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<mods:title>The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups.</mods:title>
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<mods:title id="093F87EEE977CBD23BDE39659E041CBB">The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region - taxonomic revision of the T. kelleri and T. tortuosum species groups.</mods:title>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6172621" ID-GBIF-Taxon="100105350" ID-Zenodo-Dep="6172621" LSID="urn:lsid:plazi:treatment:E1C056AC2D88E0B22FE4FA2E19F8BD0A" httpUri="http://treatment.plazi.org/id/E1C056AC2D88E0B22FE4FA2E19F8BD0A" lastPageNumber="40" pageNumber="37">
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<paragraph box="[118,447,428,448]" pageId="36" pageNumber="37" type="mainText">
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<taxonomicName authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">
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<taxonomicName id="213678485E5CB9ED86B39CF4844C8D66" ID-CoL="55PNN" authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">
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Tetramorium isectum
|
||||
<authority>Bolton</authority>
|
||||
<authority id="32F3A1DEB8272E2043422AE20F9741F2">Bolton</authority>
|
||||
,
|
||||
</taxonomicName>
|
||||
1979
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="description">
|
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<paragraph box="[118,345,455,474]" pageId="36" pageNumber="37" type="mainText">(Figs. 52, 90, 91, 92, 141)</paragraph>
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<subSubSection id="03753A56F0DAF39D3E3D7FF6DBA6F742" type="description">
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<paragraph id="544903387E64FAE7128DDB2703BBD8D6" box="[118,345,455,474]" pageId="36" pageNumber="37" type="mainText">(Figs. 52, 90, 91, 92, 141)</paragraph>
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</subSubSection>
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<subSubSection type="reference_group">
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<paragraph box="[118,1122,511,552]" pageId="36" pageNumber="37" type="mainText">
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<taxonomicName authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">
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<subSubSection id="F05377646910B3F9F2970FECF0EDE327" type="reference_group">
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<paragraph id="5F1A56A6B5309221047AC87A11516B35" box="[118,1122,511,552]" pageId="36" pageNumber="37" type="mainText">
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<taxonomicName id="95C0226C9656F897F8931E2C07468AEC" authorityName="Bolton" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">
|
||||
Tetramorium isectum
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||||
<authority>Bolton</authority>
|
||||
<authority id="C551A3F506EA9C09A7962386DDD747C5">Bolton</authority>
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||||
,
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</taxonomicName>
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||||
1979: 145. Holotype worker, MADAGASCAR, Beforona, 500 m, forest humus and litter, IX.1974 (A. Peyrieras) (MCZ: CASENT0172829) [examined].
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||||
</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection type="diagnosis">
|
||||
<paragraph box="[118,208,587,605]" pageId="36" pageNumber="37" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph box="[118,1121,615,719]" pageId="36" pageNumber="37" type="mainText">
|
||||
<subSubSection id="69F2DBB8DBC4EABECFE532ECB8C4C49A" type="diagnosis">
|
||||
<paragraph id="1D786AEBAEAC662B1B828CA1E3A7F04C" box="[118,208,587,605]" pageId="36" pageNumber="37" type="mainText">Diagnosis</paragraph>
|
||||
<paragraph id="2CD88C8920F13B4B659F12F9FF3E4FCD" box="[118,1121,615,719]" pageId="36" pageNumber="37" type="mainText">
|
||||
Within the
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||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
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||||
<taxonomicName id="A49574B81ED444D691E77564C7D162E0" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
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complex
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
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||||
<taxonomicName id="1C6DE6ACD235B8237B703AADF7BEF2C1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
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is distinguishable by the following characters: very small eyes (OI 15-16); long propodeal spines (PSLI 28-33); petiolar node with the anterodorsal margin slightly situated higher and more angular than the posterodorsal margin; posterodorsal corner of petiole not strongly protruding posteriorly; body of uniform bright orange colour.
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||||
</paragraph>
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||||
</subSubSection>
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||||
<subSubSection type="description">
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||||
<paragraph box="[118,226,756,774]" pageId="36" pageNumber="37" type="mainText">Description</paragraph>
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||||
<paragraph box="[118,1121,784,916]" pageId="36" pageNumber="37" type="mainText">HL 0.90-1.10 (0.96); HW 0.85-1.50 (0.93); SL 0.65-0.83 (0.72); EL 0.14-0.18 (0.15); PH 0.42-0.54 (0.47); PW 0.58-0.76 (0.68); WL 1.08-1.39 (1.17); PSL 0.26-0.36 (0.29); PTL 0.25-0.34 (0.28); PTH 0.33-0.40 (0.35); PTW 0.24-0.29 (0.26); PPL 0.27-0.35 (0.31); PPH 0.31-0.39 (0.34); PPW 0.32-0.40 (0.34); CI 94-99 (97); SI 74-79 (77); OI 15-16 (16); DMI 54-60 (58); LMI 39-42 (40); PSLI 28-33 (30); PeNI 36-42 (38); LPeI 71-85 (81); DPeI 86-98 (92); PpNI 49-54 (50); LPpI 82-97 (91); DPpI 103-120 (111); PPI 128-136 (132) (12 measured).</paragraph>
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||||
<paragraph box="[118,1122,925,1563]" pageId="36" pageNumber="37" type="mainText">Head usually weakly longer than wide (CI 94-99). Posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes short, not reaching posterior head margin (SI 74-79). Eyes very small (OI 15-16). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 39-42). Propodeal spines long, spinose, and acute (PSLI 28-33); propodeal lobes short, triangular to elongate-triangular, and acute. Petiolar node in profile rectangular nodiform, around 1.2 to 1.4 times higher than long (LPeI 71-85), anterior and posterior faces approximately parallel, anterodorsal margin situated weakly higher and more angulate than posterodorsal margin, dorsum tapering weakly backwards posteriorly; node in dorsal view between 1.0 to 1.2 times longer than wide (DPeI 86-98). Postpetiole in profile globular, approximately 1.0 to 1.2 times higher than long (LPpI 82-97); in dorsal view between 1.0 and 1.2 times wider than long (DPpI 103-120). Postpetiole in profile appearing approximately as voluminous as petiolar node, in dorsal view approximately 1.3 to 1.4 times wider than petiolar node (PPI 128-136). Mandibles distinctly longitudinally rugose; clypeus generally with three distinct longitudinal rugae/rugulae, sometimes with few more but much weaker rugulae present, and very rarely clypeus with irregular rugulation; cephalic dorsum between frontal carinae with 8 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae very rarely with cross-meshes; lateral and ventral head longitudinally rugose with very few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured, smooth, and shining. Waist segments irregularly rugulose with distinct punctate ground sculpture, laterally ground sculpture better developed and rugulae weaker. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with decumbent to suberect standing hairs. Body colour bright yellow to orange.</paragraph>
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||||
<subSubSection id="75DAA377735017CF04B34DC74CEA90C4" type="description">
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||||
<paragraph id="5852B454FBE4CCA1C0E73B62D0BDE8A5" box="[118,226,756,774]" pageId="36" pageNumber="37" type="mainText">Description</paragraph>
|
||||
<paragraph id="1698A0C6D62B64F9C690ED750B218E32" box="[118,1121,784,916]" pageId="36" pageNumber="37" type="mainText">HL 0.90-1.10 (0.96); HW 0.85-1.50 (0.93); SL 0.65-0.83 (0.72); EL 0.14-0.18 (0.15); PH 0.42-0.54 (0.47); PW 0.58-0.76 (0.68); WL 1.08-1.39 (1.17); PSL 0.26-0.36 (0.29); PTL 0.25-0.34 (0.28); PTH 0.33-0.40 (0.35); PTW 0.24-0.29 (0.26); PPL 0.27-0.35 (0.31); PPH 0.31-0.39 (0.34); PPW 0.32-0.40 (0.34); CI 94-99 (97); SI 74-79 (77); OI 15-16 (16); DMI 54-60 (58); LMI 39-42 (40); PSLI 28-33 (30); PeNI 36-42 (38); LPeI 71-85 (81); DPeI 86-98 (92); PpNI 49-54 (50); LPpI 82-97 (91); DPpI 103-120 (111); PPI 128-136 (132) (12 measured).</paragraph>
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<paragraph id="7288CA8D83C0717E47D1D0E0346D5B47" box="[118,1122,925,1563]" pageId="36" pageNumber="37" type="mainText">Head usually weakly longer than wide (CI 94-99). Posterior head margin concave. Anterior clypeal margin medially impressed. Frontal carinae strongly developed, diverging posteriorly, and ending at corners of posterior head margin. Antennal scrobes weakly developed, shallow, narrow, and without defined posterior and ventral margins. Antennal scapes short, not reaching posterior head margin (SI 74-79). Eyes very small (OI 15-16). Mesosomal outline in profile flat to weakly convex, moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma comparatively stout and high (LMI 39-42). Propodeal spines long, spinose, and acute (PSLI 28-33); propodeal lobes short, triangular to elongate-triangular, and acute. Petiolar node in profile rectangular nodiform, around 1.2 to 1.4 times higher than long (LPeI 71-85), anterior and posterior faces approximately parallel, anterodorsal margin situated weakly higher and more angulate than posterodorsal margin, dorsum tapering weakly backwards posteriorly; node in dorsal view between 1.0 to 1.2 times longer than wide (DPeI 86-98). Postpetiole in profile globular, approximately 1.0 to 1.2 times higher than long (LPpI 82-97); in dorsal view between 1.0 and 1.2 times wider than long (DPpI 103-120). Postpetiole in profile appearing approximately as voluminous as petiolar node, in dorsal view approximately 1.3 to 1.4 times wider than petiolar node (PPI 128-136). Mandibles distinctly longitudinally rugose; clypeus generally with three distinct longitudinal rugae/rugulae, sometimes with few more but much weaker rugulae present, and very rarely clypeus with irregular rugulation; cephalic dorsum between frontal carinae with 8 to 11 longitudinal rugae, most rugae running unbroken from posterior head margin to posterior clypeus, rugae very rarely with cross-meshes; lateral and ventral head longitudinally rugose with very few cross-meshes. Mesosoma laterally and dorsally distinctly longitudinally rugose. Forecoxae unsculptured, smooth, and shining. Waist segments irregularly rugulose with distinct punctate ground sculpture, laterally ground sculpture better developed and rugulae weaker. First gastral tergite unsculptured, smooth, and shining. All dorsal surfaces of head, mesosoma, waist segments, and gaster with abundant, long, and fine standing hairs. Anterior edges of antennal scapes with decumbent to suberect standing hairs. Body colour bright yellow to orange.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="multiple">
|
||||
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|
||||
<paragraph box="[118,1122,1525,1570]" pageId="37" pageNumber="38" type="caption">
|
||||
<pageBreakToken pageId="37" pageNumber="38" start="start">FIGURES</pageBreakToken>
|
||||
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|
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<caption id="20388B3731DE8A28437EB1AE7BA04A1A" box="[118,1122,1525,1570]" pageId="37" pageNumber="38" type="caption">
|
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<paragraph id="79927E0FF09691EF57B38B376CB0EFB3" box="[118,1122,1525,1570]" pageId="37" pageNumber="38" type="caption">
|
||||
<pageBreakToken id="A5824958492D9CCF67BDDCCA6FAEAA52" pageId="37" pageNumber="38" start="start">FIGURES</pageBreakToken>
|
||||
90-92.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="isectum">T. isectum,</taxonomicName>
|
||||
<taxonomicName id="B3B0DF5249C3669C36442BC5550E485E" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="isectum">T. isectum,</taxonomicName>
|
||||
holotype (CASENT0172829). 90. Body in profile. 91. Body in dorsal view. 92. Head in full-face view.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection type="discussion">
|
||||
<paragraph box="[118,170,118,136]" pageId="38" pageNumber="39" type="mainText">
|
||||
<pageBreakToken pageId="38" pageNumber="39" start="start">Notes</pageBreakToken>
|
||||
<subSubSection id="A487C56331972A89305DA7FCA6C3550E" type="discussion">
|
||||
<paragraph id="7B3006589BCCBFBA78CC24EF2122692B" box="[118,170,118,136]" pageId="38" pageNumber="39" type="mainText">
|
||||
<pageBreakToken id="8E4D14A9987BEB9BC332676E2B027A69" pageId="38" pageNumber="39" start="start">Notes</pageBreakToken>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1121,147,362]" pageId="38" pageNumber="39" type="mainText">
|
||||
<paragraph id="21921888898690914ED3D50E352B95C7" box="[118,1121,147,362]" pageId="38" pageNumber="39" type="mainText">
|
||||
Prior to our study
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="351F29FAD3F9C9D0638A339BC0A9361D" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
was only known from the holotype from Beforona. Fortunately, we had more material available because the species has been collected several times since the original description of Bolton (1979).
|
||||
<taxonomicName genus="Tetramoirum" pageId="38" pageNumber="39" rank="species" species="isectum">Tetramoirum isectum</taxonomicName>
|
||||
<taxonomicName id="F94C2BF37F6FAF96169297F04F4DE34F" genus="Tetramoirum" pageId="38" pageNumber="39" rank="species" species="isectum">Tetramoirum isectum</taxonomicName>
|
||||
does not seem to be as rare as might be expected from the original description. We were able to examine material from a number of localities, although the species is admittedly never very common or abundant.
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
<taxonomicName id="61ED479FF4DE1B8F510BE99B399FB935" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
is found in a strip in eastern Madagascar, which ranges from central-eastern to northeastern Madagascar. The southernmost localities are Ambatovy, Andasibe-Mantadia, and Sahafina, and the northernmost are Montagne d'Anjanaharibe and Amparihibe. The species prefers rainforests and montane rainforests at elevations of 125 to 1040 m, and was mostly collected from leaf litter.
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,372,728]" pageId="38" pageNumber="39" type="mainText">
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
<paragraph id="403341234E60E4662100282CD297761D" box="[118,1122,372,728]" pageId="38" pageNumber="39" type="mainText">
|
||||
<taxonomicName id="9C5124EB74BBC566A724236C7AF02BAD" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">Tetramorium isectum</taxonomicName>
|
||||
is morphologically fairly close to
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
<taxonomicName id="843F4416B66EE9D92DC0EF4BEFF462BE" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei,</taxonomicName>
|
||||
and during this revision we considered the synonymisation of the first under the latter. Both differ mainly in eye size, which is very small in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="877355A2AB8B83D879BA271807EB0060" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
(OI 15-16) versus small to moderately large in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="811B372E662B56278CC425C0B4082EDA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
(OI 19-25), but there a few more supporting characters found in both species, although not consistently. All specimens of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="06DAA201F7C9D6CA3E409332AC9C79C3" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
have, in addition to very small eyes, a petiolar node shape with the anterodorsal margin higher and sharper than the posterodorsal with a dorsum that tapers weakly posteriorly, as well as a bright orange body colour. However, within the vast
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="55A0DC9BEF03CE1933F87B56B068CB33" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
material available, there are specimens with a node shape like the one of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum,</taxonomicName>
|
||||
<taxonomicName id="C396902A1C8B9A6F5412C296BC9A77DF" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum,</taxonomicName>
|
||||
although these are mainly restricted to northwestern Madagascar where
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="4BE17086F63A25E9C88481EDBDDFA009" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
does not occur. Many specimens of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="08325B7BEBF7DFA2DEABC7D1C435B4A5" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
are also bright orange in colour. Nevertheless,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="A3B909D97B3A95DAB6B2DF9EA764DB45" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
was found to live in sympatry with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="32BEC5488391B4ABEFCA61CED5E9F903" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
throughout most of its distribution range, and remained remarkably recognisable. The diagnostic characters of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="3572E00FEDC090DCB1C56E7C397C4174" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
provided above are very consistent throughout all the material studied, and no intermediate forms seem to exist. With these facts in mind, we keep
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="08918AEE76278936F366EE132575F6C6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
a separate though relatively uncommon species which co-occurs regularly with the much more common and abundant
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
<taxonomicName id="D918A150CB97665437CC6DD5AA6E2D23" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei.</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph box="[118,1122,737,1037]" pageId="38" pageNumber="39" type="mainText">
|
||||
<paragraph id="D03E14D2490FA35F98A43C45F9981D36" box="[118,1122,737,1037]" pageId="38" pageNumber="39" type="mainText">
|
||||
Apart from the similarities with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
<taxonomicName id="185628BE3EFD54A0F2E8428E1DD6C303" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
|
||||
mentioned above,
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="99A081D95297E027E0F31DD315FD24C7" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
is easily recognisable within the species complex. The very small eyes (OI 15-16) are very conspicuous with the caveat that
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="76D024EBB9D58D4C23A873775467311C" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="electrum">
|
||||
T. electrum
|
||||
<authority>,</authority>
|
||||
<authority id="3BBEAAAB52CD33FE1C38AE10A98890EA">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="elf">
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="9CD2214E08655019673019E7ABA8D99F" authorityName="," genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="elf">
|
||||
<taxonomicName id="7977012DDD0FBF8A39A3557FF0689D47" genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
elf
|
||||
<authority>,</authority>
|
||||
<authority id="03C1E398E6A54237FFDAFC033C8ED946">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
<taxonomicName id="454CB6B0ACC502771D5E22A64F8204B4" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum,</taxonomicName>
|
||||
and
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="D9D322B6A75842620FB60F4B28A1D238" genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="nify">T. nify</taxonomicName>
|
||||
also have generally smaller eyes (OI 16-19). However,
|
||||
<taxonomicName authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="electrum">
|
||||
<taxonomicName id="93BCEFA86F6A09ADA83967CF75E679D8" authorityName="," class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="electrum">
|
||||
T. electrum
|
||||
<authority>,</authority>
|
||||
<authority id="373AAB24F803C7EFBDBA0051E79C4EBA">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="elf">
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="E697927B9D190AD7381FD3022FE8CA39" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="elf">
|
||||
<taxonomicName id="DEFC62175E226AAD9DA2D37A53F53BF5" genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
elf,
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
<taxonomicName id="E12734C3D1EE9FFB3514FC8FF76259BA" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isoelectrum">T. isoelectrum</taxonomicName>
|
||||
have very long to extremely long propodeal spines (PSLI 46-64), whereas the spines of
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="F3695E61417AA2350D4C9F3513AA5469" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
are much shorter (PSLI 28-35).
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="nify">Tetramorium nify</taxonomicName>
|
||||
<taxonomicName id="E1DD211AD79A64DC6B77023C58796033" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="nify">Tetramorium nify</taxonomicName>
|
||||
is also unlikely to be misidentified with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="7EC99C167F1E88F252D8BC95CFE78E8A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
since the latter is bright orange in colour and the former is very dark brown to black in colour. In addition, they have fairly differently shaped petiolar nodes. The node of
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="nify">T. nify</taxonomicName>
|
||||
<taxonomicName id="172434FB7B8A17413D3132465F9EDC6B" genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="nify">T. nify</taxonomicName>
|
||||
has antero- and posterodorsal margins at the same height and is only faintly higher than long (LPeI 91-100), whereas in
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
<taxonomicName id="FDF46B6E2637DA57D54E6F1BAA7E2FC6" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum</taxonomicName>
|
||||
the anterodorsal margin is usually weakly higher and the node is between 1.2 to 1.4 times higher than long (LPeI 71-85). The remaining species, T.
|
||||
<taxonomicName authorityName="," genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="ala">
|
||||
<taxonomicName id="A8DD140993A37F488705AAA5C0594D8A" authorityName="," genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="ala">
|
||||
ala
|
||||
<authority>,</authority>
|
||||
<authority id="11D6E0C9B3B562C052D21F3278C8BEF8">,</authority>
|
||||
</taxonomicName>
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
<taxonomicName id="B58B64CEFF31AFD532854DA75E3618BA" genus="Tetamorium" pageId="38" pageNumber="39" rank="genus">T.</taxonomicName>
|
||||
<taxonomicName id="0DFA1D7899671029E8292A14A6D104C1" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="andohahela">andohahela</taxonomicName>
|
||||
, and
|
||||
<taxonomicName genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="voasary">T. voasary,</taxonomicName>
|
||||
<taxonomicName id="7E766B1DDE082CCE17170821132FF590" genus="Tetamorium" pageId="38" pageNumber="39" rank="species" species="voasary">T. voasary,</taxonomicName>
|
||||
have much larger eyes (OI 20-24) and differently shaped petiolar nodes, and are thus not easy to confuse with
|
||||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum.</taxonomicName>
|
||||
<taxonomicName id="4EABE52593B43293D6E5965188D7DF2A" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="isectum">T. isectum.</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection type="materials_examined">
|
||||
<paragraph box="[118,295,1074,1093]" pageId="38" pageNumber="39" type="mainText">Material examined</paragraph>
|
||||
<paragraph box="[118,1122,1103,1572]" pageId="38" pageNumber="39" type="mainText">
|
||||
<subSubSection id="9486CF626BA2143AC4E2D5F042C3BC44" type="materials_examined">
|
||||
<paragraph id="85DE7BD6C7AE6EF31B7217260A79D9C5" box="[118,295,1074,1093]" pageId="38" pageNumber="39" type="mainText">Material examined</paragraph>
|
||||
<paragraph id="BBD9583164F27F58D03AD0D571B097BE" box="[118,1122,1103,1572]" pageId="38" pageNumber="39" type="mainText">
|
||||
MADAGASCAR: Mahajanga,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="8C2CCD241F43F58CF0F7BE0CD1D71CEF" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="A9B2DE73710F9D4A5B3D4FCC2D6EC5FF" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Marotandrano, Marotandrano 48.3 km S Mandritsara, 16.2832 S, 48.8144 E, 865 m, transition humid forest, 6.-8.XII.2007 (B.L. Fisher et al.); Toamasina,
|
||||
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
|
||||
<normalizedToken id="52975E2303C37DFF6746A61EDE8C28A6" originalValue="Réserve">Reserve</normalizedToken>
|
||||
<normalizedToken id="FA715865D6EEDF7A6C9A3B7CC7CFD241" originalValue="Spéciale">Speciale</normalizedToken>
|
||||
Ambatovaky, Sandrangato river, 16.8175 S, 49.295 E, 360 m, rainforest, 25.-27.II.2010 (B.L. Fisher et al.); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.8496 S, 48.2947 E, 1010 m, montane rainforest, 3.-6.III.2007 (B.L. Fisher et al.); Toamasina, Ambatovy, 12.4 km NE Moramanga, 18.8477 S, 48.2947 E, 1000 m, montane rainforest, 5.-8.III.2007 (B.L. Fisher et al.); Toamasina, Res. Ambodiriana, 4.8 km 306° Manompana, along Manompana river, 16.6723 S, 49.7012 E, 125 m, rainforest, 18.XI.2005 (B.L. Fisher et al.); Toamasina, Amparihibe, 15° 2' S, 49° 34' E, II.-III.2003 (K.A. Jackson & D. Carpenter); Toamasina, Station
|
||||
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
|
||||
<normalizedToken id="C597BFDB6D86CF6DA3DF8BC595A75EFC" originalValue="forestière">forestiere</normalizedToken>
|
||||
Analamazaotra, Analamazaotra 1.3 km S Andasibe, 18.3847 S, 48.4127 E, 980 m, montane rainforest, 11.-13.XII.2007 (B.L. Fisher et al.); Toamasina, Parc National
|
||||
<normalizedToken originalValue="d´Andasibe-Mantadia">d'Andasibe-Mantadia</normalizedToken>
|
||||
<normalizedToken id="AABB6A1FD52CA26639D03EC037DC5534" originalValue="d´Andasibe-Mantadia">d'Andasibe-Mantadia</normalizedToken>
|
||||
,
|
||||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||||
<normalizedToken id="F686914F259631198D09C61762F98A04" originalValue="Forêt">Foret</normalizedToken>
|
||||
de Mantadia, 25.7 km 248° Moramanga, 18.814 S, 48.4303 E, 1040 m, rainforest, 14.VII.2006 (F.N. Raharimalala & B. Blaimer); Toamasina, Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo, 15.1883 S, 49.615 E, 470 m, rainforest, 8.-12.III.2003 (B.L Fisher, C. Griswold et al.); Toamasina, Beforona, 500 m, IX.1974 (A. Peyrieras); Toamasina, Parc National Mananara-Nord, 7.1 km 261° Antanambe, 16.455 S, 49.7875 E, 225 m, rainforest, 14.XI.2005 (B.L. Fisher et al.); Toamasina, Sahafina forest 11.4 km W Brickaville, 18.8144 S, 48.9621 E, 140 m, rainforest, 13.-14.XII.2007 (B.L. Fisher et al.); Toamasina, Parc National de Zahamena, Onibe River, 17.7591 S, 48.8547 E, 780 m, rainforest, rainforest, 21.-23.II.2009 (B.L. Fisher et al.); Toamasina, Parc National de Zahamena,
|
||||
</paragraph>
|
||||
<paragraph box="[118,1120,118,166]" pageId="39" pageNumber="40" type="mainText">
|
||||
<pageBreakToken pageId="39" pageNumber="40" start="start">Tetezambatana</pageBreakToken>
|
||||
<paragraph id="625674598FE6C82000DF5269440692E2" box="[118,1120,118,166]" pageId="39" pageNumber="40" type="mainText">
|
||||
<pageBreakToken id="CB048EFDF65377527946AFB81E615196" pageId="39" pageNumber="40" start="start">Tetezambatana</pageBreakToken>
|
||||
forest, near junction of Nosivola and Manakambahiny Rivers, 17.74298 S, 48.72936 E, 860 m, rainforest, 18.-19.II.2009 (B.L. Fisher et al.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
Loading…
Add table
Add a link
Reference in a new issue