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<figureCitation id="1325DFF91362531FFD29FB87FCDDFB1B" box="[728,859,1149,1176]" captionStart="Figs 4550" captionStartId="43.[189,241,1887,1913]" captionTargetBox="[271,1311,274,1831]" captionTargetId="figure@43.[264,1324,265,1845]" captionTargetPageId="43" captionText="Figs 4550. Dyscolus spp., habitus (4547) and aedeagus, median lobe in lateral view (4850). 45. D. denigratus (Bates, 1891), male specimen from Guamaní. 46. D. denigratus, male specimen from Guagua Pichincha (type locality). 47. D. palatus Moret, 1998, male paratype from Atacazo. 48 50. D. denigratus. 48. From Guamaní. 49. From Cotacachi. 50. From Guagua Pichincha." figureDoi="http://doi.org/10.5281/zenodo.3829708" httpUri="https://zenodo.org/record/3829708/files/figure.png" pageId="42" pageNumber="43">Figs 4550</figureCitation>
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<emphasis id="B96A1F6E1362531FFF4CFA72FE2AFA21" box="[189,428,1416,1442]" italics="true" pageId="42" pageNumber="43">Dyscolus denigratus</emphasis>
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complex cannot be resolved from the COI sequences alone.
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We thus take a conservative approach, with no taxonomic decision regarding the specific or subspecific status of the components of the
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<mods:nameIdentifier id="E738F48FE9585F79DD4612DCC717CF3B" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
<mods:namePart id="B50E7EC5F6EFF3C6C670DC9D11A19D59">Moret, Pierre</mods:namePart>
<mods:nameIdentifier id="B6995FC5B9E81149FB07A399C2FB2923" type="LSID">2E194645-D255-438B-819E-A2B6C39AD699</mods:nameIdentifier>
<mods:affiliation id="6B71C92CB4EE787063E1746FB9F6EFC8">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="52F457C62BE8358F06D8E9454E4D8FFF" type="email">pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="5C94C8E513C1FCB4342991D133520D8D" type="personal">
<mods:role id="AD78C7ADDF3436969B6F8FCF1944D7B0">
<mods:roleTerm id="00D514746E100E0A0274F77F11E5B7B5">Author</mods:roleTerm>
<mods:name id="7138E8375F5721B5D92AFEC18CE7CCBF" type="personal">
<mods:role id="5A9D360C6CB78D36B8CAC22DA6345509">
<mods:roleTerm id="857D31C77B810C134D4792E6A4040B2C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8BAFD6094BF1FD2753AFDF76C4D3DC4A">Murienne, Jérôme</mods:namePart>
<mods:nameIdentifier id="6D4F3B1E7B957DEC8F6B2F9802712DC7" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
<mods:affiliation id="16008C9EED72FF49D929B68D1CA0A203">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
<mods:nameIdentifier id="4D623EEF6422D05D935D32947AC98607" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
<mods:namePart id="39D71E5A6FFF43B59323642FD004E88E">Murienne, Jérôme</mods:namePart>
<mods:nameIdentifier id="DF0A69B79F3F4EC180DD43985A766460" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
<mods:affiliation id="97E23F5E49D368F12913E7A07ED9F4D1">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
<mods:nameIdentifier id="15A3071BF5481A3BF97B31439A85DBA3" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="C4A619AA94ECBA64B0671B6D83356925">text</mods:typeOfResource>
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<mods:titleInfo id="8B79F66EB5C6073A33390E02DA856366">
<mods:title id="B9253E394DB329D4C1A94BC523DCFB42">European Journal of Taxonomy</mods:title>
<mods:typeOfResource id="4AC55D4F165D05B23A975A906789F02A">text</mods:typeOfResource>
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<mods:titleInfo id="12DA830578FC7DB2D0646DC1BE44C202">
<mods:title id="257217143F1529C29DD1FEDA2E35544B">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="A8AFAA1DA0C21BB0AB0E5723913B3A10">
<mods:date id="AA28D8130D692268B53AD615A8ADF97E">2020</mods:date>
<mods:detail id="DFCD56E22885DD66E6B32051DA94AD34" type="pubDate">
<mods:number id="EC901A2F6DD5B40B0F03BB368075ACFE">2020-05-15</mods:number>
<mods:part id="97255E7B53BCCC3860CC90548ECDD79D">
<mods:date id="8A508CCB0EA92137F9C92B8EC1922D30">2020</mods:date>
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<mods:number id="0DF50973722BD3F4CB6942CAF2060DA3">2020-05-15</mods:number>
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<mods:number id="5DC1EDECD8A8387680592FFB8C75A484">646</mods:number>
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<treatment id="03B7726A13685314FD9AFC5CFEEBFA9E" ID-DOI="http://doi.org/10.5281/zenodo.3848409" ID-GBIF-Taxon="164160396" ID-Zenodo-Dep="3848409" LSID="urn:lsid:zoobank.org:act:811C080F-1087-49A8-BB75-7B71DA04C593" httpUri="http://treatment.plazi.org/id/03B7726A13685314FD9AFC5CFEEBFA9E" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<subSubSection id="C30490F713685315FD9AFC5CFB47FC66" pageId="32" pageNumber="33" type="nomenclature">
<treatment id="03B7726A13685314FD9AFC5CFEEBFA9E" ID-DOI="http://doi.org/10.5281/zenodo.3848409" ID-GBIF-Taxon="164160396" ID-Zenodo-Dep="3848409" LSID="urn:lsid:zoobank.org:act:811C080F-1087-49A8-BB75-7B71DA04C593" httpUri="http://treatment.plazi.org/id/03B7726A13685314FD9AFC5CFEEBFA9E" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" scope_class="Insecta" scope_family="Carabidae" scope_order="Coleoptera" scope_subFamily="Harpalinae" scope_tribe="Platynini">
<subSubSection id="C30490F713685315FD9AFC5CFC4EFC42" box="[619,968,934,961]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BA1C37C13685315FD9AFC5CFC4EFC42" blockId="32.[370,1217,934,1032]" box="[619,968,934,961]" pageId="32" pageNumber="33">
<heading id="D0E9741013685315FD9AFC5CFC4EFC42" box="[619,968,934,961]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="33" reason="2">
<taxonomicName id="4C1EB8FF13685315FD9AFC5CFCE1FC42" ID-CoL="8M569" authority="Moret" authorityName="Moret" authorityYear="2020" box="[619,871,934,961]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ruizi" status="sp. nov.">
@ -58,14 +58,14 @@ Moret
<taxonomicNameLabel id="A259A21513685315FC9FFC5DFC4EFC42" box="[878,968,935,961]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30490F713685315FE83FC30FCDDFB8B" pageId="32" pageNumber="33" type="description">
<paragraph id="8BA1C37C13685315FE83FC30FB47FC66" blockId="32.[370,1217,934,1032]" box="[370,1217,970,997]" pageId="32" pageNumber="33">
<uri id="FF8FCF7E13685315FE83FC30FB47FC66" box="[370,1217,970,997]" pageId="32" pageNumber="33">
urn:lsid:zoobank.org:act:
<uuid id="FFB8F9A913685315FD65FC30FB47FC66" box="[660,1217,970,997]" pageId="32" pageNumber="33">811C080F-1087-49A8-BB75-7B71DA04C593</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30490F713685315FD29FC17FCDDFB8B" box="[728,859,1005,1032]" pageId="32" pageNumber="33" type="description">
<paragraph id="8BA1C37C13685315FD29FC17FCDDFB8B" blockId="32.[370,1217,934,1032]" box="[728,859,1005,1032]" pageId="32" pageNumber="33">
<figureCitation id="1325DFF913685315FD29FC17FCDDFB8B" box="[728,859,1005,1032]" captionStart="Figs 3235. 3233" captionStartId="34.[189,241,1915,1941]" captionTargetBox="[199,1386,351,1780]" captionTargetId="figure@34.[189,1398,334,1792]" captionTargetPageId="34" captionText="Figs 3235. 3233. Dyscolus salazarae Moret sp. nov. 32. Habitus of the male holotype (MNHN, COI voucher PM392-09). 33. Aedeagus, median lobe in lateral view. 3435. D. ruizi Moret sp. nov. 34. Habitus of a male paratype from Saraguro (CPM). 35. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829700" httpUri="https://zenodo.org/record/3829700/files/figure.png" pageId="32" pageNumber="33">Figs 3435</figureCitation>
</paragraph>
@ -86,7 +86,7 @@ the biodiversity of carabid beetles along an elevational gradient between
and Zamora.
</paragraph>
</subSubSection>
<subSubSection id="C30490F713685314FF4CFB1FFEEBFA9E" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="materials_examined">
<subSubSection id="C30490F713685315FF4CFB1FFA92F86A" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BA1C37C13685315FF4CFB1FFEEAFB7C" blockId="32.[189,364,1253,1279]" box="[189,364,1253,1279]" pageId="32" pageNumber="33">
<heading id="D0E9741013685315FF4CFB1FFEEAFB7C" bold="true" box="[189,364,1253,1279]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B96A1F6E13685315FF4CFB1FFEEAFB7C" bold="true" box="[189,364,1253,1279]" pageId="32" pageNumber="33">Type material</emphasis>
@ -100,7 +100,9 @@ and Zamora.
<paragraph id="8BA1C37C13685315FF4CFACDFDACFA14" blockId="32.[189,1399,1300,1432]" pageId="32" pageNumber="33">
<materialsCitation id="3B76C92113685315FF4CFACDFDACFA14" ID-GBIF-Occurrence="2620303419" collectingDate="2015-03-17" collectionCode="QCAZ" collectorName="P. Moret &amp; C. Ruiz" country="Ecuador" county="Parque Nacional Podocarpus" elevation="2900" latitude="-4.1173334" location="Waypoint 397" longLatPrecision="1" longitude="-79.16841" municipality="Cajanuma" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Loja Province" typeStatus="holotype">
<collectingCountry id="F30983EC13685315FF4CFACDFECDFAD2" box="[189,331,1335,1361]" name="Ecuador" pageId="32" pageNumber="33">ECUADOR</collectingCountry>
• ♂;
<specimenCount id="9D1808F513685315FE98FACFFE06FAD2" box="[361,384,1333,1361]" pageId="32" pageNumber="33" type="male"></specimenCount>
;
<collectingRegion id="49DA0D9E13685315FE60FACDFDBAFAD2" box="[401,572,1335,1361]" country="Ecuador" name="Loja" pageId="32" pageNumber="33">Loja Province</collectingRegion>
,
<collectingCounty id="62C0BBF013685315FDBBFACDFC23FAD2" box="[586,933,1335,1361]" pageId="32" pageNumber="33">Parque Nacional Podocarpus</collectingCounty>
@ -139,7 +141,7 @@ leg.; COI voucher PM392-68, BOLD sequence SUM056-18;
)
</paragraph>
<paragraph id="8BA1C37C13685315FF4CFA18FA92F86A" blockId="32.[189,1399,1469,2025]" pageId="32" pageNumber="33">
<materialsCitation id="3B76C92113685315FF4CFA18FD2DF99C" ID-GBIF-Occurrence="2620303423" collectingDate="2013-09-24" collectionCode="CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2920" latitude="-4.113" location="Cajanuma Sendero" longLatPrecision="78" longitude="-79.174" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Loja Province">
<materialsCitation id="3B76C92113685315FF4CFA18FD2DF99C" ID-GBIF-Occurrence="2620303423" collectingDate="2013-09-24" collectionCode="CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2920" latitude="-4.113" location="Cajanuma Sendero" longLatPrecision="78" longitude="-79.174" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Loja Province" typeStatus="paratype">
<collectingCountry id="F30983EC13685315FF4CFA18FECDFA7F" box="[189,331,1506,1532]" name="Ecuador" pageId="32" pageNumber="33">ECUADOR</collectingCountry>
<emphasis id="B96A1F6E13685315FE9DFA18FD98FA7F" bold="true" box="[364,542,1506,1532]" pageId="32" pageNumber="33">
@ -170,16 +172,14 @@ leg.;
<collectionCode id="ED0F5BB913685315FDA9F9FFFD2DF99C" box="[600,683,1541,1567]" pageId="32" pageNumber="33">CISEC</collectionCode>
</materialsCitation>
<materialsCitation id="3B76C92113685315FD38F9FFFD69F9C1" ID-GBIF-Occurrence="2620303407" collectingDate="2013-09-24" collectionCode="GBNP, QCAZ" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2920" latitude="-4.113" location="Cajanuma Sendero" longLatPrecision="78" longitude="-79.174" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F513685315FD38F9FFFD7FF99C" box="[713,761,1539,1567]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
; same collection data as for preceding; COI voucher CR120, BOLD sequence
<collectionCode id="ED0F5BB913685315FE1CF9D2FDC7F9C1" box="[493,577,1576,1602]" collectionName="GBNP" pageId="32" pageNumber="33">GBNP</collectionCode>
068-17;
<materialsCitation id="3B76C92113685315FD38F9FFFD69F9C1" ID-GBIF-Occurrence="2620303407" collectingDate="2013-09-24" collectionCode="QCAZ" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2920" latitude="-4.113" location="Cajanuma Sendero" longLatPrecision="78" longitude="-79.174" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Loja" typeStatus="paratype">
<specimenCount id="9D1808F513685315FD38F9FFFD7FF99C" box="[713,761,1539,1567]" count="1" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
; same collection data as for preceding; COI voucher CR120, BOLD sequence GBNP068-17;
<collectionCode id="ED0F5BB913685315FD6EF9D2FD69F9C1" box="[671,751,1576,1602]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="32" pageNumber="33" type="Museum">QCAZ</collectionCode>
</materialsCitation>
<materialsCitation id="3B76C92113685315FCFAF9D2FAE5F9E6" ID-GBIF-Occurrence="2620303345" collectingDate="2013-07-11" collectionCode="GBNP, CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2838" latitude="-3.993" location="El Tiro" longLatPrecision="78" longitude="-79.147" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F513685315FCFAF9D2FCBFF9C1" box="[779,825,1574,1602]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
<materialsCitation id="3B76C92113685315FCFAF9D2FAE5F9E6" ID-GBIF-Occurrence="2620303345" collectingDate="2013-07-11" collectionCode="CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2838" latitude="-3.993" location="El Tiro" longLatPrecision="78" longitude="-79.147" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Loja" typeStatus="paratype">
<specimenCount id="9D1808F513685315FCFAF9D2FCBFF9C1" box="[779,825,1574,1602]" count="1" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
;
<location id="8EC195A713685315FCB8F9D2FC27F9C1" LSID="urn:lsid:plazi:treatment:03B7726A13685314FD9AFC5CFEEBFA9E:8EC195A713685315FCB8F9D2FC27F9C1" box="[841,929,1576,1602]" country="Ecuador" latitude="-3.993" longLatPrecision="78" longitude="-79.147" name="El Tiro" pageId="32" pageNumber="33" stateProvince="Loja">El Tiro</location>
;
@ -199,16 +199,14 @@ a.s.l.
<collectorName id="26EBA6AA13685315FECDF9B6FE2CF9E6" box="[316,426,1611,1638]" pageId="32" pageNumber="33">D. Marín</collectorName>
and
<collectorName id="26EBA6AA13685315FE17F9B1FDC7F9E6" box="[486,577,1611,1637]" pageId="32" pageNumber="33">C. Ruiz</collectorName>
leg.; COI voucher CR008, BOLD sequence
<collectionCode id="ED0F5BB913685315FBACF9B1FB36F9E6" box="[1117,1200,1611,1637]" collectionName="GBNP" pageId="32" pageNumber="33">GBNP</collectionCode>
006-17;
leg.; COI voucher CR008, BOLD sequence GBNP006-17;
<collectionCode id="ED0F5BB913685315FAE1F9B1FAE5F9E6" box="[1296,1379,1611,1637]" collectionName="CISEC" pageId="32" pageNumber="33">CISEC</collectionCode>
</materialsCitation>
<materialsCitation id="3B76C92113685315FF4CF995FAF1F90A" ID-GBIF-Occurrence="2620303396" box="[189,1399,1645,1673]" collectingDate="2005-12-10" collectionCode="CISEC" collectorName="I. Mas" country="Ecuador" elevation="2920" location="Huashapamba" municipality="Saraguro" pageId="32" pageNumber="33" specimenCode="CISEC 0003240, 0003271" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F513685315FF4CF995FF6FF90A" box="[189,233,1645,1673]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
<materialsCitation id="3B76C92113685315FF4CF995FAF1F90A" ID-GBIF-Occurrence="2620303396" box="[189,1399,1645,1673]" collectingDate="2005-12-10" collectionCode="CISEC" collectorName="I. Mas" country="Ecuador" elevation="2920" location="Huashapamba" municipality="Saraguro" pageId="32" pageNumber="33" specimenCode="CISEC 0003240, 0003271" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loja" typeStatus="paratype">
<specimenCount id="9D1808F513685315FF4CF995FF6FF90A" box="[189,233,1645,1673]" count="1" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1808F513685315FF06F995FEA2F90A" box="[247,292,1647,1673]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
<specimenCount id="9D1808F513685315FF06F995FEA2F90A" box="[247,292,1647,1673]" count="1" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="6BC5590613685315FEC2F995FE26F90A" box="[307,416,1647,1673]" pageId="32" pageNumber="33">Saraguro</collectingMunicipality>
,
@ -230,20 +228,20 @@ leg.;
<specimenCode id="DBB86B0713685315FAE1F995FAF1F90A" box="[1296,1399,1647,1673]" collectionCode="CISEC" pageId="32" pageNumber="33">0003271</specimenCode>
</materialsCitation>
<materialsCitation id="3B76C92113685315FF21F968FC9BF92F" ID-GBIF-Occurrence="2620303389" box="[208,797,1680,1708]" collectingDate="2005-12-10" collectionCode="CPM" collectorName="I. Mas" country="Ecuador" elevation="2920" location="Huashapamba" municipality="Saraguro" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<specimenCount id="9D1808F513685315FF21F968FF79F92F" box="[208,255,1680,1708]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
<materialsCitation id="3B76C92113685315FF21F968FC9BF92F" ID-GBIF-Occurrence="2620303389" box="[208,797,1680,1708]" collectingDate="2005-12-10" collectionCode="CPM" collectorName="I. Mas" country="Ecuador" elevation="2920" location="Huashapamba" municipality="Saraguro" pageId="32" pageNumber="33" specimenCode="CISEC 0003240, 0003271" specimenCount="1" specimenCount-male="1" stateProvince="Loja" typeStatus="paratype">
<specimenCount id="9D1808F513685315FF21F968FF79F92F" box="[208,255,1680,1708]" count="1" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED0F5BB913685315FD2AF968FC9BF92F" box="[731,797,1682,1708]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="32" pageNumber="33">CPM</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B76C92113685315FCB0F968FB25F94C" ID-GBIF-Occurrence="2620303324" collectingDate="2013-09-05" collectionCode="CISEC" collectorName="El Tiro-Cima &amp; D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2838" latitude="-3.993" location="Zamora Province" longLatPrecision="78" longitude="-79.147" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Loja">
<materialsCitation id="3B76C92113685315FCB0F968FB25F94C" ID-GBIF-Occurrence="2620303324" collectingDate="2013-09-05" collectionCode="CISEC" collectorName="El Tiro-Cima &amp; D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2838" latitude="-3.993" location="Zamora Province" longLatPrecision="78" longitude="-79.147" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Loja" typeStatus="paratype">
<emphasis id="B96A1F6E13685315FCB0F968FBF7F92F" bold="true" box="[833,1137,1682,1708]" pageId="32" pageNumber="33">
<collectingRegion id="49DA0D9E13685315FCB0F968FCFAF92F" box="[833,892,1682,1708]" country="Ecuador" name="Loja" pageId="32" pageNumber="33">Loja</collectingRegion>
/
<location id="8EC195A713685315FC64F968FBF7F92F" LSID="urn:lsid:plazi:treatment:03B7726A13685314FD9AFC5CFEEBFA9E:8EC195A713685315FC64F968FBF7F92F" box="[917,1137,1682,1708]" country="Ecuador" latitude="-3.993" longLatPrecision="78" longitude="-79.147" name="Zamora Province" pageId="32" pageNumber="33" stateProvince="Loja">Zamora Province</location>
</emphasis>
<specimenCount id="9D1808F513685315FB7FF968FB3AF92F" box="[1166,1212,1680,1708]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
<specimenCount id="9D1808F513685315FB7FF968FB3AF92F" box="[1166,1212,1680,1708]" count="1" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
;
<collectorName id="26EBA6AA13685315FB3DF968FAF5F92F" box="[1228,1395,1682,1708]" pageId="32" pageNumber="33">El TiroCima</collectorName>
;
@ -267,16 +265,16 @@ leg.;
<collectionCode id="ED0F5BB913685315FBA1F94FFB25F94C" box="[1104,1187,1717,1743]" collectionName="CISEC" pageId="32" pageNumber="33">CISEC</collectionCode>
</materialsCitation>
<materialsCitation id="3B76C92113685315FB4EF94FFD06F971" ID-GBIF-Occurrence="2620303343" collectingDate="2013-09-05" collectionCode="CPM" collectorName="El Tiro-Cima &amp; D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2838" latitude="-3.993" location="Zamora Province" longLatPrecision="78" longitude="-79.147" pageId="32" pageNumber="33" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Zamora Province">
<specimenCount id="9D1808F513685315FB4EF94FFB6AF94C" box="[1215,1260,1715,1743]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
<materialsCitation id="3B76C92113685315FB4EF94FFD06F971" ID-GBIF-Occurrence="2620303343" collectingDate="2013-09-05" collectionCode="CPM" collectorName="El Tiro-Cima &amp; D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2838" latitude="-3.993" location="Zamora Province" longLatPrecision="78" longitude="-79.147" pageId="32" pageNumber="33" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Zamora Province" typeStatus="paratype">
<specimenCount id="9D1808F513685315FB4EF94FFB6AF94C" box="[1215,1260,1715,1743]" count="1" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1808F513685315FB0DF94FFAACF953" box="[1276,1322,1717,1744]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
<specimenCount id="9D1808F513685315FB0DF94FFAACF953" box="[1276,1322,1717,1744]" count="1" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED0F5BB913685315FDB3F922FD06F971" box="[578,640,1752,1778]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="32" pageNumber="33">CPM</collectionCode>
</materialsCitation>
<materialsCitation id="3B76C92113685315FD6AF922FC8BF895" ID-GBIF-Occurrence="2620303327" collectingDate="2013-07-11" collectionCode="MNHN" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2838" latitude="-3.993" location="El Tiro-Cima" longLatPrecision="78" longitude="-79.147" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Zamora Province">
<specimenCount id="9D1808F513685315FD6AF922FD4FF971" box="[667,713,1750,1778]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
<materialsCitation id="3B76C92113685315FD6AF922FC8BF895" ID-GBIF-Occurrence="2620303327" collectingDate="2013-07-11" collectionCode="MNHN" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2838" latitude="-3.993" location="El Tiro-Cima" longLatPrecision="78" longitude="-79.147" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Zamora Province" typeStatus="paratype">
<specimenCount id="9D1808F513685315FD6AF922FD4FF971" box="[667,713,1750,1778]" count="1" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
; LZ03
<location id="8EC195A713685315FCD2F922FC4FF971" LSID="urn:lsid:plazi:treatment:03B7726A13685314FD9AFC5CFEEBFA9E:8EC195A713685315FCD2F922FC4FF971" box="[803,969,1752,1778]" country="Ecuador" latitude="-3.993" longLatPrecision="78" longitude="-79.147" name="El Tiro-Cima" pageId="32" pageNumber="33" stateProvince="Zamora Province">El TiroCima</location>
;
@ -300,8 +298,8 @@ leg.;
<collectionCode id="ED0F5BB913685315FD42F906FC8BF895" box="[691,781,1788,1814]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="32" pageNumber="33">MNHN</collectionCode>
</materialsCitation>
<materialsCitation id="3B76C92113685315FCDAF901FDE9F8BA" ID-GBIF-Occurrence="2620303307" collectingDate="2008-03-26" collectionCode="CISEC" collectorName="Marin Rosero" country="Ecuador" elevation="2813" location="El Tiro" pageId="32" pageNumber="33" specimenCode="CISEC 0002573" specimenCount="1" specimenCount-female="1" stateProvince="Zamora Province">
<specimenCount id="9D1808F513685315FCDAF901FCDDF895" box="[811,859,1787,1814]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
<materialsCitation id="3B76C92113685315FCDAF901FDE9F8BA" ID-GBIF-Occurrence="2620303307" collectingDate="2008-03-26" collectionCode="CISEC" collectorName="Marin Rosero" country="Ecuador" elevation="2813" location="El Tiro" pageId="32" pageNumber="33" specimenCode="CISEC 0002573" specimenCount="1" specimenCount-female="1" stateProvince="Zamora Province" typeStatus="paratype">
<specimenCount id="9D1808F513685315FCDAF901FCDDF895" box="[811,859,1787,1814]" count="1" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
; LZ03
<location id="8EC195A713685315FC46F901FB89F896" LSID="urn:lsid:plazi:treatment:03B7726A13685314FD9AFC5CFEEBFA9E:8EC195A713685315FC46F901FB89F896" box="[951,1039,1787,1813]" country="Ecuador" name="El Tiro" pageId="32" pageNumber="33" stateProvince="Zamora Province">El Tiro</location>
;
@ -319,12 +317,12 @@ leg.;
<specimenCode id="DBB86B0713685315FE56F8E5FDE9F8BA" box="[423,623,1822,1849]" collectionCode="CISEC" pageId="32" pageNumber="33">CISEC 0002573</specimenCode>
</materialsCitation>
.
<materialsCitation id="3B76C92113685315FD65F8E4FDBDF8FC" ID-GBIF-Occurrence="2620303380" collectingDate="2013-07-18" collectionCode="GBNP, CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2528" latitude="-3.988" location="Carretera Loja-Zamora" longLatPrecision="78" longitude="-79.118" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Zamora-Chinchipe Province">
<materialsCitation id="3B76C92113685315FD65F8E4FDBDF8FC" ID-GBIF-Occurrence="2620303380" collectingDate="2013-07-18" collectionCode="CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2528" latitude="-3.988" location="Carretera Loja-Zamora" longLatPrecision="78" longitude="-79.118" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Zamora-Chinchipe Province" typeStatus="paratype">
<emphasis id="B96A1F6E13685315FD65F8E4FC7DF8BB" bold="true" box="[660,1019,1822,1848]" pageId="32" pageNumber="33">
<collectingRegion id="49DA0D9E13685315FD65F8E4FC7DF8BB" box="[660,1019,1822,1848]" country="Ecuador" name="Zamora-Chinchipe" pageId="32" pageNumber="33">Zamora-Chinchipe Province</collectingRegion>
</emphasis>
<specimenCount id="9D1808F513685315FBE9F8E5FBC1F8BA" box="[1048,1095,1823,1849]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
<specimenCount id="9D1808F513685315FBE9F8E5FBC1F8BA" box="[1048,1095,1823,1849]" count="1" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
;
<location id="8EC195A713685315FBA9F8E5FAF5F8BB" LSID="urn:lsid:plazi:treatment:03B7726A13685314FD9AFC5CFEEBFA9E:8EC195A713685315FBA9F8E5FAF5F8BB" box="[1112,1395,1822,1849]" country="Ecuador" latitude="-3.988" longLatPrecision="78" longitude="-79.118" name="Carretera Loja-Zamora" pageId="32" pageNumber="33" stateProvince="Zamora-Chinchipe Province">Carretera LojaZamora</location>
;
@ -344,20 +342,18 @@ a.s.l.
<collectorName id="26EBA6AA13685315FD00F8B8FCDDF8DF" box="[753,859,1858,1884]" pageId="32" pageNumber="33">D. Marín</collectorName>
and
<collectorName id="26EBA6AA13685315FC60F8B8FC6FF8DF" box="[913,1001,1858,1884]" pageId="32" pageNumber="33">C. Ruiz</collectorName>
leg.; COI voucher CR002, BOLD sequence
<collectionCode id="ED0F5BB913685315FEC2F89FFE00F8FC" box="[307,390,1893,1919]" collectionName="GBNP" pageId="32" pageNumber="33">GBNP</collectionCode>
002-17;
leg.; COI voucher CR002, BOLD sequence GBNP002-17;
<collectionCode id="ED0F5BB913685315FE19F89FFDBDF8FC" box="[488,571,1893,1919]" collectionName="CISEC" pageId="32" pageNumber="33">CISEC</collectionCode>
</materialsCitation>
<materialsCitation id="3B76C92113685315FDAAF89FFD11F821" ID-GBIF-Occurrence="2620303399" collectingDate="2013-07-18" collectionCode="CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2528" latitude="-3.988" location="Carretera Loja-Zamora" longLatPrecision="78" longitude="-79.118" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Zamora-Chinchipe">
<specimenCount id="9D1808F513685315FDAAF89FFD0DF803" box="[603,651,1893,1920]" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
<materialsCitation id="3B76C92113685315FDAAF89FFD11F821" ID-GBIF-Occurrence="2620303399" collectingDate="2013-07-18" collectionCode="CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2528" latitude="-3.988" location="Carretera Loja-Zamora" longLatPrecision="78" longitude="-79.118" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
<specimenCount id="9D1808F513685315FDAAF89FFD0DF803" box="[603,651,1893,1920]" count="1" pageId="32" pageNumber="33" type="female">1 ♀</specimenCount>
; same collection data as for preceding; COI voucher CR003, BOLD sequence GBNP003-17;
<collectionCode id="ED0F5BB913685315FDB5F872FD11F821" box="[580,663,1928,1954]" collectionName="CISEC" pageId="32" pageNumber="33">CISEC</collectionCode>
</materialsCitation>
<materialsCitation id="3B76C92113685315FD49F872FB59F846" ID-GBIF-Occurrence="2620303303" collectingDate="2013-12-05" collectionCode="CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2528" latitude="-3.988" location="Carretera Loja-Zamora" longLatPrecision="78" longitude="-79.118" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Zamora-Chinchipe">
<specimenCount id="9D1808F513685315FD49F872FD6FF821" box="[696,745,1926,1954]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
<materialsCitation id="3B76C92113685315FD49F872FB59F846" ID-GBIF-Occurrence="2620303303" collectingDate="2013-12-05" collectionCode="CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2528" latitude="-3.988" location="Carretera Loja-Zamora" longLatPrecision="78" longitude="-79.118" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
<specimenCount id="9D1808F513685315FD49F872FD6FF821" box="[696,745,1926,1954]" count="1" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<date id="FFA0E5BC13685315FB29F872FAF4F821" box="[1240,1394,1928,1955]" pageId="32" pageNumber="33" value="2013-12-05">
<collectingDate id="EFE41C5413685315FB29F872FAF4F821" box="[1240,1394,1928,1955]" pageId="32" pageNumber="33" value="2013-12-05">5 Dec. 2013</collectingDate>
@ -370,13 +366,15 @@ leg.; COI voucher CR001, BOLD sequence GBNP001-17;
<collectionCode id="ED0F5BB913685315FB7DF851FB59F846" box="[1164,1247,1963,1989]" collectionName="CISEC" pageId="32" pageNumber="33">CISEC</collectionCode>
</materialsCitation>
<materialsCitation id="3B76C92113685315FB0DF851FA96F86A" ID-GBIF-Occurrence="2620303371" collectingDate="2013-12-05" collectionCode="CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2528" latitude="-3.988" location="Carretera Loja-Zamora" longLatPrecision="78" longitude="-79.118" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Zamora-Chinchipe">
<specimenCount id="9D1808F513685315FB0DF851FAACF845" box="[1276,1322,1962,1990]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
<materialsCitation id="3B76C92113685315FB0DF851FA96F86A" ID-GBIF-Occurrence="2620303371" collectingDate="2013-12-05" collectionCode="CISEC" collectorName="D. Marin &amp; C. Ruiz" country="Ecuador" elevation="2528" latitude="-3.988" location="Carretera Loja-Zamora" longLatPrecision="78" longitude="-79.118" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Zamora-Chinchipe" typeStatus="paratype">
<specimenCount id="9D1808F513685315FB0DF851FAACF845" box="[1276,1322,1962,1990]" count="1" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
; same collection data as for preceding; COI voucher CR055, BOLD sequence GBNP029-17;
<collectionCode id="ED0F5BB913685315FB48F835FA96F86A" box="[1209,1296,1999,2025]" collectionName="CISEC" pageId="32" pageNumber="33">CISEC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30490F713695314FF4CFEF1FC8CFD54" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BA1C37C13695314FF4CFEF1FE52FEA6" blockId="33.[189,468,267,293]" box="[189,468,267,293]" pageId="33" pageNumber="34">
<heading id="D0E9741013695314FF4CFEF1FE52FEA6" bold="true" box="[189,468,267,293]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B96A1F6E13695314FF4CFEF1FE52FEA6" bold="true" box="[189,468,267,293]" pageId="33" pageNumber="34">Diagnostic description</emphasis>
@ -397,6 +395,8 @@ Habitus:
<emphasis id="B96A1F6E13695314FE48FD47FD06FD54" box="[441,640,701,727]" italics="true" pageId="33" pageNumber="34">Female genitalia</emphasis>
: unstudied.
</paragraph>
</subSubSection>
<subSubSection id="C30490F713695314FF4CFD04FA9DFC74" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8BA1C37C13695314FF4CFD04FEE5FC9B" blockId="33.[189,355,766,792]" box="[189,355,766,792]" pageId="33" pageNumber="34">
<heading id="D0E9741013695314FF4CFD04FEE5FC9B" bold="true" box="[189,355,766,792]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B96A1F6E13695314FF4CFD04FEE5FC9B" bold="true" box="[189,355,766,792]" pageId="33" pageNumber="34">Comparisons</emphasis>
@ -463,6 +463,8 @@ differs from
</taxonomicName>
by its elongate pronotum and the absence of the middle seta on the third interval.
</paragraph>
</subSubSection>
<subSubSection id="C30490F713695314FF4CFBE4FA82FB09" pageId="33" pageNumber="34" type="biology_ecology">
<paragraph id="8BA1C37C13695314FF4CFBE4FE9BFBBB" blockId="33.[189,285,1054,1080]" box="[189,285,1054,1080]" pageId="33" pageNumber="34">
<heading id="D0E9741013695314FF4CFBE4FE9BFBBB" bold="true" box="[189,285,1054,1080]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B96A1F6E13695314FF4CFBE4FE9BFBBB" bold="true" box="[189,285,1054,1080]" pageId="33" pageNumber="34">Habitat</emphasis>
@ -475,6 +477,8 @@ Upper montane forest and subpáramo on the eastern slope of the
<quantity id="4CE66E9913695314FB65FBB7FABEFBE4" box="[1172,1336,1101,1127]" metricMagnitude="3" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="33" pageNumber="34" unit="m" value="2900.0" valueMax="3000.0" valueMin="2800.0">28003000 m</quantity>
a.s.l. The specimens from SaraguroHuashapamba were collected in pitfall traps baited with dung.
</paragraph>
</subSubSection>
<subSubSection id="C30490F713695314FF4CFB4BFEEBFA9E" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BA1C37C13695314FF4CFB4BFE6DFB48" blockId="33.[189,491,1201,1227]" box="[189,491,1201,1227]" pageId="33" pageNumber="34">
<heading id="D0E9741013695314FF4CFB4BFE6DFB48" bold="true" box="[189,491,1201,1227]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B96A1F6E13695314FF4CFB4BFE6DFB48" bold="true" box="[189,491,1201,1227]" pageId="33" pageNumber="34">Geographic distribution</emphasis>

View file

@ -1,54 +1,54 @@
<document id="EAA70F4CDE4E06F712ADCDA62E53B07D" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A136B5311FDA9F92FFBE5F935" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus placitus Moret 2020, sp. nov." docType="treatment" docUuid="02B83689-1D33-4C9B-85D2-8D6316CBD0FE" docUuidSource="ZooBank" docVersion="8" lastPageNumber="37" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="36" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EC6E7F3416CA4D2F7618F6694A5CE1E4">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
<document id="BF708E1F4FB3C27A4871623407720456" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A136B5311FDA9F92FFBE5F935" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus placitus Moret 2020, sp. nov." docType="treatment" docUuid="02B83689-1D33-4C9B-85D2-8D6316CBD0FE" docUuidSource="ZooBank" docVersion="12" lastPageNumber="37" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="36" updateTime="1738077108164" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A520B9B8D37A7C4408A3D89288B076B7">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
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<mods:namePart id="865AB5B23C66659B61E98EEF141337D9">Moret, Pierre</mods:namePart>
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<mods:affiliation id="056191B789263220766631963D11D677">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="A44C22E99C00CEA512E2268D03382478" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
<mods:namePart id="F82E4BC59A71AD83D5A2FF748EDEE25A">Moret, Pierre</mods:namePart>
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<mods:affiliation id="E21767ACD93D9710C877D5F6B134BB5A">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
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<mods:namePart id="6727112737BE2522C9100A42BE59AB0C">Murienne, Jérôme</mods:namePart>
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<mods:affiliation id="F041791B607DBCED23EED1DAF1ECBA15">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
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<treatment id="03B7726A136B5311FDA9F92FFBE5F935" ID-DOI="http://doi.org/10.5281/zenodo.3848403" ID-GBIF-Taxon="164160394" ID-Zenodo-Dep="3848403" LSID="urn:lsid:zoobank.org:act:02B83689-1D33-4C9B-85D2-8D6316CBD0FE" httpUri="http://treatment.plazi.org/id/03B7726A136B5311FDA9F92FFBE5F935" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
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<treatment id="03B7726A136B5311FDA9F92FFBE5F935" ID-DOI="http://doi.org/10.5281/zenodo.3848403" ID-GBIF-Taxon="164160394" ID-Zenodo-Dep="3848403" LSID="urn:lsid:zoobank.org:act:02B83689-1D33-4C9B-85D2-8D6316CBD0FE" httpUri="http://treatment.plazi.org/id/03B7726A136B5311FDA9F92FFBE5F935" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" scope_class="Insecta" scope_family="Carabidae" scope_order="Coleoptera" scope_subFamily="Harpalinae" scope_tribe="Platynini">
<subSubSection id="C30490F7136B5316FDA9F92FFC5AF973" box="[600,988,1749,1776]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BA1C37C136B5316FDA9F92FFC5AF973" blockId="35.[365,1222,1749,1847]" box="[600,988,1749,1776]" pageId="35" pageNumber="36">
<heading id="D0E97410136B5316FDA9F92FFC5AF973" box="[600,988,1749,1776]" centered="true" fontSize="11" level="2" pageId="35" pageNumber="36" reason="2">
<taxonomicName id="4C1EB8FF136B5316FDA9F92FFCFCF973" ID-CoL="8MDGD" authority="Moret" authorityName="Moret" authorityYear="2020" box="[600,890,1749,1776]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="placitus" status="sp. nov.">
@ -58,14 +58,14 @@ Moret
<taxonomicNameLabel id="A259A215136B5316FC70F92CFC5AF973" box="[897,988,1750,1776]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BA1C37C136B5316FD59F8E6FC0AF8B4" blockId="35.[365,1222,1749,1847]" box="[680,908,1820,1847]" pageId="35" pageNumber="36">
<figureCitation id="1325DFF9136B5316FD59F8E6FCADF8B5" box="[680,811,1820,1846]" captionStart="Figs 3641" captionStartId="38.[189,241,1854,1880]" captionTargetBox="[288,1297,277,1813]" captionTargetId="figure@38.[278,1309,265,1818]" captionTargetPageId="38" captionText="Figs 3641. Dyscolus spp., habitus (3638) and aedeagus, median lobe in lateral view (3940) and apex of the median lobe (41). 36. Dyscolus arauzae Moret sp. nov., male holotype (QCAZ). 37. D. piscator Moret sp. nov., male holotype (QACZ). 38. D. placitus Moret sp. nov., male holotype (MNHN). 39. D. arauzae Moret sp. nov. 40. D. piscator Moret sp. nov. 41. D. placitus Moret sp. nov. (above) and D. piscator Moret sp. nov. (below)." figureDoi="http://doi.org/10.5281/zenodo.3829702" httpUri="https://zenodo.org/record/3829702/files/figure.png" pageId="35" pageNumber="36">Figs 38, 41</figureCitation>
(above)
@ -94,7 +94,7 @@ urn:lsid:zoobank.org:act:
</paragraph>
<paragraph id="8BA1C37C136B5316FF4CF834FD1DF86B" blockId="35.[189,667,1998,2024]" box="[189,667,1998,2024]" pageId="35" pageNumber="36">Latin adjective meaning nice, pleasant.</paragraph>
</subSubSection>
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<subSubSection id="C30490F7136C5311FF4CFEF1FD9DFDE2" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="8BA1C37C136C5311FF4CFEF1FEEAFEA6" blockId="36.[189,364,267,293]" box="[189,364,267,293]" pageId="36" pageNumber="37">
<heading id="D0E97410136C5311FF4CFEF1FEEAFEA6" bold="true" box="[189,364,267,293]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B96A1F6E136C5311FF4CFEF1FEEAFEA6" bold="true" box="[189,364,267,293]" pageId="36" pageNumber="37">Type material</emphasis>
@ -149,22 +149,22 @@ leg.;
)
</paragraph>
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<materialsCitation id="3B76C921136C5311FF4CFE27FC04FE74" ID-GBIF-Occurrence="2620303417" box="[189,898,475,503]" collectingDate="2017-03-11" collectionCode="CPM" collectorName="Paso de la Virgen &amp; P. Moret &amp; M. Gobbi" country="Ecuador" county="Guamani" elevation="3890" latitude="-0.35016665" location="ECUADOR" longLatPrecision="1" longitude="-78.19789" municipality="Paso de la Virgen" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Napo">
<materialsCitation id="3B76C921136C5311FF4CFE27FC04FE74" ID-GBIF-Occurrence="2620303417" box="[189,898,475,503]" collectingDate="2017-03-11" collectionCode="CPM" collectorName="Paso de la Virgen &amp; P. Moret &amp; M. Gobbi" country="Ecuador" county="Guamani" elevation="3890" latitude="-0.35016665" location="ECUADOR" longLatPrecision="1" longitude="-78.19789" municipality="Paso de la Virgen" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<collectingCountry id="F30983EC136C5311FF4CFE27FECDFE74" box="[189,331,477,503]" name="Ecuador" pageId="36" pageNumber="37">ECUADOR</collectingCountry>
<specimenCount id="9D1808F5136C5311FE91FE27FE0AFE74" box="[352,396,475,503]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
<specimenCount id="9D1808F5136C5311FE91FE27FE0AFE74" box="[352,396,475,503]" count="1" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB9136C5311FCB5FE27FC04FE74" box="[836,898,477,503]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="36" pageNumber="37">CPM</collectionCode>
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<specimenCount id="9D1808F5136C5311FC69FE27FC45FE74" box="[920,963,477,503]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
<materialsCitation id="3B76C921136C5311FC69FE27FEEDFD98" ID-GBIF-Occurrence="2620303353" collectingDate="2017-03-11" collectionCode="QCAZ" collectorName="Paso de la Virgen &amp; P. Moret &amp; M. Gobbi" country="Ecuador" county="Guamani" elevation="3890" latitude="-0.35016665" location="Waypoint" longLatPrecision="1" longitude="-78.19789" municipality="Paso de la Virgen" pageId="36" pageNumber="37" specimenCode="QCAZ 256725" specimenCount="1" specimenCount-female="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="9D1808F5136C5311FC69FE27FC45FE74" box="[920,963,477,503]" count="1" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBB86B07136C5311FF4CFDFAFEEDFD98" box="[189,363,512,539]" collectionCode="QCAZ" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="36" pageNumber="37" type="Museum">QCAZ 256725</specimenCode>
</materialsCitation>
<materialsCitation id="3B76C921136C5311FE70FDFAFD91FDE2" ID-GBIF-Occurrence="2620303334" collectingDate="1982-05-13" collectionCode="UASM" collectorName="H. Frania &amp; F. Sperling" country="Ecuador" elevation="3810" location="Paso de Papallacta" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Napo Province" typeStatus="paratype">
<specimenCount id="9D1808F5136C5311FE70FDFAFE2AFD99" box="[385,428,510,538]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
<specimenCount id="9D1808F5136C5311FE70FDFAFE2AFD99" box="[385,428,510,538]" count="1" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
(red label: “
<taxonomicName id="4C1EB8FF136C5311FDB3FDFAFBADFD98" authority="Moret" authorityName="Moret" authorityYear="2005" box="[578,1067,512,539]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="hirsutus" subGenus="Hydrodyscolus">
<emphasis id="B96A1F6E136C5311FDB3FDFAFD28FD99" box="[578,686,512,538]" italics="true" pageId="36" pageNumber="37">Dyscolus</emphasis>
@ -174,7 +174,9 @@ leg.;
<emphasis id="B96A1F6E136C5311FC8DFDFAFC5DFD99" box="[892,987,512,538]" italics="true" pageId="36" pageNumber="37">hirsutus</emphasis>
Moret
</taxonomicName>
, Paratype”);
,
<typeStatus id="54A57DDE136C5311FBC7FDFAFB23FD99" box="[1078,1189,512,538]" pageId="36" pageNumber="37">Paratype</typeStatus>
”);
<collectingRegion id="49DA0D9E136C5311FB30FDFBFAF4FD99" box="[1217,1394,512,539]" country="Ecuador" name="Napo" pageId="36" pageNumber="37">Napo Province</collectingRegion>
,
<location id="8EC195A7136C5311FF4CFDDEFE11FDBE" LSID="urn:lsid:plazi:treatment:03B7726A136B5311FDA9F92FFBE5F935:8EC195A7136C5311FF4CFDDEFE11FDBE" box="[189,407,547,574]" country="Ecuador" name="Paso de Papallacta" pageId="36" pageNumber="37" stateProvince="Napo Province">Paso de Papallacta</location>
@ -196,6 +198,8 @@ leg.; under stones and logs near stream;
</materialsCitation>
.
</paragraph>
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<subSubSection id="C30490F7136C5311FF4CFD7AFB10FBEC" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="8BA1C37C136C5311FF4CFD7AFE52FD19" blockId="36.[189,468,640,666]" box="[189,468,640,666]" pageId="36" pageNumber="37">
<heading id="D0E97410136C5311FF4CFD7AFE52FD19" bold="true" box="[189,468,640,666]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B96A1F6E136C5311FF4CFD7AFE52FD19" bold="true" box="[189,468,640,666]" pageId="36" pageNumber="37">Diagnostic description</emphasis>
@ -216,6 +220,8 @@ Habitus:
<emphasis id="B96A1F6E136C5311FCB4FBAFFB8BFBEC" box="[837,1037,1109,1135]" italics="true" pageId="36" pageNumber="37">Female genitalia</emphasis>
: unstudied.
</paragraph>
</subSubSection>
<subSubSection id="C30490F7136C5311FF4CFB6DFAF1FA31" pageId="36" pageNumber="37" type="discussion">
<paragraph id="8BA1C37C136C5311FF4CFB6DFEE5FB32" blockId="36.[189,355,1175,1201]" box="[189,355,1175,1201]" pageId="36" pageNumber="37">
<heading id="D0E97410136C5311FF4CFB6DFEE5FB32" bold="true" box="[189,355,1175,1201]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B96A1F6E136C5311FF4CFB6DFEE5FB32" bold="true" box="[189,355,1175,1201]" pageId="36" pageNumber="37">Comparisons</emphasis>
@ -251,6 +257,8 @@ Moret
<taxonomicNameLabel id="A259A215136C5311FCF0FA63FCDDFA30" box="[769,859,1433,1459]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
are set forth in the diagnoses of these species.
</paragraph>
</subSubSection>
<subSubSection id="C30490F7136C5311FF4CFA20FA93F9C5" pageId="36" pageNumber="37" type="biology_ecology">
<paragraph id="8BA1C37C136C5311FF4CFA20FE9BFA77" blockId="36.[189,285,1498,1524]" box="[189,285,1498,1524]" pageId="36" pageNumber="37">
<heading id="D0E97410136C5311FF4CFA20FE9BFA77" bold="true" box="[189,285,1498,1524]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B96A1F6E136C5311FF4CFA20FE9BFA77" bold="true" box="[189,285,1498,1524]" pageId="36" pageNumber="37">Habitat</emphasis>
@ -265,6 +273,8 @@ a.s.l. The specimens collected in 2017 were found in the understory of a
</taxonomicName>
forest, beneath soaked mosses on the ground.
</paragraph>
</subSubSection>
<subSubSection id="C30490F7136C5311FF4CF997FBE5F935" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8BA1C37C136C5311FF4CF997FE6DF904" blockId="36.[189,491,1645,1671]" box="[189,491,1645,1671]" pageId="36" pageNumber="37">
<heading id="D0E97410136C5311FF4CF997FE6DF904" bold="true" box="[189,491,1645,1671]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B96A1F6E136C5311FF4CF997FE6DF904" bold="true" box="[189,491,1645,1671]" pageId="36" pageNumber="37">Geographic distribution</emphasis>

View file

@ -1,54 +1,54 @@
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<subSubSection id="C30490F7136F5312FD93FB87FB40FB38" pageId="39" pageNumber="40" type="nomenclature">
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<subSubSection id="C30490F7136F5312FD93FB87FC54FB1B" box="[610,978,1149,1176]" pageId="39" pageNumber="40" type="nomenclature">
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<taxonomicName id="4C1EB8FF136F5312FD93FB87FCF7FB1B" ID-CoL="8M7XZ" authority="Moret" authorityName="Moret" authorityYear="2020" box="[610,881,1149,1176]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="gobbii" status="sp. nov.">
@ -58,14 +58,14 @@ Moret
<taxonomicNameLabel id="A259A215136F5312FC89FB84FC54FB1B" box="[888,978,1150,1176]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BA1C37C136F5312FD01FB39FCC5FB5D" blockId="39.[366,1222,1149,1246]" box="[752,835,1219,1246]" pageId="39" pageNumber="40">
<figureCitation id="1325DFF9136F5312FD01FB39FCC5FB5D" box="[752,835,1219,1246]" captionStart="Figs 4244. 42" captionStartId="40.[189,241,1923,1949]" captionTargetBox="[232,1381,485,1873]" captionTargetId="figure@40.[192,1396,473,1882]" captionTargetPageId="40" captionText="Figs 4244. 42. Dyscolus gobbii Moret sp. nov., habitus of the female holotype (QACZ, COI voucher PM210-16). 4344. D. barragani Moret sp. nov. 43. Habitus of the male paratype (CPM, COI voucher PM417-02). 44. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829704" httpUri="https://zenodo.org/record/3829704/files/figure.png" pageId="39" pageNumber="40">Fig. 42</figureCitation>
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<paragraph id="8BA1C37C136F5312FF4CFACEFCF8FAF2" blockId="39.[189,1399,1331,1393]" pageId="39" pageNumber="40">Noun in the genitive case, dedicated to Mauro Gobbi, specialist of the ecology of glacier-foreland invertebrates, who took part in the discovery of this species.</paragraph>
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<emphasis id="B96A1F6E136F5312FF4CFA62FEEAFA31" bold="true" box="[189,364,1432,1458]" pageId="39" pageNumber="40">Type material</emphasis>
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<materialsCitation id="3B76C921136F5312FF4CFA10FD07F9C9" ID-GBIF-Occurrence="2620303330" collectingDate="2017-02-27" collectionCode="QCAZ" collectorName="P. Moret &amp; M. Gobbi" country="Ecuador" county="Guamani" elevation="4230" latitude="-0.32175" location="Waypoint 210" longLatPrecision="1" longitude="-78.198555" municipality="Paso de la Virgen" pageId="39" pageNumber="40" specimenCode="SUM228" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha Province" typeStatus="holotype">
<collectingCountry id="F30983EC136F5312FF4CFA10FECDF987" box="[189,331,1514,1540]" name="Ecuador" pageId="39" pageNumber="40">ECUADOR</collectingCountry>
<specimenCount id="9D1808F5136F5312FE80FA10FE0EF987" box="[369,392,1514,1540]" pageId="39" pageNumber="40" type="female"></specimenCount>
@ -119,7 +119,9 @@ a.s.l.
<collectorName id="26EBA6AA136F5312FD14F9F4FCCAF9AB" box="[741,844,1550,1576]" pageId="39" pageNumber="40">P. Moret</collectorName>
and
<collectorName id="26EBA6AA136F5312FC7DF9F4FB87F9A4" box="[908,1025,1549,1576]" pageId="39" pageNumber="40">M. Gobbi</collectorName>
leg.; COI voucher PM210-16, BOLD sequence SUM228-18;
leg.; COI voucher PM210-16, BOLD sequence
<specimenCode id="DBB86B07136F5312FE76F9CAFE7FF9C9" box="[391,505,1584,1610]" pageId="39" pageNumber="40">SUM228</specimenCode>
-18;
<collectionCode id="ED0F5BB9136F5312FDD8F9CAFDFBF9C9" box="[553,637,1584,1610]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="39" pageNumber="40" type="Museum">QCAZ</collectionCode>
.
</materialsCitation>
@ -130,14 +132,14 @@ leg.; COI voucher PM210-16, BOLD sequence SUM228-18;
</emphasis>
</paragraph>
<paragraph id="8BA1C37C136F5312FF4CF96FFE23F976" blockId="39.[189,1399,1650,1781]" pageId="39" pageNumber="40">
<materialsCitation id="3B76C921136F5312FF4CF96FFE27F976" ID-GBIF-Occurrence="2620303431" collectingDate="2016-04-22" collectionCode="CPM" collectorName="Volcan Antisana NW &amp; E. Moreno" country="Ecuador" elevation="4401" latitude="-0.45778" location="Pichincha Province" longLatPrecision="1" longitude="-78.16797" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha Province" typeStatus="paratype">
<materialsCitation id="3B76C921136F5312FF4CF96FFE27F976" ID-GBIF-Occurrence="2620303431" collectingDate="2016-04-22" collectionCode="CPM" collectorName="E. Moreno" country="Ecuador" elevation="4401" latitude="-0.45778" location="Volcan Antisana NW" longLatPrecision="1" longitude="-78.16797" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha Province" typeStatus="paratype">
<collectingCountry id="F30983EC136F5312FF4CF96FFECDF92C" box="[189,331,1685,1711]" name="Ecuador" pageId="39" pageNumber="40">ECUADOR</collectingCountry>
<specimenCount id="9D1808F5136F5312FE9FF96FFE26F92C" box="[366,416,1685,1711]" pageId="39" pageNumber="40" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49DA0D9E136F5312FE45F96FFD19F92C" box="[436,671,1685,1711]" country="Ecuador" name="Pichincha" pageId="39" pageNumber="40">Pichincha Province</collectingRegion>
,
<collectorName id="26EBA6AA136F5312FD5EF96FFC37F92C" box="[687,945,1685,1711]" pageId="39" pageNumber="40">Volcán Antisana NW</collectorName>
<location id="8EC195A7136F5312FD5EF96FFC37F92C" LSID="urn:lsid:plazi:treatment:03B7726A136F531CFD93FB87FA88FC0C:8EC195A7136F5312FD5EF96FFC37F92C" box="[687,945,1685,1711]" country="Ecuador" latitude="-0.45778" longLatPrecision="1" longitude="-78.16797" name="Volcan Antisana NW" pageId="39" pageNumber="40" stateProvince="Pichincha Province">Volcán Antisana NW</location>
, B1C3;
<geoCoordinate id="EE2AA5BB136F5312FBEBF96FFB36F92C" box="[1050,1200,1685,1712]" degrees="0.457780" direction="south" orientation="latitude" pageId="39" pageNumber="40" precision="1" value="-0.45778">0.457780º S</geoCoordinate>
,
@ -158,6 +160,8 @@ leg.; COI voucher in ethanol PM188-01, BOLD sequence SUM183-18;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30490F7136F531CFF4CF8E6FD0CFE0C" lastPageId="41" lastPageNumber="42" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="8BA1C37C136F5312FF4CF8E6FE52F8B5" blockId="39.[189,468,1820,1846]" box="[189,468,1820,1846]" pageId="39" pageNumber="40">
<heading id="D0E97410136F5312FF4CF8E6FE52F8B5" bold="true" box="[189,468,1820,1846]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B96A1F6E136F5312FF4CF8E6FE52F8B5" bold="true" box="[189,468,1820,1846]" pageId="39" pageNumber="40">Diagnostic description</emphasis>
@ -182,6 +186,7 @@ Habitus:
<emphasis id="B96A1F6E1361531CFEC8FE8FFD86FE0C" box="[313,512,373,399]" italics="true" pageId="41" pageNumber="42">Female genitalia</emphasis>
: unstudied.
</paragraph>
</subSubSection>
<caption id="DF6193F41360531DFF4CF879FCC3F860" ID-DOI="http://doi.org/10.5281/zenodo.3829704" ID-Zenodo-Dep="3829704" httpUri="https://zenodo.org/record/3829704/files/figure.png" pageId="40" pageNumber="41" startId="40.[189,241,1923,1949]" targetBox="[232,1381,485,1873]" targetPageId="40">
<paragraph id="8BA1C37C1360531DFF4CF879FCC3F860" blockId="40.[189,1398,1923,2020]" pageId="40" pageNumber="41">
<emphasis id="B96A1F6E1360531DFF4CF879FEF6F81E" bold="true" box="[189,368,1923,1949]" pageId="40" pageNumber="41">Figs 4244. 42</emphasis>
@ -205,6 +210,7 @@ Moret
. Aedeagus, median lobe in lateral view.
</paragraph>
</caption>
<subSubSection id="C30490F71361531CFF4CFE4CFE61FDEB" pageId="41" pageNumber="42" type="discussion">
<paragraph id="8BA1C37C1361531CFF4CFE4CFEE5FE53" blockId="41.[189,355,438,464]" box="[189,355,438,464]" pageId="41" pageNumber="42">
<heading id="D0E974101361531CFF4CFE4CFEE5FE53" bold="true" box="[189,355,438,464]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="B96A1F6E1361531CFF4CFE4CFEE5FE53" bold="true" box="[189,355,438,464]" pageId="41" pageNumber="42">Comparisons</emphasis>
@ -230,6 +236,8 @@ and
</taxonomicName>
in two clades far apart from each other.
</paragraph>
</subSubSection>
<subSubSection id="C30490F71361531CFF4CFD6AFDEEFC9C" pageId="41" pageNumber="42" type="biology_ecology">
<paragraph id="8BA1C37C1361531CFF4CFD6AFE9BFD29" blockId="41.[189,285,656,682]" box="[189,285,656,682]" pageId="41" pageNumber="42">
<heading id="D0E974101361531CFF4CFD6AFE9BFD29" bold="true" box="[189,285,656,682]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="B96A1F6E1361531CFF4CFD6AFE9BFD29" bold="true" box="[189,285,656,682]" pageId="41" pageNumber="42">Habitat</emphasis>
@ -240,6 +248,8 @@ Humid superpáramo, from
<quantity id="4CE66E991361531CFE0AFD45FD34FD5A" box="[507,690,703,729]" metricMagnitude="3" metricUnit="m" metricValue="4.3" metricValueMax="4.4" metricValueMin="4.2" pageId="41" pageNumber="42" unit="m" value="4300.0" valueMax="4400.0" valueMin="4200.0">4200 to 4400 m</quantity>
a.s.l. The Guamaní specimen was collected under a stone on the shore of a small lake; the Antisana specimen was collected by pitfall trapping in a mesic environment, 20 metres from a glacier-fed stream.
</paragraph>
</subSubSection>
<subSubSection id="C30490F71361531CFF4CFCBCFA88FC0C" pageId="41" pageNumber="42" type="distribution">
<paragraph id="8BA1C37C1361531CFF4CFCBCFE6DFCE3" blockId="41.[189,491,838,864]" box="[189,491,838,864]" pageId="41" pageNumber="42">
<heading id="D0E974101361531CFF4CFCBCFE6DFCE3" bold="true" box="[189,491,838,864]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="B96A1F6E1361531CFF4CFCBCFE6DFCE3" bold="true" box="[189,491,838,864]" pageId="41" pageNumber="42">Geographic distribution</emphasis>