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<document id="1E2BB0A59A2C2C76C2846C52C3DAD480" ID-DOI="10.4081/nhs.2019.398" ID-ISSN="2385-0922" ID-Zenodo-Dep="12753895" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1721184903258" checkinUser="felipe" docAuthor="Pieroni, Vittorio" docDate="2019" docId="0C4687BCFFE3FFECFFBD1886FD44FDF7" docLanguage="en" docName="NatHistSci.6.1.37-40.pdf" docOrigin="Natural History Sciences 6 (1)" docSource="http://dx.doi.org/10.4081/nhs.2019.398" docStyle="DocumentStyle:926963A4B76E17DFFD817EF231EB8DA9.3:NatHistSci.2014-.journal_article" docStyleId="926963A4B76E17DFFD817EF231EB8DA9" docStyleName="NatHistSci.2014-.journal_article" docStyleVersion="3" docTitle="Wortheniella pygmaea Stoppani 1860" docType="treatment" docVersion="3" lastPageNumber="39" masterDocId="F07FFFC4FFE2FFEFFFCC1B10FFB1FFDE" masterDocTitle="The Norian Worthenia-like gastropods (Main Dolomite, Upper Triassic): reclassification of the specimens housed at the “ Antonio Stoppani ” Natural Museum, Italy" masterLastPageNumber="40" masterPageNumber="37" pageNumber="38" updateTime="1721893293113" updateUser="tatiana" zenodo-license-document="CC-BY-NC-4.0">
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<mods:titleInfo id="1AC210E9B1ABED17E01C0871632C8C56">
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<mods:title id="8C737D15F72C230B983103FA593BF3E6">The Norian Worthenia-like gastropods (Main Dolomite, Upper Triassic): reclassification of the specimens housed at the “ Antonio Stoppani ” Natural Museum, Italy</mods:title>
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<mods:namePart id="0F35A83D29D119678DA61796F1D49062">Pieroni, Vittorio</mods:namePart>
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<mods:typeOfResource id="59FD8B522E3CB2CC5ADC2D0FE5BC0B67">text</mods:typeOfResource>
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<mods:title id="8A079AFB9D94A65092AC5E75475712DB">Natural History Sciences</mods:title>
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<mods:part id="74947283070EFC1EABBF33CF01335098">
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<mods:date id="2780F77559E52EF74C61240AE5FD4B62">2019</mods:date>
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<mods:number id="EAF4E3DACEF264516632B74DE7BF3A57">2019-03-18</mods:number>
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<mods:url id="7D14F9DFB600BBC3940A9E51FF5AF2A0">http://dx.doi.org/10.4081/nhs.2019.398</mods:url>
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<mods:classification id="CCF31D766446FCE6E90A212D7B03050C">journal article</mods:classification>
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<mods:identifier id="D2479423F4A6208E59CF033D4E992BCC" type="DOI">10.4081/nhs.2019.398</mods:identifier>
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<mods:identifier id="FF6FF750B591103B56CB98459E198E47" type="ISSN">2385-0922</mods:identifier>
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<treatment id="0C4687BCFFE3FFECFFBD1886FD44FDF7" LSID="urn:lsid:plazi:treatment:0C4687BCFFE3FFECFFBD1886FD44FDF7" httpUri="http://treatment.plazi.org/id/0C4687BCFFE3FFECFFBD1886FD44FDF7" lastPageId="3" lastPageNumber="39" pageId="1" pageNumber="38">
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<subSubSection id="CCF56521FFE3FFEEFFBD1886FD44FC6F" box="[113,757,918,946]" pageId="1" pageNumber="38" type="nomenclature">
|
||||
<paragraph id="845036AAFFE3FFEEFFBD1886FD44FC6F" blockId="1.[113,757,918,1970]" box="[113,757,918,946]" pageId="1" pageNumber="38">
|
||||
<heading id="DF1881C6FFE3FFEEFFBD1886FD44FC6F" box="[113,757,918,946]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="38" reason="6">
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||||
<taxonomicName id="43EF4D29FFE3FFEEFFBD1886FDD1FC6C" authority="Stoppani, 1860" authorityName="Stoppani" authorityYear="1860" box="[113,608,918,946]" class="Gastropoda" family="Wortheniellidae" genus="Wortheniella" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="pygmaea">
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||||
<emphasis id="B69BEAB8FFE3FFEEFFBD1886FEC3FC6F" bold="true" box="[113,370,918,945]" italics="true" pageId="1" pageNumber="38">Wortheniella pygmaea</emphasis>
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(Stoppani, 1860-65)
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</taxonomicName>
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Comb. nov.
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</heading>
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||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CCF56521FFE3FFEEFF5B18A5FE15FBB8" pageId="1" pageNumber="38" type="description">
|
||||
<paragraph id="845036AAFFE3FFEEFF5B18A5FEA2FC11" blockId="1.[113,757,918,1970]" box="[151,275,949,975]" pageId="1" pageNumber="38">
|
||||
(
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||||
<figureCitation id="1CD42A2FFFE3FFEEFF5318A5FF6BFC11" box="[159,218,949,975]" captionStart="Fig" captionStartId="2.[189,222,906,928]" captionTargetBox="[189,1399,136,892]" captionTargetId="figure-688@2.[189,1399,136,892]" captionTargetPageId="2" captionText="Fig. 1 - A) Wortheniella pygmaea (Stoppani) AS 41/43, lateral view, showing the pattern of the growth lines, which curve backwards above and below the nodose adapical carina (corresponding to the selenizone).Songavazzo (BG), Norian,Main Dolomite. B) Wortheniella pygmaea (Stoppani) AS 41/43, latero-basal view, showing the very conspicuous threads on the base. C) Wortheniella pygmaea (Stoppani) AS 41/43, apical view of the early whorls of the spire. D) Wortheniella pygmaea (Stoppani) AS 41/43, lateral view of the early whorls of the spire. E) Wortheniella pygmaea (Stoppani) AS 41/43a, lateral view. F) Wortheniella pygmaea (Stoppani) AS 41/43a, latero-basal view, showing the region of the umbilicus, which is largely occupied by a columellar lip." figureDoi="http://doi.org/10.5281/zenodo.12753897" httpUri="https://zenodo.org/record/12753897/files/figure.png" pageId="1" pageNumber="38">Fig. 1</figureCitation>
|
||||
A-F)
|
||||
</paragraph>
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||||
<paragraph id="845036AAFFE3FFEEFFBD18C3FF4EFBD5" blockId="1.[113,757,918,1970]" pageId="1" pageNumber="38">
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1860-65
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<taxonomicName id="43EF4D29FFE3FFEEFF1918C3FD94FC33" authority="Stoppani" authorityName="Stoppani" box="[213,549,979,1005]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="pygmaea">
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||||
<emphasis id="B69BEAB8FFE3FFEEFF1918C3FE08FC33" box="[213,441,979,1005]" italics="true" pageId="1" pageNumber="38">Delphinula pygmaea</emphasis>
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Stoppani
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</taxonomicName>
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, p. 256-57, pl. 59, fig. 16-17
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</paragraph>
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||||
<paragraph id="845036AAFFE3FFEEFFBD1F1FFD45FBF4" blockId="1.[113,757,918,1970]" box="[113,756,1039,1066]" pageId="1" pageNumber="38">
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||||
1860-65
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||||
<taxonomicName id="43EF4D29FFE3FFEEFF031F1FFDA8FBF7" authority="Stoppani" authorityName="Stoppani" box="[207,537,1039,1065]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="regazzoni">
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||||
<emphasis id="B69BEAB8FFE3FFEEFF031F1FFE02FBF7" box="[207,435,1039,1065]" italics="true" pageId="1" pageNumber="38">Delphinula regazzoni</emphasis>
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Stoppani
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</taxonomicName>
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, p. 257, pl. 59, fig. 18
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</paragraph>
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<paragraph id="845036AAFFE3FFEEFFBD1F3EFE15FBB8" blockId="1.[113,757,918,1970]" pageId="1" pageNumber="38">
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1903
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||||
<taxonomicName id="43EF4D29FFE3FFEEFF781F3DFDBFFB96" authority="(Stoppani)" baseAuthorityName="Stoppani" box="[180,526,1069,1096]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFF781F3DFE20FB99" box="[180,401,1069,1095]" italics="true" pageId="1" pageNumber="38">Worthenia pygmaea</emphasis>
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||||
(Stoppani)
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</taxonomicName>
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– Tommasi, p. 114 [20], pl. 18 [3], fig. 14a-e
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</paragraph>
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</subSubSection>
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||||
<subSubSection id="CCF56521FFE3FFEEFF5B1F7AFDF1F996" pageId="1" pageNumber="38" type="materials_examined">
|
||||
<paragraph id="845036AAFFE3FFEEFF5B1F7AFDC2FB01" blockId="1.[113,757,918,1970]" pageId="1" pageNumber="38">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFF5B1F7AFEB8FB5A" bold="true" box="[151,265,1130,1156]" pageId="1" pageNumber="38">
|
||||
<typeStatus id="5B548808FFE3FFEEFF5B1F7AFEB5FB5A" box="[151,260,1130,1156]" pageId="1" pageNumber="38" type="holotype">Holotype</typeStatus>
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||||
:
|
||||
</emphasis>
|
||||
The
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||||
<typeStatus id="5B548808FFE3FFEEFE841F7AFE01FB5A" box="[328,432,1130,1156]" pageId="1" pageNumber="38" type="holotype">holotypes</typeStatus>
|
||||
of
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||||
<taxonomicName id="43EF4D29FFE3FFEEFE111F7AFD73FB5A" box="[477,706,1130,1156]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFE111F7AFD73FB5A" box="[477,706,1130,1156]" italics="true" pageId="1" pageNumber="38">Delphinula pygmaea</emphasis>
|
||||
</taxonomicName>
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||||
and
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<taxonomicName id="43EF4D29FFE3FFEEFFBD1F98FEEAFB7C" box="[113,347,1160,1186]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="regazzoni">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFFBD1F98FEEAFB7C" box="[113,347,1160,1186]" italics="true" pageId="1" pageNumber="38">Delphinula regazzoni</emphasis>
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</taxonomicName>
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, like the rest of the specimens from Main Dolomite published by Stoppani in Series III of “Paléontologie Lombarde”, are lost (see above).
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</paragraph>
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<paragraph id="845036AAFFE3FFEEFF5B1FF2FD20FB22" blockId="1.[113,757,918,1970]" box="[151,657,1250,1276]" pageId="1" pageNumber="38">
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<emphasis id="B69BEAB8FFE3FFEEFF5B1FF2FE8FFB22" bold="true" box="[151,318,1250,1276]" pageId="1" pageNumber="38">Locus typicus:</emphasis>
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||||
Caino (near Brescia, BS),
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<collectingCountry id="FCF8763AFFE3FFEEFD971FF2FD3FFB22" box="[603,654,1250,1276]" name="Italy" pageId="1" pageNumber="38">Italy</collectingCountry>
|
||||
.
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</paragraph>
|
||||
<paragraph id="845036AAFFE3FFEEFF5B1E10FDC8FAC5" blockId="1.[113,757,918,1970]" box="[151,633,1280,1307]" pageId="1" pageNumber="38">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFF5B1E10FED4FAC4" bold="true" box="[151,357,1280,1306]" pageId="1" pageNumber="38">Stratum typicum:</emphasis>
|
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Main Dolomite (Norian).
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</paragraph>
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<paragraph id="845036AAFFE3FFEEFF5B1E0EFDF1F996" blockId="1.[113,757,918,1970]" pageId="1" pageNumber="38">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFF5B1E0EFEB5FAE6" bold="true" box="[151,260,1310,1336]" pageId="1" pageNumber="38">Material:</emphasis>
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||||
<materialsCitation id="34873CF7FFE3FFEEFEC61E0FFD8DF996" collectorName="Seminario Arcivescovile Pio XI" country="American Samoa" location="Diocese of Milan" pageId="1" pageNumber="38" specimenCount="2">
|
||||
<specimenCount id="92E9FD23FFE3FFEEFEC61E0FFE01FAE6" box="[266,432,1310,1337]" count="2" pageId="1" pageNumber="38" type="generic">Two specimens</specimenCount>
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[catalogue numbers AS 41/43 (
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<figureCitation id="1CD42A2FFFE3FFEEFFB51E2DFF04FA89" box="[121,181,1341,1367]" captionStart="Fig" captionStartId="2.[189,222,906,928]" captionTargetBox="[189,1399,136,892]" captionTargetId="figure-688@2.[189,1399,136,892]" captionTargetPageId="2" captionText="Fig. 1 - A) Wortheniella pygmaea (Stoppani) AS 41/43, lateral view, showing the pattern of the growth lines, which curve backwards above and below the nodose adapical carina (corresponding to the selenizone).Songavazzo (BG), Norian,Main Dolomite. B) Wortheniella pygmaea (Stoppani) AS 41/43, latero-basal view, showing the very conspicuous threads on the base. C) Wortheniella pygmaea (Stoppani) AS 41/43, apical view of the early whorls of the spire. D) Wortheniella pygmaea (Stoppani) AS 41/43, lateral view of the early whorls of the spire. E) Wortheniella pygmaea (Stoppani) AS 41/43a, lateral view. F) Wortheniella pygmaea (Stoppani) AS 41/43a, latero-basal view, showing the region of the umbilicus, which is largely occupied by a columellar lip." figureDoi="http://doi.org/10.5281/zenodo.12753897" httpUri="https://zenodo.org/record/12753897/files/figure.png" pageId="1" pageNumber="38">Fig. 1</figureCitation>
|
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A-D) and AS 41/43a (
|
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<figureCitation id="1CD42A2FFFE3FFEEFE651E2DFDA3FA89" box="[425,530,1340,1367]" captionStart="Fig" captionStartId="2.[189,222,906,928]" captionTargetBox="[189,1399,136,892]" captionTargetId="figure-688@2.[189,1399,136,892]" captionTargetPageId="2" captionText="Fig. 1 - A) Wortheniella pygmaea (Stoppani) AS 41/43, lateral view, showing the pattern of the growth lines, which curve backwards above and below the nodose adapical carina (corresponding to the selenizone).Songavazzo (BG), Norian,Main Dolomite. B) Wortheniella pygmaea (Stoppani) AS 41/43, latero-basal view, showing the very conspicuous threads on the base. C) Wortheniella pygmaea (Stoppani) AS 41/43, apical view of the early whorls of the spire. D) Wortheniella pygmaea (Stoppani) AS 41/43, lateral view of the early whorls of the spire. E) Wortheniella pygmaea (Stoppani) AS 41/43a, lateral view. F) Wortheniella pygmaea (Stoppani) AS 41/43a, latero-basal view, showing the region of the umbilicus, which is largely occupied by a columellar lip." figureDoi="http://doi.org/10.5281/zenodo.12753897" httpUri="https://zenodo.org/record/12753897/files/figure.png" pageId="1" pageNumber="38">Fig. 1E,F</figureCitation>
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)], almost totally preserved as pseudoshells recrystallised in dolomite, from Songavazzo (BG), Italy. The specimens are part of the historic Stoppani Collection housed at the Museo Scientifico Naturalistico “Antonio Stoppani” in the
|
||||
<collectorName id="291A537CFFE3FFEEFE201EA5FF0BFA30" pageId="1" pageNumber="38">Seminario Arcivescovile Pio XI</collectorName>
|
||||
,
|
||||
<location id="81306071FFE3FFEEFF091EC3FE30FA33" LSID="urn:lsid:plazi:treatment:0C4687BCFFE3FFECFFBD1886FD44FDF7:81306071FFE3FFEEFF091EC3FE30FA33" box="[197,385,1491,1517]" country="American Samoa" name="Diocese of Milan" pageId="1" pageNumber="38">Diocese of Milan</location>
|
||||
, Venegono Inferiore,
|
||||
<collectingCountry id="FCF8763AFFE3FFEEFDAA1EC3FD28FA33" box="[614,665,1491,1517]" name="Italy" pageId="1" pageNumber="38">Italy</collectingCountry>
|
||||
.Although not the typical site for the species, Songavazzo is one of the localities where Stoppani collected the Norian gastropods he described in “Paléontologie Lombarde”
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CCF56521FFE3FFEEFF5B1D5CFBEAFC11" pageId="1" pageNumber="38" type="description">
|
||||
<paragraph id="845036AAFFE3FFEEFF5B1D5CFD42F95B" blockId="1.[113,757,918,1970]" pageId="1" pageNumber="38">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFF5B1D5CFE97F9B8" bold="true" box="[151,294,1612,1638]" pageId="1" pageNumber="38">Dimensions:</emphasis>
|
||||
<collectingCountry id="FCF8763AFFE3FFEEFEFF1D5CFEE7F9B8" box="[307,342,1612,1638]" name="American Samoa" pageId="1" pageNumber="38">AS</collectingCountry>
|
||||
41/43: H=
|
||||
<quantity id="43179B4FFFE3FFEEFE161D5CFDF9F9B8" box="[474,584,1612,1638]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="38" unit="mm" value="6.8">6.80 mm</quantity>
|
||||
; W=
|
||||
<quantity id="43179B4FFFE3FFEEFD4F1D5CFD41F9B8" box="[643,752,1612,1639]" metricMagnitude="-3" metricUnit="m" metricValue="7.45" pageId="1" pageNumber="38" unit="mm" value="7.45">7.45 mm</quantity>
|
||||
; PA =62°.
|
||||
<collectingCountry id="FCF8763AFFE3FFEEFF1A1D7AFF48F95A" box="[214,249,1642,1668]" name="American Samoa" pageId="1" pageNumber="38">AS</collectingCountry>
|
||||
41/43a: H=
|
||||
<quantity id="43179B4FFFE3FFEEFEB51D7AFE5BF95A" box="[377,490,1642,1668]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="1" pageNumber="38" unit="mm" value="10.6">10.60 mm</quantity>
|
||||
; W=
|
||||
<quantity id="43179B4FFFE3FFEEFDD01D7AFD3DF95A" box="[540,652,1642,1668]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="1" pageNumber="38" unit="mm" value="11.8">11.80 mm</quantity>
|
||||
; PA=75°.
|
||||
</paragraph>
|
||||
<paragraph id="845036AAFFE3FFEEFF5B1D98FBEAFC11" blockId="1.[113,757,918,1970]" lastBlockId="1.[832,1475,135,1969]" pageId="1" pageNumber="38">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFF5B1D98FE94F97C" bold="true" box="[151,293,1672,1698]" pageId="1" pageNumber="38">Description:</emphasis>
|
||||
Specimen
|
||||
<collectingCountry id="FCF8763AFFE3FFEEFE6C1D98FE72F97C" box="[416,451,1672,1698]" name="American Samoa" pageId="1" pageNumber="38">AS</collectingCountry>
|
||||
41/43 has a gradate trochiform shell consisting of just over 5 rapidly grown whorls. Adult whorls have an adapical and an abapical spiral carina. The former is at the angle between the ramp and the flank; the latter is at the angle between the flank and the base. The flank is flattened and almost perpendicular to the coil axis. The two early teleoconch whorls are rounded and lack evident ornamentation. The next three whorls have a distinct adapical carina. The abapical carina is exposed only on the last whorl, whereas it corresponds to the suture on the preceding two whorls. The first preserved whorl is almost planispiral. The apical part is strongly recrystallized and thus the characters of the protoconch are not identifiable. A selenizone is present in correspondence with the robust adapical spiral carina separating the sutural ramp, which is inclined and slightly swollen, from the flank of the whorl at an obtuse angle. This carina is sculptured with well-spaced, longitudinally elongate nodules. The abapical carina, thinner than the former, does not have evident nodules. The whole surface of the shell is sculptured with more or less fine threads crossed by growth lines. Three threads are visible on the sutural ramp, of which the two adapical ones are significantly more pronounced and closer together, whereas the abapical one is thinner and located further away from the two adapical threads and from the adapical carina. Three threads are present also on the flank. They are more or less equidistant between the two carinae and well developed. The base, which for the most part is embedded in matrix, is weakly rounded, convex and sculptured with more than five evident threads. The growth lines are prosocline and prosocyrt on the ramp, prosocyrt on the flank below the selenizone, prosocline on the base. The aperture, distinctly wider than high, is polygonal in shape being angulated at the two spiral carinae.The specimen
|
||||
<collectingCountry id="FCF8763AFFE3FFEEFCBF182CFC27FC88" box="[883,918,828,854]" name="American Samoa" pageId="1" pageNumber="38">AS</collectingCountry>
|
||||
41/43a – which has been completely extracted from the matrix, but has an eroded pseudoshell – shows a quite well developed columellar lip that covers almost completely the rather small umbilicus. Evident threads are visible on the base of this specimen, too.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CCF56521FFE3FFECFCA918C3FD44FDF7" lastPageId="3" lastPageNumber="40" pageId="1" pageNumber="38" type="discussion">
|
||||
<paragraph id="845036AAFFE3FFEEFCA918C3FA72FA4D" blockId="1.[832,1475,135,1969]" pageId="1" pageNumber="38">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFCA918C3FC57FC33" bold="true" box="[869,998,979,1005]" pageId="1" pageNumber="38">Discussion:</emphasis>
|
||||
This species is described by Stoppani (1860- 65) from Norian Main Dolomite of Caino (BS). In the fauna from some Prealpine Lombard localities in the provinces of Bergamo and Brescia, Stoppani described
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFAC71F3DFC53FBB8" authority="Stoppani, 1860" authorityName="Stoppani" authorityYear="1860" class="Gastropoda" family="Turbinidae" genus="Turbo" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="songavatii">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFAC71F3DFA73FB99" box="[1291,1474,1069,1095]" italics="true" pageId="1" pageNumber="38">Turbo songavatii</emphasis>
|
||||
Stoppani, 1860
|
||||
</taxonomicName>
|
||||
-65 (=
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFBEF1F5BFAC5FBBB" authority="Benecke, 1866" authorityName="Benecke" authorityYear="1866" box="[1059,1396,1099,1125]" class="Gastropoda" family="Turbinidae" genus="Turbo" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="solitarius">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFBEF1F5BFB7FFBBB" box="[1059,1230,1099,1125]" italics="true" pageId="1" pageNumber="38">Turbo solitarius</emphasis>
|
||||
Benecke, 1866
|
||||
</taxonomicName>
|
||||
, which according to De Stefani 1880 was a junior synonym of
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFA4E1F79FC1FFB7F" authorityName="Stoppani" authorityYear="1860" class="Gastropoda" family="Turbinidae" genus="Turbo" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="songavatii">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFA4E1F79FC1FFB7F" italics="true" pageId="1" pageNumber="38">Turbo songavatii</emphasis>
|
||||
</taxonomicName>
|
||||
,whichHaas1953consideredasynonymof
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFAA11F98FC78FB1E" class="Gastropoda" family="Trochidae" genus="Trochus" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="contabulatus">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFAA11F98FC78FB1E" italics="true" pageId="1" pageNumber="38">Trochus contabulatus</emphasis>
|
||||
</taxonomicName>
|
||||
Costa, 1864, p. 232, pl. 5, fig. 4),
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFAE61FB6FC55FB01" authority="Stoppani, 1860" authorityName="Stoppani" authorityYear="1860" class="Gastropoda" family="Turbinidae" genus="Turbo" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="pusillus">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFAE61FB6FA73FB1E" box="[1322,1474,1190,1216]" italics="true" pageId="1" pageNumber="38">Turbo pusillus</emphasis>
|
||||
Stoppani, 1860
|
||||
</taxonomicName>
|
||||
-65,
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFBDE1FD4FA2AFB01" authority="Stoppani, 1860" authorityName="Stoppani" authorityYear="1860" box="[1042,1435,1220,1247]" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="inzini">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFBDE1FD4FB41FB00" box="[1042,1264,1220,1246]" italics="true" pageId="1" pageNumber="38">Pleurotomaria inzini</emphasis>
|
||||
Stoppani, 1860
|
||||
</taxonomicName>
|
||||
-65,
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFC8C1FF2FB09FB23" authority="Stoppani, 1860" authorityName="Stoppani" authorityYear="1860" box="[832,1208,1250,1277]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="escheri">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFC8C1FF2FBBDFB22" box="[832,1036,1250,1276]" italics="true" pageId="1" pageNumber="38">Delphinula escheri</emphasis>
|
||||
Stoppani, 1860
|
||||
</taxonomicName>
|
||||
-65,
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFB2B1FF2FC53FAC5" authority="Stoppani, 1860" authorityName="Stoppani" authorityYear="1860" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="diadema">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFB2B1FF2FA72FB22" box="[1255,1475,1250,1276]" italics="true" pageId="1" pageNumber="38">Delphinula diadema</emphasis>
|
||||
Stoppani, 1860
|
||||
</taxonomicName>
|
||||
-65,
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFBC31E10FA2AFAC5" authority="Stoppani, 1860" authorityName="Stoppani" authorityYear="1860" box="[1039,1435,1280,1307]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="regazzoni">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFBC31E10FB43FAC4" box="[1039,1266,1280,1306]" italics="true" pageId="1" pageNumber="38">Delphinula regazzoni</emphasis>
|
||||
Stoppani, 1860
|
||||
</taxonomicName>
|
||||
-65,
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFC8C1E0EFB05FAE7" authority="Stoppani, 1860" authorityName="Stoppani" authorityYear="1860" box="[832,1204,1310,1337]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="meriani">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFC8C1E0EFBBEFAE6" box="[832,1039,1310,1336]" italics="true" pageId="1" pageNumber="38">Delphinula meriani</emphasis>
|
||||
Stoppani, 1860
|
||||
</taxonomicName>
|
||||
-65, and
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFAC71E0EFC57FA89" authority="Stoppani, 1860" authorityName="Stoppani" authorityYear="1860" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="inzini">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFAC71E0EFA73FAE6" box="[1291,1474,1310,1336]" italics="true" pageId="1" pageNumber="38">Delphinula inzini</emphasis>
|
||||
Stoppani, 1860
|
||||
</taxonomicName>
|
||||
-65. In the same publication, he described a species from the Rhaetian of
|
||||
<collectingRegion id="462BF848FFE3FFEEFBB71E4AFB5AFAAA" box="[1147,1259,1370,1396]" country="Italy" name="Lombardia" pageId="1" pageNumber="38">Lombardy</collectingRegion>
|
||||
–
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFACB1E4BFAC2FAAA" authorityName="Grateloup" authorityYear="1832" box="[1287,1395,1371,1396]" class="Gastropoda" family="Neritopsidae" genus="Neritopsis" kingdom="Animalia" order="Cycloneritida" pageId="1" pageNumber="38" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFACB1E4BFAC2FAAA" box="[1287,1395,1371,1396]" italics="true" pageId="1" pageNumber="38">Neritopsis</emphasis>
|
||||
</taxonomicName>
|
||||
?
|
||||
<emphasis id="B69BEAB8FFE3FFEEFA4B1E4AFA73FAAA" box="[1415,1474,1370,1396]" italics="true" pageId="1" pageNumber="38">oldae</emphasis>
|
||||
Stoppani, 1860-65 – which was closely related to the former.
|
||||
</paragraph>
|
||||
<paragraph id="845036AAFFE3FFEDFCA91E87FD44FA73" blockId="1.[832,1475,135,1969]" lastBlockId="2.[113,757,1126,1967]" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
|
||||
In the revision proposed by Tommasi (1903) who compared Stoppani’s types with new material collected specifically in the type locality (specimens housed at the Museo di Storia Naturale dell’Università di Pavia), all these forms were redescribed with the following generic designations:
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFC8C1D3DFB97F999" box="[832,1062,1581,1607]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="songavatii">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFC8C1D3DFB97F999" box="[832,1062,1581,1607]" italics="true" pageId="1" pageNumber="38">Worthenia songavatii</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFBFB1D3DFB46F999" box="[1079,1271,1581,1607]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="pusilla">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFBFB1D3DFB46F999" box="[1079,1271,1581,1607]" italics="true" pageId="1" pageNumber="38">Worthenia pusilla</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFAC51D3DFA0AF999" box="[1289,1467,1581,1607]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="inzini">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFAC51D3DFA0AF999" box="[1289,1467,1581,1607]" italics="true" pageId="1" pageNumber="38">Worthenia inzini</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFC8C1D5BFBA4F9BB" box="[832,1045,1611,1637]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFC8C1D5BFBA4F9BB" box="[832,1045,1611,1637]" italics="true" pageId="1" pageNumber="38">Worthenia pygmaea</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFBE81D5BFB5AF9BB" box="[1060,1259,1611,1637]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="meriani">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFBE81D5BFB5AF9BB" box="[1060,1259,1611,1637]" italics="true" pageId="1" pageNumber="38">Worthenia meriani</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFB361D5BFA3FF9B8" box="[1274,1422,1611,1638]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="undetermined">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFB361D5BFAD9F9BB" box="[1274,1384,1611,1637]" italics="true" pageId="1" pageNumber="38">Worthenia</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
, and the new species
|
||||
<emphasis id="B69BEAB8FFE3FFEEFC261D7AFA27F95A" box="[1002,1430,1642,1668]" italics="true" pageId="1" pageNumber="38">
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFC261D7AFB75F95D" box="[1002,1220,1642,1668]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="stoppanii">Worthenia stoppanii</taxonomicName>
|
||||
.
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFB001D7AFA27F95A" authorityName="Stoppani" authorityYear="1860" box="[1228,1430,1642,1668]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="escheri">Delphinula escheri</taxonomicName>
|
||||
</emphasis>
|
||||
and
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFC8C1D98FBABF97C" authorityName="Stoppani" authorityYear="1860" box="[832,1050,1672,1698]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="diadema">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFC8C1D98FBABF97C" box="[832,1050,1672,1698]" italics="true" pageId="1" pageNumber="38">Delphinula diadema</emphasis>
|
||||
</taxonomicName>
|
||||
were tentatively attributed to the genus
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFC8C1DB6FBD9F91E" authority="Koken, 1889" authorityName="Koken" authorityYear="1889" box="[832,1128,1702,1728]" class="Gastropoda" family="Schizogoniidae" genus="Schizogonium" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFC8C1DB6FC64F91E" box="[832,981,1702,1728]" italics="true" pageId="1" pageNumber="38">Schizogonium</emphasis>
|
||||
Koken, 1889
|
||||
</taxonomicName>
|
||||
, considering possible attribution also to the genus
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFC391DD4FAA5F900" authority="De Stefani, 1880" authorityName="De Stefani" authorityYear="1880" box="[1013,1300,1732,1759]" class="Gastropoda" family="Trochidae" genus="Guidonia" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFC391DD4FBEBF900" box="[1013,1114,1732,1758]" italics="true" pageId="1" pageNumber="38">Guidonia</emphasis>
|
||||
De Stefani, 1880
|
||||
</taxonomicName>
|
||||
. This distinction regarding the genera
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFBD21DF2FB59F922" authorityName="Stoppani" authorityYear="1860" box="[1054,1256,1762,1788]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="escheri">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFBD21DF2FB59F922" box="[1054,1256,1762,1788]" italics="true" pageId="1" pageNumber="38">Delphinula escheri</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFAD61DF2FCCAF8C4" authorityName="Stoppani" authorityYear="1860" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="diadema">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFAD61DF2FCCAF8C4" italics="true" pageId="1" pageNumber="38">Delphinula diadema</emphasis>
|
||||
</taxonomicName>
|
||||
is questionable: the specimens described in Tommasi 1903 display great uniformity in their main characters, so should have been attributed to a single genus. In particular, the
|
||||
<specimenCount id="92E9FD23FFE3FFEEFCAA1C4BFC4BF8AB" box="[870,1018,1883,1909]" count="6" pageId="1" pageNumber="38" type="generic">six specimens</specimenCount>
|
||||
classified as
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFB481C4BFAA8F8AB" authorityName="Koken" authorityYear="1889" box="[1156,1305,1883,1909]" class="Gastropoda" family="Schizogoniidae" genus="Schizogonium" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFB481C4BFAA8F8AB" box="[1156,1305,1883,1909]" italics="true" pageId="1" pageNumber="38">Schizogonium</emphasis>
|
||||
</taxonomicName>
|
||||
(?)
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFA8E1C4BFA3EF8AB" authorityName="Stoppani" authorityYear="1860" box="[1346,1423,1883,1909]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="escheri">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFA8E1C4BFA3EF8AB" box="[1346,1423,1883,1909]" italics="true" pageId="1" pageNumber="38">escheri</emphasis>
|
||||
</taxonomicName>
|
||||
(cat. n.
|
||||
<collectionCode id="E2FEAE6FFFE3FFEEFC941C69FC11F84D" box="[856,928,1913,1939]" country="Italy" httpUri="http://grbio.org/cool/epb5-mbff" name="Museo di Scienze Naturali" pageId="1" pageNumber="38">MSNP</collectionCode>
|
||||
20814) – the best preserved of which was illustrated by Tommasi in Table 3, Fig. 19 – do not display any substantial differences with the other forms attributed by him to the genus
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFF2E1F94FEFEFB40" box="[226,335,1156,1182]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFF2E1F94FEFEFB40" box="[226,335,1156,1182]" italics="true" pageId="2" pageNumber="39">Worthenia</emphasis>
|
||||
</taxonomicName>
|
||||
. Indeed, the description of the growth lines and upper carina of
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFE4E1FB2FDA6FB62" authorityName="Koken" authorityYear="1889" box="[386,535,1186,1212]" class="Gastropoda" family="Schizogoniidae" genus="Schizogonium" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFE4E1FB2FDA6FB62" box="[386,535,1186,1212]" italics="true" pageId="2" pageNumber="39">Schizogonium</emphasis>
|
||||
</taxonomicName>
|
||||
(?)
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFD8F1FB2FD21FB62" authorityName="Stoppani" authorityYear="1860" box="[579,656,1186,1212]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="escheri">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFD8F1FB2FD21FB62" box="[579,656,1186,1212]" italics="true" pageId="2" pageNumber="39">escheri</emphasis>
|
||||
</taxonomicName>
|
||||
made by Tommasi implicitly suggests the presence of a selenizone. The author considered
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFEAB1FCFFDFAFB26" box="[359,587,1247,1273]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="regazzoni">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFEAB1FCFFDFAFB26" box="[359,587,1247,1273]" italics="true" pageId="2" pageNumber="39">Delphinula regazzoni</emphasis>
|
||||
</taxonomicName>
|
||||
synonym of
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFD151FCFFF63FAC9" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFD151FCFFF63FAC9" italics="true" pageId="2" pageNumber="39">W. pygmaea</emphasis>
|
||||
</taxonomicName>
|
||||
and deemed Stoppani’s drawing of
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFD9D1FEEFD69FAC9" box="[593,728,1277,1303]" class="Gastropoda" family="Trochidae" genus="Delphinula" kingdom="Animalia" order="Trochida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="regazzoni">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFD9D1FEEFDD4FAC9" box="[593,613,1278,1303]" italics="true" pageId="2" pageNumber="39">D</emphasis>
|
||||
.
|
||||
<emphasis id="B69BEAB8FFE0FFEDFDBE1FEEFD69FAC9" box="[626,728,1278,1303]" italics="true" pageId="2" pageNumber="39">regazzoni</emphasis>
|
||||
</taxonomicName>
|
||||
to be based on a single, poorly preserved specimen that, in reality, is very similar to
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFE851E2AFE7BFA8D" box="[329,458,1338,1363]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFE851E2AFE7BFA8D" box="[329,458,1338,1363]" italics="true" pageId="2" pageNumber="39">W. pygmaea</emphasis>
|
||||
</taxonomicName>
|
||||
. Even
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFDD91E29FD6DFA8D" box="[533,732,1337,1363]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="meriani">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFDD91E29FD6DFA8D" box="[533,732,1337,1363]" italics="true" pageId="2" pageNumber="39">Worthenia meriani</emphasis>
|
||||
</taxonomicName>
|
||||
is very similar to
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFEC21E48FE3FFAAF" box="[270,398,1368,1393]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFEC21E48FE3FFAAF" box="[270,398,1368,1393]" italics="true" pageId="2" pageNumber="39">W. pygmaea</emphasis>
|
||||
</taxonomicName>
|
||||
, but Tommasi thought
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFDB31E48FD45FAAF" box="[639,756,1368,1394]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="meriani">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFDB31E48FD27FAAF" box="[639,662,1368,1393]" italics="true" pageId="2" pageNumber="39">W</emphasis>
|
||||
.
|
||||
<emphasis id="B69BEAB8FFE0FFEDFD6D1E48FD45FAAF" box="[673,756,1368,1393]" italics="true" pageId="2" pageNumber="39">meriani</emphasis>
|
||||
</taxonomicName>
|
||||
to be distinct in that, in contrast with the figure in Stoppani (1860-65), the
|
||||
<typeStatus id="5B548808FFE0FFEDFEC01E83FED8FA73" box="[268,361,1427,1453]" pageId="2" pageNumber="39" type="holotype">holotype</typeStatus>
|
||||
had a base that lacked ornamentation.
|
||||
</paragraph>
|
||||
<caption id="D0906622FFE0FFEDFF71189AFB2DFBC1" ID-DOI="http://doi.org/10.5281/zenodo.12753897" ID-Zenodo-Dep="12753897" httpUri="https://zenodo.org/record/12753897/files/figure.png" pageId="2" pageNumber="39" startId="2.[189,222,906,928]" targetBox="[189,1399,136,892]" targetPageId="2" targetType="figure">
|
||||
<paragraph id="845036AAFFE0FFEDFF71189AFB2DFBC1" blockId="2.[189,1399,906,1055]" pageId="2" pageNumber="39">
|
||||
Fig. 1 - A)
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFEEB189AFDC8FC7E" authority="(Stoppani) AS" authorityName="AS" baseAuthorityName="Stoppani" box="[295,633,906,928]" class="Gastropoda" family="Wortheniellidae" genus="Wortheniella" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFEEB189AFE42FC7E" box="[295,499,906,928]" italics="true" pageId="2" pageNumber="39">Wortheniella pygmaea</emphasis>
|
||||
(Stoppani) AS
|
||||
</taxonomicName>
|
||||
41/43, lateral view, showing the pattern of the growth lines, which curve backwards above and below the nodose adapical carina (corresponding to the selenizone). Songavazzo (BG), Norian, Main Dolomite. B)
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFAEF18B4FE76FC0C" authority="(Stoppani) AS" authorityName="AS" baseAuthorityName="Stoppani" class="Gastropoda" family="Wortheniellidae" genus="Wortheniella" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFAEF18B4FE8FFC0D" italics="true" pageId="2" pageNumber="39">Wortheniella pygmaea</emphasis>
|
||||
(Stoppani) AS
|
||||
</taxonomicName>
|
||||
41/43, latero-basal view, showing the very conspicuous threads on the base. C)
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFB6618ADFEF1FC32" authority="(Stoppani) AS" authorityName="AS" baseAuthorityName="Stoppani" class="Gastropoda" family="Wortheniellidae" genus="Wortheniella" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFB6618ADFAC6FC0D" box="[1194,1399,957,979]" italics="true" pageId="2" pageNumber="39">Wortheniella pygmaea</emphasis>
|
||||
(Stoppani) AS
|
||||
</taxonomicName>
|
||||
41/43, apical view of the early whorls of the spire. D)
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFC8118C6FB17FC32" authority="(Stoppani) AS" authorityName="AS" baseAuthorityName="Stoppani" box="[845,1190,982,1004]" class="Gastropoda" family="Wortheniellidae" genus="Wortheniella" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFC8118C6FBAAFC32" box="[845,1051,982,1004]" italics="true" pageId="2" pageNumber="39">Wortheniella pygmaea</emphasis>
|
||||
(Stoppani) AS
|
||||
</taxonomicName>
|
||||
41/43, lateral view of the early whorls of the spire. E)
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFE2518E0FC8FFBDB" authority="(Stoppani) AS" authorityName="AS" baseAuthorityName="Stoppani" box="[489,830,1008,1030]" class="Gastropoda" family="Wortheniellidae" genus="Wortheniella" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFE2518E0FD07FBD8" box="[489,694,1008,1030]" italics="true" pageId="2" pageNumber="39">Wortheniella pygmaea</emphasis>
|
||||
(Stoppani) AS
|
||||
</taxonomicName>
|
||||
41/43a, lateral view. F)
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFBED18E0FAC7FBDB" authority="(Stoppani) AS" authorityName="AS" baseAuthorityName="Stoppani" box="[1057,1398,1008,1030]" class="Gastropoda" family="Wortheniellidae" genus="Wortheniella" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="pygmaea">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFBED18E0FB5FFBD8" box="[1057,1262,1008,1030]" italics="true" pageId="2" pageNumber="39">Wortheniella pygmaea</emphasis>
|
||||
(Stoppani) AS
|
||||
</taxonomicName>
|
||||
41/43a, latero-basal view, showing the region of the umbilicus, which is largely occupied by a columellar lip.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="845036AAFFE0FFEDFF5B1EA2FB5CFA51" blockId="2.[113,757,1126,1967]" lastBlockId="2.[832,1475,1126,1966]" pageId="2" pageNumber="39">
|
||||
The general appearance of these Norian species is always the same, with ornaments that are more or less accentuated. Haas (1953) extensively discussed these forms (pp. 56–63), but he did not have access to Stoppani’s or Costa’s species
|
||||
<typeStatus id="5B548808FFE0FFEDFFBD1D3BFF2EF99B" box="[113,159,1579,1605]" pageId="2" pageNumber="39">type</typeStatus>
|
||||
specimens of
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFE891D3AFD9FF99A" box="[325,558,1578,1604]" class="Gastropoda" family="Trochidae" genus="Trochus" kingdom="Animalia" order="Trochida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="contabulatus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFE891D3AFD9FF99A" box="[325,558,1578,1604]" italics="true" pageId="2" pageNumber="39">Trochus contabulatus</emphasis>
|
||||
</taxonomicName>
|
||||
Costa, 1864, and
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFFBD1D58FED3F9BC" baseAuthorityName="Costa" baseAuthorityYear="1964" box="[113,354,1608,1634]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="contabulata">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFFBD1D58FED3F9BC" box="[113,354,1608,1634]" italics="true" pageId="2" pageNumber="39">Worthenia contabulata</emphasis>
|
||||
</taxonomicName>
|
||||
, which had probably already been lost during
|
||||
<collectionCode id="E2FEAE6FFFE0FFEDFF711D77FEB3F95F" box="[189,258,1639,1665]" pageId="2" pageNumber="39">WWII</collectionCode>
|
||||
(p. 62). Not being able to study the
|
||||
<typeStatus id="5B548808FFE0FFEDFD4A1D77FD70F95F" box="[646,705,1639,1665]" pageId="2" pageNumber="39">types</typeStatus>
|
||||
, Haas compared his Peruvian specimens with the material from Main Dolomite of Esztergár Valley,
|
||||
<collectingCountry id="FCF8763AFFE0FFEDFE301DB3FDEBF963" box="[508,602,1699,1725]" name="Hungary" pageId="2" pageNumber="39">Hungary</collectingCountry>
|
||||
, illustrated by Kittl (1900). According to Haas, in that material the growth lines, which in the figures in Kittl (1900) suggest the presence of a selenizone, are in reality deformed and rather axial on the flanks indicating that the shell lacks a true selenizone. Haas attributed those forms to the genus
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFE171C29FD45F88D" authority="De Stefani, 1880" authorityName="De Stefani" authorityYear="1880" box="[475,756,1849,1876]" class="Gastropoda" family="Trochidae" genus="Guidonia" kingdom="Animalia" order="Trochida" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFE171C29FDF1F88D" box="[475,576,1849,1875]" italics="true" pageId="2" pageNumber="39">Guidonia</emphasis>
|
||||
De Stefani, 1880
|
||||
</taxonomicName>
|
||||
(family
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFF081C47FE30F8AC" authorityName="Zittel" authorityYear="1895" box="[196,385,1879,1906]" class="Gastropoda" family="Trochonematidae" kingdom="Animalia" pageId="2" pageNumber="39" phylum="Mollusca" rank="family">Trochonematidae</taxonomicName>
|
||||
), amending the diagnosis of
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFD641C47FF22F851" authorityName="De Stefani" authorityYear="1880" class="Gastropoda" family="Trochidae" genus="Guidonia" kingdom="Animalia" order="Trochida" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFD641C47FF22F851" italics="true" pageId="2" pageNumber="39">Guidonia</emphasis>
|
||||
</taxonomicName>
|
||||
by identifying a
|
||||
<typeStatus id="5B548808FFE0FFEDFE801C66FECBF84E" box="[332,378,1910,1936]" pageId="2" pageNumber="39">type</typeStatus>
|
||||
species (
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFE121C65FF18F871" authority="Stoliczka, 1861" authorityName="Stoliczka" authorityYear="1861" class="Gastropoda" family="Trochidae" genus="Trochus" kingdom="Animalia" order="Trochida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="rotulus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFE121C65FD35F851" box="[478,644,1909,1935]" italics="true" pageId="2" pageNumber="39">Trochus rotulus</emphasis>
|
||||
Stoliczka, 1861
|
||||
</taxonomicName>
|
||||
), because De Stefani formally did not indicate it (for a history of the genus see: Gatto & Monari, 2010). In contrast, the specimens described here, like those in Tommasi (1903), display evidently prosocyrt growth lines on the sutural ramp and on the flank, with a marked edge in correspondence with the adapical carina, which is very much like the typical selenizone of
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFC601FECFBA8FAC8" box="[940,1049,1276,1302]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFC601FECFBA8FAC8" box="[940,1049,1276,1302]" italics="true" pageId="2" pageNumber="39">Worthenia</emphasis>
|
||||
</taxonomicName>
|
||||
. The presence of a true selenizone was hypothesised also in Tichy (1975) on specimens of
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFA981E0AFC0EFA8C" baseAuthorityName="Costa" baseAuthorityYear="1964" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="contabulata">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFA981E0AFC0EFA8C" italics="true" pageId="2" pageNumber="39">Worthenia contabulata</emphasis>
|
||||
</taxonomicName>
|
||||
from Main Dolomite from
|
||||
<collectingCountry id="FCF8763AFFE0FFEDFB111E28FA9FFA8C" box="[1245,1326,1336,1362]" name="Austria" pageId="2" pageNumber="39">Austria</collectingCountry>
|
||||
. However, Tichy did not have well-preserved specimens at his disposal, so was not able to observe the growth lines.
|
||||
</paragraph>
|
||||
<paragraph id="845036AAFFE0FFECFCA91E83FD44FDF7" blockId="2.[832,1475,1126,1966]" lastBlockId="3.[113,757,135,554]" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39">
|
||||
Thus, giving as certain the attribution of these specimens to the “
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFC541EA1FBB7FA15" box="[920,1030,1457,1483]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFC541EA1FBB7FA15" box="[920,1030,1457,1483]" italics="true" pageId="2" pageNumber="39">Worthenia</emphasis>
|
||||
</taxonomicName>
|
||||
group”, these forms must be classified more precisely at the genus level. A possible solution derives from the study of the first whorls that appear fairly well preserved in
|
||||
<collectingCountry id="FCF8763AFFE0FFEDFC091D1CFC56F9F8" box="[965,999,1548,1574]" name="American Samoa" pageId="2" pageNumber="39">AS</collectingCountry>
|
||||
41/43. Research on the juvenile stages of the “
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFC871D39FC08F99D" box="[843,953,1577,1603]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFC871D39FC08F99D" box="[843,953,1577,1603]" italics="true" pageId="2" pageNumber="39">Worthenia</emphasis>
|
||||
</taxonomicName>
|
||||
group” has brought to the designation of new genera that are rather similar in relation to their teleoconchs. As mentioned by Nuetzel & Senowbari-Darian (1999), Chronic (1952) erected the genus
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFB981D94FB49F940" authorityName="Chronic" authorityYear="1952" box="[1108,1272,1668,1694]" class="Gastropoda" family="Lophospiridae" genus="Platyworthenia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFB981D94FB49F940" box="[1108,1272,1668,1694]" italics="true" pageId="2" pageNumber="39">Platyworthenia</emphasis>
|
||||
</taxonomicName>
|
||||
for some Permian
|
||||
<collectionCode id="E2FEAE6FFFE0FFEDFC8C1DB2FCD2F962" box="[832,867,1698,1724]" country="South Africa" name="University of Stellenbosch" pageId="2" pageNumber="39">US</collectionCode>
|
||||
forms that differ from
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFB811DB2FAEAF962" authority="Koninck, 1883" authorityName="De Koninck" authorityYear="1883" box="[1101,1371,1698,1724]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFB811DB2FB0AF962" box="[1101,1211,1698,1724]" italics="true" pageId="2" pageNumber="39">Worthenia</emphasis>
|
||||
Koninck,
|
||||
<quantity id="43179B4FFFE0FFEDFAEA1DB2FAC5F962" box="[1318,1396,1698,1724]" metricMagnitude="1" metricUnit="m" metricValue="4.782819999999999" pageId="2" pageNumber="39" unit="in" value="1883.0">1883 in</quantity>
|
||||
</taxonomicName>
|
||||
having very low early whorls. Subsequently, Knight
|
||||
<emphasis id="B69BEAB8FFE0FFEDFAE91DD1FAEAF904" box="[1317,1371,1728,1754]" italics="true" pageId="2" pageNumber="39">et al.</emphasis>
|
||||
(1960, p. I209) considered
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFBC91DCEFB18F926" authorityName="Chronic" authorityYear="1952" box="[1029,1193,1758,1784]" class="Gastropoda" family="Lophospiridae" genus="Platyworthenia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFBC91DCEFB18F926" box="[1029,1193,1758,1784]" italics="true" pageId="2" pageNumber="39">Platyworthenia</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFB251DCEFAE6F926" authorityName="De Koninck" authorityYear="1883" box="[1257,1367,1758,1784]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFB251DCEFAE6F926" box="[1257,1367,1758,1784]" italics="true" pageId="2" pageNumber="39">Worthenia</emphasis>
|
||||
</taxonomicName>
|
||||
as synonyms, on account of their great similarity. Later Yoo (1994) illustrated the protoconch and the first teleoconch whorls of the Carboniferous specimens attributed by him to
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFA981C29FA73F88D" authorityName="De Koninck" authorityYear="1883" box="[1364,1474,1849,1875]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFA981C29FA73F88D" box="[1364,1474,1849,1875]" italics="true" pageId="2" pageNumber="39">Worthenia</emphasis>
|
||||
</taxonomicName>
|
||||
demonstrating that the early shell is trochiform. Schwardt (1992) erected the genus
|
||||
<taxonomicName id="43EF4D29FFE0FFEDFB981C65FB6DF851" authorityName="Schwardt" authorityYear="1992" box="[1108,1244,1909,1935]" class="Gastropoda" family="Wortheniellidae" genus="Wortheniella" kingdom="Animalia" order="Pleurotomariida" pageId="2" pageNumber="39" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE0FFEDFB981C65FB6DF851" box="[1108,1244,1909,1935]" italics="true" pageId="2" pageNumber="39">Wortheniella</emphasis>
|
||||
</taxonomicName>
|
||||
for specimens of the Upper Triassic St. Cassian Formation characterised by planispiral juvenile whorls. Nuetzel & Senowbari-Darian (1999, p. 97) assigned the Norian species from the Iranian Nayband Formation to the genus
|
||||
<taxonomicName id="43EF4D29FFE1FFECFEBF1BD4FE4BFF00" authorityName="Schwardt" authorityYear="1992" box="[371,506,196,222]" class="Gastropoda" family="Wortheniellidae" genus="Wortheniella" kingdom="Animalia" order="Pleurotomariida" pageId="3" pageNumber="40" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE1FFECFEBF1BD4FE4BFF00" box="[371,506,196,222]" italics="true" pageId="3" pageNumber="40">Wortheniella</emphasis>
|
||||
</taxonomicName>
|
||||
, and discussed the differences between the genera
|
||||
<taxonomicName id="43EF4D29FFE1FFECFE611BF2FD40FF22" authority="De Koninck, 1883" authorityName="De Koninck" authorityYear="1883" box="[429,753,226,252]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="3" pageNumber="40" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE1FFECFE611BF2FDAAFF22" box="[429,539,226,252]" italics="true" pageId="3" pageNumber="40">Worthenia</emphasis>
|
||||
De Koninck, 1883
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="43EF4D29FFE1FFECFFBD1A10FE1EFEC4" authority="Schwardt, 1992" authorityName="Schwardt" authorityYear="1992" box="[113,431,256,282]" class="Gastropoda" family="Wortheniellidae" genus="Wortheniella" kingdom="Animalia" order="Pleurotomariida" pageId="3" pageNumber="40" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE1FFECFFBD1A10FF48FEC4" box="[113,249,256,282]" italics="true" pageId="3" pageNumber="40">Wortheniella</emphasis>
|
||||
Schwardt, 1992
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="43EF4D29FFE1FFECFE261A10FF18FEE7" authority="Chronic, 1952" authorityName="Chronic" authorityYear="1952" class="Gastropoda" family="Lophospiridae" genus="Platyworthenia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="40" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE1FFECFE261A10FD3FFEC4" box="[490,654,256,282]" italics="true" pageId="3" pageNumber="40">Platyworthenia</emphasis>
|
||||
Chronic, 1952
|
||||
</taxonomicName>
|
||||
, which are certainly related to each other. They concluded that this distinction depends upon the importance one wants to bestow upon the morphology of first teleoconch whorls. Conversely, Nuetzel & Erwin (2004) assigned the species of this group from the Norian deposits of
|
||||
<collectingRegion id="462BF848FFE1FFECFD7B1A87FD45FE6F" box="[695,756,407,433]" country="United States of America" name="Idaho" pageId="3" pageNumber="40">Idaho</collectingRegion>
|
||||
(
|
||||
<collectingCountry id="FCF8763AFFE1FFECFFB61AA5FF75FE11" box="[122,196,437,463]" name="United States of America" pageId="3" pageNumber="40">U.S.A.</collectingCountry>
|
||||
) to
|
||||
<taxonomicName id="43EF4D29FFE1FFECFF3C1AA5FEECFE11" authorityName="De Koninck" authorityYear="1883" box="[240,349,437,463]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="3" pageNumber="40" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE1FFECFF3C1AA5FEECFE11" box="[240,349,437,463]" italics="true" pageId="3" pageNumber="40">Worthenia</emphasis>
|
||||
</taxonomicName>
|
||||
. Finally, Bandel (2009, p. 17) erected the family
|
||||
<taxonomicName id="43EF4D29FFE1FFECFF2E1AC3FE3DFE33" authorityName="Bandel" authorityYear="2009" box="[226,396,467,493]" class="Gastropoda" family="Wortheniellidae" kingdom="Animalia" order="Pleurotomariida" pageId="3" pageNumber="40" phylum="Mollusca" rank="family">Wortheniellidae</taxonomicName>
|
||||
, describing the protoconchs of the Carnian forms from the St. Cassian Formation as sinistral and situated a little under the first whorl of the teleoconch.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
65
data/0C/46/87/0C4687BCFFE3FFEEFFBD182DFD33FCAB.xml
Normal file
65
data/0C/46/87/0C4687BCFFE3FFEEFFBD182DFD33FCAB.xml
Normal file
|
|
@ -0,0 +1,65 @@
|
|||
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||||
<subSubSection id="CCF56521FFE3FFEEFFBD182DFE5FFC89" box="[113,494,828,855]" pageId="1" pageNumber="38" type="nomenclature">
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||||
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||||
Genus
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFF71182CFE5FFC89" ID-CoL="7PL4Q" authority="Schwardt 1992" authorityName="Schwardt" authorityYear="1992" box="[189,494,828,855]" class="Gastropoda" family="Wortheniellidae" genus="Wortheniella" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="genus">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFF71182CFEF7FC88" box="[189,326,828,854]" italics="true" pageId="1" pageNumber="38">Wortheniella</emphasis>
|
||||
Schwardt 1992
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CCF56521FFE3FFEEFFBD184BFD33FCAB" box="[113,642,859,885]" pageId="1" pageNumber="38" type="materials_examined">
|
||||
<paragraph id="845036AAFFE3FFEEFFBD184BFD33FCAB" blockId="1.[113,756,708,885]" box="[113,642,859,885]" pageId="1" pageNumber="38">
|
||||
<typeStatus id="5B548808FFE3FFEEFFBD184BFF16FCAB" box="[113,167,859,885]" pageId="1" pageNumber="38">Type</typeStatus>
|
||||
species:
|
||||
<taxonomicName id="43EF4D29FFE3FFEEFEC4184BFD33FCAB" authority="Kittl, 1891" authorityName="Kittl" authorityYear="1891" box="[264,642,859,885]" class="Gastropoda" family="Lophospiridae" genus="Worthenia" kingdom="Animalia" order="Pleurotomariida" pageId="1" pageNumber="38" phylum="Mollusca" rank="species" species="coralliophila">
|
||||
<emphasis id="B69BEAB8FFE3FFEEFEC4184BFDB8FCAB" box="[264,521,859,885]" italics="true" pageId="1" pageNumber="38">Worthenia coralliophila</emphasis>
|
||||
Kittl, 1891
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
Loading…
Reference in a new issue