||
<paragraph id="7A367835560C4E841BCFDA5E1F8801D2" box="[118,208,1113,1132]" pageId="60" pageNumber="61" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="381424BFED9B0D93965CCD7B097E6C75" box="[118,1121,1142,1273]" pageId="60" pageNumber="61" type="mainText">
|
||||
The following character set distinguishes
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="1845EDD0208C7D04E82D7420C943BD04" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
from the remainder of the species group: head distinctly longer than wide (CI 92-95); propodeal spines very long to extremely long (PSLI 38-48); petiolar node rectangular nodiform with the posterodorsal margin situated higher than the anterodorsal margin, and the dorsum convex; postpetiole in lateral view approximately 1.4 to 1.5 times higher than long (LPpI 68-72), and in dorsal view around 1.2 to 1.3 times wider than long (DPpI 119-129).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,1310,1329]" pageId="60" pageNumber="61" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,1339,1470]" pageId="60" pageNumber="61" type="mainText">HL 0.85-1.00 (0.95); HW 0.79-0.94 (0.89); SL 0.64-0.77 (0.72); EL 0.18-0.22 (0.21); PH 0.45-0.55 (0.50); PW 0.65-0.79 (0.71); WL 1.11-1.44 (1.24); PSL 0.33-0.46 (0.41); PTL 0.32-0.39 (0.36); PTH 0.38-0.47 (0.43); PTW 0.26-0.32 (0.30); PPL 0.31-0.35 (0.33); PPH 0.43-0.50 (0.46); PPW 0.37-0.44 (0.41); CI 92-95 (94); SI 79-85 (81); OI 22-24 (23); DMI 55-59 (57); LMI 37-43 (57); PSLI 38-48 (43); PeNI 39-46 (42); LPeI 80-88 (84); DPeI 78-86 (82); PpNI 53-62 (57); LPpI 68-72 (71); DPpI 119-129 (124); PPI 132-145 (137) (15 measured).</paragraph>
|
||||
<paragraph box="[118,1123,1479,1580][118,1124,118,683]" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="mainText">
|
||||
<subSubSection id="76CD86E367872184C191080AFE27BC37" type="description">
|
||||
<paragraph id="4E5F51EC0E43EAE10F1E28BF38396E8D" box="[118,226,1310,1329]" pageId="60" pageNumber="61" type="mainText">Description</paragraph>
|
||||
<paragraph id="66B1A6AC9AF14B9CCF2164DFF9AA2CD1" box="[118,1121,1339,1470]" pageId="60" pageNumber="61" type="mainText">HL 0.85-1.00 (0.95); HW 0.79-0.94 (0.89); SL 0.64-0.77 (0.72); EL 0.18-0.22 (0.21); PH 0.45-0.55 (0.50); PW 0.65-0.79 (0.71); WL 1.11-1.44 (1.24); PSL 0.33-0.46 (0.41); PTL 0.32-0.39 (0.36); PTH 0.38-0.47 (0.43); PTW 0.26-0.32 (0.30); PPL 0.31-0.35 (0.33); PPH 0.43-0.50 (0.46); PPW 0.37-0.44 (0.41); CI 92-95 (94); SI 79-85 (81); OI 22-24 (23); DMI 55-59 (57); LMI 37-43 (57); PSLI 38-48 (43); PeNI 39-46 (42); LPeI 80-88 (84); DPeI 78-86 (82); PpNI 53-62 (57); LPpI 68-72 (71); DPpI 119-129 (124); PPI 132-145 (137) (15 measured).</paragraph>
|
||||
<paragraph id="9B7DA185E2282C6438CDDAB76CB10CA5" box="[118,1123,1479,1580][118,1124,118,683]" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="mainText">
|
||||
Head distinctly longer than wide (CI 92-95); posterior head margin moderately concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, moderately deep, but narrow and without defined posterior and ventral margins. Antennal scapes short to moderate, not reaching posterior head margin (SI 79-85). Eyes of
|
||||
<pageBreakToken pageId="61" pageNumber="62" start="start">moderate</pageBreakToken>
|
||||
<pageBreakToken id="58BFF5CCF252C0E2A24DAB6DCD7F5774" pageId="61" pageNumber="62" start="start">moderate</pageBreakToken>
|
||||
size (OI 22-24). Mesosomal outline in profile weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-43). Propodeal spines very long, spinose and acute (PSLI 38-48); propodeal lobes well-developed, triangular to elongate-triangular, and acute. Petiolar node in profile rectangular nodiform but comparatively high, approximately 1.1 to 1.3 times higher than long (LPeI 80-88), anterior and posterior faces approximately parallel, posterodorsal margin situated higher than anterodorsal, dorsum convex; node in dorsal view approximately 1.2 to 1.3 times longer than wide (DPeI 78-86). Postpetiole in profile comparatively high and anteroposteriorly compressed, approximately 1.4 to 1.5 times higher than long (LPpI 68-72); in dorsal view around 1.2 to 1.3 times wider than long (DPpI 119-129). Postpetiole in profile noticeably higher and less voluminous than petiolar node, in dorsal view approximately 1.3 to 1.5 times wider than petiolar node (PPI 132-145). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose, with three to eight rugae, median ruga always present and distinct, remaining rugae variably developed; cephalic dorsum between frontal carinae with 6 to 12 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted and none with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose to reticulate-rugose. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured, smooth, and shining, with weak, superficial ground sculpture. Waist segments strongly and very regularly longitudinally rugose, rugae absent from anterior face, but running unbroken from one side to the other through posterior face. Base of first gastral tergite distinctly costulate, remainder of gaster unsculptured, smooth and shining. Ground sculpture generally faint to absent everywhere on body. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs; first gastral tergite without appressed pubescence. Anterior edges of antennal scapes suberect to erect hairs. Body a uniform very dark brown to black colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,719,737]" pageId="61" pageNumber="62" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1121,746,847]" pageId="61" pageNumber="62" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">Tetramorium noeli</taxonomicName>
|
||||
<subSubSection id="90A621ACB088754FDFA6C9DFFD10B2ED" type="discussion">
|
||||
<paragraph id="AF5FBD958FDFB3F505BFB2D549E9C31F" box="[118,170,719,737]" pageId="61" pageNumber="62" type="mainText">Notes</paragraph>
|
||||
<paragraph id="E295C999A2A69EF2D52BBD376B9B6D57" box="[118,1121,746,847]" pageId="61" pageNumber="62" type="mainText">
|
||||
<taxonomicName id="0098E67BF90291B202C1590BD4612E06" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">Tetramorium noeli</taxonomicName>
|
||||
appears to be moderately common in the tropical dry forests and rainforests of the northeastern tip of Madagascar. Currently its distribution extends from Ambanitaza north to Montagne des
|
||||
<normalizedToken originalValue="Français">Francais</normalizedToken>
|
||||
<normalizedToken id="1F1CD435BAA07F57218AE0EED717B138" originalValue="Français">Francais</normalizedToken>
|
||||
, but it might well occur in other forest localities in northeastern Madagascar not yet explored. The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="620642B77201D0D984974DA4BAFF668E" rank="species">new species</taxonomicNameLabel>
|
||||
appears to be a leaf litter inhabitant on the basis of available collection data.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,855,1147]" pageId="61" pageNumber="62" type="mainText">
|
||||
<paragraph id="DC3C858908506D09B424B8C0F5CD382B" box="[118,1122,855,1147]" pageId="61" pageNumber="62" type="mainText">
|
||||
Within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="DFD930699A6E6E4ACC41CE21C484A1D4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
species complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="B0C5332F6AF49899CADEB1B0B62D87B1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
cannot be confused with either
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
<taxonomicName id="577056FDE358441A60D8F9E534FCC442" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
or
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
<taxonomicName id="6351791B8B324F7530F1F91D97AF18AC" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
since these have a petiolar node shape with the anterodorsal and posterodorsal margins at about the same height, whereas the node of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="50CC1DAFAAE8906F0C2332AA4218ACBD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
has a posterodorsal margin which is situated much higher than the anterodorsal. The fourth species of the complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana,</taxonomicName>
|
||||
<taxonomicName id="E0C4907126A04028A63025CDF6F8659F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana,</taxonomicName>
|
||||
has a node shape similar to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli,</taxonomicName>
|
||||
<taxonomicName id="A36FD8FA7D43EBB8E2630DA605D5A0BC" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli,</taxonomicName>
|
||||
and both species are morphologically very similar. Indeed, as noted in the description of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana,</taxonomicName>
|
||||
<taxonomicName id="760B74F10A93F24A5CA136E8A3FCDD58" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana,</taxonomicName>
|
||||
it is possible that both are conspecific. However, we treat them as distinct species here, and both differ in the length of the propodeal spines and the shape of the head and postpetiole. In
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="6518F2DE78828AABB33960568F3CE8B1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
the head is distinctly longer than wide (CI 92-95), whereas the head of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
<taxonomicName id="D989E342488C703543D8DA3E16A68C01" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
is only weakly longer than wide (CI 97-98). Additionally, despite both species having extremely long propodeal spines, this character is better developed in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
<taxonomicName id="FBA45FDB86723804ED9D2DC6DC4BE823" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
(PSLI 60-65) than in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="BA6306DFD05F59D5EFDEB1FA28E0B090" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
(PSLI 38-48). Also, the postpetiole of the latter species is relatively higher (LPpI 68-75) and broader (DPpI 119-129) than in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
<taxonomicName id="04D1C0FCAE493E60DA6B73CB17BF5445" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
(LPpI 77-81; DPpI 104-112).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,1183,1201]" pageId="61" pageNumber="62" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,795,1210,1229]" pageId="61" pageNumber="62" type="mainText">
|
||||
<subSubSection id="05F43CA8A2BC8F9A646EC7B1E021E74F" type="etymology">
|
||||
<paragraph id="02EBBE44989019631128DD8A5F272CD1" box="[118,219,1183,1201]" pageId="61" pageNumber="62" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="C001274B645C8F95FABF2942E053121F" box="[118,795,1210,1229]" pageId="61" pageNumber="62" type="mainText">
|
||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="D76F3C052D47A1820D6A53AEDC9D4636" rank="species">new species</taxonomicNameLabel>
|
||||
is dedicated to Noel Hita Garcia from Remscheid, Germany.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,1264,1283]" pageId="61" pageNumber="62" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1128,1292,1584]" pageId="61" pageNumber="62" type="mainText">
|
||||
<subSubSection id="E1FDBBF917B7F92B7F8E837A4B5967DE" type="materials_examined">
|
||||
<paragraph id="46CC4E7D40EBCA7DB020630C724D736B" box="[118,295,1264,1283]" pageId="61" pageNumber="62" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="3EA13501DBE20C65F5F41CED45B9EAE3" box="[118,1128,1292,1584]" pageId="61" pageNumber="62" type="mainText">
|
||||
MADAGASCAR: Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="19C2CF886182D9FDE5E322E34A759348" originalValue="Forêt">Foret</normalizedToken>
|
||||
Ambanitaza, 26.1 km 347° Antalaha, 14.67933 S, 50.18367 E, 240 m, rainforest, 26.XI.2004 (B.L. Fisher); Antsiranana, Parc National Montagne d'Ambre, 3.6 km 235° SW Joffreville, 12.53444 S, 49.1795 E, 925 m, montane rainforest, 20.-26.I.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="FDB3E6D33750B51EDD8237ED998985BB" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="EB311A0E05BC05CF3E222AD430A414A0" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
de l'Ankarana, 13.6 km 192° SSW Anivorano Nord, 12.86361 S, 49.22583 E, 210 m, tropical dry forest, 16.-21.II.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="8250BB8058A3B1C27DF335244498893E" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Binara, 9.1 km 233° SW Daraina, 13.26333 S, 49.60333 E, 800 m, rainforest, 3.XII.2003 (B.L. Fisher); Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="DA0B0F9B56567D4F761835DC720D80D4" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Binara, 9.1 km 233° SW Daraina, 13.26333 S, 49.60333 E, 800 m, rainforest, 20.XI.2004 (B.L. Fisher); Antsiranana, Montagne des
|
||||
<normalizedToken originalValue="Français">Francais</normalizedToken>
|
||||
<normalizedToken id="C25F502E809550FA38AE3864CF7B97E6" originalValue="Français">Francais</normalizedToken>
|
||||
, 7.2 km 142° SE Antsiranana, 12.32278 S, 49.33817 E, 180 m, tropical dry forest, 22.-28.II.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana, Makirovana forest, 14.1707 S, 49.9541 E, 415 m, rainforest, 28.-29.IV.2011(B.L. Fisher et al.); Antsiranana, Makirovana forest, 14.1651 S, 49.9477 E, 900 m, montane rainforest, 30.IV.-1.V.2011 (B.L. Fisher et al.); Antsiranana, Makirovana forest, 14.16044 S, 49.95216 E, 550 m, rainforest, 1.-2.V.2011 (B.L. Fisher et al.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,119 +1,118 @@
|
|||
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|
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|
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<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="aherni">
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|
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|
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|
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<taxonomicName id="2CF8C25546B2C2C3E8EAD2804377B61B" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="aherni">
|
||||
Tetramorium aherni
|
||||
<authority>Hita</authority>
|
||||
<authority id="8A059F4F82FFDD91756D62AFE4FBEA29">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="EE2227E7EED2BDFEAF6E1DD5DF858C5B" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,440,1298,1317]" pageId="55" pageNumber="56" type="mainText">(Figs. 34, 36, 37, 111, 112, 113, 142)</paragraph>
|
||||
<subSubSection id="840228B70D3D6736A2E9977E6A999B94" type="description">
|
||||
<paragraph id="F911624A7A20D6694BF356866D671782" box="[118,440,1298,1317]" pageId="55" pageNumber="56" type="mainText">(Figs. 34, 36, 37, 111, 112, 113, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1125,1349,1536]" pageId="55" pageNumber="56" type="mainText">Holotype worker, MADAGASCAR, Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, sifted litter (leaf mold, rotten wood), collection code BLF08872, 15.-18.XI.2003 (B.L. Fisher et al.) (CASC: CASENT0045755). Paratypes, 15 workers with same data as holotype (BMNH: CASENT0045607; CASC: CASENT0045595; CASENT0045603; CASENT0045675; CASENT0045679; CASENT0045687; CASENT0045696; CASENT0045750; CASENT0045766; CASENT0045771; CASENT0045774; CASENT0045779; MCZ: CASENT0045599; MHNG: CASENT0045761; NHMB: CASENT0045746).</paragraph>
|
||||
<subSubSection id="F6C755D5BE5A6D9F56D618E853B1F982" type="materials_examined">
|
||||
<paragraph id="BAB51E6E9B0848C34A01783DD6ABE32B" box="[118,1125,1349,1536]" pageId="55" pageNumber="56" type="mainText">Holotype worker, MADAGASCAR, Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, sifted litter (leaf mold, rotten wood), collection code BLF08872, 15.-18.XI.2003 (B.L. Fisher et al.) (CASC: CASENT0045755). Paratypes, 15 workers with same data as holotype (BMNH: CASENT0045607; CASC: CASENT0045595; CASENT0045603; CASENT0045675; CASENT0045679; CASENT0045687; CASENT0045696; CASENT0045750; CASENT0045766; CASENT0045771; CASENT0045774; CASENT0045779; MCZ: CASENT0045599; MHNG: CASENT0045761; NHMB: CASENT0045746).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,118,137]" pageId="57" pageNumber="58" type="mainText">
|
||||
<pageBreakToken pageId="57" pageNumber="58" start="start">Diagnosis</pageBreakToken>
|
||||
<subSubSection id="F031138FB31613AAD068FD42E60314A3" type="diagnosis">
|
||||
<paragraph id="ADA8750E848F6A70A1A0CA8BF7B955AF" box="[118,208,118,137]" pageId="57" pageNumber="58" type="mainText">
|
||||
<pageBreakToken id="2BACBDE9FC1D6995AC6A27E99E25D4D3" pageId="57" pageNumber="58" start="start">Diagnosis</pageBreakToken>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,147,222]" pageId="57" pageNumber="58" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="aherni">Tetramorium aherni</taxonomicName>
|
||||
<paragraph id="1266343B91B9F08EECCBD2291887EB03" box="[118,1121,147,222]" pageId="57" pageNumber="58" type="mainText">
|
||||
<taxonomicName id="3B98F703B1E43361C0371403DFA44FBF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="aherni">Tetramorium aherni</taxonomicName>
|
||||
is easily distinguishable within the species group due to the following character combination: eyes moderate to large (OI 22- 24); propodeal spines very long (PSLI 41-45); petiolar node with anterodorsal and posterodorsal margins at about the same height, dorsum flat to weakly convex.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,254,272]" pageId="57" pageNumber="58" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,282,413]" pageId="57" pageNumber="58" type="mainText">HL 0.91-0.98 (0.93); HW 0.82-0.89 (0.84); SL 0.67-0.74 (0.71); EL 0.19-0.21 (0.20); PH 0.44-0.52 (0.46); PW 0.62-0.69 (0.64); WL 1.11-1.21 (1.15); PSL 0.37-0.42 (0.40); PTL 0.30-0.35 (0.32); PTH 0.34-0.38 (0.36); PTW 0.27-0.31 (0.28); PPL 0.31-0.34 (0.32); PPH 0.37-0.40 (0.38); PPW 0.34-0.39 (0.36); CI 90-92 (90); SI 81-86 (84); OI 22-24 (23); DMI 54-57 (56); LMI 39-43 (40); PSLI 41-45 (43); PeNI 42-46 (44); LPeI 84-93 (89); DPeI 86-94 (89); PpNI 53-60 (56); LPpI 82-87 (84); DPpI 106-117 (113); PPI 121-133 (128) (twelve measured).</paragraph>
|
||||
<paragraph box="[118,1122,423,1089]" pageId="57" pageNumber="58" type="mainText">Head distinctly longer than wide (CI 90-92); posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, moderately deep and broad but without defined ventral margins. Antennal scapes of moderate length, not reaching posterior head margin (SI 81-86). Eyes small to moderate (OI 22-24). Mesosomal outline in profile weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 39-43). Propodeal spines very long, spinose, and acute (PSLI 41-45); propodeal lobes well-developed, triangular to elongate-triangular, and acute. Petiolar node in profile rectangular nodiform, approximately 1.1 to 1.2 times higher than long (LPeI 84-93), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins situated at about same height, dorsum flat to weakly convex; node in dorsal view approximately 1.1 times longer than wide (DPeI 86-94). Postpetiole in profile subglobular, approximately 1.1 to 1.2 times higher than long (LPpI 82-87); in dorsal view around 1.1 to 1.2 times wider than long (DPpI 106-117). Postpetiole in profile higher than petiolar node and generally appearing slightly more voluminous, in dorsal view approximately 1.2 to 1.3 times wider than petiolar node (PPI 121-133). Mandibles longitudinally rugose/rugulose, sometimes weakly so, sometimes partly sculptured and partly unsculptured; clypeus longitudinally rugose, with three to eight rugae, median ruga always present, remaining rugae variably developed; cephalic dorsum between frontal carinae with 8 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted and none with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose with very few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured, smooth, and shining with very weak, superficial ground sculpture (punctate or rugulose). Waist segments strongly longitudinally rugose, more irregular dorsally than laterally. Base of first gastral tergite distinctly costulate, remainder of gaster unsculptured, smooth and shining. Ground sculpture generally faint to absent everywhere on body. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with suberect to erect hairs. Body of uniform dark brown to black colour.</paragraph>
|
||||
<subSubSection id="B0B08DC99725A287B8F83A71D3B4229F" type="description">
|
||||
<paragraph id="055942E503B1E46119751878119C10C0" box="[118,226,254,272]" pageId="57" pageNumber="58" type="mainText">Description</paragraph>
|
||||
<paragraph id="9F42FEBC10B0A0813111A99A89DABA3B" box="[118,1121,282,413]" pageId="57" pageNumber="58" type="mainText">HL 0.91-0.98 (0.93); HW 0.82-0.89 (0.84); SL 0.67-0.74 (0.71); EL 0.19-0.21 (0.20); PH 0.44-0.52 (0.46); PW 0.62-0.69 (0.64); WL 1.11-1.21 (1.15); PSL 0.37-0.42 (0.40); PTL 0.30-0.35 (0.32); PTH 0.34-0.38 (0.36); PTW 0.27-0.31 (0.28); PPL 0.31-0.34 (0.32); PPH 0.37-0.40 (0.38); PPW 0.34-0.39 (0.36); CI 90-92 (90); SI 81-86 (84); OI 22-24 (23); DMI 54-57 (56); LMI 39-43 (40); PSLI 41-45 (43); PeNI 42-46 (44); LPeI 84-93 (89); DPeI 86-94 (89); PpNI 53-60 (56); LPpI 82-87 (84); DPpI 106-117 (113); PPI 121-133 (128) (twelve measured).</paragraph>
|
||||
<paragraph id="44B10E13BD2476186BB1D927A7A83FE2" box="[118,1122,423,1089]" pageId="57" pageNumber="58" type="mainText">Head distinctly longer than wide (CI 90-92); posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, moderately deep and broad but without defined ventral margins. Antennal scapes of moderate length, not reaching posterior head margin (SI 81-86). Eyes small to moderate (OI 22-24). Mesosomal outline in profile weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 39-43). Propodeal spines very long, spinose, and acute (PSLI 41-45); propodeal lobes well-developed, triangular to elongate-triangular, and acute. Petiolar node in profile rectangular nodiform, approximately 1.1 to 1.2 times higher than long (LPeI 84-93), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins situated at about same height, dorsum flat to weakly convex; node in dorsal view approximately 1.1 times longer than wide (DPeI 86-94). Postpetiole in profile subglobular, approximately 1.1 to 1.2 times higher than long (LPpI 82-87); in dorsal view around 1.1 to 1.2 times wider than long (DPpI 106-117). Postpetiole in profile higher than petiolar node and generally appearing slightly more voluminous, in dorsal view approximately 1.2 to 1.3 times wider than petiolar node (PPI 121-133). Mandibles longitudinally rugose/rugulose, sometimes weakly so, sometimes partly sculptured and partly unsculptured; clypeus longitudinally rugose, with three to eight rugae, median ruga always present, remaining rugae variably developed; cephalic dorsum between frontal carinae with 8 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted and none with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose with very few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured, smooth, and shining with very weak, superficial ground sculpture (punctate or rugulose). Waist segments strongly longitudinally rugose, more irregular dorsally than laterally. Base of first gastral tergite distinctly costulate, remainder of gaster unsculptured, smooth and shining. Ground sculpture generally faint to absent everywhere on body. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with suberect to erect hairs. Body of uniform dark brown to black colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,1126,1144]" pageId="57" pageNumber="58" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1120,1154,1229]" pageId="57" pageNumber="58" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="aherni">Tetramorium aherni</taxonomicName>
|
||||
<subSubSection id="1E05E5661996A0F81F95B70018198936" type="discussion">
|
||||
<paragraph id="CEF9439783ED29306CA98DE0089B3980" box="[118,170,1126,1144]" pageId="57" pageNumber="58" type="mainText">Notes</paragraph>
|
||||
<paragraph id="F51DE6D6E55EC40B66E8DCFD94E8E111" box="[118,1120,1154,1229]" pageId="57" pageNumber="58" type="mainText">
|
||||
<taxonomicName id="D5FC429F02EF8D37DA4D0DD139247C01" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="aherni">Tetramorium aherni</taxonomicName>
|
||||
is known from Befingotra, Marojejy, and Makirovana in the north-eastern part of Madagascar, but also from Ambalagoavy, which is situated much further south in eastern Madagascar. The species is only found in lowland or montane rainforests ranging from 450 to 900 m elevation, and seems to prefer the leaf litter stratum.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,1238,1370]" pageId="57" pageNumber="58" type="mainText">
|
||||
<paragraph id="E0C4CE39C87B140CA53DB867C8488183" box="[118,1122,1238,1370]" pageId="57" pageNumber="58" type="mainText">
|
||||
Inside the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="DAD9845B4524BAE9A9C0B2ACC37D117A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
<taxonomicName id="A6D59CA41F9239CED196C74D04EA32D5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
can be easily distinguished from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="noeli">T. ambanizana,</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="noeli">T. noeli,</taxonomicName>
|
||||
<taxonomicName id="BE2322153F56AB6723D24B55E360CB63" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="noeli">T. ambanizana,</taxonomicName>
|
||||
<taxonomicName id="276EB4326CCD5F96968CEA260DF3A54B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="noeli">T. noeli,</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae.</taxonomicName>
|
||||
<taxonomicName id="C4B090C462053F4F67FBAE3D68C2D731" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae.</taxonomicName>
|
||||
The latter species has much smaller eyes (OI 16-17) and shorter propodeal spines (PSLI 27- 29) than
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
<taxonomicName id="A5D4ED018F7EF5B277B79B3110FD5560" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
(OI 22-24; PSLI 41-45). In addition,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
<taxonomicName id="032EF0604C5C711623CD52D9C11468D7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ambanizana">T. ambanizana</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="682D0B652BA719B503F232AD934CB0A6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
have a differently shaped petiolar node with the posterodorsal margin situated much higher than the anterodorsal margin, whereas
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
<taxonomicName id="B4251B639B06D19D598DD0CB14FC63E4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="aherni">T. aherni</taxonomicName>
|
||||
has the anterodorsal and posterodorsal margins at about the same height.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,1407,1425]" pageId="57" pageNumber="58" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,908,1435,1454]" pageId="57" pageNumber="58" type="mainText">
|
||||
<subSubSection id="56CE57AE503A0C673BC13A8D7048A80A" type="etymology">
|
||||
<paragraph id="18646D9EF192D7C62E6759151CAA90BC" box="[118,219,1407,1425]" pageId="57" pageNumber="58" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="AB0B2A740077017B426520E170F21513" box="[118,908,1435,1454]" pageId="57" pageNumber="58" type="mainText">
|
||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="A3AA5FDA0242796D3862CD30A4A65DF8" rank="species">new species</taxonomicNameLabel>
|
||||
is dedicated to Dan Ahern for his contribution to the Madagascar Project.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,1486,1504]" pageId="57" pageNumber="58" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1123,1514,1590]" pageId="57" pageNumber="58" type="mainText">
|
||||
<subSubSection id="87F8177366EA6A9668120D7CDF1BE258" type="materials_examined">
|
||||
<paragraph id="1CDFBE14B6FC99EA38EFB92C803DB7B6" box="[118,295,1486,1504]" pageId="57" pageNumber="58" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="938AC1D86F9E8FC8A177A753B3E75CAB" box="[118,1123,1514,1590]" pageId="57" pageNumber="58" type="mainText">
|
||||
MADAGASCAR: Antsiranana, 6.5 km SSW Befingotra,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="49C39B7C0CF318E45577DB3210FB2F0C" originalValue="Rés">Res</normalizedToken>
|
||||
. Anjanaharibe-Sud, 14.75 S, 49.5 E, 875 m, rainforest, 19.-20.X.1994 (B.L. Fisher); Antsiranana, Makirovana Forest, 14.1651 S, 49.9477 E, montane rainforest, 30.IV.-1.V.2011 (B.L. Fisher et al.); Antsiranana, Makirovana Forest, 14.16044 S, 49.95216 E, 550 m, rainforest,
|
||||
</paragraph>
|
||||
<paragraph box="[118,1123,118,278]" pageId="58" pageNumber="59" type="mainText">
|
||||
<pageBreakToken pageId="58" pageNumber="59" start="start">1</pageBreakToken>
|
||||
<paragraph id="51BE881DE2100D0EF1C5784B7966DD38" box="[118,1123,118,278]" pageId="58" pageNumber="59" type="mainText">
|
||||
<pageBreakToken id="3B0DE9AAE2FF95E389855A6A3200032D" pageId="58" pageNumber="59" start="start">1</pageBreakToken>
|
||||
.-2.V.2011 (B.L. Fisher et al.); Antsiranana, Makirovana Forest, 14.16666 S, 49.95 E, 715 m, rainforest, 1.-2.V.2011 (B.L. Fisher et al.); Antsiranana, Parc National de Marojejy, Manantenina River, 28.0 km 38° NE Andapa, 8.2 km 333° NNW Manantenina, 14.43667 S, 49.775 E, 450 m, rainforest, 12.-15.XI.2003 (B.L. Fisher et al.); Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, 15.-18.XI.2003 (B.L. Fisher et al.); Fianarantsoa, Foret d'Ambalagoavy Nord, Ikongo, Ambatombe, 21.8275 S, 47.33889 E, 625 m, 1.XII.2000 (R. Harin'Hala & M.E. Irwin).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,124 +1,123 @@
|
|||
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<mods:roleTerm id="A6B6270F9CF66C61897ECED853841DB2">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>B. L. Fisher</mods:namePart>
|
||||
<mods:namePart id="93192D8CEC16E71B8A9E5EA19CDAD1CC">B. L. Fisher</mods:namePart>
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<mods:title>Zootaxa</mods:title>
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<mods:title id="9BFF4F022684A7808C86CBC97313E3DF">Zootaxa</mods:title>
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<mods:part>
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||||
<mods:date>2012</mods:date>
|
||||
<mods:detail type="volume">
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||||
<mods:number>3592</mods:number>
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<mods:part id="36D20794D5D63CA969F18B4F888864FA">
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<mods:date id="28406BD3623A9B17D93B4280C9845EC3">2012</mods:date>
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||||
<mods:detail id="8FE1059A52DE067CC9D3E4048F1A9262" type="volume">
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||||
<mods:number id="DBBE610EA25FC33F9A1EC936BAE71479">3592</mods:number>
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||||
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||||
<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end id="292A2D808EE92B08A30C98687B64B85D">85</mods:end>
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<mods:url>http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="HNS-Pub">26064</mods:identifier>
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<mods:identifier type="ISSN">175-5326</mods:identifier>
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<mods:identifier type="ZooBank">A2D9C9ED-C0BA-4B5F-A330-C9AB7D625704</mods:identifier>
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<mods:identifier id="60CE11D27F4BBE601419B91B55E79F3E" type="ISSN">175-5326</mods:identifier>
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||||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172645" ID-GBIF-Taxon="100105389" ID-Zenodo-Dep="6172645" LSID="urn:lsid:plazi:treatment:D92D4323A796E729EA6D8F8551D63BA2" httpUri="http://treatment.plazi.org/id/D92D4323A796E729EA6D8F8551D63BA2" lastPageNumber="66" pageNumber="64">
|
||||
<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,626,118,138]" pageId="63" pageNumber="64" type="mainText">
|
||||
<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="singletonae">
|
||||
<pageBreakToken pageId="63" pageNumber="64" start="start">Tetramorium</pageBreakToken>
|
||||
<treatment id="D92D4323A796E729EA6D8F8551D63BA2" ID-DOI="http://doi.org/10.5281/zenodo.6172645" ID-GBIF-Taxon="100105389" ID-Zenodo-Dep="6172645" LSID="urn:lsid:plazi:treatment:D92D4323A796E729EA6D8F8551D63BA2" httpUri="http://treatment.plazi.org/id/D92D4323A796E729EA6D8F8551D63BA2" lastPageNumber="66" pageNumber="64" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="75463AAFCD3CB69A969A5B155DEE19D7" type="nomenclature">
|
||||
<paragraph id="494351BDA2DE402C63BD59F0C966CCAE" box="[118,626,118,138]" pageId="63" pageNumber="64" type="mainText">
|
||||
<taxonomicName id="64463A618A569F3C01A70629D7EB2313" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="singletonae">
|
||||
<pageBreakToken id="1AFA95E2D6A387B1775D7EB12C600E68" pageId="63" pageNumber="64" start="start">Tetramorium</pageBreakToken>
|
||||
singletonae
|
||||
<authority>Hita</authority>
|
||||
<authority id="4C5CFD26EB3B549DF06C5C449BBD163D">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="00F498FECA7DED38462A17E219D47832" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,476,146,165]" pageId="63" pageNumber="64" type="mainText">(Figs. 30, 32, 38, 39, 120, 121, 122, 142)</paragraph>
|
||||
<subSubSection id="9795DF2BB653DE6C90BA9EEA7023CBD4" type="description">
|
||||
<paragraph id="900B7DF4556BFD356EE237CE27ACBDDC" box="[118,476,146,165]" pageId="63" pageNumber="64" type="mainText">(Figs. 30, 32, 38, 39, 120, 121, 122, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1122,202,305]" pageId="63" pageNumber="64" type="mainText">Holotype worker, MADAGASCAR, Toamasina, 5.3 km SSE Ambanizana, Andranobe, 15.66667 S, 49.96667 E, 425 m, rainforest, pitfall trap, collection code BLF00906, 19.XI.1993 (B.L. Fisher) (CASC: CASENT0247161). Paratypes, five workers with same data as holotype (CASC: CASENT0188733; CASENT0247159; CASENT0270778).</paragraph>
|
||||
<subSubSection id="14ADCFDABA2CBD1E7ECCDAC077918A7B" type="materials_examined">
|
||||
<paragraph id="418296577BCB286882F6C890DF101C02" box="[118,1122,202,305]" pageId="63" pageNumber="64" type="mainText">Holotype worker, MADAGASCAR, Toamasina, 5.3 km SSE Ambanizana, Andranobe, 15.66667 S, 49.96667 E, 425 m, rainforest, pitfall trap, collection code BLF00906, 19.XI.1993 (B.L. Fisher) (CASC: CASENT0247161). Paratypes, five workers with same data as holotype (CASC: CASENT0188733; CASENT0247159; CASENT0270778).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,342,361]" pageId="63" pageNumber="64" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1122,371,446]" pageId="63" pageNumber="64" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="singletonae">Tetramorium singletonae</taxonomicName>
|
||||
<subSubSection id="DCA621622E06F084C0B425599D5F8921" type="diagnosis">
|
||||
<paragraph id="EACB4312E46AFFC2353404478E229059" box="[118,208,342,361]" pageId="63" pageNumber="64" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="294B24843C917B0C10FF623F297FA3DD" box="[118,1122,371,446]" pageId="63" pageNumber="64" type="mainText">
|
||||
<taxonomicName id="8669395519EBB6C11269C1BC07D44F38" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="singletonae">Tetramorium singletonae</taxonomicName>
|
||||
is easily recognisable within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="609C70A168112D3D8B0B726BCE651433" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
complex by the following combination of characters: eyes very small (OI 16-17); propodeal spines long, but very short for the species group (PSLI 27-29); petiolar node with anterodorsal and posterodorsal margins at about the same height, dorsum flat to weakly convex.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,483,501]" pageId="63" pageNumber="64" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,511,643]" pageId="63" pageNumber="64" type="mainText">HL 0.90-0.92 (0.91); HW 0.87-0.90 (0.89); SL 0.62-0.66 (0.64); EL 0.14-0.16 (0.15); PH 0.42-0.49 (0.47); PW 0.67-0.69 (0.68); WL 1.10-1.16 (1.13); PSL 0.25-0.27 (0.25); PTL 0.33-0.35 (0.34); PTH 0.33-0.36 (0.33); PTW 0.27-0.29 (0.28); PPL 0.30-0.31 (0.30); PPH 0.34-0.35 (0.34); PPW 0.35-0.37 (0.36); CI 97-98 (98); SI 70-74 (72); OI 16-17 (17); DMI 59-62 (60); LMI 38-44 (41); PSLI 27-29 (28); PeNI 40-42 (41); LPeI 94-106 (101); DPeI 81-85 (83); PpNI 51-54 (53); LPpI 86-91 (89); DPpI 115-122 (118); PPI 126-130 (128) (eight measured).</paragraph>
|
||||
<paragraph box="[118,1122,652,1261]" pageId="63" pageNumber="64" type="mainText">Head weakly longer than wider (CI 97-98). Anterior clypeal margin with median impression, sometimes weak, but always distinct. Frontal carinae well-developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, but shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short, not reaching posterior head margin (SI 70-74). Eyes very small (OI 16-17). Mesosomal outline in profile flat, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-44). Propodeal spines with very broad base, up-curved, elongate-triangular, and moderately long (PSLI 27-29); propodeal lobes well-developed, triangular and acute. Petiolar node in profile rectangular nodiform, ranging from weakly longer than high to weakly higher than long (LPeI 94-106), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum flat to weakly convex; node in dorsal view approximately 1.2 times longer than wide (DPeI 81-85). Postpetiole in profile globular, approximately 1.1 times higher than long (LPpI 86-91); in dorsal view around 1.2 times wider than long (DPpI 115-122). Postpetiole in profile appearing approximately as voluminous as petiolar node, in dorsal view approximately 1.2 to 1.3 times wider than petiolar node (PPI 126-130). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose, with three to five rugae; cephalic dorsum between frontal carinae with 8 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, few rugae interrupted or with cross-meshes; lateral and ventral head longitudinally rugose, rarely with cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured. Waist segments strongly irregularly longitudinally rugose. Base of first gastral tergite with costulate sculpture, remainder of gaster unsculptured, smooth, and shiny. Ground sculpture generally faint to absent everywhere on body. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. First gastral tergite without distinct, appressed pubescence. Anterior edges of antennal scapes with subdecumbent to erect, standing hairs. Body a uniform brown colour.</paragraph>
|
||||
<subSubSection id="1D49CCA814486AF78AF2B5D84CCAA4E1" type="description">
|
||||
<paragraph id="5AE27777BB9A04AB8A1358A14A798D9F" box="[118,226,483,501]" pageId="63" pageNumber="64" type="mainText">Description</paragraph>
|
||||
<paragraph id="AC594664EEE773B4AF9ABCDD02F948EC" box="[118,1121,511,643]" pageId="63" pageNumber="64" type="mainText">HL 0.90-0.92 (0.91); HW 0.87-0.90 (0.89); SL 0.62-0.66 (0.64); EL 0.14-0.16 (0.15); PH 0.42-0.49 (0.47); PW 0.67-0.69 (0.68); WL 1.10-1.16 (1.13); PSL 0.25-0.27 (0.25); PTL 0.33-0.35 (0.34); PTH 0.33-0.36 (0.33); PTW 0.27-0.29 (0.28); PPL 0.30-0.31 (0.30); PPH 0.34-0.35 (0.34); PPW 0.35-0.37 (0.36); CI 97-98 (98); SI 70-74 (72); OI 16-17 (17); DMI 59-62 (60); LMI 38-44 (41); PSLI 27-29 (28); PeNI 40-42 (41); LPeI 94-106 (101); DPeI 81-85 (83); PpNI 51-54 (53); LPpI 86-91 (89); DPpI 115-122 (118); PPI 126-130 (128) (eight measured).</paragraph>
|
||||
<paragraph id="0835145680430CF5E35EEED1590383D4" box="[118,1122,652,1261]" pageId="63" pageNumber="64" type="mainText">Head weakly longer than wider (CI 97-98). Anterior clypeal margin with median impression, sometimes weak, but always distinct. Frontal carinae well-developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, but shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short, not reaching posterior head margin (SI 70-74). Eyes very small (OI 16-17). Mesosomal outline in profile flat, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-44). Propodeal spines with very broad base, up-curved, elongate-triangular, and moderately long (PSLI 27-29); propodeal lobes well-developed, triangular and acute. Petiolar node in profile rectangular nodiform, ranging from weakly longer than high to weakly higher than long (LPeI 94-106), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum flat to weakly convex; node in dorsal view approximately 1.2 times longer than wide (DPeI 81-85). Postpetiole in profile globular, approximately 1.1 times higher than long (LPpI 86-91); in dorsal view around 1.2 times wider than long (DPpI 115-122). Postpetiole in profile appearing approximately as voluminous as petiolar node, in dorsal view approximately 1.2 to 1.3 times wider than petiolar node (PPI 126-130). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose, with three to five rugae; cephalic dorsum between frontal carinae with 8 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, few rugae interrupted or with cross-meshes; lateral and ventral head longitudinally rugose, rarely with cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured. Waist segments strongly irregularly longitudinally rugose. Base of first gastral tergite with costulate sculpture, remainder of gaster unsculptured, smooth, and shiny. Ground sculpture generally faint to absent everywhere on body. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. First gastral tergite without distinct, appressed pubescence. Anterior edges of antennal scapes with subdecumbent to erect, standing hairs. Body a uniform brown colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,1298,1317]" pageId="63" pageNumber="64" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1122,1327,1402]" pageId="63" pageNumber="64" type="mainText">
|
||||
<subSubSection id="C08E7F5955F6E15EB506732DC7B8FC81" type="discussion">
|
||||
<paragraph id="EC860F2AC465F1B6E47E4F596B371D36" box="[118,170,1298,1317]" pageId="63" pageNumber="64" type="mainText">Notes</paragraph>
|
||||
<paragraph id="D93E4163123D1B541A42C73B5EDCC683" box="[118,1122,1327,1402]" pageId="63" pageNumber="64" type="mainText">
|
||||
The available material from this
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="76A6C435E7F2BFD4211EE881ADE4B778" rank="species">new species</taxonomicNameLabel>
|
||||
was sampled from Ambanizana and Amparihibe. Both localities are rainforests located in the northeast of Madagascar at elevations of 425 to approximately 1000 m.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="singletonae">Tetramorium singletonae</taxonomicName>
|
||||
<taxonomicName id="AF0F5BE6E519CE545BBC8753E3164005" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="singletonae">Tetramorium singletonae</taxonomicName>
|
||||
appears to live and/or forage on the ground since most specimens were collected from pitfall traps.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,1411,1543]" pageId="63" pageNumber="64" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="singletonae">Tetramorium singletonae</taxonomicName>
|
||||
<paragraph id="00B74D901BF05ADAD8E5DCF7E521AD1E" box="[118,1122,1411,1543]" pageId="63" pageNumber="64" type="mainText">
|
||||
<taxonomicName id="350A88057559DDFA191FD805862C8A82" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="singletonae">Tetramorium singletonae</taxonomicName>
|
||||
is the species with the shortest antennal scapes (SI 70-74 vs. SI 78-86), smallest eyes (OI 16-17 vs. OI 21-24), and shortest propodeal spines (PSLI 27-29 vs. PSLI 38-65) encountered in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
<taxonomicName id="9A611E01927E1C04DDDD305D457B7E78" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="noeli">T. noeli</taxonomicName>
|
||||
species complex, and is thus easily recognisable. At first glance
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
<taxonomicName id="BA147208A71E718141289CD69D13EB88" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae</taxonomicName>
|
||||
is also morphologically close to
|
||||
<taxonomicName genus="Tetamorium" pageId="63" pageNumber="64" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="8114ACBFD69E24AEAE218501376AB8F4" genus="Tetamorium" pageId="63" pageNumber="64" rank="species" species="nify">T. nify</taxonomicName>
|
||||
from the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="CB282CA7AB31350C9B5A5C6C975577DE" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
species complex since both have shorter propodeal spines than most other
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="D767458A98584DEDC3C84841401D7508" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group species and similarly shaped waist segments, but the base of the first gastral tergite is unsculptured in
|
||||
<taxonomicName genus="Tetamorium" pageId="63" pageNumber="64" rank="species" species="nify">T. nify.</taxonomicName>
|
||||
<taxonomicName id="087AF68E4074EF7E246D1C817805053A" genus="Tetamorium" pageId="63" pageNumber="64" rank="species" species="nify">T. nify.</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="multiple">
|
||||
<caption box="[118,1122,1539,1584]" pageId="64" pageNumber="65" type="caption">
|
||||
<paragraph box="[118,1122,1539,1584]" pageId="64" pageNumber="65" type="caption">
|
||||
<pageBreakToken pageId="64" pageNumber="65" start="start">FIGURES</pageBreakToken>
|
||||
<subSubSection id="5A9A5D99DA7C81835D23B91071464A2F" type="multiple">
|
||||
<caption id="EA293458D96C899D0BDD60400EA86E53" box="[118,1122,1539,1584]" pageId="64" pageNumber="65" type="caption">
|
||||
<paragraph id="E29E94C5DDB4C64DEB96E7D90537BD8E" box="[118,1122,1539,1584]" pageId="64" pageNumber="65" type="caption">
|
||||
<pageBreakToken id="89A97AA845FFE039E4F635C0E6FB4E25" pageId="64" pageNumber="65" start="start">FIGURES</pageBreakToken>
|
||||
120-122.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae,</taxonomicName>
|
||||
<taxonomicName id="0914697A4EE53F9687C5AEE17ADE1DBB" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="singletonae">T. singletonae,</taxonomicName>
|
||||
holotype (CASENT0247161). 120. Body in profile. 121. Body in dorsal view. 122. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,118,136]" pageId="65" pageNumber="66" type="mainText">
|
||||
<pageBreakToken pageId="65" pageNumber="66" start="start">Etymology</pageBreakToken>
|
||||
<subSubSection id="79DDE49A4914C4CF4E0153CB8E53F9FC" type="etymology">
|
||||
<paragraph id="D6EA42E80C40E7883ACED4675A6EB127" box="[118,219,118,136]" pageId="65" pageNumber="66" type="mainText">
|
||||
<pageBreakToken id="F05CE376641068DE24BCA6B317A32F86" pageId="65" pageNumber="66" start="start">Etymology</pageBreakToken>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1120,147,194]" pageId="65" pageNumber="66" type="mainText">
|
||||
<paragraph id="E82E90792F3302846FA16AC12EC13AC8" box="[118,1120,147,194]" pageId="65" pageNumber="66" type="mainText">
|
||||
The
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="71C1A47B3F72D2991F7C4CE09856A50A" rank="species">new species</taxonomicNameLabel>
|
||||
is dedicated to Sarah Singleton for her support to promote the discovery and identification of life on earth.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,231,249]" pageId="65" pageNumber="66" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,259,306]" pageId="65" pageNumber="66" type="mainText">MADAGASCAR: Toamasina, 5.3 km SSE Ambanizana, Andranobe, 15.66667 S, 49.96667 E, 425 m, rainforest, 19.XI.1993 (B.L. Fisher); Toamasina, Amparihibe, 15° 2' S, 49° 34' E, II.-III.2003 (K.A. Jackson & D. Carpenter).</paragraph>
|
||||
<subSubSection id="AC52AD291AEE35FEE8B41BB76FE9DC6B" type="materials_examined">
|
||||
<paragraph id="EF5EA1B4401F15B3F79DFC849977A7AB" box="[118,295,231,249]" pageId="65" pageNumber="66" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="C5A086E2E572A72D2B661DE921320088" box="[118,1122,259,306]" pageId="65" pageNumber="66" type="mainText">MADAGASCAR: Toamasina, 5.3 km SSE Ambanizana, Andranobe, 15.66667 S, 49.96667 E, 425 m, rainforest, 19.XI.1993 (B.L. Fisher); Toamasina, Amparihibe, 15° 2' S, 49° 34' E, II.-III.2003 (K.A. Jackson & D. Carpenter).</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,187 +1,186 @@
|
|||
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172635" ID-GBIF-Taxon="100105385" ID-Zenodo-Dep="6172635" LSID="urn:lsid:plazi:treatment:DB23C78D4E61D83D7F17A6F90D04A2FA" httpUri="http://treatment.plazi.org/id/DB23C78D4E61D83D7F17A6F90D04A2FA" lastPageNumber="56" pageNumber="53">
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<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,463,585,605]" pageId="52" pageNumber="53" type="mainText">
|
||||
<taxonomicName authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pleganon">
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<treatment id="DB23C78D4E61D83D7F17A6F90D04A2FA" ID-DOI="http://doi.org/10.5281/zenodo.6172635" ID-GBIF-Taxon="100105385" ID-Zenodo-Dep="6172635" LSID="urn:lsid:plazi:treatment:DB23C78D4E61D83D7F17A6F90D04A2FA" httpUri="http://treatment.plazi.org/id/DB23C78D4E61D83D7F17A6F90D04A2FA" lastPageNumber="56" pageNumber="53" scope_family="Formicidae" scope_order="Hymenoptera">
|
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<subSubSection id="E1256A5A9DDECBB5F7FA3E032144805F" type="nomenclature">
|
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|
||||
<taxonomicName id="A52D37CDF5C21A48BFB9D85A0F3B4A89" ID-CoL="55PRQ" authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pleganon">
|
||||
Tetramorium pleganon
|
||||
<authority>Bolton</authority>
|
||||
<authority id="973BF01E6DEB3AB3263035B16E9F38F7">Bolton</authority>
|
||||
,
|
||||
</taxonomicName>
|
||||
1979
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,377,612,631]" pageId="52" pageNumber="53" type="mainText">(Figs. 48, 108, 109, 110, 142)</paragraph>
|
||||
<subSubSection id="2FA6D2968DA31D0404C1CA29A0781ED5" type="description">
|
||||
<paragraph id="4D1A6E29C9188B05FEF9663D5FE1107C" box="[118,377,612,631]" pageId="52" pageNumber="53" type="mainText">(Figs. 48, 108, 109, 110, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1122,668,758]" pageId="52" pageNumber="53" type="mainText">
|
||||
<taxonomicName authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pleganon">
|
||||
<subSubSection id="F8A5F46013D9CCAB4B5A81FF6AF20001" type="materials_examined">
|
||||
<paragraph id="B9A3CC8FAE6AB5FA9F8177F8044E7BA3" box="[118,1122,668,758]" pageId="52" pageNumber="53" type="mainText">
|
||||
<taxonomicName id="0EBD4393694AE78E07D6718CF2905234" authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pleganon">
|
||||
Tetramorium pleganon
|
||||
<authority>Bolton</authority>
|
||||
<authority id="57378FEB38A120BCA2F169DE92FB3A86">Bolton</authority>
|
||||
,
|
||||
</taxonomicName>
|
||||
1979:146. Holotype worker, MADAGASCAR, Antsiranana, 84 km SW Sambava on road to Andapa, 70-160 m, degraded forest, strays on path, AB 43, 17.II.1977 (W.L. & D.E. Brown) (MCZ: CASENT0280587) [examined]. Paratypes, three workers with same data as holotype (BMNH: CASENT0102399; MCZ: CASENT0280588) [examined].
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,792,811]" pageId="52" pageNumber="53" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,822,928]" pageId="52" pageNumber="53" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pleganon">Tetramorium pleganon</taxonomicName>
|
||||
<subSubSection id="EC889E75DDD922F973040107C9B944EF" type="diagnosis">
|
||||
<paragraph id="1F0C28ECC71A7818BB0BD6B4F9E60731" box="[118,208,792,811]" pageId="52" pageNumber="53" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="19BC47698F0AA9C05292E52F4032064C" box="[118,1121,822,928]" pageId="52" pageNumber="53" type="mainText">
|
||||
<taxonomicName id="0F096E0F21F08CCFD59421717868B5F7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pleganon">Tetramorium pleganon</taxonomicName>
|
||||
is easily recognisable within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="D150162E5CC3CD39EACF1F550DFF6DF9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group in the Malagasy region due to the character combination of: propodeal spines long to very long (PSLI 37-44); petiolar node wider than long (DPeI 111-118); dorsum of petiolar node strongly rugose; first gastral tergite with superficial, fine, reticulate-punctate sculpture, ranging from basal third to more than half of tergite.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,967,986]" pageId="52" pageNumber="53" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1117,997,1161]" pageId="52" pageNumber="53" type="mainText">HL 0.87-1.05 (0.93); HW 0.85-1.02 (0.91); SL 0.61-0.73 (0.66); EL 0.18-0.23 (0.19); PH 0.43-0.56 (0.48); PW 0.59-0.79 (0.69); WL 1.07-1.31 (1.17); PSL 0.32-0.43 (0.37); PTL 0.22-0.31 (0.25); PTH 0.34-0.44 (0.37); PTW 0.26-0.34 (0.29); PPL 0.27-0.34 (0.29); PPH 0.31-0.43 (0.35); PPW 0.34-0.43 (0.38); CI 96-99 (97); SI 71-74 (73); OI 20-23 (21); DMI 52-61 (59); LMI 38-43 (41); PSLI 37-44 (39); PeNI 40-44 (42); LPeI 63-73 (67); DPeI 111-118 (115); PpNI 52-61 (55); LPpI 80-89 (83); DPpI 122-137 (129); PPI 126-138 (130) (14 measured).</paragraph>
|
||||
<paragraph box="[118,1121,1172,1570][118,1119,118,517]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="mainText">
|
||||
<subSubSection id="01133DC7357CED89F2BDFB7D4527544E" type="description">
|
||||
<paragraph id="DCA94263C7D22856E8AD5880B5D1776E" box="[118,226,967,986]" pageId="52" pageNumber="53" type="mainText">Description</paragraph>
|
||||
<paragraph id="26EF9B9475D9F3ED9EA8943F4773400C" box="[118,1117,997,1161]" pageId="52" pageNumber="53" type="mainText">HL 0.87-1.05 (0.93); HW 0.85-1.02 (0.91); SL 0.61-0.73 (0.66); EL 0.18-0.23 (0.19); PH 0.43-0.56 (0.48); PW 0.59-0.79 (0.69); WL 1.07-1.31 (1.17); PSL 0.32-0.43 (0.37); PTL 0.22-0.31 (0.25); PTH 0.34-0.44 (0.37); PTW 0.26-0.34 (0.29); PPL 0.27-0.34 (0.29); PPH 0.31-0.43 (0.35); PPW 0.34-0.43 (0.38); CI 96-99 (97); SI 71-74 (73); OI 20-23 (21); DMI 52-61 (59); LMI 38-43 (41); PSLI 37-44 (39); PeNI 40-44 (42); LPeI 63-73 (67); DPeI 111-118 (115); PpNI 52-61 (55); LPpI 80-89 (83); DPpI 122-137 (129); PPI 126-138 (130) (14 measured).</paragraph>
|
||||
<paragraph id="C7B833EE5738CE688E3409F224BDBAFF" box="[118,1121,1172,1570][118,1119,118,517]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="mainText">
|
||||
Head longer than wide (CI 96-99); posterior head margin moderately concave. Anterior clypeal margin medially impressed, often weakly so. Frontal carinae st rongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, shallow to moderately deep, and broad, without defined posterior and ventral margins. Antennal scapes short, not reaching posterior head margin (SI 71-74). Eyes small to moderate in size (OI 20-23). Mesosomal outline in profile weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-43). Propodeal spines very long, spinose and acute (PSLI 37-44); propodeal lobes short, triangular, and acute. Petiolar node in profile rectangular nodiform, approximately 1.1 to 1.2 times higher than long (LPeI 63-73), anterior and posterior faces approximately parallel, anterodorsal margin situated higher than posterodorsal, dorsum weakly tapering backwards posteriorly; node in dorsal view approximately 1.1 to 1.2 times wider than long (DPeI 111-118). Postpetiole in profile rounded, approximately 1.1 to 1.3 times higher than long (LPpI 80-89); in dorsa l view around 1.2 to 1.4 times wider than long (DPpI 122-137). Postpetiole in profile approximately as voluminous as petiolar node, in dorsal view 1.2 to 1.4 times wider than petiolar node (PPI 126-138). Mandibles generall y finely to strongly striate; clypeus longitudinally
|
||||
<pageBreakToken pageId="53" pageNumber="54" start="start">rugose</pageBreakToken>
|
||||
<pageBreakToken id="9FE4C6F75D39F1B0C7F485AA2E92B146" pageId="53" pageNumber="54" start="start">rugose</pageBreakToken>
|
||||
, with four to nine rugae, median ruga always present and distinct, remaining rugae variably developed, usually weaker; cephalic dorsum between frontal carinae with 11 to 14 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted and none with cross-meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose, very rarely with cross-meshes. Ground sculpture on head generally feeble. Mesosoma laterally and dorsally distinctly longitudinally rugose, lateral mesosoma sometimes weaker sculptured than dorsum. Forecoxae generally unsculptured, smooth, and shining, sometimes with superficial scul pture. Ground sculpture on mesosoma generally faint to absent. Waist segments strongly rugose dorsally, laterally rugose sculpture much weaker; both waist segments with very conspicuous reticulate-punctate ground sculpture. First gastral tergite with fine, dense, reticulate-punctate ground sculpture, ranging from basal third to mo re than half of tergite. All dorsal surfaces of body with abundant, long, and fine standing hairs; first gast ral tergite with mix of moderately long appressed to decumbent hairs and more abundant and longer suberect to erect hairs. Anterior edges of antennal scapes with decumbent to suberect hairs. Body a uniform very dark brown to black colour, appendages often lighter in colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,556,574]" pageId="53" pageNumber="54" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1121,585,779]" pageId="53" pageNumber="54" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">Tetramorium pleganon</taxonomicName>
|
||||
<subSubSection id="66146135E9D3A9396F01A073BE53F245" type="discussion">
|
||||
<paragraph id="ABDED0E90C78DA66FB0C1FC1CBC5149F" box="[118,170,556,574]" pageId="53" pageNumber="54" type="mainText">Notes</paragraph>
|
||||
<paragraph id="3FB76DAE11AE303478C7EE415CDC1F9F" box="[118,1121,585,779]" pageId="53" pageNumber="54" type="mainText">
|
||||
<taxonomicName id="B94876B7472BABE3308A4AA9AA462D41" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">Tetramorium pleganon</taxonomicName>
|
||||
possesses a comparatively wide distribution in Madagascar since it is found in many localities from Zombitse and Kalambatrita in the south up to Ambato and Antsahabe in the north. It appears to be comparatively flexible in its habitat preferences. Most of the localities where it was encountered consisted of rainforest, montane rainforest, or tropical dry forest, but the number of specimens collected in these habitats was comparatively low. In contrast, the largest number of examined specimens were sampled in Uapaca woodland, savannah grassland, and degraded forests, indicating that
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="44F6FA27D48F5BEC9EDDEC4FAF54D136" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
might be more successful in disturbed and open habitats than in dense forests.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,789,1362]" pageId="53" pageNumber="54" type="mainText">
|
||||
<paragraph id="5D41766274DD2F6E4E86D01B6860EC2D" box="[118,1122,789,1362]" pageId="53" pageNumber="54" type="mainText">
|
||||
In Bolton's (1979) revision
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="A2F3D719625ADBB67826B421AA1E6183" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
was easily diagnosable and recognisable from all other Malagasy
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
<taxonomicName id="99DFDD204ABFA148D960150986202E22" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
due to its 11-segmented antennae and the sculptured first gastral tergite. However, our ongoing revision has revealed several species with 11-segmented antennae and sculpture on the first gastral tergite, indicating this character combination is more common than previously thought, and is certainly not unique to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon.</taxonomicName>
|
||||
<taxonomicName id="F35711D202DE46A7DC4938D1A4FC6AD4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon.</taxonomicName>
|
||||
Nevertheless, within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="75BDD3A5E7AAF61311639D386D09CF00" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group there are different types of sculpture observable on the first gastral tergite, and the reticulate-punctate ground sculpture seen in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="EB52C4418272D80081E8583B015490BF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
places it well within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="CED99AA05697F7A784DB67209EBF265B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
complex. Within this complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="DA10080153756D607AAFFB97D0AD4F8B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
is unlikely to be misidentified with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="1107AD1AE725987053E71607E625E8F8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
since the latter species has a completely sculptured tergite with very strongly developed and conspicuous reticulate-punctate sculpture. In
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="5975EC2DE739AE570DCBD4386EF3E832" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
the sculpture is well-developed, too, but much more superficial, and does not cover the whole tergite. In addition, the petiolar node of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="ECF4D2F84A815CE4B7D55C7650E9F32A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
is distinctly wider than long (DPeI 111-188), which contrasts with the clearly longer than wide node of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="9FAD7D5491E2BAF8EAE69C923E022613" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
(DPeI 79-85).
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra,</taxonomicName>
|
||||
<taxonomicName id="94F53C8436F565A80D7E26D07C141F19" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra,</taxonomicName>
|
||||
the third species of the complex, is morphologically much closer to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon,</taxonomicName>
|
||||
<taxonomicName id="46685627A5DF3C59222493922AEC84D2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon,</taxonomicName>
|
||||
but differs in propodeal spine length, petiolar node shape, and general development of sculpture on waist segments and gaster.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra</taxonomicName>
|
||||
<taxonomicName id="9053F1E91FA1EC8896A2924BF2381B43" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">Tetramorium avaratra</taxonomicName>
|
||||
has generally much shorter propodeal spines (PSLI 27-34 without the Nosy Be specimens, PSLI 27-37 with the Nosy Be material) than
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="B963939A68CAEB498667B323642CEE97" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
(PSLI 37-44). Also, the petiolar node is higher and wider in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="93CC62B2271774081F1874C04D163189" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
(LPeI 54-66; DPeI 126-137) than in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="C7F78438F92689D8BE89641602CF9B3C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
(LPeI 63-73; DPeI 111-118), and the latter species has much better developed sculpture on the waist segments and the first gastral tergite. As noted in the description of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra,</taxonomicName>
|
||||
<taxonomicName id="4013F8D66FD7908C11C0DD07726BD272" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra,</taxonomicName>
|
||||
both species overlap in their distribution, and are found living in sympatry in Ankarana. However, both species display their species-characteristics in this locality, and can be well separated from each other by the diagnostics provided above.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,1373,1508]" pageId="53" pageNumber="54" type="mainText">
|
||||
<paragraph id="B84F01EACDEF3D61F678F67AF756A13A" box="[118,1122,1373,1508]" pageId="53" pageNumber="54" type="mainText">
|
||||
Despite this wide distribution range and relative flexibility in habitat preference,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="17A6723A7D1E5C9644B34EF983CF82E1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
is morphologically very stable throughout its range with very little variation in morphometry, shape, or colour. The morphological similarities with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum,</taxonomicName>
|
||||
<taxonomicName id="4D3D6A779990BEEB0947FD97A88431CE" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="dysalum">T. dysalum,</taxonomicName>
|
||||
as well as the potentially erroneous placement of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
<taxonomicName id="3071D9437275D5B8130F508B63E8E1AE" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
<taxonomicName id="94AF475670933054AF0FDFE0A21BC200" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon</taxonomicName>
|
||||
within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="7251BC2E032A99E349DB256B96577558" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
species group, are discussed in detail in the description of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra.</taxonomicName>
|
||||
<taxonomicName id="AFC506E6F2368A0E2F4C751C1F37D0C0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">T. avaratra.</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="multiple">
|
||||
<caption box="[118,1122,1545,1590]" pageId="54" pageNumber="55" type="caption">
|
||||
<paragraph box="[118,1122,1545,1590]" pageId="54" pageNumber="55" type="caption">
|
||||
<pageBreakToken pageId="54" pageNumber="55" start="start">FIGURES</pageBreakToken>
|
||||
<subSubSection id="A19D519A3C56AA9499E35E44634F5246" type="multiple">
|
||||
<caption id="361EA394E6F76B4FD46480C000056A9F" box="[118,1122,1545,1590]" pageId="54" pageNumber="55" type="caption">
|
||||
<paragraph id="A8D97571CC5D518632E6168AED609F1B" box="[118,1122,1545,1590]" pageId="54" pageNumber="55" type="caption">
|
||||
<pageBreakToken id="B3E5093B0402553F45742C813530EC91" pageId="54" pageNumber="55" start="start">FIGURES</pageBreakToken>
|
||||
108-110.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon,</taxonomicName>
|
||||
<taxonomicName id="CBE91A0BA156DCDCF43DC90BD86426BF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pleganon">T. pleganon,</taxonomicName>
|
||||
holotype (CASENT0280587). 108. Body in profile. 109. Body in dorsal view. 110. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,118,137]" pageId="55" pageNumber="56" type="mainText">
|
||||
<pageBreakToken pageId="55" pageNumber="56" start="start">Material</pageBreakToken>
|
||||
<subSubSection id="4630FA6934DE5DB4F5FC29C63BAFF942" type="materials_examined">
|
||||
<paragraph id="EC8D47D6FDDBB320DFE06C2BFB70ECCD" box="[118,295,118,137]" pageId="55" pageNumber="56" type="mainText">
|
||||
<pageBreakToken id="BCC467A3A18929C7A66082E861AD82E3" pageId="55" pageNumber="56" start="start">Material</pageBreakToken>
|
||||
examined
|
||||
</paragraph>
|
||||
<paragraph box="[118,1123,146,953]" pageId="55" pageNumber="56" type="mainText">
|
||||
<paragraph id="D3919B4722B660555A0AE757EBE29263" box="[118,1123,146,953]" pageId="55" pageNumber="56" type="mainText">
|
||||
MADAGASCAR: Antananarivo,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="1EEBD8B59CDDF13F294EEAE7D1910707" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="FDE71976B05DBF1F0B989755408FAD3E" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
d'Ambohitantely,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="2FBECAF4413849A15CA58E5C9613A127" originalValue="Forêt">Foret</normalizedToken>
|
||||
d Ambohitantely, 20.9 km 72° NE d Ankazobe, 18.22528 S, 47.28683 E, 1410 m, montane rainforest, 17.-22.IV.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="0532479E0365FEBB200C80D7767BB09B" originalValue="Forêt">Foret</normalizedToken>
|
||||
Ambato, 26.6 km 33° Ambanja, 13.4645 S, 48.55167 E, 150 m, rainforest, 8.-10.XII.2004 (B.L. Fisher); Antsiranana, Ampasindava,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="E9B0C16209B9BDB1D2D6A2924E6575FE" originalValue="Forêt">Foret</normalizedToken>
|
||||
d'Ambilanivy, 3.9 km 181° S Ambaliha, 13.79861 S, 48.16167 E, 600 m, rainforest, 4.-9.III.2001 (B.L. Fisher, C. Griswold et al.); Antsiranana, Andapa, 14° 40' S, 49° 39' E, 500 m, degraded forest, 26.I.1991 (G.D. Alpert); Antsiranana, 84 km SW Sambava on road to Andapa, 70-160 m, degraded forest, 17.II.1977 (W.L. & D.E. Brown); Antsiranana, Res. Ankarana, 12° 54' S, 49° 6' E, 100 m, tropical dry forest, 22.VIII.1992 (G.D. Alpert); Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="9C460AC00C4B11A15EACA7890711FE1E" originalValue="Forêt">Foret</normalizedToken>
|
||||
d' Antsahabe, 11.4 km 275° W Daraina, 13.21167 S, 49.55667 E, 550 m, tropical dry forest, 12.XII.2003 (B.L. Fisher); Antsiranana, R.S. Manongarivo, 10.8 km 229° SW Antanambao, 13.96167 S, 48.43333 E, 400 m, rainforest, 8.XI.1998 (B.L. Fisher); Antsiranana, R.S. Manongarivo, 12.8 km 228° SW Antanambao, 13.97667 S, 48.42333 E, 780 m, rainforest, 11.-17.X.1998 (B.L. Fisher); Antsiranana, Makirovana forest, 14.17066 S, 49.95409 E, 415 m, rainforest, 28.-29.IV.2011 (B.L. Fisher et al.); Fianarantsoa, Antapia II Non Protected Area, 26.47 km SW Ambositra, 20.71889 S, 47.0867 E, 1494 m, Uapaca woodland, 4.-6.II.2010 (A. Ravelomanana); Fianarantsoa, Antohatsahomby I Non Protected Area, 22.77 km NW Ambatofinandrahana, 20.55056 S, 46.58562 E, 1550 m, Uapaca woodland, 15.-17.III.2010 (A. Ravelomanana); Fianarantsoa, Mampiarika III Non Protected Area, 28.93 km SW Ambositra, 20.73583 S, 47.08399 E, 1487 m, Uapaca woodland, 1.-2.II.2010 (A. Ravelomanana); Fianarantsoa, Soanierenana I Non Protected Area, 25.33 km SW Ambositra, 20.72139 S, 47.10994 E, 1723 m, savannah grassland, 6.-8.II.2010 (A. Ravelomanana); Mahajanga, Parc National d'Ankarafantsika,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="5FDBAEAF2AD0C34675C04A733AD63F81" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Tsimaloto, 18.3 km 46° NE de Tsaramandroso, 16.22806 S, 47.14361 E, 135 m, tropical dry forest, 2.-8.IV.2001 (B.L. Fisher, C. Griswold et al.); Mahajanga,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="910D61F5427227EF6229088AC5C6F20B" originalValue="Réserve">Reserve</normalizedToken>
|
||||
d'Ankoririka, 10.6 km 13° NE de Tsaramandroso, 16.26722 S, 47.04861 E, 210 m, tropical dry forest, 9.-14.IV.2001 (B.L. Fisher, C. Griswold et al.); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.84773 S, 48.29568 E, 1000 m, montane rainforest, 5.-8.III.2007 (B.L. Fisher et al.); Toamasina, Manakambahiny, near Vavatenina Forest, 17.46667 S, 49.35 E, 9.II.1995 (A. Pauly); Toamasina, MoraranoChrome, 17.75 S, 47.98333 E, forest, 1.I.1992 (A. Pauly); Toamasina, S.F. Tampolo, 10 km NNE Fenoarivo Atn., 17.2825 S, 49.43 E, 10 m, littoral rainforest, 10.IV.1997 (B.L. Fisher); Toliara,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="86D7AA0D254A105D4B8FAC11B91DF66B" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="05D68478F57F21ADF76DB4F62D1A40D8" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
d'Ambohijanahary,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="7518A9C1B073816B40500271A506EE58" originalValue="Forêt">Foret</normalizedToken>
|
||||
d'Ankazotsihitafototra, 35.2 km 312° NW Ambaravaranala, 18.26667 S, 45.40667 E, 1050 m, montane rainforest, 13.-17.I.2003 (B.L. Fisher et al.); Toliara,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="909BD44FBECA5A6BACAC565163E0B73A" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="3CA894EAAC252076DD21205752F0B5BC" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Kalambatritra, 23.4185 S, 46.4583 E, 1365 m, grassland, 8.II.2009 (B.L. Fisher et al.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,206 +1,205 @@
|
|||
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|
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|
||||
<taxonomicName authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">
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|
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|
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<taxonomicName id="213678485E5CB9ED86B39CF4844C8D66" ID-CoL="55PNN" authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">
|
||||
Tetramorium isectum
|
||||
<authority>Bolton</authority>
|
||||
<authority id="32F3A1DEB8272E2043422AE20F9741F2">Bolton</authority>
|
||||
,
|
||||
</taxonomicName>
|
||||
1979
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,345,455,474]" pageId="36" pageNumber="37" type="mainText">(Figs. 52, 90, 91, 92, 141)</paragraph>
|
||||
<subSubSection id="03753A56F0DAF39D3E3D7FF6DBA6F742" type="description">
|
||||
<paragraph id="544903387E64FAE7128DDB2703BBD8D6" box="[118,345,455,474]" pageId="36" pageNumber="37" type="mainText">(Figs. 52, 90, 91, 92, 141)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="reference_group">
|
||||
<paragraph box="[118,1122,511,552]" pageId="36" pageNumber="37" type="mainText">
|
||||
<taxonomicName authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">
|
||||
<subSubSection id="F05377646910B3F9F2970FECF0EDE327" type="reference_group">
|
||||
<paragraph id="5F1A56A6B5309221047AC87A11516B35" box="[118,1122,511,552]" pageId="36" pageNumber="37" type="mainText">
|
||||
<taxonomicName id="95C0226C9656F897F8931E2C07468AEC" authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">
|
||||
Tetramorium isectum
|
||||
<authority>Bolton</authority>
|
||||
<authority id="C551A3F506EA9C09A7962386DDD747C5">Bolton</authority>
|
||||
,
|
||||
</taxonomicName>
|
||||
1979: 145. Holotype worker, MADAGASCAR, Beforona, 500 m, forest humus and litter, IX.1974 (A. Peyrieras) (MCZ: CASENT0172829) [examined].
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,587,605]" pageId="36" pageNumber="37" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,615,719]" pageId="36" pageNumber="37" type="mainText">
|
||||
<subSubSection id="69F2DBB8DBC4EABECFE532ECB8C4C49A" type="diagnosis">
|
||||
<paragraph id="1D786AEBAEAC662B1B828CA1E3A7F04C" box="[118,208,587,605]" pageId="36" pageNumber="37" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="2CD88C8920F13B4B659F12F9FF3E4FCD" box="[118,1121,615,719]" pageId="36" pageNumber="37" type="mainText">
|
||||
Within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="A49574B81ED444D691E77564C7D162E0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
complex
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="1C6DE6ACD235B8237B703AADF7BEF2C1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
is distinguishable by the following characters: very small eyes (OI 15-16); long propodeal spines (PSLI 28-33); petiolar node with the anterodorsal margin slightly situated higher and more angular than the posterodorsal margin; posterodorsal corner of petiole not strongly protruding posteriorly; body of uniform bright orange colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,756,774]" pageId="36" pageNumber="37" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,784,916]" pageId="36" pageNumber="37" type="mainText">HL 0.90-1.10 (0.96); HW 0.85-1.50 (0.93); SL 0.65-0.83 (0.72); EL 0.14-0.18 (0.15); PH 0.42-0.54 (0.47); PW 0.58-0.76 (0.68); WL 1.08-1.39 (1.17); PSL 0.26-0.36 (0.29); PTL 0.25-0.34 (0.28); PTH 0.33-0.40 (0.35); PTW 0.24-0.29 (0.26); PPL 0.27-0.35 (0.31); PPH 0.31-0.39 (0.34); PPW 0.32-0.40 (0.34); CI 94-99 (97); SI 74-79 (77); OI 15-16 (16); DMI 54-60 (58); LMI 39-42 (40); PSLI 28-33 (30); PeNI 36-42 (38); LPeI 71-85 (81); DPeI 86-98 (92); PpNI 49-54 (50); LPpI 82-97 (91); DPpI 103-120 (111); PPI 128-136 (132) (12 measured).</paragraph>
|
||||
<paragraph box="[118,1122,925,1563]" pageId="36" pageNumber="37" type="mainText">Head usually weakly longer than wide (CI 94-99). Posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes short, not reaching posterior head margin (SI 74-79). Eyes very small (OI 15-16). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 39-42). Propodeal spines long, spinose, and acute (PSLI 28-33); propodeal lobes short, triangular to elongate-triangular, and acute. Petiolar node in profile rectangular nodiform, around 1.2 to 1.4 times higher than long (LPeI 71-85), anterior and posterior faces approximately parallel, anterodorsal margin situated weakly higher and more angulate than posterodorsal margin, dorsum tapering weakly backwards posteriorly; node in dorsal view between 1.0 to 1.2 times longer than wide (DPeI 86-98). Postpetiole in profile globular, approximately 1.0 to 1.2 times higher than long (LPpI 82-97); in dorsal view between 1.0 and 1.2 times wider than long (DPpI 103-120). Postpetiole in profile appearing approximately as voluminous as petiolar node, in dorsal view approximately 1.3 to 1.4 times wider than petiolar node (PPI 128-136). Mandibles distinctly longitudinally rugose; clypeus generally with three distinct longitudinal rugae/rugulae, sometimes with few more but much weaker rugulae present, and very rarely clypeus with irregular rugulation; cephalic dorsum between frontal carinae with 8 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae very rarely with cross-meshes; lateral and ventral head longitudinally rugose with very few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured, smooth, and shining. Waist segments irregularly rugulose with distinct punctate ground sculpture, laterally ground sculpture better developed and rugulae weaker. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with decumbent to suberect standing hairs. Body colour bright yellow to orange.</paragraph>
|
||||
<subSubSection id="75DAA377735017CF04B34DC74CEA90C4" type="description">
|
||||
<paragraph id="5852B454FBE4CCA1C0E73B62D0BDE8A5" box="[118,226,756,774]" pageId="36" pageNumber="37" type="mainText">Description</paragraph>
|
||||
<paragraph id="1698A0C6D62B64F9C690ED750B218E32" box="[118,1121,784,916]" pageId="36" pageNumber="37" type="mainText">HL 0.90-1.10 (0.96); HW 0.85-1.50 (0.93); SL 0.65-0.83 (0.72); EL 0.14-0.18 (0.15); PH 0.42-0.54 (0.47); PW 0.58-0.76 (0.68); WL 1.08-1.39 (1.17); PSL 0.26-0.36 (0.29); PTL 0.25-0.34 (0.28); PTH 0.33-0.40 (0.35); PTW 0.24-0.29 (0.26); PPL 0.27-0.35 (0.31); PPH 0.31-0.39 (0.34); PPW 0.32-0.40 (0.34); CI 94-99 (97); SI 74-79 (77); OI 15-16 (16); DMI 54-60 (58); LMI 39-42 (40); PSLI 28-33 (30); PeNI 36-42 (38); LPeI 71-85 (81); DPeI 86-98 (92); PpNI 49-54 (50); LPpI 82-97 (91); DPpI 103-120 (111); PPI 128-136 (132) (12 measured).</paragraph>
|
||||
<paragraph id="7288CA8D83C0717E47D1D0E0346D5B47" box="[118,1122,925,1563]" pageId="36" pageNumber="37" type="mainText">Head usually weakly longer than wide (CI 94-99). Posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes short, not reaching posterior head margin (SI 74-79). Eyes very small (OI 15-16). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 39-42). Propodeal spines long, spinose, and acute (PSLI 28-33); propodeal lobes short, triangular to elongate-triangular, and acute. Petiolar node in profile rectangular nodiform, around 1.2 to 1.4 times higher than long (LPeI 71-85), anterior and posterior faces approximately parallel, anterodorsal margin situated weakly higher and more angulate than posterodorsal margin, dorsum tapering weakly backwards posteriorly; node in dorsal view between 1.0 to 1.2 times longer than wide (DPeI 86-98). Postpetiole in profile globular, approximately 1.0 to 1.2 times higher than long (LPpI 82-97); in dorsal view between 1.0 and 1.2 times wider than long (DPpI 103-120). Postpetiole in profile appearing approximately as voluminous as petiolar node, in dorsal view approximately 1.3 to 1.4 times wider than petiolar node (PPI 128-136). Mandibles distinctly longitudinally rugose; clypeus generally with three distinct longitudinal rugae/rugulae, sometimes with few more but much weaker rugulae present, and very rarely clypeus with irregular rugulation; cephalic dorsum between frontal carinae with 8 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae very rarely with cross-meshes; lateral and ventral head longitudinally rugose with very few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured, smooth, and shining. Waist segments irregularly rugulose with distinct punctate ground sculpture, laterally ground sculpture better developed and rugulae weaker. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with decumbent to suberect standing hairs. Body colour bright yellow to orange.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="multiple">
|
||||
<caption box="[118,1122,1525,1570]" pageId="37" pageNumber="38" type="caption">
|
||||
<paragraph box="[118,1122,1525,1570]" pageId="37" pageNumber="38" type="caption">
|
||||
<pageBreakToken pageId="37" pageNumber="38" start="start">FIGURES</pageBreakToken>
|
||||
<subSubSection id="5C0328EA6371E39C0794F93AB8F8E164" type="multiple">
|
||||
<caption id="20388B3731DE8A28437EB1AE7BA04A1A" box="[118,1122,1525,1570]" pageId="37" pageNumber="38" type="caption">
|
||||
<paragraph id="79927E0FF09691EF57B38B376CB0EFB3" box="[118,1122,1525,1570]" pageId="37" pageNumber="38" type="caption">
|
||||
<pageBreakToken id="A5824958492D9CCF67BDDCCA6FAEAA52" pageId="37" pageNumber="38" start="start">FIGURES</pageBreakToken>
|
||||
90-92.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="isectum">T. isectum,</taxonomicName>
|
||||
<taxonomicName id="B3B0DF5249C3669C36442BC5550E485E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="isectum">T. isectum,</taxonomicName>
|
||||
holotype (CASENT0172829). 90. Body in profile. 91. Body in dorsal view. 92. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,118,136]" pageId="38" pageNumber="39" type="mainText">
|
||||
<pageBreakToken pageId="38" pageNumber="39" start="start">Notes</pageBreakToken>
|
||||
<subSubSection id="A487C56331972A89305DA7FCA6C3550E" type="discussion">
|
||||
<paragraph id="7B3006589BCCBFBA78CC24EF2122692B" box="[118,170,118,136]" pageId="38" pageNumber="39" type="mainText">
|
||||
<pageBreakToken id="8E4D14A9987BEB9BC332676E2B027A69" pageId="38" pageNumber="39" start="start">Notes</pageBreakToken>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,147,362]" pageId="38" pageNumber="39" type="mainText">
|
||||
<paragraph id="21921888898690914ED3D50E352B95C7" box="[118,1121,147,362]" pageId="38" pageNumber="39" type="mainText">
|
||||
Prior to our study
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="351F29FAD3F9C9D0638A339BC0A9361D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
was only known from the holotype from Beforona. Fortunately, we had more material available because the species has been collected several times since the original description of Bolton (1979).
|
||||
<taxonomicName genus="Tetramoirum" pageId="38" pageNumber="39" rank="species" species="isectum">Tetramoirum isectum</taxonomicName>
|
||||
<taxonomicName id="F94C2BF37F6FAF96169297F04F4DE34F" genus="Tetramoirum" pageId="38" pageNumber="39" rank="species" species="isectum">Tetramoirum isectum</taxonomicName>
|
||||
does not seem to be as rare as might be expected from the original description. We were able to examine material from a number of localities, although the species is admittedly never very common or abundant.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
<taxonomicName id="61ED479FF4DE1B8F510BE99B399FB935" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
is found in a strip in eastern Madagascar, which ranges from central-eastern to northeastern Madagascar. The southernmost localities are Ambatovy, Andasibe-Mantadia, and Sahafina, and the northernmost are Montagne d'Anjanaharibe and Amparihibe. The species prefers rainforests and montane rainforests at elevations of 125 to 1040 m, and was mostly collected from leaf litter.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,372,728]" pageId="38" pageNumber="39" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
<paragraph id="403341234E60E4662100282CD297761D" box="[118,1122,372,728]" pageId="38" pageNumber="39" type="mainText">
|
||||
<taxonomicName id="9C5124EB74BBC566A724236C7AF02BAD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
is morphologically fairly close to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
<taxonomicName id="843F4416B66EE9D92DC0EF4BEFF462BE" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
and during this revision we considered the synonymisation of the first under the latter. Both differ mainly in eye size, which is very small in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="877355A2AB8B83D879BA271807EB0060" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
(OI 15-16) versus small to moderately large in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="811B372E662B56278CC425C0B4082EDA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
(OI 19-25), but there a few more supporting characters found in both species, although not consistently. All specimens of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="06DAA201F7C9D6CA3E409332AC9C79C3" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
have, in addition to very small eyes, a petiolar node shape with the anterodorsal margin higher and sharper than the posterodorsal with a dorsum that tapers weakly posteriorly, as well as a bright orange body colour. However, within the vast
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="55A0DC9BEF03CE1933F87B56B068CB33" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
material available, there are specimens with a node shape like the one of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum,</taxonomicName>
|
||||
<taxonomicName id="C396902A1C8B9A6F5412C296BC9A77DF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum,</taxonomicName>
|
||||
although these are mainly restricted to northwestern Madagascar where
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="4BE17086F63A25E9C88481EDBDDFA009" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
does not occur. Many specimens of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="08325B7BEBF7DFA2DEABC7D1C435B4A5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
are also bright orange in colour. Nevertheless,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="A3B909D97B3A95DAB6B2DF9EA764DB45" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
was found to live in sympatry with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="32BEC5488391B4ABEFCA61CED5E9F903" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
throughout most of its distribution range, and remained remarkably recognisable. The diagnostic characters of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="3572E00FEDC090DCB1C56E7C397C4174" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
provided above are very consistent throughout all the material studied, and no intermediate forms seem to exist. With these facts in mind, we keep
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="08918AEE76278936F366EE132575F6C6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
a separate though relatively uncommon species which co-occurs regularly with the much more common and abundant
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
<taxonomicName id="D918A150CB97665437CC6DD5AA6E2D23" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,737,1037]" pageId="38" pageNumber="39" type="mainText">
|
||||
<paragraph id="D03E14D2490FA35F98A43C45F9981D36" box="[118,1122,737,1037]" pageId="38" pageNumber="39" type="mainText">
|
||||
Apart from the similarities with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="185628BE3EFD54A0F2E8428E1DD6C303" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
mentioned above,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="99A081D95297E027E0F31DD315FD24C7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
is easily recognisable within the species complex. The very small eyes (OI 15-16) are very conspicuous with the caveat that
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="76D024EBB9D58D4C23A873775467311C" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="electrum">
|
||||
T. electrum
|
||||
<authority>,</authority>
|
||||
<authority id="3BBEAAAB52CD33FE1C38AE10A98890EA">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="elf">
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="9CD2214E08655019673019E7ABA8D99F" authorityName="," genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="elf">
|
||||
<taxonomicName id="7977012DDD0FBF8A39A3557FF0689D47" genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
elf
|
||||
<authority>,</authority>
|
||||
<authority id="03C1E398E6A54237FFDAFC033C8ED946">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
<taxonomicName id="454CB6B0ACC502771D5E22A64F8204B4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
and
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="D9D322B6A75842620FB60F4B28A1D238" genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="nify">T. nify</taxonomicName>
|
||||
also have generally smaller eyes (OI 16-19). However,
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="93BCEFA86F6A09ADA83967CF75E679D8" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="electrum">
|
||||
T. electrum
|
||||
<authority>,</authority>
|
||||
<authority id="373AAB24F803C7EFBDBA0051E79C4EBA">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="elf">
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="E697927B9D190AD7381FD3022FE8CA39" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="elf">
|
||||
<taxonomicName id="DEFC62175E226AAD9DA2D37A53F53BF5" genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
elf,
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="E12734C3D1EE9FFB3514FC8FF76259BA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
have very long to extremely long propodeal spines (PSLI 46-64), whereas the spines of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="F3695E61417AA2350D4C9F3513AA5469" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
are much shorter (PSLI 28-35).
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="nify">Tetramorium nify</taxonomicName>
|
||||
<taxonomicName id="E1DD211AD79A64DC6B77023C58796033" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="nify">Tetramorium nify</taxonomicName>
|
||||
is also unlikely to be misidentified with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="7EC99C167F1E88F252D8BC95CFE78E8A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
since the latter is bright orange in colour and the former is very dark brown to black in colour. In addition, they have fairly differently shaped petiolar nodes. The node of
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="172434FB7B8A17413D3132465F9EDC6B" genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="nify">T. nify</taxonomicName>
|
||||
has antero- and posterodorsal margins at the same height and is only faintly higher than long (LPeI 91-100), whereas in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="FDF46B6E2637DA57D54E6F1BAA7E2FC6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
the anterodorsal margin is usually weakly higher and the node is between 1.2 to 1.4 times higher than long (LPeI 71-85). The remaining species, T.
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="ala">
|
||||
<taxonomicName id="A8DD140993A37F488705AAA5C0594D8A" authorityName="," genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="ala">
|
||||
ala
|
||||
<authority>,</authority>
|
||||
<authority id="11D6E0C9B3B562C052D21F3278C8BEF8">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
<taxonomicName id="B58B64CEFF31AFD532854DA75E3618BA" genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="0DFA1D7899671029E8292A14A6D104C1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
, and
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="voasary">T. voasary,</taxonomicName>
|
||||
<taxonomicName id="7E766B1DDE082CCE17170821132FF590" genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="voasary">T. voasary,</taxonomicName>
|
||||
have much larger eyes (OI 20-24) and differently shaped petiolar nodes, and are thus not easy to confuse with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum.</taxonomicName>
|
||||
<taxonomicName id="4EABE52593B43293D6E5965188D7DF2A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum.</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,1074,1093]" pageId="38" pageNumber="39" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,1103,1572]" pageId="38" pageNumber="39" type="mainText">
|
||||
<subSubSection id="9486CF626BA2143AC4E2D5F042C3BC44" type="materials_examined">
|
||||
<paragraph id="85DE7BD6C7AE6EF31B7217260A79D9C5" box="[118,295,1074,1093]" pageId="38" pageNumber="39" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="BBD9583164F27F58D03AD0D571B097BE" box="[118,1122,1103,1572]" pageId="38" pageNumber="39" type="mainText">
|
||||
MADAGASCAR: Mahajanga,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="8C2CCD241F43F58CF0F7BE0CD1D71CEF" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="A9B2DE73710F9D4A5B3D4FCC2D6EC5FF" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Marotandrano, Marotandrano 48.3 km S Mandritsara, 16.2832 S, 48.8144 E, 865 m, transition humid forest, 6.-8.XII.2007 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="52975E2303C37DFF6746A61EDE8C28A6" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="FA715865D6EEDF7A6C9A3B7CC7CFD241" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato river, 16.8175 S, 49.295 E, 360 m, rainforest, 25.-27.II.2010 (B.L. Fisher et al.); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.8496 S, 48.2947 E, 1010 m, montane rainforest, 3.-6.III.2007 (B.L. Fisher et al.); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.8477 S, 48.2947 E, 1000 m, montane rainforest, 5.-8.III.2007 (B.L. Fisher et al.); Toamasina, Res. Ambodiriana, 4.8 km 306° Manompana, along Manompana river, 16.6723 S, 49.7012 E, 125 m, rainforest, 18.XI.2005 (B.L. Fisher et al.); Toamasina, Amparihibe, 15° 2' S, 49° 34' E, II.-III.2003 (K.A. Jackson & D. Carpenter); Toamasina, Station
|
||||
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
|
||||
<normalizedToken id="C597BFDB6D86CF6DA3DF8BC595A75EFC" originalValue="forestière">forestiere</normalizedToken>
|
||||
Analamazaotra, Analamazaotra 1.3 km S Andasibe, 18.3847 S, 48.4127 E, 980 m, montane rainforest, 11.-13.XII.2007 (B.L. Fisher et al.); Toamasina, Parc National
|
||||
<normalizedToken originalValue="d´Andasibe-Mantadia">d'Andasibe-Mantadia</normalizedToken>
|
||||
<normalizedToken id="AABB6A1FD52CA26639D03EC037DC5534" originalValue="d´Andasibe-Mantadia">d'Andasibe-Mantadia</normalizedToken>
|
||||
,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="F686914F259631198D09C61762F98A04" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Mantadia, 25.7 km 248° Moramanga, 18.814 S, 48.4303 E, 1040 m, rainforest, 14.VII.2006 (F.N. Raharimalala & B. Blaimer); Toamasina, Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo, 15.1883 S, 49.615 E, 470 m, rainforest, 8.-12.III.2003 (B.L Fisher, C. Griswold et al.); Toamasina, Beforona, 500 m, IX.1974 (A. Peyrieras); Toamasina, Parc National Mananara-Nord, 7.1 km 261° Antanambe, 16.455 S, 49.7875 E, 225 m, rainforest, 14.XI.2005 (B.L. Fisher et al.); Toamasina, Sahafina forest 11.4 km W Brickaville, 18.8144 S, 48.9621 E, 140 m, rainforest, 13.-14.XII.2007 (B.L. Fisher et al.); Toamasina, Parc National de Zahamena, Onibe River, 17.7591 S, 48.8547 E, 780 m, rainforest, rainforest, 21.-23.II.2009 (B.L. Fisher et al.); Toamasina, Parc National de Zahamena,
|
||||
</paragraph>
|
||||
<paragraph box="[118,1120,118,166]" pageId="39" pageNumber="40" type="mainText">
|
||||
<pageBreakToken pageId="39" pageNumber="40" start="start">Tetezambatana</pageBreakToken>
|
||||
<paragraph id="625674598FE6C82000DF5269440692E2" box="[118,1120,118,166]" pageId="39" pageNumber="40" type="mainText">
|
||||
<pageBreakToken id="CB048EFDF65377527946AFB81E615196" pageId="39" pageNumber="40" start="start">Tetezambatana</pageBreakToken>
|
||||
forest, near junction of Nosivola and Manakambahiny Rivers, 17.74298 S, 48.72936 E, 860 m, rainforest, 18.-19.II.2009 (B.L. Fisher et al.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,109 +1,109 @@
|
|||
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<mods:title id="56CFFD318EE0E0371649B52FAE8D9C06">ZooKeys</mods:title>
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<mods:date>2012</mods:date>
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<mods:classification id="DA96C26CF58624385896D10FFC2A29FD">journal article</mods:classification>
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<treatment ID-GBIF-Taxon="152033753" LSID="urn:lsid:plazi:treatment:E318547193CA4D8F861C1E8B155834C9" httpUri="http://treatment.plazi.org/id/E318547193CA4D8F861C1E8B155834C9" lastPageId="4" lastPageNumber="145" pageId="3" pageNumber="144">
|
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<subSubSection pageId="3" pageNumber="144" type="nomenclature">
|
||||
<paragraph pageId="3" pageNumber="144">
|
||||
<taxonomicName LSID="http://species-id.net/wiki/Ostoma_fraterna" authority="Randall, 1838" authorityName="Randall" authorityYear="1838" class="Insecta" family="Trogossitidae" genus="Ostoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ostoma fraterna" order="Coleoptera" pageId="3" pageNumber="144" phylum="Arthropoda" rank="species" species="fraterna">Ostoma fraterna (Randall, 1838)</taxonomicName>
|
||||
<treatment id="E318547193CA4D8F861C1E8B155834C9" ID-GBIF-Taxon="152033753" LSID="urn:lsid:plazi:treatment:E318547193CA4D8F861C1E8B155834C9" httpUri="http://treatment.plazi.org/id/E318547193CA4D8F861C1E8B155834C9" lastPageId="4" lastPageNumber="145" pageId="3" pageNumber="144" scope_family="Trogossitidae" scope_order="Coleoptera" scope_subFamily="Peltinae" scope_tribe="Peltini">
|
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<subSubSection id="7A72D039AD280019D163855579FF2D84" pageId="3" pageNumber="144" type="nomenclature">
|
||||
<paragraph id="0486262BD5B2D1BF77F60BD92B5459AE" pageId="3" pageNumber="144">
|
||||
<taxonomicName id="F17031637A38EC9E807B78B33A8C287E" LSID="http://species-id.net/wiki/Ostoma_fraterna" authority="Randall, 1838" authorityName="Randall" authorityYear="1838" genus="Ostoma" lsidName="Ostoma fraterna" pageId="3" pageNumber="144" rank="species" species="fraterna">Ostoma fraterna (Randall, 1838)</taxonomicName>
|
||||
Map 2
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="4" lastPageNumber="145" pageId="3" pageNumber="144" type="material examined">
|
||||
<paragraph pageId="3" pageNumber="144">Material examined.</paragraph>
|
||||
<paragraph lastPageId="4" lastPageNumber="145" pageId="3" pageNumber="144">
|
||||
<subSubSection id="E04DCACF395B51F682AEA8BBFE2B5159" lastPageId="4" lastPageNumber="145" pageId="3" pageNumber="144" type="material examined">
|
||||
<paragraph id="4724A06DD2970815427DD3A81BBDA387" pageId="3" pageNumber="144">Material examined.</paragraph>
|
||||
<paragraph id="8BE7342B96428EB2926C9A2230E7A3D4" lastPageId="4" lastPageNumber="145" pageId="3" pageNumber="144">
|
||||
Additional New Brunswick records. Charlotte Co., St. Stephen, 17.V.1933, J. B.
|
||||
<normalizedToken originalValue="O’Donnel">O'Donnel</normalizedToken>
|
||||
<normalizedToken id="4AA5C98509DFED994693CC82E0368FA4" originalValue="O’Donnel">O'Donnel</normalizedToken>
|
||||
(8, AFC). Restigouche Co., Dionne Brook P.N.A.,
|
||||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="47.903">
|
||||
<pageBreakToken pageId="4" pageNumber="145" start="start">47.9030</pageBreakToken>
|
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<geoCoordinate id="DF94BDE820358703768198820DE77920" direction="north" orientation="latitude" precision="5" value="47.903">
|
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<pageBreakToken id="6FE84B9A73F50BBEC824E9A0234D5705" pageId="4" pageNumber="145" start="start">47.9030</pageBreakToken>
|
||||
°N
|
||||
</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-68.3503">68.3503°W</geoCoordinate>
|
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<geoCoordinate id="569A42856D61FF1BECAB81872585F8CB" direction="west" orientation="longitude" precision="5" value="-68.3503">68.3503°W</geoCoordinate>
|
||||
, 30.
|
||||
<normalizedToken originalValue="V–">V-</normalizedToken>
|
||||
<normalizedToken id="A4C2DE07E8AE9A066F391E423862DC65" originalValue="V–">V-</normalizedToken>
|
||||
15.VI.2011, M. Roy & V. Webster, old-growth northern hardwood forest, Lindgren funnel trap (1, RWC).
|
||||
</paragraph>
|
||||
<caption pageId="4" pageNumber="145">
|
||||
<paragraph pageId="4" pageNumber="145">
|
||||
<caption id="2308A76385BE663757878701D808E93B" pageId="4" pageNumber="145">
|
||||
<paragraph id="52325D47329395E6F8C569C83BED3B6A" pageId="4" pageNumber="145">
|
||||
Map 2. Collection localities in New Brunswick, Canada of
|
||||
<taxonomicName class="Insecta" family="Trogossitidae" genus="Ostoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ostoma fraterna" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="fraterna">Ostoma fraterna</taxonomicName>
|
||||
<taxonomicName id="A44BDFCA2BEF97EB19CBA3125C2128DF" genus="Ostoma" lsidName="Ostoma fraterna" pageId="4" pageNumber="145" rank="species" species="fraterna">Ostoma fraterna</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="4" pageNumber="145" type="collection and habitat data">
|
||||
<paragraph pageId="4" pageNumber="145">Collection and habitat data.</paragraph>
|
||||
<paragraph pageId="4" pageNumber="145">
|
||||
<subSubSection id="3CA820B1E910E81231213965D7B8886D" pageId="4" pageNumber="145" type="collection and habitat data">
|
||||
<paragraph id="488BA830C93D4EC7DC4A2D3C86791476" pageId="4" pageNumber="145">Collection and habitat data.</paragraph>
|
||||
<paragraph id="2F980A2394425A608B940A35904F94B9" pageId="4" pageNumber="145">
|
||||
The specimen with habitat data from New Brunswick was collected from a Lindgren funnel trap deployed in an old-growth northern hardwood forest with sugar maple and yellow birch. Specimens were collected during May and June.
|
||||
<taxonomicName class="Insecta" family="Trogossitidae" genus="Ostoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ostoma fraterna" order="Coleoptera" pageId="4" pageNumber="145" phylum="Arthropoda" rank="species" species="fraterna">Ostoma fraterna</taxonomicName>
|
||||
<taxonomicName id="0C4266624E2100BC005951CA8A726700" genus="Ostoma" lsidName="Ostoma fraterna" pageId="4" pageNumber="145" rank="species" species="fraterna">Ostoma fraterna</taxonomicName>
|
||||
has been found under bark of spruce (
|
||||
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea" order="Pinales" pageId="4" pageNumber="145" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
|
||||
<taxonomicName id="5123E12A92EE263A22B74CA36F46C398" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea" order="Pinales" pageId="4" pageNumber="145" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
|
||||
sp.) and in various
|
||||
<taxonomicName genus="Polyporaceae" lsidName="Polyporaceae" pageId="4" pageNumber="145" rank="genus">Polyporaceae</taxonomicName>
|
||||
<taxonomicName id="C0544092EFE906F8F3BA0F7585E75AA7" genus="Polyporaceae" lsidName="Polyporaceae" pageId="4" pageNumber="145" rank="genus">Polyporaceae</taxonomicName>
|
||||
species (
|
||||
<taxonomicName class="Agaricomycetes" family="Fomitopsidaceae" genus="Piptoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Piptoporus betulinus" order="Polyporales" pageId="4" pageNumber="145" phylum="Basidiomycota" rank="species" species="betulinus">Piptoporus betulinus</taxonomicName>
|
||||
<taxonomicName id="19AB1C18EECABF76A0FFC07F8E72274B" class="Agaricomycetes" family="Fomitopsidaceae" genus="Piptoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Piptoporus betulinus" order="Polyporales" pageId="4" pageNumber="145" phylum="Basidiomycota" rank="species" species="betulinus">Piptoporus betulinus</taxonomicName>
|
||||
(Fr.) Kar.,
|
||||
<taxonomicName class="Agaricomycetes" family="Fomitopsidaceae" genus="Spongiporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Spongiporus leucospongia" order="Polyporales" pageId="4" pageNumber="145" phylum="Basidiomycota" rank="species" species="leucospongia">Spongiporus leucospongia</taxonomicName>
|
||||
<taxonomicName id="24F94B4BC83C717EFEC36A95C7FB959A" class="Agaricomycetes" family="Fomitopsidaceae" genus="Spongiporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Spongiporus leucospongia" order="Polyporales" pageId="4" pageNumber="145" phylum="Basidiomycota" rank="species" species="leucospongia">Spongiporus leucospongia</taxonomicName>
|
||||
(Cke. and Hark.) Murr., and
|
||||
<taxonomicName class="Agaricomycetes" family="Polyporaceae" genus="Tyromyces" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tyromyces fragilis" order="Polyporales" pageId="4" pageNumber="145" phylum="Basidiomycota" rank="species" species="fragilis">Tyromyces fragilis</taxonomicName>
|
||||
<taxonomicName id="FD388076F291C26BED6EEABBA5365866" class="Agaricomycetes" family="Polyporaceae" genus="Tyromyces" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tyromyces fragilis" order="Polyporales" pageId="4" pageNumber="145" phylum="Basidiomycota" rank="species" species="fragilis">Tyromyces fragilis</taxonomicName>
|
||||
(Fr.) Donk) (
|
||||
<bibRefCitation author="Barron, JR" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="11" pageNumber="152" pagination="1 - 143" title="A revision of the Trogossitidae of America north of Mexico." url="10.4039/entm10375fv" volume="75" year="1971">Barron 1971</bibRefCitation>
|
||||
<bibRefCitation id="C1A73C7201A7AFC06F8CC56044DE8BE6" author="Barron, JR" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="11" pageNumber="152" pagination="1 - 143" title="A revision of the Trogossitidae of America north of Mexico." url="10.4039/entm10375fv" volume="75" year="1971">Barron 1971</bibRefCitation>
|
||||
, 1996).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="4" pageNumber="145" type="distribution">
|
||||
<paragraph pageId="4" pageNumber="145">Distribution in Canada and Alaska.</paragraph>
|
||||
<paragraph pageId="4" pageNumber="145">
|
||||
<subSubSection id="11E788818D05F710753DF02102C40226" pageId="4" pageNumber="145" type="distribution">
|
||||
<paragraph id="F9FC233D11258E3DA217DD26B69BD651" pageId="4" pageNumber="145">Distribution in Canada and Alaska.</paragraph>
|
||||
<paragraph id="33500DA400FCE90F928155117571A3CF" pageId="4" pageNumber="145">
|
||||
AK, YK, NT, BC, AB, SK, ON, QC, NB, NS, NF (
|
||||
<bibRefCitation pageId="4" pageNumber="145">Bousquet 1991</bibRefCitation>
|
||||
<bibRefCitation id="DFA5A52B7CAB0916657AEC85EAE2FE34" pageId="4" pageNumber="145">Bousquet 1991</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka 2011</bibRefCitation>
|
||||
<bibRefCitation id="22F4B7F1626C94E68FDB223A9EF80706" author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka 2011</bibRefCitation>
|
||||
). This uncommon species was first reported for New Brunswick by
|
||||
<bibRefCitation author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka (2011)</bibRefCitation>
|
||||
<bibRefCitation id="414E4000AEB073AA3E0972A9BDCC3E54" author="Majka, CG" journalOrPublisher="Journal of the Acadian Entomological Society" pageId="11" pageNumber="152" pagination="25 - 31" title="The Trogossitidae (Coleoptera) of Atlantic Canada." volume="7" year="2011">Majka (2011)</bibRefCitation>
|
||||
based on six specimens collected by W. McIntosh in Saint John during 1902. The record from Dionne Brook Protected Natural Area is the first recent record of this species from the province.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,125 +1,124 @@
|
|||
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<mods:extent id="BAAB244CE1FDA64053412B8D5DCE5305" unit="page">
|
||||
<mods:start id="164A7F654CB2B9CE99AEBCEBA59D838B">1</mods:start>
|
||||
<mods:end id="5DB4E937C92E76016D5EBEEFDD102BAE">85</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location>
|
||||
<mods:url>http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
|
||||
<mods:location id="4BC878E54084686FE4F41C1AE9B2B7F9">
|
||||
<mods:url id="7B5FB7A18081106FEA753AF128321089">http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification>journal article</mods:classification>
|
||||
<mods:identifier type="HNS-Pub">26064</mods:identifier>
|
||||
<mods:identifier type="ISSN">175-5326</mods:identifier>
|
||||
<mods:identifier type="ZooBank">A2D9C9ED-C0BA-4B5F-A330-C9AB7D625704</mods:identifier>
|
||||
<mods:classification id="B5553FCC64ABF385F2F4D39FDE257837">journal article</mods:classification>
|
||||
<mods:identifier id="497D120FE51919FF744A672C24681C69" type="HNS-Pub">26064</mods:identifier>
|
||||
<mods:identifier id="B713FF987F6FF349AF35388F78FA8B32" type="ISSN">175-5326</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172633" ID-GBIF-Taxon="100105357" ID-Zenodo-Dep="6172633" LSID="urn:lsid:plazi:treatment:E8C096AB0CC5473BCFA3B0FDA6D62124" httpUri="http://treatment.plazi.org/id/E8C096AB0CC5473BCFA3B0FDA6D62124" lastPageNumber="53" pageNumber="50">
|
||||
<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,552,1440,1460]" pageId="49" pageNumber="50" type="mainText">
|
||||
<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="jedi">
|
||||
<treatment id="E8C096AB0CC5473BCFA3B0FDA6D62124" ID-DOI="http://doi.org/10.5281/zenodo.6172633" ID-GBIF-Taxon="100105357" ID-Zenodo-Dep="6172633" LSID="urn:lsid:plazi:treatment:E8C096AB0CC5473BCFA3B0FDA6D62124" httpUri="http://treatment.plazi.org/id/E8C096AB0CC5473BCFA3B0FDA6D62124" lastPageNumber="53" pageNumber="50" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="CC19416BD50B848643B41843C533D956" type="nomenclature">
|
||||
<paragraph id="E60746759CA77B37179834792C1F52BA" box="[118,552,1440,1460]" pageId="49" pageNumber="50" type="mainText">
|
||||
<taxonomicName id="CA9828CF350FB8FED8FE46139D69EBAA" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="jedi">
|
||||
Tetramorium jedi
|
||||
<authority>Hita</authority>
|
||||
<authority id="B05419C11D30C4582171992F06A2ACFE">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="DF7888E57AAFD9586A8E12481D63C1B7" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,476,1467,1486]" pageId="49" pageNumber="50" type="mainText">(Figs. 29, 33, 44, 45, 105, 106, 107, 142)</paragraph>
|
||||
<subSubSection id="259BDCB01C8F783144F7B84DCD61F8D8" type="description">
|
||||
<paragraph id="A7B3D1CA211FD2F00BA682B6B7FA6C97" box="[118,476,1467,1486]" pageId="49" pageNumber="50" type="mainText">(Figs. 29, 33, 44, 45, 105, 106, 107, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1122,1524,1571][118,1120,118,194]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="mainText">
|
||||
<subSubSection id="9348F1089E52216772AE1656167AAB7C" type="materials_examined">
|
||||
<paragraph id="AABD155EA2592450375D701F54BE4159" box="[118,1122,1524,1571][118,1120,118,194]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="mainText">
|
||||
Holotype worker, MADAGASCAR, Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="ACA252663BFE45D46A76052F6BA3E835" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Binara, 9.1 km 233° SW Daraina, 13.26333 S, 49.60333 E, 800 m, rainforest, 3.XII.2003 (B.L. Fisher) (CASC: CASENT0043578). Paratypes, one worker with
|
||||
<pageBreakToken pageId="50" pageNumber="51" start="start">same</pageBreakToken>
|
||||
<pageBreakToken id="E7FE00DE9E4660C85DF4B5FC31EE6A1D" pageId="50" pageNumber="51" start="start">same</pageBreakToken>
|
||||
data as holotype (CASC: CASENT0043685); and nine workers from Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="08D65C672ACCD5F98D214C90112FBD61" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Binara, 9.1 km 233° SW Daraina, 13.26333 S, 49.60333 E, 725 m, rainforest, ex root mat, ground layer, collection code BLF09691, 4.XII.2003 (B.L. Fisher) (CASC: CASENT0077773, CASENT0077774, CASENT0077775).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,231,249]" pageId="50" pageNumber="51" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1122,259,334]" pageId="50" pageNumber="51" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="jedi">Tetramorium jedi</taxonomicName>
|
||||
<subSubSection id="B23EA5FD63FE2F77246C00F0165DC736" type="diagnosis">
|
||||
<paragraph id="BDDDAE8307734D2ED4BEC1FBA374E09A" box="[118,208,231,249]" pageId="50" pageNumber="51" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="5078FB2CC3508B93023ABB150D4988E4" box="[118,1122,259,334]" pageId="50" pageNumber="51" type="mainText">
|
||||
<taxonomicName id="99A65C177A28BBA0A8A8205C87ED2898" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="jedi">Tetramorium jedi</taxonomicName>
|
||||
is easily distinguished from the remainder of the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="F0AA26D965F29F1BA3503C786DC9D4FD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
species complex by the following characters: petiolar node in dorsal view distinctly longer than wide (DPeI 79-85); entire first gastral tergite conspicuously reticulate-punctate.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,371,390]" pageId="50" pageNumber="51" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1117,400,559]" pageId="50" pageNumber="51" type="mainText">HL 0.92-1.03 (0.99); HW 0.87-0.97 (0.93); SL 0.64-0.75 (0.71); EL 0.17-0.20 (0.18); PH 0.45-0.54 (0.49); PW 0.65-0.72 (0.70); WL 1.14-1.30 (1.24); PSL 0.34-0.41 (0.37); PTL 0.29-0.34 (0.32); PTH 0.34-0.40 (0.38); PTW 0.24-0.29 (0.27); PPL 0.30-0.34 (0.31); PPH 0.34-0.41 (0.39); PPW 0.36-0.40 (0.38); CI 92-95 (93); SI 74-79 (77); OI 18-22 (20); DMI 54-58 (56); LMI 38-42 (39); PSLI 35-40 (37); PeNI 37-41 (38); LPeI 80-88 (84); DPeI 79-85 (83); PpNI 53-57 (54); LPpI 76-87 (81); DPpI 116-126 (121); PPI 138-148 (142) (12 measured).</paragraph>
|
||||
<paragraph box="[118,1121,568,1234]" pageId="50" pageNumber="51" type="mainText">Head distinctly longer than wider (CI 92-95); posterior head margin moderately concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly to moderately developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short, not reaching posterior head margin (SI 74-79). Eyes small to moderate in size (OI 18-22). Mesosomal outline in profile weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-42). Propodeal spines long, spinose, and acute (PSLI 35-40); propodeal lobes short, triangular and acute. Petiolar node in profile rectangular nodiform, around 1.1 to 1.3 times higher than long (LPeI 80-88), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum straight; node in dorsal view around 1.2 to 1.3 times longer than wide (DPeI 79-85). Postpetiole in profile subglobular, approximately 1.1 to 1.3 times higher than long (LPpI 76-87); in dorsal view around 1.1 to 1.3 times wider than long (DPpI 116-126). Postpetiole in profile appearing as voluminous as petiolar node, in dorsal view approximately 1.3 to 1.5 times wider than petiolar node (PPI 138-148). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose, with three to five rugae/ rugulae, median ruga well-developed, lateral rugae/rugulae often weak and broken; cephalic dorsum between frontal carinae with 8 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, few rugae interrupted or with cross-meshes; lateral and ventral head longitudinally rugose to reticulate-rugose. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae with weak, superficial punctate ground sculpture only, otherwise unsculptured. Waist segments strongly longitudinally rugose to reticulate-rugose. Ground sculpture on head, mesosoma, and waist segments generally faint. First gastral tergite covered completely with very conspicuous punctate sculpture, appearing matt. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. First gastral tergite without appressed pubescence. Anterior edges of antennal scapes with subdecumbent to erect standing hairs. Body a uniform brown to dark brown colour.</paragraph>
|
||||
<subSubSection id="6A6AE1EC36D9D1ACD8FC4929EF126703" type="description">
|
||||
<paragraph id="D21D0241DA6623C625CA594F4107199A" box="[118,226,371,390]" pageId="50" pageNumber="51" type="mainText">Description</paragraph>
|
||||
<paragraph id="3839D5C76E4C48EBC78691255C148844" box="[118,1117,400,559]" pageId="50" pageNumber="51" type="mainText">HL 0.92-1.03 (0.99); HW 0.87-0.97 (0.93); SL 0.64-0.75 (0.71); EL 0.17-0.20 (0.18); PH 0.45-0.54 (0.49); PW 0.65-0.72 (0.70); WL 1.14-1.30 (1.24); PSL 0.34-0.41 (0.37); PTL 0.29-0.34 (0.32); PTH 0.34-0.40 (0.38); PTW 0.24-0.29 (0.27); PPL 0.30-0.34 (0.31); PPH 0.34-0.41 (0.39); PPW 0.36-0.40 (0.38); CI 92-95 (93); SI 74-79 (77); OI 18-22 (20); DMI 54-58 (56); LMI 38-42 (39); PSLI 35-40 (37); PeNI 37-41 (38); LPeI 80-88 (84); DPeI 79-85 (83); PpNI 53-57 (54); LPpI 76-87 (81); DPpI 116-126 (121); PPI 138-148 (142) (12 measured).</paragraph>
|
||||
<paragraph id="5F4C6BD946C7C7F915389FFE7B138C32" box="[118,1121,568,1234]" pageId="50" pageNumber="51" type="mainText">Head distinctly longer than wider (CI 92-95); posterior head margin moderately concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly to moderately developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short, not reaching posterior head margin (SI 74-79). Eyes small to moderate in size (OI 18-22). Mesosomal outline in profile weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-42). Propodeal spines long, spinose, and acute (PSLI 35-40); propodeal lobes short, triangular and acute. Petiolar node in profile rectangular nodiform, around 1.1 to 1.3 times higher than long (LPeI 80-88), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum straight; node in dorsal view around 1.2 to 1.3 times longer than wide (DPeI 79-85). Postpetiole in profile subglobular, approximately 1.1 to 1.3 times higher than long (LPpI 76-87); in dorsal view around 1.1 to 1.3 times wider than long (DPpI 116-126). Postpetiole in profile appearing as voluminous as petiolar node, in dorsal view approximately 1.3 to 1.5 times wider than petiolar node (PPI 138-148). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose, with three to five rugae/ rugulae, median ruga well-developed, lateral rugae/rugulae often weak and broken; cephalic dorsum between frontal carinae with 8 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, few rugae interrupted or with cross-meshes; lateral and ventral head longitudinally rugose to reticulate-rugose. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae with weak, superficial punctate ground sculpture only, otherwise unsculptured. Waist segments strongly longitudinally rugose to reticulate-rugose. Ground sculpture on head, mesosoma, and waist segments generally faint. First gastral tergite covered completely with very conspicuous punctate sculpture, appearing matt. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. First gastral tergite without appressed pubescence. Anterior edges of antennal scapes with subdecumbent to erect standing hairs. Body a uniform brown to dark brown colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,1271,1290]" pageId="50" pageNumber="51" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1122,1300,1403]" pageId="50" pageNumber="51" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="jedi">Tetramorium jedi</taxonomicName>
|
||||
<subSubSection id="F879FD093B13849F3440199EFD216855" type="discussion">
|
||||
<paragraph id="08D25BE30D587FABCE68D57987D60838" box="[118,170,1271,1290]" pageId="50" pageNumber="51" type="mainText">Notes</paragraph>
|
||||
<paragraph id="B3F55E303645F80F6DC6C446B8B40B8E" box="[118,1122,1300,1403]" pageId="50" pageNumber="51" type="mainText">
|
||||
<taxonomicName id="67AF6E886BC758BA4A2197B64ACC27C8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="jedi">Tetramorium jedi</taxonomicName>
|
||||
is mainly distributed in the northeast of the island of Madagascar from Anjanaharibe north to Binara. Surprisingly, the species is also known from Ambalagoavy, which is located much further south. The available material was sampled from lowland rainforests at elevations between 240 to 800 m. The species appears to be ground-active.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,1412,1459]" pageId="50" pageNumber="51" type="mainText">
|
||||
<paragraph id="6F8A9EF4641CA9CF9CB5BAC33A87F74C" box="[118,1121,1412,1459]" pageId="50" pageNumber="51" type="mainText">
|
||||
As mentioned in the diagnosis above,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
<taxonomicName id="11384681C6895A733ECF307AC82FE933" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="jedi">T. jedi</taxonomicName>
|
||||
is unique in the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
<taxonomicName id="1BA1693BFC9DA7DBC3A1113D5E8ABE23" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="tortuosum">T. tortuosum</taxonomicName>
|
||||
group because of the pronounced and very distinct sculpture on the first gastral tergite, which covers the whole tergite.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,1496,1515]" pageId="50" pageNumber="51" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,1120,1525,1572]" pageId="50" pageNumber="51" type="mainText">
|
||||
<subSubSection id="F7B0127BBD94876AAC1E786E6C085FB2" type="etymology">
|
||||
<paragraph id="8EC0D81DF415C5A43E31FD7FF28E9682" box="[118,219,1496,1515]" pageId="50" pageNumber="51" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="AF6E4935064C7B951CF4C6D9AF7BBC1D" box="[118,1120,1525,1572]" pageId="50" pageNumber="51" type="mainText">
|
||||
This
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="BF843AA7FA854EA104856D638B881758" rank="species">new species</taxonomicNameLabel>
|
||||
is named after the fictional, noble, and wise guardians of peace from the "Star Wars" universe created by George Lucas. The species epithet is an arbitrary combination of letters.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="multiple">
|
||||
<caption box="[118,1122,1541,1587]" pageId="51" pageNumber="52" type="caption">
|
||||
<paragraph box="[118,1122,1541,1587]" pageId="51" pageNumber="52" type="caption">
|
||||
<pageBreakToken pageId="51" pageNumber="52" start="start">FIGURES</pageBreakToken>
|
||||
<subSubSection id="C4D5D2D44DA10CCF4212D9D12A27CAB3" type="multiple">
|
||||
<caption id="5C9EE94F5E3AE7061563CF77EC434944" box="[118,1122,1541,1587]" pageId="51" pageNumber="52" type="caption">
|
||||
<paragraph id="48BF710E2812C74A377107530427B6B9" box="[118,1122,1541,1587]" pageId="51" pageNumber="52" type="caption">
|
||||
<pageBreakToken id="9317B2BF93A9E7EE2E0FC3F0FA144D5E" pageId="51" pageNumber="52" start="start">FIGURES</pageBreakToken>
|
||||
105-107.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="jedi">T. jedi,</taxonomicName>
|
||||
<taxonomicName id="464DC94386AE2389BA3EB7C1936B51E0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="jedi">T. jedi,</taxonomicName>
|
||||
holotype (CASENT0043578). 105. Body in profile. 106. Body in dorsal view. 107. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,118,137]" pageId="52" pageNumber="53" type="mainText">
|
||||
<pageBreakToken pageId="52" pageNumber="53" start="start">Material</pageBreakToken>
|
||||
<subSubSection id="FD94B67B36BF0161AEF776103C7060B4" type="materials_examined">
|
||||
<paragraph id="70DE005E2CBD971E933E39429E2E3740" box="[118,295,118,137]" pageId="52" pageNumber="53" type="mainText">
|
||||
<pageBreakToken id="9313755C33D2695751209D72494C5AAD" pageId="52" pageNumber="53" start="start">Material</pageBreakToken>
|
||||
examined
|
||||
</paragraph>
|
||||
<paragraph box="[118,1120,147,517]" pageId="52" pageNumber="53" type="mainText">
|
||||
<paragraph id="EF1687C24847BED834A8647FFDC07E1D" box="[118,1120,147,517]" pageId="52" pageNumber="53" type="mainText">
|
||||
MADAGASCAR: Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="03123D929686222D8866CDFE95C629F9" originalValue="Forêt">Foret</normalizedToken>
|
||||
Ambanitaza, 26.1 km 347° Antalaha, 14.67933 S, 50.18367 E, 240 m, rainforest, 26.XI.2004 (B.L. Fisher); Antsiranana,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="8313BA2D95A428C17070582D6C10B2B3" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Binara, 9.1 km 233° SW Daraina, 13.26333 S, 49.60333 E, 725-800 m, rainforest, 3.-4.XII.2003 (B.L. Fisher); Antsiranana, Makirovana forest, 14.17066 S, 49.95409 E, 415 m, rainforest, 29.IV.2011 (B.L. Fisher et al.); Antsiranana, Makirovana forest, 14.16666 S, 49.95 E, 715 m, rainforest, 2.V.2011 (B.L. Fisher et al.); Antsiranana, Makirovana forest, 14.17066 S, 49.95409 E, 225 m, rainforest, 4.V.2011 (B.L. Fisher et al.); Antsiranana, Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, 14.435 S, 49.76 E, 775 m, rainforest, 15.-18.XI.2003 (B.L. Fisher et al.); Antsiranana, Parc National de Marojejy, Manantenina River, 28.0 km 38° NE Andapa, 8.2 km 333° NNW Manantenina, 14.4367 S, 49.775 E, 450 m, rainforest, 12.-25.XI.2003 (B.L. Fisher et al.); Fianarantsoa, Foret d'Ambalagoavy Nord, Ikongo, Ambatombe, 21.8275 S, 47.33889 E, 625 m, 1.XII.2000 (R. Harin'Hala & M.E. Irwin); Toamasina, Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo, 15.18833 S, 49.615 E, 470 m, rainforest, 8.-12.III.2003 (B.L. Fisher & C. Griswold).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,53 +1,53 @@
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<treatment id="E8F61829A338FCCB6BD13BBAF35B47D4" ID-GBIF-Taxon="152033765" LSID="urn:lsid:plazi:treatment:E8F61829A338FCCB6BD13BBAF35B47D4" httpUri="http://treatment.plazi.org/id/E8F61829A338FCCB6BD13BBAF35B47D4" lastPageNumber="150" pageId="9" pageNumber="150" scope_family="Melyridae" scope_order="Coleoptera" scope_subFamily="Malachiinae">
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|
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<taxonomicName id="E11501F3D97BC0B7802596D2508C045B" LSID="http://species-id.net/wiki/Attalus_morulus" authority="(LeConte, 1852) **" class="Insecta" family="Melyridae" genus="Attalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Attalus morulus" order="Coleoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="morulus">Attalus morulus (LeConte, 1852)**</taxonomicName>
|
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<taxonomicName id="E11501F3D97BC0B7802596D2508C045B" LSID="http://species-id.net/wiki/Attalus_morulus" authority="(LeConte, 1852) **" genus="Attalus" lsidName="Attalus morulus" pageId="9" pageNumber="150" rank="species" species="morulus">Attalus morulus (LeConte, 1852)**</taxonomicName>
|
||||
Map 9
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
@ -71,7 +71,7 @@ New Brunswick, Northumberland Co., Blueberry Rd., off Hwy 8,
|
|||
<caption id="FD155C13A09C07B0560494DE572F848D" pageId="9" pageNumber="150">
|
||||
<paragraph id="9BFC0ACD7ABB1F1B82C2935F2CD2F8E6" pageId="9" pageNumber="150">
|
||||
Map 9. Collection localities in New Brunswick, Canada of
|
||||
<taxonomicName id="F6FA79376E5E558B4482C7C105F5644D" class="Insecta" family="Melyridae" genus="Attalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Attalus morulus" order="Coleoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="morulus">Attalus morulus</taxonomicName>
|
||||
<taxonomicName id="F6FA79376E5E558B4482C7C105F5644D" genus="Attalus" lsidName="Attalus morulus" pageId="9" pageNumber="150" rank="species" species="morulus">Attalus morulus</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</caption>
|
||||
|
|
@ -79,7 +79,7 @@ Map 9. Collection localities in New Brunswick, Canada of
|
|||
<subSubSection id="41B7933BC1A007ECE8856919E1A84E46" pageId="9" pageNumber="150" type="collection and habitat data">
|
||||
<paragraph id="3975D9FAE11A22A6E01A3BB8E02E86D7" pageId="9" pageNumber="150">Collection and habitat data.</paragraph>
|
||||
<paragraph id="D3945738ABB6485B1900B303BC44B850" pageId="9" pageNumber="150">
|
||||
<taxonomicName id="E11590E18EB065F9754081E4671E3CE4" class="Insecta" family="Melyridae" genus="Attalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Attalus morulus" order="Coleoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="morulus">Attalus morulus</taxonomicName>
|
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<taxonomicName id="E11590E18EB065F9754081E4671E3CE4" genus="Attalus" lsidName="Attalus morulus" pageId="9" pageNumber="150" rank="species" species="morulus">Attalus morulus</taxonomicName>
|
||||
were captured in Lindgren funnel traps deployed in an old-growth red pine forest. One individual was collected from foliage of jack pine (
|
||||
<taxonomicName id="CC07A9A9E6CAB124277766CCDC17D520" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus banksiana" order="Pinales" pageId="9" pageNumber="150" phylum="Tracheophyta" rank="species" species="banksiana">Pinus banksiana</taxonomicName>
|
||||
Lamb.) in a jack pine forest. Adults were captured during July and August.
|
||||
|
|
@ -91,7 +91,7 @@ Lamb.) in a jack pine forest. Adults were captured during July and August.
|
|||
BC, QC, NB (
|
||||
<bibRefCitation id="E1732CFBEA5D28A802E1D0C9492C899A" pageId="9" pageNumber="150">Bright 1991</bibRefCitation>
|
||||
).
|
||||
<taxonomicName id="B0C09C8FD9A65C2725F5A624D5279F57" class="Insecta" family="Melyridae" genus="Attalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Attalus morulus" order="Coleoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="morulus">Attalus morulus</taxonomicName>
|
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<taxonomicName id="B0C09C8FD9A65C2725F5A624D5279F57" genus="Attalus" lsidName="Attalus morulus" pageId="9" pageNumber="150" rank="species" species="morulus">Attalus morulus</taxonomicName>
|
||||
was previously reported from Connecticut and New York, in northeastern USA (
|
||||
<bibRefCitation id="8B07B509104D7D9B785C3A1AC9B84E23" pageId="9" pageNumber="150">Downie and Arnett 1996</bibRefCitation>
|
||||
).
|
||||
|
|
|
|||
|
|
@ -1,140 +1,139 @@
|
|||
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<mods:roleTerm id="DFE682D8F9A2CF1FA6D01B2AB8124998">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart>B. L. Fisher</mods:namePart>
|
||||
<mods:namePart id="9A2896A7DF10020387F95FFEEF01B391">B. L. Fisher</mods:namePart>
|
||||
</mods:name>
|
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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<mods:title id="441654C62C3D960AD5A900B5C8FF2F85">Zootaxa</mods:title>
|
||||
</mods:titleInfo>
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||||
<mods:part>
|
||||
<mods:date>2012</mods:date>
|
||||
<mods:detail type="volume">
|
||||
<mods:number>3592</mods:number>
|
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<mods:part id="A52C0E27977557F0D3AD0A29D3601C51">
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||||
<mods:date id="4FB906BDC8BE5AB2FF2B60071DEE3093">2012</mods:date>
|
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<mods:detail id="F23DDAE51DF5C3AF95D3836D8E21DAF7" type="volume">
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||||
<mods:number id="FF2C8D2AE20E50DEA45743D8561932B1">3592</mods:number>
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||||
</mods:detail>
|
||||
<mods:extent unit="page">
|
||||
<mods:start>1</mods:start>
|
||||
<mods:end>85</mods:end>
|
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<mods:extent id="E06AEDF4B9AD3071D175733653F74848" unit="page">
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<mods:start id="964A03D515BFADF8EC1FC31035484C4B">1</mods:start>
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<mods:end id="357B3B79CCFFC2612DCACEE8461BEB57">85</mods:end>
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<mods:location>
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<mods:url>http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
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<mods:location id="ECB6525B54BF3A426B4908075EC808FA">
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<mods:url id="D80563BD1088543F9FC4AF66166CD573">http://www.mapress.com/zootaxa/2012/f/zt03592p085.pdf</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="HNS-Pub">26064</mods:identifier>
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<mods:identifier type="ISSN">175-5326</mods:identifier>
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<mods:identifier type="ZooBank">A2D9C9ED-C0BA-4B5F-A330-C9AB7D625704</mods:identifier>
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<mods:classification id="ED063B90D73F6F7E56AF4A703BAC1457">journal article</mods:classification>
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<mods:identifier id="B87C17361C01A07B7AD107BC98494C8D" type="HNS-Pub">26064</mods:identifier>
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||||
<mods:identifier id="7B0648661FD3C8B3F9C559082F818439" type="ISSN">175-5326</mods:identifier>
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</mods:mods>
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||||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172623" ID-GBIF-Taxon="100105353" ID-Zenodo-Dep="6172623" LSID="urn:lsid:plazi:treatment:ECA618B7D6FFE2F06F34B9D122BE3A1D" httpUri="http://treatment.plazi.org/id/ECA618B7D6FFE2F06F34B9D122BE3A1D" lastPageNumber="42" pageNumber="40">
|
||||
<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,628,231,251]" pageId="39" pageNumber="40" type="mainText">
|
||||
<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="isoelectrum">
|
||||
<treatment id="ECA618B7D6FFE2F06F34B9D122BE3A1D" ID-DOI="http://doi.org/10.5281/zenodo.6172623" ID-GBIF-Taxon="100105353" ID-Zenodo-Dep="6172623" LSID="urn:lsid:plazi:treatment:ECA618B7D6FFE2F06F34B9D122BE3A1D" httpUri="http://treatment.plazi.org/id/ECA618B7D6FFE2F06F34B9D122BE3A1D" lastPageNumber="42" pageNumber="40" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="F3A654B0C6CF4B71D2B017319FF5AD2B" type="nomenclature">
|
||||
<paragraph id="F4882CAEDDA935E6AAFED0115B8DC1EB" box="[118,628,231,251]" pageId="39" pageNumber="40" type="mainText">
|
||||
<taxonomicName id="D207E0F538A015C1A974FAD258118B28" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="isoelectrum">
|
||||
Tetramorium isoelectrum
|
||||
<authority>Hita</authority>
|
||||
<authority id="430ACF82C097BF04421F988D083687C3">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="64CEA9A53AC90E03B871F6F90CAE89BA" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,410,258,277]" pageId="39" pageNumber="40" type="mainText">(Figs. 59, 64, 65, 93, 94, 95, 141)</paragraph>
|
||||
<subSubSection id="1BEEB35BCFA60E079A690B4AC6775E89" type="description">
|
||||
<paragraph id="98C7CFD6970A1C48F45D8C9268A8FA63" box="[118,410,258,277]" pageId="39" pageNumber="40" type="mainText">(Figs. 59, 64, 65, 93, 94, 95, 141)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1122,314,418]" pageId="39" pageNumber="40" type="mainText">Holotype worker, MADAGASCAR, Antsiranana, Betaolana Forest, along Bekona River, 14.52996 S, 49.44039 E, 880 m, rainforest, on low vegetation, collection code BLF22647, 5.III.2009 (B.L. Fisher et al.) (CASC: CASENT0152199). Paratype, one worker with same data as holotype except sampled from ground and collection code BLF22648 (CASC: CASENT0152192).</paragraph>
|
||||
<subSubSection id="5FBE297A25399E0E428CE3467B03A95B" type="materials_examined">
|
||||
<paragraph id="E3F49E731BF815F218521DAFA0DC5671" box="[118,1122,314,418]" pageId="39" pageNumber="40" type="mainText">Holotype worker, MADAGASCAR, Antsiranana, Betaolana Forest, along Bekona River, 14.52996 S, 49.44039 E, 880 m, rainforest, on low vegetation, collection code BLF22647, 5.III.2009 (B.L. Fisher et al.) (CASC: CASENT0152199). Paratype, one worker with same data as holotype except sampled from ground and collection code BLF22648 (CASC: CASENT0152192).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,455,473]" pageId="39" pageNumber="40" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1120,483,586]" pageId="39" pageNumber="40" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="isoelectrum">Tetramorium isoelectrum</taxonomicName>
|
||||
<subSubSection id="D7F7B4528B9AB7508E5375B467B016C6" type="diagnosis">
|
||||
<paragraph id="5B29BE06681375FCEC08D6883F229766" box="[118,208,455,473]" pageId="39" pageNumber="40" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="ADBF380D02F4FB501A22DE6699C55467" box="[118,1120,483,586]" pageId="39" pageNumber="40" type="mainText">
|
||||
<taxonomicName id="D287D81746EA93CC0C8AE69F6C0CE731" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="isoelectrum">Tetramorium isoelectrum</taxonomicName>
|
||||
differs from the remainder of the species group by the following character combination: propodeal spines extremely long (PSLI 58-63); petiolar node around 1.1 to 1.3 times higher than long (LPeI 79-86); posterodorsal corner of petiole not strongly protruding posteriorly; mandibles with distinct longitudinal sculpture; body dark brown to black in colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,623,642]" pageId="39" pageNumber="40" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1121,652,783]" pageId="39" pageNumber="40" type="mainText">HL 1.09-1.19 (1.16); HW 1.03-1.13 (1.10); SL 0.84-0.94 (0.91); EL 0.19-0.21 (0.20); PH 0.53-0.68 (0.61); PW 0.74-0.84 (0.81); WL 1.36-1.57 (1.50); PSL 0.63-0.75 (0.71); PTL 0.35-0.38 (0.36); PTH 0.44-0.46 (0.45); PTW 0.32-0.34 (0.33); PPL 0.38-0.40 (0.39); PPH 0.44-0.46 (0.45); PPW 0.39-0.41 (0.40); CI 93-96 (95); SI 81-84 (82); OI 17-19 (18); DMI 53-55 (54); LMI 38-45 (40); PSLI 58-63 (61); PeNI 38-46 (41); LPeI 79-86 (81); DPeI 87-93 (90); PpNI 47-54 (49); LPpI 83-91 (87); DPpI 101-105 (103); PPI 118-127 (123) (seven measured).</paragraph>
|
||||
<paragraph box="[118,1122,792,1430]" pageId="39" pageNumber="40" type="mainText">Head longer than wide (CI 93-96). Posterior head margin weakly to moderately concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short to moderately long, not reaching posterior head margin (SI 81-84). Eyes small (OI 17-19). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-45). Propodeal spines extremely long, spinose, and acute (PSLI 58-63); propodeal lobes short, triangular, and blunt. Petiolar node in profile rectangular nodiform with comparatively rounded angles, around 1.1 to 1.3 times higher than long (LPeI 79-86), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum straight to weakly convex; node in dorsal view around 1.1 times longer than wide (DPeI 87-93). Postpetiole in profile subglobular, approximately 1.1 to 1.2 times higher than long (LPpI 83-91); in dorsal view feebly wider than long (DPpI 101-105). Postpetiole in profile appearing a bit more voluminous than petiolar node, in dorsal view approximately 1.2 to 1.3 times wider than petiolar node (PPI 118-127). Mandibles distinctly longitudinally rugose; clypeus with four to six longitudinal rugulae, median rugula better developed than lateral rugulae; cephalic dorsum between frontal carinae with eight to ten longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae never with cross-meshes; lateral and ventral head longitudinally rugose with very few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose, rugae laterally more irregularly arranged. Forecoxae mainly reticulate-punctate with superficial, superimposed rugulae. Waist segments with reticulate-rugose to longitudinally rugose sculpture. Ground sculpture everywhere on body faint to absent. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. First gastral tergite without appressed pubescence. Anterior edges of antennal scapes with subdecumbent to erect standing hairs. Body of uniform very dark brown to black colour.</paragraph>
|
||||
<subSubSection id="D5F7ABBF9B5F3E03644FD9EA55DEC0C2" type="description">
|
||||
<paragraph id="1B4844E2CA144797773B8085B627A490" box="[118,226,623,642]" pageId="39" pageNumber="40" type="mainText">Description</paragraph>
|
||||
<paragraph id="7C0794A1CEAB9759284E9E7C0748A83C" box="[118,1121,652,783]" pageId="39" pageNumber="40" type="mainText">HL 1.09-1.19 (1.16); HW 1.03-1.13 (1.10); SL 0.84-0.94 (0.91); EL 0.19-0.21 (0.20); PH 0.53-0.68 (0.61); PW 0.74-0.84 (0.81); WL 1.36-1.57 (1.50); PSL 0.63-0.75 (0.71); PTL 0.35-0.38 (0.36); PTH 0.44-0.46 (0.45); PTW 0.32-0.34 (0.33); PPL 0.38-0.40 (0.39); PPH 0.44-0.46 (0.45); PPW 0.39-0.41 (0.40); CI 93-96 (95); SI 81-84 (82); OI 17-19 (18); DMI 53-55 (54); LMI 38-45 (40); PSLI 58-63 (61); PeNI 38-46 (41); LPeI 79-86 (81); DPeI 87-93 (90); PpNI 47-54 (49); LPpI 83-91 (87); DPpI 101-105 (103); PPI 118-127 (123) (seven measured).</paragraph>
|
||||
<paragraph id="D869F45419A6B47DB15B238E8C457068" box="[118,1122,792,1430]" pageId="39" pageNumber="40" type="mainText">Head longer than wide (CI 93-96). Posterior head margin weakly to moderately concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes comparatively short to moderately long, not reaching posterior head margin (SI 81-84). Eyes small (OI 17-19). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-45). Propodeal spines extremely long, spinose, and acute (PSLI 58-63); propodeal lobes short, triangular, and blunt. Petiolar node in profile rectangular nodiform with comparatively rounded angles, around 1.1 to 1.3 times higher than long (LPeI 79-86), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum straight to weakly convex; node in dorsal view around 1.1 times longer than wide (DPeI 87-93). Postpetiole in profile subglobular, approximately 1.1 to 1.2 times higher than long (LPpI 83-91); in dorsal view feebly wider than long (DPpI 101-105). Postpetiole in profile appearing a bit more voluminous than petiolar node, in dorsal view approximately 1.2 to 1.3 times wider than petiolar node (PPI 118-127). Mandibles distinctly longitudinally rugose; clypeus with four to six longitudinal rugulae, median rugula better developed than lateral rugulae; cephalic dorsum between frontal carinae with eight to ten longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae never with cross-meshes; lateral and ventral head longitudinally rugose with very few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose, rugae laterally more irregularly arranged. Forecoxae mainly reticulate-punctate with superficial, superimposed rugulae. Waist segments with reticulate-rugose to longitudinally rugose sculpture. Ground sculpture everywhere on body faint to absent. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. First gastral tergite without appressed pubescence. Anterior edges of antennal scapes with subdecumbent to erect standing hairs. Body of uniform very dark brown to black colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,1467,1485]" pageId="39" pageNumber="40" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1120,1496,1543]" pageId="39" pageNumber="40" type="mainText">
|
||||
<subSubSection id="20456C39C1F0B1E32435646AFD2E3522" type="discussion">
|
||||
<paragraph id="431540E3A6F47E82D572000605D07780" box="[118,170,1467,1485]" pageId="39" pageNumber="40" type="mainText">Notes</paragraph>
|
||||
<paragraph id="57EC8E38F5A6B8D9A0BAAABBBEA53DDB" box="[118,1120,1496,1543]" pageId="39" pageNumber="40" type="mainText">
|
||||
Currently, this
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="593679D8BE596074002255420CE83BEF" rank="species">new species</taxonomicNameLabel>
|
||||
is only known from seven specimens from Betaolana, Befingotra, and Marojejy. All three localities are rainforests found in the north-eastern part of Madagascar at elevations of 450 to 880 m.
|
||||
</paragraph>
|
||||
<caption box="[118,1122,1548,1590]" pageId="40" pageNumber="41" type="caption">
|
||||
<paragraph box="[118,1122,1548,1590]" pageId="40" pageNumber="41" type="caption">
|
||||
<pageBreakToken pageId="40" pageNumber="41" start="start">FIGURES</pageBreakToken>
|
||||
<caption id="8B6CC8286FCE040F70F048977488B7BA" box="[118,1122,1548,1590]" pageId="40" pageNumber="41" type="caption">
|
||||
<paragraph id="9E365F8A83D6D8CE15776A0EEC8DED0F" box="[118,1122,1548,1590]" pageId="40" pageNumber="41" type="caption">
|
||||
<pageBreakToken id="7AA3E96AAF68670B52EC59358113927F" pageId="40" pageNumber="41" start="start">FIGURES</pageBreakToken>
|
||||
93-95.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
<taxonomicName id="87D45B61404D5DBE26664374AE10EC4C" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
holotype (CASENT0152199). 93. Body in profile. 94. Body in dorsal view. 95. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,1120,118,362]" pageId="41" pageNumber="42" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">
|
||||
<pageBreakToken pageId="41" pageNumber="42" start="start">Tetramorium</pageBreakToken>
|
||||
<paragraph id="D3A958AAB0FA24F48FD8A15872CB2600" box="[118,1120,118,362]" pageId="41" pageNumber="42" type="mainText">
|
||||
<taxonomicName id="B9E985A8E0F6E8CA189D9A0971509C59" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">
|
||||
<pageBreakToken id="03342CA7A60F201B16B0FF9D28FF4650" pageId="41" pageNumber="42" start="start">Tetramorium</pageBreakToken>
|
||||
isoelectrum
|
||||
</taxonomicName>
|
||||
is morphologically very close to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="42E2C982274928C9B08E60A7D0994568" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="elf">T. elf,</taxonomicName>
|
||||
<taxonomicName id="E52B4ABDF2B22C1ADA2B9C1E214F2CF0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="elf">T. elf,</taxonomicName>
|
||||
as outlined in the descriptions of the latter two species.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="elf">Tetramorium elf,</taxonomicName>
|
||||
<taxonomicName id="9E6AF0A56EC4C151CCC5A82985756728" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="elf">Tetramorium elf,</taxonomicName>
|
||||
however, is yellow in colour and has mandibles that are unsculptured, smooth, and shining, whereas
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="4384F08F62247D1880113790FAA9E535" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
is very dark brown and has mandibles that are very clearly sculptured.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="E0EC669FF57FA60356FD0A896A40A7D2" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="C89F754CAFAFD61DF09087A6AE42ABC1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
are more difficult to distinguish. Both species are very close in general gestalt and easy to confuse at first glance. Nevertheless,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="C5B167E46E55F68D08DC1AA5746CD8BC" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
has a longer head (CI 93-96), longer antennal scapes (SI 81-84), and a longer and lower petiolar node (LPeI 77-86; DPeI 87-97) than
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
<taxonomicName id="BFA332FFC552A93809391195FAABDCF1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="electrum">T. electrum</taxonomicName>
|
||||
(CI 98-104; SI 67-74; LPeI 64-74; DPeI 100-114). Furthermore, due to its extremely long propodeal spines (PSLI 58-63),
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="DE5DDE459375FE7A8874639C44396C45" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
is not likely to be misidentified with the remaining species of the complex.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,399,418]" pageId="41" pageNumber="42" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,1121,428,503]" pageId="41" pageNumber="42" type="mainText">
|
||||
<subSubSection id="1C878D9537A131DFF201090B0DDBFBE4" type="etymology">
|
||||
<paragraph id="F76F96ECBE15C48077FA61BB20C82568" box="[118,219,399,418]" pageId="41" pageNumber="42" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="DCEF164E9A440A50C69002891F993ABE" box="[118,1121,428,503]" pageId="41" pageNumber="42" type="mainText">
|
||||
The name of the
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="0FB0B027C23241D85FFF791C2C48244E" rank="species">new species</taxonomicNameLabel>
|
||||
is composed of
|
||||
<normalizedToken originalValue="“iso”">"iso"</normalizedToken>
|
||||
<normalizedToken id="64DAF259DD718EAD4BF62EEACD3EC72E" originalValue="“iso”">"iso"</normalizedToken>
|
||||
, which is Old Greek meaning
|
||||
<normalizedToken originalValue="“equal”">"equal"</normalizedToken>
|
||||
<normalizedToken id="519C711B652F98BB1F46E223556AA1A8" originalValue="“equal”">"equal"</normalizedToken>
|
||||
, and
|
||||
<normalizedToken originalValue="“electrum”">"electrum"</normalizedToken>
|
||||
<normalizedToken id="B8BD83174345CA9A60AAB06EA671CFFA" originalValue="“electrum”">"electrum"</normalizedToken>
|
||||
, which is the species epithet of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="electrum">T. electrum.</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
<taxonomicName id="BB2C33D92476C4E0D841BC06EF9C1BA5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="electrum">T. electrum.</taxonomicName>
|
||||
<taxonomicName id="F8222F1BE922CE98C4CE8181A56F7AD1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="electrum">Tetramorium electrum</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="A8101FC87CBDF0024D7F586CE912FD11" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
are very similar, and the
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="C4A0F92E1E2E3266D5E70D8C80549561" rank="species">new species</taxonomicNameLabel>
|
||||
name reflects this. The species epithet is a noun in apposition, and thus invariant.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,540,559]" pageId="41" pageNumber="42" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1121,568,700]" pageId="41" pageNumber="42" type="mainText">
|
||||
<subSubSection id="0841D15831C7EEB7D81AABAC921386F3" type="materials_examined">
|
||||
<paragraph id="41EF9FB7D440C34BB5FD92A8F4178338" box="[118,295,540,559]" pageId="41" pageNumber="42" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="BBDF787ADCB78152BB0AFD32611DBC3C" box="[118,1121,568,700]" pageId="41" pageNumber="42" type="mainText">
|
||||
MADAGASCAR: Antsiranana, 6.5 km SSW Befingotra,
|
||||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||||
<normalizedToken id="EA294B3C1DA0724ABC7162D8C57E55E5" originalValue="Rés">Res</normalizedToken>
|
||||
. Anjanaharibe-Sud, 14.75 S, 49.5 E, 875 m, rainforest, 20.IX.1994 (B.L. Fisher); Antsiranana, Betaolana Forest, along Bekona River, 14.52996 S, 49.44039 E, 880 m, rainforest, 5.III.2009 (B.L. Fisher et al.); Antsiranana, Parc National de Marojejy, Manantenina River, 28.0 km 38° NE Andapa, 8.2 km 333° NNW Manantenina, 14.43667 S, 49.775 E, 450 m, rainforest, 12.-15.XI.2003 (B.L. Fisher et al.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,144 +1,143 @@
|
|||
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172655" ID-GBIF-Taxon="100105370" ID-Zenodo-Dep="6172655" LSID="urn:lsid:plazi:treatment:F009EBF7F82DEB40ABFB890DCEA41272" httpUri="http://treatment.plazi.org/id/F009EBF7F82DEB40ABFB890DCEA41272" lastPageNumber="71" pageNumber="69">
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|
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|
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<taxonomicName authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="latreillei">
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|
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<treatment id="F009EBF7F82DEB40ABFB890DCEA41272" ID-DOI="http://doi.org/10.5281/zenodo.6172655" ID-GBIF-Taxon="100105370" ID-Zenodo-Dep="6172655" LSID="urn:lsid:plazi:treatment:F009EBF7F82DEB40ABFB890DCEA41272" httpUri="http://treatment.plazi.org/id/F009EBF7F82DEB40ABFB890DCEA41272" lastPageNumber="71" pageNumber="69" scope_family="Formicidae" scope_order="Hymenoptera">
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|
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|
||||
<taxonomicName id="8C23D7EE01B5157DE8C8702E40412B80" ID-CoL="55PPF" authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
<pageBreakToken id="0D6D90F544B8E0EC9B48C47C73F07D58" pageId="68" pageNumber="69" start="start">Tetramorium</pageBreakToken>
|
||||
latreillei
|
||||
<authority>Forel</authority>
|
||||
<authority id="5811701903EDEBB9E45F2C4F2E95BC2D">Forel</authority>
|
||||
,
|
||||
</taxonomicName>
|
||||
1895
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,410,146,165]" pageId="68" pageNumber="69" type="mainText">(Figs. 16, 23, 126, 127, 128, 142)</paragraph>
|
||||
<subSubSection id="6D700D3454AE9150478D486305B1DDFA" type="description">
|
||||
<paragraph id="6F12EB080B520B594D06E4EEFFFBDEB6" box="[118,410,146,165]" pageId="68" pageNumber="69" type="mainText">(Figs. 16, 23, 126, 127, 128, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="reference_group">
|
||||
<paragraph box="[118,1122,202,267]" pageId="68" pageNumber="69" type="mainText">
|
||||
<taxonomicName authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="latreillei" subGenus="Xyphomyrmex">
|
||||
<subSubSection id="EF255DC824FC34257637C78D68DC0E65" type="reference_group">
|
||||
<paragraph id="DD7EA343FF3DBFBD0BD5C95A4E77C38A" box="[118,1122,202,267]" pageId="68" pageNumber="69" type="mainText">
|
||||
<taxonomicName id="334F75EB0AD35245C74CF75315078FB8" authorityName="Forel" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="latreillei" subGenus="Xyphomyrmex">
|
||||
Tetramorium (Xyphomyrmex) latreillei
|
||||
<authority>Forel</authority>
|
||||
<authority id="3B2D5A2E5FC3855BCA98EB53E1E3A398">Forel</authority>
|
||||
,
|
||||
</taxonomicName>
|
||||
1895:247. Lectotype worker [designated here] MADAGASCAR, est Imerina ("eastern Imerina") (Sikora) (BMNH: CASENT0102340) [examined]. Paralectotypes, two workers with same data as lectotype (MHNG: CASENT0101291, CASENT0101292) [examined].
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[155,1122,274,315]" pageId="68" pageNumber="69" type="mainText">[Note: Bolton (1979) noted that all syntypes were located in the Forel collection in MHNG. However, we also found one syntype (the newly designated lectotype) in BMNH with the additional information "remounted B. Bolton 1975".]</paragraph>
|
||||
<subSubSection id="6B09DF711C0EFE2378B958DEF7E387A9" type="discussion">
|
||||
<paragraph id="ECF53A1B110DE089CBBB09F7DEED8EC6" box="[155,1122,274,315]" pageId="68" pageNumber="69" type="mainText">[Note: Bolton (1979) noted that all syntypes were located in the Forel collection in MHNG. However, we also found one syntype (the newly designated lectotype) in BMNH with the additional information "remounted B. Bolton 1975".]</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,350,369]" pageId="68" pageNumber="69" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,379,427]" pageId="68" pageNumber="69" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="latreillei">Tetramorium latreillei</taxonomicName>
|
||||
<subSubSection id="E2C774400065754A3E015E7880CC10AA" type="diagnosis">
|
||||
<paragraph id="B80DA2E28E8360BB760503D4031650E4" box="[118,208,350,369]" pageId="68" pageNumber="69" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="6BA87387324C75E66659940AD9DE4497" box="[118,1121,379,427]" pageId="68" pageNumber="69" type="mainText">
|
||||
<taxonomicName id="AB38CF1996C4C7A349AF0D2162CD4DE9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="latreillei">Tetramorium latreillei</taxonomicName>
|
||||
is clearly recognisable within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="395431B8A234E70EB451501851116105" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
complex because it is the only species without any standing hairs on the first gastral tergite.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,465,484]" pageId="68" pageNumber="69" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1117,494,657]" pageId="68" pageNumber="69" type="mainText">HL 1.05-1.06 (1.06); HW 1.06-1.08 (1.07); SL 0.82-0.85 (0.84); EL 0.23-0.24 (0.24); PH 0.52-0.55 (0.53); PW 0.72-0.76 (0.74); WL 1.37-1.40 (1.38); PSL 0.54-0.56 (0.55); PTL 0.34-0.35 (0.35); PTH 0.41-0.43 (0.42); PTW 0.27-0.30 (0.29); PPL 0.30-0.32 (0.31); PPH 0.43-0.45 (0.44); PPW 0.40-0.44 (0.42); CI 100-101 (101); SI 77-79 (79); OI 22 (22); DMI 53-55 (54); LMI 38-39 (39); PSLI 50-53 (52); PeNI 38-39 (39); LPeI 81-85 (83); DPeI 79-86 (83); PpNI 55-58 (56); LPpI 70-72 (71); DPpI 132-138 (134); PPI 141-148 (145) (three measured).</paragraph>
|
||||
<paragraph box="[118,1119,667,1347]" pageId="68" pageNumber="69" type="mainText">Head as long as wide to weakly wider than long (CI 100-101); posterior head margin strongly concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes well-developed, moderately deep, but narrow, and without defined posterior and ventral margins. Antennal scapes short, not reaching posterior head margin (SI 77-79). Eyes of moderate size (OI 22). Mesosomal outline in profile flat, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-39). Propodeal spines massively developed with very broad base, extremely long, and acute (PSLI 50-53); propodeal lobes short and rounded. Petiolar node in profile rectangular nodiform, around 1.2 times higher than long (LPeI 81-85), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum slightly convex; node in dorsal view between 1.1 to 1.3 times longer than wide (DPeI 79-86). Postpetiole in profile subglobular, weakly anteroposteriorly compressed, approximately 1.4 times higher than long (LPpI 70-72); in dorsal view around 1.3 to 1.4 times wider than long (DPpI 132-138). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view about 1.4 to 1.5 times wider than petiolar node (PPI 141-148). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose, with three to five rugae; cephalic dorsum between frontal carinae with 9 to 12 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted or with cross-meshes; lateral and ventral head longitudinally rugose, rarely with cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae with very distinct and pronounced longitudinal rugae. Waist segments longitudinally rugose, rugae on waist segments weaker than on head and mesosoma, especially dorsally. Gaster completely unsculptured, smooth, and shining. Ground sculpture generally faint to absent everywhere on body. Head with abundant standing hairs; mesosoma with up to two pairs restricted to pronotal dorsum; remainder of mesosoma, waist segments, and first gastral tergite without standing hairs; first gastral tergite with moderately dense appressed pubescence. Anterior edges of antennal scapes with appressed hairs. Body a uniform very dark brown to black colour.</paragraph>
|
||||
<subSubSection id="8DF75F91C0319B8303007BA8FAC905D9" type="description">
|
||||
<paragraph id="5DBA4CF2E3FB5F2C60EC4C54AD4510DB" box="[118,226,465,484]" pageId="68" pageNumber="69" type="mainText">Description</paragraph>
|
||||
<paragraph id="0BC6D96AB92624917749AA94EB3A7E38" box="[118,1117,494,657]" pageId="68" pageNumber="69" type="mainText">HL 1.05-1.06 (1.06); HW 1.06-1.08 (1.07); SL 0.82-0.85 (0.84); EL 0.23-0.24 (0.24); PH 0.52-0.55 (0.53); PW 0.72-0.76 (0.74); WL 1.37-1.40 (1.38); PSL 0.54-0.56 (0.55); PTL 0.34-0.35 (0.35); PTH 0.41-0.43 (0.42); PTW 0.27-0.30 (0.29); PPL 0.30-0.32 (0.31); PPH 0.43-0.45 (0.44); PPW 0.40-0.44 (0.42); CI 100-101 (101); SI 77-79 (79); OI 22 (22); DMI 53-55 (54); LMI 38-39 (39); PSLI 50-53 (52); PeNI 38-39 (39); LPeI 81-85 (83); DPeI 79-86 (83); PpNI 55-58 (56); LPpI 70-72 (71); DPpI 132-138 (134); PPI 141-148 (145) (three measured).</paragraph>
|
||||
<paragraph id="49D5E37F7034310D80F473296FACB6DB" box="[118,1119,667,1347]" pageId="68" pageNumber="69" type="mainText">Head as long as wide to weakly wider than long (CI 100-101); posterior head margin strongly concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes well-developed, moderately deep, but narrow, and without defined posterior and ventral margins. Antennal scapes short, not reaching posterior head margin (SI 77-79). Eyes of moderate size (OI 22). Mesosomal outline in profile flat, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 38-39). Propodeal spines massively developed with very broad base, extremely long, and acute (PSLI 50-53); propodeal lobes short and rounded. Petiolar node in profile rectangular nodiform, around 1.2 times higher than long (LPeI 81-85), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins approximately at same height, dorsum slightly convex; node in dorsal view between 1.1 to 1.3 times longer than wide (DPeI 79-86). Postpetiole in profile subglobular, weakly anteroposteriorly compressed, approximately 1.4 times higher than long (LPpI 70-72); in dorsal view around 1.3 to 1.4 times wider than long (DPpI 132-138). Postpetiole in profile appearing less voluminous than petiolar node, in dorsal view about 1.4 to 1.5 times wider than petiolar node (PPI 141-148). Mandibles distinctly longitudinally rugose; clypeus longitudinally rugose, with three to five rugae; cephalic dorsum between frontal carinae with 9 to 12 longitudinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted or with cross-meshes; lateral and ventral head longitudinally rugose, rarely with cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae with very distinct and pronounced longitudinal rugae. Waist segments longitudinally rugose, rugae on waist segments weaker than on head and mesosoma, especially dorsally. Gaster completely unsculptured, smooth, and shining. Ground sculpture generally faint to absent everywhere on body. Head with abundant standing hairs; mesosoma with up to two pairs restricted to pronotal dorsum; remainder of mesosoma, waist segments, and first gastral tergite without standing hairs; first gastral tergite with moderately dense appressed pubescence. Anterior edges of antennal scapes with appressed hairs. Body a uniform very dark brown to black colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,1385,1403]" pageId="68" pageNumber="69" type="mainText">Notes</paragraph>
|
||||
<paragraph box="[118,1121,1414,1548]" pageId="68" pageNumber="69" type="mainText">Unfortunately, no modern material is available. The original type series is from "eastern Imerina" (Forel, 1895), which might be located in central-eastern Madagascar. The exact type locality remains unknown, making it difficult to resample new material. It is surprising, though, that no more specimens have been collected in the last 120 years, especially considering the large sampling effort undertaken by the Malagasy ant inventory project (Fisher, 2005).</paragraph>
|
||||
<caption box="[118,1122,1531,1577]" pageId="69" pageNumber="70" type="caption">
|
||||
<paragraph box="[118,1122,1531,1577]" pageId="69" pageNumber="70" type="caption">
|
||||
<pageBreakToken pageId="69" pageNumber="70" start="start">FIGURES</pageBreakToken>
|
||||
<subSubSection id="3E58C67F8D308BE8FE186B2606901DDA" type="discussion">
|
||||
<paragraph id="D18E44FE196814701B6DD495A1A20715" box="[118,170,1385,1403]" pageId="68" pageNumber="69" type="mainText">Notes</paragraph>
|
||||
<paragraph id="A9EE5FE2DB1453BAD9B98636996EACC3" box="[118,1121,1414,1548]" pageId="68" pageNumber="69" type="mainText">Unfortunately, no modern material is available. The original type series is from "eastern Imerina" (Forel, 1895), which might be located in central-eastern Madagascar. The exact type locality remains unknown, making it difficult to resample new material. It is surprising, though, that no more specimens have been collected in the last 120 years, especially considering the large sampling effort undertaken by the Malagasy ant inventory project (Fisher, 2005).</paragraph>
|
||||
<caption id="BA0270C0EC29A7337201BDD28AA73AB1" box="[118,1122,1531,1577]" pageId="69" pageNumber="70" type="caption">
|
||||
<paragraph id="86DD07600B8556EC10533BD854851D0B" box="[118,1122,1531,1577]" pageId="69" pageNumber="70" type="caption">
|
||||
<pageBreakToken id="7250CB4555656F6D2BE37E3AAA4CA9BE" pageId="69" pageNumber="70" start="start">FIGURES</pageBreakToken>
|
||||
126-128.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei,</taxonomicName>
|
||||
<taxonomicName id="8A70B664C8996DBC29E61C031AFBC3ED" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei,</taxonomicName>
|
||||
lectotype (CASENT0102340). 126. Body in profile. 127. Body in dorsal view. 128. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph box="[118,1122,118,475]" pageId="70" pageNumber="71" type="mainText">
|
||||
<pageBreakToken pageId="70" pageNumber="71" start="start">Within</pageBreakToken>
|
||||
<paragraph id="8BAB6C46EB867F13ADC6C8CB32BE0122" box="[118,1122,118,475]" pageId="70" pageNumber="71" type="mainText">
|
||||
<pageBreakToken id="B82626FD93D9DE4DD080C4F2245D7129" pageId="70" pageNumber="71" start="start">Within</pageBreakToken>
|
||||
the species complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="C9AA879444D44DF41C102EF7574FCE05" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
is the only species lacking standing hairs on the first gastral tergite, and is thus easily identifiable. Even without cons idering this character, it cannot be misidentified with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
<taxonomicName id="B7A04847FA0E7C6DC2C0C9C66C49B388" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="adamsi">T. adamsi</taxonomicName>
|
||||
due to the petiolar node of the latter, which has the posterodorsal margin higher than the anterodorsal margin. Shorter antennal scapes separate
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="2701B945DFB9B5249A5688176BF7CFCA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
<taxonomicName id="DF4D0AF54D7FE6882CFB56C1F18CE07F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
(SI 89-93), and the very dark brown to black colouration distinguishes it from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy,</taxonomicName>
|
||||
<taxonomicName id="C979E97B321EBC18F1C41C899F71309E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy,</taxonomicName>
|
||||
which is of orange colour. However,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="7AA464F66091B9BC8E495FDCE896F0B8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
is very close to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="D574BCC6D36C647958B1720D663C09F7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
<taxonomicName id="B1EA797B8B258FEC34B5465C610A002B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="smaug">T. smaug.</taxonomicName>
|
||||
All three species share the same morphometric range, and superficially possess a similar overall habitus. They have massively constructed, extremely long propodeal spines, reduced hairiness, and very dark brown to black colouration. This character combination clearly separates these three species from the remainder of the complex. As noted above,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="3468EA61743261E0175DF1D3DBB95A5F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
can be easily separated from
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="8C80CBC969F870E55CF445FAC59B2736" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="2FB32C5D652945FAD33F98DD087210F7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
by the pilosity/pubescence on the first gastral tergite. The latter two have only very sparse, short pubescence and few to many long, standing hairs on the first gastral tergite, whereas
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
<taxonomicName id="C2D106B26758D7A53C493196291D3DFD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">T. latreillei</taxonomicName>
|
||||
does not have a single standing hair, but instead a moderately dense, appressed pubescence.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,484,756]" pageId="70" pageNumber="71" type="mainText">
|
||||
<paragraph id="9027115C16B14CBD04DB9BBAB0306924" box="[118,1122,484,756]" pageId="70" pageNumber="71" type="mainText">
|
||||
It is possible that all three species are conspecific, and that the differences in pilosity/pubescence represent geographical variation. The overall morphological similarity might support this hypothesis, as well as the fact that none of the three species was found in sympatry. However, our own studies (Hita Garcia et al., 2010; Hita Garcia & Fisher, 2011, 2012), as well as previous studies on
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
<taxonomicName id="06014540629FE7C0694AB81DBF75D2A6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
|
||||
(Bolton, 1976, 1977, 1979, 1980), concur that pilosity and pubescence patterns are generally very species-specific. The hairs on the mesosoma and first gastral tergite represent a particularly good diagnostic character.
|
||||
<taxonomicName authorityName=", T." class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
<taxonomicName id="461F500B27D00EFDE4CF446B3D45F05E" authorityName=", T." class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
Tetramorium latreillei
|
||||
<authority>
|
||||
<authority id="D04FF95DA4C9353E89766BEB07254D5C">
|
||||
,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
<taxonomicName id="C66A09DDBDA5E9635FDD4D1FB4BC2468" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra</taxonomicName>
|
||||
</authority>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="9A22CF18A28FEDAA8327E94E0F9DFE08" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
all seem to be relatively rare, making them unlikely to be found in the same locality. The distribution ranges of the three, however, strongly overlap. We currently consider all three distinct species that can be well separated, although the species delimitations presented herein might change with additional material.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,793,812]" pageId="70" pageNumber="71" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,441,821,841]" pageId="70" pageNumber="71" type="mainText">MADAGASCAR: Imerina (Sikora).</paragraph>
|
||||
<subSubSection id="387D743A50DBA22617115EC7BCE1028A" type="materials_examined">
|
||||
<paragraph id="36F685A7AE2F60725E1AB0CA06D4C878" box="[118,295,793,812]" pageId="70" pageNumber="71" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="AF98C1CFBEAF410696CCA6F5D873FE23" box="[118,441,821,841]" pageId="70" pageNumber="71" type="mainText">MADAGASCAR: Imerina (Sikora).</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,162 +1,161 @@
|
|||
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172659" ID-GBIF-Taxon="100105376" ID-Zenodo-Dep="6172659" LSID="urn:lsid:plazi:treatment:F9CBF718942F19CE52C5561A5C39F186" httpUri="http://treatment.plazi.org/id/F9CBF718942F19CE52C5561A5C39F186" lastPageNumber="76" pageNumber="74">
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<subSubSection type="nomenclature">
|
||||
<paragraph box="[118,582,118,138]" pageId="73" pageNumber="74" type="mainText">
|
||||
<taxonomicName authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="nazgul">
|
||||
<pageBreakToken pageId="73" pageNumber="74" start="start">Tetramorium</pageBreakToken>
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<treatment id="F9CBF718942F19CE52C5561A5C39F186" ID-DOI="http://doi.org/10.5281/zenodo.6172659" ID-GBIF-Taxon="100105376" ID-Zenodo-Dep="6172659" LSID="urn:lsid:plazi:treatment:F9CBF718942F19CE52C5561A5C39F186" httpUri="http://treatment.plazi.org/id/F9CBF718942F19CE52C5561A5C39F186" lastPageNumber="76" pageNumber="74" scope_family="Formicidae" scope_order="Hymenoptera">
|
||||
<subSubSection id="1107FD04B064AD200A52A05B384BA17D" type="nomenclature">
|
||||
<paragraph id="6CE40ED005CF782BE6FCF26D0162E9F3" box="[118,582,118,138]" pageId="73" pageNumber="74" type="mainText">
|
||||
<taxonomicName id="C88824F5478B0C85CD33FDEA96FF3FE7" authorityName="Hita" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="nazgul">
|
||||
<pageBreakToken id="EFB4A929C46F39EA057CDA79D341C3E0" pageId="73" pageNumber="74" start="start">Tetramorium</pageBreakToken>
|
||||
nazgul
|
||||
<authority>Hita</authority>
|
||||
<authority id="615AF7870863B90B3526A721F6795FA1">Hita</authority>
|
||||
</taxonomicName>
|
||||
Garcia & Fisher
|
||||
<taxonomicNameLabel rank="species">sp. n.</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="C5F829D643EB26699EB085F728538D64" rank="species">sp. n.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,410,146,165]" pageId="73" pageNumber="74" type="mainText">(Figs. 19, 20, 132, 133, 134, 142)</paragraph>
|
||||
<subSubSection id="656B04545132EC5F1067C722F7BD1175" type="description">
|
||||
<paragraph id="3DB077EDD9BB6CD01623F6277D6FD324" box="[118,410,146,165]" pageId="73" pageNumber="74" type="mainText">(Figs. 19, 20, 132, 133, 134, 142)</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,1123,202,474]" pageId="73" pageNumber="74" type="mainText">
|
||||
<subSubSection id="B2104D815B5A6416C38FB00E1F8FBB16" type="materials_examined">
|
||||
<paragraph id="97B7DA52241E7E5828619409994B9C8A" box="[118,1123,202,474]" pageId="73" pageNumber="74" type="mainText">
|
||||
Holotype worker, MADAGASCAR, Toliara,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="2E6876D33EC9206392736F4B12A9FAF2" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="96D8036CE34AF288DA5BB26C33C7571C" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
d'Ambohijanahary,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="3BA5640D3E8358723481F8B49A52FF2B" originalValue="Forêt">Foret</normalizedToken>
|
||||
d'Ankazotsihitafototra, 35.2 km 312° NW Ambaravaranala, 18.26667 S, 45.40667 E, 1050 m, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF07020, 13.-17.I.2003 (B.L. Fisher, C. Griswold et al.) (CASENT: CASENT0028625). Paratypes, 31 workers with same data as holotype (BMNH: CASENT0028584; CASC: CASENT0028579; CASENT0028585; CASENT0028590; CASENT0028591; CASENT0028601; CASENT0028604; CASENT0028605; CASENT0028606; CASENT0028610; CASENT0028620; CASENT0028621; CASENT0028622; CASENT0028626; CASENT0028627; CASENT0028628; CASENT0028632; CASENT0028636; CASENT0028652; CASENT0028663; CASENT0028670; CASENT0028671; CASENT0028674; CASENT0028678; CASENT0028680; CASENT0028681; CASENT0028689; CASENT0028690; MCZ: CASENT0028595; MHNG: CASENT0028594; NHMB: CASENT0028592).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,511,529]" pageId="73" pageNumber="74" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1122,539,643]" pageId="73" pageNumber="74" type="mainText">
|
||||
<subSubSection id="1184E686AE52DA990598E79E6DF25C26" type="diagnosis">
|
||||
<paragraph id="75972B88C71AAD83E0E4B702B34B8D0A" box="[118,208,511,529]" pageId="73" pageNumber="74" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="20E7A83B79FA7D3C9FBB90C58A190DC9" box="[118,1122,539,643]" pageId="73" pageNumber="74" type="mainText">
|
||||
The following character combination renders
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
<taxonomicName id="F99831DBFEEEDEC410915AA66E63A6DA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
easily diagnosable within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="E9ACC21071BEA6F151A88FD8757EAE7F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
species complex: antennal scapes comparatively long (SI 89-92); propodeal spines very long (PSLI 39-43); anterodorsal and posterodorsal margins of petiolar node situated at about same height; first gastral tergite with numerous standing hairs; dark brown to black body colour.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
||||
<paragraph box="[118,226,680,698]" pageId="73" pageNumber="74" type="mainText">Description</paragraph>
|
||||
<paragraph box="[118,1117,708,868]" pageId="73" pageNumber="74" type="mainText">HL 0.95-1.06 (1.02); HW 0.85-0.98 (0.93); SL 0.79-0.89 (0.84); EL 0.21-0.24 (0.22); PH 0.48-0.56 (0.53); PW 0.67-0.75 (0.73); WL 1.23-1.39 (1.34); PSL 0.38-0.45 (0.42); PTL 0.31-0.36 (0.34); PTH 0.38-0.43 (0.41); PTW 0.26-0.31 (0.29); PPL 0.31-0.36 (0.33); PPH 0.38-0.42 (0.41); PPW 0.35-0.41 (0.39); CI 90-92 (91); SI 89-92 (90); OI 23-25 (24); DMI 54-57 (55); LMI 37-41 (39); PSLI 39-43 (41); PeNI 37-42 (40); LPeI 81-88 (84); DPeI 82-89 (86); PpNI 51-55 (54); LPpI 78-85 (81); DPpI 113-122 (118); PPI 127-140 (133) (12 measured).</paragraph>
|
||||
<paragraph box="[118,1120,877,1543]" pageId="73" pageNumber="74" type="mainText">Head distinctly longer than wide (CI 90-92); posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, shallow and narrow, without defined ventral margins. Antennal scapes of moderate length, not reaching posterior head margin (SI 89-92). Eyes of moderate size (OI 23-25). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-41). Propodeal spines very long, spinose and acute (PSLI 39-43); propodeal lobes well-developed, triangular, and usually acute. Petiolar node in profile rectangular nodiform, approximately 1.1 to 1.2 times higher than long (LPeI 81-88), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins situated at about same height, dorsum flat to weakly convex; node in dorsal view approximately 1.1 to 1.2 times longer than wide (DPeI 82-89). Postpetiole in profile globular, approximately 1.2 to 1.3 times higher than long (LPpI 78-85); in dorsa l view around 1.1 to 1.2 times wider than long (DPpI 113-122). Postpetiole in profile appearing approximately as voluminous as petiolar node, in dorsal view approximately 1.3 to 1.4 times wider than petiolar node (PPI 127-140). Mandibles striate; clypeus longitudinally rugose/rugulose, with three to eight rugae/rugulae, median ruga usually present and distinct, remaining rugae/rugulae variably developed, usually weaker than median ruga; cephalic dorsum between frontal carinae with 8 to 13 longit udinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted and with cross- meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose to reticulate-rugose. Mesosoma laterally and dorsally mainly longitudinally rugose. Forecoxae with well-developed and conspicuous longitudinal rugae. Waist segments strongly rugose. Gaster unsculptured, smooth, and shining. Ground sculpture generally faint to absent everywhere on body. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with suberect to er ect hairs. Body of uniform dark brown to black colour, often appendages of slightly lighter colour.</paragraph>
|
||||
<subSubSection id="D6457C7DAD3A20E34281503A1C1059A1" type="description">
|
||||
<paragraph id="C763A495C9B79D08A57E5A632A80A826" box="[118,226,680,698]" pageId="73" pageNumber="74" type="mainText">Description</paragraph>
|
||||
<paragraph id="869129C101372AC57217D2640DBDA1A0" box="[118,1117,708,868]" pageId="73" pageNumber="74" type="mainText">HL 0.95-1.06 (1.02); HW 0.85-0.98 (0.93); SL 0.79-0.89 (0.84); EL 0.21-0.24 (0.22); PH 0.48-0.56 (0.53); PW 0.67-0.75 (0.73); WL 1.23-1.39 (1.34); PSL 0.38-0.45 (0.42); PTL 0.31-0.36 (0.34); PTH 0.38-0.43 (0.41); PTW 0.26-0.31 (0.29); PPL 0.31-0.36 (0.33); PPH 0.38-0.42 (0.41); PPW 0.35-0.41 (0.39); CI 90-92 (91); SI 89-92 (90); OI 23-25 (24); DMI 54-57 (55); LMI 37-41 (39); PSLI 39-43 (41); PeNI 37-42 (40); LPeI 81-88 (84); DPeI 82-89 (86); PpNI 51-55 (54); LPpI 78-85 (81); DPpI 113-122 (118); PPI 127-140 (133) (12 measured).</paragraph>
|
||||
<paragraph id="DF6EEFE8029E585AB66ADCAC531F19AD" box="[118,1120,877,1543]" pageId="73" pageNumber="74" type="mainText">Head distinctly longer than wide (CI 90-92); posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes developed, shallow and narrow, without defined ventral margins. Antennal scapes of moderate length, not reaching posterior head margin (SI 89-92). Eyes of moderate size (OI 23-25). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 37-41). Propodeal spines very long, spinose and acute (PSLI 39-43); propodeal lobes well-developed, triangular, and usually acute. Petiolar node in profile rectangular nodiform, approximately 1.1 to 1.2 times higher than long (LPeI 81-88), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins situated at about same height, dorsum flat to weakly convex; node in dorsal view approximately 1.1 to 1.2 times longer than wide (DPeI 82-89). Postpetiole in profile globular, approximately 1.2 to 1.3 times higher than long (LPpI 78-85); in dorsa l view around 1.1 to 1.2 times wider than long (DPpI 113-122). Postpetiole in profile appearing approximately as voluminous as petiolar node, in dorsal view approximately 1.3 to 1.4 times wider than petiolar node (PPI 127-140). Mandibles striate; clypeus longitudinally rugose/rugulose, with three to eight rugae/rugulae, median ruga usually present and distinct, remaining rugae/rugulae variably developed, usually weaker than median ruga; cephalic dorsum between frontal carinae with 8 to 13 longit udinal rugae, most rugae running unbroken from posterior head margin to anterior clypeus, few rugae interrupted and with cross- meshes; scrobal area mostly unsculptured; lateral and ventral head longitudinally rugose to reticulate-rugose. Mesosoma laterally and dorsally mainly longitudinally rugose. Forecoxae with well-developed and conspicuous longitudinal rugae. Waist segments strongly rugose. Gaster unsculptured, smooth, and shining. Ground sculpture generally faint to absent everywhere on body. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with suberect to er ect hairs. Body of uniform dark brown to black colour, often appendages of slightly lighter colour.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="multiple">
|
||||
<caption box="[118,1122,1549,1591]" pageId="74" pageNumber="75" type="caption">
|
||||
<paragraph box="[118,1122,1549,1591]" pageId="74" pageNumber="75" type="caption">
|
||||
<pageBreakToken pageId="74" pageNumber="75" start="start">FIGURES</pageBreakToken>
|
||||
<subSubSection id="50E0952E8A5912EA652E0FD0EDF4AD6F" type="multiple">
|
||||
<caption id="0698330FE35FF720F9A9E527152DFC4B" box="[118,1122,1549,1591]" pageId="74" pageNumber="75" type="caption">
|
||||
<paragraph id="90DA17B8C1BBE1CF8905839D4E631DD1" box="[118,1122,1549,1591]" pageId="74" pageNumber="75" type="caption">
|
||||
<pageBreakToken id="5CEE0B35E044DDFD4E42BA37414AD256" pageId="74" pageNumber="75" start="start">FIGURES</pageBreakToken>
|
||||
132-134.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul,</taxonomicName>
|
||||
<taxonomicName id="E558C5919E6B5AA8602E7B5880464C71" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul,</taxonomicName>
|
||||
holotype (CASENT0028625). 132. Body in profile. 133. Body in dorsal view. 134. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,118,136]" pageId="75" pageNumber="76" type="mainText">
|
||||
<pageBreakToken pageId="75" pageNumber="76" start="start">Notes</pageBreakToken>
|
||||
<subSubSection id="A4D77677AE0A7D405B9FDCDB2E25195E" type="discussion">
|
||||
<paragraph id="20D9EFC7D55A29993C19C499BF99A88C" box="[118,170,118,136]" pageId="75" pageNumber="76" type="mainText">
|
||||
<pageBreakToken id="AFB920D2BF514E7F70AFC9562C1C01EB" pageId="75" pageNumber="76" start="start">Notes</pageBreakToken>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,147,222]" pageId="75" pageNumber="76" type="mainText">
|
||||
<paragraph id="E2ACF048FE93224503918F808641A76D" box="[118,1121,147,222]" pageId="75" pageNumber="76" type="mainText">
|
||||
This
|
||||
<taxonomicNameLabel rank="species">new species</taxonomicNameLabel>
|
||||
<taxonomicNameLabel id="2B140482065C4469189670218A279D78" rank="species">new species</taxonomicNameLabel>
|
||||
is known from few specimens from Analalava and Zombitse, both tropical dry forests, and a high number of specimens from the type locality Ambohijanahary, which is a montane rainforest. The altitudinal range is 700 to 1100 m, and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
<taxonomicName id="98E21861E00985BC9558F5D13333C23B" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
appears to be a leaf litter inhabitant.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,231,503]" pageId="75" pageNumber="76" type="mainText">
|
||||
<paragraph id="AC1A40C4AD90ADEC203ED9A7D200F2B0" box="[118,1122,231,503]" pageId="75" pageNumber="76" type="mainText">
|
||||
Within the
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="20816646095A6200430DFB5C84219948" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
species complex, it is easily separated from the three species
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
<taxonomicName id="A0590A8059BABD8CD559E6DA3CA488BB" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="latreillei">
|
||||
T. latreillei
|
||||
<authority>,</authority>
|
||||
<authority id="45D3BEE4068E4A73A635FCC3C13DDB8B">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
<taxonomicName id="B782B806355FBA4ECC1A4447BA5D0E59" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="sabatra">T. sabatra,</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
<taxonomicName id="1C758F81AC6A19D7F2905B2DF354D822" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="smaug">T. smaug</taxonomicName>
|
||||
which have extremely long and massively developed propodeal spines with a very broad base (PSLI 48-72), whereas the spines of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
<taxonomicName id="E5A1F63F8531B470581C2B716D890D06" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
are very long, but less massive and without such a broad base (PSLI 39-43).
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="adamsi">Tetramorium adamsi,</taxonomicName>
|
||||
<taxonomicName id="155BCA09F78271217ACBCD492BAC28B7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="adamsi">Tetramorium adamsi,</taxonomicName>
|
||||
which is only known from the area around Manongarivo, has a petiolar node shape with the posterodorsal margin situated higher than the anterodorsal, whereas in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
<taxonomicName id="8783FE3C90F574B88B429A5772965E4F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul</taxonomicName>
|
||||
both margins are about the same height. The last species of the complex,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy,</taxonomicName>
|
||||
<taxonomicName id="A5466C706630E627ECA93897719D52F9" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy,</taxonomicName>
|
||||
is also very unlikely to be confused with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul.</taxonomicName>
|
||||
<taxonomicName id="4643054AFE3C3A09E3DABAB03709C0C8" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul.</taxonomicName>
|
||||
The most obvious character is colouration (orange to light brown in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
<taxonomicName id="48D74CDA977AE9933A1AD7EACC933A63" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
versus dark brown to black in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul),</taxonomicName>
|
||||
<taxonomicName id="A9472BB6BD607C64A37495196BA1CB3E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">T. nazgul),</taxonomicName>
|
||||
but they also differ in antennal scape length and shape of the waist segments.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">Tetramorium nazgul</taxonomicName>
|
||||
<taxonomicName id="AA708D695177AE14C2B48358124E6282" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nazgul">Tetramorium nazgul</taxonomicName>
|
||||
has longer antennal scapes (SI 89-92), a relatively longer petiolar node (LPeI 81-88), and a higher and broader postpetiole (LPpI 78-85; DPpI 113-122) than
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
<taxonomicName id="ED420626370DB7B783F52DBAF5821A3F" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="marojejy">T. marojejy</taxonomicName>
|
||||
(SI 79-85; LPeI 89-97; LPpI 88-95; DPpI 101-109).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="etymology">
|
||||
<paragraph box="[118,219,540,558]" pageId="75" pageNumber="76" type="mainText">Etymology</paragraph>
|
||||
<paragraph box="[118,1120,568,616]" pageId="75" pageNumber="76" type="mainText">
|
||||
<subSubSection id="3EE5746715A6AD2583F9D671450E176B" type="etymology">
|
||||
<paragraph id="90176B421B27D956D6EF76D669BFE4CC" box="[118,219,540,558]" pageId="75" pageNumber="76" type="mainText">Etymology</paragraph>
|
||||
<paragraph id="1DF038E90CB6554A4DE81B17826096F2" box="[118,1120,568,616]" pageId="75" pageNumber="76" type="mainText">
|
||||
The species name
|
||||
<normalizedToken originalValue="“nazgul”">"nazgul"</normalizedToken>
|
||||
<normalizedToken id="9A660F760B6A505D9C432F9408F3DDB0" originalValue="“nazgul”">"nazgul"</normalizedToken>
|
||||
is taken from J.R.R. Tolkien's "The Lord of the Rings" and refers to the evil ringwraiths who serve the main villain
|
||||
<normalizedToken originalValue="“Sauron”">"Sauron"</normalizedToken>
|
||||
<normalizedToken id="B0259162C8682F4F89C84AF7A8C74A88" originalValue="“Sauron”">"Sauron"</normalizedToken>
|
||||
. The species epithet is an arbitrary combination of letters.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,653,671]" pageId="75" pageNumber="76" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,681,869]" pageId="75" pageNumber="76" type="mainText">
|
||||
<subSubSection id="268B314BE9134917DECA2FBD0AF76DF9" type="materials_examined">
|
||||
<paragraph id="5D7935CF1FCDDDCF08BF38EC86DD9441" box="[118,295,653,671]" pageId="75" pageNumber="76" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="292BEBAE55D74FF461BB6FB2A4289826" box="[118,1122,681,869]" pageId="75" pageNumber="76" type="mainText">
|
||||
MADAGASCAR: Fianarantsoa,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="E1D248380E4756BFC590C839E4C08C61" originalValue="Forêt">Foret</normalizedToken>
|
||||
d'Analalava, 29.6 km 280° W Ranohira, 22.59167 S, 45.12833 E, 700 m, tropical dry forest, 1.-5.II.2003 (B.L. Fisher, C. Griswold et al.); Toliara,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="2DF540FC85A33117C1062D2D7C08A4B7" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="DBDA394E5B01DD59BDDBEB5E1ABE4F5E" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
d'Ambohijanahary,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="2ABFDFFF70214F00582230008DEAF2A3" originalValue="Forêt">Foret</normalizedToken>
|
||||
d'Ankazotsihitafototra, 35.2 km 312° NW Ambaravaranala, 18.26667 S, 45.40667 E, 1050 m, montane rainforest, 13.-17.I.2003 (B.L. Fisher, C. Griswold et al.); Toliara,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="0386FFE0A7C0CCD3C4B4C97AA6B11A7F" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="062D2BCAC01B32557CE77749F713E198" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
d'Ambohijanahary,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="C6F2083B669C1F563A71A3C724598FE3" originalValue="Forêt">Foret</normalizedToken>
|
||||
d'Ankazotsihitafototra, 34.6 km 314° NW Ambaravaranala, 18.26 S, 45.4183 E, 1100 m, montane rainforest, 16.I.2003 (B.L. Fisher, C. Griswold et al.); Toliara, Parc National de Zombitse, 19.8 km 84° E Sakaraha, 22.84333 S, 44.71 E, 770 m, tropical dry forest, 5.-9.II.2003 (B.L. Fisher, C. Griswold et al.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
Loading…
Reference in a new issue