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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e8286" ID-PMC="PMC4867701" ID-Pensoft-Pub="1314-2828--8286" ID-PubMed="27226761" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828--e8286" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="Checklist of Serengeti Ecosystem Grasses" checkinTime="1553126254102" checkinUser="pensoft" docAuthor="Williams, Emma Victoria, Elia Ntandu, John, Ficinski, Pawel &amp; Vorontsova, Maria" docDate="2016" docId="A2E727B4AA90C3EBD5FC7F9CD9AD5C9C" docLanguage="en" docName="BiodivDatJour 4: e8286" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e8286" docTitle="Digitaria ternata A. Rich. Stapf" docType="treatment" docVersion="5" lastPageNumber="8286" masterDocId="FFD5FFC2FFE76F0DFFB3FFA2FFE6FFE5" masterDocTitle="Checklist of Serengeti Ecosystem Grasses" masterLastPageNumber="8286" masterPageNumber="8286" pageNumber="8286" updateTime="1668127305304" updateUser="ExternalLinkService">
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<mods:namePart>Ficinski, Pawel</mods:namePart>
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<mods:namePart>Vorontsova, Maria</mods:namePart>
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<taxonomicName authority="A. Rich. Stapf" class="Liliopsida" family="Poaceae" genus="Digitaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Digitaria ternata" order="Poales" pageId="0" pageNumber="8286" phylum="Tracheophyta" rank="species" species="ternata">Digitaria ternata A.Rich.Stapf</taxonomicName>
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<paragraph pageId="0" pageNumber="8286">Materials</paragraph>
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Type status:
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; recordNumber: 367; recordedBy:
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; Taxon: scientificName: Digitariaternata A.Rich.Stapf; kingdom: Plantae; family: Poaceae; genus: Digitaria; specificEpithet: ternata; scientificNameAuthorship: A.Rich.Stapf; Location: continent: Africa; country:
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; stateProvince: Arusha; county: Ngorongoro; locality:
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; ownerInstitutionCode: AAU; basisOfRecord: PreservedSpecimen
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<materialsCitation collectingDate="1973-03-13" collectionCode="NHT, Herbarium" collectorName="Frame, GW" country="Tanzania" latitude="-2.933333" location="Empakaai crater" longitude="35.816666" pageId="0" pageNumber="8286" specimenCode="1141" specimenCount="1" typeStatus="Other material">
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; recordNumber: 87; recordedBy:
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; Taxon: scientificName: Digitariaternata A.Rich.Stapf; kingdom: Plantae; family: Poaceae; genus: Digitaria; specificEpithet: ternata; scientificNameAuthorship: A.Rich.Stapf; Location: continent: Africa; country:
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; stateProvince: Arusha; county: Ngorongoro; locality:
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; verbatimLocality: Ngorongoro conservation area, Empakaai crater, top of east rim.; minimumElevationInMeters: 2400; decimalLatitude:
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; collectionCode:
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; ownerInstitutionCode: NHT; basisOfRecord: PreservedSpecimen
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<materialsCitation collectingDate="1988-04-18" collectionCode="NHT, Herbarium" collectorName="Chuwa, S" country="Tanzania" latitude="-3.183333" location="Endulen" longitude="35.15" pageId="0" pageNumber="8286" specimenCode="1142" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: catalogNumber:
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; recordNumber: 2637; recordedBy:
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; Taxon: scientificName: Digitariaternata A.Rich.Stapf; kingdom: Plantae; family: Poaceae; genus: Digitaria; specificEpithet: ternata; scientificNameAuthorship: A.Rich.Stapf; Location: continent: Africa; country:
<collectingCountry pageId="0" pageNumber="8286">Tanzania</collectingCountry>
; stateProvince: Arusha; county: Ngorongoro; locality:
<location LSID="urn:lsid:plazi:treatment:A2E727B4AA90C3EBD5FC7F9CD9AD5C9C:87E87A17DF832E6208DAB37C377F172C" country="Tanzania" latitude="-3.183333" longitude="35.15" name="Endulen" pageId="0" pageNumber="8286">Endulen</location>
; verbatimLocality: Ngorongoro conservation area.; minimumElevationInMeters: 1524; decimalLatitude:
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; decimalLongitude:
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; Event: eventDate:
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; collectionCode:
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<paragraph pageId="0" pageNumber="8286">Widespread</paragraph>
</subSubSection>
</treatment>
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.133.37773" ID-GBIF-Dataset="a6caa96b-fb3a-4877-9e16-b2f519188af6" ID-PMC="PMC6791900" ID-Pensoft-Pub="1314-2003-133-1" ID-Pensoft-UUID="99F75C9803D65C4880141BE7B6589FB5" ID-PubMed="31632179" ModsDocID="1314-2003-133-1" checkinTime="1570497209971" checkinUser="pensoft" docAuthor="Shalimov, Aleksandr Petrovich, Wu, Yu-Dong &amp; Zhang, Xian-Chun" docDate="2019" docId="A2E72B26C12558B7B2D473386870EC9B" docLanguage="en" docName="PhytoKeys 133: 1-76" docOrigin="PhytoKeys 133" docSource="http://dx.doi.org/10.3897/phytokeys.133.37773" docTitle="Selaginella aitchisonii Hieron." docType="treatment" docVersion="6" id="99F75C9803D65C4880141BE7B6589FB5" lastPageNumber="1" masterDocId="99F75C9803D65C4880141BE7B6589FB5" masterDocTitle="A taxonomic revision of the genus Selaginella (Selaginellaceae) from Nepal" masterLastPageNumber="76" masterPageNumber="1" pageNumber="1" updateTime="1668139527481" updateUser="ExternalLinkService">
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<treatment ID-GBIF-Taxon="159654788" LSID="urn:lsid:plazi:treatment:A2E72B26C12558B7B2D473386870EC9B" httpUri="http://treatment.plazi.org/id/A2E72B26C12558B7B2D473386870EC9B" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="A2E72B26-C125-58B7-B2D4-73386870EC9B" authority="Hieron." class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Selaginella aitchisonii" order="Selaginellales" pageId="0" pageNumber="1" phylum="Pteridophyta" rank="species" species="aitchisonii">Selaginella aitchisonii Hieron.</taxonomicName>
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<taxonomicName class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Selaginella aitchisonii" order="Selaginellales" pageId="0" pageNumber="1" phylum="Pteridophyta" rank="species" species="aitchisonii">Selaginella aitchisonii</taxonomicName>
Hieron., Nat. Pflanzenfam. 1(4): 674. 1902;
<bibRefCitation author="Dixit, RD" journalOrPublisher="Dehra Dun, Bishen Singh Mahendra Pal Singh" pageId="0" pageNumber="1" refId="B4" refString="Dixit, RD, 1992. Selaginellaceae of India. Dehra Dun, Bishen Singh Mahendra Pal Singh" title="Selaginellaceae of India." year="1992">Dixit 1992</bibRefCitation>
;
<bibRefCitation author="Thapa, N" journalOrPublisher="Department Plant Resources" pageId="0" pageNumber="1" pagination="1 - 175" refId="B27" refString="Thapa, N, 2002. Pteridophytes of Nepal. Bull. Department Plant Resources 19: 1 - 175" title="Pteridophytes of Nepal. Bull." volume="19" year="2002">Thapa 2002</bibRefCitation>
;
<bibRefCitation author="Fraser-Jenkins, CR" journalOrPublisher="National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" pageId="0" pageNumber="1" refId="B7" refString="Fraser-Jenkins, CR, Kandel, DR, Pariyar, S, 2015. Ferns and fern-allies of Nepal. Vol. 1. National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" title="Ferns and fern-allies of Nepal. Vol. 1." year="2015">Fraser-Jenkins et al. 2015</bibRefCitation>
;
<bibRefCitation author="Fraser-Jenkins, CR" journalOrPublisher="National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" pageId="0" pageNumber="1" refId="B8" refString="Fraser-Jenkins, CR, Gandhi, KN, Kholia, BS, Benniamin, A, 2017. An annotated checklist of Indian Pteridophytes, Part - I (Lycopodiaceae to Thelypteridace). Bishen Singh Mahendra Pal Singh Dehra Dun, 562 pp." title="An annotated checklist of Indian Pteridophytes, Part - I (Lycopodiaceae to Thelypteridace). Bishen Singh Mahendra Pal Singh Dehra Dun, 562 pp." year="2017">Fraser-Jenkins et al. 2017</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
≡
<taxonomicName authorityName="Alston" authorityYear="1945" baseAuthorityName="Hieron." class="Lycopodiopsida" family="Selaginellaceae" form="aitchisonii" genus="Selaginella" higherTaxonomySource="treatment-meta" infraspecific-rank="f." kingdom="Plantae" lsidName="Selaginella sanguinolenta" order="Selaginellales" pageId="0" pageNumber="1" phylum="Pteridophyta" rank="form" species="sanguinolenta">Selaginella sanguinolenta f. aitchisonii</taxonomicName>
(Hieron.) Alston, Proc. Nat. Inst. Sci. India 11(3): 215. 1945.
<emphasis bold="true" pageId="0" pageNumber="1">Type.</emphasis>
PAKISTAN. Kurram Valley, Shend Toi,
<emphasis italics="true" pageId="0" pageNumber="1">J.E.T. Aitchison 369</emphasis>
(syntype: B [20 0121871]; K, CAL); KYRGYZSTAN. Turkestan, Akburtaseh,
<emphasis italics="true" pageId="0" pageNumber="1">A. Regel 1878</emphasis>
(syntype: B [20 0121870]); KYRGYZSTAN. Turkestan, Musart-Thal.,
<emphasis italics="true" pageId="0" pageNumber="1">A. Regel 1877</emphasis>
(syntype: B [20 0121869]).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Stems 10-25 cm, erect, slender. Rhizophores restricted to the basal part of stems, located on ventral side. Main stems branches from near bases, lateral branches dichotomously compound. Main stems terete, reddish, in basal part main stem 1.0-1.1 mm in diam. Axillary leaves symmetrical, ovate-oblong, carinate, at base obtuse, margin hyaline, at base denticulate, in middle and upper part entire, apex acuminate. Leaves isomorphic, slightly asymmetrical, ovate-lanceolate, uniauriculate at base, obtuse, peltate, margin hyaline, denticulate at base, entire in middle and upper part, apex mucronate to acuminate. Strobili solitary, terminal, compact, tetrahedral, 6.0-10.0
<normalizedToken originalValue="×">x</normalizedToken>
1.0-2.0 mm, slightly wider branches. Sporophylls monomorphic, ovate, truncate at base, strongly keeled, apex mucronate to acute. Megaspores yellow, irregular verrucate. Microspores deep yellow, surface rugulose-tuberculate, with perispore on surface.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">On mossy rocks. Alt. 2200-3400 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution in Nepal.</paragraph>
<paragraph pageId="0" pageNumber="1">W.</paragraph>
<paragraph pageId="0" pageNumber="1">
Nepalese threatened status: VU, globally threatened (
<bibRefCitation author="Fraser-Jenkins, CR" journalOrPublisher="National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" pageId="0" pageNumber="1" refId="B7" refString="Fraser-Jenkins, CR, Kandel, DR, Pariyar, S, 2015. Ferns and fern-allies of Nepal. Vol. 1. National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" title="Ferns and fern-allies of Nepal. Vol. 1." year="2015">Fraser-Jenkins et al. 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">General distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">AFGHANISTAN, INDIA (Jammu and Kashmir), KYRGYZSTAN, PAKISTAN.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="chromosome number">
<paragraph pageId="0" pageNumber="1">Chromosome number.</paragraph>
<paragraph pageId="0" pageNumber="1">not available data.</paragraph>
<paragraph pageId="0" pageNumber="1">Selected specimens examined:</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">W Nepal</emphasis>
:
<emphasis bold="true" pageId="0" pageNumber="1">HUMLA</emphasis>
: &quot;Phal Ko Odar to Pipling, on mossy stone. 2600 m, 6 Jun 1980,
<emphasis italics="true" pageId="0" pageNumber="1">P.R. Shakya &amp; B. Roy 5514</emphasis>
&quot; (KATH).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="notes">
<paragraph pageId="0" pageNumber="1">Note.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" lsidName="Selaginella aitchisonii" order="Selaginellales" pageId="0" pageNumber="1" phylum="Pteridophyta" rank="species" species="aitchisonii">
<emphasis italics="true" pageId="0" pageNumber="1">Selaginella aitchisonii</emphasis>
</taxonomicName>
is morphologically closely related to the widespread
<taxonomicName family="Selaginellaceae" kingdom="Plantae" lsidName="S. sanguinolenta" order="Selaginellales" pageId="0" pageNumber="1" rank="species" species="sanguinolenta">
<emphasis italics="true" pageId="0" pageNumber="1">S. sanguinolenta</emphasis>
</taxonomicName>
, a species complex consists of several morphological variable forms which might be recognised as distinct species pending our molecular phylogentic analysis (data not published).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart>Sun, Yue</mods:namePart>
<mods:affiliation>School of wetland, Jiangsu Provincial Key Laboratory of Coastal Wetland Bioresources and Environmental Protection, Jiangsu Key Laboratory for Bioresources of Saline Soils, Yancheng Teachers University, Yancheng 224051, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yu, Lei</mods:namePart>
<mods:affiliation>School of wetland, Jiangsu Provincial Key Laboratory of Coastal Wetland Bioresources and Environmental Protection, Jiangsu Key Laboratory for Bioresources of Saline Soils, Yancheng Teachers University, Yancheng 224051, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Xinzheng</mods:namePart>
<mods:affiliation>Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China &amp; Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, China &amp; University of Chinese Academy of Sciences, Beijing 200049, China &amp; Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao, China</mods:affiliation>
<mods:nameIdentifier type="email">lixzh@qdio.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-11-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1068</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>11</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1068.71925</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1068.71925</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1068-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">61BF36B3B9DB5C948ACEDEEA0A905DE2</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="190303301" LSID="urn:lsid:zoobank.org:act:AAAAD262-9B46-4B96-BD38-F6FBF8F7D5A3" httpUri="http://treatment.plazi.org/id/A2E7DB986B785109BE12E30E45C24BEC" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/AAAAD262-9B46-4B96-BD38-F6FBF8F7D5A3" authority="Sun &amp; Yu &amp; Li, 2021" authorityName="Sun &amp; Yu &amp; Li" authorityYear="2021" class="Polychaeta" family="Orbiniidae" genus="Phylo" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Phylo heterochaetus" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="heterochaetus" status="sp. nov.">Phylo heterochaetus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Anterior end of Phylo heterochaetus sp. nov. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1068.71925.figure1" httpUri="https://binary.pensoft.net/fig/605877" pageId="0" pageNumber="1">Figures 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
: MBM286984: nYS, Sta. 3875-04,
<geoCoordinate degrees="38" direction="north" minutes="45" orientation="latitude" precision="925" value="38.75">38°45'N</geoCoordinate>
,
<geoCoordinate degrees="123" direction="east" minutes="30" orientation="longitude" precision="925" value="123.5">123°30'E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="6.4" unit="m" value="64.0">64 m</quantity>
,
<date value="2015-12-16">16 Dec. 2015</date>
.
<materialsCitation collectingDate="2016-12-18" elevation="31" latitude="39.116665" location="Sta." longLatPrecision="1171" longitude="122.9" specimenCount="1" typeStatus="Paratype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratype</typeStatus>
</emphasis>
: MBM286985: nYS,
<location LSID="urn:lsid:plazi:treatment:A2E7DB986B785109BE12E30E45C24BEC:C1C153E19ABE48B24AA7214C1384CFFC" latitude="39.116665" longLatPrecision="1171" longitude="122.9" name="Sta.">Sta.</location>
16,
<geoCoordinate degrees="39" direction="north" minutes="07" orientation="latitude" precision="925" value="39.116665">39°07'N</geoCoordinate>
,
<geoCoordinate degrees="122" direction="east" minutes="54" orientation="longitude" precision="925" value="122.9">122°54'E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.1" unit="m" value="31.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="3.1" unit="m" value="31.0">31 m</elevation>
</quantity>
,
<collectingDate value="2016-12-18">18 Dec. 2016</collectingDate>
</materialsCitation>
.
<materialsCitation collectingDate="2016-12-18" latitude="39.0" location="Sta." longLatPrecision="1172" longitude="122.916664" specimenCount="1" typeStatus="paratype">
MBM286986: nYS,
<location LSID="urn:lsid:plazi:treatment:A2E7DB986B785109BE12E30E45C24BEC:AC0BE992F8B3BCDED703C2DF31A786AF" latitude="39.0" longLatPrecision="1172" longitude="122.916664" name="Sta.">Sta.</location>
C3,
<geoCoordinate degrees="39" direction="north" minutes="00" orientation="latitude" precision="925" value="39.0">39°00'N</geoCoordinate>
,
<geoCoordinate degrees="122" direction="east" minutes="55" orientation="longitude" precision="925" value="122.916664">122°55'E</geoCoordinate>
, 45 meters,
<collectingDate value="2016-12-18">18 Dec. 2016</collectingDate>
</materialsCitation>
.
<materialsCitation collectingDate="2016-12-18" collectorName="Sta. A" elevation="36" latitude="39.083332" location="Sta." longLatPrecision="1171" longitude="122.583336" specimenCount="1" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:A2E7DB986B785109BE12E30E45C24BEC:1CA326D6F81BEFFA52EB3683D3022C45" latitude="39.083332" longLatPrecision="1171" longitude="122.583336" name="Sta.">Sta.</location>
MBM286987: nYS,
<collectorName>Sta. A</collectorName>
1,
<geoCoordinate degrees="39" direction="north" minutes="05" orientation="latitude" precision="925" value="39.083332">39°05'N</geoCoordinate>
,
<geoCoordinate degrees="122" direction="east" minutes="35" orientation="longitude" precision="925" value="122.583336">122°35'E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.6" unit="m" value="36.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="3.6" unit="m" value="36.0">36 m</elevation>
</quantity>
,
<collectingDate value="2016-12-18">18 Dec. 2016</collectingDate>
</materialsCitation>
.
<materialsCitation collectingDate="1959-07-11" collectorName="Mu Chen." elevation="51" latitude="122.75" location="Sta." longLatPrecision="1051" longitude="38.15" specimenCount="1" typeStatus="paratype">
MBM023255: nYS,
<location LSID="urn:lsid:plazi:treatment:A2E7DB986B785109BE12E30E45C24BEC:3EB5BB3ACF4A35F948837A5A6FDB5C7B" latitude="122.75" longLatPrecision="1051" longitude="38.15" name="Sta.">Sta.</location>
2049,
<geoCoordinate degrees="122" direction="north" minutes="45" orientation="latitude" precision="925" value="122.75">122°45'N</geoCoordinate>
,
<geoCoordinate degrees="38" direction="east" minutes="09" orientation="longitude" precision="925" value="38.15">38°09'E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="5.1" unit="m" value="51.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="5.1" unit="m" value="51.0">51 m</elevation>
</quantity>
, coll.
<collectorName>Mu Chen.</collectorName>
<collectingDate value="1959-07-11">11 Jul. 1959</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1959-04-15" collectorName="Mu Chen." elevation="63" latitude="38.5" location="Sta." longLatPrecision="1175" longitude="123.5" specimenCount="1">
MBM023221: nYS,
<location LSID="urn:lsid:plazi:treatment:A2E7DB986B785109BE12E30E45C24BEC:06059770AB2525235C10EDA99668377E" latitude="38.5" longLatPrecision="1175" longitude="123.5" name="Sta.">Sta.</location>
2056,
<geoCoordinate degrees="38" direction="north" minutes="30" orientation="latitude" precision="925" value="38.5">38°30'N</geoCoordinate>
,
<geoCoordinate degrees="123" direction="east" minutes="30" orientation="longitude" precision="925" value="123.5">123°30'E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="6.3" unit="m" value="63.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="6.3" unit="m" value="63.0">63 m</elevation>
</quantity>
,
<collectorName>Mu Chen.</collectorName>
<collectingDate value="1959-04-15">15 Apr. 1959</collectingDate>
</materialsCitation>
.
<materialsCitation collectingDate="1958-10-01" collectorName="Yuheng Cui." latitude="38.0" location="Sta." longLatPrecision="1178" longitude="123.5" specimenCount="1">
MBM009975: nYS,
<location LSID="urn:lsid:plazi:treatment:A2E7DB986B785109BE12E30E45C24BEC:723C9D5FEF4E1F9DEFE46EE7F87553B5" latitude="38.0" longLatPrecision="1178" longitude="123.5" name="Sta.">Sta.</location>
2057,
<geoCoordinate degrees="38" direction="north" minutes="00" orientation="latitude" precision="925" value="38.0">38°00'N</geoCoordinate>
,
<geoCoordinate degrees="123" direction="east" minutes="30" orientation="longitude" precision="925" value="123.5">123°30'E</geoCoordinate>
, 70.5 m, coll.
<collectorName>Yuheng Cui.</collectorName>
<collectingDate value="1958-10-01">1 Oct. 1958</collectingDate>
</materialsCitation>
.
<materialsCitation collectingDate="2016-12-18" elevation="35" latitude="39.0" location="Sta." longLatPrecision="1172" longitude="122.583336" specimenCount="1">
YCTU000004: nYS,
<location LSID="urn:lsid:plazi:treatment:A2E7DB986B785109BE12E30E45C24BEC:46694D91BEFF0D5C389395E41A2AF3E6" latitude="39.0" longLatPrecision="1172" longitude="122.583336" name="Sta.">Sta.</location>
A3,
<geoCoordinate degrees="39" direction="north" minutes="00" orientation="latitude" precision="925" value="39.0">39°00'N</geoCoordinate>
,
<geoCoordinate degrees="122" direction="east" minutes="35" orientation="longitude" precision="925" value="122.583336">122°35'E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.5" unit="m" value="35.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="3.5" unit="m" value="35.0">35 m</elevation>
</quantity>
,
<collectingDate value="2016-12-18">18 Dec. 2016</collectingDate>
</materialsCitation>
.
<materialsCitation collectingDate="2016-12-18" elevation="37" latitude="39.266666" location="Sta." longLatPrecision="1170" longitude="123.3" specimenCount="1">
YCTU000005: nYS,
<location LSID="urn:lsid:plazi:treatment:A2E7DB986B785109BE12E30E45C24BEC:070F1DB1B6808E1383C4BB1166A39D16" latitude="39.266666" longLatPrecision="1170" longitude="123.3" name="Sta.">Sta.</location>
48,
<geoCoordinate degrees="39" direction="north" minutes="16" orientation="latitude" precision="925" value="39.266666">39°16'N</geoCoordinate>
,
<geoCoordinate degrees="123" direction="east" minutes="18" orientation="longitude" precision="925" value="123.3">123°18'E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.7" unit="m" value="37.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="3.7" unit="m" value="37.0">37 m</elevation>
</quantity>
,
<collectingDate value="2016-12-18">18 Dec. 2016</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">All specimens incomplete, posterior end missing, holotype with 60 chaetigers, 34 mm long and 5 mm wide. Body elongate, about same width throughout; thorax depressed, abdomen cylindrical.</paragraph>
<paragraph pageId="0" pageNumber="1">
Prostomium short, conical, tapering to rounded tip; eyepots absent, nuchal organs narrow groove at junction with peristomium (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Anterior end of Phylo heterochaetus sp. nov. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1068.71925.figure1" httpUri="https://binary.pensoft.net/fig/605877" pageId="0" pageNumber="1">1</figureCitation>
). Peristomium a narrow asetigerous segment, distinctly separated from prostomium and chaetiger 1; mouth with anterior oral lip arising from posterior margin of peristomium, posterior oral lip from anterior margin of chaetiger 2. Holotype with proboscis everted, consisting of two large, inflated lobes (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Anterior end of Phylo heterochaetus sp. nov. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1068.71925.figure1" httpUri="https://binary.pensoft.net/fig/605877" pageId="0" pageNumber="1">1</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1068.71925.figure1" httpUri="https://binary.pensoft.net/fig/605877" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
Anterior end of
<taxonomicName authorityName="Sun &amp; Yu &amp; Li" authorityYear="2021" class="Polychaeta" family="Orbiniidae" genus="Phylo" kingdom="Animalia" lsidName="Phylo heterochaetus" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="heterochaetus">
<emphasis italics="true" pageId="0" pageNumber="1">Phylo heterochaetus</emphasis>
</taxonomicName>
sp. nov. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
Holotype with 21 thoracic chaetigers, divided into anterior and posterior sections: anterior thorax with 13 chaetigers (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Anterior end of Phylo heterochaetus sp. nov. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1068.71925.figure1" httpUri="https://binary.pensoft.net/fig/605877" pageId="0" pageNumber="1">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2A</figureCitation>
), posterior thorax with modified spines beginning on chaetiger 14 continuing over 8 chaetigers. Intersegmental annulations first present from chaetiger 1, well developed (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Anterior end of Phylo heterochaetus sp. nov. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1068.71925.figure1" httpUri="https://binary.pensoft.net/fig/605877" pageId="0" pageNumber="1">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2A</figureCitation>
). Thoracic notopodial postchaetal lobe arising from narrow base, broadly triangular, narrowing apically (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2B</figureCitation>
), becoming long and narrow in abdominal chaetigers (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2C, D</figureCitation>
). Thoracic neuropodia thick, elongate, fringed, bearing rows of uncini and several capillaries. First chaetiger with 2 rounded postchaetal lobe, increasing to 13-14 on middle thoracic chaetigers (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2A</figureCitation>
), then decreasing to 6 or 7 on last thoracic chaetigers. Subpodial lobes or stomach papillae from chaetiger 12, numbering 1 at first, then extending to ventral midline on chaetigers 18-25, then abruptly absent (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Anterior end of Phylo heterochaetus sp. nov. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1068.71925.figure1" httpUri="https://binary.pensoft.net/fig/605877" pageId="0" pageNumber="1">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2A</figureCitation>
). Abdominal neuropodia bilobed, with inner lobe blunt, outer lobe cirriform (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2C, D</figureCitation>
). First abdominal chaetiger with 6 extra lobes or ventral cirri ventral to neuropodium, then reduced to 2 on following 3 chaetigers and 1 or 2 on subsequent chaetigers (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2C, D</figureCitation>
). Interramal cirrus absent.
</paragraph>
<caption doi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName authorityName="Sun &amp; Yu &amp; Li" authorityYear="2021" class="Polychaeta" family="Orbiniidae" genus="Phylo" kingdom="Animalia" lsidName="Phylo heterochaetus" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="heterochaetus">
<emphasis italics="true" pageId="0" pageNumber="1">Phylo heterochaetus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
anterior body in lateral view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
parapodium of chaetiger 7
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
parapodium of anterior abdominal chaetiger
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
parapodium of chaetiger 62
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
); 400
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">B-D</emphasis>
); 200
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
Branchiae from chaetiger 5, on anterior chaetigers short, triangular, tapering to narrow papillate rounded apex (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2B</figureCitation>
), then becoming narrower and longer, leaf-like and ciliated (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2C, D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Thoracic notopodia with fascicles of crenulated capillaries. Abdominal notopodia with few long, thin capillaries, 2-4 furcate chaetae and 3-5 imbedded aciculae, furcate chaetae with unequal tynes, each tyne with fine needles directed medially, shaft with transverse rows of barbs (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Anterior thoracic neuropodia with 4 rows and 1 short posterior rows of yellow uncini, and posterior row of crenulate capillaries (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2E</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3A, B, E, F</figureCitation>
). Anterior 2 rows mixed with 10-12 subuluncini, last row with 7-9 uncini, curved ventrally to anterior rows (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3A, B</figureCitation>
). Uncini of first 1-4 rows short, with blunt-tipped apex, with 10-18 transverse rows of blunt barbs on convex side, with conspicuous groove apically (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3B, D</figureCitation>
); uncini of last row longer than anterior ones, only located in ventral side of neuropodium (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3A</figureCitation>
); subuluncini resemble uncini, but with distally pointed tip, with more than 30 rows of blunt barbs, surrounded by sheath (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3B</figureCitation>
). Posterior thoracic neuropodia with 4-7 modified spines (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3C</figureCitation>
), single short row of uncini and 1 or 2 rows crenulate capillaries. Modified spines large, hastate, superior one projecting from neuropodium, with glandular pouch (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3C</figureCitation>
). Abdominal neurochaetae including 4-6 thin capillaries and single acicula, thin and imbedded.
</paragraph>
<caption doi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
SEM pictures of
<taxonomicName authorityName="Sun &amp; Yu &amp; Li" authorityYear="2021" class="Polychaeta" family="Orbiniidae" genus="Phylo" kingdom="Animalia" lsidName="Phylo heterochaetus" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="heterochaetus">
<emphasis italics="true" pageId="0" pageNumber="1">Phylo heterochaetus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
neurochaetae of thorax
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
uncini and subuluncini
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
modified spines
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
uncini of last rows of thoracic neuropodia
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
furcate chaeta
<emphasis bold="true" pageId="0" pageNumber="1">F, G</emphasis>
crenulate capillaries. Scale bars: 250
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
); 200
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">A, E, F</emphasis>
); 50
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
); 30
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
); 10
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
Anterior thoracic neuropodia with 5 or 6 rows of yellow uncini and posterior row of crenulate capillaries (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylo heterochaetus sp. nov. A anterior body in lateral view B parapodium of chaetiger 7 C parapodium of anterior abdominal chaetiger D parapodium of chaetiger 62 E diagram showing arrangement of anterior thoracic neurochaetae. Scale bars: 2 mm (A); 400 μm (B-D); 200 μm (E)." figureDoi="10.3897/zookeys.1068.71925.figure2" httpUri="https://binary.pensoft.net/fig/605878" pageId="0" pageNumber="1">2E</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3A, B</figureCitation>
). Anterior 2 rows mixed with 10-12 subuluncini, last row with 7-9 uncini, curved ventrally to anterior rows (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3A, B</figureCitation>
). Uncini of anterior 4 or 5 rows short, with blunt-tipped apex, with 10-18 transverse rows of blunt barbs on convex side, with conspicuous groove apically (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3B, D</figureCitation>
); uncini of last row longer than anterior ones, only located in ventral side of neuropodium (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3A</figureCitation>
).
<taxonomicName lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Subuluncini">Subuluncini</taxonomicName>
resemble uncini but with distally pointed tip, with more than 30 rows of blunt barbs, surrounded by sheath (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3B</figureCitation>
). Posterior thoracic neuropodia with 4-7 modified spines (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3C</figureCitation>
), single short row of uncini and 1 or 2 rows crenulate capillaries. Modified spines large, hastate, superior one projecting from neuropodium (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM pictures of Phylo heterochaetus sp. nov. A neurochaetae of thorax B uncini and subuluncini C modified spines D uncini of last rows of thoracic neuropodia E furcate chaeta F, G crenulate capillaries. Scale bars: 250 μm (C); 200 μm (A, E, F); 50 μm (B); 30 μm (G); 10 μm (D)." figureDoi="10.3897/zookeys.1068.71925.figure3" httpUri="https://binary.pensoft.net/fig/605879" pageId="0" pageNumber="1">3C</figureCitation>
). Abdominal neurochaetae including 4-6 thin capillaries and single acicula, thin and imbedded.
</paragraph>
<paragraph pageId="0" pageNumber="1">Pygidium not observed.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">One paratype (MBM286986) with 13 and 14 anterior thoracic chaetigers on left and right side respectively, posterior thorax with 9 and 8 chaetigers, respectively. Holotype and other specimens with 13 anterior thoracic chaetigers, posterior thorax with 4-8.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species is named for the thoracic neuropodia with two kinds of uncini.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">Northern Yellow Sea, China.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is unusual in the genus in having anterior thoracic neuropodia with 5 or 6 rows of uncini intermixed with a few subuluncini in the first 1 or 2 rows.
<taxonomicName authorityName="Sun &amp; Yu &amp; Li" authorityYear="2021" class="Polychaeta" family="Orbiniidae" genus="Phylo" kingdom="Animalia" lsidName="Phylo heterochaetus" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="heterochaetus">
<emphasis italics="true" pageId="0" pageNumber="1">Phylo heterochaetus</emphasis>
</taxonomicName>
sp. nov. is similar with
<taxonomicName lsidName="P. ornatus" pageId="0" pageNumber="1" rank="species" species="ornatus">
<emphasis italics="true" pageId="0" pageNumber="1">P. ornatus</emphasis>
</taxonomicName>
(Verrill, 1873) for: 1) anterior thoracic neuropodia with rows of uncini, 2) lacking interramal cirri, 3) anterior thorax with 13 or 14 chaetigers. They can be easily distinguished by 1) species of
<taxonomicName lsidName="P. heterochaetus" pageId="0" pageNumber="1" rank="species" species="heterochaetus">
<emphasis italics="true" pageId="0" pageNumber="1">P. heterochaetus</emphasis>
</taxonomicName>
sp. nov. with 4-8 posterior thoracic chaetigers, while the latter species with 13 or more chaetigers; 2) species of
<taxonomicName lsidName="P. heterochaetus" pageId="0" pageNumber="1" rank="species" species="heterochaetus">
<emphasis italics="true" pageId="0" pageNumber="1">P. heterochaetus</emphasis>
</taxonomicName>
sp. nov. with 10 or 11 subuluncini intermixed with first 2 rows uncini, while the latter species lacks subuluncini; 3) modified spines hastate in
<taxonomicName lsidName="P. heterochaetus" pageId="0" pageNumber="1" rank="species" species="heterochaetus">
<emphasis italics="true" pageId="0" pageNumber="1">P. heterochaetus</emphasis>
</taxonomicName>
sp. nov. and acicular in
<taxonomicName lsidName="P. ornatus" pageId="0" pageNumber="1" rank="species" species="ornatus">
<emphasis italics="true" pageId="0" pageNumber="1">P. ornatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>