diff --git a/data/03/25/87/0325879DF232FFDDFF60FB4EFCC9D80B.xml b/data/03/25/87/0325879DF232FFDDFF60FB4EFCC9D80B.xml new file mode 100644 index 00000000000..e90b77ce212 --- /dev/null +++ b/data/03/25/87/0325879DF232FFDDFF60FB4EFCC9D80B.xml @@ -0,0 +1,1133 @@ +<document id="2BA7EB8B19EECE31EBFDF3961D91A3CA" ID-DOI="10.1093/zoolinnean/zlae146" ID-ISSN="0024-4082" ID-ZooBank="8B93307-034E-4D7F-96BC-B17F8F32B122" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1732851283380" checkinUser="plazi" docAuthor="Delfino, Massimo, Cohen, Brigeưe F., Govender, Romala, Haarhoff, Pippa, Macaluso, Loredana, Marino, Liana, Maưhews, Ŋalassa, Wencker, Lukardis C. 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Box 266, 9300, Bloemfontein, South Africa & Evolutionary Studies Institute, ºniversity of the Witwatersrand, Private Bag 3, Wits, 2050, Johannesburg, South Africa & Genus DSTI * NRF Centre of Excellence in Palaeosciences, ºniversity of the Witwatersrand, Private Bag 3, Wits, 2050, Johannesburg, South Africa</mods:affiliation> +</mods:name> +<mods:name id="190E3B4B50A7C91CB3CE2D592365A797" type="personal"> +<mods:role id="DF984086C4540B08011B1D5423128D70"> +<mods:roleTerm id="F0790838C8E33D92102B59EAB705A84F">Author</mods:roleTerm> +</mods:role> +<mods:namePart id="4EF334690A99C648B944E515DD7DD929">Govender, Romala</mods:namePart> +<mods:affiliation id="FA9D4483C45BB0551A0AAB5F6EBB2C42">Iziko South African Museum, 25 Queen Victoria Street, 8001, Cape Town, South Africa & UCTBiological Sciences, University of Cape Town, Private Bag, Rondebosch, 7701, Cape Town, South Africa</mods:affiliation> +</mods:name> +<mods:name id="BA8F6F736D4C8C04207CB0E4BE484E8F" type="personal"> +<mods:role id="5CADE4201E8D8256F5C084FE09FFDF4B"> +<mods:roleTerm id="AA73B9C3FCDD2A51EF4C14D8BCAE2B0C">Author</mods:roleTerm> +</mods:role> +<mods:namePart id="5110C0E3E82C06D12D461036A47B8AC4">Haarhoff, Pippa</mods:namePart> +<mods:affiliation id="3EADF4FF3148D506551C57355F66B668">West Coast Fossil Park, P. 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captionTargetId-7="figure-7@10.[136,1467,144,1811]" captionTargetId-8="figure-113@11.[114,1458,144,1687]" captionTargetId-9="figure-312@12.[129,1473,144,1306]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-1="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-2="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." captionText-3="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." captionText-4="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." captionText-5="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." captionText-6="Figure 7. Chersina langebaanwegi, paratype SAM-PQL-21749: nuchal in external view (A); pygal in external (B) and internal (C) views; anterior portion of the plastron (and part of the nuchal) in ventral view (D); xiphiplastra in ventral (E) and dorsal (F) views. Scale bars equal 7 cm in (A, D) and 5 cm in (B, C, E, F)." captionText-7="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." captionText-8="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." captionText-9="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="2" pageNumber="3">Figs 1–10</figureCitation> +) +</paragraph> +</subSubSection> +<subSubSection id="C3966500F232FFD1FF60FB11FF22DB05" pageId="2" pageNumber="3" type="reference_group"> +<paragraph id="8B33368BF232FFD1FF60FB11FE30D823" blockId="2.[129,778,1013,1569]" box="[156,496,1263,1287]" pageId="2" pageNumber="3"> +1973 +<taxonomicName id="4C8C4D08F232FFD1FF29FB11FE8DD823" box="[213,333,1263,1287]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="undetermined"> +<emphasis id="B9F8EA99F232FFD1FF29FB11FEECD823" box="[213,300,1263,1287]" italics="true" pageId="2" pageNumber="3">Chersina</emphasis> +sp. +</taxonomicName> +- Hendey, p. 13. +</paragraph> +<paragraph id="8B33368BF232FFD1FF60FAF1FE31D802" blockId="2.[129,778,1013,1569]" box="[156,497,1294,1319]" pageId="2" pageNumber="3"> +1974 +<taxonomicName 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Hendey, p. 31. +</paragraph> +<paragraph id="8B33368BF232FFD1FF60FAB3FD5FD841" blockId="2.[129,778,1013,1569]" box="[156,671,1357,1381]" pageId="2" pageNumber="3"> +1986 +<taxonomicName id="4C8C4D08F232FFD1FF29FAB3FE8DD841" box="[213,333,1357,1381]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="undetermined"> +<emphasis id="B9F8EA99F232FFD1FF29FAB3FEECD841" box="[213,300,1357,1381]" italics="true" pageId="2" pageNumber="3">Chersina</emphasis> +sp. +</taxonomicName> +- Meylan and Auffenberg, p. 288. +</paragraph> +<paragraph id="8B33368BF232FFD1FF60FA93FDC7D8A1" blockId="2.[129,778,1013,1569]" box="[156,519,1388,1413]" pageId="2" pageNumber="3"> +1997 +<taxonomicName id="4C8C4D08F232FFD1FF29FA92FE8DD8A0" box="[213,333,1388,1412]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="undetermined"> +<emphasis id="B9F8EA99F232FFD1FF29FA92FEECD8A0" box="[213,300,1388,1412]" italics="true" pageId="2" pageNumber="3">Chersina</emphasis> +sp. +</taxonomicName> +- Broadley, p. 111. +</paragraph> +<paragraph id="8B33368BF232FFD1FF60FA72FDA7D880" blockId="2.[129,778,1013,1569]" box="[156,615,1420,1444]" pageId="2" pageNumber="3"> +2000 +<taxonomicName id="4C8C4D08F232FFD1FF29FA72FE8DD887" box="[213,333,1420,1444]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="undetermined"> +<emphasis id="B9F8EA99F232FFD1FF29FA72FEECD880" box="[213,300,1420,1444]" italics="true" pageId="2" pageNumber="3">Chersina</emphasis> +sp. +</taxonomicName> +- Lapparent de Broin, p. 64. +</paragraph> +<paragraph id="8B33368BF232FFD1FF60FA55FDEED8E7" blockId="2.[129,778,1013,1569]" box="[156,558,1451,1475]" pageId="2" pageNumber="3"> +2007 +<taxonomicName id="4C8C4D08F232FFD1FF29FA55FE8DD8E7" box="[213,333,1451,1475]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="undetermined"> +<emphasis id="B9F8EA99F232FFD1FF29FA55FEECD8E7" box="[213,300,1451,1475]" italics="true" pageId="2" pageNumber="3">Chersina</emphasis> +sp. +</taxonomicName> +- Daniels +<emphasis id="B9F8EA99F232FFD1FE4DFA52FE24D8E7" box="[433,484,1451,1475]" italics="true" pageId="2" pageNumber="3">et al.,</emphasis> +p. 916. +</paragraph> +<paragraph id="8B33368BF232FFD1FF60FA35FDF5D8C7" blockId="2.[129,778,1013,1569]" box="[156,565,1482,1507]" pageId="2" pageNumber="3"> +2021 +<taxonomicName id="4C8C4D08F232FFD1FF29FA34FE8DD8C6" box="[213,333,1482,1506]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="undetermined"> +<emphasis id="B9F8EA99F232FFD1FF29FA34FEECD8C6" box="[213,300,1482,1506]" italics="true" pageId="2" pageNumber="3">Chersina</emphasis> +sp. +</taxonomicName> +- Georgalis +<emphasis id="B9F8EA99F232FFD1FE3AFA35FE39D8C6" box="[454,505,1482,1506]" italics="true" pageId="2" pageNumber="3">et al.,</emphasis> +p. 49. +</paragraph> +<paragraph id="8B33368BF232FFD1FF60FA14FF22DB05" blockId="2.[129,778,1013,1569]" pageId="2" pageNumber="3"> +2023 +<taxonomicName id="4C8C4D08F232FFD1FF29FA14FD08DB26" authority="(Duméril in Schweigger. 1812)" baseAuthorityName="Duméril in Schweigger" baseAuthorityYear="1812" box="[213,712,1514,1538]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F232FFD1FF29FA14FE47DB26" box="[213,391,1514,1538]" italics="true" pageId="2" pageNumber="3">Chersina angulata</emphasis> +(Duméril in Schweigger. 1812) +</taxonomicName> +- Bhat +<emphasis id="B9F8EA99F232FFD1FF7DF9F4FF73DB05" box="[129,179,1545,1569]" italics="true" pageId="2" pageNumber="3">et al.,</emphasis> +p. 2. +</paragraph> +</subSubSection> +<subSubSection id="C3966500F232FFD1FF7DF9BDFD51DA71" pageId="2" pageNumber="3" type="etymology"> +<paragraph id="8B33368BF232FFD1FF7DF9BDFD51DA71" blockId="2.[129,778,1602,1877]" pageId="2" pageNumber="3"> +<emphasis id="B9F8EA99F232FFD1FF7DF9BDFEFCDB7E" box="[129,316,1603,1627]" italics="true" pageId="2" pageNumber="3">Derivation of name:</emphasis> +The species name,meaning ‘of Langebaanweg’, derives from the site where it was collected, and is currently the only site at which it has been identified. Worth noting is that turtles represent the most common vertebrate taxon at Langebaanweg, and that this species is by far the most common taxon in the turtle assemblage. This species is dedicated to Prof. Margaretha D. Hofmeyr (1950–2020) for her commitment to the study of African extant tortoises and for having promoted the study of the fossil tortoises from Langebaanweg. +</paragraph> +</subSubSection> +<subSubSection id="C3966500F232FFD1FF7DF888FE73DA8A" pageId="2" pageNumber="3" type="description"> +<paragraph id="8B33368BF232FFD1FF7DF888FE73DA8A" blockId="2.[129,777,1910,1966]" pageId="2" pageNumber="3"> +<emphasis id="B9F8EA99F232FFD1FF7DF888FEEEDAAA" box="[129,302,1910,1934]" italics="true" pageId="2" pageNumber="3">Zoobank LSID:</emphasis> +urn:lsid:zoobank.org:pub:A8B93307-034E - 4D7F -96BC -B17F8F32B122. +</paragraph> +</subSubSection> +<subSubSection id="C3966500F232FFD0FCC5FF6EFD5BDB6A" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined"> +<paragraph id="8B33368BF232FFD1FCC5FF6EFC6CDDE3" blockId="2.[825,1474,143,199]" pageId="2" pageNumber="3"> +<materialsCitation id="3BE43CD6F232FFD1FCC5FF6EFC68DDE3" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-20734" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape" typeStatus="holotype"> +<emphasis id="B9F8EA99F232FFD1FCC5FF6EFC5ADD8C" box="[825,922,144,168]" italics="true" pageId="2" pageNumber="3"> +<typeStatus id="54378829F232FFD1FCC5FF6EFC55DD8C" box="[825,917,144,168]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus> +: +</emphasis> +<emphasis id="B9F8EA99F232FFD1FC55FF71FBADDD83" bold="true" box="[937,1133,143,167]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC55FF71FBADDD83" box="[937,1133,143,167]" pageId="2" pageNumber="3">SAM-PQL-20734</specimenCode> +</emphasis> +: nearly complete +<specimenCount id="9D8AFD02F232FFD1FAC3FF6EFA43DD8C" box="[1343,1411,144,168]" pageId="2" pageNumber="3" type="female">female</specimenCount> +shell ( +<figureCitation id="13B72A0EF232FFD1FCB8FF51FC5CDDE3" box="[836,924,175,199]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[113,178,1312,1336]" captionStartId-1="4.[129,194,1394,1418]" captionStartId-2="5.[113,178,1047,1071]" captionTargetBox-0="[117,1455,147,1281]" captionTargetBox-1="[130,1471,145,1364]" captionTargetBox-2="[147,1424,145,1017]" captionTargetId-0="figure-340@3.[114,1458,144,1284]" captionTargetId-1="figure-300@4.[129,1473,144,1366]" captionTargetId-2="figure-538@5.[146,1426,144,1019]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-1="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-2="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="2" pageNumber="3">Figs 1–3</figureCitation> +) +</materialsCitation> +. +</paragraph> +<paragraph id="8B33368BF232FFD1FCC5FF17FC1CDF93" blockId="2.[825,1474,232,695]" pageId="2" pageNumber="3"> +<materialsCitation id="3BE43CD6F232FFD1FCC5FF17FA02DC04" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-20925" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype"> +<emphasis id="B9F8EA99F232FFD1FCC5FF17FC60DC24" box="[825,928,233,256]" italics="true" pageId="2" pageNumber="3"> +<typeStatus id="54378829F232FFD1FCC5FF17FC5BDC24" box="[825,923,233,256]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus> +: +</emphasis> +<emphasis id="B9F8EA99F232FFD1FC55FF16FBABDC24" bold="true" box="[937,1131,232,256]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC55FF16FBABDC24" box="[937,1131,232,256]" pageId="2" pageNumber="3">SAM-PQL-20925</specimenCode> +</emphasis> +: partial +<specimenCount id="9D8AFD02F232FFD1FB3AFF16FB37DC24" box="[1222,1271,232,256]" pageId="2" pageNumber="3" type="male">male</specimenCount> +shell with a nearly complete plastron, several peripherals and pygal ( +<figureCitation id="13B72A0EF232FFD1FA91FEF6FA70DC04" box="[1389,1456,264,288]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="2" pageNumber="3">Fig. 4</figureCitation> +); +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FCC5FED9FA02DC7B" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-21109" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype"> +<emphasis id="B9F8EA99F232FFD1FCC5FED9FC3CDC1B" bold="true" box="[825,1020,295,319]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FCC5FED9FC3CDC1B" box="[825,1020,295,319]" pageId="2" pageNumber="3">SAM-PQL-21109</specimenCode> +</emphasis> +: partially preserved shell of a +<specimenCount id="9D8AFD02F232FFD1FAA5FED9FA4ADC1B" box="[1369,1418,295,319]" pageId="2" pageNumber="3" type="male">male</specimenCount> +spe - cimen missing most of the costals and all the neurals ( +<figureCitation id="13B72A0EF232FFD1FA89FEB8FA70DC7A" box="[1397,1456,326,350]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="2" pageNumber="3">Fig. 5</figureCitation> +); +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FCC5FE9BFB59DC99" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-21174" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype"> +<emphasis id="B9F8EA99F232FFD1FCC5FE9BFC3CDC59" bold="true" box="[825,1020,357,381]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FCC5FE9BFC3CDC59" box="[825,1020,357,381]" pageId="2" pageNumber="3">SAM-PQL-21174</specimenCode> +</emphasis> +: highly fragmentary, incomplete shell whose best -preserved elements are the nuchal, fifth neural, pygal, and, above all, both epiplastra ( +<figureCitation id="13B72A0EF232FFD1FBB1FE5AFB47DC98" box="[1101,1159,420,444]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="2" pageNumber="3">Fig. 6</figureCitation> +); +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB5CFE5AFB67DF3F" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-21749" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype"> +<emphasis id="B9F8EA99F232FFD1FB5CFE5AFAA2DC98" bold="true" box="[1184,1378,420,444]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB5CFE5AFAA2DC98" box="[1184,1378,420,444]" pageId="2" pageNumber="3">SAM-PQL-21749</specimenCode> +</emphasis> +: crushed but nearly complete shell with partially articulated elements, including well -preserved and well -exposed nuchal, pygal, epiplastra, and xiphiplastra ( +<figureCitation id="13B72A0EF232FFD1FBA0FDFCFB55DF3E" box="[1116,1173,514,538]" captionStart="Figure 7" captionStartId="9.[114,179,1706,1730]" captionTargetBox="[115,1436,145,1676]" captionTargetId="figure-121@9.[114,1458,144,1678]" captionTargetPageId="9" captionText="Figure 7. Chersina langebaanwegi, paratype SAM-PQL-21749: nuchal in external view (A); pygal in external (B) and internal (C) views; anterior portion of the plastron (and part of the nuchal) in ventral view (D); xiphiplastra in ventral (E) and dorsal (F) views. Scale bars equal 7 cm in (A, D) and 5 cm in (B, C, E, F)." pageId="2" pageNumber="3">Fig. 7</figureCitation> +); +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB51FDFCFB99DF1E" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-22076" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="paratype"> +<emphasis id="B9F8EA99F232FFD1FB51FDFCFAAFDF3E" bold="true" box="[1197,1391,514,538]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB51FDFCFAAFDF3E" box="[1197,1391,514,538]" pageId="2" pageNumber="3">SAM-PQL-22076</specimenCode> +</emphasis> +: a com - plete +<specimenCount id="9D8AFD02F232FFD1FC8EFDDCFC63DF1E" box="[882,931,546,570]" pageId="2" pageNumber="3" type="male">male</specimenCount> +plastron ( +<figureCitation id="13B72A0EF232FFD1FBF2FDDCFB87DF1E" box="[1038,1095,546,570]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="2" pageNumber="3">Fig. 8</figureCitation> +); +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB9CFDDFFC17DF93" country="South Africa" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" pageId="2" pageNumber="3" specimenCode="SAM-PQL-35023d" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype"> +<emphasis id="B9F8EA99F232FFD1FB9CFDDFFAF1DF1D" bold="true" box="[1120,1329,545,569]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB9CFDDFFAF1DF1D" box="[1120,1329,545,569]" pageId="2" pageNumber="3">SAM-PQL-35023d</specimenCode> +</emphasis> +: partially pre - served shell of a very large +<specimenCount id="9D8AFD02F232FFD1FBA8FDBFFB75DF7D" box="[1108,1205,577,601]" pageId="2" pageNumber="3" type="generic">specimen</specimenCount> +represented mostly by an epiplastra, entoplastron, left hyoplastron, three neurals, pygal with the surrounding fragmentary peripherals, and three peri - pherals ( +<figureCitation id="13B72A0EF232FFD1FC6FFD61FC0BDF93" box="[915,971,671,695]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="2" pageNumber="3">Fig. 9</figureCitation> +) +</materialsCitation> +. +</paragraph> +<paragraph id="8B33368BF232FFD1FCC5FD27FC73DB59" blockId="2.[825,1474,728,1661]" pageId="2" pageNumber="3"> +<emphasis id="B9F8EA99F232FFD1FCC5FD27FBC5DFD4" box="[825,1029,729,753]" italics="true" pageId="2" pageNumber="3">Referred specimens:</emphasis> +<materialsCitation id="3BE43CD6F232FFD1FBE5FD26FA02DFD5" box="[1049,1474,728,753]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-12740a" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FBE5FD26FB27DFD4" bold="true" box="[1049,1255,728,752]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FBE5FD26FB27DFD4" box="[1049,1255,728,752]" pageId="2" pageNumber="3">SAM-PQL-12740a</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FCC5FD06FB77DE34" box="[825,1207,760,784]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-12750c" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FCC5FD06FBC7DE34" bold="true" box="[825,1031,760,784]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FCC5FD06FBC7DE34" box="[825,1031,760,784]" pageId="2" pageNumber="3">SAM-PQL-12750c</specimenCode> +</emphasis> +: left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB42FD06FB1DDE0B" pageId="2" pageNumber="3" specimenCode="SAM-PQL-12754a" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB42FD06FA4CDE34" bold="true" box="[1214,1420,760,784]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB42FD06FA4CDE34" box="[1214,1420,760,784]" pageId="2" pageNumber="3">SAM-PQL-12754a</specimenCode> +</emphasis> +: two epiplastra sutured to the entoplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB16FCE9FC21DE6B" pageId="2" pageNumber="3" specimenCode="SAM-PQL-12754b" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB16FCE9FA7ADE0B" bold="true" box="[1258,1466,791,815]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB16FCE9FA7ADE0B" box="[1258,1466,791,815]" pageId="2" pageNumber="3">SAM-PQL-12754b</specimenCode> +</emphasis> +: left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC11FCC8FAB7DE6B" box="[1005,1399,822,847]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-12754c" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC11FCC8FB7BDE6A" bold="true" box="[1005,1211,822,846]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC11FCC8FB7BDE6A" box="[1005,1211,822,846]" pageId="2" pageNumber="3">SAM-PQL-12754c</specimenCode> +</emphasis> +: left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FA7FFCC8FB65DE4A" collectionCode="SAM" pageId="2" pageNumber="3" specimenCode="SAM- PQL-24826b" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FA7FFCC8FC0BDE4A" bold="true" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FA7FFCC8FC0BDE4A" pageId="2" pageNumber="3">SAM-PQL-24826b</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB44FCA8FC74DEA9" pageId="2" pageNumber="3" specimenCode="SAM-PQL-24827a" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB44FCA8FA46DE4A" bold="true" box="[1208,1414,854,878]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB44FCA8FA46DE4A" box="[1208,1414,854,878]" pageId="2" pageNumber="3">SAM-PQL-24827a</specimenCode> +</emphasis> +: left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC3CFC8BFA02DEA9" box="[960,1474,885,909]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-24862a" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC3CFC8BFB4EDEA9" bold="true" box="[960,1166,885,909]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC3CFC8BFB4EDEA9" box="[960,1166,885,909]" pageId="2" pageNumber="3">SAM-PQL-24862a</specimenCode> +</emphasis> +: right and left epiplastrons; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FCC5FC6AFB25DE89" box="[825,1253,916,941]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-24862b" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FCC5FC6AFBC9DE88" bold="true" box="[825,1033,916,940]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FCC5FC6AFBC9DE88" box="[825,1033,916,940]" pageId="2" pageNumber="3">SAM-PQL-24862b</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB04FC6AFC2DDEE8" pageId="2" pageNumber="3" specimenCode="SAM-PQL-52640" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB04FC6AFA7BDE88" bold="true" box="[1272,1467,916,940]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB04FC6AFA7BDE88" box="[1272,1467,916,940]" pageId="2" pageNumber="3">SAM-PQL-52640</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC09FC4AFABBDEE8" box="[1013,1403,948,972]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73745" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC09FC4AFB78DEE8" bold="true" box="[1013,1208,948,972]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC09FC4AFB78DEE8" box="[1013,1208,948,972]" pageId="2" pageNumber="3">SAM-PQL-73745</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FA7FFC4AFBE0DECF" collectionCode="SAM" pageId="2" pageNumber="3" specimenCode="SAM- PQL-73747" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FA7FFC4AFC7DDECF" bold="true" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FA7FFC4AFC7DDECF" pageId="2" pageNumber="3">SAM-PQL-73747</specimenCode> +</emphasis> +: nuchal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FBD3FC2DFA02DECF" box="[1071,1474,979,1003]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73749" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FBD3FC2DFB31DECF" bold="true" box="[1071,1265,979,1003]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FBD3FC2DFB31DECF" box="[1071,1265,979,1003]" pageId="2" pageNumber="3">SAM-PQL-73749</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FCC5FC0CFB78D92F" box="[825,1208,1010,1035]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73750" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FCC5FC0CFC3CD92E" bold="true" box="[825,1020,1010,1034]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FCC5FC0CFC3CD92E" box="[825,1020,1010,1034]" pageId="2" pageNumber="3">SAM-PQL-73750</specimenCode> +</emphasis> +, left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB3AFC0CFCABD90E" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73751" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB3AFC0CFA52D92E" bold="true" box="[1222,1426,1010,1034]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB3AFC0CFA4DD92E" box="[1222,1421,1010,1034]" pageId="2" pageNumber="3">SAM-PQL-73751</specimenCode> +: +</emphasis> +nu - chal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC8AFBECFB57D90E" box="[886,1175,1042,1066]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73754" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC8AFBECFBF8D90E" bold="true" box="[886,1080,1042,1066]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC8AFBECFBF8D90E" box="[886,1080,1042,1066]" pageId="2" pageNumber="3">SAM-PQL-73754</specimenCode> +</emphasis> +: nuchal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB5DFBECFA02D90E" box="[1185,1474,1042,1066]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73757" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB5DFBECFAA3D90E" bold="true" box="[1185,1379,1042,1066]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB5DFBECFAA3D90E" box="[1185,1379,1042,1066]" pageId="2" pageNumber="3">SAM-PQL-73757</specimenCode> +</emphasis> +: nuchal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FCC5FBCFFB7BD96D" box="[825,1211,1073,1097]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73761" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FCC5FBCFFC3BD96D" bold="true" box="[825,1019,1073,1097]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FCC5FBCFFC3BD96D" box="[825,1019,1073,1097]" pageId="2" pageNumber="3">SAM-PQL-73761</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB3DFBCFFC74D94C" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73763" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB3DFBCFFA44D96D" bold="true" box="[1217,1412,1073,1097]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB3DFBCFFA44D96D" box="[1217,1412,1073,1097]" pageId="2" pageNumber="3">SAM-PQL-73763</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC3FFBAEFB29D94C" box="[963,1257,1104,1128]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73765" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC3FFBAEFB46D94C" bold="true" box="[963,1158,1104,1128]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC3FFBAEFB46D94C" box="[963,1158,1104,1128]" pageId="2" pageNumber="3">SAM-PQL-73765</specimenCode> +</emphasis> +: nuchal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB04FBAEFC02D9AC" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73769" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB04FBAEFA7BD94C" bold="true" box="[1272,1467,1104,1128]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB04FBAEFA7BD94C" box="[1272,1467,1104,1128]" pageId="2" pageNumber="3">SAM-PQL-73769</specimenCode> +</emphasis> +: entoplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC31FB8EFB2ED9AC" box="[973,1262,1136,1160]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73777" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC31FB8EFB4FD9AC" bold="true" box="[973,1167,1136,1160]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC31FB8EFB4FD9AC" box="[973,1167,1136,1160]" pageId="2" pageNumber="3">SAM-PQL-73777</specimenCode> +</emphasis> +: nuchal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB05FB8EFC20D983" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73781" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB05FB8EFA7AD9AC" bold="true" box="[1273,1466,1136,1160]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB05FB8EFA7AD9AC" box="[1273,1466,1136,1160]" pageId="2" pageNumber="3">SAM-PQL-73781</specimenCode> +</emphasis> +: left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC17FB71FAB8D983" box="[1003,1400,1167,1191]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73789" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC17FB71FB6ED983" bold="true" box="[1003,1198,1167,1191]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC17FB71FB6ED983" box="[1003,1198,1167,1191]" pageId="2" pageNumber="3">SAM-PQL-73789</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FA7FFB71FBA8D9E2" collectionCode="SAM" pageId="2" pageNumber="3" specimenCode="SAM- PQL-73792" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FA7FFB71FC7DD9E2" bold="true" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FA7FFB71FC7DD9E2" pageId="2" pageNumber="3">SAM-PQL-73792</specimenCode> +</emphasis> +: left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB90FB50FA02D9E2" box="[1132,1474,1198,1222]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73795" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB90FB50FAEED9E2" bold="true" box="[1132,1326,1198,1222]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB90FB50FAEED9E2" box="[1132,1326,1198,1222]" pageId="2" pageNumber="3">SAM-PQL-73795</specimenCode> +</emphasis> +: entoplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FCC5FB30FB7BD9C2" box="[825,1211,1230,1254]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73799" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FCC5FB30FC3CD9C2" bold="true" box="[825,1020,1230,1254]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FCC5FB30FC3CD9C2" box="[825,1020,1230,1254]" pageId="2" pageNumber="3">SAM-PQL-73799</specimenCode> +</emphasis> +: left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB35FB30FC74D821" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73803" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB35FB30FA4CD9C2" bold="true" box="[1225,1420,1230,1254]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB35FB30FA4CD9C2" box="[1225,1420,1230,1254]" pageId="2" pageNumber="3">SAM-PQL-73803</specimenCode> +</emphasis> +: left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC34FB13FB24D821" box="[968,1252,1261,1285]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73807" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC34FB13FB4AD821" bold="true" box="[968,1162,1261,1285]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC34FB13FB4AD821" box="[968,1162,1261,1285]" pageId="2" pageNumber="3">SAM-PQL-73807</specimenCode> +</emphasis> +: pygal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB04FB13FC46D800" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73808" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB04FB13FA7BD821" bold="true" box="[1272,1467,1261,1285]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB04FB13FA7BD821" box="[1272,1467,1261,1285]" pageId="2" pageNumber="3">SAM-PQL-73808</specimenCode> +</emphasis> +: nuchal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC55FAF2FB15D800" box="[937,1237,1292,1316]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73809" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC55FAF2FBACD800" bold="true" box="[937,1132,1292,1316]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC55FAF2FBACD800" box="[937,1132,1292,1316]" pageId="2" pageNumber="3">SAM-PQL-73809</specimenCode> +</emphasis> +: pygal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB04FAF2FC38D860" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73811" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB04FAF2FA7AD800" bold="true" box="[1272,1466,1292,1316]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB04FAF2FA7AD800" box="[1272,1466,1292,1316]" pageId="2" pageNumber="3">SAM-PQL-73811</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FBF7FAD5FAB0D860" box="[1035,1392,1323,1348]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73819" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FBF7FAD5FB0DD867" bold="true" box="[1035,1229,1323,1347]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FBF7FAD5FB0DD867" box="[1035,1229,1323,1347]" pageId="2" pageNumber="3">SAM-PQL-73819</specimenCode> +</emphasis> +: entoplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FA7FFAD5FB5CD847" collectionCode="SAM" pageId="2" pageNumber="3" specimenCode="SAM- PQL-73825" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FA7FFAD5FC7DD847" bold="true" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FA7FFAD5FC7DD847" pageId="2" pageNumber="3">SAM-PQL-73825</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB4EFAB5FC74D8A6" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73828" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB4EFAB5FAB4D847" bold="true" box="[1202,1396,1355,1379]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB4EFAB5FAB4D847" box="[1202,1396,1355,1379]" pageId="2" pageNumber="3">SAM-PQL-73828</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC21FA94FA9AD8A6" box="[989,1370,1386,1410]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73829" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC21FA94FB60D8A6" bold="true" box="[989,1184,1386,1410]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC21FA94FB60D8A6" box="[989,1184,1386,1410]" pageId="2" pageNumber="3">SAM-PQL-73829</specimenCode> +</emphasis> +: entoplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FA7FFA94FBECD886" collectionCode="SAM" pageId="2" pageNumber="3" specimenCode="SAM- PQL-73831" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FA7FFA94FC7DD886" bold="true" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FA7FFA94FC7DD886" pageId="2" pageNumber="3">SAM-PQL-73831</specimenCode> +</emphasis> +: nuchal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FBBAFA77FAA9D886" box="[1094,1385,1417,1442]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73832" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FBBAFA77FAC8D885" bold="true" box="[1094,1288,1417,1441]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FBBAFA77FAC8D885" box="[1094,1288,1417,1441]" pageId="2" pageNumber="3">SAM-PQL-73832</specimenCode> +</emphasis> +: pygal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FA7FFA77FB64D8E5" collectionCode="SAM" pageId="2" pageNumber="3" specimenCode="SAM- PQL-73834" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FA7FFA77FC7DD8E5" bold="true" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FA7FFA77FC7DD8E5" pageId="2" pageNumber="3">SAM-PQL-73834</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB42FA57FC74D8C4" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73836" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB42FA57FA40D8E5" bold="true" box="[1214,1408,1449,1473]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB42FA57FA40D8E5" box="[1214,1408,1449,1473]" pageId="2" pageNumber="3">SAM-PQL-73836</specimenCode> +</emphasis> +: left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC32FA36FAA9D8C4" box="[974,1385,1480,1504]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73837" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC32FA36FB51D8C4" bold="true" box="[974,1169,1480,1504]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC32FA36FB51D8C4" box="[974,1169,1480,1504]" pageId="2" pageNumber="3">SAM-PQL-73837</specimenCode> +</emphasis> +: left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FA7FFA36FB54DB24" collectionCode="SAM" pageId="2" pageNumber="3" specimenCode="SAM- PQL-73845" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FA7FFA36FC7DD8DB" bold="true" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FA7FFA36FC7DD8DB" pageId="2" pageNumber="3">SAM-PQL-73845</specimenCode> +</emphasis> +: left epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB51FA19FC74DB3B" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73860" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB51FA19FAB0D8DB" bold="true" box="[1197,1392,1511,1535]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB51FA19FAB0D8DB" box="[1197,1392,1511,1535]" pageId="2" pageNumber="3">SAM-PQL-73860</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FC21F9F9FA9ADB3B" box="[989,1370,1543,1567]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73862" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FC21F9F9FB60DB3B" bold="true" box="[989,1184,1543,1567]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FC21F9F9FB60DB3B" box="[989,1184,1543,1567]" pageId="2" pageNumber="3">SAM-PQL-73862</specimenCode> +</emphasis> +: entoplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FA7FF9F9FBCEDB1A" collectionCode="SAM" pageId="2" pageNumber="3" specimenCode="SAM- PQL-73863" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FA7FF9F9FC7DDB1A" bold="true" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FA7FF9F9FC7DDB1A" pageId="2" pageNumber="3">SAM-PQL-73863</specimenCode> +</emphasis> +: pygal; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FBE6F9D8FAB8DB1A" box="[1050,1400,1574,1598]" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73876" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FBE6F9D8FB1CDB1A" bold="true" box="[1050,1244,1574,1598]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FBE6F9D8FB1CDB1A" box="[1050,1244,1574,1598]" pageId="2" pageNumber="3">SAM-PQL-73876</specimenCode> +</emphasis> +: entoplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FA7FF9D8FB5CDB7A" collectionCode="SAM" pageId="2" pageNumber="3" specimenCode="SAM- PQL-73877" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FA7FF9D8FC7DDB79" bold="true" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FA7FF9D8FC7DDB79" pageId="2" pageNumber="3">SAM-PQL-73877</specimenCode> +</emphasis> +: right epiplastron; +</materialsCitation> +<materialsCitation id="3BE43CD6F232FFD1FB4EF9BBFC6EDB59" pageId="2" pageNumber="3" specimenCode="SAM-PQL-73878" specimenCount="1"> +<emphasis id="B9F8EA99F232FFD1FB4EF9BBFAB4DB79" bold="true" box="[1202,1396,1605,1629]" pageId="2" pageNumber="3"> +<specimenCode id="DB2A9EF0F232FFD1FB4EF9BBFAB4DB79" box="[1202,1396,1605,1629]" pageId="2" pageNumber="3">SAM-PQL-73878</specimenCode> +</emphasis> +: right epiplastron +</materialsCitation> +. +</paragraph> +<paragraph id="8B33368BF232FFD1FCC5F961FBD3DA31" blockId="2.[825,1475,1694,1969]" pageId="2" pageNumber="3"> +<emphasis id="B9F8EA99F232FFD1FCC5F961FBCBDB92" box="[825,1035,1694,1718]" italics="true" pageId="2" pageNumber="3">Type locality and age:</emphasis> +‘E’ Quarry, +<location id="8E536050F232FFD1FB79F960FADCDB92" LSID="urn:lsid:plazi:treatment:0325879DF232FFDDFF60FB4EFCC9D80B:8E536050F232FFD1FB79F960FADCDB92" box="[1157,1308,1694,1718]" country="South Africa" county="Mya" latitude="-32.966667" longLatPrecision="1207" longitude="18.15" municipality="Cape Town" name="Langebaanweg" pageId="2" pageNumber="3" stateProvince="Western Cape">Langebaanweg</location> +( +<geoCoordinate id="EEB8504CF232FFD1FAD6F960FAB7DB92" box="[1322,1399,1694,1718]" degrees="18" direction="east" minutes="9" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="18.15">18°9’ E</geoCoordinate> +, +<geoCoordinate id="EEB8504CF232FFD1FA7CF960FC89DBF2" degrees="32" direction="south" minutes="58" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="-32.966667">32°58’ S</geoCoordinate> +), approximately +<quantity id="4C749B6EF232FFD1FC04F940FB87DBF2" box="[1016,1095,1726,1750]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="3" unit="km" value="110.0"> +<locationDeviation id="9A812FA9F232FFD1FC04F940FB87DBF2" box="[1016,1095,1726,1750]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="3" unit="km" value="110.0">110 km</locationDeviation> +</quantity> +NNW of +<collectingMunicipality id="6B57ACF1F232FFD1FB4FF940FAEEDBF1" box="[1203,1326,1726,1750]" pageId="2" pageNumber="3">Cape Town</collectingMunicipality> +, +<collectingCountry id="F39B761BF232FFD1FAC5F940FA7FDBF2" box="[1337,1471,1726,1750]" name="South Africa" pageId="2" pageNumber="3">South Africa</collectingCountry> +. +<geologicalTimeScale id="F9ACB620F232FFD1FCC5F923FAD2DBD1" box="[825,1298,1757,1781]" pageId="2" pageNumber="3">Upper Varswater Formation; Early Pliocene</geologicalTimeScale> +, about 5.1 Mya ( +<bibRefCitation id="EF1D4B7AF232FFD1FCB8F903FBC2DA31" author="Roberts DL & Mauhews T & Herries AIR" box="[836,1026,1789,1813]" pageId="2" pageNumber="3" pagination="191 - 214" refId="ref15165" refString="Roberts DL, Mauhews T, Herries AIR et al. Regional and global context of the Late Cenozoic Langebaanweg (LBW) palaeontological site: west coast of South Africa. Earth-Science Reviews 2011; 106: 191 - 214." type="journal article" year="2011"> +Roberts +<emphasis id="B9F8EA99F232FFD1FC67F903FC0ADA31" box="[923,970,1789,1813]" italics="true" pageId="2" pageNumber="3">et al.</emphasis> +2011 +</bibRefCitation> +). +</paragraph> +<paragraph id="8B33368BF232FFD0FCA9F8E2FD5BDB6A" blockId="2.[825,1475,1694,1969]" lastBlockId="3.[113,763,1496,1614]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3"> +The fossil vertebrates of this locality have been studied since the late 1950s ( +<bibRefCitation id="EF1D4B7AF232FFD1FBE4F8C5FADADA77" author="Singer R & Hooijer DA" box="[1048,1306,1851,1875]" pageId="2" pageNumber="3" pagination="101 - 2" refId="ref15307" refString="Singer R, Hooijer DA. A Stegolophodon from South Africa. Nature 1958; 182: 101 - 2. hups: // doi. org / 10.1038 / 182101 a 0" type="journal article" year="1958">Singer and Hooijer 1958</bibRefCitation> +, +<bibRefCitation id="EF1D4B7AF232FFD1FADBF8C5FA7CDA77" author="Simpson GG" box="[1319,1468,1851,1875]" pageId="2" pageNumber="3" pagination="1144 - 5" refId="ref15271" refString="Simpson GG. Fossil penguin from the Late Cenozoic of South Africa. Science 1971; 171: 1144 - 5. hups: // doi. org / 10.1126 / science. 171. 3976.1144" type="journal article" year="1971">Simpson 1971</bibRefCitation> +, +<bibRefCitation id="EF1D4B7AF232FFD1FCC5F8A5FC07DA57" author="Hendey QB" box="[825,967,1883,1907]" pageId="2" pageNumber="3" pagination="13 - 4" refId="ref14417" refString="Hendey QB. Fossil occurrences at Langebaanweg, Cape Province. Nature 1973; 244: 13 - 4. hups: // doi. org / 10.1038 / 244013 a 0" type="journal article" year="1973">Hendey 1973</bibRefCitation> +, +<bibRefCitation id="EF1D4B7AF232FFD1FC29F8A5FBD4DA57" author="Hendey QB" box="[981,1044,1883,1907]" pageId="2" pageNumber="3" pagination="1 - 369" refId="ref14449" refString="Hendey QB. Ne Late Cenozoic carnivora of the south * western Cape Province. Annals of the South Atican Museum 1974 a; 63: 1 - 369." type="journal article" year="1974">1974a</bibRefCitation> +, b, 1976) and have been still actively in - vestigated in recent years (among others see +<bibRefCitation id="EF1D4B7AF232FFD1FAD4F884FC4DDA95" author="Groenewald PA & Sealy J & Stynder D" pageId="2" pageNumber="3" refId="ref14351" refString="Groenewald PA, Sealy J, Stynder D et al. Dietary resource partitioning among three coeval proboscidean taxa (Anancus capensis, Mammuthus subplanitons, Looodonta cookei) from the South African Early Pliocene locality of Langebaanweg E Quarry. Palaeogeography, Palaeoclimatology, Palaeoecology 2020; 543: 109606. hups: // doi. org / 10.1016 / j. palaeo. 2020.109606" type="journal volume" year="2020"> +Groenewald +<emphasis id="B9F8EA99F232FFD1FA4DF885FC92DA95" italics="true" pageId="2" pageNumber="3">et al.</emphasis> +2020 +</bibRefCitation> +, +<bibRefCitation id="EF1D4B7AF232FFD1FC67F867FB31DA95" author="Valenciano A & Govender R" box="[923,1265,1945,1969]" pageId="2" pageNumber="3" refId="ref15686" refString="Valenciano A, Govender R. New fossils of Mellivora benfieldi (Mammalia, Carnivora, Mustelidae) from Langebaanweg, ' E' Quarry (South Africa, Early Pliocene): re * evaluation of the African Neogene mellivorines. Journal of Vertebrate Paleontology 2020 a; 40: e 1817754 - e 1817754. hups: // doi. org / 10.1080 / 02724634.2020.1817754" type="journal volume" year="2020">Valenciano and Govender 2020a</bibRefCitation> +, b, +<bibRefCitation id="EF1D4B7AF232FFD1FAE5F867FA7CDA95" author="Govender R" box="[1305,1468,1945,1969]" pageId="2" pageNumber="3" refId="ref14214" refString="Govender R. Shark * seal interaction off South Africa's west coast during the Early Pliocene (5 Ma). South Atican Journal of Science 2021; 117: 8837." type="journal volume" year="2021">Govender 2021</bibRefCitation> +, Nacarino -Meneses and Chinsamy 2022, +<bibRefCitation id="EF1D4B7AF233FFD0FDC9FA26FD35D8D4" author="Rabe C & Chinsamy A & Valenciano A" box="[565,757,1496,1520]" pageId="3" pageNumber="4" refId="ref15121" refString="Rabe C, Chinsamy A, Valenciano A. Taxonomic and palaeobiological implications of a large, pathological sabretooth (Carnivora, Felidae, Machairodontinae) from the Lower Pliocene of South Africa. Papers in Palaeontology 2022; 8: e 1463." type="journal volume" year="2022"> +Rabe +<emphasis id="B9F8EA99F233FFD0FD86FA27FD72D8D4" box="[634,690,1496,1520]" italics="true" pageId="3" pageNumber="4">et al.</emphasis> +2022 +</bibRefCitation> +, +<bibRefCitation id="EF1D4B7AF233FFD0FF8DFA09FEA3DB2B" author="Valenciano A & Morales J & Govender R" box="[113,355,1527,1551]" pageId="3" pageNumber="4" pagination="366 - 94" refId="ref15809" refString="Valenciano A, Morales J, Govender R. Eucyon khoikhoi sp. nov. (Carnivora: Canidae) from Langebaanweg ' E' Quarry (Early Pliocene, South Africa): the most complete African canini from the Mio * Pliocene. Zoological Journal of the Linnean Society 2022; 194: 366 - 94." type="journal article" year="2022"> +Valenciano +<emphasis id="B9F8EA99F233FFD0FF12FA06FEE4DB2B" box="[238,292,1527,1551]" italics="true" pageId="3" pageNumber="4">et al.</emphasis> +2022 +</bibRefCitation> +, +<bibRefCitation id="EF1D4B7AF233FFD0FE88FA09FD90DB2B" author="Doughty MB & Tucker RT & Govender R" box="[372,592,1527,1551]" pageId="3" pageNumber="4" refId="ref13893" refString="Doughty MB, Tucker RT, Govender R et al. Ne contribution of abiotic markers in interpreting the taphonomic weathering of Homiphoca long * bones from Langebaanweg, South Africa. Journal of Atican Earth Sciences 2023; 206: 105022. hups: // doi. org / 10.1016 / j. jafrearsci. 2023.105022" type="journal volume" year="2023"> +Doughty +<emphasis id="B9F8EA99F233FFD0FE27FA06FDD1DB2B" box="[475,529,1527,1551]" italics="true" pageId="3" pageNumber="4">et al.</emphasis> +2023 +</bibRefCitation> +, +<bibRefCitation id="EF1D4B7AF233FFD0FD9DFA09FF23DB0B" author="Govender R & Marx FG" pageId="3" pageNumber="4" refId="ref14246" refString="Govender R, Marx FG. New cetacean fossils from the Late Cenozoic of South Africa. Frontiers in Earth Science 2023; 10: 1058104." type="journal volume" year="2023">Govender and Marx 2023</bibRefCitation> +, +<bibRefCitation id="EF1D4B7AF233FFD0FF0CF9E9FDDDDB0B" author="Jannello JM & Chinsamy A" box="[240,541,1559,1583]" pageId="3" pageNumber="4" pagination="953 - 71" refId="ref14723" refString="Jannello JM, Chinsamy A. Osteohistology and palaeobiology of giraffids from the Mio * Pliocene Langebaanweg (South Africa). 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A new anuran genus from the fossil sites of Langebaanweg and Cooper's Cave, South Africa. Atican Journal of Herpetology 2023; 72: 163 - 89. hups: // doi. org / 10.1080 / 21564574.2 023.2251502" type="journal article" year="2023">Matthews and Steininger 2023</bibRefCitation> +; and literature therein). +</paragraph> +</subSubSection> +<caption id="DFF36603F233FFD0FF8DFADEFCF4D88C" pageId="3" pageNumber="4" startId="3.[113,178,1312,1336]" targetBox="[117,1455,147,1281]" targetPageId="3" targetType="figure"> +<paragraph id="8B33368BF233FFD0FF8DFADEFCF4D88C" blockId="3.[113,1459,1312,1449]" pageId="3" pageNumber="4"> +<emphasis id="B9F8EA99F233FFD0FF8DFADEFF09D81C" bold="true" box="[113,201,1312,1336]" pageId="3" pageNumber="4">Figure 1.</emphasis> +<taxonomicName id="4C8C4D08F233FFD0FF32FADEFE67D81C" box="[206,423,1312,1336]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F233FFD0FF32FADEFE67D81C" box="[206,423,1312,1336]" italics="true" pageId="3" pageNumber="4">Chersina langebaanwegi</emphasis> +</taxonomicName> +, holotype SAM -PQL -20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views. Abbreviations: Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm. +</paragraph> +</caption> +<subSubSection id="C3966500F233FFD7FF8DF991FD3ADA7B" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis"> +<paragraph id="8B33368BF233FFD0FF8DF991FE93DA20" blockId="3.[113,763,1647,1953]" pageId="3" pageNumber="4"> +<emphasis id="B9F8EA99F233FFD0FF8DF991FE86DBA3" box="[113,326,1647,1671]" italics="true" pageId="3" pageNumber="4">Differential diagnosis:</emphasis> +Member of testudinid genus +<taxonomicName id="4C8C4D08F233FFD0FD87F991FD12DBA3" authorityName="Gray" authorityYear="1830" box="[635,722,1647,1671]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="genus"> +<emphasis id="B9F8EA99F233FFD0FD87F991FD12DBA3" box="[635,722,1647,1671]" italics="true" pageId="3" pageNumber="4">Chersina</emphasis> +</taxonomicName> +be - cause of the following combination of characters: wide vertebral scutes almost equal to the vertebral scutes; single trapezoidal suprapygal, notched pygal, and distinct gular protrusion covered by fused gulars scutes. +</paragraph> +<paragraph id="8B33368BF233FFD7FF71F8F2FD3ADA7B" blockId="3.[113,763,1647,1953]" lastBlockId="4.[128,778,1550,1887]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4"> +Differing from +<taxonomicName id="4C8C4D08F233FFD0FECAF8F2FE30DA00" box="[310,496,1804,1828]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F233FFD0FECAF8F2FE30DA00" box="[310,496,1804,1828]" italics="true" pageId="3" pageNumber="4">Chersina angulata</emphasis> +</taxonomicName> +regarding the following characters (see Supporting information for the comparison with +<taxonomicName id="4C8C4D08F233FFD0FF8DF8B5FF28DA46" box="[113,232,1866,1890]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F233FFD0FF8DF8B5FF28DA46" box="[113,232,1866,1890]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> +</taxonomicName> +): the shell is distinctly domed and not relatively low and elongated as in +<taxonomicName id="4C8C4D08F233FFD0FE91F894FE20DAA6" box="[365,480,1898,1922]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F233FFD0FE91F894FE20DAA6" box="[365,480,1898,1922]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> +</taxonomicName> +( +<figureCitation id="13B72A0EF233FFD0FE0CF894FD95DAA5" box="[496,597,1898,1922]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. 1E, F</figureCitation> +; Supporting in - formation, +<figureCitation id="13B72A0EF233FFD0FF18F877FEEADA85" box="[228,298,1929,1953]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. S1</figureCitation> +); the nuchal is not as significantly arched (in anterior view) as that of +<taxonomicName id="4C8C4D08F233FFD0FBCFFA26FB68D8D4" box="[1075,1192,1496,1520]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F233FFD0FBCFFA26FB68D8D4" box="[1075,1192,1496,1520]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> +</taxonomicName> +( +<figureCitation id="13B72A0EF233FFD0FB47FA26FACAD8D4" box="[1211,1290,1496,1520]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. 1G</figureCitation> +; Supporting in - formation, +<figureCitation id="13B72A0EF233FFD0FC62FA09FC27DB2B" box="[926,999,1527,1551]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. S2</figureCitation> +) and does not develop a ventral transversal ridge with medial notch and a dorsal shelf as in +<taxonomicName id="4C8C4D08F233FFD0FAF0F9E9FABDDB0B" box="[1292,1405,1559,1583]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F233FFD0FAF0F9E9FABDDB0B" box="[1292,1405,1559,1583]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> +</taxonomicName> +( +<figureCitation id="13B72A0EF233FFD0FA71F9E9FC87DB6A" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="3" pageNumber="4">Fig. 6B</figureCitation> +; Supporting information, +<figureCitation id="13B72A0EF233FFD0FBAFF9C8FB03DB69" box="[1107,1219,1590,1614]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="3" pageNumber="4">Fig. S3A, B</figureCitation> +); the cervical is, on the dorsal surface, much broader and proportionally longer than that of +<taxonomicName id="4C8C4D08F233FFD0FC87F98BFC31DBA8" box="[891,1009,1652,1676]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F233FFD0FC87F98BFC31DBA8" box="[891,1009,1652,1676]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> +</taxonomicName> +( +<figureCitation id="13B72A0EF233FFD0FBFAF98AFBB4DBA8" box="[1030,1140,1652,1677]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. 2A, B</figureCitation> +; Supporting information, +<figureCitation id="13B72A0EF233FFD0FA71F98AFCB4DB8F" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="3" pageNumber="4">Fig. S3C, D</figureCitation> +); the first neural is hexagonal and not rectangular as in +<taxonomicName id="4C8C4D08F233FFD0FA61F96AFC40DBEF" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F233FFD0FA61F96AFC40DBEF" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> +</taxonomicName> +( +<figureCitation id="13B72A0EF233FFD0FC6DF94DFBD3DBEF" box="[913,1043,1715,1739]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Figure 1A, B</figureCitation> +; Supporting information, +<figureCitation id="13B72A0EF233FFD0FADDF94DFAA7DBEF" box="[1313,1383,1715,1739]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="3" pageNumber="4">Fig. S4</figureCitation> +); most of the neurals host the ribheads ( +<figureCitation id="13B72A0EF233FFD0FB80F92CFB0ADBCF" box="[1148,1226,1746,1771]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="3" pageNumber="4">Figs 3A</figureCitation> +, +<figureCitation id="13B72A0EF233FFD0FB2AF92DFB32DBCF" box="[1238,1266,1747,1771]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="3" pageNumber="4">9B</figureCitation> +; Supporting infor - mation, +<figureCitation id="13B72A0EF233FFD0FC7CF90CFC06DA2E" box="[896,966,1778,1802]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="3" pageNumber="4">Fig. S5</figureCitation> +) and not regularly the costals as in +<taxonomicName id="4C8C4D08F233FFD0FAC5F90CFA6CDA2E" box="[1337,1452,1778,1802]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F233FFD0FAC5F90CFA6CDA2E" box="[1337,1452,1778,1802]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> +</taxonomicName> +; the sixth marginal scute does not contact the third pleural scute as in +<taxonomicName id="4C8C4D08F233FFD0FC9DF8CFFC14DA6C" box="[865,980,1840,1864]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F233FFD0FC9DF8CFFC14DA6C" box="[865,980,1840,1864]" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> +</taxonomicName> +( +<figureCitation id="13B72A0EF233FFD0FC19F8CEFB8ADA6C" box="[997,1098,1840,1865]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="3" pageNumber="4">Fig. 1E, F</figureCitation> +; Supporting information, +<figureCitation id="13B72A0EF233FFD0FAA6F8CEFA61DA6C" box="[1370,1441,1840,1864]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="3" pageNumber="4">Fig. S6</figureCitation> +); the anterior protrusion of the plastron (the gular protrusion of the epiplastra) is comparatively more developed than in +<taxonomicName id="4C8C4D08F233FFD0FA61F88EFC40DA82" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F233FFD0FA61F88EFC40DA82" italics="true" pageId="3" pageNumber="4">C. angulata</emphasis> +</taxonomicName> +(especially in males; +<figureCitation id="13B72A0EF233FFD0FBA9F870FB7BDA82" box="[1109,1211,1934,1959]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="3" pageNumber="4">Figs 4A, B</figureCitation> +, +<figureCitation id="13B72A0EF233FFD0FB39F870FAC0DA82" box="[1221,1280,1934,1958]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="3" pageNumber="4">5C, D</figureCitation> +, +<figureCitation id="13B72A0EF233FFD0FAF6F870FA82DA82" box="[1290,1346,1934,1958]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="3" pageNumber="4">6D, E</figureCitation> +, +<figureCitation id="13B72A0EF233FFD0FAB0F871FA44DA82" box="[1356,1412,1935,1959]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="3" pageNumber="4">8A, B</figureCitation> +, +<figureCitation id="13B72A0EF233FFD7FA72F871FF53DB01" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">9C, D</figureCitation> +, +<figureCitation id="13B72A0EF234FFD7FF61F9F0FF77DB02" box="[157,183,1550,1574]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="4" pageNumber="5">10</figureCitation> +; Supporting information, +<figureCitation id="13B72A0EF234FFD7FE43F9F0FDC4DB02" box="[447,516,1550,1574]" captionStart="Figure 7" captionStartId="9.[114,179,1706,1730]" captionTargetBox="[115,1436,145,1676]" captionTargetId="figure-121@9.[114,1458,144,1678]" captionTargetPageId="9" captionText="Figure 7. Chersina langebaanwegi, paratype SAM-PQL-21749: nuchal in external view (A); pygal in external (B) and internal (C) views; anterior portion of the plastron (and part of the nuchal) in ventral view (D); xiphiplastra in ventral (E) and dorsal (F) views. Scale bars equal 7 cm in (A, D) and 5 cm in (B, C, E, F)." pageId="4" pageNumber="5">Fig. S7</figureCitation> +); the dorsal surface of the epiplastral lips is nearly flat or very slightly convex and not vari - ably concave as in +<taxonomicName id="4C8C4D08F234FFD7FEC0F9B3FE6EDB40" box="[316,430,1612,1636]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F234FFD7FEC0F9B3FE6EDB40" box="[316,430,1612,1636]" italics="true" pageId="4" pageNumber="5">C. angulata</emphasis> +</taxonomicName> +( +<figureCitation id="13B72A0EF234FFD7FE41F9B2FDC3DB41" box="[445,515,1612,1637]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="4" pageNumber="5">Fig. 10</figureCitation> +; Supporting information, +<figureCitation id="13B72A0EF234FFD7FF7DF992FF06DBA0" box="[129,198,1644,1668]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="4" pageNumber="5">Fig. S8</figureCitation> +); the epiplastral lips in most of the cases end posteriorly with a nearly straight or moderately convex profile (in most cases not producing any significant pocket) and not with a very strong convexity as in +<taxonomicName id="4C8C4D08F234FFD7FEE5F934FE4ADBC6" box="[281,394,1738,1762]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F234FFD7FEE5F934FE4ADBC6" box="[281,394,1738,1762]" italics="true" pageId="4" pageNumber="5">C. angulata</emphasis> +</taxonomicName> +(which regularly has a very deep gular pocket) ( +<figureCitation id="13B72A0EF234FFD7FF1DF917FEECDA25" box="[225,300,1769,1793]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="4" pageNumber="5">Figs 3B</figureCitation> +, +<figureCitation id="13B72A0EF234FFD7FEC4F917FE96DA25" box="[312,342,1769,1793]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="4" pageNumber="5">5D</figureCitation> +, +<figureCitation id="13B72A0EF234FFD7FE9EF917FE41DA25" box="[354,385,1769,1793]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="4" pageNumber="5">6D</figureCitation> +, +<figureCitation id="13B72A0EF234FFD7FE71F917FE69DA25" box="[397,425,1769,1793]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="4" pageNumber="5">8B</figureCitation> +, +<figureCitation id="13B72A0EF234FFD7FE48F917FE13DA25" box="[436,467,1769,1793]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="4" pageNumber="5">9D</figureCitation> +, +<figureCitation id="13B72A0EF234FFD7FE23F917FDE9DA25" box="[479,553,1769,1793]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="4" pageNumber="5">10A, D</figureCitation> +; Supporting informa - tion, +<figureCitation id="13B72A0EF234FFD7FF46F8F6FEC4DA04" box="[186,260,1800,1824]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="4" pageNumber="5">Fig. S9</figureCitation> +); the entoplastron is wider than long instead of being longer than wide, or circular, as in +<taxonomicName id="4C8C4D08F234FFD7FDEDF8D6FD42DA64" box="[529,642,1832,1856]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F234FFD7FDEDF8D6FD42DA64" box="[529,642,1832,1856]" italics="true" pageId="4" pageNumber="5">C. angulata</emphasis> +</taxonomicName> +( +<figureCitation id="13B72A0EF234FFD7FD6DF8D6FD1EDA64" box="[657,734,1832,1856]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="4" pageNumber="5">Figs 1D</figureCitation> +, +<figureCitation id="13B72A0EF234FFD7FD14F8D6FCC4DA64" box="[744,772,1832,1856]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="4" pageNumber="5">3B</figureCitation> +, +<figureCitation id="13B72A0EF234FFD7FF7CF8B9FF79DA7A" box="[128,185,1863,1887]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="4" pageNumber="5">4A, B</figureCitation> +, +<figureCitation id="13B72A0EF234FFD7FF38F8B9FEC0DA7A" box="[196,256,1863,1887]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="4" pageNumber="5">5C, D</figureCitation> +, +<figureCitation id="13B72A0EF234FFD7FEF7F8B9FE83DA7A" box="[267,323,1863,1887]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="4" pageNumber="5">8A, B</figureCitation> +, +<figureCitation id="13B72A0EF234FFD7FEB2F8B9FE4BDA7A" box="[334,395,1863,1887]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="4" pageNumber="5">9C, D</figureCitation> +; Supporting information, +<figureCitation id="13B72A0EF234FFD7FD6BF8B9FD29DA7B" box="[663,745,1863,1887]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="4" pageNumber="5">Fig. S10</figureCitation> +). +</paragraph> +</subSubSection> +<caption id="DFF36603F234FFD7FF7DFA8CFF36D8F9" pageId="4" pageNumber="5" startId="4.[129,194,1394,1418]" targetBox="[130,1471,145,1364]" targetPageId="4" targetType="figure"> +<paragraph id="8B33368BF234FFD7FF7DFA8CFF36D8F9" blockId="4.[129,1468,1394,1502]" pageId="4" pageNumber="5"> +<emphasis id="B9F8EA99F234FFD7FF7DFA8CFF18D8AE" bold="true" box="[129,216,1394,1418]" pageId="4" pageNumber="5">Figure 2.</emphasis> +<taxonomicName id="4C8C4D08F234FFD7FF21FA8CFE76D8AE" box="[221,438,1394,1418]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F234FFD7FF21FA8CFE76D8AE" box="[221,438,1394,1418]" italics="true" pageId="4" pageNumber="5">Chersina langebaanwegi</emphasis> +</taxonomicName> +, holotype SAM -PQL -20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations: An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm. +</paragraph> +</caption> +<subSubSection id="C3966500F234FFDDFF7CF87FFE67DFF8" lastPageId="14" lastPageNumber="15" pageId="4" pageNumber="5" type="description"> +<paragraph id="8B33368BF234FFD7FF7CF87FFC70DBA0" blockId="4.[128,778,1920,1976]" lastBlockId="4.[825,1473,1550,1668]" pageId="4" pageNumber="5"> +<emphasis id="B9F8EA99F234FFD7FF7CF87FFF3FDABC" box="[128,255,1921,1944]" italics="true" pageId="4" pageNumber="5">Description:</emphasis> +If not otherwise stated, the following descrip - tion is mostly based on the +<typeStatus id="54378829F234FFD7FE58F85EFDC5DA9C" box="[420,517,1952,1976]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus> +SAM -PQL -20734 ( +<figureCitation id="13B72A0EF234FFD7FD22F85EFCA2DB02" box="[734,866,1550,1976]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[113,178,1312,1336]" captionStartId-1="4.[129,194,1394,1418]" captionStartId-2="5.[113,178,1047,1071]" captionTargetBox-0="[117,1455,147,1281]" captionTargetBox-1="[130,1471,145,1364]" captionTargetBox-2="[147,1424,145,1017]" captionTargetId-0="figure-340@3.[114,1458,144,1284]" captionTargetId-1="figure-300@4.[129,1473,144,1366]" captionTargetId-2="figure-538@5.[146,1426,144,1019]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-1="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." captionText-2="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="4" pageNumber="5">Figs 1–3</figureCitation> +) as it is the only nearly complete and mounted shell so far available. There are no cranial, axial, or appendicular elements associated with the +<typeStatus id="54378829F234FFD7FB69F9B2FB36DB40" box="[1173,1270,1612,1636]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus> +and the referred specimens. +</paragraph> +<paragraph id="8B33368BF234FFD7FBACF95EFB6BDB9E" blockId="4.[825,1475,1696,1978]" box="[1104,1195,1696,1722]" pageId="4" pageNumber="5"> +<emphasis id="B9F8EA99F234FFD7FBACF95EFB6BDB9E" box="[1104,1195,1696,1722]" italics="true" pageId="4" pageNumber="5">Carapace</emphasis> +</paragraph> +<paragraph id="8B33368BF234FFD6FCC5F939FE4EDB0E" blockId="4.[825,1475,1696,1978]" lastBlockId="5.[113,763,1147,1578]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5"> +The nearly complete female shell SAM -PQL -20734 ( +<typeStatus id="54378829F234FFD7FAABF938FA7ADBFA" box="[1367,1466,1734,1758]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus> +) indicates that the carapace of this species was highly domed (length +<quantity id="4C749B6EF234FFD7FC73F8FBFC26DA39" box="[911,998,1797,1821]" metricMagnitude="-1" metricUnit="m" metricValue="2.36" pageId="4" pageNumber="5" unit="cm" value="23.6">23.6 cm</quantity> +, width 15.7, height +<quantity id="4C749B6EF234FFD7FB43F8FBFAE2DA39" box="[1215,1314,1797,1821]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="4" pageNumber="5" unit="mm" value="14.7">14.7 mm</quantity> +), with a nearly smooth dorsal profile both in lateral and anterior/posterior views. The posterior border is relatively curved inwards and not significantly flared. Very weak bosses are visible in lateral view ( +<figureCitation id="13B72A0EF234FFD7FCB8F87CFC71DABE" box="[836,945,1922,1946]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="4" pageNumber="5">Fig. 1C, D</figureCitation> +) that correspond to five vertebral scutes. In dorsal view, the anterior edge of the shell ( +<figureCitation id="13B72A0EF234FFD7FB46F85CFAE4DA9D" box="[1210,1316,1954,1978]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="4" pageNumber="5">Fig. 2A, B</figureCitation> +) has a modest notch laterally delimited by the small protrusion of the first peripherals (corresponding to the sulcus between the first and second marginal) and, moreover, the central area of the carapace, corresponding to the bridge, is somewhat swollen if compared to the areas anterior and posterior to it. The anterior opening is kidney shaped because the anterior lobe of the plastron is ele - vated in the dorsal direction ( +<figureCitation id="13B72A0EF235FFD6FE66FAC9FE22D86B" box="[410,482,1335,1359]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="5" pageNumber="6">Fig. 3A</figureCitation> +; see description of the plas - tron). The posterior opening is rather broad ( +<figureCitation id="13B72A0EF235FFD6FDAFFAA8FD01D84A" box="[595,705,1366,1390]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="5" pageNumber="6">Fig. 2C, D</figureCitation> +) and markedly much broader than in extant +<taxonomicName id="4C8C4D08F235FFD6FDF3FA8BFD05D8A9" box="[527,709,1397,1421]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F235FFD6FDF3FA8BFD05D8A9" box="[527,709,1397,1421]" italics="true" pageId="5" pageNumber="6">Chersina angulata</emphasis> +</taxonomicName> +. The carapace has no hinge and moreover there is no hinge between the carapace and plastron [as also indicated, among other, by the partial plastron SAM -PQL -20925 ( +<figureCitation id="13B72A0EF235FFD6FE19FA2DFDE1D8C8" box="[485,545,1491,1516]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="5" pageNumber="6">Fig. 4</figureCitation> +) that is still sutured with some bridge peripherals] or within the plastron (see also SAM -PQL -22076— +<figureCitation id="13B72A0EF235FFD6FEB9F9ECFEBDDB0E" box="[325,381,1554,1578]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="5" pageNumber="6">Fig. 8</figureCitation> +). +</paragraph> +<caption id="DFF36603F235FFD6FF8DFBE9FA48D96E" pageId="5" pageNumber="6" startId="5.[113,178,1047,1071]" targetBox="[147,1424,145,1017]" targetPageId="5" targetType="figure"> +<paragraph id="8B33368BF235FFD6FF8DFBE9FA48D96E" blockId="5.[113,1432,1047,1099]" pageId="5" pageNumber="6"> +<emphasis id="B9F8EA99F235FFD6FF8DFBE9FF09D90B" bold="true" box="[113,201,1047,1071]" pageId="5" pageNumber="6">Figure 3.</emphasis> +<taxonomicName id="4C8C4D08F235FFD6FF32FBE9FE67D90B" box="[206,423,1047,1071]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F235FFD6FF32FBE9FE67D90B" box="[206,423,1047,1071]" italics="true" pageId="5" pageNumber="6">Chersina langebaanwegi</emphasis> +</taxonomicName> +, holotype SAM -PQL -20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information. +</paragraph> +</caption> +<paragraph id="8B33368BF235FFD6FF8DF9B2FBF4D9D5" blockId="5.[113,762,1611,1949]" lastBlockId="5.[810,1459,1147,1954]" pageId="5" pageNumber="6"> +<emphasis id="B9F8EA99F235FFD6FF8DF9B2FF2FDB47" box="[113,239,1611,1635]" italics="true" pageId="5" pageNumber="6">Bony plates:</emphasis> +The nuchal is a hexagonal element, about 1/3 broader than long ( +<quantity id="4C749B6EF235FFD6FEC0F995FE4ADBA6" box="[316,394,1643,1667]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="6" unit="mm" value="65.0">65 mm</quantity> +wide and +<quantity id="4C749B6EF235FFD6FE0BF995FD82DBA6" box="[503,578,1643,1667]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="5" pageNumber="6" unit="mm" value="47.0">47 mm</quantity> +long in the holo - +<typeStatus id="54378829F235FFD6FF8DF975FF5FDB87" box="[113,159,1675,1699]" pageId="5" pageNumber="6">type</typeStatus> +SAM -PQL -20734), with borders that can be nearly straight (SAM -PQL -21174) or weakly concave (SAM - PQL -20734, SAM -PQL -21174) (the character has been scored as straight because of the weak concavity). The anterior edge can be straight (SAM -PQL -21174) or shallowly concave with, at least in some cases (SAM -PQL -20734), a small, acute notch at the centre. In anterior view, the anterior edge is only modestly arched (but not deeply arched as in +<taxonomicName id="4C8C4D08F235FFD6FE13F898FDA4DA59" box="[495,612,1893,1917]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F235FFD6FE13F898FDA4DA59" box="[495,612,1893,1917]" italics="true" pageId="5" pageNumber="6">C. angulata</emphasis> +</taxonomicName> +). On the ven - tral surface, the transversal ridge marking the line of maximum thickness of the element (slightly posterior to the end of the ventral fold of the scutes) is straight and medially notched as in +<taxonomicName id="4C8C4D08F235FFD6FCBAFB44FC7BD9F5" box="[838,955,1209,1233]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F235FFD6FCBAFB44FC7BD9F5" box="[838,955,1209,1233]" italics="true" pageId="5" pageNumber="6">C. angulata</emphasis> +</taxonomicName> +that develops a sort of dorsal shelf (Supporting information, +<figureCitation id="13B72A0EF235FFD6FC4DFB27FBE3D9D4" box="[945,1059,1241,1265]" captionStart="Figure 3" captionStartId="5.[113,178,1047,1071]" captionTargetBox="[147,1424,145,1017]" captionTargetId="figure-538@5.[146,1426,144,1019]" captionTargetPageId="5" captionText="Figure 3. Chersina langebaanwegi, holotype SAM-PQL-20734: CT renderings of the visceral surface of the neural series (A) and the plastron (B). Scale bar equals 2 cm in (A) and 5 cm in (B). The STL file of the CT scan of the whole shell is available in the Supporting information." pageId="5" pageNumber="6">Fig. S3A, B</figureCitation> +). +</paragraph> +<paragraph id="8B33368BF235FFD5FCB9FB06FE75DA64" blockId="5.[810,1459,1147,1954]" lastBlockId="6.[129,777,1520,1951]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6"> +There are eight neural elements and the neural formula is: 6,8,4,8,4,6A,4,6A. The first neural of SAM -PQL -20734 is anteroposteriorly elongated and hexagonal with anterolateral sides approximately equal to the posterolateral ones. Neurals from two to seven are wider than long (in absence of the proper state, the third has been scored as square but it is in fact rect - angular). The eighth is longer than wide and, at least on the better -preserved right side, it seems to be hexagonal (with a short anterolateral side). On the visceral surface of the neurals from two to five there is clear evidence for the emergence of very thin costal processes from the neural and not from the corres - ponding costals. In addition, the sixth neural may show the same condition as indicated by a small irregularity located close to the posterior border (and by the absence of a costal process on the corresponding costal). The costovertebral tunnel is, therefore, very narrow (much narrower than the width of the neurals). The two joined neurals of SAM -PQL -35023d ( +<figureCitation id="13B72A0EF235FFD6FB13F913FAEBDA21" box="[1263,1323,1773,1797]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="5" pageNumber="6">Fig. 9</figureCitation> +) have a sort of tubercle on the visceral surface close to the lateral edge. The preservation of the ventral surface of the isolated neural of SAM - PQL -21174 does not allow confirmation of this morphology in another specimen but its irregular surface could indicate the presence of costal processes ( +<figureCitation id="13B72A0EF235FFD6FB9BF874FB64DA86" box="[1127,1188,1930,1954]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="5" pageNumber="6">Fig. 6</figureCitation> +). Noteworthy is that the three preserved costals of SAM -PQL -21109 do not have a costal process on the medial area of their visceral surface. There is only one suprapygal that is trapezoidal in shape; its posterior edge is approximately straight. The female pygal of SAM -PQL -20734 is quadrangular and shows a small notch centrally located on the free, ventral edge; it is only moderately convex. The male pygal SAM -PQL -20925 is also quadrangular. The male pygal of SAM - PQL -21109 and likely also the partly preserved large male pygal SAM -PQL -35023d are hexagonal, with short anterolateral sides and long, concave posterolateral sides; the free, ventral edge is broadly concave in both cases. +</paragraph> +<caption id="DFF36603F236FFD5FF7DFA72FACED89B" pageId="6" pageNumber="7" startId="6.[129,194,1420,1444]" targetBox="[131,1471,146,1390]" targetPageId="6" targetType="figure"> +<paragraph id="8B33368BF236FFD5FF7DFA72FACED89B" blockId="6.[129,1455,1420,1472]" pageId="6" pageNumber="7"> +<emphasis id="B9F8EA99F236FFD5FF7DFA72FF18D880" bold="true" box="[129,216,1420,1444]" pageId="6" pageNumber="7">Figure 4.</emphasis> +<taxonomicName id="4C8C4D08F236FFD5FF21FA72FE76D880" box="[221,438,1420,1444]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F236FFD5FF21FA72FE76D880" box="[221,438,1420,1444]" italics="true" pageId="6" pageNumber="7">Chersina langebaanwegi</emphasis> +</taxonomicName> +, paratype SAM -PQL -20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm. +</paragraph> +</caption> +<paragraph id="8B33368BF236FFD5FF60F8B6FC3DDBE7" blockId="6.[129,777,1520,1951]" lastBlockId="6.[825,1475,1519,1951]" pageId="6" pageNumber="7">There are eight pairs of costals. Those from two to six are markedly trapezoidal with alternate broad and narrow medial and lateral ends. The sixth and, above all, the seventh and eighth costals are posteriorly bent. Only the first, seventh, and eighth costals bear costal processes on their visceral surfaces (see de - scription of neurals above). The first costals have a very stout costal process, and the rib is raised in an evident, anteriorly con - cave arched ridge. The external surface of costals from two to five shows in the medial sector (close to the neurals) marked growth grooves and ridges.</paragraph> +<paragraph id="8B33368BF236FFD4FCA9F935FD84DA4B" blockId="6.[825,1475,1519,1951]" lastBlockId="7.[113,763,1754,1903]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7"> +There are 11 peripherals. Those from three to seven are involved in the bridge; their external surface is smooth, devoid of any signifi - cant keel (that is conversely present in +<taxonomicName id="4C8C4D08F236FFD5FB3DF8F7FAAEDA05" box="[1217,1390,1801,1825]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F236FFD5FB3DF8F7FAAEDA05" box="[1217,1390,1801,1825]" italics="true" pageId="6" pageNumber="7">Chersina angulata</emphasis> +</taxonomicName> +). In the +<typeStatus id="54378829F236FFD5FCC5F8D7FC53DA65" box="[825,915,1833,1857]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus> +SAM -PQL -20734 the first and second peripherals de - velop a small, distal projection corresponding to the intermarginal sulcus, but there are no intermarginal projections or interperipheral serrations in the posterior area. Conversely, in SAM -PQL -21109 there is a marked intermarginal projection on the ninth periph - eral (best preserved on the right side). The third peripheral hosts a small axillary scute that develops on the hyoplastron buttress as well. The seventh peripheral hosts part of the large inguinal scute (that is mostly developed on the hypoplastron). +</paragraph> +<caption id="DFF36603F237FFD4FF8DF9A4FF1EDB8E" pageId="7" pageNumber="8" startId="7.[113,178,1626,1650]" targetBox="[115,1456,145,1596]" targetPageId="7" targetType="figure"> +<paragraph id="8B33368BF237FFD4FF8DF9A4FF1EDB8E" blockId="7.[113,1414,1626,1706]" pageId="7" pageNumber="8"> +<emphasis id="B9F8EA99F237FFD4FF8DF9A4FF08DB56" bold="true" box="[113,200,1626,1650]" pageId="7" pageNumber="8">Figure 5.</emphasis> +<taxonomicName id="4C8C4D08F237FFD4FF31F9A4FE67DB56" box="[205,423,1626,1650]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F237FFD4FF31F9A4FE67DB56" box="[205,423,1626,1650]" italics="true" pageId="7" pageNumber="8">Chersina langebaanwegi</emphasis> +</taxonomicName> +, paratype SAM -PQL -21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm. +</paragraph> +</caption> +<paragraph id="8B33368BF237FFDAFF8DF86BFEE9DA7D" blockId="7.[113,762,1940,1995]" lastBlockId="9.[113,762,1826,2007]" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8"> +<emphasis id="B9F8EA99F237FFD4FF8DF86BFF33DA88" box="[113,243,1941,1964]" italics="true" pageId="7" pageNumber="8">Horny scutes:</emphasis> +The cervical scute is rather large on both the dorsal and ventral surface of the nuchal. On the dorsal surface, it can be only slightly longer than broad (in SAM -PQL -20734, respect - ively 15.0 and +<quantity id="4C749B6EF237FFD4FC40F907FBD9DA35" box="[956,1049,1785,1809]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="7" pageNumber="8" unit="mm" value="14.3">14.3 mm</quantity> +; +<figureCitation id="13B72A0EF237FFD4FBDFF907FB45DA35" box="[1059,1157,1785,1809]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="7" pageNumber="8">Fig. 2A, B</figureCitation> +), and it is slightly longer than 25% of the nuchal. The cervical scute of SAM -PQL -21174 is distinctly longer than broad but its length corresponds to about 25% of the nuchal. On the ventral surface of the nuchal, the cer - vical is very long and has nearly parallel margins. The first pleural scutes slightly overlap the nuchal in SAM -PQL -20734 ( +<figureCitation id="13B72A0EF237FFD4FA9FF868FCF8DAE9" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="7" pageNumber="8">Fig. 2A, B</figureCitation> +) even less in SAM -PQL -21174 (where only the left side is well preserved) but not in SAM -PQL -21109 (that also preserves only the left side). +</paragraph> +<caption id="DFF36603F238FFDBFF7DF8D2FA45DA7B" pageId="8" pageNumber="9" startId="8.[129,194,1836,1860]" targetBox="[169,1432,146,1806]" targetPageId="8" targetType="figure"> +<paragraph id="8B33368BF238FFDBFF7DF8D2FA45DA7B" blockId="8.[129,1438,1836,1888]" pageId="8" pageNumber="9"> +<emphasis id="B9F8EA99F238FFDBFF7DF8D2FF18DA60" bold="true" box="[129,216,1836,1860]" pageId="8" pageNumber="9">Figure 6.</emphasis> +<taxonomicName id="4C8C4D08F238FFDBFF21F8D2FE76DA60" box="[221,438,1836,1860]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F238FFDBFF21F8D2FE76DA60" box="[221,438,1836,1860]" italics="true" pageId="8" pageNumber="9">Chersina langebaanwegi</emphasis> +</taxonomicName> +, paratype SAM -PQL -PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm. +</paragraph> +</caption> +<caption id="DFF36603F239FFDAFF8EF954FE14DBDE" pageId="9" pageNumber="10" startId="9.[114,179,1706,1730]" targetBox="[115,1436,145,1676]" targetPageId="9" targetType="figure"> +<paragraph id="8B33368BF239FFDAFF8EF954FE14DBDE" blockId="9.[113,1451,1706,1786]" pageId="9" pageNumber="10"> +<emphasis id="B9F8EA99F239FFDAFF8EF954FF0ADBE6" bold="true" box="[114,202,1706,1730]" pageId="9" pageNumber="10">Figure 7.</emphasis> +<taxonomicName id="4C8C4D08F239FFDAFF32F954FE68DBE6" box="[206,424,1706,1730]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F239FFDAFF32F954FE68DBE6" box="[206,424,1706,1730]" italics="true" pageId="9" pageNumber="10">Chersina langebaanwegi</emphasis> +</taxonomicName> +, paratype SAM -PQL -21749: nuchal in external view (A); pygal in external (B) and internal (C) views; anterior portion of the plastron (and part of the nuchal) in ventral view (D); xiphiplastra in ventral (E) and dorsal (F) views. Scale bars equal 7 cm in (A, D) and 5 cm in (B, C, E, F). +</paragraph> +</caption> +<paragraph id="8B33368BF239FFDAFF71F89FFF25DAF3" blockId="9.[113,762,1826,2007]" pageId="9" pageNumber="10">The first vertebral scute is as long as it is broad and has nearly parallel sides; its posterior border (the sulcus between the first and second vertebral) crosses the first neural close to its pos - terior edge.</paragraph> +<paragraph id="8B33368BF239FFDFFCB9F8DCFD88DB8D" blockId="9.[810,1459,1826,2007]" lastBlockId="12.[129,778,1462,1705]" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10"> +Intervertebral sulci are located on the first, third, fifth, and sev - enth costals, whereas the interpleural sulci on the second, fourth, sixth, and eighth costals. Since the posterior edge of the fifth ver - tebral is located only slightly posteriorly to the suprapygal -pygal suture (and is, therefore, visible +<quantity id="4C749B6EF239FFDAFB8EF85EFB08DA93" box="[1138,1224,1952,1976]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="10" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity> +posterior to the suture on the pygal of SAM -PQL -20734), this character was scored as if the sulcus coincides with the suprapygal -pygal suture (char - acter 112, state 1). With the exception of the nuchal and pygal areas, the pleuro -marginal sulcus overlaps the costo -peripheral suture. The five vertebral scutes are rather large, nearly equalling the width of the pleurals. The intervertebral sulci are located on the suture between the first and the second neurals, on the third, fifth, and eight neurals (very close to the anterior suture of the latter, not much posteriorly as in +<taxonomicName id="4C8C4D08F23BFFD8FB4BF8ABFAE9DA49" box="[1207,1321,1877,1901]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F23BFFD8FB4BF8ABFAE9DA49" box="[1207,1321,1877,1901]" italics="true" pageId="11" pageNumber="12">C. angulata</emphasis> +</taxonomicName> +). The second marginal scute does not contact the nuchal or the first vertebral. The sixth marginal corresponds entirely to the second pleural (does not contact the third pleural; unlike in +<taxonomicName id="4C8C4D08F23CFFDFFDABFA49FD0BD8EA" box="[599,715,1462,1486]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F23CFFDFFDABFA49FD0BD8EA" box="[599,715,1462,1486]" italics="true" pageId="12" pageNumber="13">C. angulata</emphasis> +</taxonomicName> +). The tenth marginal scute does not contact the fifth vertebral scute. The posterior sulcus of the fifth vertebral scute is placed slightly posterior to the suprapygal -pygal suture in SAM -PQL -20734 ( +<figureCitation id="13B72A0EF23CFFDFFF77F9CDFF32DB6F" box="[139,242,1587,1612]" captionStart="Figure 2" captionStartId="4.[129,194,1394,1418]" captionTargetBox="[130,1471,145,1364]" captionTargetId="figure-300@4.[129,1473,144,1366]" captionTargetPageId="4" captionText="Figure 2. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C) and interpretative drawings (B, D) of the nearly complete female shell in anterodorsal (A, B) and posterior (C, D) views. Abbreviations:An, anal scute; Ce, cervical scute; co, costal; ep, epiplastron; Gu, gular scute; Ma, marginal scute; ne, neural; nu, nuchal; per, peripheral; py, pygal; sp, suprapygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="12" pageNumber="13">Fig. 2C, D</figureCitation> +) and even more posteriorly (therefore, clearly on the pygal) in SAM -PQL -20925, SAM -PQL -21109, and, above all, in the very large male SAM -PQL -35023d. The twelfth marginal scutes are fused in a single supracaudal scute. +</paragraph> +<caption id="DFF36603F23AFFD9FF7DF8D1FE4EDA46" pageId="10" pageNumber="11" startId="10.[129,194,1839,1863]" targetBox="[137,1465,146,1809]" targetPageId="10" targetType="figure"> +<paragraph id="8B33368BF23AFFD9FF7DF8D1FE4EDA46" blockId="10.[129,1453,1839,1891]" pageId="10" pageNumber="11"> +<emphasis id="B9F8EA99F23AFFD9FF7DF8D1FF18DA63" bold="true" box="[129,216,1839,1863]" pageId="10" pageNumber="11">Figure 8.</emphasis> +<taxonomicName id="4C8C4D08F23AFFD9FF21F8D1FE76DA63" box="[221,438,1839,1863]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F23AFFD9FF21F8D1FE76DA63" box="[221,438,1839,1863]" italics="true" pageId="10" pageNumber="11">Chersina langebaanwegi</emphasis> +</taxonomicName> +, paratype SAM -PQL -PQL -22076: male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views. Scale bars equal 5 cm. +</paragraph> +</caption> +<caption id="DFF36603F23BFFD8FF8DF94CFCCCDBC2" pageId="11" pageNumber="12" startId="11.[113,178,1714,1738]" targetBox="[131,1423,182,1679]" targetPageId="11" targetType="figure"> +<paragraph id="8B33368BF23BFFD8FF8DF94CFCCCDBC2" blockId="11.[113,1454,1714,1767]" pageId="11" pageNumber="12"> +<emphasis id="B9F8EA99F23BFFD8FF8DF94CFF09DBEE" bold="true" box="[113,201,1714,1738]" pageId="11" pageNumber="12">Figure 9.</emphasis> +<taxonomicName id="4C8C4D08F23BFFD8FF32F94DFE67DBEF" box="[206,423,1715,1739]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F23BFFD8FF32F94DFE67DBEF" box="[206,423,1715,1739]" italics="true" pageId="11" pageNumber="12">Chersina langebaanwegi</emphasis> +</taxonomicName> +, paratype SAM -PQL -35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm. +</paragraph> +</caption> +<caption id="DFF36603F23CFFDFFF7DFAC8FC01D8A2" pageId="12" pageNumber="13" startId="12.[129,194,1334,1358]" targetBox="[132,1471,147,1304]" targetPageId="12" targetType="figure"> +<paragraph id="8B33368BF23CFFDFFF7DFAC8FC01D8A2" blockId="12.[129,1463,1334,1414]" pageId="12" pageNumber="13"> +<emphasis id="B9F8EA99F23CFFDFFF7DFAC8FF25D86A" bold="true" box="[129,229,1334,1358]" pageId="12" pageNumber="13">Figure 10.</emphasis> +<taxonomicName id="4C8C4D08F23CFFDFFF15FAC8FE03D86A" box="[233,451,1334,1358]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F23CFFDFFF15FAC8FE03D86A" box="[233,451,1334,1358]" italics="true" pageId="12" pageNumber="13">Chersina langebaanwegi</emphasis> +</taxonomicName> +, male left epiplastron SAM -PQL -12750c and female right SAM -PQL -73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm. +</paragraph> +</caption> +<paragraph id="8B33368BF23CFFDFFE61F937FE2CDBC7" blockId="12.[129,777,1737,1957]" box="[413,492,1737,1763]" pageId="12" pageNumber="13"> +<emphasis id="B9F8EA99F23CFFDFFE61F937FE2CDBC7" box="[413,492,1737,1763]" italics="true" pageId="12" pageNumber="13">Plastron</emphasis> +</paragraph> +<paragraph id="8B33368BF23CFFDEFF7DF90EFE32DDE3" blockId="12.[129,777,1737,1957]" lastBlockId="13.[113,763,144,199]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13"> +The large ( +<quantity id="4C749B6EF23CFFDFFF0FF90EFE90DA23" box="[243,336,1776,1800]" metricMagnitude="-1" metricUnit="m" metricValue="2.93" pageId="12" pageNumber="13" unit="mm" value="293.0">293 mm</quantity> +long) and complete male plastron SAM - PQL -22076 ( +<figureCitation id="13B72A0EF23CFFDFFEEDF8F1FE8DDA03" box="[273,333,1807,1831]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="12" pageNumber="13">Fig. 8</figureCitation> +) provides a unique opportunity to assess the general morphology of this shell portion that otherwise is not entirely preserved in a single specimen. The anterior lobe is somewhat trilobed, longer than the posterior lobe (93 and +<quantity id="4C749B6EF23CFFDFFD13F893FF6CDA80" metricMagnitude="-2" metricUnit="m" metricValue="6.8" pageId="12" pageNumber="13" unit="mm" value="68.0">68 mm</quantity> +, respectively) and does not host any hinge. In dorsal (or ventral) view, the gularo -humeral sulcus corresponds to a dis - tinct constriction in the lateral profile of the element. Because of the anterodorsal orientation of the hyoplastra, the epiplastra are raised from the horizontal plane and their anterior protru - sion is anteriorly directed and approximately parallel to the hori - zontal plane but about +<quantity id="4C749B6EF23CFFDFFBE3F9ADFB90DB4E" box="[1055,1104,1619,1643]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="cm" value="2.0">2 cm</quantity> +above it. Such a general morphology is also present in all the preserved plastra. The ventral plastral surface is distinctly medially concave in correspondence of the posterior region of the hyoplastra, the whole hypoplastra and at least the anterior region of the xiphyplastra, indicating that this specimen was likely a male (as SAM -PQL -20925 and the very large SAM -PQL -21109). The +<typeStatus id="54378829F23CFFDFFB8FF8F1FB0FDA03" box="[1139,1231,1807,1831]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus> +SAM -PQL -20734 pre - serves a nearly complete plastron (only the left hypoplastron is missing) that is fully congruent with the one previously de - scribed but slightly smaller, less protruding anteriorly and flat, at least the latter character indicating that this specimen could be a female. The plastron extends anteriorly up to the third peripheral and posteriorly up to the seventh one. +</paragraph> +<paragraph id="8B33368BF23DFFDEFF8DFF17FF0BDF1E" blockId="13.[113,763,233,1979]" pageId="13" pageNumber="14"> +<emphasis id="B9F8EA99F23DFFDEFF8DFF17FF28DC25" box="[113,232,233,257]" italics="true" pageId="13" pageNumber="14">Bony plates:</emphasis> +The epiplastra are characterized by anterior projec - tions that in males can be very long, accounting in most cases (SAM -PQL -21174; +<figureCitation id="13B72A0EF23DFFDEFEB4FED9FE47DC1B" box="[328,391,295,319]" captionStart="Figure 6" captionStartId="8.[129,194,1836,1860]" captionTargetBox="[169,1432,146,1806]" captionTargetId="figure-8@8.[168,1434,144,1808]" captionTargetPageId="8" captionText="Figure 6. Chersina langebaanwegi, paratype SAM-PQL-PQL 21174: nuchal in dorsal view (A); pygal in external (B) and internal (C) views; epiplastra in dorsal (D), ventral (E) and posterior views (F); right epiplastron in medial (G) and lateral (H) views. Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 6</figureCitation> +) for about half of the anteropos - terior length of this element, but in SAM -PQL -35023d they are even longer and represent nearly 80% of the total length (total length +<quantity id="4C749B6EF23DFFDEFF40FE7BFECBDCB9" box="[188,267,389,413]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="13" pageNumber="14" unit="mm" value="95.0">95 mm</quantity> +; projection length +<quantity id="4C749B6EF23DFFDEFE2AFE7BFDE5DCB9" box="[470,549,389,413]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="14" unit="mm" value="75.0">75 mm</quantity> +; +<figureCitation id="13B72A0EF23DFFDEFDCEFE7BFDAFDCB9" box="[562,623,389,413]" captionStart="Figure 9" captionStartId="11.[113,178,1714,1738]" captionTargetBox="[131,1423,182,1679]" captionTargetId="figure-113@11.[114,1458,144,1687]" captionTargetPageId="11" captionText="Figure 9. Chersina langebaanwegi, paratype SAM-PQL-35023d: two neurals in external (A) and internal (B) views; incomplete anterior lobe of the plastron in ventral (C) and dorsal (D) views. Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 9</figureCitation> +). In females, the projection is much less developed as shown by the +<typeStatus id="54378829F23DFFDEFD62FE5BFD3ADC99" box="[670,762,421,445]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus> +SAM -PQL -20734 and, among the referred materials, by SAM - PQL -73878 (compare male epiplastron SAM -PQL -12750c, +<figureCitation id="13B72A0EF23DFFDEFF8DFDFDFF2EDF3F" box="[113,238,515,539]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="13" pageNumber="14">Fig. 10A–C</figureCitation> +, with female epiplastron SAM -PQL -73878, +<figureCitation id="13B72A0EF23DFFDEFD28FDFDFF7ADF1E" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="13" pageNumber="14">Fig. 10D–F</figureCitation> +). +</paragraph> +<paragraph id="8B33368BF23DFFDEFF71FDBCFBC2DC86" blockId="13.[113,763,233,1979]" lastBlockId="13.[810,1460,143,1765]" pageId="13" pageNumber="14"> +In most cases, the projections are quite asymmetric (one of the two is much more developed than the other one), and, therefore, the interepiplastral suture is not located sagittally but deviates laterally. The long and broad dorsal surface of the lip is generally rather flat (SAM -PQL -21174) or, very rarely, modestly convex (SAM -PLQ -24826a). In dorsal view, the pos - terior edge of the gular lip is nearly straight or modestly convex but in some cases it can be moderately overhanging. There are however some exceptions such as the female right epiplastra SAM -PQL -73749 and SAM -PQL -73878 ( +<figureCitation id="13B72A0EF23DFFDEFDCDFCA5FD69DE57" box="[561,681,859,883]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="13" pageNumber="14">Fig. 10D–F</figureCitation> +) (both fitting with +<taxonomicName id="4C8C4D08F23DFFDEFF11FC84FE1ADEB6" box="[237,474,890,914]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F23DFFDEFF11FC84FE1ADEB6" box="[237,474,890,914]" italics="true" pageId="13" pageNumber="14">Chersina langebaanwegi</emphasis> +</taxonomicName> +because the golarohumeral sulcus indicates that the gular area reaches but does not enter the entoplastron), and the two epiplastra SAM -PQL -12754a (fitting with +<taxonomicName id="4C8C4D08F23DFFDEFF0AFC27FE63DED4" box="[246,419,984,1008]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F23DFFDEFF0AFC27FE63DED4" box="[246,419,984,1008]" italics="true" pageId="13" pageNumber="14">C. langebaanwegi</emphasis> +</taxonomicName> +also because of the shape of the associated entoplastron) showing a gular lip moderately convex and the male left epiplastron SAM -PQL -73837 or the female right epiplastron SAM -PQL -24826b that both have a distinctly overhanging posterior edge of the gular lip, forming a rather deep pocket (the edge of the lip reaches in the first and even sur - passes in the second the anterior margin of the epi -entoplastron suture). The visceral surface of the epiplastra, posterior to the pad, is raised in a variably developed tubercle along the suture with the entoplastron (approximately at mid -length of the an - terolateral suture of the entoplastron). The ventral surface of the epiplastra is relatively flat in the area covered by the single gular. The epiplastra thin out gradually in a lateral direction along the suture with the hyoplastron, in correspondence with the dorsal area covered by the humeral shield. The entoplastron is regu - larly wider than long but only slightly in the +<typeStatus id="54378829F23DFFDEFDABFA50FD73D8E2" box="[599,691,1454,1478]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus> +SAM - PQL -20734 where it is +<quantity id="4C749B6EF23DFFDEFE95FA33FE06D8C1" box="[361,454,1485,1509]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="13" pageNumber="14" unit="mm" value="24.5">24.5 mm</quantity> +wide and 22.0 mm long, and much more in SAM -PQL -22076 where it is +<quantity id="4C749B6EF23DFFDEFDBEFA13FD53DB20" box="[578,659,1517,1541]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="14" unit="mm" value="8.1">8.1 mm</quantity> +wide and +<quantity id="4C749B6EF23DFFDEFF8DF9F2FF10DB00" box="[113,208,1548,1572]" metricMagnitude="-2" metricUnit="m" metricValue="3.63" pageId="13" pageNumber="14" unit="mm" value="36.3">36.3 mm</quantity> +long. The visceral surface of the epiplastron hosts a Y -shaped ridge whose anterior rami originates from the above described tubercles on the epiplastra. The central depression between the anterior rami is in all cases very modest. The ven - tral surface of the element is weakly concave. The hyoplastra are slightly longer than wide. They develop a rather robust and long buttress that reaches the first costal plate. Their medial sector is sexually dimorphic, being concave in males and flat in females. The hypoplastra are approximately rectangular. They develop a very robust posterior buttress that contacts the fifth and sixth costal plate. These elements show the same sexual dimorphism of the hyoplastra. The xipiplastra are not significantly dimorphic, with the exception of the presence of a medial concavity devel - oped in hyo -, hypo -, and xiphyplastra of males (the plastron is rather flat in females). In both sexes, the lateral edge is slightly convex with nearly no (SAM -PQL -22076; +<figureCitation id="13B72A0EF23DFFDEFB0AFF51FAF4DDE3" box="[1270,1332,175,199]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 8</figureCitation> +) or a very modest (SAM -PQL -20734; +<figureCitation id="13B72A0EF23DFFDEFBB2FF31FB48DDC3" box="[1102,1160,207,231]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="13" pageNumber="14">Fig. 1</figureCitation> +) notch corresponding to the femoroanal sulcus. The anal notch is in both sexes rather shallow and delimiting an open angle of about 125–140°. However, the male xipiplastra of SAM -PQL -21109 ( +<figureCitation id="13B72A0EF23DFFDEFB49FED3FB2EDC60" box="[1205,1262,300,325]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 5</figureCitation> +) are different from those of the equally large male SAM -PQL -22076 ( +<figureCitation id="13B72A0EF23DFFDEFABFFEB2FABFDC40" box="[1347,1407,332,356]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 8</figureCitation> +) be - cause their tip is bent upwards and the anal notch is much deeper and narrower (115°). +</paragraph> +<paragraph id="8B33368BF23DFFDEFCB9FE55FAB6D888" blockId="13.[810,1460,143,1765]" pageId="13" pageNumber="14"> +<emphasis id="B9F8EA99F23DFFDEFCB9FE55FC02DCE6" box="[837,962,427,450]" italics="true" pageId="13" pageNumber="14">Horny scutes</emphasis> +: The gular scutes are fused in a single unit that, in all cases ( +<figureCitation id="13B72A0EF23DFFDEFC4BFE37FBECDCC5" box="[951,1068,457,481]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="13" pageNumber="14">Fig. 1C, D</figureCitation> +), except one, reaches posteriorly to the anterior edge of the entoplastron (without entering it). In SAM -PQL -21109 ( +<figureCitation id="13B72A0EF23DFFDEFC0BFDF6FBF3DF04" box="[1015,1075,520,544]" captionStart="Figure 5" captionStartId="7.[113,178,1626,1650]" captionTargetBox="[115,1456,145,1596]" captionTargetId="figure-177@7.[114,1458,144,1598]" captionTargetPageId="7" captionText="Figure 5. Chersina langebaanwegi, paratype SAM-PQL-21109, male: pygal area in posterior view (A); posterior portion of the shell in right lateral view (B); anterior lobe of the plastron in ventral and dorsal views (C, D); posterior lobe in ventral and dorsal views (E,F). Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 5</figureCitation> +), however, it stops anteriorly to the entoplastron as shown by the incomplete, but informative, left epiplastron. The gularohumeral sulcus is laterally concave on the dorsal surface and weakly anteromedially convex on the ven - tral surface. The angle at the junction of the two gularohumeral sulci (the right and the left) varies in relation to the form of the gular projection: from slightly less than 90° (SAM -PQL -21174), to approximately 90° (SAM -PQL -20734, 20925, 35023d), to slightly more than 90° (SAM -PQL -22076). The ratio between the length of the gulars and the humerals (the median antero - posterior length of the humerals) seems to vary according to the development of the gular projections and, therefore, the sex of the specimen: the female SAM -PQL -20734 has a gular as long as the humerals, whereas all the male plastral have humerals that are at least slightly longer than the gular. The humeropectoral su - ture regularly passes posterior to the entoplastron (that, there - fore, is crossed only by the interhumeral suture), and is regularly concave in the anterior direction but has a rather variable trend: it is much more open in the males SAM -PQL -20925 ( +<figureCitation id="13B72A0EF23DFFDEFABAFBC2FABDD970" box="[1350,1405,1084,1108]" captionStart="Figure 4" captionStartId="6.[129,194,1420,1444]" captionTargetBox="[131,1471,146,1390]" captionTargetId="figure-285@6.[129,1473,144,1392]" captionTargetPageId="6" captionText="Figure 4. Chersina langebaanwegi, paratype SAM-PQL-20925: male plastron in ventral view (A) and detail of the anterior lobe of the plastron in dorsal view (B); posterior lobe of the plastron in ventral view (C); pygal area in posterior view (D). Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 4</figureCitation> +) and SAM -PQL -22076 ( +<figureCitation id="13B72A0EF23DFFDEFC0FFBA5FBEAD957" box="[1011,1066,1115,1139]" captionStart="Figure 8" captionStartId="10.[129,194,1839,1863]" captionTargetBox="[137,1465,146,1809]" captionTargetId="figure-7@10.[136,1467,144,1811]" captionTargetPageId="10" captionText="Figure 8. Chersina langebaanwegi, paratype SAM-PQL-PQL-22076:male plastron in ventral (A), dorsal (B), left lateral (C), and anterior (D) views.Scale bars equal 5 cm." pageId="13" pageNumber="14">Fig. 8</figureCitation> +), where the medial sector of the sulcus is nearly straight and perpendicular to the sagittal axis of the plastron, than in the holotypic female shell SAM -PQL -20734 ( +<figureCitation id="13B72A0EF23DFFDEFCC9FB47FC61D9F5" box="[821,929,1209,1233]" captionStart="Figure 1" captionStartId="3.[113,178,1312,1336]" captionTargetBox="[117,1455,147,1281]" captionTargetId="figure-340@3.[114,1458,144,1284]" captionTargetPageId="3" captionText="Figure 1. Chersina langebaanwegi, holotype SAM-PQL-20734: photos (A, C, E, G) and interpretative drawings (B, D, F) of the nearly complete female shell in dorsal (A, B), ventral (C, D), left lateral (E, F), and anterior (G) views.Abbreviations:Ab, abdominal scute; An, anal scute; Ax, axillary scute; Ce, cervical scute; co, costal; en, entoplastron; ep, epiplastron; Fe, femoral scute; Gu, gular scute; Hu, humeral scute; hyo, hyoplastron; hyp, hypoplastron; In, inguinal scute; Ma, marginal scute; ne, neural; nu, nuchal; Pe, pectoral scute; per, peripheral; Pl, pleural scute; py, pygal; Ve, vertebral scute; xi, xiphiplastron. Scale bar equals 5 cm." pageId="13" pageNumber="14">Fig. 1C, D</figureCitation> +). The pectoroabdominal sulci are modestly convex in the anterior direction and reach medially the mid of the interhyoplastral suture. The abdominofemoral sulci are located close to the hypoxiphyplastral suture but do not reach it medi - ally. They are laterally slightly convex in the anterior direction and medially nearly straight. The femoroanal sulci are mod - erately sigmoid and reach, with an acute posterior angle, the interxiphyplastral suture slightly anterior to its midlength. +</paragraph> +<paragraph id="8B33368BF23DFFDEFCB9FA4AFC3CDBC1" blockId="13.[810,1460,143,1765]" pageId="13" pageNumber="14">Axillary and inguinal scutes are present. The first ones (one per side) are relatively small and develop mostly on the third peripheral but also on the hyoplastron. The large inguinal scutes develop mainly at the posterolateral corner of each hypoplastron as well as partly on the seventh peripheral. The inguinal scute contacts the femoral scute both on the ventral and (especially) on the lateral surface of the hypoplastron. In visceral view, there is a significant covering of the plastral scutes on the dorsal sur - face of the bony elements delimiting the anterior and posterior aperture of the shell.</paragraph> +<paragraph id="8B33368BF23DFFDEFCAAF8FFFA47DA3F" blockId="13.[810,1459,1792,1981]" box="[854,1415,1792,1819]" pageId="13" pageNumber="14"> +<emphasis id="B9F8EA99F23DFFDEFCAAF8FFFB50DA3F" bold="true" box="[854,1168,1793,1819]" pageId="13" pageNumber="14">Critical specimens referred to</emphasis> +<taxonomicName id="4C8C4D08F23DFFDEFB68F8FEFA47DA3F" authority="Ne" authorityName="Delfino & Cohen & Govender & Haarhoff & Macaluso & Marino & Maưhews & Wencker & Pavia" authorityYear="2024" box="[1172,1415,1792,1819]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F23DFFDEFB68F8FEFA47DA3F" bold="true" box="[1172,1415,1792,1819]" italics="true" pageId="13" pageNumber="14">Chersina langebaanwegi</emphasis> +</taxonomicName> +</paragraph> +<paragraph id="8B33368BF23DFFDDFCD6F8D6FE67DFF8" blockId="13.[810,1459,1792,1981]" lastBlockId="14.[129,778,144,732]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14"> +The tentative referral of the female right epiplastron SAM - PQL -24826b to +<taxonomicName id="4C8C4D08F23DFFDEFC2BF8B9FB44DA7B" authorityName="Delfino & Cohen & Govender & Haarhoff & Macaluso & Marino & Maưhews & Wencker & Pavia" authorityYear="2024" box="[983,1156,1863,1887]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F23DFFDEFC2BF8B9FB44DA7B" box="[983,1156,1863,1887]" italics="true" pageId="13" pageNumber="14">C. langebaanwegi</emphasis> +</taxonomicName> +, despite the dorsal convexity of the gular lip and its exceptional posterior development (somehow similar to the extant +<taxonomicName id="4C8C4D08F23DFFDEFB7DF878FB38DAB9" box="[1153,1272,1925,1949]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F23DFFDEFB7DF878FB38DAB9" box="[1153,1272,1925,1949]" italics="true" pageId="13" pageNumber="14">C. angulata</emphasis> +</taxonomicName> +), is based on the fact that the same degree of lip convexity is shared by the anterior portion of the female anterior plastral lobe SAM -PQL - 12754a that includes the two epiplastra and the entoplastron: the latter has an entoplastron wider that long, and the posterior edge of the gular lip in the epiplastron is straight, as typically shown by most of the specimens of +<taxonomicName id="4C8C4D08F23EFFDDFE0EFEF0FD5FDC01" authorityName="Delfino & Cohen & Govender & Haarhoff & Macaluso & Marino & Maưhews & Wencker & Pavia" authorityYear="2024" box="[498,671,269,293]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="langebaanwegi"> +<emphasis id="B9F8EA99F23EFFDDFE0EFEF0FD5FDC01" box="[498,671,269,293]" italics="true" pageId="14" pageNumber="15">C. langebaanwegi</emphasis> +</taxonomicName> +. A convex epiplastral lip also characterizes the male left epiplastron SAM - PQL -24827a which shows a straight posterior edge of the gular lip. The same condition (convex epiplastral lip poster - iorly straight) is shown by the right female epiplastron SAM - PQL -73749. Both SAM -PQL -24826b and SAM -PQL -24827a have the tip of the gular protrusion slightly pointing in the anteroventral direction. Among the referred material, a convex epiplastral lip associated with a variably shaped posterior edge of the lip (and, therefore, a variably developed pocket) is pre - sent in the male epiplastra SAM -PQL -73750 and SAM -PQL - 12740c. The female right epiplastron SAM -PQL -73878 shows an intermediate condition in having a rather flat gular lip that ends posteriorly with a marked convexity that gives origin to a moderate pocket ( +<figureCitation id="13B72A0EF23EFFDDFEBDFD3AFE57DFF8" box="[321,407,708,732]" captionStart="Figure 10" captionStartId="12.[129,194,1334,1358]" captionTargetBox="[132,1471,147,1304]" captionTargetId="figure-312@12.[129,1473,144,1306]" captionTargetPageId="12" captionText="Figure 10. Chersina langebaanwegi, male left epiplastron SAM-PQL-12750c and female right SAM-PQL-73878 in dorsal (A, D, respectively), ventral (B, E, respectively), and medial (C, F, respectively) views. Note the sexual dimorphism of the gular projection and the variability of the development epiplastral pad (compare also with previous figures). Scale bars equal 7 cm." pageId="14" pageNumber="15">Fig. 10F</figureCitation> +). +</paragraph> +</subSubSection> +<subSubSection id="C3966500F23EFFDDFF03FD05FCC9D80B" pageId="14" pageNumber="15" type="discussion"> +<paragraph id="8B33368BF23EFFDDFF03FD05FD40DE11" blockId="14.[128,779,763,1327]" pageId="14" pageNumber="15"> +<emphasis id="B9F8EA99F23EFFDDFF03FD05FD40DE11" bold="true" pageId="14" pageNumber="15">Phylogenetic relationships of the new extinct species from Langebaanweg</emphasis> +</paragraph> +<paragraph id="8B33368BF23EFFDDFF7DFCBCFEB7D931" blockId="14.[128,779,763,1327]" pageId="14" pageNumber="15"> +The phylogenetic analysis resulted in the consensus trees re - ported in +<figureCitation id="13B72A0EF23EFFDDFF17FC9FFE90DE5D" box="[235,336,865,889]" captionStart="Figure 11" captionStartId="15.[113,178,1447,1471]" captionTargetBox="[117,1455,147,1416]" captionTargetId="figure-213@15.[114,1458,144,1419]" captionTargetPageId="15" captionText="Figure 11. Strict consensus trees without (A) and with (B) a molecular scaffold to constrain the position of extant taxa (according to the topologies of Thomson et al.2021 and Fritz et al. 2022). Both the topologies result from maximum parsimony phylogenetic analyses under implied weights (k = 6) of 42 extant taxa and four extinct taxa (highlighted with diamonds), using 171 osteological characters.The analysis is based on the character matrix and character definitions of Vlachos and Rabi (2018) but all extinct taxa, except the African ones coming from South of the Equator (Impregnochelys pachytectis, Mesochersus orangeus, and Namibchersus namaquensis), were removed and four extant taxa (Astrochelys yniphora, Chersina angulata, Chersobius signatus, and Pyois arachnoides), scored for shell characters only, were added." pageId="14" pageNumber="15">Figure 11</figureCitation> +. The first round of the analysis without a molecular scaffold ( +<figureCitation id="13B72A0EF23EFFDDFEACFC7EFE69DEBC" box="[336,425,896,920]" captionStart="Figure 11" captionStartId="15.[113,178,1447,1471]" captionTargetBox="[117,1455,147,1416]" captionTargetId="figure-213@15.[114,1458,144,1419]" captionTargetPageId="15" captionText="Figure 11. Strict consensus trees without (A) and with (B) a molecular scaffold to constrain the position of extant taxa (according to the topologies of Thomson et al.2021 and Fritz et al. 2022). Both the topologies result from maximum parsimony phylogenetic analyses under implied weights (k = 6) of 42 extant taxa and four extinct taxa (highlighted with diamonds), using 171 osteological characters.The analysis is based on the character matrix and character definitions of Vlachos and Rabi (2018) but all extinct taxa, except the African ones coming from South of the Equator (Impregnochelys pachytectis, Mesochersus orangeus, and Namibchersus namaquensis), were removed and four extant taxa (Astrochelys yniphora, Chersina angulata, Chersobius signatus, and Pyois arachnoides), scored for shell characters only, were added." pageId="14" pageNumber="15">Fig. 11A</figureCitation> +; Consistency Index (CI) = 0.293; Retention Index (RI) = 0.620; Most Parsimonious Trees (MPT) = 47.997) found two most parsimonious trees, and the second round of tree bisection and reconnection retained one most parsimonious tree. +</paragraph> +<paragraph id="8B33368BF23EFFDDFF60FBE3FCC9D80B" blockId="14.[128,779,763,1327]" pageId="14" pageNumber="15"> +In the analysis with the molecular scaffold ( +<figureCitation id="13B72A0EF23EFFDDFD5EFBE3FCC2D911" box="[674,770,1053,1077]" captionStart="Figure 11" captionStartId="15.[113,178,1447,1471]" captionTargetBox="[117,1455,147,1416]" captionTargetId="figure-213@15.[114,1458,144,1419]" captionTargetPageId="15" captionText="Figure 11. Strict consensus trees without (A) and with (B) a molecular scaffold to constrain the position of extant taxa (according to the topologies of Thomson et al.2021 and Fritz et al. 2022). Both the topologies result from maximum parsimony phylogenetic analyses under implied weights (k = 6) of 42 extant taxa and four extinct taxa (highlighted with diamonds), using 171 osteological characters.The analysis is based on the character matrix and character definitions of Vlachos and Rabi (2018) but all extinct taxa, except the African ones coming from South of the Equator (Impregnochelys pachytectis, Mesochersus orangeus, and Namibchersus namaquensis), were removed and four extant taxa (Astrochelys yniphora, Chersina angulata, Chersobius signatus, and Pyois arachnoides), scored for shell characters only, were added." pageId="14" pageNumber="15">Fig. 11B</figureCitation> +; CI = 0.268, RI = 0.572; MPT 53.1423), the New Technology Search found two most parsimonious trees, and the second round of tree bisection and reconnection retained two most par - simonious trees. Both the analyses confirm that the new extinct species from Langebaanweg is the sister taxon of +<taxonomicName id="4C8C4D08F23EFFDDFD73FB44FCC4D9F5" box="[655,772,1209,1233]" class="Testudines" family="Testudinidae" genus="Chersina" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="angulata"> +<emphasis id="B9F8EA99F23EFFDDFD73FB44FCC4D9F5" box="[655,772,1209,1233]" italics="true" pageId="14" pageNumber="15">C. angulata</emphasis> +</taxonomicName> +. Remarkably, despite sharing with +<taxonomicName id="4C8C4D08F23EFFDDFE01FB27FD62D9D4" box="[509,674,1241,1265]" class="Testudines" family="Testudinidae" genus="Astrochelys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="undetermined"> +<emphasis id="B9F8EA99F23EFFDDFE01FB27FDA9D9D5" box="[509,617,1241,1265]" italics="true" pageId="14" pageNumber="15">Astrochelys</emphasis> +spp. +</taxonomicName> +a highly domed shell, and with +<taxonomicName id="4C8C4D08F23EFFDDFE8CFB06FE25D834" baseAuthorityName="Vaillant" baseAuthorityYear="1885" box="[368,485,1272,1296]" class="Testudines" family="Testudinidae" genus="Astrochelys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="yniphora"> +<emphasis id="B9F8EA99F23EFFDDFE8CFB06FE25D834" box="[368,485,1272,1296]" italics="true" pageId="14" pageNumber="15">A. yniphora</emphasis> +</taxonomicName> +a long gular protrusion and a single gular shield, it is not retrieved in the clade of +<taxonomicName id="4C8C4D08F23EFFDDFD6BFAE9FCC3D80B" authorityName="J.E.Gray" authorityYear="1873" box="[663,771,1303,1327]" class="Testudines" family="Testudinidae" genus="Astrochelys" kingdom="Animalia" order="Cryptodira" pageId="14" pageNumber="15" phylum="Chordata" rank="genus"> +<emphasis id="B9F8EA99F23EFFDDFD6BFAE9FCC3D80B" box="[663,771,1303,1327]" italics="true" pageId="14" pageNumber="15">Astrochelys</emphasis> +</taxonomicName> +. +</paragraph> +</subSubSection> +</treatment> +</document> \ No newline at end of file diff --git a/data/87/5D/87/875D87D0FFF8FFDAFF61FA5A089DFE0F.xml b/data/87/5D/87/875D87D0FFF8FFDAFF61FA5A089DFE0F.xml new file mode 100644 index 00000000000..eabd371ee47 --- /dev/null +++ b/data/87/5D/87/875D87D0FFF8FFDAFF61FA5A089DFE0F.xml @@ -0,0 +1,1111 @@ +<document id="7C55AADEE7FECE4FB064334C717FAC71" ID-DOI="10.11646/phytotaxa.674.3.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="14520725" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1734555491044" checkinUser="felipe" docAuthor="Wang, Zhongde, Lan, Yaoqing, Huang, Qilin & Tong, Yi" docDate="2024" docId="875D87D0FFF8FFDAFF61FA5A089DFE0F" docLanguage="en" docName="phytotaxa.674.3.252-264.pdf" docOrigin="Phytotaxa 674 (3)" docSource="https://doi.org/10.11646/phytotaxa.674.3.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Mahonia nanlingensis Y. Tong & Z. D. Wang 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="260" masterDocId="7B64FFA8FFFBFFD3FFE9FFCE0D11FFD7" masterDocTitle="Mahonia nanlingensis (Berberidaceae), a new species from China" masterLastPageNumber="264" masterPageNumber="252" pageNumber="255" updateTime="1741807922850" updateUser="guilherme" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED"> +<mods:mods id="2DF59A18E9A89091AC6FF605672D14B2" xmlns:mods="http://www.loc.gov/mods/v3"> +<mods:titleInfo id="460FECCB6E722556224A79DFA17C1846"> +<mods:title id="CC33188EADEA2299B1A737010B42F462">Mahonia nanlingensis (Berberidaceae), a new species from China</mods:title> +</mods:titleInfo> +<mods:name id="2BB7EB9673BDC4D01A9B01086255D1EC" type="personal"> +<mods:role id="AFBBB6892B3749DAADB7AB28D9079EE8"> +<mods:roleTerm id="DFE5353C8A3AD00A4CCCB0694B2D19DF">Author</mods:roleTerm> +</mods:role> +<mods:namePart id="25B2A8C2E7FE3377A352CAB017E7E156">Wang, Zhongde</mods:namePart> +<mods:nameIdentifier id="DD3891F7E9DAB2D3FE1E020DE146E338" type="ORCID">0009-0009-1085-8813</mods:nameIdentifier> +<mods:affiliation id="639EAB45E81B08B6B4535CFF89521608">School of Pharmaceutical Sciences, Guangzhou University of Chinese Medicine, Guangzhou 510006, China</mods:affiliation> +<mods:nameIdentifier id="91F735039C97CD824E72E77610326A58" type="email">wzd_mahonia@163.com</mods:nameIdentifier> +</mods:name> +<mods:name id="895BD178F61DACD1EF536F176EF088F2" type="personal"> +<mods:role id="FAEEC878E908F43E22E704A5ABA9C5D3"> +<mods:roleTerm id="7B7E0832855ECB6F361FA2E8BE1F6A8F">Author</mods:roleTerm> +</mods:role> +<mods:namePart id="770BFC4FC04B44A6ED431769F802418A">Lan, Yaoqing</mods:namePart> +<mods:nameIdentifier id="17EB64E2199E430C90E15963E8549160" type="ORCID">0009-0007-3476-5801</mods:nameIdentifier> +<mods:affiliation id="46CA4139DA4B4D6AF2381F48BB99C9DC">School of Pharmaceutical Sciences, Guangzhou University of Chinese Medicine, Guangzhou 510006, China</mods:affiliation> +<mods:nameIdentifier id="8B22E27B5887C226A4FD52686278B833" type="email">861510723@qq.com</mods:nameIdentifier> +</mods:name> +<mods:name id="922DB76363832DD7D15EB9F0F6CD6E04" type="personal"> +<mods:role id="189166C3D417F1C1689BF10D136D347C"> +<mods:roleTerm id="EB819858452996466A01DCD824B5E04C">Author</mods:roleTerm> +</mods:role> +<mods:namePart id="F326BEB80B7BFC3834108DFAAA44F760">Huang, Qilin</mods:namePart> +<mods:nameIdentifier id="33ED0E0073F10726CA8CC8761E9F3025" type="ORCID">0009-0004-1642-1093</mods:nameIdentifier> +<mods:affiliation id="C4CA057D1AB5369F57D1460C78464E39">School of Pharmaceutical Sciences, Guangzhou University of Chinese Medicine, Guangzhou 510006, China</mods:affiliation> +<mods:nameIdentifier id="3300AE67A10E03F927B5E7B8E76D8207" type="email">2651245438@qq.com</mods:nameIdentifier> +</mods:name> +<mods:name id="A641AD7F51CC6228C90DCCEE1CAEC62C" type="personal"> +<mods:role id="70579F82F98573678703EE2A8B8E9D1A"> +<mods:roleTerm id="CCDB315A5934C2416133060D5A77C3C6">Author</mods:roleTerm> +</mods:role> +<mods:namePart id="019889F9F740EDD602A0906741E5A1A9">Tong, Yi</mods:namePart> +<mods:nameIdentifier id="83107615070E106B975DBAB9841BD50F" type="ORCID">0000-0003-3928-3725</mods:nameIdentifier> +<mods:affiliation id="132D681ED5A31852E59B4A6CAABE8E34">School of Pharmaceutical Sciences, Guangzhou University of Chinese Medicine, Guangzhou 510006, China</mods:affiliation> +<mods:nameIdentifier id="8FA07ADB6B2A6A80A81A2390B3CEF61E" type="email">tongyi@gzucm.edu.cn</mods:nameIdentifier> +</mods:name> +<mods:typeOfResource id="17A7FF4DF525F9734A3D1CBD01BE3F08">text</mods:typeOfResource> +<mods:relatedItem id="6FDB2E27CAD0B4D34FEE49E20DEA7674" type="host"> +<mods:titleInfo id="19E441152263B85299A40F9737771052"> +<mods:title id="B452200C20749B0CDFD055112F3637EC">Phytotaxa</mods:title> +</mods:titleInfo> +<mods:part id="17D0445904FDF3013D2FE83074A5E098"> +<mods:date id="13874AF087167359F0756E0D0BDA8B76">2024</mods:date> +<mods:detail id="F4AC5E9B974E9938E516A5107B3B9E06" type="pubDate"> +<mods:number id="F166B823F7DA7A675C8A4DA126A58BFF">2024-11-28</mods:number> +</mods:detail> +<mods:detail id="B405C6BAA1737825B5C940B5D78CEC2A" type="volume"> +<mods:number id="3E2345DDA003036BCA64189430CF9270">674</mods:number> +</mods:detail> +<mods:detail id="E029472BB2CEC6A23B8E0FFEBC69CEB4" type="issue"> +<mods:number id="4FCDBBDD79527A987FF1138608F646BF">3</mods:number> +</mods:detail> +<mods:extent id="48D2274F9CADEF7281A29B48443289CA" unit="page"> +<mods:start id="1408EDFAD62F4D17B3E36EE2747256D0">252</mods:start> +<mods:end id="676821734949416539AD657139F189DE">264</mods:end> +</mods:extent> +</mods:part> +</mods:relatedItem> +<mods:location id="9B8B461E10E8E3BFCF300DA903B4DFF8"> +<mods:url id="28801C3E52C5D3478B9B42A228E14C6D">https://doi.org/10.11646/phytotaxa.674.3.2</mods:url> +</mods:location> +<mods:classification id="35E6AAD68BA5E83BB3E5FE656B470277">journal article</mods:classification> +<mods:identifier id="5B3B9A29CDD90AE83A902D07195FFA56" type="DOI">10.11646/phytotaxa.674.3.2</mods:identifier> +<mods:identifier id="C4D2D619124CFA31A683A83BBE1FF8A8" type="ISSN">1179-3163</mods:identifier> +<mods:identifier id="6551082D7131BC5FEF386E7CE59A2803" type="Zenodo-Dep">14520725</mods:identifier> +</mods:mods> +<treatment id="875D87D0FFF8FFDAFF61FA5A089DFE0F" LSID="urn:lsid:plazi:treatment:875D87D0FFF8FFDAFF61FA5A089DFE0F" httpUri="http://treatment.plazi.org/id/875D87D0FFF8FFDAFF61FA5A089DFE0F" lastPageId="9" lastPageNumber="260" pageId="3" pageNumber="255"> +<subSubSection id="47EE654DFFF8FFD0FF61FA5A0E47FA78" box="[136,854,1428,1455]" pageId="3" pageNumber="255" type="nomenclature"> +<paragraph id="0F4B36C6FFF8FFD0FF61FA5A0E47FA78" blockId="3.[136,1452,1428,1995]" box="[136,854,1428,1455]" pageId="3" pageNumber="255"> +<taxonomicName id="C8F44D45FFF8FFD0FF61FA5A0F63FA78" authority="Y.Tong & Z.D.Wang" authorityName="Y. Tong & Z. D. Wang" authorityYear="2024" box="[136,626,1428,1455]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="nanlingensis" status="sp. nov."> +<emphasis id="3D80EAD4FFF8FFD0FF61FA5A0C93FA79" box="[136,386,1428,1454]" italics="true" pageId="3" pageNumber="255">Mahonia nanlingensis</emphasis> +Y.Tong & Z.D.Wang +</taxonomicName> +, +<taxonomicNameLabel id="26B357AFFFF8FFD0FD97FA5B0FC5FA78" box="[638,724,1429,1455]" pageId="3" pageNumber="255" rank="species"> +<emphasis id="3D80EAD4FFF8FFD0FD97FA5B0FDCFA79" box="[638,717,1429,1454]" italics="true" pageId="3" pageNumber="255">sp. nov</emphasis> +. +</taxonomicNameLabel> +( +<figureCitation id="97CF2A43FFF8FFD0FD0DFA5A0E5FFA78" box="[740,846,1428,1455]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="5.[136,229,1953,1975]" captionStartId-1="6.[136,234,1030,1052]" captionStartId-2="6.[136,234,2027,2049]" captionStartId-3="7.[136,234,1591,1613]" captionTargetBox-0="[169,1426,194,1918]" captionTargetBox-1="[164,1421,187,1002]" captionTargetBox-2="[164,1422,1077,2003]" captionTargetBox-3="[151,1436,190,1567]" captionTargetId-0="figure-27@5.[162,1426,187,1925]" captionTargetId-1="figure-19@6.[164,1423,187,1002]" captionTargetId-2="figure-46@6.[164,1423,1077,2003]" captionTargetId-3="figure-27@7.[151,1436,190,1567]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 2. Illustration of Mahonia nanlingensis A. Habit. B. Outer sepal. C. Median sepal. D. Inner sepal. E. Petal. F. Stamen. G. Pistil. H. Berry and bract. I–J. Leaflets. Drawn by Qi-Lin Huang." captionText-1="FIGURE 3. Mahonia nanlingensis sp. nov. A. leaves. B. seedling. C. flower. D & E. habit. F. infructescence." captionText-2="FIGURE 4. Mahonia nanlingensis sp. nov. A. habit. B. tender leaf. C–D. bud scales. E. transection of stem. F–G. inflorescences. H. bud scales of inflorescence. I. infructescences." captionText-3="FIGURE 5. Mahonia nanlingensis sp. nov. A. outer sepal. B. median sepal. C. inner sepal. D. petal. E. petal with stamen. F. another shape of outer sepal. G–I. stamens. J. pistil. K. pedicel and bract. L. berry." figureDoi-0="http://doi.org/10.5281/zenodo.14520729" figureDoi-1="http://doi.org/10.5281/zenodo.14520733" figureDoi-2="http://doi.org/10.5281/zenodo.14520735" figureDoi-3="http://doi.org/10.5281/zenodo.14520737" httpUri-0="https://zenodo.org/record/14520729/files/figure.png" httpUri-1="https://zenodo.org/record/14520733/files/figure.png" httpUri-2="https://zenodo.org/record/14520735/files/figure.png" httpUri-3="https://zenodo.org/record/14520737/files/figure.png" pageId="3" pageNumber="255">Figs. 2–5</figureCitation> +) +</paragraph> +</subSubSection> +<subSubSection id="47EE654DFFF8FFD0FF54FA760E65F9CD" pageId="3" pageNumber="255" type="materials_examined"> +<paragraph id="0F4B36C6FFF8FFD0FF54FA760E65F9CD" blockId="3.[136,1452,1428,1995]" pageId="3" pageNumber="255"> +<emphasis id="3D80EAD4FFF8FFD0FF54FA760DE5FA05" bold="true" box="[189,244,1464,1490]" italics="true" pageId="3" pageNumber="255">Type</emphasis> +:— +<materialsCitation id="BF9C3C9BFFF8FFD0FEFFFA760F88F9CD" collectingDate="2023-10-22" collectionCode="IBSC" collectorName="Y. Tong & Z. D. Wang & Z. C. Fan" country="China" county="Ruyuan Yao Autonomous County" elevation="535" latitude="24.967445" location="Tianzhushan Village" longLatPrecision="1" longitude="113.18407" municipality="Daqiao Town" pageId="3" pageNumber="255" specimenCount="1" stateProvince="Guangdong Province" typeStatus="holotype"> +<collectingCountry id="77E37656FFF8FFD0FEFFFA760C65FA05" box="[278,372,1464,1490]" name="China" pageId="3" pageNumber="255">CHINA</collectingCountry> +. +<collectingRegion id="CD30F824FFF8FFD0FE6BFA760F65FA05" box="[386,628,1464,1490]" country="China" name="Guangdong" pageId="3" pageNumber="255">Guangdong Province</collectingRegion> +, +<collectingCounty id="E62A4E4AFFF8FFD0FD6BFA770914FA05" box="[642,1029,1464,1491]" pageId="3" pageNumber="255">Ruyuan Yao Autonomous County</collectingCounty> +, +<collectingMunicipality id="EF2FACBCFFF8FFD0FBFBFA7609BEFA04" box="[1042,1199,1464,1491]" pageId="3" pageNumber="255">Daqiao Town</collectingMunicipality> +, +<location id="0A2B601DFFF8FFD0FB55FA7608B9FA05" LSID="urn:lsid:plazi:treatment:875D87D0FFF8FFDAFF61FA5A089DFE0F:0A2B601DFFF8FFD0FB55FA7608B9FA05" box="[1212,1448,1464,1490]" country="China" county="Ruyuan Yao Autonomous County" latitude="24.967445" longLatPrecision="1" longitude="113.18407" municipality="Daqiao Town" name="Tianzhushan Village" pageId="3" pageNumber="255" stateProvince="Guangdong Province">Tianzhushan Village</location> +, +<geoCoordinate id="6AC05001FFF8FFD0FF61FA120C39FA20" box="[136,296,1500,1527]" degrees="24" direction="north" minutes="58" orientation="latitude" pageId="3" pageNumber="255" precision="1" seconds="2.80" value="24.967445">24°58’2.80”N</geoCoordinate> +, +<geoCoordinate id="6AC05001FFF8FFD0FEDEFA120CF1FA20" box="[311,480,1500,1527]" degrees="113" direction="east" minutes="11" orientation="longitude" pageId="3" pageNumber="255" precision="1" seconds="2.65" value="113.18407">113°11’2.65”E</geoCoordinate> +, +<elevation id="84D9D1F5FFF8FFD0FE07FA120F66FA21" box="[494,631,1500,1527]" metricMagnitude="2" metricUnit="m" metricValue="5.35" pageId="3" pageNumber="255" unit="m" value="535.0"> +<quantity id="C80C9B23FFF8FFD0FE07FA120F29FA20" box="[494,568,1500,1527]" metricMagnitude="2" metricUnit="m" metricValue="5.35" pageId="3" pageNumber="255" unit="m" value="535.0">535 m</quantity> +a.s.l. +</elevation> +, broad-leaved evergreen forest margin, +<date id="7B4A1006FFF8FFD0FBABFA1209C5FA21" box="[1090,1236,1500,1526]" pageId="3" pageNumber="255" value="2023-10-22"> +<collectingDate id="6B0EE9EEFFF8FFD0FBABFA1209C5FA21" box="[1090,1236,1500,1526]" pageId="3" pageNumber="255" value="2023-10-22">22 Oct 2023</collectingDate> +</date> +, fr., +<emphasis id="3D80EAD4FFF8FFD0FAE2FA130CC7F9CD" italics="true" pageId="3" pageNumber="255"> +<collectorName id="A2015310FFF8FFD0FAE2FA130873FA21" box="[1291,1378,1501,1526]" pageId="3" pageNumber="255">Y. Tong</collectorName> +, +<collectorName id="A2015310FFF8FFD0FA99FA130DDBF9CD" pageId="3" pageNumber="255">Z. D. Wang</collectorName> +, +<collectorName id="A2015310FFF8FFD0FF3CF9CF0C51F9CD" box="[213,320,1536,1562]" pageId="3" pageNumber="255">Z. C. Fan</collectorName> +TY23102201 +</emphasis> +( +<typeStatus id="D04F8864FFF8FFD0FE0CF9CE0F5CF9CD" box="[485,589,1536,1562]" pageId="3" pageNumber="255" type="holotype">holotype</typeStatus> +: +<collectionCode id="69E5AE03FFF8FFD0FDBEF9CE0F88F9CD" box="[599,665,1536,1562]" pageId="3" pageNumber="255">IBSC</collectionCode> +</materialsCitation> +; +<materialsCitation id="BF9C3C9BFFF8FFD0FD4DF9CE0E79F9CD" box="[676,872,1536,1562]" collectingDate="2023-10-22" collectionCode="GUCM" collectorName="Y. Tong & Z. D. Wang & Z. C. Fan" country="China" county="Ruyuan Yao Autonomous County" elevation="535" latitude="24.967445" location="Tianzhushan Village" longLatPrecision="1" longitude="113.18407" municipality="Daqiao Town" pageId="3" pageNumber="255" specimenCount="1" stateProvince="Guangdong Province" typeStatus="isotype"> +<typeStatus id="D04F8864FFF8FFD0FD4DF9CE0E15F9CD" box="[676,772,1536,1562]" pageId="3" pageNumber="255" type="isotype">isotypes</typeStatus> +: +<collectionCode id="69E5AE03FFF8FFD0FCE6F9CE0E79F9CD" box="[783,872,1536,1562]" pageId="3" pageNumber="255">GUCM</collectionCode> +</materialsCitation> +). +</paragraph> +</subSubSection> +<subSubSection id="47EE654DFFF8FFD0FF54F9EA0950F81D" pageId="3" pageNumber="255" type="diagnosis"> +<paragraph id="0F4B36C6FFF8FFD0FF54F9EA0950F81D" blockId="3.[136,1452,1428,1995]" pageId="3" pageNumber="255"> +<emphasis id="3D80EAD4FFF8FFD0FF54F9EA0C21F9E9" bold="true" box="[189,304,1572,1598]" italics="true" pageId="3" pageNumber="255">Diagnosis</emphasis> +:— +<taxonomicName id="C8F44D45FFF8FFD0FEB2F9EA0F45F9E9" authorityName="Y. Tong & Z. D. Wang" authorityYear="2024" box="[347,596,1572,1598]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="nanlingensis"> +<emphasis id="3D80EAD4FFF8FFD0FEB2F9EA0F45F9E9" box="[347,596,1572,1598]" italics="true" pageId="3" pageNumber="255">Mahonia nanlingensis</emphasis> +</taxonomicName> +is most similar to +<taxonomicName id="C8F44D45FFF8FFD0FCC9F9EB0EBDF9E9" authorityName="Gagnepain" authorityYear="1908" box="[800,940,1572,1598]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="bodinieri"> +<emphasis id="3D80EAD4FFF8FFD0FCC9F9EB0EBDF9E9" box="[800,940,1572,1598]" italics="true" pageId="3" pageNumber="255">M. bodinieri</emphasis> +</taxonomicName> +in leaves and fruit. However, +<taxonomicName id="C8F44D45FFF8FFD0FB10F9EB08BDF9E9" authorityName="Y. Tong & Z. D. Wang" authorityYear="2024" box="[1273,1452,1572,1598]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="nanlingensis"> +<emphasis id="3D80EAD4FFF8FFD0FB10F9EB08BDF9E9" box="[1273,1452,1572,1598]" italics="true" pageId="3" pageNumber="255">M. nanlingensis</emphasis> +</taxonomicName> +can be readily distinguished by its cuspidate leaflet apex, truncate stamen apex, narrowly incised petal apex, and emarginate inner sepals. Additionally, +<taxonomicName id="C8F44D45FFF8FFD0FDDDF9A30FF6F951" authorityName="Y. Tong & Z. D. Wang" authorityYear="2024" box="[564,743,1644,1670]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="nanlingensis"> +<emphasis id="3D80EAD4FFF8FFD0FDDDF9A30FF6F951" box="[564,743,1644,1670]" italics="true" pageId="3" pageNumber="255">M. nanlingensis</emphasis> +</taxonomicName> +has a shallowly toothed apex on the lower leaflets, with shorter apical teeth (approximately +<quantity id="C80C9B23FFF8FFD0FE2BF95E0F01F97D" box="[450,528,1680,1707]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="255" unit="cm" value="0.5">0.5 cm</quantity> +) and 7–9 teeth on each side of the leaflet, while +<taxonomicName id="C8F44D45FFF8FFD0FBDBF95F09AFF97D" authorityName="Gagnepain" authorityYear="1908" box="[1074,1214,1680,1706]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="bodinieri"> +<emphasis id="3D80EAD4FFF8FFD0FBDBF95F09AFF97D" box="[1074,1214,1680,1706]" italics="true" pageId="3" pageNumber="255">M. bodinieri</emphasis> +</taxonomicName> +has 4–6 deeper teeth per side, with a longer apical tooth (approximately +<quantity id="C80C9B23FFF8FFD0FD3DF97A0E03F919" box="[724,786,1716,1742]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="255" unit="cm" value="1.0">1 cm</quantity> +). It is also morphologically similar to +<taxonomicName id="C8F44D45FFF8FFD0FB20F97B086BF919" authorityName="Gagnepain" authorityYear="1908" box="[1225,1402,1716,1743]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="duclouxiana"> +<emphasis id="3D80EAD4FFF8FFD0FB20F97B09F1F919" box="[1225,1248,1717,1742]" italics="true" pageId="3" pageNumber="255">M</emphasis> +. +<emphasis id="3D80EAD4FFF8FFD0FB19F97A086BF919" box="[1264,1402,1716,1742]" italics="true" pageId="3" pageNumber="255">duclouxiana</emphasis> +</taxonomicName> +and +<taxonomicName id="C8F44D45FFF8FFD0FF61F9170C2FF925" authorityName="Takeda" authorityYear="1917" box="[136,318,1752,1778]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="hancockiana"> +<emphasis id="3D80EAD4FFF8FFD0FF61F9170C2FF925" box="[136,318,1752,1778]" italics="true" pageId="3" pageNumber="255">M. hancockiana</emphasis> +</taxonomicName> +in habit and in having 7–11 leaflets with elongate elliptical leaf blades and shallowly serrated leaf margins. However, it can be distinguished by the stout, unbranched inflorescence, broadly spoon-shaped bract, petals basally glandless, apex narrowly incised, ovules 2 per ovary, stigma broadly discoid, pericarp waxy white pruinose with obvious reticulation and carpopodium twice as long as bract. +<taxonomicName id="C8F44D45FFF8FFD0FC93F88A0964F889" authorityName="Y. Tong & Z. D. Wang" authorityYear="2024" box="[890,1141,1860,1886]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="nanlingensis"> +<emphasis id="3D80EAD4FFF8FFD0FC93F88A0964F889" box="[890,1141,1860,1886]" italics="true" pageId="3" pageNumber="255">Mahonia nanlingensis</emphasis> +</taxonomicName> +may also be confused with +<taxonomicName id="C8F44D45FFF8FFD0FF61F8A70C2BF855" authorityName="Ying & Boufford" authorityYear="2001" box="[136,314,1896,1922]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="cardiophylla"> +<emphasis id="3D80EAD4FFF8FFD0FF61F8A70C2BF855" box="[136,314,1896,1922]" italics="true" pageId="3" pageNumber="255">M. cardiophylla</emphasis> +</taxonomicName> +and +<taxonomicName id="C8F44D45FFF8FFD0FE87F8A70CC6F855" authorityName="Carriere" authorityYear="1855" baseAuthorityName="Fortune" baseAuthorityYear="1850" box="[366,471,1896,1922]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="255" phylum="Tracheophyta" rank="species" species="bealei"> +<emphasis id="3D80EAD4FFF8FFD0FE87F8A70CC6F855" box="[366,471,1896,1922]" italics="true" pageId="3" pageNumber="255">M. bealei</emphasis> +</taxonomicName> +, particularly in habit and tender leaves. However, it can be distinguished by leaflets long elliptic, blade abaxially green and not glaucous, margins from the 3rd leaflets 7 or 9 fine-toothed and 5 or 6 pairs of lateral veins from the midrib of each leaflet with apex obviously netted (see +<tableCitation id="4276037DFFF8FFD0FC37F87E0921F81D" box="[990,1072,1968,1994]" captionStart="TABLE 1" captionStartId="4.[136,228,652,678]" captionTargetPageId="4" captionText="TABLE 1. Comparison of Mahonia nanlingensis, M. duclouxiana, M. hancockiana, M. cardiophylla, M. bealei and M. bodinieri." pageId="3" pageNumber="255">Table 1</tableCitation> +). +</paragraph> +</subSubSection> +<caption id="5B8B664EFFFFFFD7FF61FD4208E3FD71" box="[136,1522,652,678]" pageId="4" pageNumber="256" startId="4.[136,228,652,678]" targetBox="[143,2054,697,1603]" targetIsTable="true" targetPageId="4" targetType="table"> +<paragraph id="0F4B36C6FFFFFFD7FF61FD4208E3FD71" blockId="4.[136,1522,652,678]" box="[136,1522,652,678]" pageId="4" pageNumber="256"> +<emphasis id="3D80EAD4FFFFFFD7FF61FD420DEEFD71" bold="true" box="[136,255,652,678]" pageId="4" pageNumber="256">TABLE 1.</emphasis> +Comparison of +<taxonomicName id="C8F44D45FFFFFFD7FE45FD420F8BFD71" authorityName="Y. Tong & Z. D. Wang" authorityYear="2024" box="[428,666,652,678]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="nanlingensis"> +<emphasis id="3D80EAD4FFFFFFD7FE45FD420F8BFD71" box="[428,666,652,678]" italics="true" pageId="4" pageNumber="256">Mahonia nanlingensis</emphasis> +</taxonomicName> +, +<emphasis id="3D80EAD4FFFFFFD7FD4EFD420820FD71" box="[679,1329,652,678]" italics="true" pageId="4" pageNumber="256"> +<taxonomicName id="C8F44D45FFFFFFD7FD4EFD420E40FD71" authorityName="Gagnepain" authorityYear="1908" box="[679,849,652,678]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="duclouxiana">M. duclouxiana</taxonomicName> +, +<taxonomicName id="C8F44D45FFFFFFD7FCB5FD42091BFD71" authorityName="Takeda" authorityYear="1917" box="[860,1034,652,678]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="hancockiana">M. hancockiana</taxonomicName> +, +<taxonomicName id="C8F44D45FFFFFFD7FBFCFD4209D3FD71" authorityName="Ying & Boufford" authorityYear="2001" box="[1045,1218,652,678]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="cardiophylla">M. cardiophylla</taxonomicName> +, +<taxonomicName id="C8F44D45FFFFFFD7FB24FD420820FD71" authorityName="Carriere" authorityYear="1855" baseAuthorityName="Fortune" baseAuthorityYear="1850" box="[1229,1329,652,678]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="bealei">M. bealei</taxonomicName> +</emphasis> +and +<taxonomicName id="C8F44D45FFFFFFD7FA8CFD4208FFFD71" authorityName="Gagnepain" authorityYear="1908" box="[1381,1518,652,678]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="bodinieri">M. bodinieri</taxonomicName> +. +</paragraph> +</caption> +<paragraph id="0F4B36C6FFFFFFD7FF66FD770511F9F6" pageId="4" pageNumber="256"> +<table id="7DF4C466FFFF002CFF66FD770517F994" box="[143,2054,697,1603]" gridcols="7" gridrows="14" pageId="4" pageNumber="256"> +<tr id="B1C43484FFFF002CFF66FD770517FD19" box="[143,2054,697,718]" gridrow="0" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FD770C87FD19" box="[143,406,697,718]" gridcol="0" gridrow="0" pageId="4" pageNumber="256"> +<emphasis id="3D80EAD4FFFFFFD7FF66FD770DABFD19" bold="true" box="[143,186,697,718]" pageId="4" pageNumber="256">Item</emphasis> +</th> +<th id="F2155DF8FFFF002CFE5DFD770FA5FD19" box="[436,692,697,718]" gridcol="1" gridrow="0" pageId="4" pageNumber="256"> +<taxonomicName id="C8F44D45FFFFFFD7FE5DFD770F52FD19" authorityName="Y. Tong & Z. D. Wang" authorityYear="2024" box="[436,579,697,718]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="nanlingensis"> +<emphasis id="3D80EAD4FFFFFFD7FE5DFD770F52FD19" bold="true" box="[436,579,697,718]" italics="true" pageId="4" pageNumber="256">M. nanlingensis</emphasis> +</taxonomicName> +</th> +<th id="F2155DF8FFFF002CFD3AFD770EC9FD19" box="[723,984,697,718]" gridcol="2" gridrow="0" pageId="4" pageNumber="256"> +<taxonomicName id="C8F44D45FFFFFFD7FD3AFD770E4FFD19" authorityName="Gagnepain" authorityYear="1908" box="[723,862,697,718]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="duclouxiana"> +<emphasis id="3D80EAD4FFFFFFD7FD3AFD770E4FFD19" bold="true" box="[723,862,697,718]" italics="true" pageId="4" pageNumber="256">M. duclouxiana</emphasis> +</taxonomicName> +</th> +<th id="F2155DF8FFFF002CFC1DFD7709FCFD19" box="[1012,1261,697,718]" gridcol="3" gridrow="0" pageId="4" pageNumber="256"> +<taxonomicName id="C8F44D45FFFFFFD7FC1DFD770992FD19" authorityName="Takeda" authorityYear="1917" box="[1012,1155,697,718]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="hancockiana"> +<emphasis id="3D80EAD4FFFFFFD7FC1DFD770992FD19" bold="true" box="[1012,1155,697,718]" italics="true" pageId="4" pageNumber="256">M. hancockiana</emphasis> +</taxonomicName> +</th> +<th id="F2155DF8FFFF002CFAEFFD7708F6FD19" box="[1286,1511,697,718]" gridcol="4" gridrow="0" pageId="4" pageNumber="256"> +<taxonomicName id="C8F44D45FFFFFFD7FAEFFD770883FD19" authorityName="Ying & Boufford" authorityYear="2001" box="[1286,1426,697,718]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="cardiophylla"> +<emphasis id="3D80EAD4FFFFFFD7FAEFFD770883FD19" bold="true" box="[1286,1426,697,718]" italics="true" pageId="4" pageNumber="256">M. cardiophylla</emphasis> +</taxonomicName> +</th> +<th id="F2155DF8FFFF002CF9E4FD770A16FD19" box="[1549,1799,697,718]" gridcol="5" gridrow="0" pageId="4" pageNumber="256"> +<taxonomicName id="C8F44D45FFFFFFD7F9E4FD770B4EFD19" authorityName="Carriere" authorityYear="1855" baseAuthorityName="Fortune" baseAuthorityYear="1850" box="[1549,1631,697,718]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="bealei"> +<emphasis id="3D80EAD4FFFFFFD7F9E4FD770B4EFD19" bold="true" box="[1549,1631,697,718]" italics="true" pageId="4" pageNumber="256">M. bealei</emphasis> +</taxonomicName> +</th> +<th id="F2155DF8FFFF002CF8F4FD770517FD19" box="[1821,2054,697,718]" gridcol="6" gridrow="0" pageId="4" pageNumber="256"> +<taxonomicName id="C8F44D45FFFFFFD7F8F4FD770A98FD19" authority="Fresh" authorityName="Fresh" box="[1821,1929,697,718]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="4" pageNumber="256" phylum="Tracheophyta" rank="species" species="bodinieri"> +<emphasis id="3D80EAD4FFFFFFD7F8F4FD770A98FD19" bold="true" box="[1821,1929,697,718]" italics="true" pageId="4" pageNumber="256">M. bodinieri</emphasis> +</taxonomicName> +</th> +</tr> +<tr id="B1C43484FFFF002CFF66FD380517FCFA" box="[143,2054,758,813]" gridrow="1" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FD380C87FCFA" box="[143,406,758,813]" gridcol="0" gridrow="1" pageId="4" pageNumber="256">Fresh leaf colour</th> +<td id="F2155DF8FFFF002CFE5DFD380FA5FCFA" box="[436,692,758,813]" gridcol="1" gridrow="1" pageId="4" pageNumber="256">abaxially yellowish green, adaxially dark green, lucid</td> +<td id="F2155DF8FFFF002CFD3AFD380EC9FCFA" box="[723,984,758,813]" gridcol="2" gridrow="1" pageId="4" pageNumber="256">abaxially yellowish green, adaxially dull to slightly shiny</td> +<td id="F2155DF8FFFF002CFC1DFD3809FCFCFA" box="[1012,1261,758,813]" gridcol="3" gridrow="1" pageId="4" pageNumber="256">abaxially pale, adaxially dull grayish green</td> +<td id="F2155DF8FFFF002CFAEFFD3808F6FCFA" box="[1286,1511,758,813]" gridcol="4" gridrow="1" pageId="4" pageNumber="256">abaxially yellowish green, adaxially dull</td> +<td id="F2155DF8FFFF002CF9E4FD380A16FCFA" box="[1549,1799,758,813]" gridcol="5" gridrow="1" pageId="4" pageNumber="256">abaxially glaucous, adaxially dull grayish green</td> +<td id="F2155DF8FFFF002CF8F4FD380517FCFA" box="[1821,2054,758,813]" gridcol="6" gridrow="1" pageId="4" pageNumber="256">abaxially yellowish green, adaxially dark green, lucid</td> +</tr> +<tr id="B1C43484FFFF002CFF66FC9B0517FC5C" box="[143,2054,853,907]" gridrow="2" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FC9B0C87FC5C" box="[143,406,853,907]" gridcol="0" gridrow="2" pageId="4" pageNumber="256">Number of leaflet margin teeth each side</th> +<td id="F2155DF8FFFF002CFE5DFC9B0FA5FC5C" box="[436,692,853,907]" gridcol="1" gridrow="2" pageId="4" pageNumber="256">7–9</td> +<td id="F2155DF8FFFF002CFD3AFC9B0EC9FC5C" box="[723,984,853,907]" gridcol="2" gridrow="2" pageId="4" pageNumber="256">2–12</td> +<td id="F2155DF8FFFF002CFC1DFC9B09FCFC5C" box="[1012,1261,853,907]" gridcol="3" gridrow="2" pageId="4" pageNumber="256">4–8</td> +<td id="F2155DF8FFFF002CFAEFFC9B08F6FC5C" box="[1286,1511,853,907]" gridcol="4" gridrow="2" pageId="4" pageNumber="256">3–8</td> +<td id="F2155DF8FFFF002CF9E4FC9B0A16FC5C" box="[1549,1799,853,907]" gridcol="5" gridrow="2" pageId="4" pageNumber="256">2–6</td> +<td id="F2155DF8FFFF002CF8F4FC9B0517FC5C" box="[1821,2054,853,907]" gridcol="6" gridrow="2" pageId="4" pageNumber="256">4–6</td> +</tr> +<tr id="B1C43484FFFF002CFF66FC6D0517FC0D" box="[143,2054,931,986]" gridrow="3" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FC6D0C87FC0D" box="[143,406,931,986]" gridcol="0" gridrow="3" pageId="4" pageNumber="256">Distance between lowest pair leaflet and base of petiole</th> +<td id="F2155DF8FFFF002CFE5DFC6D0FA5FC0D" box="[436,692,931,986]" gridcol="1" gridrow="3" pageId="4" pageNumber="256">0.5–1 cm</td> +<td id="F2155DF8FFFF002CFD3AFC6D0EC9FC0D" box="[723,984,931,986]" gridcol="2" gridrow="3" pageId="4" pageNumber="256">1 cm</td> +<td id="F2155DF8FFFF002CFC1DFC6D09FCFC0D" box="[1012,1261,931,986]" gridcol="3" gridrow="3" pageId="4" pageNumber="256">0.5–2 cm</td> +<td id="F2155DF8FFFF002CFAEFFC6D08F6FC0D" box="[1286,1511,931,986]" gridcol="4" gridrow="3" pageId="4" pageNumber="256">1–1.5 cm</td> +<td id="F2155DF8FFFF002CF9E4FC6D0A16FC0D" box="[1549,1799,931,986]" gridcol="5" gridrow="3" pageId="4" pageNumber="256">1–1.5 cm</td> +<td id="F2155DF8FFFF002CF8F4FC6D0517FC0D" box="[1821,2054,931,986]" gridcol="6" gridrow="3" pageId="4" pageNumber="256">2–2.5 cm</td> +</tr> +<tr id="B1C43484FFFF002CFF66FC3C0517FBD0" box="[143,2054,1010,1031]" gridrow="4" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FC3C0C87FBD0" box="[143,406,1010,1031]" gridcol="0" gridrow="4" pageId="4" pageNumber="256">Pairs and size of leaflets</th> +<td id="F2155DF8FFFF002CFE5DFC3C0FA5FBD0" box="[436,692,1010,1031]" gridcol="1" gridrow="4" pageId="4" pageNumber="256">5–9 pairs, 12–15 × 4.5–6 cm</td> +<td id="F2155DF8FFFF002CFD3AFC3C0EC9FBD0" box="[723,984,1010,1031]" gridcol="2" gridrow="4" pageId="4" pageNumber="256">4–9 pairs, 4.5–16 × 1.5–5 cm</td> +<td id="F2155DF8FFFF002CFC1DFC3C09FCFBD0" box="[1012,1261,1010,1031]" gridcol="3" gridrow="4" pageId="4" pageNumber="256">4–9 pairs, 3–11 × 1.8–2.8 cm</td> +<td id="F2155DF8FFFF002CFAEFFC3C08F6FBD0" box="[1286,1511,1010,1031]" gridcol="4" gridrow="4" pageId="4" pageNumber="256">8–10 pairs, 3–9 × 2–4 cm</td> +<td id="F2155DF8FFFF002CF9E4FC3C0A16FBD0" box="[1549,1799,1010,1031]" gridcol="5" gridrow="4" pageId="4" pageNumber="256">5–7 pairs, 10–15 × 10‒20 cm</td> +<td id="F2155DF8FFFF002CF8F4FC3C0517FBD0" box="[1821,2054,1010,1031]" gridcol="6" gridrow="4" pageId="4" pageNumber="256">7–9 pairs, 10–12 × 3–5 cm</td> +</tr> +<tr id="B1C43484FFFF002CFF66FBD10517FB82" box="[143,2054,1055,1109]" gridrow="5" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FBD10C87FB82" box="[143,406,1055,1109]" gridcol="0" gridrow="5" pageId="4" pageNumber="256">Outer sepals</th> +<td id="F2155DF8FFFF002CFE5DFBD10FA5FB82" box="[436,692,1055,1109]" gridcol="1" gridrow="5" pageId="4" pageNumber="256">elliptic, apex rounded or cordate</td> +<td id="F2155DF8FFFF002CFD3AFBD10EC9FB82" box="[723,984,1055,1109]" gridcol="2" gridrow="5" pageId="4" pageNumber="256">ovate to triangular-ovate, apex rounded</td> +<td id="F2155DF8FFFF002CFC1DFBD109FCFB82" box="[1012,1261,1055,1109]" gridcol="3" gridrow="5" pageId="4" pageNumber="256">ovate, apically obtuse, apex rounded</td> +<td id="F2155DF8FFFF002CFAEFFBD108F6FB82" box="[1286,1511,1055,1109]" gridcol="4" gridrow="5" pageId="4" pageNumber="256">triangular-ovate, apex rounded</td> +<td id="F2155DF8FFFF002CF9E4FBD10A16FB82" box="[1549,1799,1055,1109]" gridcol="5" gridrow="5" pageId="4" pageNumber="256">ovate, apex rounded</td> +<td id="F2155DF8FFFF002CF8F4FBD10517FB82" box="[1821,2054,1055,1109]" gridcol="6" gridrow="5" pageId="4" pageNumber="256">ovate, apex rounded</td> +</tr> +<tr id="B1C43484FFFF002CFF66FBA30517FB74" box="[143,2054,1133,1187]" gridrow="6" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FBA30C87FB74" box="[143,406,1133,1187]" gridcol="0" gridrow="6" pageId="4" pageNumber="256">Petals</th> +<td id="F2155DF8FFFF002CFE5DFBA30FA5FB74" box="[436,692,1133,1187]" gridcol="1" gridrow="6" pageId="4" pageNumber="256">base glandless, apex narrowly incised</td> +<td id="F2155DF8FFFF002CFD3AFBA30EC9FB74" box="[723,984,1133,1187]" gridcol="2" gridrow="6" pageId="4" pageNumber="256">base with distinct glands, apex emarginate</td> +<td id="F2155DF8FFFF002CFC1DFBA309FCFB74" box="[1012,1261,1133,1187]" gridcol="3" gridrow="6" pageId="4" pageNumber="256">base with distinct glands, apex emarginate</td> +<td id="F2155DF8FFFF002CFAEFFBA308F6FB74" box="[1286,1511,1133,1187]" gridcol="4" gridrow="6" pageId="4" pageNumber="256">base with distinct glands, apex incised</td> +<td id="F2155DF8FFFF002CF9E4FBA30A16FB74" box="[1549,1799,1133,1187]" gridcol="5" gridrow="6" pageId="4" pageNumber="256">base with distinct glands, apex slightly emarginate</td> +<td id="F2155DF8FFFF002CF8F4FBA30517FB74" box="[1821,2054,1133,1187]" gridcol="6" gridrow="6" pageId="4" pageNumber="256">base with indistinct glands, apex emarginate</td> +</tr> +<tr id="B1C43484FFFF002CFF66FB750517FB07" box="[143,2054,1211,1232]" gridrow="7" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FB750C87FB07" box="[143,406,1211,1232]" gridcol="0" gridrow="7" pageId="4" pageNumber="256">Stamen apex</th> +<td id="F2155DF8FFFF002CFE5DFB750FA5FB07" box="[436,692,1211,1232]" gridcol="1" gridrow="7" pageId="4" pageNumber="256">truncate</td> +<td id="F2155DF8FFFF002CFD3AFB750EC9FB07" box="[723,984,1211,1232]" gridcol="2" gridrow="7" pageId="4" pageNumber="256">truncate or rounded</td> +<td id="F2155DF8FFFF002CFC1DFB7509FCFB07" box="[1012,1261,1211,1232]" gridcol="3" gridrow="7" pageId="4" pageNumber="256">rounded</td> +<td id="F2155DF8FFFF002CFAEFFB7508F6FB07" box="[1286,1511,1211,1232]" gridcol="4" gridrow="7" pageId="4" pageNumber="256">truncate</td> +<td id="F2155DF8FFFF002CF9E4FB750A16FB07" box="[1549,1799,1211,1232]" gridcol="5" gridrow="7" pageId="4" pageNumber="256">rounded to truncate</td> +<td id="F2155DF8FFFF002CF8F4FB750517FB07" box="[1821,2054,1211,1232]" gridcol="6" gridrow="7" pageId="4" pageNumber="256">3-denticulate</td> +</tr> +<tr id="B1C43484FFFF002CFF66FB270517FB29" box="[143,2054,1257,1278]" gridrow="8" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FB270C87FB29" box="[143,406,1257,1278]" gridcol="0" gridrow="8" pageId="4" pageNumber="256">Ovules per ovary</th> +<td id="F2155DF8FFFF002CFE5DFB270FA5FB29" box="[436,692,1257,1278]" gridcol="1" gridrow="8" pageId="4" pageNumber="256">2</td> +<td id="F2155DF8FFFF002CFD3AFB270EC9FB29" box="[723,984,1257,1278]" gridcol="2" gridrow="8" pageId="4" pageNumber="256">4–7</td> +<td id="F2155DF8FFFF002CFC1DFB2709FCFB29" box="[1012,1261,1257,1278]" gridcol="3" gridrow="8" pageId="4" pageNumber="256">4 or 5</td> +<td id="F2155DF8FFFF002CFAEFFB2708F6FB29" box="[1286,1511,1257,1278]" gridcol="4" gridrow="8" pageId="4" pageNumber="256">2</td> +<td id="F2155DF8FFFF002CF9E4FB270A16FB29" box="[1549,1799,1257,1278]" gridcol="5" gridrow="8" pageId="4" pageNumber="256">3 or 4</td> +<td id="F2155DF8FFFF002CF8F4FB270517FB29" box="[1821,2054,1257,1278]" gridcol="6" gridrow="8" pageId="4" pageNumber="256">2</td> +</tr> +<tr id="B1C43484FFFF002CFF66FAD80517FAFC" box="[143,2054,1302,1323]" gridrow="9" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FAD80C87FAFC" box="[143,406,1302,1323]" gridcol="0" gridrow="9" pageId="4" pageNumber="256">Stigma</th> +<td id="F2155DF8FFFF002CFE5DFAD80FA5FAFC" box="[436,692,1302,1323]" gridcol="1" gridrow="9" pageId="4" pageNumber="256">broad and flat</td> +<td id="F2155DF8FFFF002CFD3AFAD80EC9FAFC" box="[723,984,1302,1323]" gridcol="2" gridrow="9" pageId="4" pageNumber="256">long pointed</td> +<td id="F2155DF8FFFF002CFC1DFAD809FCFAFC" box="[1012,1261,1302,1323]" gridcol="3" gridrow="9" pageId="4" pageNumber="256">long</td> +<td id="F2155DF8FFFF002CFAEFFAD808F6FAFC" box="[1286,1511,1302,1323]" gridcol="4" gridrow="9" pageId="4" pageNumber="256">short</td> +<td id="F2155DF8FFFF002CF9E4FAD80A16FAFC" box="[1549,1799,1302,1323]" gridcol="5" gridrow="9" pageId="4" pageNumber="256">short</td> +<td id="F2155DF8FFFF002CF8F4FAD80517FAFC" box="[1821,2054,1302,1323]" gridcol="6" gridrow="9" pageId="4" pageNumber="256">inconspicuous</td> +</tr> +<tr id="B1C43484FFFF002CFF66FA8D0517FA8F" box="[143,2054,1347,1368]" gridrow="10" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FA8D0C87FA8F" box="[143,406,1347,1368]" gridcol="0" gridrow="10" pageId="4" pageNumber="256">Berry</th> +<td id="F2155DF8FFFF002CFE5DFA8D0FA5FA8F" box="[436,692,1347,1368]" gridcol="1" gridrow="10" pageId="4" pageNumber="256">4–6 mm in diam.</td> +<td id="F2155DF8FFFF002CFD3AFA8D0EC9FA8F" box="[723,984,1347,1368]" gridcol="2" gridrow="10" pageId="4" pageNumber="256">5–8 mm in diam.</td> +<td id="F2155DF8FFFF002CFC1DFA8D09FCFA8F" box="[1012,1261,1347,1368]" gridcol="3" gridrow="10" pageId="4" pageNumber="256">Unknown</td> +<td id="F2155DF8FFFF002CFAEFFA8D08F6FA8F" box="[1286,1511,1347,1368]" gridcol="4" gridrow="10" pageId="4" pageNumber="256">4–7 mm in diam.</td> +<td id="F2155DF8FFFF002CF9E4FA8D0A16FA8F" box="[1549,1799,1347,1368]" gridcol="5" gridrow="10" pageId="4" pageNumber="256">10–12 mm in diam.</td> +<td id="F2155DF8FFFF002CF8F4FA8D0517FA8F" box="[1821,2054,1347,1368]" gridcol="6" gridrow="10" pageId="4" pageNumber="256">4–6 mm in diam.</td> +</tr> +<tr id="B1C43484FFFF002CFF66FABE0517FA52" box="[143,2054,1392,1413]" gridrow="11" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FABE0C87FA52" box="[143,406,1392,1413]" gridcol="0" gridrow="11" pageId="4" pageNumber="256">Flowering period</th> +<td id="F2155DF8FFFF002CFE5DFABE0FA5FA52" box="[436,692,1392,1413]" gridcol="1" gridrow="11" pageId="4" pageNumber="256">Aug–Oct</td> +<td id="F2155DF8FFFF002CFD3AFABE0EC9FA52" box="[723,984,1392,1413]" gridcol="2" gridrow="11" pageId="4" pageNumber="256">Nov–Apr</td> +<td id="F2155DF8FFFF002CFC1DFABE09FCFA52" box="[1012,1261,1392,1413]" gridcol="3" gridrow="11" pageId="4" pageNumber="256">Feb</td> +<td id="F2155DF8FFFF002CFAEFFABE08F6FA52" box="[1286,1511,1392,1413]" gridcol="4" gridrow="11" pageId="4" pageNumber="256">Feb–Apr</td> +<td id="F2155DF8FFFF002CF9E4FABE0A16FA52" box="[1549,1799,1392,1413]" gridcol="5" gridrow="11" pageId="4" pageNumber="256">Sep–Jun</td> +<td id="F2155DF8FFFF002CF8F4FABE0517FA52" box="[1821,2054,1392,1413]" gridcol="6" gridrow="11" pageId="4" pageNumber="256">Jun–Sep</td> +</tr> +<tr id="B1C43484FFFF002CFF66FA530517FA65" box="[143,2054,1437,1458]" gridrow="12" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FA530C87FA65" box="[143,406,1437,1458]" gridcol="0" gridrow="12" pageId="4" pageNumber="256">Fruiting period</th> +<td id="F2155DF8FFFF002CFE5DFA530FA5FA65" box="[436,692,1437,1458]" gridcol="1" gridrow="12" pageId="4" pageNumber="256">Oct–Nov</td> +<td id="F2155DF8FFFF002CFD3AFA530EC9FA65" box="[723,984,1437,1458]" gridcol="2" gridrow="12" pageId="4" pageNumber="256">Mar–Jun</td> +<td id="F2155DF8FFFF002CFC1DFA5309FCFA65" box="[1012,1261,1437,1458]" gridcol="3" gridrow="12" pageId="4" pageNumber="256">unknown</td> +<td id="F2155DF8FFFF002CFAEFFA5308F6FA65" box="[1286,1511,1437,1458]" gridcol="4" gridrow="12" pageId="4" pageNumber="256">May–Jun</td> +<td id="F2155DF8FFFF002CF9E4FA530A16FA65" box="[1549,1799,1437,1458]" gridcol="5" gridrow="12" pageId="4" pageNumber="256">Mar–May</td> +<td id="F2155DF8FFFF002CF8F4FA530517FA65" box="[1821,2054,1437,1458]" gridcol="6" gridrow="12" pageId="4" pageNumber="256">Aug–Dec</td> +</tr> +<tr id="B1C43484FFFF002CFF66FA040517F994" box="[143,2054,1482,1603]" gridrow="13" pageId="4" pageNumber="256"> +<th id="F2155DF8FFFF002CFF66FA040C87F994" box="[143,406,1482,1603]" gridcol="0" gridrow="13" pageId="4" pageNumber="256">Distribution</th> +<td id="F2155DF8FFFF002CFE5DFA040FA5F994" box="[436,692,1482,1603]" gridcol="1" gridrow="13" pageId="4" pageNumber="256">Guangdong, Guangxi</td> +<td id="F2155DF8FFFF002CFD3AFA040EC9F994" box="[723,984,1482,1603]" gridcol="2" gridrow="13" pageId="4" pageNumber="256">Guangxi, Sichuan, Yunnan, Guizhou</td> +<td id="F2155DF8FFFF002CFC1DFA0409FCF994" box="[1012,1261,1482,1603]" gridcol="3" gridrow="13" pageId="4" pageNumber="256">SE Yunnan</td> +<td id="F2155DF8FFFF002CFAEFFA0408F6F994" box="[1286,1511,1482,1603]" gridcol="4" gridrow="13" pageId="4" pageNumber="256">Guangxi, S Hunan, SE Sichuan, SE Yunnan</td> +<td id="F2155DF8FFFF002CF9E4FA040A16F994" box="[1549,1799,1482,1603]" gridcol="5" gridrow="13" pageId="4" pageNumber="256">Anhui, Fujian, Guangdong, Guangxi, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Zhejiang</td> +<td id="F2155DF8FFFF002CF8F4FA040517F994" box="[1821,2054,1482,1603]" gridcol="6" gridrow="13" pageId="4" pageNumber="256">Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Zhejiang</td> +</tr> +</table> +</paragraph> +<caption id="5B8B664EFFFEFFD6FF61F86F0FD3F80D" ID-DOI="http://doi.org/10.5281/zenodo.14520729" ID-Zenodo-Dep="14520729" httpUri="https://zenodo.org/record/14520729/files/figure.png" pageId="5" pageNumber="257" startId="5.[136,229,1953,1975]" targetBox="[169,1426,194,1918]" targetPageId="5" targetType="figure"> +<paragraph id="0F4B36C6FFFEFFD6FF61F86F0FD3F80D" blockId="5.[136,1452,1953,2011]" pageId="5" pageNumber="257"> +<emphasis id="3D80EAD4FFFEFFD6FF61F86F0DECF861" bold="true" box="[136,253,1953,1975]" pageId="5" pageNumber="257">FIGURE 2.</emphasis> +Illustration of +<taxonomicName id="C8F44D45FFFEFFD6FE62F86F0E04F861" authority="A. Habit. B. Outer" box="[395,789,1953,1975]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="5" pageNumber="257" phylum="Tracheophyta" rank="species" species="nanlingensis"> +<emphasis id="3D80EAD4FFFEFFD6FE62F86F0F71F861" box="[395,608,1953,1974]" italics="true" pageId="5" pageNumber="257">Mahonia nanlingensis</emphasis> +<emphasis id="3D80EAD4FFFEFFD6FD8CF86F0F6CF861" bold="true" box="[613,637,1953,1974]" pageId="5" pageNumber="257">A.</emphasis> +Habit. +<emphasis id="3D80EAD4FFFEFFD6FD2AF86F0FC8F861" bold="true" box="[707,729,1953,1974]" pageId="5" pageNumber="257">B.</emphasis> +Outer +</taxonomicName> +sepal. +<emphasis id="3D80EAD4FFFEFFD6FCBEF86F0E7EF860" bold="true" box="[855,879,1953,1975]" pageId="5" pageNumber="257">C.</emphasis> +Median sepal. +<emphasis id="3D80EAD4FFFEFFD6FBE9F86F0906F861" bold="true" box="[1024,1047,1953,1974]" pageId="5" pageNumber="257">D.</emphasis> +Inner sepal. +<emphasis id="3D80EAD4FFFEFFD6FB78F86F09B6F861" bold="true" box="[1169,1191,1953,1974]" pageId="5" pageNumber="257">E.</emphasis> +Petal. +<emphasis id="3D80EAD4FFFEFFD6FB01F86F09ECF861" bold="true" box="[1256,1277,1953,1974]" pageId="5" pageNumber="257">F.</emphasis> +Stamen. +<emphasis id="3D80EAD4FFFEFFD6FABFF86F087EF861" bold="true" box="[1366,1391,1953,1975]" pageId="5" pageNumber="257">G.</emphasis> +Pistil. +<emphasis id="3D80EAD4FFFEFFD6FF61F80B0DB0F80D" bold="true" box="[136,161,1989,2010]" pageId="5" pageNumber="257">H.</emphasis> +Berry and bract. +<emphasis id="3D80EAD4FFFEFFD6FEA1F80B0C7EF80D" bold="true" box="[328,367,1989,2010]" pageId="5" pageNumber="257">I–J.</emphasis> +Leaflets. Drawn by Qi-Lin Huang. +</paragraph> +</caption> +<caption id="5B8B664EFFFDFFD5FF61FBC809BAFBCB" ID-DOI="http://doi.org/10.5281/zenodo.14520733" ID-Zenodo-Dep="14520733" box="[136,1195,1030,1052]" httpUri="https://zenodo.org/record/14520733/files/figure.png" pageId="6" pageNumber="258" startId="6.[136,234,1030,1052]" targetBox="[164,1421,187,1002]" targetPageId="6" targetType="figure"> +<paragraph id="0F4B36C6FFFDFFD5FF61FBC809BAFBCB" blockId="6.[136,1195,1030,1052]" box="[136,1195,1030,1052]" pageId="6" pageNumber="258"> +<emphasis id="3D80EAD4FFFDFFD5FF61FBC80C12FBCC" bold="true" box="[136,259,1030,1052]" pageId="6" pageNumber="258">FIGURE 3.</emphasis> +<taxonomicName id="C8F44D45FFFDFFD5FEE1FBC80CCFFBCC" authorityName="Wang & Lan & Huang & Tong" authorityYear="2024" box="[264,478,1030,1051]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="6" pageNumber="258" phylum="Tracheophyta" rank="species" species="nanlingensis" status="sp. nov."> +<emphasis id="3D80EAD4FFFDFFD5FEE1FBC80CCFFBCC" box="[264,478,1030,1051]" italics="true" pageId="6" pageNumber="258">Mahonia nanlingensis</emphasis> +</taxonomicName> +<taxonomicNameLabel id="26B357AFFFFDFFD5FE0DFBC80F3FFBCC" box="[484,558,1030,1051]" pageId="6" pageNumber="258" rank="species"> +<emphasis id="3D80EAD4FFFDFFD5FE0DFBC80F38FBCC" box="[484,553,1030,1051]" italics="true" pageId="6" pageNumber="258">sp. nov</emphasis> +. +</taxonomicNameLabel> +<emphasis id="3D80EAD4FFFDFFD5FDDDFBC80F5DFBCC" bold="true" box="[564,588,1030,1051]" pageId="6" pageNumber="258">A.</emphasis> +leaves. +<emphasis id="3D80EAD4FFFDFFD5FD73FBC80FA1FBCC" bold="true" box="[666,688,1030,1051]" pageId="6" pageNumber="258">B.</emphasis> +seedling. +<emphasis id="3D80EAD4FFFDFFD5FCFBFBC80E38FBCB" bold="true" box="[786,809,1030,1052]" pageId="6" pageNumber="258">C.</emphasis> +flower. +<emphasis id="3D80EAD4FFFDFFD5FC91FBC80E98FBCC" bold="true" box="[888,905,1030,1051]" pageId="6" pageNumber="258">D</emphasis> +& +<emphasis id="3D80EAD4FFFDFFD5FC41FBC80EAFFBCC" bold="true" box="[936,958,1030,1051]" pageId="6" pageNumber="258">E.</emphasis> +habit. +<emphasis id="3D80EAD4FFFDFFD5FBE9FBC80902FBCC" bold="true" box="[1024,1043,1030,1051]" pageId="6" pageNumber="258">F.</emphasis> +infructescence. +</paragraph> +</caption> +<caption id="5B8B664EFFFDFFD5FF61F8250F0FF7F1" ID-DOI="http://doi.org/10.5281/zenodo.14520735" ID-Zenodo-Dep="14520735" httpUri="https://zenodo.org/record/14520735/files/figure.png" pageId="6" pageNumber="258" startId="6.[136,234,2027,2049]" targetBox="[164,1422,1077,2003]" targetPageId="6" targetType="figure"> +<paragraph id="0F4B36C6FFFDFFD5FF61F8250F0FF7F1" blockId="6.[136,1451,2027,2086]" pageId="6" pageNumber="258"> +<emphasis id="3D80EAD4FFFDFFD5FF61F8250C13F7D6" bold="true" box="[136,258,2027,2049]" pageId="6" pageNumber="258">FIGURE 4.</emphasis> +<emphasis id="3D80EAD4FFFDFFD5FEE1F8220F3DF7D6" box="[264,556,2028,2049]" italics="true" pageId="6" pageNumber="258"> +<taxonomicName id="C8F44D45FFFDFFD5FEE1F8220CCCF7D6" authorityName="Wang & Lan & Huang & Tong" authorityYear="2024" box="[264,477,2028,2049]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="6" pageNumber="258" phylum="Tracheophyta" rank="species" species="nanlingensis" status="sp. nov.">Mahonia nanlingensis</taxonomicName> +<taxonomicNameLabel id="26B357AFFFFDFFD5FE0AF8220F3DF7D6" box="[483,556,2028,2049]" pageId="6" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel> +</emphasis> +<emphasis id="3D80EAD4FFFDFFD5FDD9F8250F59F7D7" bold="true" box="[560,584,2027,2048]" pageId="6" pageNumber="258">A.</emphasis> +habit. +<emphasis id="3D80EAD4FFFDFFD5FD60F8220F8EF7D6" bold="true" box="[649,671,2028,2049]" pageId="6" pageNumber="258">B.</emphasis> +tender leaf. +<emphasis id="3D80EAD4FFFDFFD5FCFFF8250E5BF7D6" bold="true" box="[790,842,2027,2049]" pageId="6" pageNumber="258">C–D.</emphasis> +bud scales. +<emphasis id="3D80EAD4FFFDFFD5FC57F8220EC5F7D6" bold="true" box="[958,980,2028,2049]" pageId="6" pageNumber="258">E.</emphasis> +transection of stem. +<emphasis id="3D80EAD4FFFDFFD5FB72F82209DEF7D6" bold="true" box="[1179,1231,2027,2049]" pageId="6" pageNumber="258">F–G.</emphasis> +inflorescences. +<emphasis id="3D80EAD4FFFDFFD5FA80F8220893F7D6" bold="true" box="[1385,1410,2028,2049]" pageId="6" pageNumber="258">H.</emphasis> +bud scales of inflorescence. +<emphasis id="3D80EAD4FFFDFFD5FE84F7DE0C6CF7F2" bold="true" box="[365,381,2064,2085]" pageId="6" pageNumber="258">I.</emphasis> +infructescences. +</paragraph> +</caption> +<caption id="5B8B664EFFFCFFD4FF61F9F90E0AF9A6" ID-DOI="http://doi.org/10.5281/zenodo.14520737" ID-Zenodo-Dep="14520737" httpUri="https://zenodo.org/record/14520737/files/figure.png" pageId="7" pageNumber="259" startId="7.[136,234,1591,1613]" targetBox="[151,1436,190,1567]" targetPageId="7" targetType="figure"> +<paragraph id="0F4B36C6FFFCFFD4FF61F9F90E0AF9A6" blockId="7.[136,1452,1591,1649]" pageId="7" pageNumber="259"> +<emphasis id="3D80EAD4FFFCFFD4FF61F9F90C13F99B" bold="true" box="[136,258,1591,1613]" pageId="7" pageNumber="259">FIGURE 5.</emphasis> +<emphasis id="3D80EAD4FFFCFFD4FEEFF9F90F38F99A" box="[262,553,1591,1613]" italics="true" pageId="7" pageNumber="259"> +<taxonomicName id="C8F44D45FFFCFFD4FEEFF9F90CCAF99B" authorityName="Wang & Lan & Huang & Tong" authorityYear="2024" box="[262,475,1591,1612]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="7" pageNumber="259" phylum="Tracheophyta" rank="species" species="nanlingensis" status="sp. nov.">Mahonia nanlingensis</taxonomicName> +<taxonomicNameLabel id="26B357AFFFFCFFD4FE09F9F60F38F99A" box="[480,553,1592,1613]" pageId="7" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel> +</emphasis> +<emphasis id="3D80EAD4FFFCFFD4FDC4F9F90F55F99B" bold="true" box="[557,580,1591,1612]" pageId="7" pageNumber="259">A.</emphasis> +outer sepal. +<emphasis id="3D80EAD4FFFCFFD4FD52F9F90FC3F99B" bold="true" box="[699,722,1591,1612]" pageId="7" pageNumber="259">B.</emphasis> +median sepal. +<emphasis id="3D80EAD4FFFCFFD4FCB7F9F90E67F99A" bold="true" box="[862,886,1591,1613]" pageId="7" pageNumber="259">C.</emphasis> +inner sepal. +<emphasis id="3D80EAD4FFFCFFD4FC04F9F90914F99B" bold="true" box="[1005,1029,1591,1612]" pageId="7" pageNumber="259">D.</emphasis> +petal. +<emphasis id="3D80EAD4FFFCFFD4FBAAF9F90948F99B" bold="true" box="[1091,1113,1591,1612]" pageId="7" pageNumber="259">E.</emphasis> +petal with stamen. +<emphasis id="3D80EAD4FFFCFFD4FAF9F9F90832F99B" bold="true" box="[1296,1315,1591,1612]" pageId="7" pageNumber="259">F.</emphasis> +another shape of outer sepal. +<emphasis id="3D80EAD4FFFCFFD4FEFEF9950C54F9A7" bold="true" box="[279,325,1627,1649]" pageId="7" pageNumber="259">G–I.</emphasis> +stamens. +<emphasis id="3D80EAD4FFFCFFD4FE4DF9950CA7F9A7" bold="true" box="[420,438,1627,1648]" pageId="7" pageNumber="259">J.</emphasis> +pistil. +<emphasis id="3D80EAD4FFFCFFD4FE11F9950F00F9A7" bold="true" box="[504,529,1627,1648]" pageId="7" pageNumber="259">K.</emphasis> +pedicel and bract. +<emphasis id="3D80EAD4FFFCFFD4FD2EF9950FCCF9A7" bold="true" box="[711,733,1627,1648]" pageId="7" pageNumber="259">L.</emphasis> +berry. +</paragraph> +</caption> +<subSubSection id="47EE654DFFFCFFDBFF54F96E0E0BFEBB" lastPageId="8" lastPageNumber="260" pageId="7" pageNumber="259" type="description"> +<paragraph id="0F4B36C6FFFCFFDBFF54F96E0E0BFEBB" blockId="7.[136,1452,1696,2083]" lastBlockId="8.[136,1452,158,365]" lastPageId="8" lastPageNumber="260" pageId="7" pageNumber="259"> +<emphasis id="3D80EAD4FFFCFFD4FF54F96E0C52F96D" bold="true" box="[189,323,1696,1722]" italics="true" pageId="7" pageNumber="259">Description</emphasis> +:—Shrubs or small trees, +<quantity id="C80C9B23FFFCFFD4FD8CF96E0FD3F96D" box="[613,706,1696,1723]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="7" pageNumber="259" unit="m" value="1.75" valueMax="3.0" valueMin="0.5">0.5–3 m</quantity> +tall. Bark cork-like, deeply furrowed. Bud scales triangular, ca. 3 × +<quantity id="C80C9B23FFFCFFD4FF49F90A0DE1F909" box="[160,240,1732,1759]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="259" unit="cm" value="0.5">0.5 cm</quantity> +, green and then becoming reddish brown when flowering. Leaves abaxially yellowish green, not glaucous, adaxially dark green, lucid, ovate-oblong to lanceolate, 50–60 × +<quantity id="C80C9B23FFFCFFD4FC8CF9260EC9F8D5" box="[869,984,1768,1794]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="7" pageNumber="259" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 cm</quantity> +, with (6)7–8 pairs of leaflets, lowest pair +<quantity id="C80C9B23FFFCFFD4FF61F8C20DFEF8F1" box="[136,239,1804,1831]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="7" pageNumber="259" unit="cm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 cm</quantity> +above base of petiole; venations distinctly reticulate on both surfaces, 5-veined from base with 1 vein usually obscure abaxially, lateral veins 5 or 6 pairs, anastomosing near margin; rachis stout, +<quantity id="C80C9B23FFFCFFD4FBD2F8FE0988F89D" box="[1083,1177,1840,1866]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="7" pageNumber="259" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity> +in diam.; internodes (2–) +<quantity id="C80C9B23FFFCFFD4FF79F89A0DF5F8B9" box="[144,228,1876,1903]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="7" pageNumber="259" unit="cm" value="7.0" valueMax="9.0" valueMin="5.0">5–9 cm</quantity> +long, the uppermost internode shorter, lowermost internode longest, middle internodes nearly equal. Leaflets subsessile, lowest suborbicular, 3–3.5 × +<quantity id="C80C9B23FFFCFFD4FDA0F8B60FA5F845" box="[585,692,1912,1939]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="7" pageNumber="259" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 cm</quantity> +, those above becoming oblong to broadly lanceolate, 12–15 × +<quantity id="C80C9B23FFFCFFD4FA85F8B60DBCF861" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="7" pageNumber="259" unit="cm" value="5.25" valueMax="6.0" valueMin="4.5">4.5–6 cm</quantity> +, base oblique, truncate to cuneate, apex cuspidate, margins of 1st–2nd leaflets with 3 or 5 coarse teeth, 3 +<superScript id="F8819B8EFFFCFFD4FA9CF8550892F87E" attach="none" box="[1397,1411,1947,1961]" fontSize="6" pageId="7" pageNumber="259">rd</superScript> +–8 +<superScript id="F8819B8EFFFCFFD4FA76F85508BDF87E" attach="right" box="[1439,1452,1947,1961]" fontSize="6" pageId="7" pageNumber="259">th</superScript> +leaflets with 7 or 9 smaller teeth, upper leaflets with few (sometimes 5–7) fine teeth +<quantity id="C80C9B23FFFCFFD4FBBFF80E09A5F80D" box="[1110,1204,1984,2010]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="7" pageNumber="259" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity> +long; terminal leaflet slightly smaller than middle leaflets, 10–15 × +<quantity id="C80C9B23FFFCFFD4FD66F82A0FEAF829" box="[655,763,2020,2047]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="7" pageNumber="259" unit="cm" value="4.25" valueMax="5.0" valueMin="3.5">3.5–5 cm</quantity> +, petiolule +<quantity id="C80C9B23FFFCFFD4FC99F82A0EC8F829" box="[880,985,2020,2047]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="7" pageNumber="259" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 cm</quantity> +long. Tender leaves red. Inflorescence of 9 to 15 ascending racemes, 15–20(–25) cm long. Pedicel +<quantity id="C80C9B23FFFCFFD4FCE5F7C60E6FF7F5" box="[780,894,2056,2083]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="7" pageNumber="259" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity> +long; floral bracts narrowly ovate to ovate, 1–1.5 × +<quantity id="C80C9B23FFF3FFDBFF49FF500C04FF6E" box="[160,277,158,185]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="260" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity> +, apically pointed. Sepals yellow; outer sepals elliptic, purplish red, apex rounded or cordate, ca. 2 × +<quantity id="C80C9B23FFF3FFDBFA61FF500DA6FF0A" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="260" unit="mm" value="1.5">1.5 mm</quantity> +; median sepals elliptic, ca. 2.5 × +<quantity id="C80C9B23FFF3FFDBFDC0FF0C0F93FF0A" box="[553,642,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="260" unit="mm" value="1.5">1.5 mm</quantity> +; inner sepals narrowly elliptic, apical emarginate, ca. 3 × +<quantity id="C80C9B23FFF3FFDBFAECFF0C084FFF0A" box="[1285,1374,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="260" unit="mm" value="1.5">1.5 mm</quantity> +. Petals narrowly elliptic, subequal to median sepals, ca. 3 × +<quantity id="C80C9B23FFF3FFDBFD3AFF280E3AFED6" box="[723,811,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="260" unit="mm" value="1.5">1.5 mm</quantity> +, basally glandless, apex narrowly incised. Stamens +<quantity id="C80C9B23FFF3FFDBFA85FF280DA5FEF2" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="8" pageNumber="260" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity> +long; apex of anthers truncate; anther connective not prolonged. Ovary oblong-ovoid, ca. +<quantity id="C80C9B23FFF3FFDBFB47FEC40815FEF2" box="[1198,1284,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="260" unit="mm" value="1.5">1.5 mm</quantity> +long; ovules 2 per locule; stigma broad, flat. Berry initially green and then becoming purplish black when mature, white pruinose, oval or nearly spherical, +<quantity id="C80C9B23FFF3FFDBFE77FE9C0CEAFEBA" box="[414,507,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="260" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity> +in diam., style persistent. +</paragraph> +</subSubSection> +<caption id="5B8B664EFFF3FFDBFF61FBD1089EFBE2" ID-DOI="http://doi.org/10.5281/zenodo.14520739" ID-Zenodo-Dep="14520739" box="[136,1423,1055,1077]" httpUri="https://zenodo.org/record/14520739/files/figure.png" pageId="8" pageNumber="260" startId="8.[136,234,1055,1077]" targetBox="[151,1436,389,1030]" targetPageId="8" targetType="figure"> +<paragraph id="0F4B36C6FFF3FFDBFF61FBD1089EFBE2" blockId="8.[136,1423,1055,1077]" box="[136,1423,1055,1077]" pageId="8" pageNumber="260"> +<emphasis id="3D80EAD4FFF3FFDBFF61FBD10C12FBE3" bold="true" box="[136,259,1055,1077]" pageId="8" pageNumber="260">FIGURE 6.</emphasis> +Species similar to +<emphasis id="3D80EAD4FFF3FFDBFE50FBD10F87FBE3" box="[441,662,1055,1076]" italics="true" pageId="8" pageNumber="260"> +<taxonomicName id="C8F44D45FFF3FFDBFE50FBD10F83FBE3" authorityName="Wang & Lan & Huang & Tong" authorityYear="2024" box="[441,658,1055,1076]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="260" phylum="Tracheophyta" rank="species" species="nanlingensis">Mahonia nanlingensis</taxonomicName> +. +</emphasis> +<emphasis id="3D80EAD4FFF3FFDBFD72FBD10FDFFBE3" bold="true" box="[667,718,1055,1076]" pageId="8" pageNumber="260">A–B.</emphasis> +<emphasis id="3D80EAD4FFF3FFDBFD3DFBEE0E27FBE3" box="[724,822,1055,1077]" italics="true" pageId="8" pageNumber="260"> +<taxonomicName id="C8F44D45FFF3FFDBFD3DFBEE0E23FBE3" authorityName="Carriere" authorityYear="1855" baseAuthorityName="Fortune" baseAuthorityYear="1850" box="[724,818,1055,1077]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="260" phylum="Tracheophyta" rank="species" species="bealei">M. bealei</taxonomicName> +. +</emphasis> +<emphasis id="3D80EAD4FFF3FFDBFCD2FBD10E7EFBE2" bold="true" box="[827,879,1055,1077]" pageId="8" pageNumber="260">C–D.</emphasis> +<emphasis id="3D80EAD4FFF3FFDBFC9FFBEE0905FBE3" box="[886,1044,1055,1077]" italics="true" pageId="8" pageNumber="260"> +<taxonomicName id="C8F44D45FFF3FFDBFC9FFBEE0901FBE3" authorityName="Gagnepain" authorityYear="1908" box="[886,1040,1055,1077]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="260" phylum="Tracheophyta" rank="species" species="duclouxiana">M. duclouxiana</taxonomicName> +. +</emphasis> +<emphasis id="3D80EAD4FFF3FFDBFBF1FBD10956FBE3" bold="true" box="[1048,1095,1055,1076]" pageId="8" pageNumber="260">E–F.</emphasis> +<emphasis id="3D80EAD4FFF3FFDBFBA4FBEE09FFFBE3" box="[1101,1262,1055,1077]" italics="true" pageId="8" pageNumber="260"> +<taxonomicName id="C8F44D45FFF3FFDBFBA4FBEE09FBFBE3" authorityName="Ying & Boufford" authorityYear="2001" box="[1101,1258,1055,1077]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="260" phylum="Tracheophyta" rank="species" species="cardiophylla">M. cardiophylla</taxonomicName> +. +</emphasis> +<emphasis id="3D80EAD4FFF3FFDBFB1AFBD1081AFBE3" bold="true" box="[1267,1291,1055,1077]" pageId="8" pageNumber="260">G.</emphasis> +<taxonomicName id="C8F44D45FFF3FFDBFAF8FBEE0898FBE3" box="[1297,1417,1055,1077]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="260" phylum="Tracheophyta" rank="species" species="bodinieri"> +<emphasis id="3D80EAD4FFF3FFDBFAF8FBEE0898FBE3" box="[1297,1417,1055,1077]" italics="true" pageId="8" pageNumber="260">M. bodinieri</emphasis> +</taxonomicName> +. +</paragraph> +</caption> +<subSubSection id="47EE654DFFF3FFDBFF61FBAD0872FB12" pageId="8" pageNumber="260" type="etymology"> +<paragraph id="0F4B36C6FFF3FFDBFF61FBAD0C12FBA9" blockId="8.[136,1451,1123,1222]" box="[136,259,1123,1150]" pageId="8" pageNumber="260"> +<heading id="540381AAFFF3FFDBFF61FBAD0C12FBA9" bold="true" box="[136,259,1123,1150]" fontSize="11" level="1" pageId="8" pageNumber="260" reason="1"> +<emphasis id="3D80EAD4FFF3FFDBFF61FBAD0C12FBA9" bold="true" box="[136,259,1123,1150]" italics="true" pageId="8" pageNumber="260">Etymology</emphasis> +</heading> +</paragraph> +<paragraph id="0F4B36C6FFF3FFDBFF61FB490872FB12" blockId="8.[136,1451,1123,1222]" pageId="8" pageNumber="260"> +The specific epithet +<taxonomicName id="C8F44D45FFF3FFDBFE8EFB460F13FB76" authorityName="Wang & Lan & Huang & Tong" authorityYear="2024" box="[359,514,1159,1186]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="260" phylum="Tracheophyta" rank="species" species="nanlingensis"> +“ +<emphasis id="3D80EAD4FFF3FFDBFE9DFB490F13FB76" box="[372,514,1159,1185]" italics="true" pageId="8" pageNumber="260">nanlingensis</emphasis> +</taxonomicName> +” refers to the current known distribution area of the new species, namely the Nanling Mountains. The Chinese name is given as “ +<emphasis id="3D80EAD4FFF3FFDBFD87FB620E07FB11" bold="true" box="[622,790,1196,1222]" pageId="8" pageNumber="260">南*十大功劳</emphasis> +” (Chinese Pronunciation: Nán lǐng shí dà gōng láo). +</paragraph> +</subSubSection> +<subSubSection id="47EE654DFFF3FFDBFF61FB3D0FAAFAAE" pageId="8" pageNumber="260" type="biology_ecology"> +<paragraph id="0F4B36C6FFF3FFDBFF61FB3D0F3DFAD9" blockId="8.[136,1451,1267,1402]" box="[136,556,1267,1294]" pageId="8" pageNumber="260"> +<heading id="540381AAFFF3FFDBFF61FB3D0F3DFAD9" bold="true" box="[136,556,1267,1294]" fontSize="11" level="1" pageId="8" pageNumber="260" reason="1"> +<emphasis id="3D80EAD4FFF3FFDBFF61FB3D0F3DFAD9" bold="true" box="[136,556,1267,1294]" italics="true" pageId="8" pageNumber="260">Habitat, distribution, and phenology</emphasis> +</heading> +</paragraph> +<paragraph id="0F4B36C6FFF3FFDBFF61FAD90FAAFAAE" blockId="8.[136,1451,1267,1402]" pageId="8" pageNumber="260"> +<taxonomicName id="C8F44D45FFF3FFDBFF61FAD90C97FAE6" authorityName="Wang & Lan & Huang & Tong" authorityYear="2024" box="[136,390,1303,1329]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="260" phylum="Tracheophyta" rank="species" species="nanlingensis"> +<emphasis id="3D80EAD4FFF3FFDBFF61FAD90C97FAE6" box="[136,390,1303,1329]" italics="true" pageId="8" pageNumber="260">Mahonia nanlingensis</emphasis> +</taxonomicName> +is mainly distributed in +<collectingRegion id="CD30F824FFF3FFDBFD4EFAD90E3BFAE6" box="[679,810,1303,1329]" country="China" name="Guangdong" pageId="8" pageNumber="260">Guangdong</collectingRegion> +and +<collectingRegion id="CD30F824FFF3FFDBFC8EFAD90EDBFAE6" box="[871,970,1303,1329]" country="China" name="Guangxi" pageId="8" pageNumber="260">Guangxi</collectingRegion> +, +<collectingCountry id="77E37656FFF3FFDBFC30FAD9090DFAE6" box="[985,1052,1303,1329]" name="China" pageId="8" pageNumber="260">China</collectingCountry> +( +<figureCitation id="97CF2A43FFF3FFDBFBC7FAD90967FAE5" box="[1070,1142,1303,1330]" captionStart="FIGURE 7" captionStartId="9.[136,234,1638,1660]" captionTargetBox="[192,1395,503,1601]" captionTargetId="figure-203@9.[183,1404,494,1610]" captionTargetPageId="9" captionText="FIGURE 7. Distribution of Mahonia nanlingensis. Wild distribution (●); cultivated site (☆)." figureDoi="http://doi.org/10.5281/zenodo.14520741" httpUri="https://zenodo.org/record/14520741/files/figure.png" pageId="8" pageNumber="260">Fig. 7</figureCitation> +). +<location id="0A2B601DFFF3FFDBFB66FAD609B1FAE5" LSID="urn:lsid:plazi:treatment:875D87D0FFF8FFDAFF61FA5A089DFE0F:0A2B601DFFF3FFDBFB66FAD609B1FAE5" box="[1167,1184,1304,1330]" country="China" name="It" pageId="8" pageNumber="260" stateProvince="Guangdong">It</location> +grows in broad-leaved evergreen forests and mixed broad-leaved evergreen forests and forest margins at elevations of ca. +<quantity id="C80C9B23FFF3FFDBFB57FAF5083EFA81" box="[1214,1327,1339,1366]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="5.0" metricValueMin="0.5" pageId="8" pageNumber="260" unit="m" value="275.0" valueMax="500.0" valueMin="50.0"> +<elevation id="84D9D1F5FFF3FFDBFB57FAF5083EFA81" box="[1214,1327,1339,1366]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="5.0" metricValueMin="0.5" pageId="8" pageNumber="260" unit="m" value="275.0" valueMax="500.0" valueMin="50.0">50–500 m</elevation> +</quantity> +. Flowering August to October; fruiting October to November. +</paragraph> +</subSubSection> +<subSubSection id="47EE654DFFF3FFDBFF61FA690981F94D" pageId="8" pageNumber="260" type="conservation"> +<paragraph id="0F4B36C6FFF3FFDBFF61FA690C7DFA15" blockId="8.[136,1452,1447,2086]" box="[136,364,1447,1474]" pageId="8" pageNumber="260"> +<heading id="540381AAFFF3FFDBFF61FA690C7DFA15" bold="true" box="[136,364,1447,1474]" fontSize="11" level="1" pageId="8" pageNumber="260" reason="1"> +<emphasis id="3D80EAD4FFF3FFDBFF61FA690C7DFA15" bold="true" box="[136,364,1447,1474]" italics="true" pageId="8" pageNumber="260">Conservation status</emphasis> +</heading> +</paragraph> +<paragraph id="0F4B36C6FFF3FFDBFF61FA050981F94D" blockId="8.[136,1452,1447,2086]" pageId="8" pageNumber="260"> +<taxonomicName id="C8F44D45FFF3FFDBFF61FA050C93FA32" authorityName="Wang & Lan & Huang & Tong" authorityYear="2024" box="[136,386,1483,1509]" class="Magnoliopsida" family="Berberidaceae" genus="Mahonia" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="260" phylum="Tracheophyta" rank="species" species="nanlingensis"> +<emphasis id="3D80EAD4FFF3FFDBFF61FA050C93FA32" box="[136,386,1483,1509]" italics="true" pageId="8" pageNumber="260">Mahonia nanlingensis</emphasis> +</taxonomicName> +has been collected from over thirty localities and occurs across a vast area covering more than +<quantity id="C80C9B23FFF3FFDBFF61FA3E0C11F9DB" box="[136,256,1520,1548]" metricMagnitude="7" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="260" unit="km" value="20000.0">20,000 km</quantity> +<quantity id="C80C9B23FFF3FFDBFEE9FA3F0C34F9DE" box="[256,293,1519,1548]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="260" unit="in" value="2.0">² in</quantity> +the Nanling Mountains of northern +<collectingRegion id="CD30F824FFF3FFDBFD50FA210E2DF9DE" box="[697,828,1519,1545]" country="China" name="Guangdong" pageId="8" pageNumber="260">Guangdong</collectingRegion> +provinces and +<collectingRegion id="CD30F824FFF3FFDBFC0FFA21095BF9DE" box="[998,1098,1519,1545]" country="China" name="Guangxi" pageId="8" pageNumber="260">Guangxi</collectingRegion> +. There are 100–150 individuals at the +<typeStatus id="D04F8864FFF3FFDBFF3DF9DA0C15F9F9" box="[212,260,1556,1582]" pageId="8" pageNumber="260">type</typeStatus> +locality. However, due to medicinal and ornamental uses, as well as a reduction of habitat caused by continuous expansion of land use, the threat risk of this new species seems to be high. Therefore, the species should be regarded as “Vulnerable” (VU) according to A4, B1b and C1 criteria of IUCN Red List Category and Criteria ( +<bibRefCitation id="6B654B37FFF3FFDBFA8CF9920DD4F94D" author="IUCN" pageId="8" pageNumber="260" refId="ref6724" refString="IUCN (2022) Guidelines for using the IUCN Red List categories and criteria (Version 15.1). Prepared by the IUCN Standards and Petitions Committee." type="book" year="2022">IUCN 2022</bibRefCitation> +). It is necessary to protect its habitat and collect and preserve its germplasm resources. +</paragraph> +</subSubSection> +<subSubSection id="47EE654DFFF3FFDAFF54F96D089DFE0F" lastPageId="9" lastPageNumber="261" pageId="8" pageNumber="260" type="materials_examined"> +<paragraph id="0F4B36C6FFF3FFDAFF54F96D089DFE0F" blockId="8.[136,1452,1447,2086]" lastBlockId="9.[136,1452,158,473]" lastPageId="9" lastPageNumber="261" pageId="8" pageNumber="260"> +<materialsCitation id="BF9C3C9BFFF3FFDBFF54F96D09F4F935" collectingDate="2019-09-01" collectionCode="CMMI" collectorName="Y. 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