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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.885.38980" ID-GBIF-Dataset="5fe8e574-2edb-4095-8b3e-c6d856bb07f2" ID-PMC="PMC6848233" ID-Pensoft-Pub="1313-2970-885-1" ID-Pensoft-UUID="870C0613A13A5C1EA54511B2C64DEBE4" ID-PubMed="31736615" ID-ZooBank="A950D3CB4C914E62AF6631B76D73880C" ModsDocID="1313-2970-885-1" checkinTime="1572912889973" checkinUser="pensoft" docAuthor="Sutcharit, Chirasak, Naggs, Fred, Ablett, Jonathan, Sang, Pham Van, Luong Van Hao, &amp; Panha, Somsak" docDate="2019" docId="5EE0ABFD67EB56EC837E193293F933D4" docLanguage="en" docName="ZooKeys 885: 1-14" docOrigin="ZooKeys 885" docSource="http://dx.doi.org/10.3897/zookeys.885.38980" docTitle="Bertia Ancey 1887" docType="treatment" docVersion="4" id="870C0613A13A5C1EA54511B2C64DEBE4" lastPageNumber="1" masterDocId="870C0613A13A5C1EA54511B2C64DEBE4" masterDocTitle="Notes on the sinistral helicoid snail Bertia cambojiensis (Reeve, 1860) from Vietnam (Eupulmonata, Dyakiidae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="1" updateTime="1668168000113" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Notes on the sinistral helicoid snail Bertia cambojiensis (Reeve, 1860) from Vietnam (Eupulmonata, Dyakiidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naggs, Fred</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ablett, Jonathan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sang, Pham Van</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Luong Van Hao,</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>885</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>14</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.885.38980</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.885.38980</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-885-1</mods:identifier>
<mods:identifier type="ZooBank">A950D3CB4C914E62AF6631B76D73880C</mods:identifier>
<mods:identifier type="Pensoft-UUID">870C0613A13A5C1EA54511B2C64DEBE4</mods:identifier>
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<treatment ID-GBIF-Taxon="160134099" LSID="urn:lsid:plazi:treatment:5EE0ABFD67EB56EC837E193293F933D4" httpUri="http://treatment.plazi.org/id/5EE0ABFD67EB56EC837E193293F933D4" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
Genus
<taxonomicName LSID="5EE0ABFD-67EB-56EC-837E-193293F933D4" authority="Ancey, 1887" authorityName="Ancey" authorityYear="1887" class="Gastropoda" family="Dyakiidae" genus="Bertia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bertia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Bertia Ancey, 1887</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Bertia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bertia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Bertia</taxonomicName>
Ancey, 1887: 53.
<bibRefCitation author="Baker, HB" journalOrPublisher="Bishop Museum Bulletin" pageId="0" pageNumber="1" pagination="205 - 346" refId="B5" refString="Baker, HB, 1941. Zonitid snails from Pacific Islands, Part 3. Genera other the Microcystinae. Bernice P. Bishop Museum Bulletin 166: 205 - 346" title="Zonitid snails from Pacific Islands, Part 3. Genera other the Microcystinae. Bernice P." volume="166" year="1941">Baker 1941</bibRefCitation>
: 320, 321.
<bibRefCitation author="Zilch, A" editor="Schindewolf, OH" journalOrPublisher="Academy of Natural Science Philadelphia, Philadelphia" pageId="0" pageNumber="1" refId="B48" refString="Zilch, A, Schindewolf, OH, 1959. Gastropoda, Euthyneura. In: SchindewolfOH (Ed.) Handbuch der Palaeozoologie, Band 6, Gastropoda.Gebrueder Borntraeger, Berlin, 1-400." title="Gastropoda, Euthyneura. In: SchindewolfOH (Ed.) Handbuch der Palaeozoologie, Band 6, Gastropoda. Gebrueder Borntraeger, Berlin, 1 - 400." year="1959">Zilch 1959</bibRefCitation>
: 275, 276.
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Annals and Magazine of Natural History, Series" pageId="0" pageNumber="1" refId="B37" refString="Schileyko, AA, 2003. Treatise on recent terrestrial pulmonated mollusks. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica, Supplement 2, Part 10: 1309-1466." title="Treatise on recent terrestrial pulmonated mollusks. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica, Supplement 2, Part 10: 1309 - 1466." year="2003">Schileyko 2003</bibRefCitation>
: 1362.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Ariophanta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ariophanta (Bertia)" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subGenus" subGenus="Bertia">Ariophanta (Bertia)</taxonomicName>
Thiele, 1931: 1001.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix cambojiensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cambojiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Helix cambojiensis</emphasis>
</taxonomicName>
Reeve, 1860, by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Thach, NN" journalOrPublisher="The Festivus" pageId="0" pageNumber="1" pagination="240 - 242" refId="B43" refString="Thach, NN, 2015. Bertiasetzeri, a new species of land snail from Vietnam (Gastropoda: Stylommatophora: Dyakiidae). The Festivus 47: 240 - 242" title="Bertiasetzeri, a new species of land snail from Vietnam (Gastropoda: Stylommatophora: Dyakiidae)." volume="47" year="2015">Thach (2015)</bibRefCitation>
recognised four nominal species in
<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Bertia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bertia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Bertia</emphasis>
</taxonomicName>
: &quot;
<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Bertia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bertia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Bertia</emphasis>
</taxonomicName>
&quot;
<taxonomicName lsidName="brookei" pageId="0" pageNumber="1" rank="species" species="brookei">
<emphasis italics="true" pageId="0" pageNumber="1">brookei</emphasis>
</taxonomicName>
(Adam &amp; Reeve, 1848),
<taxonomicName lsidName="B. cambojiensis" pageId="0" pageNumber="1" rank="species" species="cambojiensis">
<emphasis italics="true" pageId="0" pageNumber="1">B. cambojiensis</emphasis>
</taxonomicName>
(type species),
<taxonomicName lsidName="B. pergrandis" pageId="0" pageNumber="1" rank="species" species="pergrandis">
<emphasis italics="true" pageId="0" pageNumber="1">B. pergrandis</emphasis>
</taxonomicName>
(Smith, 1893) and
<taxonomicName lsidName="B. setzeri" pageId="0" pageNumber="1" rank="species" species="setzeri">
<emphasis italics="true" pageId="0" pageNumber="1">B. setzeri</emphasis>
</taxonomicName>
Thach, 2015. Nevertheless, he had overlooked key characters of the genitalia published by
<bibRefCitation DOI="https://doi.org/10.1111/j.1469-7998.1891.tb06805.x" author="Godwin-Austen, HH" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="22 - 47" refId="B14" refString="Godwin-Austen, HH, 1891. On a collection of land shells made in Borneo by Mr. Everett, with descriptions of supposed new species. Part II. Zonitidae and Helicinidae. Proceedings of the Zoological Society of London 59: 22 - 47" title="On a collection of land shells made in Borneo by Mr. Everett, with descriptions of supposed new species. Part II. Zonitidae and Helicinidae." url="https://doi.org/10.1111/j.1469-7998.1891.tb06805.x" volume="59" year="1891">Godwin-Austen (1891</bibRefCitation>
: pl. 6, fig. 6) and
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Annals and Magazine of Natural History, Series" pageId="0" pageNumber="1" refId="B37" refString="Schileyko, AA, 2003. Treatise on recent terrestrial pulmonated mollusks. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica, Supplement 2, Part 10: 1309-1466." title="Treatise on recent terrestrial pulmonated mollusks. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae. Ruthenica, Supplement 2, Part 10: 1309 - 1466." year="2003">Schileyko (2003</bibRefCitation>
: 1345, 1346, fig. 1758). The Bornean endemic species,
<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix brookei" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="brookei">
<emphasis italics="true" pageId="0" pageNumber="1">Helix brookei</emphasis>
</taxonomicName>
Adam &amp; Reeve, 1848, exhibits a long and cylindrical gametolytic sac, lacks an epiphallus and amatorial organ complex, and a caudal foss and a caudal horn is absent; characters that unequivocally distinguish this species from the
<taxonomicName family="Dyakiidae" lsidName="" pageId="0" pageNumber="1" rank="family">Dyakiidae</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1111/j.1469-7998.1891.tb06805.x" author="Godwin-Austen, HH" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="22 - 47" refId="B14" refString="Godwin-Austen, HH, 1891. On a collection of land shells made in Borneo by Mr. Everett, with descriptions of supposed new species. Part II. Zonitidae and Helicinidae. Proceedings of the Zoological Society of London 59: 22 - 47" title="On a collection of land shells made in Borneo by Mr. Everett, with descriptions of supposed new species. Part II. Zonitidae and Helicinidae." url="https://doi.org/10.1111/j.1469-7998.1891.tb06805.x" volume="59" year="1891">Godwin-Austen 1891</bibRefCitation>
,
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="190 - 201" refId="B19" refString="Laidlaw, FF, 1931. On a new subfamily Dyakiinae of the Zonitidae. Proceedings of the Malacological Society of London 19: 190 - 201" title="On a new subfamily Dyakiinae of the Zonitidae." volume="19" year="1931">Laidlaw 1931</bibRefCitation>
,
<bibRefCitation author="Sutcharit, C" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="279 - 287" refId="B42" refString="Sutcharit, C, Tongkerd, P, Tan, AS-H, Panha, S, 2012. Taxonomic revision of Dyakiajanus from Peninsular Malaysia (Pulmonata: Dyakiidae), with notes on other sinistrally coiled helicarionids. The Raffles Bulletin of Zoology 60: 279 - 287" title="Taxonomic revision of Dyakiajanus from Peninsular Malaysia (Pulmonata: Dyakiidae), with notes on other sinistrally coiled helicarionids." volume="60" year="2012">Sutcharit et al. 2012</bibRefCitation>
). It is clear that
<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix brookei" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="brookei">
<emphasis italics="true" pageId="0" pageNumber="1">Helix brookei</emphasis>
</taxonomicName>
Adam &amp; Reeve, 1848 does not belong in
<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Bertia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bertia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Bertia</emphasis>
</taxonomicName>
and we recognise the following three as
<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Bertia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bertia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Bertia</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>