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<mods:title id="95DF7A69FC06105D0141D06E40597DEA">Pterosaur teeth from the Southern Neuquén Basin (Patagonia, Argentina): New insights on the reconstruction of ornithocheiriform dental anatomy</mods:title>
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<paragraph id="8BB42E56FFE22530374DF8E0FDBFF8A0" blockId="3.[98,537,1694,1884]" box="[98,537,1818,1847]" pageId="3" pageNumber="76">
<taxonomicName id="4C0B55D5FFE22530374DF8E0FDBFF8A0" authority="Andres, 2021" authorityName="Andres" authorityYear="2021" box="[98,537,1818,1847]" kingdom="Animalia" order="Pterosauria" pageId="3" pageNumber="76">
Ornithocheiriformes
<bibRefCitation id="EF9A53A7FFE225303642F8E0FDBFF8A0" author="Andres, B." box="[365,537,1818,1847]" pageId="3" pageNumber="76" pagination="203 - 2017" refId="ref9426" refString="Andres, B. 2021. Phylogenetic systematics of Quetzalcoatlus Lawson 1975 (Pterodactyloidea: Azhdarchoidea). Journal of Vertebrate Paleontology 41: 203 - 2017." type="journal article" year="2021">Andres, 2021</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="C3117DDDFFE22530374DF8BCFF65F8CB" box="[98,195,1862,1884]" pageId="3" pageNumber="76" type="description">
<paragraph id="8BB42E56FFE22530374DF8BCFF65F8CB" blockId="3.[98,537,1694,1884]" box="[98,195,1862,1884]" pageId="3" pageNumber="76">
<figureCitation id="133032D3FFE22530374DF8BCFF19F8CB" box="[98,191,1862,1884]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="4.[151,184,981,1003]" captionStartId-1="5.[97,130,967,989]" captionStartId-2="5.[98,131,1822,1844]" captionStartId-3="6.[151,184,914,936]" captionTargetBox-0="[157,1535,166,951]" captionTargetBox-1="[101,768,166,937]" captionTargetBox-2="[101,767,1126,1795]" captionTargetBox-3="[154,821,163,887]" captionTargetId-0="figure-302@4.[152,1535,161,956]" captionTargetId-1="figure-436@5.[98,766,176,938]" captionTargetId-2="figure-525@5.[99,770,1123,1796]" captionTargetId-3="figure-779@6.[153,821,173,888]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Fig. 3. Isolated pterosaur teeth from the Cerro de los Leones locality, Albian, Lower Cretaceous, Picún Leufú, Argentina. Morphotype 1 (H, J, L, N), Morphotype 2 (A, C, M, P, Q, R, S), and Morphotype 3 (B, D, E, G, I, K, O, T). A. MCF-PVPH-739-2. B. MCF-PVPH-741. C. MCF-PVPH-743. D. MCFPVPH-875. E. MCF-PVPH-879-1. F. MCF-PVPH-879-2. G. MCF-PVPH-879-3. H. MCF-PVPH-879-4. I. MCF-PVPH-879-5. J. MCF- PVPH-879-6. K. MCF-PVPH-879-7. L. MCF-PVPH-879-8. M. MCF-PVPH-879-9. N. MCF-PVPH-879-10. O. MCF-PVPH-879-11. P. MCF- PVPH-880-1. Q. MCFPVPH-880-2. R. MCF-PVPH-880-3. S. MCF-PVPH-880-4. T. MCF-PVPH-880-5. In labial (AG, I, K, OQ, S, T), lingual (H, M, N, R), and mesial (J, L) views." captionText-1="Fig.4.Isolated tooth (Morphotype 1) from the Cerro de los Leones locality, Albian, Lower Cretaceous, Picún Leufú, Argentina. MCF-PVPH-879-8 in labial (A1, A2), lingual (A3, A4), mesial (A5, A6), and distal (A7,A8) views. The basal cross section (A9, A10) is visible due to the natural breaking of the crown." captionText-2="Fig. 5. Isolated tooth (Morphotype 2) from the Cerro de los Leones locality,Albian, Lower Cretaceous, Picún Leufú,Argentina.MCF-PVPH-880-1 in labial (A1, A2), lingual (A3, A4), distal (A5, A6), and mesial (A7, A8) views. The basal cross section (A9,A10) is visible due to the natural breaking of the crown." captionText-3="Fig. 6. Isolated tooth (Morphotype 3) from the Cretaceous Cerro de los Leones locality, Picún Leufú, Argentina. MCF-PVPH-879-2 in labial (A1, A2), lingual (A3, A4), distal (A5, A6), and mesial (A7, A8) views. The basal cross section (A9, A10) is visible due to the natural breaking of the crown." figureDoi-0="http://doi.org/10.5281/zenodo.12194864" figureDoi-1="http://doi.org/10.5281/zenodo.12194866" figureDoi-2="http://doi.org/10.5281/zenodo.12194870" figureDoi-3="http://doi.org/10.5281/zenodo.12194872" httpUri-0="https://zenodo.org/record/12194864/files/figure.png" httpUri-1="https://zenodo.org/record/12194866/files/figure.png" httpUri-2="https://zenodo.org/record/12194870/files/figure.png" httpUri-3="https://zenodo.org/record/12194872/files/figure.png" pageId="3" pageNumber="76">Figs. 36</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3117DDDFFE22530374DF897FA19F945" pageId="3" pageNumber="76" type="materials_examined">
<paragraph id="8BB42E56FFE22530374DF897FA19F945" blockId="3.[98,771,1901,1959]" lastBlockId="3.[808,1481,1619,1747]" pageId="3" pageNumber="76">
<emphasis id="B97FF244FFE22530374DF897FF62F810" box="[98,196,1901,1927]" italics="true" pageId="3" pageNumber="76">Material</emphasis>
.—
<materialsCitation id="3B63240BFFE2253037CBF894FA19F945" pageId="3" pageNumber="76">
Twenty isolated teeth: MCF-PVPH-879-4, 879- 6, 879-8, 879-10 (
<figureCitation id="133032D3FFE225303617F877FE24F830" box="[312,386,1933,1959]" captionStart="Fig" captionStartId="5.[97,130,967,989]" captionTargetBox="[101,768,166,937]" captionTargetId="figure-436@5.[98,766,176,938]" captionTargetPageId="5" captionText="Fig.4.Isolated tooth (Morphotype 1) from the Cerro de los Leones locality, Albian, Lower Cretaceous, Picún Leufú, Argentina. MCF-PVPH-879-8 in labial (A1, A2), lingual (A3, A4), mesial (A5, A6), and distal (A7,A8) views. The basal cross section (A9, A10) is visible due to the natural breaking of the crown." figureDoi="http://doi.org/10.5281/zenodo.12194866" httpUri="https://zenodo.org/record/12194866/files/figure.png" pageId="3" pageNumber="76">Fig. 4</figureCitation>
); MCF-PVPH-739-2, 743, 879-9, 880-1, 880-2, 880-3, and 880-4 (
<figureCitation id="133032D3FFE225303386F9A9FB56F9F9" box="[1193,1264,1619,1646]" captionStart="Fig" captionStartId="5.[98,131,1822,1844]" captionTargetBox="[101,767,1126,1795]" captionTargetId="figure-525@5.[99,770,1123,1796]" captionTargetPageId="5" captionText="Fig. 5. Isolated tooth (Morphotype 2) from the Cerro de los Leones locality,Albian, Lower Cretaceous, Picún Leufú,Argentina.MCF-PVPH-880-1 in labial (A1, A2), lingual (A3, A4), distal (A5, A6), and mesial (A7, A8) views. The basal cross section (A9,A10) is visible due to the natural breaking of the crown." figureDoi="http://doi.org/10.5281/zenodo.12194870" httpUri="https://zenodo.org/record/12194870/files/figure.png" pageId="3" pageNumber="76">Fig. 5</figureCitation>
); MCF-PVPH-741, 875, 879-1, 879-2, 879-3, 879-5, 879-7, 879-11, and 880-5 (
<figureCitation id="133032D3FFE22530341FF96DFCD1F926" box="[816,887,1687,1713]" captionStart="Fig" captionStartId="6.[151,184,914,936]" captionTargetBox="[154,821,163,887]" captionTargetId="figure-779@6.[153,821,173,888]" captionTargetPageId="6" captionText="Fig. 6. Isolated tooth (Morphotype 3) from the Cretaceous Cerro de los Leones locality, Picún Leufú, Argentina. MCF-PVPH-879-2 in labial (A1, A2), lingual (A3, A4), distal (A5, A6), and mesial (A7, A8) views. The basal cross section (A9, A10) is visible due to the natural breaking of the crown." figureDoi="http://doi.org/10.5281/zenodo.12194872" httpUri="https://zenodo.org/record/12194872/files/figure.png" pageId="3" pageNumber="76">Fig. 6</figureCitation>
). All from the Cretaceous Cerro de los Leones locality, Albian, Lower Cretaceous, Picún Leufú,
<collectingCountry id="F31C6EC6FFE22530326AF942FA1DF945" box="[1349,1467,1720,1746]" name="Argentina" pageId="3" pageNumber="76">Argentina</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3117DDDFFE225363407F91CFBE3FADF" lastPageId="5" lastPageNumber="78" pageId="3" pageNumber="76" type="description">
<paragraph id="8BB42E56FFE225373407F91CFE0FFA18" blockId="3.[808,1481,1765,1959]" lastBlockId="4.[151,825,1164,1423]" lastPageId="4" lastPageNumber="77" pageId="3" pageNumber="76">
<emphasis id="B97FF244FFE225303407F91CFC16F968" box="[808,944,1766,1791]" italics="true" pageId="3" pageNumber="76">Description</emphasis>
.—All specimens here described are represented by isolated tooth crowns with reabsorbed or lacking roots (
<figureCitation id="133032D3FFE22530345BF8D2FC1DF8D5" box="[884,955,1832,1858]" captionStart="Fig" captionStartId="4.[151,184,981,1003]" captionTargetBox="[157,1535,166,951]" captionTargetId="figure-302@4.[152,1535,161,956]" captionTargetPageId="4" captionText="Fig. 3. Isolated pterosaur teeth from the Cerro de los Leones locality, Albian, Lower Cretaceous, Picún Leufú, Argentina. Morphotype 1 (H, J, L, N), Morphotype 2 (A, C, M, P, Q, R, S), and Morphotype 3 (B, D, E, G, I, K, O, T). A. MCF-PVPH-739-2. B. MCF-PVPH-741. C. MCF-PVPH-743. D. MCFPVPH-875. E. MCF-PVPH-879-1. F. MCF-PVPH-879-2. G. MCF-PVPH-879-3. H. MCF-PVPH-879-4. I. MCF-PVPH-879-5. J. MCF- PVPH-879-6. K. MCF-PVPH-879-7. L. MCF-PVPH-879-8. M. MCF-PVPH-879-9. N. MCF-PVPH-879-10. O. MCF-PVPH-879-11. P. MCF- PVPH-880-1. Q. MCFPVPH-880-2. R. MCF-PVPH-880-3. S. MCF-PVPH-880-4. T. MCF-PVPH-880-5. In labial (AG, I, K, OQ, S, T), lingual (H, M, N, R), and mesial (J, L) views." figureDoi="http://doi.org/10.5281/zenodo.12194864" httpUri="https://zenodo.org/record/12194864/files/figure.png" pageId="3" pageNumber="76">Fig. 3</figureCitation>
). In this sense, the apical wear facets, when present, and the lack of the roots do not seem to have been caused by weathering events and/or taphonomic processes, suggesting that the teeth could have been lost in life. The tooth-crowns are conidont (cone-shaped, sensu
<bibRefCitation id="EF9A53A7FFE5253735EFFB76FEB3FB5F" author="Hendrickx, C. &amp; Mateus, O. &amp; Araujo, R." pageId="4" pageNumber="77" pagination="982797" refId="ref11070" refString="Hendrickx, C., Mateus, O., and Araujo, R. 2015. A proposed terminology of theropod teeth (Dinosauria, Saurischia), Journal of Vertebrate Paleontology 35 (5): e 982797." type="journal article" year="2015">Hendrickx et al. 2015</bibRefCitation>
), apicobasally elongated and slightly recurved. The cross-sections are diversified in shape, subcircular, oval or elliptical. Taking into account the general morphology of the teeth, we identify three morphotypes, which are defined on the basis of three morphological features (
<tableCitation id="C6891BEDFFE5253735FAFAC8FC8EFADB" box="[725,808,1330,1356]" captionStart="Table 1" captionStartId="4.[152,205,1757,1779]" captionTargetId="graphics-470@4.[152,1535,1796,1952]" captionTargetPageId="4" captionText="Table 1. Most inclusive morphological characterization of the Cerro de Los Leones teeth, divided in three morphotypes." pageId="4" pageNumber="77">Table 1</tableCitation>
): tooth crown curvature, shape of the crown cross-section, and presence of carinae.
</paragraph>
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<paragraph id="8BB42E56FFE5253737B8FC2FFE68FBE0" blockId="4.[151,1537,981,1143]" pageId="4" pageNumber="77">
Fig. 3. Isolated pterosaur teeth from the Cerro de los Leones locality, Albian, Lower Cretaceous, Picún Leufú, Argentina.
<taxonomicName id="4C0B55D5FFE52537322DFC2FFAD4FC7C" box="[1282,1394,981,1003]" class="Mammalia" family="Amphicyonidae" genus="Morphotype" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="79" phylum="Chordata" rank="genus">Morphotype</taxonomicName>
1 (H, J, L, N),
<taxonomicName id="4C0B55D5FFE5253737B8FC0BFEA1FB90" box="[151,263,1009,1031]" class="Mammalia" family="Amphicyonidae" genus="Morphotype" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="79" phylum="Chordata" rank="genus">Morphotype</taxonomicName>
2 (A, C, M, P, Q, R, S), and
<taxonomicName id="4C0B55D5FFE52537350AFC0BFD33FB90" box="[549,661,1009,1031]" class="Mammalia" family="Amphicyonidae" genus="Morphotype" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="79" phylum="Chordata" rank="genus">Morphotype</taxonomicName>
3 (B, D, E, G, I, K, O, T).
<emphasis id="B97FF244FFE52537348EFC0BFC17FB91" bold="true" box="[929,945,1009,1030]" pageId="4" pageNumber="77">A</emphasis>
. MCF-PVPH-739-2.
<emphasis id="B97FF244FFE525373354FC0BFB2CFB91" bold="true" box="[1147,1162,1009,1030]" pageId="4" pageNumber="77">B</emphasis>
. MCF-PVPH-741.
<emphasis id="B97FF244FFE52537326FFC0BFAF6FB90" bold="true" box="[1344,1360,1009,1031]" pageId="4" pageNumber="77">C</emphasis>
. MCF-PVPH-743.
<emphasis id="B97FF244FFE5253737B8FBF7FF01FBB5" bold="true" box="[151,167,1037,1058]" pageId="4" pageNumber="77">D</emphasis>
. MCFPVPH-875.
<emphasis id="B97FF244FFE525373674FBF7FECCFBB5" bold="true" box="[347,362,1037,1058]" pageId="4" pageNumber="77">E</emphasis>
. MCF-PVPH-879-1.
<emphasis id="B97FF244FFE525373516FBF7FDE1FBB5" bold="true" box="[569,583,1037,1058]" pageId="4" pageNumber="77">F</emphasis>
. MCF-PVPH-879-2.
<emphasis id="B97FF244FFE52537343AFBF7FC81FBB4" bold="true" box="[789,807,1037,1059]" pageId="4" pageNumber="77">G</emphasis>
. MCF-PVPH-879-3.
<emphasis id="B97FF244FFE5253734DAFBF7FBA1FBB5" bold="true" box="[1013,1031,1037,1058]" pageId="4" pageNumber="77">H</emphasis>
. MCF-PVPH-879-4.
<emphasis id="B97FF244FFE5253733FAFBF7FB78FBB5" bold="true" box="[1237,1246,1037,1058]" pageId="4" pageNumber="77">I</emphasis>
. MCF-PVPH-879-5.
<emphasis id="B97FF244FFE525373283FBF7FA11FBB5" bold="true" box="[1452,1463,1037,1058]" pageId="4" pageNumber="77">J</emphasis>
. MCFPVPH-879-6.
<emphasis id="B97FF244FFE52537360FFBD3FE94FBA9" bold="true" box="[288,306,1065,1086]" pageId="4" pageNumber="77">K</emphasis>
. MCF-PVPH-879-7.
<emphasis id="B97FF244FFE52537352CFBD3FDB4FBA9" bold="true" box="[515,530,1065,1086]" pageId="4" pageNumber="77">L</emphasis>
. MCF-PVPH-879-8.
<emphasis id="B97FF244FFE5253735CAFBD3FD5CFBA9" bold="true" box="[741,762,1065,1086]" pageId="4" pageNumber="77">M</emphasis>
. MCF-PVPH-879-9.
<emphasis id="B97FF244FFE5253734E3FBD3FC7AFBA9" bold="true" box="[972,988,1065,1086]" pageId="4" pageNumber="77">N</emphasis>
. MCF-PVPH-879-10.
<emphasis id="B97FF244FFE525373395FBD3FB6AFBA8" bold="true" box="[1210,1228,1065,1087]" pageId="4" pageNumber="77">O</emphasis>
. MCF-PVPH-879-11.
<emphasis id="B97FF244FFE525373287FBD3FA10FBA9" bold="true" box="[1448,1462,1065,1086]" pageId="4" pageNumber="77">P</emphasis>
. MCFPVPH-880-1.
<emphasis id="B97FF244FFE525373635FBBFFE8AFBCC" bold="true" box="[282,300,1093,1115]" pageId="4" pageNumber="77">Q</emphasis>
. MCFPVPH-880-2.
<emphasis id="B97FF244FFE5253736C5FBBFFE5CFBCD" bold="true" box="[490,506,1093,1114]" pageId="4" pageNumber="77">R</emphasis>
. MCF-PVPH-880-3.
<emphasis id="B97FF244FFE5253735EFFBBEFD6BFBCD" bold="true" box="[704,717,1092,1114]" pageId="4" pageNumber="77">S</emphasis>
. MCF-PVPH-880-4.
<emphasis id="B97FF244FFE5253734BCFBBFFC04FBCD" bold="true" box="[915,930,1093,1114]" pageId="4" pageNumber="77">T</emphasis>
. MCF-PVPH-880-5. In labial (AG, I, K, OQ, S, T), lingual (H, M, N, R), and mesial (J, L) views.
</paragraph>
</caption>
<paragraph id="8BB42E56FFE5253737B8FA5BFA58FA46" blockId="4.[151,824,1441,1733]" lastBlockId="4.[862,1535,1164,1490]" pageId="4" pageNumber="77">
<emphasis id="B97FF244FFE5253737B8FA5BFE9FFA2C" box="[151,313,1441,1467]" italics="true" pageId="4" pageNumber="77">
<taxonomicName id="4C0B55D5FFE5253737B8FA5BFE84FA2C" box="[151,290,1441,1467]" class="Mammalia" family="Amphicyonidae" genus="Morphotype" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="77" phylum="Chordata" rank="genus">Morphotype</taxonomicName>
1
</emphasis>
.—MCF-PVPH-879-4, 879-6, 879-8, 879-10 (
<figureCitation id="133032D3FFE5253737B0FA38FF45FA4B" box="[159,227,1474,1500]" captionStart="Fig" captionStartId="5.[97,130,967,989]" captionTargetBox="[101,768,166,937]" captionTargetId="figure-436@5.[98,766,176,938]" captionTargetPageId="5" captionText="Fig.4.Isolated tooth (Morphotype 1) from the Cerro de los Leones locality, Albian, Lower Cretaceous, Picún Leufú, Argentina. MCF-PVPH-879-8 in labial (A1, A2), lingual (A3, A4), mesial (A5, A6), and distal (A7,A8) views. The basal cross section (A9, A10) is visible due to the natural breaking of the crown." figureDoi="http://doi.org/10.5281/zenodo.12194866" httpUri="https://zenodo.org/record/12194866/files/figure.png" pageId="4" pageNumber="77">Fig. 4</figureCitation>
). These tooth crowns are apicobasally elongated and strongly lingually recurved. The basal cross-section is subcircular, and weakly labiolingually compressed. All specimens lack carinae. In MCF-PVPH-879-4, 879-8, and 879-10 the enamel layer is fragmentary, exposing the underlying dentine layer. The dentine surface is mainly smooth, but several transversal and longitudinal fractures are present, especially on the lingual and apical surfaces. The preserved enamel surface is slightly wrinkled, showing a set of anastomosed, apicobasally extended ridges and grooves, which are more pronounced basally and progressively weaker apically. On the other hand, MCF-PVPH-879-6 preserves small fragments of enamel on the labial and lingual surfaces, resulting in an almost smooth surface. The specimens where the apical portion is preserved (MCF-PVPH-879-6 and 879-8) show wear facets in apical, labial or mesial position. In MCF-PVPH-879-8 the apical wear facet is wide and rounded, extending basally to the half of the crown (
<figureCitation id="133032D3FFE525373282FA4DFA57FA46" box="[1453,1521,1463,1489]" captionStart="Fig" captionStartId="5.[97,130,967,989]" captionTargetBox="[101,768,166,937]" captionTargetId="figure-436@5.[98,766,176,938]" captionTargetPageId="5" captionText="Fig.4.Isolated tooth (Morphotype 1) from the Cerro de los Leones locality, Albian, Lower Cretaceous, Picún Leufú, Argentina. MCF-PVPH-879-8 in labial (A1, A2), lingual (A3, A4), mesial (A5, A6), and distal (A7,A8) views. The basal cross section (A9, A10) is visible due to the natural breaking of the crown." figureDoi="http://doi.org/10.5281/zenodo.12194866" httpUri="https://zenodo.org/record/12194866/files/figure.png" pageId="4" pageNumber="77">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BB42E56FFE525363471FA19FC50FDED" blockId="4.[862,1536,1507,1733]" lastBlockId="5.[808,1482,158,634]" lastPageId="5" lastPageNumber="78" pageId="4" pageNumber="77">
<emphasis id="B97FF244FFE525373471FA19FC5EFA6A" box="[862,1016,1507,1533]" italics="true" pageId="4" pageNumber="77">
<taxonomicName id="4C0B55D5FFE525373471FA19FC40FA6A" box="[862,998,1507,1533]" class="Mammalia" family="Amphicyonidae" genus="Morphotype" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="77" phylum="Chordata" rank="genus">Morphotype</taxonomicName>
2
</emphasis>
.—MCF-PVPH-739-2, 743, 879-9, 880-1, 880-2, 880-3, and 880-4 (
<figureCitation id="133032D3FFE525373308F9FEFBC1F988" box="[1063,1127,1540,1567]" captionStart="Fig" captionStartId="5.[98,131,1822,1844]" captionTargetBox="[101,767,1126,1795]" captionTargetId="figure-525@5.[99,770,1123,1796]" captionTargetPageId="5" captionText="Fig. 5. Isolated tooth (Morphotype 2) from the Cerro de los Leones locality,Albian, Lower Cretaceous, Picún Leufú,Argentina.MCF-PVPH-880-1 in labial (A1, A2), lingual (A3, A4), distal (A5, A6), and mesial (A7, A8) views. The basal cross section (A9,A10) is visible due to the natural breaking of the crown." figureDoi="http://doi.org/10.5281/zenodo.12194870" httpUri="https://zenodo.org/record/12194870/files/figure.png" pageId="4" pageNumber="77">Fig. 5</figureCitation>
). These tooth crowns are slightly lingually recurved and almost straight in mesial view. The basal cross-section is elliptical, with a labiolingual compression. The dentine surface is almost smooth. MCF-PVPH-739-2 is incomplete, lacking part of the basal and apical portion, whereas small and smooth patches of enamel are preserved on the labial surface of the crown. Conversely, the surface of MCF-PVPH-743 is slightly wrinkled on the lingual side of the crown base, whereas in MCF-PVPH-880-1 it has several longitudinal grooves that fade apically (
<figureCitation id="133032D3FFE4253633C5FF04FA8BFE8E" box="[1258,1325,254,281]" captionStart="Fig" captionStartId="5.[98,131,1822,1844]" captionTargetBox="[101,767,1126,1795]" captionTargetId="figure-525@5.[99,770,1123,1796]" captionTargetPageId="5" captionText="Fig. 5. Isolated tooth (Morphotype 2) from the Cerro de los Leones locality,Albian, Lower Cretaceous, Picún Leufú,Argentina.MCF-PVPH-880-1 in labial (A1, A2), lingual (A3, A4), distal (A5, A6), and mesial (A7, A8) views. The basal cross section (A9,A10) is visible due to the natural breaking of the crown." figureDoi="http://doi.org/10.5281/zenodo.12194870" httpUri="https://zenodo.org/record/12194870/files/figure.png" pageId="5" pageNumber="78">Fig. 5</figureCitation>
). The crowns show different wear facets, of which the apical is rounded and wide, whereas the mesial and distal ones are narrower and apicobasally extended. All the specimens, except MCFPVPH-879-9, have both distal and mesial carinae, without denticles. MCF-PVPH-879-9 is incomplete, lacking part of the basal and apical portions. Moreover, it seems to have only one distal carina; however, taking into account that the crown is heavily weathered, without most of the dentine and the enamel surface, especially on the mesial surface, the loss of the mesial carina due to diagenetic process should be taken into consideration.
</paragraph>
<caption id="DF747EDEFFE5253737B7F927FABEF964" box="[152,1304,1757,1779]" pageId="4" pageNumber="77" startId="4.[152,205,1757,1779]" targetBox="[162,1523,1804,1949]" targetIsTable="true" targetPageId="4" targetType="table">
<paragraph id="8BB42E56FFE5253737B7F927FABEF964" blockId="4.[152,1304,1757,1779]" box="[152,1304,1757,1779]" pageId="4" pageNumber="77">Table 1. Most inclusive morphological characterization of the Cerro de Los Leones teeth, divided in three morphotypes.</paragraph>
</caption>
<paragraph id="8BB42E56FFE52537364AF8E0FA55F80A" pageId="4" pageNumber="77">
<table id="F90BDCF6FFE5DACC378DF8F6FA55F80A" box="[162,1523,1804,1949]" gridcols="5" gridrows="4" pageId="4" pageNumber="77">
<tr id="353B2C14FFE5DACC378DF8F6FA55F8A9" box="[162,1523,1804,1854]" gridrow="0" pageId="4" pageNumber="77" rowspan-0="1">
<th id="76EA4568FFE5DACC366BF8F6FD8EF8A9" box="[324,552,1804,1854]" gridcol="1" gridrow="0" pageId="4" pageNumber="77">Crown curvature</th>
<th id="76EA4568FFE5DACC3515F8F6FD1BF8A9" box="[570,701,1804,1854]" gridcol="2" gridrow="0" pageId="4" pageNumber="77">Cross-section shape</th>
<th id="76EA4568FFE5DACC35FEF8F6FCBAF8A9" box="[721,796,1804,1854]" gridcol="3" gridrow="0" pageId="4" pageNumber="77">Carinae</th>
<th id="76EA4568FFE5DACC3417F8F6FA55F8A9" box="[824,1523,1804,1854]" gridcol="4" gridrow="0" pageId="4" pageNumber="77">Teeth</th>
</tr>
<tr id="353B2C14FFE5DACC378DF8B2FA55F8C9" box="[162,1523,1864,1886]" gridrow="1" pageId="4" pageNumber="77">
<th id="76EA4568FFE5DACC378DF8B2FE8DF8C9" box="[162,299,1864,1886]" gridcol="0" gridrow="1" pageId="4" pageNumber="77">
<taxonomicName id="4C0B55D5FFE52537378DF8B2FEBFF8C9" box="[162,281,1864,1886]" class="Mammalia" family="Amphicyonidae" genus="Morphotype" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="79" phylum="Chordata" rank="genus">Morphotype</taxonomicName>
1
</th>
<td id="76EA4568FFE5DACC366BF8B2FD8EF8C9" box="[324,552,1864,1886]" gridcol="1" gridrow="1" pageId="4" pageNumber="77">more lingual than distal</td>
<td id="76EA4568FFE5DACC3515F8B2FD1BF8C9" box="[570,701,1864,1886]" gridcol="2" gridrow="1" pageId="4" pageNumber="77">subcircular</td>
<td id="76EA4568FFE5DACC35FEF8B2FCBAF8C9" box="[721,796,1864,1886]" gridcol="3" gridrow="1" pageId="4" pageNumber="77">absent</td>
<td id="76EA4568FFE5DACC3417F8B2FA55F8C9" box="[824,1523,1864,1886]" gridcol="4" gridrow="1" pageId="4" pageNumber="77">MCF-PVPH-879-4, 879-6, 879-8, 879-10</td>
</tr>
<tr id="353B2C14FFE5DACC378DF892FA55F8E9" box="[162,1523,1896,1918]" gridrow="2" pageId="4" pageNumber="77">
<th id="76EA4568FFE5DACC378DF892FE8DF8E9" box="[162,299,1896,1918]" gridcol="0" gridrow="2" pageId="4" pageNumber="77">
<taxonomicName id="4C0B55D5FFE52537378DF892FEBFF8E9" box="[162,281,1896,1918]" class="Mammalia" family="Amphicyonidae" genus="Morphotype" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="79" phylum="Chordata" rank="genus">Morphotype</taxonomicName>
2
</th>
<td id="76EA4568FFE5DACC366BF892FD8EF8E9" box="[324,552,1896,1918]" gridcol="1" gridrow="2" pageId="4" pageNumber="77">more lingual than distal</td>
<td id="76EA4568FFE5DACC3515F892FD1BF8E9" box="[570,701,1896,1918]" gridcol="2" gridrow="2" pageId="4" pageNumber="77">elliptical</td>
<td id="76EA4568FFE5DACC35FEF892FCBAF8E9" box="[721,796,1896,1918]" gridcol="3" gridrow="2" pageId="4" pageNumber="77">present</td>
<td id="76EA4568FFE5DACC3417F892FA55F8E9" box="[824,1523,1896,1918]" gridcol="4" gridrow="2" pageId="4" pageNumber="77">MCF-PVPH-739-2, 743, 879-9, 880-1, 880-2, 880-3, 880-4</td>
</tr>
<tr id="353B2C14FFE5DACC378DF87DFA55F80A" box="[162,1523,1927,1949]" gridrow="3" pageId="4" pageNumber="77">
<th id="76EA4568FFE5DACC378DF87DFE8DF80A" box="[162,299,1927,1949]" gridcol="0" gridrow="3" pageId="4" pageNumber="77">
<taxonomicName id="4C0B55D5FFE52537378DF87DFEBFF80A" box="[162,281,1927,1949]" class="Mammalia" family="Amphicyonidae" genus="Morphotype" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="79" phylum="Chordata" rank="genus">Morphotype</taxonomicName>
3
</th>
<td id="76EA4568FFE5DACC366BF87DFD8EF80A" box="[324,552,1927,1949]" gridcol="1" gridrow="3" pageId="4" pageNumber="77">more distal than lingual</td>
<td id="76EA4568FFE5DACC3515F87DFD1BF80A" box="[570,701,1927,1949]" gridcol="2" gridrow="3" pageId="4" pageNumber="77">oval</td>
<td id="76EA4568FFE5DACC35FEF87DFCBAF80A" box="[721,796,1927,1949]" gridcol="3" gridrow="3" pageId="4" pageNumber="77">absent</td>
<td id="76EA4568FFE5DACC3417F87DFA55F80A" box="[824,1523,1927,1949]" gridcol="4" gridrow="3" pageId="4" pageNumber="77">MCF-PVPH-741, 875, 879-1, 879-2, 879-3, 879-5, 978-7, 879-11, 880-5</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB42E56FFE425363407FD71FBA3FB70" blockId="5.[808,1481,650,1352]" pageId="5" pageNumber="78">
<emphasis id="B97FF244FFE425363407FD71FC6EFD32" box="[808,968,651,677]" italics="true" pageId="5" pageNumber="78">
<taxonomicName id="4C0B55D5FFE425363407FD71FC15FD32" box="[808,947,651,677]" class="Mammalia" family="Amphicyonidae" genus="Morphotype" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="78" phylum="Chordata" rank="genus">Morphotype</taxonomicName>
3
</emphasis>
.—MCF-PVPH-741, 875, 879-1, 879-2, 879-3, 879-5, 879-7, 879-11, and 880-5 (
<figureCitation id="133032D3FFE4253633BAFD51FB7EFD52" box="[1173,1240,683,709]" captionStart="Fig" captionStartId="6.[151,184,914,936]" captionTargetBox="[154,821,163,887]" captionTargetId="figure-779@6.[153,821,173,888]" captionTargetPageId="6" captionText="Fig. 6. Isolated tooth (Morphotype 3) from the Cretaceous Cerro de los Leones locality, Picún Leufú, Argentina. MCF-PVPH-879-2 in labial (A1, A2), lingual (A3, A4), distal (A5, A6), and mesial (A7, A8) views. The basal cross section (A9, A10) is visible due to the natural breaking of the crown." figureDoi="http://doi.org/10.5281/zenodo.12194872" httpUri="https://zenodo.org/record/12194872/files/figure.png" pageId="5" pageNumber="78">Fig. 6</figureCitation>
). These tooth crowns are more recurved distally than lingually, with oval, labiolingually compressed cross-sections, and smooth dentine surfaces. The apical wear facet is rounded, worn at the apex from multiple directions, especially mesially and distally. All the specimens lack carinae. MCF-PVPH-875 preserves two portions of smooth enamel on the labial and distal surfaces, whereas in MCF-PVPH-879-2 the enamel is well preserved, especially on the basal half of the crown, and wrinkled, showing several apicobasally oriented and anastomosed ridges and grooves, slightly deeper on the lingual surface. A similar condition is observed on the preserved enamel of MCF-PVPH-879-1, 879-5, and 880-5, whereas in MCF-PVPH-879-3 the enamel is apically worn, with apicobasally oriented grooves on the basal half of the crown. The crowns of MCF-PVPH-875, 879-2, and 879-3 feature labial wear on their apexes and mesial and distal wear that visibly rounds their apices.
</paragraph>
<caption id="DF747EDEFFE42536374EFC3DFF64FBDB" ID-DOI="http://doi.org/10.5281/zenodo.12194866" ID-Zenodo-Dep="12194866" httpUri="https://zenodo.org/record/12194866/files/figure.png" pageId="5" pageNumber="78" startId="5.[97,130,967,989]" targetBox="[101,768,166,937]" targetPageId="5" targetType="figure">
<paragraph id="8BB42E56FFE42536374EFC3DFF64FBDB" blockId="5.[97,771,967,1101]" pageId="5" pageNumber="78">
Fig. 4. Isolated tooth (
<taxonomicName id="4C0B55D5FFE425363608FC3DFE3FFC4A" box="[295,409,967,989]" class="Mammalia" family="Amphicyonidae" genus="Morphotype" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="79" phylum="Chordata" rank="genus">Morphotype</taxonomicName>
1) from the Cerro de los Leones locality, Albian, Lower Cretaceous, Picún Leufú, Argentina. MCF-PVPH-879-8 in labial (A
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, A
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), lingual (A
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, A
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), mesial (A
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, A
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), and distal (A
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, A
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) views. The basal cross section (A
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, A
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) is visible due to the natural breaking of the crown.
</paragraph>
</caption>
<paragraph id="8BB42E56FFE425363461FB17FBE3FADF" blockId="5.[808,1481,650,1352]" pageId="5" pageNumber="78">MCF-PVPH-741 shows a wide and smooth groove, slightly deeper basally and apicobasally extended to the apical third of the crown.</paragraph>
</subSubSection>
</treatment>
</document>