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<document id="D8BA7181D062BCF34936EC77F9F6A0B9" ID-DOI="10.3897/zse.100.120207" ID-ZooBank="55DBF63A-85CF-42C0-8218-15F310FB177A" ID-publisher-id="120207" URI-arpha="123D4283-0C8F-5E2E-9E65-4D0700BDBA20" URI-zoobank="http://zoobank.org/55DBF63A-85CF-42C0-8218-15F310FB177A" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1718641467983" checkinUser="pensoft" docAuthor="Villarreal, Osvaldo, Ahumada-C., Daniela &amp; Navas-S., Gabriel R." docDate="2024" docId="A2A72C0A27B3578D80E5BD65E4D16D5E" docLanguage="en" docName="Zoosystematics and Evolution 100(3): 803-820" docOrigin="Zoosystematics and Evolution 100 (3)" docSource="https://zse.pensoft.net/article/120207/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Eutimesius ephippiatus" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="820" masterDocId="123D42830C8F5E2E9E654D0700BDBA20" masterDocTitle="Exploring the diversity of Eutimesius Roewer, 1913: new species and records from Colombia and Venezuela (Opiliones, Gonyleptoidea, Stygnidae)" masterLastPageNumber="820" masterPageNumber="803" pageNumber="803" updateTime="1718646022642" updateUser="admin">
<mods:mods id="91C94AFA4A3B43B2F194E7D33C4FDBC8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9E2E92536908FF1EBA764B805242743F">
<mods:title id="3D99155EB924A7D694040E69E4375E92">Exploring the diversity of Eutimesius Roewer, 1913: new species and records from Colombia and Venezuela (Opiliones, Gonyleptoidea, Stygnidae)</mods:title>
</mods:titleInfo>
<mods:name id="11A48102F9EF987EDD4E81595E613E33" type="personal">
<mods:role id="B8AE905799DB4D651AC2D56D57A21A59">
<mods:roleTerm id="2D37683A52FE7C34D2FCD42FBB1C8C3D">Author</mods:roleTerm>
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<mods:namePart id="68652F6F85F308C7278D72070E5D859A">Villarreal, Osvaldo</mods:namePart>
<mods:nameIdentifier id="46700ED70B4BC9C103BC732CAC790F97" type="ORCID">0000-0001-5355-3723</mods:nameIdentifier>
<mods:affiliation id="F41B084255D1248047609247D4928B6F">Centro de Ecología, Instituto Venezolano de Investigaciones Científicas (IVIC), km 11 carretera Panamericana, Altos de Pipe, edo. Miranda 1204 - A, Venezuela &amp; Instituto y Museo del Instituto de Zoología Agrícola, Facultad de Agronomía, Universidad Central de Venezuela, Apartado 4579, Maracay 2101, Aragua, Venezuela &amp; Departamento de Invertebrados, Museu Nacional / UFRJ, Quinta da Boa Vista, São Cristóvão, 20.940 - 040, Rio de Janeiro – RJ, Brazil</mods:affiliation>
</mods:name>
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<mods:role id="E6B4D79B640A9F55216854CCCEC6743C">
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<mods:namePart id="F155C88B9DA37F849782AD97B4F2A0B8">Ahumada-C., Daniela</mods:namePart>
<mods:nameIdentifier id="D40B41E884C2CF471933CA7103228D86" type="ORCID">0000-0002-4182-5143</mods:nameIdentifier>
<mods:affiliation id="A87962686032A8174140ED1655CF5445">Grupo de Investigación Biología Descriptiva y Aplicada, Programa de Biología, Universidad de Cartagena, Zaragocilla Cra. 50 # 24 - 120, Cartagena de Indias, Bolívar, Colombia</mods:affiliation>
</mods:name>
<mods:name id="4D28906EF87C24C2DC81B8E9CBF8E13D" type="personal">
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<mods:namePart id="ACC088754771E4D0FE0931B248B41CF3">Navas-S., Gabriel R.</mods:namePart>
<mods:nameIdentifier id="D723AA1E7776FC3B658EADFBBFE13328" type="ORCID">0000-0001-9554-6345</mods:nameIdentifier>
<mods:affiliation id="9C6914403FDAEFC706F828004D359E06">Grupo de Investigación Biología Descriptiva y Aplicada, Programa de Biología, Universidad de Cartagena, Zaragocilla Cra. 50 # 24 - 120, Cartagena de Indias, Bolívar, Colombia</mods:affiliation>
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<mods:title id="9BEB12CA0B100C70E5208BA857EE7FAA">Zoosystematics and Evolution</mods:title>
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<mods:date id="4882B0C5204890DF154BCC7C94084015">2024</mods:date>
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<mods:number id="7E3016C5340B20CC985C46FA9C4538F0">2024-06-17</mods:number>
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<mods:number id="8B009B7C324CFFD4C79086CD25B10930">100</mods:number>
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<mods:number id="608A01F8DCE0ACA05B199F8BFCCC1268">3</mods:number>
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<treatment id="A2A72C0A27B3578D80E5BD65E4D16D5E" ID-DOI="http://doi.org/10.5281/zenodo.11975173" ID-Zenodo-Dep="11975173" ID-arpha="A2A72C0A-27B3-578D-80E5-BD65E4D16D5E" LSID="urn:lsid:plazi:treatment:A2A72C0A27B3578D80E5BD65E4D16D5E" httpUri="http://treatment.plazi.org/id/A2A72C0A27B3578D80E5BD65E4D16D5E">
<subSubSection id="14D3A861783489CC4D465A41AE1C99D4" type="nomenclature">
<paragraph id="C9FAD5520E49B612ADDC0F8DAD8F04B2">
<taxonomicName id="D8E03BE30FCA1230938E5BD7C8A8AAE0" ID-CoL="7W5WS" authority="(Roewer, 1915)" baseAuthorityName="Roewer" baseAuthorityYear="1915" class="Arachnida" family="Stygnidae" genus="Eutimesius" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="2C2502FB6A77DDC745616464445B2ECC" italics="true">Eutimesius ephippiatus</emphasis>
(
<bibRefCitation id="69991C9674CAE5C270D97185623A39DE" author="Roewer C-F" issue="3" journalOrPublisher="Archiv für Naturgeschichte" pagination="1-152" refId="B14" refString="Roewer C-F (1915) 106 neue Opilioniden. Archiv fur Naturgeschichte 81 (3): 1 - 152." title="106 neue Opilioniden." volume="81" year="1915">Roewer, 1915</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="877C45865FD8866B76C13D65EE293852" type="description">
<paragraph id="153F3414FEEF9D8CE8771EC2F4B2621E">
<figureCitation id="6C26162758BF6660C3668ED095A78DD8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Eutimesius ephippiatus: A. Male habitus, dorsal view (ICN-Ao - 1492); B. Female habitus, dorsal view (ICN-Ao - 18); C. Male habitus, lateral view (ICN-Ao - 1492). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure9" httpUri="https://binary.pensoft.net/fig/1072637">Figs 9</figureCitation>
,
<figureCitation id="69094527C0CFF46620202BEB3328E405" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Eutimesius ephippiatus male (ICN-Ao - 18): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure10" httpUri="https://binary.pensoft.net/fig/1072638">10</figureCitation>
,
<figureCitation id="BF30FC4476A520C2720CD2073CBA5B39" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eutimesius spp., penis, distal portion in dorsal, ventral, and lateral view: A – C. E. aroa sp. nov., (MIZA 0105934); D – F. E. canoabo sp. nov., (MIZA 0105945); G – I. E. ephippiatus, (ICN-Ao - 1492); J – L. E. guaichia sp. nov., (MIZA 0105935); M – O. E. simoni, (ICN-Ao - 328). A – I. Unscaled." figureDoi="10.3897/zse.100.120207.figure14" httpUri="https://binary.pensoft.net/fig/1072642">14 G – I</figureCitation>
,
<figureCitation id="897589D6CF0A7C31CF7C34F4F3565671" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map showing the distribution of Eutimesius." figureDoi="10.3897/zse.100.120207.figure2" httpUri="https://binary.pensoft.net/fig/1072630">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="829F4A64830CBE17DECD97E8BF6014CE" type="reference_group">
<paragraph id="533E52522CBDAD9501A6B6BB45998D3B">
<treatmentCitationGroup id="A319E3DDD63A914433DB44EC79F8C4D1">
<treatmentCitation id="8B3278E51D952381C764BD25268D3072">
<taxonomicName id="F8C2C060C7DF1923EADA2D1B420E2421" ID-CoL="9X4YR" authority="Roewer" authorityName="Roewer" authorityYear="1915" class="Arachnida" family="Stygnidae" genus="Dichobunistygnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="32FA79BE08CA4588491C20B1FC0D0ACB" italics="true">Dichobunistygnus ephippiatus</emphasis>
Roewer
</taxonomicName>
C-F, (1915): 105, figs 57 a – b.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="B39ECE31C96DB7B54F9309FD76501C0E">
<treatmentCitationGroup id="4FD3A96F03D2690539F80F4EE2D3145E">
<treatmentCitation id="7FD03F212846E44FEE831F732F071303">
<taxonomicName id="D8841CD852179C0CC1C7728ECE67447F" ID-CoL="7W5WS" baseAuthorityName="Roewer" baseAuthorityYear="1915" class="Arachnida" family="Stygnidae" genus="Eutimesius" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="953D845904B7586B4285F3749E0BE8B6" italics="true">Eutimesius ephippiatus</emphasis>
</taxonomicName>
<comment id="308653CD560E24AC1858875DE1C0ED73">
:
<treatmentCitation id="2266ED11E80D6D02DAEC3DD54442ED4B" author="Pinto-da-Rocha R" page="187" year="1997">
<bibRefCitation id="519BB2DE89EABAD80A521F50859D1740" DOI="10.11606/issn.2176-7793.v33i4p163-342" author="Pinto-da-Rocha R" firstAuthor="Pinto-da-Rocha" issue="4" journalOrPublisher="Arquivos de Zoologia" pagination="163-342" refId="B12" refString="Pinto-da-Rocha R (1997) Systematic review of the neotropical family Stygnidae (Opiliones: Laniatores, Gonyleptoidea). Arquivos de Zoologia 33 (4): 163–342. https://doi.org/10.11606/issn.2176-7793.v33i4p163-342" title="Systematic review of the neotropical family Stygnidae (Opiliones: Laniatores, Gonyleptoidea)." volume="33" year="1997">Pinto-da-Rocha (1997)</bibRefCitation>
: 187
</treatmentCitation>
, figs 54–56.
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUYAG" type="material">
<paragraph id="25454CC2E3FB58D6DD68D7955FF847FA">
<heading id="E253BAA236B6A8B93ED308FD91436DEC" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="4C3F3C61C21EBF1350BB971A1D05D98B">
<materialsCitation id="68F64F2B41984F032E69934ACF2F29A2" collectedFrom="forest edge roadside" collectingDate="2016-08" collectionCode="MPUJ" collectorName="Rodriguez, C." country="Colombia" county="Santa Maria" latitude="4.89811" location="Hyca Quye" longLatPrecision="1" longitude="-73.29344" municipality="Santa Maria" specimenCode="ENT 004 8876" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Boyaca">
<collectingCountry id="24EB9037299736170052EF8481D29458" name="Colombia">Colombia</collectingCountry>
•
<specimenCount id="BE5DD3813ED713555697B6FCDDE22794" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="004CE0CE095ED2610AFD639CA4E5FE75" count="1" type="female">1 ♀</specimenCount>
;
<collectingRegion id="EF5F149D92ACBF864E078C31E27D7DA1" country="Colombia" name="Boyaca">Boyacá</collectingRegion>
(New department record),
<collectingMunicipality id="6108ACCD96B2B21FC134FA37918920D8">Santa María</collectingMunicipality>
, sendero
<location id="81A683EEB63E7E579C933FE9D94AE49F" LSID="urn:lsid:plazi:treatment:A2A72C0A27B3578D80E5BD65E4D16D5E:81A683EEB63E7E579C933FE9D94AE49F" country="Colombia" county="Santa Maria" latitude="4.89811" longLatPrecision="1" longitude="-73.29344" municipality="Santa Maria" name="Hyca Quye" stateProvince="Boyaca">Hyca Quye</location>
,
<locationDeviation id="DD9498F1E98E0977E119F483CD196799">
<quantity id="4ACF52B9C10D8B8D1B9B8CCC8AF2C731" metricMagnitude="3" metricUnit="m" metricValue="5.5" unit="km" value="5.5">5.5 km</quantity>
NW from Santa María
</locationDeviation>
,
<collectedFrom id="C2E8239196DFEB429E69E8420EED1944">forest edge roadside</collectedFrom>
; (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4YAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.293440,4.898110]}">
<geoCoordinate id="56DEFC8D295B4C7E4B068EF8393059CC" degrees="4.89811" direction="north" orientation="latitude" precision="1" value="4.89811">4.89811 ° N</geoCoordinate>
,
<geoCoordinate id="635E3C9B03C44EA327E77D845B08E2D2" degrees="73.29344" direction="west" orientation="longitude" precision="1" value="-73.29344">73.29344 ° W</geoCoordinate>
</named-content>
);
<date id="C1CD9BC4F7A1C0D43770671E4C532B2C" value="2016-08">
<collectingDate id="B8BE8AE6B6F08F3E4DBA7CAE6F7C6472" value="2016-08">Ago. 2016</collectingDate>
</date>
; (
<collectorName id="1D211FDCAC8ADC549263A335F117B5F8">Rodríguez, C.</collectorName>
leg.) (
<abbrev content-type="institution" id="ABBRID0ECZAG" xlink_title="Colección entomólogica, Museo Javeriano de Historia Natural">
<collectionCode id="6F210EA2A9357D868DC263E91544D220">MPUJ</collectionCode>
_
<specimenCode id="9CC3CE4BE540566F141B802DF614B609">ENT 004 8876</specimenCode>
</abbrev>
)
</materialsCitation>
. •
<materialsCitation id="62ED66C5BC5D957A96DEC21EC259A087" collectedFrom="forest edge" collectingDate="2015-08" collectingDateMax="2015-08" collectingDateMin="2015-08" collectingMethod="manual capture" collectionCode="MPUJ" collectorName="Guzman, K." country="Argentina" county="Santa Maria" elevation="850" latitude="4.85673" location="Quebrada La Argentina" longLatPrecision="1" longitude="-73.26375" municipality="Santa Maria" specimenCode="ENT 0039466" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<specimenCount id="C0B8FA9610E8EF03EE388821F3DDDC4C" count="1" type="male">1 ♂</specimenCount>
,
<collectingRegion id="BEC8D3561B4318B3D991E84379052F80" country="Colombia" name="Boyaca">Boyacá</collectingRegion>
,
<collectingMunicipality id="013D1F7AA27D6F8442244802A24F4D26">Santa María</collectingMunicipality>
,
<location id="340D4732F0F0C459ABDC448B90F85CC8" LSID="urn:lsid:plazi:treatment:A2A72C0A27B3578D80E5BD65E4D16D5E:340D4732F0F0C459ABDC448B90F85CC8" country="Argentina" county="Santa Maria" latitude="4.85673" longLatPrecision="1" longitude="-73.26375" municipality="Santa Maria" name="Quebrada La Argentina" stateProvince="Boyaca">Quebrada La Argentina</location>
,
<collectedFrom id="B1DEBA6C1A5E29820AF14AD793C7A3ED">forest edge</collectedFrom>
,
<locationDeviation id="F64AAB2EC7B630E3C5CB35EED3214600">
<quantity id="1EB4897DC7EAFBB3A6EBD6A44D91198F" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="km" value="0.5">0.5 km</quantity>
SSW from Santa María
</locationDeviation>
; (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKZAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.263750,4.856730]}">
<geoCoordinate id="F12D7EBD21D4B5C45AD6A1B91EF52494" degrees="4.85673" direction="north" orientation="latitude" precision="1" value="4.85673">4.85673 ° N</geoCoordinate>
,
<geoCoordinate id="A000AE957F7682192B5EFF7FA664726F" degrees="73.26375" direction="west" orientation="longitude" precision="1" value="-73.26375">73.26375 ° W</geoCoordinate>
</named-content>
);
<collectingMethod id="4A7461EE7ECA746BB966D249D9BBA1B8">manual capture</collectingMethod>
;
<elevation id="F412A8771EA17C760C734C8EF64023D0" metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">
<quantity id="1A1B17E874D25C7E6780D03ABB5E2FC5" metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">850 m</quantity>
a. s. l.
</elevation>
;
<collectingDate id="88E7F9E906C2A4E3B94180785FBC88CA">Ago-Sep. 2015</collectingDate>
; (
<collectorName id="21907BD6250048F44243D0F6F4338FCF">Guzmán, K.</collectorName>
leg.) (
<abbrev content-type="institution" id="ABBRID0EPZAG" xlink_title="Colección entomólogica, Museo Javeriano de Historia Natural">
<collectionCode id="C3235B7F70391098BFE43A9E477B653E">MPUJ</collectionCode>
_
<specimenCode id="AA27C47B2423B49C503F92CAEE82A013" collectionCode="ENT" country="Uganda" lsid="urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" type="Herbarium">ENT 0039466</specimenCode>
</abbrev>
)
</materialsCitation>
. •
<materialsCitation id="EC4CCF887DB85B4DD88A55D4DF70BBE5" collectingDate="1978-10-14" collectorName="La Rotta, C." country="Argentina" latitude="5.1866" location="Agua Azul" longLatPrecision="7" longitude="-72.5558" municipality="Casanare" specimenCode="ICN-Ao - 18" specimenCount="1" specimenCount-female="1" stateProvince="Boyaca">
<specimenCount id="2C639A91A7688E6D41678B75D0EC3ED6" count="1" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="E3C336AE14826A808AF106A68CE92685">Casanare</collectingMunicipality>
(new department record),
<location id="673D429B1855B266C77F9C568C7EC786" LSID="urn:lsid:plazi:treatment:A2A72C0A27B3578D80E5BD65E4D16D5E:673D429B1855B266C77F9C568C7EC786" country="Argentina" latitude="5.1866" longLatPrecision="7" longitude="-72.5558" municipality="Casanare" name="Agua Azul" stateProvince="Boyaca">Agua Azul</location>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXZAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-72.555800,5.186600]}">
<geoCoordinate id="DCD1D117160661189E42B688DCC3DA0E" degrees="5.1866" direction="north" orientation="latitude" precision="5" value="5.1866">5.1866 ° N</geoCoordinate>
,
<geoCoordinate id="9F02A2AE8E905810D0E4944FBA6015E0" degrees="72.5558" direction="west" orientation="longitude" precision="5" value="-72.5558">72.5558 ° W</geoCoordinate>
</named-content>
];
<date id="02E966C198C3307BCC91985759036CE4" value="1978-10-14">
<collectingDate id="1932C36F0558C34590B38CD878353A4A" value="1978-10-14">14 Oct. 1978</collectingDate>
</date>
; (
<collectorName id="42869158CE45CF5401A28CE0AD1A3720">La Rotta, C.</collectorName>
leg.) (
<specimenCode id="DADD1314FF3929065CD5737C34E6EAD1">
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- 18
</specimenCode>
)
</materialsCitation>
. •
<materialsCitation id="F7E730B6791F7D7AB4AFCB69E9FBDCF8" collectingDate="1998-04" collectingMethod="manual capture" collectionCode="V" collectorName="Rodriguez, V. &amp; Sierra, S. &amp; Florez, E. &amp; Varon, A." country="Argentina" county="Cundinamarca" elevation="500" latitude="4.7133" location="left bank of the Rio Zaquea" longLatPrecision="7" longitude="-73.3016" municipality="Inspeccion San Pedro de Jagua" specimenCode="ICN-Ao - 1491" specimenCount="1" specimenCount-female="1" stateProvince="Boyaca">
<specimenCount id="75086CD494B06DCF7480C0A88B559B02" count="1" type="female">1 ♀</specimenCount>
;
<collectingCounty id="3734A5B034FDC1AFD9118F544CCA42D4">Cundinamarca</collectingCounty>
, Ubalá, Ubalá B.,
<collectingMunicipality id="6BD7CA60587B3AA3130299CD443B1279">Inspección San Pedro de Jágua</collectingMunicipality>
,
<location id="0DD069C321231872D259E695CA6FCD3E" LSID="urn:lsid:plazi:treatment:A2A72C0A27B3578D80E5BD65E4D16D5E:0DD069C321231872D259E695CA6FCD3E" country="Argentina" county="Cundinamarca" latitude="4.7133" longLatPrecision="7" longitude="-73.3016" municipality="Inspeccion San Pedro de Jagua" name="left bank of the Rio Zaquea" stateProvince="Boyaca">left bank of the Rio Zaquea</location>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EE1AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.301600,4.713300]}">
<geoCoordinate id="FAC2FC027AE4DF8EC23EA87475C8A01A" degrees="4.7133" direction="north" orientation="latitude" precision="5" value="4.7133">4.7133 ° N</geoCoordinate>
,
<geoCoordinate id="8673B5724DA5791199CB667E8479F9F7" degrees="73.3016" direction="west" orientation="longitude" precision="5" value="-73.3016">73.3016 ° W</geoCoordinate>
</named-content>
];
<elevation id="2754BF5F52982D2BE112A52F63211E6B" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<quantity id="D3A4F872494054BCC0DDC8033F0AB86E" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</quantity>
a. s. l.
</elevation>
;
<collectingMethod id="958A96A865E0D8512D93BFAD75ECFE76">manual capture</collectingMethod>
;
<date id="9B69FF75D31C15B0210559170176F6BA" value="1998-04">
<collectingDate id="644AF22A19F002C059F543D713E712D9" value="1998-04">Apr. 1998</collectingDate>
</date>
; (
<collectorName id="6AA09BB5D559793C87C836B7E22BB14D">Rodríguez, V.</collectorName>
,
<collectorName id="0DE56F379C5C2E173B059E313EFB3235">Sierra, S.</collectorName>
,
<collectorName id="09C8AE310C4518A07CA25F7533295CEF">Flórez, E.</collectorName>
,
<collectorName id="BE950F2C44AA58B03E66B12310923A3F">Varón, A.</collectorName>
leg.) (
<specimenCode id="C26CFB8AAF0E38680CECB5FA705D7BAB">
<named-content id="3BF123F5886D94DB46B8867A75120FF7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-nacional-de-colombia" xlink_title="Instituto de Ciencias Naturales of the Universidad Nacional de Colombia, Bogotá, Colombia">ICN-Ao</named-content>
- 1491
</specimenCode>
)
</materialsCitation>
. •
<materialsCitation id="6E94F57DFA80D77C2C53A20120FE6589" collectingDate="1998-04" collectingMethod="manual capture" collectionCode="V" collectorName="Rodriguez, V. &amp; Sierra, S. &amp; Florez, E. &amp; Varon, A." country="Argentina" county="Cundinamarca" elevation="1000" latitude="4.743" location="Vda. Boca de Monte" longLatPrecision="7" longitude="-73.3016" municipality="Inspeccion Mambita" specimenCode="ICN-Ao - 1492" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca">
<specimenCount id="5DCC5056078C19B5E5F3062F884221E5" count="1" type="male">1 ♂</specimenCount>
;
<collectingCounty id="0ACBF597528082B759314370A80FD54D">Cundinamarca</collectingCounty>
, Ubalá, Ubalá B.,
<collectingMunicipality id="040EF46F50A6EED14B19245F96992630">Inspección Mambita</collectingMunicipality>
,
<location id="73ECB31D8AC66B207DE4AA2F278668A2" LSID="urn:lsid:plazi:treatment:A2A72C0A27B3578D80E5BD65E4D16D5E:73ECB31D8AC66B207DE4AA2F278668A2" country="Argentina" county="Cundinamarca" latitude="4.743" longLatPrecision="7" longitude="-73.3016" municipality="Inspeccion Mambita" name="Vda. Boca de Monte" stateProvince="Boyaca">Vda. Boca de Monte</location>
, surroundings Boca de Monte school; (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ER1AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.301600,4.743000]}">
<geoCoordinate id="63EAC511A7C77159F41207594FC58808" degrees="4.7430" direction="north" orientation="latitude" precision="5" value="4.743">4.7430 ° N</geoCoordinate>
,
<geoCoordinate id="46E67B472F51060693255ACF334C3681" degrees="73.3016" direction="west" orientation="longitude" precision="5" value="-73.3016">73.3016 ° W</geoCoordinate>
</named-content>
);
<elevation id="1B100E70439E370D3DDD72BDB26EBD6F" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<quantity id="678E9AFC833D8FE8962632AD46D413C1" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1,000 m</quantity>
a. s. l.
</elevation>
;
<collectingMethod id="C826EFB2C80EE4E31521A734D3493DEE">manual capture</collectingMethod>
;
<date id="401481942BE90090B39AC0F950276DAC" value="1998-04">
<collectingDate id="5006F0DFF880CEA291DC71FD007140E1" value="1998-04">Apr. 1998</collectingDate>
</date>
; (
<collectorName id="A58E588DF6FA1DD509FABD13CE6F2D7E">Rodríguez, V.</collectorName>
,
<collectorName id="BEE0B4BC855C5B8E7A379E8311105F60">Sierra, S.</collectorName>
,
<collectorName id="EDD9129BA8D944C16D00FB671DCC65D3">Flórez, E.</collectorName>
,
<collectorName id="502D78D5E7F5AE24320922B20D15F54E">Varón, A.</collectorName>
leg.) (
<specimenCode id="2A2F24CEB779F0A25DF6928A5299EB0A">
<named-content id="2D8207DDF0BCD0C04B06DBCB05C89E0E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-nacional-de-colombia" xlink_title="Instituto de Ciencias Naturales of the Universidad Nacional de Colombia, Bogotá, Colombia">ICN-Ao</named-content>
- 1492
</specimenCode>
)
</materialsCitation>
. •
<materialsCitation id="82361D08243297AC5D972A9B374DECC9" collectingDate="1998-04" collectingMethod="manual capture" collectionCode="V" collectorName="Rodriguez, V. &amp; Sierra, S. &amp; Florez, E. &amp; Varon, A." country="Argentina" county="Inspeccion San Pedro de Jagua" elevation="500" latitude="4.7133" location="left bank of the Rio Zaquea" longLatPrecision="7" longitude="-73.3392" municipality="Vda. Soya" specimenCode="ICN-Ao - 1511" specimenCount="1" specimenCount-female="1" stateProvince="Boyaca">
<specimenCount id="B3F6DCAC2001CFE169251E8E34F121F5" count="1" type="female">1 ♀</specimenCount>
; Cundinamarca, Ubalá, Ubalá B.,
<collectingCounty id="8EB55A2D801E01FDF182B42165BE42E9">Inspección San Pedro de Jágua</collectingCounty>
,
<collectingMunicipality id="01989E92819960BAB76F9545DC02F43A">Vda. Soya</collectingMunicipality>
,
<location id="7E768F2FA44022FAEE41CE14BF7FB607" LSID="urn:lsid:plazi:treatment:A2A72C0A27B3578D80E5BD65E4D16D5E:7E768F2FA44022FAEE41CE14BF7FB607" country="Argentina" county="Inspeccion San Pedro de Jagua" latitude="4.7133" longLatPrecision="7" longitude="-73.3392" municipality="Vda. Soya" name="left bank of the Rio Zaquea" stateProvince="Boyaca">left bank of the Rio Zaquea</location>
; (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E51AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-73.339200,4.713300]}">
<geoCoordinate id="CF7A42148650B20B1B2188B6712A67A3" degrees="4.7133" direction="north" orientation="latitude" precision="5" value="4.7133">4.7133 ° N</geoCoordinate>
,
<geoCoordinate id="2B0BA8B98EFD3925206611582B5D9AE5" degrees="73.3392" direction="west" orientation="longitude" precision="5" value="-73.3392">73.3392 ° W</geoCoordinate>
</named-content>
);
<elevation id="DF7D5A471B8F09ADAB5E73D1DEBB4565" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<quantity id="802404A097BD61C632834E3916D43A23" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</quantity>
a. s. l.
</elevation>
;
<collectingMethod id="3ADB5FB02283829A0875B1A42D04B0E4">manual capture</collectingMethod>
;
<date id="901D8E10376051327CFCF39FCC781E55" value="1998-04">
<collectingDate id="A812B74E20C84BE51CD0F1AE37BA4520" value="1998-04">Apr. 1998</collectingDate>
</date>
; (
<collectorName id="5C427F9FACF33BB9D813E2CE93CBE76A">Rodríguez, V.</collectorName>
,
<collectorName id="C52A4CBA62E692014608F920DB7D5DA8">Sierra, S.</collectorName>
,
<collectorName id="A3611ECD20CE896F686594D87B4DDFEA">Flórez, E.</collectorName>
,
<collectorName id="78DA02FC1969599A09B146844D275DE2">Varón, A.</collectorName>
leg.) (
<specimenCode id="0532D18E277D0FA18E3E69069E9DED71">
<named-content id="3B86129475D20C69388A9022569F3654" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-nacional-de-colombia" xlink_title="Instituto de Ciencias Naturales of the Universidad Nacional de Colombia, Bogotá, Colombia">ICN-Ao</named-content>
- 1511
</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EI2AG" type="diagnosis">
<paragraph id="C36B2854EBAEF957EF47E5F8B6EF0842">
<heading id="7B80B2523CB599709F2416E281D646B2" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="4D2A35758A96A4526723E7F3AE20BFEC">
It is distinguishable from all other species in the genus by the pattern of dry white spots that encircled completely the eyes (Figs
<figureCitation id="BAECBE1689785B6F3B4F9082B0855810" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Eutimesius ephippiatus: A. Male habitus, dorsal view (ICN-Ao - 1492); B. Female habitus, dorsal view (ICN-Ao - 18); C. Male habitus, lateral view (ICN-Ao - 1492). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure9" httpUri="https://binary.pensoft.net/fig/1072637">9</figureCitation>
,
<figureCitation id="AC6FE89B4AB173B75A91BF0859742281" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Eutimesius ephippiatus male (ICN-Ao - 18): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure10" httpUri="https://binary.pensoft.net/fig/1072638">10 A, B</figureCitation>
), and the proximal half of femora and tibiae IV without tubercles or with only minute granules (Figs
<figureCitation id="CFC3B602CA0B4F9ABB77625791D445AA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Eutimesius ephippiatus male (ICN-Ao - 18): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure10" httpUri="https://binary.pensoft.net/fig/1072638">10 F, G</figureCitation>
).
</paragraph>
<caption id="68711075F545AE2CFD4B9D7CBAB5DA99" ID-DOI="10.3897/zse.100.120207.figure9" ID-arpha="290D3584-0F74-5C19-9239-30612CE73E01" httpUri="https://binary.pensoft.net/fig/1072637" startId="F9">
<paragraph id="4ED1331EF084AF3D80B879325162D171">
<label id="1640656FD134515D10253A782C438648">Figure 9.</label>
</paragraph>
<paragraph id="B9250BDC8F226827F4E39ACB7C7C1069">
<taxonomicName id="3E21C665AAB1D583E69DD8955A097982" baseAuthorityName="Roewer" baseAuthorityYear="1915" class="Arachnida" family="Stygnidae" genus="Eutimesius" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="97AB00EF9EC01717E343C230F625C7EE" italics="true">Eutimesius ephippiatus</emphasis>
</taxonomicName>
:
<emphasis id="5F9A030B31D96D0F625CBFEB44F3601F" bold="true">A.</emphasis>
Male habitus, dorsal view (
<specimenCode id="115B0244C27EEEE434BC940E9FE3F3CD">
<named-content id="5B9FC8415F80BF3C5220DF56A0471F6D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-nacional-de-colombia" xlink_title="Instituto de Ciencias Naturales of the Universidad Nacional de Colombia, Bogotá, Colombia">ICN-Ao</named-content>
- 1492
</specimenCode>
);
<emphasis id="97C587CAC926481C8EE54BE2BB66CFD8" bold="true">B.</emphasis>
Female habitus, dorsal view (
<specimenCode id="253B228EF4F0594CBCFCC5B6BF52C8AD">
<named-content id="8C4998EAB3BAB622FCDFC80743010B16" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-nacional-de-colombia" xlink_title="Instituto de Ciencias Naturales of the Universidad Nacional de Colombia, Bogotá, Colombia">ICN-Ao</named-content>
- 18
</specimenCode>
);
<emphasis id="2D127A16E6BCEA157B6845E2A910C027" bold="true">C.</emphasis>
Male habitus, lateral view (
<specimenCode id="8BF3EDD63D06DA1016F1D37CF480A140">
<named-content id="7FB47CB8716A93D2EDA5A4F22361F41A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-nacional-de-colombia" xlink_title="Instituto de Ciencias Naturales of the Universidad Nacional de Colombia, Bogotá, Colombia">ICN-Ao</named-content>
- 1492
</specimenCode>
). Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="FB62E3B93844DC33D720E5EDCEAE5F93" ID-DOI="10.3897/zse.100.120207.figure10" ID-arpha="7B50CA3E-CD98-558B-A9BE-9C4F71AB0DF2" httpUri="https://binary.pensoft.net/fig/1072638" startId="F10">
<paragraph id="31AA208DEC5B995808D187A81262B931">
<label id="FB0551DF4225618D39044FCE0DC20530">Figure 10.</label>
</paragraph>
<paragraph id="39FD75F6533A73F6D68E7B783FD2CAEE">
<taxonomicName id="E22E87ED4878BC4FDB02CD79FAB1A225" baseAuthorityName="Roewer" baseAuthorityYear="1915" class="Arachnida" family="Stygnidae" genus="Eutimesius" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="7B1D5C778A3267F83C64F4DEEF156338" italics="true">Eutimesius ephippiatus</emphasis>
</taxonomicName>
male (
<specimenCode id="74B8D7F0BE7E4EFDDFE08A307BE19710">
<named-content id="F8F5879056C43E58586EDB2ADB5CDE13" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-nacional-de-colombia" xlink_title="Instituto de Ciencias Naturales of the Universidad Nacional de Colombia, Bogotá, Colombia">ICN-Ao</named-content>
- 18
</specimenCode>
):
<emphasis id="97F7D0855A65A855339D81A685131A03" bold="true">A.</emphasis>
Habitus, dorsal view;
<emphasis id="815A342AEA28950DC3257933C4F05EFF" bold="true">B.</emphasis>
Ditto, lateral view;
<emphasis id="2F44A87A41E97B536207B165DA806CCC" bold="true">C.</emphasis>
Right chelicera, frontal view;
<emphasis id="67F143B26FAAB305C09DB5A9E8FFA660" bold="true">D.</emphasis>
Right pedipalp, tibia, and tarsus, ectal view;
<emphasis id="A914CEA493209324D0FBA576432BD5EB" bold="true">E.</emphasis>
Ditto, mesal view;
<emphasis id="353B6CA84990D8852F57B8F6E9AFAD49" bold="true">F.</emphasis>
Right leg IV, dorsal view;
<emphasis id="0EE2534DAFB90F86BECBB26ACAE7D271" bold="true">G.</emphasis>
Ditto, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ES5AG" type="description">
<paragraph id="AC63142B2A2FDF185091B37C433F62BE">
<heading id="C6E469A99AEBF88830EF138414C6A771" reason="title">Complementary description.</heading>
</paragraph>
<paragraph id="A0E8C927DC10DD3431CDAA6A8570AAF9">
<emphasis id="6267D96A6F391A4D94176EABD1452A83" bold="true">
Male.
<emphasis id="66967F7D60D6F87D0CC2F291B260F172" bold="true" italics="true">Measurements</emphasis>
.
</emphasis>
<abbrev id="ABBRID0E45AG" xlink_title="dorsal scutum length">DSL</abbrev>
: 1.4;
<abbrev id="ABBRID0EB6AG" xlink_title="dorsal scutum width">DSW</abbrev>
: 2.8;
<abbrev id="ABBRID0EF6AG" xlink_title="maximum abdominal scutum length">AL</abbrev>
: 1.7;
<abbrev id="ABBRID0EJ6AG" xlink_title="maximum abdominal scutum width">AW</abbrev>
: 3.0;
<abbrev id="ABBRID0EN6AG" xlink_title="interocular distance">IOD</abbrev>
: 1.6; pedipalp:
<abbrev id="ABBRID0ER6AG" xlink_title="pedipalpal coxa">CoPp</abbrev>
0.7, TrPp 0.7,
<abbrev id="ABBRID0EV6AG" xlink_title="pedipalpal femur">FePp</abbrev>
2.9,
<abbrev id="ABBRID0EZ6AG" xlink_title="pedipalpal patella">PaPp</abbrev>
1.3,
<abbrev id="ABBRID0E46AG" xlink_title="pedipalpal tibia">TiPp</abbrev>
1.5,
<abbrev id="ABBRID0EBABG" xlink_title="pedipalpal tarsus">TaPp</abbrev>
1.3,
<abbrev id="ABBRID0EFABG" xlink_title="pedipalp claw">ClPp</abbrev>
0.8; total 9.2; Leg IV:
<abbrev id="ABBRID0EJABG" xlink_title="femur length">FeL</abbrev>
7.6,
<abbrev id="ABBRID0ENABG" xlink_title="tibia length">TiL</abbrev>
2.6.
<emphasis id="E9130A44239B1E05887645AB0C5CC31C" bold="true" italics="true">Dorsum</emphasis>
(Figs
<figureCitation id="48377314A207044E17E9D568CEB65EFF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Eutimesius ephippiatus: A. Male habitus, dorsal view (ICN-Ao - 1492); B. Female habitus, dorsal view (ICN-Ao - 18); C. Male habitus, lateral view (ICN-Ao - 1492). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure9" httpUri="https://binary.pensoft.net/fig/1072637">9 A, C</figureCitation>
,
<figureCitation id="914E10D7C383334FA02A3B66473900DD" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Eutimesius ephippiatus male (ICN-Ao - 18): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure10" httpUri="https://binary.pensoft.net/fig/1072638">10 A, B</figureCitation>
).
<abbrev id="ABBRID0E3ABG" xlink_title="dorsal scutum">DS</abbrev>
outline Epsilon
<typeStatus id="0809F7C02C937F6CAFDA018DFE7662A9">type</typeStatus>
. Anterior margin of
<abbrev id="ABBRID0EABBG" xlink_title="dorsal scutum">DS</abbrev>
with four anterolateral large tubercles. Anteromedial and lateral process of the cheliceral sockets short. Eyes separated into two small smooth mounds, placed slightly posteriorly on the carapace. Interocular region with small granules and one central elevated and irregular mound that terminates in two short tubercles. Lateral margins with a row of seven tubercles. Mesotergum divided into four areas, III – IV mostly fused: I divided medially into two oval halves, with one conspicuous tubercle on each side and some lateral granules; two entire, with seven-eight conspicuous tubercles; III / IV with one pair of paramedian very large spines projected backward (three times the length of the interocular eminence); and granulated, posterior region with a row of small granules. Posterior margin smooth, free tergites with a row of small granules. Anal operculum and sternites with small granules.
<emphasis id="7CA503E5FF848D8DA249E98F873D3EC4" bold="true" italics="true">Venter</emphasis>
(Figs
<figureCitation id="4FBA29E4BD3CD3A202391F9CB441A90F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Eutimesius ephippiatus: A. Male habitus, dorsal view (ICN-Ao - 1492); B. Female habitus, dorsal view (ICN-Ao - 18); C. Male habitus, lateral view (ICN-Ao - 1492). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure9" httpUri="https://binary.pensoft.net/fig/1072637">9 C</figureCitation>
,
<figureCitation id="BEDEAA74B48618206D47F85400A5D5FD" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Eutimesius ephippiatus male (ICN-Ao - 18): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure10" httpUri="https://binary.pensoft.net/fig/1072638">10 B</figureCitation>
). Coxa I with three distal tubercles and a medial row of five tubercles; II with three distal tubercles, a median row of nine tubercles, and a posterior row of three-five small tubercles; III with three distal tubercles, three-five anterior, eight medial, and five posterior tubercles; IV with four intercoxal tubercles, one median row with eight tubercles aligned, some scattered anterior tubercles, and about eight-ten posterior tubercles aligned. Genital operculum and stigmatic area with scattered granules. Free sternites with a row of small granules.
<emphasis id="E1BD3A877A1DE22E284AA901FF727C9C" bold="true" italics="true">Chelicerae</emphasis>
(Figs
<figureCitation id="241B4601E4D30B46C7988C94B2DB7BB3" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Eutimesius ephippiatus: A. Male habitus, dorsal view (ICN-Ao - 1492); B. Female habitus, dorsal view (ICN-Ao - 18); C. Male habitus, lateral view (ICN-Ao - 1492). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure9" httpUri="https://binary.pensoft.net/fig/1072637">9 A, C</figureCitation>
,
<figureCitation id="54B8972BD19B185CF47D4D894AB1E5C1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Eutimesius ephippiatus male (ICN-Ao - 18): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure10" httpUri="https://binary.pensoft.net/fig/1072638">10 A – C</figureCitation>
). Segment I smooth with well-defined bulla with tiny granules. Segment II swollen. Fixed finger with one medial tooth, mobile finger with one sub-proximal large tooth, one medial large tooth and two small subdistal teeth.
<emphasis id="431530D56A8266639E9B7588A3C26B06" bold="true" italics="true">Pedipalps</emphasis>
(Figs
<figureCitation id="1B59BF6BFC5813125504957456E46D75" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Eutimesius ephippiatus: A. Male habitus, dorsal view (ICN-Ao - 1492); B. Female habitus, dorsal view (ICN-Ao - 18); C. Male habitus, lateral view (ICN-Ao - 1492). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure9" httpUri="https://binary.pensoft.net/fig/1072637">9 A</figureCitation>
,
<figureCitation id="E16A931DC8E500C3647B0BB3CF36942D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Eutimesius ephippiatus male (ICN-Ao - 18): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure10" httpUri="https://binary.pensoft.net/fig/1072638">10 A, D, F</figureCitation>
). Coxa with some granules. Trochanter with three dorsodistal tubercles and one dorsal tubercle in the medial position, ventrally with some minute granules. Femur and patella dorsally and ventrally, with some scattered granules. Patella distally swollen. Tibia dorsal with some minute granules, ventrally with a row of minute granules, mesal IIII, ectal IIiIiI. Tarsus dorsally and ventrally smooth; mesal IiIiIi, ectal IiiIiii.
<emphasis id="A06031E236D93E7627046D4D47007A94" bold="true" italics="true">Legs</emphasis>
(Figs
<figureCitation id="3ADAD9FDCCAE29890FD8CB24FD84D1C3" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Eutimesius ephippiatus: A. Male habitus, dorsal view (ICN-Ao - 1492); B. Female habitus, dorsal view (ICN-Ao - 18); C. Male habitus, lateral view (ICN-Ao - 1492). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure9" httpUri="https://binary.pensoft.net/fig/1072637">9 A, C</figureCitation>
,
<figureCitation id="8BCD1F8465F80966C065D037E6C663DD" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Eutimesius ephippiatus male (ICN-Ao - 18): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure10" httpUri="https://binary.pensoft.net/fig/1072638">10 A, B, F, G</figureCitation>
). Coxae I – III unarmed, IV with one large prodorsal tubercle in medial position. Trochanter II with one dorsodistal tubercle, III – IV ventrally granulated, III with one row of 4–5 dorsodistal tubercles and one row of four proventral tubercles, IV with 4–5 prodorsal tubercles (the two distalmost larger), one large retrodorsal sub-distal tubercle, and a group of small prodorsal proximal tubercles. Femora I – II with longitudinal rows of granules dorsally and ventrally, III with one prodorsal distal conical tubercle and ventral row of conical tubercles increasing in size distally, IV with one prolateral and one retrolateral row of small tubercles, dorsally with two distal conical tubercles, and ventrally with two longitudinal rows of conical tubercles increasing in size distally. Patellae III – IV granulated, IV dorsally tuberculated, with one retrodorsal distal conical tubercle, one retroventral row of tubercles (the distalmost larger), two conical ventral tubercles in medial portion (the distalmost larger), and one proventral conical tubercle in distal portion. Tibia III with some granules, IV with one large retrodorsal distal tubercle and some scattered granules and small tubercles, two ventral rows of tubercles. Tarsal claws III and IV opposites and pectinated. Tarsal counts: 7 (3) - 7 (3) / 17-19 / 9-10 / 9 - 9.
<emphasis id="C31DBBEB42941C7576E6823A95C16869" bold="true" italics="true">Penis</emphasis>
(Fig.
<figureCitation id="02B2F27BD7756B11C868F2D7F9524311" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eutimesius spp., penis, distal portion in dorsal, ventral, and lateral view: A – C. E. aroa sp. nov., (MIZA 0105934); D – F. E. canoabo sp. nov., (MIZA 0105945); G – I. E. ephippiatus, (ICN-Ao - 1492); J – L. E. guaichia sp. nov., (MIZA 0105935); M – O. E. simoni, (ICN-Ao - 328). A – I. Unscaled." figureDoi="10.3897/zse.100.120207.figure14" httpUri="https://binary.pensoft.net/fig/1072642">14 G – I</figureCitation>
).
<taxonomicName id="7189E35FF25DB19E6A00B2851774912B" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Stygnidae" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="subFamily" subFamily="Heterostygninae">Heterostygninae</taxonomicName>
general pattern. Malleus swollen.
<abbrev id="ABBRID0E3CBG" xlink_title="lamina parva">LP</abbrev>
with a shallow neck and a deep distal cleft.
<abbrev id="ABBRID0EADBG" xlink_title="macrosetae of penis">MS</abbrev>
- A located on the malleus, two pair dorsally and one pair more ventrally;
<abbrev id="ABBRID0EEDBG" xlink_title="macrosetae of penis">MS</abbrev>
- B pair located ventrally;
<abbrev id="ABBRID0EIDBG" xlink_title="macrosetae of penis">MS</abbrev>
- C pair located medially on the
<abbrev id="ABBRID0EMDBG" xlink_title="lamina parva">LP</abbrev>
, distally to the neck;
<abbrev id="ABBRID0EQDBG" xlink_title="macrosetae of penis">MS</abbrev>
- D 1 large, located sligthly proximal to
<abbrev id="ABBRID0EUDBG" xlink_title="macrosetae of penis">MS</abbrev>
- E;
<abbrev id="ABBRID0EYDBG" xlink_title="macrosetae of penis">MS</abbrev>
- D 2 located basally, near the base of the glans;
<abbrev id="ABBRID0E3DBG" xlink_title="macrosetae of penis">MS</abbrev>
- E apparently with only one pair of setae visible. Glans globose, with short and dorsally curved stylus with small dorsal process.
<emphasis id="CB7D2530AD123A64B0A082492D3AD4EA" bold="true" italics="true">Color</emphasis>
(Fig.
<figureCitation id="90E5E6289BBC670A6CCD14A5F12973F7" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Eutimesius ephippiatus: A. Male habitus, dorsal view (ICN-Ao - 1492); B. Female habitus, dorsal view (ICN-Ao - 18); C. Male habitus, lateral view (ICN-Ao - 1492). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure9" httpUri="https://binary.pensoft.net/fig/1072637">9 A, C</figureCitation>
).
<abbrev id="ABBRID0EHEBG" xlink_title="dorsal scutum">DS</abbrev>
, and free tergites Dark orange yellow (72). Spots on carapace and mesotergum Vivid orange yellow (66). Pair of paramedian spines in area III Brownish black (65). Free sternites Dark yellowish brown (78). Chelicerae reticulated and pedipalps Light greenish yellow (101). Coxae I – IV and trochanters I – IV brilliant yellow (83), Femora I – II reticulated light greenish yellow (101), and III and
<collectionCode id="311D612C37BA32D73FA46023ABC0E15D" country="Norway" name="Mykotektet, National Veterinary Institute">VI</collectionCode>
brilliant yellow (83).
</paragraph>
<paragraph id="C0C42CD834CB9F19D85FB539CEFFD49E">
<emphasis id="10CBC5DE90431358330E0CB0474C4781" bold="true">
Female.
<emphasis id="3887AEF9963BC3F244D1B310B8AA63CA" bold="true" italics="true">Measurements</emphasis>
.
</emphasis>
<abbrev id="ABBRID0ESEBG" xlink_title="dorsal scutum length">DSL</abbrev>
: 1.2–1.5;
<abbrev id="ABBRID0EWEBG" xlink_title="dorsal scutum width">DSW</abbrev>
: 2.5–3.2;
<abbrev id="ABBRID0E1EBG" xlink_title="maximum abdominal scutum length">AL</abbrev>
: 1.5–2.2;
<abbrev id="ABBRID0E5EBG" xlink_title="maximum abdominal scutum width">AW</abbrev>
: 2.9–3.5;
<abbrev id="ABBRID0ECFBG" xlink_title="interocular distance">IOD</abbrev>
: 1.3–1.7; pedipalp:
<abbrev id="ABBRID0EGFBG" xlink_title="pedipalpal coxa">CoPp</abbrev>
0.4–0.8, TrPp 0.5–0.6,
<abbrev id="ABBRID0EKFBG" xlink_title="pedipalpal femur">FePp</abbrev>
1.9–2.6,
<abbrev id="ABBRID0EOFBG" xlink_title="pedipalpal patella">PaPp</abbrev>
1.0–1.4,
<abbrev id="ABBRID0ESFBG" xlink_title="pedipalpal tibia">TiPp</abbrev>
0.8–1.4,
<abbrev id="ABBRID0EWFBG" xlink_title="pedipalpal tarsus">TaPp</abbrev>
0.8–1.3,
<abbrev id="ABBRID0E1FBG" xlink_title="pedipalp claw">ClPp</abbrev>
0.7–0 - 9, total 4.3–8.0; Leg IV:
<abbrev id="ABBRID0E5FBG" xlink_title="femur length">FeL</abbrev>
6.2–8.2,
<abbrev id="ABBRID0ECGBG" xlink_title="tibia length">TiL</abbrev>
3.4–4.1.
<emphasis id="B770C88A205F9478F63D113CCD2631D4" bold="true" italics="true">Description of female</emphasis>
(Fig.
<figureCitation id="49FC6069F1AA0A067B1C8E8EABBC5C6B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Eutimesius ephippiatus: A. Male habitus, dorsal view (ICN-Ao - 1492); B. Female habitus, dorsal view (ICN-Ao - 18); C. Male habitus, lateral view (ICN-Ao - 1492). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure9" httpUri="https://binary.pensoft.net/fig/1072637">9 B</figureCitation>
). Similar to male, except by abdominal scutum wider posteriorly; spots on area II, anterior and lateral margins; chelicerae not swollen; ornamentation of leg IV conspicuously less developed; basitarsus I not swollen. Chelicerae dark yellow (88), pedipalps, trochanters I – III, and femora I – III strong yellow (84), Trochanter IV strong yellowish brown (74), and femur IV strong yellow (84).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENGBG" type="distribution">
<paragraph id="83414E18EDCE5269D84BFB2BB1271E8E">
<heading id="856078C9E218236FAA83A92A79571E32" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="12D9DC1FCB0DC1A3DDE6D57DAF55B2D7">
<collectingCountry id="939F505C7D3AB6586EDD6B81967BB9B2" name="Colombia">Colombia</collectingCountry>
,
<collectingRegion id="3A34388680EF17A9AA267F7F54A3B724" country="Colombia" name="Casanare">Casanare</collectingRegion>
,
<collectingRegion id="9EA0EF55FE89399C391B05CFFBBA4426" country="Colombia" name="Cundinamarca">Cundinamarca</collectingRegion>
,
<collectingRegion id="8775CBFFD5931B4A7778372765A3C607" country="Colombia" name="Quindio">Quindio</collectingRegion>
(Fig.
<figureCitation id="6A5894745760D953B6162117FAFBAC3F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map showing the distribution of Eutimesius." figureDoi="10.3897/zse.100.120207.figure2" httpUri="https://binary.pensoft.net/fig/1072630">2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

177
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<mods:title>DNA barcodes reveal 63 overlooked species of Canadian beetles (Insecta, Coleoptera)</mods:title>
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<mods:namePart>Pentinsaari, Mikko</mods:namePart>
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<mods:namePart>Douglas, Hume</mods:namePart>
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<treatment ID-GBIF-Taxon="160631320" LSID="urn:lsid:plazi:treatment:A2A7C7EFBD70555C9C87705C991415B2" httpUri="http://treatment.plazi.org/id/A2A7C7EFBD70555C9C87705C991415B2" lastPageNumber="53" pageId="0" pageNumber="53">
<subSubSection pageId="0" pageNumber="53" type="nomenclature">
<paragraph pageId="0" pageNumber="53">
<taxonomicName LSID="A2A7C7EF-BD70-555C-9C87-705C991415B2" authority="Kraatz, 1853" authorityName="Kraatz" authorityYear="1853" class="Insecta" family="Staphylinidae" genus="Myllaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myllaena infuscata" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="infuscata">Myllaena infuscata Kraatz, 1853</taxonomicName>
<figureCitation captionStart="Figures 7, 8" captionStartId="F5" captionText="Figures 7, 8. 7 A Atheta vaga (Heer), habitus 7 B Atheta vaga, aedeagus, lateral view 7 C Atheta fanatica Casey, aedeagus, lateral view 7 D Atheta fanatica, spermatheca 8 Myllaena infuscata Kraatz 8 A habitus 8 B aedeagus, ventral view 8 C aedeagus, lateral view 8 D spermatheca 8 B-D V. Assing. Scale bars: 1.0 mm (7 A), 0.2 mm (7 B-D), 0.5 mm (8 A), 0.1 mm (8 B-D)." figureDoi="10.3897/zookeys.894.37862.figures7-8" httpUri="https://binary.pensoft.net/fig/362141" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures7-8">Figure 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="distribution">
<paragraph pageId="0" pageNumber="53">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="53">
Native to the western Palaearctic region, widely distributed in Europe but rare in the north (
<bibRefCitation author="Palm, T" journalOrPublisher="Entomologiska Foereningen i Stockholm, Stockholm" pageId="0" pageNumber="53" refId="B160" refString="Palm, T, 1968. Svensk Insektfauna 9. Skalbaggar. Coleoptera, Kortvingar: Fam. Staphylinidae, Haefte 5. Underfam. Aleocharinae (Deinopsis-Trichomicra). Entomologiska Foereningen i Stockholm, Stockholm" title="Svensk Insektfauna 9. Skalbaggar. Coleoptera, Kortvingar: Fam. Staphylinidae, Haefte 5. Underfam. Aleocharinae (Deinopsis-Trichomicra)." year="1968">Palm 1968</bibRefCitation>
) and also reported from India (Kashmir) (
<bibRefCitation DOI="https://doi.org/10.1163/9789004296855" author="Schuelke, M" editor="Loebl, I" journalOrPublisher="PhD Thesis, Michigan State University, Department of Entomology" pageId="0" pageNumber="53" refId="B187" refString="Schuelke, M, Smetana, A, Loebl, I, Loebl, D, 2015. 2). Brill, Leiden.https://doi.org/10.1163/9789004296855" title="2). Brill, Leiden" url="https://doi.org/10.1163/9789004296855" year="2015">
<normalizedToken originalValue="Schülke">Schuelke</normalizedToken>
and Smetana 2015
</bibRefCitation>
), though many of these records need confirmation (
<bibRefCitation author="Assing, V" journalOrPublisher="Linzer biologische Beitraege" pageId="0" pageNumber="53" pagination="1015 - 1032" refId="B11" refString="Assing, V, 2018. On some Myllaena species in the East Mediterranean and Caucasus regions (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beitraege 50: 1015 - 1032" title="On some Myllaena species in the East Mediterranean and Caucasus regions (Coleoptera: Staphylinidae: Aleocharinae)." volume="50" year="2018">Assing 2018</bibRefCitation>
). It has previously been reported as occurring in the Nearctic region by
<bibRefCitation DOI="https://doi.org/10.1163/9789004296855" author="Schuelke, M" editor="Loebl, I" journalOrPublisher="PhD Thesis, Michigan State University, Department of Entomology" pageId="0" pageNumber="53" refId="B187" refString="Schuelke, M, Smetana, A, Loebl, I, Loebl, D, 2015. 2). Brill, Leiden.https://doi.org/10.1163/9789004296855" title="2). Brill, Leiden" url="https://doi.org/10.1163/9789004296855" year="2015">
<normalizedToken originalValue="Schülke">Schuelke</normalizedToken>
and Smetana (2015)
</bibRefCitation>
but this is probably based on an erroneous synonymy of North American
<taxonomicName lsidName="M. immunda" pageId="0" pageNumber="53" rank="species" species="immunda">
<emphasis italics="true" pageId="0" pageNumber="53">M. immunda</emphasis>
</taxonomicName>
Casey with this species in older literature.
<bibRefCitation DOI="https://doi.org/10.4039/Ent114181-3" author="Klimaszewski, J" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="53" pagination="181 - 242" refId="B106" refString="Klimaszewski, J, 1982. Studies of Myllaenini (Coleoptera: Staphylinidae, Aleocharinae). 1. Systematics, phylogeny, and zoogeography of Nearctic Myllaena Erichson. The Canadian Entomologist 114: 181 - 242" title="Studies of Myllaenini (Coleoptera: Staphylinidae, Aleocharinae). 1. Systematics, phylogeny, and zoogeography of Nearctic Myllaena Erichson." url="https://doi.org/10.4039/Ent114181-3" volume="114" year="1982">Klimaszewski (1982)</bibRefCitation>
corrected the synonymy, and
<taxonomicName lsidName="M. immunda" pageId="0" pageNumber="53" rank="species" species="immunda">
<emphasis italics="true" pageId="0" pageNumber="53">M. immunda</emphasis>
</taxonomicName>
is now considered a synonym of
<taxonomicName lsidName="M. arcana" pageId="0" pageNumber="53" rank="species" species="arcana">
<emphasis italics="true" pageId="0" pageNumber="53">M. arcana</emphasis>
</taxonomicName>
Casey, 1911. The true
<taxonomicName lsidName="M. infuscata" pageId="0" pageNumber="53" rank="species" species="infuscata">
<emphasis italics="true" pageId="0" pageNumber="53">M. infuscata</emphasis>
</taxonomicName>
is reported here as adventive in the Nearctic region (Ontario, Canada).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="canadian records">
<paragraph pageId="0" pageNumber="53">Canadian records.</paragraph>
<paragraph pageId="0" pageNumber="53">Ontario: Rouge National Urban Park, 15-Sep-2013 (1 ex, CBG).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="diagnostic information">
<paragraph pageId="0" pageNumber="53">Diagnostic information.</paragraph>
<paragraph pageId="0" pageNumber="53">
Body length 1.2-1.5 mm. Habitus as in
<figureCitation captionStart="Figures 7, 8" captionStartId="F5" captionText="Figures 7, 8. 7 A Atheta vaga (Heer), habitus 7 B Atheta vaga, aedeagus, lateral view 7 C Atheta fanatica Casey, aedeagus, lateral view 7 D Atheta fanatica, spermatheca 8 Myllaena infuscata Kraatz 8 A habitus 8 B aedeagus, ventral view 8 C aedeagus, lateral view 8 D spermatheca 8 B-D V. Assing. Scale bars: 1.0 mm (7 A), 0.2 mm (7 B-D), 0.5 mm (8 A), 0.1 mm (8 B-D)." figureDoi="10.3897/zookeys.894.37862.figures7-8" httpUri="https://binary.pensoft.net/fig/362141" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures7-8">Fig. 8A</figureCitation>
. Aedeagus as in
<figureCitation captionStart="Figures 7, 8" captionStartId="F5" captionText="Figures 7, 8. 7 A Atheta vaga (Heer), habitus 7 B Atheta vaga, aedeagus, lateral view 7 C Atheta fanatica Casey, aedeagus, lateral view 7 D Atheta fanatica, spermatheca 8 Myllaena infuscata Kraatz 8 A habitus 8 B aedeagus, ventral view 8 C aedeagus, lateral view 8 D spermatheca 8 B-D V. Assing. Scale bars: 1.0 mm (7 A), 0.2 mm (7 B-D), 0.5 mm (8 A), 0.1 mm (8 B-D)." figureDoi="10.3897/zookeys.894.37862.figures7-8" httpUri="https://binary.pensoft.net/fig/362141" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures7-8">Fig. 8B, C</figureCitation>
. Spermatheca as in
<figureCitation captionStart="Figures 7, 8" captionStartId="F5" captionText="Figures 7, 8. 7 A Atheta vaga (Heer), habitus 7 B Atheta vaga, aedeagus, lateral view 7 C Atheta fanatica Casey, aedeagus, lateral view 7 D Atheta fanatica, spermatheca 8 Myllaena infuscata Kraatz 8 A habitus 8 B aedeagus, ventral view 8 C aedeagus, lateral view 8 D spermatheca 8 B-D V. Assing. Scale bars: 1.0 mm (7 A), 0.2 mm (7 B-D), 0.5 mm (8 A), 0.1 mm (8 B-D)." figureDoi="10.3897/zookeys.894.37862.figures7-8" httpUri="https://binary.pensoft.net/fig/362141" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures7-8">Fig. 8D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="bionomic notes">
<paragraph pageId="0" pageNumber="53">Bionomic notes.</paragraph>
<paragraph pageId="0" pageNumber="53">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Myllaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myllaena infuscata" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis italics="true" pageId="0" pageNumber="53">Myllaena infuscata</emphasis>
</taxonomicName>
occurs in both exposed and shaded microhabitats along the margins of still and running water (
<bibRefCitation author="Reid, CAM" journalOrPublisher="Entomologist's Monthly Magazine" pageId="0" pageNumber="53" pagination="1 - 24" refId="B175" refString="Reid, CAM, 1991. Notes on distribution of Myllaena spp. (Col., Staphylinidae) in Buckinghamshire and north-east England. Entomologist's Monthly Magazine 127: 1 - 24" title="Notes on distribution of Myllaena spp. (Col., Staphylinidae) in Buckinghamshire and north-east England." volume="127" year="1991">Reid 1991</bibRefCitation>
). The Canadian specimen was extracted from soil and leaf litter collected near the mouth of Rouge River.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="53" type="comments">
<paragraph pageId="0" pageNumber="53">Comments.</paragraph>
<paragraph pageId="0" pageNumber="53">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Myllaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myllaena infuscata" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis italics="true" pageId="0" pageNumber="53">Myllaena infuscata</emphasis>
</taxonomicName>
is distinctive in the Nearctic fauna for its spermathecal shape, which forms concentric circular coils (
<figureCitation captionStart="Figures 7, 8" captionStartId="F5" captionText="Figures 7, 8. 7 A Atheta vaga (Heer), habitus 7 B Atheta vaga, aedeagus, lateral view 7 C Atheta fanatica Casey, aedeagus, lateral view 7 D Atheta fanatica, spermatheca 8 Myllaena infuscata Kraatz 8 A habitus 8 B aedeagus, ventral view 8 C aedeagus, lateral view 8 D spermatheca 8 B-D V. Assing. Scale bars: 1.0 mm (7 A), 0.2 mm (7 B-D), 0.5 mm (8 A), 0.1 mm (8 B-D)." figureDoi="10.3897/zookeys.894.37862.figures7-8" httpUri="https://binary.pensoft.net/fig/362141" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures7-8">Fig. 8D</figureCitation>
). The median lobe of the aedeagus in lateral view is also distinctive among species in North America (
<figureCitation captionStart="Figures 7, 8" captionStartId="F5" captionText="Figures 7, 8. 7 A Atheta vaga (Heer), habitus 7 B Atheta vaga, aedeagus, lateral view 7 C Atheta fanatica Casey, aedeagus, lateral view 7 D Atheta fanatica, spermatheca 8 Myllaena infuscata Kraatz 8 A habitus 8 B aedeagus, ventral view 8 C aedeagus, lateral view 8 D spermatheca 8 B-D V. Assing. Scale bars: 1.0 mm (7 A), 0.2 mm (7 B-D), 0.5 mm (8 A), 0.1 mm (8 B-D)." figureDoi="10.3897/zookeys.894.37862.figures7-8" httpUri="https://binary.pensoft.net/fig/362141" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures7-8">Fig. 8B, C</figureCitation>
). Within the western Palaearctic fauna, only
<taxonomicName lsidName="M. minuta" pageId="0" pageNumber="53" rank="species" species="minuta">
<emphasis italics="true" pageId="0" pageNumber="53">M. minuta</emphasis>
</taxonomicName>
has similar genitalia but differs in the shape of the median lobe in lateral view. The single female from Ontario bears the characteristic spermatheca and its corresponding barcode sequence falls within the BIN associated with
<taxonomicName lsidName="M. infuscata" pageId="0" pageNumber="53" rank="species" species="infuscata">
<emphasis italics="true" pageId="0" pageNumber="53">M. infuscata</emphasis>
</taxonomicName>
, rather than
<taxonomicName lsidName="M. minuta" pageId="0" pageNumber="53" rank="species" species="minuta">
<emphasis italics="true" pageId="0" pageNumber="53">M. minuta</emphasis>
</taxonomicName>
, its similar sister species. It is unknown whether this is a recent introduction to North America or if its small size has impeded its detection.
</paragraph>
</subSubSection>
</treatment>
</document>